Callmander, M. W., R. Bolliger, R. M. Hanitrarivo & L. Nusbaumer (2013). Pandanus tsingycola Callm. & Nusb. (Pandanaceae), a new species endemic to western Madagascar. Candollea 68: 229–235. In English, English and French abstracts.
A recent intensive inventory in the Beanka region in western Madagascar has led to the discovery of a new species of Pandanaceae, described here as Pandanus tsingycola Callm. & Nusb. This remarkable species can be easily distinguished from all other known species by the unique shape and configuration of its syncarps. These possess few phalanges (6–7) that are incompletely united (with up to 38 carpels) with each of these carpels bearing an erect to curved, acute, thorny stigma. Pandanus tsingycola seems to be endemic to the limestone “tsingy” formation of Beanka.