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1 July 2016 Hepaticites iporangae Ricardi-Branco, Faria, Jasper, and Guerra-Sommer, 2011 from the early Permian of the Paraná Basin, Brazil, is not a liverwort but a tracheophyte
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Abstract

From new and more complete material, which includes frond fragments with casts of tracheid remains of the rachis, it can be demonstrated that the putative liverwort Hepaticites iporangae Ricardi-Branco, Faria, Jasper, and Guerra-Sommer, 2011, from the early Permian Rio Bonito Formation (Sakmarian) of the Paraná Basin, Brazil, is not a bryophyte but a tracheophyte. The new material was collected from the same locality and layer as the type material, in the Quitéria outcrop in the municipality of Encruzilhada do Sul, state of Rio Grande do Sul, Brazil. From the morphology of the deeply dissected pinnatifid pinnules with narrow laminar lobes, the taxon is provisionally reassigned to the genus Rhodeopteridium. Thus we propose the new combination ‘Rhodeopteridiumiporangae new combination for this taxon. This new systematic interpretation contributes to our understanding of early liverworts (by removing Hepaticites iporangae as a possible taxon thereof) and clarifies an issue of diversity of the flora of the early Permian Rio Bonito Formation resulting from the original misidentification.

© 2016, The Paleontological Society
Dieter Uhl, André Jasper, Talyssa Valerius, Joseline Manfroi, Margot Guerra-Sommer, and Fresia Soledad Ricardi-Branco "Hepaticites iporangae Ricardi-Branco, Faria, Jasper, and Guerra-Sommer, 2011 from the early Permian of the Paraná Basin, Brazil, is not a liverwort but a tracheophyte," Journal of Paleontology 90(4), (1 July 2016). https://doi.org/10.1017/jpa.2016.82
Accepted: 1 March 2016; Published: 1 July 2016
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