Clinton D. Francis,^1,2 Nathan J. Kleist,¹ Benjamin J. Davidson,¹ Catherine P. Ortega,^3,4 Alexander Cruz<sup>1

1</sup>Ecology and Evolutionary Biology, University of Colorado, Boulder, Colorado 80309, USA
²Present address: National Evolutionary Synthesis Center, 2024 W Main Street, Suite A200, Box 104403, Durham, North Carolina 27705, USA. E-mail: clinton.francis@nescent.org
³San Juan Institute of Natural and Cultural Resources, Fort Lewis College, 1000 Rim Drive, Durango, Colorado 81301, USA
⁴Present address: Ecosphere Environmental Services, 776 E. Second Avenue, Durango, Colorado 81301, USA.