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1 December 2012 A New Monofenestratan Pterosaur from the Kimmeridge Clay Formation (Kimmeridgian, Upper Jurassic) of Dorset, England
David M. Martill, Steve Etches
Author Affiliations +
Fig. 1.
Simplified geological map showing the distribution of the Kimmeridge Clay Formation on the Isle of Purbeck and the location of Kimmeridge Bay where Cuspicephalus scarfi was discovered.

Fig. 2.
A monofenestratan pterosaur Cuspicephalus scarfi gen. et sp. nov. from the Upper Jurassic of Kimmeridge Bay, England; MJML K1918. A. Original skull on slab of mudstone, lacking the mandible and dentition. B. Outline diagram of preserved bone. Light grey is bone, dark grey is fibrous bone of sagittal crest, black is dental alveoli where unambiguous. C. Restoration of skull outline, with hypothetical lower jaw. Scale bar 50 mm.

Table 1.
Key measurements of holotype skull of Cuspicephalus scarfi. Naof, nasoantorbital fenestra.

Fig. 3.
A monofenestratan pterosaur Cuspicephalus scarfi gen. et sp. nov. from the Upper Jurassic, Kimmeridge Bay, Dorset; MJML K1918. A. Posterior of skull showing near complete orbit. B. Distal tip of rostrum showing anteriorly directed alveolus 1. C. Rostrum anterior of nasoantorbital fenestra. D. Sagittal crest. Scale bars 10 mm.

Fig. 4.
A monofenestratan pterosaur Cuspicephalus scarfi gen et sp. nov. from the Upper Jurassic, Kimmeridge Bay, Dorset; MJML K1918. A. Left exoccipital and occipital condyle. Scale bar 10 mm. B. Outline of right jugal showing triangular shape of anteroventral border of orbit.

Table 2.
Rostral indices (RI) of selected non-pterodactyloid monofenestratans, germanodactylid pterosaurs, and the crested “rhamphorhynchoid” Pterorhynchus.

Fig. 5.
Sagittal crest of Cuspicephalus scarfi gen. et sp. nov. from the Upper Jurassic, Kimmeridge Bay, Dorset (A) compared with Darwinopterus modularis (Lü, Unwin, Jin, Liu, and Ji, 2010) (B) (from Lü et al. 2009b), Kunpengopterus sinensis Wang, Kellner, Jiang, Cheng, Meng, and Rodrigues, 2010 (C) (based on Wang et al. 2010), Germanodactylus cristatus (Wiman, 1925) (D) (based on Wellnhofer 1970), and the elaborate crest-bearing Pterorhynchus wellnhoferi Czerkas and Ji, 2002 (E) (based on Czerkas and Ji 2002). Drawings not to scale.

Fig. 6.
Variation in cranial angle (subtended between the squamosal-quadrate and jugal-maxilla-premaxilla jaw line from a selection of Jurassic monofenestratans. A. Cuspicephalus scarfi gen et sp. nov. from the Upper Jurassic, Kimmeridge Bay, Dorset. B. Darwinopterus robustodens Lü, Xu, Chang, and Zhang, 2011b. C. Darwinopterus linglongtaensis (Lü, Unwin, Jin, Liu, and Ji, 2010). D. Kunpengopterus sinensis Wang, Kellner, Jiang, Cheng, Meng, and Rodrigues, 2010. E. Pterodactylus antiquus Sömmerring, 1812. F. Germanodactylus cristatus (Wiman, 1925). G. Gnathosaurus subulatus Meyer, 1834. H. “Germanodactylus” rhamphastinus (Wagner, 1851). B after Lü et al. (2011b); C-D after Wang et al. (2010); E-H after Wellnhofer (1970).

KEYWORDS
Cuspicephalus
Jurassic
Kimmeridgian
Monofenestrata
Pterosauria
United Kingdom
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