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1 May 2013 New Information on Amphibamids (Tetrapoda, Temnospondyli) from Richards Spur (Fort Sill), Oklahoma
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A nearly complete amphibamid skull from the Richards Spur locality in Oklahoma is demonstrated to be a new species of Tersomius. This new species has a mosaic of features seen in other amphibamids, and appears to fall on the Amphibamus side of the amphibamid basal dichotomy of some authors. This specimen also calls into question the validity of the Tersomius assignment of a large specimen from the Texas Red Beds. One feature of note is the presence of a weakly bicuspid fang on one of the vomers. This poorly developed feature is present only on an unattached replacement cusp, and appears to have been rapidly worn away through use. This is the first potential bicuspid palatal fang known from an amphibamid. A second partial skull is assigned to the recently described species Pasawioops mayi. A micro-computed tomographic reconstruction shows a prominent ventral flange of the nasal that meets a counterpart from the prefrontal and palatine to wall the nasal capsule posteriorly. The nasolacrimal canal shows the three openings in the orbit quickly converge to pass rostrally as a single canal except for a mysterious lateral diverticulation that has a superficial exposure on the lacrimal. Both skulls preserve a pars palatina of the premaxilla, in common with batrachians, and the Pasawioops specimen also preserves a prominent palatal process of the premaxilla, a frog synapomorphy. Richards Spur preserves one of the most diverse amphibamid faunas in the world, with three species described so far.

Jason S. Anderson and John R. Bolt "New Information on Amphibamids (Tetrapoda, Temnospondyli) from Richards Spur (Fort Sill), Oklahoma," Journal of Vertebrate Paleontology 33(3), (1 May 2013).
Received: 23 April 2012; Accepted: 30 August 2012; Published: 1 May 2013

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