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The clerine checkered beetle genus Dologenitus Opitz, new replacement name for PoecilochroaChevrolat, 1876 (Insecta: Coleoptera: Cleridae), a junior primary homonym of PoecilochroaWestring, 1874 (Arachnida: Araneae: Gnaphosidae), is proposed and described to include Clerus cyanipennis Klug, Clerus nigrocinctus Klug, and Enoclerus paludatus (Vaurie). The new genus is based on four synapotypic characters that distinguish these beetles from other New World clerines and establish the monophyletic status of the genus. The characters are: stout dorsal setae particularly notable on the elytral disc; females with a dagger-like ovipositor; absence of an interspicular plate; and the female reproductive system without a saccular bursa copulatrix. Lectotypes are designated for D. cyanipennis (Klug), and Eurycranus pulchellus Wolcott and Placopterus obscuripes Pic are considered its junior synonyms. Clerus retrocinctus Corporaal and Clerus latefasciatus Wolcott are synonymized with D. nigrocinctus (Klug). Differences in the aedeagus, integument color, and general body form were used as criteria to predict what magnitude of anatomical difference represents a biological species. Field observations, and laboratory dissections of the digestive tract, revealed that these beetles are omnivorous: they feed on pollen and other flower-visiting insects. It is suggested that Dologenitus represents an ancient line of clerines that does not have closely related relatives among extant New World Clerinae. A fully resolved intrageneric hypothesis of phylogeny is provided. The treatment includes a halftone frontispiece, 37 line drawings, six color photographs depicting adult beetles, a character matrix, a phylogenetic tree, and three distribution maps.
A key to the 18 genera of hispine beetles (Coleoptera: Chrysomelidae: Cassidinae) of the Caribbean Basin as well as a key to the seven genera and species of hispines known from Hispaniola are presented. Each species is briefly redescribed, known biological information is summarized, distributional information is listed, and a habitus photo of the adult of each Caribbean genus and all Hispaniola species is provided.
Leptarctus mummorum, new species, from the late Clarendonian of northwestern Kansas is described for the unusual mustelid genus LeptarctusLeidy, 1856. It is distinguished from other species of the genus by its larger size, temporal crests meeting along the central axis of the skull, relatively longer rostrum and bullar processes, depression anterior to the orbit, oblique ridge along the orbital wall, upper third premolar (P3) with an accessory internal accessory cusp, and upper fourth premolar (P4) with a greatly enlarged parastyle. It appears that previous suggestions that Leptarctus was strictly herbivorous and arboreal are not supportable based on the morphologies of the skull and dentition.
Fifteen Recent species and three Miocene species of Rhysodini (Coleoptera: Carabidae: Scaritinae) are named and described as new species: Yamatosa phuka (Thailand), Arrowina nan (Thailand), Arrowina taksar (Nepal), Omoglymmius (Omoglymmius) truncatus (Thailand), Omoglymmius (Omoglymmius) batantae (Irian Jaya), Rhyzodiastes (Temoana) orestes (Tibet), Rhyzodiastes (Rhyzostrix) ininius (Guyane), Rhyzodiastes (Rhyzostrix) exsequiae (Brazil), Clinidium (Clinidium) ashei (Panama), Clinidium (Clinidium) brusteli (Ecuador), Clinidium (Clinidium) chandleri (Costa Rica), Clinidium (Clinidium) dux (Philippines), Clinidium (Clinidium) erwiniCosta Rica), Clinidium (Clinidium) felix (Panama), and Clinidium (Clinidium) talamanca (Costa Rica), and the Dominican Republic amber fossils, Clinidium (Clinidium) poinari, Plesioglymmius (Ameroglymmius) zayasi, and Clinidium (Tainoa) grimaldii. The males of Rhyzodiastes (Temoana) denticauda R. Bell and J. Bell and Clinidium (Clinidium) segne R. Bell and J. Bell, previously known only from females, are described. The females of Yamatosa schawalleri R. Bell and J. Bell and Clinidium (Clinidium) impressum R. Bell and J. Bell, previously known from males, are described. Range extensions or clarifications are given for the following 30 species: Leoglymmius lignarius (Olliff), Medisores abditus R. Bell and J. Bell, Dhysores thoreyi (Grouvelle), Rhysodes sulcatus Fabricius, Yamatosa boysi (Arrow), Yamatosa reitteri (Bell), Yamatosa sinensis R. Bell and J. Bell, Plesioglymmius (Juxtaglymmius) jugatus R. Bell and J. Bell, Omoglymmius (Orthoglymmius) cavifrons (Grouvelle), Omoglymmius (Orthoglymmius) crenatus (Grouvelle), Omoglymmius (Omoglymmius) laticeps R. Bell, Omoglymmius (Omoglymmius) germari Ganglbauer, Omoglymmius (Omoglymmius) vicinus (Grouvelle), Grouvellina tubericeps (Fairemaire), Grouvellina radama R. Bell and J. Bell, Rhyzodiastes (Temoana) preorbitalis R. Bell and J. Bell, Rhyzodiastes (Temoana) denticauda R. Bell and J. Bell, Clinidium (Mexiclinidium) championi R. Bell and J. Bell, Clinidium (Mexiclinidium) newtoni R. Bell and J. Bell, Clinidium (Clinidium) centrale Grouvelle, Clinidium (Clinidium) granatense Chevrolat, Clinidium (Clinidium) insigne Grouvelle, Clinidium (Clinidium) integrum Grouvelle, Clinidium (Clinidium) jolyi R. Bell and J. Bell, Clinidium (Clinidium) mathani Grouvelle, Clinidium (Clinidium) moldenkei R. Bell and J. Bell, Clinidium (Clinidium) oberthueri Grouvelle, Clinidium (Clinidium) rossi R. Bell, Clinidium (Clinidium) segne R. Bell and J. Bell, and Clinidium (Clinidium) spatulatum R. Bell and J. Bell. Four revised keys are presented: Key to Species of Genus Arrowina; Key to Genera of Subtribe Clinidiina; Key to Species of Subgenus Rhyzodiastes (Rhyzostrix); and Key to Species of genus Clinidium (Clinidium). A discussion of the phylogeny of Clinidium (Clinidium) is given. The terminology for the antennal setae of Omoglymmius americanus (Laporte), as used by Bell and Bell, is compared with that of Baker (2001).