Tomohiro Tsuji,^1,* Hiroki Gotoh,^1,* Shinichi Morita,² Junya Hirata,¹ Yohei Minakuchi,³ Toshinobu Yaginuma,¹ Atsushi Toyoda,^3,4 Teruyuki Niimi<sup>1,2,5,**

1</sup>1Graduate School of Bioagricultural Sciences, Nagoya University, Chikusa, Nagoya 464-8601, Japan
²2Division of Evolutionary Developmental Biology, National Institute for Basic Biology, 38, Nishigona
³3Comparative Genomics Laboratory, National Institute of Genetics, Mishima, Shizuoka 411-8540, Japan
⁴4Advanced Genomics Center, National Institute of Genetics, Mishima, Shizuoka 411-8540, Japan
⁵5Department of Basic Biology, School of Life Science, SOKENDAI (The Graduate University for Advanced

^* These authors equally contributed to this work.
^**Corresponding author. E-mail: niimi@nibb.ac.jp