Harpactocarcinus punctulatus istriensis Bachmayer and Nosan, 1959 is elevated to species level. Analysis of the larger foraminiferans associated with specimens of H. istriensis suggests a habitat preference for off-shore, clear, shelf environments below fair-weather wave base and an age of early to middle Lutetian (Eocene). A review of the paleoenvironmental indicators for nearly all species within the genera referred to the Zanthopsidae Via, 1959 suggests that all exhibit similar habitat preferences. Description of the paleoenvironmental preference for an entire extinct decapod family has not before been possible.
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