The Diptera fauna of Ozark caves is fairly typical of that found elsewhere in the United States east of the Great Plains. The sphaerocerid Spelobia tenebrarum is the only troglobitic dipteran found in Ozark caves. Probable troglophiles include the mycetophilid Macrocera nobilis, the psychodids Psychoda spp., and the phorid Megaselia cavernicola. The tipulids commonly found in Ozark caves, such as Dactylolabis montana, Dolichopeza spp. and Limonia spp. are probably all trogloxenes, as are the Trichoceridae and Heleomyzidae. The mycetophilids, Exechiopsis umbratica and Rymosia sp., and the culicids are among the most common cave dipterans in the study area. The culicids overwinter in caves and other protected places; they are properly classified as trogloxenes. The main sources of food for cave Diptera are other insects, carrion, guano, and allochthanous plant debris.