On the basis of seed morphology, Echinocactus mandragora Berger is seen as unrelated to Turbinicarpus, but a closer ally of Sclerocactus, Echinomastus and Pediocactus. It differs from these genera in its floral and fruit morphology, therefore we propose a new genus, Lodia Mosco & Zanovello, to accommodate this species.
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Vol. 2000 • No. 18