An adult greater white-toothed shrew (Crocidura russula) had both a quadricuspid aortic valve and a single coronary artery arising from the aorta. The shrew was caught on 10 May 1994 in the environs of Málaga, southern Spain. Both congenital anomalies may be potential causes of cardiac dysfunction, but apparently produced no significant cardiac complication in the shrew. This is the first report of a quadricuspid aortic valve in a wild-living mammal.