Tomoko Naganuma,^1,* Shinsuke Koike,¹ Rumiko Nakashita,² Chinatsu Kozakai,³ Koji Yamazaki,⁴ Shino Furusaka,¹ Koichi Kaji<sup>5

1</sup>Laboratory of Forest Conservation Biology, Faculty of Agriculture, Tokyo University of Agriculture and Technology, Fuchu, Tokyo 183-8509, Japan
²Wildlife Ecology Laboratory, Forestry and Forest Products Research Institute, Tsukuba, Ibaraki 305-8687, Japan
³Wildlife Management Laboratory, Central Region Agricultural Research Center, National Agriculture and Food Research Organization, Tsukuba, Ibaraki 305-8666, Japan
⁴Laboratory of Forest Ecology, Department of Forest Science, Faculty of Regional Environmental Science, Tokyo University of Agriculture, Setagaya, Tokyo 156-8502, Japan
⁵Laboratory of Wildlife Management, Faculty of Agriculture, Tokyo University of Agriculture and Technology, Fuchu, Tokyo 183-8509, Japan

^*To whom correspondence should be addressed. E-mail: tama.827dx@gmail.com