This catalog (published in seven parts, all released on the same day) is based on only the published literature for the Staphylinidae. Of the 32 subfamilies, the following 28 are included herein: Apateticinae, Dasycerinae, Empelinae, Euaesthetinae, Glypholomatinae, Habrocerinae, Leptotyphlinae, Megalopsidiinae, Micropeplinae, Microsilphinae, Neophoninae, Olisthaerinae, Omaliinae, Osoriinae, Oxyporinae, Oxytelinae, Phloeocharinae, Piestinae, Protactinae†, Proteininae, Protopselaphinae, Pseudopsinae, Solieriinae, Staphylininae, Steninae, Tachyporinae, Trichophyinae, and Trigonurinae. The Aleocharinae, Paederinae, Pselphinae, and Scaphidiinae are excluded from this edition of the catalog. References to the original citation or description are given for available family-group, genus-group, and species-group names of both extant and extinct forms. The type genus is cited for each family-group name, the type species for each genus-group name, and the type locality for each species-group name. Where appropriate, all subgenera, subspecies, or synonyms are listed for each valid name. Annotated subsequent references are presented for all names. Distributional summaries are given for each valid taxon. Full bibliographic citations are in Part VII. A short historical review, coauthored with Aleš Smetana, follows the Introduction (Part I), with the main focus on biographical sketches that include many photographs. The goal of this catalog is to summarize the current state of knowledge of the family and to stimulate worldwide monographic studies.
STAPHYLININAE
[294 genera; 6665 species; All Regions except the Antarctic]
Staphylininae Latreille, 1802.
— Eichelbaum, 1909: 159 (list of genera with references; generic synonyms; number of genera; number of species in each genus; general distribution of genera).
— Reitter, 1909: 105 (key to tribus).
— Cameron, 1921a (characters; key to Singapore species and genera).
— Šulc, 1922 (osmeterium; anatomy; histology; physiology).
— Porta, 1926: 85 (characters; species of Italy).
— Böving and Craighead, 1930: 30 (larval characters).
— Cameron, 1932: 1 (characters; key to tribes).
— Joy, 1932: 105 (key to British tribes).
— Scheerpeltz, 1933: 1297 (world catalog supplement).
— Voris, 1940 (phylogenetic relationships based on larvae).
— Voris, 1939a: 289 (egg, larval and pupal characters).
— Bernet Kempers, 1941: 35 (larval characters).
— Scheerpeltz, 1940: 38 (key to Palaearctic genera).
— Blackwelder, 1943: 393 (notes; key to tribes of West Indies).
— Blackwelder, 1944: 131 (checklist of species from Mexico, Central and South America, and the West Indies).
— V. Hansen, 1952: 1 (characters; Danish species).
— Crowson, 1955: 34 (adult and larval characters in key).
— Hatch, 1957: 48, 172 (characters; key to tribes and genera of Pacific Northwest).
— Arnett, 1963: 246, 274 (key to North American genera; characters; list of North American genera).
— Székessy, 1963: 46 (characters; key to genera of Hungary).
— Palm, 1963b: 38 (characters; key to tribe, genera, and species of Sweden).
— Steel, 1963: 54 (list of species of Malham Tarn; collecting notes; Britain).
— Lohse, 1964: 169 (characters; key to central European species).
— Moore, 1965: 97 (key to Nearctic genera).
— Seevers, 1965: 323 (myrmecophilus species; notes on classification).
— Kasule, 1966: 266, 274 (larval characters included Xantholinini; notes).
— Pototskaia, 1967: 12 (larval characters; key to tribe).
— Coiffait and Saiz, 1968: 346 (key to tribes and genera of Chile).
— Steel and Woodroffe, 1969: 136 (list of species collected from Isle of Rhum; collecting notes).
— Britton, 1970: 545 (characters of adults; Australia; notes).
— Kasule, 1970: 56 (larval characters).
— Moore and Legner, 1973c: 212 (key to genera of United States and Canada).
— Moore and Legner, 1974g: 560 (key to genera of America north of Mexico).
— Moore and Legner, 1974h: 470 (catalog of the taxonomy, biology, and ecology of developmental stages of species of America north of Mexico).
— Coiffait, 1974: 1 (characters; key to tribes, genera and species of western Palaearctic region).
— Moore and Legner, 1975: 6 (catalog; North America north of Mexico).
— Topp, 1978: 307, 311 (larval characters; key to genera).
— Moore and Legner, 1979: 14 (characters; tabular key to genera of North America north of Mexico).
— Muona, 1979: 14 (list of Scandinavian species).
— Burakowski, Mroczkowski, and Stefańska, 1980: 29 (catalog; Poland).
— Spahr, 1981: 93 (references to specimens from amber and copal).
— Watt and McColl, 1982: 19, 20 (characters; notes on New Zealand taxa).
— Pajni and Kohli, 1982: 10 (comparative morphology of aedeagus).
— E. Matthews, 1982: 7, 24 (notes; key to genera of Australia).
— Tóth, 1983: 183 (list of species of the Hortobágy National Park, Hungary).
— Tóth, 1984 (characters; key to genera and species of Hungary).
— Outerelo and Gamarra, 1985: 47 (key to genera of Iberian Peninsula).
— Tóth, 1985a: 88 (list of species of Bakony Mountains, Hungary).
— Dettner, 1987: 30 (defensive glands).
— Drugmand 1987b (comments on distribution of species in Belgium).
— Gusarov, 1989 (list of species from the Crimea; ecological and habitat notes).
— Frank, 1991: 346 (larval characters).
— Frania, 1992 (studies on ecdysis and metamorphosis).
— Newton and Thayer, 1992: 13 (summary of current classification; discussion).
— Ashe and Newton, 1993: 281 (related to Paederinae and Pseudopsinae).
— Dettner, 1993a (summary and discussion of defensive secretions and exocrine glands and their bearing on phylogeny).
— Hayashi, 1993b: 282 (terms for taxonomic characters).
— Welch, 1993 (ovariole structure and development).
— Lawrence and Newton, 1995: 830 (notes).
— Sparacio, 1995: 160 (characters; species of Sicily).
— Klimaszewski, Newton, and Thayer, 1996: 150 (review of subfamily for New Zealand; notes; list of New Zealand genera; number of New Zealand species).
— M. Hansen, 1996: 37 (listed in Staphylinine group of subfamilies).
— M. Hansen, 1996: 104 (list of Danish species).
— Staniec, 1996: 117 (key to known pupae of Poland).
— Downie and Arnett, 1996: 371 (characters; key to genera of Northeastern North America).
— Ádám, 1996: 244 (list of species collected in Bükk National Park, Hungary).
— Hayashi, 1997: 27 (characters of Eucibdelus group; genera included: Philetaerius, Trichocosmetes, Eucibdelus, Phytolinus, Rhychocheilus).
— M. Hansen, 1997: 83, 134, 176, 177, 180 (characters supporting clade; autapomorphic features; key adult and larval characters; notes).
— Anderson, Nash, and O'Connor, 1997: 18 (annotated checklist; Ireland).
— Solodovnikov, 1998: 337 (list of species in northwest Caucasus; Russia).
— Staniec, 1999a: 55 (key to known pupae of Poland).
— Schmidt, 1999 (methods for rearing species).
— Smetana and Davies, 2000: 18 (key to genera and subgenera associated with Staphylinus sensu lato).
— Newton, Thayer, Ashe, and Chandler, 2000: 328, 389 (key to Nearctic genera; characters; notes).
— Klimaszewski, 2000: 23, 67 (characters; notes on classification, natural history, distribution, and key to tribes in Canada and Alaska).
Staphylinitae:
Tribe Diochini
Diochini Casey, 1906.
— Smetana, 1982: 26 (characters; revision of genera and species of North America north of Mexico).
— Outerelo and Gamarra, 1985: 47 (genera of Iberian Peninsula).
— Newton and Thayer, 1992: 64 (tribe of Staphylininae).
— Newton, Thayer, Ashe, and Chandler, 2000: 390 (characters; notes).
Diochi Casey, 1906: 359, 429 (genera included: Diochus). Type genus: Diochus (see Newton and Thayer, 1992: 64).
— Blackwelder, 1944: 143 (checklist of species from Mexico, Central and South America, and the West Indies).
Diochina Casey, 1906.
Diochinae Moore, 1964a: 89 (genera included: Diochus, Ophioomma). Type genus: Diochus (see Newton and Thayer, 1992: 64).
— Newton and Thayer, 1992: 64 (synonym of Diochini).
— Moore and Legner, 1974g: 559 (key to genera of America north of Mexico).
— Moore and Legner, 1975: 69 (catalog; North America north of Mexico).
— Moore and Legner, 1979: 49 (characters).
— Lawrence and Newton, 1995: 830 (included in Staphylininae).
Diochini Coiffait, 1972: 127, 368 (tribe of Xantholininae; genera included: Diochus). Type genus: Diochus (see Newton and Thayer, 1992: 64).
Antarctothius
[2 species; Neotropical Region]
Antarctothius Coiffait and Saiz, 1970: 279 (species included: antarcticus, fuegius). Type species: Othius antarcticus Fairmaire, fixed by original designation.
antarcticus Fairmaire, 1885: 12 (Othius; Type locality: Terre de Feu).
— Bernhauer and Schubert, 1914: 317 (Othius; catalog).
— Scheerpeltz, 1933: 1321 (Othius; catalog).
— Blackwelder, 1944: 145 (Othius; Argentina).
— Coiffait, Saiz, 1970: 281 (Antarctothius; characters; Tierra del Fuego).
— Saiz, 1970b: 390 (Antarctothius; Chile).
▴ Distribution: Chile, Argentina.
fuegius Coiffait and Saiz, 1970: 285 (Antarctothius; Type locality: Bahia Orange, Terre de Feu).
Coomania
[1 species; Oriental Region]
Coomania Cameron, 1939a: 22 (species included: tonkinensis). Type species: Coomania tonkinensis Cameron, fixed by monotypy.
tonkinensis Cameron, 1939a: 23 (Coomania; Type locality: Tonkin: Hoa Binh).
Diochus
[38 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Oriental, Australian, and Oceanic Regions]
Diochus Erichson, 1839b: 300 (species included: nanus). Type species: Diochus nanus Erichson, fixed by monotypy.
— Lacordaire, 1854: 65 (characters; notes; list of species).
— LeConte, 1861a: 65 (characters).
— LeConte, 1863: 24 (list of North American species).
— Gemminger and Harold, 1868: 608 (catalog).
— Redtenbacher, 1874: 223 (characters).
— Fauvel, 1875a: XXIV [= 1875b: 226] (catalog).
— Sharp, 1876: 184 (notes).
— Duvivier, 1883: 160 (catalog).
— Lynch, 1884: 171 (characters).
— Sharp, 1885: 466 (notes; Mexican and Central American species).
— Olliff, 1887: 472 (characters).
— Heyden, Reitter, and Weise, 1891: 105 (list of species of Europe and Caucasus).
— Casey, 1906: 430 (characters; key to species).
— Heyden, Reitter, and Weise, 1906: 163 (list of species of Europe and Caucasus).
— Reitter, 1908: 103 (characters).
— Blatchley, 1910: 400 (characters; key to Indiana species).
— Bernhauer and Schubert, 1914: 319 (world catalog; 21 species).
— Leng, 1920: 106 (catalog of North American species).
— Cameron, 1921a: 354, 404 (characters; key to Singapore species; catalog).
— Winkler, 1925: 373 (catalog for Palaearctic region).
— Cameron, 1925: 53 (catalog of Indian species).
— Bruch, 1928a: 427 (catalog; Argentina).
— Cameron, 1932: 44 (characters; key to species of British India).
— Scheerpeltz, 1933: 1323 (world catalog supplement).
— Normand, 1935: 370 (list of species of Tunisia).
— Blackwelder, 1943: 455 (characters; type species: nanus).
— Blackwelder, 1944: 143 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 128 (type species: nanus).
— Coiffait, 1956a: 37 (note).
— Kocher, 1958: 142 (checklist of species; Morocco).
— Moore, 1963a: 94 (characters).
— Coiffait, 1968: 156 (key to species of Gabon; notes).
— Coiffait, 1972: 368 (characters; type species: nanus; key to species of western Palaearctic region; references).
— Shibata, 1973c: 55 (checklist of species of Taiwan).
— Tikhomirova, 1973a: 180 (checklist of species of USSR).
— Moore and Legner, 1975: 69 (catalog; North America north of Mexico).
— Moore and Legner, 1979: 49 (characters; notes).
— Smetana, 1982: 27 (characters; habitus; revision species of North America north of Mexico).
— Shibata, 1983: 86 (checklist of species of Japan).
— Hammond, 1984: 205 (checklist of species of Borneo).
— Frank, 1986a: 369 (checklist of species of Florida).
— Downie and Arnett, 1996: 395 (characters).
— Lundgren, 1998: 50 (list of species from Florida; records for other States listed for each species).
— Newton, Thayer, Ashe, and Chandler, 2000: 390 (1 Nearctic species; notes; characters in key).
Rhegmatocerus Motschulsky, 1858: 657 (species included: conicollis, punctipennis, antennatus). Type species: Rhegmatocerus punctipennis Motschulsky, fixed by subsequent designation by Blackwelder, 1943: 455.
— Gemminger and Harold, 1868: 608 (synonym of Diochus).
— Lynch, 1884: 171 (synonym of Diochus).
— Bernhauer and Schubert, 1914: 319 (synonym of Diochus).
— Cameron, 1932: 44 (synonym of Diochus).
— Blackwelder, 1943: 455 (synonym of Diochus; type species: punctipennis).
— Blackwelder, 1952: 339 (type species).
— Coiffait, 1972: 368 (synonym of Diochus).
— Smetana, 1982: 27 (synonym of Diochus).
antennalis Cameron, 1922, see: nanus Erichson, 1839.
antennatus Motschulsky, 1858: 659 (Rhegmatocerus; Type locality: Indes orientales).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Bernhauer, 1922c: 231 (Diochus; Formosa).
— Bernhauer, 1928: 15 (Diochus; Sumatra).
— Cameron, 1932: 46 (Diochus; characters; Penang; Malay Peninsula).
— Scheerpeltz, 1933: 1323 (Diochus; catalog).
— Cameron, 1933d: 385 (Diochus; Sumatra).
— Cameron, 1934d: 79 (type examined).
— Cameron, 1937: 4 (Diochus; Java).
— Shibata, 1973a: 130 (Diochus; Taiwan).
— Coiffait, 1982: 27 (Diochus; characters; Nepal).
▴ Distribution: “East Indies”, Nepal, Malaysia, Indonesia, New Guinea, Taiwan.
indicus Kraatz, 1859: 113 (Diochus; Type locality: India orientalis).
— Bernhauer and Schubert, 1914: 319 (Diochus; synonym of antennatus).
— Cameron, 1931: 360 (Diochus; New Guinea).
— Cameron, 1932: 46 (Diochus; synonym of antennatus).
— Cameron, 1934d: 79 (Diochus; synonym of antennatus).
apicipennis Cameron, 1922, see: nanus Erichson, 1839.
astutus Casey, 1906: 433 (Diochus; Type locality: South Africa: Wellington).
bicolor Scheerpeltz, 1974: 131 (Diochus; Type locality: Cape Prov.: Cape Peninsula, Cape Point Nature Reserve).
bicoloripennis Coiffait, 1981f: 333 (Diochus; Type locality: South Andaman, Jolly Boy Island).
borneensis Cameron, 1933b: 344 (Diochus; Type locality: British North Borneo: Kenokok, 3300 feet).
brevipennis Casey, 1906, see: schaumii Kraatz, 1860.
brunniventris Cameron, 1938: 8 (Diochus; Type locality: Belgian Congo: Eala).
communis Coiffait, 1968: 159 (Diochus; Type locality: Gabon: Belinga).
conicollis Motschulsky, 1858: 658 (Rhegmatocerus; Type locality: Indes orientales).
— Bernhauer, 1902b: 32 (Diochus; Ceylon).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Bernhauer, 1922c: 231 (Diochus; Formosa).
— Cameron, 1932: 45 (Diochus; characters; Ceylon; India).
— Scheerpeltz, 1933: 1324 (Diochus; catalog).
— Shibata, 1973a: 131 (Diochus; Taiwan).
— Coiffait, 1977b: 206 (Diochus; Bhutan).
— Coiffait, 1981f: 329 (Diochus; Andaman Islands).
— Coiffait, 1982: 27 (Diochus; Nepal).
— Frank, 1982: 35 (Diochus; list of parasites).
▴ Distribution: “East Indies”, Sri Lanka, India, Bhutan, Andaman Islands, Taiwan.
major Kraatz, 1859: 113 (Diochus; Type locality: India orientalis; Ceylan).
— Bernhauer, 1902b: 32 (Diochus; synonym of conicollis).
— Bernhauer and Schubert, 1914: 319 (Diochus; synonym of conicollis).
— Cameron, 1932: 45 (Diochus; synonym of conicollis).
— Cameron, 1934d: 79 (Diochus; synonym of conicollis).
cristobalensis Coiffait, 1981g: 295 (Diochus; Type locality: Ile Galapagos: Ile San Cristobal, montagne près Progreso).
debilis Coiffait, 1968: 159 (Diochus; Type locality: Gabon: Belinga).
dilutus Reitter, 1908, see: staudingeri Kraatz, 1860.
divisus Fauvel, 1877: 235 (Diochus; Type locality: Australie, Nouvelle Galles du Sud).
— Olliff, 1887: 473 (Diochus; characters; Australia).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Lea, 1925: 220 (Diochus; characters; Australia).
▴ Distribution: Australia.
dundoensis Cameron, 1959: 115 (Diochus; Type locality: Angola: Dundo; Sierra Leone: Freetown).
flandrianus Bernhauer, 1932b: 145 (Diochus; Type locality: Equateur: Flandria).
flavicans Sharp, 1876, see: nanus Erichson, 1839.
formicetorum Bernhauer, 1927e: 244 (Diochus; Type locality: Argentinien: Prov. Santa Fe, Fives Lilles).
franzi Coiffait, 1981g: 294 (Diochus; Type locality: Ile Galapagos: Ile Santiago).
gabonicus Coiffait, 1968: 157 (Diochus; Type locality: Gabon: Makokou).
indicus Kraatz, 1859, see: antennatus Motschulsky, 1858.
inornatus Sharp, 1885, see: nanus Erichson, 1839.
isabelae Coiffait, 1981g: 293 (Diochus; Type locality: Ile Galapagos: Ile Isabela, Sierra Negra).
japonicus Cameron, 1930c: 206 (Diochus; Type locality: Kobe).
— Scheerpeltz, 1933: 1324 (Diochus; catalog).
— Nakane, 1963a: 90 (Diochus; characters; habitus photograph; Japan).
— J. Li, 1993: 32 (Diochus; China).
▴ Distribution: Japan, China.
libanoticus Fagel, 1966a: 43 (Diochus; Type locality: Liban: Kartaba, vallée du Nahr Ibrahim, 1000 m).
longicornis Sharp, 1876, see: nanus Erichson, 1839.
longicornis Cameron, 1952, see: petilus Herman, 2001.
longus Lea, 1929: 203 (Diochus; Type locality: Western Australia: Bunbury).
machadoi Cameron, 1959: 115 (Diochus; Type locality: Angola: Dundo; Vila Luso; Luachimo Forest).
maculicollis Fauvel, 1891, see: nanus Erichson, 1839.
major Kraatz, 1859, see: conicollis Motschulsky, 1858.
montguilloni Coiffait, 1987: 497 (Diochus; replacement name for testaceus Coiffait).
testaceus Coiffait, 1981g: 296 (Diochus; [preoccupied]; Type locality: Ile Galapagos: Ile Santiago, Montagne).
nanus Erichson, 1839b: 301 (Diochus; Type locality: Columbia; [Note: See lectotype designation by Smetana, 1977c]. Also cited from Americae meridionalis regiones).
— Chevrolat and Fauvel, 1863: 430 (Diochus; characters; Cuba, Colombia).
— Solsky, 1872: 308 (Diochus; Cayenne).
— Lynch, 1884: 172 (Diochus; characters; Argentina).
— Sharp, 1885: 466 (Diochus; notes; Guatemala; Panama; Colombia).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Blackwelder, 1943: 455 (Diochus; characters; Cuba; Jamaica; Dominican Republic; Puerto Rico; St. Croix; Guadeloupe; St. Lucia; Barbados; St. Vincent; Grenadines; Grenada; Trinidad).
— Smetana, 1977c: 347 (Diochus; lectotype designation: Colombia; characters; notes).
— Frank, 1982: 35 (Diochus; list of parasites).
▴ Distribution: Argentina, Brazil, French Guiana, Venezuela, Colombia, Panama, Nicaragua, Guatemala, Mexico, West Indies.
parvulus Kraatz, 1860: 27 (Diochus; Type locality: Bahia).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Blackwelder, 1943: 456 (Diochus; synonym of nanus).
longicornis Sharp, 1876: 184 (Diochus; Type locality: Tapajos).
— Sharp, 1887: 791 (Diochus; Mexico).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Blackwelder, 1943: 456 (Diochus; synonym of nanus).
vicinus Sharp, 1876: 185 (Diochus; Type locality: Tapajos).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Blackwelder, 1943: 456 (Diochus; synonym of nanus).
tarsalis Sharp, 1876: 185 (Diochus; Type locality: Tapajos).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Scheerpeltz, 1933: 1324 (Diochus; catalog).
— Blackwelder, 1943: 456 (Diochus; synonym of nanus).
flavicans Sharp, 1876: 185 (Diochus; Type locality: Tapajos).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Blackwelder, 1943: 456 (Diochus; synonym of nanus).
inornatus Sharp, 1885: 466 (Diochus; Type locality: Nicaragua, Chontales; Panama, Bugaba 800–1500 feet, Tolé, Peña Blanca 3000–4000 feet).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Blackwelder, 1943: 456 (Diochus; synonym of nanus).
vilis Sharp, 1885: 467 (Diochus; Type locality: Guatemala, near the city).
— Bernhauer and Schubert, 1914: 320 (Diochus; catalog).
— Blackwelder, 1943: 456 (Diochus; synonym of nanus).
maculicollis Fauvel, 1891: 106 (Diochus; proposed for the species Kraatz misidentified as nanus; Type locality: Caracas; Colonie Tovar).
— Kraatz, 1860: 27 (Diochus; [Note: Misidentification: Bernhauer and Schubert, 1914: 319, cited nanus Kraatz as a synonym of maculicollis; Kraatz did not describe the species as new, he attributed it to Erichson]; characters; Venezuela).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Blackwelder, 1943: 456 (Diochus; synonym of nanus).
perplexus Cameron, 1922: 116 (Diochus; Type locality: Grenada).
apicipennis Cameron, 1922: 116 (Diochus; Type locality: St. Vincent).
antennalis Cameron, 1922: 117 (Diochus; Type locality: Grenada).
pumilio Bernhauer, 1929h: 193 (Diochus; Type locality: Mexiko: Veracruz).
ochraceus Cameron, 1926b: 347 (Diochus; Type locality: Siwaliks, Mothronwala).
— Cameron, 1932: 47 (Diochus; characters; India).
— Scheerpeltz, 1933: 1324 (Diochus; catalog).
▴ Distribution: India.
octavii Fauvel, 1877: 235 (Diochus; Type locality: Australe, Victoria).
— Fauvel, 1878g: 535 (Diochus; Australia).
— Olliff, 1887: 472 (Diochus; characters; Australia).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Bernhauer, 1920: 16 (Diochus; Australia).
— Lea, 1925: 220 (Diochus; characters; Australia).
▴ Distribution: Australia.
pallidiceps Casey, 1906, see: schaumii Kraatz, 1860.
papuanus Cameron, 1939b: 145 (Diochus; Type locality: New Guinea: Kokoda, altitude 1200 feet).
parvulus Kraatz, 1860, see: nanus Erichson, 1839.
perplexus Cameron, 1922, see: nanus Erichson, 1839.
petilus Herman, 2001: 43 (Diochus; replacement name for longicornis Cameron).
longicornis Cameron, 1952a: 328 (Diochus; [preoccupied]; Type locality: Belgian Congo: Katanga, galerie forestière de la Kisanga).
proximus Cameron, 1952a: 328 (Diochus; Type locality: Belgian Congo: Massif des Kundelungu).
pubiventris Lea, 1929: 203 (Diochus; Type locality: North Australia: Darwin; New South Wales: Tamworth).
— Scheerpeltz, 1933: 1324 (Diochus; catalog).
— Scheerpeltz, 1935a: 649 (Diochus; Australia).
▴ Distribution: Australia.
pulchellus Cameron, 1918: 84 (Diochus; Type locality: Singapore: Sembawang).
— Cameron, 1921a: 375, 404 (Diochus; characters; Singapore).
— Scheerpeltz, 1933: 1324 (Diochus; catalog).
▴ Distribution: Singapore.
pumilio Bernhauer, 1929, see: nanus Erichson, 1839.
punctipennis Motschulsky, 1858: 659 (Rhegmatocerus; Type locality: Indes orientales).
— Fauvel, 1898a: 117 (Diochus; Seychelles).
— H. Kolbe, 1910: 20 (Diochus; Seychelles).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Bernhauer, 1922b: 176 (Diochus; Seychelles).
— Cameron, 1932: 46 (Diochus; characters; India).
— Scheerpeltz, 1933: 1324 (Diochus; catalog).
— Cameron, 1934d: 79 (Diochus; type examined).
— Cameron, 1937: 4 (Diochus; Java).
— Scheerpeltz, 1957c: 233 (Diochus; Sumba).
— Coiffait, 1981f: 329 (Diochus; Andaman Islands).
— Lecoq, 1990: 189 (Diochus; Mascarene Islands).
▴ Distribution: Mascarene Islands, Seychelles, Andaman Islands, India, Indonesia.
santiagus Coiffait, 1981g: 294 (Diochus; Type locality: Ile Galapagos: Ile Santiago).
schaumii Kraatz, 1860: 27 (Diochus; Type locality: Louisiana; [Note: See lectotype designation by Smetana, 1982]. Also cited from Georgia).
— Sharp, 1885: 467 (Diochus; Louisiana; Guatemala; Nicaragua).
— Casey, 1906: 431 (Diochus; characters; Rhodes Island; North Carolina; Missouri; Texas).
— Blatchley, 1910: 400 (Diochus; characters; Indiana).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Notman, 1920b: 704 (Diochus; Florida).
— Blackwelder, 1943: 456 (Diochus; synonym of nanus).
— Smetana, 1977c: 348 (Diochus; valid species; characters).
— Smetana, 1982: 29 (Diochus; lectotype designation: Louisiana; characters; Ontario; Quebec; Alabama; Arkansas; Connecticut; Florida; Georgia; Illinois; Indiana; Iowa; Kansas; Kentucky; Louisiana; Maine; Maryland; Massachusetts; Michigan; Mississippi; Missouri; New Hampshire; New Jersey; New York; North Carolina; Ohio; Oklahoma; Pennsylvania; Rhode Island; South Carolina; Texas; Virginia; West Virginia; Wisconsin).
— Campbell and Davies, 1991: 123 (Diochus; checklist of species; Canada).
— Downie and Arnett, 1996: 395 (Diochus; characters; USA).
— Schmidt, 1999: 108 (Diochus; food for adults and larvae used for rearing species; rearing methods).
▴ Distribution: Canada, USA, Guatemala, Nicaragua.
brevipennis Casey, 1906: 431 (Diochus; Type locality: Missouri; [Note: See lectotype designation by Smetana, 1982]. Also cited from Virginia; North Carolina, Asheville).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Blackwelder, 1943: 456 (Diochus; synonym of nanus).
— Smetana, 1982: 29 (Diochus; lectotype designation: Mo. [Missouri]; synonym of schaumii).
pallidiceps Casey, 1906: 432 (Diochus; Type locality: Texas, Austin; [Note: See lectotype designation by Smetana, 1982]).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Blackwelder, 1943: 456 (Diochus; synonym of nanus).
— Smetana, 1982: 29 (Diochus; lectotype designation; synonym of schaumii).
thoracicus Casey, 1906: 432 (Diochus; Type locality: New York, Long Island; [Note: See lectotype designation by Smetana, 1982]).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Blackwelder, 1943: 456 (Diochus; synonym of nanus).
— Smetana, 1982: 29 (Diochus; lectotype designation; synonym of schaumii).
senegalensis Fauvel, 1900b: 72 (Diochus; Type locality: Congo: Kinchassa; Boma. Haut-Senegal: Badumbe).
staudingeri Kraatz, 1860: 27 (Diochus; Type locality: Andalusia).
— Redtenbacher, 1874: 223 (Diochus; characters).
— Fauvel, 1878: 115 [= 1878a: 35] (Diochus; Algeria).
— Fauvel, 1886: 47 [= 1886a: 39] (Diochus; Algeria).
— Fauvel, 1897d: 289 (Diochus; Algeria).
— Fauvel, 1902b: 98 (Diochus; Algeria; Tunisia).
— Reitter, 1908: 108 (Baptolinus; characters; Andalusia).
— Bernhauer and Schubert, 1914: 319 (Diochus; catalog).
— Gridelli, 1931a: 157 (Diochus; characters; Algeria).
— Scheerpeltz, 1933: 1324 (Diochus; catalog).
— Coiffait, 1972: 369 (Diochus; characters; references; North Africa; Spain; France).
▴ Distribution: Portugal, Spain, Algeria, Tunisia.
dilutus Reitter, 1908: 108 [= 1908a: 11] (Diochus; variety of staudingeri; Type locality: Andalusien).
— Scheerpeltz, 1933: 1324 (Diochus; cited as dilutior; synonym of staudingeri).
— Coiffait, 1972: 369 (Diochus; cited as dilutior; variety of staudingeri; characters; Portugal).
tahitiensis Coiffait, 1976: 225 (Diochus; Type locality: Tahiti: Vallée Papenoo).
tarsalis Sharp, 1876, see: nanus Erichson, 1839.
testaceus Bernhauer, 1932b: 145 (Diochus; Type locality: Mulongo: Malinge; Niunzu).
testaceus Coiffait, 1981, see: montguilloni Coiffait, 1987.
thoracicus Casey, 1906, see: schaumii Kraatz, 1860.
ventralis Coiffait, 1968: 160 (Diochus; Type locality: Gabon: Belinga).
vicinus Sharp, 1876, see: nanus Erichson, 1839.
vilis Sharp, 1885, see: nanus Erichson, 1839.
Tribe Maorothiini
Maorothiini Assing, 2000: 16 (genera included: Maorothius; characters; phylogenetic analysis of tribal relationships). Type genus: Maorothius Assing, fixed by original designation.
Maorothius
[20 species; Australian Region]
Maorothius Assing, 2000: 22, 58 (species included: adustus, hamifer, setiger, longispinosus, brevispinosus, insulanus, pubescens, brookesi, effeminatus, hammondi, pectinatus, puncticeps, coalitus, tridens, volans, dispar, brouni, torquatus, tonsor, solus; phylogenetic analysis; key to species). Type species: Othius adustus Broun, fixed by original designation.
adustus Broun, 1880: 101 (Othius; Type locality: Parua; [Note: See lectotype designation by Assing, 2000]).
— Bernhauer and Schubert, 1914: 317 (Othius; catalog).
— Assing, 2000: 24 (Maorothius; lectotype designation: Parua, Whangarei, New Zealand; characters; notes; New Zealand).
▴ Distribution: New Zealand.
angustus Broun, 1893, see: brouni Steel, 1949.
brevispinosus Assing, 2000: 32, 63 (Maorothius; Type locality: AK: Ahuroa forest reserve).
brookesi Cameron, 1952c: 42 (Othius; Type locality: New Zealand: North Island: Maku River, Bay of Plenty).
brouni Steel, 1949a: 270 (Gyrohypnus; replacement name for angustus Broun).
angustus Broun, 1893: 1026 (Othius; [preoccupied]; Type locality: Taieri).
— Bernhauer and Schubert, 1914: 317 (Othius; catalog).
— Steel, 1949a: 270 (Gyrohypnus; synonym of brouni).
— Assing, 2000: 51 (Maorothius; synonym of brouni).
coalitus Assing, 2000: 45 (Maorothius; Type locality: New Zealand: NN, Beebys Knob, 1220 m).
dispar Assing, 2000: 49 (Maorothius; Type locality: L. Rotoroa, 1200′).
effeminatus Assing, 2000: 39, 63 (Maorothius; Type locality: N. Z.: Whangarei).
hammondi Assing, 2000: 41 (Maorothius; Type locality: New Zealand: WO: Kauri Flat, 14 km. SW Waingaro).
insulanus Assing, 2000: 33 (Maorothius; Type locality: New Zealand: Lt Barrier).
hamifer Assing, 2000: 27 (Maorothius; Type locality: Waipu).
longispinosus Assing, 2000: 30, 63 (Maorothius; Type locality: N-Zealand: Aukland [sic]).
pectinatus Assing, 2000: 42 (Maorothius; Type locality: New Zealand: ND, Russell SF, Runaruku).
pubescens Assing, 2000: 34 (Maorothius; Type locality: Waitotara, Wanganui).
puncticeps Broun, 1894: 419 (Othius; Type locality: Mount Pirongia).
— Bernhauer and Schubert, 1914: 318 (Othius; catalog).
— Assing, 2000: 43 (Maorothius; characters; notes; New Zealand).
▴ Distribution: New Zealand.
setiger Assing, 2000: 29, 62 (Maorothius; Type locality: N. Zeal.: Waipoua For., Waipoua Strm., 70 m).
solus Assing, 2000: 57 (Maorothius; Type locality: New Zealand WN, Kowhai Park, Brooklyn).
tonsor Assing, 2000: 56, 63 (Maorothius; Type locality: New Zeal.: S Isl., Banks Penins).
torquatus Assing, 2000: 55 (Maorothius; Type locality: Oaro, N. Canterbury).
tridens Assing, 2000: 46 (Maorothius; Type locality: WN: Desert Rd, S. Turangi, Mangatawai Stm.).
volans Assing, 2000: 47 (Maorothius; Type locality: Maungatapere Bush).
Tribe Othiini
Othiini Thomson, 1859.
— Reitter, 1908: 102 (characters).
— Portevin, 1929: 367 (characters; key to French genera).
— Scheerpeltz, 1933: 1297 (synonym of Xantholinini).
— Jeannel and Jarrige, 1949: 371 (cave species).
— Jarrige, 1952a: 131 (characters).
— Coiffait, 1956a: 37 (review of French taxa; characters).
— Lohse, 1964: 166 (characters; key to central European species).
— Scheerpeltz, 1968a: 63 (catalog of Austrian species).
— Coiffait, 1972: 127, 326 (tribe of Xantholininae; characters; key to genera and species of western Palaearctic region).
— Muona, 1979: 16 (list of Scandinavian species).
— Smetana, 1982: 35 (characters; revision of and key to genera and species of North America north of Mexico).
— Bordoni, 1982: 378 (key to genera of Italy).
— Newton, 1985: 208 (notes on austral distribution and taxonomic notes).
— Outerelo and Gamarra, 1985: 43 (key to genera of Iberian Peninsula).
— Newton and Thayer, 1992: 64 (tribe of Staphylininae).
— Klimaszewski, Newton, and Thayer, 1996: 150 (list of New Zealand genera; number of New Zealand species).
— M. Hansen, 1996: 108 (list of Danish species).
— Downie and Arnett, 1996: 396 (characters).
— Newton, Thayer, Ashe, and Chandler, 2000: 328, 390 (key to Nearctic genera).
Othiides Thomson, 1859: 26 (genera included: Othius, Gyrohypnus). Type genus: Othius (see Newton and Thayer, 1992: 64).
Othii Thomson, 1859.
Othiina Thomson, 1859.
Atrecini Hatch, 1957: 172, 231 (key to genera of Pacific Northwest; genera included: Atrecus, Parothius). Type genus: Atrecus (see Newton and Thayer, 1992: 64).
Atrecus
[14 species; Nearctic and Palaearctic Regions]
Atrecus Jacquelin du Val, 1856: 31 (species included: pilicornis). Type species: Staphylinus pilicornis Paykull, fixed by original designation by monotypy.
— Schaum, 1859: 27 (synonym of Baptolinus).
— Gemminger and Harold, 1868: 606 (synonym of Baptolinus).
— Jacquelin du Val, 1868: 65 (catalog; European species).
— Fauvel, 1873b: 86 [= 1873a: 372] (synonym of Baptolinus).
— Mulsant and Rey, 1877a: 237 (synonym of Baptolinus).
— Ganglbauer, 1895: 472 (synonym of Baptolinus).
— Bernhauer and Schubert, 1914: 316 (synonym of Baptolinus).
— Tottenham, 1949: 370, 417 (type species: affinis Paykull = pilicornis Jacquelin du Val; notes; checklist of British species).
— Blackwelder, 1952: 69 (type species: pilicornis).
— Hatch, 1957: 232 (characters; key to species of Pacific Northwest).
— Smetana, 1958: 117 (characters; habitus of one species; type species: pilicornis; key to Czechoslovakian species).
— Moore, 1963a: 94 (characters).
— Székessy, 1963: 51 (characters).
— Lohse, 1964: 167 (synonym of Baptolinus).
— Pototskaia, 1967: 47 (larval characters).
— Tikhomirova, 1973a: 180 (checklist of species of USSR).
— Moore and Legner, 1975: 72 (catalog; North America north of Mexico).
— Shilov, 1975a: 375 [= 1975b: 97] (Atrecus; Komi region of Russia).
— Pototskaia, 1976: 171 (larval characters).
— Pope, 1977: 28 (list of British species).
— Moore and Legner, 1979: 55 (characters; notes).
— Muona, 1979: 16 (list of Scandinavian species).
— Smetana, 1982: 36 (characters; revision of and key to species of North America north of Mexico).
— Bordoni, 1982: 397 (synonym of Baptolinus).
— Shibata, 1983: 85 (checklist of species of Japan).
— Muona and Viramo, 1986: 14 (list of species of northeastern Finland).
— Lohse and Lucht, 1989: 159 (valid name).
— Angelini, 1991: 190 (list of species from high elevations of Calabria, Italy).
— Campbell and Davies, 1991: 123 (checklist of species; Alaska and Canada).
— Silfverberg, 1992: 19 (list of species of Fennoscandia, Denmark, and the Baltic States).
— Welch, 1993: 228 (cited as Atrechus; ovariole structure and development).
— Janák, 1993a: 47 (list of species of Czech Republic and Slovakia).
— Ciceroni and Zanetti, 1995: 26 (list of species of Italy).
— M. Hansen, 1996: 108 (list of Danish species).
— M. Hansen, Mahler, Palm, and Pedersen, 1996: 243 (additions to list of Danish species).
— Downie and Arnett, 1996: 396 (characters; key to species of Northeastern North America).
— Assing, 1998c: 349 (key to species of Palaearctic region).
— Newton, Thayer, Ashe, and Chandler, 2000: 390 (5 Nearctic species; notes; characters in key).
Baptolinus Kraatz, 1857d: 659 (species included: alternans, pilicornis). Type species: Staphylinus pilicornis Paykull, fixed by subsequent designation by Tottenham, 1945: 70.
— Redtenbacher, 1857: 982 (characters).
— G. Waterhouse, 1858: 26 (catalog of British species).
— Schaum, 1859: 27 (catalog; European species).
— LeConte, 1861a: 65 (characters).
— Gredler, 1863: 108 (list of species from Tirol).
— LeConte, 1863: 24 (list of North American species).
— Wencker and Silbermann, 1866: 30 (list of species; collecting notes; France).
— Gemminger and Harold, 1868: 606 (catalog).
— Sharp, 1871d: 12 (list of British species).
— Fauvel, 1873b: 86 [= 1873a: 372] (characters; key to species).
— Redtenbacher, 1874: 221 (characters; key to Austrian species).
— Fauvel, 1875a: XXIV [= 1875b: 226] (catalog).
— Mulsant and Rey, 1877a: 237 (characters; key to French species).
— Provancher, 1877: 251 (characters).
— Heyden, 1880: 75 (list of species of Siberian region).
— Duvivier, 1883: 159 (catalog).
— Fowler, 1888: 294 (notes).
— Seidlitz, 1891: 379 (key to some European species).
— Seidlitz, 1891a: 404 (key to some European species).
— Heyden, Reitter, and Weise, 1891: 105 (list of species of Europe and Caucasus).
— Heyden, 1893: 43 (list of species of Siberian region).
— Ganglbauer, 1895: 472 (characters; key to middle European species).
— Heyden, 1896: 33 (list of species of Siberian region).
— Casey, 1906: 424 (characters; key to species).
— Heyden, Reitter, and Weise, 1906: 162 (list of species of Europe and Caucasus).
— Reitter, 1908: 103, 107 (characters; key to species).
— Reitter, 1909: 132, 133 (characters; key to species).
— Petri, 1912: 70 (list of species of Siebenbürgen).
— Bernhauer and Schubert, 1914: 316 (world catalog; 9 species).
— Johansen, 1914: 428 (characters; key to Danish species).
— Leng, 1920: 106 (catalog of North American species).
— Winkler, 1925: 373 (catalog for Palaearctic region).
— Porta, 1926: 92 (characters; key to species of Italy).
— Portevin, 1929: 368 (key to French species).
— Roubal, 1930: 364 (catalog; Slovakia).
— Joy, 1932: 127 (key to British species).
— Scheerpeltz, 1933: 1320 (world catalog supplement).
— Tottenham, 1945: 70 (synonym of Atrecus; type species designation: Staphylinus pilicornis Paykull).
— Tottenham, 1949: 370 (synonym of Atrecus; type species: pilicornis).
— Blackwelder, 1952: 72 (type species: pilicornis).
— V. Hansen, 1952: 7, 21 (characters; key to Danish species).
— Coiffait, 1956a: 42 (key to French species).
— Smetana, 1958: 117 (synonym of Atrecus).
— Palm, 1963b: 53 (characters; key to species of Sweden).
— Lohse, 1964: 167 (characters; key to central European species).
— Scheerpeltz, 1968a: 63 (catalog of Austrian species).
— Kasule, 1970: 74 (larval characters).
— Coiffait, 1972: 358 (characters; larval characters; type species: pilicornis; key to species of western Palaearctic region; references).
— Pope, 1977: 28 (synonym of Atrecus).
— Uhlig, 1979: 252 (some species collected in Germany).
— Burakowski, Mroczkowski, and Stefańska, 1980: 22 (catalog; Poland).
— Smetana, 1982: 36 (synonym of Atrecus).
— Bordoni, 1982: 397 (characters; habitus; key to species of Italy).
— Tóth, 1989: 39 (characters; key to species of Hungary).
— Janák, 1993a: 47 (synonym of Atrecus).
affinis Paykull, 1789: 24 (Staphylinus; Type locality: Suecia).
— Erichson, 1839a: 422 (Othius; synonym of pilicornis).
— Erichson, 1839b: 297 (Othius; synonym of pilicornis).
— Jacquelin du Val, 1856: pl. 11, fig. 55 (Atrecus; [Note: Misidentification: Bernhauer and Schubert, 1914: 316, cited pilicornis Jacquelin du Val as a synonym of affinis; Jacquelin du Val did not describe the species as new, he attributed it to Paykull]; characters).
— Kraatz, 1857d: 661 (Baptolinus; synonym of alternans).
— Fauvel, 1873b: 87 [= 1873a: 373] (Baptolinus; characters; France; Scandinavia; Britain; Germany; Switzerland; Austria; Italy; Sicily; Sardinia; Caucasus).
— Fauvel, 1874: 338 [= 1874b: 44] (Baptolinus; Spain; Russia).
— Ganglbauer, 1895: 474 (Baptolinus; characters; Europe).
— Reitter, 1908: 108 (Baptolinus; characters; Europe; Caucasus).
— Reitter, 1909: 134 (Baptolinus; characters; Germany).
— Bernhauer and Schubert, 1914: 316 (Baptolinus; catalog).
— Johansen, 1914: 429 (Baptolinus; characters; Denmark).
— Portevin, 1929: 368 (Baptolinus; characters; France).
— Scheerpeltz, 1933: 1320 (Baptolinus; catalog).
— Paulian, 1941: 221 (Baptolinus; larval characters; France).
— V. Hansen, 1952: 22 (Baptolinus; characters; Denmark).
— Coiffait, 1956a: 43 (Baptolinus; characters in key).
— Smetana, 1958: 121 (Atrecus; characters; ecology; Czechoslovakia).
— Palm, 1963b: 54 (Baptolinus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 167 (Baptolinus; characters; central Europe).
— Smetana, 1964d: 73 (Atrecus; Czechoslovakia).
— Coiffait, 1965e: 120 (Othius; Turkey).
— Horion, 1965: 103 (Baptolinus; Caucasus; Siberia; Finland; Sweden; Norway; Denmark; Britain; widespread in central and southern Europe; Spain; Italy; Dalmatia; Croatia; Bosnia; Albania; Greece).
— Coiffait, 1966: 22 (Baptolinus; Turkey).
— Pototskaia, 1967: 49 (Atrecus; larval characters).
— Szujecki, 1968a: 726 (Atrecus; cited as afiinis; Poland).
— Bolov, 1969a: 328 (Atrecus; collecting notes; Russia).
— Coiffait, 1972: 363 (Baptolinus; characters; references; Europe; Scandinavia; Britain; Siberia; Turkey).
— Bordoni, 1973a: 192 (Baptolinus; Turkey).
— Bordoni, 1976: 236 (Baptolinus; Turkey).
— Pototskaia, 1976: 172 (Atrecus; larval characters).
— Legner and Moore, 1977: 176 (Atrecus; found under bark).
— Pope, 1977: 28 (Atrecus; Britain).
— Bordoni, 1982: 404 (Baptolinus; characters; Italy).
— Bordoni, 1986b: 386 (Baptolinus; Slovakia; Czechoslovakia; Croatia; Serbia; Austria; Macedonia; Bulgaria; Carniola).
— Segers, 1986: 41 (Baptolinus; checklist of species; Belgium).
— Lucht, 1987: 103 (Baptolinus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Tóth, 1989: 39 (Baptolinus; characters; Hungary).
— Ciceroni, 1993: 77 (Atrecus; Italy).
— Welch, 1993: 228 (Atrecus; ovariole number and ovary structure).
— Janák, 1993a: 47 (Atrecus; Czech Republic; Slovakia).
— Sparacio, 1995: 159 (Atrecus; characters; notes; Sicily).
— Ciceroni and Zanetti, 1995: 26 (Atrecus; Italy).
— Luff, 1996: 30 (Atrecus; collecting notes; Isle of Man).
— Assing, 1998c: 345, 350 (Atrecus; characters; discussion of variation; Georgia; Turkey; Greece; Bulgaria).
— Owen, 1999b: 61 (Atrecus; captured underground at roots of old trees; Britain).
— Fowles, Alexander, and Key, 1999: 129 (Atrecus; saproxylic species; status as endangered or threatened species; Britain).
— Lott, 1999: 72 (Atrecus; saproxylic species; habitat; endangered status; Britain).
— Owen, 2000: 251 (Atrecus; pitfall trap collections from underground at roots of trees; Britain).
▴ Distribution: Europe, Russia, Caucasus, Turkey.
alternans Gravenhorst, 1802: 48 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).
— Latreille, 1804: 333 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 107 (Staphylinus).
— Gyllenhal, 1810: 359 (Staphylinus; synonym of pilicornis Paykull).
— Curtis, 1829: 26 (Xantholinus; catalog; Britain).
— C. Sahlberg, 1830: 335 (Staphylinus; synonym of pilicornis Paykull).
— Mannerheim, 1830: 34 (Gyrohypnus; Finland).
— Mannerheim, 1831: 448 (Gyrohypnus; Finland).
— Griffith and Pidgeon, 1832: 292 (Xantholinus).
— Stephens, 1833: 254 (Othius; characters; England).
— Nordmann, 1837: 120 (Gyrohypnus; Germany; Sweden; Tauria).
— Erichson, 1839a: 422 (Othius; synonym of pilicornis).
— Heer, 1839: 248 (Othius; characters; Switzerland).
— Erichson, 1839b: 297 (Othius; synonym of pilicornis).
— Stephens, 1839: 402 (Othius; characters; England).
— Laporte, 1840: 180 (Gyrohypnus; characters; France).
— Fairmaire and Laboulbène, 1856: 499 (Othius; synonym of pilicornis).
— Kraatz, 1857d: 660 (Baptolinus; characters; Germany).
— Redtenbacher, 1857: 982 (Baptolinus; characters; Austria).
— Thomson, 1860: 187 (Gyrohypnus; characters; Scandinavia).
— Fauvel, 1865a: 298 [= 1865b: 48] (Baptolinus; habitat; France).
— Fauvel, 1867c: 114 [= 1868b: 213] (Baptolinus; synonym of affinis).
— Fauvel, 1873b: 87 [= 1873a: 373] (Baptolinus; synonym of affinis).
— Redtenbacher, 1874: 222 (Baptolinus; characters; Austria).
— Seidlitz, 1875: 281 (Baptolinus; characters; Baltic region).
— J. Sahlberg, 1876: 33 (Baptolinus; Finland).
— Mulsant and Rey, 1877a: 243 (Baptolinus; characters; France).
— Fowler, 1888: 294 (Baptolinus; characters; Britain).
— Ganglbauer, 1895: 474 (Baptolinus; synonym of affinis).
— Reitter, 1908: 108 (Baptolinus; synonym of affinis).
— Bernhauer and Schubert, 1914: 316 (Baptolinus; synonym of affinis).
— Scheerpeltz, 1933: 1321 (Baptolinus; synonym of affinis).
— Smetana, 1958: 121 (Atrecus; synonym of affinis).
— Coiffait, 1972: 363 (Baptolinus; synonym of affinis).
— Legner and Moore, 1977: 176 (Atrecus; found under bark).
— Pope, 1977: 28 (Atrecus; synonym of affinis).
— Bordoni, 1982: 404 (Baptolinus; synonym of affinis).
nigriceps Mannerheim, 1830: 34 (Gyrohypnus; Type locality: Suecia: Kinnekulle).
— Mannerheim, 1831: 448 (Gyrohypnus; Sweden).
— Nordmann, 1837: 120 (Gyrohypnus; Sweden).
— Erichson, 1839b: 297 (Othius; synonym of pilicornis).
— Kraatz, 1857d: 660 (Baptolinus; synonym of alternans).
— Mulsant and Rey, 1877a: 244 (Baptolinus; synonym of pilicornis).
— Ganglbauer, 1895: 474 (Baptolinus; synonym of affinis).
— Reitter, 1908: 108 (Baptolinus; synonym of affinis).
— Bernhauer and Schubert, 1914: 316 (Baptolinus; synonym of affinis).
— Smetana, 1958: 121 (Atrecus; synonym of affinis).
— Coiffait, 1972: 363 (Baptolinus; synonym of affinis).
— Bordoni, 1982: 404 (Baptolinus; synonym of affinis).
glabricornis Stephens, 1833: 254 (Othius; Type locality: near London; Crwmlyn sand-hills).
— Stephens, 1839: 402 (Othius; characters; England).
— Hardy, 1851: 37 (Othius; synonym of pilicornis).
— G. Waterhouse, 1858: 26 (Othius; synonym of alternans).
— Ganglbauer, 1895: 474 (Baptolinus; synonym of affinis).
— Reitter, 1908: 108 (Baptolinus; synonym of affinis).
— Bernhauer and Schubert, 1914: 316 (Baptolinus; synonym of affinis).
— Smetana, 1958: 121 (Atrecus; synonym of affinis).
— Coiffait, 1972: 363 (Baptolinus; synonym of affinis).
— Bordoni, 1982: 404 (Baptolinus; synonym of affinis).
frigidus Dufour, 1843: 35 (Xantholinus; Type locality: vallée d'Ossau).
— Fauvel, 1869: 490 (Baptolinus; synonym of pilicornis).
— Fauvel, 1873b: 87 [= 1873a: 373] (Baptolinus; synonym of affinis).
— Ganglbauer, 1895: 474 (Baptolinus; synonym of affinis).
— Reitter, 1908: 108 (Baptolinus; synonym of affinis).
— Bernhauer and Schubert, 1914: 316 (Baptolinus; synonym of affinis).
— Smetana, 1958: 121 (Atrecus; synonym of affinis).
— Coiffait, 1972: 363 (Baptolinus; synonym of affinis).
— Bordoni, 1982: 404 (Baptolinus; synonym of affinis).
dimidiatus Motschulsky, 1860: 565 (Othius; Type locality: Sibérie orientale).
— Hochhuth, 1862: 69 (Othius; characters; Siberia).
— Fauvel, 1873b: 87 [= 1873a: 373] (Baptolinus; synonym of affinis).
— Bernhauer and Schubert, 1914: 316 (Baptolinus; synonym of affinis).
— Smetana, 1958: 121 (Atrecus; synonym of affinis).
— Coiffait, 1972: 363 (Baptolinus; synonym of affinis).
— Bordoni, 1982: 404 (Baptolinus; synonym of affinis).
caucasicus Roubal, 1933: 77 (Baptolinus; subspecies of affinis; Type locality: Ca. occ.: Krasnaja Poljiana; Teberda).
— Coiffait, 1972: 362, 363 (Baptolinus; subspecies of affinis; characters; Caucasus).
— Coiffait, 1978d: 163 (Baptolinus; subspecies of affinis; Turkey).
— Assing, 1998c: 345 (Atrecus; discussion; synonym of affinis).
alternans Gravenhorst, 1802, see: affinis Paykull, 1789.
americanus Casey, 1906: 426 (Baptolinus; Type locality: Lake Superior; [Note: See lectotype designation by Smetana, 1982]).
— Bernhauer and Schubert, 1914: 316 (Baptolinus; catalog).
— Notman, 1920: 17 (Baptolinus; New York).
— Smetana, 1982: 44 (Atrecus; lectotype designation; characters; Nova Scotia; Ontario; Quebec; Maine; Massachusetts; Michigan; New Hampshire; New York; North Carolina; Pennsylvania; Vermont; West Virginia).
— Smetana, 1990c: 83 (Atrecus; Arkansas; Tennessee).
— Downie and Arnett, 1996: 395 (Atrecus; characters; USA).
▴ Distribution: Canada, USA.
ardeanus Ciceroni, 1990: 55 (Atrecus; Type locality: Italia: Massiccio del Pollino, Piana del Pollino, 1700 m).
— Ciceroni and Zanetti, 1995: 26 (Atrecus; Italy).
— Assing, 1998c: 350 (Atrecus; characters).
▴ Distribution: Italy.
brevicornis Smetana, 1967d: 199 (Atrecus; Type locality: Mongolei: Central aimak, 12 km SO v. Ulan-Baator, Nucht in Bogdo ul, 1500 m).
— Coiffait, 1972: 364 (Baptolinus; characters).
— Assing, 1998c: 350 (Atrecus; characters).
▴ Distribution: Mongolia.
casalei Bordoni, 1987: 568 (Baptolinus; Type locality: Grecia orientale: M.te Pangaeon, Akrovunion, Kavala, 1300 m).
— Assing, 1998c: 349 (Atrecus; characters).
— Assing and Wunderle, 1999a: 481 (Atrecus; characters; notes; Greece).
▴ Distribution: Greece.
caucasicus Roubal, 1933, see: affinis Paykull, 1789.
dimidiatus Motschulsky, 1860, see: affinis Paykull, 1789.
fraternus Casey, 1906, see: punctiventris Fall, 1901.
frigidus Dufour, 1843, see: affinis Paykull, 1789.
fulvicollis Fairmaire and Germain, 1861, see: Cheilocolpus.
glabricornis Stephens, 1833, see: affinis Paykull, 1789.
longiceps Fauvel, 1872: 374 [= 1873b: 88] (Baptolinus; Type locality: Nancy; Savoie, Rhonnes; Grande Chartreuse).
— Mulsant and Rey, 1877a: 247 (Baptolinus; characters; France).
— Ganglbauer, 1895: 474 (Baptolinus; characters; France; Switzerland; Germany; Austria; Tuscany; North America; [Note: According to Smetana, 1982: 37, 47, longiceps does not occur in North America, most specimens so identified are americanus]).
— Reitter, 1908: 107 (Baptolinus; characters; central Europe).
— Reitter, 1909: 133 (Baptolinus; characters; Germany).
— Bernhauer and Schubert, 1914: 316 (Baptolinus; catalog).
— Portevin, 1929: 368 (Baptolinus; characters; France).
— Scheerpeltz, 1933: 1321 (Baptolinus; catalog).
— V. Hansen, 1952: 22 (Baptolinus; characters; Denmark).
— Coiffait, 1956a: 43 (Baptolinus; characters in key).
— Smetana, 1958: 120 (Atrecus; characters; ecology; Czechoslovakia).
— Palm, 1963b: 54 (Baptolinus; characters; Sweden; Denmark; Norway; Finland).
— Szujecki, 1963a: 33 (Atrecus; Poland).
— Lohse, 1964: 167 (Baptolinus; characters; central Europe).
— Horion, 1965: 104 (Baptolinus; Finland; Sweden; Norway; Denmark; France; Switzerland; Italy; Slovenia; Dalmatia; Bosnia; Slovakia; Czech Republic; Poland; Austria; Germany).
— Smetana, 1966: 330 (Atrecus; notes; Czechoslovakia).
— Coiffait, 1972: 364 (Baptolinus; characters; references; Europe; Scandinavia; Italy; Siberia; Britain).
— Legner and Moore, 1977: 176 (Atrecus; found under bark).
— Bordoni, 1982: 405 (Baptolinus; characters; Italy).
— Lucht, 1987: 103 (Baptolinus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Denmark; Sweden).
— Bordoni, 1986b: 387 (Baptolinus; Slovakia; Slovenia; Carniola).
— Tóth, 1989: 40 (Baptolinus; characters; Hungary).
— Janák, 1993a: 47 (Atrecus; Czech Republic; Slovakia).
— Ciceroni, 1993: 77 (Atrecus; Italy).
— Ciceroni and Zanetti, 1995: 26 (Atrecus; Italy).
— Assing, 1998c: 350 (Atrecus; characters).
▴ Distribution: Europe.
macrocephalus Nordmann, 1837: 124 (Gyrohypnus; Type locality: insula Sitka).
— Erichson, 1839b: 297 (Othius; characters; America borealis occidentalis, ad littora ins. Sitchae).
— Mannerheim, 1843b: 227 (Othius; characters; Sitka Island).
— Mannerheim, 1852: 312 (Othius; characters; notes; Sitka).
— Provancher, 1877: 252 (Baptolinus; cited as melanocephalus; characters; Quebec).
— Casey, 1906: 426 (Baptolinus; characters; New Hampshire; British Columbia; Queen Charlotte Island; Prince of Wales Island).
— Bernhauer and Schubert, 1914: 316 (Baptolinus; catalog).
— Van Dyke, 1924: 17 (Baptolinus; Alaska).
— Fall, 1926: 149 (Baptolinus; Alaska).
— Hatch, 1957: 232 (Atrecus; characters; British Columbia; Washington; Oregon; Idaho).
— Smetana, 1982: 38 (Atrecus; characters; Alberta; British Columbia; New Brunswick; Newfoundland; Northwest Territories; Nova Scotia; Ontario; Quebec; Alaska; California; Idaho; Maine; Montana; New Hampshire; New York; Oregon; Vermont; Washington).
— Dearborn and Donahue, 1994: 14 (Atrecus; Maine).
— Downie and Arnett, 1996: 396 (Atrecus; characters; USA).
▴ Distribution: USA, Canada.
newtoni Smetana, 1982: 53 (Atrecus; Type locality: Washington: Pierce Co., Mt. Rainier N. P., 4.7 mi. W Longmire, 2200 feet. Also cited from Oregon).
nigriceps Mannerheim, 1830, see: affinis Paykull, 1789.
parvioculatus Assing, 1998c: 346 (Atrecus; Type locality: Turquie: S. Vit, Ayder, 1400 m, Vallée de la Firtina, Rize).
pilicornis Paykull, 1790: 135 (Staphylinus; Type locality: Not cited but presumably Suecia).
— Schrank, 1798: 642 (Staphylinus; characters; notes).
— Paykull, 1800: 379 (Staphylinus; characters; Sweden).
— Gyllenhal, 1810: 359 (Staphylinus; characters; Sweden).
— Zetterstedt, 1828: 81 (Xantholinus; characters; Lapland).
— Curtis, 1829: 26 (Xantholinus; catalog; Britain).
— C. Sahlberg, 1830: 335 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 34 (Gyrohypnus; Finland; Sweden).
— Mannerheim, 1831: 448 (Gyrohypnus; Finland; Sweden).
— Stephens, 1833: 254 (Othius; characters; England).
— Nordmann, 1837: 119 (Gyrohypnus; Europe).
— Zetterstedt, 1838: 66 (Xantholinus; characters; habitat; Lapland).
— Erichson, 1839a: 422 (Othius; characters; Germany).
— Heer, 1839: 248 (Othius; characters; Switzerland).
— Erichson, 1839b: 296 (Othius; characters; Sweden; Germany).
— Stephens, 1839: 401 (Othius; characters; England).
— Märkel, 1845: 235 (Othius; with Formica fuliginosa).
— Jacobsen, 1849: 675 (Othius; characters; notes; Denmark).
— Hochhuth, 1849: 100 (Othius; characters; Lenkoran).
— Redtenbacher, 1849: 690 (Othius; characters; Austria).
— Hardy, 1851: 37 (Othius; notes; England).
— Fairmaire and Laboulbène, 1856: 499 (Othius; characters; France).
— Kraatz, 1857d: 661 (Baptolinus; characters; Germany).
— Redtenbacher, 1857: 179 (Othius; characters; Austria).
— Redtenbacher, 1857: 982 (Baptolinus; characters; Austria).
— Thomson, 1859: 27 (type species of Gyrohypnus).
— Thomson, 1860: 187 (Gyrohypnus; characters; Scandinavia).
— Fauvel, 1865a: 298 [= 1865b: 48] (Baptolinus; habitat; France).
— Fauvel, 1873b: 86 [= 1873a: 372] (Baptolinus; characters; France; Lapland; Scandinavia; Russia; Germany; Switzerland; Austria; Italy).
— Redtenbacher, 1874: 222 (Baptolinus; characters; Austria).
— Seidlitz, 1875: 279 (Baptolinus; characters; Baltic region).
— J. Sahlberg, 1876: 33 (Baptolinus; Finland).
— Mulsant and Rey, 1877a: 241 (Baptolinus; characters; France).
— Sharp, 1889: 249 (Baptolinus; Japan).
— Ganglbauer, 1895: 473 (Baptolinus; characters; north and middle Europe).
— Reitter, 1908: 107 (Baptolinus; characters; northern and central Europe).
— Reitter, 1909: 133 (Baptolinus; characters; Germany).
— Bernhauer and Schubert, 1914: 316 (Baptolinus; catalog).
— Johansen, 1914: 429 (Baptolinus; characters; Denmark).
— Šulc, 1922 (Baptolinus; osmeterium; anatomy; histology; physiology).
— Bernhauer, 1926g: 2 (Baptolinus; Kamchatka region).
— Portevin, 1929: 368 (Baptolinus; characters; France).
— Scheerpeltz, 1933: 1321 (Baptolinus; catalog).
— V. Hansen, 1952: 23 (Baptolinus; characters).
— Coiffait, 1956a: 42 (Baptolinus; characters in key).
— Smetana, 1958: 119 (Atrecus; characters; ecology; Czechoslovakia).
— Palm, 1963b: 54 (Baptolinus; characters; Sweden; Norway; Finland).
— Nakane, 1963a: 90 (Atrecus; characters; habitus photograph; Japan).
— Lohse, 1964: 167 (Baptolinus; characters; central Europe).
— Horion, 1965: 106 (Baptolinus; Russia; Finland; Sweden; Norway; Ukraine; Poland; Czechoslovakia; Hungary; Romania; Italy; Switzerland; France; Dalmatia; Bosnia Hercegovina; Croatia; Slovenia; Spain ?; Austria; Germany).
— Pototskaia, 1967: 49 (Atrecus; larval characters).
— Coiffait, 1972: 362 (Baptolinus; characters; references; Europe; Siberia; Caucasus).
— Smetana 1975c: 168 (Baptolinus; Mongolia).
— Pototskaia, 1976: 172 (Atrecus; larval characters).
— Legner and Moore, 1977: 176 (Atrecus; found under bark).
— Bordoni, 1982: 402 (Baptolinus; characters; Italy).
— Bordoni, 1986b: 386 (Baptolinus; Slovakia).
— Lucht, 1987: 103 (Baptolinus; checklist; Germany; Poland; Czechoslovakia; Austria; France; Sweden).
— Tóth, 1989: 41 (Baptolinus; characters).
— Lohse and Lucht, 1989: 159 (Atrecus; characters).
— J. Li, 1993: 33 (Atrecus; China).
— Janák, 1993a: 47 (Atrecus; Czech Republic; Slovakia).
— Muona and Viramo, 1995: 9 (Atrecus; collected from dung of Ursus arctos; Finland).
— Ciceroni and Zanetti, 1995: 26 (Atrecus; Italy).
— Raitschev, 1997a: 100 (Baptolinus; notes; Bulgaria).
— Assing, 1998c: 350 (Atrecus; characters).
— Ryabukhin, 1999: 56 (Atrecus; catalog; habitat; distribution; northeastern Russia).
— Naomi, Kuranishi, Saito, and Maruyama, 2000: 109 (Atrecus; Kamchatka).
— Olberg and Andersen, 2000: 223 (Atrecus; attracted to trunk-window traps baited with Fomes fomentarius and Phellinus spp.; Norway).
▴ Distribution: Europe, Russia, Azerbaijan, Mongolia, China, Japan.
suturalis Motschulsky, 1845b: 359 (Othius; Type locality: Kamtschatka; [Note: See lectotype designation by Assing, 1998c]).
— Bernhauer and Schubert, 1914: 316 (Baptolinus; catalog).
— Assing, 1998c: 344 (Atrecus; lectotype designation; characters; Russia; synonym of pilicornis).
— Ryabukhin, 1999: 56 (Atrecus; cited as valid species; catalog; habitat; distribution; northeastern Russia).
punctiventris Fall, 1901: 221 (Baptolinus; Type locality: Lake Tahoe, California; [Note: See lectotype designation by Smetana, 1982]. Also cited from Utah and Vancouver Island [Note: According to Smetana, 1982: 48, the Canadian specimen is probably a misidentification of quadripennis).
— Casey, 1906: 427 (Baptolinus; characters; California).
— Bernhauer and Schubert, 1914: 316 (Baptolinus; catalog).
— Hatch, 1957: 233 (Atrecus; characters; British Columbia; Washington; Oregon; [Note: According to Smetana, 1982: 51, Hatch's Canadian record may be a misidentification of quadripennis]).
— Smetana, 1982: 47 (Atrecus; lectotype designation: Lake Tahoe, Cal; characters; California; Nevada; Oregon; Washington).
▴ Distribution: USA.
fraternus Casey, 1906: 428 (Baptolinus; Type locality: California, Siskyou Co.; [Note: See lectotype designation by Smetana, 1982]).
— Bernhauer and Schubert, 1914: 316 (Baptolinus; catalog).
— Smetana, 1982: 47 (Atrecus; lectotype designation; synonym of punctiventris).
quadripennis Casey, 1906: 427 (Baptolinus; Type locality: Idaho, Coeur d'Alene; [Note: See lectotype designation by Smetana, 1982]).
— Bernhauer and Schubert, 1914: 316 (Baptolinus; catalog).
— Hatch, 1957: 233 (Atrecus; characters; British Columbia; Washington; Oregon; Idaho).
— Smetana, 1982: 51 (Atrecus; lectotype designation; characters; Alberta; British Columbia; Idaho; Oregon; Utah; Washington; Wyoming).
— Smetana, 1988a: 526 (Atrecus; collecting notes; Oregon).
▴ Distribution: Canada, USA.
schuelkei Assing, 1998c: 348 (Atrecus; Type locality: China: W-Sichuan, Daxue Shan, Hailuogou Glacier Park Camp 1, 2100 m, 29.36.00N, 102.03.35E).
suturalis Motschulsky, 1845, see: pilicornis Paykull, 1790.
yunnanus Assing, 2000c: 75 (Atrecus; Type locality: China: Yunnan Prov., Heishui, 35 km N. Lijiang).
Othius
[105 species (4 doubtful); Palaearctic, Oriental, and Australian Regions]
Othius Stephens, 1829a: 23 (species included: fulgidus, pilicornis, ustulatus). Type species: Staphylinus punctulatus Goeze, fixed by action of the Commission in Opinion 1250 (ICZN, 1983).
— Stephens, 1829: 284 (catalog; Britain).
— Stephens, 1832: 200 (characters).
— Stephens, 1833: 253 (characters; habitat).
— Westwood, 1838a: 16 (characters; type species).
— Erichson, 1839a: 420 (characters).
— Heer, 1839: 247 (characters).
— Erichson, 1839b: 294 (characters).
— Stephens, 1839: 385 (characters).
— Redtenbacher, 1849: 690 (characters).
— Schaum, 1852: 25 (list of species; Europe).
— Lacordaire, 1854: 63 (characters; notes; list of species).
— Wollaston, 1854: 574 (characters; notes).
— Fairmaire and Laboulbène, 1856: 498 (characters).
— Jacquelin du Val, 1856: 30 (characters).
— Gistel, 1856: 387 (list of species).
— Kraatz, 1857d: 653 (characters).
— Redtenbacher, 1857: 179 (characters; key to Austrian species).
— Thomson, 1858: 30 (characters).
— G. Waterhouse, 1858: 26 (catalog of British species).
— Thomson, 1859: 26 (characters; type species: fulvipennis).
— Schaum, 1859: 27 (catalog; European species).
— Thomson, 1860: 185 (characters).
— LeConte, 1861a: 65 (characters).
— Gredler, 1863: 108 (list of species from Tirol).
— LeConte, 1863: 24 (list of North American species).
— Fauvel, 1866a: 347 [= 1866b: 98] (characters).
— Wencker and Silbermann, 1866: 31 (list of species; collecting notes; France).
— Gemminger and Harold, 1868: 607 (catalog).
— Jacquelin du Val, 1868: 65 (catalog; European species).
— Sharp, 1871d: 12 (list of British species).
— Fauvel, 1873b: 81 [= 1873a: 367] (Cafius; characters; France).
— Redtenbacher, 1874: 222 (characters; key to Austrian species).
— Fauvel, 1875a: XXIV [= 1875b: 226] (catalog).
— Mulsant and Rey, 1877a: 219 (characters; key to French species).
— Broun, 1880: 101 (characters).
— Heyden, 1880: 76 (list of species of Siberian region).
— Duvivier, 1883: 159 (catalog).
— Fowler, 1888: 294 (notes; key to British species).
— Seidlitz, 1891: 379 (key to some European species).
— Seidlitz, 1891a: 404 (key to some European species).
— Heyden, Reitter, and Weise, 1891: 105 (list of species of Europe and Caucasus).
— Heyden, 1893: 43 (list of species of Siberian region).
— Ganglbauer, 1895: 468 (larval and adult characters; key to middle European species).
— Fauvel, 1897d: 289 (catalog of species of Barbary; Madeira; Canaries).
— Casey, 1906: 422 (characters).
— Heyden, Reitter, and Weise, 1906: 162 (list of species of Europe and Caucasus).
— Reitter, 1908: 103, 104 (characters; key to species).
— Reitter, 1909: 132 (characters; key to species).
— Petri, 1912: 70 (list of species of Siebenbürgen).
— Bernhauer and Schubert, 1914: 317 (world catalog; 28 species).
— Johansen, 1914: 426 (characters; key to Danish species).
— Winkler, 1925: 373 (catalog for Palaearctic region).
— Porta, 1926: 91 (characters; key to species of Italy).
— Portevin, 1929: 367 (key to French species).
— Roubal, 1930: 365 (catalog; Slovakia).
— Cameron, 1932: 41 (characters; key to species of British India).
— Joy, 1932: 127 (key to British species).
— Scheerpeltz, 1933: 1321 (world catalog supplement).
— Normand, 1935: 369 (list of species of Tunisia).
— Blackwelder, 1944: 145 (checklist of species from Mexico, Central and South America, and the West Indies).
— Tottenham, 1949: 371 (synonym of Gyrohypnus; type species: punctulatus Goeze).
— Blackwelder, 1952: 279 (type species: fulgidus; discussion).
— V. Hansen, 1952: 23 (characters; key to Danish species).
— Coiffait, 1956a: 37, 38 (characters; key to species; cited as synonym of Gyrohypnus but believes rules of priority should be suspended for this case).
— Kocher, 1958: 141 (checklist of species; Morocco).
— Smetana, 1958: 122 (characters; habitus of one species; type species: laeviusculus; key to Czechoslovakian species).
— Smetana, 1959a: 209 (list of species collected in Albania).
— Ferreira, 1962a: 16 (list of Portuguese species).
— Székessy, 1963: 51 (characters).
— Palm, 1963b: 54 (characters; key to species of Sweden).
— Lohse, 1964: 167 (characters; key to central European species).
— Pototskaia, 1967: 46 (larval characters).
— Coiffait and Saiz, 1968: 345 (list of Chilean species).
— Scheerpeltz, 1968a: 63 (catalog of Austrian species).
— Kasule, 1970: 74, 76 (larval characters).
— Coiffait, 1972: 327 (characters; larval characters; type species: punctulatus; key to species of western Palaearctic region; references).
— Bordoni, 1973j: 691 (list of species from islands near Sicily).
— Tikhomirova, 1973a: 180 (checklist of species of USSR).
— Palm, 1976: 211 (key to species of Canary Islands).
— Pope, 1977: 28 (list of British species).
— Topp, 1978: 311 (larval characters).
— Palm, 1979: 269 (key to species of Madeira; notes).
— Muona, 1979: 16 (list of Scandinavian species).
— Uhlig, 1979: 252 (some species collected in Germany).
— M. Dvoř, 1979: 115 (some species collected in Slovakia).
— Smetana, 1979: 50 (requested that International Commission on Zoological Nomenclature set aside all type species designations and designate Staphylinus punctulatus Goeze as type species and place Othius on the Official list of Generic Names in Zoology).
— Uhlig, Vogel, and Sieber, 1980: 246 (some species collected in Germany).
— Burakowski, Mroczkowski, and Stefańska, 1980: 25 (catalog; Poland).
— Uhlig and Vogler, 1981: 109 (list of some species collected in Germany; habitat and collecting notes).
— Bordoni, 1982: 378 (characters; habitus; key to species of Italy).
— ICZN, 1983: 85 (Othius placed on Official List of Generic Names in Zoology and Staphylinus punctulatus is fixed as the type species).
— Shibata, 1983: 81 (checklist of species of Japan).
— Shibata, 1986a: 123 (supplemental list of species from Taiwan).
— Muona and Viramo, 1986: 14 (list of species of northeastern Finland).
— Segers, 1986: 41 (checklist of species; Belgium).
— Tóth, 1989: 34 (characters; key to species of Hungary).
— Lohse and Lucht, 1989: 159 (supplement to key to species of Lohse, 1964).
— Angelini, 1991: 192 (list of species from high elevations of Calabria, Italy).
— Silfverberg, 1992: 18 (list of species of Fennoscandia, Denmark, and the Baltic States).
— Siitonen, 1993: 230 (list of Finnish species).
— Welch, 1993: 228 (ovariole structure and development).
— Janák, 1993a: 47 (list of species of Czech Republic and Slovakia).
— Hernández, Outerelo, and Gamarra, 1994: 190 (checklist of species; Canary Islands).
— Assing and Wunderle, 1995a (revision of and key to species of Madeira; notes).
— Ciceroni and Zanetti, 1995: 26 (list of species of Italy).
— M. Hansen, 1996: 108 (list of Danish species).
— Assing, 1997a (review of and key to species of Canary Islands).
— Assing, 1997 (revision of and key to species of western Palaearctic; characters for species groups).
— Assing, 1998 (revision of and key to species of Himalayan region).
— Assing, 1998a (new records for, colonization by, and phylogeny of species of Madeira Canary Islands).
— Assing, 1999 (revision of and key to eastern Palaearctic species east of Himalaya).
— Assing and Schülke, 1999: 8 (key to species of central Europe).
Othiellus Casey, 1906: 422, 423 (species included: laeviusculus, melanocephalus). Type species: Othius laeviusculus Stephens, fixed by subsequent designation by Tottenham, 1939d: 339.
— Bernhauer and Schubert, 1914: 317 (synonym of Othius).
— Cameron, 1932: 41 (synonym of Othius).
— Tottenham, 1939d: 229 (type species designation: laeviusculus).
— Tottenham, 1949: 371 (synonym of Gyrohypnus; type species: laeviusculus Stephens).
— Blackwelder, 1952: 279 (type species).
— Smetana, 1958: 122 (synonym of Othius).
— Coiffait, 1972: 327 (synonym of Othius).
— Tikhomirova, 1973a: 180 (subgenus of Othius; checklist of species of USSR).
Othiogeiton Scheerpeltz, 1976: 31 (species included: nepalensis). Type species: Othiogeiton nepalensis Scheerpeltz, fixed by original designation and monotypy.
acifer Assing, 1998: 314 (Othius; Type locality: Nepal: Taplejung Distr., upper Simbua Khola Valley, near Tseram, 3250–3350 m).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).
▴ Distribution: Nepal.
acutus Assing, 1999: 72 (Othius; Type locality: Taiwan: Taitung, Kuanshan Yakou).
adustus Broun, 1880, see: Maorothius.
aequabilis Assing, 1999: 68 (Othius; Type locality: Taiwan: Nantou Hsien, Yushan N. P., 1.8 km W Pai-Yun Hut, 3375 m).
albanicus Reitter, 1918, see: lapidicola Märkel and Kiesenwetter, 1848.
algarvensis Coiffait, 1964, see: subuliformis Stephens, 1833.
amamianus Ito, 1993a: 144 (Othius; Type locality: Japan: Kagoshima Pref.: Shimmura, Amami-Oshima Is.).
angustus Stephens, 1833: 255 (Othius; Type locality: Bristol; Edinburgh; [Note: See lectotype designation by Assing, 1997]).
— Hardy, 1851: 36 (Othius; synonym of melanocephalus).
— G. Waterhouse, 1858: 26 (Othius; synonym of melanocephalus).
— Reitter, 1908: 106 (Othius; synonym of melanocephalus).
— Bernhauer and Schubert, 1914: 318 (Othius; synonym of melanocephalus).
— Smetana, 1958: 126 (Othius; cited as valid species; characters; ecology; Czechoslovakia).
— Lohse, 1964: 168 (Othius; synonym of melanocephalus).
— Coiffait, 1972: 348 (Othius; characters; references; Europe; Scandinavia; Britain; Caucasus).
— Pope, 1977: 28 (Othius; Britain).
— Bordoni, 1982: 394 (Othius; characters; Italy).
— Frank, 1982: 31 (Othius; cited as angustatus; list of parasites).
— Tóth, 1989: 36 (Othius; cited as angustatus; characters).
— Nowosad, 1990: 101 (Othius; collected from mole nest; notes; Poland).
— Janák, 1993a: 47 (Othius; Czech Republic; Slovakia).
— Sadler and Dugmore, 1995: 144 (Othius; habitat notes; Iceland).
— Ciceroni and Zanetti, 1995: 26 (Othius; Italy).
— Assing, 1997: 24 (Othius; lectotype designation: [Note: Locality not cited]; characters; notes; Norway; Sweden; Denmark; Portugal; Spain; France; Germany; Switzerland; Austria; Poland; Czech Republic; Slovakia; Romania).
— Assing and Schülke, 1999: 11 (Othius; nominate subspecies; characters).
— Assing, 1999c: 666 (Othius; subspecies angustus; Norway; Finland; Denmark; Russian Federation; Poland; Spain; Andorra; France; Switzerland; Swiss-Austria territory; Austria; Italy; Czech Republic; Slovakia; Polish, Czech or Slovakian territory; Bosnia Herzegovina).
▴ Distribution: Iceland, Europe, Russia, Georgia, Turkey.
melanocephalus Gravenhorst, 1806: 107 (Staphylinus; [preoccupied]; Type locality: Brunsvigae; [Note: See lectotype designation by Assing, 1997]).
— Gyllenhal, 1810: 360 (Staphylinus; characters; Sweden).
— Curtis, 1829: 27 (Gyrohypnus; catalog; Britain).
— C. Sahlberg, 1830: 336 (Staphylinus; characters; Finland).
— Griffith and Pidgeon, 1832: 292 (Xantholinus).
— Nordmann, 1837: 120 (Gyrohypnus; Sweden; Germany; France).
— Erichson, 1839a: 421 (Othius; characters; Germany).
— Heer, 1839: 248 (Othius; characters; Switzerland).
— Erichson, 1839b: 295 (Othius; characters; Europe).
— Stephens, 1839: 402 (Othius; characters; England).
— Grimm, 1845: 134 (Othius; with Formica fuliginosa; Germany).
— Jacobsen, 1849: 674 (Othius; characters; notes; Denmark).
— Redtenbacher, 1849: 690 (Othius; characters; Austria).
— Hardy, 1851: 36 (Othius; notes; England).
— Fairmaire and Laboulbène, 1856: 498 (Othius; characters; France).
— Kraatz, 1857d: 656 (Othius; characters; Germany).
— Redtenbacher, 1857: 179 (Othius; characters; Austria).
— Thomson, 1860: 185 (Othius; characters; Scandinavia).
— Hochhuth, 1862: 69 (Othius; Russia).
— Fauvel, 1865a: 298 [= 1865b: 48] (Othius; habitat; France).
— Fauvel, 1873b: 85 [= 1873a: 371] (Othius; characters; France; Europe).
— Redtenbacher, 1874: 223 (Othius; characters; Austria).
— Seidlitz, 1875: 279 (Othius; characters; Baltic region).
— J. Sahlberg, 1876: 31 (Othius; Finland).
— Mulsant and Rey, 1877a: 231 (Othius; characters; France).
— Fowler, 1888: 295 (Othius; characters; Britain).
— Ganglbauer, 1895: 470 (Othius; characters; north and middle Europe).
— Poppius, 1905: 10 (Othius; Orkney Islands; Shetland Islands).
— Casey, 1906: 423 (Othiellus).
— Reitter, 1908: 106 (Othius; characters; northern and central Europe).
— Reitter, 1909: 133 (Othius; characters; Germany).
— Sharp and Muir, 1912: 499 (Othius; characters of male genitalia).
— Bernhauer and Schubert, 1914: 318 (Othius; catalog).
— Johansen, 1914: 427 (Othius; characters; Denmark).
— Šulc, 1922 (Othius; osmeterium; anatomy; histology; physiology).
— Portevin, 1929: 368 (Othius; characters; France).
— Scheerpeltz, 1933: 1322 (Othius; catalog).
— West, 1937: 41 (Othius; Faroe Islands).
— V. Hansen, 1952: 25 (Othius; characters; Denmark).
— Coiffait, 1956a: 40 (Othius; characters in key).
— Smetana, 1958: 126 (Othius; synonym of angustus).
— Larsson and Gígja, 1959: 81 (Othius; biological notes; Iceland).
— Palm, 1963b: 55 (Othius; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 168 (Othius; characters; central Europe).
— Horion, 1965: 110 (Othius; widespread in northern Europe, in central Europe south to Spain and Italy).
— Pototskaia, 1967: 48 (Othius; larval characters).
— Kasule, 1970: 76 (Othius; larval characters).
— Coiffait, 1972: 348 (Othius; synonym of angustus).
— Pope, 1977: 28 (Othius; synonym of angustus).
— Lucht, 1987: 103 (Othius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Lohse and Lucht, 1989: 160 (Othius; cited as valid name; characters).
— Assing, 1993: 33 (Othius; bionomics; Germany).
— Ciceroni and Zanetti, 1995: 26 (Othius; synonym of angustus).
— Gerend and Braunert, 1997: 197 (Othius; cited as valid species; notes; Luxembourg).
— Assing, 1997: 24 (Othius; lectotype designation; synonym of angustus).
— Uhlig, 1998: 136 (Othius; synonym of angustus).
sexpunctatus Haliday, 1841: 187 (Othius; Type locality: Holywood).
— Kraatz, 1857d: 656 (Othius; synonym of melanocephalus).
— Ganglbauer, 1895: 470 (Othius; synonym of melanocephalus).
— Reitter, 1908: 106 (Othius; synonym of melanocephalus).
— Bernhauer and Schubert, 1914: 318 (Othius; synonym of melanocephalus).
— Smetana, 1958: 127 (Othius; synonym of angustus).
— Coiffait, 1972: 348 (Othius; synonym of angustus).
stenocephalus Eppelsheim, 1881: 506 (Othius; Type locality: Helenendorf; [Note: See lectotype designation by Assing, 1997]).
— Reitter, 1908: 106 (Othius; characters; Caucasus).
— Bernhauer and Schubert, 1914: 317 (Othius; variety of laeviusculus).
— Scheerpeltz, 1933: 1322 (Othius; variety of laeviusculus).
— Coiffait, 1972: 348 (Othius; characters; references; Balkans; Caucasus).
— Coiffait, 1978d: 163 (Othius; Turkey).
— Assing, 1997: 30 (Othius; subspecies of angustus; lectotype designation; characters; notes; Turkey; Caucasus, Russia, Georgia).
— Assing, 1999c: 667 (Othius; subspecies of angustus; Turkey; Georgia; Caucasus).
— Solodovnikov, 2000a: 325 (Othius; subspecies of angustus; notes; Turkey).
sipeki Roubal, 1919: 15 (Othius; aberration of melanocephalus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from [Bohemia]: Pribram).
— Scheerpeltz, 1933: 1322 (Othius; aberration of melanocephalus).
— Smetana, 1958: 127 (Othius; aberration of angustus).
angustus Broun, 1893, see: Maorothius.
antarcticus Fairmaire, 1885, see: Antarctothius.
apicalis Cameron, 1926b: 347 (Othius; Type locality: Chakrata District, Chulli Khud, 6500 ft.; [Note: See lectotype designation by Assing, 1998]. Also cited from Mussorie District; Arni Gad).
— Cameron, 1932: 43 (Othius; characters; India).
— Scheerpeltz, 1933: 1321 (Othius; catalog).
— Paulian, 1941: 219 (Othius; larval characters; India).
— Assing, 1998: 295 (Othius; characters; lectotype designation: Chakrata District, Chulli Khud, 6500 ft.; Nepal; India; Sikkim).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).
▴ Distribution: India, Nepal.
arieiroensis Palm, 1979: 272 (Othius; Type locality: Madeira: between Pico Arieiro and Poiso, alt. about 1600 m).
— Assing and Wunderle, 1995a: 60 (Othius; characters; bionomics; Madeira).
— Assing, 1998a: 215 (Othius; Madeira).
▴ Distribution: Madeira.
arisanus Shibata, 1973, see: punctatus Bernhauer, 1922.
atavisticus Rüschkamp, 1932, see: subuliformis Stephens, 1833.
atavus Assing, 1999: 32 (Othius; Type locality: China: Yunnan prov., Gaoligongshan Mts., 90 km W of Baoshan).
ater Stephens, 1833, see: Gyrohypnus.
baculifer Assing and Wunderle, 1995a: 61 (Othius; Type locality: Madeira: Bica da Cana, 1550 m).
badius Zheng, 1995: 341, 346 (Othius; Type locality: Sichuan, Mt. Emei, alt. 1752–2070 m).
bhutanensis Assing, 1999c: 681 (Othius; Type locality: Bhutan: Paro District, Chiley-La, 3000–3500 m).
bordonii Assing, 1999: 13 (Othius; Type locality: Kouy-Tchéou, Rég. de Pin-Fa).
bovinus Coquerel, 1860, see: laeviusculus Stephens, 1833.
brachypterus Wollaston, 1864: 580 (Othius; Type locality: Gomera).
— Wollaston, 1865: 499 (Othius; habitat; Canary Islands).
— Fauvel, 1902b: 99 (Othius; Canary Islands).
— Bernhauer and Schubert, 1914: 317 (Othius; catalog).
— Coiffait, 1954: 165 (Othius; Canary Islands).
— Coiffait, 1956a: 41 (Othius; characters in key).
— Coiffait, 1972: 354 (Othius; characters; references; Canary Islands).
— Palm, 1976: 212 (Othius; characters; Canary Islands).
— Assing, 1997a: 104, 112 (Othius; characters; notes; Canary Islands).
— Assing, 1998a: 215 (Othius; Canary Islands).
— Assing, 1999a: 346 (Othius; Canary Islands).
— Assing, 1999c: 668 (Othius; Canary Islands).
▴ Distribution: Canary Islands.
brevicornis Wollaston, 1857, see: jansoni Wollaston, 1854.
brevipennis Kraatz, 1857d: 657 (Othius; Type locality: Oesterreich; [Note: See lectotype designation by Assing, 1997]).
— Redtenbacher, 1857: 982 (Othius; characters; Austria).
— Fauvel, 1873b: 85 [= 1873a: 371] (Othius; characters; Austria).
— Redtenbacher, 1874: 223 (Othius; characters; Austria).
— Ganglbauer, 1895: 472 (Othius; characters; eastern Alps).
— Reitter, 1908: 107 (Othius; characters; Austrian Alps).
— Bernhauer and Schubert, 1914: 317 (Othius; catalog).
— Scheerpeltz, 1933: 1321 (Othius; catalog).
— Walles, 1938: 67, 70 (Othius; Poland).
— Scheerpeltz and Höfler, 1948: 156 (Othius; fungus inhabitant).
— Smetana, 1958: 130 (Othius; characters; ecology; Czechoslovakia).
— Lohse, 1964: 169 (Othius; characters; central Europe).
— Horion, 1965: 114 (Othius; Germany; Austria; Italy; Slovenia; Poland; Slovakia).
— Coiffait, 1972: 352 (Othius; characters; references; Europe).
— Bordoni, 1982: 388 (Othius; characters; Italy).
— Frank, 1982: 31 (Othius; list of parasites).
— Lucht, 1987: 103 (Othius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland).
— Tóth, 1989: 39 (Othius; characters; Hungary).
— Janák, 1993a: 47 (Othius; Slovakia).
— Ciceroni and Zanetti, 1995: 27 (Othius; Italy).
— Assing, 1997: 93 (Othius; lectotype designation; characters; notes; Germany; Austria; Czech Republic; Slovakia; Hungary; Slovenia; Croatia; Bosnia-Herzegovina).
— Assing and Solodovnikov, 1998: 305 (Othius; characters; Poland).
— Assing and Schülke, 1999: 12 (Othius; characters).
— Assing, 1999c: 677 (Othius; notes; Austria; Slovakia; Slovenia; Italy; Bosnia Herzegovina).
▴ Distribution: Poland, Germany, Austria, Czech Republic, Slovakia, Hungary, Slovenia, Croatia, Bosnia Hercegovina.
dilutus Motschulsky, 1858a: 210 (Othius; Type locality: Styrie; [Note: See lectotype designation by Assing, 1997]).
— Fauvel, 1873b: 83 [= 1873a: 369] (Othius; synonym of myrmecophilus).
— Mulsant and Rey, 1877a: 228 (Othius; synonym of mymecophilus).
— Ganglbauer, 1895: 472 (Othius; synonym of brevipennis).
— Reitter, 1908: 107 (Othius; synonym of brevipennis).
— Bernhauer and Schubert, 1914: 317 (Othius; synonym of brevipennis).
— Smetana, 1958: 130 (Othius; synonym of brevipennis).
— Coiffait, 1972: 352 (Othius; synonym of brevipennis).
— Bordoni, 1982: 388 (Othius; synonym of brevipennis).
— Assing, 1997: 93 (Othius; lectotype designation; synonym of brevipennis).
brookesi Cameron, 1952, see: Maorothius.
cachemiricus Coiffait, 1982a: 252 (Othius; Type locality: Cachemire: Daksum, 2400–2700 m).
californicus Mannerheim, 1843, see: Parothius.
casarum Gistel, 1857: 79 (Othius; [Note: Nomen dubium]; Type locality: Alpes Tyrolis, Salisburgi, et Helvetiae).
castaneipennis Gistel, 1857: 69 (Othius; [Note: Nomen dubium]; Type locality: Tyrolis alpes).
chongqingensis Zheng, 1995, see: latus Sharp, 1874.
chrysurus Reitter, 1891b: 195 (Othius; Type locality: Taschkent; [Note: See lectotype designation by Assing, 1997]).
— Reitter, 1908: 105 (Othius; characters; Tashkent).
— Bernhauer and Schubert, 1914: 317 (Othius; catalog).
— Scheerpeltz, 1933: 1321 (Othius; catalog).
— Coiffait, 1972: 341 (Othius; characters; Turkestan).
— Assing, 1997: 104 (Othius; lectotype designation; characters; notes; Tajikistan; Uzbekistan; Kazakhstan).
— Assing, 1999c: 679 (Othius; correct measurements for TiL and TaL cited in Assing, 1997, by factor of 2.02; collecting notes; Tajikistan; Uzbekistan; Kirghizia).
▴ Distribution: Kazakhstan, Uzbekistan, Tajikistan, Kyrgyzstan.
clavifer Assing, 1998: 319 (Othius; Type locality: Nepal: Sankhua Sabha Distr., Thudam).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).
▴ Distribution: Nepal.
coiffaiti Lohse, 1963, see: philonthoides Wollaston, 1864.
collapsus Assing, 1999: 78 (Othius; Type locality: Taiwan: Nantou Hsien, near Tsifeng, 2300 m).
coniceps Assing, 1999: 83 (Othius; Type locality: Taiwan: Nantou Hsien, near Hohuanshan, 3100 m).
conifer Assing, 1998: 315 (Othius; Type locality: Nepal: Taplejung Distr., pass Deorali W Yamputhin, 3400 m).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).
▴ Distribution: Nepal.
contumax Assing, 1999: 81 (Othius; Type locality: Taiwan: Nantou Hsien, Nenkaoshan, Tenchi Hut, 2890 m).
corniger Assing, 1998: 321 (Othius; Type locality: Nepal: Taplejung Distr., upper Simbua Khola Valley, near Tseram, 3250–3350 m).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).
▴ Distribution: Nepal.
corpulentus Coiffait, 1972: 340, 343 (Othius; subspecies of transsilvanicus; Type locality: Mt. Sarco, Hu. merid.; [Note: See lectotype designation by Assing, 1997]. Originally cited from sud des Karpathes).
— Assing, 1997: 87, 129 (Othius; lectotype designation: Mt. Sarco, Hu. merid.; valid species; characters; notes; Romania).
— Assing, 1999c: 676 (Othius; Romania; Serbia ?).
▴ Distribution: Romania.
crassus Motschulsky, 1858a: 210 (Othius; Type locality: Alpes de la Carniole, Doga-Niva; [Note: See lectotype designation by Assing, 1997]).
— Fauvel, 1873b: 83 [= 1873a: 369] (Othius; synonym of lapidicola).
— Ganglbauer, 1895: 472 (Othius; characters; eastern Alps; north Carpathians).
— Reitter, 1908: 107 (Othius; characters; Carpathians; Caucasus).
— Bernhauer and Schubert, 1914: 317 (Othius; catalog).
— Scheerpeltz, 1933: 1322 (Othius; catalog).
— Smetana, 1958: 130 (Othius; characters; ecology; Czechoslovakia).
— Lohse, 1964: 169 (Othius; characters; central Europe).
— Horion, 1965: 113 (Othius; Austria; Italy; Poland; Slovakia; Romania).
— Coiffait, 1972: 351 (Othius; characters; references; Alps; Carpathians; France; Caucasus).
— Bordoni, 1982: 386 (Othius; characters; Italy).
— Lucht, 1987: 103 (Othius; checklist; Germany; Poland; Czechoslovakia; Austria).
— Tóth, 1989: 38 (Othius; characters; Hungary).
— Janák, 1993a: 47 (Othius; Slovakia).
— Ciceroni and Zanetti, 1995: 27 (Othius; Italy).
— Assing, 1997: 73 (Othius; lectotype designation; characters; notes; Austria; Slovenia; probably restricted range and citations from France, the Caucasus and other regions and countries are probably misidentifications).
— Assing and Schülke, 1999: 14 (Othius; characters).
— Assing, 1999c: 674 (Othius; notes; Austria; Slovenia).
▴ Distribution: Austria, Slovenia.
pallidus Brancsik, 1874: 135 (Othius; Type locality: Styria; [Note: See lectotype designation by Assing, 1997]. Originally cited from Bachergebirge bei Marburg in Steiermark).
— Ganglbauer, 1895: 472 (Othius; synonym of crassus).
— Reitter, 1908: 107 (Othius; synonym of pallidus).
— Bernhauer and Schubert, 1914: 317 (Othius; synonym of crassus).
— Smetana, 1958: 130 (Othius; synonym of crassus).
— Coiffait, 1972: 352 (Othius; synonym of crassus).
— Bordoni, 1982: 386 (Othius; synonym of crassus).
— Assing, 1997: 73 (Othius; lectotype designation: Styria; synonym of crassus).
deceptivus Cuvier, 1833, see: punctulatus Goeze, 1777.
deharvengi Coiffait, 1982: 53 (Othius; Type locality: Nepal: versant sud-est du Col de Goropani, ouest de Pokhara, 2150 m).
— Assing, 1998: 329 (Othius; characters; Nepal).
— Assing, 1999c: 669 (Othius; Nepal).
▴ Distribution: Nepal.
discrepans Assing, 1999: 45 (Othius; Type locality: China: Sichuan, Ziling Mt., 1600–2400 m).
dilutus Motschulsky, 1858, see: brevipennis Kraatz, 1857.
dimidiatus Motschulsky, 1860, see: Atrecus.
donisthorpei Chitty, 1903 [Note: Nomen nudum], see: punctulatus Goeze, 1777.
elongatus Coiffait, 1956, see: lapidicola Märkel and Kiesenwetter, 1848.
excisus Assing, 1999: 46 (Othius; Type locality: China: S. Sichuan, S. Xichang, Mt. Luoji, 2300–2500 m).
extraordinarius Assing, 1998: 303 (Othius; Type locality: N-Indien: Himach. Pr., Umg. Kufri, ca. 2600 m).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).
▴ Distribution: India.
fastigatus Assing and Solodovnikov, 1998: 301 (Othius; Type locality: Georgia: Caucasus Swanetien).
fibulifer Assing, 1999: 53 (Othius; Type locality: China: W Sichuan, Daxue Shan, E Theto-La [recte: Tsheto-La] Pass, W. Kangding, 3500 m, 30.00.42 N, 101.51.47 E).
flavocaudatus Cameron, 1926b: 346 (Othius; Type locality: Chakrata District, Sijla Gad, 5000 feet; [Note: See lectotype designation by Assing, 1998]).
— Cameron, 1932: 42 (Othius; characters; India).
— Scheerpeltz, 1933: 1322 (Othius; catalog).
— Assing, 1998: 301 (Othius; characters; lectotype designation: Chakrata District, Sijla Gad, 5000 feet; Nepal; India).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).
▴ Distribution: Nepal, India.
fulminans Gravenhorst, 1802, see: punctulatus Goeze, 1777.
fulvipennis Fabricius, 1793, see: punctulatus Goeze, 1777.
fulvipes Motschulsky, 1858, see: Gabrius.
fulvopterus Geoffroy, 1785, see: punctulatus Goeze, 1777.
fuscicornis Heer, 1841, see: laeviusculus Stephens, 1833.
gansuensis Assing, 1999, see: latus Sharp, 1874.
glabricornis Stephens, 1833, see: Atrecus.
goui Zheng, 1995: 342, 346 (Othius; Type locality: Sichuan: Wolong, 1830 m).
grandis Hochhuth, 1849: 99 (Othius; Type locality: Cauc. Gotsch.; [Note: See lectotype designation by Gusarov, 1992a]. Originally cited from Lenkoran; Transkaukasien; Taurien).
— Hochhuth, 1862: 69 (Othius; Russia).
— Reitter, 1891b: 195 (Othius; characters; Caucasus).
— Reitter, 1908: 105 (Othius; characters; Caucasus; Lenkoran; Crimea).
— Bernhauer and Schubert, 1914: 317 (Othius; catalog).
— Scheerpeltz, 1933: 1322 (Othius; catalog).
— Coiffait, 1965e: 120 (Othius; Turkey).
— Pototskaia, 1967: 48 (Othius; larval characters).
— Bolov, 1969a: 328 (Othius; collecting notes; Russia).
— Coiffait, 1972: 342 (Othius; characters; references; Russia; Turkey).
— Petrenko, 1980: 81 (Othius; Caucasus).
— Gusarov, 1992a: 784 [= 1993a: 73] (Othius; lectotype designation: Cauc. Gotsch.).
— Assing, 1997: 55 (Othius; characters; notes; Turkey; Georgia; Russia).
— Solodovnikov, 1998a: 16 (Othius; notes; Russia, Krasnodar territory).
— Assing, 1999c: 672 (Othius; correct measurements for TiL and TaL cited in Assing, 1997, by factor of 2.02; Caucasus).
— Solodovnikov, 2000a: 324 (Othius; notes; Turkey).
▴ Distribution: Turkey, Azerbaijan, Georgia, Russia, Ukraine.
hamatus Assing, 1999: 70 (Othius; Type locality: Taiwan: Kaohsiung Hsien, Peinantashan trail, 2250 m).
hebes Assing and Solodovnikov, 1998: 299 (Othius; Type locality: Armenia: Caucasus, Armen. Geb. Also cited from Georgia and Azerbaijan).
ibericus Coiffait, 1956, see: punctulatus Goeze, 1777.
infestus Assing, 1998: 311 (Othius; Type locality: Nepal: Taplejung Distr., upper Simbua Khola Valley, near Tseram, 3250–3350 m).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).
▴ Distribution: Nepal.
intermedius Korge, 1962: 151 (Othius; Type locality: Kanarische Inseln: Teneriffa: Montana Cagancho, ca. 1600 m).
— Coiffait, 1972: 357 (Othius; characters; Canary Islands).
— Palm, 1976: 214 (Othius; characters; Canary Islands).
— Assing, 1997a: 107, 113 (Othius; characters; notes; Canary Islands).
— Assing, 1998a: 216 (Othius; Canary Islands).
▴ Distribution: Canary Islands.
jadwigae Roubal, 1911: 5 (Othius; Type locality: Kr. Poljana; [Note: See lectotype designation by Assing, 1997]).
— Bernhauer and Schubert, 1914: 317 (Othius; cited as jadvigae; catalog).
— Coiffait, 1972: 356 (Othius; characters; Caucasus).
— Assing, 1997: 61 (Othius; lectotype designation; characters; notes; Caucasus).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1997, by factor of 1.54).
— Solodovnikov, 2000a: 328 (Othius; notes; Georgia).
▴ Distribution: Russia, Caucasus.
jaegeri Assing, 1998: 334 (Othius; Type locality: Nepal: Telbrung Danda, 3700 m, SE Annapurna mts.).
jansoni Wollaston, 1854: 576 (Othius; Type locality: Madeira: Ribeiro de Santa Luzia; Pico d'Arribentao above Funchal; [Note: See lectotype designation by Assing and Wunderle, 1995a]).
— Wollaston, 1857: 187 (Othius; characters; habitat; Madeira Islands).
— Wollaston, 1865: 499 (Othius; habitat; notes; Madeira Islands).
— Fauvel, 1902b: 99 (Othius; Madeira).
— Bernhauer and Schubert, 1914: 317 (Othius; catalog).
— Smetana, 1963b: 36 (Othius; Madeira).
— Coiffait, 1972: 354 (Othius; characters; references; Madeira).
— Palm, 1979: 271 (Othius; characters; collecting notes; Madeira).
— Assing and Wunderle, 1995a: 58 (Othius; lectotype designation: [Note: Locality not cited]; bionomics; characters; Madeira).
— Assing, 1998a: 215 (Othius; Madeira).
— Assing, 1999c: 672 (Othius; Madeira).
▴ Distribution: Madeira.
vestitus Wollaston, 1857: 186 (Othius; Type locality: Madeira: Lombo dos Pecegueiros; Cruzinhas at more than 4000 ft.).
— Fauvel, 1902b: 99 (Othius; synonym of jansoni).
— Bernhauer and Schubert, 1914: 317 (Othius; synonym of jansoni).
— Coiffait, 1972: 354 (Othius; synonym of jansoni).
— Palm, 1979: 272 (Othius; valid species; collecting notes; characters; Madeira).
— Assing and Wunderle, 1995a: 58 (Othius; synonym of jansoni).
brevicornis Wollaston, 1857: 187 (Othius; Type locality: Madeira: Dezerta Grande).
— Wollaston, 1865: 499 (Othius; notes; Madeira Islands).
— Fauvel, 1902b: 99 (Othius; synonym of jansoni).
— Bernhauer and Schubert, 1914: 317 (Othius; synonym of jansoni).
— Coiffait, 1972: 354 (Othius; synonym of jansoni).
— Palm, 1979: 271 (Othius; synonym of jansoni).
— Assing and Wunderle, 1995a: 58 (Othius; synonym of jansoni).
javanus Cameron, 1937, see: Metolinus.
jumlaensis Assing, 1999c: 669 (Othius; Type locality: Nepal: Jumla Distr., Ghurchi-Lagna Pass, 3500 m).
kashmiricus Cameron, 1943: 127 (Othius; Type locality: Kashmir: Gulmarg).
kastamonuensis Bordon, 1976, see: lapidicola Märkel and Kiesenwetter, 1848.
korgei Coiffait, 1965e: 123 (Othius; Type locality: Turquie d'Asie: Akkus près Ünye).
— Coiffait, 1972: 351 (Othius; characters; Turkey).
— Bordoni, 1976: 234 (Othius; Turkey).
— Assing, 1997: 100 (Othius; characters; notes; Turkey).
— Assing and Solodovnikov, 1998: 305 (Othius; characters).
— Assing, 1999c: 678 (Othius; Anatolia).
▴ Distribution: Turkey.
kusuii Ito, 1993, see: medius Sharp, 1874.
laeviusculus Stephens, 1833: 255 (Othius; Type locality: London; Suffolk; Crwmlyn sand-hills; [Note: See lectotype designation by Assing, 1997]).
— Stephens, 1839: 402 (Othius; characters; England).
— Fauvel, 1873b: 84 [= 1873a: 370] (Othius; characters; France; Britain; Switzerland; Austria; Algeria; Morocco; Cyprus; Caucasus; Iran).
— Fauvel, 1878: 116 [= 1878a: 36] (Othius; Barbary).
— Fauvel, 1886: 48 [= 1886a: 40] (Othius; North Africa).
— Fowler, 1888: 295 (Othius; characters; Britain).
— Ganglbauer, 1895: 470 (Othius; characters; middle Europe; Mediterranean region).
— Fauvel, 1902b: 99 (Othius; Barbary).
— Casey, 1906: 423 (Othiellus).
— Sainte-Claire Deville, 1906: 94 (Othius; Corsica).
— Reitter, 1908: 106 (Othius; characters; central and southern Europe).
— Reitter, 1909: 133 (Othius; characters; Germany).
— Bernhauer and Schubert, 1914: 317 (Othius; catalog).
— Portevin, 1929: 368 (Othius; characters; France).
— Scheerpeltz, 1933: 1322 (Othius; catalog).
— Coiffait, 1956a: 40 (Othius; characters in key; Greece; Cyprus; Turkey; France).
— Scheerpeltz, 1956c: 1099 (Othius; Cyprus).
— Smetana, 1958: 125 (Othius; characters; ecology; Czechoslovakia).
— Scheerpeltz, 1961e: 136 (Othius; Rhodes).
— Scheerpeltz, 1963a: 419 (Othius; Greece).
— Lohse, 1964: 168 (Othius; characters; central Europe).
— Focarile, 1964: 58 (Othius; Italy).
— Scheerpeltz, 1965c: 505 (Othius; Sicily).
— Horion, 1965: 108 (Othius; Morocco; Tunisia; Turkey; Iberian Peninsula; Balearics; Corsica; Corsica; Sardinia; Sicily; Italy; Greece; Crete; Dalmatia; Bosnia Hercegovina; Albania; Cyprus; France; Switzerland; Belgium; Britain; Romania; Hungary; Czech Republic; Poland; Austria; Germany).
— Coiffait, 1966: 22 (Othius; Turkey).
— Coiffait, 1970e: 62 (Othius; Cyprus).
— Coiffait, 1972: 344 (Othius; characters; references; Europe; Britain; North Africa).
— Pope, 1977: 28 (Othius; Britain).
— Bordoni, 1982: 391 (Othius; characters; Italy).
— Lucht, 1987: 103 (Othius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).
— Tóth, 1989: 35 (Othius; characters; Hungary).
— Ciceroni, 1993: 76 (Othius; Italy).
— Scheerpeltz, 1964a: 299 (Othius; Crete).
— Welch, 1993: 228 (Othius; ovariole number and ovary structure).
— Janák, 1993a: 47 (Othius; Czech Republic; Slovakia).
— Sparacio, 1995: 159 (Othius; characters; notes; Sicily).
— Ciceroni and Zanetti, 1995: 27 (Othius; Italy).
— Vorst, 1996 (Othius; notes on biology and distribution; Netherlands).
— Assing, 1997: 34 (Othius; lectotype designation: [Note: Locality not cited]; characters; notes; Great Britain; Morocco; Algeria; Tunisia; Portugal; Spain; Balearic Islands; France; Germany; Switzerland; Austria; Italy; Poland; Czech Republic; Slovakia; Hungary; Slovenia; Croatia; Bosnia-Herzegovina; Bulgaria; Macedonia; Greece; Cyprus; Turkey; Syria; Caucasus).
— Assing and Schülke, 1999: 9 (Othius; characters).
— Assing, 1999c: 667 (Othius; Morocco; Algeria; Spain; France; Germany; Italy; Austria; Czech Republic; Bosnia Herzegovina; Yugoslavia; Macedonia; Greece; Cyprus; Turkey; Iran).
— Solodovnikov, 2000a: 325 (Othius; notes; Russia).
— Assing and Wunderle, 2001: 36 (Othius; Cyprus).
▴ Distribution: Tunisia, Algeria, Morocco, Europe, Russia, Azerbaijan, Turkey, Syria, Iran.
punctipennis Lacordaire, 1835: 409 (Staphylinus; Type locality: Paris).
— Erichson, 1839b: 296 (Othius; characters; Austria; Switzerland; Italy; France; England).
— Heer, 1841: 581 (Othius; characters; Switzerland).
— Hochhuth, 1849: 100 (Othius; characters; Lenkoran).
— Redtenbacher, 1849: 690 (Othius; characters; Austria).
— Rosenhauer, 1856: 70 (Othius; Malaga).
— Fairmaire and Laboulbène, 1856: 498 (Othius; characters; France).
— Kraatz, 1857d: 655 (Othius; characters; Germany).
— Redtenbacher, 1857: 179 (Othius; characters; Austria).
— Kraatz, 1858a: 65 (Othius; Greece).
— G. Waterhouse, 1858: 26 (Othius; synonym of laeviusculus).
— Hochhuth, 1862: 69 (Othius; Russia).
— Fauvel, 1869: 490 (Othius; synonym of laeviusculus).
— Fauvel, 1873b: 84 [= 1873a: 370] (Othius; synonym of laeviusculus).
— Redtenbacher, 1874: 222 (Othius; characters; Austria).
— Mulsant and Rey, 1877a: 234 (Othius; characters; France).
— Reitter, 1908: 106 (Othius; synonym of laeviusculus).
— Bernhauer and Schubert, 1914: 317 (Othius; synonym of laeviusculus).
— Scheerpeltz, 1933: 1322 (Othius; synonym of laeviusculus).
— Smetana, 1958: 126 (Othius; synonym of laeviusculus).
— Coiffait, 1972: 344 (Othius; synonym of laeviusculus).
— Bordoni, 1982: 391 (Othius; synonym of laeviusculus).
— Assing, 1997: 34 (Othius; synonym of laeviusculus).
fuscicornis Heer, 1841: 581 (Othius; Type locality: Aigle).
— Kraatz, 1857d: 655 (Othius; synonym of punctipennis).
— Reitter, 1908: 106 (Othius; synonym of laeviusculus).
— Bernhauer and Schubert, 1914: 317 (Othius; synonym of laeviusculus).
— Smetana, 1958: 126 (Othius; synonym of laeviusculus).
— Coiffait, 1972: 344 (Othius; synonym of laeviusculus).
— Bordoni, 1982: 391 (Othius; synonym of laeviusculus).
— Assing, 1997: 34 (Othius; synonym of laeviusculus).
bovinus Coquerel, 1860: 154 (Quedius; Type locality: Mers-el-Kébir).
— Fauvel, 1869: 486 (Othius; synonym of laeviusculus).
— Fauvel, 1873b: 84 [= 1873a: 370] (Othius; synonym of laeviusculus).
— Fauvel, 1902b: 99 (Othius; synonym of laeviusculus).
— Bernhauer and Schubert, 1914: 317 (Othius; synonym of laeviusculus).
— Scheerpeltz, 1933: 1322 (Othius; synonym of laeviusculus).
— Coiffait, 1972: 344 (Othius; synonym of laeviusculus).
— Bordoni, 1982: 391 (Othius; synonym of laeviusculus).
— Assing, 1997: 34 (Othius; synonym of laeviusculus).
lapidicola Märkel and Kiesenwetter, 1848a: 321 (Othius; Type locality: Kärnthner Alpen; [Note: See lectotype designation by Assing, 1997]).
— Fairmaire and Laboulbène, 1856: 498 (Othius; characters; France).
— Kraatz, 1857d: 657 (Othius; characters; Germany).
— Fauvel, 1865a: 299 [= 1865b: 49] (Othius; habitat; France).
— Fauvel, 1873b: 83 [= 1873a: 369] (Othius; characters; France; Germany; Switzerland; Austria; Italy; Scandinavia).
— Fauvel, 1874: 338 [= 1874b: 44] (Othius; Lapland; Spain).
— Redtenbacher, 1874: 222 (Othius; characters; Austria).
— J. Sahlberg, 1876: 31 (Othius; characters; Finland).
— Mulsant and Rey, 1877a: 225 (Othius; characters; France).
— Ganglbauer, 1895: 471 (Othius; characters; north and middle Europe).
— Reitter, 1909: 133 (Othius; characters; Germany).
— Bernhauer and Schubert, 1914: 318 (Othius; catalog).
— Portevin, 1929: 368 (Othius; characters; France).
— Scheerpeltz, 1933: 1322 (Othius; catalog).
— V. Hansen, 1952: 26 (Othius; characters).
— Smetana, 1958: 127 (Othius; characters; ecology; Czechoslovakia).
— Palm, 1963b: 56 (Othius; characters; Sweden; Norway; Finland).
— Lohse, 1964: 168 (Othius; characters; central Europe).
— Horion, 1965: 111 (Othius; Canaries; Turkey; Russia; Finland; Sweden; Norway; France; Switzerland; Czech Republic; Slovakia; Poland; Bosnia Hercegovina; Montenegro; Serbia; Macedonia; Bulgaria; Greece; Italy; Spain; Austria; Germany).
— Coiffait, 1966: 22 (Othius; Turkey).
— Coiffait, 1972: 347 (Othius; characters; references; Europe; Caucasus; Spain; Italy; Balkans; Turkey).
— Coiffait, 1978d: 163 (Othius; Turkey).
— Bordoni, 1982: 392 (Othius; characters; Italy).
— Frank, 1982: 31 (Othius; list of parasites).
— Lucht, 1987: 103 (Othius; checklist; Germany; Poland; Czechoslovakia; Austria; Sweden).
— Tóth, 1989: 36 (Othius; characters).
— Nowosad, 1990: 101 (Othius; collected from mole nest; notes; Poland).
— Ciceroni, 1993: 76 (Othius; Italy).
— Wittwer, 1993: 249 (Othius; Switzerland).
— Janák, 1993a: 47 (Othius; Czech Republic; Slovakia).
— Hodge and Jones, 1995: 37 (Othius; characters; Britain).
— Ciceroni and Zanetti, 1995: 27 (Othius; Italy).
— Assing, 1997: 8 (Othius; lectotype designation; characters; notes; Norway; Sweden; Finland; Russia; France; Germany; Switzerland; Austria; Italy; Czech Republic; Slovenia; Hungary; Romania; Croatia; Bosnia-Herzegovina; Yugoslavia; Macedonia; Albania; Bulgaria; Greece; Turkey; Armenia).
— Uhlig, 1998: 136 (Othius; note).
— Assing and Schülke, 1999: 10 (Othius; characters).
— Assing, 1999c: 663 (Othius; notes on elevation of collections; Finland; Russian Federation; France; Germany; Switzerland; Austria; Italy; Czech Republic; Slovakia; Romania; Croatia; Bosnia Herzegovina; Bulgaria; Greece; Turkey; Georgia; Caucasus).
— Solodovnikov, 2000a: 325 (Othius; notes; Turkey).
▴ Distribution: Europe, Georgia, Armenia, Turkey.
suturalis Motschulsky, 1858a: 211 (Othius; [preoccupied]; Type locality: Alpes de la Carniole, Koschutna).
— Fauvel, 1873b: 83 [= 1873a: 369] (Othius; synonym of lapidicola).
— Ganglbauer, 1895: 471 (Othius; synonym of lapidicola).
— Bernhauer and Schubert, 1914: 318 (Othius; synonym of lapidicola).
— Smetana, 1958: 127 (Othius; synonym of lapidicola).
— Coiffait, 1972: 347 (Othius; synonym of lapidicola).
— Bordoni, 1982: 392 (Othius; synonym of lapidocola).
— Assing, 1997: 8 (Othius; synonym of lapidicola).
longicornis Thomson, 1871: 369 (Othius; Type locality: Norrland).
— Fauvel, 1873b: 83 [= 1873a: 369] (Othius; synonym of lapidicola).
— J. Sahlberg, 1876: 31 (Othius; synonym of lapidicola).
— Ganglbauer, 1895: 471 (Othius; synonym of lapidicola).
— Bernhauer and Schubert, 1914: 318 (Othius; synonym of lapidicola).
— Smetana, 1958: 127 (Othius; synonym of lapidicola).
— Coiffait, 1972: 347 (Othius; synonym of lapidicola).
— Bordoni, 1982: 392 (Othius; synonym of lapidicola).
— Assing, 1997: 8 (Othius; synonym of lapidicola).
albanicus Reitter, 1918: 154 (Othius; Type locality: Albanien: Miloti; [Note: See lectotype designation by Assing, 1997]).
— Scheerpeltz, 1933: 1321 (Othius; catalog).
— Coiffait, 1968b: 93 (Othius; notes; Greece).
— Coiffait, 1972: 346 (Othius; characters; Albania; Greece; Turkey; Bulgaria).
— Assing, 1997: 8, 128 (Othius; lectotype designation; synonym of lapidicola).
elongatus Coiffait, 1956b: 139 (Othius; Type locality: Turquie d'Asie, Villayet d'Afyon: Dinar. Also cited from Slovakia).
— Lohse, 1963: 177 (Othius; synonym of lapidicola).
— Lohse, 1964: 168 (Othius; synonym of lapidicola).
— Coiffait, 1965e: 120 (Othius; cited as valid species; Turkey).
— Coiffait, 1972: 343 (Othius; characters; Turkey).
— Assing, 1997: 8 (Othius; synonym of lapidicola).
rhodicus Coiffait, 1976b: 85 (Othius; Type locality: Rhodes: Monte Profitis Ilias, 800 m).
kastamonuensis Bordoni, 1976: 234 (Othius; Type locality: Turchia: vil. Kastamonu, Ilgaz dag gecidi, 1800–2300 m).
latus Sharp, 1874: 51 (Othius; Type locality: Japan; [Note: See lectotype designation by Assing, 1999]. Originally cited from Copper Temple, Nagasaki; northern China).
— Bernhauer and Schubert, 1914: 318 (Othius; catalog).
— Scheerpeltz, 1933: 1322 (Othius; catalog).
— Ito, 1993a: 141 (Othius; characters; Japan).
— J. Li, 1993: 32 (Othius; China).
— Assing, 1999: 20 (Othius; lectotype designation: Japan; characters; Russia; Japan; China).
▴ Distribution: Japan, China, Russia.
stoetzneri Bernhauer, 1931: 1 (Othius; Type locality: Szetschwan: Kwanshien; [Note: See lectotype designation by Assing, 1999]).
— Scheerpeltz, 1933: 1323 (Othius; catalog).
— Assing, 1999: 20 (Othius; lectotype designation; synonym of latus latus).
ozakii Ito, 1993a: 143 (Othius; subspecies of latus; Type locality: Japan: Aomori Pref.: Jyogakura, Mt. Hakkoda).
chongqingensis Zheng, 1995: 343, 346 (Othius; Type locality: Sichuan: Mt. Jin-yun, Chongqing City, 800–919 m).
gansuensis Assing, 1999: 27 (Othius; subspecies of latus; Type locality: China: Gansu,
28 km S. Lanzhou, 2200 m).
laufferi Bernhauer, 1909, see: piceus Scriba, 1870.
linkei Bernhauer, 1915, see: subuliformis Stephens, 1833.
loeffleri Scheerpeltz, 1976a: 124 (Othius; Type locality: Nepal: Beding, Rolwaling, 3940 m).
— Coiffait, 1981d: 323 (Othius; Nepal).
— Assing, 1998: 336 (Othius; not examined; notes).
— Assing, 1999c: 680 (Othius; characters of holotype; holotype male, not female; notes; Nepal).
▴ Distribution: Nepal.
nepalensis Scheerpeltz, 1976: 32 (Othiogeiton; Type locality: Nepal: Unterhalb Almsied-lung Taboche, ca. 4500 m [Note: This is the only locality cited for the species, but the allotype is the holotype since no other specimens were mentioned; see Assing, 1998: 325, 326]).
— Assing, 1998: 325 (Othius; characters; Nepal).
— Assing, 1999c: 680, 684 (Othius; synonym of loeffleri; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).
longicornis Thomson, 1871, see: lapidicola Märkel and Kiesenwetter, 1848.
longicuneatus Assing, 1998: 323 (Othius; Type locality: Gorkha Dist., Darondi Khola bis oberhalb Barpak, 3600–3450 [sic] m).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).
▴ Distribution: Nepal.
lubricus Assing, 1999: 41 (Othius; Type locality: China: Yunnan, Kunming, Western Hills, 1800 m, 25°04′ N, 102°41′ E).
maculativentris Zheng, 1995: 344, 346 (Othius; Type locality: Sichuan: Wolong, 1920–2400 m).
— Assing, 1999: 38 (Othius; characters; China).
— Assing, 1999c: 683 (Othius; Sichuan).
▴ Distribution: China.
medius Sharp, 1874: 50 (Othius; Type locality: Japan; [Note: See lectotype designation by Assing, 1999]. Originally cited from Hiogo; Nagasaki).
— Bernhauer and Schubert, 1914: 318 (Othius; catalog).
— Nakane, 1963a: 90 (Othiellus; characters; habitus photograph; Japan).
— Yuh, Paik, Kwon, and Lee, 1985: 239 (Othius; Korea).
— Hasegawa and Kanie, 1992: 20 (Othius; seashore collection; Japan).
— Ito, 1993a: 146 (Othius; characters; Japan).
— J. Li, 1993: 33 (Othius; China).
— Watanabe, 1995d: 256 (Othius; Okushiri-tô Island).
— Watanabe, 1996a: 12 (Othius; mine collection; Japan).
— Assing, 1999: 7 (Othius; lectotype designation: Japan; characters; discussions; Japan; China; South Korea).
— Assing, 1999c: 682 (Othius; characters; Japan).
▴ Distribution: Japan, Korea, China.
kusuii Ito, 1993a: 147 (Othius; subspecies of medius; Type locality: Japan: Hokkaido: Tamaura, Okushiri Is.).
yakushimanus Ito, 1993a: 148 (Othius; subspecies of medius; Type locality: Japan: Kagoshima Pref.: Miyanoura, Yakushima Is.).
melanocephalus Gravenhorst, 1806, see: angustus Stephens, 1833.
microphallus Assing, 1998a: 217 (Othius; Type locality: Teneriffa, Pico del Ingles).
microphtalmus Coiffait, 1954: 169 (Othius; Type locality: Bosque del Cedro, île de Gomera, Canaries).
— Coiffait, 1956a: 42 (Othius; characters in key).
— Coiffait, 1972: 358 (Othius; characters; references; Canary Islands).
— Palm, 1976: 214 (Othius; characters; Canary Islands).
— Assing, 1997a: 105, 113 (Othius; characters; notes; Canary Islands).
— Assing, 1998a: 216 (Othius; Canary Islands).
— Assing, 1999a: 346 (Othius; Canary Islands).
— Assing, 1999c: 669 (Othius; Canary Islands).
▴ Distribution: Canary Islands.
monticola Cameron, 1943: 128 (Othius; Type locality: Darjeeling: Ghum district: Tiger Hill, altitude 8500–10,000 feet).
— Assing, 1998: 305 (Othius; characters; India).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).
▴ Distribution: India.
myrmecophilus Kiesenwetter, 1843, see: subuliformis Stephens, 1833.
neglectus Palm, 1976: 212 (Othius; Type locality: Gran Canaria: Teror, 450 m).
— Assing, 1997a: 106, 113 (Othius; characters; notes; Canary Islands).
— Assing, 1998a: 216 (Othius; characters; Canary Islands).
— Assing, 1999c: 669 (Othius; Canary Islands).
▴ Distribution: Canary Islands.
nepalensis Scheerpeltz, 1976, see: loeffleri Scheerpeltz, 1976.
obscurus Stephens, 1833, see: Gyrohypnus.
opacipennis Cameron, 1939c: 162 (Othius; Type locality: Yannanfou).
— Assing, 1999: 50 (Othius; characters; China).
— Assing, 1999c: 683 (Othius; characters; Sichuan).
▴ Distribution: China.
ozakii Ito, 1993, see: latus Sharp, 1874.
pallidus Brancsik, 1874, see: crassus Motschulsky, 1858.
palmaensis Assing, 1997a: 110 (Othius; Type locality: Isl. Can., La Palma, Cumbre Nueva, O-Seite, Fayal-Brezal, 1320 m).
— Assing and Wunderle, 1999: 435 (Othius; collecting notes; Canary Islands).
— Assing, 1999c: 679 (Othius; collecting notes; Canary Islands).
▴ Distribution: Canary Islands.
paralleliceps Quedenfeldt, 1882: 183 (Othius; Type locality: Hispania mer.: Algeciras in Andalusia; [Note: See lectotype designation by Assing, 1997]. Also cited from Marocco: Tetuan).
— Quedenfeldt, 1884: 375 (Othius; characters; Spain; Morocco).
— Fauvel, 1886: 47 [= 1886a: 39] (Othius; notes; Morocco; Spain).
— Fauvel, 1902b: 99 (Othius; Morocco).
— Reitter, 1908: 105 (Othius; characters; Andalusia).
— Bernhauer and Schubert, 1914: 318 (Othius; catalog).
— Scheerpeltz, 1933: 1323 (Othius; catalog).
— C. Koch, 1941: 61 (Othius; Morocco).
— Coiffait, 1972: 342 (Othius; characters; references; Morocco; Spain).
— Assing, 1997: 53 (Othius; lectotype designation: Algeciras; characters; notes; Morocco; Gibraltar).
— Assing, 1999c: 672 (Othius; notes; correct measurements for TiL and TaL cited in Assing, 1997, by factor of 2.02; Morocco).
▴ Distribution: Morocco, Spain.
xantholinoides Fairmaire, 1883: cviii (Othius; Type locality: Tetuan, Maroc; [Note: See lectotype designation by Assing, 1997]).
— Fauvel, 1886: 47 [= 1886a: 39] (Othius; Morocco).
— Fauvel, 1902b: 99 (Othius; synonym of paralleliceps).
— Bernhauer and Schubert, 1914: 318 (Othius; synonym of paralleliceps).
— C. Koch, 1941: 61 (Othius; characters; Morocco).
— Coiffait, 1972: 331, 342 (Othius; variety of paralleliceps; characters).
— Assing, 1997: 53 (Othius; lectotype designation: Tetuan, Maroc; synonym of paralleliceps).
parentium Solodovnikov, 1996: 300 (Othius; Type locality: Krasnodarskii Krai, 30 km sev.-zap. Dagomysa, gora Khuko, 1600 m).
— Solodovnikov, 1996a: 347 (Othius; translation of Solodovnikov, 1996).
— Assing, 1997: 102 (Othius; characters; notes; Russia).
▴ Distribution: Russia.
parvipennis Shibata, 1973a: 128 (Othiellus; Type locality: Taiwan: Alishan, about 2200 m alt., Chiai Hsien).
— Ito, 1989: 27 (Othius; characters; Taiwan).
— Assing, 1999: 64 (Othius; characters; Taiwan).
▴ Distribution: Taiwan.
permutatus Assing, 1997: 83, 129 (Othius; Type locality: Romania mer., Munţii Bucegi, Valea Cerbului, 900–1000 m. Also cited from Poland-Slovakia and Ukraine).
— Assing and Schülke, 1999: 12 (Othius; characters).
— Assing, 1999c: 675 (Othius; Poland; Slovakia; Ukraine; Romania).
— Burakowski, Mroczkowski, and Stefańska, 2000: 48 (Othius; catalog; Poland).
▴ Distribution: Romania, Poland, Slovakia, Ukraine.
perreaui Coiffait, 1984e: 376 (Othius; Type locality: Nepal oriental: Milke Himal, 2000 m).
— Assing, 1998: 325 (Othius; characters; Nepal).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).
▴ Distribution: Nepal.
philonthoides Wollaston, 1864: 581 (Othius; Type locality: Grand Canary: El Monte).
— Wollaston, 1865: 499 (Othius; notes; Canary Islands).
— Fauvel, 1869: 490 (Othius; synonym of brachypterus).
— Fauvel, 1869a: 152 (Othius; synonym of brachypterus).
— Wollaston, 1871: 305 (Othius; notes; valid species).
— Fauvel, 1902b: 99 (Othius; synonym of brachypterus).
— Bernhauer and Schubert, 1914: 317 (Othius; synonym of brachypterus).
— Coiffait, 1972: 354 (Othius; synonym of brachypterus).
— Palm, 1976: 214 (Othius; cited as valid species; characters; Canary Islands).
— Assing, 1997a: 110, 113 (Othius; characters; notes; Canary Islands).
— Assing, 1998a: 216 (Othius; notes; synonyms; Canary Islands).
— Assing, 1999c: 678 (Othius; wing length; Canary Islands).
▴ Distribution: Canary Islands.
coiffaiti Lohse, 1963: 177 (Othius; proposed for Coiffait's misidentification of lapidicola; Type locality: Kanarische Inseln).
— Coiffait, 1956a: 41 (Othius; [Note: Cited as lapidicola; misidentification according to Lohse, 1963]; characters in key; France; Turkey; Caucasus; Spain; Canary Islands; [Note: I assume the Continental localities are erroneous]).
— Coiffait, 1972: 353 (Othius; characters; references; Canary Islands).
— Palm, 1976: 216 (Othius; characters; Canary Islands).
— Hernández, Outerelo, and Gamarra, 1994: 193 (Othius; Canary Islands).
— Assing, 1997a: 112, 113 (Othius; characters; notes; Canary Islands).
— Assing, 1998a: 216 (Othius; synonym of philonthoides).
pseudolapidicola Coiffait, 1964a: 508 (Othius; proposed for Coiffait's 1954: 165 misidentification of lapidicola; Type locality: Canaries, Ile de Hierro).
— Coiffait, 1954: 165 (Othius; [Note: Misidentification: cited as lapidicola according to Coiffait, 1964a: 508]; Canary Islands).
— Coiffait, 1972: 353 (Othius; synonym of coiffaiti).
— Assing, 1998a: 216 (Othius; synonym of philonthoides).
piceus Scriba, 1870: 82 (Othius; Type locality: Sierra de la Nieve bei Ronda; [Note: See lectotype designation by Assing, 1997]).
— Fauvel, 1873b: 83 [= 1873a: 369] (Othius; synonym of lapidicola).
— Ganglbauer, 1895: 471 (Othius; synonym of lapidicola).
— Bernhauer and Schubert, 1914: 318 (Othius; synonym of lapidicola).
— Smetana, 1958: 127 (Othius; synonym of lapidicola).
— Coiffait, 1972: 347 (Othius; may be the same as pilifer).
— Assing, 1997: 20 (Othius; lectotype designation; valid species; characters; notes; Portugal; Spain; Morocco; Algeria).
— Assing, 1999c: 665 (Othius; Spain; Morocco).
▴ Distribution: Portugal, Spain, Morocco, Algeria.
pilifer Quedenfeldt, 1882: 182 (Othius; Type locality: Marocco: Tetuan).
— Quedenfeldt, 1884: 375 (Othius; characters; Morocco).
— Fauvel, 1886: 48 [= 1886a: 40] (Othius; Morocco; Algeria).
— Fauvel, 1902b: 99 (Othius; Morocco; Algeria).
— Bernhauer and Schubert, 1914: 318 (Othius; catalog).
— Scheerpeltz, 1933: 1323 (Othius; catalog).
— Coiffait, 1956a: 40 (Othius; characters in key; North Africa; Spain).
— Coiffait, 1972: 345 (Othius; characters; references; North Africa; Spain).
— Coiffait, 1972: 347 (Othius; may be synonym of piceus).
— Coiffait, 1973a: 273 (Othius; Morocco).
— Frank, 1982: 32 (Othius; list of parasites).
— Assing, 1997: 20 (Othius; synonym of piceus).
reitteri Fuente, 1898: 83 (Othius; Type locality: Pozuelo, Ciudad Real, Espagne; [Note: See lectotype designation by Assing, 1997]. Originally cited from Pozuelo de Calatrava).
— Reitter, 1908: 106 (Othius; characters; Spain).
— Bernhauer and Schubert, 1914: 319 (Othius; catalog).
— Scheerpeltz, 1933: 1323 (Othius; catalog).
— C. Koch, 1941: 61 (Othius; Morocco).
— Coiffait, 1972: 356 (Othius; characters; references; Portugal; France; Spain).
— Assing, 1997: 20 (Othius; lectotype designation: Pozuelo, Ciudad Real, Espagne; synonym of piceus).
laufferi Bernhauer, 1909b: 199 (Othius; Type locality: Spanien, Prov. Madrid).
— Bernhauer and Schubert, 1914: 318 (Othius; cited as lauferi; catalog).
— Scheerpeltz, 1933: 1322 (Othius; catalog).
— Coiffait, 1972: 345 (Othius; characters; Spain; France).
— Assing, 1997: 20 (Othius; synonym of piceus).
pilifer Quedenfeldt, 1882, see: piceus Scriba, 1870.
pokharensis Coiffait, 1983d: 163 (Othius; Type locality: Nepal: Dhumpus Kara, environs de Pokhara).
— Assing, 1998: 298 (Othius; characters; Nepal; Sikkim; India).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).
▴ Distribution: Nepal, India.
ponticus Coiffait, 1978d: 167 (Othius; subspecies of transsilvanicus, cited as transylvanicus; Type locality: Col de Ciftecoprüler, 1000 m).
— Assing, 1997: 92 (Othius; valid species; characters; notes; Turkey).
— Assing and Solodovnikov, 1998: 305 (Othius; characters).
▴ Distribution: Turkey.
praecisus Assing, 1999: 76 (Othius; Type locality: Taiwan: Kaohsiung Hsien, Peinantashan trail, 2500 m).
proximus Gistel, 1857: 25 (Othius; [Note: Nomen dubium]; Type locality: Bayern).
pseudolapidicola Coiffait, 1964, see: philonthoides Wollaston, 1864.
pulchellus Perroud and Montrousier, 1864, see: Zeteotomus.
punctatus Bernhauer, 1923a: 124 (Othius; replacement name for puncticeps Bernhauer).
— Scheerpeltz, 1933: 1323 (Othius; catalog).
— Assing, 1999: 33 (Othius; characters; China; Taiwan).
▴ Distribution: China, Taiwan.
puncticeps Bernhauer, 1916d: 26 (Othius; [preoccupied]; Type locality: China: Kiautschou).
— Bernhauer, 1923a;124 (Othius; synonym of punctatus).
— Scheerpeltz, 1933: 1323 (Othius; synonym of punctatus).
arisanus Shibata, 1973a: 126 (Othiellus; Type locality: Taiwan: Alishan, about 2200 m alt., Chiai Hsien).
puncticeps Broun, 1894, see: Maorothius.
puncticeps Bernhauer, 1916, see: punctatus Bernhauer, 1923.
punctipennis Lacordaire, 1835, see: laeviusculus Stephens, 1833.
punctulatus Goeze, 1777: 730 (Staphylinus; Type locality: Not cited).
— Paykull, 1789: 22 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 318 cited fulgidus as a synonym of punctulatus; Paykull did not describe the species as new, he attributed it to Fabricius]; redescription; Sweden).
— Gmelin, 1790: 2035 (Staphylinus; characters; France).
— Turton, 1802: 518 (Staphylinus; cited as pustulatus; characters; France).
— Fauvel, 1886: 47 [= 1886a: 39] (Othius; Algeria).
— Ganglbauer, 1895: 469 (Othius; characters; Europe).
— Fauvel, 1902b: 99 (Othius; Algeria; Tunisia).
— Poppius, 1905: 10 (Othius; Shetland Islands).
— Sainte-Claire Deville, 1906: 94 (Othius; Corsica).
— Reitter, 1908: 105 (Othius; characters; Europe).
— Reitter, 1909: 133 (Othius; characters; Germany).
— Bernhauer and Schubert, 1914: 318 (Othius; catalog).
— Johansen, 1914: 427 (Othius; characters; Denmark).
— Eichelbaum, 1915a: 95 (Othius; abdominal segmentation; apical segements).
— Šulc, 1922 (Othius; osmeterium; anatomy; histology; physiology).
— Portevin, 1929: 367 (Othius; characters; France).
— Scheerpeltz, 1933: 1323 (Othius; catalog).
— West, 1937: 40 (Othius; Faroe Islands).
— Jeannel and Jarrige, 1949: 372 (Othius; cave species; France).
— V. Hansen, 1952: 24 (Othius; characters; Denmark).
— Coiffait, 1956a: 39 (Othius; characters in key).
— Smetana, 1958: 124 (Othius; characters; ecology; Czechoslovakia).
— Szujecki, 1960a (egg, larval and pupal characters; biological notes).
— Scheerpeltz, 1963a: 419 (Othius; Greece).
— Palm, 1963b: 55 (Othius; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 167 (Othius; characters; central Europe).
— Coiffait, 1965e: 120 (Othius; Turkey).
— Horion, 1965: 108 (Othius; widespread in northern and central Europe south to Portugal, Spain, Sardinia, Sicily, Italy, Croatia, Bosnia and Serbia).
— Coiffait, 1966: 22 (Othius; Turkey).
— Pototskaia, 1967: 48 (Othius; larval characters).
— Szujecki, 1968a: 726 (Othius; Poland).
— Kasule, 1970: 76 (Othius; larval characters).
— Kasule, 1970a: 55 (Othius; life history).
— Coiffait, 1972: 340 (Othius; characters; references; Europe; Scandinavia; Britain; North Africa; Siberia).
— Coiffait, 1973a: 273 (Othius; Morocco).
— Pope, 1977: 28 (Othius; Britain).
— Coiffait, 1978d: 163 (Othius; Turkey).
— Topp, 1979: 23 (Othius; development; diapause).
— Smetana, 1979: 51 (requested that International Commission on Zoological Nomenclature place punctulatus on the Official List of Specific Names in Zoology).
— Outerelo, 1980a: 57 (Othius; Spain).
— Bordoni, 1982: 389 (Othius; characters; Italy).
— Frank, 1982: 32 (Othius; list of parasites).
— ICZN, 1983: 85 (Othius; placed on Official List of Species Names in Zoology and fixed as the type species of Othius).
— Ádám, 1987: 148 (Othiellus; collecting notes; Hungary).
— Lucht, 1987: 103 (Othius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Tóth, 1989: 34 (Othius; characters; Hungary).
— Nowosad, 1990: 99 (Othius; collected from mole nest; notes; Poland).
— Assing, 1993: 35 (Othius; bionomics; Germany).
— Ciceroni, 1993: 76 (Othius; Italy).
— Welch, 1993: 228 (Othius; ovariole number and ovary structure).
— J. Li, 1993: 33 (Othius; China).
— Janák, 1993a: 47 (Othius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 27 (Othius; Italy).
— Assing, 1997: 46 (Othius; characters; notes; Sweden; Denmark; Great Britain; Morocco; Algeria; Tunisia; Canary Islands; Portugal; Spain; France; Germany; Switzerland; Austria; Italy; Poland; Czech Republic; Slovakia; Croatia; Bosnia-Herzegovina; Macedonia; Bulgaria; Romania; Greece; Turkey; Russia; Georgia; Armenia; Ukraine).
— Owen, 1999b: 61 (Othius; captured underground at roots of old trees; Britain).
— Assing and Wunderle, 1999: 437 (Othius; Canary Islands).
— Assing and Schülke, 1999: 8 (Othius; characters).
— Assing, 1999c: 670 (Othius; notes; correct measurements for TiL and TaL cited in Assing, 1997, by factor of 2.02; Morocco; Algeria; Spain; Canary Islands; France; Austria; Italy; Poland; Slovenia; Croatia; Greece; Anatolia; Moldavia; Ukraine; Kazakhstan).
— Owen, 2000: 251 (Othius; pitfall trap collections from underground at roots of trees; Britain).
— Assing, 2001: 75 (Othius; collected from xerothermous site; Germany).
▴ Distribution: Tunisia, Algeria, Canary Islands, Europe, Russia, Georgia, Armenia, Turkey, Kazakhstan.
fulvopterus Geoffroy, 1785: 167 (Staphylinus; Type locality: Paris).
— Villers, 1789: 421 (Staphylinus; characters; France).
— Gravenhorst, 1802: 33 (Staphylinus; synonym of nitidus Fabricius).
— Erichson, 1839b: 295 (Othius; synonym of fulvipennis).
— Kraatz, 1857d: 655 (Othius; synonym of fulvipennis).
— Ganglbauer, 1895: 469 (Othius; synonym of punctulatus).
— Reitter, 1908: 105 (Othius; synonym of punctulatus).
— Bernhauer and Schubert, 1914: 318 (Othius; synonym of punctulatus).
— Smetana, 1958: 124 (Othius; synonym of punctulatus).
— Coiffait, 1972: 340 (Othius; synonym of punctulatus).
— Bordoni, 1982: 389 (Othius; synonym of punctulatus).
— Assing, 1997: 46 (Othius; synonym of punctulatus).
fulvipennis Fabricius, 1793: 537 (Paederus; Type locality: Kiliae).
— Panzer, 1795: 362 (Paederus; characters; Germany).
— Turton, 1802: (Staphylinus; characters;).
— Gravenhorst, 1806: 106 (Staphylinus; synonym of fulminans Gravenhorst).
— Panzer, 1808: 24 (Staphylinus; characters; illustrations).
— Gyllenhal, 1810: 365 (Lathrobium; characters; Lapland).
— Zetterstedt, 1828: 82 (Lathrobium; characters; Lapland).
— C. Sahlberg, 1830: 335 (Staphylinus; synonym of fulminans Gravenhorst).
— Erichson, 1839a: 420 (Othius; characters; Germany).
— Heer, 1839: 247 (Othius; characters; Switzerland).
— Erichson, 1839b: 295 (Othius; characters; Europe).
— Stephens, 1839: 401 (Othius; characters; England).
— Jacobsen, 1849: 674 (Othius; notes; Denmark).
— Redtenbacher, 1849: 690 (Othius; characters; Austria).
— Hardy, 1851: 36 (Othius; notes; England).
— Fairmaire and Laboulbène, 1856: 498 (Othius; characters; France).
— Kraatz, 1857d: 654 (Othius; characters; Germany).
— Redtenbacher, 1857: 180 (Othius; characters; Austria).
— Thomson, 1859: 26 (type species of Othius).
— Thomson, 1860: 185 (Othius; characters; Scandinavia).
— Hochhuth, 1862: 69 (Othius; Russia).
— Fauvel, 1873b: 82 [= 1873a: 368] (Othius; characters; France; Europe).
— Fauvel, 1874: 338 [= 1874b: 44] (Othius; Siberia).
— Redtenbacher, 1874: 222 (Othius; characters; Austria).
— Seidlitz, 1875: 279 (Othius; characters; Baltic region).
— J. Sahlberg, 1876: 31 (Othius; Finland).
— Mulsant and Rey, 1877a: 222 (Othius; cited as fulvipenis; characters; France).
— Sharp, 1889: 250 (Othius; Japan).
— Fowler, 1888: 295 (Othius; characters; Britain).
— Reitter, 1891b: 195 (Othius; characters; Europe).
— Ganglbauer, 1895: 469 (Othius; synonym of punctulatus).
— Reitter, 1908: 105 (Othius; synonym of punctulatus).
— Sharp and Muir, 1912: 499 (Othius; characters of male genitalia).
— Bernhauer and Schubert, 1914: 318 (Othius; synonym of punctulatus).
— Scheerpeltz, 1933: 1323 (Othius; synonym of punctulatus).
— Smetana, 1958: 124 (Othius; synonym of punctulatus).
— Coiffait, 1972: 340 (Othius; synonym of punctulatus).
— Pope, 1977: 28 (Othius; synonym of punctulatus).
— Bordoni, 1982: 389 (Othius; synonym of punctulatus).
— Assing, 1997: 46 (Othius; synonym of punctulatus).
ustulatus Gravenhorst, 1802: 46 (Staphylinus; Type locality: Brunsvigae).
— Latreille, 1804: 332 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 106 (Staphylinus; variety of fulminans Gravenhorst).
— Curtis, 1829: 26 (Xantholinus; catalog; Britain).
— Stephens, 1833: 255 (Othius; characters; England).
— Erichson, 1839b: 295 (Othius; synonym of fulvipennis).
— Stephens, 1839: 402 (Othius; characters; England).
— Fairmaire and Laboulbène, 1856: 498 (Othius; synonym of fulvipennis).
— Kraatz, 1857d: 655 (Othius; synonym of fulvipennis).
— Mulsant and Rey, 1877a: 223 (Othius; synonym of fulvipennis).
— Ganglbauer, 1895: 469 (Othius; synonym of punctulatus).
— Bernhauer and Schubert, 1914: 318 (Othius; synonym of punctulatus).
— Smetana, 1958: 124 (Othius; synonym of punctulatus).
— Coiffait, 1972: 340 (Othius; synonym of punctulatus).
— Bordoni, 1982: 389 (Othius; synonym of punctulatus).
— Assing, 1997: 46 (Othius; synonym of punctulatus).
fulminans Gravenhorst, 1802: 47 (Staphylinus; Type locality: Brunsvigae).
— Latreille, 1804: 332 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 105 (Staphylinus; characters).
— Gyllenhal, 1810: 357 (Staphylinus; characters; Sweden).
— C. Sahlberg, 1830: 334 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 33 (Gyrohypnus; Sweden).
— Mannerheim, 1831: 447 (Gyrohypnus; Sweden).
— Griffith and Pidgeon, 1832: 292 (Xantholinus; cited as fulmineus).
— Runde, 1835: 11 (Gyrohypnus; characters; Germany).
— Lacordaire, 1835: 410 (Cafius; characters; France).
— Nordmann, 1837: 119 (Gyrohypnus; Sweden; Germany; Tauria).
— Erichson, 1839a: 420 (Othius; synonym of fulvipennis).
— Erichson, 1839b: 295 (Othius; synonym of fulvipennis).
— Hardy, 1851: 36 (Othius; synonym of fulvipennis).
— Fairmaire and Laboulbène, 1856: 498 (Othius; synonym of fulvipennis).
— Kraatz, 1857d: 655 (Othius; synonym of fulvipennis).
— Thomson, 1860: 185 (Othius; synonym of fulvipennis).
— J. Sahlberg, 1876: 31 (Othius; synonym of fulvipennis).
— Mulsant and Rey, 1877a: 223 (Othius; synonym of fulvipennis).
— Ganglbauer, 1895: 469 (Othius; synonym of punctulatus).
— Bernhauer and Schubert, 1914: 318 (Othius; synonym of punctulatus).
— Smetana, 1958: 124 (Othius; synonym of punctulatus).
— Coiffait, 1972: 340 (Othius; synonym of punctulatus).
— Bordoni, 1982: 389 (Othius; synonym of punctulatus).
— Assing, 1997: 46 (Othius; synonym of punctulatus).
deceptivus Cuvier, 1833: 196 (Staphylinus; Type locality: Not cited).
— Bernhauer and Schubert, 1914: 318 (Othius; synonym of punctulatus).
— Smetana, 1958: 124 (Othius; synonym of punctulatus).
— Coiffait, 1972: 340 (Othius; synonym of punctulatus).
— Bordoni, 1982: 389 (Othius; synonym of punctulatus).
— Assing, 1997: 46 (Othius; synonym of punctulatus).
donisthorpei Chitty, 1903: 151 (Othius; aberration of fulvipennis; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from New Forest).
— Scheerpeltz, 1933: 1323 (Othius; variety of punctulatus).
— Coiffait, 1972: 333, 340 (Othius; variety of punctulatus; characters).
— Pope, 1977: 28 (Othius; synonym of punctulatus).
ibericus Coiffait, 1956a: 39 (Othius; Type locality: Espagne: Rio Valsain, 1600 m, Pr. de Ségovie).
— Coiffait, 1972: 332, 340 (Othius; subspecies of punctulatus; characters).
— Assing, 1997: 46 (Othius; synonym of punctulatus).
quadratus Zheng, 1995: 342, 346 (Othius; Type locality: Sichuan, Wolong, 2400 m).
reitteri Fuente, 1898, see: piceus Scriba, 1870.
reticulatus Assing, 1999: 74 (Othius; Type locality: Taiwan: Nantou Hsien, near Tsifeng, 2200 m).
rhodicus Coiffait, 1976, see: lapidicola Märkel and Kiesenwetter, 1848.
rosti Bernhauer, 1907: 385 (Othius; Type locality: Nemoro und Sapporo auf Jesso; [Note: See lectotype designation by Assing, 1999]).
— Bernhauer and Schubert, 1914: 319 (Othius; catalog).
— Watanabe and Shibata, 1965: 321 (Othius, cited as Otiellus; Japan).
— Watanabe, 1994 (Othius; Rebun-tô Island).
— Watanabe, 1995d: 256 (Othius; Okushiri-tô Island).
— Watanabe, 1996a: 12 (Othius; cave collection; Japan).
— Assing, 1999: 16 (Othius; lectotype designation: Jesso, Nemoro, Japan; characters; Russia; Japan).
— Assing, 1999c: 682 (Othius; Russian Federation).
▴ Distribution: Japan, Russia.
ussuriensis Coiffait, 1974a: 200 (Othius; Type locality: Vladivostok, Chamora).
rudis Zheng, 1995: 343, 346 (Othius; Type locality: Sichuan: Mt. Emei, 1500 m).
ruficornis Cameron, 1928c: 561 (Othius; Type locality: Sikkim: Karponang, 10,000 feet [Note: See lectotype designation by Assing, 1998]).
— Cameron, 1932: 44 (Othius; characters; India).
— Scheerpeltz, 1933: 1323 (Othius; catalog).
— Assing, 1998: 309 (Othius; characters; lectotype designation; India).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).
▴ Distribution: India.
rufipennis Sharp, 1874: 49 (Othius; Type locality: Orakami Marsh; [Note: See lectotype designation by Assing, 1999]).
— Bernhauer and Schubert, 1914: 319 (Othius; catalog).
— Nakane, 1963a: 90 (Othiellus; characters; habitus photograph; Japan).
— Yuh, Paik, Kwon, and Lee, 1985: 239 (Othius; Korea).
— Assing, 1999: 29 (Othius; lectotype designation; characters; China).
— Assing, 1999c: 683 (Othius; Japan; China).
▴ Distribution: Japan, Korea, China.
ruivomontis Assing and Wunderle, 1995a: 62 (Othius; Type locality: Madeira: Achada do Teixeira, 1580 m).
schawalleri Assing, 1998: 332 (Othius; Type locality: Nepal: Myagdi Distr., Myagdi Khola, N. Dobang, 2800–3100 m).
scoticus Stephens, 1833: 256 (Othius; [Note: Nomen dubium]; Type locality: Scotland).
sculptipennis Assing, 1999: 49 (Othius; Type locality: China: Sichuan, 28.07 N, 101.05 E, 30 km NW Muli/Bowa).
semipunctatus Fairmaire and Germain, 1861, see: Neobisnius.
serratus Assing, 1997: 81 (Othius; Type locality: Kaukasus, Suram).
sexpunctatus Haliday, 1841, see: angustus Stephens, 1833.
shibatai Ito, 1989: 28 (Othius; Type locality: Taiwan: Chiayi Hsien: Alishan).
sinuosus Assing, 1998: 307 (Othius; Type locality: Karakorum: Haramosch-Massiv, Iskere, Kaltaro Bach).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).
▴ Distribution: India.
sipeki Roubal, 1919 [Note: Nomen nudum], see: angustus Stephens, 1833.
smetanai Assing, 1999: 67 (Othius; Type locality: Taiwan: Pingtung Hsien, Peitawushan, Kuai-Ku Hut, 2325 m).
solodovnikovi Assing, 1997: 43 (Othius; Type locality: W. Caucasus, Urup river near Urup mountain, 2000 m).
— Solodovnikov, 1996: 297 (Othius; [Note: Misidentified as jadwigae according to Assing, 1997: 43]; characters; Russia).
— Solodovnikov, 1996a: 345 (Othius; [Note: Misidentified as jadwigae according to Assing, 1997: 43]; translation of Solodovnikov, 1996).
— Assing, 1999c: 669 (Othius; Caucasus).
▴ Distribution: Russia, Caucasus.
stenocephalus Eppelsheim, 1881, see: angustus Stephens, 1833.
stoetzneri Bernhauer, 1931, see: latus Sharp, 1874.
strigulosus Wollaston, 1854: 575 (Othius; Type locality: Madeira: Ribeiro de Santa Luzia; Ribeiro Frio; [Note: See lectotype designation by Assing and Wunderle, 1995a]).
— Wollaston, 1857: 186 (Othius; habitat; Madeira).
— Wollaston, 1865: 498 (Othius; habitat; Madeira).
— Fauvel, 1902b: 99 (Othius; Madeira).
— Bernhauer and Schubert, 1914: 319 (Othius; catalog).
— Smetana, 1963b: 36 (Othius; Madeira).
— Coiffait, 1972: 349 (Othius; characters; Madeira).
— Palm, 1979: 269 (Othius; characters; collecting notes; Madeira).
— Assing and Wunderle, 1995a: 56 (Othius; lectotype designation: [Note: Locality not cited]; characters; bionomics; Madeira).
— Assing, 1998a: 214 (Othius; Madeira).
— Assing, 1999c: 672 (Othius; Madeira).
▴ Distribution: Madeira.
subuliformis Stephens, 1833: 256 (Othius; Type locality: London; Suffolk; [Note: See lectotype designation by Assing, 1997]).
— Stephens, 1839: 402 (Othius; characters; England).
— Reitter, 1908: 106 (Othius; synonym of melanocephalus).
— Bernhauer and Schubert, 1914: 318 (Othius; synonym of melanocephalus).
— Smetana, 1958: 127 (Othius; synonym of angustus).
— Coiffait, 1972: 348 (Othius; synonym of angustus).
— Bordoni, 1982: 394 (Othius; synonym of angustus).
— Assing, 1997: 62, 128 (Othius; lectotype designation: [Note: Locality not cited]; valid species; characters; notes; Sweden; Norway; Denmark; Great Britain; Belgium; Germany; Portugal; Spain; France; Italy; Switzerland; Austria; Poland; Czech Republic; Slovakia; Latvia; Russia; Romania; records from Slovenia, Hungary, Romania, Balkans, and the Caucasus are misidentifications).
— Assing and Schülke, 1999: 10, 11 (Othius; characters).
— Assing, 1999c: 673 (Othius; notes; Finland; Russian Federation; Denmark; Portugal; Spain; Germany; France; Switzerland; Swiss or Austrian territory; Austria; Czech Republic; Poland; Czech or Polish territory; Italy).
— Assing, 2001: 75 (Othius; collected from xerothermous site; Germany).
▴ Distribution: Europe.
myrmecophilus Kiesenwetter, 1843: 308 (Othius; Type locality: Not cited).
— Mannerheim, 1830: 34 (Gyrohypnus; [Note: Misidentification: Bernhauer and Schubert, 1914: 318 cited melanocephalus Mannerheim as a synonym of myrmecophilus Kiesenwetter; Mannerheim did not describe the species as new, he attributed it to Gyllenhal who cited Gravenhorst as the author]; Finland).
— Mannerheim, 1831: 448 (Gyrohypnus; [Note: Misidentification: Bernhauer and Schubert, 1914: 318 cited melanocephalus Mannerheim as a synonym of myrmecophilus Kiesenwetter; Mannerheim did not describe the species as new, he attributed it to Gyllenhal who cited Gravenhorst as the author]; Finland).
— Märkel, 1845: 235 (Othius; notes; with Formica fuliginosa).
— Redtenbacher, 1849: 824 (Xantholinus; characters; Austria).
— Kraatz, 1857d: 658 (Othius; characters; Germany).
— Redtenbacher, 1857: 179 (Othius; characters; Austria).
— Thomson, 1860: 186 (Othius; characters; Scandinavia).
— Fauvel, 1865a: 299 [= 1865b: 49] (Othius; habitat; France).
— Fauvel, 1873b: 83 [= 1873a: 369] (Othius; characters; France; Europe).
— Redtenbacher, 1874: 222 (Othius; characters; Austria).
— Seidlitz, 1875: 279 (Othius; characters; Baltic region).
— J. Sahlberg, 1876: 32 (Othius; Finland).
— Mulsant and Rey, 1877a: 228 (Othius; characters; France).
— Fowler, 1888: 296 (Othius; characters; Britain).
— Ganglbauer, 1895: 471 (Othius; characters; Europe).
— Reitter, 1908: 107 (Othius; characters; Europe).
— Reitter, 1909: 133 (Othius; characters; Germany).
— Poppius, 1913: 10 (Othius; Faroes).
— Bernhauer and Schubert, 1914: 318 (Othius; catalog).
— Johansen, 1914: 428 (Othius; characters; Denmark).
— Portevin, 1929: 368 (Othius; characters; France).
— Scheerpeltz, 1933: 1322 (Othius; catalog).
— West, 1937: 41 (Othius; Faroe Islands).
— Jeannel and Jarrige, 1949: 373 (Othius; cave species; France).
— V. Hansen, 1952: 26 (Othius; characters; Denmark).
— Coiffait, 1956a: 41 (Othius; characters in key).
— Smetana, 1958: 128 (Othius; characters; ecology; Czechoslovakia).
— Coiffait, 1958: 24 (Othius; valid species; [Note: Misidentified as stenocephalus according to Coiffait, 1972: 350]; characters).
— Palm, 1963b: 56 (Othius; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 168 (Othius; characters; central Europe).
— Smetana, 1964d: 73 (Othius; Czechoslovakia).
— Horion, 1965: 113 (Othius; widespread in northern and central Europe south to Portugal, Spain, and Italy).
— Pototskaia, 1967: 48 (Othius; larval characters).
— Szujecki, 1968a: 726 (Othius; Poland).
— Kasule, 1970: 76 (Othius; larval characters).
— Kasule, 1970a: 59 (Othius; life history).
— Coiffait, 1972: 350 (Othius; characters; references; Europe; Britain; Scandinavia; Caucasus).
— Pope, 1977: 28 (Othius; Britain).
— Schminke, 1978: 6 (Othius; development and temperature).
— Topp, 1979: 21 (Othius; development; diapause).
— Outerelo, 1980a: 57 (Othius; Spain).
— Bordoni, 1982: 385 (Othius; characters; Italy).
— Frank, 1982: 32 (Othius; list of parasites).
— Lucht, 1987: 103 (Othius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Tóth, 1989: 37 (Othius; characters; Hungary).
— Nowosad, 1990: 103 (Othius; collected from mole nest; notes; Poland).
— Assing, 1993: 31 (Othius; bionomics; Germany).
— Welch, 1993: 228 (Othius; ovariole number and ovary structure).
— Janák, 1993a: 47 (Othius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 27 (Othius; Italy).
— Assing, 1997: 62 (Othius; synonym of subuliformis).
— Outerelo, Gamarra, and Salgado, 1998: 128 (Othius; cited as valid species; collected from cave; Spain; Portugal).
— Owen, 1999a: 15 (Othius; cited as valid species; subterranean capture; Britain).
— Owen, 1999b: 61 (Othius; cited as valid species; captured underground at roots of old trees; Britain).
— Owen, 2000: 251 (Othius; cited as valid species; pitfall trap collections from underground at roots of trees; Britain).
linkei Bernhauer, 1915j: 267 (Othius; variety of myrmecophilus; Type locality: Umgebung Leipzigs).
— Bernhauer, 1923d: 181 (Othius; characters; Sachsen).
— Scheerpeltz, 1933: 1322 (Othius; aberration of myrmecophilus).
— Smetana, 1958: 128 (Othius; synonym of myrmecophilus).
— Coiffait, 1972: 333, 338, 350 (Othius; variety of myrmecophilus; characters).
— Assing, 1997: 62 (Othius; synonym of subuliformis).
atavisticus Rüschkamp, 1932: 156 (Othius; form of myrmecophilus; Type locality: Rheinufer bei Bonn).
— Smetana, 1958: 128 (Othius; synonym of myrmecophilus).
— Coiffait, 1972: 350 (Othius; synonym of myrmecophilus).
algarvensis Coiffait, 1964a: 507 (Othius; subspecies of myrmecophilus; Type locality: Portugal: Algarve, Barranco do Veilho).
— Coiffait, 1972: 338, 350 (Othius; subspecies of myrmecophilus; characters).
— Assing, 1997: 62 (Othius; synonym of subuliformis).
suturalis Motschulsky, 1845, see: Atrecus.
suturalis Motschulsky, 1858, see: lapidicola Märkel and Kiesenwetter, 1848.
svaneticus Assing, 1999c: 676 (Othius; Type locality: Caucasus, Swanetien).
taiwanus Ito, 1989: 25 (Othius; Type locality: Taiwan: Chiayi Hsien: Alishan).
transsilvanicus Ganglbauer, 1895: 471 (Othius; Type locality: Coodgebirge; [Note: See lectotype designation by Assing, 1997]. Originally cited from Transsilvanische Alpen).
— Reitter, 1908: 107 (Othius; characters; Transylvania).
— Bernhauer and Schubert, 1914: 319 (Othius; catalog).
— Scheerpeltz, 1933: 1323 (Othius; catalog).
— Smetana, 1958: 129 (Othius; characters; ecology).
— Szujecki, 1969a: 254 (Othius; notes; Poland).
— Coiffait, 1972: 343 (Othius; characters; references; Romania).
— Frank, 1982: 32 (Othius; list of parasites).
— Tóth, 1989: 37 (Othius; characters).
— Janák, 1993a: 47 (Othius; cited as transilvanicus; Slovakia).
— Assing, 1997: 78, 128 (Othius; lectotype designation: Coodgebirge; characters; notes; Romania; Ukraine; Poland).
— Assing and Schülke, 1999: 14 (Othius; characters).
— Assing, 1999c: 675 (Othius; Poland; Romania; Ukraine).
▴ Distribution: Romania, Slovakia, Poland, Uraine.
truncatus Assing, 1998: 316 (Othius; Type locality: Nepal: Taplejung Distr., pasture Lassetham, NW Yamputhin, 3300–3500 m).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).
▴ Distribution: Nepal.
tuberipennis Assing, 1999: 52 (Othius; Type locality: China: Yunnan prov., Dali).
turcmenus Fauvel, 1900d: 228 (Othius; Type locality: Turkestan: Verni; [Note: See lectotype designation by Assing, 1997]).
— Reitter, 1908: 105 (Othius; characters; Turkestan).
— Bernhauer and Schubert, 1914: 319 (Othius; catalog).
— Scheerpeltz, 1933: 1323 (Othius; catalog).
— Coiffait, 1972: 355 (Othius; characters; references; Turkestan).
— Assing, 1997: 107 (Othius; lectotype designation; characters; notes; Kazakhstan).
— Assing, 1999c: 679 (Othius; characters; notes; correct measurements for TiL and TaL cited in Assing, 1997, by factor of 2.02; Kazakhstan; Kyrgyzstan).
▴ Distribution: Kazakhstan, Kyrgyzstan.
ushakovi Assing and Solodovnikov, 1998: 303 (Othius; Type locality: Georgia: Caucasus, Kodorsky range, Bakhundjara SE mt. Khodjali, h 1750 m).
ussuriensis Coiffait, 1974, see: rosti Bernhauer, 1907.
ustulatus Gravenhorst, 1802, see: punctulatus Goeze, 1777.
vestitus Wollaston, 1857, see: jansoni Wollaston, 1854.
virgifer Assing, 1998: 320 (Othius; Type locality: O. Nepal: Numbug [= Mumbug], O. Makalu, 3500–3700 m).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).
▴ Distribution: Nepal.
volans J. Sahlberg, 1876: 32 (Othius; Type locality: Helsingfors; [Note: See lectotype designation by Assing, 1997]. Also cited from Yläne).
— Reitter, 1908: 106 (Othius; characters; Finland).
— Bernhauer and Schubert, 1914: 319 (Othius; catalog).
— Scheerpeltz, 1933: 1323 (Othius; catalog).
— Renkonen, 1939: 64 (Othius; notes; Finland).
— V. Hansen, 1952: 26 (Othius; characters).
— Palm, 1963b: 56 (Othius; characters; Sweden; Denmark; Norway; Finland).
— Coiffait, 1972: 357 (Othius; characters; references; Finland).
— Lohse and Lucht, 1989: 160 (Othius; characters).
— Ødegaard and Andersen, 1994 (Othius; Norway).
— Assing, 1997: 40 (Othius; lectotype designation: H:fors [Helsingfors]; characters; notes; Finland; Germany; Italy; Switzerland reported by others).
— Uhlig, 1998: 136 (Othius; notes; Germany; Switzerland).
— Assing and Schülke, 1999: 11 (Othius; characters).
— Assing, 1999c: 668 (Othius; Switzerland; Italy).
— Burakowski, Mroczkowski, and Stefańska, 2000: 48 (Othius; catalog; Poland).
▴ Distribution: Finland, Norway, Germany, Switzerland, Italy.
wrasei Assing, 1999: 56 (Othius; Type locality: China: W. Sichuan, Daxue Shan, Hailuogou Glacier Park [Gongga Shan] Camp 1, 2100 m, 29.36 N, 102.04 E).
wunderlei Assing, 1997: 70 (Othius; Type locality: E. Andalusien, Sierra de Bermeja, Umg. Ronda, 1000 m).
xantholinoides Fairmaire, 1883, see: paralleliceps Quedenfeldt, 1882.
yakushimanus Ito, 1993, see: medius Sharp, 1874.
yushanus Ito, 1989: 29 (Othius; Type locality: Taiwan: Chiayi Hsien: Mt. Yushan).
yusmargensis Coiffait, 1982a: 250 (Othius; Type locality: Cachemire: Yusmarg, 2300–2400 m).
zerchei Assing, 1997a: 108 (Othius; Type locality: Tenerife, Teno, Erjos, 900 m).
ziganensis Solodovnikov, 2000a: 325 (Othius; Type locality: Turkey: Trubzon, NW slopes of Kalkanli Daglari, 7–10 km NE of Zigana pass, 1700–1800 m).
zolotarevi Roubal, 1911b: 401 (Othius; Type locality: Caucase occidental, montes Gagrenses; [Note: See lectotype designation by Assing, 1997]).
— Scheerpeltz, 1933: 1323 (Othius; catalog).
— Coiffait, 1972: 355 (Othius; characters; Caucasus).
— Solodovnikov, 1996: 295 (Othius; characters; Georgia).
— Solodovnikov, 1996a: 343 (Othius; translation of Solodovnikov, 1996).
— Assing, 1997: 59 (Othius; lectotype designation: Abchasia, Mont. Gagrenses, 6000′; characters; notes; Georgia).
— Assing, 1999c: 684 (Othius; correct measurements for TiL and TaL cited in Assing, 1997, by factor of 1.54).
▴ Distribution: Georgia.
Parothius
[2 species; Nearctic Region]
Parothius Casey, 1906: 422, 423 (species included: californicus). Type species: Othius californicus Mannerheim, fixed by monotypy.
— Bernhauer and Schubert, 1914: 317 (world catalog; 1 species).
— Leng, 1920: 106 (catalog of North American species).
— Blackwelder, 1952: 295 (type species: californicus).
— Hatch, 1957: 232 (characters; species of Pacific Northwest).
— Moore, 1963a: 94 (characters).
— Moore and Legner, 1975: 89 (catalog; North America north of Mexico).
— Moore and Legner, 1979: 72 (characters; notes).
— Smetana, 1982: 54 (characters; habitus; revision of and key to genera and species of North America north of Mexico).
— Campbell and Davies, 1991: 123 (checklist of species; Canada).
— Newton, Thayer, Ashe, and Chandler, 2000: 390 (2 Nearctic species; notes; characters in key).
californicus Mannerheim, 1843b: 228 (Othius; Type locality: California; [Note: See lectotype designation by Smetana, 1982]).
— Casey, 1906: 424 (Parothius; characters; California).
— Bernhauer and Schubert, 1914: 317 (Parothius; catalog).
— Hatch, 1957: 232 (Parothius; characters; British Columbia; Washington; Oregon).
— Smetana, 1982: 54 (Parothius; lectotype designation; characters; British Columbia; California; Oregon; Washington).
▴ Distribution: Canada, USA.
punctatus Smetana, 1982: 58 (Parothius; Type locality: California; Mendocino. Also cited from British Columbia and Oregon).
Tribe Platyprosopini
Platyprosopini Lynch, 1884.
— Eichelbaum, 1909: 159 (generic references; number of species in genus; general distribution of genus).
— Pototskaia, 1967: 45 (larval characters).
— Smetana, 1982: 22 (revision of genera and species of North America north of Mexico; characters).
— Newton and Thayer, 1992: 64 (tribe of Staphylininae).
— Newton, Thayer, Ashe, and Chandler, 2000: 389 (characters; notes).
Platyprosoparia Lynch, 1884: 165 (genera included: Platyprosopus). Type genus: Platyprosopus (see Newton and Thayer, 1992: 64).
Platyprosopina Sharp, 1887: 783 (genera included: Platyprosopus). Type genus: Platy-porsopus (see Newton and Thayer, 1992: 64).
Platyprosopi:
Platyprosopinae Moore, 1964a: 89 (characters; notes; genera included: Platyprosopus). Type genus: Platyprosopus (see Newton and Thayer, 1992: 64).
— Moore and Legner, 1979: 51 (characters).
— Newton and Thayer, 1992: 64 (synonym of Platyprosopini).
— Lawrence and Newton, 1995: 830 (included in Staphylininae).
Platyprosopini Coiffait, 1972: 126, 371 (genera included: Platyprosopus). Type genus: Platyprosopus (see Newton and Thayer, 1992: 65).
Platyprosopus
[50 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, and Oriental Regions]
Platyprosopus Mannerheim, 1830: 8, 36 (species included: elongatus). Type species: Platyprosopus elongatus Mannerheim, fixed by monotypy.
— Mannerheim, 1831: 422, 450 (characters).
— Dejean, 1833: 64 (list of species).
— Nordmann, 1837: 133 (characters).
— Erichson, 1839b: 292 (characters).
— Laporte, 1840: 181 (characters).
— Lacordaire, 1854: 63 (characters; notes; list of species).
— Schaum, 1859: 27 (catalog; European species).
— Gemminger and Harold, 1868: 608 (catalog).
— Redtenbacher, 1874: 223 (characters).
— Sharp, 1876: 101 (notes; characters).
— Heyden, 1880: 76 (list of species of Siberian region).
— Duvivier, 1883: 160 (catalog).
— Lynch, 1884: 166 (characters).
— Heyden, Reitter, and Weise, 1891: 105 (list of species of Europe and Caucasus).
— Heyden, 1893: 43 (list of species of Siberian region).
— Heyden, 1896: 33 (list of species of Siberian region).
— Heyden, Reitter, and Weise, 1906: 163 (list of species of Europe and Caucasus).
— Reitter, 1908: 103 (characters; key to species).
— Bernhauer and Schubert, 1914: 320 (world catalog; 28 species).
— Winkler, 1925: 374 (catalog for Palaearctic region).
— Cameron, 1925: 53 (catalog of Indian species).
— Bruch, 1928a: 427 (catalog; Argentina).
— Cameron, 1932: 47 (characters; key to species of British India).
— Scheerpeltz, 1933: 1324 (world catalog supplement).
— Normand, 1935: 370 (list of species of Tunisia).
— Blackwelder, 1944: 145 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 313 (type species: elongatus).
— Coiffait, 1956a: 37 (note).
— Moore, 1963a: 94, 95 (characters; notes).
— Coiffait, 1972: 371 (characters; larval characters; type species: elongatus; key to species of western Palaearctic region; references).
— Shibata, 1973c: 56 (checklist of species of Taiwan).
— Tikhomirova, 1973a: 180 (checklist of species of USSR).
— Moore and Legner, 1975: 71 (catalog; North America north of Mexico).
— Irmler, 1977: 58, 68 (key to Neotropical species; larval characters).
— Moore and Legner, 1979: 51 (characters; notes).
— Spahr, 1981: 96 (references to specimens from amber and copal).
— Smetana, 1982: 22 (characters; habitus; revision of species of North America north of Mexico).
— Newton, Thayer, Ashe, and Chandler, 2000: 390 (1 Nearctic species; notes; characters in key).
Metopius Mannerheim, 1830: 36 ([Note: Nomen nudum (Article 11.6), cited as a synonym of Platyprosopus; preoccupied by Panzer, 1806]).
— Mannerheim, 1831: 450 (synonym of Platyprosopus).
— Blackwelder, 1952: 241 (synonym of Platyprosopus; type species).
— Smetana, 1982: 22 (synonym of Platyprosopus).
Megaprosopus Reitter, 1908: 104 [= 1908a: 7] (species included: beduinus; [preoccupied by Macquart, 1843]). Type species: Platyprosopus beduinus Nordmann, fixed by monotypy.
— Bernhauer and Schubert, 1914: 320 (synonym of Platyprosopus).
— Cameron, 1932: 47 (subgenus of Platyprosopus).
— Scheerpeltz, 1933: 1324 (subgenus of Platyprosopus).
— Blackwelder, 1952: 235 (type species: beduinus).
— Coiffait, 1972: 371 (synonym of Platyprosopus subgenus Megaprosopoda).
Megaprosopoda E. Strand, 1935: 297 (replacement name for Megaprosopus Reitter). Type species: Platyprosopus beduinus Nordmann, fixed by objective synonymy with Megaprosopus.
— Blackwelder, 1952: 235 (type species).
— Coiffait, 1972: 371, 373 (subgenus of Platyprosopus; characters; type species: beduinus; key to species of western Palaearctic region; references).
aequalis Bernhauer and Schubert, 1914: 320 (Platyprosopus; replacement name for parallelus Bernhauer).
— Cameron, 1932: 52 (Platyprosopus; characters; Bengal).
— Scheerpeltz, 1933: 1324 (Platyprosopus; catalog).
— Shibata, 1973a: 131 (Platyprosopus; Taiwan).
▴ Distribution: India, Taiwan.
parallelus Bernhauer, 1911: 86 (Platyprosopus; [preoccupied]; Type locality: N. Bengal: Katihar, Purneah District).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; synonym of aequalis).
— Cameron, 1932: 53 (Platyprosopus; synonym of aequalis).
aethiopicus Bernhauer, 1939a: 81 (Platyprosopus; Type locality: Südl. Aethiopien: Nanoropus, bords du Rodolphe, 565 m).
amazonicus Irmler, 1977: 62, 69 (Platyprosopus; Type locality: Brazil: Rio Branco, Igarapé Aripuaña, Territorio de Roraima).
araxis Reitter, 1891, see: bagdadensis Stierlin, 1867.
argentinus Irmler, 1977: 64 (Platyprosopus; Type locality: Chaco de Santiago del Estero, bords du Rio Salado, environs d'Icano).
arrowi Tottenham, 1961: 205 (Platyprosopus; Type locality: Nigeria: Onitsha).
bagdadensis Stierlin, 1867: 218 (Platyprosopus; Type locality: Bagdad).
— Fauvel, 1897d: 289 (Platyprosopus; Algeria).
— Fauvel, 1902b: 98 (Platyprosopus; Algeria; Tunisia).
— Reitter, 1908: 104 (Platyprosopus; characters; Syria; Transcaucasus; Arax valley).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; catalog).
— Scheerpeltz, 1933: 1325 (Platyprosopus; catalog).
— Scheerpeltz, 1961: 6 (Platyprosopus; Iran).
— Coiffait, 1972: 375 (Platyprosopus; characters; references; Transcaucasus; Middle East; North Africa).
— H. Franz, 1987: 69 (Platyprosopus; Cyprus).
— Gusarov, 1997: 286 (Platyprosopus; Saudi Arabia).
▴ Distribution: Azerbaijan, Iran, Iraq, Syria, Saudi Arabia, Cyprus, Tunisia, Algeria.
araxis Reitter, 1891a: 138 (Platyprosopus; Type locality: Ufer des Araxes bei Ordubad).
— Fauvel, 1902b: 98 (Platyprosopus; synonym of bagdadensis).
— Reitter, 1908: 104 (Platyprosopus; synonym of bagdadensis).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; synonym of bagdadensis).
— Coiffait, 1972: 375 (Platyprosopus; synonym of bagdadensis).
beduinus Nordmann, 1837: 133 (Platyprosopus; Type locality: Aegyptus).
— Erichson, 1839b: 293 (Platyprosopus; characters; Nubia).
— Fauvel, 1886: 47 [= 1886a: 39] (Platyprosopus; Egypt).
— Fauvel, 1902b: 98 (Platyprosopus; Egypt).
— Reitter, 1908: 7 (Platyprosopus; subgenus Megaprosopus; characters; Egypt; Nubia; Senegal).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; catalog).
— Scheerpeltz, 1933: 1325 (Platyprosopus; catalog).
— C. Koch, 1936b: 163 (Platyprosopus; characters; Egypt).
— Coiffait, 1972: 380 (Platyprosopus; subgenus Megaprosopoda; characters; references; Egypt; Erythraea; Chad; Senegal).
— Scheerpeltz, 1974c: 20 (Platyprosopus; Sudan).
— Frank, 1982: 45 (Platyprosopus; list of parasites).
▴ Distribution: Egypt, Eritrea, Sudan, Chad, Senegal.
bernhaueri Cerruti, 1950: 49 (Platyprosopus; Type locality: Somalia: territorio dei Rahanuim).
bhutanicus Coiffait, 1977b: 220 (Platyprosopus; subgenus Megaprosopoda; Type locality: Bhutan: Phuntsholing, 200–400 m).
biblus Bordoni, 1984: 337 (Platyprosopus; Type locality: Libano: Les Cèdres, Biblos, 2300 m).
bilineatus Fauvel, 1895: 239 (Platyprosopus; Type locality: Bírmanie, Bhamo).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; catalog).
— Cameron, 1932: 50 (Platyprosopus; characters; Burma).
— Scheerpeltz, 1933: 1325 (Platyprosopus; catalog).
▴ Distribution: Myanmar.
biseriatus Eppelsheim, 1895b: 400 (Platyprosopus; Type locality: Calcutta).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; catalog).
— Cameron, 1932: 52 (Platyprosopus; characters; Burma; India).
— Scheerpeltz, 1933: 1325 (Platyprosopus; catalog).
— Last, 1961: 306 (Platyprosopus; Cambodia).
▴ Distribution: India, Myanmar, Cambodia.
bruchi Bernhauer, 1923b: 58 (Platyprosopus; Type locality: Argentinien: Prov. Santa Fé, Rosario).
— Scheerpeltz, 1933: 1325 (Platyprosopus; catalog).
— Irmler, 1977: 62 (Platyprosopus; characters; Argentina).
▴ Distribution: Argentina.
bucephalus Lynch, 1884: 168 (Platyprosopus; Type locality: Provincia Bonaerensi [Baradero] et Territorio Missionum).
consularis Fauvel, 1895: 239 (Platyprosopus; Type locality: Pegu, Palon; Bírmanie; Siam, Cambodge; Cochinchine; Tonkin).
— Fauvel, 1904a: 69 (Platyprosopus; Burma; Siam; Cambodia; Indochina; Tonkin; Sumatra; Java).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; catalog).
— Bernhauer, 1922a: 150 (Platyprosopus; Cambodia).
— Cameron, 1932: 49 (Platyprosopus; characters; Burma; India).
— Scheerpeltz, 1933: 1325 (Platyprosopus; catalog).
▴ Distribution: India, Myanmar, Thailand, Cambodia, Vietnam, Indonesia.
daressalamensis Bernhauer, 1937: 300 (Platyprosopus; Type locality: Tanganyika: Dar-es-salam; Ukerewe-Insel im Victoria-Nyanza, Muruntuguru).
elatus Fauvel, 1895: 239 (Platyprosopus; replacement name for indicus Kraatz).
— Fauvel, 1904a: 70 (Platyprosopus; India; Burma).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; [Note: Fauvel, 1904 cited as the date of publication]; catalog).
— Cameron, 1932: 49 (Platyprosopus; characters; Burma; India).
— Scheerpeltz, 1933: 1325 (Platyprosopus; catalog).
— Coiffait, 1982a: 232 (Platyprosopus; India).
▴ Distribution: India, Myanmar, Cambodia.
indicus Kraatz, 1859: 114 (Platyprosopus; [preoccupied]; Type locality: India borealis).
— Fauvel, 1904a: 70 (Platyprosopus; synonym of elatus).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; synonym of elatus).
— Cameron, 1932: 49 (Platyprosopus; synonym of elatus).
— Last, 1961: 306 (Platyprosopus; cited as valid species; Cambodia).
elongatus Mannerheim, 1830: 36 (Platyprosopus; Type locality: Caucasus; Armenia).
— Mannerheim, 1831: 450 (Platyprosopus; characters; Caucasus; Armenia).
— Nordmann, 1837: 134 (Platyprosopus; characters; Caucasus; Grusiniae).
— Erichson, 1839b: 294 (Platyprosopus; characters; Grusia; Chersonesus Taurica).
— Laporte, 1840: 181 (Platyprosopus; characters; Caucasus; Crimea; Armenia).
— Hochhuth, 1849: 98 (Platyprosopus; characters).
— Solsky, 1870: 462 (Platyprosopus; Astrakhan).
— Redtenbacher, 1874: 223 (Platyprosopus; characters).
— Solsky, 1874: 184 (Philonthus; characters; Turkestan).
— Eppelsheim, 1892: 334 (Platyprosopus; Turkestan).
— Reitter, 1908: 104 (Platyprosopus; characters; southern Russia; Transcaucasus; Transcaspian region; Turkestan).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; catalog).
— Scheerpeltz, 1933: 1325 (Platyprosopus; catalog).
— Scheerpeltz, 1961b: 34 (Platyprosopus; Afghanistan).
— Scheerpeltz, 1961h: 252 (Platyprosopus; subgenus Platyprosopus; Afghanistan).
— Scheerpeltz, 1963: 11 (Platyprosopus; Afghanistan).
— Pototskaia, 1967: 45 (Platyprosopus; larval characters).
— Coiffait, 1972: 377 (Platyprosopus; characters; references; Russia; Caucasus; Transcaucasus; region south of Caspian Sea; Turkestan).
— Bordoni, 1975a: 151 (Platyprosopus; Afghanistan).
— Petrenko, 1980: 81 (Platyprosopus; Caucasus).
▴ Distribution: Ukraine, Russia, Georgia, Armenia, Afghanistan, Turkmenistan, Uzbekistan.
fraternus Bernhauer, 1922a: 150 (Platyprosopus; Type locality: Kambodja).
frontalis Sharp, 1876: 107 (Platyprosopus; Type locality: Tapajos).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; catalog).
— Irmler, 1977: 62 (Platyprosopus; characters; Brazil).
▴ Distribution: Brazil.
fuliginosus Erichson, 1840: 923 (Platyprosopus; Type locality: Bengala).
— Kraatz, 1859: 115 (Platyprosopus; characters; Bengal).
— Fauvel, 1904a: 70 (Platyprosopus; Burma; Bengal; Cambodia; Tonkin).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; catalog).
— Bernhauer, 1922a: 150 (Platyprosopus; Indochina; Cambodia).
— Cameron, 1932: 51 (Platyprosopus; characters; Bihar).
— Scheerpeltz, 1933: 1325 (Platyprosopus; catalog).
— Last, 1961: 306 (Platyprosopus; Cambodia).
— Scheerpeltz, 1976a: 86 (Platyprosopus; Nepal).
▴ Distribution: India, Nepal, Myanmar, Cambodia, Vietnam, “East Indies”.
orientalis Motschulsky, 1858a: 212 (Platyprosopus; Type locality: Indes orientales).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; synonym of fuliginosus).
— Cameron, 1932: 51 (Platyprosopus; synonym of fuliginosus).
fulvicollis Motschulsky, 1858a: 212 (Platyprosopus; Type locality: Indes orientales).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; catalog).
— Cameron, 1932: 51 (Platyprosopus; characters; Bihar).
— Scheerpeltz, 1933: 1325 (Platyprosopus; catalog).
— Coiffait, 1982a: 232 (Platyprosopus; India).
▴ Distribution: India, “East Indies”.
linearis Kraatz, 1859: 115 (Platyprosopus; Type locality: India orientalis).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; synonym of fulvicollis).
— Cameron, 1932: 51 (Platyprosopus; synonym of fulvicollis).
ghidinii Cerruti, 1950: 49 (Platyprosopus; Type locality: Gambela, Galla-Sidamo).
grandiceps Irmler, 1977: 65 (Platyprosopus; Type locality: Estancia la Noria, Rio San Javier, Santa Fé, Argentine).
gridellii Cerruti, 1950: 50 (Platyprosopus; Type locality: Eritrea: Tessenei).
hierichonticus Reiche and Saulcy, 1856: 361 (Platyprosopus; Type locality: Jourdain; Chypre).
— Kraatz, 1858a: 65 (Platyprosopus; characters; Greece).
— Peyron, 1858: 420 (Platyprosopus; Turkey).
— Reitter, 1908: 104 (Platyprosopus; cited as hierochonticus; characters; Syria; Palestine; Cyprus; Greece).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; catalog).
— Scheerpeltz, 1933: 1325 (Platyprosopus; catalog).
— Coiffait, 1972: 378 (Platyprosopus; cited as hierochonticus; characters; references; Palestine; Syria; Asia Minor; Cyprus; Greece).
— Assing and Wunderle, 2001: 36 (Platyprosopus; Cyprus).
▴ Distribution: Israel, Jordan, Syria, Cyprus, Turkey, Greece.
indicus Motschulsky, 1858a: 212 (Platyprosopus; Type locality: Indes orientales).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; catalog).
— Bernhauer, 1928: 15 (Platyprosopus; Sumatra).
— Cameron, 1932: 49 (Platyprosopus; characters; India).
— Scheerpeltz, 1933: 1325 (Platyprosopus; catalog).
— Cameron, 1933d: 386 (Platyprosopus; Sumatra).
— Paulian, 1941: 217 (Platyprosopus; larval characters; India).
▴ Distribution: India, Indonesia.
laevicollis Kraatz, 1859: 114 (Platyprosopus; Type locality: India borealis).
— Fauvel, 1895: 239 (Platyprosopus; synonym of indicus).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; synonym of indicus).
— Cameron, 1932: 49 (Platyprosopus; synonym of indicus).
indicus Kraatz, 1859, see: elatus Fauvel, 1895.
keiseri Scheerpeltz, 1961g: 238, 257 (Platyprosopus; subgenus Platyprosopus; Type locality: West-Madagaskar: Tsaramandroso).
kraatzi Bernhauer, 1908d: 326 (Platyprosopus; Type locality: Paraguay).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; catalog).
— Irmler, 1977: 60 (Platyprosopus; characters; Paraguay; Argentina).
▴ Distribution: Paraguay, Argentina.
laevicollis Kraatz, 1859, see: indicus Motschulsky, 1858.
laticeps Sharp, 1876: 103 (Platyprosopus; Type locality: Ega).
linearis Kraatz, 1859, see: fulvicollis Motschulsky, 1858.
longicollis Eppelsheim, 1885: 113 (Platyprosopus; Type locality: Goldküste: Adda).
major Sharp, 1876: 102 (Platyprosopus; Type locality: Ega).
— Bernhauer and Schubert, 1914: 321 (Platyprosopus; catalog).
— Irmler, 1977: 58 (Platyprosopus; characters; Brazil; Peru).
▴ Distribution: Brazil, Peru.
marshalli Bernhauer, 1937: 300 (Platyprosopus; Type locality: Kenya: Homa, South Kavirondo).
mateui Coiffait, 1966b: 341 (Platyprosopus; Type locality: Mauritanie, Tamour Tartafet, Tamchakett).
mexicanus Sharp, 1887: 783 (Platyprosopus; Type locality: Mexico, San Juan Bautista in Tabasco, Tlacotalpam; [Note: See lectotype designation by Smetana, 1982]).
— Bernhauer and Schubert, 1914: 321 (Platyprosopus; catalog).
— Irmler, 1977: 60 (Platyprosopus; characters; Mexico).
— Smetana, 1982: 24 (Platyprosopus; lectotype designation: San Juan Bautista; characters; Mexico; Texas).
▴ Distribution: Mexico, USA.
texanus Moore, 1963a: 96 (Platyprosopus; Type locality: Texas: Brownsville).
minor Sharp, 1876: 106 (Platyprosopus; Type locality: Ega).
— Bernhauer and Schubert, 1914: 321 (Platyprosopus; catalog).
— Irmler, 1977: 62 (Platyprosopus; characters; Brazil).
▴ Distribution: Brazil.
nigriventris Cameron, 1947a: 55 (Platyprosopus; Type locality: Abyssinia: Nargi).
niloticus Fauvel, 1904c: 275 (Platyprosopus; Type locality: Haute-Egypte: Assouan. Abyssinie).
— Bernhauer and Schubert, 1914: 321 (Platyprosopus; catalog).
— C. Koch, 1936b: 163 (Platyprosopus; characters; Egypt).
— Coiffait, 1972: 378 (Platyprosopus; characters; Egypt; Ethiopia).
▴ Distribution: Ethiopia, Egypt.
ochraceus Fauvel, 1900b: 72 (Platyprosopus; Type locality: Congo: Kinchassa).
opacifrons Sharp, 1876: 107 (Platyprosopus; Type locality: Ega).
— Bernhauer and Schubert, 1914: 321 (Platyprosopus; catalog).
— Irmler, 1977: 62 (Platyprosopus; characters; Brazil).
▴ Distribution: Brazil.
orientalis Motschulsky, 1858, see: fuliginosus Erichson, 1840.
pallescens Tottenham, 1961: 206 (Platyprosopus; Type locality: Senegal: Parc National du Niokolo-Koba, Badi).
parallelus Sharp, 1876: 104 (Platyprosopus; Type locality: Ega).
— Bernhauer and Schubert, 1914: 321 (Platyprosopus; catalog).
— Irmler, 1977: 60 (Platyprosopus; characters; Brazil).
▴ Distribution: Brazil.
parallelus Bernhauer, 1911, see: aequalis Bernhauer and Schubert, 1914.
puncticeps Sharp, 1876, see: rectus Sharp, 1876.
rectus Sharp, 1876: 105 (Platyprosopus; Type locality: Manaos).
— Bernhauer and Schubert, 1914: 321 (Platyprosopus; catalog).
— Irmler, 1977: 58, 60, 69 (Platyprosopus; synonym of puncticeps; characters; Brazil).
▴ Distribution: Brazil.
puncticeps Sharp, 1876: 104 (Platyprosopus; Type locality: Tapajos).
— Bernhauer and Schubert, 1914: 321 (Platyprosopus; catalog).
— Irmler, 1977: 57, 58 (Platyprosopus; synonym of puncticeps).
rufescens Sharp, 1876: 106 (Platyprosopus; Type locality: Ega; Tapajos; St. Paulo).
— Bernhauer and Schubert, 1914: 321 (Platyprosopus; catalog).
— Irmler, 1977: 62 (Platyprosopus; characters; Brazil).
▴ Distribution: Brazil.
similis Sharp, 1876: 108 (Platyprosopus; Type locality: Manaos).
socius Bernhauer, 1922a: 150 (Platyprosopus; Type locality: Kambodja; Pnom-Penh).
tamulus Erichson, 1840: 923 (Platyprosopus; Type locality: Bengala).
— Kraatz, 1859: 115 (Platyprosopus; characters; Bengal; India orientalis).
— Bernhauer and Schubert, 1914: 320 (Platyprosopus; variety of fuliginosus).
— Cameron, 1932: 50 (Platyprosopus; characters; Burma; Bihar).
— Scheerpeltz, 1933: 1325 (Platyprosopus; catalog; Indochina).
— Yadav and Pillai, 1975 (Platyprosopus, cited as Elatyprosophus; chromosomal structures).
— Scheerpeltz, 1976a: 86 (Platyprosopus; Nepal).
— Coiffait, 1982a: 232 (Platyprosopus; India).
— Barrion, Litsinger, and Last, 1991: 1078 (Platyprosopus; associated with rice agroecosystem).
▴ Distribution: India, Nepal, Myanmar, Vietnam.
tarumaensis Irmler, 1977: 69 (Platyprosopus; Type locality: Manaus, Rio Taruma Mirîm).
tchadicus Coiffait, 1968a: 135, 141 (Platyprosopus; Type locality: Tchad: Amdjeria).
texanus Moore, 1963, see: mexicanus Sharp, 1887.
tongai Bernhauer, 1939c: 116 (Platyprosopus; Type locality: Siam: Swarnkalok).
Tribe Staphylinini
Staphylinini Latreille, 1802.
— Reitter, 1908: 101, 106 (characters).
— Jakobson, 1908: 447 (characters).
— Eichelbaum, 1909: 169 (list of genera with references; generic synonyms; number of genera; number of species in each genus; general distribution of genera).
— Bernhauer and Schubert, 1914: 321 (world catalog).
— Johansen, 1914: 341 (characters).
— Porta, 1926: 92 (characters; species of Italy).
— Portevin, 1929: 346 (key to French genera).
— Cameron, 1932: 2, 53 (characters in key; characters; key to subtribes).
— Joy, 1932: 113 (key to British genera).
— Scheerpeltz, 1933: 1325 (world catalog supplement).
— Scheerpeltz, 1940: 42 (key to Palaearctic genera).
— Blackwelder, 1943: 394 (characters; key to genera of West Indies).
— Blackwelder, 1944: 131 (checklist of species from Mexico, Central and South America, and the West Indies).
— V. Hansen, 1952: 27 (characters; key to Danish genera).
— Coiffait, 1956e: 183, 186 (key to genera).
— Hatch, 1957: 173 (key to genera of Pacific Northwest).
— Smetana, 1958: 131 (characters; key to Czechoslovakian genera and species).
— Arnett, 1963: 275 (characters; list of North American genera).
— Palm, 1963b: 57 (characters; key to genera and species of Sweden).
— Lohse, 1964: 169 (characters; key to central European species).
— Pototskaia, 1967: 63 (larval characters).
— Coiffait and Saiz, 1968: 347, 363 (characters; Chilean species).
— Scheerpeltz, 1968a: 79 (catalog of Austrian species).
— Kasule, 1970: 56 (larval characters).
— Coiffait, 1974: 353 (characters; key to subtribes, genera, and species of western Palaearctic region).
— Muona, 1979: 15 (list of Scandinavian species).
— Burakowski, Mroczkowski, and Stefańska, 1980: 29 (catalog; Poland).
— Newton, 1985: 209 (notes on austral distribution and taxonomic notes).
— Outerelo and Gamarra, 1985: 53 (key to genera of Iberian Peninsula).
— Smetana, 1995e: 47, 48 (notes; key to subtribes).
— Klimaszewski, Newton, and Thayer, 1996: 151 (list of New Zealand genera; number of New Zealand species).
— M. Hansen, 1996: 104 (list of Danish species).
— Smetana and Davies, 2000: 4, 12, 37 (discussion of characters; characters and key to subtribes of north temperate region; checklist of genera and species).
— Smetana and Davies, 2000: 18 (key to genera and subgenera associated with Staphylinus sensu lato).
— Newton, Thayer, Ashe, and Chandler, 2000: 330, 392 (key to Nearctic subtribes; characters; notes).
Subtribe Amblyopinina
Amblyopinina Seevers, 1944.
Amblyopininae Seevers, 1944: 157 (genera included: Amblyopinus). Type genus: Amblyopinus (see Newton and Thayer, 1992: 65).
— Seevers, 1955 (revision; ecology; distribution; host relationships and list; evolution; key to genera).
— Coiffait and Saiz, 1968: 365 (notes; key to Chilean genera).
— Lawrence and Newton, 1995: 830 (included in Staphylininae).
Amblyopinini Seever, 1944.
— Machado-Allison, 1963: 380 (key to genera).
— Machado-Allison and Barrera, 1964: 182 (discussion of characters of some genera).
— Barrera, 1972: 79, 82 (key to genera of Peru; discussion).
— Newton, 1985: 209 (notes on austral distribution and taxonomic notes).
— Ashe and Timm, 1988: 48, 54–56 (key to genera; notes on interaction with hosts and on phylogenetic relationships).
— Timm and Ashe, 1989: 379 (comments on systematics of group).
Amblyopinodes
[12 species; Neotropical Region]
Amblyopinodes Seevers, 1955: 247 (species included: gahani, piceus, travassosi, claviger, peruvianus, similis). Type species: Amblyopinus gahani Fauvel, fixed by original designation.
— Machado-Allison, 1962: 313 (key to species).
— Machado-Allison, 1962a (notes; key to species; checklist of species with hosts).
— Machado-Allison, 1963 (characters; revision and classification of and key to species; list rodent hosts; distribution).
— Machado-Allison and Barrera, 1964: 181 (characters).
— Martinez, Barrera, and Machado-Allison, 1970: 132 (characters).
— Ashe and Timm, 1988: 49 (characters).
Streptopinodes Martinez, Barrera, and Machado-Allison, 1970: 133 (subgenus of Amblyopinodes; species included: longus). Type species: Amblyopinus longus E. Franz, fixed by original designation and monotypy.
adae Machado-Allison, 1962a: 83 (Amblyopinodes; Type locality: Brasil: Estado de São Paulo, Estación Biolólgica de Boraceia).
— Machado-Allison, 1963: 386 (Amblyopinodes; characters; notes; host rodent).
▴ Distribution: Brazil.
amazonicus Guerrero, 1996: 660 (Amblyopinodes; Type locality: Peru: Madre de Dios, Pakitza).
barrerai Machado-Allison, 1962, see: caviae Martínez and Prosen, 1958.
caviae Martínez and Prosen, 1958: 27 (Amblyopinodes; Type locality: Paraguay: Dpto. Boquerón, Gran Chaco, Guarnición Oruro).
— Machado-Allison, 1962a: 82 (Amblyopinodes; characters).
— Machado-Allison, 1963: 389 (Amblyopinodes; characters; notes; host rodent).
— Martinez, Barrera, and Machado-Allison, 1970: 136 (Amblyopinodes; subgenus Amblyopinodes; characters; Argentina; Bolivia; Brazil; Paraguay).
▴ Distribution: Argentina, Paraguay, Bolivia, Brazil.
barrerai Machado-Allison, 1962: 324 (Amblyopinodes; Type locality: Brasil: Viçosa, Estado de Ceará).
— Machado-Allison, 1962a: 82 (Amblyopinodes; characters).
— Machado-Allison, 1963: 386 (Amblyopinodes; characters; notes; host rodent).
— Martinez, Barrera, and Machado-Allison, 1970: 136 (Amblyopinodes; synonym of caviae).
claviger E. Franz, 1930: 71 (Amblyopinus; Type locality: Hansa, Sta. Catharina, Brasilien).
— Scheerpeltz, 1933: 1469 (Amblyopinus; catalog).
— Costa Lima, 1936: 58, 59 (Amblyopinus; synonym of gahani).
— Seevers, 1955: 249 (Amblyopinodes; characters; notes; Brazil).
— Machado-Allison, 1962a: 82 (Amblyopinodes; characters).
— Machado-Allison, 1963: 389 (Amblyopinodes; characters; notes; host rodent).
▴ Distribution: Brazil.
distinctus Machado-Allison, 1962, see: piceus Brèthes, 1926.
gahani Fauvel, 1901b: 6 (Amblyopinus; Type locality: La Plata).
— Bernhauer and Schubert, 1916: 441 (Amblyopinus; catalog).
— Scheerpeltz, 1933: 1469 (Amblyopinus; catalog).
— Costa Lima, 1936: 59 (Amblyopinus; characters; notes).
— Seevers, 1955: 248 (Amblyopinodes; characters; notes; Argentina).
— Machado-Allison, 1962: 320 (Amblyopinodes; characters; Argentina; Brazil).
— Machado-Allison, 1962a: 82 (Amblyopinodes; characters).
— Machado-Allison, 1963: 392 (Amblyopinodes; characters; notes; host rodent).
— Machado-Allison and Barrera, 1964: 176, 178 (Amblyopinodes; characters; notes).
— Martinez, Barrera, and Machado-Allison, 1970: 139 (Amblyopinodes; subgenus Amblyopinodes; notes; Argentina).
▴ Distribution: Argentina, Uruguay, Brazil, Peru.
similis Seevers, 1955: 251 (Amblyopinodes; Type locality: Peru: Hacienda Cadena, Marcapata, Cuzco, 1000 meters).
— Machado-Allison, 1962a: 82 (Amblyopinodes; characters).
— Machado-Allison, 1963: 402 (Amblyopinodes; characters; notes; host rodent).
— Machado-Allison and Antequera, 1968: 129 (Amblyopinodes; subspecies of gahani; characters; Peru).
— Barrera, 1972: 81 (Amblyopinodes; subspecies of gahani; notes; Peru).
munoai Mañé Garzón and San Martín, 1961: 103 (Amblyopinodes; Type locality: Uruguay: Bañado Tropa Vieja, Dto. Canelones).
— San Martin, 1963 (Amblyopinodes; characters of male genitalia).
— Machado-Allison, 1963: 396 (Amblyopinodes; characters; notes; host rodent).
— Machado-Allison and Barrera, 1964: 177, 178 (Amblyopinodes; subspecies of gahani; characters; notes; Uruguay).
nordestinus Machado-Allison, 1962: 327 (Amblyopinodes; Type locality: Brasil: São Benedito, Estado de Ceará).
— Machado-Allison, 1962a: 82 (Amblyopinodes; characters).
— Machado-Allison, 1963: 398 (Amblyopinodes; characters; notes).
— Machado-Allison and Barrera, 1964: 178 (Amblyopinodes; subspecies of gahani; characters; notes).
guimaraesi Machado-Allison, 1962a: 86 (Amblyopinodes; Type locality: Brasil: Estado de São Paulo, Estación Biológica de Boraceia).
— Machado-Allison, 1963: 390 (Amblyopinodes; characters; notes; host rodent).
— Martinez, Barrera, and Machado-Allison, 1970: 139 (Amblyopinodes; subgenus Amblyopinodes; characters; notes; Brazil).
▴ Distribution: Brazil.
longus E. Franz, 1930: 74 (Amblyopinus; Type locality: Hansa, Sta. Catharina, Brasilien).
— Scheerpeltz, 1933: 1469 (Amblyopinus; catalog).
— Costa Lima, 1936: 62 (Amblyopinus; characters).
— Seevers, 1955: 234 (Amblyopinus; characters; notes; Brazil).
— Martinez, Barrera, and Machado-Allison, 1970: 133 (Amblyopinodes; subgenus Streptopinodes; characters; Brazil).
▴ Distribution: Brazil.
major Machado-Allison and Barrera, 1972: 11 (Amblyopinodes; Type locality: Bolivar State, 85 Km SSE El Dorado, 1032 m).
munoai Mañé Garzón and San Martín, 1961, see: gahani Fauvel, 1901.
nordestinus Machado-Allison, 1962, see: gahani Fauvel, 1901.
peruvianus Seevers, 1955: 250 (Amblyopinodes; Type locality: Peru: Quince Mil, Cuzco, 680 meters).
— Machado-Allison, 1962a: 82 (Amblyopinodes; characters).
— Machado-Allison, 1963: 399 (Amblyopinodes; characters; notes; host rodent).
— Barrera, 1972: 81 (Amblyopinodes; notes; Peru).
▴ Distribution: Peru.
piceus Brèthes, 1926: 18 (Omaloxenus; Type locality: Misiones, par Isidoro Escalera, de San Ignacio, Argentina).
— Scheerpeltz, 1933: 1469 (Omaloxenus; catalog).
— Costa Lima, 1936: 58 (Amblyopinus; synonym of gahani).
— Seevers, 1955: 248 (Amblyopinodes; characters; notes; Argentina).
— Machado-Allison, 1962: 313 (Amblyopinodes; characters; Brazil; Argentina).
— Machado-Allison, 1962a: 82 (Amblyopinodes; characters).
— Machado-Allison, 1963: 399 (Amblyopinodes; characters; notes; host rodent; Brazil; Argentina).
▴ Distribution: Argentina, Brazil.
distinctus Machado-Allison, 1962a: 88 (Amblyopinodes; subspecies of piceus; Type locality: Brasil: Estado de Pernambuco, Municipio de Garanhuns).
similis Seevers, 1955, see: gahani Fauvel, 1901.
travassosi Costa Lima, 1927: 842 (Amblyopinus; Type locality: Japuhyba, Etat de Rio de Janeiro).
— Costa Lima, 1927a: 382 (Amblyopinus; cited as new species; characters; Brazil).
— Scheerpeltz, 1933: 1469 (Amblyopinus; catalog).
— Costa Lima, 1936: 61 (Amblyopinus; characters; Brazil).
— Seevers, 1955: 249 (Amblyopinodes; characters; Brazil).
— Machado-Allison, 1962: 322 (Amblyopinodes; characters; Brazil).
— Machado-Allison, 1962a: 82 (Amblyopinodes; characters).
— Machado-Allison, 1963: 401 (Amblyopinodes; characters; notes; host rodent).
▴ Distribution: Brazil.
venezolanus Machado-Allison and Barrera, 1972: 8 (Amblyopinodes; Type locality: Bolivar State, 85 km SSE El Dorado, 1032 m).
Amblyopinus
[37 species; Neotropical Region]
Amblyopinus Solsky, 1875: 10 (species included: jelskii, mniszechi). Type species: Amblyopinus jelskii Solsky, fixed by subsequent designation by R. Lucas, 1920: 88.
— Duvivier, 1883: 130 (catalog).
— A. Matthews, 1884: 93 (characters).
— Bernhauer and Schubert, 1916: 441 (world catalog; 8 species).
— Scheerpeltz, 1933: 1469 (world catalog supplement).
— Bruch, 1936: 94 (characters).
— Costa Lima, 1936: 66 (key to species).
— Fonseca, 1939 (parasitic species).
— Blackwelder, 1944: 150 (checklist of species from Mexico, Central and South America, and the West Indies).
— Seevers, 1944: 160 (key to species).
— Blackwelder, 1952: 48 (type species).
— Seevers, 1955: 221 (characters; notes; key to species; species group classification).
— Machado-Allison and Barrera, 1964: 181 (characters).
— Coiffait and Saiz, 1968: 366, 367 (characters).
— Ashe and Timm, 1988: 49 (characters).
— Ashe and Timm, 1995: 374, 376, 392 (notes on natural history; key to species of Central America; list of host records; distribution).
Omaloxenus Notman, 1923: 1 (species included: bequaerti). Type species: Omaloxenus bequaerti Notman, fixed by original designation and monotypy.
— Scheerpeltz, 1933: 1469 (world catalog supplement).
— Bruch, 1936: 58 (synonym of Amblyopinus).
— Blackwelder, 1952: 274 (type species: bequaerti).
— Seevers, 1955: 222 (synonym of Amblyopinus).
akodoni Seevers, 1955: 228 (Amblyopinus; Type locality: Peru: San Miguel, Tambo, Polanco, Ayacucho).
ancashi Seevers, 1955: 232 (Amblyopinus; Type locality: Peru: Quilcayhanca, Huaras, Ancash, 4000 meters).
angustus Arrow, 1907: 125 (Amblyopinus; Type locality: British Guiana, Mt. Roraima).
— Bernhauer and Schubert, 1916: 441 (Amblyopinus; catalog).
— Bruch, 1936: 59 (Amblyopinus; characters; British Guiana).
— Seevers, 1955: 247 (Amblyopinus).
▴ Distribution: Guyana.
barrerai Zaragoza Caballero and Hernández, 1993: 28 (Amblyopinus; Type locality: Oaxaca: 4.5 km NO Campamento a Santa María Totontepec, 2480 m).
bequaerti Notman, 1923: 2 (Omaloxenus; Type locality: Alto Itatiaya, Setto do Itatiaya, Brazil, on Monodelphis).
— Scheerpeltz, 1933: 1469 (Omaloxenus; catalog).
— Costa Lima, 1936: 58, 64 (Amblyopinus; synonym of henseli).
— Seevers, 1944: 168 (Amblyopinus; valid species; characters).
— Seevers, 1955: 247 (Amblyopinus).
▴ Distribution: Brazil.
bolivari Barrera, Machado-Allison, and Muñiz, 1960, see: schmidti Seevers, 1944.
brandesi Kraatz, 1900: 212 (Amblyopinus; Type locality: Peru, Callanga).
— Fauvel, 1900a: 64 (Amblyopinus; synonym of jelskii).
— Bernhauer and Schubert, 1916: 441 (Amblyopinus; catalog).
— Costa Lima, 1936: 57, 62 (Amblyopinus; synonym of jelskii).
— Seevers, 1955: 227 (Amblyopinus; valid species; characters; notes; Peru).
— Barrera, 1972: 79 (Amblyopinus; notes; Peru).
▴ Distribution: Peru.
chilomysi Seevers, 1955: 239 (Amblyopinus; Type locality: Colombia: Rio Balcones, Guasca, Cundinamarca, 2750 meters).
claviger E. Franz, 1930, see: Amblyopinodes.
colombiae Seevers, 1955: 231 (Amblyopinus; Type locality: Colombia: Las Frias, 10 km. east of Sonsón, Antioquia, 2400 meters).
delicatus Barrera and Machado-Allison, 1968: 90 (Amblyopinus; Type locality: Colombia: departamento del Valle, Calle, Pichinde, La Flora, 1900 m).
emarginatus Seevers, 1955: 239 (Amblyopinus; Type locality: Colombia: San Antonio, San Agustin, Huila).
— Machado-Allison and Barrera, 1964: 175 (Amblyopinus; collecting notes; Panama; Venezuela).
— Barrera, 1966: 284 (Amblyopinus; Panama).
— Barrera, and Machado-Allison, 1968: 96 (Amblyopinus; characters; Colombia).
— Machado-Allison and Barrera, 1972: 8 (Amblyopinus; Venezuela).
— Ashe and Timm, 1987 (Amblyopinus; interaction with rodent host; Costa Rica).
— Ashe and Timm, 1987a (Amblyopinus; interaction with rodent host; Costa Rica).
— Timm and Ashe, 1987 (Amblyopinus; comments on host and elevational specificity; Panama).
— Ashe and Timm, 1995: 376, 394 (Amblyopinus; characters; host cited; notes; Colombia; Costa Rica; Panama).
▴ Distribution: Venezuela, Colombia, Panama, Costa Rica.
fuegensis Arrow, 1907: 126 (Amblyopinus; Type locality: Tierra del Fuego, Useless Bay).
— Bernhauer and Schubert, 1916: 441 (Amblyopinus; cited as fulgensis; catalog).
— Scheerpeltz, 1933: 1469 (Amblyopinus; catalog).
— Bruch, 1936: 92 (Amblyopinus; probably synonym of jelskii).
— Costa Lima, 1936: 58, 63 (Amblyopinus; probably synonym of jelskii).
— Seevers, 1944: 163 (Amblyopinus; characters; Chile).
— Seevers, 1955: 237 (Amblyopinus; characters; Chile).
— Coiffait and Saiz, 1968: 367 (Amblyopinus; Chile).
— Martinez, Barrera, and Machado-Allison, 1970: 128 (Amblyopinus; characters; Chile; Argentina).
▴ Distribution: Argentina, Chile.
gahani Fauvel, 1901, see: Amblyopinodes.
germaini Fauvel, 1900, see: Megamblyopinus.
henseli H. Kolbe, 1911: 117 (Amblyopinus; Type locality: Süd Brasilien, Rio Grande do Sul oder Rio de Janeiro).
— Scheerpeltz, 1933: 1469 (Amblyopinus; catalog).
— Costa Lima, 1936: 58, 63, 64 (Amblyopinus; notes; probably synonym of waterhousei).
— Seevers, 1944: 167 (Amblyopinus; valid species; characters).
— Seevers, 1955: 247 (Amblyopinus; Brazil).
▴ Distribution: Brazil.
hershkovitzi Seevers, 1955: 238 (Amblyopinus; Type locality: Colombia: Valdivia, Antioquia, 1400 meters).
huilae Seevers, 1955: 242 (Amblyopinus; Type locality: Colombia: San Antonio, San Agustin, Huila, 2250 meters).
intermedius Machado-Allison and Barrera, 1972: 6 (Amblyopinus; Type locality: Trujillo State, 15 km e. Trujillo, hacienda La Misisi, 2360 m).
isabelae Barrera, 1966: 284 (Amblyopinus; Type locality: Mexico: Omiltemi, State of Guerrero).
— Ashe and Timm, 1995: 380, 394 (Amblyopinus; cited as isabellae; characters; host cited; notes; Mexico).
▴ Distribution: Mexico.
jansoni A. Matthews, 1878, see: Myotyphlus.
jelskii Solsky, 1875: 11 (Amblyopinus; Type locality: Montana de Chanchamago, haienda Amable Maria).
— A. Matthews, 1884: 92, 96 (Amblyopinus; notes; characters; Peru).
— Fauvel, 1900a: 64 (Amblyopinus; Peru).
— H. Kolbe, 1911: 117 (Amblyopinus; notes).
— Bernhauer and Schubert, 1916: 441 (Amblyopinus; catalog).
— Scheerpeltz, 1933: 1469 (Amblyopinus; catalog).
— Bruch, 1936: 93 (Ambylopinus; illustration).
— Costa Lima, 1936: 62 (Amblyopinus; characters).
— Seevers, 1944: 161 (Amblyopinus; characters; Peru).
— Seevers, 1955: 224 (Amblyopinus; notes).
— Barrera, 1972: 79 (Amblyopinus; notes; Peru).
▴ Distribution: Peru.
kalinowskii Seevers, 1955: 243 (Amblyopinus; Type locality: Peru: Maracapata, Limapunco, Cuzco, 2400 meters).
kofordi Seevers, 1955: 231 (Amblyopinus; Type locality: Peru: 17 miles south of Masocruz, Puno, 13,500 feet).
longus E. Franz, 1930, see: Amblyopinodes.
machadoi Barrera, 1972: 81 (Amblyopinus; Type locality: 14 km M la Oroya, Yauli, Junín, 4640 m, Peru).
marmosae Seevers, 1944: 169 (Amblyopinus; Type locality: Brazil: Apiahy, São Paulo).
metasternalis Machado-Allison and Barrera, 1972: 4 (Amblyopinus; Type locality: Trujillo State, 15 km E Trujillo, Hacienda La Misisi, 2360 m).
mniszechi Solsky, 1875, see: Megamblyopinus.
monticolus Seevers, 1955: 228 (Amblyopinus; Type locality: Peru: Lago Suche, Moquegua, 14,600 feet).
montivagans Seevers, 1955: 240 (Amblyopinus; Type locality: Colombia: San Cristobal, Bogota, Cundinamarca).
oryzomysi Seevers, 1955: 240 (Amblyopinus; Type locality: Colombia: Las Frias, 10 km. east of Sonson, Antioquia, 2400 meters).
pacae Seevers, 1955: 244 (Amblyopinus; Type locality: Peru: Marcapata, Cuzco, 3100 meters).
piceus Brèthes, 1926, see: Amblyopinodes.
piurae Seevers, 1955: 232 (Amblyopinus; Type locality: Peru: Tambo, Huancabamba, Pirua, 2880 meters).
proximus Machado-Allison and Barrera, 1972: 1 (Amblyopinus; Type locality: Amazonas, Cerro Duida, Cabecera del Caño Culebra, 1400 m).
punae Seevers, 1955: 227 (Amblyopinus; Type locality: Cailloma, Arequipa, Peru).
sanborni Seevers, 1944: 166 (Amblyopinus; Type locality: Peru: Segrario, Puno).
— Seevers, 1955: 238 (Amblyopinus; characters; notes; Peru).
— Barrera, 1972: 81 (Amblyopinus; notes; Peru).
▴ Distribution: Peru.
schmidti Seevers, 1944: 164 (Amblyopinus; Type locality: Guatemala: Santa Elena, Chimal).
— Seevers, 1955: 231 (Amblyopinus; characters; Guatemala).
— Barrera, Machado-Allison, and Muñiz, 1960: 128 (Amblyopinus; characters).
— Machado-Allison and Barrera, 1972: 8 (Amblyopinus; notes).
— Ashe and Timm, 1995: 385, 395 (Amblyopinus; characters; host cited; notes; Mexico; Guatemala).
▴ Distribution: Guatemala, Mexico.
bolivari Barrera, Machado-Allison, and Muñiz, 1960: 127 (Amblyopinus; Type locality: México: D.F.: El Zarco, a 3100 m).
— Machado-Allison and Barrera, 1964: 174 (Amblyopinus; collecting notes; Mexico).
— Machado-Allison and Barrera, 1972: 8 (Amblyopinus; subspecies of schmidti).
— Ashe and Timm, 1995: 388, 394 (Amblyopinus; subspecies of schmidti; characters; host cited; Mexico).
seeversi Machado-Allison and Barrera, 1964: 179 (Megamblyopinus; Type locality: Venezuela: Serranía del Avila, 2200 m alt.).
— Machado-Allison and Barrera, 1972: 6 (Amblyopinus; characters; Venezuela).
▴ Distribution: Venezuela.
spinipennis Seevers, 1955: 234 (Amblyopinus; Type locality: Peru: Maracapata, Limapunco, Cuzco).
tiptoni Barrera, 1966: 282 (Amblyopinus; Type locality: Panama: Bambito, Chiriqui, 5800 feet).
— Barrera, 1966a (Amblyopinus; characters; Costa Rica).
— Ashe and Timm, 1987a (Amblyopinus; interaction with rodent host; Costa Rica).
— Timm and Ashe, 1987 (Amblyopinus; comments on host and elevational specificity; Panama).
— Ashe and Timm, 1995: 389, 395 (Amblyopinus; characters; host cited; Costa Rica; Panama).
▴ Distribution: Panama, Costa Rica.
trapidoi Barrera and Machado-Allison, 1968: 93 (Amblyopinus; Type locality: Colombia: Departamento del Cauca, vecindad del Cerro Munchique, Cordillera Occidental, 60 Km W Popayán, 2500 m).
travassosi Costa Lima, 1927, see: Amblyopinodes.
waterhousei Fauvel, 1900a: 64 (Amblyopinus; Type locality: Equateur).
— Bernhauer and Schubert, 1916: 441 (Amblyopinus; catalog).
— Costa Lima, 1936: 63 (Amblyopinus; characters).
— Seevers, 1944: 165 (Amblyopinus; characters; Colombia).
— Seevers, 1955: 243 (Amblyopinus; characters; Colombia).
— Machado-Allison and Barrera, 1964: 176 (Amblyopinus; Ecuador; Colombia; Venezuela; collecting notes).
— Barrera, and Machado-Allison, 1968: 98 (Amblyopinus; notes; Colombia).
— Machado-Allison and Barrera, 1972: 2 (Amblyopinus; Venezuela).
▴ Distribution: Venezuela, Colombia, Ecuador.
wenzeli Martínez, Barrera, and Machado-Allison, 1970: 130 (Amblyopinus; Type locality: Brasil: Santa Catarina, Xixim).
Chilamblyopinus
[1 species; Neotropical Region]
Chilamblyopinus Ashe and Timm, 1988: 49 (species included: piceus). Type species: Chilamblyopinus piceus Ashe and Timm, fixed by monotypy and original designation.
piceus Ashe and Timm, 1988: 52 (Chilamblyopinus; Type locality: Chile: Osorno Prov., Valle de La Picada).
Edrabius
[11 species; Neotropical Region]
Edrabius Fauvel, 1900a: 63, 65 (species included: philippianus). Type species: Edrabius philippianus Fauvel, fixed by monotypy.
— Bernhauer and Schubert, 1916: 441 (world catalog; 1 species).
— Bruch, 1928a: 429 (catalog; Argentina).
— Scheerpeltz, 1933: 1469 (world catalog supplement).
— Bruch, 1936: 94 (characters).
— Blackwelder, 1944: 150 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 145 (type species: philippianus).
— Seevers, 1955: 254 (characters; notes; type species: philippianus).
— Coiffait and Saiz, 1968: 365, 366 (characters; key to Chilean species).
— Ashe and Timm, 1988: 48 (characters).
— Timm and Ashe, 1989 (larval characters for an undescribed species; comments on systematic position).
— Ashe, Timm, and Gallardo, 1996 (summary of distribution of species; host associations).
alticolus Seevers, 1955: 259 (Edrabius; Type locality: Peru: Pampa de Titire, 29 km. northeast of Tarata, Tacna, 14,600 feet).
— Barrera, 1972: 82 (Edrabius; notes; Peru).
— Ashe, Timm and Gallardo, 1996: 738 (Edrabius; host; Bolivia).
▴ Distribution: Peru, Bolivia.
argentinus Seevers, 1955: 258 (Edrabius; Type locality: Argentina: Hualfin, Catamarca, near Tucuman).
— Coiffait and Saiz, 1968: 366 (Edrabius; characters; Chile).
— Ashe, Timm and Gallardo, 1996: 739 (Edrabius; host; notes on variation; Argentina).
▴ Distribution: Argentina, Chile.
australis Ashe, Timm and Gallardo, 1996: 735 (Edrabius; Type locality: Chile: Bío-Bío Prov.: Laguna Laja; also host cited).
chilensis Scheerpeltz, 1957a: 222 (Edrabius; Type locality: Chile: Hoch-Anden, Curico, Las Tablas).
— Ashe, Timm and Gallardo, 1996: 739 (Edrabius; host; notes on variation; Chile).
▴ Distribution: Chile.
chilensiformis Ashe, Timm and Gallardo, 1996: 737 (Edrabius; Type locality: Chile: coquimob Prov.: 10 km N Puente Los Molles, 32°09′S, 71°31′W; also host cited).
grandis Ashe, Timm and Gallardo, 1996: 733 (Edrabius; Type locality: Argentina: Prov. Río Negro: 13 km WSW Comallo, el. 1150 m. Also cited from Chile; also host cited).
kuscheli Scheerpeltz, 1957a: 220 (Edrabius; Type locality: Chile: Hoch-Anden, Arica-Lipiche, 4400 m).
pearsoni Seevers, 1955: 258 (Edrabius; Type locality: Peru: Rio Santa Rosa, Puno, 13,300 feet).
peruanus Seevers, 1955: 259 (Edrabius; Type locality: Peru: Pichupichuni, 8 km. north of Huacullani, Puno, 12,600 feet).
— Barrera, 1972: 82 (Edrabius; notes; Peru).
— Ashe, Timm and Gallardo, 1996: 739 (Edrabius; host; Chile).
▴ Distribution: Peru, Chile.
philippianus Fauvel, 1900a: 65 (Edrabius; Type locality: Chili: desert d'Atacama, Antofagasta).
— H. Kolbe, 1911: 117 (Edrabius; notes).
— Bernhauer and Schubert, 1916: 441 (Edrabius; catalog).
— Scheerpeltz, 1933: 1469 (Edrabius; catalog).
— Bruch, 1936: 93, 94 (Edrabius; charactes; illustrations).
— Costa Lima, 1936: 65, 66 (Edrabius; characters).
— Seevers, 1955: 255 (Edrabius; characters; notes; Chile).
— Scheerpeltz, 1957a: 219 (Edrabius; characters; Chile; Argentina).
— Coiffait and Saiz, 1968: 366, 367 (Edrabius; characters; Chile).
— Ashe, Timm and Gallardo, 1996: 739 (Edrabius; type host identification corrected).
▴ Distribution: Chile, Argentina.
weiseri Seevers, 1955: 255 (Edrabius; Type locality: Argentina: Troquero, Jujuy).
Megamblyopinus
[3 species; Neotropical Region]
Megamblyopinus Seevers, 1955: 251 (species included: mniszechi, germaini). Type species: Amblyopinus mniszechi Solsky, fixed by original designation.
— Machado-Allison and Barrera, 1964: 181 (characters).
— Coiffait and Saiz, 1968: 366, 367 (characters).
— Ashe and Timm, 1988: 48 (characters).
chilensis Coiffait and Saiz, 1968: 367 (Megamblyopinus; Type locality: Chili: Camarones).
germaini Fauvel, 1900a: 63 (Amblyopinus; Type locality: Pérou).
— Bernhauer and Schubert, 1916: 441 (Amblyopinus; catalog).
— Bruch, 1936: 57 (Amblyopinus).
— Bruch, 1936: 58 (Amblyopinus).
— Seevers, 1955: 254 (Megamblyopinus; characters; notes; Peru).
— Barrera, 1972: 82 (Megamblyopinus; notes; Peru).
▴ Distribution: Peru.
mniszechi Solsky, 1875: 11 (Amblyopinus; Type locality: Montana de Chanchamago, haienda Amable Maria).
— Fauvel, 1900a: 63 (Amblyopinus; Peru).
— H. Kolbe, 1911: 117 (Amblyopinus; notes).
— Bernhauer and Schubert, 1916: 441 (Amblyopinus; catalog).
— Scheerpeltz, 1933: 1469 (Amblyopinus; catalog).
— Bruch, 1936: 58 (Amblyopinus; characters).
— Seevers, 1955: 251 (Megamblyopinus; characters; notes; Peru).
— Barrera, 1972: 82 (Megamblyopinus; notes; Peru).
▴ Distribution: Peru.
seeversi Machado-Allison and Barrera, 1964, see: Amblyopinus.
Myotyphlus
[1 species; Australian Region]
Myotyphlus Fauvel, 1883: 40 (species included: jansoni). Type species: Amblyopinus jansoni A. Matthews, fixed by monotypy.
— Duvivier, 1883: 137 (catalog).
— Bernhauer and Schubert, 1916: 440 (world catalog; 1 species).
— Scheerpeltz, 1933: 1468 (world catalog supplement).
— Blackwelder, 1952: 253 (type species: jansoni).
— Seevers, 1955: 259 (type species: jansoni).
— Hamilton-Smith and Adams, 1966 (may be in Quediini).
— Ashe and Timm, 1988: 48 (characters).
Cryptommatus A. Matthews, 1884: 88 (species included: jansoni). Type species: Amblyopinus jansoni A. Matthews, fixed by monotypy.
— Bernhauer and Schubert, 1916: 440 (synonym of Myotyphlus).
— Blackwelder, 1952: 113 (synonym of Myotyphlus; type species: jansoni).
jansoni A. Matthews, 1878: 278 (Amblyopinus; Type locality: Tasmania).
— Fauvel, 1883: 40 (Myotyphlus).
— A. Matthews, 1884: 91 (Cryptommatus; characters; Tasmania).
— H. Kolbe, 1911: 117 (Myotyphlus; notes).
— Bernhauer and Schubert, 1916: 440 (Myotyphlus; catalog).
— Lea, 1925: 253 (Cryptommatus; characters; Australia).
— Seevers, 1955: 260 (Myotyphlus; characters; notes; Australia).
— Hamilton-Smith and Adams, 1966 (Myotyphlus; not an obligate ectoparasite; found in guano in bat cave; Victoria).
▴ Distribution: Australia.
Subtribe Anisolinina
Anisolinina Hayashi, 1993b: 284 (genera included: Anisolinus, Amichrotus, Tympanophorus, Hesperosoma). Type genus: Anisolinus, by original designation.
Amichrotus[11 species; Palaearctic and Oriental Regions]
Amichrotus Sharp, 1889: 114 (species included: apicipennis). Type species: Amichrotus apicipennis Sharp, fixed by monotypy.
— Bernhauer and Schubert, 1914: 367 (world catalog; 4 species).
— Winkler, 1925: 381 (catalog for Palaearctic region).
— Cameron, 1925: 64 (catalog of Indian species).
— Scheerpeltz, 1933: 1376 (world catalog supplement).
— Blackwelder, 1952: 49 (type species: apicipennis).
— Sawada, 1961: 4 (characters in key).
— Shibata, 1973c: 63 (checklist of species of Taiwan).
— Tikhomirova, 1973a: 186 (checklist of species of USSR).
— Naomi, 1982: 128 (characters).
— Naomi, 1982a: 37 (characters; key to subgenera).
— Naomi, 1983: 49 (characters; species of Japan).
— Hammond, 1984: 205 (checklist of species of Borneo).
— Shibata, 1984: 109 (checklist of species of Japan).
— Shibata, 1986a: 125 (supplemental list of species from Taiwan).
— Hayashi, 2000: 24 (notes on empodial setae).
apicipennis Sharp, 1889: 115 (Amichrotus; Type locality: Miyanoshita to Junsai).
— Bernhauer and Schubert, 1914: 367 (Amichrotus; catalog).
— Cameron, 1949b: 468 (Amichrotus; China, Japan, Formosa).
— Nakane, 1963a: 92 (Amichrotus; characters; habitus photograph; Japan).
— Frank, 1982: 33 (Amichrotus; list of parasites).
— Naomi, 1983: 51 (Amichrotus; characters; Japan).
— J. Li, 1993: 41 (Amichrotus; China).
▴ Distribution: Taiwan, China, Japan.
bryanti Cameron, 1937: 23 (Thoracostrongylus; Type locality: Java; Borneo: Quop, W. Sarawak).
costatus Fauvel, 1895, see: Thoracostrongylus.
distinctus Cameron, 1932: 215 (Thoracostrongylus; Type locality: Burma: Ruby Mines).
— Scheerpeltz, 1933: 1407 (Thoracostrongylus; catalog).
— Hammond, 1984: 194, 195 (Amichrotus; Borneo).
▴ Distribution: Myanmar, Borneo.
doriae Gridelli, 1924f: 207 (Ontholestes; subgenus Thoracostrongylus; Type locality: Borneo, Sarawak).
— Scheerpeltz, 1933: 1407 (Thoracostrongylus; catalog).
— Hammond, 1984: 195 (Amichrotus; Borneo).
▴ Distribution: Borneo.
elegans Cameron, 1920a: 218 (Amichrotus; Type locality: Sidapur, Coorg).
— Cameron, 1932: 214 (Thoracostrongylus; characters; India).
— Scheerpeltz, 1933: 1407 (Thoracostrongylus; catalog).
— Cameron, 1933d: 391 (Thoracostrongylus; Sumatra).
— Hammond, 1984: 195 (Amichrotus; Borneo).
— Biswas and Biswas, 1985: 146 (Thoracostrongylus; India).
▴ Distribution: India, Indonesia, Borneo.
excellens Bernhauer, 1939d: 100 (Amichrotus; Type locality: Japan: Kamikochi).
— Naomi, 1983: 52 (Amichrotus; notes).
— Hayashi, 1995: 47 (Amichrotus; characters; review of type).
▴ Distribution: Japan.
formosanus Shibata, 1976a: 12 (Amichrotus; Type locality: Taiwan: near Tsifeng, about 2200 m alt., Nantou Hsien).
jacobsoni Bernhauer, 1915h: 231 (Amichrotus; Type locality: Java: Nongkodjadjar).
merritti Bernhauer, 1912g: 253 (Amichrotus; Type locality: Luzon: Laguna, Mt. Banajao).
miwai Bernhauer, 1943, see: Hesperosoma.
picticollis Cameron, 1937: 22 (Thoracostrongylus; Type locality: Java: Batoerraden).
ruficollis Cameron, 1932: 216 (Thoracostrongylus; Type locality: Burma: Ruby Mines).
— Scheerpeltz, 1933: 1407 (Thoracostrongylus; catalog).
— Hammond, 1984: 194, 195 (Amichrotus; Borneo).
▴ Distribution: Myanmar, Borneo.
sarawakensis Bernhauer, 1915, see: Thoracostrongylus.
semicyaneus Fauvel, 1895, see: Thoracostrongylus.
Anisolinus
[11 species; Palaearctic and Oriental Regions]
Anisolinus Sharp, 1889: 113 (species included: picticornis, elegans). Type species: Anisolinus picticornis Sharp, fixed by subsequent designation by R. Lucas, 1920: 98.
— Bernhauer and Schubert, 1914: 365 (world catalog; 4 species).
— Winkler, 1925: 381 (catalog for Palaearctic region).
— Cameron, 1925: 64 (catalog of Indian species).
— Blackwelder, 1952: 53 (type species).
— Sawada, 1961: 4 (characters in key).
— Tikhomirova, 1973a: 185 (checklist of species of USSR).
— Naomi, 1981: 109 (characters; key to species of Japan).
— Naomi, 1982: 128 (characters).
— Naomi, 1983: 49 (notes; list of species of Japan).
— Shibata, 1984: 107 (checklist of species of Japan).
— Naomi, 1991: 2 (characters; revision of species of Japan; phylogeny and biogeography of species of Japan).
— Hayashi, 1993b: 284 (characters).
— Schillhammer, 1999b: 93 (tribe Anisolinus).
— Hayashi, 2000: 24 (notes on empodial setae).
Hesperodes Scheerpeltz, 1965: 265 (species included: malaisei, nitidus; preoccupied by Coquillett, 1900). Type species: Hesperodes malaisei Scheerpeltz, 1965: 267, fixed by original designation.
— Lundgren, 1987: 14 (synonym of Blackwelderella).
— Schillhammer, 1999b: 93 (synonym of Anisolinus).
Amaurochlamys Scheerpeltz, 1965: 273 (species included: malaisei). Type species: Amaurochlamys malaisei Scheerpeltz, 1965: 275, fixed by original designation and monotypy.
— Schillhammer, 1999b: 93 (synonym of Anisolinus).
— Hayashi, 2000: 24 (cited as valid genus; notes on empodial setae).
Blackwelderella Lundgren, 1987: 14 (replacement name for Hesperodes Scheerpeltz). Type species: Hesperodes malaisei Scheerpeltz, fixed by objective synonymy with Hesperodes for which the type had been fixed.
abdominalis Schubert, 1911, see: Diatrechus.
aethiopicus Fauvel, 1907, see: Diatrechus.
anthracinus Fauvel, 1905, see: Diatrechus.
auronotatus Fauvel, 1895, see: Tolmerinus.
bicolor Bernhauer, 1906, see: Diatrechus.
birmanus Schillhammer, 1993b: 93 (Anisolinus; replacement name for malaisei Scheerpeltz, 1956: 275).
malaisei Scheerpeltz, 1965: 275 (Amaurochlamys; [preoccupied nec Scheerpeltz, 1965: 267]; Type locality: N.E. Burma, Kambaiti, 7000 ft.).
caeruleipennis Cameron, 1932: 250 (Craspedomerus; Type locality: Sikkim: Rungbong Valley).
— Scheerpeltz, 1933: 1410 (Craspedomerus; catalog).
— Scheerpeltz, 1976a: 148 (Craspedomerus; characters).
— Coiffait, 1984e: 374 (Craspedomerus; cited as coeruleipennis; Nepal).
— Schillhammer, 1999b: 94 (Anisolinus; notes).
▴ Distribution: India, Nepal.
elegans Sharp, 1889: 114 (Anisolinus; Type locality: Miyanoshita).
— Bernhauer and Schubert, 1914: 365 (Anisolinus; catalog).
— Nakane, 1963a: 92 (Anisolinus; characters; habitus photograph; Japan).
— Naomi, 1981: 110, 111 (Anisolinus; characters; Japan).
— Naomi, 1991: 2 (Anisolinus; characters; Japan).
— Hayashi, 1993b: 284 (Anisolinus; Japan).
▴ Distribution: Japan.
fasciiventris Fauvel, 1895, see: Tolmerinus.
femoratus Schubert, 1911, see: Diatrechus.
hayashii Sawada, 1961: 4 (Anisolinus; Type locality: Japan: Honshu: Mt. Koya, Wakayama).
— Naomi, 1981: 110, 111 (Anisolinus; characters; Japan).
— Naomi, 1991: 5 (Anisolinus; characters; Japan).
— Hayashi, 1993b: 284 (Anisolinus; Japan).
▴ Distribution: Japan.
humeralis Fauvel, 1907, see: Diatrechus.
kawanoi Sawada, 1963, see: tsurugiensis Sawada, 1961.
malaisei Scheerpeltz, 1965: 267 (Hesperodes; Type locality: N.E. Burma, Kambaiti, 1800 m).
— Lundgren, 1987: 14 (Blackwelderella).
— Schillhammer, 1999b: 93 (Anisolinus).
▴ Distribution: Myanmar.
malaisei Scheerpeltz, 1965: 275, see: birmanus Schillhammer, 1999.
nitidus Scheerpeltz, 1965: 269 (Hesperodes; Type locality: N.E. Burma, Kambaiti, 2000 m).
— Lundgren, 1987: 14 (Blackwelderella).
— Schillhammer, 1999b: 93 (Anisolinus).
▴ Distribution: Myanmar.
picticornis Sharp, 1889: 113 (Anisolinus; Type locality: Miyanoshita).
— Bernhauer and Schubert, 1914: 365 (Anisolinus; catalog).
— Nakane, 1963a: 92 (Anisolinus; characters; habitus photograph; Japan).
— Naomi, 1981: 110, 112 (Anisolinus; characters; Japan).
— Naomi, 1991: 6 (Anisolinus; characters; Japan).
— Hayashi, 1993b: 284 (Anisolinus; Japan).
— Watanabe, 1996a: 13 (Anisolinus; cited as pilicornis; cave collection; Japan).
▴ Distribution: Japan.
raffrayi Fauvel, 1905, see: Diatrechus.
rougemonti Naomi, 1993: 185 (Anisolinus; Type locality: Nepal: Sawa, 2500 m).
rufipennis Fauvel, 1895, see: Tolmerinus.
taoi Naomi, 1981: 113 (Anisolinus; Type locality: Japan: Yamanashi Pref.: Daibosatsu-pass).
— Naomi, 1991: 5 (Anisolinus; characters; Japan).
— Hayashi, 1993b: 284 (Anisolinus; Japan).
▴ Distribution: Japan.
tsurugiensis Sawada, 1961: 5 (Anisolinus; Type locality: Japan: Shikoku: Tokushima, Mt. Tsurugi).
— Nakane, 1963a: 92 (Anisolinus; [Note: Misidentified as kawanoi Sawada according to Shibata, 1984: 109]; characters; habitus photograph; Japan).
— Naomi, 1981: 110, 112 (Anisolinus; characters; Japan).
— Naomi, 1991: 5 (Anisolinus; characters; Japan).
▴ Distribution: Japan.
kawanoi Sawada, 1963: 92 (Anisolinus; Type locality: Shikoku).
wittmeri Coiffait, 1982a: 269 (Philonthopsis; [preoccupied]; Type locality: Nepal: Thodung via Those, 3100 m).
Hesperosoma
[3 species; Palaearctic and Oriental Regions]
Hesperosoma Scheerpeltz, 1965: 270 (species included: malaisei). Type species: Hesperosoma malaisei Scheerpeltz, fixed by original designation and monotypy.
Paramichrotus Naomi, 1982a: 37, 38 (subgenus of Amichrotus; species included: miwai).
Type species: Amichrotus miwai Bernhauer, fixed by original designation and monotypy.
malaisei Scheerpeltz, 1965: 271 (Hesperosoma; Type locality: N.E. Burma, Kambaiti, 7000 ft.).
miwai Bernhauer, 1943a: 177 (Amichrotus; Type locality: Formosa: Arisan).
— Shibata, 1976a: 10 (Amichrotus; characters; Taiwan).
— Naomi, 1982a: 38 (Amichrotus; subgenus Paramichrotus; characters; Taiwan).
— Hayashi, 1993b: 290 (Hesperosoma; characters; Taiwan).
▴ Distribution: Taiwan.
nanshanchiana Hayashi, 1993a: 123 (Hesperosoma; subspecies of miwai; Type locality: Taiwan: Nanshanchi, Nantou Hsien).
nanshanchiana Hayashi, 1993, see: miwai Bernhauer, 1943.
sakoi Hayashi, 1993a: 124 (Hesperosoma; Type locality: Taiwan: Fungchiifo, Chiai Hsien).
Misantlius
[7 species; Neotropical Region]
Misantlius Sharp, 1885: 393 (species included: optatus, torquatus, carinulatus, discedens). Type species: Misantlius carinulatus Sharp, fixed by subsequent designation by R. Lucas, 1920: 420.
— Bernhauer and Schubert, 1914: 367 (world catalog; 6 species).
— Blackwelder, 1944: 136 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 249 (type species).
carinulatus Sharp, 1885: 394 (Misantlius; Type locality: Mexico; Toxpam).
discedens Sharp, 1885: 395 (Misantlius; Type locality: Mexico, Yolotepec).
gebieni Bernhauer, 1942: 26 (Misantlius; Type locality: Kostarika: Vara Blanca, 1700 m).
jalappae Bernhauer, 1910b: 378 (Misantlius; Type locality: Mexiko: Jalapa).
optatus Sharp, 1885: 394 (Misantlius; Type locality: Mexico; Misantla).
rufipennis Sharp, 1887: 786 (Misantlius; Type locality: Mexico, Yolotepec, Juquila).
torquatus Sharp, 1885: 394 (Misantlius; Type locality: Guatemala, Calderas).
Montguillonius
[1 species; Palaearctic Region]
Montguillonius Coiffait, 1987: 497 (replacement name for Hesperopsis Coiffait). Type species: Hesperopsis sikkimensis Coiffait, fixed by objective synonymy with Hesperopsis for which a type had been fixed.
Hesperopsis Coiffait, 1982a: 268 (species included: sikkimensis; preoccupied by Dyar, 1905). Type species: Hesperopsis sikkimensis Coiffait, fixed by original designation and monotypy.
sikkimensis Coiffait, 1982a: 268 (Hesperopsis; Type locality: Sikkim: Dzongri, 3000 m).
Philomyceta
[2 species; Palaearctic Region]
Philomyceta Cameron, 1944: 12 (species included: caeruleipennis, affinis). Type species: Philomyceta caeruleipennis Cameron, by original designation.
affinis Cameron, 1944: 13 (Philomyceta; Type locality: Darjeeling: Ghum district).
caeruleipennis Cameron, 1944: 12 (Philomyceta; Type locality: Darjeeling: Ghum district).
Subtribe Hyptiomina
Hyptiomina Casey, 1906.
— Newton and Thayer, 1992: 65 (subtribe of Staphylinini).
— Smetana, 1995e: 48 (characters).
— Newton, Thayer, Ashe, and Chandler, 2000: 397 (characters; notes).
Hyptiomae Casey, 1906: 359, 361 (genera included: Hyptioma). Type genus: Hyptioma (see Newton and Thayer, 1992: 65).
Holisi Blackwelder, 1944: 143 (checklist of species from Mexico, Central and South America, and the West Indies).
Holisini Blackwelder, 1944.
Holisina Newton, 1988: 259.
Holisus
[31 species; Neotropical, Nearctic, and Ethiopian Regions]
Holisus Erichson, 1839b: 298 (species included: humilis, analis, debilis, guildingii). Type species: Holisus analis Erichson, fixed by subsequent designation by Duponchel, 1841: 57.
— Lacordaire, 1854: 64 (characters; notes; list of species).
— Gemminger and Harold, 1868: 596 (catalog).
— Sharp, 1876: 180 (notes).
— Duvivier, 1883: 153 (catalog).
— Sharp, 1885: 462 (notes; Mexican and Central American species).
— Bernhauer and Schubert, 1914: 321 (world catalog; 23 species).
— Bruch, 1928a: 427 (catalog; Argentina).
— Scheerpeltz, 1933: 1326 (world catalog supplement).
— Blackwelder, 1943: 460 (characters; type species: analis; key to species of West Indies).
— Blackwelder, 1944: 143 (checklist of species from Mexico, Central and South America, and the West Indies).
— Scheerpeltz, 1956b: 19 (key to African species).
— Newton, 1988: 261 (checklist).
— Newton, Thayer, Ashe, and Chandler, 2000: 397 (1 unidentified Nearctic species; notes; characters in key).
Piestomorphus Motschulsky, 1858: 666 (species included: ater). Type species: Piestomorphus ater Motschulsky, fixed by monotypy.
— Gemminger and Harold, 1868: 596 (synonym of Holisus).
— Sharp, 1885: 462 (synonym of Holisus).
— Bernhauer and Schubert, 1914: 321 (synonym of Holisus).
— Blackwelder, 1943: 460 (synonym of Holisus; type species: ater).
— Blackwelder, 1952: 309 (synonym of Holisus; type species: ater).
Hyptioma Casey, 1906: 362 (species included: cubensis). Type species: Hyptioma cubensis Casey, fixed by monotypy.
— Bernhauer and Schubert, 1914: 320 (world catalog; 1 species).
— Scheerpeltz, 1933: 1326 (synonym of Holisus).
— Blackwelder, 1943: 460 (synonym of Holisus; type species: cubensis).
— Blackwelder, 1952: 197 (synonym of Holisus; type species: cubensis).
abdominalis Wendeler, 1956a: 230 (Holisus; Type locality: Brasilien: Espirito Santo, Sta. Theresa).
africanus Bernhauer, 1932, see: Medon.
analis Erichson, 1839b: 299 (Holisus; Type locality: Caracas Columbiae).
— Fauvel, 1891: 119 (Holisus; Venezuela).
— Bernhauer and Schubert, 1914: 321 (Holisus; catalog; Brazil; Peru).
▴ Distribution: Peru, Brazil, Venezuela.
ater Motschulsky, 1858: 667 (Piestomorphus; Type locality: Colombie).
— Fauvel, 1891: 118 (Holisus; Venezuela).
— Bernhauer and Schubert, 1914: 321 (Holisus; catalog).
▴ Distribution: Venezuela, Colombia.
atratulus Sharp, 1885: 464 (Holisus; Type locality: Guatemala, El Tumbador 2500 feet, Capetillo).
cingulatus Cameron, 1918, see: Holisomimus.
collaris Baudi, 1848: 136 (Lithocharis; Type locality: Venezuela).
— Kraatz, 1857b: 176 (Holisus; transferred from Lithocharis).
— Fauvel, 1891: 119 (Holisus; Venezuela).
— Bernhauer and Schubert, 1914: 321 (Holisus; catalog).
▴ Distribution: Venezuela.
condei Wendeler, 1956a: 229 (Holisus; Type locality: Brasilien: Espirito Santo, Leopoldina Col., Rio Bonito).
confinis Sharp, 1885: 462 (Holisus; Type locality: Panama, Bugaba).
cubensis Casey, 1906, see: debilis Erichson, 1839.
debilis Erichson, 1839b: 299 (Holisus; Type locality: Americae meridionalis ins. St. Johannis).
— Bernhauer and Schubert, 1914: 321 (Holisus; catalog).
— Blackwelder, 1943: 461 (Holisus; characters; Bahamas; Cuba; Haiti; Puerto Rico; St John; St. Lucia; St. Vincent).
▴ Distribution: West Indies.
cubensis Casey, 1906: 362 (Hyptioma; Type locality: Cuba, Havana).
— Bernhauer and Schubert, 1914: 320 (Hyptioma; catalog).
— Scheerpeltz, 1933: 1326 (Holisus; catalog).
— Blackwelder, 1943: 461 (Holisus; synonym of debilis).
rufoniger Cameron, 1922: 117 (Holisus; Type locality: Haiti; St. Vincent).
— Scheerpeltz, 1933: 1326 (Holisus; catalog).
— Blackwelder, 1943: 461 (Holisus; synonym of debilis).
depressus Sharp, 1876: 180 (Holisus; Type locality: Ega).
discedens Sharp, 1876: 183 (Holisus; Type locality: Not cited).
elatus Sharp, 1885: 465 (Holisus; Type locality: Guatemala, San Geronimo; Panama, Bugaba, Volcan de Chiriqui).
eleusoides Sharp, 1885: 466 (Holisus; Type locality: Guatemala, Las Mercedes 3000 feet, Cerro Zunil 4000–5000 feet).
excavatus Sharp, 1876: 182 (Holisus; Type locality: Ega).
filum Sharp, 1885: 464 (Holisus; Type locality: Guatemala, Teleman in the Polochic Valley).
guildingii Erichson, 1839b: 300 (Holisus; Type locality: Americae meridionalis ins. St. Vincentii).
— Bernhauer and Schubert, 1914: 322 (Holisus; catalog).
— Blackwelder, 1943: 462 (Holisus; characters; Puerto Rico; Antigua; St. Lucia; St. Vincent).
— Legner and Moore, 1977: 176 (Holisus; found under bark).
▴ Distribution: West Indies.
humilis Erichson, 1839b: 299 (Holisus; Type locality: Brasilia).
laevis Blackwelder, 1943: 463 (Holisus; Type locality: Jamaica: 1 mile east of Bath in parish of St. Thomas. Also cited from Cuba).
nigriceps Wendeler, 1956a: 229 (Holisus; Type locality: Brasilien: Nova Teutonia).
oblitus Sharp, 1885: 465 (Holisus; Type locality: Guatemala, near the city, Cerro Zunil 4000 to 5000 feet, Purula).
ogloblini Bernhauer, 1934b: 117 (Holisus; Type locality: Argentinien: Misiones, Loreto).
pallens Sharp, 1885: 464 (Holisus; Type locality: Panama, Volcan de Chiriqui 2000 to 3000 feet).
parvus Cameron, 1918, see: Holisomimus.
picipes Sharp, 1876: 181 (Holisus; Type locality: Ega).
planus Schubert, 1909: 295 (Holisus; Type locality: südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d'Orizaba).
prolixus Sharp, 1885: 463 (Holisus; Type locality: Guatemala, Cahabon, San Juan in Vera Paz).
protensus Sharp, 1885: 462 (Holisus; Type locality: Guatemala, San Geronimo, Rio Maria Linda).
rufipes Sharp, 1885: 464 (Holisus; Type locality: Guatemala, San Geronimo 3000 feet).
rufoniger Cameron, 1922, see: debilis Erichson, 1839.
rufopygus Bernhauer, 1916g: 265 (Holisus; Type locality: Bolivien: Yuracares).
schedli Scheerpeltz, 1956b: 19, 20 (Holisus; Type locality: Belgisch-Kongo: Hembe Bitale [see Scheerpeltz, 1956a: 484).
solskyi Sharp, 1885: 463 (Holisus; Type locality: Mexico, Cordova).
— Solsky, 1868a: 141 (Holius; [Note: Misidentification: Bernhauer and Schubert, 1914: 322, cited ater Solsky as a synonym of solskyi; Solsky did not describe the species as new, he attributed it to Motschulsky]; Mexico).
— Bernhauer and Schubert, 1914: 322 (Holisus; catalog).
▴ Distribution: Mexico.
sparsiceps Bernhauer, 1942: 24 (Holisus; Type locality: Kostarika).
umbra Sharp, 1876: 182 (Holisus; Type locality: Ega).
Subtribe Philonthina
Philonthina Kirby, 1837.
— Lohse, 1964: 170 (characters; key to central European species).
— Dettner, 1983 (comparative anatomy of defensive glands and defensive secretions).
— Dettner, 1987: 30 (defensive glands).
— Newton and Thayer, 1992: 65 (subtribe of Staphylinini).
— Dettner, 1993a (summary and discussion of defensive secretions and exocrine glands and their bearing on phylogeny).
— Smetana, 1995e (characters; revision of and key to species of USA and Canada; key to holarctic genera; revision of genera of USA and Canada; phylogeny; historical discussion; natural history; habitat; life cycle; immature stages; fossils).
— Klimaszewski, Newton, and Thayer, 1996: 151 (list of New Zealand genera; number of New Zealand species).
— Smetana and Davies, 2000: 12, 13 (characters).
— Newton, Thayer, Ashe, and Chandler, 2000: 332, 395 (key to Nearctic genera; characters; notes).
— Hayashi, 2000: 23 (notes on empodial setae).
Philonthidae Kirby, 1837: 91 (genera included: Philonthus). Type genus: Philonthus (see Newton and Thayer, 1992: 65).
Rémates Mulsant and Rey, 1877: 452 (genera included: Remus). Type genus: Remus (see Newton and Thayer, 1992: 65).
Craspedomerina Bernhauer, 1911.
— Newton and Thayer, 1992: 65 (subtribe of Staphylinini).
— Smetana, 1995e: 48 (characters).
— Smetana and Davies, 2000: 12 (synonym of Philonthina).
Craspedomeri Bernhauer, 1911: 88 (genera included: Craspedomerus). Type genus: Craspedomerus (see Newton and Thayer, 1992: 65).
— Cameron, 1932: 54, 248 (characters).
— Scheerpeltz, 1933: 1410 (world catalog supplement).
— Scheerpeltz, 1940: 47 (key to Palaearctic genera).
— Moore, 1960: 100 (key to genera).
Philonthini Coiffait, 1956e: 183 (genera included: none cited). Type genus: Philonthus (see Newton and Thayer, 1992: 65).
— Coiffait, 1963: 6 (characters; key to genera).
— Pototskaia, 1966: 152 (larval characters; key to some genera using larval characters).
— Pototskaia, 1967: 52 (larval characters).
— Coiffait and Saiz, 1968: 347 (key to Chilean genera).
— Scheerpeltz, 1968a: 64 (catalog of Austrian species).
— Coiffait, 1974: 2 (characters; key to genera and species of western Palaearctic region).
— Smetana, 1977b: 178 (synonym of Quediini).
— Muona, 1979: 14 (list of Scandinavian species).
— Outerelo and Gamarra, 1985: 48 (key to genera of Iberian Peninsula).
— Newton and Thayer, 1992: 65 (synonym of Philonthina).
Actinomorphus
[1 species; Oriental Region]
Actinomorphus Schillhammer, 1996: 59 (species included: lativentris). Type species: Actinomorphus lativentris Schillhammer, fixed by original designation and monotypy.
lativentris Schillhammer, 1996: 62 (Actinomorphus; Type locality: Sarawak: 4th Division GN. Mulu N.P., 100–500 m).
Actinus
[2 species; Australian Region]
Actinus Fauvel, 1878f: 250 (species included: imperialis). Type species: Actinus imperialis Fauvel, fixed by monotypy.
— Duvivier, 1883: 149 (catalog).
— Olliff, 1887: 495 (characters; Australian species).
— Bernhauer and Schubert, 1914: 373 (world catalog; 2 species).
— Blackwelder, 1952: 37 (type species: imperialis).
cupreosplendens Cameron, 1943, see: Mysolius.
macleayi Olliff, 1887: 495 (Actinus; Type locality: Johnstone River, Cairns, North Queensland).
— Bernhauer and Schubert, 1914: 373 (Actinus; catalog).
— Bernhauer, 1920: 16 (Actinus; Australia).
— Lea, 1925: 228 (Actinus; characters; Australia).
▴ Distribution: Australia.
imperialis Fauvel, 1878f: 250 (Actinus; Type locality: Nouvelle-Guinée méridionale, Fly River).
— Fauvel, 1879: 96 (Actinus; New Guinea).
— Bernhauer and Schubert, 1914: 373 (Actinus; catalog).
— Cameron, 1924: 87 (Actinus; New Guinea).
— Lea, 1925: 228 (Actinus; characters; Australia).
— Cameron, 1931: 363 (Actinus; New Guinea).
— Cameron, 1937a: 104 (Actinus; New Guinea).
— Last, 1987: 55 (Actinus; Papua New Guinea).
▴ Distribution: New Guinea, Australia.
Afrorabigus
[7 species; Ethiopian Region]
Afrorabigus Levasseur, 1965: 57 (species included: robusticornis, belingaensis, kolweziensis, muscicola, basilewskyi). Type species: Philonthus basilewskyi Tottenham, fixed by original designation.
basilewskyi Tottenham, 1962: 195 (Philonthus; Type locality: Uganda: Bugiri, 1400 m).
belingaensis Levasseur, 1965: 61 (Afrorabigus; Type locality: Gabon: Mission biologique au Gabon, Belinga).
kolweziensis Levasseur, 1965: 60 (Afrorabigus; Type locality: Katanga: Kolwezi).
manikaensis Levasseur, 1967b: 1628 (Afrorabigus; Type locality: région de Kolwezi, Katanga, Manika).
muscicola Levasseur, 1965: 60 (Afrorabigus; Type locality: Katanga: Kolwezi).
robusticornis Bernhauer, 1915d: 137 (Philonthus; Type locality: Deutsch-Ostafrika: Arusha-Ju; Moshi).
— Scheerpeltz, 1933: 1358 (Philonthus; catalog).
— Bernhauer and Chapman, 1939: 68 (Philonthus; Kenya).
— Tottenham, 1949b: 328 (Philonthus; characters).
— Tottenham, 1962: 137 (Philonthus; Tanganyika).
— Levasseur, 1965: 62 (Afrorabigus; characters; Zaire; Kenya; Sudan; Kilimanjaro).
▴ Distribution: Sudan, Kenya, Tanzania, Congo.
toumodiensis Levasseur, 1967b: 1629 (Afrorabigus; Type locality: Côte d'Ivoire, Lamto, Toumodi).
Agacerus
[1 species; Oriental Region]
Agacerus Fauvel, 1895: 245 (replacement name for Eurycerus Fauvel). Type species: Agacerus pectinatus Fauvel, fixed by objective synonymy with Eurycerus.
— Bernhauer and Schubert, 1914: 396 (world catalog; 1 species).
— Cameron, 1925: 71 (catalog of Indian species).
— Cameron, 1932: 255 (characters).
— Scheerpeltz, 1933: 1411 (world catalog supplement).
— Blackwelder, 1952: 40 (type species: pectinatus).
— Scheerpeltz, 1965: 278 (characters; type species: pectinatus).
Eurycerus Fauvel, 1895: 244 (species included: pectinatus; preoccupied by Illiger, 1807, Dejean, 1833, and Kaup, 1844; [Note: Eurycerus was validated on page 244 and replaced on page 245. It might be argued that Eurycerus was originally published as a junior synonym and is therefore an unavailable name (Article 11.6). However, since it was published on one page and synonymized on the next, it can be argued that the name was not originally published as a synonym]). Type species: Agacerus pectinatus Fauvel, fixed by monotypy.
— Bernhauer and Schubert, 1914: 396 (synonym of Agacerus).
— Cameron, 1932: 255 (synonym of Agacerus).
— Blackwelder, 1952: 158 (type species).
pectinatus Fauvel, 1895: 245 (Agacerus; Type locality: Bírmanie, Carin Cheba, 900–1100 m).
Atopocentrum
[1 species; Neotropical]
Atopocentrum Bernhauer, 1906c: 327 (species included: mirabile). Type species: Atopocentrum mirabile Bernhauer, fixed by monotypy.
— Bernhauer and Schubert, 1914: 365 (world catalog; 1 species).
— Blackwelder, 1944: 138 (in checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 69 (type species: mirabile).
mirabile Bernhauer, 1906c: 327 (Atopocentrum; Type locality: Peru: Cushi, 1800 m).
Belonuchus
[196 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Oriental, Australian, and Oceanic Regions]
Belonuchus Nordmann, 1837: 129 (species included: haemorrhoidalis). Type species: Staphylinus haemorrhoidalis Fabricius, fixed by monotypy.
— Erichson, 1840: 419 (characters).
— Lacordaire, 1854: 80 (characters; notes; list of species).
— LeConte, 1861a: 65 (characters).
— LeConte, 1863: 24 (list of North American species).
— Gemminger and Harold, 1868: 595 (catalog).
— Sharp, 1876: 156 (characters; notes).
— Fauvel, 1878f: 269 (characters; notes; key to New Guinea species).
— Duvivier, 1883: 152 (catalog).
— Sharp, 1885: 428 (characters; Mexican and Central American species).
— Blatchley, 1910: 379 (characters).
— Bernhauer and Schubert, 1914: 369 (world catalog; 93 species).
— Casey, 1915: 445 (key to species of USA).
— Bernhauer, 1916g: 265 (characters; notes).
— Leng, 1920: 108 (catalog of North American species).
— Cameron, 1921a: 354, 405 (characters; key to Singapore species; catalog).
— Cameron, 1925: 65 (catalog of Indian species).
— Bruch, 1928a: 428 (catalog; Argentina).
— Scheerpeltz, 1933: 1377 (world catalog supplement).
— Blackwelder, 1943: 420 (characters; type species: haemorrhoidalis; key to species of West Indies).
— Blackwelder, 1944: 136 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 74 (type species: haemorrhoidalis).
— Moore, 1965: 98, 101 (characters; notes).
— Shibata, 1973c: 61 (checklist of species of Taiwan).
— Tikhomirova, 1973a: 186 (checklist of species of USSR).
— Moore and Legner, 1975: 6 (catalog; North America north of Mexico).
— Moore and Legner, 1979: 16 (characters; notes).
— Shibata, 1983: 139 (checklist of species of Japan).
— Hammond, 1984: 205 (checklist of species of Borneo).
— Frank, 1986a: 369 (checklist of species of Florida).
— Smetana, 1995e: 32, 722 (characters; key to North American species; phylogenetic relationships; notes).
— Downie and Arnett, 1996: 383 (characters; key to species of Northeastern North America).
— Lundgren, 1998: 50 (list of species from Florida; records for other States listed for each species).
— Newton, Thayer, Ashe, and Chandler, 2000: 395 (9 Nearctic species; notes; characters in key).
— Hayashi, 2000: 23 (notes on empodial setae).
Trapeziderus Motschulsky, 1860a: 77 (species included: bicolor). Type species: Trapeziderus bicolor Motschulsky, fixed by monotypy.
— Gemminger and Harold, 1868: 599 (catalog).
— Fauvel, 1895: 266 (synonym of Belonuchus).
— Bernhauer and Schubert, 1914: 369 (synonym of Belonuchus).
— Cameron, 1932: 170 (synonym of Belonuchus).
— Blackwelder, 1943: 420 (synonym of Belonuchus; type species: bicolor).
— Blackwelder, 1952: 392 (synonym of Belonuchus; type species: bicolor).
— Smetana, 1995e: 722 (synonym of Belonuchus).
Trapezinotus Motschulsky, 1870: 49 (species included: bicolor; cited with Trapeziderus as junior synonym; [Note: This name was cited as the senior synonym of Trapeziderus. The reason for this action is unclear, but presumably it is a replacement name.
Although Motschulsky used the name Trapezidera in Helotidae in 1860, the names are not homonyms]). Type species: Trapeziderus bicolor Motschulsky, fixed by objective synonymy with Trapeziderus.
Musicoderus Sharp, 1885: 455 (species included: cephalotes, gracilis). Type species: Musicoderus cephalotes Sharp, fixed by subsequent designation by Blackwelder, 1943: 420 [Note: Blackwelder, 1952: 251, stated that R. Lucas, 1920: 425, designated the type species but the designation was ambiguous because two species were cited as “cephalotes et gracilis”].
— Bernhauer and Schubert, 1914: 369 (world catalog; 3 species).
— Scheerpeltz, 1933: 1377 (subgenus of Belonuchus).
— Blackwelder, 1943: 420 (synonym of Belonuchus; type species: cephalotes).
— Blackwelder, 1952: 251 (type species).
— Smetana, 1995e: 722 (synonym of Belonuchus).
abdominalis Cameron, 1937: 20 (Belonuchus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft).
abyssinus Fauvel, 1881, see: Philonthus.
aeneipennis Fauvel, 1895: 268 (Belonuchus; Type locality: Nord-ouest de Sumatra, Tebing-tinggi; Tandjong Morawa, Serdang; Borneo, Sarawak; Malacca; Singapore).
— Fauvel, 1904a: 60 (Belonuchus; Nilghiris; Bírmanie; Malacca; Singapore; Borneo; Sumatra).
— Bernhauer and Schubert, 1914: 369 (Belonuchus; catalog).
— Cameron, 1921a: 378, 405 (Belonuchus; characters; Singapore).
— Scheerpeltz, 1933: 1377 (Belonuchus; catalog).
▴ Distribution: Indonesia, Singapore, Borneo, Malaysia, Myanmar, India.
aeneiventris Fauvel, 1878f: 270 (Belonuchus; Type locality: Nouvelle-Guinée, Andai, Ile Yule; Ternate, Acquiconora).
aequalis Sharp, 1876: 161 (Belonuchus; Type locality: Para, Ega, St. Paulo; Conceicao, Rio Mauhes).
agilis Erichson, 1840: 423 (Belonuchus; Type locality: Cuba).
— Guérin-Méneville, 1857: 37 (Belonuchus; characters).
— Chevrolat and Fauvel, 1863: 433 (Belonuchus; characters; Cuba).
— Fauvel, 1901: 87 (Belonuchus; Jamaica; Haiti; Cuba; Colombia).
— Bernhauer and Schubert, 1914: 369 (Belonuchus; catalog).
— Blackwelder, 1943: 427 (Belonuchus; characters; Cuba; Jamaica).
▴ Distribution: West Indies, Colombia.
chevrolati Fauvel, 1863: 432 (Belonuchus; Type locality: La Havane, Cuba).
— Fauvel, 1869: 489 (Belonuchus; synonym of agilis).
— Fauvel, 1901: 87 (Belonuchus; synonym of agilis).
— Bernhauer and Schubert, 1914: 369 (Belonuchus; synonym of agilis).
— Blackwelder, 1943: 427 (Belonuchus; synonym of agilis).
albovariegatus Bernhauer, 1916g: 276 (Belonuchus; Type locality: West-Columbien: Umgebung Cali am Rio Cauca: St. Antonio, 2000 m; Alto de las cruces, 2200–2400 m).
alternatus Sharp, 1885: 430 (Belonuchus; Type locality: Guatemala, near the City, San Geronimo; Nicaragua, Chontales; Panama, Bugaba).
— Bernhauer and Schubert, 1914: 369 (Belonuchus; catalog).
▴ Distribution: Panama, Nicaragua, Guatemala.
altissimus Bernhauer, 1916g: 275 (Belonuchus; Type locality: West-Columbien: Am Monte Socorro in den West-Cordilleren, 3800 m).
altivagans Bernhauer, 1916g: 272 (Belonuchus; Type locality: West-Columbien: Pacho in den Ost-Cordilleren, 2000 m).
amplipennis Fauvel, 1891: 117 (Belonuchus; Type locality: Colonie Tovar; San Esteban).
amplus Blackwelder, 1943: 425 (Belonuchus; Type locality: St. Lucia: one-half mile northwest of St. Remy on road to Soufriere, 2.5 miles southeast of Soufriere).
analis Schubert, 1911: 29 (Belonuchus; Type locality: Lombok).
andinus Bernhauer, 1916g: 278 (Belonuchus; Type locality: Bolivien: Yuracares).
angusticeps Scheerpeltz, 1960c: 117 (Belonuchus; subgenus Musicoderus; Type locality: Bolivien: Cochabamba, Yungas de Palmar, 2000 m).
angusticollis Bernhauer, 1906c: 330 (Belonuchus; Type locality: Peru: Chanchamayo).
anthracinus Schubert, 1911: 30 (Belonuchus; Type locality: Transvaal, Zoutpansberg, 800 m).
antiguae Blackwelder, 1943: 434 (Belonuchus; Type locality: Antigua: near summit of Boggy Peak, parish of St. Mary, elevation 1300 feet).
aphaobius Smetana, 1995e: 740 (Belonuchus; Type locality: Texas: Austin).
apicicornis Bernhauer, 1916g: 270 (Belonuchus; subgenus Musicoderus; Type locality: West-Columbien: Cali am Rio Cauca: Rio Aquatal, 1800 m; S. Antonio, 2000 m; Rio Vitaco, 2200 m).
apiciventris Sharp, 1885: 419 (Philonthus; Type locality: Mexico, Juquila).
— Bernhauer and Schubert, 1914: 329 (Philonthus; catalog).
— Navarrete-Heredia and Márquez-Luna, 1993 (Philonthus; cave collections; Mexico).
— Navarrete-Heredia, 1995: 84, 89, 100 (Philonthus; cited as apaciventris; characters; habits; natural history notes; host mushrooms; Mexico).
— Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 61 (Belonuchus; Mexico).
▴ Distribution: Mexico.
argentinus Bernhauer, 1916g: 278 (Belonuchus; Type locality: Argentinien: Misiones; Brasilien: Espirito Santo, Rio de Janeiro).
arizonicus Casey, 1915: 448 (Belonuchus; Type locality: Arizona: Benson and Sta. Rita Mts.; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1377 (Belonuchus; catalog).
— Smetana, 1995e: 743 (Belonuchus; lectotype designation: Arizona: Benson and Sta. Rita Mts.; characters; notes; Arizona; California; New Mexico; Texas).
▴ Distribution: USA.
texanus Casey, 1915: 447 (Belonuchus; Type locality: Texas: Brownsville; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1379 (Belonuchus; catalog).
— Smetana, 1995e: 743 (Belonuchus; lectotype designation; synonym of arizonicus).
armatus Sharp, 1876: 163 (Belonuchus; Type locality: Abacaxis).
arrogans Bernhauer, 1916g: 274 (Belonuchus; Type locality: West-Columbien: Ost-Cordilleren, Pacho, 2000 m).
asperatus Bernhauer, 1906c: 336 (Belonuchus; Type locality: Peru: Pachitea).
— Bernhauer and Schubert, 1914: 369 (Belonuchus; catalog).
— Scheerpeltz, 1960c: 120 (Belonuchus; subgenus Musicoderus; Bolivia).
▴ Distribution: Peru, Bolivia.
assamensis Cameron, 1932: 176 (Belonuchus; Type locality: Assam: Naga Hills).
associatus Cameron, 1938b: 150 (Belonuchus; Type locality: Not cited, presumably Java).
morosus Cameron, 1938b: 150 (Belonuchus; variety of associatus; Type locality: E. Java: Res. Kediri, Toempoek).
avidus Erichson, 1840: 423 (Belonuchus; Type locality: Novae Valenciae in Columbia).
— Fauvel, 1891: 117 (Belonuchus; Venezuela).
— Bernhauer and Schubert, 1914: 369 (Belonuchus; catalog).
▴ Distribution: Colombia, Venezuela.
bafutensis Levasseur, 1968: 1381 (Belonuchus; Type locality: Cameroun: forêt de Bafut, Nguemba).
bakeri Bernhauer, 1915a: 25 (Belonuchus; Type locality: Philippinen: Mts. Banahao).
— Scheerpeltz, 1933: 1377 (Belonuchus; catalog).
— Jarrige, 1951: 134 (Belonuchus; Tonkin).
▴ Distribution: Philippines, Vietnam.
barbicornis Bernhauer, 1916g: 271 (Belonuchus; subgenus Musicoderus; Type locality: West-Columbien: Alto de las Cruces, 2200 m; S. Antonio, 2200 m).
basiventris Sharp, 1885: 418 (Philonthus; Type locality: Mexico, Cordova, Puebla Yolos, Oaxaca, Jalapa).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Marquez-Luna and Navarrete Heredia, 1995: 35 (Philonthus; in detritus of Atta mexicana; Mexico).
— Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 61 (Belonuchus; necrophilous species; distribution in Mexico).
▴ Distribution: Mexico.
batesi Sharp, 1876: 157 (Belonuchus; Type locality: Ega).
bicolor Motschulsky, 1860, see: grandiceps Kraatz, 1859.
bicoloratus Schubert, 1911: 30 (Belonuchus; Type locality: Lombok).
bicoloripennis Bernhauer, 1928g: 121 (Belonuchus; Type locality: Belg.-Kongo: Haute Uele: Moto).
bidens Sharp, 1885: 430 (Belonuchus; Type locality: Mexico, Jalapa, Oaxaca, Mirador, Tuxtla; Nicaragua, Chontales).
biformis Cameron, 1951b: 127 (Belonuchus; Type locality: Nigeria: Jos, Plateau Bauchi).
rufopygus Cameron, 1951b: 128 (Belonuchus; variety of biformis; Type locality: not stated but possibly Nigeria: Jos, Plateau Bauchi).
bipunctatus Bernhauer, 1935c: 90 (Belonuchus; Type locality: Kolumbien [ohne nähere Fundortangabe]; Cali, Villa Elvira).
birmanus Cameron, 1932: 174 (Belonuchus; Type locality: Burma: Ruby Mines).
boops Sharp, 1885: 435 (Belonuchus; Type locality: Panama, Bugaba).
borneensis Cameron, 1933b: 346 (Belonuchus; Type locality: British North Borneo: Kenokok, 3300 feet).
boxi Bernhauer, 1934b: 212 (Belonuchus; Type locality: Argentinien: Tucuman).
brasilianus Bernhauer, 1917c: 19 (Belonuchus; Type locality: Brasilien: S. Catharina).
breviceps Fauvel, 1891: 117 (Belonuchus; Type locality: Colonie Tovar).
brevicollis Fauvel, 1877: 267 (Belonuchus; Type locality: Australie orientale, Queensland, Wide Bay).
— Bernhauer and Schubert, 1914: 369 (Belonuchus; catalog).
— Lea, 1925: 228 (Belonuchus; characters; Australia).
▴ Distribution: Australia.
bruchi Bernhauer, 1912e: 175 (Belonuchus; Type locality: Argentinien: Prov. Buenos Aires).
bruchi Bernhauer, 1934, see: duplicatus Blackwelder, 1944.
bruchianus Bernhauer, 1911a: 413 (Belonuchus; Type locality: Argentinien, Gob. Misiones).
bugnioni Fauvel, 1901: 86 (Belonuchus; Type locality: Martinique).
— Bernhauer and Schubert, 1914: 369 (Belonuchus; catalog).
— Blackwelder, 1943: 428 (Belonuchus; characters; Guadeloupe; Martinique).
▴ Distribution: West Indies.
eximius Bernhauer, 1917i: 105 (Belonuchus; Type locality: Guadeloupe).
— Scheerpeltz, 1933: 1378 (Belonuchus; catalog).
— Blackwelder, 1943: 428 (Belonuchus; synonym of bugnioni).
cacao Blackwelder, 1943: 434 (Belonuchus; Type locality: Trinidad: one-half mile south-southwest of Manzanilla junction on the Plum Road).
candens Erichson, 1840: 460 (Philonthus; Type locality: Bogotae, Columbia).
— Guérin-Méneville, 1855: 583 (Philonthus; Brazil).
— Bernhauer and Schubert, 1914: 369 (Belonuchus; catalog).
▴ Distribution: Colombia, Brazil.
cariniventris Bernhauer, 1917k: 225 (Belonuchus; Type locality: Mexiko: Jalappa).
cavipennis Bernhauer, 1916g: 267 (Belonuchus; subgenus Musicoderus; Type locality: Bolivien: Yuracares).
cephalotes Sharp, 1885: 455 (Musicoderus; Type locality: Panama, Bugaba 800 to 1500 feet).
— Bernhauer and Schubert, 1914: 368 (Musicoderus; catalog).
— Scheerpeltz, 1933: 1377 (Belonuchus; catalog).
▴ Distribution: Panama.
ceylonicus Cameron, 1932: 176 (Belonuchus; Type locality: Ceylon: Kotte).
— Scheerpeltz, 1933: 1377 (Belonuchus; catalog).
— Legner and Moore, 1977: 176 (Belonuchus; found under bark).
▴ Distribution: Sri Lanka.
chevrolati Fauvel, 1863, see: agilis Erichson, 1840.
clypeatus Sharp, 1876: 160 (Belonuchus; Type locality: Ega).
coelestinus Bernhauer, 1908d: 330 (Belonuchus; Type locality: St. Vincent).
— Bernhauer and Schubert, 1914: 369 (Belonuchus; catalog).
— Blackwelder, 1943: 426 (Belonuchus; characters; St. Vincent).
▴ Distribution: West Indies.
coeruleus Cameron, 1922: 122 (Belonuchus; Type locality: St. Vincent).
— Scheerpeltz, 1933: 1377 (Belonuchus; catalog).
— Blackwelder, 1943: 426 (Belonuchus; synonym of coelestinus).
coelestis Bernhauer, 1916g: 266 (Belonuchus; subgenus Musicoderus; Type locality: Bolivien: Yuracares).
coeruleispennis Motschulsky, 1858: 668 (Belonuchus; Type locality: Brésil intérieur).
— Motschulsky, 1870: 49 (Belonuchus; cited as coeruleipennis).
— Bernhauer and Schubert, 1914: 369 (Belonuchus; cited as coeruleipennis; catalog).
▴ Distribution: Brazil.
coeruleus Cameron, 1922, see: coelestinus Bernhauer, 1908.
cognatus Sharp, 1885: 429 (Belonuchus; Type locality: British Honduras, Belize; Guatemala, El Reposo, Las Mercedes, San Isidro, Mirandilla; Nicaragua, Chontales; Panama, Bugaba).
— Bernhauer and Schubert, 1914: 369 (Belonuchus; catalog).
— Scheerpeltz, 1933: 1377 (Belonuchus; catalog; Argentina).
— Blackwelder, 1943: 427 (Belonuchus; characters; St. Vincent; Trinidad; Colombia; Venezuela).
— Frank, 1982: 34 (Belonuchus; list of parasites).
▴ Distribution: West Indies, Belize, Guatemala, Nicaragua, Panama, Colombia, Venezuela, Argentina.
coloratus Last, 1989: 882 (Belonuchus; Type locality: Onerunka, near Kainantu).
columbicus Bernhauer, 1916g: 271 (Belonuchus; subgenus Musicoderus; Type locality: West-Columbien: Umgebung von Cali, St. Antonio, 2000 m).
connexus Say, 1830: 35 [= 1834: 448] (Staphylinus; Type locality: Mexico).
— Erichson, 1840: 927 (Belonuchus; characters; Mexico).
— Sharp, 1885: 433 (Belonuchus; Mexico).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog).
▴ Distribution: Mexico.
convexus Bernhauer, 1908d: 331 (Musicoderus; Type locality: Peru: Pozuzo).
— Bernhauer and Schubert, 1914: 368 (Musicoderus; catalog).
— Scheerpeltz, 1933: 1377 (Belonuchus; catalog; Nicaragua).
▴ Distribution: Peru, Nicaragua.
cordilleranus Bernhauer, 1916g: 270 (Belonuchus; subgenus Musicoderus; Type locality: West-Columbien: Cali am Rio Cauca: Rio Aquatal, 1800 m; S. Antonio, 2000 m; Rio Vitaco, 2200 m).
cribriceps Bernhauer, 1921c: 172 (Belonuchus; Type locality: Corrientes: San Roque).
danforthi Blackwelder, 1943: 429 (Belonuchus; Type locality: Puerto Rico: about 13 miles east of Maricao and north of Yauco, elevation about 3000 feet).
decipiens Sharp, 1876: 159 (Belonuchus; Type locality: Ega).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog).
— Scheerpeltz, 1933: 1378 (Belonuchus; catalog).
— Silvestri, 1945: 325 (Belonuchus; characters; Brazil).
▴ Distribution: Brazil.
dejectus Sharp, 1885, see: viridipennis Baudi, 1848.
dichrous Erichson, 1840: 420 (Belonuchus; Type locality: Orizaba, Mexico).
— Solsky, 1868a: 137 (Belonuchus; Mexico).
— Sharp, 1885: 429 (Belonuchus; notes; Mexico; Guatemala).
— Fauvel, 1891: 116 (Belonuchus; Venezuela).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog).
▴ Distribution: Mexico, Guatemala, Venezuela.
diversicornis Bernhauer, 1917i: 104 (Belonuchus; Type locality: Brasilien: Espirito Santo).
diversus Bernhauer, 1928g: 121 (Belonuchus; Type locality: Belg.-Kongo: Haute Uele: Moto).
docilis Sharp, 1885: 435 (Belonuchus; Type locality: Guatemala, Panteleon; Panama, Bugaba, Tolé).
dohrni Fauvel, 1877, see: Sternotoxus.
dominicus Blackwelder, 1943: 431 (Belonuchus; Type locality: Dominica: Hillsboro Estate, at mouth of the Layou River, Leeward side).
duplicatus Blackwelder, 1944: 137 (Belonuchus; replacement name for bruchi Bernhauer).
bruchi Bernhauer, 1934b: 213 (Belonuchus; [preoccupied]; Type locality: Argentinien: Prov. Tucuman).
ecuadorensis Bernhauer, 1935c: 91 (Belonuchus; Type locality: Ecuador).
ephippiatus Say, 1830: 35 [= 1834: 448] (Staphylinus; Type locality: [Note: See neotype designation by Smetana, 1995e]. Originally cited from Mexico).
— Erichson, 1840: 927 (Belonuchus; characters; Mexico).
— LeConte, 1859: 563 (Belonuchus; California).
— Fauvel, 1901: 86 (Belonuchus; Colombia; California; Arizona; New Mexico; Mexico; Venezuela).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog).
— Nishida, 1994: 70 (Belonuchus; Hawaii).
— Smetana, 1995e: 746 (Belonuchus; neotype designation: [Note: Locality not cited; specimen selected from LeConte collection with “golden round disc”]; characters; notes; Arizona; California; New Mexico; Texas).
— Nishida, 1997: 60 (Belonuchus; Hawaii).
▴ Distribution: Venezuela, Colombia, Mexico, USA, Hawaii.
simplex Sharp, 1885: 432 (Belonuchus; Type locality: Mexico, Oaxaca; [Note: See lectotype designation by Smetana, 1995e]. Also cited from Mexico, Vera Cruz).
— Sharp, 1887: 789 (Belonuchus; Mexico).
— Fauvel, 1901: 86 (Belonuchus; synonym of ephippiatus).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; synonym of ephippiatus).
— Smetana, 1995e: 746 (Belonuchus; lectotype designation: Oxaca, Mexico; synonym of ephippiatus).
punctiventris Casey, 1915: 446 (Belonuchus; Type locality: Arizona: Benson; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1379 (Belonuchus; catalog).
— Smetana, 1995e: 746 (Belonuchus; lectotype designation; synonym of ephippiatus).
laticeps Casey, 1915: 446 (Belonuchus; Type locality: California: near Indio; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1378 (Belonuchus; catalog).
— Smetana, 1995e: 746 (Belonuchus; lectotype designation; synonym of ephippiatus).
jacobianus Casey, 1915: 447 (Belonuchus; Type locality: California: San Diego; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1378 (Belonuchus; catalog).
— Smetana, 1995e: 746 (Belonuchus; lectotype designation; synonym of ephippiatus).
schaefferi Cooper, 1933a: 545 (Belonuchus; Type locality: Texas: Brownsville).
erichsoni Bernhauer, 1917k: 223 (Belonuchus; Type locality: Mexiko).
erythropterus Solsky, 1868a: 137 (Belonuchus; Type locality: Mexique).
— Sharp, 1885: 432 (Belonuchus; characters; Mexico; Venezuela).
— Sharp, 1887: 789 (Belonuchus; Mexico).
— Fauvel, 1901: 86 (Belonuchus; synonym of ephippiatus).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; synonym of ephippiatus).
— Smetana, 1995e: 751, 752 (Belonuchus; valid species; characters; notes; Texas; Mexico).
▴ Distribution: Mexico, USA.
excelsus Bernhauer, 1917c: 19 (Belonuchus; Type locality: West-Columbien: Umgebung von Cali: St. Antonio, 2000 m).
eximius Bernhauer, 1917, see: bugnioni Fauvel, 1901.
falsus Bernhauer, 1917i: 105 (Belonuchus; Type locality: Venezuela).
fassli Bernhauer, 1917d: 37 (Belonuchus; Type locality: West-Columbien: Umgebung von Cali am Rio Cauca, Cañon del Mt. Tolima, 1700 m).
ferrugatus Erichson, 1840: 931 (Philonthus; Type locality: Java).
— Kraatz, 1859: 96 (Philonthus; characters; Java).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog).
— Scheerpeltz, 1933: 1378 (Belonuchus; catalog).
— Cameron, 1933d: 391 (Belonuchus; Sumatra).
— Cameron, 1937: 19 (Belonuchus; Java).
▴ Distribution: Indonesia, Philippines.
rufotestaceus Cameron, 1941c: 381 (Belonuchus; variety of ferrugatus; Type locality: Philippines: Luzon: Los Banos).
flavipennis Solsky, 1870a: 263 (Belonuchus; Type locality: Mexique).
— Sharp, 1885: 430 (Belonuchus; characters; Mexico).
— Sharp, 1887: 789 (Belonuchus; Mexico).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog).
▴ Distribution: Mexico.
flavocaudatus Bernhauer, 1921c: 170 (Belonuchus; Type locality: Argentinien: Prov. Jujuy, Hornadita, 3400 m).
formosus Gravenhorst, 1806, see: rufipennis Fabricius, 1801.
forsteri Scheerpeltz, 1960c: 115 (Belonuchus; subgenus Musicoderus; Type locality: Bolivien: Yungas de Arepucho, Sihuencas, 2200–2500 m).
fossulatus Wasmann, 1887: 414 (Belonuchus; Type locality: Blumenau, Prov. St. Catharina, Brasil).
frater Bernhauer, 1906c: 333 (Belonuchus; Type locality: Peru).
fuscipes Fauvel, 1878f: 270 (Belonuchus; Type locality: Nouvelle-Guinée, Andai; Hatam, Salvatti; Iles Arou).
— Fauvel, 1879: 107 (Belonuchus; Dorei; Celebes).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog).
— Cameron, 1931: 363 (Belonuchus; New Guinea).
— Frank, 1982: 34 (Belonuchus; list of parasites).
▴ Distribution: New Guinea, Indonesia.
gagates Erichson, 1840: 424 (Belonuchus; Type locality: Cuba; [Note: See lectotype designation by Smetana, 1995e]. Also cited from Puerto Rico).
— Guérin-Méneville, 1857: 38 (Belonuchus; characters; Puerto Rico; Cuba).
— Chevrolat and Fauvel, 1863: 433 (Belonuchus; characters; Cuba, Santo Domingo, Puerto Rico).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog).
— Scheerpeltz, 1933: 1379 (Belonuchus; catalog; Jamaica).
— Blackwelder, 1943: 424 (Belonuchus; characters; Bahamas Islands; Cuba; Jamaica; Hispaniola; Puerto Rico; st. Thomas; St. John; Montserrat).
— Smetana, 1995e: 737 (Belonuchus; lectotype designation: Cuba; characters; notes; Florida).
▴ Distribution: Indonesia, USA.
minax Erichson, 1840: 424 (Belonuchus; Type locality: Americae insulae St. Thoma et St. Johannis).
— Bernhauer and Schubert, 1914: 371 (Belonuchus; catalog).
— Blackwelder, 1943: 424 (Belonuchus; cited as minar; synonym of gagates).
nigrocaeruleus Cameron, 1913a: 175 (Musicoderus; Type locality: Jamaica: Newcastle district, 3000 feet).
— Blackwelder, 1943: 424 (Belonuchus; synonym of gagates).
— Smetana, 1995e: 737 (Belonuchus; synonym of gagates).
gowdeyi Bernhauer, 1935c: 91 (Belonuchus; Type locality: Jamaica: Hope Gardens).
— Blackwelder, 1943: 424 (Belonuchus; synonym of gagates).
— Smetana, 1995e: 737 (Belonuchus; synonym of gagates).
ganglbaueri Bernhauer, 1906c: 331 (Belonuchus; Type locality: Bolivia).
gardneri Cameron, 1932: 173 (Belonuchus; Type locality: Bengal: Rangirum, alt. 6000 feet, near Darjeeling).
— Scheerpeltz, 1933: 1378 (Belonuchus; catalog).
— Legner and Moore, 1977: 176 (Belonuchus; cited as gardeneri; found under bark).
▴ Distribution: India.
gedyei Cameron, 1951c: 403 (Belonuchus; Type locality: Kenya Colony: Mutha district, 4000 feet).
gowdeyi Bernhauer, 1935, see: gagates Erichson, 1839.
gracilis Sharp, 1885: 456 (Musicoderus; Type locality: Nicaragua, Chontales).
— Bernhauer and Schubert, 1914: 368 (Musicoderus; catalog).
— Scheerpeltz, 1933: 1378 (Belonuchus; catalog).
▴ Distribution: Nicaragua.
grandiceps Sharp, 1876, see: sharpi Bernhauer, 1907.
grandiceps Kraatz, 1859: 95 (Philonthus; Type locality: Ceylan).
— Fauvel, 1902a: 41 (Belonuchus).
— Bernhauer, 1902b: 28 (Philonthus; Ceylon).
— Fauvel, 1903: 159 (Belonuchus; Ceylon).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog).
— Bernhauer, 1928: 17 (Belonuchus; Sumatra).
— Cameron, 1932: 171 (Belonuchus; characters; Ceylon; India).
— Scheerpeltz, 1933: 1378 (Belonuchus; catalog).
— Cameron, 1933d: 391 (Belonuchus; Sumatra).
▴ Distribution: India, Sri Lanka, Indonesia.
bicolor Motschulsky, 1860a: 79 (Trapeziderus; Type locality: montagnes de Noura-Ellia à Ceylan).
— Fauvel, 1903: 159 (Belonuchus; synonym of grandiceps).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; synonym of grandiceps).
— Cameron, 1932: 171 (Belonuchus; synonym of grandiceps).
thoracicus Fauvel, 1903: 159 (Belonuchus; variety of grandiceps; Type locality: Iles Mentawei).
grandis Bernhauer, 1928g: 120 (Belonuchus; Type locality: Hervorkommen).
haemorrhoidalis Fabricius, 1801: 596 (Staphylinus; [preoccupied]; Type locality: America meridionalis).
— Nordmann, 1837: 130 (Belonuchus; cited as haemoroidalis; characters; Brazil).
— Erichson, 1840: 420 (Belonuchus; characters; Brazil; Colombia).
— Solsky, 1872: 303 (Belonuchus; Cayenne).
— Sharp, 1876: 157 (Belonuchus; Brazil).
— Fauvel, 1891: 116 (Belonuchus; [Note: Misidentification: Bernhauer and Schubert, 1914: 370, cited cognatus Fauvel as a synonym of haemorrhoidalis; Fauvel did not describe the species as new, he attributed it to Sharp]; Venezuela).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog).
— Scheerpeltz, 1933: 1378 (Belonuchus; catalog).
— Frank, 1982: 34 (Belonuchus; list of parasites).
— Herman, 2001: 42 (Belonuchus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Brazil, French Guiana, Venezuela, Colombia.
hispaniolus Blackwelder, 1943: 422 (Belonuchus; Type locality: Dominican Republic: San Francisco Mountains).
holisinus Fauvel, 1904, see: Gabrius.
holisoides Sharp, 1876: 161 (Belonuchus; Type locality: Ega).
imitator Cameron, 1932: 176 (Belonuchus; Type locality: Burma: Schwego-myo).
imperialis Cameron, 1946: 11 (Belonuchus; Type locality: Philippines: Mt. Banahao).
impressifrons Sharp, 1876: 162 (Belonuchus; Type locality: Ega).
iridescens Solsky, 1868a: 140 (Belonuchus; Type locality: Mexique).
irregularis Bernhauer, 1916g: 275 (Belonuchus; Type locality: West-Columbien: Pacho in den Ost-Cordilleren, 2000 m).
jacobianus Casey, 1915, see: ephippiatus Say, 1830.
jalappensis Bernhauer, 1917k: 224 (Belonuchus; Type locality: Mexiko: Jalappa).
javanus Cameron, 1938b: 150 (Belonuchus; Type locality: E. Java: Res. Kediri: Besoeki, Samberbajem).
jelskii Solsky, 1872: 296 (Philonthus; Type locality: Puna, Lima).
kedirianus Cameron, 1938b: 149 (Belonuchus; Type locality: E. Java: Res. Kediri, Toempoek).
laevipennis Cameron, 1942: 25 (Belonuchus; Type locality: Kostarika: Cartago, Pino, 1000 m).
laticeps Casey, 1915, see: ephippiatus Say, 1830.
latro Erichson, 1840: 422 (Belonuchus; Type locality: Not cited).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog; locality cited as South America but the basis for this information is unknown to me).
▴ Distribution: South America ?.
limbatus Fauvel, 1878f: 271 (Belonuchus; Type locality: Nouvelle-Guinée, Andai).
lividipes Fauvel, 1878f: 271 (Belonuchus; Type locality: Nouvelle-Guinée, Andai).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog).
— Cameron, 1931: 363 (Belonuchus; New Guinea).
▴ Distribution: New Guinea.
longiceps Bernhauer, 1906c: 332 (Belonuchus; Type locality: Peru).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog).
— Scheerpeltz, 1933: 1378 (Belonuchus; catalog).
▴ Distribution: Peru.
maculipennis Fauvel, 1895: 267 (Belonuchus; Type locality: Bírmanie, Carin Cheba, 900–1100 m).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog).
— Cameron, 1932: 174 (Belonuchus; characters; Burma).
— Scheerpeltz, 1933: 1378 (Belonuchus; catalog).
▴ Distribution: Myanmar.
magnificus Bernhauer, 1916g: 277 (Belonuchus; Type locality: West-Columbien: Umgebung von Cali: S. Antonio, 2000 m).
marginicollis Lea, 1931: 374 (Belonuchus; Type locality: New Guinea: Komba).
meridionalis Bernhauer, 1916g: 269 (Belonuchus; subgenus Musicoderus; Type locality: Bolivien: Chaco).
mexicanus Solsky, 1864: 440 (Belonuchus; Type locality: Mexique).
— Sharp, 1885: 436 (Belonuchus; Mexico).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog).
— Scheerpeltz, 1933: 1378 (Belonuchus; catalog; Argentina).
— Silvestri, 1945: 324 (Belonuchus; characters; Brazil).
▴ Distribution: Mexico, Brazil, Argentina.
militaris Bernhauer, 1921c: 170 (Belonuchus; Type locality: Brasilien: São Paulo).
mimeticus Sharp, 1885: 436 (Belonuchus; Type locality: Panama, Bugaba 800 to 1500 feet).
minax Erichson, 1840, see: gagates Erichson, 1840.
miripennis Bernhauer, 1916g: 267 (Belonuchus; subgenus Musicoderus; Type locality: Columbien: Umgebung von Cali am Cauca, Cañon del Mte. Tolima, 1700 m).
modestus Fauvel, 1891: 118 (Belonuchus; Type locality: Colonie Tovar).
montanellus Bernhauer, 1916g: 272 (Belonuchus; Type locality: columbische Cordilleren: Cañon del Mte. Tolima, 1700 m; Alto de las cruces, 2200 m, in der Umgebung von Cali am Cauca).
montanicus Blackwelder, 1944: 137 (Belonuchus; replacement name for montanus Bernhauer).
montanus Bernhauer, 1916g: 273 (Belonuchus; [preoccupied]; Type locality: West-Columbien: Cañon del Mte. Tolima, 1700 m; Rio Vitaco, 2200 m; Villa Elvira, 1800 m).
— Scheerpeltz, 1933: 1378 (Belonuchus; catalog).
— Blackwelder, 1944: 137 (Belonuchus; synonym of montanicus).
montanus Bernhauer, 1916, see: montanicus Blackwelder, 1944.
montanus Fauvel, 1895, see: rufoniger Fauvel, 1895.
monticola Bernhauer, 1917c: 18 (Belonuchus; Type locality: West-Columbien: Umgebung von Cali am Rio Cauca: Rio Vitaco, 2200 m).
moquinus Casey, 1885: 125 (Belonuchus; Type locality: Arizona, Morrison; [Note: See lectotype designation by Smetana, 1995e]).
— Horn, 1885a: 109 (Belonuchus; synonym of xanthomelas).
— Bernhauer and Schubert, 1914: 372 (Belonuchus; synonym of xanthomelas).
— Casey, 1915: 447 (Belonuchus; characters; Arizona).
— Scheerpeltz, 1933: 1378 (Belonuchus; catalog).
— Smetana, 1995e: 733 (Belonuchus; lectotype designation; characters; notes; Arizona; California; Texas).
▴ Distribution: USA.
quadrifer Casey, 1915: 448 (Belonuchus; Type locality: Arizona: Prescott; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1379 (Belonuchus; catalog).
— Smetana, 1995e: 733 (Belonuchus; lectotype designation; synonym of moquinus).
mordens Erichson, 1840: 422 (Belonuchus; Type locality: Brasilia).
— Guérin-Méneville, 1855: 583 (Belonuchus; Brazil).
— Bernhauer and Schubert, 1914: 371 (Belonuchus; catalog).
— Scheerpeltz, 1933: 1379 (Belonuchus; catalog; Argentina).
▴ Distribution: Brazil, Argentina.
moritzi Bernhauer, 1906c: 331 (Belonuchus; Type locality: Venezuela).
morosus Cameron, 1938, see: associatus Cameron, 1938.
motoensis Cameron, 1929: 65 (Belonuchus; Type locality: Haut Uele: Moto).
mundus Erichson, 1840: 425 (Belonuchus; Type locality: Americae meridionalis ins. St. Vincentii).
— Bernhauer and Schubert, 1914: 371 (Belonuchus; catalog).
— Blackwelder, 1943: 430 (Belonuchus; characters; St. Lucia; St. Vincent; Grenada).
▴ Distribution: West Indies.
mutator Fauvel, 1879: 106 (Belonuchus; Type locality: Gilolo; Célèbes; Malacca; Pulo Pinang).
— Bernhauer and Schubert, 1914: 371 (Belonuchus; catalog).
— Bernhauer, 1928: 17 (Belonuchus; Sumatra).
— Scheerpeltz, 1933: 1379 (Belonuchus; catalog; Singapore).
— Cameron, 1933d: 391 (Belonuchus; Sumatra).
— Cameron, 1937: 20 (Belonuchus; Java).
▴ Distribution: Sri Lanka, Indonesia, Singapore.
nalandensis Bernhauer, 1902b: 30 (Belonuchus; Type locality: Nalanda).
— Bernhauer and Schubert, 1914: 371 (Belonuchus; catalog).
— Bernhauer, 1922a: 151 (Belonuchus; Indochina).
— Cameron, 1932: 173 (Belonuchus; characters; Ceylon).
— Scheerpeltz, 1933: 1379 (Belonuchus; catalog).
▴ Distribution: Sri Lanka, “Indochina”.
nigerrimus Bernhauer, 1917k: 225 (Belonuchus; Type locality: Mexiko: Jalappa).
nigripennis Cameron, 1928, see: quadratus Kraatz, 1859.
nigripes Bernhauer, 1922, see: rufoniger Fauvel, 1895.
nigrocaeruleus Cameron, 1913, see: gagates Erichson, 1839.
nigrorufus Fauvel, 1907: 38 (Belonuchus; Type locality: Kilimandjaro: Kibonoto).
nilgiriensis Cameron, 1932: 175 (Belonuchus; Type locality: Nilgiri Hills).
nullicedo Dohrn, 1892: 74 (Belonuchus; Type locality: Philippinen).
oakleyi Blackwelder, 1943: 422 (Belonuchus; Type locality: Puerto Rico: Mayaguez).
obscuricornis Bernhauer, 1936a: 23 (Belonuchus; Type locality: Lulua: Kapanga).
oxyporinus Sharp, 1885: 419 (Philonthus; Type locality: Mexico, Oaxaca).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Frank, 1982: 42 (Philonthus; list of parasites).
— Navarrete-Heredia, 1995: 86, 89, 100 (Philonthus; characters; habits; natural history notes; host mushrooms; Mexico).
— Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 61 (Belonuchus; Mexico).
▴ Distribution: Mexico.
pallidus Casey, 1915: 449 (Belonuchus; Type locality: Florida: New River; [Note: See lectotype designation by Smetana, 1995e]. Also cited from Florida: Palm Beach).
— Scheerpeltz, 1933: 1379 (Belonuchus; catalog).
— Smetana, 1995e: 735 (Belonuchus; lectotype designation: New River, Florida; characters; notes; Florida; Louisiana).
▴ Distribution: USA.
pallipes Melsheimer, 1844, see: rufipennis Fabricius, 1801.
palustris Bernhauer, 1916g: 268 (Belonuchus; subgenus Musicoderus; Type locality: columbische Westcordilleren, Alto de las cruces, am Kamm des Gebirgszuges in 2200 m).
panamensis Sharp, 1885: 434 (Belonuchus; Type locality: Nicaragua, Chontales; Panama, Volcan de Chiriqui, Bugaba, Tolé).
paradoxipunctatus Bernhauer, 1921c: 171 (Belonuchus; Type locality: Corrientes: San Roque).
paradoxus Bernhauer, 1906c: 333 (Belonuchus; Type locality: Peru: Pachitea).
parcus Bernhauer, 1928g: 119 (Belonuchus; Type locality: Belg.-Kongo: Stanleyville a Kilo; Haute-Uele: Moto; Haute-Uele: Watsa; Ituri: Madyu).
— Scheerpeltz, 1933: 1379 (Belonuchus; catalog).
— Scheerpeltz, 1971c: 170 (Belonuchus; Cameroon).
▴ Distribution: Congo, Cameroon.
pectinipes Bernhauer, 1910b: 379 (Belonuchus; Type locality: Mexiko: Cordoba).
penetrans Silvestri, 1946a: 522 (Belonuchus; Type locality: Brasilia: Campo Grande, Matto Grosso).
pexus Motschulsky, 1858, see: Xanthopygus.
piceolus Bernhauer, 1935c: 90 (Belonuchus; Type locality: Brasilien).
picticollis Cameron, 1937: 20 (Belonuchus; Type locality: Java: G. Boerengrang, Preanger, alt. 1600 m; G. Tangkoeban Prahoe, alt. 4000–5000 ft).
pictipennis Sharp, 1885: 436 (Belonuchus; Type locality: Panama, Bugaba).
planiceps Erichson, 1840: 425 (Belonuchus; Type locality: Columbia).
— Fauvel, 1891: 118 (Belonuchus; Venezuela).
— Bernhauer and Schubert, 1914: 371 (Belonuchus; catalog).
▴ Distribution: Colombia, Venezuela.
pollens Sharp, 1885: 431 (Belonuchus; Type locality: North America; Mexico, Oaxaca; Guatemala, near the city; Honduras).
— Bernhauer and Schubert, 1914: 371 (Belonuchus; catalog).
— Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 61 (Belonuchus; necrophilous species; distribution in Mexico).
▴ Distribution: “North America”, Mexico, Guatemala, Honduras.
posticalis Laporte, 1840: 176 (Staphylinus; Type locality: Brésil).
prasinipennis Bernhauer, 1917c: 18 (Belonuchus; Type locality: Bolivien: Yuracares).
pulchripennis Bernhauer, 1908d: 330 (Belonuchus; Type locality: Peruvia centralis: Montana de Nancho, 7000 ft.).
punctiventris Casey, 1915, see: ephippiatus Say, 1830.
punctus Schubert, 1911: 32 (Belonuchus; Type locality: Zagnando, Dahomey).
purpuripennis Bernhauer, 1916g: 278 (Belonuchus; Type locality: West-Columbien: Umgebung von Cali im Caucatal: St. Antonio, 2200 m; Bolivien: Yuracares).
quadratus Kraatz, 1859: 96 (Philonthus; Type locality: India borealis).
— Bernhauer and Schubert, 1914: 371 (Belonuchus; catalog).
— Cameron, 1932: 172 (Belonuchus; characters; India).
— Scheerpeltz, 1933: 1379 (Belonuchus; catalog).
— Cameron, 1937: 19 (Belonuchus; Java).
— Scheerpeltz, 1965: 108 (Belonuchus; Burma).
▴ Distribution: India, Myanmar, Vietnam, Singapore, Indonesia.
nigripennis Cameron, 1928a: 38 (Belonuchus; variety of quadratus; Type locality: Tonkin. Also cited from Singpore and Celebes).
— Cameron, 1921a: 378, 405 (Belonuchus; [Note: Misidentified as mutator according to Cameron, 1928a: 38]; characters; Singapore).
— Scheerpeltz, 1933: 1379 (Belonuchus; variety of quadratus).
quadriceps Fauvel, 1904d: 287 (Belonuchus; Type locality: Usambara: Derema. Zanzibar. Natal).
— Bernhauer and Schubert, 1914: 371 (Belonuchus; catalog).
— Scheerpeltz, 1933: 1379 (Belonuchus; catalog).
▴ Distribution: Tanzania, South Africa.
quadrifer Casey, 1915, see: moquinus Casey, 1885.
richteri Bernhauer, 1922: 15 (Belonuchus; Type locality: Argentinien: Misiones).
ruficeps Cameron, 1925a: 187 (Belonuchus; variety of ferrugatus; Type locality: Tjibodas, 1500 m).
— Scheerpeltz, 1933: 1378 (Belonuchus; variety of ferrugatus).
— Cameron, 1937: 19 (Belonuchus; cited as a valid species; Java).
▴ Distribution: Indonesia.
rufipennis Fabricius, 1801: 597 (Staphylinus; Type locality: America meridionalis).
— Erichson, 1840: 426 (Belonuchus; synonym of formosus).
— Fauvel, 1877a: xxviii (Belonuchus; Brazil).
— Fauvel, 1877a: xxviii (Belonuchus; Brazil).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; synonym of formosus).
— Blackwelder, 1943: 433 (Belonuchus; characters; Tobago; Trinidad).
— Coiffait, 1981g: 289 (Belonuchus; Galapagos Islands).
— Frank, 1982: 34 (Belonuchus; list of parasites).
— Hunter, Kincher, Bay, Beerwinkle, 1991: 168 (Belonuchus; seasonal distribution and diel flight activity; Texas).
— Campbell and Davies, 1991: 119 (Belonuchus; checklist of species; Canada).
— Marquez-Luna and Navarrete Heredia, 1995: 35 (Belonuchus; in detritus of Atta mexicana; Mexico).
— Smetana, 1995e: 726 (Belonuchus; characters; notes; Ontario; Quebec; Alabama; Arkansas; Colorado; Connecticut; Delaware; Florida; Georgia; Illinois; Indiana; Kansas; Kentucky; Louisiana; Maryland; Massachusetts; Michigan; Mississippi; Missouri; Nebraska; New Jersey; New Mexico; New York; North Carolina; Ohio; Oklahoma; Pennsylvania; Tennessee; Texas; Virginia; West Virginia; also Jamaica and El Salvador).
— Downie and Arnett, 1996: 383 (Belonuchus; characters; USA).
— Schmidt, 1999: 108 (Belonuchus; food for adults and larvae used for rearing species; rearing methods).
▴ Distribution: Hawaii, Galapagos Islands, Argentina, Brazil, French Guiana, Venezuela, Colombia, Panama, Nicaragua, El Salvador, Guatemala, Mexico, West Indies, USA, Canada.
formosus Gravenhorst, 1806: 72 (Staphylinus; Type locality: Carolina; [Note: See lectotype designation by Smetana, 1995e]).
— Erichson, 1840: 426 (Belonuchus; characters; Colombia; Cayenne; Carolina).
— Solsky, 1875: 15 (Belonuchus; Peru).
— Sharp, 1876: 160 (Belonuchus; Brazil).
— Fauvel, 1877a: xxviii (Belonuchus; synonym of rufipennis).
— Sharp, 1885: 431 (Belonuchus; characters; Carolina; Pennsylvania; Mexico; British Honduras; Guatemala; Nicaragua; Panama; Colombia; Brazil).
— Fauvel, 1891: 118 (Belonuchus; Venezuela).
— Fauvel, 1901: 87 (Belonuchus; Venezuela; eastern United States to Brazil).
— Fall and Cockerell, 1907: 166 (Belonuchus; New Mexico).
— Blatchley, 1910: 379 (Belonuchus; characters; Indiana).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog).
— Notman, 1920b: 706 (Belonuchus; Florida).
— Scheerpeltz, 1933: 1378 (Belonuchus; catalog; Argentina).
— Blackwelder, 1943: 432 (Belonuchus; synonym of rufipennis).
— Silvestri, 1945: 313 (Belonuchus; characters of adults, first instar and mature larvae, and pupa; life history notes).
— Silvestri, 1945a (Belonuchus; anomalous adult).
— Scheerpeltz, 1975a: 243 (Belonuchus; collecting notes; Brazil).
— Legner and Moore, 1977: 176 (Belonuchus; found under bark).
— Marcuzzi, 1977: 13 (Belonuchus, cited as Belonichus; Venezuela).
— Coiffait, 1981g: 289 (Belonuchus; synonym of rufipennis).
— Smetana, 1995e: 726 (Belonuchus; lectotype designation; synonym of rufipennis).
— Nishida, 1997: 60 (Belonuchus; Hawaii).
pallipes Melsheimer, 1844: 35 (Belonuchus; Type locality: Pennsylvania).
— Fauvel, 1877a: xxviii (Belonuchus; synonym of rufipennis).
— Fauvel, 1891: 118 (Belonuchus; synonym of formosus).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; synonym of formosus).
— Blackwelder, 1943: 432 (Belonuchus; synonym of rufipennis).
— Smetana, 1995e: 727 (Belonuchus; synonym of rufipennis).
rufoniger Fauvel, 1895: 266 (Belonuchus; Type locality: Bírmanie, Carin Cheba, 900–1100 m).
— Bernhauer and Schubert, 1914: 371 (Belonuchus; catalog).
— Bernhauer, 1928: 17 (Belonuchus; Sumatra).
— Cameron, 1932: 172 (Belonuchus; characters; Burma; India).
— Scheerpeltz, 1933: 1379 (Belonuchus; catalog).
— Cameron, 1933d: 391 (Belonuchus; Sumatra).
— Legner and Moore, 1977: 176 (Belonuchus; found under bark).
▴ Distribution: India, Myanmar, Indonesia, Taiwan, Japan.
montanus Fauvel, 1895: 267 (Belonuchus; variety of rufoniger; Type locality: Java oriental, monts Tengger, 1300 m; Java occidental, Sukabumi, 700 m).
— Bernhauer and Schubert, 1914: 371 (Belonuchus; variety of rufoniger).
— Cameron, 1937: 20 (Belonuchus; variety of rufoniger; Java).
nigripes Bernhauer, 1922c: 233 (Belonuchus; variety of rufoniger; Type locality: Hoozan, Taihorin).
— Scheerpeltz, 1933: 1379 (Belonuchus; variety of rufoniger).
— Shibata, 1974: 31 (Belonuchus; variety of rufoniger; Japan).
rufopygus Cameron, 1951, see: biformis Cameron, 1951.
rufotestaceus Cameron, 1941, see: ferrugatus Erichson, 1840.
rugicollis Bernhauer, 1916g: 276 (Belonuchus; Type locality: Peru).
satyrus Erichson, 1840: 421 (Belonuchus; Type locality: Columbia).
— Fauvel, 1891: 116 (Belonuchus; Venezuela).
— Fauvel, 1901: 86 (Belonuchus; Colombia; Venezuela).
— Blackwelder, 1943: 431 (Belonuchus; characters; Trinidad).
▴ Distribution: Colombia, Venezuela, West Indies.
schaefferi Cooper, 1933, see: ephippiatus Say, 1830.
semitestaceus Schubert, 1906: 374 (Belonuchus; Type locality: Kwai, Usambara).
sericeicollis Cameron, 1950g: 43 (Belonuchus; Type locality: Belgian Congo: Rutshuru, alt. 1285 m).
setiger Sharp, 1876: 164 (Belonuchus; Type locality: Ega).
sharpi Bernhauer, 1907a: 285 (Belonuchus; replacement name for grandiceps Sharp. Also cited from Peru).
grandiceps Sharp, 1876: 158 (Belonuchus; [preoccupied]; Type locality: Ega; Tapajos, St. Paulo).
silvestris Bernhauer, 1917c: 17 (Belonuchus; Type locality: West-Columbien: Cali am Rio Cauca, Alto de las cruces, 2200 m).
similis Sharp, 1885: 436 (Belonuchus; Type locality: Panama, Volcan de Chiriqui between 2500 and 6000 feet).
simillimus Bernhauer, 1917i: 106 (Belonuchus; Type locality: Westkolumbien: Villa Elvira, 1800 m).
simplex Sharp, 1885, see: ephippiatus Say, 1830.
skalitzkyi Bernhauer, 1906c: 334 (Belonuchus; Type locality: Peru: Pumananea).
spinicornis Champion, 1913: 4 (Musicoderus; Type locality: Costa Rica, Pitahaya, 1400 m).
stenoderus Sharp, 1885: 434 (Belonuchus; Type locality: Panama, Bugaba).
subaeneipennis Bernhauer, 1916g: 269 (Belonuchus; subgenus Musicoderus; Type locality: Umgebung von Cali in West-Columbien: Cañon del Mte. Tolima, 1700 m).
subaeneus Bernhauer, 1916g: 268 (Belonuchus; subgenus Musicoderus; Type locality: Bolivien: Yuracares).
subconvexus Bernhauer, 1937: 302 (Belonuchus; Type locality: Kenya: Homa, South Kavirondo).
subdentatus Eppelsheim, 1895a: 130 (Belonuchus; Type locality: Togoland).
subsellatus Bernhauer, 1916g: 276 (Belonuchus; Type locality: Bolivien: Yuracares).
superbus Bernhauer, 1916g: 273 (Belonuchus; Type locality: West-Cordilleren im Morast unter Bananenköder: Alto de las cruces, 2200 m; Rio Vitaco, 2000 m; Rio Aquatal, 1800 m; S. Antonio, 2000 m; Villa Elvira, 1800 m).
taczanovskii Solsky, 1872: 303 (Belonuchus; Type locality: Monte Rico, Perou).
— Bernhauer and Schubert, 1914: 371 (Belonuchus; cited as taczanowskii; catalog).
▴ Distribution: Peru.
taprobanus Cameron, 1932: 175 (Belonuchus; Type locality: Ceylon).
terminalis Laporte, 1840: 176 (Staphylinus; [preoccupied]; Type locality: Brésil).
— Bernhauer and Schubert, 1914: 371 (Belonuchus; catalog).
— Herman, 2001: 42 (Belonuchus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Brazil.
texanus Casey, 1915, see: arizonicus Casey, 1915.
thoracicus Fauvel, 1903, see: grandiceps Kraatz, 1859.
tremolerasi Bernhauer, 1912d: 40 (Belonuchus; Type locality: Uruguay).
— Bernhauer and Schubert, 1914: 372 (Belonuchus; catalog).
— Scheerpeltz, 1933: 1379 (Belonuchus; catalog; Argentina, Paraguay).
▴ Distribution: Uruguay, Paraguay, Argentina.
trinitatis Blackwelder, 1943: 423 (Belonuchus; Type locality: Trinidad: Department of Africulture Stock Farm at St. Joseph, 6 miles east of Port of Spain).
tripunctatus Fauvel, 1895: 267 (Belonuchus; Type locality: Bírmanie, Carin Cheba, 900–1100 m).
— Bernhauer and Schubert, 1914: 372 (Belonuchus; catalog).
— Cameron, 1932: 176 (Belonuchus; characters; Burma).
— Scheerpeltz, 1933: 1379 (Belonuchus; catalog).
— Cameron, 1933d: 391 (Belonuchus; Sumatra).
▴ Distribution: Myanmar, Indonesia.
troglophilus Smetana, 1995e: 742 (Belonuchus; Type locality: Arizona: Buckelew Cave, Cochise Co. Also cited from Oklahoma and Texas).
tucumanensis Bernhauer, 1934b: 213 (Belonuchus; Type locality: Argentinien: Tucuman).
ussuriensis Tikhomirova, 1973: 170 (Belonuchus; Type locality: Suputinskii zapovednik).
variolosus Solsky, 1870a: 263 (Belonuchus; Type locality: Cayenne).
— Solsky, 1872: 305 (Belonuchus; cited as variolaris; Cayenne).
— Bernhauer and Schubert, 1914: 372 (Belonuchus; catalog).
▴ Distribution: French Guiana.
venezolanus Bernhauer, 1917i: 104 (Belonuchus; Type locality: Venezuela; Kolumbien).
— Scheerpeltz, 1933: 1380 (Belonuchus; catalog).
— Marcuzzi, 1977: 13 (Belonuchus, cited as Belonichus; Venezuela).
▴ Distribution: Venezuela, Colombia.
vestitus R. Sahlberg, 1847, see: Xanthodermus.
viridipennis Baudi, 1848: 125 (Belonuchus; Type locality: Novae Valenciae, Columbia).
— Fauvel, 1891: 118 (Belonuchus; Venezuela).
— Bernhauer and Schubert, 1914: 372 (Belonuchus; catalog).
▴ Distribution: Venezuela, Colombia, Panama, Guatemala, Mexico.
dejectus Sharp, 1885: 433 (Belonuchus; Type locality: Mexico, Cordova, Jalapa, Las Vigas, Mirador; Guatemala, Capetillo, Cerro Zunil, Balheu, Tamahu, San Joaquin, Purula, El Tumbador, San Geronimo; Panama, Volcan de Chiriqui 2500 to 4000 feet).
— Sharp, 1887: 789 (Belonuchus; synonym of viridipennis).
— Fauvel, 1891: 118 (Belonuchus; synonym of viridipennis).
— Bernhauer and Schubert, 1914: 372 (Belonuchus; synonym of viridipennis).
vulneratus Bernhauer, 1906c: 335 (Belonuchus; Type locality: Peru: Pachitea).
xantholinoides Bernhauer, 1917i: 106 (Belonuchus; Type locality: Westkolumbien: Villa Elvira, 1800 m; S. Antonio, 2200 m, Umgebung von Cali am Rio Cauca).
xanthomelas Solsky, 1868a: 139 (Belonuchus; Type locality: Mexique).
— Sharp, 1885: 417 (Philonthus; Mexico; Arizona).
— Bernhauer and Schubert, 1914: 372 (Belonuchus; catalog).
— Scheerpeltz, 1933: 1380 (Belonuchus; catalog).
— Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 61 (Belonuchus; necrophilous species; distribution in Mexico).
▴ Distribution: Mexico, USA.
xanthopterus Nordmann, 1837: 99 (Philonthus; Type locality: Parae in America meridionalis).
— Erichson, 1840: 421 (Belonuchus; characters; Brazil).
— Sharp, 1876: 157 (Belonuchus; Brazil).
— Bernhauer and Schubert, 1914: 372 (Belonuchus; catalog).
▴ Distribution: Brazil.
xanthopus Solsky, 1868a: 139 (Belonuchus; Type locality: Mexique).
Bisnius
[66 species; Neotropical, Nearctic, Palaearctic, Oriental, and Australian Regions]
Bisnius Stephens, 1829a: 23 (species included: cephalotes and four unavailable names; [Note: The same questions and controversy concerning the authorship and date of publication for Philonthus apply to Bisnius. The issue is discussed in my “note” attached to the citation of the original description for Philonthus]). Type species: Staphylinus cephalotes Gravenhorst, fixed by monotypy.
— Curtis, 1829: 26 (catalog; Britain).
— Stephens, 1829: 283 (catalog; Britain).
— Stephens, 1832: 200 (characters).
— Stephens, 1833: 247 (characters).
— Westwood, 1838a: 16 (characters; type species).
— Stephens, 1839: 385 (characters).
— Erichson, 1840: 426 (synonym of Philonthus).
— Lacordaire, 1854: 80 (synonym of Philonthus).
— Kraatz, 1857d: 565 (synonym of Philonthus).
— G. Waterhouse, 1858: 24 (synonym of Philonthus).
— Gemminger and Harold, 1868: 584 (synonym of Philonthus).
— Fauvel, 1874: 207 [= 1874a: 433] (synonym of Philonthus).
— Mulsant and Rey, 1876: 583 [= 1877: 439] (characters; key to French species).
— Heyden, Reitter, and Weise, 1891: 102 (list of species of Europe and Caucasus).
— Bernhauer and Schubert, 1914: 326 (synonym of Philonthus).
— Johansen, 1914: 423 (characters; key to Danish species).
— Cameron, 1932: 62 (subgenus of Philonthus).
— Tottenham, 1939d: 227 (discussion of nomenclature).
— Blackwelder, 1943: 399 (subgenus of Philonthus; type species: cephalotes).
— Tottenham, 1949: 372, 418 (type species: cephalotes Gravenhorst; subgenus of Philonthus; checklist of British species).
— Blackwelder, 1952: 76 (type species: cephalotes).
— Smetana, 1995e: 513 (valid genus; characters; notes; key to North American species; transfer of Old World species to Bisnius; phylogenetic relationships).
— Lundgren, 1998: 50 (list of species from Florida; records for other States listed for each species).
— Newton, Thayer, Ashe, and Chandler, 2000: 396 (34 Nearctic species; notes; characters in key).
— Hayashi, 2000: 23 (notes on empodial setae).
Gefyrobius Thomson, 1859: 24 (species included: nitidulus). Type species: Staphylinus nitidulus Gravenhorst, fixed by original designation and monotypy.
— Thomson, 1860: 166 (characters).
— Gemminger and Harold, 1868: 584 (synonym of Philonthus).
— Fauvel, 1874: 207 [= 1874a: 433] (synonym of Philonthus).
— Mulsant and Rey, 1876: 530 [= 1877: 386] (characters).
— Heyden, Reitter, and Weise, 1891: 104 (subgenus of Philonthus; list of species of Europe and Caucasus).
— Heyden, 1893: 43 (subgenus of Philonthus; list of species of Siberian region).
— Heyden, 1896: 33 (subgenus of Philonthus; list of species of Siberian region).
— Bernhauer and Schubert, 1914: 326 (synonym of Philonthus).
— Cameron, 1932: 62 (subgenus of Philonthus).
— Scheerpeltz, 1933: 1330 (subgenus of Philonthus).
— Blackwelder, 1943: 399 (synonym of Bisnius; type species: nitidulus).
— Tottenham, 1949: 372 (synonym of Bisnius; type species: denigrator Gravenhorst = nitidulus Gravenhorst).
— Blackwelder, 1952: 168 (type species: nitidulus).
— Smetana, 1958: 143 (synonym of Philonthus).
— Coiffait, 1974: 116, 120, 192 (subgenus of Philonthus; characters; key to species of western Palaearctic region).
— Smetana, 1995e: 513 (synonym of Bisnius).
alaskensis Smetana, 1965, see: hyperboreus J. Sahlberg, 1880.
agenor Smetana, 1995e: 591 (Bisnius; Type locality: Washington: Mt. Rainier N. P., Tahoma Crk., 2400 ft. Also cited from California and Oregon).
agyrtes Smetana, 1973a: 46 (Philonthus; replacement name for negligens Smetana).
— Coiffait, 1974: 285 (Philonthus; characters; Italy).
— Brunne, 1976: 87 (Philonthus; characters).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; cited as “specie dubbia”; Italy).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: Italy.
negligens Smetana, 1966a: 18 (Philonthus; subgenus Philonthus; [preoccupied]; Type locality: Italien: Süd-Tirol, Brixen-Bressanone).
— Coiffait, 1967a: 408 (Philonthus; characters; Italy).
— Brunne, 1976: 87 (Philonthus; synonym of agyrtes).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; synonym of agyrtes).
— Smetana, 1995e: 516 (Bisnius).
alaiensis Fauvel, 1900d: 231 (Philonthus; Type locality: Turkestan: Monts Alai; Djizak).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog).
— Scheerpeltz, 1933: 1331 (Philonthus; catalog).
— Coiffait, 1967a: 398, 407 (Philonthus; characters; Turkestan; Transcaspian region).
— Coiffait, 1974: 211 (Philonthus; characters; Turkestan; Transcaspian region).
— Shilov, 1976: 150 (Philonthus; Mongolia).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: Kazakhstan, Mongolia.
tantulus Luze, 1904: 94 (Philonthus; Type locality: Seravschan: Turusch-dara-Pass; Jagnot: Schach-sara, Varsaut, Fl. Dshidshigrut).
— Bernhauer and Schubert, 1914: 357 (Philonthus; cited as tantillus; catalog).
— Scheerpeltz, 1933: 1363 (Philonthus; catalog).
— Coiffait, 1974: 211 (Philonthus; synonym of alaiensis).
albionicus Mannerheim, 1843b: 231 (Philonthus; Type locality: California; [Note: See lectotype designation by Smetana, 1995e]).
— Horn, 1884: 217 (Philonthus; characters; Alaska; Washington Territory; Vancouver; Arizona).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog).
— Hatch, 1957: 183 (Philonthus; characters; British Columbia; Washington; Oregon).
— Smetana, 1995e: 596 (Bisnius; lectotype designation; characters; notes; California).
▴ Distribution: Canada, USA.
arenarius Kirshenblat, 1950: 240 (Philonthus; Type locality: Turkmeniia: Ashkhabad).
— Coiffait, 1967a: 401 (Philonthus; characters; Turkmenia).
— Coiffait, 1974: 281 (Philonthus; characters; Turkmenia).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: Turkmenistan.
atrolucens Casey, 1915, see: filicornis Horn, 1884.
attiguus Herman, 2001: 43 (Bisnius; replacement name for propinquus Kirshenblat).
propinquus Kirshenblat, 1950: 238 (Philonthus; [preoccupied]; Type locality: Buriat-Mongol'skiia ASSR: Ulan-ude; Chitinskaia oblast': plato Kurtus, Dauriia; Gorulevskaia-Tataurova; Chita, r. Kaidalovka; Iakutskaia ASSR: Iakusk).
— Coiffait, 1967a: 410 (Philonthus; characters; central Asia).
— Coiffait, 1974: 287 (Philonthus; characters; Siberia).
— Smetana, 1995e: 516 (Bisnius).
— Herman, 2001: 43 (Bisnius; synonym of attiguus).
baicalensis Bernhauer, 1927c: 123 (Philonthus; Type locality: Baikalgebiet: Turansk; Kultuk).
— Scheerpeltz, 1933: 1333 (Philonthus; catalog).
— Coiffait, 1967a: 408 (Philonthus; characters; Baikal).
— Coiffait, 1974: 284 (Philonthus; characters; Baikal region).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: Russia.
bassus Smetana, 1995e: 586 (Bisnius; Type locality: California: Eldorado Co.: 5 mi. SW Kyburz, 4000 ft. Also cited from Oregon).
batkhizensis Coiffait, 1967a: 442 (Philonthus; subgenus Philonthus; Type locality: réserve d'État Batkhis S.S.R. de Turkménie).
— Coiffait, 1974: 235 (Philonthus; characters; Turkestan).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: Turkmenistan.
bion Smetana, 1995e: 572 (Bisnius; Type locality: British Columbia: Mt. Garibaldi, 14 mi. N. Squamish, 4000 ft. Also cited from Oregon).
blandus Gravenhorst, 1806: 72 (Staphylinus; Type locality: America septentr.; [Note: See lectotype designation by Smetana, 1995e]).
— Say, 1830: 35 [= 1834: 449] (Staphylinus; characters).
— Erichson, 1840: 458 (Philonthus; characters; America septentrionalis).
— Melsheimer, 1844: 36 (Philonthus; distinct from laetulus Say).
— Provancher, 1877: 249 (Philonthus; characters; Quebec).
— Horn, 1884: 208 (Philonthus; characters; Canada to Virginia and Illinois).
— Blatchley, 1910: 387 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Notman, 1920a: 180 (Philonthus; New York).
— Smetana, 1995e: 579 (Bisnius; lectotype designation; characters; notes; Nova Scotia; Ontario; Quebec; Alabama; Arkansas; Connecticut; Delaware; Georgia; Illinois; Indiana; Iowa; Kansas; Kentucky; Maine; Maryland; Massachusetts; Michigan; Minnesota; Nebraska; New Hampshire; New Jersey; New York; North Carolina; Ohio; Oklahoma; Pennsylvania; South Carolina; Tennessee; Vermont; Virginia; West Virginia; Wisconsin).
— Levesque and Levesque, 1996: 292 (Philonthus; seasonal abundance; Quebec).
— Downie and Arnett, 1996: 380 (Philonthus; characters; USA).
▴ Distribution: Canada, USA.
pulchellus Melsheimer, 1844: 36 (Philonthus; Type locality: Not cited).
— Fauvel, 1869: 488 (Philonthus; synonym of blandus).
— Bernhauer and Schubert, 1914: 330 (Philonthus; synonym of blandus).
— Smetana, 1995e: 579 (Bisnius; synonym of blandus).
thoracicus Melsheimer, 1844: 36 (Philonthus; [Note: Preoccupied secondary homonym in Philonthus]; variety of pulchellus; Type locality: Not cited).
paederinus Sachse, 1852: 126 (Philonthus; Type locality: Not cited).
— Fauvel, 1869: 488 (Philonthus; synonym of blandus).
— Bernhauer and Schubert, 1914: 330 (Philonthus; synonym of blandus).
— Smetana, 1995e: 579 (Bisnius; synonym of blandus).
brouni Duvivier, 1883, see: sordidus Gravenhorst, 1802.
bucharensis Coiffait, 1967a: 442 (Philonthus; subgenus Philonthus; Type locality: Buchara, S.S.R. d'Ouzbekistan. Also cited from Kirghizistan).
— Coiffait, 1974: 236 (Philonthus; characters; Kirghistan; Uzbekistan).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: Uzbekistan, Kyrgyzstan.
cephalicus Casey, 1915: 438 (Philonthus; Type locality: Manitoba: Aweme).
— Scheerpeltz, 1933: 1335 (Philonthus; catalog).
— Moore, 1965: 100 (Gabrius).
— Smetana, 1995e: 567 (Bisnius; characters; notes; Alberta; Manitoba).
▴ Distribution: Canada.
cephalotes Gravenhorst, 1802: 22 (Staphylinus; [preoccupied]; Type locality: Brunsvigae; [Note: See lectotype designation by Smetana, 1995e]. No locality cited originally).
— Latreille, 1804: 311 (Staphylinus; characters; France; Germany).
— Gravenhorst, 1806: 71 (Staphylinus; characters; France).
— Gyllenhal, 1810: 320 (Staphylinus; characters; Sweden).
— Curtis, 1829: 26 (Bisnius; catalog; Britain).
— C. Sahlberg, 1830: 321 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 28 (Staphylinus; Finland).
— Mannerheim, 1831: 442 (Staphylinus; Finland).
— Stephens, 1833: 247 (Bisnius; characters; England).
— Lacordaire, 1835: 390 (Staphylinus; characters; France).
— Nordmann, 1837: 82 (Philonthus; characters; Sweden; Russia; Germany).
— Westwood, 1838a: 16 (Bisnius; characters; type species).
— Zetterstedt, 1838: 63 (Staphylinus; characters; habitat; Lapland).
— Erichson, 1839a: 459 (Philonthus; characters; Germany).
— Heer, 1839: 263 (Philonthus; characters; Switzerland).
— Stephens, 1839: 399 (Bisnius; characters; England).
— Erichson, 1840: 455 (Philonthus; characters; Europe; Syria; Egypt).
— Laporte, 1840: 178 (Staphylinus; characters; France).
— Jacobsen, 1849: 695 (Philonthus; characters; Denmark).
— Redtenbacher, 1849: 701 (Philonthus; characters; Austria).
— Hardy, 1851: 28 (Philonthus; notes; England).
— Fairmaire and Laboulbène, 1856: 519 (Philonthus; characters; France).
— Staudinger, 1857: 286 (Philonthus; notes; Iceland).
— Kraatz, 1857d: 592 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 190 (Philonthus; characters; Austria).
— Thomson, 1860: 158 (Philonthus; characters; Scandinavia).
— Fauvel, 1865a: 296 [= 1865b: 46] (Philonthus; habitat; France).
— Fauvel, 1874: 220 [= 1874a: 446] (Philonthus; characters; France; Algeria; Egypte; Cyprus; Syria; North America).
— Redtenbacher, 1874: 209 (Philonthus; characters; Austria).
— Seidlitz, 1875: 274 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 427 [= 1877: 283] (Philonthus; characters; France).
— J. Sahlberg, 1876: 16 (Philonthus; Finland).
— Fauvel, 1878: 122 [= 1878a: 42] (Philonthus; Algeria; Egypt).
— Horn, 1884: 209 (Philonthus; characters; Massachusetts; middle States).
— Fauvel, 1886: 53 [= 1886a: 45] (Philonthus; Egypt; Algeria).
— Fowler, 1888: 267 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 450 (Philonthus; characters; Europe; Mediterranean region).
— Xambeu, 1900: 4 (Philonthus; larval characters).
— Fauvel, 1902b: 108 (Philonthus; Algeria).
— Poppius, 1909: 26 (Philonthus; subgenus Gefyrobius; Lena Valley, Siberia).
— Reitter, 1909: 127 (Philonthus; characters; Germany).
— Blatchley, 1910: 388 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 331 (Philonthus; catalog).
— Johansen, 1914: 402 (Philonthus; characters; Denmark).
— Scheerpeltz, 1929a: 117 (Philonthus; notes; Lake Baikal).
— Portevin, 1929: 359 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1335 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— West, 1937: 40 (Philonthus; Faroe Islands).
— Paulian, 1941: 225 (Philonthus; larval characters).
— Tottenham, 1949: 418 (Philonthus; checklist; Britain).
— Jeannel and Jarrige, 1949: 375 (Philonthus; notes; cave species; Romania; France).
— V. Hansen, 1952: 61 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1955b: 227 (Philonthus; characters).
— Hatch, 1957: 194 (Philonthus; characters; British Columbia; Washington; Oregon).
— Scheerpeltz, 1958a: 16 (Philonthus; Turkey).
— Smetana, 1958: 201 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 154 (Philonthus; characters).
— Smetana, 1959a: 211 (Philonthus; Albania).
— Larsson and Gígja, 1959: 84 (Philonthus; biological notes; Iceland).
— Ferreira, 1962: 17 (Philonthus; Portugal).
— Palm, 1963b: 77 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 180 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 150 (Philonthus; all Europe).
— Pototskaia, 1966: 156 (Philonthus; larval characters).
— Pototskaia, 1967: 57 (Philonthus; larval characters).
— Coiffait, 1967a: 408 (Philonthus; characters).
— Scheerpeltz, 1968a: 67 (Philonthus; catalog; Austria).
— Coiffait, 1970e: 62 (Philonthus; Cyprus).
— Coiffait, 1974: 240 (Philonthus; characters; Europe; Britain; Scandinavia; Siberia; North Africa).
— Shilov, 1975a: 376 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Shilov, 1976: 151 (Philonthus; Mongolia).
— Pope, 1977: 29 (Philonthus; Britain).
— Bordoni, 1978a: 59 (Philonthus; cave collection; Turkey).
— Muona, 1979: 14 (Philonthus; Finland; Russia; Sweden; Norway; Denmark; Baltic States).
— Waterston, Owen, Welch, Bacchus and Hammond, 1981: 274 (Philonthus; Outer Hebrides).
— Frank, 1982: 39 (Philonthus; list of parasites).
— Tóth, 1984: 60 (Philonthus; characters; Hungary).
— Muona and Viramo, 1986: 13 (Philonthus; Finland).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Gusarov, 1989: 10 (Philonthus; habitat notes; Ukraine).
— Silfverberg, 1992: 16 (Philonthus; Finland; Russia; Sweden; Norway; Denmark; Estonia; Latvia; Lithuania).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Sadler and Dugmore, 1995: 144 (Philonthus; habitat notes; Iceland).
— Smetana, 1995e: 527 (Bisnius; lectotype designation: Brunsuigae; characters; notes; British Columbia; Manitoba; Newfoundland; Nova Scotia; Ontario; Quebec; Arizona; California; Colorado; Florida; Idaho; Iowa; Maryland; Massachusetts; Michigan; New Jersey; New York; North Carolina; Oregon; Pennsylvania; Washington; Wisconsin).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; Italy).
— Downie and Arnett, 1996: 381 (Philonthus; characters; USA).
— M. Hansen, 1996: 105 (Philonthus; checklist; Denmark).
— T. Wagner, 1997: 235 (Philonthus; collecting notes; Germany).
— Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 36 (Philonthus; Latvia).
— Anderson, Nash, and O'Connor, 1997: 18 (Philonthus; Ireland).
— Owen, 1997a: 149 (Philonthus; collecting sites; Ireland).
— Bordoni and Oromi, 1998: 1157 (Philonthus; facultative cave species; Bulgaria; France; Roumania).
— Silfverberg, 1999: 50 (Bisnius; Estonia).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
— Herman, 2001: 43 (Bisnius; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5 and the older homonym is a nomen dubium).
▴ Distribution: Egypt, Algeria, Iceland, Europe, Turkey, Syria, Russia, Kazakhstan, Mongolia, Canada, USA.
megacephalus Heer, 1839: 263 (Philonthus; [Note: Heer cited his 1834 paper as the original reference for this name but all the species group names in both 1834 articles are unavailable]; Type locality: Matt).
— Heer, 1841: 582 (Philonthus; notes).
— Kraatz, 1857d: 593 (Philonthus; synonym of cephalotes).
— Fauvel, 1874: 220 [= 1874a: 446] (Philonthus; synonym of cephalotes).
— Mulsant and Rey, 1876: 430 [= 1877: 286] (Philonthus; characters; France).
— J. Sahlberg, 1876: 16 (Philonthus; synonym of cephalotes).
— Ganglbauer, 1895: 450 (Philonthus; synonym of cephalotes).
— Bernhauer and Schubert, 1914: 331 (Philonthus; synonym of cephalotes).
— Smetana, 1958: 202 (Philonthus; synonym of cephalotes).
— Coiffait, 1974: 240 (Philonthus; synonym of cephalotes).
— Smetana, 1995e: 527 (Bisnius; synonym of cephalotes).
subrugosus Mulsant and Rey, 1876: 424 [= 1877: 280] (Philonthus; Type locality: Provence).
— Ganglbauer, 1895: 450 (Philonthus; synonym of cephalotes).
— Bernhauer and Schubert, 1914: 331 (Philonthus; synonym of cephalotes).
— Smetana, 1958: 202 (Philonthus; synonym of cephalotes).
— Smetana, 1995e: 527 (Bisnius; synonym of cephalotes).
foveifrons J. Sahlberg, 1913: 66 (Philonthus; aberration of cephalotes; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Hierosolyma, Palaestina).
— Scheerpeltz, 1933: 1336 (Philonthus; aberration of cephalotes).
— Smetana, 1958: 202 (Philonthus; synonym of cephalotes).
— Coiffait, 1967a: 408 (Philonthus; aberration of cephalotes; characters).
— Coiffait, 1974: 240 (Philonthus; variety of cephalotes).
— Smetana, 1995e: 527 (Bisnius; synonym of cephalotes).
rufipennis Delahon, 1914: 621 (Philonthus; aberration of cephalotes; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; preoccupied]; cited from Mark Brandenburg).
— Scheerpeltz, 1933: 1336 (Philonthus; synonym of cephalotes).
— Coiffait, 1974: 241 (Philonthus; synonym of cephalotes).
— Smetana, 1995e: 527 (Bisnius; synonym of cephalotes).
synonymicus Roubal, 1915b: 102 (Philonthus; replacement name for rufipennis Delahon; [Note: Nomen nudum; the preoccupied name, rufipennis Delahon, is unavailable and so therefore is synonymicus]).
— Scheerpeltz, 1933: 1336 (Philonthus; aberration of cephalotes).
— Smetana, 1959: 154 (Philonthus; aberration; characters).
— Smetana, 1958: 202 (Philonthus; synonym of cephalotes).
— Coiffait, 1967a: 410 (Philonthus; aberration of cephalotes; characters).
— Coiffait, 1974: 241 (Philonthus; variety of cephalotes).
— Tóth, 1984: 61 (Philonthus; cited as synonimicus; aberration of cephalotes).
— Smetana, 1995e: 527 (Bisnius; synonym of cephalotes).
thessalonicus Coiffait, 1963: 23 (Philonthus; aberration of cephalotes; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Salonique).
— Coiffait, 1967a: 408 (Philonthus; variety of cephalotes; characters).
— Coiffait, 1974: 241 (Philonthus; variety of cephalotes).
chilensis Solier, 1849, see: sordidus Gravenhorst, 1802.
chloropterus Solier, 1849, see: sordidus Gravenhorst, 1802.
coloripennis Bernhauer, 1910, see: sordidus Gravenhorst, 1802.
crestonensis Hatch, 1957: 193 (Philonthus; Type locality: British Columbia: Creston).
— Legner and Moore, 1977: 176 (Philonthus; found under bark).
— Smetana, 1995e: 540 (Bisnius; characters; notes; British Columbia).
▴ Distribution: Canada.
crito Smetana, 1995e: 574 (Bisnius; Type locality: Washington: Olympic Nat. Pk., Hoh Ranger Stn., 600 ft.).
denigrator Gravenhorst, 1806, see: nitidulus Gravenhorst, 1802.
diversipennis Bernhauer, 1899: 23 (Philonthus; Type locality: Tschavanga, Kola; Levander in Varsuga; Eversmann in Orenburg; Jaroslaw in Russland).
— Bernhauer and Schubert, 1914: 336 (Philonthus; catalog).
— Scheerpeltz, 1933: 1340 (Philonthus; catalog).
— Kirshenblat, 1951: 240 (Philonthus; characters in key).
— Palm, 1963b: 79 (Philonthus; characters; Finland).
— Pototskaia, 1966: 149, 156 (Philonthus; larval characters).
— Pototskaia, 1967: 58 (Philonthus; larval characters).
— Coiffait, 1967a: 407 (Philonthus; characters; Russia; Turkestan).
— Coiffait, 1974: 213 (Philonthus; characters; Russia; Turkestan).
— Shilov, 1975a: 376 [= 1975b: 99] (Philonthus; Komi region of Russia).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: Finland, Russia, “Turkestan”.
eidmanni Scheerpeltz, 1929a: 117 (Philonthus; Type locality: Nord-Mandschurei).
— Scheerpeltz, 1933: 1341 (Philonthus; catalog).
— Kirshenblat, 1950: 241 (Philonthus; characters in key).
— Coiffait, 1967a: 410 (Philonthus; characters; Manchuria).
— Coiffait, 1974: 286 (Philonthus; characters).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: China.
epeus Smetana, 1995e: 578 (Bisnius; Type locality: British Columbia: 2 mi. S. Salmo).
filicornis Horn, 1884: 210 (Philonthus; Type locality: Lake Tahoe, Nevada).
— Bernhauer and Schubert, 1914: 338 (Philonthus; catalog).
— Smetana, 1995e: 584 (Bisnius; characters; notes; Arizona; California).
▴ Distribution: USA.
atrolucens Casey, 1915: 434 (Philonthus; Type locality: California: Gilroy Hot Springs, Sta. Clara Co.; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1333 (Philonthus; catalog).
— Smetana, 1995e: 584 (Bisnius; lectotype designation; synonym of filicornis).
fimetarius Gravenhorst, 1802: 175 (Staphylinus; Type locality: Brunsvigae; [Note: See lectotype designation by Smetana, 1995e]).
— Latreille, 1804: 325 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 80 (Staphylinus; characters).
— Gyllenhal, 1810: 324 (Staphylinus; characters; Sweden).
— Zetterstedt, 1828: 75 (Staphylinus; characters; Lapland).
— Curtis, 1829: 25 (Philonthus; catalog; Britain).
— C. Sahlberg, 1830: 322 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 29 (Staphylinus; Finland; Sweden; Lapland; Russia).
— Mannerheim, 1831: 443 (Staphylinus; Finland; Sweden; Lapland; Russia).
— Stephens, 1832: 232 (Philonthus; characters; England).
— Ménétriés, 1832: 146 (Staphylinus; Caucasus-Persia region).
— Lacordaire, 1835: 406 (Staphylinus; characters; France).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Nordmann, 1837: 84 (Philonthus; Sweden; Germany; France).
— Zetterstedt, 1838: 63 (Staphylinus; characters; habitat; Lapland).
— Erichson, 1839a: 460 (Philonthus; characters; Germany).
— Stephens, 1839: 396 (Philonthus; characters; England).
— Erichson, 1840: 456 (Philonthus; characters; Europe).
— Dufour, 1843: 34 (Staphylinus; characters; France).
— Jacobsen, 1849: 696 (Philonthus; characters; Denmark).
— Hochhuth, 1849: 137 (Philonthus; characters).
— Redtenbacher, 1849: 703 (Philonthus; characters; Austria).
— Kiesenwetter, 1851a: 416 (Philonthus; Mont Serrat).
— Hardy, 1851: 28 (Philonthus; notes; England).
— Fairmaire and Laboulbène, 1856: 521 (Philonthus; characters; France).
— Kraatz, 1857d: 592 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 193 (Philonthus; characters; Austria).
— Thomson, 1860: 159 (Philonthus; characters; Scandinavia).
— Fauvel, 1865a: 296 [= 1865b: 46] (Philonthus; habitat; France).
— Fauvel, 1874: 240 [= 1874a: 466] (Philonthus; characters; France; Algeria; Cyprus; Turkey; Caucasus).
— Redtenbacher, 1874: 211 (Philonthus; characters; Austria).
— Seidlitz, 1875: 274 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 430 [= 1877: 286] (Philonthus; characters; larval characters; France).
— J. Sahlberg, 1876: 16 (Philonthus; Finland).
— Fauvel, 1878: 124 [= 1878a: 44] (Philonthus; Algeria).
— Fauvel, 1886: 55 [= 1886a: 47] (Philonthus; Morocco; Algeria).
— Fowler, 1888: 268 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 450 (Philonthus; characters; Europe; Mediterranean region).
— Fauvel, 1902b: 111 (Philonthus; Morocco; Algeria; Tunisia; Egypt).
— Poppius, 1905: 10 (Philonthus; Orkney Islands; Shetland Islands).
— Sainte-Claire Deville, 1906: 97 (Philonthus; Corsica).
— Reitter, 1909: 127 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 338 (Philonthus; catalog).
— Johansen, 1914: 401 (Philonthus; characters; Denmark).
— Portevin, 1929: 360 (Philonthus; characters; France).
— Gridelli, 1930: 73 (Philonthus; Libya).
— Cameron, 1932: 92 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1342 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— West, 1937: 40 (Philonthus; Faroe Islands).
— Jeannel and Jarrige, 1949: 374 (Philonthus; cave species; Dalamatia; Hercegowina).
— V. Hansen, 1952: 60 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1958: 200 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 151 (Philonthus; characters).
— Scheerpeltz, 1961d: 129 (Philonthus; Italy).
— Palm, 1963b: 76 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 179 (Philonthus; subgenus Philonthus; characters; central Europe).
— Smetana, 1964d: 75 (Philonthus; subgenus Philonthus; Czechoslovakia).
— Scheerpeltz, 1965b: 402 (Philonthus; Sicily).
— Smetana, 1965: 17 (Philonthus; Newfoundland).
— Horion, 1965: 146 (Philonthus; widespread in southern and central Europe; Finland; Sweden; Norway).
— Coiffait, 1967a: 410 (Philonthus; synonym of rigidicornis).
— Raianu, 1968a: 467 (Philonthus; Romania).
— Szujecki, 1968a: 730 (Philonthus; Poland).
— Kasule, 1970: 68 (Philonthus; larval characters).
— Coiffait, 1974: 217 (Philonthus; synonym of rigidicornis).
— Shilov, 1975a: 376 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Pope, 1977: 29 (Philonthus; Britain).
— Muona, 1979: 14 (Philonthus; synonym of rigidicornis).
— Hinton, 1981: 660 (Philonthus; characters of egg).
— Frank, 1982: 40 (Philonthus; list of parasites).
— Boller, 1983: 120 (Philonthus; larval characters).
— Tóth, 1984: 57 (Philonthus; characters; Hungary).
— Biswas and Biswas, 1985: 145 (Philonthus; India).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Pilon and Zanetti, 1991: 58 (Philonthus; Italy).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Smetana, 1995e: 556 (Bisnius; lectotype designation: Brunsvigae; characters; notes; Newfoundland; Quebec; introduced into North America).
— Ádám, 1996: 245 (Bisnius; synonym of rigidicornis).
— Owen, 1997a: 149 (Philonthus; collecting sites; Ireland).
— Kashcheev, 1999a: 58 (Philonthus; Kazakhstan).
▴ Distribution: Egypt, Libya, Tunisia, Algeria, Morocco, Europe, Russia, “Caucasus-Iranian region”, Turkey, Kazakhstan, India, “East Indies”, Canada.
rigidicornis Gravenhorst, 1802: 23 (Staphylinus; Type locality: Brunsvigae; [Note: See lectotype designation by Smetana, 1995e]. No locality cited originally).
— Latreille, 1804: 312 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 69 (Staphylinus).
— Mannerheim, 1830: 28 (Staphylinus; Lapland).
— Mannerheim, 1831: 442 (Staphylinus; Lapland).
— Nordmann, 1837: 82 (Philonthus; Prussia; Lapland).
— Zetterstedt, 1838: 63 (Staphylinus; characters; habitat; Lapland).
— Erichson, 1839a: 460 (Philonthus; synonym of fimetarius).
— Heer, 1839: 263 (Philonthus; characters; Switzerland).
— Erichson, 1840: 456 (Philonthus; synonym of fimetarius).
— Kraatz, 1857d: 592 (Philonthus; synonym of fimetarius).
— Ganglbauer, 1895: 450 (Philonthus; synonym of fimetarius).
— Bernhauer and Schubert, 1914: 338 (Philonthus; synonym of fimetarius).
— Cameron, 1932: 92 (Philonthus; synonym of fimetarius).
— Smetana, 1958: 200 (Philonthus; synonym of fimetarius).
— Coiffait, 1967a: 410 (Philonthus; cited as valid name; characters).
— Coiffait, 1974: 217 (Philonthus; valid species; characters; Palaearctic region).
— Pace and Zanetti, 1977: 296 (Philonthus; Italy).
— Coiffait, 1978d: 164 (Philonthus; Turkey).
— Muona, 1979: 14 (Philonthus; cited as valid name).
— Outerelo, 1980a: 56 (Philonthus; Spain).
— Smetana, 1993b: 48 (Philonthus; synonym of fimetarius).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; synonym of fimetarius).
— Smetana, 1995e: 556 (Bisnius; lectotype designation: Brunsvigae; synonym of fimetarius).
— Ádám, 1996: 245 (Bisnius; valid name; Hungary).
— Outerelo, Gamarra, and Salgado, 1998: 129 (Philonthus; subgenus Philonthus; cited as valid species; collected from cave; Spain).
longulus Motschulsky, 1858: 662 (Gabrius; Type locality: Indes orientales).
— Bernhauer and Schubert, 1914: 338 (Philonthus; synonym of fimetarius).
— Cameron, 1932: 92 (Philonthus; synonym of fimetarius).
— Smetana, 1958: 200 (Philonthus; synonym of fimetarius).
— Coiffait, 1974: 218 (Philonthus; synonym of rigidicornis).
— Smetana, 1995e: 556 (Bisnius; synonym of fimetarius).
xanthomerus Kraatz, 1859: 83 (Philonthus; Type locality: India orientalis).
— Bernhauer and Schubert, 1914: 338 (Philonthus; synonym of fimetarius).
— Cameron, 1932: 92 (Philonthus; synonym of fimetarius).
— Scheerpeltz, 1933: 1368 (Philonthus; catalog; male in original series is xathomerus; female in series is fimetarius).
— Smetana, 1958: 200 (Philonthus; synonym of fimetarius).
— Coiffait, 1974: 218 (Philonthus; synonym of rigidicornis).
interpunctatus Hochhuth, 1860: 566 (Philonthus; Type locality: Russie méridionale aux environs de Kharkov).
— Hochhuth, 1862: 57 (Philonthus; characters; Süden des Poltawschen Gouvernements; bei Ekatherinoslaw).
— Hochhuth, 1872: 137 (Philonthus; characters; Ukraine).
— Fauvel, 1874: 240 [= 1874a: 466] (Philonthus; synonym of fimetarius).
— Ganglbauer, 1895: 450 (Philonthus; synonym of fimetarius).
— Bernhauer and Schubert, 1914: 338 (Philonthus; synonym of fimetarius).
— Cameron, 1932: 92 (Philonthus; synonym of fimetarius).
— Smetana, 1958: 200 (Philonthus; synonym of fimetarius).
— Coiffait, 1974: 218 (Philonthus; synonym of rigidicornis).
— Smetana, 1995e: 556 (Bisnius; synonym of fimetarius).
stenoderus Reiche, 1861: 204 (Philonthus; Type locality: Corsica).
— Fauvel, 1874: 240 [= 1874a: 466] (Philonthus; synonym of fimetarius).
— Ganglbauer, 1895: 450 (Philonthus; synonym of fimetarius).
— Bernhauer and Schubert, 1914: 338 (Philonthus; synonym of fimetarius).
— Cameron, 1932: 92 (Philonthus; synonym of fimetarius).
— Smetana, 1958: 200 (Philonthus; synonym of fimetarius).
— Coiffait, 1974: 218 (Philonthus; synonym of rigidicornis).
— Smetana, 1995e: 556 (Bisnius; synonym of fimetarius).
finmarkicus Munster, 1921, see: scoticus Joy and Tomlin, 1913.
foveifrons J. Sahlberg, 1913 [Note: Nomen nudum (Articles 1.3.4, 45.5)], see: cephalotes Gravenhorst, 1802.
fragilis Gravenhorst, 1802, see: subuliformis Gravenhorst, 1802.
friebi Bernhauer, 1927b: 95 (Philonthus; Type locality: Transbaikalien: Tschita).
— Scheerpeltz, 1933: 1343 (Philonthus; catalog).
— Coiffait, 1967a: 410 (Philonthus; characters; Transbaikal).
— Smetana, 1967d: 206 (Philonthus; characters; notes; Mongolia).
— Coiffait, 1974: 286 (Philonthus; characters; Transbaikal; Mongolia).
— Shilov, 1976: 151 (Philonthus; Mongolia).
— Schillhammer, 1991: 54 (Philonthus; difficult to distinguish from protenus).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: Russia, Mongolia.
fuentei Roubal, 1909: 100 (Philonthus; Type locality: Španělsko [Spain]: Pozuelo de Calatrava, Ciudad-Real.).
— Bernhauer and Schubert, 1914: 339 (Philonthus; catalog).
— Scheerpeltz, 1933: 1343 (Philonthus; catalog).
— Smetana, 1955b: 227 (Philonthus; characters).
— Coiffait, 1967a: 408 (Philonthus; characters; Spain).
— Coiffait, 1974: 285 (Philonthus; characters; Spain).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: Spain.
fulvipes Stephens, 1833, see: Philonthus.
fuscicornis Stephens, 1833, see: Philonthus.
fuscus Gravenhorst, 1802, see: subuliformis Gravenhorst, 1802.
germanus Sharp, 1874: 41 (Philonthus; Type locality: Nagasaki).
— Bernhauer and Schubert, 1914: 340 (Philonthus; catalog).
— Scheerpeltz, 1933: 1344 (Philonthus; catalog).
— Watanabe and Shibata, 1972: 64 (Philonthus; Yaku-shima Island).
— J. Li, 1993: 37 (Philonthus; China).
— Kanehisa, Tsumuki, and Kawazu, 1994 (Philonthus; chemical secretions).
— Smetana, 1995e: 516 (Bisnius).
— Watanabe, 1998: 313 (Philonthus; Japan).
▴ Distribution: Japan, China.
geryon Smetana, 1995e: 598 (Bisnius; Type locality: California: Upper Canyon Cr. Mdws., 16 mi. N. Junction City, 4800 feet, Trinity Co.).
hanae Vávra and Schillhammer, 1993, see: palmi Smetana, 1955.
heidenreichi Hubenthal, 1911, see: varipennis Scriba, 1864.
hesperidum Smetana, 1995e: 597 (Bisnius; Type locality: British Columbia: Goldstream Pk., 5 mi. N. Victoria. Also cited from Oregon, Washington).
howdeni Smetana, 1995e: 571 (Bisnius; Type locality: North Dakota: 5 mi. SW. McLeod).
hyperboreus J. Sahlberg, 1880: 70 (Philonthus; Type locality: Dudinka).
— Poppius, 1909: 24 (Philonthus; subgenus Gefyrobius; Lena Valley, Siberia).
— Bernhauer and Schubert, 1914: 341 (Philonthus; catalog).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Smetana, 1965f: 254 (Philonthus; characters; notes).
— Coiffait, 1967a: 411 (Philonthus; characters; central Asia; Siberia).
— Coiffait, 1974: 218 (Philonthus; characters; Siberia).
— Smetana 1975c: 171 (Philonthus; Mongolia).
— Shilov, 1976: 151 (Philonthus; Mongolia).
— Smetana, 1995e: 516 (Bisnius).
— Ryabukhin, 1999: 64 (Philonthus; catalog; habitat; distribution; northeastern Russia).
▴ Distribution: Russia, Mongolia, Canada, USA.
alaskensis Smetana, 1965a: 4 (Philonthus; subgenus Philonthus; subspecies of hyperboreus; Type locality: Alaska: Umiat).
— Smetana, 1995e: 754 (Bisnius; subspecies of hyperboreus; characters; notes; Manitoba; Northwest Territories; Alaska).
impressicollis Stephens, 1835, see: puella Nordmann, 1837.
impressifrons Broun, 1880, see: sordidus Gravenhorst, 1802.
inquietus Erichson, 1840: 928 (Philonthus; Type locality: America septentrionalis; [Note: See lectotype designation by Smetana, 1995e]).
— Horn, 1884: 210 (Philonthus; characters; South Carolina to Texas).
— Fauvel, 1891: 112 (Philonthus; Venezuela).
— Bernhauer and Schubert, 1914: 342 (Philonthus; catalog).
— Hunter, Kincher, Bay, Beerwinkle, 1991: 168 (Philonthus; seasonal distribution and diel flight activity; Texas).
— Smetana, 1995e: 552 (Bisnius; lectotype designation; characters; notes; Ontario; Quebec; Alabama; Arizona; Arkansas; Florida; Georgia; Illinois; Kansas; Kentucky; Louisiana; Maryland; Michigan; Mississippi; Missouri; New Jersey; North Carolina; Ohio; Oklahoma; South Carolina; Tennessee; Texas; Virginia).
▴ Distribution: Canada, USA, Venezuela.
instabilis Horn, 1884: 218 (Philonthus; Type locality: Colorado; [Note: See lectotype designation by Smetana, 1995e]. Also cited from Wyoming).
— Fall and Cockerell, 1907: 166 (Philonthus; New Mexico).
— Bernhauer and Schubert, 1914: 342 (Philonthus; catalog).
— Hatch, 1957: 183 (Philonthus; characters; British Columbia; Washington; Oregon).
— Smetana, 1995e: 599 (Bisnius; lectotype designation: Colorado; characters; notes; Alberta; British Columbia; Manitoba; Saskatchewan; Yukon Territory; Arizona; California; Colorado; Montana; Nebraska; Nevada; New Mexico; Oregon; Utah; Washington; Wyoming).
▴ Distribution: Canada, USA.
interpunctatus Hochhuth, 1860, see: fimetarius Gravenhorst, 1802.
intrudens Tottenham, 1949, see: longicollis Bernhauer, 1908.
inversus Horn, 1884: 239 (Philonthus; Type locality: Garland, Colorado; [Note: See lectotype designation by Smetana, 1995e]).
— Bernhauer and Schubert, 1914: 342 (Philonthus; catalog).
— Smetana, 1995e: 575 (Bisnius; lectotype designation; characters; notes; Colorado).
▴ Distribution: USA.
lautus Casey, 1915: 433 (Philonthus; Type locality: Wyoming: Cheyenne; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1348 (Philonthus; catalog).
— Smetana, 1995e: 538 (Bisnius; lectotype designation; characters; notes; Alberta; Ontario; Saskatchewan; Quebec; California; Colorado; Idaho; Oregon; Washington).
▴ Distribution: Canada, USA.
longicollis Bernhauer, 1908b: 33 (Philonthus; Type locality: Siberien).
— Bernhauer and Schubert, 1914: 344 (Philonthus; catalog).
— Scheerpeltz, 1933: 1349 (Philonthus; catalog).
— Coiffait, 1967a: 410 (Philonthus; characters; Siberia).
— Coiffait, 1974: 288 (Philonthus; characters; Siberia).
— Schillhammer, 1999a: 61 (Bisnius).
▴ Distribution: Russia, Czech Republic.
intrudens Tottenham, 1949a: 43 (Philonthus; Type locality: Beskid Mts.).
— Lohse, 1956: 88 (Philonthus; characters; notes).
— Smetana, 1958: 201 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 152 (Philonthus; characters).
— Lohse, 1964: 179 (Philonthus; subgenus Philonthus; characters; central Europe).
— Coiffait, 1967a: 410 (Philonthus; characters; Czechoslovakia).
— Coiffait, 1974: 289 (Philonthus; characters; Czechoslovakia).
— Tóth, 1984: 57 (Philonthus; characters; Hungary).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic ?).
— Smetana, 1995e: 516 (Bisnius).
— Schillhammer, 1999a: 61 (Bisnius; synonym of longicollis).
longipennis Provancher, 1877, see: sordidus Gravenhorst, 1802.
longulus Motschulsky, 1858, see: fimetarius Gravenhorst, 1802.
macies Sharp, 1874: 41 (Philonthus; Type locality: Nagasaki).
— Bernhauer and Schubert, 1914: 345 (Philonthus; catalog).
— Scheerpeltz, 1933: 1349 (Philonthus; catalog).
— Nakane, 1963a: 91 (Philonthus; characters; habitus photograph; Japan).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: Japan.
megacephalus Heer, 1839, see: cephalotes Gravenhorst, 1802.
microtophilus Coiffait, 1967a: 444 (Philonthus; subgenus Philonthus; Type locality: Mets Ichtkkan Sar, S.S.R. d'Armenia).
— Coiffait, 1974: 238 (Philonthus; characters; Armenia; Caucasus).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: Armenia.
minax Stephens, 1833, see: puella Nordmann, 1837.
negligens Smetana, 1966, see: agyrtes Smetana, 1973.
niger Melsheimer, 1844, see: sordidus Gravenhorst, 1802.
nigriventris Thomson, 1867: 147 (Philonthus; Type locality: Stehag och Båstad i Skåne).
— Fauvel, 1874: 220 [= 1874a: 446] (Philonthus; characters; Scandinavia; Scotland).
— Fowler, 1888: 268 (Philonthus; characters; Britain).
— Bernhauer and Schubert, 1914: 348 (Philonthus; catalog).
— Scheerpeltz, 1933: 1353 (Philonthus; catalog).
— Wüsthoff, 1937: 32 (Philonthus; illustration of aedeagus).
— V. Hansen, 1952: 61 (Philonthus; subgenus Philonthus; characters).
— Smetana, 1955b: 227 (Philonthus; characters).
— Lohse, 1956: 89 (Philonthus; characters; notes).
— Smetana, 1958: 206 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 153 (Philonthus; characters).
— Palm, 1963b: 78 (Philonthus; characters; Sweden; Norway; Finland).
— Lohse, 1964: 180 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 150 (Philonthus; Russia; Finland; Sweden; Norway; Britain; Germany; Austria; Czechoslovakia; Switzerland).
— Coiffait, 1967a: 405, 407 (Philonthus; characters).
— Coiffait, 1974: 239 (Philonthus; characters; Europe; Britain; Siberia).
— Shilov, 1975a: 376 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Pope, 1977: 29 (Philonthus; Britain).
— Tóth, 1984: 60 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Czechoslovakia; Austria; Denmark; Sweden).
— Wittwer, 1993: 250 (Philonthus; Switzerland).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Smetana, 1995e: 516 (Bisnius).
— Muona and Viramo, 1995: 9 (Philonthus; collected in dung of dog; Finland).
— Burakowski, Mroczkowski, and Stefańska, 2000: 50 (Philonthus; subgenus Philonthus; catalog; Poland).
▴ Distribution: Europe, Russia.
nitidulus Gravenhorst, 1802: 27 (Staphylinus; [Note: Preoccupied; although this name is a junior homonym of a species described in Staphylinus that is now in Tachyporus and although Tottenham, 1939d: 234, elected to list it as a junior synonym of Philonthus denigrator, no one has accepted this synonymy. However, Article 23.9.5 permits continued used of B. nitidulus (Gravenhorst) as a valid name]; Type locality: Not cited, but presumably Brunsvigae).
— Latreille, 1804: 315 (Staphylinus; characters; France; Germany).
— Gravenhorst, 1806: 69 (Staphylinus).
— Gyllenhal, 1810: 330 (Staphylinus; characters; Sweden).
— Gyllenhal, 1827: 479 (Staphylinus; characters).
— C. Sahlberg, 1830: 323 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 29 (Staphylinus; Finland).
— Mannerheim, 1831: 443 (Staphylinus; Finland).
— Runde, 1835: 8 (Staphylinus; characters; Germany).
— Nordmann, 1837: 85 (Philonthus; Germany; Sweden).
— Erichson, 1839a: 458 (Philonthus; characters; Germany).
— Heer, 1839: 262 (Philonthus; characters; Switzerland).
— Erichson, 1840: 450 (Philonthus; characters; Germany; Sweden).
— Redtenbacher, 1849: 702 (Philonthus; characters; Austria).
— Fairmaire and Laboulbène, 1856: 520 (Philonthus; characters; France).
— Kraatz, 1857d: 589 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 192 (Philonthus; characters; Austria).
— Thomson, 1859: 24 (type species of Gefyrobius).
— Thomson, 1860: 167 (Gefyrobius; characters; Scandinavia).
— Hochhuth, 1862: 57 (Philonthus; Russia).
— Fauvel, 1874: 249 [= 1874a: 475] (Philonthus; characters; France; Scandinavia; Germany; Switzerland; Austria; Tirol; Russia).
— Redtenbacher, 1874: 211 (Philonthus; characters; Austria).
— Seidlitz, 1875: 275 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 533 [= 1877: 389] (Gefyrobius; characters; France).
— J. Sahlberg, 1876: 22 (Gefyrobius; Finland).
— Ganglbauer, 1895: 449 (Philonthus; characters; north and middle Europe).
— Reitter, 1909: 128 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 348 (Philonthus; catalog).
— Johansen, 1914: 399 (Philonthus; characters; Denmark).
— Portevin, 1929: 361 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1353 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Tottenham, 1939d: 234 (Philonthus; synonym of denigrator; preoccupied).
— V. Hansen, 1952: 57 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1955b: 229 (Philonthus; cited as valid species; notes).
— Smetana, 1958: 188 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 149 (Philonthus; characters).
— Palm, 1963b: 73 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 179 (Philonthus; subgenus Philonthus; characters; central Europe).
— M. Dvořák, 1965: 89 (Philonthus; Slovakia).
— Horion, 1965: 141 (Philonthus; Finland; Sweden; Norway; Denmark; Russia; Ukraine; Baltics; Poland; Czechoslovakia; Hungary; Romania; Bulgaria; Italy; Switzerland ?; France ?; Germany; Austria).
— Coiffait, 1967a: 434 (Philonthus; subgenus Gefyrobius; characters).
— Szujecki, 1969a: 256 (Philonthus; notes; Poland).
— Tikhomirova, 1973: 183 (Philonthus; USSR).
— Coiffait, 1974: 192 (Philonthus; subgenus Gefyrobius; characters; Europe; Siberia; France).
— Shilov, 1976: 150 (Philonthus; Mongolia).
— Muona, 1979: 14 (Philonthus; Finland; Russia; Sweden; Norway; Denmark; Baltics).
— Burakowski, Mroczkowski, and Stefańska, 1980: 74 (Philonthus; catalog; Poland).
— Tóth, 1984: 54 (Philonthus; characters; Hungary).
— Drugmand, 1987a: 311 (Philonthus; characters; Belgium).
— Ádám, 1987: 151 (Gefyrobius; cited as valid species; collecting notes; Hungary).
— Drugmand, 1987b: 14, 43 (Philonthus; subgenus Gefyrobius; Belgium).
— Lucht, 1987: 106 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Denmark; Sweden).
— Nowosad, 1990: 111 (Philonthus; collected from mole nest; notes; Poland).
— Silfverberg, 1992: 16 (Philonthus; Finland; Russia; Sweden; Norway; Denmark; Estonia; Latvia; Lithuania).
— J. Li, 1993: 36 (Philonthus; China).
— Smetana, 1993b: 48 (Philonthus; subgenus Gefyrobius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy ?).
— Smetana, 1995e: 513 (Bisnius; cited as type species of Gefyrobius that is cited as a synonym of Bisnius).
— M. Hansen, 1996: 105 (Philonthus; Denmark).
— Herman, 2001: 43 (Bisnius; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Europe, Russia, Mongolia, China.
denigrator Gravenhorst, 1806: 92 (Staphylinus; [Note: This name was elevated to replace nitidulus by Tottenham, 1939d]; Type locality: Brunsvigae).
— Nordmann, 1837: 85 (Philonthus; Germany).
— Erichson, 1839a: 458 (Philonthus; synonym of nitidulus).
— Heer, 1839: 262 (Philonthus; synonym of nitidulus).
— Erichson, 1840: 450 (Philonthus; synonym of nitidulus).
— Kraatz, 1857d: 589 (Philonthus; synonym of nitidulus).
— Ganglbauer, 1895: 449 (Philonthus; synonym of nitidulus).
— Bernhauer and Schubert, 1914: 348 (Philonthus; synonym of nitidulus).
— Tottenham, 1939d: 234 (Philonthus; valid name for preoccupied nitidulus).
— Smetana, 1958: 188 (Philonthus; synonym of nitidulus).
— Coiffait, 1974: 192 (Philonthus; synonym of nitidulus).
— Muona, 1979: 14 (Philonthus; synonym of nitidus [Note: This synonymy may be an error; it was cited in a checklist without discussion and denigrator and nitidus have never been associated to my knowledge]).
noualhieri Fauvel, 1898: 99 (Philonthus; Type locality: Biskra).
— Fauvel, 1902b: 110 (Philonthus; Algeria).
— Bernhauer and Schubert, 1914: 348 (Philonthus; catalog).
— Scheerpeltz, 1933: 1353 (Philonthus; catalog).
— Coiffait, 1967a: 403 (Philonthus; characters; Algeria).
— Coiffait, 1974: 234 (Philonthus; characters; Algeria; Morocco).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: Algeria, Morocco.
pachycephalus Nordmann, 1837, see: sordidus Gravenhorst, 1802.
paederinus Sachse, 1852, see: blandus Gravenhorst, 1806.
palmi Smetana, 1955b: 224 (Philonthus; Type locality: Sicilien, Caltagirone).
— Coiffait, 1967a: 407 (Philonthus; characters; Sicily).
— Coiffait, 1974: 284 (Philonthus; characters; Sicily).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; introduced in Italy).
— Smetana, 1995e: 535 (Bisnius; characters; notes; apparently introduced to Sicily; Nova Scotia; Ontario; Quebec; Connecticut; Florida; Idaho; Illinois; Iowa; Maryland; Massachusetts; New Hampshire; New Jersey; New York; Oklahoma; Pennsylvania).
▴ Distribution: Italy, Czech Republic, Canada, USA.
hanae Vávra and Schillhammer, 1993: 35 (Philonthus; Type locality: CR: Moravia mer., Břeclav).
palousi Hatch, 1957: 193 (Philonthus; subspecies of pugetensis; Type locality: Washington: Pullman. Also cited from Oregon and Idaho).
— Smetana, 1995e: 570 (Bisnius; valid species; characters; notes; New Mexico; Oregon; Washington).
▴ Distribution: USA.
parcus Sharp, 1874: 40 (Philonthus; Type locality: Hiogo).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Scheerpeltz, 1933: 1354 (Philonthus; catalog).
— Smetana 1975c: 171 (Philonthus; Mongolia).
— Brunne, 1976: 86 (Philonthus; characters).
— Shilov, 1976: 151 (Philonthus; Mongolia).
— Pope, 1977: 29 (Philonthus; Britain).
— Mahler and Pritzl, 1980: 122 (Philonthus; characters).
— Lohse and Lucht, 1989: 166 (Philonthus; characters).
— Kuschel, 1990: 26 (Philonthus; introduced, Palaearctic species; winged; habitat; New Zealand).
— Schillhammer, 1991: 54 (Philonthus; difficult to distinguish from protenus).
— Schillhammer, 1993: 326 (Philonthus; Austria).
— Pilon and Zanetti, 1991: 58 (Philonthus; notes; Italy).
— J. Li, 1993: 33 (Philonthus; China).
— J. Li, 1993: 36 (Philonthus; China).
— Hodge and Jones, 1995: 39 (Philonthus; characters; Britain).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; introduced in Italy).
— Smetana, 1995e: 532 (Bisnius; characters; notes; British Columbia; Ontario; Quebec; Arizona; California; Colorado; Illinois; Indiana; Missouri; Nevada; New Jersey; New York; New Mexico; North Carolina; Oklahoma; Oregon; Pennsylvania; Tennessee; Utah; Washington).
— Watanabe, 1996a: 12 (Philonthus; cave collection; Japan).
— Tronquet, 1998: 10 (Philonthus; notes; France).
— Schillhammer, 1999a: 61 (Bisnius).
— Ødegaard, 1999: 72 (Philonthus; invasive species associated with compost heaps in Sweden, Norway, Denmark, Finland).
— Burakowski, Mroczkowski, and Stefańska, 2000: 50 (Philonthus; subgenus Philonthus; catalog; Poland).
▴ Distribution: Britain, Austria, Italy, Mongolia, China, Japan, New Zealand, Canada, USA.
subaereipennis Bernhauer, 1939d: 97 (Philonthus; Type locality: Nordwestl. China: Chinkiang).
parumpunctatus Erichson, 1840, see: puella Nordmann, 1837.
pentheri Ganglbauer, 1905: 273 (Philonthus; Type locality: Erdschias-Gebiet).
— Bernhauer and Schubert, 1914: 350 (Philonthus; catalog).
— Scheerpeltz, 1933: 1355 (Philonthus; catalog).
— Kirshenblat, 1950: 241 (Philonthus; characters in key).
— Coiffait, 1967a: 401 (Philonthus; characters; Asia Minor; Armenia).
— Coiffait, 1974: 233 (Philonthus; characters; Asia Minor; Armenia).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: Turkey, Armenia.
pholeophilus Kirshenblat, 1950: 237 (Philonthus; Type locality: Amurskaia oblast': okr. Blagoveshchenska).
— Smetana, 1963c: 296 (Philonthus; subgenus Philonthus; characters; Mongolia).
— Coiffait, 1967a: 405 (Philonthus; characters; Siberia; Mongolia).
— Coiffait, 1974: 282 (Philonthus; characters; East Asia).
— Smetana 1975c: 172 (Philonthus; Mongolia).
— Shilov, 1976: 152 (Philonthus; Mongolia).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: Mongolia, Russia.
picicornis Horn, 1884: 218 (Philonthus; Type locality: Lake Tahoe, Nevada; [Note: See lectotype designation by Smetana, 1995e]).
— Bernhauer and Schubert, 1914: 350 (Philonthus; catalog).
— Hatch, 1957: 184 (Philonthus; characters; British Columbia; Washington; Idaho; Oregon).
— Smetana, 1995e: 562 (Bisnius; lectotype designation; characters; notes; British Columbia; Nevada; Washington).
▴ Distribution: Canada, USA.
zetes Smetana, 1995e: 563 (Bisnius; subspecies of picicornis; Type locality: Arizona: Chuska Mts., 2600 m, Apache Co. Also cited from New Mexico).
placidus Erichson, 1840, see: sordidus Gravenhorst, 1802.
plagiatus Fauvel, 1874: 222 [= 1874a: 448] note (Philonthus; Type locality: Algérie).
— Fauvel, 1876a: 73 [= 1876: 258] (Philonthus; Tunisia).
— Fauvel, 1878: 122 [= 1878a: 42] (Philonthus; Algeria; Tunisia).
— Fauvel, 1886: 54 [= 1886a: 46] (Philonthus; Tunisia; Algeria).
— Fauvel, 1902b: 109 (Philonthus; Algeria; Tunisia; Tripolitania).
— Bernhauer and Schubert, 1914: 350 (Philonthus; catalog).
— Gridelli, 1930: 73 (Philonthus; Libya).
— Scheerpeltz, 1933: 1355 (Philonthus; catalog).
— C. Koch, 1936a: 75 (Philonthus; Libya).
— Kirshenblat, 1950: 241 (Philonthus; characters in key).
— Coiffait, 1967a: 401 (Philonthus; characters; North Africa).
— Coiffait, 1974: 233 (Philonthus; characters; North Africa).
— Smetana, 1995e: 516 (Bisnius).
— Schillhammer, 1999a: 61 (Bisnius).
▴ Distribution: Algeria, Tunisia, Libya, Turkmenistan, Uzbekistan.
sogdianus Kirshenblat, 1950: 239 (Philonthus; Type locality: Turkmeniia: Ashkhabad; st. Dzhebel; Uzbekistan: Golodnaia step'; Kuropatkino).
— Coiffait, 1967a: 401 (Philonthus; characters; Turkmenia; Uzbekistan).
— Coiffait, 1974: 234 (Philonthus; characters; Turkestan; Uzbekistan).
— Smetana, 1995e: 516 (Bisnius).
— Schillhammer, 1999a: 61 (Bisnius; synonym of plagiatus).
poppaeus Smetana, 1995e: 593 (Bisnius; Type locality: California: Inverness).
potanini Eppelsheim, 1889a: 177 (Philonthus; subgenus Gefyrobius; Type locality: Kan-ssu).
— Bernhauer and Schubert, 1914: 351 (Philonthus; catalog).
— Scheerpeltz, 1933: 1356 (Philonthus; catalog; Tibet).
— Coiffait, 1967a: 403 (Philonthus; characters; China; Tibet).
— Coiffait, 1974: 212 (Philonthus; characters; China; Tibet).
— Shilov, 1976: 150 (Philonthus; notes).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: China.
propinquus Kirschenblat, see: attiguus Herman, 2001.
pseudoparcus Brunne, 1976: 87 (Philonthus; Type locality: Trittau bei Hamburg, Forst Hahnheide).
— Lundberg, 1972: 173 (Philonthus; [Note: The name is a nomen nudum here]; Sweden).
— Mahler and Pritzl, 1980: 121 (Philonthus; notes; characters; Denmark).
— Lohse and Lucht, 1989: 166 (Philonthus; characters; Norway).
— Schillhammer, 1993: 326 (Philonthus; Austria).
— Hodge and Jones, 1995: 40 (Philonthus; characters; Britain).
— Smetana, 1995e: 516 (Bisnius).
— Assing, Frisch, Kahlen, et al., 1998: 138 (Philonthus; Austria).
— Ødegaard and Ligaard, 2000: 10 (Philonthus; notes; Norway).
— Burakowski, Mroczkowski, and Stefańska, 2000: 51 (Philonthus; subgenus Philonthus; catalog; Poland).
▴ Distribution: Britain, Denmark, Austria, Germany.
psyllophagus Kirshenblat, 1933: 102 (Philonthus; Type locality: Transbaikalien: Adun-Tshelon).
— Kirshenblat, 1950: 240 (Philonthus; characters in key).
— Smetana, 1963c: 296 (Philonthus; subgenus Philonthus; Mongolia).
— Coiffait, 1967a: 407 (Philonthus; characters; Transbaikal; Mongolia).
— Smetana, 1967d: 207 (Philonthus; Mongolia).
— Coiffait, 1974: 283 (Philonthus; characters; Transbaikal; Mongolia).
— Smetana 1975c: 172 (Philonthus; Mongolia).
— Shilov, 1976: 152 (Philonthus; Mongolia).
— Boháč, 1988: 551 [= 1989: 34] (Philonthus; cited as psyllophaqus; characters).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: Russia, Mongolia.
puella Nordmann, 1837: 101 (Philonthus; Type locality: Berolinum).
— Nordmann, 1837: 106 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 351 cited punctus Nordmann as a synonym of puella Nordmann; Nordmann did not describe the species as new, he attributed it to Gyllenhal who cited Gravenhorst]; characters; Sweden; Finland).
— Erichson, 1839a: 472 (Philonthus; characters; Germany).
— Erichson, 1840: 493 (Philonthus; characters; Germany).
— Kraatz, 1857d: 620 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 198 (Philonthus; characters; Germany).
— Thomson, 1860: 162 (Philonthus; characters; Scandinavia).
— Fauvel, 1874: 220 [= 1874a: 446] (Philonthus; characters; France; Scandinavia; Britain; Germany; Switzerland; Austria; Russia).
— Redtenbacher, 1874: 215 (Philonthus; characters; Austria).
— Seidlitz, 1875: 278 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 518 [= 1877: 374] (Philonthus; characters; France).
— J. Sahlberg, 1876: 19 (Philonthus; Finland).
— Fowler, 1888: 280 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 458 (Philonthus; characters; north and middle Europe).
— Reitter, 1909: 131 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 351 (Philonthus; catalog).
— Johansen, 1914: 414 (Philonthus; characters; Denmark).
— Bernhauer, 1926g: 2 (Philonthus; Kamchatka region).
— Portevin, 1929: 364 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1356 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Tottenham, 1949: 419 (Philonthus; checklist; Britain).
— V. Hansen, 1952: 78 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1958: 223 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 170 (Philonthus; characters).
— Palm, 1963b: 96 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 186 (Philonthus; subgenus Philonthus; characters; central Europe).
— Raianu, 1964: 103 (Philonthus; Romania).
— Horion, 1965: 173 (Philonthus; Russia; Finland; Sweden; Norway; Britain; Denmark; Baltics; Holland; Urals; Poland; Czechslovakia; Romania; Bosnia; Austria; Italy; Switzerland; France; Germany).
— Coiffait, 1967a: 432 (Philonthus; characters; Europe).
— Raianu, 1968a: 468 (Philonthus; Romania).
— Scheerpeltz, 1968a: 73 (Philonthus; catalog; Austria).
— Kasule, 1970: 68 (Philonthus; larval characters).
— Coiffait, 1974: 219 (Philonthus; characters; Europe; Russia; Britain).
— Shilov, 1975a: 376 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Pope, 1977: 29 (Philonthus; Britain).
— Muona, 1979: 14 (Philonthus; Finland; Russia; Sweden; Norway; Denmark; Baltic States).
— Boháč, 1980: 86 (Philonthus; Caucasus).
— Burakowski, Mroczkowski, and Stefańska, 1980: 62 (Philonthus; catalog; Poland).
— Hinton, 1981: 660 (Philonthus; characters of egg).
— Boller, 1983: 120 (Philonthus; larval characters).
— Tóth, 1984: 77 (Philonthus; characters).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Allen, 1988: 194 (Philonthus; collecting notes; Britain).
— Drugmand, 1992: 33 (Philonthus; characters; notes; Belgium).
— Silfverberg, 1992: 16 (Philonthus; Finland; Russia; Sweden; Norway; Denmark; Estonia; Latvia; Lithuania).
— Wittwer, 1993: 250 (Philonthus; Switzerland).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Siitonen, 1993: 230 (Philonthus; collecting notes; Finland).
— Smetana, 1995e: 516 (Bisnius).
— Muona and Viramo, 1995: 9 (Philonthus; collected from dung of Ursus arctos; Finland).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— M. Hansen, Mahler, Palm, and Pedersen, 1996: 244 (Philonthus; Denmark).
— M. Hansen, 1996: 106 (Philonthus; checklist; Denmark).
— Sterrenburg and Schülke, 1997: 18 (Philonthus; Netherlands).
— Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 37 (Philonthus; Latvia).
— Anderson, Nash, and O'Connor, 1997: 19 (Philonthus; Ireland).
— Herman, 2001: 24 (Bisnius; nomen protectum).
▴ Distribution: Europe, Russia, Caucasus, Turkey.
watsoni Stephens, 1832: 240 (Philonthus; [Note: This name is older than puella]; Type locality: London; Yorkshire; Suffolk).
— Curtis, 1829: 26 (Philonthus; nomen nudum; catalog; Britain).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Stephens, 1839: 398 (Philonthus; characters; England).
— Hardy, 1851: 31 (Philonthus; synonym of minax).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of puella).
— Bernhauer and Schubert, 1914: 351 (Philonthus; synonym of puella).
— Smetana, 1958: 223 (Philonthus; synonym of puella).
— Coiffait, 1974: 219 (Philonthus; synonym of puella).
— Herman, 2001: 24 (Bisnius; nomen oblitum).
minax Stephens, 1833: 241 (Philonthus; [Note: This name is older than puella]; Type locality: near Hull, in Yorkshire).
— Curtis, 1829: 26 (Philonthus; nomen nudum; catalog; Britain).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Stephens, 1839: 398 (Philonthus; characters; England).
— Hardy, 1851: 31 (Philonthus; notes; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of puella).
— Bernhauer and Schubert, 1914: 351 (Philonthus; synonym of puella).
— Smetana, 1958: 223 (Philonthus; synonym of puella).
— Coiffait, 1974: 219 (Philonthus; synonym of puella).
— Herman, 2001: 24 (Bisnius; nomen oblitum).
impressicollis Stephens, 1835: 436 (Philonthus; [Note: This name is older than puella]; Type locality: New Forest).
— Stephens, 1839: 398 (Philonthus; characters; England).
— Hardy, 1851: 31 (Philonthus; synonym of minax).
— Herman, 2001: 24 (Bisnius; nomen oblitum).
parumpunctatus Erichson, 1840: 499 (Philonthus; Type locality: Sweden; montes Uralenses).
— Jacobsen, 1849: 705 (Philonthus; characters; Denmark).
— Hardy, 1851: 31 (Philonthus; synonym of minax).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of puella).
— Fairmaire and Laboulbène, 1856: 529 (Philonthus; characters; France).
— Kraatz, 1857d: 620 (Philonthus; synonym of puella).
— Redtenbacher, 1857: 197 (Philonthus; characters; Austria).
— Peyron, 1858: 427 (Philonthus; Turkey).
— Fauvel, 1874: 220 [= 1874a: 446] (Philonthus; synonym of puella).
— Ganglbauer, 1895: 458 (Philonthus; synonym of puella).
— Bernhauer and Schubert, 1914: 351 (Philonthus; synonym of puella).
— Scheerpeltz, 1933: 1356 (Philonthus; synonym of puella).
— Smetana, 1958: 223 (Philonthus; synonym of puella).
— Coiffait, 1974: 219 (Philonthus; synonym of puella).
pugetensis Hatch, 1957: 192 (Philonthus; Type locality: Washington: Seattle. Also cited from Oregon).
— Smetana, 1995e: 567 (Bisnius; characters; notes; British Columbia; Manitoba; Ontario; Quebec; Saskatchewan; Indiana; Massachusetts; New Hampshire; New York; Oregon; Pennsylvania; Washington).
▴ Distribution: Canada, USA.
pulchellus Melsheimer, 1844, see: blandus Gravenhorst, 1806.
puncticollis Stephens, 1835, see: Philonthus.
quediinus Horn, 1884: 209 (Philonthus; Type locality: Detroit; [Note: See lectotype designation by Smetana, 1995e]. Also cited from Kansas).
— Bernhauer and Schubert, 1914: 352 (Philonthus; catalog).
— Smetana, 1995e: 541 (Bisnius; lectotype designation: Detroit; characters; notes; Quebec; Kansas; Massachusetts; Michigan; New Hamphire; South Dakota).
— Downie and Arnett, 1996: 380 (Philonthus; characters; USA).
▴ Distribution: Canada, USA.
rainieri Hatch, 1957: 194 (Philonthus; Type locality: Washington: Mt. Rainier, Paradise Val., 5–9000 ft. Also cited from Oregon).
— Smetana, 1995e: 592 (Bisnius; characters; notes; British Columbia; Oregon; Washington).
▴ Distribution: USA.
reitteri Eppelsheim, 1889d: 18 (Philonthus; Type locality: Circassien).
— Bernhauer and Schubert, 1914: 353 (Philonthus; catalog).
— Scheerpeltz, 1933: 1358 (Philonthus; catalog).
— Smetana, 1955: 206 (Philonthus; notes).
— Coiffait, 1967a: 411 (Philonthus; characters; Caucasus).
— Fagel, 1968a: 193 (Philonthus; Anatolia).
— Korge, 1971a: 7 (Philonthus; Turkey).
— Coiffait, 1974: 215 (Philonthus; characters; Caucasus; Transcaucasus).
— Smetana, 1977: 794 (Philonthus; Turkey).
— Smetana, 1995e: 516 (Bisnius).
— Solodovnikov, 1998a: 15 (Bisnius; notes; Russia, Krasnodar territory).
▴ Distribution: Russia, Caucasus, Turkey.
rigidicornis Gravenhorst, 1802, see: fimetarius Gravenhorst, 1802.
rotundiceps Stephens, 1833, see: Philonthus.
rubrithorax Kirshenblat, 1950, see: subuliformis Gravenhorst, 1802.
rufipennis Delahon, 1914 [Note: Nomen nudum], see: cephalotes Gravenhorst, 1802.
scoticus Joy and Tomlin, 1913: 100 (Philonthus; Type locality: Creag Dhu, 2581 feet, Kingussie, Inverness-shire).
— Donisthorpe, 1930: 96 (Philonthus; characters; Britain).
— Scheerpeltz, 1933: 1360 (Philonthus; catalog).
— Wüsthoff, 1937: 32 (Philonthus; illustration of aedeagus).
— Smetana, 1955b: 227 (Philonthus; characters).
— Palm, 1963b: 78 (Philonthus; characters; Sweden; Norway; Finland).
— Coiffait, 1967a: 407 (Philonthus; characters; Scandinavia; north British Islands).
— Coiffait, 1974: 211 (Philonthus; characters; Scandinavia; Scotland).
— Pope, 1977: 30 (Gabrius; Britain).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: Britain, Norway, Sweden, Finland.
finmarkicus Munster, 1921: 55 (Philonthus; Type locality: Norvegia arctica [many localities]).
— Scheerpeltz, 1933: 1360 (Philonthus; synonym of scoticus).
— Coiffait, 1974: 211 (Philonthus; synonym of scoticus).
— Pope, 1977: 30 (Gabrius; synonym of scoticus).
scribae Fauvel, 1867, see: varipennis Scriba, 1864.
siegwaldii Mannerheim, 1843b: 230 (Philonthus; Type locality: insula Sitkha; [Note: See lectotype designation by Smetana, 1995e]).
— Horn, 1884: 213 (Philonthus; characters; Alaska south to California).
— Bernhauer and Schubert, 1914: 355 (Philonthus; catalog).
— Van Dyke, 1924: 17 (Philonthus; Alaska).
— Fall, 1926: 150 (Philonthus; Alaska).
— Hatch, 1957: 183 (Philonthus; characters; British Columbia; Washington; Idaho).
— Smetana, 1964: 35 (Philonthus; subgenus Philonthus; hind wings; Alaska).
— Smetana, 1965: 27 (Philonthus; subgenus Philonthus; characters; Labrador; Newfoundland).
— Smetana, 1965a: 8 (Philonthus; Alaska).
— Miller and Morgan, 1982: 264 (Philonthus; Postglacial fossils; New York).
— Smetana, 1995e: 547 (Bisnius; lectotype designation; characters; notes; Alberta; British Columbia; Manitoba; Newfoundland; Labrador; New Brunswick; Northwest Territories; Nova Scotia; Ontario; Quebec; Saskatchewan; Yukon Territory; Alaska; California; Connecticut; Maine; Michigan; Montana; New Hampshire; New Jersey; New York; North Carolina; Oregon; Tennessee; Vermont; Washington; West Virginia; Wisconsin).
— Downie and Arnett, 1996: 381 (Philonthus; characters; USA).
— Ryabukhin, 1999: 64 (Philonthus; catalog; habitat; distribution; northeastern Russia).
▴ Distribution: Canada, USA, Russia.
simplex Stephens, 1833, see: Philonthus.
sogdianus Kirshenblat, 1950, see: plagiatus Fauvel, 1874.
sordidus Gravenhorst, 1802: 176 (Staphylinus; Type locality: Brunsvigae; [Note: See lectotype designation by Smetana, 1995e]).
— Latreille, 1804: 325 (Staphylinus; characters).
— Gyllenhal, 1810: 326 (Staphylinus; characters; Sweden).
— Zetterstedt, 1828: 75 (Staphylinus; characters; Lapland).
— C. Sahlberg, 1830: 322 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 29 (Staphylinus; Finland; Sweden).
— Mannerheim, 1831: 443 (Staphylinus; Finland; Sweden).
— Stephens, 1832: 233 (Philonthus; characters; England).
— Lacordaire, 1835: 396 (Staphylinus; characters; France).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Nordmann, 1837: 84 (Philonthus; characters; Sweden; Germany).
— Zetterstedt, 1838: 63 (Staphylinus; characters; habitat; Lapland).
— Erichson, 1839a: 459 (Philonthus; characters; Germany).
— Heer, 1839: 262 (Philonthus; characters; Switzerland).
— Stephens, 1839: 396 (Philonthus; characters; England).
— Erichson, 1840: 456 (Philonthus; characters; Europe).
— P. Lucas, 1846: 112 (Philonthus; notes; Algeria).
— Jacobsen, 1849: 695 (Philonthus; characters; Denmark).
— Redtenbacher, 1849: 825 (Philonthus; characters; Austria).
— Kiesenwetter, 1851a: 417 (Philonthus; France).
— Hardy, 1851: 28 (Philonthus; notes; England).
— Wollaston, 1854: 582 (Philonthus; characters; notes; Madeira Islands).
— Rosenhauer, 1856: 71 (Philonthus; Malaga).
— Fairmaire and Laboulbène, 1856: 519 (Philonthus; characters; France).
— Staudinger, 1857: 286 (Philonthus; notes; Iceland).
— Wollaston, 1857: 189 (Philonthus; habitat; Madeira Islands).
— Kraatz, 1858a: 61 (Philonthus; Greece).
— Redtenbacher, 1857: 190 (Philonthus; characters; Austria).
— Thomson, 1860: 159 (Philonthus; characters; Scandinavia).
— Hochhuth, 1862: 58 (Philonthus; Siberia; Dauria).
— Wollaston, 1864: 569 (Philonthus; notes; Canary Islands).
— Wollaston, 1865: 491 (Philonthus; notes; Madeira Islands; Canary Islands).
— Fauvel, 1866a: 344 [= 1866b: 95] (Philonthus; characters; Chile).
— Redtenbacher, 1867: 29 (Philonthus; Madeira).
— Crotch, 1867a: 383 (Philonthus; Azores).
— Fauvel, 1874: 222 [= 1874a: 448] (Philonthus; characters; France; Barbary; Cyprus; Turkey; Caucasus; Iran; Siberia; Dauria; Amur region; North America; Chile).
— Redtenbacher, 1874: 209 (Philonthus; characters; Austria).
— Seidlitz, 1875: 274 (Philonthus; characters; Baltic region).
— Fauvel, 1876a: 73 [= 1876: 258] (Philonthus; Egypt; Syria).
— Mulsant and Rey, 1876: 434 [= 1877: 290] (Philonthus; larval characters; characters; France).
— J. Sahlberg, 1876: 16 (Philonthus; Finland).
— Fauvel, 1878: 122 [= 1878a: 42] (Philonthus; Algeria; Tunisia).
— Fauvel, 1878g: 546 (Philonthus; characters; Australia).
— Horn, 1884: 209 (Philonthus; characters; Canada; Michigan; Colorado; Vancouver; California).
— Fauvel, 1886: 53 [= 1886a: 45] (Philonthus; North Africa).
— Fowler, 1888: 268 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 451 (Philonthus; characters; Palaearctic region; North America; Chile).
— Fauvel, 1902b: 109 (Philonthus; Barbary; Egypt; Azores; Madeira; Canaries).
— Fall and Cockerell, 1907: 166 (Philonthus; New Mexico).
— Sainte-Claire Deville, 1906: 96 (Philonthus; Corsica).
— Montandon, 1908: 80 (Philonthus; Romania).
— Poppius, 1909: 26 (Philonthus; subgenus Gefyrobius; Lena Valley, Siberia).
— Reitter, 1909: 127 (Philonthus; characters; Germany).
— Blatchley, 1910: 388 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 355 (Philonthus; catalog).
— Johansen, 1914: 402 (Philonthus; characters; Denmark).
— Bernhauer, 1920: 16 (Philonthus; Australia).
— Gridelli, 1924e: 177 (Philonthus; Persia).
— Lea, 1925: 220 (Philonthus; Australian introduction).
— Dodero, 1925: 5 (Philonthus; Libya).
— Fall, 1926: 150 (Philonthus; Alaska).
— Portevin, 1929: 359 (Philonthus; characters; France).
— Gridelli, 1930: 73 (Philonthus; Libya).
— Cameron, 1932: 93 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1361 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Bernhauer, 1936: 5 (Philonthus; Mongolia).
— Gridelli, 1934: 74 (Philonthus; Karakoram region).
— Bernhauer, 1940c: 4 (Philonthus; Azores).
— Méquignon, 1942a: 20 (Philonthus; Azores).
— Hinton, 1945: 75 (Philonthus; characters).
— Scheerpeltz and Höfler, 1948: 156 (Philonthus; characters; fungus inhabitant).
— V. Hansen, 1952: 62 (Philonthus; subgenus Philonthus; characters; Denmark).
— Lee and Ryckman, 1954: 24 (Philonthus; reared from burrowing owl's nest; California).
— Coiffait, 1955e: 426 (Philonthus; cave collection; Lebanon).
— Smetana, 1955b: 227 (Philonthus; characters).
— Hatch, 1957: 194 (Philonthus; characters; British Columbia; Idaho; Washington; Oregon).
— Smetana, 1958: 203 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 154 (Philonthus; characters).
— Larsson and Gígja, 1959: 85 (Philonthus; biological notes; Iceland).
— Scheerpeltz, 1961d: 129 (Philonthus; Italy).
— Scheerpeltz, 1961e: 137 (Philonthus; subgenus Gefyrobius; Rhodes).
— Scheerpeltz, 1961h: 252 (Philonthus; Afghanistan).
— Smetana, 1963b: 36 (Philonthus; subgenus Philonthus; Madeira).
— Smetana, 1963e: 73 (Philonthus; characters; notes).
— Scheerpeltz, 1963: 13 (Philonthus; Afghanistan).
— Palm, 1963b: 78 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 181 (Philonthus; subgenus Philonthus; characters; central Europe).
— Scheerpeltz, 1964a: 300 (Philonthus; subgenus Philonthus; Crete).
— Scheerpeltz, 1965b: 402 (Philonthus; Sicily).
— Scheerpeltz, 1965c: 506 (Philonthus; Sicily).
— Smetana, 1965: 17 (Philonthus; St. Pierre; Newfoundland).
— Horion, 1965: 152 (Philonthus; all Europe; Madeira; Canaries; Azores).
— Smetana, 1967d: 207 (Philonthus; Mongolia).
— Raianu, 1968a: 467 (Philonthus; Romania).
— Szujecki, 1968a: 731 (Philonthus; Poland).
— Smetana, 1970: 59 (Philonthus; distributional summary; Azores).
— Scheerpeltz, 1972: 18 (Philonthus; Argentina).
— Coiffait, 1974: 239 (Philonthus; synonym of pachycephalus).
— Shilov, 1975a: 376 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Brunne, 1976: 87 (Philonthus; characters).
— Shilov, 1976: 152 (Philonthus; Mongolia).
— Pope, 1977: 29 (Philonthus; Britain).
— Smetana, 1978: 117 (Philonthus; collecting notes; Israel).
— Muona, 1979: 14 (Philonthus; synonym of pachycephalus).
— Mahler and Pritzl, 1980: 122 (Philonthus; characters).
— Burakowski, Mroczkowski, and Stefańska, 1980: 66 (Philonthus; catalog; Poland).
— Hinton, 1981: 660 (Philonthus; characters of egg).
— Coiffait, 1982: 29 (Philonthus; synonym of pachycephalus).
— Frank, 1982: 43 (Philonthus; list of parasites).
— Boller, 1983: 117 (Philonthus; larval characters).
— Tóth, 1984: 62 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Lohse and Lucht, 1989: 168 (Philonthus; characters).
— Nowosad, 1990: 107 (Philonthus; collected from mole nest; notes; Poland).
— Kuschel, 1990: 26 (Philonthus; introduced, Palaearctic species; winged; habitat; New Zealand).
— Welch, 1993: 228 (Philonthus; ovariole number and ovary structure).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Dearborn and Donahue, 1994: 15 (Philonthus; Maine).
— Ciceroni and Zanetti, 1995: 29 (Philonthus; Italy).
— Smetana, 1995e: 523 (Bisnius; lectotype designation; characters; notes; Alberta; British Columbia; Manitoba; New Brunswick; Newfoundland; Nova Scotia; Ontario; Quebec; Saskatchewan; St. Pierre; Alaska; Arizona; Arkansas; California; Colorado; Connecticut; Georgia; Idaho; Illinois; Indiana; Iowa; Kansas; Kentucky; Maine; Maryland; Massachusetts; Michigan; Minnesota; Missouri; Montana; Nebraska; Nevada; New Hampshire; New Jersey; New Mexico; New York; North Carolina; Ohio; Oklahoma; Oregon; Pennsylvania; Rhode Island; Utah; Virginia; Washington; Wisconsin; Wyoming).
— Ádám, 1996: 245 (Bisnius; synonym of pachycephalus).
— Owen, 1997a: 149 (Philonthus; collecting sites; Ireland).
— Emberson, 1998: 36 (Philonthus; collecting notes; Chatham Islands).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
▴ Distribution: Syria, Lebanon, Israel, Egypt, Libya, Tunisia, Algeria, Canary Islands, Madeira, Azores, Iceland, Europe, Russia, Caucasus, Turkey, Iran, Kazakhstan, Afghanistan, “Karakorum region”, India, Nepal, Mongolia, Australia, New Zealand, Chatham Islands, Canada, USA, Argentina, Chile.
pachycephalus Nordmann, 1837: 82 (Philonthus; Type locality: Borussia).
— Erichson, 1839a: 459 (Philonthus; synonym of sordidus).
— Heer, 1839: 263 (Philonthus; synonym of sordidus).
— Erichson, 1840: 457 (Philonthus; synonym of sordidus).
— Kraatz, 1857d: 591 (Philonthus; synonym of sordidus).
— Fauvel, 1866a: 344 [= 1866b: 95] (Philonthus; synonym of sordidus).
— Ganglbauer, 1895: 451 (Philonthus; synonym of sordidus).
— Bernhauer and Schubert, 1914: 355 (Philonthus; synonym of sordidus).
— Cameron, 1932: 92 (Philonthus; synonym of sordidus).
— Smetana, 1958: 203 (Philonthus; synonym of sordidus).
— Coiffait, 1967a: 408 (Philonthus; characters).
— Coiffait, 1968c: 502 (Philonthus; cave collection; Corsica).
— Coiffait, 1974a: 198 (Philonthus; Siberia).
— Coiffait, 1974: 239 (Philonthus; characters; Holarctic region; Australia).
— Pace and Zanetti, 1977: 297 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; synonym of sordidus).
— Bordoni, 1978a: 59 (Philonthus; cave collection; Turkey).
— Coiffait, 1979d: 552 (Philonthus; Afghanistan).
— Muona, 1979: 14 (Philonthus; cited as valid species).
— Coiffait, 1982: 29 (Philonthus; cited as valid species; India; Nepal).
— Coiffait, 1982a: 232 (Philonthus; cited as valid species; India).
— Coiffait, 1983: 274 (Philonthus; cited as valid species; Kashmir).
— Coiffait, 1983e: 477 (Philonthus; Afghanistan).
— Lohse and Lucht, 1989: 168 (Philonthus; synonym of sordidus).
— Borges, 1990: Table II (Philonthus; Azores).
— Smetana, 1993b: 48 (Philonthus; synonym of sordidus).
— Sadler and Dugmore, 1995: 144 (Philonthus; habitat notes; Iceland).
— Ciceroni and Zanetti, 1995: 29 (Philonthus; synonym of sordidus).
— Smetana, 1995e: 523 (Bisnius; synonym of sordidus).
— Ádám, 1996: 245 (Bisnius; valid name; Hungary).
placidus Erichson, 1840: 458 (Philonthus; Type locality: Chersonesus Tauricus; Hungaria).
— Kraatz, 1857d: 591 (Philonthus; synonym of sordidus).
— Hochhuth, 1862: 58 (Philonthus; Dauria).
— Ganglbauer, 1895: 451 (Philonthus; synonym of sordidus).
— Bernhauer and Schubert, 1914: 356 (Philonthus; synonym of sordidus).
— Cameron, 1932: 92 (Philonthus; synonym of sordidus).
— Smetana, 1958: 203 (Philonthus; synonym of sordidus).
— Coiffait, 1974: 240 (Philonthus; synonym of pachycephalus).
— Smetana, 1995e: 523 (Bisnius; synonym of sordidus).
niger Melsheimer, 1844: 38 (Philonthus; [Note: Preoccupied secondary homonym in Philonthus]; Type locality: Pennsylvania; [Note: See lectotype designation by Smetana, 1995e]).
— Ganglbauer, 1895: 454 (Philonthus; synonym of agilis).
— Bernhauer and Schubert, 1914: 327 (Philonthus; synonym of agilis).
— Smetana, 1958: 193 (Philonthus; synonym of agilis).
— Smetana, 1995e: 523 (Bisnius; lectotype designation; synonym of sordidus).
chilensis Solier, 1849: 315 (Staphylinus; Type locality: Chili; [Note: See lectotype designation by Coiffait and Saiz, 1968]. Originally cited from Chile: San Carlos; Coquimbo; Longotomo).
— Kraatz, 1859a: 4 (Philonthus; notes).
— Fairmaire and Germain, 1861: 434 (Philonthus; characters; Chile).
— Fauvel, 1864a: 125 (Philonthus; notes; Chile).
— Fauvel, 1866a: 344 [= 1866b: 95] (Philonthus; synonym of sordidus).
— Fauvel, 1869: 489 (Philonthus; synonym of sordidus).
— Bernhauer and Schubert, 1914: 355 (Philonthus; synonym of sordidus).
— Cameron, 1932: 92 (Philonthus; synonym of sordidus).
— Smetana, 1958: 203 (Philonthus; synonym of sordidus).
— Coiffait and Saiz, 1968: 353 (Philonthus; lectotype designation: Chili; valid species; characters; Chile).
— Saiz, 1969a: 9 (Philonthus; notes).
— Saiz, 1971b: 341 (Philonthus; Chile).
— Smetana, 1995e: 523 (Bisnius; synonym of sordidus).
chloropterus Solier, 1849: 319 (Staphylinus; [preoccupied]; Type locality: Chile).
— Kraatz, 1859a: 5 (Philonthus; notes).
— Fairmaire and Germain, 1861: 432 (Philonthus; characters; Chile).
— Fauvel, 1866a: 344 [= 1866b: 95] (Philonthus; synonym of sordidus).
— Fauvel, 1869: 489 (Philonthus; synonym of sordidus).
— Bernhauer and Schubert, 1914: 355 (Philonthus; synonym of sordidus).
— Cameron, 1932: 92 (Philonthus; synonym of sordidus).
— Smetana, 1958: 203 (Philonthus; synonym of sordidus).
— Coiffait and Saiz, 1968: 353 (Philonthus; synonym of chilensis).
— Saiz, 1969a: 9 (Philonthus; synonym of chilensis).
— Smetana, 1995e: 523 (Bisnius; synonym of sordidus).
longipennis Provancher, 1877: 730 (Philonthus; Type locality: Quebec; [Note: See lectotype designation by Smetana, 1995e]. No locality cited originally).
— Provancher, 1877a: 308 (Philonthus; synonym of sordidus).
— Ganglbauer, 1895: 451 (Philonthus; synonym of sordidus).
— Bernhauer and Schubert, 1914: 355 (Philonthus; synonym of sordidus).
— Cameron, 1932: 92 (Philonthus; synonym of sordidus).
— Smetana, 1958: 203 (Philonthus; synonym of sordidus).
— Smetana, 1995e: 523 (Bisnius; lectotype designation: Quebec; synonym of sordidus).
impressifrons Broun, 1880: 110 (Philonthus; Type locality: Parua).
— Bernhauer and Schubert, 1914: 355 (Philonthus; synonym of sordidus).
— Cameron, 1932: 92 (Philonthus; synonym of sordidus).
— Smetana, 1958: 203 (Philonthus; synonym of sordidus).
brouni Duvivier, 1883: 145 (Philonthus; replacement name for impressifrons Broun; [Note: Unnecessary replacement name; I find no impressifrons in Philonthus that predates that of Broun]).
— Bernhauer and Schubert, 1914: 355 (Philonthus; synonym of sordidus).
— Cameron, 1932: 92 (Philonthus; synonym of sordidus).
— Smetana, 1958: 203 (Philonthus; synonym of sordidus).
— Smetana, 1995e: 523 (Bisnius; synonym of sordidus).
coloripennis Bernhauer, 1910b: 257 (Philonthus; variety of sordidus; Type locality: Ala-Tau, Syr-Darja).
— Bernhauer and Schubert, 1914: 356 (Philonthus; synonym of sordidus).
— Cameron, 1932: 92 (Philonthus; synonym of sordidus).
— Hodge and Jones, 1995: 40 (Philonthus; aberration of sordidus; characters; Britain).
testaceipennis Roubal, 1948: 123 (Philonthus; aberration of sordidus; cited as testa-ceipenennis on page 123 and testaceipennis on page 124; [Note: Nomen nudum (Articles 1.3.4, 45.5)); preoccupied]; cited from Slovakia: Zadiel; Bohemia: Praha-Stresovice).
— Smetana, 1958: 203 (Philonthus; aberration of sordidus).
— Smetana, 1959: 154 (Philonthus; aberration of sordidus; characters).
— Coiffait, 1967a: 408 (Philonthus; aberration of pachycephalus; characters).
— Coiffait, 1974: 240 (Philonthus; variety of pachycephalus).
— Smetana, 1995e: 523 (Bisnius; synonym of sordidus).
sparsus P. Lucas, 1846: 112 (Philonthus; Type locality: Algeria: environs du cercle de Lacalle).
— Kraatz, 1857d: 574 (Philonthus; characters).
— Fauvel, 1869: 489 (Philonthus; synonym of sordidus).
— Fauvel, 1874: 222 [= 1874a: 448] (Philonthus; synonym of sordidus).
— Mulsant and Rey, 1876: 439 [= 1877: 295] (Philonthus; characters; France).
— Ganglbauer, 1895: 451 (Philonthus; synonym of sordidus).
— Fauvel, 1902b: 109 (Philonthus; synonym of sordidus).
— Bernhauer and Schubert, 1914: 356 (Philonthus; synonym of sordidus).
— Cameron, 1932: 92 (Philonthus; synonym of sordidus).
— Tottenham, 1949a: 42 (Philonthus; valid species).
— Lohse, 1956: 88 (Philonthus; notes).
— Lohse, 1964: 181 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 152 (Philonthus; Algeria; Tunisia; France; Britain; Holland; Germany).
— Coiffait, 1967a: 408 (Philonthus; characters; North Africa; France; central Europe).
— Coiffait, 1974: 272 (Philonthus; characters; North Africa; France; Germany; Gold Coast).
— Brunne, 1976: 86 (Philonthus; characters).
— Mahler and Pritzl, 1980: 122 (Philonthus; characters).
— Tóth, 1984: 63 (Philonthus; characters).
— Lucht, 1987: 104 (Philonthus; checklist; Germany; Austria; Benelux).
— Drugmand, 1988: 62 (Philonthus; Belgium).
— Lohse and Lucht, 1989: 166 (Philonthus; characters).
— Smetana, 1995e: 516 (Bisnius).
— Assing, Frisch, Kahlen, et al., 1998: 137 (Philonthus; note).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
▴ Distribution: Ghana, Algeria, Tunisia, France, Germany, Holland, Belgium, Britain, Kazakhstan.
speculicollis Solodovnikov and Schillhammer, 2000: 167 (Bisnius; Type locality: Abkhazia: near Avadkhara, confluence of Avadkhara and Mzimna rivers, 1700 m).
spermophili Ganglbauer, 1897: 568 (Philonthus; Type locality: Wien).
— Bernhauer, 1899: 22 (Philonthus; valid species; notes).
— Reitter, 1909: 127 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 356 (Philonthus; catalog).
— Scheerpeltz, 1933: 1361 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Smetana, 1955b: 227 (Philonthus; characters).
— Smetana, 1958: 205 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 154 (Philonthus; characters).
— Lohse, 1964: 180 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 151 (Philonthus; Russia; Ukraine; Poland; Czechoslovakia; Hungary; Austria; Germany; Italy; France; Belgium; Holland).
— Coiffait, 1967a: 407 (Philonthus; characters; central and western Europe).
— Coiffait, 1974: 238 (Philonthus; characters; Europe; Russia).
— Brunne, 1976: 86 (Philonthus; characters).
— Mahler and Pritzl, 1980: 121 (Philonthus; characters).
— Tóth, 1984: 61 (Philonthus; cited as spermophilii; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; France; Benelux).
— Lohse and Lucht, 1989: 166 (Philonthus; characters).
— Nowosad, 1990: 109 (Philonthus; collected from mole nest; notes; Poland).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Smetana, 1995e: 516 (Bisnius).
▴ Distribution: Europe, Russia.
stacesmithi Hatch, 1957: 194 (Philonthus; Type locality: British Columbia: Copper Mt.).
— Smetana, 1995e: 586 (Bisnius; characters; notes; Alberta; British Columbia; Montana; Oregon; Utah; Washington).
▴ Distribution: Canada, USA.
stenoderus Reiche, 1861, see: fimetarius Gravenhorst, 1802.
subaereipennis Bernhauer, 1939, see: parcus Sharp, 1874.
subrugosus Mulsant and Rey, 1876, see: cephalotes Gravenhorst, 1802.
subuliformis Gravenhorst, 1802: 29 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).
— Latreille, 1804: 317 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 61 (Staphylinus; characters).
— Gyllenhal, 1810: 312 (Staphylinus; characters; Sweden).
— C. Sahlberg, 1830: 319 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 27 (Staphylinus; Russia).
— Mannerheim, 1831: 441 (Staphylinus; Russia).
— Zetterstedt, 1838: 62 (Staphylinus; characters; habitat; Lapland).
— Erichson, 1839a: 461 (Philonthus; synonym of fuscus).
— Heer, 1839: 264 (Philonthus; synonym of fuscus).
— Erichson, 1840: 457 (Philonthus; synonym of fuscus).
— Kraatz, 1857d: 593 (Philonthus; synonym of fuscus).
— Ganglbauer, 1895: 451 (Philonthus; synonym of fuscus).
— Bernhauer and Schubert, 1914: 340 (Philonthus; synonym of fuscus).
— Smetana, 1958: 208 (Philonthus; synonym of fuscus).
— Coiffait, 1967a: 405 (Philonthus; characters; Europe).
— Coiffait, 1974: 236 (Philonthus; characters; Europe; England; Russia; Scandinavia).
— Pope, 1977: 29 (Philonthus; Britain).
— Lohse and Lucht, 1989: 165 (Philonthus; note).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Philonthus; Italy).
— Smetana, 1995e: 516 (Bisnius).
— Martikainen, Kaila, and Haila, 1998: 296 (Philonthus; possibly threatened species; Finland).
— Fowles, Alexander, and Key, 1999: 129 (Philonthus; saproxylic species; status as endangered or threatened species; Britain).
▴ Distribution: Europe, Russia, Uzbekistan.
fuscus Gravenhorst, 1802: 29 (Staphylinus; preoccupied; Type locality: Not cited, but presumably Brunsvigae).
— Latreille, 1804: 317 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 59 (Staphylinus; characters).
— Gyllenhal, 1810: 329 (Staphylinus; characters; Sweden).
— Mannerheim, 1830: 29 (Staphylinus; Finland).
— Mannerheim, 1831: 443 (Staphylinus; Finland).
— Nordmann, 1837: 90 (Philonthus; Germany; Sweden).
— Erichson, 1839a: 461 (Philonthus; characters; Germany).
— Heer, 1839: 264 (Philonthus; characters; Switzerland).
— Erichson, 1840: 457 (Philonthus; characters; Germany; Sweden).
— Redtenbacher, 1849: 702 (Philonthus; characters; Austria).
— Kraatz, 1857d: 593 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 192 (Philonthus; characters; Austria).
— Thomson, 1860: 159 (Philonthus; characters; Scandinavia).
— Fauvel, 1874: 224 [= 1874a: 450] (Philonthus; characters; France; Scandinavia; Britain; Germany; Switzerland; Austria; Italy).
— Redtenbacher, 1874: 210 (Philonthus; characters; Austria).
— Seidlitz, 1875: 274 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 440 [= 1877: 296] (Philonthus; characters; France).
— Fowler, 1888: 269 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 451 (Philonthus; characters; north and middle Europe).
— Reitter, 1909: 126 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 340 (Philonthus; catalog).
— Johansen, 1914: 403 (Philonthus; characters; Denmark).
— Portevin, 1929: 359 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1344 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— V. Hansen, 1952: 62 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1955b: 227 (Philonthus; cited as valid species; characters).
— Smetana, 1958: 208 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 153 (Philonthus; characters).
— Lohse, 1964: 180 (Philonthus; subgenus Philonthus; characters; central Europe).
— Palm, 1963b: 77 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Horion, 1965: 149 (Philonthus; Russia; Finland; Sweden; Norway; Denmark; Britain; France to Poland and Russia; Pyrenees; Italy; Romania; Germany; Austria).
— Last, 1974a: 225 (Philonthus; collecting notes; England).
— Coiffait, 1974: 236 (Philonthus; synonym of subuliformis).
— Pope, 1977: 29 (Philonthus; synonym of subuliformis).
— Boller, 1983: 118 (Philonthus; larval characters).
— Tóth, 1984: 60 (Philonthus; characters; Hungary).
— Lucht, 1987: 104 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Boháč, 1988: 551 [= 1989: 34] (Philonthus; synonym of subuliformis).
— Lohse and Lucht, 1989: 165 (Philonthus; synonym of subuliformis).
— Smetana, 1993b: 48 (Philonthus; synonym of subuliformis).
— Ciceroni and Zanetti, 1995: 29 (Philonthus; synonym of subuliformis).
fragilis Gravenhorst, 1802: 30 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).
— Latreille, 1804: 317 (Staphylinus; characters; Germany).
— Gyllenhal, 1810: 313 (Staphylinus; synonym of subuliformis).
— Erichson, 1839a: 461 (Philonthus; synonym of fuscus).
— Heer, 1839: 264 (Philonthus; synonym of fuscus).
— Erichson, 1840: 457 (Philonthus; synonym of fuscus).
— Kraatz, 1857d: 593 (Philonthus; synonym of fuscus).
— Ganglbauer, 1895: 451 (Philonthus; synonym of fuscus).
— Bernhauer and Schubert, 1914: 340 (Philonthus; synonym of fuscus).
— Smetana, 1958: 208 (Philonthus; synonym of fuscus).
— Coiffait, 1974: 236 (Philonthus; synonym of subuliformis).
— Boháč, 1988: 551 [= 1989: 34] (Philonthus; synonym of subuliformis).
terminatus Gravenhorst, 1802: 30 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).
— Latreille, 1804: 317 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 61 (Staphylinus).
— Erichson, 1839a: 461 (Philonthus; synonym of fuscus).
— Heer, 1839: 264 (Philonthus; synonym of fuscus).
— Erichson, 1840: 457 (Philonthus; synonym of fuscus).
— Kraatz, 1857d: 593 (Philonthus; synonym of fuscus).
— Ganglbauer, 1895: 451 (Philonthus; synonym of fuscus).
— Bernhauer and Schubert, 1914: 340 (Philonthus; synonym of fuscus).
— Smetana, 1958: 208 (Philonthus; cited as terminalis; synonym of fuscus).
— Coiffait, 1974: 236 (Philonthus; cited as terminalis; synonym of subuliformis).
— Boháč, 1988: 551 [= 1989: 34] (Philonthus; cited as terminalis; synonym of subuliformis).
syrovatkai Roubal, 1913: 109 (Philonthus; aberration of fuscus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Chudenice).
— Scheerpeltz, 1933: 1344 (Philonthus; aberration of fuscus).
— Smetana, 1959: 153 (Philonthus; aberration of fuscus; characters).
— Smetana, 1955b: 227 (Philonthus; aberration of fuscus; characters).
— Smetana, 1958: 208 (Philonthus; synonym of fuscus).
— Coiffait, 1967a: 405 (Philonthus; aberration of subuliformis; characters).
— Coiffait, 1974: 236 (Philonthus; variety of subuliformis).
— Pope, 1977: 29 (Philonthus; synonym of subuliformis).
— Hodge and Jones, 1995: 40 (Philonthus; aberration of subuliformis; characters; Britain).
rubrithorax Kirshenblat, 1950: 237 (Philonthus; Type locality: Uzbekistan: Gul'dzha).
— Coiffait, 1967a: 390 (Philonthus; characters; Uzbekistan).
— Coiffait, 1974: 277 (Philonthus; characters; Uzbekistan).
— Boháč, 1988: 549, 551 [= 1989: 32, 34] (Philonthus; characters; synonym of subuliformis).
summanus Smetana, 1995e: 590 (Bisnius; Type locality: Oregon: Wallowa Co.: Eagles Cap Prim. Area Moccasin Lk., 7500 ft. Also cited from Montana).
synonymicus Roubal, 1915, see: cephalotes Gravenhorst, 1802.
syrovatkai Roubal, 1913 [Note: Nomen nudum], see: subuliformis Gravenhorst, 1802.
tantulus Luze, 1904, see: alaiensis Fauvel, 1900.
tereus Smetana, 1995e: 566 (Bisnius; Type locality: British Columbia: 21 mi. W. Creston. Also cited from Alberta and Montana).
terminatus Gravenhorst, 1802, see: subuliformis Gravenhorst, 1802.
testaceipennis Roubal, 1948 [Note: Nomen nudum], see: sordidus Gravenhorst, 1802.
thessalonicus Coiffait, 1963 [Note: Nomen nudum], see: cephalotes Gravenhorst, 1802.
thoracicus Melsheimer, 1884, see: blandus Gravenhorst, 1806.
varipennis Scriba, 1864a: 379 (Philonthus; [Note: This name is older than scribae; name said to be preoccupied by variipennis Kraatz but the two are not homonyms. The variant spelling difference is not covered by Article 58]; Type locality: Mehadia).
— Fauvel, 1867c: 112 [= 1868b: 211] (Philonthus; cited as variipennis; synonym of scribae).
— Fauvel, 1874: 224 [= 1874a: 450] (Philonthus; synonym of scribae).
— Redtenbacher, 1874: 217 (Philonthus; characters).
— Ganglbauer, 1895: 451 (Philonthus; synonym of scribae).
— Bernhauer and Schubert, 1914: 354 (Philonthus; synonym of scribae).
— Scheerpeltz, 1933: 1360 (Philonthus; synonym of scribae).
— Smetana, 1958: 207 (Philonthus; aberration of scribae).
— Coiffait, 1974: 237 (Philonthus; synonym of scribae).
— Burakowski, Mroczkowski, and Stefańska, 1980: 71 (Philonthus; cited as valid name; catalog; Poland).
— Ádám, 1987: 151 (Philonthus; cited as valid name; collecting notes; Hungary).
— Nowosad, 1990: 111 (Philonthus; cited as valid name; collected from mole nest; notes; Poland).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
▴ Distribution: Europe, Russia, Kazakhstan, Caucasus.
scribae Fauvel, 1867c: 112 [= 1868b: 211] (Philonthus; [Note: Unnecessary replacement name; proposed to replace “variipennis” Scriba that was said to be preoccupied by variipennis Kraatz; however Scriba cited the name as varipennis; see the previous “note” at varipennis]).
— Erichson, 1840: 458 (Philonthus; [Note: Bernhauer and Schubert, 1914: 354 cited placidus variety Erichson as a synonym of scribae; there is no available name]; characters).
— Kraatz, 1857d: 591 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 354 cited sordidus variety Kraatz as a synonym of scribae; there is no available name]; characters; Germany).
— Fauvel, 1874: 224 [= 1874a: 450] (Philonthus; characters; Germany; Hungary; Austria; Russia; Dauria; Netherland; Belgium).
— Eppelsheim, 1892: 332 (Philonthus; Turkestan).
— Ganglbauer, 1895: 451 (Philonthus; characters; Europe).
— Reitter, 1909: 126 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 354 (Philonthus; catalog).
— Scheerpeltz, 1933: 1360 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Kirshenblat, 1950: 241 (Philonthus; characters in key).
— Smetana, 1955b: 229 (Philonthus; notes).
— Smetana, 1958: 207 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 153 (Philonthus; characters).
— Lohse, 1964: 180 (Philonthus; subgenus Philonthus; cited as scribai; characters; central Europe).
— Horion, 1965: 147 (Philonthus; Russia; Hungary; Romania; Bosnia; Czechoslovakia; Austria; Germany; Holland).
— Coiffait, 1967a: 405 (Philonthus; characters; central Europe; Caucasus; Kazakhstan).
— Coiffait, 1974: 237 (Philonthus; characters; Europe; Russia; Kazakhstan).
— Tóth, 1984: 60 (Philonthus; cited as scribai; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; cited as scribai; checklist; Germany; Poland; Czechoslovakia; Austria; Benelux).
— Callot, 1993: 14 (Philonthus; notes; France).
— Smetana, 1993b: 48 (Philonthus; synonym of varipennis).
— Smetana, 1993b: 48 (Philonthus; synonym of varipennis).
— Smetana, 1995e: 516 (Bisnius).
heidenreichi Hubenthal, 1911a: 189 (Philonthus; variety of scribae; Type locality: Not cited).
— Bernhauer and Schubert, 1914: 354 (Philonthus; aberration of scribae).
— Smetana, 1959: 153 (Philonthus; aberration of heidenreichi; characters).
— Coiffait, 1967a: 405 (Philonthus; variety of scribae; characters).
— Coiffait, 1974: 237 (Philonthus; variety of scribae).
— Tóth, 1984: 60 (Philonthus; aberration of scribae).
versutus Horn, 1884: 211 (Philonthus; Type locality: California; [Note: See lectotype designation by Smetana, 1995e]).
— Bernhauer and Schubert, 1914: 360 (Philonthus; catalog).
— Smetana, 1995e: 559 (Bisnius; lectotype designation; characters; notes; British Columbia; California; Idaho; Nevada; New Mexico; Oregon; Utah; Washington).
▴ Distribution: Canada, USA.
watsoni Stephens, 1832, see: puella Nordmann, 1837.
xanthomerus Kraatz, 1859, see: fimetarius Gravenhorst, 1802.
zetes Smetana, 1995, see: picicornis Horn, 1884.
Cafius
[45 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions]
Cafius Curtis, 1829: 26, 245 (species included: xantholoma and two unavailable names). Type species: Staphylinus xantholoma Gravenhorst, fixed by monotypy.
— Stephens, 1829a: 23 (catalog; Britain).
— Stephens, 1829: 283 (catalog; Britain).
— Curtis, 1830: pl. 322 (characters; type species; Britain).
— Mannerheim, 1830: 7, 31 (characters).
— Mannerheim, 1831: 421, 445 (characters).
— Stephens, 1832: 200 (characters).
— Stephens, 1833: 245 (characters).
— Dejean, 1833: 63 (list of species).
— Lacordaire, 1835: 410 (characters).
— Westwood, 1838a: 16 (characters; type species).
— Stephens, 1839: 385 (characters).
— Erichson, 1840: 426 (synonym of Philonthus).
— Laporte, 1840: 179 (characters).
— Solier, 1849: 313 (characters).
— Lacordaire, 1854: 80 (synonym of Philonthus).
— Thomson, 1859: 24 (characters; type species: xantholoma).
— Thomson, 1860: 152 (characters).
— Gemminger and Harold, 1868: 584 (synonym of Philonthus).
— Fauvel, 1874: 195 [= 1874a: 421] (characters; key to species).
— Fauvel, 1875a: XXVIII [= 1875b: 230] (catalog).
— Mulsant and Rey, 1876: 564 [= 1877: 420] (characters; key to French species).
— Fauvel, 1877: 250 (notes; key to species).
— Fauvel, 1878f: 251 (characters; notes).
— Duvivier, 1883: 149 (catalog).
— Horn, 1884: 234 (characters; notes; key to North American species).
— Olliff, 1887: 498 (characters; Australian species).
— Fowler, 1888: 281 (notes; key to British species).
— Heyden, Reitter, and Weise, 1891: 102 (list of species of Europe and Caucasus).
— Ganglbauer, 1895: 462 (characters; key to middle European species).
— Fauvel, 1897d: 297 (catalog of species of Barbary; Canaries).
— Heyden, Reitter, and Weise, 1906: 163 (list of species of Europe and Caucasus).
— Reitter, 1909: 116, 123 (characters).
— Bernhauer and Schubert, 1914: 361 (world catalog; 43 species).
— Johansen, 1914: 420 (characters).
— Leng, 1920: 109 (catalog of North American species).
— Cameron, 1921a: 354, 405 (characters; key to Singapore species; catalog).
— Winkler, 1925: 380 (catalog for Palaearctic region).
— Cameron, 1925: 63 (catalog of Indian species).
— Porta, 1926: 108 (characters; key to species of Italy).
— Portevin, 1929: 354 (characters; key to French species).
— Gridelli, 1931: 122 (key to European and Mediterranean species).
— Cameron, 1932: 154 (characters; key to species of British India).
— Joy, 1932: 115 (key to British species).
— Scheerpeltz, 1933: 1368 (world catalog supplement).
— Normand, 1935: 372 (list of species of Tunisia).
— C. Koch, 1936b: 172 (key to species).
— Blackwelder, 1943: 435 (characters; type species: fucicola; key to species of West Indies).
— Blackwelder, 1944: 135 (checklist of species from Mexico, Central and South America, and the West Indies).
— Kloet and Hincks, 1945: 166 (list of British species).
— Tottenham, 1949: 373, 419 (type species: xantholoma Gravenhorst; notes; checklist of British species).
— Blackwelder, 1952: 88 (type species: xantholoma).
— V. Hansen, 1952: 34 (characters; Danish species).
— Hatch, 1957: 173, 210 (characters; key to species of Pacific Northwest).
— Kocher, 1958: 151 (checklist of species; Morocco).
— Smetana, 1959a: 214 (list of species collected in Albania).
— Ferreira, 1962a: 17 (list of Portuguese species).
— Coiffait, 1963: 8, 20 (characters; key to subgenera).
— Palm, 1963b: 60 (characters; species of Sweden).
— Lohse, 1964: 173 (characters; key to central European species).
— Moore, 1965: 98, 101 (characters; notes).
— Pototskaia, 1966: 152 (larval characters).
— Pototskaia, 1967: 53 (larval characters).
— Coiffait and Saiz, 1968: 348, 361 (characters; notes; key to Chilean species).
— Kasule, 1970: 65 (larval characters).
— James, Moore, and Legner, 1971: 279 (notes on habitat, predation, feeding, mating; larval characters; key to known larvae and key to pupae of American species).
— Bordoni, 1973j: 710 (list of species from islands near Sicily).
— Tikhomirova, 1973a: 182 (checklist of species of USSR).
— Coiffait, 1974: 338 (adult and larval characters; key to species of western Palaearctic region).
— Moore, 1975a: 140 (key to Nearctic species).
— Moore and Legner, 1975: 9 (catalog; North America north of Mexico).
— Moore and Legner, 1976: 540 (includes seashore species; notes).
— Pope, 1977: 30 (list of British species).
— Topp, 1978: 312 (larval characters).
— Moore and Legner, 1979: 17 (characters; notes).
— Muona, 1979: 14 (list of Scandinavian species).
— Orth and Moore, 1980: 181, 198 (notes; key to species of eastern and western North America; characters).
— Burakowski, Mroczkowski, and Stefańska, 1980: 33 (catalog; Poland).
— Uhlig and Vogler, 1981: 110 (list of some species collected in Germany; habitat and collecting notes).
— Tóth, 1984: 39 (characters; habitus).
— Frank, Carlysle, and Rey, 1986: 155 (discussion of habitat, community, and adaptations to environment).
— Shibata, 1986a: 123 (supplemental list of species from Taiwan).
— Drugmand 1987b (comments on distribution of Belgian species).
— Campbell and Davies, 1991: 119 (checklist of species; Alaska and Canada).
— Silfverberg, 1992: 17 (list of species of Fennoscandia, Denmark, and the Baltic States).
— Welch, 1993: 229 (ovariole structure and development).
— Kanehisa, Tsumuki, and Kawazu, 1994 (chemical secretions).
— Smetana, 1995e: 50 (characters).
— Ciceroni and Zanetti, 1995: 27 (list of species of Italy).
— M. Hansen, 1996: 104 (list of Danish species).
— Lundgren, 1998: 51 (list of species from Florida; records for other States listed for each species).
— Newton, Thayer, Ashe, and Chandler, 2000: 396 (13 Nearctic species; notes; characters in key).
— Hayashi, 2000: 23 (notes on empodial setae).
Platygonium Motschulsky, 1845a: 40 (without species). Type species: Platygonium sculticeps Motschulsky, fixed by Motschulsky, 1858: 649, by subsequent monotypy.
— Motschulsky, 1858: 649 (description; first included species: sculticeps).
— Gemminger and Harold, 1868: 615 (catalog).
— Bernhauer and Schubert, 1912: 273 (catalog).
— Winkler, 1925: 369 (catalog; Palaearctic region).
— Blackwelder, 1939: 120 (type species).
— Blackwelder, 1952: 313 (type species: sculticeps).
— Gusarov, 1991a: 8 (subgenus of Cafius).
Bryonomus Casey, 1885a: 313 (species included: canescens, seminitens). Type species: Philonthus canescens Mäklin, fixed by subsequent designation by Blackwelder, 1943: 435.
— Bernhauer and Schubert, 1914: 361 (subgenus of Cafius).
— Cameron, 1932: 154 (subgenus of Cafius).
— Blackwelder, 1943: 435 (subgenus of Cafius; type species: canescens).
— Blackwelder, 1952: 86 (type species).
Euremus Blackwelder, 1943: 435 (subgenus of Cafius; Bierig did not designate a type species, so Pseudoremus Koch is unavailable. Blackwelder's designation of a type species made the name available so he is the author [see Herman, 2001]; see Bierig, 1934a for species included). Type species: Philonthus bistriatus Erichson, fixed by original designation.
— Bierig, 1934a: 68 (subgenus of Cafius; species included: rufifrons, bistriatus, fonticola cited as fronticola, pacificus, lithocharinus, nauticus cited as nautucus; characters; [Note: Bierig did not designate a type species so Euremus Bierig is unavailable, but was made available by Blackwelder]).
— Blackwelder, 1943: 435 (subgenus of Cafius; type species: bistriatus).
— Blackwelder, 1952: 158 (type species).
— Hatch, 1957: 210 (subgenus of Cafius; characters; species of Pacific Northwest).
— Coiffait, 1974: 338, 341 (subgenus of Cafius; characters; key to species of western Palaearctic region).
— Herman, 2001: 9 (type species; discussion; Blackwelder is author of name, not Bierig).
Philonthopsis Blackwelder, 1943: 435 (subgenus of Cafius; [preoccupied by Cameron, 1932; Koch did not designate a type species, so Philonthopsis Koch is unavailable. Blackwelder's designation of a type species made the name available so he is the author]). Type species: Cafius sabulosus Fauvel, fixed by subsequent designation by Blackwelder, 1943: 435 (see Herman, 2001).
— C. Koch, 1936b: 173, 178 (subgenus of Cafius; species included: australis, areolatus, litoreus, sabulosus; [Note: Koch did not designate a type species so Philonthopsis Koch is unavailable but was made available by Blackwelder]).
— Blackwelder, 1952: 301 (type species).
— Herman, 2001: 10 (type species; discussion; Blackwelder is author of name, not Koch).
Pseudoremus Blackwelder, 1943: 435 (subgenus of Cafius; [Koch did not designate a type species, so Pseudoremus C. Koch is unavailable. Blackwelder's designation of a type species made the name available so he is the author]). Type species: Philonthus lithocharinus LeConte, fixed by original designation (see Herman, 2001).
— C. Koch, 1936b: 175, 179 (subgenus of Cafius; species included: opacus, vestitus, rufescens, algarum, histrio, mimulus, lithocharinus, ragazzii, nauticus; [Note: Koch did not designate a type species so Pseudoremus Koch is unavailable, but was made available by Blackwelder]).
— Blackwelder, 1952: 330 (type species).
— Hatch, 1957: 210 (subgenus of Cafius; characters; species of Pacific Northwest).
— Coiffait, 1974: 339, 341 (subgenus of Cafius; characters; key to species of western Palaearctic region).
— Herman, 2001: 10 (type species; discussion; Blackwelder is author of name, not Koch).
Ifacus Blackwelder, 1952: 198 (subgenus of Cafius; replacement name for Philonthopsis Koch [Note: As stated above Blackwelder is the author of the name]). Type species: Cafius sabulosus Fauvel, fixed by objective synonymy with Philonthopsis.
Suborthidus Coiffait, 1963: 20 (subgenus of Cafius; species included: cicatricosus). Type species: Cafius cicatricosus Erichson, fixed by monotypy.
— Coiffait, 1974: 338, 341 (subgenus of Cafius; characters; key to species of western Palaearctic region).
— Gusarov, 1991a: 8 (synonym of Platygonium).
aegyptiacus Motschulsky, 1858, see: Remus.
aguayoi Bierig, 1934a: 66 (Cafius; subgenus Remus; Type locality: USA: Woods Hole, Massachusetts).
— C. Koch, 1936b: 192 (Cafius; characters).
— Orth and Moore, 1980: 199 (Cafius; characters).
— Frank, Carlysle, and Rey, 1986: 149 (Cafius; notes).
▴ Distribution: USA.
algarum Sharp, 1874: 38 (Philonthus; subgenus Cafius; Type locality: Nagasaki).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Bernhauer and Schubert, 1914: 361 (Cafius; catalog).
— Scheerpeltz, 1933: 1369 (Cafius; catalog).
— C. Koch, 1936b: 191, 193 (Cafius; characters in key).
— Watanabe, 1961: 354 (Cafius; Izu Islands).
— Watanabe, 1993d (Cafius; cited as alganum; Sarushima Island).
— Kanehisa, Tsumuki, and Kawazu, 1994 (Cafius; chemical secretions).
▴ Distribution: Japan.
algophilus Broun, 1894: 419 (Cafius; Type locality: Mokohinou Island).
— Bernhauer and Schubert, 1914: 361 (Cafius; catalog).
— Kuschel, 1990: 26 (Cafius; endemic; winged; habitat; New Zealand).
— Emberson, 1998: 35 (Cafius; collecting notes; Chatham Islands).
▴ Distribution: New Zealand, Chatham Islands.
amblyterus Olliff, 1887, see: Philonthus.
anchora Fauvel, 1877, see: Hesperus.
andamanensis Coiffait, 1981f: 337 (Cafius; Type locality: South Andaman, Ile Rutland).
areolatus Fauvel, 1877, see: australis Redtenbacher, 1867.
arrowi Bernhauer, 1932, see: martini Cameron, 1927.
australis Redtenbacher, 1867: 28 (Ocypus; Type locality: Sidney).
— Fauvel, 1877: 251 (Cafius; characters; Australia).
— Olliff, 1887: 500 (Cafius; characters; Australia).
— Bernhauer and Schubert, 1914: 361 (Cafius; catalog).
— Lea, 1925: 225 (Cafius; characters; Australia).
— Scheerpeltz, 1933: 1369 (Cafius; catalog).
— C. Koch, 1936b: 178 (Cafius; characters in key; Australia).
▴ Distribution: Australia.
areolatus Fauvel, 1877: 251 (Cafius; Type locality: Australie, Victoria).
— Fauvel, 1878g: 541 (Cafius; characters).
— Olliff, 1887: 499 (Cafius; characters; Australia).
— Lea, 1899: 540 (Cafius; Australia).
— Bernhauer and Schubert, 1914: 361 (Cafius; catalog).
— Lea, 1925: 225 (Cafius; synonym of australis).
— C. Koch, 1936b: 176, 178 (Cafius; synonym of australis).
bilineatus Erichson, 1840, see: bistriatus Erichson, 1840.
bistriatus Erichson, 1840: 502 (Philonthus; Type locality: Americae septentrionalis Insula Longa [Long-Island]).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Horn, 1884: 237 (Cafius; characters; Atlantic coast and on margin of tide-water streams).
— Bernhauer and Schubert, 1914: 361 (Cafius; catalog).
— Notman, 1920b: 705 (Cafius; Florida).
— Scheerpeltz, 1933: 1369 (Cafius; catalog).
— Bierig, 1934a: 68 (Cafius; subgenus Euremus).
— C. Koch, 1936b: 187 (Cafius; characters in key; New York; Massachusetts; Lesser Antilles; Jamaica; Florida).
— Blackwelder, 1943: 439 (Cafius; characters; Bahamas; Cuba; Jamaica; Mona Island; Puerto Rico; St. Thomas; St. John; St. Croix; St. Kitts; Antigua; Montserrat; Guadeloupe; Dominica; St. Lucia; Barbados; Grenadines; Grenada; Tobago; Trinidad).
— Ramos, 1946: 33 (Cafius; Mona Island, West Indies).
— Smetana, 1965: 32 (Cafius; Newfoundland).
— Moore, 1975a: 140, 142 (Cafius; [Note: Misidentified as sulcicollis according to Orth and Moore, 1980: 195]; adult and larval characters).
— Orth and Moore, 1980: 195, 199 (Cafius; characters; notes; California; Sonora; Baja California Norte; Baja California Sur; Long Island Sound; Chesapeake Bay).
— Frank, 1985: 61 (Cafius; Florida).
— Frank, Carlysle, and Rey, 1986: 149,158 (Cafius; characters; discussion of biogeography; Quebec; New Brunswick; Nova Scotia; Maine; Massachusetts; New York; New Jersey; Maryland; Virginia; Florida; Texas; Antigua; Virgin Islands; Jamaica; Venezuela; Mexico).
▴ Distribution: Canada, USA, Mexico, West Indies, Venezuela.
bilineatus Erichson, 1840: 503 (Philonthus; Type locality: Americae meridionalis ins. St. Johannis).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Scheerpeltz, 1933: 1369 (Cafius; catalog).
— Blackwelder, 1943: 438 (Cafius; synonym of bistriatus).
— Orth and Moore, 1980: 195 (Cafius; synonym of bistriatus).
— Frank, Carlysle, and Rey, 1986: 149 (Cafius; synonym of bistriatus).
fulgens Frank, 1986: 153 (Cafius; subspecies of bistriatus; Type locality: Mexico: Baja California Sur: Mulege. Also cited from California).
bisulcatus Solier, 1849: 314 (Staphylinus; Type locality: Chile).
— Kraatz, 1859a: 4 (Staphylinus; subgenus Cafius; notes).
— Fairmaire and Germain, 1861: 432 (Philonthus; characters; Chile).
— Fauvel, 1864a: 126 (Philonthus; notes; Chile).
— Fauvel, 1866a: 342 [= 1866b: 93] (Philonthus; characters; Chile).
— Redtenbacher, 1867: 29 (Philonthus; Chile; Australia).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Scheerpeltz, 1933: 1369 (Cafius; catalog).
— C. Koch, 1936b: 183 (Cafius; characters in key; Chile).
— Coiffait and Saiz, 1968: 362 (Cafius; characters; Chile).
— Saiz, 1969a: 8 (Cafius; notes).
— Saiz, 1971b: 349 (Cafius; notes; Chile).
— Frank, 1982: 34 (Cafius; list of parasites).
▴ Distribution: Chile, Australia.
bryanti Cameron, 1943a: 343 (Cafius; Type locality: S. Australia: Adelaide).
canescens Mäklin, 1852: 313 (Philonthus; Type locality: insula Edgecombe; [Note: See lectotype designation by Orth and Moore, 1980]).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Horn, 1884: 235 (Cafius; characters; Pacific seacoast).
— Casey, 1885a: 314 (Bryonomus; characters).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Scheerpeltz, 1933: 1369 (Cafius; catalog).
— C. Koch, 1936b: 183 (Cafius; characters in key; California; Washington; Alaska).
— Hatch, 1957: 211 (Cafius; subgenus Bryonomus; characters; British Columbia; Washington; Oregon).
— R. Dvořák, 1957a (Cafius; subgenus Bryonomus; characters).
— James, Moore, and Legner, 1971: 284, 287, 288 (Cafius; larval and pupal characters; California).
— Moore, 1975a: 140 (Cafius; characters).
— Orth, Moore, and Fisher, 1978: 185 (Cafius; characters; collecting notes; California).
— Orth and Moore, 1980: 185 (Cafius; lectotype designation; characters; British Columbia; Alaska; Washington; Oregon; California; Baja California Norte).
— Frank, 1982: 34 (Cafius; list of parasites).
▴ Distribution: Canada, USA, Mexico.
caribeanus Bierig, 1934a: 68 (Cafius; variety of corallicola; Type locality: Plaia de Marianao).
— Blackwelder, 1943: 437 (Cafius; cited as valid species; characters; Cuba; Jamaica; Puerto Rico; St. Croix; Antigua; Guadeloupe; Dominica; Grenadines; Grenada).
— Orth and Moore, 1980: 197, 199 (Cafius; characters; Mexico; West Indies).
— Frank, 1985: 61 (Cafius; Florida; Venezuela).
— Frank, Carlysle, and Rey, 1986: 149 (Cafius; characters).
▴ Distribution: Mexico, West Indies, Venezuela.
catenatus Fauvel, 1877: 256 (Cafius; Type locality: Australie, Nouvelle Galles du Sud).
— Fauvel, 1878g: 542 (Cafius; characters; Australia).
— Olliff, 1887: 506 (Cafius; characters; Australia).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— C. Koch, 1936b: 185 (Cafius; characters in key; Australia).
— Frank, 1982: 34 (Cafius; list of parasites).
▴ Distribution: Australia.
caviceps Broun, 1886: 942 (Cafius; Type locality: Mokohinau Island).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— C. Koch, 1936b: 180 (Cafius; characters in key; New Zealand).
▴ Distribution: New Zealand.
puncticeps White, 1846: 6 (Staphylinus; subgenus Cafius; [Note: This name is older than caviceps]; Type locality: New Zealand).
— Fairmaire, 1847: 91 (Staphylinus; translation of original description).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Fauvel, 1877: 254 (Cafius; characters; Nouvelle Zélande).
— Broun, 1880: 105 (Xantholinus; subgenus Cafius; position unknown).
— Bernhauer and Schubert, 1914: 363 (Cafius; catalog).
— Scheerpeltz, 1933: 1370 (Cafius; catalog).
— Steel, 1949a: 270 (Cafius; synonym of caviceps; [Note: Steel cited the date of White's description as 1874 and listed the species in Xantholinus but it was originally described in Staphylinus and later moved to Xantholinus where it was a secondary homonym]).
— Frank, 1982: 34 (Cafius; list of parasites).
ceylonicus Bernhauer, 1902b: 29 (Cafius; Type locality: Weligama).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Cameron, 1932: 155 (Cafius; characters; Ceylon).
— Scheerpeltz, 1933: 1369 (Cafius; catalog).
▴ Distribution: Sri Lanka.
cicatricosus Erichson, 1840: 454 (Philonthus; Type locality: Sicilia).
— Fairmaire and Laboulbène, 1856: 521 (Philonthus; characters; France).
— Kraatz, 1857d: 575 (Philonthus).
— Fauvel, 1874: 197 [= 1874a: 423] (Cafius; characters; France; England; Sardinia; Sicily).
— Mulsant and Rey, 1876: 567 [= 1877: 423] (Cafius; characters; France).
— Fowler, 1888: 281 (Cafius; characters; Britain).
— Ganglbauer, 1895: 462 (Cafius; characters; England; France; Italy; Sardinia; Sicily).
— Fauvel, 1902b: 106 (Cafius; Tunisia).
— Sainte-Claire Deville, 1906: 95 (Cafius; Corsica).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Portevin, 1929: 355 (Cafius; characters; France).
— Gridelli, 1931: 123 (Cafius; characters in key).
— Scheerpeltz, 1933: 1369 (Cafius; catalog).
— C. Koch, 1936b: 181 (Cafius; characters in key; England; France; Spain; Italy; Corsica; Dalmatia; Tunisia).
— Coiffait, 1974: 350 (Cafius; subgenus Suborthidus; characters; Mediterranean coast; England).
— Pope, 1977: 30 (Cafius; Britain).
— Gusarov, 1991a: 8 (Cafius; subgenus Platygonium; notes).
— Ciceroni and Zanetti, 1995: 27 (Cafius; Italy).
▴ Distribution: Britain, France, Italy, Tunisia, Russia.
sculticeps Motschulsky, 1858: 649 (Platygonium; Type locality: Kharov, Rus. mer.; [Note: See lectotype designation by Gusarov, 1991a]).
— Gemminger and Harold, 1868: 615 (Platygonium; cited as sculpticeps Motschulsky).
— Bernhauer and Schubert, 1912: 273 (Platygonium; cited as sculpticeps; catalog).
— Gusarov, 1991a: 8 (Cafius; lectotype designation: Kharov, Rus. mer.; synonym of cicatricosus).
decipiens LeConte, 1863a: 40 (Philonthus; Type locality: San Diego; [Note: See lectotype designation by Orth and Moore, 1980]).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Horn, 1884: 239 (Cafius; characters; California).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Scheerpeltz, 1933: 1369 (Cafius; catalog).
— C. Koch, 1936b: 186 (Cafius; characters in key; California).
— Orth and Moore, 1980: 192 (Cafius; lectotype designation; characters; California; Baja California Norte).
▴ Distribution: Mexico, USA.
densiventris Fauvel, 1877, see: nauticus Fairmaire, 1849.
dubius LeConte, 1863, see: opacus LeConte, 1863.
expuncticollis C. Koch, 1936 [Note: Nomen nudum], see: quadriimpressus White, 1846.
femoralis Mäklin, 1853, see: mutatus Gemminger and Harold, 1868.
flicki Vauloger, 1897: 238 (Cafius; Type locality: iles Kerkenna, Tunisia).
— Fauvel, 1902b: 106 (Cafius; Tunisia).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Gridelli, 1931: 120, 124 (Cafius; characters; Tunisia).
— C. Koch, 1936b: 182 (Cafius; characters in key; Tunisia; Tripolitania).
— Coiffait, 1974: 349 (Cafius; characters; Tunisia).
— Ciceroni and Zanetti, 1995: 27 (Cafius; Italy).
▴ Distribution: Tunisia, Libya, Italy.
fonticola Erichson, 1840: 501 (Philonthus; Type locality: montes Sinai; mons S. Catharinae).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Bierig, 1934a: 68 (Cafius; subgenus Euremus; cited as fronticola).
— C. Koch, 1936b: 168 (Cafius; notes; Egypt).
— C. Koch, 1936b: 183 (Cafius; characters in key; Red Sea).
— Coiffait, 1974: 347 (Cafius; subgenus Euremus; characters; Red Sea).
▴ Distribution: Egypt.
fucicola Curtis, 1830: 322 (Cafius; Type locality: England: Mount Edgecombe, near Plymouth).
— Curtis, 1829: 30 (Cafius; nomen nudum; catalog; Britain).
— Stephens, 1833: 246 (Cafius; characters; England).
— Nordmann, 1837: 84 (Philonthus; [Note: Nordmann attributed this name to Leach who did not describe it; however Curtis described fucicola in Cafius and I assume it is this species to which Nordmann is referring]; characters; England).
— Stephens, 1839: 399 (Cafius; characters; England).
— Erichson, 1840: 454 (Philonthus; characters; England).
— Hardy, 1851: 28 (Philonthus; notes; England).
— Kraatz, 1857d: 575 (Philonthus).
— Fauvel, 1874: 197 [= 1874a: 423] (Cafius; characters; France; England).
— Fowler, 1888: 282 (Cafius; characters; Britain).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Portevin, 1929: 354 (Cafius; characters; France).
— Gridelli, 1931: 124 (Cafius; characters in key).
— Scheerpeltz, 1933: 1369 (Cafius; catalog).
— C. Koch, 1936b: 182 (Cafius; characters in key; England; France).
— Coiffait, 1974: 348 (Cafius; characters; France; England).
— Pope, 1977: 30 (Cafius; Britain).
▴ Distribution: Britain, France.
fulgens Frank, 1986, see: bistriatus Erichson, 1840.
gigas Lea, 1929: 204 (Cafius; Type locality: Lord Howe Island: Mount Ledgbird).
heroopoliticus C. Koch, 1936, see: xantholoma Gravenhorst, 1806.
histrio Sharp, 1874: 37 (Philonthus; subgenus Cafius; Type locality: Amakusa and Iwosima).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Scheerpeltz, 1933: 1369 (Cafius; catalog).
— C. Koch, 1936b: 191 (Cafius; characters in key; Hong Kong; Korea).
— Watanabe, 1961: 354 (Cafius; Izu Islands).
— Watanabe and Shibata, 1972: 64 (Cafius; Yaku-shima Island).
▴ Distribution: Japan, Korea, Hong Kong.
horni Bernhauer, 1902, see: Phucobius.
johnsoni Fall, 1916, see: Philonthus.
keysianus Donisthorpe, 1930 [Note: Nomen nudum], see: xantholoma Gravenhorst, 1806.
laetabilis Olliff, 1887, see: Philonthus.
laeus Olliff, 1887, see: Philonthus.
lateralis Stephens, 1833, see: xantholoma Gravenhorst, 1806.
lithocharinus LeConte, 1863a: 38 (Philonthus; Type locality: San Diego, California).
— Horn, 1884: 236 (Cafius; characters; California sea-coast).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Scheerpeltz, 1933: 1370 (Cafius; catalog).
— Bierig, 1934a: 68 (Cafius; subgenus Euremus).
— C. Koch, 1936b: 189 (Cafius; characters in key; California).
— Hatch, 1957: 210 (Cafius; subgenus Pseudoremus; characters; Washington).
— Leech and Moore, 1971 (Cafius; flight records; notes; California).
— James, Moore, and Legner, 1971: 283, 287, 288 (Cafius; larval and pupal characters; California).
— Moore, 1975a: 140 (Cafius; characters).
— Orth, Moore, and Fisher, 1978: 186 (Cafius; characters; collecting notes; California).
— Moore and Andrews, 1985: 221 (Cafius; notes; Baja California Sur).
▴ Distribution: Mexico, USA.
littoralis Stephens, 1833, see: xantholoma Gravenhorst, 1806.
littoralis Fauvel, 1877, see: pacificus Erichson, 1840.
litoreus Broun, 1880: 108 (Staphylinus; [preoccupied by littoreus Linné, 1758 (Article 58.7)]; Type locality: Tairua).
— Broun, 1886: 942 (Cafius).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— C. Koch, 1936b: 178 (Cafius; characters in key; New Zealand).
— Klimaszewski and Watt, 1997: 171 (Cafius; characters; collecting notes; New Zealand).
— Herman, 2001: 43 (Cafius; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: New Zealand.
longipennis Walker, 1858, see: nauticus Fairmaire, 1849.
luteipennis Horn, 1884: 237 (Cafius; Type locality: California sea-coast; [Note: See lectotype designation by Orth and Moore, 1980]).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Scheerpeltz, 1933: 1370 (Cafius; catalog).
— C. Koch, 1936b: 187 (Cafius; characters in key; California; British Columbia).
— Hatch, 1957: 211 (Cafius; subgenus Euremus; characters; British Columbia; Washington; Oregon).
— Leech and Moore, 1971 (Cafius; notes; California).
— James, Moore, and Legner, 1971: 282, 287, 288 (Cafius; larval and pupal characters; California).
— Moore, 1975a: 140 (Cafius; characters).
— Orth, Moore, and Fisher, 1978: 186 (Cafius; characters; collecting notes; California).
— Orth and Moore, 1980: 191 (Cafius; lectotype designation; characters; British Columbia; Washington; Oregon; California; Baja California Norte).
— Moore and Andrews, 1985: 222 (Cafius; Baja California Sur).
▴ Distribution: Canada, USA, Mexico.
maritimus Broun, 1880: 109 (Staphylinus; Type locality: Tairua).
— Broun, 1886: 942 (Cafius).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— C. Koch, 1936b: 184 (Cafius; characters in key; New Zealand).
▴ Distribution: New Zealand.
martini Cameron, 1927d: 251 (Cafius; Type locality: Jeddah; Kamaran Island).
— Scheerpeltz, 1933: 1370 (Cafius; catalog).
— C. Koch, 1936b: 188 (Cafius; characters in key; Red Sea region).
— Coiffait, 1974: 347 (Cafius; subgenus Euremus; characters; Red Sea).
▴ Distribution: Saudi Arabia, Yemen.
arrowi Bernhauer, 1932c: 234 (Cafius; Type locality: Perim).
— Scheerpeltz, 1933: 1369 (Cafius; catalog).
— Bernhauer, 1936g: 244 (Cafius; synonym of martini).
— Coiffait, 1974: 347 (Cafius; synonym of martini).
mimulus Sharp, 1874: 38 (Philonthus; subgenus Cafius; [preoccupied]; Type locality: Amakusa and Iwosima).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Scheerpeltz, 1933: 1370 (Cafius; catalog).
— C. Koch, 1936b: 191 (Cafius; characters in key).
— Watanabe, 1993d (Cafius; Sarushima Island).
— Herman, 2001: 43 (Cafius; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Japan.
mutatus Gemminger and Harold, 1868: 590 (Philonthus; replacement name for femoralis Mäklin).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Scheerpeltz, 1933: 1370 (Cafius; catalog).
— Smetana, 1965a: 9 (Cafius; synonym of femoralis; said to be unnecessary replacement name).
▴ Distribution: Canada, USA.
femoralis Mäklin, 1853: 189 (Philonthus; [preoccupied]; Type locality: insula Kadjak).
— Horn, 1884: 238 (Cafius; characters; Alaska).
— Bernhauer and Schubert, 1914: 362 (Cafius; synonym of mutatus).
— C. Koch, 1936b: 193 (Cafius; characters).
— Hatch, 1957: 210 (Cafius; subgenus Remus; characters; British Columbia; Washington).
— Smetana, 1965a: 9 (Cafius; subgenus Remus).
nasutus Fauvel, 1877: 257 (Cafius; Type locality: Iles Fidji, Viti).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Lea, 1925: 225 (Cafius; needs confirmation as Australian species).
▴ Distribution: Fiji.
nauticus Fairmaire, 1849: 288 (Philonthus; Type locality: Taïti).
— Fauvel, 1874c: 438 (Cafius; characters; New Caledonia; Tahiti).
— Blackburn and Sharp, 1885: 222 (Cafius; habitat; Hawaii).
— Fauvel, 1889b: 262 (Cafius; New Caledonia).
— Fauvel, 1898a: 117 (Cafius; Seychelles).
— Fauvel, 1903d: 268 (Cafius; New Caledonia; Australia; Tahiti; Aru; Java; China; Ceylon; Seychelles; Madagascar; Perim; Berbera; Obock; Sikitra; Erythrea; Hawaii).
— Sharp, 1908: 548 (Cafius; Hawaii).
— H. Kolbe, 1910: 19 (Cafius; Seychelles).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Cameron, 1921a: 378, 405 (Cafius; characters; Singapore).
— Bernhauer, 1922b: 177 (Cafius; Seychelles).
— Lea, 1925: 226 (Cafius; characters; Australia).
— Cameron, 1927a: 26 (Cafius; Samoa).
— Cameron, 1927c: 264 (Cafius; Fiji; New Caledonia).
— Cameron, 1932: 155 (Cafius; characters; Ceylon).
— Scheerpeltz, 1933: 1370 (Cafius; catalog).
— Bierig, 1934a: 68 (Cafius; subgenus Euremus; cited as nautucus).
— C. Koch, 1936b: 190 (Cafius; characters in key).
— Scheerpeltz, 1954: 114 (Cafius; Ceylon).
— Jarrige, 1964a: 177 (Cafius; New Caledonia).
— Shibata, 1973b: 132 (Cafius; subgenus Remus; Taiwan; Ryukyus).
— Coiffait, 1974: 345 (Cafius; subgenus Pseudoremus; characters; coast of Indian Ocean and Red Sea).
— Shibata, 1974: 31 (Cafius; Japan).
— Coiffait, 1976: 217 (Cafius; Tahiti).
— Coiffait, 1977a: 171 (Cafius; Austral Islands).
— Bordoni and Bordoni, 1989: 745 (Cafius; subgenus Pseudoremus; introduced to Greece).
— Lecoq, 1990: 205 (Cafius; Mascarene Islands).
— Ikudome and Hayakawa, 1994: 187 (Cafius; cattle dung collections; Ryukyu Islands).
— Nishida, 1994: 71 (Cafius; Hawaii).
— Ito, 1996: 66 (Cafius; Japan).
— Nishida, 1997: 60 (Cafius; Hawaii).
▴ Distribution: Hawaii, Japan, China, Taiwan, Society Islands, Austral Islands, Samoa, Fiji, New Caledonia, Australia, Indonesia, Singapore, Sri Lanka, Madagascar, Seychelles, Mascarene Islands, Yemen, Eritrea, Djibouti, Somalia, Greece.
longipennis Walker, 1858: 205 (Ocypus; Type locality: Ceylon).
— Fauvel, 1902a: 41 (Cafius; synonym of nauticus).
— Fauvel, 1903d: 268 (Cafius; synonym of nauticus).
— Bernhauer and Schubert, 1914: 362 (Cafius; synonym of nauticus).
— Cameron, 1932: 155 (Cafius; synonym of nauticus).
— Coiffait, 1974: 345 (Cafius; synonym of nauticus).
puncticollis Boheman, 1858: 31 (Philonthus; [preoccupied]; Type locality: China).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Fauvel, 1902a: 41 (Cafius; synonym of nauticus).
— Fauvel, 1903d: 268 (Cafius; synonym of nauticus).
— Bernhauer and Schubert, 1914: 362 (Cafius; synonym of nauticus).
— Cameron, 1932: 155 (Cafius; synonym of nauticus).
— Coiffait, 1974: 345 (Cafius; synonym of nauticus).
parallelus Kraatz, 1859: 99 (Philonthus; Type locality: Ceylan).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Fauvel, 1902a: 41 (Cafius; synonym of nauticus).
— Bernhauer, 1902b: 29 (Cafius; Ceylon).
— Bernhauer and Schubert, 1914: 362 (Cafius; synonym of nauticus).
— Cameron, 1932: 155 (Cafius; synonym of nauticus).
densiventris Fauvel, 1877: 258 (Cafius; Type locality: Australie orientale, Port Makay. Also cited from Iles Arou).
— Fauvel, 1878f: 252 (Cafius; characters; Aru Islands; Australia).
— Olliff, 1887: 507 (Cafius; characters; Australia).
— Fauvel, 1903d: 268 (Cafius; synonym of nauticus).
— Bernhauer and Schubert, 1914: 362 (Cafius; synonym of nauticus).
— Lea, 1925: 225 (Cafius; synonym of nauticus).
— Bernhauer, 1926f: 317 (Cafius; Buru).
— Cameron, 1932: 155 (Cafius; synonym of nauticus).
— Coiffait, 1974: 345 (Cafius; synonym of nauticus).
obscuricornis C. Koch, 1936, see: Remus.
occidentalis Olliff, 1887, see: Remus.
opacus LeConte, 1863a: 40 (Philonthus; [Note: Preoccupied secondary homonym in Philonthus]; Type locality: San Diego; [Note: See lectotype designation by Orth and Moore, 1980]).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Horn, 1884: 239 (Cafius; characters; California).
— Bernhauer and Schubert, 1914: 363 (Cafius; catalog).
— Scheerpeltz, 1933: 1370 (Cafius; catalog).
— C. Koch, 1936b: 188 (Cafius; characters in key; California).
— Moore and Orth, 1977a: 287 (Cafius; range extension; California; Baja California).
— Orth, Moore, and Fisher, 1978: 187 (Cafius; characters; collecting notes; California).
— Orth and Moore, 1980: 187 (Cafius; lectotype designation; characters; British Columbia; Washington; California; Baja California Norte).
— Moore and Andrews, 1985: 222 (Cafius; Baja California Norte).
▴ Distribution: Canada, USA, Mexico.
dubius LeConte, 1863a: 39 (Philonthus; [Note: Preoccupied in Philonthus]; Type locality: San Diego, California).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Bernhauer and Schubert, 1914: 363 (Cafius; synonym of opacus).
— Orth and Moore, 1980: 187 (Cafius; synonym of opacus).
pacificus Erichson, 1840: 501 (Philonthus; Type locality: Terra Van Diemenii).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Fauvel, 1877: 254 (Cafius; characters; Tasmania).
— Olliff, 1887: 504 (Cafius; characters; Australia).
— Bernhauer and Schubert, 1914: 363 (Cafius; catalog).
— Lea, 1925: 226 (Cafius; characters; Australia).
— Scheerpeltz, 1933: 1370 (Cafius; catalog).
— Bierig, 1934a: 68 (Cafius; subgenus Euremus).
— C. Koch, 1936b: 185 (Cafius; characters in key; Australia).
▴ Distribution: Australia.
littoralis Fauvel, 1877: 254 (Cafius; [preoccupied]; Type locality: Australie, Victoria, Nouvelle Galles du Sud, Queensland, Wide Bay).
— Olliff, 1887: 504 (Cafius; characters; Australia).
— Lea, 1899: 540 (Cafius; Australia).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Lea, 1925: 226 (Cafius; synonym of pacificus).
— Bierig, 1934a: 68 (Cafius; synonym ? of pacificus).
— C. Koch, 1936b: 185 (Cafius; synonym of pacificus).
parallelus Kraatz, 1859, see: nauticus Fairmaire, 1849.
postseriatus C. Koch, 1936 [Note: Nomen nudum], see: sabulosus Fauvel, 1877.
pumilus Mannerheim, 1830, see: Gabrius.
puncticeps White, 1846, see: caviceps Broun, 1886.
puncticollis Boheman, 1858, see: nauticus Fairmaire, 1849.
quadriimpressus White, 1846: 6 (Staphylinus; subgenus Gyrohypnus ?; Type locality: New Zealand).
— Fairmaire, 1847: 91 (Staphylinus; translation of original description).
— Fauvel, 1877: 247 (Xantholinus; characters; New Zealand).
— Broun, 1880: 105 (Xantholinus; subgenus Gyrohypnus; position unknown).
— Bernhauer and Schubert, 1914: 307 (Xantholinus; catalog).
— Scheerpeltz, 1933: 1314 (Xantholinus; catalog).
— C. Koch, 1936b: 180 (Cafius; characters in key; New Zealand).
— Kuschel, 1990: 26 (Cafius; endemic; winged; habitat; New Zealand).
— Emberson, 1998: 35 (Cafius; collecting notes; Chatham Islands).
▴ Distribution: New Zealand, Chatham Islands.
expuncticollis C. Koch, 1936b: 180 (Cafius; aberration of quadriimpressus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from New Zealand).
ragazzii Gestro, 1889: 32 (Cafius; Type locality: Assab).
— Bernhauer and Schubert, 1914: 363 (Cafius; catalog).
— C. Koch, 1936b: 189 (Cafius; characters in key; Red Sea region).
— Coiffait, 1974: 345 (Cafius; subgenus Pseudoremus; characters; coast of Red Sea).
▴ Distribution: “Red Sea region”.
rufescens Sharp, 1889: 44 (Cafius; Type locality: Yokohama; Niigata; Hakodate; Hong Kong).
— Bernhauer and Schubert, 1914: 363 (Cafius; catalog).
— Scheerpeltz, 1933: 1370 (Cafius; catalog).
— Nakane, 1963a: 92 (Cafius; characters; habitus photograph; Japan).
— Hasegawa and Kanie, 1992: 20 (Cafius; seashore collection; Japan).
— Watanabe, 1993d (Cafius; Sarushima Island).
— Kanehisa, Tsumuki, and Kawazu, 1994 (Cafius; chemical secretions).
▴ Distribution: Japan, China.
rufifrons Bierig, 1934a: 68 (Cafius; subgenus Euremus; Type locality: Cuba: Playa de Marianao).
— Blackwelder, 1943: 438 (Cafius; synonym of bistriatus).
— Orth and Moore, 1980: 198 (Cafius; valid species; characters; Florida).
— Frank, Carlysle, and Rey, 1986: 149, 153, 158 (Cafius; characters; discussion of biogeography; Florida; Cuba).
▴ Distribution: USA, West Indies.
sabulosus Fauvel, 1877: 253 (Cafius; Type locality: Australie septentrionale et orientale; Cap York, Port Denison, Port Makay, Gayndah, Sydney).
— Olliff, 1887: 501 (Cafius; characters; Australia).
— Lea, 1899: 540 (Cafius; Australia).
— Bernhauer and Schubert, 1914: 363 (Cafius; catalog).
— C. Koch, 1936b: 179 (Cafius; characters in key; Australia).
▴ Distribution: Australia.
postseriatus C. Koch, 1936b: 179 (Cafius; aberration of sabulosus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Australia).
sculticeps Motschulsky, 1858, see: cicatricosus Erichson, 1840.
seminitens Horn, 1884: 236 (Cafius; Type locality: California sea-coast; [Note: See lectotype designation by Orth and Moore, 1980]).
— Casey, 1885a: 314 (Bryonomus; characters).
— Bernhauer and Schubert, 1914: 363 (Cafius; catalog).
— Scheerpeltz, 1933: 1370 (Cafius; catalog).
— C. Koch, 1936b: 183 (Cafius; characters in key; California).
— Hatch, 1957: 211 (Cafius; subgenus Bryonomus; characters; British Columbia; Washington; Oregon).
— R. Dvořák, 1957a (Cafius; subgenus Bryonomus; characters).
— Smetana, 1965a: 9 (Cafius; subgenus Bryonomus; British Columbia).
— James, Moore, and Legner, 1971: 285, 287, 288 (Cafius; larval and pupal characters; California).
— Moore, 1975a: 140 (Cafius; characters).
— Orth, Moore, and Fisher, 1978: 186 (Cafius; characters; collecting notes; California).
— Orth and Moore, 1980: 183 (Cafius; lectotype designation; characters; British Columbia; Washington; Oregon; California; Baja California Norte).
— Frank, 1982: 34 (Cafius; list of parasites).
▴ Distribution: Canada, USA, Mexico.
seriatus Fauvel, 1877: 255 (Cafius; Type locality: Australie, Victoria).
— Fauvel, 1878g: 541 (Cafius; characters; Australia).
— Olliff, 1887: 506 (Cafius; characters; Australia).
— Lea, 1899: 540 (Cafius; notes; Pelsart Island).
— Bernhauer and Schubert, 1914: 363 (Cafius; catalog).
— C. Koch, 1936b: 185 (Cafius; characters in key; Australia).
▴ Distribution: Australia.
sharpi C. Koch, 1936, see: Philonthus.
speculifrons Fauvel, 1877, see: Paracraspedomerus.
splendoris Last, 1987: 52 (Cafius; Type locality: Papua New Guinea: New Guinea NE: Morobe Dist., Mt. Amingwiwa, 1000–2300 m).
subtilis Cameron, 1922: 121 (Cafius; Type locality: Jamaica).
— Scheerpeltz, 1933: 1370 (Cafius; catalog).
— Bierig, 1934a: 65 (Cafius; characters; Cuba).
— C. Koch, 1936b: 192 (Cafius; characters).
— Blackwelder, 1943: 436 (Cafius; characters; Cuba; Jamaica; Puerto Rico; St. Croix; St. Kitts; Antigua; Montserrat; Guadeloupe; Dominica).
— Ramos, 1946: 33 (Cafius; Mona Island, West Indies).
— Orth and Moore, 1980: 199 (Cafius; notes; characters).
— Frank, 1985: 62 (Cafius; Florida).
— Frank, Carlysle, and Rey, 1986: 149 (Cafius; characters).
▴ Distribution: West Indies, USA.
sulcicollis LeConte, 1863a: 40 (Philonthus; Type locality: San Diego; [Note: See lectotype designation by Orth and Moore, 1980]).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Horn, 1884: 237 (Cafius; characters; California sea-coast).
— Bernhauer and Schubert, 1914: 363 (Cafius; catalog).
— Scheerpeltz, 1933: 1370 (Cafius; catalog).
— C. Koch, 1936b: 187 (Cafius; characters in key; California).
— Leech and Moore, 1971 (Cafius; notes; California).
— Moore and Legner, 1973d (Cafius; distributional notes).
— Moore 1974a: 119 (Cafius; notes; Mexico).
— Moore and Legner, 1977: 459 (Cafius; Mexico).
— Orth, Moore, and Fisher, 1978: 187 (Cafius; characters; collecting notes; California).
— Orth and Moore, 1980: 194 (Cafius; lectotype designation; characters; California; Baja California Norte).
▴ Distribution: Mexico, USA.
tessellatus Stephens, 1833, see: xantholoma Gravenhorst, 1806.
variegatus Erichson, 1840, see: xantholoma Gravenhorst, 1806.
variolosus Sharp, 1871, see: xantholoma Gravenhorst, 1806.
velutinus Fauvel, 1877: 256 (Cafius; Type locality: Australie, Nouvelle Galles du Sud).
— Fauvel, 1878g: 542 (Cafius; characters; Australia).
— Olliff, 1887: 506 (Cafius; characters; Australia).
— Lea, 1899: 540 (Cafius; Australia).
— Bernhauer and Schubert, 1914: 363 (Cafius; catalog).
— Lea, 1925: 226 (Cafius; characters; Australia).
▴ Distribution: Australia.
vestitus Sharp, 1874: 37 (Philonthus; subgenus Cafius; Type locality: Amakusa and Iwosima).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Bernhauer and Schubert, 1914: 363 (Cafius; catalog).
— Scheerpeltz, 1933: 1370 (Cafius; catalog).
— C. Koch, 1936b: 189 (Cafius; characters in key; Japan; Korea).
— Nakane, 1963a: 92 (Cafius; characters; habitus photograph; Japan).
— Watanabe and Shibata, 1965: 321 (Cafius; subgenus Remus; Japan).
— Watanabe and Shibata, 1972: 64 (Cafius; Yaku-shima Island).
— Hasegawa and Kanie, 1992: 20 (Cafius; seashore collection; Japan).
— Watanabe, 1993d (Cafius; Sarushima Island).
— J. Li, 1993: 40 (Cafius; China).
— Kanehisa, Tsumuki, and Kawazu, 1994 (Cafius; chemical secretions).
▴ Distribution: Japan, Korea, China.
xantholoma Gravenhorst, 1806: 41 (Staphylinus; Type locality: Rostochii).
— Gyllenhal, 1810: 323 (Staphylinus; characters; Sweden).
— Germar, 1827: 3 (Staphylinus; characters; Baltic).
— Curtis, 1829: 26 (Cafius; catalog; Britain).
— Curtis, 1830: pl. 322 (Cafius; characters; Britain).
— Mannerheim, 1830: 31 (Cafius; Sweden).
— Mannerheim, 1831: 445 (Cafius; Sweden).
— Stephens, 1833: 246 (Cafius; characters; England; Isle of Wight).
— Nordmann, 1837: 126 (Gyrohypnus; Germany; Sweden).
— Westwood, 1838a: 16 (Cafius; type species).
— Zetterstedt, 1838: 64 (Staphylinus; characters; habitat; Lapland).
— Stephens, 1839: 399 (Cafius; characters; England).
— Erichson, 1840: 452 (Philonthus; characters; Germaniae septentrionalis; Sweden; England; Portugal).
— Laporte, 1840: 179 (Cafius; characters; Germany; England).
— Jacobsen, 1849: 693 (Philonthus; notes; characters; Denmark).
— Redtenbacher, 1849: 825 (Philonthus; characters; Austria).
— Kiesenwetter, 1851a: 416 (Philonthus; France).
— Küster, 1853a: 58 (Philonthus; characters; Lapland; Sweden; England; France; Spain; Portugal; Italy).
— Hardy, 1851: 27 (Philonthus; notes; England).
— Fairmaire and Laboulbène, 1856: 521 (Philonthus; characters; France).
— Staudinger, 1857: 286 (Philonthus; notes; Iceland).
— Kraatz, 1857d: 595 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 192 (Philonthus; characters; Austria).
— Peyron, 1858: 425 (Philonthus; Turkey).
— Thomson, 1859: 24 (type species of Cafius).
— Thomson, 1860: 152 (Cafius; characters; Scandinavia).
— Wollaston, 1864: 570 (Philonthus; characters; Canary Islands).
— Wollaston, 1865: 491 (Philonthus; habitat; notes; Canary Islands).
— Fauvel, 1874: 198 [= 1874a: 424] (Cafius; characters; France; Algeria; Turkey).
— Redtenbacher, 1874: 211 (Philonthus; characters; Austria).
— Seidlitz, 1875: 275 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 571 [= 1877: 427] (Gabrius; characters; France).
— Fauvel, 1878: 120 [= 1878a: 40] (Cafius; Morocco; Algeria).
— Fauvel, 1886: 52 [= 1886a: 44] (Cafius; Morocco; Tunisia; Algeria).
— Fowler, 1888: 282 (Cafius; characters; Britain).
— Seidlitz, 1891: 390 (Cafius; characters; Europe).
— Seidlitz, 1891a: 415 (Cafius; characters; Europe).
— Ganglbauer, 1895: 463 (Cafius; characters; Europe).
— Fauvel, 1902b: 106 (Cafius; Barbary; Egypt; Canaries).
— Poppius, 1905: 10 (Cafius; Orkney Islands).
— Sainte-Claire Deville, 1906: 95 (Cafius; Corsica).
— Montandon, 1908: 79 (Cafius; Romania).
— Reitter, 1909: 124 (Cafius; characters; Germany).
— Bernhauer and Schubert, 1914: 363 (Cafius; catalog).
— Johansen, 1914: 420 (Cafius; characters; Denmark).
— Portevin, 1929: 354 (Cafius; characters; France).
— Gridelli, 1930: 72 (Cafius; Libya).
— Gridelli, 1931: 123 (Cafius; characters in key).
— Scheerpeltz, 1933: 1370 (Cafius; catalog).
— C. Koch, 1936b: 168, 169 (Cafius; characters; Egypt).
— C. Koch, 1936b: 181 (Cafius; characters in key; Europe; North Africa; Canary Islands).
— Paulian, 1941: 231 (Cafius; larval characters).
— V. Hansen, 1952: 34 (Cafius; characters; Denmark).
— Cameron, 1951: 283 (Cafius; Egypt).
— Larsson and Gígja, 1959: 88 (Cafius; biological notes; Iceland).
— Scheerpeltz, 1960d: 136 (Cafius; subgenus Cafius; Portugal).
— Palm, 1963b: 60 (Cafius; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 173 (Cafius; characters; central Europe).
— Scheerpeltz, 1965c: 508 (Cafius; Sicily).
— Horion, 1965: 120 (Cafius; sea coast of Europe and North Africa).
— Pototskaia, 1967: 53 (Cafius; larval characters).
— Coiffait and Saiz, 1968: 362, 363 (Cafius; characters; Chile).
— James, Moore, and Legner, 1971: 282 (Cafius; larval characters).
— Saiz, 1971b: 349 (Cafius; Chile).
— Coiffait, 1974: 347 (Cafius; characters; Europe; Scandinavia; North Africa; Canary Islands; southern Africa; Red Sea).
— Pope, 1977: 30 (Cafius; Britain).
— Hinton, 1981: 661 (Cafius; characters of egg).
— Frank, 1982: 34 (Cafius; list of parasites).
— Elliott, King, and Fordy, 1983 (Cafius; habitat; natural history).
— Tóth, 1984: 40 (Cafius; characters).
— Elliott and King, 1986 (Cafius; adaptations for intertidal living).
— Segers, 1986: 46 (Cafius; checklist of species; Belgium).
— Lucht, 1987: 104 (Cafius; checklist; Germany; Poland; France; Benelux; Denmark; Sweden).
— Haghebaert, 1989a: 278 (Cafius; notes; Camargue).
— Israelson, 1990: 2 (Cafius; Azores).
— Borges, 1990: Table II (Cafius; Azores).
— Welch, 1993: 229 (Cafius; ovariole number and ovary structure).
— Hernández, Outerelo, and Gamarra, 1994: 187 (Cafius; checklist of species; Canary Islands).
— Sparacio, 1995: 161 (Cafius; characters; notes; Sicily).
— Ciceroni and Zanetti, 1995: 27 (Cafius; Italy).
— Dauphin and Tamisier, 1997: 94 (Cafius; notes; France).
— Owen, 1997a: 149 (Cafius; collecting sites; Ireland).
— Assing and Wunderle, 2001: 36 (Cafius; Cyprus).
▴ Distribution: Iceland, Europe, Azores, Canary Islands, Morocco, Algeria, Tunisia, Libya, Egypt, Turkey, Chile.
lateralis Stephens, 1833: 246 (Cafius; Type locality: Essex; Kent; Sussex).
— Curtis, 1829: 26 (Cafius; nomen nudum; catalog; Britain).
— Holme, 1837: 63 (Cafius; notes; Scilly Islands).
— Stephens, 1839: 399 (Cafius; variety of xantholoma; characters; England).
— Hardy, 1851: 28 (Philonthus; synonym of xantholoma).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of xantholoma).
— Bernhauer and Schubert, 1914: 363 (Cafius; synonym of xantholoma).
— Coiffait, 1974: 348 (Cafius; synonym of xantholoma).
littoralis Stephens, 1833: 247 (Cafius; variety of lateralis; Type locality: Not cited, presumably Britain).
tessellatus Stephens, 1833: 247 (Cafius; variety of lateralis; Type locality: Not cited, presumably Britain).
variegatus Erichson, 1840: 453 (Philonthus; Type locality: Sicilia).
— Fauvel, 1869: 489 (Philonthus; synonym of xantholoma).
— Fauvel, 1874: 198 [= 1874a: 424] (Staphylinus; synonym of xantholoma).
— Ganglbauer, 1895: 463 (Cafius; variety of xantholoma).
— Bernhauer and Schubert, 1914: 363 (Cafius; variety of xantholoma).
— Donisthorpe, 1930: 96 (Cafius; aberration of xantholoma; characters; Britain).
— Scheerpeltz, 1933: 1370 (Cafius; variety of xantholoma).
— Coiffait, 1974: 348 (Cafius; variety of xantholoma).
— Pope, 1977: 30 (Cafius; synonym of xantholoma).
variolosus Sharp, 1871: 181 (Philonthus; form of xantholoma; Type locality: England and Scotland).
— Fauvel, 1874: 198 [= 1874a: 424] (Staphylinus; synonym of xantholoma).
— Ganglbauer, 1895: 463 (Cafius; variety of xantholoma).
— Scheerpeltz, 1933: 1371 (Cafius; variety of xantholoma).
— Coiffait, 1974: 348 (Cafius; variety of xantholoma).
— Pope, 1977: 30 (Cafius; synonym of xantholoma).
keysianus Donisthorpe, 1930: 97 (Cafius; aberration of xantholoma; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Plymouth district; Tregantle; Isle of Wight).
heroopoliticus C. Koch, 1936b: 169 (Cafius; subspecies of xantholoma; Type locality: Sinai: Tor).
zealandicus Cameron, 1947d: 705 (Cafius; Type locality: New Zealand: Auckland: Turgirangi, Waikanera Range).
Chroaptomus
[3 species; Neotropical Region]
Chroaptomus Sharp, 1885: 437 (species included: flagrans, regalis). Type species: Philonthus flagrans Erichson, fixed by subsequent designation by R. Lucas, 1920: 184.
— Bernhauer and Schubert, 1914: 366 (world catalog; 3 species).
— Scheerpeltz, 1933: 1376 (world catalog supplement).
— Blackwelder, 1952: 101 (type species).
— Hayashi, 2000: 23 (notes on empodial setae).
coelites Erichson, 1840, see: flagrans Erichson, 1840.
flagrans Erichson, 1840: 441 (Philonthus; Type locality: Mexico; Columbia).
— Solsky, 1868a: 127 (Philonthus; Mexico).
— Sharp, 1885: 437 (Chroaptomus; characters; Mexico; Guatemala; Costa Rica; Panama; Colombia; Venezuela).
— Fauvel, 1891: 111 (Philonthus; Venezuela).
— Bernhauer and Schubert, 1914: 366 (Chroaptomus; catalog; Ecuador, Peru, Bolivia).
— Scheerpeltz, 1933: 1376 (Chroaptomus; catalog; Argentina; Brazil).
— Marcuzzi, 1977: 12 (Chroaptomus; Venezuela).
— Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 61 (Chroaptomus; necrophilous species; distribution in Mexico).
▴ Distribution: Mexico, Guatemala, Costa Rica, Panama, Colombia, Venezuela, Brazil, Ecuador, Peru, Bolivia, Argentina.
coelites Erichson, 1840: 440 (Philonthus; Type locality: Brasilia).
— Bernhauer and Schubert, 1914: 332 (Philonthus; catalog).
— Scheerpeltz, 1933: 1376 (Chroaptomus; variety of coelites).
regalis Sharp, 1885: 438 (Chroaptomus; Type locality: Panama, Bugaba 800 to 1500 feet, Volcan de Chiriqui 4000 to 6000 feet, Caldera 1200 feet).
richteri Bernhauer, 1912d: 44 (Philonthus; subgenus Chroaptomus; Type locality: Argentinien: Prov. Tucuman).
Craspedomerus
[13 species; Palaearctic and Oriental Regions]
Craspedomerus Bernhauer, 1911: 88 (species included: glenoides). Type species: Philonthus glenoides Schubert, fixed by monotypy.
— Bernhauer and Schubert, 1914: 400 (world catalog; 1 species).
— Winkler, 1925: 387 (catalog for Palaearctic region).
— Cameron, 1925: 71 (catalog of Indian species).
— Cameron, 1932: 248 (characters; key to species of British India).
— Scheerpeltz, 1933: 1410 (world catalog supplement).
— Blackwelder, 1952: 110 (type species: glenoides).
— Moore, 1960: 100 (characters in key).
— Scheerpeltz, 1976a: 148 (key to species of Himalayan region).
— Hayashi, 2000: 23 (notes on empodial setae).
beckeri Bernhauer, 1934: 11 (Craspedomerus; Type locality: West-China: Prov. Szechuan, Kinfushan).
bernhaueri Cameron, 1926b: 364 (Craspedomerus; Type locality: Chakrata district).
— Cameron, 1932: 249 (Craspedomerus; characters; India).
— Scheerpeltz, 1933: 1410 (Craspedomerus; catalog).
— Scheerpeltz, 1976: 43 (Craspedomerus; Nepal).
— Scheerpeltz, 1976a: 150 (Craspedomerus; characters).
— Coiffait, 1982: 32 (Craspedomerus; Nepal).
— Coiffait, 1982a: 233 (Craspedomerus; Nepal).
▴ Distribution: India, Nepal.
birmanus Scheerpeltz, 1965: 257 (Craspedomerus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).
caeruleipennis Cameron, 1932, see: Anisolinus.
cyanipennis Scheerpeltz, 1976a: 144 (Craspedomerus; Type locality: Nepal: Thodung, 3100 m).
— Coiffait, 1982: 32 (Craspedomerus; Nepal).
— Coiffait, 1982a: 233 (Craspedomerus; Nepal).
▴ Distribution: Nepal.
ganeshensis Coiffait, 1983d: 169 (Craspedomerus; Type locality: Nepal: Ganesh Himal, entre Salmi et Rupchet, 1800 à 3400 m).
glenoides Schubert, 1908: 622 (Philonthus; Type locality: Kulu, Himalaya).
— Bernhauer, 1911: 88 (Craspedomerus).
— Bernhauer and Schubert, 1914: 400 (Craspedomerus; catalog).
— Cameron, 1932: 249 (Craspedomerus; characters; Himalayan region).
— Scheerpeltz, 1933: 1410 (Craspedomerus; catalog).
— Scheerpeltz, 1976a: 149 (Craspedomerus; characters).
▴ Distribution: India.
malaisei Scheerpeltz, 1965: 259 (Craspedomerus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).
montanus Coiffait, 1982: 76 (Craspedomerus; Type locality: Nepal: westl. Dhaulagiri, zw. Pelma und Pass Jungla Banjyang, 2100–2700 m).
nepalensis Scheerpeltz, 1976a: 146 (Craspedomerus; Type locality: Nepal: Salung-Phuleli, 2400–3000 m).
nepalicus Coiffait, 1977: 246 (Philonthus; Type locality: Environs de Maharigaon, 3000–3500 m, région de Jumla, Népal occidental).
sinetuber Coiffait, 1977b: 223 (Philonthus; Type locality: Bhutan: Dechhi Paka, 3300 m).
— Coiffait, 1979c: 319 (Craspedomerus).
— Schillhammer, 1992: 66 (Craspedomerus).
▴ Distribution: Bhutan.
tricoloricornis Coiffait, 1977b: 222 (Philonthus; Type locality: Bhutan: Dechhi Paka).
— Coiffait, 1979c: 319 (Craspedomerus).
— Schillhammer, 1992: 66 (Craspedomerus).
▴ Distribution: Bhutan.
violaceipennis Cameron, 1928c: 566 (Craspedomerus; Type locality: Sikkim: Karponang, 10,000 feet; Phadam Chen, 9000 feet. Tibet: Chumbi Valley, 11,000 feet; Yatung, 10,000 feet).
Diplostictus
[1 species; Oceanic Region]
Diplostictus Fauvel, 1874c: 437 (species included: chenui). Type species: Staphylinus chenui Perroud and Montrousier, fixed by monotypy.
— Fauvel, 1877: 262 (note).
— Duvivier, 1883: 149 (catalog).
— Bernhauer and Schubert, 1914: 322 (world catalog; 1 species).
— Scheerpeltz, 1933: 1326 (world catalog supplement).
— Blackwelder, 1952: 129 (type species: chenui).
buergersi Bernhauer, 1915, see: Pachypelmopus.
chenui Perroud and Montrousier, 1864: 86 (Staphylinus; Type locality: Nouvelle Calédonie: Kanala).
— Fauvel, 1869: 486 (Philonthus).
— Fauvel, 1874c: 437 (Diplostictus; characters; New Caledonia).
— Fauvel, 1877: 262 (Diplostictus).
— Fauvel, 1889b: 263 (Diplostictus; New Caledonia).
— Fauvel, 1903d: 270 (Diplostictus; New Caledonia).
— Bernhauer and Schubert, 1914: 322 (Diplostictus; catalog).
— Scheerpeltz, 1966e: 406 (Diplostictus; characters; New Caledonia).
— Legner and Moore, 1977: 176 (Diplostictus; found under bark).
▴ Distribution: New Caledonia.
Dorcophilonthus
[1 species; Oriental Region]
Dorcophilonthus Hayashi, 2000: 15 (species included: clavicornis). Type species: Dorcophilonthus clavicornis Hayashi, fixed by original designation and monotypy.
clavicornis Hayashi, 2000: 22 (Dorcophilonthus; Type locality: NE India: Megalaya: Jowai env., 1400 m, 25°30′N, 92°10′E).
Ecitophytes
[1 species; Neotropical Region]
Ecitophytes Wasmann, 1923: lxii (species included: coniceps). Type species: Ecitophytes coniceps Wasmann, fixed by monotypy.
coniceps Wasmann, 1923: lxii (Ecitophytes; Type locality: Minas Geraës).
Endeius
[8 species; Neotropical Region]
Endeius Coiffait and Saiz, 1968: 355 (species included: nitidipennis, loensis, punctipennis, subpunctipennis). Type species: Endeius loensis Coiffait and Saiz, fixed by original designation.
franzi Saiz, 1971b: 345 (Endeius; Type locality: Chile: El Arrayán, Santiago).
loensis Coiffait and Saiz, 1968: 357 (Endeius; Type locality: Chili: Guatin, Antofagasta).
lugubris Saiz, 1971b: 347 (Endeius; Type locality: Chile: Río Los Molles, Coquimbo).
multipunctatus Coiffait, 1981g: 297 (Endeius; Type locality: Ile Galapagos: Ile Santa Cruz, versant sud-est, basse altitude).
nitidipennis Solier, 1849: 316 (Staphylinus; Type locality: Chile).
— Kraatz, 1859a: 4 (Philonthus; notes).
— Fairmaire and Germain, 1861: 432 (Philonthus; characters; Chile).
— Fauvel, 1866a: 344 [= 1866b: 95] (Philonthus; characters; Chile).
— Bernhauer and Schubert, 1914: 348 (Philonthus; catalog).
— Bernhauer, 1921a: 42 (Philonthus; Juan Fernandez Islands).
— Scheerpeltz, 1933: 1353 (Philonthus; catalog; Argentina).
— Coiffait and Saiz, 1968: 359 (Endeius; characters; Chile).
— Saiz, 1969a: 9 (Endeius; notes).
— Saiz, 1971b: 343, 345 (Endeius; characters; Chile).
▴ Distribution: Argentina, Chile, Juan Fernandez.
ovaliceps Coiffait, 1981g: 296 (Endeius; Type locality: Ile Galapagos: Ile San Cristobal, région de Progreso).
punctipennis Solier, 1849: 319 (Staphylinus; [preoccupied]; Type locality: Quillota; [Note: See lectotype designation by Coiffait and Saiz, 1968]. Originally cited from Chile: Santa Rosa).
— Kraatz, 1859a: 5 (Philonthus; notes).
— Fairmaire and Germain, 1861: 432 (Philonthus; characters; Chile).
— Fauvel, 1864a: 125 (Philonthus; notes; Chile).
— Fauvel, 1866a: 345 [= 1866b: 96] (Philonthus; characters; Chile).
— Bernhauer and Schubert, 1914: 352 (Philonthus; catalog).
— Scheerpeltz, 1933: 1357 (Philonthus; catalog).
— Coiffait and Saiz, 1968: 358 (Endeius; lectotype designation: Quillota; characters; Chile).
— Saiz, 1969a: 9 (Endeius; notes).
— Saiz, 1971b: 344 (Endeius; characters; Chile; Argentina).
— Scheerpeltz, 1972: 20 (Philonthus; Argentina).
— Saiz, Solervicens, and Ojeda, 1989: 52 (Endeius; collecting notes; habitus; Chile; Argentina).
— Herman, 2001: 43 (Endeius; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Chile, Argentina.
subpunctipennis Coiffait and Saiz, 1968: 358 (Endeius; Type locality: Chili).
Erichsonius
[155 species; Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, and Australian Regions]
Erichsonius Schubert, 1911, see: Pancarpius.
Erichsonius Fauvel, 1874: 201 [= 1874a: 427] (species included: agnatus, filiformis, gratus, humilis, ludicrus, paederoides, praelongus, pumilus, sobrinus, terminalis, umbripennis, cinerascens, rivularis, signaticornis, villosulus, semipunctatus, procerulus, prolixus, orbus). Type species: Staphylinus cinerascens Gravenhorst, fixed by subsequent designation by Lucas, 1920: 73, who designated the type species for the objective synonym Actobius.
— Thomson, 1858: 29 (cited as Rembus; [Note: Misidentification of Erichsonius]; characters; species included: cinerascens).
— Thomson, 1859: 25 (cited as Remus; [Note: Thomson's use of Remus is a misidentification. Tottenham, 1949: 371 cited it with a type species and as a synonym of Erichsonius as though it were an available name]; type species).
— Thomson, 1860: 167 (cited as Remus; [Note: Thomson's use of Remus is a Misidentification. Tottenham, 1949: 371 cited it with a type species and as a synonym of Erichsonius as though it were an available name]; characters).
— Fauvel, 1875a: XXIX [= 1875b: 231] (synonym of Actobius).
— Fauvel, 1876a: 72 [= 1876: 257] (synonym of Actobius).
— Duvivier, 1883: 151 (catalog).
— Bernhauer and Schubert, 1914: 325 (synonym of Actobius).
— Cameron, 1932: 59 (synonym of Actobius).
— Blackwelder, 1943: 440 (characters; type species: cinerascens; key to species of West Indies).
— Blackwelder, 1944: 131 (checklist of species from Mexico, Central and South America, and the West Indies).
— Kloet and Hincks, 1945: 165 (list of British species; type species: cinerascens).
— Tottenham, 1949: 371, 417 (type species: cinerascens Gravenhorst; notes; checklist of British species).
— Blackwelder, 1952: 151 (type species).
— Tottenham, 1956: 260 (characters; notes; key to African species).
— Smetana, 1958: 140 (characters; habitus of one species; type species: cinerascens; key to Czechoslovakian species).
— Smetana, 1959a: 209 (list of species collected in Albania).
— Watanabe, 1963: 7 (key to Japanese species).
— Coiffait, 1963: 7, 9 (characters; key to subgenera).
— Székessy, 1963: 53 (characters).
— Lohse, 1964: 172 (characters; key to central European species).
— Moore, 1965: 98, 100 (characters; notes).
— Coiffait, 1965c: 843 (characters; key to some species).
— Scheerpeltz, 1968a: 65 (catalog of Austrian species).
— Shibata, 1973c: 57 (checklist of species of Taiwan).
— Tikhomirova, 1973a: 182 (checklist of species of USSR).
— Coiffait, 1974: 6 (characters; key to species of western Palaearctic region).
— Frank, 1975 (characters; revision of and key to New World species).
— Moore and Legner, 1975: 12 (catalog; North America north of Mexico).
— Pope, 1977: 29 (list of British species).
— Topp, 1978: 312 (larval characters).
— Moore and Legner, 1979: 18 (characters; notes).
— Muona, 1979: 14 (list of Scandinavian species).
— Uhlig, 1979: 252 (some species collected in Germany).
— M. Dvořák, 1979: 115 (some species collected in Slovakia).
— Uhlig, Vogel, and Sieber, 1980: 246 (some species collected in Germany).
— Burakowski, Mroczkowski, and Stefańska, 1980: 31 (catalog; Poland).
— Uhlig and Vogler, 1981: 110 (list of some species collected in Germany; habitat and collecting notes).
— Frank, 1981b: 100 (discussion of characters and phylogenetic relationships; partial subgeneric classification).
— Shibata, 1983: 89 (checklist of species of Japan).
— Hammond, 1984: 205 (checklist of species of Borneo).
— Tóth, 1984: 37 (characters; habitus; key to species of Hungary).
— Frank, 1986a: 370 (checklist of species of Florida).
— Muona and Viramo, 1986: 13 (list of species of northeastern Finland).
— Segers, 1986: 42 (checklist of species; Belgium).
— Drugmand 1987b (comments on distribution of Belgian species).
— Lohse and Lucht, 1989: 160 (supplement to key to species of Lohse, 1964).
— Masch and Uhlig, 1990: 232 (revision and key to Ethiopian region species of the robustus group).
— Uhlig and Sterrenburg, 1990: 167 (key to species of Benelux countries).
— Uhlig, 1991b (summary of research on genus; estimate of number of species).
— Campbell and Davies, 1991: 117 (checklist of species; Alaska and Canada).
— Uhlig and Watanabe, 1992 (redescription; revision of and key to Japanese species; Zoogeography).
— Silfverberg, 1992: 16 (list of species of Fennoscandia, Denmark, and the Baltic States).
— Masch, 1993 (phylogeny of of the Ethiopian robustus group).
— Smetana, 1995e: 50 (characters).
— Ciceroni and Zanetti, 1995: 27 (list of species of Italy).
— M. Hansen, 1996: 104 (list of Danish species).
— Schmidt, 1996: 211 (characters of egg, larvae, and pupa).
— Cho, 1996a: 162 (characters; species of Korea).
— Downie and Arnett, 1996: 372 (characters; key to species of Northeastern North America).
— Lundgren, 1998: 51 (list of species from Florida; records for other States listed for each species).
— Assing, Frisch, Kahlen, et al., 1998: 137 (note).
— Newton, Thayer, Ashe, and Chandler, 2000: 396 (19 Nearctic species; notes; characters in key).
— Hayashi, 2000: 23 (notes on empodial setae).
Actobius Fauvel, 1875a: XXIX [= 1875b: 231] (replacement name for Erichsonius Fauvel; [Note: An unnecessary replacement name]). Type species: Staphylinus cinerascens Gravenhorst, fixed by subsequent designation by Lucas, 1920: 72.
— Horn, 1884: 225 (characters; key to North American species).
— Sharp, 1885: 457 (characters; Mexican and Central American species).
— Fowler, 1888: 283 (notes; key to British species).
— Heyden, Reitter, and Weise, 1891: 102 (list of species of Europe and Caucasus).
— Ganglbauer, 1895: 465 (characters).
— Fauvel, 1897d: 298 (catalog of species of Barbary; Canaries; Madeira; Azores).
— Heyden, Reitter, and Weise, 1906: 163 (list of species of Europe and Caucasus).
— Reitter, 1909: 116, 123 (characters; key to species).
— Blatchley, 1910: 391 (characters; key to Indiana species).
— Petri, 1912: 71 (list of species of Siebenbürgen).
— Bernhauer and Schubert, 1914: 324 (world catalog; 19 species).
— Johansen, 1914: 422 (characters).
— Leng, 1920: 106 (catalog of North American species).
— Cameron, 1921a: 354, 404 (characters; key to Singapore species; catalog).
— Winkler, 1925: 374 (catalog for Palaearctic region).
— Cameron, 1925: 54 (catalog of Indian species).
— Porta, 1926: 99 (characters; key to species of Italy).
— Portevin, 1929: 355 (characters; key to French species).
— Roubal, 1930: 367 (catalog; Slovakia).
— Cameron, 1932: 59 (characters; key to species of British India).
— Joy, 1932: 126 (key to British species).
— Scheerpeltz, 1933: 1328 (world catalog supplement).
— Normand, 1935: 370 (list of species of Tunisia).
— Tottenham, 1939d: 228 (type species designation; genus properly called Erichsonius).
— Blackwelder, 1943: 440 (synonym of Erichsonius; type species: cinerascens).
— Bernhauer, 1944: 148 (key to Palaearctic species).
— Kloet and Hincks, 1945: 165 (synonym of Erichsonius).
— Tottenham, 1949: 371 (synonym of Erichsonius; type species: cinerascens Gravenhorst).
— V. Hansen, 1952: 32 (characters; Danish species).
— Blackwelder, 1952: 37 (type species).
— Hatch, 1957: 173, 174 (characters; key to species of Pacific Northwest).
— Kocher, 1958: 143 (checklist of species; Morocco).
— Ferreira, 1962a: 16 (list of Portuguese species).
— Palm, 1963b: 59 (characters; species of Sweden).
— Lohse, 1964: 172 (synonym of Erichsonius).
— Coiffait, 1974: 6 (synonym of Erichsonius).
— Frank, 1975: 179 (synonym of Erichsonius).
— Pope, 1977: 29 (synonym of Erichsonius).
— Smetana, 1993b: 48 (synonym of Erichsonius).
Sectophilonthus Tottenham, 1949b: 358 (subgenus of Philonthus; species included: biparamerosus). Type species: Philonthus (Sectophilonthus) biparamerosus Tottenham, fixed by monotypy.
— Blackwelder, 1952: 349 (type species: biparamerosus).
— Coiffait, 1974: 7 (synonym of Erichsonius).
— Abdullah, 1975 (discussion of aedeagal characters).
— Lohse and Lucht, 1989: 161 (subgenus of Erichsonius; characters; key to species of central Europe).
Parerichsonius Coiffait, 1963: 9 (subgenus of Erichsonius; species included: signaticornis). Type species: Philonthus signaticornis Mulsant and Rey, fixed by original designation and monotypy.
— Coiffait, 1965c: 843, 845 (subgenus of Erichsonius; characters; key to species).
— Coiffait, 1974: 7, 8, 10 (subgenus of Erichsonius; characters; key to species of western Palaearctic region).
— Frank, 1981b: 103 (synonym of Erichsonius [Sectophilonthus]).
adustus Sharp, 1887, see: Neobisnius.
aequiventris Tottenham, 1956: 265 (Erichsonius; [Note: Although Tottenham attributed this name to Cameron as a name to be published, Tottenham validated it in a key]; Type locality: Angola).
affinis Cameron, 1926b: 350 (Actobius; Type locality: Dehra Dun and Mussorie Districts).
— Cameron, 1932: 61 (Actobius; characters; India).
— Scheerpeltz, 1933: 1328 (Actobius; catalog).
— Bernhauer, 1944: 148 (Actobius; characters in key).
— Coiffait, 1965c: 844 (Erichsonius; subgenus Erichsonius; characters; Himalaya).
— Coiffait, 1974: 14 (Erichsonius; subgenus Erichsonius; characters; Himalayan region).
— Frank, 1981b: 103 (Erichsonius; subgenus Sectophilonthus).
▴ Distribution: India.
afghanicus Tronquet, 1981: 77 (Erichsonius; Type locality: Afghanistan: Sairobi).
algericus Coiffait, 1965: 845, 847 (Erichsonius; subgenus Parerichsonius; Type locality: Algérie: Mouzaïa près de Médéa).
— Coiffait, 1974: 16 (Erichsonius; subgenus Parerichsonius; characters; Algeria; Morocco; Iberian Peninsula?).
— Frank, 1981b: 103 (Erichsonius; subgenus Sectophilonthus).
▴ Distribution: Algeria, Morocco, “Iberian Peninsula” ?.
allardi Levasseur, 1969: 883 (Erichsonius; Type locality: région de Kolwezi, Zilo, Katanga).
alternans Sharp, 1885, see: Neobisnius.
altissimus Bernhauer, 1934g: 235 (Actobius; Type locality: Ruwenzori, 4200 m).
alumnus Frank, 1975: 196 (Erichsonius; Type locality: Massachusetts: Middlesex Co.: Natick. Also cited from Quebec, Ontario, Massachusetts, New York, New Jersey, Maryland, District of Columbia, Virginia, North Carolina and Georgia).
— Frank, 1981b: 99, 103 (Erichsonius; subgenus Erichsonius; South Dakota; Wisconsin; Georgia; Florida).
— Schmidt, 1996: 206, 213 (Erichsonius; characters of egg, larvae, and pupa; natural history; Nebraska).
— Downie and Arnett, 1996: 374 (Erichsonius; characters; USA).
— Schmidt, 1999: 108 (Erichsonius; food for adults and larvae used for rearing species; rearing methods).
▴ Distribution: Canada, USA.
ampliceps Tottenham, 1962: 148 (Erichsonius; Type locality: Tanganyika Terr.: Ngorongoro, région bocagère, 2300 m. Also cited from Kenya).
bicolor Tottenham, 1962: 149 (Erichsonius; variety of ampliceps; [Note: Nomen nudum (Article 15.2)]; cited from Tanganyika Terr.: Mt. Oldeani, versant Est, 2350–2500 m).
andringitranus Jarrige, 1978: 283 (Erichsonius; subgenus Parerichsonius; Type locality: Andringitra Est, Anjavidilava, 1850–1950 m).
antiquus Frank, 1981b: 97 (Erichsonius; Type locality: Arizona: Cochise Co.: Chiricahua Mts., Rucker Canyon).
arduus Sharp, 1885, see: Neobisnius.
armeniacus Coiffait, 1965: 844, 846 (Erichsonius; subgenus Erichsonius; Type locality: Kafan, R.S.S. d'Arménie).
— Coiffait, 1974: 15 (Erichsonius; subgenus Erichsonius; characters; Armenia).
— Frank, 1981b: 103 (Erichsonius; subgenus Sectophilonthus).
▴ Distribution: Armenia.
assamensis Cameron, 1932: 61 (Actobius; Type locality: Assam: Naga Hills, alt. 5000 feet).
bamakoensis Levasseur, 1969: 893 (Erichsonius; Type locality: Mali: Bamako, cascade d'Oyanoko).
basalis Motschulsky, 1858: 664 (Philonthus; Type locality: Indes orientales).
— Bernhauer and Schubert, 1914: 322 (Neobisnius; catalog).
— Bernhauer, 1922c: 231 (Actobius; Formosa).
— Cameron, 1932: 60 (Actobius; characters; Ceylon; India).
— Scheerpeltz, 1933: 1328 (Actobius; catalog).
— Bernhauer, 1944: 148 (Actobius; characters in key).
— Frank, 1982: 35 (Erichsonius; list of parasites).
▴ Distribution: India, Sri Lanka, Taiwan.
castaneipennis Kraatz, 1859: 96 (Philonthus; Type locality: India orientalis; Ceylan).
— Fauvel, 1902a: 41 (Philonthus; synonym of basalis).
— Bernhauer, 1902b: 28 (Philonthus; Ceylon).
— Bernhauer and Schubert, 1914: 322 (Neobisnius; synonym of basalis).
— Cameron, 1932: 60 (Actobius; synonym of basalis).
— Scheerpeltz, 1933: 1328 (Actobius; synonym of basalis).
humeralis Cameron, 1920a: 214 (Actobius; Type locality: Nilgiri Hills).
— Cameron, 1932: 60 (Actobius; synonym of basalis).
— Scheerpeltz, 1933: 1328 (Actobius; variety of basalis).
— Cameron, 1934: 19 (Actobius; variety of basalis; India).
basilewskyi Tottenham, 1962: 149 (Erichsonius; Type locality: Tanganyika Terr.: Uluguru Mts., sommet du Kidunda, 1800–1950 m).
berei Tottenham, 1953b: 493 (Actobius; Type locality: Ruwenzori: above Lake Bujuku, 12,750 ft.).
bicolor Tottenham, 1962 [Note: Nomen nudum], see: ampliceps Tottenham, 1962.
biparamerosus Tottenham, 1949, see: orphanulus Erichson, 1840.
blanci Jarrige, 1978: 280 (Erichsonius; subgenus Parerichsonius; Type locality: Andringitra Centre, zone sommitale, N-O du pic Boby, rivière Sahanambo, 2500 m).
bolgius Tottenham, 1953b: 494 (Actobius; Type locality: Uganda: Kawanda, near Kampala).
boops Fauvel, 1905, see: Neobisnius.
brachycephalus Frank, 1975: 186 (Erichsonius; Type locality: New Jersey: Cape May Co., Anglesea. Also cited from Maine, Massachusetts, New Jersey, Illinois, and Texas).
— Frank, 1981b: 100, 103 (Erichsonius; subgenus Sectophilonthus; Virginia).
— Downie and Arnett, 1996: 373 (Erichsonius; characters; USA).
▴ Distribution: USA.
brincki Scheerpeltz, 1974: 138 (Erichsonius; Type locality: Natal: Royal Natal National Park, The Cascades, 5200 ft.).
brunneicornis Jarrige, 1970: 46 (Erichsonius; subgenus Parerichsonius; Type locality: Madagascar: Mt. Tsaratanana, 1500 m).
capensis Cameron, 1945c: 714 (Actobius; Type locality: South Africa: Simon's Town).
— Tottenham, 1956: 264 (Erichsonius; characters in key).
— Uhlig, 1997: 275 (Erichsonius; subgenus Sectophilonthus; characters; South Africa).
▴ Distribution: South Africa.
capribarbatus Masch and Uhlig, 1990: 257 (Erichsonius; subgenus Sectophilonthus; Type locality: Malawi: Viphya-Mts., Chamambo Forest).
— Masch, 1993: 187, 206 (Erichsonius; subgenus Sectophilonthus; phylogeny of robustus species group).
▴ Distribution: Malawi.
castaneipennis Kraatz, 1859, see: basalis Motschulsky, 1858.
chinensis Bernhauer, 1939k: 592 (Actobius; Type locality: N. W. China: Tienmuschan).
cinerascens Gravenhorst, 1802: 49 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).
— Latreille, 1804: 334 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 117 (Staphylinus; characters; Germany).
— Gyllenhal, 1810: 361 (Staphylinus; characters; Sweden).
— Gyllenhal, 1827: 482 (Staphylinus; note).
— Curtis, 1829: 26 (Gabrius; catalog; Britain).
— C. Sahlberg, 1830: 336 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 31 (Staphylinus; Russia; Sweden).
— Mannerheim, 1831: 445 (Staphylinus; Russia; Sweden).
— Nordmann, 1837: 108 (Philonthus; Germany; Sweden).
— Erichson, 1839a: 478 (Philonthus; characters; Germany).
— Heer, 1839: 272 (Philonthus; characters; Switzerland).
— Erichson, 1840: 509 (Philonthus; characters; Europe).
— Laporte, 1840: 179 (Staphylinus; cited as cineraceus; characters; Germany).
— Jacobsen, 1849: 705 (Philonthus; characters; Denmark).
— Kiesenwetter, 1849: 20 (Philonthus; characters; Europe).
— Redtenbacher, 1849: 707 (Philonthus; characters; Austria).
— Fairmaire and Laboulbène, 1856: 531 (Philonthus; characters; France).
— Kraatz, 1857d: 622 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 197 (Philonthus; characters; Austria).
— Thomson, 1858: 29 (Remus, cited as Rembus; type species).
— Thomson, 1859: 25 (type species of Remus).
— Thomson, 1860: 167 (Remus; characters; Scandinavia).
— Fauvel, 1874: 202 [= 1874a: 428] (Erichsonius; characters; France; North America [Note: According to Frank, 1975: 201 this North American record of cinerascens may be a misidentification of nanus]).
— Redtenbacher, 1874: 215 (Philonthus; characters; Austria).
— Seidlitz, 1875: 279 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 603 [= 1877: 459] (Remus; characters; France).
— J. Sahlberg, 1876: 22 (Remus; Finland).
— Horn, 1884: 225 (Actobius; characters; Michigan to Florida [Note: According to Frank, 1975: 201 this North American record of cinerascens may be a misidentification of nanus]).
— Fowler, 1888: 284 (Actobius; characters; Britain).
— Ganglbauer, 1895: 465 (Actobius; characters; Europe; North America [Note: According to Frank, 1975: 201 this North American record of cinerascens may be a misidentification of nanus]).
— Sainte-Claire Deville, 1906: 94 (Actobius; Corsica).
— Reitter, 1909: 123 (Actobius; characters; Germany).
— Blatchley, 1910: 392 (Actobius; characters; Indiana [Note: According to Frank, 1975: 201 this North American record of cinerascens may be a misidentification of nanus]).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
— Johansen, 1914: 423 (Bisnius; characters; Denmark).
— Notman, 1919: 136 (Actobius; New York [Note: According to Frank, 1975: 201 this North American record of cinerascens may be a misidentification of nanus]).
— Notman, 1920b: 705 (Actobius; Florida [Note: According to Frank, 1975: 201 this North American record of cinerascens may be a misidentification of nanus]).
— Portevin, 1929: 355 (Actobius; characters; France).
— Scheerpeltz, 1933: 1328 (Actobius; catalog).
— Peyerimhoff, 1937: 264 (Actobius; Morocco).
— Bernhauer, 1944: 148 (Actobius; characters in key).
— V. Hansen, 1952: 32 (Actobius; characters; Denmark).
— Hatch, 1957: 174 (Actobius; characters; British Columbia; Washington [Note: According to Frank, 1975: 201 this North American record of cinerascens may be a misidentification of nanus]).
— Smetana, 1958: 141 (Erichsonius; characters; ecology; Czechoslovakia).
— Palm, 1963b: 59 (Actobius; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 173 (Erichsonius; characters; central Europe).
— Smetana, 1964d: 73 (Erichsonius; Czechoslovakia).
— Focarile, 1964: 66–68 (Actobius; characters; notes; Italy).
— Coiffait, 1965c: 844 (Erichsonius; subgenus Erichsonius; characters; Europe, Scandinavia; Caucasus; Transcaucasus).
— Horion, 1965: 118 (Erichsonius; widespread in Europe from high north to central Europe to southern Europe in Spain, Corsica, Italy, Dalmatia, Bosnia Hercegovina and Ukraine).
— Szujecki, 1968a: 727 (Neobisnius; Poland).
— Coiffait, 1974: 12 (Erichsonius; subgenus Erichsonius; characters; France; central Europe; Corsica).
— Pope, 1977: 29 (Erichsonius; Britain).
— Hinton, 1981: 659 (Erichsonius; characters of egg).
— Frank, 1981b: 103 (Erichsonius; subgenus Erichsonius).
— Frank, 1982: 35 (Erichsonius; list of parasites).
— Tóth, 1984: 39 (Erichsonius; characters; Hungary).
— Ádám, 1987: 149 (Erichsonius; collecting notes; Hungary).
— Lucht, 1987: 104 (Erichsonius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Lohse and Lucht, 1989: 162 (Erichsonius; subgenus Erichsonius; characters).
— Uhlig and Sterrenburg, 1990: 168, 171 (Erichsonius; characters; Netherlands).
— Pilon and Zanetti, 1991: 55 (Erichsonius; Italy).
— Smetana, 1993b: 48 (Erichsonius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 27 (Erichsonius; Italy).
— Bordoni, 1995: 88 (Erichsonius; ecological notes; Italy).
— Owen, 1997a: 149 (Erichsonius; collecting sites; Ireland).
▴ Distribution: Morocco, Europe, Russia, Armenia.
melanocephalus Hochhuth, 1851: 25 (Philonthus; [preoccupied]; Type locality: Armenien).
— Fauvel, 1875a: XXIX [= 1875b: 231] (Actobius).
— Bernhauer and Schubert, 1914: 325 (Actobius; synonym of nigriceps).
— Bernhauer, 1944: 148 (Actobius; characters in key).
— Coiffait, 1965c: 844 (Erichsonius; synonym of cinerascens).
— Coiffait, 1974: 12 (Erichsonius; synonym of cinerascens).
nigriceps Gemminger and Harold, 1868: 590 (Philonthus; replacement name for melanocephalus Hochhuth).
— Fauvel, 1875a: XXIX [= 1875b: 231] (Actobius).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
robustus W. Kolbe, 1918: 205 (Actobius; variety of cinerascens; [Note: Gabriel cited as author]; [preoccupied]; Type locality: Not cited).
— Scheerpeltz, 1933: 1328 (Actobius; synonym of kolbei).
— Smetana, 1958: 141 (Erichsonius; synonym of cinerascens).
— Strassen, 1964: 102 (Actobius; synonym of subopacus).
— Coiffait, 1974: 12 (Erichsonius; synonym of cinerascens).
kolbei Scheerpeltz, 1933: 1328 (Actobius; replacement name for robustus W. Kolbe).
— Smetana, 1958: 141 (Erichsonius; synonym of cinerascens).
— Strassen, 1964: 102 (Actobius; synonym of subopacus).
— Coiffait, 1974: 12 (Erichsonius; variety of cinerascens).
obsoletus Méquignon, 1948: 76 (Actobius; variety of cinerascens; Type locality: Seine-et-Oise: Saint-Germain-en-Laye).
— Coiffait, 1965c: 844 (Erichsonius; subgenus Erichsonius; variety of cinerascens; characters).
— Coiffait, 1974: 12 (Erichsonius; variety of cinerascens; characters).
civicus Frank, 1975: 201 (Erichsonius; Type locality: Massachusetts: Norfolk Co.: Wellesley. Also cited from Ontario, New Hampshire, Massachusetts, Rhode Island, New York, Michigan, Illinois, Wisconsin, Maryland, North Carolina, Georgia, and Louisiana).
— Frank, 1981b: 100 (Erichsonius; subgenus Erichsonius; Quebec; Florida; Wisconsin).
— Downie and Arnett, 1996: 374 (Erichsonius; characters; USA).
▴ Distribution: Canada, USA.
coloratus Tottenham, 1956: 262 (Erichsonius; [Note: Although Tottenham attributed this name to Cameron as a name to be published, Tottenham validated it in a key]; Type locality: Angola).
coloratus Cameron, 1959: 115 (Actobius [Note: preoccupied; name validated first by Tottenham, 1956]; Type locality: Angola: Dundo).
coloratus Cameron, 1959, see: coloratus Tottenham, 1956.
comorensis Uhlig, 1991: 311 (Erichsonius; subgenus Sectophilonthus?; Type locality: Grande Comore, Lac Hantsongoma).
— Lecoq, 1996: 147 (Erichsonius; indentification of these specimens uncertain; aedeagus).
▴ Distribution: Comoros.
concolor Sharp, 1885, see: Neobisnius.
confusus Levasseur, 1971: 294 (Erichsonius; Type locality: monts Loma, Sierra Leone, 1600 m).
commixtus Tottenham, 1956: 274 (Erichsonius; Type locality: South Africa: Mashonaland).
creberrimus Tottenham, 1956: 268 (Erichsonius; Type locality: Ruanda: terr. Nyanza, Kinazi, 1600 m).
crebriceps Bernhauer, 1933d: 296 (Actobius; Type locality: Leopoldville).
crescenti Frank, 1975: 182 (Erichsonius; Type locality: U.S.A.: Massachusetts, Middlesex Co., Framingham. Also cited from Illinois, North Carolina, Florida, and Louisiana).
— Frank, 1981b: 103 (Erichsonius; subgenus Sectophilonthus).
— Downie and Arnett, 1996: 373 (Erichsonius; characters; USA).
▴ Distribution: USA.
curtipennis Bernhauer, 1932b: 146 (Actobius; Type locality: Katanga: Katompe).
— Scheerpeltz, 1933: 1329 (Actobius; catalog).
— Tottenham, 1956: 265 (Erichsonius; comparative notes).
▴ Distribution: Congo.
curtus Tottenham, 1962: 145 (Erichsonius; Type locality: Tanganyika: Kilimanjaro, Bismarck Hügel, 3100 m).
deceptivus Tottenham, 1956: 267 (Erichsonius; Type locality: Ruanda: terr. Nyanza, Kinazi, 1600 m. Also cited from Urundi).
delicatulus Sharp, 1885, see: Neobisnius.
descarpentriesi Jarrige, 1978: 281 (Erichsonius; subgenus Parerichsonius; Type locality: Andringitra Centre, éboulis Sud du plateau Andohariana, 2150 m).
dundoensis Tottenham, 1956: 225, 263 (Erichsonius; [Note: Although Tottenham attributed this name to Cameron as a name to be published, Tottenham validated it in a key]; Type locality: Urundi: terr. Bururi, Nyamurenbe, 900 m; Angola).
dundoensis Cameron, 1959: 115 (Actobius; [Note: Preoccupied; name first validated by Tottenham, 1956]; Type locality: Angola: Dundo: Dundundo River; Chiumbe River).
dundoensis Cameron, 1959, see: dundoensis Tottenham, 1956.
dux Smetana, 1967i: 552 (Erichsonius; Type locality: Türkei: Bürücek, Toros).
— Coiffait, 1974: 14 (Erichsonius; characters; notes).
— Petrenko, 1980: 82 (Erichsonius; Azerbaijan; Armenia).
▴ Distribution: Turkey, Azerbaijan, Armenia.
elegantulus Horn, 1884, see: Neobisnius.
elgonensis Bernhauer, 1939a: 90 (Actobius; Type locality: Kenya: Mt. Elgon, Osthang, Suam fishing hut, 2400 m).
— Bernhauer and Chapman, 1939: 64 (Actobius; Kenya).
— Tottenham, 1956: 266 (Erichsonius; comparative notes).
▴ Distribution: Kenya.
facilis Sharp, 1885, see: Neobisnius.
fallax Tottenham, 1953b: 495 (Actobius; Type locality: Ruwenzori: above Lake Bujuku, 12,750 ft).
felix Tottenham, 1962: 151 (Erichsonius; Type locality: Tanganyika Terr.: Uluguru Mts., vallée de l'Ululu-Ndongo, 1500 m).
flavicornis Fauvel, 1895, see: Neobisnius.
floridanus Frank, 1975: 198 (Erichsonius; Type locality: Florida: Orange Co.: Winter Park).
formosus Fall, 1901, see: Neobisnius.
fortis Sharp, 1885, see: Neobisnius.
franzi Lecoq, 1990: 191 (Erichsonius; Type locality: La Réunion: Maïdo près du Grand Bénard).
fraterculus Horn, 1884: 226 (Actobius; Type locality: northern Illinois; Michigan).
— Blatchley, 1910: 392 (Actobius; characters; Indiana).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
— Frank, 1975: 192 (Erichsonius; characters; Quebec; Ontario; Maine; New Hampshire; Massachusetts; New York; Michigan; Illinois).
— Legner and Moore, 1977: 176 (Erichsonius; found under bark).
— Frank, 1981b: 103 (Erichsonius; subgenus Erichsonius).
— Downie and Arnett, 1996: 373 (Erichsonius; characters; USA).
▴ Distribution: Canada, USA.
fulvicornis Notman, 1920, see: Neobisnius.
gabonicus Levasseur, 1980: 352 (Erichsonius; Type locality: Gabon: Belinga).
gabrieli Horion, 1935, see: subopacus Hochhuth, 1851.
gentyi Levasseur, 1969: 894 (Erichsonius; Type locality: Côte d'Ivoire, la Mé près Bingerville).
girardi Levasseur, 1969: 897 (Erichsonius; Type locality: Côte d'Ivoire, Lamto).
glabratus Kirshenblat, 1951: 543 (Actobius; Type locality: Primorskii krai: st. Grodekovo).
gracilicornis Scheerpeltz, 1974: 140 (Erichsonius; Type locality: Natal: Tugela river, 12 miles WNW Bergville).
griveaudi Jarrige, 1978: 284 (Erichsonius; subgenus Parerichsonius; Type locality: Andringitra Est, Anjavidilava, 1850–1950 m).
hannemanni Uhlig, 1991a: 97 (Erichsonius; subgenus Sectophilonthus; Type locality: Madagaskar, Namoroka).
helgae Masch and Uhlig, 1990: 268 (Erichsonius; subgenus Sectophilonthus; Type locality: Zaire: Provinz Kivu: Territorium Uvira, 800 m).
— Masch, 1993: 187, 208 (Erichsonius; subgenus Sectophilonthus; phylogeny of robustus species group).
▴ Distribution: Congo.
hergeri Uhlig, 1990: 137 (Erichsonius; subgenus Sectophilonthus; Type locality: Madagascar, Massif de l'Itremo).
hiekei Uhlig, 1995: 287 (Erichsonius; subgenus Sectophilonthus; Type locality: Zimbabwe: 18°17′S, 32°43′E, Nyanga N. P., Rhodes Dam).
hokkaidoensis Uhlig and Watanabe, 1992: 24 (Erichsonius; subgenus Sectophilonthus; Type locality: Japan: Hokkaido: Hokkaido Prefecture: Iburi, [Muroran city]).
horni Bernhauer, 1922c: 231 (Actobius; Type locality: Kankau).
— Scheerpeltz, 1933: 1329 (Actobius; catalog).
— Bernhauer, 1944: 149 (Actobius; characters in key).
▴ Distribution: Taiwan.
humeralis Cameron, 1920, see: basalis Motschulsky, 1858.
hummleri Bernhauer, 1944: 149 (Actobius; Type locality: Herzegowina, Tovarnica, Bojanatal).
— Coiffait, 1965: 843 (Erichsonius; translation of original description; Hercegovina).
— Coiffait, 1974: 13 (Erichsonius; subgenus Erichsonius; characters; Herzegowina).
— Frank, 1981b: 103 (Erichsonius; subgenus Erichsonius).
▴ Distribution: Bosnia Hercegovina.
ibericus Coiffait, 1965, see: rivularis Kiesenwetter, 1858.
impeditus Levasseur, 1971: 295 (Erichsonius; Type locality: monts Loma, Sierra Leone, 1600 m).
infimus Horn, 1884, see: Neobisnius.
inornatus Sharp, 1889, see: Neobisnius.
inutilis Horn, 1884: 225 (Actobius; Type locality: Michigan; New England States).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
— Frank, 1975: 194 (Erichsonius; characters; Quebec; Ontario; Maine; New Hampshire; Massachusetts; New York; Michigan; Illinois).
— Frank, 1981b: 103 (Erichsonius; subgenus Erichsonius).
— Downie and Arnett, 1996: 373 (Erichsonius; characters; USA).
▴ Distribution: Canada, USA.
itoi Uhlig and Watanabe, 1992: 39, 92 (Erichsonius; subgenus Sectophilonthus; Type locality: Japan: Honshu: Nara Prefecture: Mt. Ohdaigahara).
japonicus Cameron, 1933a: 170 (Actobius; Type locality: Japan: Kobe).
— Bernhauer, 1944: 149 (Actobius; characters in key).
— Watanabe, 1963: 7, 9 (Erichsonius; characters; Japan).
— Tikhomirova, 1973: 182 (Erichsonius; China; Japan).
— Uhlig and Watanabe, 1992: 17 (Erichsonius; subgenus Sectophilonthus; lectotype designation; redescription; natural history; phylogenetic relationships; Japan).
— Cho, 1996a: 163 (Erichsonius; characters; Korea).
▴ Distribution: Japan, Korea, China.
jarrigei Levasseur, 1969: 890 (Erichsonius; Type locality: région de Kolwezi, Manika, Katanga).
jocosus Horn, 1884, see: Neobisnius.
jocosus Tottenham, 1962: 146 (Erichsonius; [Note: Preoccupied as a secondary homonym]; Type locality: Tanganyika Terr.: Kilimanjaro, Marangu, 2400 m).
jucundus Horn, 1884, see: Neobisnius.
kamerunensis Bernhauer, 1915k: 300 (Actobius; Type locality: Kamerun: Musake).
— Scheerpeltz, 1933: 1329 (Actobius; catalog).
— Bernhauer, 1942c: 361 (Actobius; Cameroon).
— Tottenham, 1956: 265 (Erichsonius; comparative notes).
▴ Distribution: Cameroon.
katanganus Levasseur, 1969: 888 (Erichsonius; Type locality: Kolwezi, Katanga).
kiboshensis Bernhauer, 1915d: 135 (Actobius; Type locality: Deutsch-Ostafrika: Kibosho).
— Scheerpeltz, 1933: 1329 (Actobius; catalog).
— Tottenham, 1956: 265 (Erichsonius; comparative notes).
▴ Distribution: Tanzania.
kobensis Cameron, 1933a: 169 (Actobius; Type locality: Mayasan, Hyogo Prefecture, Honshu, Japan; [Note: See lectotype designation by Uhlig and Watanabe, 1992]. Originally cited from Japan: Kobe, Mayasan. Kyushu, Mt. Unzen).
— Bernhauer, 1944: 148 (Actobius; [Note: Misinterpretation of species according to Uhlig and Watanabe, 1992]; characters in key;).
— Watanabe, 1963: 7, 8 (Erichsonius; [Note: Misinterpretation of species according to Uhlig and Watanabe, 1992]; characters; Japan).
— Nakane, 1963a: 90 (Erichsonius; characters; habitus photograph; Japan).
— Uhlig and Watanabe, 1992: 49 (Erichsonius; subgenus Sectophilonthus; lectotype designation: Mayasan, Hyogo Prefecture, Honshu, Japan; characters; natural history; phylogenetic relationships; Japan).
▴ Distribution: Japan.
kolbei Scheerpeltz, 1933, see: cinerascens Gravenhorst, 1802.
kolweziensis Levasseur, 1969: 892 (Erichsonius; Type locality: région de Kolwezi, Manika, Katanga).
lamtoensis Levasseur, 1969: 896 (Erichsonius; Type locality: Côte d'Ivoire, Lamto).
laticeps Cameron, 1918: 86 (Actobius; Type locality: Singapore: Mandai).
— Cameron, 1921a: 376, 404 (Actobius; characters; Singapore).
— Scheerpeltz, 1933: 1329 (Actobius; catalog).
▴ Distribution: Singapore.
lecordieri Levasseur, 1969: 887 (Erichsonius; Type locality: région de Kolwezi, Musonoie).
leleupi Tottenham, 1962: 152 (Erichsonius; Type locality: Tanganyika Terr.: Kilimanjaro, Marangu, 1550 m).
leonensis Tottenham, 1956: 272 (Erichsonius; Type locality: Sierra Leone: Freetown).
leptocerus Fauvel, 1905b: 177 (Actobius; Type locality: Madagascar sud: forêts de la côte est; Fort Dauphin; Andrahomana).
lohsei Uhlig, 1990a: 138 (Erichsonius; subgenus Sectophilonthus; Type locality: Zaire: Kasai Oriental, Gandajika).
lomaensis Levasseur, 1971: 292 (Erichsonius; Type locality: monts Loma, Sierra Leone, 1600 m).
longipennis Jarrige, 1970: 47 (Erichsonius; subgenus Parerichsonius; Type locality: Madagascar: Mt. Tsaratanana, 1500 m).
loxatus Horn, 1884, see: patella Horn, 1884.
maghrebicus Coiffait, 1965: 846, 849 (Erichsonius; subgenus Parerichsonius; Type locality: Algérie: Mouzaïa, près de Médéa).
— Coiffait, 1973a: 274 (Erichsonius; Morocco).
— Coiffait, 1974: 16 (Erichsonius; subgenus Parerichsonius; characters; Algeria; Morocco).
— Outerelo, 1976: 265 (Erichsonius; Spain).
— Frank, 1981b: 103 (Erichsonius; subgenus Sectophilonthus).
▴ Distribution: Algeria, Morocco, Spain.
major Cameron, 1943: 128 (Actobius; Type locality: Darjeeling: Ghum district: Mangpu).
malheurensis Hatch, 1957, see: puncticeps Horn, 1884.
mangpuensis Cameron, 1943: 128 (Actobius; Type locality: Darjeeling: Ghum district: Mangpu).
manikaensis Levasseur, 1969: 889 (Erichsonius; Type locality: région de Kolwezi, Manika, Katanga).
mauriticus Uhlig, 1988: 83 (Erichsonius; subgenus Sectophilonthus; Type locality: Mauritius: Piton Grand Bassin).
mauritiensis Uhlig, 1988: 79 (Erichsonius; subgenus Sectophilonthus; Type locality: Mauritius: Piton Grand Bassin).
maximus Tottenham, 1956: 270 (Erichsonius; Type locality: Ruanda: Tshuruyaga, forêt du Rugege, 2400 m).
— Masch and Uhlig, 1990: 235 (Erichsonius; subgenus Sectophilonthus; characters; Zaire; Rwanda).
— Masch, 1993: 187, 215 (Erichsonius; subgenus Sectophilonthus; phylogeny of robustus species group).
▴ Distribution: Rwanda, Congo.
melanocephalus Hochhuth, 1851, see: cinerascens Gravenhorst, 1802.
meurgeusae Coiffait, 1979d: 555 (Erichsonius; Type locality: Nouristan: Darah i Nour).
mixtus Sharp, 1885, see: Neobisnius.
modestus Tottenham, 1956: 264 (Erichsonius; [Note: Although Tottenham attributed this name to Cameron as a name to be published, Tottenham validated it in a key]; Type locality: Angola).
moestus Sharp, 1887, see: Neobisnius.
montanellus Levasseur, 1971: 293 (Erichsonius; Type locality: monts Loma, Sierra Leone, 1600 m).
monticola Cameron, 1943: 128 (Actobius; Type locality: Darjeeling: Ghum district: Mangpu).
muscicola Jarrige, 1970: 48 (Erichsonius; subgenus Parerichsonius; Type locality: Madagascar: Mt. Tsaratanana, 1500 m).
musidentatus Masch and Uhlig, 1990: 253 (Erichsonius; subgenus Sectophilonthus; Type locality: Malawi: Viphya-Mts., Chamambo Forest).
— Masch, 1993: 187, 212 (Erichsonius; subgenus Sectophilonthus; phylogeny of robustus species group).
▴ Distribution: Malawi.
musonoiensis Levasseur, 1969: 885 (Erichsonius; Type locality: région de Kolwezi, Musonoie, Katanga).
myops Jarrige, 1978: 283 (Erichsonius; subgenus Parerichsonius; Type locality: Andringitra Centre, zone sommitale, cirque Boby, 2500 m).
nanus Horn, 1884: 225 (Actobius; Type locality: Canada; Michigan; New England States).
— Blatchley, 1910: 392 (Actobius; characters; Indiana).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
— Hatch, 1957: 174 (Actobius; characters; Idaho).
— Smetana, 1965: 11 (Erichsonius; characters; St. Pierre; Nova Scotia; Newfoundland).
— Smetana, 1965a: 3 (Erichsonius; Alaska).
— Frank, 1975: 198 (Erichsonius; characters; St. Pierre & Miquelon; Newfoundland; Nova Scotia; New Brunswick; Quebec; Ontario; Northwest Territories; British Columbia; Alaska; New Hampshire; Maine; Massachusetts; New York; New Jersey; Illinois; Wisconsin; Washington).
— Frank, 1981b: 100, 103 (Erichsonius; subgenus Erichsonius; Virginia).
— Frank, 1982: 36 (Erichsonius; list of parasites).
— Downie and Arnett, 1996: 374 (Erichsonius; characters; USA).
▴ Distribution: Canada, USA.
naomii Uhlig and Watanabe, 1992: 66 (Erichsonius; subgenus Sectophilonthus; Type locality: Japan: Okinawa Prefecture: Nansei-Shotô: Ishigaki Island: Mt. Omoto).
nepalicus Coiffait, 1981d: 328 (Acylophorus; Type locality: Kluihakani, Halambu, Népal central).
— Smetana, 1988: 339 (Erichsonius; most likely identical with basalis Motschulsky).
▴ Distribution: Nepal.
nigriceps Gemminger and Harold, 1868, see: cinerascens Gravenhorst, 1802.
niloticus Scheerpeltz, 1963b: 439 (Erichsonius; Type locality: Wadi-Halfa).
nitidicollis Jarrige, 1970: 45 (Erichsonius; subgenus Parerichsonius; Type locality: Madagascar: Mt. Tsaratanana, 15–1600 m).
nitidulus Sharp, 1885, see: Neobisnius.
nouristanicus Coiffait, 1979d: 554 (Erichsonius; Type locality: Nouristan: vallée du Pech).
nukurensis Fauvel, 1907: 37 (Actobius; Type locality: Nakuro, Rift Valley).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
— Bernhauer and Chapman, 1939: 64 (Actobius; Kenya).
— Tottenham, 1956: 264 (Erichsonius; characters in key).
— Scheerpeltz, 1974: 138, 141 (Erichsonius; characters; South Africa).
▴ Distribution: Kenya, South Africa.
obsoletus Méquignon, 1948, see: cinerascens Gravenhorst, 1802.
ocreatus Horn, 1884, see: Neobisnius.
oculatus Fauvel, 1905, see: Neobisnius.
omissus Levasseur, 1971: 297 (Erichsonius; Type locality: monts Loma, Sierra Leone, 1600 m).
ophthalmicus Tottenham, 1956: 262, 269 (Erichsonius; Type locality: Ruanda: terr. Kisenyi, Gishwati, 2000 m).
orphanulus Erichson, 1840: 931 (Philonthus; replacement name for orphanus Erichson, 1840: 499).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Coiffait, 1972: 50 (Erichsonius; synonymic note).
— Coiffait, 1974: 5, 7 (Erichsonius; synonymic note).
— Frank, 1981b: 103 (Erichsonius; subgenus Sectophilonthus).
▴ Distribution: Madagascar.
orphanus Erichson, 1840: 499 (Philonthus; [preoccupied]; Type locality: Madagascar).
biparamerosus Tottenham, 1949b: 358 (Philonthus; subgenus Sectophilonthus; Type locality: Madagascar: Tananarive).
— Coiffait, 1972: 50 (Erichsonius; synonym of orphanulus).
— Coiffait, 1974: 5, 7 (Erichsonius; synonymic note).
— Frank, 1981b: 103 (Erichsonius; synonym of orphanulus).
orphanus Erichson, 1840, see: orphanulus Erichson, 1840.
papuanus Cameron, 1937a: 100 (Actobius; Type locality: New Guinea: Kokoda, alt. 1200 ft.).
parcior Tottenham, 1956: 274 (Erichsonius; Type locality: Natal: Escourt).
parcus Horn, 1884: 230 (Actobius; Type locality: Michigan; District of Columbia).
— Blatchley, 1910: 393 (Actobius; characters; Indiana).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
— Notman, 1920b: 705 (Actobius; Florida).
— Frank, 1975: 187 (Erichsonius; characters; Massachusetts; District of Columbia; Virginia; South Carolina; Louisiana).
— Frank, 1981b: 100l, 103 (Erichsonius; subgenus Sectophilonthus; Florida).
— Downie and Arnett, 1996: 373 (Erichsonius; characters; USA).
▴ Distribution: USA.
parvus Tottenham, 1961: 207 (Erichsonius; Type locality: Senegal: Parc National du Niokolo-Koba, Badi).
patella Horn, 1884: 229 (Actobius; Type locality: Michigan; New York).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
— Frank, 1975: 185 (Erichsonius; characters; Quebec; Ontario; New Hampshire; New York; Massachusetts; Pennsylvania; Indiana; Illinois; Michigan; New Jersey; District of Columbia; Kentucky; Tennessee; West Virginia; Virginia; North Carolina; Louisiana).
— Frank, 1981b: 103 (Erichsonius; subgenus Sectophilonthus).
— Downie and Arnett, 1996: 373 (Erichsonius; characters; USA).
▴ Distribution: Canada, USA.
loxatus Horn, 1884: 229 (Actobius; Type locality: District of Columbia).
— Blatchley, 1910: 393 (Actobius; characters; Indiana).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
— Frank, 1975: 185 (Erichsonius; synonym of patella).
pauliani Jarrige, 1970: 48 (Erichsonius; subgenus Parerichsonius; Type locality: Madagascar: Mt. Tsaratanana, 1800 m).
picinus Jarrige, 1978: 279 (Erichsonius; subgenus Parerichsonius; Type locality: Andringitra Est, Marositry, Est rivière Antsifotra, 2000 m).
politus Sharp, 1885, see: Neobisnius.
pomonae Frank, 1975: 188 (Erichsonius; Type locality: Mt. Wilson, Los Angeles Co., California).
potor Tottenham, 1956: 262, 275 (Erichsonius; Type locality: Tanganyika: Kilimandjaro, 13,000 ft.).
proximalis Tottenham, 1956: 262, 276 (Erichsonius; Type locality: Tanganyika: Kilimandjaro, 13,000 feet).
pseudomaximus Masch and Uhlig, 1990: 265 (Erichsonius; subgenus Sectophilonthus; Type locality: Zaire: Provinz Kivu: Territorium Lubero, rés. forest. Biena, 1800 m).
— Masch, 1993: 187, 215 (Erichsonius; subgenus Sectophilonthus; phylogeny of robustus species group).
▴ Distribution: Congo.
pubidentatus Masch and Uhlig, 1990: 244 (Erichsonius; subgenus Sectophilonthus; Type locality: Zaire: Territorium Albertville, Mont Kabobo, Haute Kiymbi, 1700 m).
— Masch, 1993: 187, 213 (Erichsonius; subgenus Sectophilonthus; phylogeny of robustus species group).
▴ Distribution: Congo.
pullus Zheng, 1993b: 491, 492 (Erichsonius; Type locality: China: Sichuan, Xide).
puncticeps Horn, 1884: 230 (Actobius; Type locality: southern part of California).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
— Frank, 1975: 191 (Erichsonius; characters; Arizona, California).
— Frank, 1981b: 103 (Erichsonius; subgenus Sectophilonthus).
▴ Distribution: USA.
malheurensis Hatch, 1957: 182 (Philonthus; Type locality: Oregon: Malheur Co.: Sucker Creek Canyon).
puncticulosus Zheng, 1993b: 490, 492 (Erichsonius; Type locality: China: Sichuan, Mt. Emei, alt. 710–1120 m).
pusio Horn, 1884: 230 (Actobius; Type locality: Garland, Colorado).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
— Frank, 1975: 190 (Erichsonius; characters; Quebec, Ontario, British Columbia, Massachusetts, Michigan, Illinois, Colorado, Utah, North Carolina).
— Frank, 1981b: 100, 103 (Erichsonius; subgenus Sectophilonthus; Virginia).
— Schmidt, 1996: 211, 213 (Erichsonius; characters of egg, larvae, and pupa; natural history; Nebraska).
— Downie and Arnett, 1996: 373 (Erichsonius; characters; USA).
— Schmidt, 1999: 108 (Erichsonius; food for adults and larvae used for rearing species; rearing methods).
▴ Distribution: Canada, USA.
rivularis Kiesenwetter, 1858: 61 (Philonthus; Type locality: Hochgebirge um Vrachori in Aetolien).
— Mulsant and Rey, 1876: 602 [= 1877: 458] (Remus; characters; Greece).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
— Bernhauer, 1944: 149 (Actobius; characters in key).
— Coiffait, 1965c: 844 (Erichsonius; subgenus Erichsonius; characters; Greece; Asia Minor; Armenia).
— Coiffait, 1974: 14 (Erichsonius; subgenus Erichsonius; characters; Greece; Asia Minor; Armenia; Persia).
— Frank, 1981b: 103 (Erichsonius; subgenus Sectophilonthus).
▴ Distribution: Spain, Greece, Turkey, Armenia, Iran.
ibericus Coiffait, 1965: 844, 847 (Erichsonius; subgenus Erichsonius; subspecies of rivularis; Type locality: Espagne: La Musara, Alcover, Prov. de Tarragone).
robustus Bernhauer, 1915d: 135 (Actobius; Type locality: Deutsch-Ostafrika: Utzurungwe).
— Scheerpeltz, 1933: 1329 (Actobius; catalog).
— Tottenham, 1956: 266 (Erichsonius; comparative notes).
— Masch and Uhlig, 1990: 240 (Erichsonius; subgenus Sectophilonthus; characters; Tanzania).
— Masch, 1993: 187, 215 (Erichsonius; subgenus Sectophilonthus; phylogeny of robustus species group).
▴ Distribution: Tanzania.
robustus W. Kolbe, 1918, see: cinerascens Gravenhorst, 1802.
rosellus Frank, 1975: 195 (Erichsonius; Type locality: New Jersey: Union Co.: Roselle Park. Also cited from Quebec, Ontario, Maryland, Massachusetts, New York, New Jersey).
— Frank, 1981b: 100, 103 (Erichsonius; subgenus Erichsonius; Florida).
— Downie and Arnett, 1996: 373 (Erichsonius; characters; USA).
▴ Distribution: Canada, USA.
rotundodentatus Masch and Uhlig, 1990: 247 (Erichsonius; subgenus Sectophilonthus; Type locality: Tansania: Mt. Mbize, 12 miles N.E. Sumbawanga).
— Masch, 1993: 187, 215 (Erichsonius; subgenus Sectophilonthus; phylogeny of robustus species group).
▴ Distribution: Tanzania.
rufipennis Bernhauer, 1932b: 146 (Actobius; Type locality: Elisabethville; Kansenia).
— Scheerpeltz, 1933: 1329 (Actobius; catalog).
— Bernhauer and Chapman, 1939: 65 (Actobius; Kenya).
— Tottenham, 1956: 263 (Erichsonius; characters in key; Belgian Congo; Angola).
▴ Distribution: Kenya, Congo, Angola.
rusticus Frank, 1975: 194 (Erichsonius; Type locality: Illinois: Marion Co.: Kinmundy. Also cited from Illinois, Maryland and South Carolina).
— Frank, 1981b: 103 (Erichsonius; subgenus Erichsonius).
— Downie and Arnett, 1996: 373 (Erichsonius; characters; USA).
▴ Distribution: USA.
ruwenzoriensis Bernhauer, 1934g: 236 (Actobius; Type locality: Ruwenzori, 4200 m).
— Bernhauer and Chapman, 1939: 65 (Actobius; Kenya).
— Tottenham, 1956: 265 (Erichsonius; comparative notes).
▴ Distribution: “Ruwenzori region”, Kenya.
sawadai Uhlig and Watanabe, 1992: 45 (Erichsonius; subgenus Sectophilonthus; Type locality: Japan: Hokkaido: Hokkaido Prefecture: Oonuma Park).
scabrosus Tottenham, 1956: 271 (Erichsonius; Type locality: Ruanda: Tshuruyaga, forêt du Rugege, 2400 m).
— Masch and Uhlig, 1990: 262 (Erichsonius; subgenus Sectophilonthus; characters; Rwanda).
— Masch, 1993: 187, 209 (Erichsonius; subgenus Sectophilonthus; phylogeny of robustus species group).
▴ Distribution: Rwanda.
scotti Bernhauer, 1931d: 583 (Actobius; Type locality: Abyssinia: Mt. Chillalo, ca. 9,000 ft.; Jem-Jem Forest, 8,000–9,000 ft.).
— Scheerpeltz, 1933: 1329 (Actobius; catalog).
— Tottenham, 1956: 261 (Erichsonius; characters in key).
▴ Distribution: Ethiopia.
rossicus Coiffait, 1965, see: Philonthus.
senilis Horn, 1884, see: Neobisnius.
shibatai Watanabe and Uhlig, 1992: 28 (Erichsonius; subgenus Sectophilonthus; Type locality: Japan: Honshu: Kanagawa Prefecture: Ohkura near Tanzawa).
signaticornis Mulsant and Rey, 1853: 62 [= 1853a: 75] (Philonthus; Type locality: Lyonnais, Bugey).
— Fairmaire and Laboulbène, 1856: 531 (Philonthus; characters; France).
— Kraatz, 1855: 24 (Xantholinus; notes).
— Kraatz, 1857d: 623 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 198 (Philonthus; characters; Germany).
— Fauvel, 1874: 203 [= 1874a: 429] (Erichsonius; characters; France; Britain; Switzerland; Tirol; Italy).
— Redtenbacher, 1874: 217 (Philonthus; characters; Germany).
— Fauvel, 1876a: 72 [= 1876: 257] (Actobius; Spain; Algeria).
— Mulsant and Rey, 1876: 600 [= 1877: 456] (Remus; characters; France).
— Fauvel, 1878: 121 [= 1878a: 41] (Actobius; Algeria).
— Fauvel, 1886: 52 [= 1886a: 44] (Actobius; Morocco; Algeria).
— Fowler, 1888: 284 (Actobius; characters; Britain).
— Ganglbauer, 1895: 466 (Neobisnius; characters; western Europe; Tyrol; Italy).
— Fauvel, 1902b: 107 (Actobius; Morocco; Algeria; Tunisia).
— Sainte-Claire Deville, 1906: 94 (Actobius; Corsica).
— Reitter, 1909: 123 (Actobius; characters; Germany).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
— Portevin, 1929: 356 (Actobius; characters; France).
— Scheerpeltz, 1933: 1329 (Actobius; catalog).
— Bernhauer, 1944: 148 (Actobius; characters in key).
— V. Hansen, 1952: 33 (Actobius; characters).
— Smetana, 1958: 142 (Erichsonius; characters; ecology; Czechoslovakia).
— Smetana, 1960b: 259 (Erichsonius; Czechoslovakia).
— Coiffait, 1963: 9 (Erichsonius; subgenus Parerichsonius).
— Lohse, 1964: 172 (Erichsonius; characters; central Europe).
— Coiffait, 1965c: 845 (Erichsonius; subgenus Parerichsonius; characters; Europe).
— Horion, 1965: 119 (Erichsonius; Britain; Holland; Belgium; France; Switzerland; Czech Republic; Slovakia; Poland; Portugal; Spain; Corsica; Italy; Germany; Austria).
— Smetana, 1967i: 554 (Erichsonius; characters).
— Coiffait, 1974: 15 (Erichsonius; subgenus Parerichsonius; characters; central Europe; France; Corsica).
— Osella and Zanetti, 1975: 108 (Erichsonius; collected from nest of Talpa europaea; notes; Italy).
— Pope, 1977: 29 (Erichsonius; Britain).
— Bordoni and Carfi, 1978 (Erichsonius; note on peg setae of aedeagus).
— Frank, 1981b: 103 (Erichsonius; subgenus Sectophilonthus).
— Frank, 1982: 36 (Erichsonius; list of parasites).
— Tóth, 1984: 38 (Erichsonius; characters).
— Lucht, 1987: 104 (Erichsonius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).
— Lohse and Lucht, 1989: 161 (Erichsonius; subgenus Sectophilonthus; characters).
— Uhlig and Sterrenburg, 1990: 167, 169 (Erichsonius; characters; Netherlands).
— Smetana, 1993b: 48 (Erichsonius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 27 (Erichsonius; Italy).
— Eyre, Luff, and Lott, 2000: 33 (Erichsonius; Britain).
▴ Distribution: Tunisia, Algeria, Morocco, Europe.
silvaticus Bernhauer, 1931d: 584 (Actobius; Type locality: Abyssinia: Mt. Chillalo, ca. 9,000 ft.).
silvestris Bernhauer, 1931d: 584 (Actobius; Type locality: Abyssinia: Jem-Jem Forest, Jem-Jem river, ca. 8,000 ft.; Wouramboulchi, ca. 9,000 ft.; Mt. Chillalo, ca. 9,000 ft.).
simplex Sharp, 1885, see: Neobisnius.
sinusidentatus Masch and Uhlig, 1990: 250 (Erichsonius; subgenus Sectophilonthus; Type locality: Zaire: Territorium Albertville, Mont Kabobo, Haute Kiymbi, 1800 m).
— Masch, 1993: 187, 214 (Erichsonius; subgenus Sectophilonthus; phylogeny of robustus species group).
▴ Distribution: Congo.
smetanai Frank, 1975: 197 (Erichsonius; Type locality: Louisiana: St. John the Baptist Par., Manchac).
sobosanus Watanabe, 1963: 8 (Erichsonius; Type locality: Japan: Kyushu: Oita Pref.: Mt. Sobo).
— Tikhomirova, 1973: 182 (Erichsonius; in checklist).
— Uhlig and Watanabe, 1992: 32 (Erichsonius; subgenus Sectophilonthus; redescription; natural history; phylogenetic relationships; Japan).
▴ Distribution: Japan.
subniger Mulsant and Rey, 1876, see: subopacus Hochhuth, 1851.
subopacus Hochhuth, 1851: 27 (Philonthus; Type locality: Tiflis).
— Fauvel, 1874: 202 [= 1874a: 428] (Erichsonius; synonym of rivularis).
— Bernhauer and Schubert, 1914: 325 (Actobius; synonym of cinerascens).
— Bernhauer, 1944: 148 (Actobius; characters in key; Armenia; Asia Minor; Balkan Peninsula; France; Poland).
— Scheerpeltz, 1956c: 1100 (Actobius; Cyprus).
— Méquignon, 1948: 76 (Actobius; variety of cinerascens; characters; France).
— Smetana, 1958: 142 (Erichsonius; characters; ecology; Czechoslovakia).
— Smetana, 1960b: 259 (Erichsonius; Czechoslovakia).
— Lohse, 1964: 173 (Erichsonius; characters; central Europe).
— Focarile, 1964: 51, 66 (Actobius; characters; notes; Italy).
— Coiffait, 1965c: 844 (Erichsonius; subgenus Erichsonius; characters; Caucasus; Armenia; Asia Minor; Balkans; central Europe; France).
— Horion, 1965: 118 (Erichsonius; Turkey; Armenia; France; Balkans; Czech Republic; Slovakia; Poland ?; Austria).
— Smetana, 1967i: 552 (Erichsonius; Turkey).
— Coiffait, 1974: 13 (Erichsonius; subgenus Erichsonius; characters; Caucasus; Transcaucasus; Asia Minor; Balkans; central Europe; France).
— Petrenko, 1980: 82 (Erichsonius; Azerbaijan).
— Frank, 1981b: 103 (Erichsonius; subgenus Erichsonius).
— Tóth, 1984: 39 (Erichsonius; characters).
— Lucht, 1987: 104 (Erichsonius; checklist; Germany; Poland; Czechoslovakia; Austria).
— Lohse and Lucht, 1989: 163 (Erichsonius; subgenus Erichsonius; characters).
— Uhlig and Sterrenburg, 1990: 168 (Erichsonius; characters).
— Smetana, 1993b: 48 (Erichsonius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 27 (Erichsonius; Italy).
— Assing and Wunderle, 2001: 36 (Erichsonius; Cyprus).
▴ Distribution: Europe, Turkey, Georgia, Armenia, Azerbaijan.
subniger Mulsant and Rey, 1876: 605 [= 1877: 461] (Remus; variety of cinerascens; Type locality: Saint-Raphael).
— Ganglbauer, 1895: 465 (Actobius; variety of cinerascens).
— Bernhauer and Schubert, 1914: 325 (Actobius; synonym of cinerascens).
— Smetana, 1958: 142 (Erichsonius; synonym of subopacus).
— Coiffait, 1974: 13 (Erichsonius; synonym of subopacus).
gabrieli Horion, 1935: 144 (Actobius; variety of cinerascens; Type locality: Schlesien).
superbus Tottenham, 1962: 145 (Erichsonius; Type locality: Tanganyika Terr.: Uluguru Mts., vallée de l'Ululu-Ndogo, 1500 m).
— Masch and Uhlig, 1990: 260 (Erichsonius; subgenus Sectophilonthus; characters; Tanzania).
— Masch, 1993: 187, 209 (Erichsonius; subgenus Sectophilonthus; phylogeny of robustus species group).
▴ Distribution: Tanzania.
tenuicornis Fauvel, 1907: 37 (Actobius; Type locality: Voi).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
— Bernhauer and Chapman, 1939: 65 (Actobius; Kenya).
— Tottenham, 1956: 264 (Erichsonius; characters in key).
— Tottenham, 1962: 134 (Erichsonius; Tanganyika).
— Scheerpeltz, 1974: 138 (Erichsonius; South Africa; northern Rhodesia).
▴ Distribution: Kenya, Tanzania, Zambia, South Africa.
terebrans Tottenham, 1956: 263, 267 (Erichsonius; Type locality: Urundi: terr. Bururi, Nyamurembe).
texanus Frank, 1975: 188 (Erichsonius; Type locality: N. Braunfels, Texas).
triangularis Tottenham, 1961: 203, 208 (Erichsonius; Type locality: Senegal: Parc National du Niokolo-Koba: Badi).
tristis Cameron, 1932b: 133 (Actobius; Type locality: Blukwa, Nizi).
— Scheerpeltz, 1933: 1329 (Actobius; catalog).
— Tottenham, 1956: 264 (Erichsonius; characters in key).
▴ Distribution: Congo.
tsaratananus Jarrige, 1970: 45 (Erichsonius; Type locality: Madagascar: Mt. Tsaratanana, 1500 m).
tuberculatus Uhlig and Masch, 1990: 90 (Erichsonius; subgenus Sectophilonthus; Type locality: Zaire: Provinz Kuvu: Territorium Uvira, Haute Luvubu, 2675 m).
uhligi Lecoq, 1996: 147 (Erichsonius; Type locality: l'archipel des Comores: Grande-Comore, La Convalescence, 1750 m).
validus Cameron, 1928g: 444 (Actobius; Type locality: Borneo: Mt. Murud, 5000–6000 feet).
— Scheerpeltz, 1933: 1329 (Actobius; catalog).
— Hammond, 1984: 205 (Erichsonius; Borneo).
▴ Distribution: Borneo.
variolosus Tottenham, 1956: 274 (Erichsonius; Type locality: Tanganyika: Marangu).
venustus Tottenham, 1956: 262, 275 (Erichsonius; Type locality: Zanzibar).
vinsoni Cameron, 1947c: 116 (Philonthus; subgenus Gabrius; Type locality: Mauritius: Mt. Cocotte).
— Uhlig, 1988: 74 (Erichsonius; subgenus Sectophilonthus; characters; notes Mauritius).
— Lecoq, 1990: 191 (Erichsonius; Mascarene Islands).
▴ Distribution: Mauritius.
vulgaris Uhlig and Watanabe, 1992: 58 (Erichsonius; subgenus Sectophilonthus; Type locality: Japan: Honshu: Gunma Prefecture: Yunokoya).
watanabei Uhlig, 1992: 72 (Erichsonius; subgenus Sectophilonthus; Type locality: Japan: Honshu: Aomori Prefecture: Mt. Hakkôda).
ytenensis Sharp, 1913b: 101 (Actobius; Type locality: New Forest).
— Donisthorpe, 1930: 97 (Actobius; characters; Britain).
— Scheerpeltz, 1933: 1329 (Actobius; catalog).
— Coiffait, 1965c: 845 (Erichsonius; subgenus Parerichsonius; characters; England; France).
— Smetana, 1967i: 554 (Erichsonius; characters).
— Coiffait, 1974: 17 (Erichsonius; subgenus Parerichsonius; characters; England; France).
— Outerelo, 1976: 264 (Erichsonius; notes; Spain).
— Pope, 1977: 29 (Erichsonius; Britain).
— Frank, 1981b: 103 (Erichsonius; subgenus Sectophilonthus).
— Lohse and Lucht, 1989: 162 (Erichsonius; subgenus Sectophilonthus; characters).
— Uhlig and Sterrenburg, 1990: 167, 172 (Erichsonius; characters; Netherlands).
— Bruge, Haghebaert, and Drugmand, 1991: 47 (Erichsonius; notes; characters; Belgium).
▴ Distribution: Britain, Netherlands, Belgium, France, Spain.
zapfi Uhlig, 1997: 266 (Erichsonius; Type locality: Republic of South Africa: Cape Province: De Hoop Nature Reserve, De Hoop Vlei, 34°27.2′ [S], 20°24.2′E).
ziloensis Levasseur, 1969: 884 (Erichsonius; Type locality: région de Kolwezi, Zilo, Katanga).
Flohria
[2 species; Neotropical and Nearctic Regions]
Flohria Sharp, 1884: 391 (species included: laticornis). Type species: Flohria laticornis Sharp, fixed by monotypy.
— Bernhauer and Schubert, 1914: 366 (world catalog; 1 species).
— Blackwelder, 1944: 136 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 166 (type species: laticornis).
— Newton, Thayer, Ashe, and Chandler, 2000: 396 (1 Nearctic species; notes; characters in key).
laticornis Sharp, 1884: 392 (Flohria; Type locality: Mexico, Almolonga, Jalcomulco, Uruapan, Ciudad in Durango).
subcoeruleus LeConte, 1863a: 35 (Trigonophorus; Type locality: Texas).
Gabriopalpus
[1 species; Palaearctic Region]
Gabriopalpus Schillhammer, 2000a: 35 (species included: ecitolycoides). Type species: Gabriopalpus ecitoycoides Schillhammer, fixed by original designation and monotypy.
ecitolycoides Schillhammer, 2000a: 37 (Gabriopalpus; Type locality: Cachemire: Sonamarg).
Gabrius
[337 species (1 doubtful); Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions]
Gabrius Stephens, 1829a: 23 (species included: villosulus, semipunctatus, cinerascens, ventralis, attenuatus, nanus, pallipes, aterrimus, suaveolens; [Note: The same questions and controversy concerning the authorship and date of publication for Philonthus apply to Gabrius. The issue is discussed in my “note” attached to the citation of the original description for Philonthus]). Type species: Staphylinus aterrimus Gravenhorst, fixed by subsequent designation by Westwood, 1838a: 16.
— Curtis, 1829: 26 (catalog; Britain).
— Stephens, 1829: 283 (catalog; Britain).
— Stephens, 1832: 200 (characters).
— Stephens, 1833: 249 (characters; habitat).
— Westwood, 1838a: 16 (characters; type species).
— Stephens, 1839: 385 (characters).
— Erichson, 1840: 426 (synonym of Philonthus).
— Lacordaire, 1854: 80 (synonym of Philonthus).
— Kraatz, 1857d: 565 (synonym of Philonthus).
— G. Waterhouse, 1858: 24 (synonym of Philonthus).
— Thomson, 1860: 169 (characters).
— Motschulsky, 1860: 567 (characters).
— Gemminger and Harold, 1868: 584 (synonym of Philonthus).
— Fauvel, 1874: 207 [= 1874a: 433] (synonym of Philonthus).
— Mulsant and Rey, 1876: 535 [= 1877: 391] (characters; key to French species).
— Seidlitz, 1891: 389 (key to some European species).
— Seidlitz, 1891a: 414 (key to some European species).
— Heyden, Reitter, and Weise, 1891: 104 (subgenus of Philonthus; list of species of Europe and Caucasus).
— Heyden, 1893: 43 (subgenus of Philonthus; list of species of Siberian region).
— Heyden, 1896: 33 (subgenus of Philonthus; list of species of Siberian region).
— Heyden, Reitter, and Weise, 1906: 168 (subgenus of Philonthus; list of species of Europe and Caucasus).
— Reitter, 1909: 124 (subgenus of Philonthus; characters).
— Bernhauer and Schubert, 1914: 326 (subgenus of Philonthus).
— Gridelli, 1920 (revision; valid genus; key to species; illustrations).
— Winkler, 1925: 379 (catalog for Palaearctic region; subgenus of Philonthus).
— Cameron, 1925: 63 (catalog of Indian species).
— Gridelli, 1928: 41 (notes).
— Cameron, 1932: 62 (subgenus of Philonthus).
— Joy, 1932: 115 (key to British species).
— Scheerpeltz, 1933: 1329 (subgenus of Philonthus).
— Blackwelder, 1943: 399 (subgenus of Philonthus; type species: suaveolens).
— Fagel, 1945: 37, 38 (characters; notes; key to some species of nigritulus group).
— Jeannel and Paulian, 1945: 79 (type species: aterrimus).
— Kloet and Hincks, 1945: 166 (list of British species).
— Tottenham, 1949: 372, 419 (type species: nigritulus = aterrimus; notes; checklist of British species).
— Coiffait, 1951 (key to French species).
— Smetana, 1952b (revision of species of Czechoslovakia; characters; key to species).
— V. Hansen, 1952: 81 (subgenus of Philonthus; characters; key to Danish species).
— Blackwelder, 1952: 166 (type species).
— Smetana, 1953a: 167, 170, 175 (key to species of three species groups).
— Smetana, 1954a: 114 (key to central European species of splendidulus group).
— Hatch, 1957: 180, 198 (subgenus of Philonthus; characters; key to species of Pacific Northwest).
— Smetana, 1957 (key to European species; aedeagal illustrations).
— Smetana, 1958: 228 (characters; habitus of one species; type species: nigritulus; key to Czechoslovakian species).
— Smetana, 1959a: 214 (list of species collected in Albania).
— Smetana, 1960c (revision of and key to species of nigritulus group).
— Smetana, 1961: 36 (key to species of vernalis group).
— Tottenham, 1962: 235 (characters; notes; African species).
— Coiffait, 1963: 9 (characters; key to species groups and list of included French species for each).
— Székessy, 1963: 56 (characters).
— Palm, 1963b: 98 (subgenus of Philonthus; characters; key to species of Sweden).
— Lohse, 1964: 187 (characters; key to central European species).
— Coiffait, 1964c: 165 (hesperius species group; characters; key to species).
— Moore, 1965: 97, 100 (characters; notes).
— Pototskaia, 1966: 152, 154 (larval characters; key to some species).
— Pototskaia, 1967: 52 (larval characters).
— Coiffait and Saiz, 1968: 347, 348 (characters; notes; Chilean species).
— Scheerpeltz, 1968a: 76 (catalog of Austrian species).
— Bolov, 1969a: 328 (collecting notes for some species; Russia).
— Kasule, 1970: 64, 65 (larval characters).
— Shibata, 1973c: 61 (checklist of species of Taiwan).
— Bordoni, 1973j: 708 (list of species from islands near Sicily).
— Tikhomirova, 1973a: 185 (checklist of species of USSR).
— Coiffait, 1974: 17 (adult and larval characters; key to species of western Palaearctic region).
— Moore and Legner, 1975: 13 (catalog; North America north of Mexico).
— Pototskaia, 1976: 171 (larval characters).
— Pope, 1977: 30 (list of British species).
— Topp, 1978: 312 (larval characters).
— Moore and Legner, 1979: 19 (characters; notes).
— Muona, 1979: 15 (list of Scandinavian species).
— Uhlig, 1979: 254 (some species collected in Germany).
— M. Dvořák, 1979: 115 (some species collected in Slovakia).
— Uhlig, Vogel, and Sieber, 1980: 247 (some species collected in Germany).
— Burakowski, Mroczkowski, and Stefańska, 1980: 79 (catalog; Poland).
— Uhlig and Vogler, 1981: 117 (list of some species collected in Germany; habitat and collecting notes).
— Dettner, 1983 (comparative anatomy of defensive glands and defensive secretions).
— Shibata, 1983: 127 (checklist of species of Japan).
— Smetana, 1984 (review of and key to species of Japan).
— Hammond, 1984: 205 (checklist of species of Borneo).
— Tóth, 1984: 82 (characters; habitus; key to species of Hungary).
— Frank, 1986a: 370 (checklist of species of Florida).
— Muona and Viramo, 1986: 13 (list of species of northeastern Finland).
— Segers, 1986: 42 (checklist of species; Belgium).
— Drugmand 1987b (comments on distribution of Belgian species).
— Ádám, 1987: 151 (list of four species with collecting notes; Hungary).
— Lecoq, 1990: 194 (key to species of Mascarene Islands).
— Angelini, 1991: 191 (list of species from high elevations of Calabria, Italy).
— M. Hansen, Kristensen, Mahler, Pedersen, 1991: 107 (list of Danish species).
— Campbell and Davies, 1991: 119 (checklist of species; Alaska and Canada).
— Silfverberg, 1992: 16 (list of species of Fennoscandia, Denmark, and the Baltic States).
— Siitonen, 1993: 230 (list of Finnish species).
— Welch, 1993: 228 (ovariole structure and development).
— Smetana, 1993b: 49 (list of species of Czech Republic and Slovakia).
— Hernández, Outerelo, and Gamarra, 1994: 187 (checklist of species; Canary Islands).
— Smetana, 1995e: 603 (characters; notes; phylogenetic relationships; key to North American species).
— M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 29 (additional records to Danish list of species).
— Ciceroni and Zanetti, 1995: 29 (list of species of Italy).
— M. Hansen, 1996: 37, 106 (probably belongs in Philonthus; list of Danish species).
— M. Hansen, Mahler, Palm, and Pedersen, 1996: 244 (additions to list of Danish species).
— Downie and Arnett, 1996: 382 (characters; key to species of Northeastern North America).
— Gillerfors, 1997: 85 (key to species of northern Europe).
— Schillhammer, 1997 (revision of and species groups for species of Oriental region; checklist of species for world).
— Cho and Lee, 1997: 272 (key to species of Korea).
— Assing, Frisch, Kahlen, et al., 1998: 138 (supplement to key to species in Lohse, 1964; note).
— Newton, Thayer, Ashe, and Chandler, 2000: 396 (42 Nearctic species; notes; characters in key).
— Hayashi, 2000: 23 (notes on empodial setae).
abas Smetana, 1984: 135 (Gabrius; Type locality: Japan: Gumma Pref.: 5 km E Usui Pass, 900 m).
aberdarensis Bernhauer, 1939a: 90 (Philonthus; subgenus Gabrius; Type locality: Kenya: Mt. Aberdare, Westhang, Mais. forest. Kinangop, 2600 m).
abscissus Tottenham, 1962: 248 (Gabrius; Type locality: Kenya: Molo, Mau Escarpment, 2150–2200 m).
acutiphallus Schillhammer, 1997, see: sagittifer Schillhammer, 1997.
adustus Casey, 1915: 440 (Philonthus; Type locality: California: Sta. Cruz Mts.; [Note: See lectotype designation by Smetana, 1995e]. Also cited from San Francisco, California).
— Scheerpeltz, 1933: 1330 (Philonthus; catalog).
— Smetana, 1995e: 720 (Gabrius; lectotype designation: Santa Cruz Mts, California; characters; notes; California; Oregon).
▴ Distribution: USA.
capizzii Hatch, 1957: 201 (Philonthus; subgenus Gabrius; Type locality: Oregon: Gold Beach, Doug.).
aegon Smetana, 1995e: 671 (Gabrius; Type locality: California: S. Francisco Co.).
aelius Smetana, 1995e: 719 (Gabrius; Type locality: California:: S. Diego Co.: Cleveland N. F., Mt. Laguna, 5900 feet).
afghanicus Coiffait, 1979, see: furtivus Cameron, 1932.
agiliformis Tottenham, 1956: 293 (Gabrius; Type locality: Belgian Congo: Nialopul).
ajmonis Gridelli, 1934, see: furtivus Cameron, 1932.
albipes Stephens, 1833, see: splendidulus Gravenhorst, 1802
aliicornis Bernhauer, 1939i: 260 (Philonthus; Type locality: Kamerun: Mt. Cameron: Musake, 6350 ft.).
— Bernhauer, 1942c: 362 (Philonthus; Cameroon).
— Tottenham, 1956b: 207, 211 (Gabrius; characters).
▴ Distribution: Cameroon.
allardi Levasseur, 1967: 655 (Gabrius; Type locality: région de Kolwezi, Katanga, Ruwé, Lualaba).
almorensis Cameron, 1932: 144 (Philonthus; Type locality: Almora, Dhauli Ganga).
— Scheerpeltz, 1933: 1332 (Philonthus; catalog).
— Schillhammer, 1997: 24 (Gabrius; characters; Afghanistan).
▴ Distribution: India, Afghanistan.
altissimus Bernhauer, 1939a: 89 (Philonthus; subgenus Gabrius; Type locality: Kenya: Camp III de Elgon, zone de Bruyères, Est, 3500 m; Cratère de l'Elgon, pied du Kottobbos, 4000 m).
— Jeannel and Paulian, 1945: 84 (Philonthus; subgenus Gabrius; adult and larval characters; Kenya).
— Schillhammer, 1997: 75 (Gabrius).
▴ Distribution: Kenya.
amanensis Schillhammer, 1990: 90 (Gabrius; Type locality: Yarpuz, bei Osmaniye, 900 m).
amans Tottenham, 1962: 236 (Gabrius; Type locality: Tanganyika Terr.: Uluguru Mts., vallee de l'Ululu-Ndogo, 1500 m. Also cited from Kenya).
amasiensis Coiffait, 1980, see: tokatensis Smetana, 1977.
amulius Smetana, 1995e: 655 (Gabrius; Type locality: Ohio: 5 mi. E W. Liberty, Logan Co. Also cited from New York).
analis Heer, 1839, see: splendidulus Gravenhorst, 1802.
anatolicus Smetana, 1953: 119 (Gabrius; Type locality: Anatolia: Yeniköy, Toros, 1000 m).
— Smetana, 1961: 39 (Gabrius; characters; Turkey).
— Coiffait, 1974: 68 (Gabrius; characters; Anatolia).
▴ Distribution: Turkey.
ancoripenis Cho and Lee, 1997: 273 (Gabrius; Type locality: Korea: Dongchonri, Yeanmyon, Andong, Kyǔngsangpukdo).
angusticeps Bernhauer, 1933b: 57 (Philonthus; subgenus Gabrius; Type locality: Eritrea: Adi Caie).
apomontis Schillhammer, 2000c: 135 (Gabrius; Type locality: Philippines: Mindanao: Davao Prov.: Lk. Linau, N. slope Mt. Apo, 7900′).
appendiculatus Sharp, 1910: 131 (Gabrius; Type locality: Anglia et Scotia).
— Bernhauer and Schubert, 1914: 329 (Philonthus; catalog).
— Gridelli, 1920: 150 (Gabrius; characters; England; Scotland; Austria; Moravia; Bohemia; Italy).
— Scheerpeltz, 1933: 1332 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Wüsthoff, 1937: 31 (Philonthus; illustration of aedeagus).
— Fagel, 1945: 42 (Gabrius; characters; Belgium).
— Coiffait, 1951: 113, 115 (Gabrius; characters in key; France).
— V. Hansen, 1952: 90 (Philonthus; subgenus Gabrius; characters; Denmark).
— Smetana, 1952a: 91 (Gabrius; illustrations).
— Smetana, 1952b: 126 (Gabrius; characters; Czechoslovakia).
— Smetana, 1957: 66 (Gabrius; synonym of subnigritulus).
— Smetana, 1958: 255 (Gabrius; synonym of subnigritulus).
— Smetana, 1960c: 345 (Gabrius; synonym of subnigritulus).
— Lohse, 1964: 190 (Gabrius; synonym of subnigritulus).
— Coiffait, 1974: 58 (Gabrius; synonym of subnigritulus).
— Pope, 1977: 30 (Gabrius; synonym of subnigritulus).
— Hinton, 1981: 660 (Gabrius; characters of egg).
— Nowosad, 1990: 114 (Gabrius; collected from mole nest; notes; Poland).
— Silfverberg, 1991: 21 (Gabrius; appendiculatus is the valid name).
— Smetana, 1993b: 49 (Gabrius; synonym of subnigritulus).
— Smetana, 1995e: 628 (Gabrius; synonym of subnigritulus Reitter).
— Schillhammer, 1997: 82 (Gabrius; synonym of subnigritulus Reitter).
— Herman, 2001: 25 (Gabrius; valid name).
▴ Distribution: Europe, Russia, Canada.
subnigritulus Reitter, 1909: 132 (Philonthus; subgenus Gabrius; aberration of nigritulus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]).
— Scheerpeltz, 1933: 1353 (Philonthus; cited as subnigritulus Reitter; variety of nigritulus).
— Smetana, 1957: 66, 69 (Gabrius; cited as subnigritulus Reitter; cited as senior synonym of appendiculatus; characters).
— Smetana, 1958: 255 (Gabrius; cited as subnigritulus Reitter; characters; ecology; Czechoslovakia).
— Smetana, 1960c: 345 (Gabrius; cited as subnigritulus Reitter; characters; England; Scotland).
— Szujecki, 1963a: 34 (Gabrius; cited as subnigritulus Reitter; Poland).
— Palm, 1963b: 106 (Philonthus; subgenus Gabrius; cited as subnigritulus Reitter; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 190 (Gabrius; cited as subnigritulus Reitter; characters; central Europe).
— Horion, 1965: 192 (Gabrius; cited as subnigritulus Reitter; widespread in southern part of northern Europe and in central Europe; Dalmatia; Greece).
— Bordoni, 1973i: 329 (Gabrius; cited as subnigritulus Reitter; characters).
— Coiffait, 1974: 58 (Gabrius; cited as subnigritulus Reitter; characters; Britain; Scandinavia; Alps; Pyrenees).
— Shilov, 1975a: 376 [= 1975b: 99] (Gabrius; cited as subnigritulus Reitter; Komi region of Russia).
— Pope, 1977: 30 (Gabrius; cited as subnigritulus Reitter; Britain).
— Muona, 1979: 15 (Gabrius; cited as subnigritulus Reitter and as a valid name).
— Tóth, 1984: 88 (Gabrius; cited as subnigritulus Reitter; characters; Hungary).
— Lucht, 1987: 106 (Gabrius; cited as subnigritulus Reitter; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Smetana, 1990: 175 (Gabrius; cited as subnigritulus Reitter; characters; collecting notes; introduced into North America; British Columbia; Newfoundland; Quebec).
— Silfverberg, 1991: 21 (Gabrius; subnigritulus was made available by Smetana when using Reitter's unavailable name).
— Smetana, 1993b: 49 (Gabrius; cited as subnigritulus Reitter; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; cited as subnigritulus Reitter; Italy).
— Smetana, 1995e: 628 (Gabrius; cited as subnigritulus Reitter; [Note: Cited in the present work as as a synonym of appendiculatus because the Reitter usage was an aberration therefore an unavailable name and appendiculatus had been cited as a synonym of subnigritulus Reitter by various authors]; characters; notes; British Columbia; Newfoundland; Quebec).
— Levesque and Levesque, 1996: 291 (Gabrius; cited as subnigritulus Reitter; seasonal abundance; Quebec).
subnigritulus Smetana, 1956b: 171 (Gabrius; [Note: preoccupied; Reitter cited as author but Reitter's usage was as an aberration and so is unavailable but the name was made available by Smetana, 1956]; cited as senior synonym of appendiculatus; Type locality: Central Europe).
approximatus Hatch, 1957, see: crassulus Horn, 1884.
armeniacus Coiffait, 1966c: 506 (Gabrius; Type locality: Arménie russe, environ d'Erevan).
arnoldii Smetana, 1961: 33 (Gabrius; Type locality: UdSSR, Nordwestlicher Kaukasus, Umgebung von Tuapse).
— Coiffait, 1974: 70 (Gabrius; characters; Caucasus).
— Schillhammer, 1997: 75 (Gabrius; cited as arnoldi).
▴ Distribution: Russia.
astutoides A. Strand, 1946: 171 (Philonthus; subgenus Gabrius; Type locality: AK 13: Røa just outside of Oslo; [Note: Two other uncited localities in Norway are mentioned]).
— Smetana, 1953a: 173 (Gabrius; notes on identification; Czechoslovakia).
— Smetana, 1957: 62 (Gabrius; characters; Norway; Czechoslovakia).
— Smetana, 1958: 239 (Gabrius; characters; ecology; Czechoslovakia).
— Palm, 1963b: 105 (Philonthus; subgenus Gabrius; characters; Norway).
— Lohse, 1964: 189 (Gabrius; characters; central Europe).
— Horion, 1965: 187 (Gabrius; Norway; Germany; Czech Republic; Germany).
— Lundberg, 1972: 173 (Philonthus; Sweden).
— Coiffait, 1974: 78 (Gabrius; characters; Scandinavia; Europe; Pyrenees).
— Pope, 1977: 30 (Gabrius; Britain).
— Tóth, 1984: 87 (Gabrius; characters; Hungary).
— Lucht, 1987: 106 (Gabrius; checklist; Germany; Czechoslovakia Sweden).
— Smetana, 1991: 89 (Gabrius; characters; introduced in North America; New Brunswick; Nova Scotia; Ontario; Quebec; New Hampshire; Pennsylvania).
— Smetana, 1993b: 49 (Gabrius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
— Smetana, 1995e: 644 (Gabrius; characters; notes; Manitoba; New Brunswick; Nova Scotia; Ontario; Quebec; New Hampshire; Pennsylvania).
— Renner, 1998: 92 (Gabrius; Germany).
— Burakowski, Mroczkowski, and Stefańska, 2000: 52 (Gabrius; catalog; Poland).
▴ Distribution: Europe, Canada, USA.
astutus Erichson, 1840: 492 (Philonthus; Type locality: Austria).
— Kolenati, 1846: 21 (Philonthus; Caucasi declivitate septentrionalis).
— Hochhuth, 1849: 144 (Philonthus; characters).
— Redtenbacher, 1849: 706 (Philonthus; characters; Austria).
— Fairmaire and Laboulbène, 1856: 528 (Philonthus; characters; France).
— Kraatz, 1857d: 614 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 196 (Philonthus; characters; Austria).
— Hochhuth, 1862: 63 (Philonthus; Russia).
— Fauvel, 1865a: 297 [= 1865b: 47] (Philonthus; habitat; France).
— Fauvel, 1874: 241 [= 1874a: 467] (Philonthus; characters; France; Germany; Austria; Switzerland; Russia; Armenia; Caucasus; Lake Baikal).
— Redtenbacher, 1874: 215 (Philonthus; characters; Austria).
— Mulsant and Rey, 1876: 551 [= 1877: 407] (Gabrius; characters; France).
— Fowler, 1888: 278 (Philonthus; characters; Britain).
— Eppelsheim, 1892: 334 (Philonthus; Turkestan).
— Ganglbauer, 1895: 460 (Philonthus; subgenus Gabrius; characters; middle Europe; Caucasus; Siberia).
— Sainte-Claire Deville, 1906: 98 (Philonthus; Corsica).
— Reitter, 1909: 132 (Philonthus; subgenus Gabrius; characters; Germany).
— Bernhauer and Schubert, 1914: 329 (Philonthus; catalog).
— Gridelli, 1920: 139 (Gabrius; characters).
— Bernhauer, 1926g: 2 (Philonthus; Kamchatka region).
— Portevin, 1929: 366 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1333 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Smetana, 1953: 121 (Gabrius; Turkey).
— Smetana, 1957: 62 (Gabrius; characters; central Europe; Caucasus; Siberia).
— Smetana, 1958: 238 (Gabrius; characters; ecology; Czechoslovakia).
— Smetana, 1961a: 44 (Gabrius; Bulgaria).
— Lohse, 1964: 189 (Gabrius; characters; central Europe).
— Smetana, 1964c: 167 (Gabrius; Czechoslovakia).
— Horion, 1965: 186 (Gabrius; Britain; Holland; Belgium; Switzerland; France; Spain; Corsica; Sardinia; Italy; Dalmatia; Bosnia Hercegovina; Romania; Hungary; Czech Republic; Slovakia; Poland; Ukraine; Austria; Germany).
— Smetana, 1966: 331 (Gabrius; notes; Czechoslovakia).
— Smetana, 1967i: 556 (Gabrius; Turkey).
— Szujecki, 1969a: 257 (Gabrius; notes; Poland).
— Coiffait, 1974: 78 (Gabrius; characters; Europe; Caucasus; Asia Minor; Lebanon).
— Tóth, 1984: 86 (Gabrius; characters; Hungary).
— Lucht, 1987: 106 (Gabrius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).
— Lohse and Lucht, 1989: 168 (Gabrius; note).
— Pilon and Zanetti, 1991: 56 (Gabrius; Italy).
— Smetana, 1993b: 49 (Gabrius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
— Kashcheev, 1999a: 58 (Gabrius; Kazakhstan).
▴ Distribution: Europe, Russia, Armenia, Turkey, Lebanon, Turkmenistan, Uzbekistan, Kazakhstan.
erythrostomus Hochhuth, 1851: 22 (Philonthus; Type locality: Armenien).
— Fauvel, 1874: 241 [= 1874a: 467] (Philonthus; synonym of astutus).
— Ganglbauer, 1895: 460 (Philonthus; synonym of astutus).
— Bernhauer and Schubert, 1914: 329 (Philonthus; synonym of astutus).
— Smetana, 1958: 238 (Gabrius; synonym of astutus).
— Coiffait, 1974: 78 (Gabrius; synonym of astutus).
mendosus Bernhauer, 1928b: 12 (Philonthus; subgenus Gabrius; Type locality: Kaukasus: Borshom).
— Scheerpeltz, 1933: 1350 (Philonthus; catalog).
— Smetana, 1973b: 128 (Gabrius; notes; synonym of astutus).
— Coiffait, 1974: 78 (Gabrius; synonym of astutus).
delendus Coiffait, 1972a: 82 (Gabrius; variety of astutus; [Note: Nomen nudum (Article 15.2)]; cited from Ariège, Saint-Lary).
aterrimus Gravenhorst, 1802, see: nigritulus Gravenhorst, 1802.
atlasicus Coiffait, 1973: 111 (Gabrius; Type locality: Tachdirt, Djebel Toubkal, 2500 m, Grand Atlas marocain).
attenuatus Stephens, 1832, see: nigritulus Gravenhorst, 1802.
aureus Drugmand, 1990: 209 (Gabrius; Type locality: Turkmenistan: Repetek).
austriacus Scheerpeltz, 1947, see: velox Sharp, 1910.
autumnalis Cameron, 1932: 145 (Philonthus; Type locality: Mussoorie district: Keyarkuli, alt. 7000 feet; [Note: See lectotype designation by Schillhammer, 1997]).
— Scheerpeltz, 1933: 1333 (Philonthus; catalog).
— Schillhammer, 1997: 23 (Gabrius; lectotype designation; characters; Nepal; India; China).
▴ Distribution: India, Nepal, China.
badilikus Tottenham, 1956: 298 (Gabrius; Type locality: Urundi: Bururi, 1800–2000 m).
badius Motschulsky, 1858, see: Philonthus.
badius Kiesenwetter, 1858: 61 (Philonthus; Type locality: Sicilien).
— Fauvel, 1874: 242 [= 1874a: 468] (Philonthus; characters; Corsica; Sicily; Antolia).
— Mulsant and Rey, 1876: 543 [= 1877: 399] (Gabrius; characters; Corsica).
— Sainte-Claire Deville, 1906: 98 (Philonthus; Corsica).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Gridelli, 1920: 137 (Gabrius; characters; Greece; Corsica; Sardinia; Sicily).
— Portevin, 1929: 365 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1333 (Philonthus; catalog).
— Coiffait, 1951: 109, 114 (Gabrius; characters in key; France).
— Smetana, 1953a: 178 (Gabrius; Corsica).
— Smetana, 1957: 59 (Gabrius; characters; Greece; Sicily; Corsica).
— Smetana, 1961: 38 (Gabrius; characters; Turkey; Greece; Sicily; Sardinia; Corsica).
— Coiffait, 1974: 68 (Gabrius; characters; Corsica; Sardinia; Sicily; Greece; Asia Minor).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
▴ Distribution: France, Italy, Greece, Turkey.
insularis Gautier, 1868: 326 (Philonthus; Type locality: Corse).
— Fauvel, 1874: 242 [= 1874a: 468] (Philonthus; synonym of badius).
— Bernhauer and Schubert, 1914: 330 (Philonthus; synonym of badius).
— Coiffait, 1974: 68 (Gabrius; synonym of badius).
bafutensis Levasseur, 1967a: 957 (Gabrius; Type locality: forêt de Bafut, Cameroun, N'guemba).
basalis Stephens, 1833, see: nigritulus Gravenhorst, 1802.
balthasari Smetana, 1956, see: Gabronthus.
bernardinensis Fall, 1930, see: crassulus Horn, 1884.
beroni Raitschev, 1997: 29 (Gabrius; Type locality: Bulgaria: Osogovo mountain, in the area of mount Ruen, 2000–2151 m altitude).
bescidicus Smetana, 1954a: 116 (Gabrius; Type locality: Mähren, Beskydy, Bílá).
— Palm, 1956: 41 (Philonthus; subgenus Gabrius; characters).
— Smetana, 1957: 60 (Gabrius; characters; Czechoslovakia).
— Smetana, 1958: 243 (Gabrius; characters; ecology; Czechoslovakia).
— Palm, 1963b: 104 (Philonthus; subgenus Gabrius; characters; Finland).
— Lohse, 1964: 188 (Gabrius; characters; central Europe).
— Coiffait, 1974: 67 (Gabrius; characters; Czechoslovakia).
— Tóth, 1984: 85 (Gabrius; characters).
— Lucht, 1987: 106 (Gabrius; checklist; Czechoslovakia).
— Silfverberg, 1992: 16 (Gabrius; Sweden; Finland).
— Smetana, 1993b: 49 (Gabrius; Czech Republic).
▴ Distribution: Czech Republic, Finland, Sweden.
besucheti Schillhammer, 1997: 42 (Gabrius; Type locality: India: Meghalaya Khasi Hills Weiloi, 1700 m).
bidentatus Horn, 1884: 214 (Philonthus; Type locality: California, Vancouver, Washington Territory).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Hatch, 1957: 204 (Philonthus; subgenus Gabrius; characters; British Columbia; Washington; Oregon).
— Moore, 1965: 100 (Gabrius).
— Smetana, 1995e: 710 (Gabrius; characters; notes; British Columbia; California; Oregon; Washington).
▴ Distribution: Canada, USA.
bifidus Tottenham, 1962: 244 (Gabrius; Type locality: Kenya: Molo, Mau Escarpment, 2150–2200 m).
birmanus Schillhammer, 1997: 44 (Gabrius; Type locality: Burma: Carin Asciuü Ghecù, 1400–1500 m. Also cited from Thailand).
bishopi Sharp, 1910: 131 (Gabrius; Type locality: Scotia: Beattock and Thornhill).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Gridelli, 1920: 148 (Gabrius; characters; England).
— Scheerpeltz, 1933: 1334 (Philonthus; catalog).
— V. Hansen, 1952: 91 (Philonthus; subgenus Gabrius; characters; Denmark).
— Coiffait, 1953a: 266 (Gabrius; [Note: Misidentified as suffragani Joy according to Schillhammer, 1997: 75 who cited it as though a synonym of bishopi]; characters; France).
— Smetana, 1957: 64, 69 (Gabrius; characters; England; Scandinavia; Germany; Austria; France).
— Smetana, 1960c: 336 (Gabrius; characters; Scotland; Britain; Germany; Austria; France).
— Lohse, 1964: 190 (Gabrius; characters; central Europe).
— Horion, 1965: 192 (Gabrius; Britain; France; Austria; Germany).
— Coiffait, 1974: 61 (Gabrius; characters; Britain; Europe; Alps; Corsica).
— Pope, 1977: 30 (Gabrius; Britain).
— Tóth, 1984: 89 (Gabrius; characters).
— Lucht, 1987: 106 (Gabrius; checklist; Germany; Austria).
— Eyre, Luff, and Lott, 2000: 33 (Gabrius; Britain).
▴ Distribution: Europe.
biturigensis Coiffait, 1951, see: breviventer Sperk, 1835.
brevipennis Horn, 1884: 212 (Philonthus; Type locality: Pt. Pins LS [Lake Superior]; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from north shore of Lake Superior).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Smetana, 1995e: 613 (Gabrius; lectotype designation: Pt. Pins LS [Lake Superior]; characters; notes; Alberta; British Columbia; Manitoba; New Brunswick; Newfoundland; Northwest Territories; Nova Scotia; Ontario; Quebec; Saskatchewan; Alaska; Maine; Massachusetts; Michigan; New Hampshire; New York; Vermont; Washington; Wisconsin).
— Levesque and Levesque, 1996: 292 (Gabrius; seasonal abundance; Quebec).
▴ Distribution: Canada, USA.
malkini Hatch, 1957: 200 (Philonthus; subgenus Gabrius; Type locality: Oregon: Coos Co.: Marshfield, now Coos Bay).
— Moore, 1965: 100 (Gabrius).
— Smetana, 1965: 29 (Gabrius; characters; Newfoundland).
— Smetana, 1965a: 8 (Gabrius; Alaska).
— Smetana, 1995e: 613 (Gabrius; synonym of brevipennis).
brevissimus Tottenham, 1956: 297 (Gabrius; Type locality: Ruanda: Tshuruyaga, forêt du Rugege, 2400 m).
breviventer Sperk, 1835: 153 (Xantholinus; Type locality: südliches Russland).
— Solsky, 1874: 191 (Scopaeus; characters; Turkestan).
— Ganglbauer, 1895: 460 (Philonthus; synonym of thermarum).
— Bernhauer and Schubert, 1912: 248 (Scopaeus; synonym of gracilis).
— Bernhauer and Schubert, 1914: 357 (Philonthus; synonym of thermarum).
— Cameron, 1932: 134 (Philonthus; synonym of thermarum).
— Scheerpeltz, 1957c: 234 (Philonthus; synonym of thermarum).
— Smetana, 1958: 217 (Philonthus; synonym of thermarum).
— Coiffait, 1974: 114 (Gabronthus; synonym of thermarum).
— Schillhammer, 1997: 16 (Gabrius; characters; synonyms).
▴ Distribution: Tunisia, Algeria, Europe, Russia, Caucasus, Turkey, India, USA.
coxalus Hochhuth, 1872: 142 (Philonthus; Type locality: Not cited, but presumably from Gouvernements Kiew and/or Volhynien; [Note: See lectotype designation by Gusarov, 1991a]).
— Bernhauer and Schubert, 1914: 347 (Philonthus; synonym of nigritulus).
— Cameron, 1932: 147 (Philonthus; cited as coxalis; synonym of nigritulus).
— Smetana, 1958: 249 (Gabrius; synonym of nigritulus).
— Coiffait, 1974: 54 (Gabrius; synonym of nigritulus).
— Gusarov, 1991a: 7 (Gabrius; valid species; lectotype designation: [Note: Locality not cited]; notes).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
— Smetana, 1995e: 624 (Gabrius; characters; notes; California).
— Schillhammer, 1997: 16 (Gabrius; synonym of breviventer).
pennatus Sharp, 1910: 130 (Gabrius; Type locality: Anglia et Scotia; [Note: See lectotype designation by Gusarov, 1991a]).
— Bernhauer and Schubert, 1914: 350 (Philonthus; catalog).
— Gridelli, 1920: 149 (Gabrius; characters; England; Scotland; Austria; Moravia; Germany; Dalmatia; Italy).
— Scheerpeltz, 1933: 1354 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Wüsthoff, 1937: 31 (Philonthus; illustration of aedeagus).
— Fagel, 1945: 41 (Gabrius; characters; Belgium).
— Jeannel and Jarrige, 1949: 376 (Philonthus; subgenus Gabrius; cave species; Spain).
— Coiffait, 1951: 111, 114 (Gabrius; characters in key; France).
— V. Hansen, 1952: 90 (Philonthus; subgenus Gabrius; characters; Denmark).
— Smetana, 1952b: 125 (Gabrius; characters).
— Smetana, 1953: 122 (Gabrius; Turkey).
— Smetana, 1957: 63, 70 (Gabrius; characters; Europe; Caucasus; Turkey; Algeria; Tunisia).
— Smetana, 1958: 254 (Gabrius; characters; ecology; Czechoslovakia).
— Smetana, 1960c: 340 (Gabrius; characters; Caucasus; Turkey; Algeria; Tunisia).
— Smetana, 1961a: 45 (Gabrius; Bulgaria).
— Palm, 1963b: 106 (Philonthus; subgenus Gabrius; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 191 (Gabrius; characters; central Europe).
— Smetana, 1964d: 78 (Gabrius; Czechoslovakia).
— Focarile, 1964: 56 (Gabrius; Italy).
— Scheerpeltz, 1965b: 403 (Gabrius; Sicily).
— Scheerpeltz, 1965c: 507 (Gabrius; Sicily).
— Horion, 1965: 197 (Gabrius; Algeria; Tunisia; Turkey; Caucasus; Russia; Finland; Sweden; Norway; Britain; Denmark; widespread in central and southern Europe).
— Raianu, 1969a: 325 (Gabrius; characters; ecological notes; Romania).
— Bordoni, 1973i: 327 (Gabrius; characters).
— Coiffait, 1974: 65 (Gabrius; characters; Europe; Britain; Asia Minor).
— Pope, 1977: 30 (Gabrius; Britain).
— Frank, 1982: 36 (Gabrius; list of parasites).
— Tóth, 1984: 93 (Gabrius; characters; Hungary).
— Lucht, 1987: 106 (Gabrius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Pilon and Zanetti, 1991: 56 (Gabrius; Italy).
— Gusarov, 1991a: 7 (Gabrius; lectotype designation: [Note: Locality not cited]; synonym of coxalus).
— Smetana, 1993b: 49 (Gabrius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; synonym of coxalus).
— Smetana, 1995e: 624 (Gabrius; synonym of coxalus).
— Schillhammer, 1997: 16 (Gabrius; synonym of breviventer).
— Assing, Frisch, Kahlen, et al., 1998: 138 (Gabrius; synonym of breviventer).
coxatus Bernhauer, 1914d: 9 (Philonthus; unjustified emendation of coxalus Hochhuth [Articles 32 and 33]; [preoccupied]).
— Scheerpeltz, 1933: 1352 (Philonthus; synonym of nigritulus).
— Smetana, 1960c: 325 (Gabrius; synonym of nigritulus; [Note: This name is cited as a synonym of breviventer rather than nigritulus because it is an emendation of coxalus which is a synonym of breviventer]).
nigrituloides Cameron, 1932: 148 (Philonthus; Type locality: India).
— Scheerpeltz, 1933: 1352 (Philonthus; catalog).
— Yadav, Chand, and Yadav, 1995: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
— Schillhammer, 1997: 16 (Gabrius; synonym of breviventer).
biturigensis Coiffait, 1951: 110, 113 (Gabrius; Type locality: Indre: Châteauroux).
— Smetana, 1960c: 352 (Gabrius; characters from original description; species uncertain).
— Coiffait, 1974: 65 (Gabrius; synonym of pennatus).
— Smetana, 1995e: 624 (Gabrius; synonym of coxalus).
hublei Coiffait and Segers, 1985: 298 (Gabrius; Type locality: Belgique: Commune de Gamaarden, province de Flandre orientale).
— Lohse, 1988: 42 (Gabrius; synonym of velox).
— Schillhammer, 1990: 92 (Gabrius; characters; notes).
— Smetana, 1995e: 624 (Gabrius; synonym of coxalus).
— Assing, Frisch, Kahlen, et al., 1998: 138 (Gabrius; synonym of breviventer).
bulbifer Schillhammer, 1997: 20 (Gabrius; Type locality: Pakistan: Swat, Malam Jabba, 2300–2400 m).
bulgaricus Coiffait, 1971b: 279 (Gabrius; Type locality: Bulgarie: Zabârdo, Rodopes).
burckhardti Schillhammer, 1997: 54 (Gabrius; Type locality: Thailand: Chiang Mai, Doi Inthanon, 2450 m).
burgeoni Bernhauer, 1928g: 107 (Philonthus; Type locality: Belg.-Kongo: Manyema).
— Cameron, 1929: 64 (Philonthus; characters; Belgian Congo).
— Scheerpeltz, 1933: 1335 (Philonthus; catalog).
— Tottenham, 1949b: 353 (Philonthus; characters).
— Tottenham, 1956b: 220 (Gabrius; characters).
▴ Distribution: Congo.
canariensis Fauvel, 1898: 100 (Philonthus; named for simulans as misidentified by Wollaston, 1864; Type locality: Canaries).
— Wollaston, 1864: 574 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 331 cited simulans Wollaston as a synonym of canariensis; Wollaston attributed it to his earlier description of the species]; notes; Canary Islands).
— Fauvel, 1902b: 111 (Philonthus; Canaries).
— Bernhauer and Schubert, 1914: 331 (Philonthus; catalog).
— Scheerpeltz, 1933: 1335 (Philonthus; catalog).
— Smetana, 1960c: 327 (Gabrius; characters; Canary Islands).
— Smetana, 1960c: 307 (Gabrius; [Note: Misidentified as simulans according to Smetana, 1962: 97]; characters; Madeira and Canary Islands).
— Smetana, 1962: 97 (Gabrius; characters; Canary Islands).
— Coiffait, 1974: 62 (Gabrius; characters; Canary Islands).
— Assing, 1999a: 346 (Gabrius; Canary Islands).
— Assing and Wunderle, 1999: 437 (Gabrius; Canary Islands).
▴ Distribution: Canary Islands.
capizzii Hatch, 1957, see: adustus Casey, 1915.
caporiaccoi Gridelli, 1934: 81 (Gabrius; Type locality: Valle Punmah: Jula di Punmah, 3100 m).
— Schillhammer, 1997: 30 (Gabrius; characters; India; Pakistan).
▴ Distribution: Afghanistan, Pakistan, India.
darahensis Coiffait, 1979d: 557 (Gabrius; Type locality: Nouristan: Darah i Nour, 3000 m).
capucinus Schillhammer, 1997: 41 (Gabrius; Type locality: India: Uttar Pradesh, Garhwal, au-dessus Pauri, 1900 m. Also cited from Pakistan and Nepal).
cascadensis Scheerpeltz, 1974: 161 (Gabrius; Type locality: Natal: Royal Natal National Park, The Cascades, 5200 ft.).
chapmani Bernhauer, 1939a: 69, 89 (Philonthus; Type locality: Kenya: Mt. Elgon, Osthang, Suam fishing hut, 2400 m).
— Jeannel and Paulian, 1945: 87 (Philonthus; subgenus Gabrius; Kenya).
— Tottenham, 1949b: 354 (Philonthus; characters).
— Schillhammer, 1997: 76 (Gabrius).
▴ Distribution: Kenya.
chiliensis Coiffait and Saiz, 1968: 348 (Gabrius; Type locality: Chile: Romeral, Curicó).
— Fauvel, 1866a: 346 [= 1866b: 97] (Philonthus; [Note: Cited as nigritulus; misidentification of chiliensis according to Coiffait and Saiz, 1968: 348 who cited the name as nigritus]; characters; Chile).
— Saiz, 1971b: 340 (Gabrius; cited as chilensis; Chile).
▴ Distribution: Chile.
chiriquiensis Herman, 2001: 44 (Gabrius; replacement name for pilipes Bierig).
pilipes Bierig, 1940a: 142 (Philonthus; subgenus Gabrius; [preoccupied]; Type locality: Panamá: Volcán de Chiriqui, 1500 m).
chungkingensis Bernhauer, 1943, see: disjunctus Bernhauer and Schubert, 1914.
cinctiventris Scheerpeltz, 1965: 229 (Gabrius; Type locality: N.E. Burma, Kambaiti, 7000 ft.).
cinerascens Stephens, 1833, see: Neobisnius.
claviger Tottenham, 1956: 294 (Gabrius; Type locality: Ruanda: Rutovu, forêt du Rugege, 2350 m).
cleon Smetana, 1995e: 635 (Gabrius; Type locality: Washington: 10 mi. S. Sequim).
clunalis Horn, 1884: 213 (Philonthus; Type locality: California, probably in the southern portion; [Note: See lectotype designation by Smetana, 1995e]).
— Bernhauer and Schubert, 1914: 332 (Philonthus; catalog).
— Smetana, 1995e: 702 (Gabrius; lectotype designation; characters; notes; California).
▴ Distribution: USA.
cobosi Coiffait, 1964c: 171 (Gabrius; Type locality: Espagne: sumidero C. Navas, Sierra de Ronda, Province de Malaga).
coeruleipennis Coiffait, 1983: 276 (Gabrius; Type locality: Cachemire: Gangria, 3500 m).
collartianus Cameron, 1935a: 374 (Philonthus; Type locality: Mahagi: Nialopul; Nizi: Buba).
— Tottenham, 1956: 228, 292, 304 (Gabrius; characters; notes; Urundi).
— Levasseur, 1971: 299 (Gabrius; Sierra Leone).
▴ Distribution: Sierra Leone, Congo, Burundi.
colossus Schillhammer, 1997: 67 (Gabrius; Type locality: Bali: Penulisan).
colubrinus Fauvel, 1868, see: fauvelii Coquerel, 1866.
communis Motschulsky, 1860, see: Philonthus.
confusoides Schillhammer, 2000c: 120 (Gabrius; Type locality: Nepal: Parbat Distr., Ghoropani Pass, N slope, 2700 m).
confusus Cameron, 1932: 148 (Philonthus; Type locality: Kansar, Chakrata dist., 7050′; [Note: See lectotype designation by Schillhammer, 1997]. Originally cited from Chakrata, Mussoorie, and Simla districts, alt. 7000–8000 feet).
— Scheerpeltz, 1933: 1337 (Philonthus; catalog).
— Schillhammer, 1997: 21 (Gabrius; lectotype designation: Kansar, Chakrata dist., 7050′; characters; India).
— Schillhammer, 2000c: 120 (Gabrius; India; Nepal).
▴ Distribution: India, Nepal.
conicus Bernhauer, 1939, see: sharpianus Cameron, 1930.
contiguus Motschulsky, 1860, see: Philonthus.
convexiceps Scheerpeltz, 1965: 232 (Gabrius; Type locality: N.E. Burma, Kambaiti, 7000 ft.).
coryndoni Cameron, 1942b: 329 (Philonthus; subgenus Gabrius; Type locality: Chyulu Hills, altitude 5600 feet).
coxalus Hochhuth, 1872, see: breviventer Sperk, 1835.
coxatus Bernhauer, 1914, see: breviventer Sperk, 1835.
crassulus Horn, 1884: 213 (Philonthus; Type locality: Nevada).
— Bernhauer and Schubert, 1914: 333 (Philonthus; catalog).
— Smetana, 1995e: 658 (Gabrius; characters; notes; California; Nevada; Oregon; Washington).
▴ Distribution: USA.
bernardinensis Fall, 1930: 253 (Philonthus; Type locality: southern California, San Bernardino Mts.).
— Scheerpeltz, 1933: 1334 (Philonthus; catalog).
— Moore, 1965: 100 (Gabrius; cited as bernardensis).
— Smetana, 1995e: 658 (Gabrius; synonym of crassulus).
approximatus Hatch, 1957: 200 (Philonthus; subgenus Gabrius; Type locality: Washington: Kooskooskie, Walla Walla Co. Also cited from Oregon).
curvipenis Schillhammer, 1997: 21 (Gabrius; Type locality: E. Nepal: [Sankhua Sabha District], Kosi, Val. Induwa Khola, 2000 m).
cushmani Hatch, 1957: 204 (Philonthus; subgenus Gabrius; Type locality: Washington: Lake Cushman).
— Moore, 1965: 100 (Gabrius).
— Smetana, 1995e: 709 (Gabrius; characters; notes; British Columbia; Oregon; Washington).
▴ Distribution: Canada, USA.
femineus Hatch, 1957: 204 (Philonthus; subgenus Gabrius; Type locality: Washington: Easton).
cyphonotus Joy, 1913: 25 (Gabrius; Type locality: Caucasus and the borders of the Caspian Sea; Daghestan, Lenkoran, etc.).
— Scheerpeltz, 1933: 1339 (Philonthus; catalog).
— Smetana, 1953a: 179 (Gabrius; Daghestan; Transcaucasia).
— Smetana, 1957: 65 (Gabrius; characters; Turkestan; Caucasus).
— Smetana, 1960c: 330 (Gabrius; characters; Caucasus; Daghestan; Transcaucasian region).
— Coiffait, 1974: 60 (Gabrius; characters; Caucasus; Transcaucasus; region of Caspian Sea).
▴ Distribution: Russia, Azerbaijan, “Turkestan”.
damon Smetana, 1984: 135 (Gabrius; Type locality: Japan: Yamagata Pref.: Tachiyzawa).
darahensis Coiffait, 1979, see: caporiaccoi Gridelli, 1934.
darjeelingensis Schillhammer, 1997: 53 (Gabrius; Type locality: India: W. Bengal, Darjeeling distr., Tigerhill, 2500–2600 m).
deceptivus Cameron, 1943: 132 (Philonthus; subgenus Gabrius; Type locality: Kashmir: Gulmarg; [Note: See lectotype designation by Schillhammer, 1997]).
— Schillhammer, 1997: 31 (Gabrius; lectotype designation; characters; India; Pakistan).
▴ Distribution: Pakistan, India.
deceptor Cameron, 1932: 146 (Philonthus; Type locality: Simla Hills: Gahan; [Note: See lectotype designation by Schillhammer, 1997]. Also cited from Mussoorie district, alt. 6000–7000 feet).
— Scheerpeltz, 1933: 1339 (Philonthus; catalog).
— Coiffait, 1982: 27 (Gabrius; India).
— Schillhammer, 1997: 18 (Gabrius; lectotype designation: Gahan, Simla Hills, 7000′; characters; India; Nepal; Bhutan; China).
— Schillhammer, 2000c: 120 (Gabrius; Nepal).
▴ Distribution: India, Nepal, Bhutan, Myanmar, China.
tensicornis Scheerpeltz, 1965: 230 (Gabrius; Type locality: N.E. Burma, Kambaiti, 7000 ft.).
deceptoroides Schillhammer, 1997: 19 (Gabrius; Type locality: Nepal: 412 Sankhua Sabha Dist., Arun Valley betw. Mure and Hurure, 2050–2150 m).
decipiens Horn, 1884, see: picipennis Mäklin, 1852.
deflexus Tottenham, 1956b: 223 (Gabrius; Type locality: Nyassaland: Qodeni).
delamarei Jarrige, 1970: 49 (Gabrius; Type locality: Madagascar: Haut Sambirano, 1000 m).
delendus Coiffait, 1972 [Note: Nomen nudum], see: astutus Erichson, 1840.
demades Smetana, 1984: 138 (Gabrius; Type locality: Japan: Toyama Pref.: Tateyama Mts., Bijodaira, 1000 m).
demarcatus Tottenham, 1940, see: sharpianus Cameron, 1930.
dentipes Eppelsheim, 1889: 172 (Philonthus; Type locality: Arragonien).
— Bernhauer and Schubert, 1914: 335 (Philonthus; catalog).
— Gridelli, 1920: 130 (Gabrius; characters; Spain).
— Scheerpeltz, 1933: 1339 (Philonthus; catalog).
— Peyerimhoff, 1937: 265 (Philonthus; characters; notes; Morocco).
— C. Koch, 1941: 64 (Philonthus; subgenus Gabrius; Morocco).
— Smetana, 1957: 57 (Gabrius; characters; Spain; North Africa).
— Smetana, 1961: 36 (Gabrius; characters; Spain; North Africa).
— Coiffait, 1964c: 166 (Gabrius; characters; Morocco).
— Coiffait, 1974: 74 (Gabrius; characters; Morocco).
▴ Distribution: Spain, Morocco.
desperatus Schillhammer, 1997: 44 (Gabrius; Type locality: Burma: Carin Asciuü Checù, 1400–1500 m).
dieckmanni Smetana, 1957: 70 (Gabrius; Type locality: Deutschland: Thüringen, Salzstelle bei Esperstedt-Frankenhausen, Kyffhäuser).
— Smetana, 1960c: 315 (Gabrius; characters; Germany).
— Lohse, 1964: 189 (Gabrius; characters; central Europe).
— Horion, 1965: 192 (Gabrius; Germany).
— Coiffait, 1974: 64 (Gabrius; characters; central Europe).
— Lucht, 1987: 106 (Gabrius; checklist; Germany).
▴ Distribution: Germany.
diengensis Schillhammer, 1997: 65 (Gabrius; Type locality: C-Java: Dieng).
disjunctus Bernhauer and Schubert, 1914: 336 (Philonthus; replacement name for distans Fauvel).
— Cameron, 1932: 135 (Philonthus; characters; Burma).
— Scheerpeltz, 1933: 1340 (Philonthus; catalog).
— Schillhammer, 1997: 48 (Gabrius; characters; India; Nepal; Thailand; China).
— Schillhammer, 2000c: 132 (Gabrius; China; Nepal).
▴ Distribution: India, Nepal, Myanmar, Thailand, China.
distans Fauvel, 1895: 265 (Philonthus; [preoccupied]; Type locality: Bírmanie, Carin Asciuii Ghécu, 1400–1500 m).
— Cameron, 1932: 135 (Philonthus; synonym of disjunctus).
— Schillhammer, 1997: 48 (Gabrius; synonym of disjunctus).
disjunctus Cameron, 1925: 59 (Philonthus; [preoccupied]; replacement name for distans Fauvel).
fluviatilis Cameron, 1932: 140 (Philonthus; Type locality: Assam: Naga Hills, alt. 5000 feet).
— Scheerpeltz, 1933: 1342 (Philonthus; catalog).
— Schillhammer, 1997: 48 (Gabrius; synonym of disjunctus).
chungkingensis Bernhauer, 1943: 75 (Philonthus; subgenus Gabrius; Type locality: China: Prov. Szechuan, Gebirge b. Chungking).
— Smetana, 1973b: 127 (Gabrius; characters; notes).
— Schillhammer, 1997: 48 (Gabrius; synonym of disjunctus).
disjunctus Cameron, 1925, see: disjunctus Bernhauer and Schubert, 1914.
distans Fauvel, 1895, see: disjunctus Bernhauer and Schubert, 1914.
distans Horn, 1884: 211 (Philonthus; Type locality: L. Tahoe, Nev.; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from Vancouver; Nevada).
— Bernhauer and Schubert, 1914: 336 (Philonthus; catalog).
— Fall, 1926: 150 (Philonthus; Alaska).
— Smetana, 1995e: 687 (Gabrius; lectotype designation: L. Tahoe, Nev.; characters; notes; California; Nevada; Oregon).
▴ Distribution: USA.
molliculus Casey, 1915: 436 (Philonthus; Type locality: California: Truckee, 6000 ft.).
— Scheerpeltz, 1933: 1351 (Philonthus; catalog).
— Smetana, 1995e: 687 (Gabrius; lectotype designation; synonym of distans).
distinguendus Cameron, 1950g: 41 (Philonthus; subgenus Gabrius; Type locality: Belgian Congo: lac N'Gando, pied volc. Karisimbi, alt. 2400 m).
divisifrons Cameron, 1950g: 39 (Philonthus; subgenus Gabrius; Type locality: Belgian Congo: mont Sesero, près Bitashimva, alt. 2000 m).
doderoi Gridelli, 1920: 135 (Gabrius; Type locality: Ficuzza, Sicilia; S. Biase, Vallo Lucano).
— Scheerpeltz, 1933: 1340 (Philonthus; catalog).
— Smetana, 1957: 59, 61 (Gabrius; [Note: Misidentified as laticollis according to Smetana, 1965f: 258]; characters; Spain; France; Sicily).
— Smetana, 1961: 38, 39 (Gabrius; [Note: Misidentified as laticollis according to Smetana, 1965f: 258] characters; Spain; France; Sicily; Malta).
— Scheerpeltz, 1961d: 130 (Gabrius; Italy).
— Coiffait, 1963: 21 (Gabrius; notes).
— Smetana, 1965f: 258 (Gabrius; notes).
— Coiffait, 1974: 77 (Gabrius; characters; Italy; Sicily).
— Sparacio, 1995: 160 (Gabrius; characters; notes; Sicily).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
▴ Distribution: Italy, France, Spain.
satanellus Bordoni, 1973i: 332 (Gabrius; Type locality: Lazio, Promontorio del Circeo).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; probable synonym of doderoi; Italy).
— Schillhammer, 1996a: 82 (Gabrius; synonym of doderoi).
dolosus Tottenham, 1956b: 219 (Gabrius; Type locality: Kenya: Muguga, near Kikuyu).
dorjulensis Coiffait, 1977, see: fimetarioides Scheerpeltz, 1976.
downiei Hatch, 1957, see: punctatellus Horn, 1884.
dubiosus Schillhammer, 1997, see: rufocinctus Schillhammer, 1997.
dufbergi Gillerfors, 1997: 84 (Gabrius; Type locality: Sweden: Skåne, Vomb).
egens Sharp, 1874: 44 (Philonthus; Type locality: Hiogo).
— Duvivier, 1883: 145 (Philonthus; cited as egeus; [Note: Misspelling: Bernhauer and Schubert, 1914: 336 cited egeus as a synonym of egens]; checklist).
— Bernhauer and Schubert, 1914: 336 (Philonthus; catalog).
— Scheerpeltz, 1933: 1341 (Philonthus; catalog).
— Smetana, 1960c: 303 (Gabrius; characters; Japan).
— Smetana, 1984: 130 (Gabrius; characters; Japan).
— Cho and Lee, 1997: 272 (Gabrius; characters; Korea).
▴ Distribution: Japan.
egnatius Smetana, 1995e: 662 (Gabrius; Type locality: California: S. Mateo Co.).
ellipticeps Scheerpeltz, 1965: 231 (Gabrius; Type locality: N.E. Burma, Kambaiti, 7000 ft.).
emeishanus Schillhammer, 1996: 66 (Gabrius; Type locality: China: Sichuan Emei Shan).
eremius Herman, 2001: 44 (Gabrius; replacement name for horni Bernhauer and Schubert).
horni Bernhauer and Schubert, 1914: 341 (Philonthus; replacement name for parvus Horn; [preoccupied]).
— Moore, 1965: 100 (Gabrius).
— Smetana, 1995e: 638 (Gabrius; characters; notes; Arizona; California; Colorado; Nevada; Nevada; New Mexico; Utah).
— Downie and Arnett, 1996: 382 (Gabrius; characters; USA).
— Herman, 2001: 44 (Gabrius; synonym of eremius).
parvus Horn, 1884: 214 (Philonthus; [Note: Preoccupied as a secondary homonym; when this name was replaced it was a secondary homonym of parvus Solier, a species that was in Philonthus at the time but which was described in and is currently in Staphylinus]; Type locality: Arizona; [Note: See lectotype designation by Smetana, 1995e]).
— Blatchley, 1910: 388 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 341 (Philonthus; synonym of horni).
— Smetana, 1995e: 638 (Gabrius; lectotype designation; synonym of horni).
erythrostomus Hochhuth, 1851, see: astutus Erichson, 1840.
eumolpus Smetana, 1995e: 642 (Gabrius; Type locality: Arizona: Sta. Rita Mts., Mt Wright-son, 9000–9453 ft.).
excelsoides Schillhammer, 1997: 36 (Gabrius; Type locality: Nepal: Manang Distr. For. W Bagarchhap, 2250 m).
excelsus Cameron, 1932, see: perexcelsus Tottenham, 1939.
excisicauda Bernhauer, 1939a: 87 (Philonthus; Type locality: Kenya: Mt. Aberdare, Westhang, Mais. forest. Kinangop, 2600 m; Mt. Elgon, Osthang, Suam fishing hut, 2400 m).
exiguus Nordmann, 1837: 105 (Philonthus; Type locality: Berolinum).
— Erichson, 1839a: 475 (Philonthus; characters; Germany).
— Heer, 1839: 271 (Philonthus; characters; Switzerland).
— Erichson, 1840: 491 (Philonthus; characters; Germany).
— Redtenbacher, 1849: 706 (Philonthus; characters; Austria).
— Kraatz, 1857d: 615 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 196 (Philonthus; characters; Austria).
— Thomson, 1867a: 43 (Philonthus; characters; Scandinavia).
— Fauvel, 1874: 262 [= 1874a: 488] (Philonthus; characters; France; Scandinavia; Germany; Switzerland; Austria; Russia; Italy; Corsica; Spain).
— Redtenbacher, 1874: 214 (Philonthus; characters; Austria).
— Seidlitz, 1875: 278 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 549 [= 1877: 405] (Gabrius; characters; France).
— Ragusa, 1892: 167 (Philonthus; Sicily).
— Ganglbauer, 1895: 461 (Philonthus; subgenus Gabrius; characters; Europe).
— Reitter, 1909: 132 (Philonthus; subgenus Gabrius; characters; Germany).
— Bernhauer and Schubert, 1914: 337 (Philonthus; catalog).
— Gridelli, 1920: 133 (Gabrius; characters; Corsica; Italy).
— Portevin, 1929: 367 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1341 (Philonthus; catalog).
— Español, 1945: 93 (Philonthus; subgenus Gabrius; Spain).
— Coiffait, 1951: 110, 114 (Gabrius; characters in key; France).
— V. Hansen, 1952: 88 (Philonthus; subgenus Gabrius; characters; Denmark).
— Smetana, 1957: 61 (Gabrius; characters; Germany; Czechoslovakia; Austria).
— Smetana, 1958: 237 (Gabrius; characters; ecology; Czechoslovakia).
— Smetana, 1961: 39 (Gabrius; characters; Europe).
— Smetana, 1961a: 43 (Gabrius; Bulgaria).
— Palm, 1963b: 104 (Philonthus; subgenus Gabrius; characters; Sweden; Denmark).
— Lohse, 1964: 188 (Gabrius; characters; central Europe).
— Horion, 1965: 185 (Gabrius; Denmark; Sweden; Holland; Belgium; Switzerland; France; Spain; Corsica; Sardinia; Italy; Hercegovina; Romania; Hungary; Czechoslovakia; Poland; Ukraine; Germany; Austria).
— Korge, 1971a: 7 (Gabrius; Turkey).
— Coiffait, 1974: 75 (Gabrius; characters; Europe; Corsica; France).
— Osella and Zanetti, 1975: 110 (Gabrius; collected from nest of Talpa europaea; notes; Italy).
— Pope, 1977: 30 (Gabrius; Britain).
— Frank, 1982: 36 (Gabrius; list of parasites).
— Tóth, 1984: 85 (Gabrius; characters; Hungary).
— Lucht, 1987: 106 (Gabrius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Pilon and Zanetti, 1991: 55 (Gabrius; Italy).
— Smetana, 1993b: 49 (Gabrius; Czech Republic; Slovakia).
— Hodge and Jones, 1995: 41 (Gabrius; characters; Britain).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
▴ Distribution: Europe, Russia, Turkey.
nigricornis Mulsant and Rey, 1876: 551 [= 1877: 407 (Gabrius; Type locality: environs de Lyon).
exspectatus Smetana, 1952: 121 (Gabrius; Type locality: Ost-Slowakei, Gebirge Vihorlat, Remetské Hámry).
— Smetana, 1954a: 118 (Gabrius; characters; Bulgaria; Bosnia; Slovakia; Balkan region).
— Palm, 1956: 41 (Philonthus; subgenus Gabrius; characters).
— Smetana, 1957: 60 (Gabrius; characters; Ukraine; Carpathian region; Bulgaria; Bosnia; Czechoslovakia; Austria; Sweden).
— Smetana, 1958: 241 (Gabrius; characters; ecology; Czechoslovakia).
— Smetana, 1961a: 44 (Gabrius; Bulgaria).
— Palm, 1963b: 104 (Philonthus; subgenus Gabrius; characters; Sweden; Norway; Finland).
— Lohse, 1964: 187 (Gabrius; cited as expectatus; characters; central Europe).
— Horion, 1965: 189 (Gabrius; Russia; Ukraine; Bulgaria; Bosnia; Albania; Czech Republic; Slovakia; Germany; Finland; Sweden; Norway).
— Pototskaia, 1966: 142, 154 (Gabrius; cited as expectatus; larval characters).
— Pototskaia, 1967: 56 (Gabrius; larval characters).
— Raianu, 1969a: 324 (Gabrius; characters; Romania).
— Szujecki, 1969a: 257 (Gabrius; notes; Poland).
— Korge, 1971a: 7 (Gabrius; cited as expectatus; Turkey).
— Coiffait, 1974: 66 (Gabrius; cited as expectatus; characters; Europe; Balkans; Pyrenees).
— Shilov, 1975a: 376 [= 1975b: 99] (Gabrius; cited as expectatus; Komi region of Russia).
— Pototskaia, 1976: 173 (Gabrius; cited as expectatus; larval characters).
— Tóth, 1984: 84 (Gabrius; cited as expectatus; characters; Hungary).
— Lucht, 1987: 106 (Gabrius; checklist; Germany; Poland; Czechoslovakia; Austria).
— Lohse and Lucht, 1989: 168 (Gabrius; note).
— Gusarov, 1991a: 6 (Gabrius; notes).
— Smetana, 1993b: 49 (Gabrius; Czech Republic; Slovakia).
— Olberg and Andersen, 2000: 223 (Gabrius; cited as expectatus; attracted to trunk-window traps baited with Fomes fomentarius and Phellinus spp.; Norway).
▴ Distribution: Europe, Russia, Turkey.
tichomirovae Bogdanov, 1984: 458 (Gabrius; Type locality: Zakarpat'e, Uzhgorodskii r-n, Khrebet Makovitsa).
fageli Drugmand, 1990: 210 (Gabrius; Type locality: Anatolie occidentale: Abant Dagh).
fallaciosus Horn, 1884: 212 (Philonthus; Type locality: Pennsylvania; [Note: See lectotype designation by Smetana, 1995e]. Also cited from Massachusetts).
— Bernhauer and Schubert, 1914: 337 (Philonthus; catalog).
— Smetana, 1995e: 683 (Gabrius; lectotype designation: Pennsylvania; characters; notes; Ontario; Quebec; Connecticut; Indiana; Iowa; Maine; Massachusetts; Michigan; New Hampshire; New Jersey; New York; North Carolina; Ohio; Pennsylvania; Tennessee; Vermont; Virginia).
— Downie and Arnett, 1996: 381 (Philonthus; characters; USA).
▴ Distribution: Canada, USA.
falsimitator Schillhammer, 1997: 39 (Gabrius; Type locality: India: Uttar Pradesh, Garhwal, 2 km E de Dhanolti, 2250 m. Also cited from Nepal and Pakistan).
fauvelii Coquerel, 1866: 316 (Philonthus; Type locality: La Réunion: Bourbon).
— Fauvel, 1867b: 66 (Philonthus; characters; Réunion).
— Bernhauer and Schubert, 1914: 337 (Philonthus; cited as fauveli; catalog).
— Jarrige, 1957: 110 (Gabrius; cited as fauveli; Réunion).
— Lecoq, 1990: 192 (Gabrius; cited as fauveli; Mascarene Islands).
▴ Distribution: Réunion.
colubrinus Fauvel, 1868a: 70 (Philonthus; Type locality: La Réunion).
— Fauvel, 1867b: 67 (Philonthus; synonym of fauvelii).
— Bernhauer and Schubert, 1914: 337 (Philonthus; synonym of fauvelii).
— Jarrige, 1957: 110 (Gabrius; synonym of fauvelii).
— Lecoq, 1990: 192 (Gabrius; synonym of fauvelii).
femineus Hatch, 1957, see: cushmani Hatch, 1957.
femoralis Hochhuth, 1851: 19 (Philonthus; Type locality: Kaukasien, Fluss Kupan; [Note: See lectotype designation by Gusarov, 1992a]).
— Fauvel, 1874: 261 [= 1874a: 487] (Philonthus; synonym of vernalis).
— Eppelsheim, 1892: 334 (Philonthus; Turkestan).
— Bernhauer and Schubert, 1914: 337 (Philonthus; catalog).
— Gridelli, 1920: 128 (Gabrius; characters; Austria; Bosnia; Italy).
— Gridelli, 1928: 45 (Gabrius; characters; notes; Poland, Austria; Croatia; Bosnia; Albania; Italy).
— Scheerpeltz, 1933: 1342 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Wüsthoff, 1937: 32 (Philonthus; illustration of aedeagus).
— Coiffait, 1951: 105, 113 (Gabrius; characters in key; France).
— Horion, 1956: 3 (Gabrius; characters; notes).
— Smetana, 1957: 58 (Gabrius; characters; Caucasus; Balkans; Italy; France; Spain; Austria; Czechoslovakia; Hungary; Germany).
— Smetana, 1958: 236 (Gabrius; characters; ecology; Czechoslovakia).
— Smetana, 1961: 37 (Gabrius; characters; Caucasus; Balkans; central Europe; Italy; France; Spain).
— Smetana, 1961a: 43 (Gabrius; Bulgaria).
— Lohse, 1964: 187 (Gabrius; characters; central Europe).
— Horion, 1965: 184 (Gabrius; Russia; Romania; Macedonia; Bosnia Hercegovina; Serbia; Dalmatia; Albania; Croatia; Istria; Slovenia; Italy; Switzerland; France; Spain ?; Hungary; Poland; Czech Republic; Slovakia; Austria; Germany).
— Smetana, 1966: 331 (Gabrius; notes; Czechoslovakia).
— Korge, 1971a: 7 (Gabrius; Turkey).
— Coiffait, 1974: 71 (Gabrius; characters; Italy; Balkans).
— Bordoni, 1974b: 9 (Gabrius; Italy).
— Smetana, 1977: 791 (Gabrius; Turkey).
— Petrenko, 1980: 82 (Gabrius; Caucasus).
— Tóth, 1984: 83 (Gabrius; characters; Hungary).
— Lucht, 1987: 106 (Gabrius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland).
— Gusarov, 1992a: 785 [= 1993a: 74] (Gabrius; lectotype designation).
— Smetana, 1993b: 49 (Gabrius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
▴ Distribution: Europe, Caucasus, Turkey, Uzbekistan.
fulvipes Motschulsky, 1858a: 211 (Othius; Type locality: Carniole).
— Bernhauer and Schubert, 1914: 337 (Philonthus; synonym of femoralis).
— Scheerpeltz, 1933: 1342 (Philonthus; synonym of femoralis).
— Smetana, 1958: 236 (Gabrius; synonym of femoralis).
— Coiffait, 1974: 71 (Gabrius; synonym of femoralis).
fenderi Hatch, 1957: 201 (Philonthus; subgenus Gabrius; Type locality: Oregon: McMinnville. Also cited from Washington and Idaho).
— Moore, 1965: 100 (Gabrius).
— Smetana, 1995e: 712 (Gabrius; characters; notes; California; Idaho; Oregon; Utah; Washington).
▴ Distribution: USA.
feretrius Smetana, 1995e: 715 (Gabrius; Type locality: California: Eureka).
fimbriolatus Erichson, 1840: 486 (Philonthus; Type locality: Madasgascar).
— Fauvel, 1898a: 118 (Philonthus; Seychelles).
— H. Kolbe, 1910: 20 (Philonthus; Seychelles).
— Bernhauer and Schubert, 1914: 338 (Philonthus; catalog).
— Bernhauer, 1931d: 587 (Philonthus; Abyssinia).
— Bernhauer and Chapman, 1939: 70 (Philonthus; Kenya).
— Tottenham, 1949b: 352 (Philonthus; characters).
— Tottenham, 1956b: 223 (Gabrius; may be restricted to Madagascar; characters).
— Lecoq, 1990: 192 (Gabrius; Mascarene Islands).
▴ Distribution: Ethiopia, Kenya, Seychelles, Madagascar, Mascarene Islands.
fimetarioides Scheerpeltz, 1976a: 125 (Philonthus; Type locality: Nepal: Khumjung, 3800 m).
— Coiffait, 1982: 28 (Philonthus; Nepal).
— Coiffait, 1982a: 232 (Philonthus; India).
— Schillhammer, 1992: 64 (Gabrius; characters; Nepal; India; Pakistan).
— Schillhammer, 1997: 10 (Gabrius; characters; Pakistan; India; Nepal; China).
— Schillhammer, 2000c: 118 (Gabrius; India; Nepal; Bhutan).
▴ Distribution: China, Bhutan, Nepal, India, Pakistan.
dorjulensis Coiffait, 1977b: 226 (Philonthus; Type locality: Bhutan: Dorjula, 3100 m).
fissicauda Tottenham, 1962: 242 (Gabrius; Type locality: Kilimanjaro, Bismarck Hügel, 3100 m).
fissiceps Schillhammer, 2000c: 133 (Gabrius; Type locality: Philippines: Mindanao, Seliban Riv., Mt. Apo, alt. 7000 ft.).
flavimanus Gemminger and Harold, 1868, see: osseticus Kolenati, 1846.
flavipes Motschulsky, 1860, see: osseticus Kolenati, 1846.
flexuellus Tottenham, 1956b: 218 (Gabrius; Type locality: N. W. Rhodesia: Mamwala).
forcipatus Hatch, 1957: 202 (Philonthus; subgenus Gabrius; Type locality: British Columbia: Needles. Also cited from Washington, Oregon, and Idaho).
— Smetana, 1995e: 716 (Gabrius; characters; notes; British Columbia; California; Idaho; Oregon; Washington).
▴ Distribution: Canada, USA.
formosus Motschulsky, 1860, see: Rabigus.
franzi Smetana, 1967b: 153 (Gabrius; Type locality: Hisp. bor., Prov. Lugo, Valle de Lozera).
fulvipes Motschulsky, 1858, see: femoralis Hochhuth, 1851.
furtivus Cameron, 1932: 149 (Philonthus; Type locality: India, Uttar Pradesh, Chakrata Dist., Chulli Khud, 8000′; [Note: See lectotype designation by Schillhammer, 1997]. Originally cited from Chakrata, Mussoorie, and Simla districts, alt. 6000–8000 feet).
— Scheerpeltz, 1933: 1343 (Philonthus; catalog).
— Schillhammer, 1997: 27 (Gabrius; lectotype designation: India, Uttar Pradesh, Chakrata Dist., Chulli Khud, 8000′; characters; India; Pakistan; Afghanistan).
— Schillhammer, 2000c: 128 (Gabrius; Nepal).
▴ Distribution: Afghanistan, Pakistan, India, Nepal.
ajmonis Gridelli, 1934: 80 (Gabrius; Type locality: Valle del Braldo: Chokpiong, 2600 m).
afghanicus Coiffait, 1979d: 556 (Gabrius; Type locality: Nouristan: entre Camdech et Kouchtous, 1400–2000 m).
schawalleri Coiffait, 1982: 55 (Gabrius; Type locality: Inde: Cachemire, Gulmarg, Pir Panjal Gebirge, 2800 m).
fuscolaterus Motschulsky, 1860, see: Gabronthus.
fuscus Last, 1989, see: Philonthus.
gabrieli Schillhammer, 1997: 58 (Gabrius; Type locality: Nepal: [Sankhua Sabha District]: Kosi, Forêt S. Mangsingma, 2250 m).
gabrioides Bernhauer, 1913: 131 (Philonthus; Type locality: Ostsibirien: Schipka-Gora, Sotka-Gora).
— Bernhauer and Schubert, 1914: 340 (Philonthus; catalog).
— Scheerpeltz, 1933: 1344 (Philonthus; catalog).
— Schillhammer, 1997: 77 (Gabrius).
— Schillhammer, 2000c: 125 (Gabrius; characters; Primorskiy Kray).
▴ Distribution: Russia.
gagliardii Gridelli, 1936a: 153 (Gabrius; Type locality: Viareggio sul litorale della Pineta Reale).
— Smetana, 1957: 64 (Gabrius; characters; Italy).
— Smetana, 1960c: 317 (Gabrius; characters; Italy).
— Smetana, 1967b: 158 (Gabrius; characters).
— Coiffait, 1974: 64 (Gabrius; cited as gagliardi; characters; Italy).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
▴ Distribution: Italy.
gavoyi Bernhauer, 1928, see: pisciformis Fauvel, 1874.
gedyei Cameron, 1951, see: marakwetensis Bernhauer, 1939.
gelo Smetana, 1984a: 649 (Gabrius; Type locality: Japon: Gunma Nikko Nat. Park ss/Konsei Pass, 1500 m).
gladipenis Cho and Lee, 1997: 275 (Gabrius; Type locality: Korea: Dongcheonri, Yeanmyon, Andong, Kyǔngsangpukdo).
gomyi Lecoq, 1990: 194 (Gabrius; Type locality: La Réunion: Plaines des Chicots, près du gîte, 1850 m).
graecus Bordoni, 1994: 31 (Gabrius; Type locality: Grecia: Patras).
grassei Levasseur, 1980: 352 (Gabrius; Type locality: Gabon: Belinga).
gratellus Cameron, 1932: 146 (Philonthus; Type locality: India, Uttar Pradesh, Chakrata Dist., Dodora Khud, 8000′; [Note: See lectotype designation by Schillhammer, 1997]. Originally cited from Mussoorie, Chakrata, and Simla districts, alt. 6000–8000 feet).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Coiffait, 1982: 27 (Gabrius; characters; India).
— Schillhammer, 1997: 33 (Gabrius; lectotype designation: India, Uttar Pradesh, Chakrata Dist., Dodora Khud, 8000′; characters; India).
▴ Distribution: India.
gunungapi Schillhammer, 1997: 70 (Gabrius; Type locality: Sumatra: Jambi Mt. Kerinci, 3300 m).
habilis Tottenham, 1956b: 221 (Gabrius; Type locality: Cape Colony: Pondoland, Port St. John).
halisius Tottenham, 1956b: 209 (Gabrius; Type locality: Kenya: Muguga, near Kikuyu).
hammondi Schillhammer, 1997: 72 (Gabrius; Type locality: Indonesia: Sulawesi Utara Gng. Ambang F.R. nr. Kotamobagu, 1400–1600 m).
haptsui Schillhammer, 2000c: 129 (Gabrius; Type locality: Nepal: Kaski Distr., about Pothana, 2000 m).
hartmanni Schillhammer, 2000c: 119 (Gabrius; Type locality: Nepal: Prov. Karnali, Distr. Jumla, 2 km E Churta, 3100 m).
hazarahensis Tronquet, 1981: 79 (Gabrius; Type locality: Afghanistan: massif du Kakrak, Koh-I-Baba, 2600 m).
— Schillhammer, 1997: 29 (Gabrius; characters; Afghanistan; Pakistan).
▴ Distribution: Afghanistan, Pakistan.
hercules Schillhammer, 1997: 67 (Gabrius; Type locality: Indonesia: N-Sumatra: Toba, Samosir, Tomok).
heres Smetana, 1962: 100 (Gabrius; Type locality: Kanarische Inseln: Gran Canaria).
hesperius Fauvel, 1878: 124 [= 1878a: 44] (Philonthus; Type locality: Edough, près Bône; Espagne: Andalousie, Algésiras; Portugal).
— Duvivier, 1883: 146 (Philonthus; cited as hesperinus; [Note: Misspelling: Bernhauer and Schubert, 1914: 341 cited hesperinus Duvivier as a synonym of hesperius Fauvel; Duvivier did not propose an emendation or replacement name, he misspelled herperius]; catalog).
— Fauvel, 1886: 55 [= 1886a: 47] (Philonthus; Algeria).
— Fauvel, 1902b: 111 (Philonthus; Morocco; Algeria; Tunisia).
— Bernhauer and Schubert, 1914: 341 (Philonthus; catalog).
— Gridelli, 1920: 130 (Gabrius; characters; Algeria; Spain; Portugal).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Jeannel and Jarrige, 1949: 376 (Philonthus; subgenus Gabrius; cave species; Algeria).
— Smetana, 1957: 57 (Gabrius; characters; Algeria; Tunisia; Morocco; Spain).
— Smetana, 1961: 36 (Gabrius; characters; Algeria; Tunisia; Morocco; Spain).
— Coiffait, 1964c: 167 (Gabrius; characters; Algeria; Tunisia).
— Coiffait, 1974: 72 (Gabrius; characters; Algeria; Tunisia).
▴ Distribution: Tunisia, Algeria, Morocco, Spain, Portugal.
heterocerus Cameron, 1950g: 42 (Philonthus; subgenus Gabrius; Type locality: Belgian Congo: Rweru, volc. Mikeno, alt. 2400 m).
himalayicus Bernhauer, 1915b: 52 (Philonthus; Type locality: Zentral-Himalaya: Kulu, 10,000 Fuss).
— Cameron, 1932: 144 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Schillhammer, 1997: 26 (Gabrius; characters; India).
▴ Distribution: India.
hippisleyae Smetana, 1995e: 661 (Gabrius; Type locality: British Columbia: Terrace. Also cited from Alaska and Washington).
— Hatch, 1957: 200 (Philonthus; subgenus Gabrius; [Note: Misidentified as microphthalmus according to Smetana, 1995e: 661]; characters; British Columbia; Washington).
▴ Distribution: Canada, USA.
hirthei Schillhammer, 2000c: 124 (Gabrius; Type locality: Nepal: Annapurna Mts., Larjung, 2550–2800 m).
hoberlandti Smetana, 1953: 122 (Gabrius; Type locality: Anatolia: Ulukisla).
— Smetana, 1960c: 329 (Gabrius; characters; Turkey; Bulgaria).
— Smetana, 1961a: 45 (Gabrius; notes; Bulgaria).
— Coiffait, 1974: 62 (Gabrius; characters; Asia Minor; Bulgaria).
▴ Distribution: Turkey, Bulgaria.
hoggarensis Coiffait, 1966b: 343 (Gabrius; subspecies of nigritulus; Type locality: Hoggar, Tamanrasset).
— Coiffait, 1966c: 506 (Gabrius; subspecies of nigritulus; [Note: Cited here as a new species but described in earlier paper in same journal]; Hoggar).
— Coiffait, 1974: 54 (Gabrius; characters; subspecies of nigritulus).
— Schillhammer, 1996a: 81 (Gabrius; valid species; France).
▴ Distribution: Algeria, France.
krampfbiegeri Schillhammer, 1990: 89 (Gabrius; Type locality: Algerien: Bou Saada).
holisinus Fauvel, 1904d: 288 (Belonuchus; Type locality: Usambara: Derema. Gabon: Libreville; Loango; Monts de Cristal. Loanda: Golungo Alto. Natal).
— Eichelbaum, 1913: 130 (Belonuchus; cited as holosinus; Tanzania).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog).
— Levasseur, 1968: 1384 (Gabrius).
▴ Distribution: South Africa, Angola, Tanzania, Congo Republic, Gabon, Ivory Coast.
pujoli Levasseur, 1967: 653 (Gabrius; Type locality: Côte-d'Ivoire: Eloka).
horni Bernhauer and Schubert, 1914, see: eremius Herman, 2001.
hublei Coiffait and Segers, 1985, see: breviventer Sperk, 1835.
huendus Tottenham, 1956b: 215 (Gabrius; Type locality: Transvaal: Rosettenville).
humidulus Cameron, 1932: 147 (Philonthus; Type locality: India, Uttar Pradesh, Chakrata Distr., Manjgaon, 6500′; [Note: See lectotype designation by Schillhammer, 1997]. Originally cited from Chakrata district: Manjgaon and Chulli Khuds, Mohna and Bangar, alt. 6000–8000 feet).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Schillhammer, 1997: 30 (Gabrius; lectotype designation: India, Uttar Pradesh, Chakrata Distr., Manjgaon, 6500′; characters; India).
▴ Distribution: India.
hyas Smetana, 1995e: 636 (Gabrius; Type locality: Texas: Chisos Mts.).
hydrophilus Schillhammer, 1997: 66 (Gabrius; Type locality: Indonesia: N-Sumatra: Toba, Samosir, Tomok).
ignobilis Cameron, 1943: 132 (Philonthus; subgenus Gabrius; Type locality: Kashmir: Gulmarg, altitude 8000–10,000 feet; [Note: See lectotype designation by Schillhammer, 1997]).
— Schillhammer, 1997: 16 (Gabrius; lectotype designation; characters; India).
— Schillhammer, 2000c: 120 (Gabrius; India).
▴ Distribution: India.
ladakhensis Coiffait, 1982: 54 (Gabrius; Type locality: Inde: Ladakh, Kargil, 2950 m).
imitator Fauvel, 1895: 265 (Philonthus; Type locality: Carin Asciuü Checù, 1400–1500 m; [Note: See lectotype designation by Schillhammer, 1997]. Originally cited from Bírmanie, Carin Asciuii Ghécu, 1400–1500 m; Sikkim, Darjeeling).
— Bernhauer and Schubert, 1914: 341 (Philonthus; catalog).
— Cameron, 1932: 149 (Philonthus; characters; Burma; India).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Coiffait, 1977b: 207 (Philonthus; Bhutan).
— Legner and Moore, 1977: 176 (Philonthus; found under bark).
— Coiffait, 1982: 27 (Gabrius; characters; Nepal).
— Schillhammer, 1997: 45 (Gabrius; lectotype designation: Carin Asciuü Checù, 1400–1500 m; characters; Burma; Thailand).
▴ Distribution: Thailand, Myanmar.
immutatus Motschulsky, 1860, see: Philonthus.
incubens Schillhammer, 1991: 53 (Gabrius; Type locality: USSR: Tschita, Transbaikalien).
ingratus Fauvel, 1905b: 179 (Philonthus; Type locality: forêt de Moramanga; Tananarive).
— Bernhauer and Schubert, 1914: 342 (Philonthus; catalog).
— Schillhammer, 1997: 78 (Gabrius).
▴ Distribution: Madagascar.
insidiosus Tottenham, 1962: 240 (Gabrius; Type locality: Tanganyika Terr.: Ngorongoro, 2300 m).
insignis Luze, 1904: 98 (Philonthus; subgenus Gabrius; Type locality: Seravschan: Kschtut: Artutsch, Putschin-Pass, Kumar, Postigau, Urmitan; Varsaut; Iskander-kul).
— Bernhauer and Schubert, 1914: 342 (Philonthus; catalog).
— Gridelli, 1928: 49 (Gabrius; characters; notes; Turkestan).
— Scheerpeltz, 1933: 1346 (Philonthus; catalog).
— Smetana, 1953a: 178 (Gabrius; Caucasus).
— Smetana, 1961: 37 (Gabrius; characters; Turkestan; Uzbekistan; Caucasus).
— Coiffait, 1970d: 143 (Gabrius; Uzbekistan).
— Coiffait, 1974: 70 (Gabrius; characters; Turkestan; Uzbekistan; Caucasus).
— Tronquet, 1981: 70 (Gabrius; Afghanistan).
▴ Distribution: Caucasus, Afghanistan, Tajikistan, Uzbekistan.
insularis Gautier, 1868, see: badius Kiesenwetter, 1858.
interceptionis Schillhammer, 1997: 71 (Gabrius; Type locality: Indonesia: S-Sulawesi, E. Malino, Kanre Apia, 1400 m).
io Smetana, 1984: 133 (Gabrius; Type locality: Japan: Kyoto Pref.: Seryo-toge, 500–700 m).
jacksoni Cameron, 1951c: 401 (Philonthus; Type locality: Uganda: Kalinzu F., 4000 ft.).
— Tottenham, 1955: 173 (Philonthus; characters).
— Schillhammer, 1997: 78 (Gabrius).
▴ Distribution: Uganda.
jaechi Schillhammer, 1997: 50 (Gabrius; Type locality: Indonesia: Java centr.: Dieng Plateau).
japonicus Shibata, 1991a: 90 (Gabrius; Type locality: Japan: Kanagawa Pref.: near Bodai, Tanzawa).
javanicus Schillhammer, 1997: 65 (Gabrius; Type locality: Indonesia: Java: S. Bogor, Cibodas).
jeannelianus Bernhauer, 1939a: 87 (Philonthus; Type locality: British-Ostafrika: Mt. Kenya, Westhang, obere Waldzone, 3200–3300 m).
jendeki Schillhammer, 1997: 12 (Gabrius; Type locality: China: Yunnan, Lijiang, Yulongshan Nat. Res.).
kabisus Tottenham, 1956: 301 (Gabrius; Type locality: Urundi: terr. Bururi, Nyamurembe, 900 m).
kabulensis Tronquet, 1981: 80 (Gabrius; Type locality: Afghanistan: Sairobi).
— Schillhammer, 1997: 14 (Gabrius; characters; Afghanistan; Pakistan; India).
▴ Distribution: Afghanistan, Pakistan, India.
kahleni Schillhammer, 1992: 61 (Gabrius; Type locality: N. Italy: Südtirol, Kalterer See, 230 m).
kainpitensis Last, 1989, see: Philonthus.
kambaitiensis Scheerpeltz, 1965: 228 (Gabrius; Type locality: N.E. Burma, Kambaiti, 7000 ft.).
katonae Bernhauer, 1915d: 143 (Philonthus; subgenus Gabrius; Type locality: Arusha-Ju; Kibosho; West-Usambara).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
— Bernhauer and Chapman, 1939: 66 (Philonthus; Kenya).
— Tottenham, 1962: 142, 252 (Gabrius; specimens of type series represent several species; Tanganyika).
▴ Distribution: Kenya, Tanzania.
tenganus Tottenham, 1956b: 212 (Gabrius; Type locality: Kenya: Muguga, near Kikuyu).
kerinciensis Schillhammer, 1997: 69 (Gabrius; Type locality: Indonesia: Sumatra: Jambi Mt. Kerinci, 2100 m).
kessleri Blatchley, 1936, see: ovaliceps Fall, 1930.
keysianus Sharp, 1910: 130 (Gabrius; Type locality: Anglia: Devon, Slapton Ley).
— Bernhauer and Schubert, 1914: 343 (Philonthus; catalog).
— Gridelli, 1920: 145 (Gabrius; characters; Sardinia).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
— Fagel, 1945: 42 (Gabrius; characters; Belgium).
— Coiffait, 1951: 111, 114 (Gabrius; characters in key; France).
— V. Hansen, 1952: 91 (Philonthus; subgenus Gabrius; characters; Denmark).
— Smetana, 1952b: 117 (Gabrius; characters; Czechoslovakia).
— Smetana, 1957: 64, 69 (Gabrius; characters; England; Scandinavia; Holland; Belgium; France; Italy; Sardinia; Germany).
— Smetana, 1960c: 313 (Gabrius; characters; England; Sweden; Denmark; Germany; Holland; Belgium; France; Sardinia).
— Palm, 1963b: 107 (Philonthus; subgenus Gabrius; characters).
— Lohse, 1964: 189 (Gabrius; characters; central Europe).
— Horion, 1965: 191 (Gabrius; Denmark; Britain; Holland; Belgium; France; Sardinia; Germany).
— Bordoni, 1973i: 327 (Gabrius; characters).
— Coiffait, 1974: 63 (Gabrius; characters; coastal region of Europe, Britain, Sicily, Sardinia).
— Pope, 1977: 30 (Gabrius; Britain).
— Lucht, 1987: 106 (Gabrius; checklist; Germany; Benelux; Denmark).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy ?).
— Luff, 1996: 30 (Gabrius; collecting notes; Isle of Man).
▴ Distribution: Europe.
kivulius Tottenham, 1956: 300 (Gabrius; Type locality: Ruanda: terr. Nyanza, Gitarama, 1850 m).
kleebergi Schillhammer, 1997: 58 (Gabrius; Type locality: Ost-Nepal Umg.: [Solukhumbu District]: Sete Lamjura Pass).
kobayashii Smetana, 1984: 128 (Gabrius; Type locality: Japan: Gumma Pref.: 7 km E Usui Pass, 850 m).
krampfbiegeri Schillhammer, 1990, see: hoggarensis Coiffait, 1966.
kuanshanensis Bernhauer, 1914b: 66 (Philonthus; subgenus Gabrius; Type locality: Südliche Mandschurei).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
— Smetana, 1973b: 132 (Gabrius; characters; notes).
— Coiffait, 1974: 79 (Gabrius; characters; Manchuria).
— Smetana, 1984: 142 (Gabrius; characters; Japan).
— J. Li, 1993: 38 (Gabrius; China).
▴ Distribution: China, Japan.
kuliabensis Bernhauer, 1915c: 71 (Philonthus; subgenus Gabrius; Type locality: Buchara: Provinz Kuljab, Ak-son-Tal).
— Gridelli, 1920: 141 (Gabrius; characters).
— Scheerpeltz, 1933: 1347 (Philonthus; cited kuljabensis; catalog).
— Smetana, 1973b: 135 (Gabrius; characters; notes).
— Coiffait, 1974: 80 (Gabrius; characters; Transcaspian region).
— Tronquet, 1981: 70 (Gabrius; Afghanistan).
▴ Distribution: Afghanistan, “Transcaspian region”, Uzbekistan.
laciades Smetana, 1995e: 689 (Gabrius; Type locality: California: Mt. Shasta, McBride Springs, 4750 ft. Also cited from Oregon).
ladakhensis Coiffait, 1982, see: ignobilis Cameron, 1943.
laelaps Smetana, 1995e: 647 (Gabrius; Type locality: British Columbia: Herald Provincial Park, Reienecker Creek, below Reienecker Falls, 380–410 m).
lapidicola Cameron, 1932: 145 (Philonthus; Type locality: Kashmir: Gulmarg, Ferozepur Nala, alt. 6500 feet).
— Scheerpeltz, 1933: 1348 (Philonthus; catalog).
— Coiffait, 1982: 27 (Gabrius; India).
— Schillhammer, 1997: 26 (Gabrius; characters; India; Pakistan; Afghanistan).
▴ Distribution: India, Pakistan, Afghanistan.
laticollis Fauvel, 1874: 262 [= 1874: 488] (Philonthus; Type locality: Tarbes; Espagne).
— Fauvel, 1876a: 75 [= 1876: 260] (Philonthus; Sicily).
— Mulsant and Rey, 1876: 557 [= 1877: 413] (Gabrius; characters).
— Bernhauer and Schubert, 1914: 343 (Philonthus; catalog).
— Gridelli, 1920: 132 (Gabrius; characters; Spain; France).
— Portevin, 1929: 366 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1348 (Philonthus; catalog).
— Coiffait, 1951: 110, 114 (Gabrius; characters in key; France).
— Fagel, 1959a: 96 (Gabrius; Spain).
— Smetana, 1967b: 153 (Gabrius; Spain).
— Coiffait, 1974: 77 (Gabrius; characters; Pyrenees).
▴ Distribution: France, Spain.
latro Joy, 1913: 26 (Gabrius; Type locality: Caucasus, Lenkoran).
— Gridelli, 1920: 144 (Gabrius; characters; Caucasus).
— Scheerpeltz, 1933: 1348 (Philonthus; catalog).
— Smetana, 1957: 67 (Gabrius; characters; Caucasus; Greece; France; southern Europe).
— Smetana, 1960c: 321 (Gabrius; characters; Caucasus; Greece; France; Bulgaria).
— Smetana, 1961a: 44 (Gabrius; Bulgaria).
— Lohse, 1964: 190 (Gabrius; characters; central Europe).
— Coiffait, 1974: 56 (Gabrius; characters; Caucasus; Lebanon; Balkans; Maritime Alps).
— Bordoni, 1975a: 154 (Gabrius; characters; Afghanistan).
— Smetana, 1977: 794 (Gabrius; Turkey).
— Smetana, 1978: 116 (Gabrius; Israel).
— Tóth, 1984: 91 (Gabrius; characters).
— Schillhammer, 1997: 14 (Gabrius; characters; notes; Afghanistan; Pakistan).
▴ Distribution: Europe, Armenia, Azerbaijan, Turkey, Syria, Lebanon, Israel, Afghanistan, Pakistan.
latroides Coiffait, 1963: 21 (Gabrius; Type locality: Syrie: environs de Damas).
— Coiffait, 1974: 56 (Gabrius; characters; Syria; Turkey; Armenia).
— Schillhammer, 1992: 66 (Gabrius; synonym of latro).
— Schillhammer, 1997: 14 (Gabrius; synonym of latro).
latroides Coiffait, 1963, see: latro Joy, 1913. lebedevi Bernhauer, 1910a: 72 (Philonthus; subgenus Gabrius; Type locality: Ala-Tau, Syr Darja).
— Bernhauer and Schubert, 1914: 343 (Philonthus; catalog).
— Smetana, 1973b: 134 (Gabrius; characters; notes).
— Coiffait, 1974: 79 (Gabrius; characters; Turkestan; Uzbekistan).
▴ Distribution: “Turkestan”, Uzbekistan.
tschinganus Coiffait, 1970d: 145 (Gabrius; subspecies of lebedevi; Type locality: Ouzbekistan, Monts Tschingan, 1500 m environ).
leleupi Tottenham, 1962: 238 (Gabrius; Type locality: Tanganyika Terr.: Mt. Hanang, versant Sud, 2400–2600 m).
lemur Jarrige, 1978: 286 (Gabrius; Type locality: Andringitra Centre, plateau Andohariana, 2000–2100 m).
lethifer Tottenham, 1956b: 215 (Gabrius; Type locality: Kenya: Muguga near Kikuyu).
linauensis Schillhammer, 2000c: 136 (Gabrius; Type locality: Philippines: Davao Prov.: Lake Linau, N. Slope Mt. Apo, 7900′).
linearis Casey, 1915, see: shulli Hatch, 1957.
linearis Schillhammer, 2000c: 138 (Gabrius; [preoccupied]; Type locality: Philippines: Luzon: Balbalan).
lividipes Baudi, 1848: 129 (Philonthus; Type locality: Sardinia).
— Kraatz, 1857b: 176 (Philonthus; synonym of rubripennis).
— Kraatz, 1857d: 1060 (Philonthus; synonym of rubripennis).
— Fauvel, 1874: 242 [= 1874a: 468] (Philonthus; synonym of rubripennis).
— Mulsant and Rey, 1876: 512 [= 1877: 368] (Philonthus; synonym of rubripennis).
— Ganglbauer, 1895: 461 (Philonthus; synonym of rubripennis).
— Bernhauer and Schubert, 1914: 344 (Philonthus; catalog).
— Gridelli, 1920: 141 (Gabrius; characters; Italy; Sardinia).
— Scheerpeltz, 1933: 1349 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Coiffait, 1951: 109, 114 (Gabrius; characters in key; France).
— V. Hansen, 1952: 88 (Philonthus; subgenus Gabrius; characters; Denmark).
— Smetana, 1953: 121 (Gabrius; Turkey).
— Smetana, 1953a: 178 (Gabrius; Carpathian region; Moravia; Bohemia).
— Smetana, 1957: 61 (Gabrius; characters; England; Caucasus; Turkey; Germany; Czechoslovakia; Austria).
— Smetana, 1958: 240 (Gabrius; characters; ecology; Czechoslovakia).
— Palm, 1963b: 104 (Philonthus; subgenus Gabrius; characters; Finland).
— Lohse, 1964: 188 (Gabrius; characters; central Europe).
— Smetana, 1964c: 168 (Gabrius; Czechoslovakia).
— Horion, 1965: 188 (Gabrius; Russia; Finland; Georgia; Ukraine; Poland; Czechoslovakia; Balkans; Italy; Spain; Portugal; France; Switzerland; Belgium ?; Holland; Austria; Germany).
— Coiffait, 1974: 80 (Gabrius; characters; Europe; Britain; Scandinavia; Finland; Asia Minor; Caucasus; France).
— Tóth, 1984: 85 (Gabrius; characters; Hungary).
— Lundberg, 1986: 148 (Gabrius; Sweden).
— Schülke and Uhlig, 1988: 11 (Gabrius; notes; Germany).
— Lucht, 1987: 106 (Gabrius; checklist; Germany; Poland; Czechoslovakia; Austria; France; Benelux).
— Wittwer, 1993: 250 (Gabrius; Switzerland).
— Smetana, 1993b: 49 (Gabrius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
— Schülke, 1998g: 90 (Gabrius; Germany).
▴ Distribution: Europe, Caucasus, Turkey.
rubripennis Kiesenwetter, 1844: 346 (Philonthus; [preoccupied]; Type locality: Leipzig).
— Kiesenwetter, 1848a: 24 (Philonthus; characters).
— Redtenbacher, 1849: 825 (Philonthus; characters).
— Fairmaire and Laboulbène, 1856: 529 (Philonthus; characters; France).
— Kraatz, 1857d: 612 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 198 (Philonthus; characters; Germany).
— Fauvel, 1874: 241 [= 1874a: 467] (Philonthus; characters; France; Germany; Italy; Sardinia).
— Redtenbacher, 1874: 217 (Philonthus; characters; Germany).
— Mulsant and Rey, 1876: 512 [= 1877: 368] (Philonthus; characters; France).
— Ganglbauer, 1895: 461 (Philonthus; subgenus Gabrius; characters; Germany; France; Piemont; Sardinia; Caucasus).
— Reitter, 1909: 132 (Philonthus; subgenus Gabrius; characters; Germany).
— Bernhauer and Schubert, 1914: 344 (Philonthus; synonym of lividipes).
— Portevin, 1929: 366 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1349 (Philonthus; synonym of lividipes).
— Wüsthoff, 1934: 61 (Philonthus; synonym of lividipes).
— Smetana, 1957: 61 (Gabrius; synonym of lividipes).
— Smetana, 1958: 240 (Gabrius; synonym of lividipes).
— Lohse, 1964: 188 (Gabrius; synonym of lividipes).
— Coiffait, 1974: 80 (Gabrius; synonym of lividipes).
loebli Schillhammer, 1997: 45 (Gabrius; Type locality: India: Uttar Pradesh, Garhwal, 4 km au Sud de Bhatwari, 1400 m. Also cited from Nepal).
lomaensis Levasseur, 1971: 299 (Gabrius; Type locality: monts Loma, Sierra Leone, 1600 m).
londianus Cameron, 1951, see: marakwetensis Bernhauer, 1939.
longulus Motschulsky, 1858, see: Bisnius.
lopezi Hatch, 1957, see: picipennis Mäklin, 1852.
lysippus Smetana, 1995e: 651 (Gabrius; Type locality: Quebec: Meach L., Gatineau Park. Also cited from New Hampshire, New York).
macer Lecoq, 1990: 193 (Gabrius; Type locality: La Réunion: Plaine des Cafres, Notre Dame de la Paix, 1700 m).
maderensis Coiffait, 1963, see: simulans Wollaston, 1857.
magniceps Cameron, 1948a: 229 (Philonthus; subgenus Gabrius; Type locality: Ivory Coast: Reserve du Banco).
magniceps Smetana, 1952, see: nigritulus Gravenhorst, 1802.
magnicollis Bernhauer, 1915d: 141 (Philonthus; Type locality: Kilimandscharo).
— Scheerpeltz, 1933: 1350 (Philonthus; catalog).
— Schillhammer, 1997: 79 (Gabrius).
▴ Distribution: Tanzania.
malaysianus Schillhammer, 1997: 49 (Gabrius; Type locality: Malaysia: Cameron Highlands).
malickyi Schillhammer, 2000c: 128 (Gabrius; Type locality: Nepal: Godawri [Godavari], 1700 m, 27°35′, 85°23′).
malkini Hatch, 1957, see: brevipennis Horn, 1884.
mandli Schillhammer, 1992: 62 (Gabrius; Type locality: USSR: Werchne-Udinsk, Transbaikal).
mandschuricus Bernhauer, 1914b: 66 (Philonthus; subgenus Gabrius; Type locality: Südliche Mandschurei: Chi-Kuan-Shan).
— Gridelli, 1920: 128 (Gabrius; characters; Manchuria).
— Scheerpeltz, 1933: 1350 (Philonthus; catalog).
— Smetana, 1953a: 171 (Gabrius; characters; eastern Siberia).
— Smetana, 1961: 38 (Gabrius; characters; Manchuria).
— Smetana, 1973b: 126 (Gabrius; notes).
— Coiffait, 1974: 70 (Gabrius; characters; Siberia; Manchuria).
— J. Li, 1993: 38 (Gabrius; China).
▴ Distribution: Russia, China.
marakwetensis Bernhauer, 1939a: 88 (Philonthus; Type locality: Kenya: Marakwet, Campi Cherangani, 3000 m).
gedyei Cameron, 1951c: 403 (Philonthus; Type locality: Kenya Colony: Kinangop, 11,000 feet).
londianus Cameron, 1951c: 403 (Philonthus; Type locality: Kenya Colony: Londiani).
maritimus Motschulsky, 1858, see: Gabronthus.
marshalli Cameron, 1951c: 402 (Philonthus; [preoccupied]; Type locality: South Africa: Natal, Frere).
— Schillhammer, 1997: 79 (Gabrius).
— Herman, 2001: 44 (Gabrius; may be conspecific with senior homonym so use of junior homonym maintained pending further study).
▴ Distribution: South Africa.
martensi Schillhammer, 1997: 41 (Gabrius; Type locality: Nepal: 374 Taplejung Distr., upper Tamur Valley, from Lungthung waterfall to bamboo brige, 1800–2150 m).
martialis Cameron, 1932: 431 (Philonthus; Type locality: Waziristan).
— Scheerpeltz, 1933: 1350 (Philonthus; catalog).
— Schillhammer, 1997: 28 (Gabrius; characters; Pakistan).
▴ Distribution: Pakistan.
martialoides Schillhammer, 1997: 28 (Gabrius; Type locality: NE-Afghanistan: Qulatak, 1950 m, Salangtal).
matatielensis Scheerpeltz, 1974: 163 (Gabrius; Type locality: Cape Prov.: Border, 20 miles N Natatiele).
mauretanicus Peyerimhoff, 1937, see: primigenius Joy, 1913.
mcneilli Schillhammer, 1997: 11 (Gabrius; Type locality: Nepal: [Solukhumbu District], Ost-Nepal Umg. Junbesi Tognosa, 3800 m).
mediocris Cameron, 1943: 129 (Philonthus; Type locality: Kashmir: Gulmarg; [Note: See lectotype designation by Schillhammer, 1997]).
meghalayensis Schillhammer, 1997: 44 (Gabrius; Type locality: India: Meghalaya Khasi Hills, Cherrapunjee, 1200 m).
megophthalmus Cameron, 1948a: 229 (Philonthus; Type locality: Ivory Coast: Reserve du Banco).
mendosus Bernhauer, 1928, see: astutus Erichson, 1840.
mengensis Smetana, 1967b: 157 (Gabrius; Type locality: Hisp. bor., Prov. Avila, Puerto de Menga, cca 1500 m).
meridies Smetana, 1995e: 668 (Gabrius; Type locality: Arizona: Graham Co., Pinaleno Mts., Post Ck., 9000 ft. Also cited from Colorado, New Mexico, Utah, Wyoming).
metagenes Tottenham, 1956: 300 (Gabrius; Type locality: Urundi: Bururi, 1800–2000 m).
methalius Tottenham, 1956: 295 (Gabrius; Type locality: Ruanda: Tshuruyaga, forêt du Rugege, 2400 m).
microphthalmus Horn, 1884: 216 (Philonthus; Type locality: N.J. [New Jersey]; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from Vancouver eastward to the New England States, and as far south as Virginia).
— Blatchley, 1910: 388 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 346 (Philonthus; catalog).
— Fall, 1930: 252, 253 (Philonthus; characters; differentiation from nigritulus; New Hampshire, Massachusetts, Michigan, Illinois, British Columbia).
— Scheerpeltz, 1933: 1351 (Philonthus; catalog).
— Moore, 1965: 100 (Gabrius).
— Frank, 1982: 42 (Philonthus; list of parasites).
— Smetana, 1995e: 663 (Gabrius; lectotype designation: N.J. [New Jersey]; characters; notes; Manitoba; New Brunswick; Nova Scotia; Ontario; Quebec; Connecticut; Indiana; Iowa; Maine; Massachusetts; Michigan; New Hampshire; New Jersey; New York; North Carolina; Ohio; Pennsylvania; Rhode Island; South Dakota; Vermont; Virginia; West Virginia; Wisconsin).
— Levesque and Levesque, 1996: 292 (Gabrius; seasonal abundance; Quebec).
— Downie and Arnett, 1996: 383 (Gabrius; characters; USA).
▴ Distribution: Canada, USA.
miltiades Smetana, 1995, see: punctatellus Horn, 1884.
minahasa Schillhammer, 1997: 73 (Gabrius; Type locality: Indonesia: Sulawesi Utara, Gng. Ambang F.R. nr. Kotamobagu, 1300 m).
minax Tottenham, 1962: 245 (Gabrius; Type locality: Tanganyika Terr.: Mt. Oldeani, versant Est, 2300 m).
mirei Levasseur, 1967a: 955 (Gabrius; Type locality: mont Cameroun, forêt Antoir, point 1,200 m).
molliculus Casey, 1915, see: distans Horn, 1884.
montanus Bernhauer, 1934g: 237 (Philonthus; [preoccupied]; Type locality: Ruwenzori: Kalonge, 2050 m; Kivu: Mts. Nyamukubi, 2600 m).
— Tottenham, 1956b: 207 (Gabrius; characters).
— Herman, 2001: 44 (Gabrius; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Congo.
monticola Bernhauer, 1931d: 586 (Philonthus; subgenus Gabrius; Type locality: Abyssinia: Mt. Zukwala, ca. 9000 ft. and 9600 ft.; Mt. Chillalo, ca. 9000 ft.; Jem-Jem Forest, ca. 9,000 ft.).
— Scheerpeltz, 1933: 1351 (Philonthus; catalog).
— Bernhauer and Chapman, 1939: 71 (Philonthus; Kenya).
— Tottenham, 1949b: 362 (Gabrius; characters).
— Tottenham, 1956: 305 (Gabrius; characters in key).
— Tottenham, 1956b: 206, 207, 211 (Gabrius; characters).
▴ Distribution: Ethiopia, Kenya.
morator Schillhammer, 1997: 68 (Gabrius; Type locality: Indonesia: W-Sumatra, Singgalang, 2100–2800 m).
mubilibanus Cameron, 1950g: 42 (Philonthus; subgenus Gabrius; Type locality: Belgian Congo: Mubiliba, volc. Nyamuragira, alt. 2000 m).
muelleri Gridelli, 1928: 47 (Gabrius; Type locality: Lenkoran, Caucaso).
— Scheerpeltz, 1933: 1352 (Philonthus; catalog).
— Smetana, 1961: 37 (Gabrius; characters; Daghestan; Caspian region).
— Coiffait, 1974: 69 (Gabrius; characters; Daghestan; Transcaspian region).
▴ Distribution: Russia, Azerbaijan, “Transcaspian region”.
musonoiensis Levasseur, 1967: 655 (Gabrius; Type locality: région de Kolwezi, Katanga, Musonoie).
multipunctatus Blatchley, 1910, see: ovaliceps Fall, 1930.
multipunctatus Shibata, 1991, see: schillhammeri Shibata, 1996.
nanellus Casey, 1915, see: nigritulus Gravenhorst, 1802.
nanus Gravenhorst, 1806, see: splendidulus Gravenhorst, 1802.
negus Tottenham, 1956b: 217 (Gabrius; Type locality: Abyssinia: Djem-Djem Forest).
neobisniformis Bernhauer, 1915d: 144 (Philonthus; subgenus Gabrius; Type locality: Abessinien: Dire Daoua).
— Scheerpeltz, 1933: 1352 (Philonthus; catalog).
— Schillhammer, 1997: 79 (Gabrius).
▴ Distribution: Ethiopia.
nepalensis Schillhammer, 1997: 11 (Gabrius; Type locality: Nepal: [Patan District], Kathmandu District, Phulcoki, 2600 m).
nepos Smetana, 1984: 146 (Gabrius; Type locality: Japan: Gumma Pref.: Usui Bypass, 700 m).
neptunus Schillhammer, 1997: 43 (Gabrius; Type locality: India: W. Bengal, Darjeeling Distr. Ghoom-Lopchu, 2000 m. Also cited from Nepal).
nguembaensis Levasseur, 1968: 1383 (Gabrius; Type locality: Cameroun: forêt de Bafut, Nguemba).
nigricornis Mulsant and Rey, 1876, see: exiguus Nordmann, 1837.
nigrituloides Cameron, 1932, see: breviventer Sperk, 1835.
nigrituloides Coiffait, 1963, see: ravasinii Gridelli, 1920.
nigritulus Gravenhorst, 1802: 41 (Staphylinus; Type locality: Brunsvigae; [Note: See lectotype designation by Smetana, 1962c]).
— Latreille, 1804: 328 (Staphylinus; characters; France; Germany).
— Gravenhorst, 1806: 94 (Staphylinus; characters).
— Mannerheim, 1830: 32 (Cafius; Finland).
— Mannerheim, 1831: 446 (Cafius; Finland).
— Nordmann, 1837: 126 (Gyrohypnus; Germania; Tauria).
— Erichson, 1839a: 476 (Philonthus; synonym of aterrimus).
— Erichson, 1840: 492 (Philonthus; synonym of aterrimus).
— P. Lucas, 1846: 113 (Philonthus; synonym of aterrimus).
— Küster, 1854: 16 (Philonthus; synonym of aterrimus).
— Fairmaire and Laboulbène, 1856: 528 (Philonthus; synonym of aterrimus).
— Wollaston, 1857: 191 (Philonthus; habitat; Madeira Islands).
— Kraatz, 1857d: 616 (Philonthus; characters; Germany).
— Hochhuth, 1862: 63 (Philonthus; Russia).
— Wollaston, 1864: 574 (Philonthus; notes; Canary Islands).
— Fauvel, 1865a: 297 [= 1865b: 47] (Philonthus; habitat; France).
— Wollaston, 1865: 494 (Philonthus; habitat; Madeira Islands; Canary Islands).
— Crotch, 1867a: 383 (Philonthus; Azores).
— Baudi, 1870: 386 (Philonthus; characters).
— Fauvel, 1874: 243 [= 1874a: 469] (Philonthus; characters; France; Barbary; Turkey; Caucasus; Transcaucasia; Iran; Siberia; North America; Chile; Australia).
— Solsky, 1874: 181 (Philonthus; characters; Turkestan).
— Seidlitz, 1875: 278 (Philonthus; characters; Baltic region).
— Fauvel, 1876a: 74 [= 1876: 259] (Philonthus; Canary Islands; Syria; Turkestan).
— Mulsant and Rey, 1876: 554 [= 1877: 410] (Gabrius; characters; France).
— J. Sahlberg, 1876: 24 (Gabrius; Finland).
— Wollaston, 1877: 29 (Philonthus; characters; St. Helena).
— Fauvel, 1878: 124 [= 1878a: 44] (Philonthus; Barbary).
— Fauvel, 1878g: 548 (Philonthus; characters; Australia).
— Horn, 1884: 216 (Philonthus; characters; Vancouver to Arizona, Lake Superior; New England States).
— Blackburn and Sharp, 1885: 223 (Philonthus; habitat; Hawaii).
— Fauvel, 1886: 55 [= 1886a: 47] (Philonthus; North Africa).
— Sharp, 1889: 40 (Philonthus; notes; Japan).
— Fowler, 1888: 278 (Philonthus; characters; Britain).
— Eppelsheim, 1892: 334 (Philonthus; Turkestan).
— Ganglbauer, 1895: 460 (Philonthus; subgenus Gabrius; characters; Europe; Mediterranean region; Siberia; North America; Chile; Australia).
— Fauvel, 1902b: 111 (Philonthus; Barbary; Egypt; Azores; Madeira; Canaries).
— Fall and Cockerell, 1907: 166 (Philonthus; New Mexico).
— Sainte-Claire Deville, 1906: 97 (Philonthus; Corsica).
— Sharp, 1908: 548 (Philonthus; Hawaii).
— Montandon, 1908: 80 (Philonthus; Romania).
— Poppius, 1909: 27 (Philonthus; subgenus Gabrius; Lena Valley, Siberia).
— Reitter, 1909: 132 (Philonthus; subgenus Gabrius; characters; Germany).
— Blatchley, 1910: 389 (Philonthus; characters; Indiana).
— Eichelbaum, 1913: 130 (Philonthus; Tanzania).
— Poppius, 1913: 10 (Philonthus; subgenus Gabrius; Iceland).
— Bernhauer and Schubert, 1914: 347 (Philonthus; catalog).
— Johansen, 1914: 418 (Philonthus; subgenus Gabrius; characters; Denmark).
— Notman, 1919a: 95 (Philonthus; Ontario).
— Gridelli, 1920: 142 (Gabrius; characters; Italy).
— Šulc, 1922 (Gabrius; osmeterium; anatomy; histology; physiology).
— Lea, 1925: 220 (Philonthus; Australian introduction).
— Fall, 1926: 150 (Philonthus; Alaska, Yukon Territory).
— Cameron and Scott, 1928: 272 (Philonthus; Rodriguez Island).
— Portevin, 1929: 366 (Philonthus; characters; France).
— Gridelli, 1930: 74 (Gabrius; Libya).
— Bernhauer, 1931d: 588 (Philonthus; Abyssinia).
— Peyerimhoff, 1931: 32 (Philonthus; subgenus Gabrius; notes; Algeria).
— Cameron, 1932: 147 (Philonthus; characters; Ceylon).
— Scheerpeltz, 1933: 1352 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Scheerpeltz, 1935a: 654 (Philonthus; Australia).
— Uyttenboogaart, 1937: 82 (Philonthus; notes; Canary Islands).
— Bernhauer, 1940c: 4 (Philonthus; Azores; Madeira).
— C. Koch, 1941: 64 (Philonthus; subgenus Gabrius; Morocco).
— Méquignon, 1942a: 21 (Gabrius; Azores).
— Fagel, 1945: 40 (Gabrius; characters; Belgium).
— Jeannel and Jarrige, 1949: 376 (Philonthus; subgenus Gabrius; cave species; France; Serbia; Romania).
— Coiffait, 1951: 113, 115 (Gabrius; characters in key; France).
— V. Hansen, 1952: 89 (Philonthus; subgenus Gabrius; characters; Denmark).
— Smetana, 1952b: 120 (Gabrius; characters; Czechoslovakia).
— Smetana, 1953: 122 (Gabrius; Turkey).
— Coiffait, 1953a: 265 (Gabrius; [Note: Misidentified as latro Joy according to Schillhammer, 1997: 80]; characters; France).
— Coiffait, 1954: 165 (Gabrius; Canary Islands).
— Scheerpeltz, 1956a: 485 (Gabrius; Belgian Congo).
— Smetana, 1957: 67, 69 (Gabrius; characters).
— Smetana, 1958: 248 (Gabrius; characters; ecology; Czechoslovakia).
— Larsson and Gígja, 1959: 87 (Philonthus; biological notes; Iceland).
— Smetana, 1960c: 325 (Gabrius; characters; wide-spread species).
— Scheerpeltz, 1961d: 130 (Gabrius; Italy).
— Smetana, 1961a: 44 (Gabrius; Bulgaria).
— Smetana, 1962c: 236 (Gabrius; notes on specimens in Gravenhorst collection; lectotype designation).
— Smetana, 1963b: 38 (Gabrius; Madeira).
— Palm, 1963b: 105 (Philonthus; subgenus Gabrius; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 191 (Gabrius; characters; central Europe).
— Scheerpeltz, 1964a: 300 (Gabrius; Crete).
— Focarile, 1964: 56 (Gabrius; Italy).
— Moore, 1965: 100 (Gabrius).
— Scheerpeltz, 1965b: 403 (Gabrius; Sicily).
— Scheerpeltz, 1965c: 507 (Gabrius; Sicily).
— Horion, 1965: 194 (Gabrius; all Europe).
— Pototskaia, 1966: 154 (Gabrius; larval characters).
— Pototskaia, 1967: 57 (Gabrius; larval characters).
— Smetana, 1967b: 159 (Gabrius; Spain).
— Smetana, 1967i: 556 (Gabrius; Turkey).
— Smetana, 1970: 59 (Gabrius; distributional summary; Azores).
— Fagel, 1972: 103 (Gabrius; notes; Sainte Helene).
— Scheerpeltz, 1972: 22 (Gabrius; Argentina).
— Coiffait, 1973a: 274 (Gabrius; Morocco).
— Bordoni, 1973i: 329 (Gabrius; characters).
— Coiffait, 1974: 54 (Gabrius; characters; Europe; North Africa; Anatolia).
— Bordoni, 1974b: 9 (Gabrius; notes; Italy).
— Osella and Zanetti, 1975: 110 (Gabrius; collected from nest of Talpa europaea; notes; Italy).
— Scheerpeltz, 1976: 35 (Gabrius; Nepal).
— Pope, 1977: 30 (Gabrius; Britain).
— Smetana, 1978: 115 (Gabrius; Israel).
— Outerelo, 1980a: 56 (Gabrius; Spain).
— Tronquet, 1981: 70 (Gabrius; Afghanistan).
— Coiffait, 1981e: 349 (Gabrius; Baltistan).
— Coiffait, 1982: 27 (Gabrius; India).
— Bordoni, 1982a: 140 (Gabrius; cave collection; Sardinia).
— Frank, 1982: 36 (Gabrius; list of parasites).
— Israelson, 1984: 145 (Gabrius; Azores).
— Tóth, 1984: 91 (Gabrius; characters; Hungary).
— Lucht, 1987: 106 (Gabrius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Smetana, 1990: 178 (Gabrius; characters).
— Borges, 1990: Table II (Gabrius; Azores).
— Kuschel, 1990: 26 (Gabrius; introduced, Palaearctic species; winged; habitat; New Zealand).
— Nowosad, 1990: 114 (Gabrius; collected from mole nest; notes; Poland).
— Pilon and Zanetti, 1991: 56 (Gabrius; Italy).
— Welch, 1993: 228 (Gabrius; ovariole number and ovary structure).
— J. Li, 1993: 38 (Gabrius; China).
— Smetana, 1993b: 49 (Gabrius; Czech Republic; Slovakia).
— Nishida, 1994: 71 (Gabrius; Hawaii).
— Smetana, 1995e: 619 (Gabrius; characters; notes; Alberta; British Columbia; Manitoba; Ontario; Quebec; California; Connecticut; Idaho; Illinois; Indiana; Kentucky; Maine; Maryland; Massachusetts; Michigan; Montana; Nebraska; Nevada; New Hampshire; New Jersey; New Mexico; New York; North Dakota; Ohio; Oregon; Pennsylvania; Rhode Island; Texas; Utah; Vermont; Virginia; Washington; West Virginia; Wisconsin; Mexico).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
— Sparacio, 1995: 160 (Gabrius; characters; notes; Sicily).
— Downie and Arnett, 1996: 382 (Gabrius; characters; USA).
— Schillhammer, 1997: 14 (Gabrius; characters; Australia).
— Nishida, 1997: 61 (Gabrius; Hawaii).
— Ryabukhin, 1999: 66 (Gabrius; catalog; habitat; distribution; northeastern Russia).
— Kashcheev, 1999a: 57 (Gabrius; Kazakhstan).
▴ Distribution: St. Helena, Congo, Tanzania, Ethiopia, Egypt, Libya, Algeria, Morocco, Canary Islands, Madeira, Azores, Iceland, Europe, Russia, Caucasus, Turkey, Israel, Syria, Iran, Kazakhstan, “Turkestan”, Afghanistan, Nepal, India, Rodriguez Island, Sri Lanka, China, Japan, Hawaii, Australia, New Zealand, Canada, USA, Mexico, Argentina, Chile.
aterrimus Gravenhorst, 1802: 41 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).
— Latreille, 1804: 328 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 94 (Staphylinus; characters).
— Gyllenhal, 1810: 349 (Staphylinus; characters; Sweden).
— Gyllenhal, 1827: 479 (Staphylinus; characters).
— Zetterstedt, 1828: 79 (Staphylinus; characters; Lapland).
— Curtis, 1829: 26 (Gabrius; catalog; Britain).
— C. Sahlberg, 1830: 331 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 32 (Cafius; Finland; Sweden).
— Mannerheim, 1831: 446 (Cafius; Finland; Sweden).
— Stephens, 1833: 250 (Gabrius; characters; England).
— Runde, 1835: 11 (Staphylinus; characters; Germany).
— Lacordaire, 1835: 408 (Staphylinus; cited as atterimus; characters; France).
— Nordmann, 1837: 126 (Gyrohypnus; Europe).
— Westwood, 1838a: 16 (Gabrius; characters; type species).
— Zetterstedt, 1838: 65 (Staphylinus; characters; habitat; Lapland).
— Erichson, 1839a: 476 (Philonthus; characters; Germany).
— Stephens, 1839: 400 (Gabrius; characters; England).
— Erichson, 1840: 492 (Philonthus; characters; Europe; America septentrionalis).
— Laporte, 1840: 179 (Cafius; characters; France).
— P. Lucas, 1846: 113 (Philonthus; notes; Algeria).
— Jacobsen, 1849: 703 (Philonthus; characters; Denmark).
— Hochhuth, 1849: 144 (Philonthus; characters).
— Redtenbacher, 1849: 706 (Philonthus; characters; Austria).
— LeConte, 1850: 220 (Philonthus; Lake Superior).
— Hardy, 1851: 31 (Philonthus; notes; England).
— Mäklin, 1853: 188 (Philonthus; characters; Kadjak Island).
— Wollaston, 1854: 584 (Philonthus; characters; notes; Madeira Islands).
— Küster, 1854: 16 (Philonthus; characters; Europe).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of nigritulus).
— Rosenhauer, 1856: 71 (Philonthus; notes; Algeciras; Granada).
— Fairmaire and Laboulbène, 1856: 528 (Philonthus; characters; France).
— Kraatz, 1857d: 616 (Philonthus; synonym of nigritulus).
— Redtenbacher, 1857: 196 (Philonthus; characters; Austria).
— Peyron, 1858: 427 (Philonthus; Turkey).
— Thomson, 1860: 170 (Gabrius; characters; Scandinavia).
— Gemminger and Harold, 1868: 590 (Philonthus; synonym of nigritulus).
— Redtenbacher, 1874: 215 (Philonthus; characters; Austria).
— Ganglbauer, 1895: 460 (Philonthus; synonym of nigritulus).
— Bernhauer and Schubert, 1914: 347 (Philonthus; synonym of nigritulus).
— Cameron, 1932: 147 (Philonthus; synonym of nigritulus).
— Scheerpeltz, 1933: 1352 (Philonthus; synonym of nigritulus).
— Smetana, 1958: 249 (Gabrius; synonym of nigritulus).
— Smetana, 1960c: 325 (Gabrius; synonym of nigritulus).
— Fagel, 1972: 103 (Gabrius; synonym of nigritulus).
— Coiffait, 1974: 54 (Gabrius; synonym of nigritulus).
— Smetana, 1995e: 619 (Gabrius; synonym of nigritulus).
pumilus Mannerheim, 1830: 32 (Cafius; Type locality: Ostrogothia Sveciae).
— Mannerheim, 1831: 446 (Cafius; Sweden).
— Erichson, 1840: 492 (Philonthus; synonym of aterrimus).
— Hochhuth, 1849: 144 (Philonthus; synonym of aterrimus).
— Küster, 1854: 16 (Philonthus; synonym of aterrimus).
— Kraatz, 1857d: 616 (Philonthus; synonym of nigritulus).
— Motschulsky, 1858: 661 (Gabrius; characters; Germany).
— Gemminger and Harold, 1868: 591 (Philonthus; variety of nigritulus).
— Fauvel, 1874: 243 [= 1874a: 469] (Philonthus; synonym of nigritulus).
— Ganglbauer, 1895: 460 (Philonthus; synonym of nigritulus).
— Bernhauer and Schubert, 1914: 348 (Philonthus; synonym of nigritulus).
— Cameron, 1932: 147 (Philonthus; synonym of nigritulus).
— Smetana, 1958: 249 (Gabrius; synonym of nigritulus).
— Smetana, 1960c: 325 (Gabrius; synonym of nigritulus).
— Coiffait, 1974: 54 (Gabrius; synonym of nigritulus).
— Smetana, 1995e: 619 (Gabrius; synonym of nigritulus).
pallipes Stephens, 1833: 250 (Gabrius; Type locality: Norfolk; Suffolk).
— Stephens, 1839: 400 (Gabrius; characters; England).
— Hardy, 1851: 31 (Philonthus; synonym of aterrimus).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of nigritulus).
pygmaeus Stephens, 1833: 250 (Gabrius; Type locality: Suffolk; Norfolk; Devonshire).
— Stephens, 1839: 400 (Gabrius; characters; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of nigritulus).
— Gemminger and Harold, 1868: 591 (Philonthus; synonym of nigritulus).
phaeopus Stephens, 1833: 250 (Gabrius; Type locality: London; Hants.: New Forest).
— Stephens, 1839: 400 (Gabrius; characters; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of nigritulus).
— Gemminger and Harold, 1868: 591 (Philonthus; synonym of nigritulus).
— Bernhauer and Schubert, 1914: 347 (Philonthus; synonym of nigritulus).
— Cameron, 1932: 147 (Philonthus; synonym of nigritulus).
— Smetana, 1958: 249 (Gabrius; synonym of nigritulus).
— Smetana, 1960c: 325 (Gabrius; synonym of nigritulus).
— Coiffait, 1974: 54 (Gabrius; synonym of nigritulus).
— Smetana, 1995e: 619 (Gabrius; synonym of nigritulus).
attenuatus Stephens, 1833: 251 (Gabrius; Type locality: Battersea-fields; Devonshire).
— Stephens, 1839: 401 (Gabrius; characters; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of nigritulus).
— Gemminger and Harold, 1868: 590 (Philonthus; synonym of nigritulus).
— Bernhauer and Schubert, 1914: 347 (Philonthus; synonym of nigritulus).
— Cameron, 1932: 147 (Philonthus; synonym of nigritulus).
— Smetana, 1958: 249 (Gabrius; synonym of nigritulus).
— Smetana, 1960c: 325 (Gabrius; synonym of nigritulus).
— Coiffait, 1974: 54 (Gabrius; synonym of nigritulus).
— Smetana, 1995e: 619 (Gabrius; synonym of nigritulus).
basalis Stephens, 1833: 251 (Gabrius; Type locality: London).
— Stephens, 1839: 400 (Gabrius; characters; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of nigritulus).
— Gemminger and Harold, 1868: 590 (Philonthus; synonym of nigritulus).
— Bernhauer and Schubert, 1914: 347 (Philonthus; synonym of nigritulus).
— Cameron, 1932: 147 (Philonthus; synonym of nigritulus).
— Smetana, 1958: 249 (Gabrius; synonym of nigritulus).
— Smetana, 1960c: 325 (Gabrius; synonym of nigritulus).
— Coiffait, 1974: 54 (Gabrius; synonym of nigritulus).
— Smetana, 1995e: 619 (Gabrius; synonym of nigritulus).
pusillus Heer, 1839: 271 (Philonthus; Type locality: Bern).
— Kraatz, 1857d: 610 (Philonthus; characters; Germany).
— Fauvel, 1874: 243 [= 1874a: 469] (Philonthus; synonym of nigritulus).
— Ganglbauer, 1895: 460 (Philonthus; synonym of nigritulus).
— Bernhauer and Schubert, 1914: 348 (Philonthus; synonym of nigritulus).
— Cameron, 1932: 147 (Philonthus; synonym of nigritulus).
— Smetana, 1958: 249 (Gabrius; synonym of nigritulus).
— Smetana, 1960c: 325 (Gabrius; synonym of nigritulus).
— Coiffait, 1974: 54 (Gabrius; synonym of nigritulus).
— Smetana, 1995e: 619 (Gabrius; synonym of nigritulus).
pygmaeus Snellen, 1854: 70 (Staphylinus; [preoccupied]; Type locality: Nederland).
— Gemminger and Harold, 1868: 591 (Philonthus; variety of nigritulus).
— Fauvel, 1869: 488 (Philonthus; synonym of nigritulus).
— Fauvel, 1874: 243 [= 1874a: 469] (Philonthus; synonym of nigritulus).
— Ganglbauer, 1895: 460 (Philonthus; synonym of nigritulus).
— Bernhauer and Schubert, 1914: 348 (Philonthus; synonym of nigritulus).
— Cameron, 1932: 147 (Philonthus; synonym of nigritulus).
— Smetana, 1958: 249 (Gabrius; synonym of nigritulus).
— Smetana, 1960c: 325 (Gabrius; synonym of nigritulus).
— Coiffait, 1974: 54 (Gabrius; synonym of nigritulus).
— Smetana, 1995e: 620 (Gabrius; synonym of nigritulus).
ovicollis Broun, 1880: 109 (Staphylinus; Type locality: Whangarei Harbour).
— Bernhauer and Schubert, 1914: 347 (Philonthus; synonym of nigritulus).
— Cameron, 1932: 147 (Philonthus; synonym of nigritulus).
— Cameron, 1945d: 786 (Philonthus; subgenus Gabrius; near nigritulus).
— Smetana, 1958: 249 (Gabrius; synonym of nigritulus).
— Smetana, 1960c: 325 (Gabrius; synonym of nigritulus).
— Smetana, 1995e: 620 (Gabrius; synonym of nigritulus).
nanellus Casey, 1915: 440 (Philonthus; Type locality: northern Illinois: Highland Park; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1352 (Philonthus; catalog).
— Moore, 1965: 100 (Gabrius).
— Smetana, 1995e: 620 (Gabrius; lectotype designation; synonym of nigritulus).
— Downie and Arnett, 1996: 382 (Gabrius; characters; USA).
magniceps Smetana, 1952b: 122, 132 (Gabrius; variety of nigritulus; [preoccupied]; Type locality: Boh., Roudnice n. L.).
— Smetana, 1958: 249 (Gabrius; synonym of nigritulus).
— Smetana, 1960c: 325 (Gabrius; synonym of nigritulus).
— Coiffait, 1974: 54 (Gabrius; synonym of nigritulus).
— Smetana, 1995e: 620 (Gabrius; synonym of nigritulus).
signiceps Smetana, 1954: 41 (Gabrius; variety of nigritulus; replacement name for magniceps Smetana).
— Smetana, 1957: 67, 69 (Gabrius; characters).
— Smetana, 1958: 249 (Gabrius; synonym of nigritulus).
— Smetana, 1960c: 325 (Gabrius; synonym of nigritulus).
— Scheerpeltz, 1965b: 403 (Gabrius; variety of nigritulus; Sicily).
— Scheerpeltz, 1965c: 507 (Gabrius; variety of nigritulus; Sicily).
— Coiffait, 1974: 54 (Gabrius; characters; variety of nigritulus).
— Tóth, 1984: 91 (Gabrius; variety of nigritulus).
— Smetana, 1995e: 620 (Gabrius; synonym of nigritulus).
nitetis Tottenham, 1956: 301 (Gabrius; Type locality: Ruanda: terr. Ruhengeri, Kagogo, 1900 m).
noei Coiffait, 1966c: 506 (Gabrius; Type locality: Turquie d'Asie: Mont Ararat, 3000 m).
normandi Smetana, 1956: 78 (Gabrius; Type locality: Nord-Afrika, Algerien, Edough).
— Smetana, 1957: 57 (Gabrius; characters; Algeria).
— Smetana, 1961: 36 (Gabrius; characters; Algeria).
— Coiffait, 1964c: 168 (Gabrius; characters; Algeria; Morocco; Portugal?).
— Coiffait, 1974: 73 (Gabrius; characters; Algeria; Morocco; Portugal ?).
▴ Distribution: Algeria, Morocco, Portugal ?.
novellus Tottenham, 1956b: 208, 213 (Gabrius; [Note: Cited as novus Cameron on pages 208 and 213 but renamed as a new species according to footnote on page 213]; Type locality: Angola).
novus Cameron, 1935a: 375 (Philonthus; subgenus Gabrius; Type locality: Mahagi: Nialopul).
obenbergeri Smetana, 1953a: 165 (Gabrius; Type locality: Yugoslavia: Sarajevo).
— Smetana, 1953: 119 (Gabrius; notes on species group placement).
— Smetana, 1957: 59 (Gabrius; characters; Yugoslavia).
— Smetana, 1961: 39 (Gabrius; characters; Yugoslavia; Bulgaria).
— Smetana, 1961a: 44 (Gabrius; Bulgaria).
— Coiffait, 1974: 68 (Gabrius; characters; Yugoslavia; Bulgaria).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
▴ Distribution: Italy, Bosnia Hercegovina, Bulgaria.
oblidens Tottenham, 1956b: 221 (Gabrius; Type locality: Natal: Mouth of Umkomaas River. Also cited from Tanganyika).
obnixus Tottenham, 1956b: 217 (Gabrius; Type locality: S. Rhodesia: Mashonaland).
obscuratus Cameron, 1950g: 41 (Philonthus; subgenus Gabrius; Type locality: Belgian Congo: lac N'Gando, pied volc. Karisimbi, alt. 2400 m).
oceanicus Tottenham, 1956b: 223 (Gabrius; Type locality: Seychelles).
— Bernhauer, 1922b: 176 (Philonthus; [Note: Misidentified as fimbriolatus according to Tottenham, 1956b: 223]; Seychelles).
▴ Distribution: Seychelles.
oceanus Hatch, 1957: 204 (Philonthus; subgenus Gabrius; Type locality: Washington: Seaview. Also cited from British Columbia).
— Moore, 1965: 100 (Gabrius).
— Smetana, 1995e: 707 (Gabrius; characters; notes; British Columbia).
▴ Distribution: Canada, USA.
okamotoi Shibata, 1991a: 85 (Gabrius; Type locality: Japan: Hiroshima Pref.: Anjûkyô, Kake-chô, Yamagata-gun).
ophion Smetana, 1984: 146 (Gabrius; Type locality: Japan: Gumma Pref.: 7 km E Usui Pass, 850 m).
osseticus Kolenati, 1846: 20 (Philonthus; Type locality: Caucasus centralis ad montem Kasbek in Ossetia).
— Hochhuth, 1849: 140 (Philonthus; synonym of vernalis).
— Bernhauer and Schubert, 1914: 360 (Philonthus; synonym of vernalis).
— Smetana, 1958: 235 (Gabrius; synonym of vernalis).
— Coiffait, 1974: 71 (Gabrius; synonym of vernalis).
— Pope, 1977: 30 (Gabrius; Britain).
— Frank, 1982: 36 (Gabrius; list of parasites).
— Lohse and Lucht, 1989: 168 (Gabrius; note).
— Pilon and Zanetti, 1991: 55 (Gabrius; Italy).
— Smetana, 1993b: 49 (Gabrius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
— Ádám, 1996: 245 (Gabrius; Hungary).
— Gillerfors, 1997: 85 (Gabrius; characters).
— Gerend and Braunert, 1997: 198 (Gabrius; notes; Luxembourg).
— Owen, 1997a: 149 (Gabrius; collecting sites; Ireland).
— Ryabukhin, 1999: 65 (Gabrius; catalog; habitat; distribution; northeastern Russia).
— Eyre, Luff, and Lott, 2000: 33 (Gabrius; Britain).
— Herman, 2001: 25 (Gabrius; synonymy; protected status proposed).
▴ Distribution: Europe, Russia, Georgia, Caucasus, Turkey.
vernalis Gravenhorst, 1806: 75 (Staphylinus; [preoccupied]; Type locality: Göttingae; Brunsvigae).
— Gyllenhal, 1810: 332 (Staphylinus; characters; Sweden).
— C. Sahlberg, 1830: 324 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 29 (Staphylinus; Finland; Sweden).
— Mannerheim, 1831: 443 (Staphylinus; Finland; Sweden).
— Lacordaire, 1835: 404 (Staphylinus; characters; France).
— Nordmann, 1837: 93 (Philonthus; Europe).
— Erichson, 1839a: 470 (Philonthus; characters; Germany).
— Heer, 1839: 268 (Philonthus; characters; Switzerland).
— Stephens, 1839: 400 (Gabrius; characters; England).
— Erichson, 1840: 478 (Philonthus; characters; Europe).
— Laporte, 1840: 178 (Staphylinus; characters; France).
— Grimm, 1845: 135 (Philonthus; with Formica rufa; Germany).
— Jacobsen, 1849: 701 (Philonthus; characters; Denmark).
— Hochhuth, 1849: 140 (Philonthus; characters).
— Redtenbacher, 1849: 705 (Philonthus; characters; Austria).
— Küster, 1854: 13 (Philonthus; characters; Germany).
— Fairmaire and Laboulbène, 1856: 523 (Philonthus; characters; France).
— Kraatz, 1857d: 606 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 195 (Philonthus; characters; Austria).
— Peyron, 1858: 426 (Philonthus; Turkey).
— Thomson, 1860: 170 (Gabrius; characters; Scandinavia).
— Hochhuth, 1862: 62 (Philonthus; Russia).
— Fauvel, 1874: 261 [= 1874a: 487] (Philonthus; characters; France; Sardinia; Turkey; Georgia; Caucasus; Lake Baikal).
— Redtenbacher, 1874: 214 (Philonthus; characters; Austria).
— Seidlitz, 1875: 276 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 538 [= 1877: 394] (Gabrius; characters; France).
— J. Sahlberg, 1876: 23 (Gabrius; Finland).
— Fowler, 1888: 274 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 459 (Philonthus; subgenus Gabrius; characters; Europe; Siberia).
— Montandon, 1908: 80 (Philonthus; Romania).
— Reitter, 1909: 131 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 360 (Philonthus; catalog).
— Johansen, 1914: 416 (Philonthus; subgenus Gabrius; characters; Denmark).
— Gridelli, 1920: 127 (Gabrius; characters; Italy).
— Gridelli, 1928: 50 (Gabrius; characters in key).
— Portevin, 1929: 366 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1367 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— V. Hansen, 1952: 84 (Philonthus; subgenus Gabrius; characters; Denmark).
— Smetana, 1957: 58 (Gabrius; characters; Europe to Siberia).
— Smetana, 1958: 235 (Gabrius; characters; ecology; Czechoslovakia).
— Smetana, 1961: 38 (Gabrius; characters).
— Smetana, 1961a: 43 (Gabrius; Bulgaria).
— Palm, 1963b: 103 (Philonthus; subgenus Gabrius; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 187 (Gabrius; characters; central Europe).
— Horion, 1965: 183 (Gabrius; Russia; Finland; Sweden; Norway; Britain; Denmark; widespread in all central Europe; Spain; Italy; Dalmatia; Bosnia).
— Pototskaia, 1966: 154 (Gabrius; larval characters).
— Pototskaia, 1967: 56 (Gabrius; larval characters).
— Szujecki, 1968a: 732 (Gabrius; Poland).
— Coiffait, 1974: 71 (Gabrius; characters; Europe; Scandinavia; Siberia; France; Corsica).
— Pope, 1977: 30 (Gabrius; synonym of osseticus).
— Tóth, 1984: 83 (Gabrius; characters; Hungary).
— Lucht, 1987: 106 (Gabrius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Lohse and Lucht, 1989: 168 (Gabrius; synonym of osseticus).
— Nowosad, 1990: 113 (Gabrius; collected from mole nest; notes; Poland).
— Smetana, 1993b: 49 (Gabrius; synonym of osseticus).
— Drugmand, 1994b (Gabrius; aberrant aedeagus).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; synonym of osseticus).
— Ádám, 1996: 245 (Gabrius; synonym of osseticus).
suaveolens Stephens, 1833: 249 (Gabrius; [Note: This name is older than osseticus]; Type locality: Hertford, banks of Lea River).
— Stephens, 1839: 400 (Gabrius; characters; England).
— Hardy, 1851: 29 (Philonthus; synonym of varians).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of vernalis).
— Bernhauer and Schubert, 1914: 360 (Philonthus; synonym of vernalis).
— Tottenham, 1939d: 231 (Gabrius; synonym of ventralis).
— Smetana, 1958: 235 (Gabrius; synonym of vernalis).
— Coiffait, 1974: 71 (Gabrius; synonym of vernalis).
— Herman, 2001: 25 (Gabrius; synonym of osseticus).
flavipes Motschulsky, 1860: 567 (Gabrius; [Note: Bernhauer and Schubert, 1914 cited this name as a synonym of femoralis (p. 337) and vernalis (p. 360). Most recently Schillhammer, 1997: 80 listed it as a synonym of osseticus]; Type locality: Géorgie méridionale près de la frontière de Perse).
— Hochhuth, 1862: 61 (Philonthus; characters; Georgia).
— Fauvel, 1874: 261 [= 1874a: 487] (Philonthus; synonym of vernalis).
— Bernhauer and Schubert, 1914: 337 (Philonthus; synonym of femoralis).
— Bernhauer and Schubert, 1914: 360 (Philonthus; synonym of vernalis).
— Scheerpeltz, 1933: 1342 (Philonthus; synonym of femoralis).
— Coiffait, 1951: 107, 114 (Gabrius; cited as valid species; characters in key; France).
— Smetana, 1958: 235, 236 (Gabrius; synonym of vernalis [235] and femoralis [236]).
— Coiffait, 1974: 71 (Gabrius; synonym of vernalis).
— Schillhammer, 1997: 80 (Gabrius; synonym of osseticus).
flavimanus Gemminger and Harold, 1868: 588 (Philonthus; replacement for flavipes Motschulsky).
— Fauvel, 1874: 261 [= 1874a: 487] (Philonthus; synonym of vernalis).
— Bernhauer and Schubert, 1914: 360 (Philonthus; synonym of vernalis).
— Scheerpeltz, 1933: 1342 (Philonthus; synonym of femoralis).
— Smetana, 1958: 235, 236 (Gabrius; synonym of vernalis [235] and femoralis [236]).
— Coiffait, 1974: 71 (Gabrius; synonym of femoralis).
— Schillhammer, 1997: 77 (Gabrius; synonym of femoralis).
ottawensis Casey, 1915, see: punctatellus Horn, 1884.
ovaliceps Fall, 1930: 253 (Philonthus; Type locality: Three Mile Island, Lake Winnipesaukee, N.H.; also listed from Massachusett and Virginia).
— Scheerpeltz, 1933: 1354 (Philonthus; catalog).
— Moore, 1965: 100 (Gabrius).
— Smetana, 1995e: 676 (Gabrius; characters; notes; Ontario; Connecticut; Illinois; Indiana; Kentucky; Maine; Maryland; Massachusetts; Michigan; Missouri; New Hampshire; New Jersey; New York; North Carolina; Ohio; Pennsylvania; Rhode Island; Texas; Vermont; Virginia; West Virginia).
▴ Distribution: Canada, USA.
multipunctatus Blatchley, 1910: 389 (Philonthus; [Note: Preoccupied as a secondary homonym in Philonthus]; Type locality: Lawrence County, Indiana).
— Bernhauer and Schubert, 1914: 347 (Philonthus; catalog).
— Blatchley, 1936: 256 (Philonthus; synonym of kessleri).
— Smetana, 1995e: 676 (Gabrius; synonym of ovaliceps).
kessleri Blatchley, 1936: 256 (Philonthus; replacement name for multipunctatus Blatchley).
— Smetana, 1995e: 676 (Gabrius; synonym of ovaliceps).
— Downie and Arnett, 1996: 381 (Philonthus; characters; USA).
ovicollis Broun, 1880, see: nigritulus Gravenhorst, 1802.
paederoides Motschulsky, 1858, see: Philonthus.
pakistanicus Schillhammer, 1997: 19 (Gabrius; Type locality: Pakistan: Chitral, Lawarai Pass, 2600 m).
pallipes Stephens, 1833, see: nigritulus Gravenhorst, 1802.
palmi Schillhammer, 1992: 62 (Gabrius; Type locality: Italy: Sardinien: S. Teresa).
parabadilikus Tottenham, 1962: 253 (Gabrius; Type locality: Tanganyika Terr.: Ngorongoro, 2300 m).
parkeri Cameron, 1932: 146 (Philonthus; Type locality: India, Uttar Pradesh, Konain, 7800′, Chakrata; [Note: See lectotype designation by Schillhammer, 1997]. Originally cited from Chakrata and Simla districts, alt. 6000–8000 feet).
— Scheerpeltz, 1933: 1354 (Philonthus; catalog).
— Schillhammer, 1997: 38 (Gabrius; lectotype designation: India, Uttar Pradesh, Konain, 7800′, Chakrata; characters; India; Nepal).
— Schillhammer, 2000c: 129 (Gabrius; Nepal).
▴ Distribution: India, Nepal.
sodalis Cameron, 1943: 131 (Philonthus; subgenus Gabrius; Type locality: United Provinces: Chakrata district: Sainj Khud, altitude 6500 feet).
parvulus Gravenhorst, 1802, see: splendidulus Gravenhorst, 1802.
parvus Horn, 1884, see: eremius Herman, 2001.
pelzelmayeri Bernhauer, 1939k: 590 (Philonthus; subgenus Gabrius; Type locality: N. W. China: Tienmuschan).
— Smetana, 1973b: 133 (Gabrius; cited as pelzelmeyeri; characters; notes).
— Coiffait, 1974: 84 (Gabrius; characters; China).
▴ Distribution: China.
peninsula Tottenham, 1962: 247 (Gabrius; Type locality: Tanganyika Terr.: Uluguru Mts., sommet du Kidunda, 1800–1950 m).
pennatus Sharp, 1910, see: breviventer Sperk, 1835.
perexcelsus Tottenham, 1939d: 233 (Philonthus; replacement name for excelsus Cameron).
— Schillhammer, 2000c: 129 (Gabrius; Nepal).
— Herman, 2001: 14 (Gabrius).
▴ Distribution: India, Nepal.
excelsus Cameron, 1932: 147 (Philonthus; [preoccupied]; Type locality: India, Himachal Pradesh, Gahan, 7000′; [Note: See lectotype designation by Schillhammer, 1997]. Also cited from Chakrata district: Bangar, alt. 6000 feet).
— Scheerpeltz, 1933: 1341 (Philonthus; catalog).
— Schillhammer, 1997: 34 (Gabrius; lectotype designation: India, Himachal Pradesh, Gahan, 7000′; characters; India; Nepal).
— Schillhammer, 2000c: 129 (Gabrius; synonym of perexcelsus).
perflexus Schillhammer, 1997: 37 (Gabrius; Type locality: Nepal: [Sankhua Sabha District]: Khandbari District, For. NE Kuwapani, 2500 m).
peyerimhoffi Coiffait, 1964c: 170 (Gabrius; Type locality: versant nord du Mont Mahmel, Massif de l'Aurès, Algérie).
phaeopus Stephens, 1833, see: nigritulus Gravenhorst, 1802.
philo Smetana, 1984: 140 (Gabrius; Type locality: Japan: Nikko N. P., Ryuzu).
philomimus Schillhammer, 2000c: 125 (Gabrius; Type locality: Russia: Primorskiy Kray: Sikhote-Alin, Biol. Stat., 30 km SE Chuguyevka, 44.05N, 134.12E, 650 m).
philonthoides Schillhammer, 1992: 64 (Gabrius; Type locality: China: Gansu, Meijishan).
picipennis Mäklin, 1852: 313 (Philonthus; [preoccupied]; Type locality: insula Sitkha).
— Fauvel, 1874: 237 [= 1874a: 463] (Philonthus; synonym of quisquiliarius).
— Ganglbauer, 1895: 460 (Philonthus; synonym of nigritulus).
— Bernhauer and Schubert, 1914: 350 (Philonthus; catalog).
— Hatch, 1957: 199 (Philonthus; subgenus Gabrius; characters; British Columbia; Washington; Oregon).
— Moore, 1965: 100 (Gabrius).
— Smetana, 1965: 29 (Gabrius; characters; notes; Labrador; St. Pierre; Newfoundland).
— Smetana, 1990: 178 (Gabrius; characters).
— Smetana, 1995e: 629 (Gabrius; characters; notes; all Provinces and Territories of Canada except Prince Edward Island; Alaska; Arizona; California; Colorado; Connecticut; Idaho; Illinois; Maine; Massachusetts; Michigan; Montana; Nebraska; Nevada; New Hampshire; New Mexico; New Mexico; New York; North Dakota; Oregon; Pennsylvania; South Dakota; Utah; Washington; Wisconsin; Wyoming).
— Herman, 2001: 44 (Gabrius; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Canada, USA.
decipiens Horn, 1884: 217 (Philonthus; [preoccupied]; Type locality: California).
— Bernhauer and Schubert, 1914: 334 (Philonthus; catalog).
— Fall, 1926: 150 (Philonthus; Alaska).
— Moore, 1965: 100 (Gabrius).
— Smetana, 1995e: 629 (Gabrius; synonym of picipennis).
lopezi Hatch, 1957: 199 (Philonthus; subgenus Gabrius; Type locality: Washington: Lopez Is. Also cited from British Columbia, Oregon, and Idaho).
— Moore, 1965: 100 (Gabrius).
— Smetana, 1965: 57 (Gabrius; synonym of decipiens).
— Smetana, 1995e: 629 (Gabrius; synonym of picipennis).
piger Schillhammer, 1997: 32 (Gabrius; Type locality: NW-India [Kashmir], Cachmire, Pahalgam Kolakoi, 3400 m).
piliger Mulsant and Rey, 1876: 553 [= 1877: 409] (Gabrius; Type locality: Corte Saint-François, en Corse).
— Bernhauer and Schubert, 1914: 347 (Philonthus; synonym of nigritulus).
— Cameron, 1932: 147 (Philonthus; synonym of nigritulus).
— Scheerpeltz, 1933: 1353 (Philonthus; synonym of nigritulus).
— Coiffait, 1951: 110, 114 (Gabrius; characters in key; France).
— Smetana, 1957: 63, 68 (Gabrius; characters; England; Scandinavia; Italy; Corsica; Caucasus; Turkey; Germany; Czechoslovakia; Austria).
— Smetana, 1958: 247 (Gabrius; characters; ecology; Czechoslovakia).
— Smetana, 1960c: 311 (Gabrius; characters; Turkey; Sweden; Denmark; Norway; Germany; Holland; Belgium; England; France; Italy; Corsica; Austria; Czechoslovakia; Yugoslovia; Siberia; Caucasus).
— Palm, 1963b: 106 (Philonthus; subgenus Gabrius; characters; Sweden).
— Lohse, 1964: 189 (Gabrius; characters; central Europe).
— Horion, 1965: 190 (Gabrius; Europe).
— Coiffait, 1974: 82 (Gabrius; characters; Europe; Asia Minor; Caucasus; Daghestan; Sweden; Britain; Italy; France; Corsica).
— Pope, 1977: 30 (Gabrius; Britain).
— Petrenko, 1980: 82 (Gabrius; Caucasus).
— Tóth, 1984: 87 (Gabrius; characters; Hungary).
— Lucht, 1987: 106 (Gabrius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; Benelux; Sweden).
— Welch, 1993: 229 (Gabrius; ovariole number and ovary structure).
— Smetana, 1993b: 49 (Gabrius; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
— Gerend and Braunert, 1997: 198 (Gabrius; notes; Luxembourg).
▴ Distribution: Europe, Russia, Caucasus, Turkey.
stipes Sharp, 1910: 129 (Gabrius; Type locality: Anglia: Plymouth; Mickleham; Cambridge).
— Bernhauer and Schubert, 1914: 356 (Philonthus; catalog).
— Gridelli, 1920: 152 (Gabrius; characters; England; Daghestan; Caucasus).
— Sainte-Claire Deville, 1926: 119 (Philonthus; subgenus Gabrius; notes; Corsica).
— Scheerpeltz, 1933: 1362 (Philonthus; catalog).
— Wüsthoff, 1937: 32 (Philonthus; illustration of aedeagus).
— Méquignon, 1944: 20 (Gabrius; synonym of piliger).
— Fagel, 1945: 40 (Gabrius; characters; Belgium).
— Smetana, 1952b: 119 (Gabrius; characters; Czechoslovakia).
— Smetana, 1953: 121 (Gabrius; Turkey).
— Smetana, 1953a: 178 (Gabrius; Savinian Alps).
— Smetana, 1957: 63 (Gabrius; synonym of piliger).
— Smetana, 1958: 247 (Gabrius; synonym of piliger).
— Smetana, 1960c: 311 (Gabrius; synonym of piliger).
— Lohse, 1964: 189 (Gabrius; synonym of piliger).
— Pope, 1977: 30 (Gabrius; synonym of piliger).
pilipes Bierig, 1940, see: chiriquiensis Herman, 2001.
pisciformis Fauvel, 1874: 242 [= 1874a: 468] (Philonthus; Type locality: Var, Entraigues).
— Fauvel, 1874: 340 [= 1874b: 46] (Philonthus; Spain).
— Mulsant and Rey, 1876: 542 [= 1877: 398] (Gabrius; characters; France).
— Eppelsheim, 1887b: 432 (Philonthus; synonym of femoralis).
— Ganglbauer, 1895: 459 (Philonthus; subgenus Gabrius; characters; France; Spain).
— Bernhauer, 1899: 24 (Philonthus; notes; Siebenbürgen; Hungary; Bosnia; Austria).
— Bernhauer and Schubert, 1914: 337 (Philonthus; synonym of femoralis).
— Portevin, 1929: 365 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1342 (Philonthus; catalog).
— Smetana, 1957: 58 (Gabrius; synonym of femoralis).
— Smetana, 1958: 236 (Gabrius; synonym of femoralis).
— Smetana, 1961: 37 (Gabrius; synonym of femoralis).
— Smetana, 1967b: 156 (Gabrius; Spain).
— Coiffait, 1974: 67 (Gabrius; characters; Spain; France; Pyrenees).
▴ Distribution: France, Spain.
tibialis Mulsant and Rey, 1876: 540 [= 1877: 396] (Gabrius; Type locality: environs de Fréjus et dans le Languedoc).
— Duvivier, 1883: 148 (Philonthus; catalog; synonym of pisciformis).
— Ganglbauer, 1895: 459 (Philonthus; synonym of pisciformis).
— Bernhauer and Schubert, 1914: 337 (Philonthus; synonym of femoralis).
— Coiffait, 1951: 105, 114 (Gabrius; cited as valid species; characters in key; France).
— Smetana, 1957: 59 (Gabrius; cited as valid species; characters; France; Spain).
— Fagel, 1959a: 96 (Gabrius; notes; Spain).
— Smetana, 1961: 39 (Gabrius; characters; Spain; France).
— Smetana, 1967b: 156 (Gabrius; synonym of pisciformis).
— Coiffait, 1974: 67 (Gabrius; synonym of pisciformis).
gavoyi Bernhauer, 1928b: 12 (Philonthus; subgenus Gabrius; Type locality: Gavoy in Südfrankreich; Granada in Spanien).
— Scheerpeltz, 1933: 1344 (Philonthus; catalog).
— Smetana, 1967b: 156 (Gabrius; synonym of pisciformis).
— Smetana, 1973b: 126 (Gabrius; notes; synonym of pisciformis).
— Coiffait, 1974: 67 (Gabrius; synonym of pisciformis).
plumosus Last, 1981: 125 (Gabrius; Type locality: Marifuanga, Asaro-Chimbu Divide).
— Last, 1987: 55 (Gabrius; Papua New Guinea).
— Schillhammer, 1997: 64 (Gabrius; characters; New Guinea).
▴ Distribution: New Guinea.
praedictus Schillhammer, 1997: 57 (Gabrius; Type locality: Nepal: 328 Panchthar Distr., Paniporua, 2300 m).
primigenius Joy, 1913: 25 (Gabrius; Type locality: N.W. Spain, Pontevedra).
— Scheerpeltz, 1933: 1367 (Philonthus; synonym of velox).
— Smetana, 1957: 65 (Gabrius; synonym of velox).
— Smetana, 1958: 251 (Gabrius; synonym of velox).
— Smetana, 1960c: 331 (Gabrius; characters; Spain; North Africa; France).
— Smetana, 1967b: 159 (Gabrius; cited as promigenius; Spain).
— Coiffait, 1974: 60 (Gabrius; characters; North Africa; Iberian Peninsula; France).
— Pope, 1977: 30 (Gabrius; synonym of velox).
— Smetana, 1995e: 627 (Gabrius; valid species).
— Outerelo, Gamarra, and Salgado, 1998: 129 (Gabrius; collected from cave; Spain).
▴ Distribution: France, Spain, Morocco.
mauretanicus Peyerimhoff, 1937a: 266 (Philonthus; subgenus Gabrius; Type locality: Causse du Mahmel, Aurès, vers 1800 m).
— Coiffait, 1951: 112, 115 (Gabrius; characters in key; France).
— Smetana, 1957: 65 (Gabrius; characters; North Africa; France).
— Smetana, 1960c: 331 (Gabrius; synonym of primigenius).
— Smetana, 1967b: 159 (Gabrius; synonym of primigenius).
— Coiffait, 1973a: 274 (Gabrius; Morocco).
— Coiffait, 1974: 60 (Gabrius; synonym of primigenius).
propinquus Cameron, 1933d: 389 (Philonthus; subgenus Gabrius; [preoccupied]; Type locality: Sumatra: Gunung Singgalang, 1000 m; [Note: See lectotype designation by Schillhammer, 1997]).
— Cameron, 1937: 6 (Philonthus; Java).
— Schillhammer, 1997: 49 (Gabrius; lectotype designation; characters; Sumatra; Java; Bali).
— Schillhammer, 2000c: 132 (Gabrius; Java; Bali).
— Herman, 2001: 44 (Gabrius; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Indonesia.
pseudimitator Schillhammer, 1997: 46 (Gabrius; Type locality: Thailand: Chiang Mai, Doi Suthep, 1550 m).
pseudopennatus Cameron, 1945c: 716 (Philonthus; subgenus Gabrius; Type locality: South Africa: Simon's Town).
— Tottenham, 1956: 305 (Gabrius; characters in key).
— Tottenham, 1956b: 207, 208 (Gabrius; characters).
▴ Distribution: South Africa.
pubens Mulsant and Rey, 1878, see: Gabronthus.
pullatus Herman, 2001: 44 (Gabrius; replacement name for turneri Cameron).
turneri Cameron, 1951c: 403 (Philonthus; subgenus Gabrius; [preoccupied]; Type locality: Kenya Colony: Londiani).
punctatellus Horn, 1884: 215 (Philonthus; [preoccupied]; Type locality: Ft. McLeod Brit. Amer.; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from Hudson's Bay region; Lake Tahoe).
— Bernhauer and Schubert, 1914: 352 (Philonthus; catalog).
— Moore, 1965: 100 (Gabrius).
— Smetana, 1995e: 692 (Gabrius; subspecies punctatellus; lectotype designation: Ft. McLeod Brit. Amer.; characters; notes; Alberta; British Columbia; Saskatchewan; Yukon Territory; Colorado; Idaho; Montana; Oregon; Utah; Washington).
— Herman, 2001: 44 (Gabrius; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Canada, USA.
ottawensis Casey, 1915: 438 (Philonthus; Type locality: Canada: Ottawa; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1354 (Philonthus; catalog).
— Moore, 1965: 100 (Gabrius; cited as ottawaensis).
— Smetana, 1995e: 693, 695 (Gabrius; subspecies of punctatellus; lectotype designation; characters; notes; Manitoba; Ontario; Quebec; Massachusetts; New Hampshire; New York; Wisconsin).
downiei Hatch, 1957: 203 (Philonthus; subgenus Gabrius; Type locality: Washington: Pullman. Also cited from British Columbia and Idaho).
— Moore, 1965: 100 (Gabrius; cited as downei).
— Smetana, 1995e: 692 (Gabrius; synonym of punctatellus punctatellus).
wawawai Hatch, 1957: 203 (Philonthus; subgenus Gabrius; Type locality: Washington; Wawawai. Also cited from Oregon and British Columbia).
miltiades Smetana, 1995e: 693, 695 (Gabrius; subspecies of punctatellus; Type locality: New Mexico: Lincoln N.F., 1 mi. SE Cloudcroft, 8750 feet. Also cited from Arizona, Nevada, Nebraska; labelled as intermediate between miltiades and punctatellus: Colorado and Utah).
pujoli Levasseur, 1967, see: holisinus Fauvel, 1904.
pumilus Mannerheim, 1830, see: nigritulus Gravenhorst, 1802.
pusillus Heer, 1839, see: nigritulus Gravenhorst, 1802.
puthzi Schillhammer, 1997: 35 (Gabrius; Type locality: Nepal [Sindhupalchok District]: Prov. Bagmati, Malemchi, 2800 m).
pygmaeus Stephens, 1833, see: nigritulus Gravenhorst, 1802.
pygmaeus Snellen, 1854, see: nigritulus Gravenhorst, 1802.
quadriceps Jarrige, 1978: 285 (Gabrius; Type locality: Andringitra Est, Anjavidilava, 2000 m).
quadripunctatus Schillhammer, 1992: 64 (Gabrius; Type locality: China: Gansu, Meijishan).
rabigoides Jarrige, 1957: 110 (Gabrius; Type locality: Réunion: forêt de Bélouve).
ramifer Tottenham, 1956: 303 (Gabrius; Type locality: Ruanda: Shangugu, 1500 m).
ravasinii Gridelli, 1920: 147 (Gabrius; Type locality: Carinzia).
— Scheerpeltz, 1933: 1358 (Philonthus; catalog).
— Fagel, 1955a: 85 (Gabrius; Greece).
— Smetana, 1960c: 351 (Gabrius; characters from original description; species uncertain).
— Horion, 1965: 195 (Gabrius; Austria; Slovenia; Greece; Holland).
— Bordoni, 1973i: 329 (Gabrius; characters).
— Coiffait, 1974: 55 (Gabrius; characters; Europe; Asia Minor; Caucasus; Pyrenees).
— Smetana, 1977: 794 (Gabrius; Turkey).
— Tóth, 1984: 92 (Gabrius; characters; Hungary).
— Schillhammer, 1990: 91 (Gabrius; characters; notes).
— Smetana, 1993b: 49 (Gabrius; Czech Republic; Slovakia).
— Schillhammer, 1993: 326 (Gabrius; Austria).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
— Assing, Frisch, Kahlen, et al., 1998: 138 (Gabrius; note; Austria).
— Tronquet, 1998: 11 (Gabrius; notes; France).
▴ Distribution: Europe, Caucasus, Turkey.
spurius Smetana, 1954b: 178 (Gabrius; [Note: This name was validated by Smetana, 1954b; comparative characters were cited and an illustration of the aedeagus was published; the author stated the species was to be described later]; Type locality: Moravia: Prostějov [Note: Localities taken from Smetana, 1956b: 175. Also cited from Anatolia, Bohemia, Czechoslovakia, Austria, Hungary and the Caucasus).
— Smetana, 1952b: 122 (Gabrius; [Note: Misidentified as subnigritulus according to Smetana, 1958: 250]; characters; Czechoslovakia).
— Smetana, 1957: 67, 69 (Gabrius; characters; Czechoslovakia; Austria; Hungary; Transylvania; Bulgaria; Caucasus; Turkey; Germany).
— Smetana, 1958: 250 (Gabrius; characters; ecology; Czechoslovakia).
— Smetana, 1960c: 323 (Gabrius; characters; Caucasus; Turkey; Czechoslovakia; Austria; Hungary; Transylvania; Bulgaria).
— Smetana, 1961a: 44 (Gabrius; Bulgaria).
— Lohse, 1964: 191 (Gabrius; characters; central Europe).
— Horion, 1965: 196 (Gabrius; Caucasus; Turkey; Bulgaria; Romania; Czech Republic; Slovakia; Austria).
— Smetana, 1967b: 159 (Gabrius; Spain).
— Coiffait, 1974: 55 (Gabrius; synonym of ravasinii).
— Smetana, 1977: 794 (Gabrius; synonym of ravasinii).
— Lucht, 1987: 106 (Gabrius; checklist; Czechoslovakia; Austria).
— Schillhammer, 1990: 91 (Gabrius; notes).
— Smetana, 1993b: 49 (Gabrius; synonym of ravasinii).
— Assing, Frisch, Kahlen, et al., 1998: 138 (Gabrius; synonym of ravasinii).
spurius Smetana, 1956b: 171, 175 (Gabrius; [Note: Cited as a new species but the name was validated in Smetana, 1954b]; characters; Moravia. Also cited from Anatolia, Bohemia, Austria, Hungary, Transylvania, Bulgaria, and Caucasus).
nigrituloides Coiffait, 1963: 20 (Gabrius; [preoccupied]; Type locality: Forêt de Lente, Drôme).
— Coiffait, 1974: 55 (Gabrius; characters; France).
— Schillhammer, 1996a: 81 (Gabrius; synonym of ravasinii).
riedeli Schillhammer, 1997: 70 (Gabrius; Type locality: Indonesia: N-Sumatra: Kutacane Gg. Sinabung, 2000 m).
robustus Smetana, 1953a: 168 (Gabrius; Type locality: Macedonia: Kistac).
— Smetana, 1957: 66 (Gabrius; characters; Macedonia).
— Smetana, 1960c: 349 (Gabrius; characters; Macedonia).
— Coiffait, 1974: 59 (Gabrius; characters; Macedonia).
— Schuh, Schillhammer, and Zettel, 1992: 220 (Gabrius; characters; notes; Austria).
— Assing, Frisch, Kahlen, et al., 1998: 138 (Gabrius; characters; Macedonia; Austria).
▴ Distribution: Macedonia, Austria.
roubali Smetana, 1952a: 89 (Gabrius; Type locality: Rossia mer., Caucasus occ., Adler).
— Smetana, 1960c: 343 (Gabrius; characters; Caucasus).
— Coiffait, 1974: 59 (Gabrius; characters; Caucasus).
▴ Distribution: Russia.
roubalioides Schillhammer, 1997: 15 (Gabrius; Type locality: O-Afghanistan: Do-Schak, Hindukusch, Khinjantal, 2500 m).
rougemonti Schillhammer, 1997: 51 (Gabrius; Type locality: Bali: Penulisan).
rubripennis Kiesenwetter, 1844, see: lividipes Baudi, 1848.
rufocinctus Schillhammer, 1997: 55 (Gabrius; Type locality: Nepal [Sindhupalchok District]: Prov. Bagmati, Malemchi, 2800 m).
dubiosus Schillhammer, 1997: 56 (Gabrius; subspecies of rufocinctus; Type locality: India: W. Bengal, Darjeeling dist., Tigerhill. Also cited from Nepal).
rupicola Cameron, 1943: 131 (Philonthus; subgenus Gabrius; Type locality: Darjeeling: Ghum district; [Note: See lectotype designation by Schillhammer, 1997]).
— Schillhammer, 1997: 29 (Gabrius; lectotype designation; characters; India; Nepal).
▴ Distribution: India, Nepal.
sacerdotalis Joy, 1913: 26 (Gabrius; Type locality: Caucasus, Swanetia).
— Gridelli, 1920: 149 (Gabrius; characters; Caucasus).
— Scheerpeltz, 1933: 1359 (Philonthus; catalog).
— Smetana, 1960c: 339 (Gabrius; characters; Caucasus).
— Korge, 1971a: 7 (Gabrius; Turkey).
— Coiffait, 1974: 62 (Gabrius; characters; Caucasus).
▴ Distribution: Turkey, Georgia.
sagittifer Schillhammer, 1997: 60 (Gabrius; Type locality: Nepal: [Sindhupalchok District]: Prov. Bagmati, Malemchi Khola near Malemchi, 2100 m. Also cited from India).
acutiphallus Schillhammer, 1997: 61 (Gabrius; subspecies of sagittifer; Type locality: China: Yunnan, 100 km W. Baoshan, Gaoligongshan Nat. Res.).
sallustius Smetana, 1995e: 691 (Gabrius; Type locality: Oregon: Fly Cr. Valley, SE Umatilla County. Also cited from Washington).
— Hatch, 1957: 202 (Philonthus; subgenus Gabrius; [Note: Misidentified as distans according to Smetana, 1995e: 689, 691]; characters; British Columbia; Washington).
▴ Distribution: Canada, USA.
satanellus Bordoni, 1973, see: doderoi Gridelli, 1920.
schawalleri Coiffait, 1982, see: furtivus Cameron, 1932.
schawallerianus Schillhammer, 1997: 56 (Gabrius; Type locality: Nepal: 328 Panchthar Distr., Paniporua, 2300 m).
schillhammeri Shibata, 1996: 146 (Gabrius; replacement name for multipunctatus Shibata).
multipunctatus Shibata, 1991a: 88 (Gabrius; [preoccupied]; Type locality: Japan: Yamanashi Pref.: Akazawa Spa, Masuho).
schmidti Schillhammer, 2000c: 123 (Gabrius; Type locality: Nepal Himalaya: Manaslu-Mts., Dudh Pokhari Lekh zw. Simia Kharka and Malamche Kharka, 33–3500 m).
schoedli Schillhammer, 1997: 72 (Gabrius; Type locality: Indonesia: S-Sulawesi, E. Malino, Kanre Apia, 1400 m).
schwendingeri Schillhammer, 1997: 46 (Gabrius; Type locality: Thailand: Mae Hong Son - Doi Chang, 20 km E Pai).
scrutandus Schillhammer, 1997: 35 (Gabrius; Type locality: Nepal: [Sankhua Sabha District]: Khandbari Distr., Induwa Khola Valley, 2850 m).
seattlensis Hatch, 1957: 201 (Philonthus; subgenus Gabrius; Type locality: Washington: Seattle. Also cited from British Columbia and Oregon).
— Moore, 1965: 100 (Gabrius).
— Smetana, 1995e: 669 (Gabrius; characters; notes; British Columbia; Oregon; Washington).
▴ Distribution: Canada, USA.
semipunctatus Stephens, 1833: 252 (Gabrius; Type locality: Suffolk).
senegalensis Cameron, 1949a: 320 (Gabrius; Type locality: Guinée: Mt. Nimba, alt. 700–900 m. Also cited from Ivory Coast).
septempunctatus Cameron, 1930c: 207 (Philonthus; subgenus Gabrius; Type locality: Kobe: Gomo).
— Scheerpeltz, 1933: 1360 (Philonthus; catalog).
— Cameron, 1949b: 468 (Philonthus; Formosa).
— Smetana, 1984: 122 (Gabrius; species unknown; Japan).
▴ Distribution: Japan, Taiwan.
serpentis Schillhammer, 1997: 60 (Gabrius; Type locality: Nepal: [Sindhupalchok District]: Prov. Bagmati, Malemchi, 2800 m).
sesengeanus Tottenham: [Note: Nomen nudum (Article 13.1.1); Tottenham, 1962: 143 cited this name in Gabrius, attributed it to himself, cited it from Uganda but without characters and without a reference to an earlier description].
sexualis Smetana, 1954: 42 (Gabrius; Type locality: Gallia mer., Alpes Maritimes, In. Vesubie. Also cited from Italy).
— Smetana, 1957: 63 (Gabrius; characters; France; Italy).
— Smetana, 1960c: 342 (Gabrius; characters; France; Italy).
— Bordoni, 1973i: 327, 331 (Gabrius; characters; Italy).
— Coiffait, 1974: 65 (Gabrius; characters; Alps; Italy; North Africa).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; probable subspecies of coxalus; Italy).
▴ Distribution: Italy, France, “North Africa”.
sharpianus Cameron, 1930c: 207 (Philonthus; subgenus Gabrius; Type locality: Settsu; Katsuogi: Nagasaki; Miyanoshita).
— Scheerpeltz, 1933: 1360 (Philonthus; catalog).
— Smetana, 1960c: 305 (Gabrius; characters; Japan).
— Smetana, 1984: 142 (Gabrius; characters; Japan).
▴ Distribution: Japan, China.
conicus Bernhauer, 1939d: 99 (Philonthus; subgenus Gabrius; Type locality: Japan: Tsushima. Nordwestl. China: Chinkiang).
demarcatus Tottenham, 1940c: 68 (Gabrius; Type locality: Japan: Nagasaki).
— Smetana, 1960c: 305 (Gabrius; synonym of sharpianus).
— Smetana, 1984: 142 (Gabrius; synonym of sharpianus).
shulli Hatch, 1957: 202 (Philonthus; subgenus Gabrius; Type locality: Idaho: Pierce, alt. 3200 [feet]).
— Moore, 1965: 100 (Gabrius).
— Smetana, 1995e: 700 (Gabrius; characters; notes; British Columbia; Idaho; Oregon; Washington).
▴ Distribution: Canada, USA.
linearis Casey, 1915: 439 (Philonthus; [preoccupied]; Type locality: British Columbia: Metlakatla; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1349 (Philonthus; catalog).
— Hatch, 1957: 203 (Philonthus; subgenus Gabrius; characters; British Columbia).
— Moore, 1965: 100 (Gabrius).
— Smetana, 1995e: 700 (Gabrius; lectotype designation; synonym of shulli).
siamensis Schillhammer, 1997: 52 (Gabrius; Type locality: Thailand: prov. Chiang Mai, Doi Inthanon, 1780 m).
signiceps Smetana, 1954, see: nigritulus Gravenhorst, 1802.
simulans Wollaston, 1857: 190 (Philonthus; Type locality: Madeira Islands).
— Wollaston, 1865: 494 (Philonthus; habitat; Madeira Islands; Canary Islands).
— Fauvel, 1902b: 111 (Philonthus; Madeira).
— Bernhauer and Schubert, 1914: 331 (Philonthus; synonym of canariensis).
— Bernhauer and Schubert, 1914: 355 (Philonthus; catalog).
— Coiffait, 1954: 165 (Gabrius; cited as “caneriensis Fauvel simulans Wollaston”; [Note: Misidentified as canariensis according to Smetana, 1960c: 307]; Canary Islands).
— Smetana, 1962: 98 (Gabrius; characters; Canary Islands).
— Smetana, 1963b: 37 (Gabrius; notes; characters; Madeira).
— Coiffait, 1974: 62 (Gabrius; characters; Madeira).
▴ Distribution: Madeira.
maderensis Coiffait, 1963: 21 (Gabrius; Type locality: Madère: Rabaçal).
smetanai Schillhammer, 1997: 22 (Gabrius; Type locality: Nepal: [Sankhua Sabha District], Khandbari District, above Tashigaon, 3550 m).
sodalis Cameron, 1943, see: parkeri Cameron, 1932.
solomonicus Schillhammer, 1997: 64 (Gabrius; Type locality: Solomon Islands: Guadalcanal, Popamanasiu, 5700 feet).
spatulipenis Schillhammer, 1997: 57 (Gabrius; Type locality: Nepal: Manang District, Chame, 2200–2650 m).
sphagnicola Sjöberg, 1950: 159 (Philonthus; subgenus Gabrius; Type locality: Schweden: Waldsees in Loos, Hälsingland).
— Smetana, 1957: 66 (Gabrius; characters; Sweden; central Europe).
— Smetana, 1960c: 348 (Gabrius; characters; Sweden; Scandinavia).
— Szujecki, 1963 (Gabrius; characters; habitat; Poland).
— Palm, 1963b: 105 (Philonthus; subgenus Gabrius; characters; Sweden; Finland).
— Lohse, 1964: 189 (Gabrius; characters; central Europe).
— Horion, 1965: 190 (Gabrius; Ireland; Britain; Sweden; Holland; Belgium; France; Switzerland; Corsica; Italy; Romania; Slovakia; Ukraine; Germany; Austria).
— Huggert and Ulefors, 1971: 58 (Philonthus; notes; Sweden).
— Coiffait, 1974: 58 (Gabrius; characters; Scandinavia).
— Lucht, 1987: 107 (Gabrius; checklist; Germany; Poland; Sweden).
— Lohse and Lucht, 1989: 168 (Gabrius; note).
▴ Distribution: Europe.
splendidulus Gravenhorst, 1802: 41 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae; [Note: See lectotype designation by Smetana, 1962c]).
— Latreille, 1804: 327 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 93 (Staphylinus; characters).
— Gyllenhal, 1810: 348 (Staphylinus; characters; Sweden).
— C. Sahlberg, 1830: 330 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 32 (Cafius; Finland).
— Mannerheim, 1831: 446 (Cafius; Finland).
— Nordmann, 1837: 126 (Gyrohypnus; Germany; Sweden; France).
— Erichson, 1839a: 470 (Philonthus; characters; Germany).
— Heer, 1839: 269 (Philonthus; characters; Switzerland).
— Erichson, 1840: 478 (Philonthus; characters; Europe).
— Jacobsen, 1849: 701 (Philonthus; characters; Denmark).
— Hochhuth, 1849: 142 (Philonthus; characters; Iberia).
— Redtenbacher, 1849: 705 (Philonthus; characters; Austria).
— Hardy, 1851: 30 (Philonthus; notes; England).
— Rosenhauer, 1856: 71 (Philonthus; Cartama).
— Fairmaire and Laboulbène, 1856: 522 (Philonthus; characters; France).
— Kraatz, 1857d: 608 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 195 (Philonthus; characters; Austria).
— Kraatz, 1858a: 61 (Philonthus; Greece).
— Thomson, 1860: 170 (Gabrius; characters; Scandinavia).
— Hochhuth, 1862: 62 (Philonthus; Russia).
— Fauvel, 1874: 239 [= 1874a: 465] (Philonthus; characters; France; Caucasus).
— Redtenbacher, 1874: 214 (Philonthus; characters; Austria).
— Seidlitz, 1875: 277 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 544 [= 1877: 400] (Gabrius; characters; France).
— J. Sahlberg, 1876: 23 (Gabrius; Finland).
— Fowler, 1888: 275 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 459 (Philonthus; subgenus Gabrius; characters; Europe; Caucasus).
— Reitter, 1909: 131 (Philonthus; subgenus Gabrius; characters; Germany).
— Bernhauer and Schubert, 1914: 356 (Philonthus; catalog).
— Johansen, 1914: 416 (Philonthus; subgenus Gabrius; characters; Denmark).
— Gridelli, 1920: 134 (Gabrius; characters; Italy).
— Šulc, 1922 (Gabrius; osmeterium; anatomy; histology; physiology).
— Gridelli, 1928: 50 (Gabrius; characters in key).
— Portevin, 1929: 365 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1362 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Scheerpeltz and Höfler, 1948: 157 (Gabrius; fungus inhabitant).
— Coiffait, 1951: 108, 114 (Gabrius; characters in key; France).
— Smetana, 1952: 122 (Gabrius; illustrations).
— V. Hansen, 1952: 86 (Philonthus; subgenus Gabrius; characters; Denmark).
— Palm, 1956: 40, 41 (Philonthus; subgenus Gabrius; characters; notes).
— Smetana, 1957: 61 (Gabrius; characters; Caucasus to Siberia; central Europe).
— Smetana, 1958: 242 (Gabrius; characters; ecology; Czechoslovakia).
— Smetana, 1962c: 237 (Gabrius; lectotype designation; notes on specimens in Gravenhorst collection).
— Palm, 1963b: 103 (Philonthus; subgenus Gabrius; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 188 (Gabrius; characters; central Europe).
— Horion, 1965: 189 (Gabrius; all Europe).
— Pototskaia, 1966: 154 (Gabrius; larval characters).
— Smetana, 1967b: 156 (Gabrius; Spain).
— Pototskaia, 1967: 53 (Gabrius; larval characters).
— Szujecki, 1968a: 732 (Gabrius; Poland).
— Coiffait, 1974: 66 (Gabrius; characters; Europe; Scandinavia; Caucasus; Siberia).
— Pototskaia, 1976: 173 (Gabrius; larval characters).
— Pope, 1977: 30 (Gabrius; Britain).
— Coiffait, 1978d: 164 (Gabrius; Turkey).
— Tóth, 1984: 85 (Gabrius; characters; Hungary).
— Lucht, 1987: 106 (Gabrius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Janák and Vysoky, 1992: 137 (Gabrius; collected with Formica rufa and truncorum; Bohemia).
— Welch, 1993: 229 (Gabrius; ovariole number and ovary structure).
— Smetana, 1993b: 49 (Gabrius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
— Smetana, 1995e: 673 (Gabrius; characters; notes; British Columbia; Maryland).
— Fowles, Alexander, and Key, 1999: 129 (Gabrius; saproxylic species; status as endangered or threatened species; Britain).
— Lott, 1999: 72 (Gabrius; saproxylic species; habitat; endangered status; Britain).
▴ Distribution: Europe, Russia, Caucasus, Canada, USA.
parvulus Gravenhorst, 1802: 42 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).
— Latreille, 1804: 328 (Staphylinus; characters).
— Gravenhorst, 1806: 95 (Staphylinus; characters; Germany).
— Mannerheim, 1830: 34 (Gyrohypnus; Finland).
— Mannerheim, 1831: 448 (Gyrohypnus; Finland).
— Nordmann, 1837: 120 (Gyrohypnus; Germany; Russia).
— Erichson, 1839a: 470 (Philonthus; synonym of splendidulus).
— Erichson, 1840: 479 (Philonthus; synonym of splendidulus).
— Fairmaire and Laboulbène, 1856: 522 (Philonthus; synonym of splendidulus).
— Kraatz, 1857d: 608 (Philonthus; synonym of splendidulus).
— Ganglbauer, 1895: 459 (Philonthus; synonym of splendidulus).
— Bernhauer and Schubert, 1914: 356 (Philonthus; synonym of splendidulus).
— Smetana, 1958: 242 (Gabrius; synonym of splendidulus).
— Coiffait, 1974: 66 (Gabrius; synonym of splendidulus).
— Smetana, 1995e: 673 (Gabrius; synonym of splendidulus).
nanus Gravenhorst, 1806: 96 (Staphylinus; [preoccupied]; Type locality: Not cited).
— Gyllenhal, 1810: 347 (Staphylinus; characters; Sweden).
— Zetterstedt, 1828: 79 (Staphylinus; characters; Lapland).
— Curtis, 1829: 26 (Gabrius; catalog; Britain).
— C. Sahlberg, 1830: 330 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 32 (Cafius; Finland; Sweden).
— Mannerheim, 1831: 446 (Cafius; Finland; Sweden).
— Stephens, 1833: 251 (Gabrius; characters; England).
— Runde, 1835: 10 (Cafius; characters; Germany).
— Nordmann, 1837: 126 (Gyrohypnus; Germany; Sweden; France).
— Zetterstedt, 1838: 65 (Staphylinus; characters; habitat; Lapland).
— Erichson, 1839a: 470 (Philonthus; synonym of splendidulus).
— Stephens, 1839: 400 (Gabrius; characters; England).
— Erichson, 1840: 479 (Philonthus; synonym of splendidulus).
— Laporte, 1840: 179 (Cafius; characters; France).
— Hardy, 1851: 30 (Philonthus; synonym of splendidulus).
— Fairmaire and Laboulbène, 1856: 522 (Philonthus; synonym of splendidulus).
— Kraatz, 1857d: 608 (Philonthus; synonym of splendidulus).
— Thomson, 1860: 170 (Gabrius; synonym of splendidulus).
— Ganglbauer, 1895: 459 (Philonthus; synonym of splendidulus).
— Bernhauer and Schubert, 1914: 356 (Philonthus; synonym of splendidulus).
— Smetana, 1958: 242 (Gabrius; synonym of splendidulus).
— Coiffait, 1974: 66 (Gabrius; synonym of splendidulus).
— Smetana, 1995e: 673 (Gabrius; synonym of splendidulus).
albipes Stephens, 1833: 252 (Gabrius; Type locality: near London).
— Stephens, 1839: 400 (Gabrius; characters; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of nigritulus).
— Hardy, 1851: 30 (Philonthus; synonym of splendidulus).
analis Heer, 1839: 268 (Philonthus; Type locality: Matt; Zürich; Schaffhausen).
— Kraatz, 1857d: 596 (Philonthus).
— Ganglbauer, 1895: 459 (Philonthus; synonym of splendidulus).
— Bernhauer and Schubert, 1914: 356 (Philonthus; synonym of splendidulus).
— Smetana, 1958: 242 (Gabrius; synonym of splendidulus).
— Coiffait, 1974: 66 (Gabrius; synonym of splendidulus).
— Smetana, 1995e: 673 (Gabrius; synonym of splendidulus).
spurius Smetana, 1954, see: ravasinii Gridelli, 1920.
stipes Sharp, 1910, see: piliger Mulsant and Rey, 1876.
suaveolens Stephens, 1833, see: osseticus Kolenati, 1846.
subdepressus Bernhauer, 1939d: 97 (Philonthus; Type locality: Nordwestl. China: Chinkiang. Japan: Unzen bei Shimabara).
— Smetana, 1973b: 130 (Gabrius; characters; notes).
— Smetana, 1984: 125 (Gabrius; characters; Japan).
▴ Distribution: China, Japan.
submetallicus Cameron, 1933b: 346 (Philonthus; subgenus Gabrius; Type locality: British North Borneo: Lumu Lumu, 5500 feet; [Note: See lectotype designation by Schillhammer, 1997]).
— Hammond, 1984: 194, 195 (Gabrius; Borneo).
— Schillhammer, 1997: 62 (Gabrius; lectotype designation; characters; Malaysia).
▴ Distribution: Borneo, Malaysia.
subnigrituloides Scheerpeltz, 1933, see: subnigritulus Joy, 1913.
subnigritulus Reitter, 1909 [Note: Nomen nudum], see: appendiculatus Sharp, 1910.
subnigritulus Joy, 1913: 26 (Gabrius; Type locality: Caucasus).
— Scheerpeltz, 1933: 1363 (Philonthus; synonym of subnigrituloides).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Smetana, 1953: 122 (Gabrius; Turkey).
— Smetana, 1953a: 178 (Gabrius; Caucasus; Austria; Bulgaria; Hungary; Moravia).
— Coiffait, 1974: 57 (Gabrius; synonym of subnigrituloides).
▴ Distribution: Europe, Russia, Caucasus, Turkey, Kazakhstan.
subnigrituloides Scheerpeltz, 1933: 1363 (Philonthus; replacement name for subnigritulus Joy).
— Smetana, 1952b: 118 (Gabrius; [Note: Misidentified as tornus according to Smetana, 1958: 246]; characters; Czechoslovakia).
— Smetana, 1957: 67, 68 (Gabrius; characters; Caucasus; Transcaucasia; Slovakia).
— Smetana, 1958: 246 (Gabrius; characters; ecology; Czechoslovakia).
— Smetana, 1960c: 318 (Gabrius; characters; Caucasus; Transcaucasian region; Daghestan; Slovakia).
— Lohse, 1964: 190 (Gabrius; characters; central Europe).
— Smetana, 1965f: 258 (Gabrius; Aulie Atta).
— Horion, 1965: 194 (Gabrius; Slovakia).
— Coiffait, 1974: 57 (Gabrius; characters; Caucasus; Transcaucasus; Daghestan; central Europe; Maritime Alpes; Corsica).
— Tóth, 1984: 90 (Gabrius; characters).
— Lucht, 1987: 106 (Gabrius; checklist; Czechoslovakia; Austria).
— Silfverberg, 1991: 21 (Gabrius; unnecessary replacement name that was proposed by Scheerpeltz who thought that Reitter's subnigritulus was available; Reitter's name was described as an aberration and is unavailable).
— Smetana, 1993b: 49 (Gabrius; Slovakia).
subnigritulus Smetana, 1956, see: appendiculatus Sharp, 1910.
successorius Schillhammer, 1997: 40 (Gabrius; Type locality: Nepal: [Sindhupalchok District], Prov. Bagmati, Malemchi, 2800 m).
suffragani Joy, 1913: 26 (Gabrius; Type locality: Dalmatia).
— Gridelli, 1920: 147 (Gabrius; characters; Transylvania; Dalmatia).
— Scheerpeltz, 1933: 1363 (Philonthus; catalog).
— Wüsthoff, 1937: 31 (Philonthus; illustration of aedeagus).
— Smetana, 1952b: 124 (Gabrius; characters; Czechoslovakia).
— Smetana, 1957: 64, 70 (Gabrius; characters; Dalmatia; Bosnia; Romania; Hungary; Austria; Czechoslovakia).
— Smetana, 1958: 253 (Gabrius; characters; ecology; Czechoslovakia).
— Smetana, 1960c: 338 (Gabrius; characters; Austria; Albania; Dalmatia; Bosnia; Romania; Czechoslovakia; Austria; Hungary).
— Lohse, 1964: 191 (Gabrius; characters; central Europe).
— Horion, 1965: 196 (Gabrius; Dalmatia; Bosnia; Romania; Hungary; Slovakia; Austria; Albania; Switzerland; Germany).
— Smetana, 1966: 331 (Gabrius; Czechoslovakia).
— Raianu, 1969a: 326 (Gabrius; characters; ecological notes; Romania).
— Korge, 1971a: 7 (Gabrius; Turkey).
— Coiffait, 1974: 61 (Gabrius; characters; Europe; Balkans; France).
— Tóth, 1984: 92 (Gabrius; characters; Hungary).
— Lucht, 1987: 106 (Gabrius; checklist; Czechoslovakia; Austria).
— Smetana, 1993b: 49 (Gabrius; Slovakia).
— Burakowski, Mroczkowski, and Stefańska, 2000: 52 (Gabrius; catalog; Poland).
▴ Distribution: Europe, Turkey.
talangensis Schillhammer, 1997: 69 (Gabrius; Type locality: Indonesia: W-Sumatra: 70 km SE Padang Gg. Talang, 1500 m).
taplejungensis Schillhammer, 1997: 53 (Gabrius; Type locality: Nepal: 356 Taplejung Dist., Omje Kharka, NW Yamputhin, 2300–2500 m).
tenganominus Tottenham, 1962: 251 (Gabrius; Type locality: Tanganyika Terr.: Mt. Hanang, versant Sud, gorge du Himit, 2200–2300 m; [Note: Cited on page 142 as tenganomimus]).
tenganus Tottenham, 1956, see: katonae Bernhauer, 1915.
tensicornis Scheerpeltz, 1965, see: deceptor Cameron, 1932.
thailandicus Schillhammer, 1997: 47 (Gabrius; Type locality: Thailand: Chiang Mai, Doi Inthanon, 1650 m).
thermicola Cameron, 1947: 3 (Philonthus; subgenus Gabrius; Type locality: Nizi: Blukwa).
tibialis Mulsant and Rey, 1876, see: pisciformis Fauvel, 1874.
tichomirovae Bogdanov, 1984, see: exspectatus Smetana, 1952.
tilakholensis Schillhammer, 1997: 33 (Gabrius; Type locality: Nepal [Jumla District], Ufer des Tila Khola b. Jumla, Gebiet von Jumla, Westnepal).
tirolensis Luze, 1903d: 386 (Philonthus; subgenus Gabrius; Type locality: Osttirol, Taufers, 850 m Seehöhe).
— Bernhauer and Schubert, 1914: 329 (Philonthus; variety of astutus).
— Scheerpeltz, 1933: 1364 (Philonthus; catalog).
— Wüsthoff, 1937: 32 (Philonthus; illustration of aedeagus).
— Coiffait, 1951: 107, 114 (Gabrius; characters in key; France).
— Coiffait, 1953a: 265 (Gabrius; characters; Spain).
— Smetana, 1953a: 178 (Gabrius; Bavaria; Savinian Alps; Croatia).
— Smetana, 1957: 60, 62 (Gabrius; characters; Pyrenees; Switzerland; Italy; Austria; Czech Republic; Slovakia; Germany).
— Smetana, 1958: 240 (Gabrius; characters; ecology; Czechoslovakia).
— Lohse, 1964: 187 (Gabrius; characters; central Europe).
— Smetana, 1964c: 167 (Gabrius; Czechoslovakia).
— Horion, 1965: 187 (Gabrius; Slovenia; France; Switzerland; Italy; Germany; Austria; Slovakia).
— Smetana, 1967b: 156 (Gabrius; Spain).
— Coiffait, 1970d: 143 (Gabrius; Uzbekistan).
— Coiffait, 1974: 81 (Gabrius; characters; Alps; Pyrenees).
— Tóth, 1984: 83 (Gabrius; characters).
— Lucht, 1987: 106 (Gabrius; checklist; Germany; Poland; Czechoslovakia; Austria; Sweden).
— Pilon and Zanetti, 1991: 56 (Gabrius; Italy).
— Smetana, 1993b: 49 (Gabrius; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
— Sagvolden and Hansen, 1996: 91 (Gabrius; Norway).
— Gillerfors, 1997: 85 (Gabrius; characters).
— Renner, 1998: 92 (Gabrius; Germany).
▴ Distribution: Europe, Uzbekistan.
tokatensis Smetana, 1977: 792 (Gabrius; Type locality: Turquie Tokat, Tokat-Almus).
amasiensis Coiffait, 1980: 252 (Gabrius; Type locality: Turquie: environs de Amasia).
tolkieni Schillhammer, 1997: 36 (Gabrius; Type locality: Nepal: [Sankhua Sabha District]: Khandbari Distr., Induwa Khola Valley, 2000 m. Also cited from India).
tornus Joy, 1913: 26 (Gabrius; Type locality: Caucasus, Helenendorf).
— Gridelli, 1920: 144 (Gabrius; characters; Caucasus).
— Scheerpeltz, 1933: 1364 (Philonthus; catalog).
— Smetana, 1953a: 178 (Gabrius; Aulie Atta).
— Smetana, 1960c: 320 (Gabrius; characters; Caucasus).
— Coiffait, 1974: 57 (Gabrius; characters; Caucasus).
▴ Distribution: Caucasus, Kazakhstan.
toxotes Joy, 1913: 26 (Gabrius; Type locality: Dalmatia).
— Gridelli, 1920: 152 (Gabrius; characters; Transylvania).
— Scheerpeltz, 1933: 1364 (Philonthus; catalog).
— Wüsthoff, 1937: 31 (Philonthus; cited as toxotus; illustration of aedeagus).
— Fagel, 1945: 42 (Gabrius; characters; Belgium).
— Coiffait, 1951: 113, 115 (Gabrius; characters in key; France).
— Smetana, 1952b: 127 (Gabrius; characters; Czechoslovakia).
— Smetana, 1953a: 179 (Gabrius; eastern Siberia; Savinian Alps).
— Smetana, 1957: 65, 69 (Gabrius; characters; France; England; Scandinavia; Belgium; Holland; Germany; Czechoslovakia; Austria; Italy; Dalmatia; Romania; eastern Asia).
— Smetana, 1958: 256 (Gabrius; characters; ecology; Czechoslovakia).
— Smetana, 1960c: 347 (Gabrius; characters; Dalmatia; Belgium; Holland; Czechoslovakia; Germany; Austria; Italy; Romania; Siberia).
— Lohse, 1964: 191 (Gabrius; characters; central Europe).
— Horion, 1965: 195 (Gabrius; Dalmatia; Romania; Czech Republic; Slovakia; Italy; Switzerland; France; Belgium; Holland; Austria; Germany).
— Smetana, 1966: 331 (Gabrius; notes; Czechoslovakia).
— Lundberg, 1972: 173 (Philonthus; Sweden).
— Bordoni, 1973i: 329 (Gabrius; characters).
— Coiffait, 1974: 58 (Gabrius; characters; Europe; Siberia; Balkans; France; Belgium; Holland).
— Tóth, 1984: 92 (Gabrius; characters).
— Lucht, 1987: 107 (Gabrius; checklist; Germany; Czechoslovakia; Austria; Switzerland; Benelux; Sweden).
— Pilon and Zanetti, 1991: 56 (Gabrius; Italy).
— Smetana, 1993b: 49 (Gabrius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
▴ Distribution: Europe, Russia.
trifidus Schillhammer, 1992: 64 (Gabrius; Type locality: China: Gansu, Meijishan).
trigonalis Tottenham, 1956: 296 (Gabrius; Type locality: Ruanda: Tshuruyaga, forêt du Rugege, 2400 m).
trossuloides Cameron, 1933d: 389 (Philonthus; subgenus Gabrius; Type locality: Sumatra: Fort de Kock; [Note: See lectotype designation by Schillhammer, 2000c]).
— Schillhammer, 1999a: 62 (Gabrius).
— Schillhammer, 2000c: 134 (Gabrius; lectotype designation: characters; Sumatra; may also be from Taiwan).
▴ Distribution: Indonesia.
trossuloides Coiffait, 1966, see: trossuliformis Schillhammer, 1999.
trossuliformis Schillhammer, 1999a: 62 (Gabrius; replacement name for trossuloides Coiffait).
trossuloides Coiffait, 1966c: 507 (Gabrius; [preoccupied]; Type locality: Arménie russe, Sevan, 3000 m).
— Coiffait, 1974: 83 (Gabrius; characters; Armenia).
— Schillhammer, 1999a: 62 (Gabrius; synonym of trossuliformis).
trossulus Nordmann, 1837: 102 (Philonthus; Type locality: Berolinum).
— Erichson, 1839a: 476 (Philonthus; synonym of aterrimus).
— Erichson, 1840: 492 (Philonthus; synonym of aterrimus).
— P. Lucas, 1846: 113 (Philonthus; synonym of aterrimus).
— Staudinger, 1857: 286 (Philonthus; notes; Iceland).
— Kraatz, 1857d: 616 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 985 (Philonthus; characters; Germany).
— Kraatz, 1858a: 61 (Philonthus; Greece).
— Thomson, 1860: 171 (Gabrius; characters; Scandinavia).
— Fauvel, 1865a: 297 [= 1865b: 47] (Philonthus; habitat; France).
— Fauvel, 1874: 243 [= 1874a: 469] (Philonthus; synonym of nigritulus).
— Redtenbacher, 1874: 217 (Philonthus; characters).
— Seidlitz, 1875: 278 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 557 [= 1877: 413] (Gabrius; characters; Sweden; Germany).
— Fowler, 1888: 279 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 460 (Philonthus; subgenus Gabrius; characters; north Europe; Germany; Bohemia).
— Reitter, 1909: 132 (Philonthus; subgenus Gabrius; characters; Germany).
— Bernhauer and Schubert, 1914: 348 (Philonthus; variety of nigritulus).
— Johansen, 1914: 419 (Philonthus; subgenus Gabrius; characters; Denmark).
— Gridelli, 1920: 143 (Gabrius; characters).
— Scheerpeltz, 1933: 1365 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Wüsthoff, 1937: 31 (Philonthus; illustration of aedeagus).
— Fagel, 1945: 40 (Gabrius; characters; Belgium).
— Coiffait, 1951: 110, 114 (Gabrius; characters in key; France).
— Smetana, 1952b: 116 (Gabrius; characters; Czechoslovakia).
— V. Hansen, 1952: 88 (Philonthus; subgenus Gabrius; characters; Denmark).
— Smetana, 1957: 62, 68 (Gabrius; characters; Iceland; Scandinavia; western and central Europe).
— Smetana, 1958: 245 (Gabrius; characters; ecology; Czechoslovakia).
— Larsson and Gígja, 1959: 86 (Philonthus; biological notes; Iceland).
— Smetana, 1960c: 308 (Gabrius; characters; Scandinavia; Iceland; England; Scotland; France; Belgium; Holland; Germany; Czechoslovakia; Poland; Austria; Hungary).
— Smetana, 1961a: 44 (Gabrius; Bulgaria).
— Szujecki, 1963a: 34 (Gabrius; Poland).
— Palm, 1963b: 105 (Philonthus; subgenus Gabrius; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 190 (Gabrius; characters; central Europe).
— Smetana, 1964d: 77 (Gabrius; Czechoslovakia).
— Horion, 1965: 194 (Gabrius; Russia; Lapland; North Cape; Britain; Holland; Belgium; France; Germany; Austria; Switzerland; Italy; Czech Republic; Slovakia; Poland; Hungary; Romania).
— Pototskaia, 1966: 154 (Gabrius; larval characters).
— Pototskaia, 1967: 57 (Gabrius; larval characters).
— Coiffait, 1974: 83 (Gabrius; characters; Europe; Britain; Scandinavia).
— Pototskaia, 1976: 173 (Gabrius; larval characters).
— Pope, 1977: 30 (Gabrius; Britain).
— Frank, 1982: 36 (Gabrius; list of parasites).
— Tóth, 1984: 89 (Gabrius; characters; Hungary).
— Lucht, 1987: 106 (Gabrius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Welch, 1993: 229 (Gabrius; ovariole number and ovary structure).
— Smetana, 1993b: 49 (Gabrius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Gabrius; Italy).
— Sadler and Dugmore, 1995: 144 (Gabrius; habitat notes; Iceland).
▴ Distribution: Iceland, Europe.
tsaratananus Jarrige, 1970: 49 (Gabrius; Type locality: Madagascar: Mt. Tsaratanana, 15–1600 m).
tschinganus Coiffait, 1970, see: lebedevi Bernhauer, 1910.
turcmenus Fauvel, 1900d: 232 (Philonthus; Type locality: Buchara; [Note: See lectotype designation by Schillhammer, 1997]).
— Bernhauer and Schubert, 1914: 358 (Philonthus; catalog).
— Scheerpeltz, 1933: 1365 (Philonthus; catalog).
— Gridelli, 1934: 77 (Gabrius; characters; notes; Karakoram region).
— Smetana, 1973b: 128 (Gabrius; characters).
— Coiffait, 1974: 63 (Gabrius; characters; Turkestan).
— Schillhammer, 1997: 17 (Gabrius; lectotype designation; characters; Afghanistan; Pakistan; Tadjikistan; Uzbekistan).
▴ Distribution: Uzbekistan, Tajikistan, Afghanistan, Pakistan.
turneri Cameron, 1951, see: pullatus Herman, 2001.
ulpius Smetana, 1995e: 653 (Gabrius; Type locality: Quebec: Rigaud. Also cited from Nova Scotia, Ontario, Quebec, Georgia, Michigan, New Hampshire, New York, North Carolina, Ohio, West Virginia).
ultimus Schillhammer, 1997: 12 (Gabrius; Type locality: Nepal: [Patan District], Kathmandu District, Phulcoki, 2600 m).
unculus Tottenham, 1956b: 222 (Gabrius; Type locality: S. Rhodesia: Mashonaland, Salisbury).
unzenensis Bernhauer, 1939e: 153 (Philonthus; subgenus Gabrius; Type locality: Japan: Unzen bei Shimabara).
— Smetana, 1973b: 129 (Gabrius; characters; notes).
— Smetana, 1984: 125 (Gabrius; characters; Japan).
— Cho and Lee, 1997: 272 (Gabrius; characters; Korea).
▴ Distribution: Japan.
ussuriensis Schillhammer, 1991: 53 (Gabrius; Type locality: USSR: Nikolsk Ussurijsk, Ussurigebirge).
velox Sharp, 1910: 130 (Gabrius; Type locality: Anglia: ad flumen Lymington).
— Bernhauer and Schubert, 1914: 359 (Philonthus; catalog).
— Gridelli, 1920: 140 (Gabrius; characters; Englnad).
— Scheerpeltz, 1933: 1367 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Wüsthoff, 1937: 32 (Philonthus; illustration of aedeagus).
— Fagel, 1945: 41 (Gabrius; characters; Belgium).
— Coiffait, 1951: 111, 115 (Gabrius; characters in key; France).
— V. Hansen, 1952: 90 (Philonthus; subgenus Gabrius; characters; Denmark).
— Smetana, 1952b: 123 (Gabrius; characters; Czechoslovakia).
— Smetana, 1957: 65, 69 (Gabrius; characters; England; France; Belgium; Holland; Germany; Czechoslovakia; Austria; Scandinavia).
— Smetana, 1958: 251 (Gabrius; characters; ecology; Czechoslovakia).
— Smetana, 1960c: 333 (Gabrius; characters; England; Germany; Czechoslovakia; Austria).
— Palm, 1963b: 106 (Philonthus; subgenus Gabrius; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 190 (Gabrius; characters; central Europe).
— Horion, 1965: 192 (Gabrius; Britain; Norway; Sweden; Finland; Russia; Denmark; Holland; Belgium; France; Czechoslovakia; Hungary; Austria; Germany).
— Smetana, 1966: 331 (Gabrius; Czechoslovakia).
— Coiffait, 1974: 60 (Gabrius; characters; Europe; Britain; France).
— Pope, 1977: 30 (Gabrius; Britain).
— Tóth, 1984: 89 (Gabrius; characters).
— Lucht, 1987: 106 (Gabrius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Lohse and Lucht, 1989: 169 (Gabrius; note).
— Smetana, 1993b: 49 (Gabrius; Czech Republic; Slovakia).
— Smetana, 1995e: 626 (Gabrius; characters; notes; Maryland).
▴ Distribution: Europe, USA.
austriacus Scheerpeltz, 1947: 283 (Gabrius; Type locality: Österreich: Marchfeld, Niederösterreich).
— Smetana, 1952b: 124 (Gabrius; characters).
— Smetana, 1957: 65, 69 (Gabrius; characters; Austria).
— Smetana, 1958: 253 (Gabrius; characters; ecology; Czechoslovakia).
— Smetana, 1960c: 335 (Gabrius; characters; Austria).
— Lohse, 1964: 190 (Gabrius; characters; central Europe).
— Horion, 1965: 193 (Gabrius; Austria).
— Coiffait, 1974: 59 (Gabrius; characters; Austria).
— Tóth, 1984: 89 (Gabrius; characters).
— Lucht, 1987: 106 (Gabrius; checklist; Austria).
— Lohse and Lucht, 1989: 169 (Gabrius; synonym of velox).
— Smetana, 1995e: 626 (Gabrius; synonym of velox).
ventralis Stephens, 1833: 252 (Gabrius; [Note: Nomen dubium]; Type locality: Suffolk, and on the shores of the Severn, near Bristol).
vernalis Gravenhorst, 1806, see: osseticus Kolenati, 1846.
vibius Smetana, 1995e: 681 (Gabrius; Type locality: Quebec: Rigaud. Also cited from New Hampshire, New York, Ohio, Pennsylvania, Vermont, West Virginia).
vicinus Cameron, 1932: 149 (Philonthus; Type locality: Simla Hills: Kotgarh, alt. 7000 feet; [Note: See lectotype designation by Schillhammer, 1997]).
— Scheerpeltz, 1933: 1367 (Philonthus; catalog).
— Schillhammer, 1997: 47 (Gabrius; lectotype designation; characters; India; Nepal; Pakistan).
▴ Distribution: Pakistan, India, Nepal.
viduus Cameron, 1933b: 346 (Philonthus; subgenus Gabrius; [preoccupied]; Type locality: British North Borneo, Mt. Kinabalu, Kamborangah, 7200 feet; [Note: See lectotype designation by Schillhammer, 1997]. Also cited from Lumu Lumu, 5500 feet).
— Cameron, 1937: 6 (Philonthus; Java).
— Hammond, 1984: 194, 195 (Gabrius; Borneo).
— Schillhammer, 1997: 63 (Gabrius; lectotype designation: British North Borneo, Mt. Kinabalu, Kamborangah, 7200 feet; characters; Malaysia).
— Herman, 2001: 44 (Gabrius; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Borneo, Malaysia, Indonesia.
villosulus Stephens, 1833, see: Neobisnius.
vindex Smetana, 1995e: 648 (Gabrius; Type locality: Quebec: Gatineau Pk., Blind Lake. Also cited from Manitoba, Alaska, and Minnesota).
— Smetana, 1965: 29 (Gabrius; [Note: Misidentified as seattlensis according to Smetana, 1995e: 648]; characters; Newfoundland).
▴ Distribution: Canada, USA.
virilis Horn, 1884: 215 (Philonthus; Type locality: Vancouver; [Note: See lectotype designation by Smetana, 1995e]).
— Bernhauer and Schubert, 1914: 361 (Philonthus; catalog).
— Hatch, 1957: 200 (Philonthus; subgenus Gabrius; characters; British Columbia; Washington; Oregon).
— Moore, 1965: 100 (Gabrius).
— Smetana, 1995e: 705 (Gabrius; lectotype designation; characters; notes; British Columbia; Oregon; Washington).
▴ Distribution: Canada, USA.
visokeanus Cameron, 1950g: 39 (Philonthus; subgenus Gabrius; Type locality: Belgian Congo: Visoke, alt. 2800–3300 m).
vulcanicola Schillhammer, 1997: 65 (Gabrius; Type locality: Java: W Java, Mt. Gede, 50 km SE Bogor, 2600 m).
vulcanus Cameron, 1950g: 40 (Philonthus; subgenus Gabrius; Type locality: Belgian Congo: Shamuheru, volc. Nyamuragira, alt. 1843 m).
wawawai Hatch, 1957, see: punctatellus Horn, 1884.
xanthipes Schillhammer, 1997: 43 (Gabrius; Type locality: Nepal: [Sindhupalchok District], Prov. Bagmati, Gul Bhanjyang, 2600 m).
yakinius Tottenham, 1956b: 210 (Gabrius; Type locality: Cape Colony: Albany dist.).
yamanei Smetana, 1984: 130 (Gabrius; Type locality: Japan: Gumma Pref.: 5 km E Usui Pass, 900 m).
zerchei Schillhammer, 1996: 65 (Gabrius; Type locality: Russia: Primorskiy Krai Ryazanovka, 14 km SW Slavyanka, 42.48 N, 131.12 E, 50 m).
Gabronthus
[39 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions]
Gabronthus Tottenham, 1955: 180 (species included: maritimus, alluaudanus, pluvialis, thermarum, kisiranus, oribates, tenuissimus, minutissimus, turneri, gombanus, kosanus chyuluensis, fundator, mgogoricus, badalus, meridionalis, styx, inclinans, corrugatus, thermaroides, sulcifrons, pygmaeus, minusculus). Type species: Gabrius maritimus Motschulsky, fixed by original designation.
— Tottenham, 1955: 181, 190 (keys to African and Oriental species).
— Smetana, 1959: 172 (characters).
— Coiffait, 1963: 7, 12 (characters; notes; thermarum cited as type species).
— Lohse, 1964: 186 (characters; key to central European species).
— Moore, 1965: 97, 99 (characters; notes).
— Shibata, 1973c: 60 (checklist of species of Taiwan).
— Bordoni, 1973j: 707 (list of species from islands near Sicily).
— Coiffait, 1974: 111 (characters; key to species of western Palaearctic region).
— Moore and Legner, 1975: 15 (catalog; North America north of Mexico).
— Pope, 1977: 30 (list of British species).
— Moore and Legner, 1979: 21 (characters; notes).
— Muona, 1979: 15 (list of Scandinavian species).
— Burakowski, Mroczkowski, and Stefańska, 1980: 79 (catalog; Poland).
— Shibata, 1983: 124 (checklist of species of Japan).
— Tóth, 1984: 80 (characters; habitus; key to species of Hungary).
— Frank, 1986a: 370 (checklist of species of Florida).
— Drugmand 1987b (comments on distribution of Belgian species).
— Ádám, 1987: 151 (Gabronthus; collecting notes; Hungary).
— M. Hansen, Kristensen, Mahler, Pedersen, 1991: 107 (list of Danish species).
— Silfverberg, 1992: 16 (list of species of Fennoscandia, Denmark, and the Baltic States).
— Smetana, 1993b: 48 (list of species of Czech Republic and Slovakia).
— Smetana, 1995e: 478 (characters; key to North American species; notes).
— M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 29 (additional records to Danish list of species).
— Ciceroni and Zanetti, 1995: 29 (list of species of Italy).
— M. Hansen, 1996: 37, 106 (probably belongs within Philonthus; list of Danish species).
— Downie and Arnett, 1996: 383 (characters).
— M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (list of species collected in 1997; Denmark).
— Lundgren, 1998: 51 (list of species from Florida; records for other States listed for each species).
— Newton, Thayer, Ashe, and Chandler, 2000: 396 (2 Nearctic species; notes; characters in key).
allardi Levasseur, 1964: 291 (Gabronthus; Type locality: Luena, Haut Lomami, Katanga).
alluaudanus Jeannel and Paulian, 1945: 90 (Philonthus; subgenus Gabrius; Type locality: Diego Suarez).
angustatus Kraatz, 1859, see: thermarum Aubé, 1850.
australicus Cameron, 1943a: 343 (Philonthus; subgenus Gabrius; Type locality: Australia: Port Darwin).
badalus Tottenham, 1955: 186 (Gabronthus; Type locality: S. Nigeria).
balthasari Smetana, 1956, see: limbatus Fauvel, 1900.
chejuensis Cho, 1996: 28 (Gabronthus; Type locality: Korea: Chejudo, Kyoraeri).
chyuluensis Cameron, 1942b: 329 (Philonthus; subgenus Gabrius; Type locality: Chyulu Hills, altitude 5600 feet).
— Cameron, 1950g: 42 (Philonthus; subgenus Gabrius; Belgian Congo).
— Tottenham, 1955: 189 (Gabronthus; characters).
— Tottenham, 1962: 141 (Gabronthus; Tanganyika).
▴ Distribution: Kenya, Tanzania, Congo.
corrugatus Tottenham, 1955: 194 (Gabronthus; Type locality: Philippines Islands: Luzon, Los Banos).
— Last, 1968: 368 (Philonthus; [Note: Misidentified as thermarum according to Last, 1981: 125]; New Guinea).
— Last, 1981: 125 (Gabronthus; New Guinea).
▴ Distribution: Philippines, New Guinea.
exilis Kraatz, 1851, see: thermarum Aubé, 1850.
flavicollis Motschulsky, 1858: 660 (Heterothops; Type locality: Indes orientales).
— Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).
— Cameron, 1932: 266 (Heterothops; characters; Indes orientales).
— Scheerpeltz, 1933: 1422 (Heterothops; catalog).
— Cameron, 1934d: 79 (Heterothops; type examined; synonym of Gabrius pulchellus).
— Smetana, 1988: 319 (Gabronthus).
▴ Distribution: “East Indies”, Sri Lanka, Bhutan, Malaysia, Singapore, Indonesia.
pulchellus Kraatz, 1859: 92 (Philonthus; [preoccupied]; Type locality: India orientalis; Ceylan).
— Bernhauer and Schubert, 1914: 351 (Philonthus; catalog).
— Cameron, 1921a: 378, 405 (Philonthus; characters; Singapore).
— Cameron, 1932: 136 (Philonthus; characters; Ceylon).
— Scheerpeltz, 1933: 1356 (Philonthus; catalog).
— Cameron, 1933d: 387 (Philonthus; Sumatra).
— Coiffait, 1977b: 206 (Gabronthus; Bhutan).
— Smetana, 1988: 319 (Gabronthus; synonym of flavicollis).
foveifrons Cameron, 1932a: 142 (Philonthus; subgenus Gabrius; Type locality: Selangor: Bukit Kutu. Pahang: Raub, Bentong).
fundator Tottenham, 1955: 190 (Gabronthus; Type locality: Kenya: Muguga).
fuscolaterus Motschulsky, 1860, see: thermarum Aubé, 1850.
gombanus Tottenham, 1955: 188 (Gabronthus; Type locality: Abyssinia: Djem-Djem Forest).
gracilis Coiffait, 1983c: 181 (Gabronthus; Type locality: Tahiti: Mont Arue).
inclinans Walker, 1859: 51 (Xantholinus; Type locality: Ceylon).
— Fauvel, 1903d: 271 (Philonthus; synonym of thermarum).
— Fauvel, 1905e: 83 (Philonthus; synonym of thermarum).
— Bernhauer and Schubert, 1914: 357 (Philonthus; synonym of thermarum).
— Cameron, 1932: 134 (Philonthus; synonym of thermarum).
— Blackwelder, 1943: 403 (Philonthus; synonym of thermarum).
— Tottenham, 1955: 191 (Gabronthus; valid species; characters; angustatus Kraatz may be synonym; Ceylon; Sumatra; Java).
— Scheerpeltz, 1957c: 234 (Philonthus; [Note: I assume the author overlooked Tottenham's elevation of the species]; synonym of thermarum).
— Smetana, 1958: 217 (Philonthus; [Note: I assume the author overlooked Tottenham's elevation of the species]; synonym of thermarum).
— Coiffait, 1974: 114 (Gabronthus; [Note: I assume the author overlooked Tottenham's elevation of the species]; synonym of thermarum).
— Scheerpeltz, 1978: 224 (Philonthus; [Note: I assume the author overlooked Tottenham's elevation of the species]; synonym of thermarum).
▴ Distribution: Sri Lanka, Indonesia.
kisiranus Tottenham, 1955: 187 (Gabronthus; Type locality: Senegal).
kosanus Tottenham, 1955: 189 (Gabronthus; Type locality: Kenya: Muguga).
levasseuri Jarrige, 1978: 288 (Gabronthus; Type locality: Andringitra Centre, plateau Andohariana, 2000–2100 m).
libanicus Saulcy, 1864, see: maritimus Motschulsky, 1858.
limbatus Fauvel, 1900d: 230 (Philonthus; Type locality: Baical, rivière Sludianka).
— Bernhauer and Schubert, 1914: 344 (Philonthus; catalog).
— Scheerpeltz, 1933: 1349 (Philonthus; catalog).
— Coiffait, 1974: 116 (Gabronthus; characters; Siberia).
— Schuh, Schillhammer, and Zettel, 1992: 220 (Gabronthus; notes; Austria).
— Smetana, 1993b: 48 (Gabronthus; Slovakia).
— Assing, Frisch, Kahlen, et al., 1998: 138 (Gabronthus; note; Austria).
▴ Distribution: Russia, Slovakia, Austria.
balthasari Smetana, 1956b: 171, 173 (Gabrius; Type locality: Ostslowakei bei Královský Chl'mec).
— Smetana, 1957: 58 (Gabrius; characters; Slovakia).
— Smetana, 1959e (Gabronthus; characters).
— Smetana, 1958: 257 (Gabrius; characters; ecology; Czechoslovakia).
— Lohse, 1964: 187 (Gabronthus; characters; central Europe).
— Coiffait, 1974: 115 (Gabronthus; cited as balthazari; characters; Slovakia).
— Tóth, 1984: 82 (Gabronthus; cited as balthazari; characters).
— Lucht, 1987: 106 (Gabronthus; checklist; Czechoslovakia).
— Schillhammer, 1990: 92 (Gabronthus; synonym of limbatus).
— Schuh, Schillhammer, and Zettel, 1992: 220 (Gabronthus; synonym of limbatus).
— Smetana, 1993b: 48 (Gabronthus; synonym of limbatus).
— Assing, Frisch, Kahlen, et al., 1998: 138 (Gabronthus; synonym of limbatus).
lineifrons Cameron, 1926b: 359 (Philonthus; subgenus Gabrius; Type locality: Simla and Chakrata Districts, 7000 to 8000 feet).
— Cameron, 1932: 135 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1349 (Philonthus; catalog).
— Schillhammer, 1999a: 62 (Gabronthus).
▴ Distribution: India.
maritimus Motschulsky, 1858: 661 (Gabrius; Type locality: Alexandrie en Egypte).
— Gemminger and Harold, 1868: 594 (Philonthus; synonym of thermarum).
— Fauvel, 1869: 489 (Philonthus; valid species).
— Fauvel, 1874: 245 [= 1874a: 471] (Philonthus; race of thermarum; characters; France; Sicily; Corsica; Algeria; Cyprus; Egypt; Syria).
— Fauvel, 1878: 125 [= 1878a: 45] (Philonthus; variety of thermarum; Algeria; Egypt).
— Fauvel, 1886: 56 [= 1886a: 48] (Philonthus; variety of thermarum; notes; Morocco; Egypt; Algeria).
— Ganglbauer, 1895: 460 (Philonthus; subgenus Gabrius; characters; middle Europe).
— Fauvel, 1902b: 111 (Philonthus; Morocco; Algeria; Egypt).
— Sainte-Claire Deville, 1906: 98 (Philonthus; Corsica).
— Bernhauer and Schubert, 1914: 345 (Philonthus; catalog).
— Cameron, 1921a: 378, 405 (Philonthus; characters; Singapore).
— Portevin, 1929: 366 (Philonthus; subspecies of thermarum; characters; France).
— Gridelli, 1930: 74 (Philonthus; Libya).
— Bernhauer, 1931d: 587 (Philonthus; Abyssinia).
— Scheerpeltz, 1933: 1350 (Philonthus; catalog).
— C. Koch, 1941: 63 (Philonthus; Morocco).
— Bernhauer, 1942c: 361 (Philonthus; Cameroon).
— Jarrige, 1951: 134 (Philonthus; Annam; Indochina).
— Smetana, 1953: 119 (Philonthus; Turkey).
— Tottenham, 1955: 185 (Gabronthus; characters; many species are misidentified as maritimus).
— Smetana, 1959: 173 (Gabronthus; characters).
— Scheerpeltz, 1963b: 429 (Philonthus; subgenus Gabronthus; Egypt).
— Smetana, 1967i: 555 (Gabronthus; Turkey).
— Coiffait, 1973a: 274 (Philonthus; Morocco).
— Coiffait, 1974: 113 (Gabronthus; characters; Mediterranean region).
— Watanabe and Shibata, 1976: 332 (Philonthus; subgenus Gabronthus; Malaysia).
— Scheerpeltz, 1978: 224 (Philonthus; subgenus Gabronthus; Timor).
— Smetana, 1978: 116 (Gabronthus; Israel).
— Coiffait, 1981b: 236 (Gabronthus; Saudi Arabia).
— Coiffait, 1986: 393 (Gabronthus; Saudi Arabia).
— Lecoq, 1990: 196 (Gabronthus; Mascarene Islands).
— Ito, 1992a: 64 (Gabronthus; Yakushima Island).
— Shibata, 1993a: 318 (Gabronthus; Taiwan).
— Ciceroni and Zanetti, 1995: 29 (Gabronthus; Italy).
— Gusarov, 1997: 287 (Gabronthus; Saudi Arabia).
— Assing, 1998e: 140 (Gabronthus; Canary Islands).
— Assing, 2000b: 115 (Gabronthus; collecting notes; Canary Islands).
▴ Distribution: France, Italy, Cyprus, Turkey, Cameroon, Canary Islands, Morocco, Algeria, Libya, Egypt, Ethiopia, Saudi Arabia, Israel, Lebanon, Syria, Mascarene Islands, Malaysia, Vietnam, Singapore, Indonesia, Taiwan, Japan.
libanicus Saulcy, 1864: 639 (Philonthus; Type locality: Beïroût).
— Baudi, 1870: 386 (Philonthus; characters; Cyprus; Lebanon).
— Ganglbauer, 1895: 460 (Philonthus; synonym of maritimus).
— Bernhauer and Schubert, 1914: 346 (Philonthus; synonym of maritimus).
— Coiffait, 1974: 113 (Gabronthus; synonym of maritimus).
— Scheerpeltz, 1978: 224 (Philonthus; synonym of maritimus).
mimulus Rottenberg, 1870: 30 (Philonthus; Type locality: Sicily: Catania).
— Fauvel, 1874: 245 [= 1874a: 471] (Philonthus; synonym of thermarum maritimus).
— Ganglbauer, 1895: 460 (Philonthus; synonym of maritimus).
— Bernhauer and Schubert, 1914: 346 (Philonthus; synonym of maritimus).
— Coiffait, 1974: 113 (Gabronthus; synonym of maritimus).
— Scheerpeltz, 1978: 224 (Philonthus; synonym of maritimus).
pubens Mulsant and Rey, 1878b: 259 (Gabrius; Type locality: la Garonne, à Saint-Raphaël, Var).
— Ganglbauer, 1895: 460 (Philonthus; synonym of maritimus).
— Bernhauer and Schubert, 1914: 346 (Philonthus; synonym of maritimus).
— Coiffait, 1974: 113 (Gabronthus; synonym of maritimus).
— Scheerpeltz, 1978: 224 (Philonthus; synonym of maritimus).
meridionalis Cameron, 1945c: 716 (Philonthus; subgenus Gabrius; Type locality: South Africa: Simon's Town).
mgogoricus Tottenham, 1955: 185 (Gabronthus; Type locality: Rhodesia: Mashonaland, Salisbury; also listed from South Africa and Tanganyika).
— Frank, 1984: 476 (Gabronthus; characters; collecting notes; Jamaica; Haiti; Puerto Rico; St. Croix; Florida).
— Smetana, 1995e: 485 (Gabronthus; characters; notes; introduced to: Jamaica, Haiti, Puerto Rico, U.S. Virgin Islands, Florida, Mississippi, North Carolina).
▴ Distribution: Zimbabwe, South Africa, Tanzania, West Indies, USA.
mimulus Rottenberg, 1870, see: maritimus Motschulsky, 1858.
minusculus Tottenham, 1955: 193 (Gabronthus; Type locality: Ceylon).
minutissimus Bernhauer, 1915d: 143 (Philonthus; subgenus Gabrius; Type locality: Deutsch-Ostafrika: Unjanjembe).
— Bernhauer, 1931d: 587 (Philonthus; subgenus Gabrius; Abyssinia).
— Scheerpeltz, 1933: 1351 (Philonthus; catalog).
— Tottenham, 1955: 188 (Gabronthus; characters).
▴ Distribution: Tanzania, Ethiopia.
musonoiensis Levasseur, 1964: 290 (Gabronthus; Type locality: Région de Kolwezi, Katanga, Musonoie).
oribates Jeannel and Paulian, 1945: 88 (Philonthus; subgenus Gabrius; Type locality: camp II de l'Elgon, Elgon Sae Mill, alt. 2470 m. Also cited from Cameroon).
pluvialis Tottenham, 1955: 185 (Gabronthus; Type locality: Kenya: Muguga).
pubens Mulsant and Rey, 1878, see: maritimus Motschulsky, 1858.
pulchellus Kraatz, 1859, see: flavicollis Motschulsky, 1858.
pygmaeus Kraatz, 1859: 93 (Philonthus; Type locality: Ceylan; India orientalis).
— Fauvel, 1878f: 268 (Philonthus; synonym of thermarum).
— Sharp, 1889: 39 (Philonthus; notes; Japan).
— Ganglbauer, 1895: 460 (Philonthus; synonym of thermarum).
— Fauvel, 1903d: 271 (Philonthus; synonym of thermarum).
— Fauvel, 1905e: 83 (Philonthus; synonym of thermarum).
— Bernhauer and Schubert, 1914: 357 (Philonthus; synonym of thermarum).
— Cameron, 1932: 134 (Philonthus; synonym of thermarum).
— Blackwelder, 1943: 403 (Philonthus; synonym of thermarum).
— Jarrige, 1951: 134 (Philonthus; characters; Indochina).
— Tottenham, 1955: 192 (Gabronthus; valid species; characters; Ceylon; Philippines).
— Scheerpeltz, 1957c: 234 (Philonthus; [Note: I assume the author overlooked Tottenham's elevation of the species]; synonym of thermarum).
— Smetana, 1958: 217 (Philonthus; [Note: I assume the author overlooked Tottenham's elevation of the species]; synonym of thermarum).
— Coiffait, 1974: 114 (Gabronthus; [Note: I assume the author overlooked Tottenham's elevation of the species]; synonym of thermarum).
— Scheerpeltz, 1978: 224 (Philonthus; [Note: I assume the author overlooked Tottenham's elevation of the species]; synonym of thermarum).
— Lecoq, 1990: 196 (Gabronthus; Mascarene Islands).
▴ Distribution: Mascarene Islands, Sri Lanka, India, “Indochina”, Philippines, Japan.
quadriceps Cameron, 1950d: 20 (Philonthus; subgenus Gabrius; Type locality: Pahang: Cameron Highlands, Ginting Kial, 5000 feet).
rufoflavus Cameron, 1936b: 44 (Philonthus; subgenus Gabrius; Type locality: Selangor: Kuala Lumpur).
sincerus Tottenham, 1962: 233 (Gabronthus; Type locality: Tanganyika Terr.: Ngorongoro, 2400–2500 m).
styx Tottenham, 1955: 194 (Gabronthus; Type locality: Siberia: Amur).
sulcifrons Sharp, 1889: 39 (Philonthus; Type locality: Nagasaki).
— Fauvel, 1903d: 271 (Philonthus; synonym of thermarum).
— Fauvel, 1905e: 83 (Philonthus; synonym of thermarum).
— Bernhauer and Schubert, 1914: 357 (Philonthus; synonym of thermarum).
— Bernhauer, 1922c: 232 (Philonthus; valid species; characters; Formosa).
— Scheerpeltz, 1933: 1363 (Philonthus; catalog).
— Cameron, 1933d: 387 (Philonthus; Sumatra).
— Tottenham, 1955: 191 (Gabronthus; valid species; characters; Japan).
— Watanabe and Shibata, 1972: 64 (Philonthus; Yaku-shima Island).
— Kuschel, 1990: 26 (Gabronthus; introduced, Palaearctic species; winged; habitat; New Zealand).
— J. Li, 1993: 40 (Gabronthus; China).
— Cho, 1996: 26 (Gabronthus; characters; Korea).
▴ Distribution: China, Korea, Japan, Taiwan, Indonesia, Sri Lanka, New Zealand.
tahitiensis Coiffait, 1976: 225 (Gabronthus; Type locality: Tahiti: Papenoo).
tenuicorpus Cho, 1996: 27 (Gabronthus; Type locality: Korea: Kyongsangbukdo, Taegu, Sankyukdong, Campus of Kyungpook National University).
tenuissimus Tottenham, 1949b: 344 (Philonthus; Type locality: Natal: Kloof).
thermaroides Tottenham, 1955: 193 (Gabronthus; Type locality: Philippines Islands: Mindanao, Momungan).
— Last, 1968: 368 (Philonthus; [Note: Misidentified as thermarum according to Last, 1981: 125]; New Guinea).
— Last, 1981: 125 (Gabronthus; New Guinea).
▴ Distribution: Philippines, New Guinea.
thermarum Aubé, 1850: 316 (Philonthus; Type locality: Paris, Jardin des Plantes).
— Fairmaire and Laboulbène, 1856: 522 (Philonthus; characters; France).
— Kraatz, 1857d: 608 (Philonthus; characters; Germany).
— Wollaston, 1860: 102 (Philonthus; characters; notes Madeira).
— Wollaston, 1865: 491 (Philonthus; Madeira Islands).
— Baudi, 1870: 386 (Philonthus; characters).
— Solsky, 1872: 303 (Philonthus; Cayenne).
— Fauvel, 1874: 245 [= 1874a: 470] (Philonthus; characters; France; Britain; Germany; Italy; Ceylon; East Indies; China; Japan; Java; Madeira; North America; Guyana).
— Sharp, 1874: 44 (Philonthus; characters; Japan; China).
— Seidlitz, 1875: 277 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 546 [= 1877: 402] (Gabrius; characters; France).
— Fauvel, 1878f: 268 (Philonthus; characters; New Guinea; Java; Japan; China).
— Horn, 1884: 196 (Philonthus; characters; Missouri; District of Columbia).
— Fowler, 1888: 276 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 460 (Philonthus; subgenus Gabrius; characters; middle Europe; Italy; Madeira; East Africa; Ceylon; India; China; Japan; North America; Guyana).
— Fauvel, 1902b: 111 (Philonthus; Algeria; Madeira).
— Fauvel, 1903d: 271 (Philonthus; New Caledonia).
— Fauvel, 1905e: 83 (Philonthus; Java).
— Reitter, 1909: 131 (Philonthus; subgenus Gabrius; characters; Germany).
— Eichelbaum, 1913: 130 (Philonthus; Tanzania).
— Bernhauer and Schubert, 1914: 357 (Philonthus; catalog).
— Johansen, 1914: 417 (Philonthus; subgenus Gabrius; characters; Denmark).
— Bernhauer, 1922b: 176 (Philonthus; Seychelles).
— Bernhauer, 1922c: 232 (Philonthus; Formosa).
— Lea, 1925: 221 (Philonthus; not recorded in Australia).
— Bernhauer, 1928: 17 (Philonthus; Sumatra).
— Portevin, 1929: 365 (Philonthus; characters; France).
— Cameron, 1931: 363 (Philonthus; New Guinea).
— Cameron, 1932: 134 (Philonthus; characters; Ceylon; India).
— Scheerpeltz, 1933: 1364 (Philonthus; catalog).
— Cameron, 1933d: 387 (Philonthus; Sumatra).
— Cameron, 1934a: 22 (Philonthus; New Hebrides).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Cameron, 1937: 6 (Philonthus; Java).
— Bernhauer and Chapman, 1939: 69 (Philonthus; Kenya).
— Méquignon, 1942a: 21 (Philonthus; Azores).
— Blackwelder, 1943: 403 (Philonthus; characters; Jamaica; Haiti; Puerto Rico; St. Croix; Antigua; Guadeloupe; Grenada).
— Cameron, 1951: 282 (Philonthus; subgenus Gabrius; Egypt).
— V. Hansen, 1952: 87 (Philonthus; subgenus Gabrius; characters; Denmark).
— Tottenham, 1955: 183 (Gabronthus; characters; some names listed as synonyms are different species).
— Scheerpeltz, 1957c: 234 (Philonthus; Sumba).
— Smetana, 1958: 217 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 172 (Gabronthus; characters).
— Palm, 1963b: 103 (Philonthus; subgenus Gabrius; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 187 (Gabronthus; characters; central Europe).
— Horion, 1965: 181 (Philonthus; subgenus Gabronthus; Ireland; Britain; Norway; Sweden; Finland; Denmark; Holland; Belgium; Switzerland; France; Spain; Portugal; Italy; Czech Republic; Slovakia; Poland; Austria; Germany).
— Levasseur, 1968a: 53 (Gabronthus; Central African Republic).
— Scheerpeltz, 1971c: 170 (Philonthus; Cameroon).
— Coiffait, 1974: 114 (Gabronthus; characters; Europe).
— Scheerpeltz, 1974: 161 (Philonthus; Basutoland; South Africa).
— Scheerpeltz, 1974c: 24 (Philonthus; Ethiopia; Sudan).
— Scheerpeltz, 1976a: 89 (Gabronthus; Nepal).
— Pope, 1977: 30 (Gabronthus; Britain).
— Scheerpeltz, 1978: 224 (Philonthus; Timor).
— Frank, 1984: 475 (Gabronthus; characters; collecting notes; Jamaica; Puerto Rico; Antigua; St. Kitts; Guadeloupe; Costa Rica; Florida).
— Tóth, 1984: 81 (Gabronthus; characters; Hungary).
— Segers, 1986: 43 (Gabronthus; checklist of species; Belgium).
— Lucht, 1987: 106 (Gabronthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; Benelux; Denmark; Sweden).
— Borges, 1990: Table II (Gabronthus; Azores).
— Lecoq, 1990: 197 (Gabronthus; Mascarene Islands).
— Campbell and Davies, 1991: 119 (Gabronthus; checklist of species; Canada).
— J. Li, 1993: 40 (Gabronthus; China).
— Smetana, 1993b: 48 (Gabronthus; Czech Republic; Slovakia).
— Hernández, Outerelo, and Gamarra, 1994: 188 (Gabronthus; checklist of species; Canary Islands).
— Ciceroni and Zanetti, 1995: 29 (Gabronthus; Italy ?).
— Smetana, 1995e: 481 (Gabronthus; characters; notes; Ontario; Quebec; Connecticut; Florida; Illinois; Kansas; Massachusetts; Michigan; New Hampshire; New Jersey; North Carolina; Ohio; Oklahoma; Pennsylvania).
— Downie and Arnett, 1996: 383 (Gabronthus; characters; USA).
— Sterrenburg and Schülke, 1997: 20 (Gabronthus; Netherlands).
— Ødegaard, 1999: 71 (Gabronthus; invasive species associated with compost heaps in Nordic countries).
▴ Distribution: Wide-spread in Africa, Europe, Asia, North America, West Indies.
exilis Kraatz, 1851: 293 (Philonthus; Type locality: Not cited).
— Fairmaire and Laboulbène, 1856: 522 (Philonthus; synonym of thermarum).
— Kraatz, 1857d: 608 (Philonthus; synonym of thermarum).
— Redtenbacher, 1857: 198 (Philonthus; characters; Germany).
— Redtenbacher, 1874: 217 (Philonthus; characters; Germany).
— Ganglbauer, 1895: 460 (Philonthus; synonym of thermarum).
— Bernhauer and Schubert, 1914: 357 (Philonthus; synonym of thermarum).
— Cameron, 1932: 134 (Philonthus; synonym of thermarum).
— Scheerpeltz, 1933: 1364 (Philonthus; synonym of thermarum).
— Blackwelder, 1943: 403 (Philonthus; synonym of thermarum).
— Scheerpeltz, 1957c: 234 (Philonthus; synonym of thermarum).
— Smetana, 1958: 217 (Philonthus; synonym of thermarum).
— Coiffait, 1974: 114 (Gabronthus; synonym of thermarum).
— Scheerpeltz, 1978: 224 (Philonthus; synonym of thermarum).
— Smetana, 1995e: 481 (Gabronthus; synonym of thermarum).
angustatus Kraatz, 1859: 92 (Philonthus; Type locality: Ceylan; India orientalis).
— Fauvel, 1878f: 268 (Philonthus; synonym of thermarum).
— Sharp, 1889: 39 (Philonthus; notes; China; Japan).
— Ganglbauer, 1895: 460 (Philonthus; synonym of thermarum).
— Fauvel, 1903d: 271 (Philonthus; synonym of thermarum).
— Fauvel, 1905e: 83 (Philonthus; synonym of thermarum).
— Bernhauer and Schubert, 1914: 357 (Philonthus; synonym of thermarum).
— Cameron, 1932: 134 (Philonthus; synonym of thermarum).
— Blackwelder, 1943: 403 (Philonthus; synonym of thermarum).
— Scheerpeltz, 1957c: 234 (Philonthus; synonym of thermarum).
— Smetana, 1958: 217 (Philonthus; synonym of thermarum).
— Coiffait, 1974: 114 (Gabronthus; synonym of thermarum).
— Scheerpeltz, 1978: 224 (Philonthus; synonym of thermarum).
fuscolaterus Motschulsky, 1860a: 76 (Gabrius; Type locality: montagnes de Noura-Ellia à Ceylan).
— Fauvel, 1903d: 271 (Philonthus; synonym of thermarum).
— Fauvel, 1905e: 83 (Philonthus; synonym of thermarum).
— Bernhauer and Schubert, 1914: 357 (Philonthus; synonym of thermarum).
— Cameron, 1932: 134 (Philonthus; synonym of thermarum).
— Blackwelder, 1943: 403 (Philonthus; synonym of thermarum).
— Scheerpeltz, 1957c: 234 (Philonthus; synonym of thermarum).
— Smetana, 1958: 217 (Philonthus; synonym of thermarum).
— Coiffait, 1974: 114 (Gabronthus; synonym of thermarum).
— Scheerpeltz, 1978: 224 (Philonthus; synonym of thermarum).
— Smetana, 1995e: 481 (Gabronthus; synonym of thermarum).
turneri Tottenham, 1949b: 344 (Philonthus; Type locality: Natal: Van Reengen, Drakensberg).
ziloanus Levasseur, 1964: 292 (Gabronthus; Type locality: env. de Kolwezi, Katanga, Zilo).
Glyphesthus
[12 species; Ethiopian Region]
Glyphesthus Kraatz, 1858: 364 (species included: rufipennis, niger). Type species: Glyphesthus rufipennis Kraatz, fixed by subsequent designation by Blackwelder, 1952: 171.
— Gemminger and Harold, 1868: 573 (catalog; cited as Glyphestes).
— Bernhauer and Schubert, 1916: 411 (world catalog; 2 species).
— Bernhauer, 1931b: 85 (key to species).
— Scheerpeltz, 1933: 1421 (world catalog supplement).
Aleioglyphesthus Herman, 2001: 8 (subgenus of Glyphesthus; species included: see Scheerpeltz, 1975). Type species: Glyphesthus (Aleioglyphesthus) zimmeri Scheerpeltz, fixed by original designation).
— Scheerpeltz, 1975: 110 (subgenus of Glyphesthus; species included: congoensis, neavei, hauseri, zimmeri; [Note: No type species designation so name was unavailable (Article 13.3)]).
congoensis Bernhauer, 1931b: 85, 87 (Glyphesthus; Type locality: Belgisch-Kongo: Elisabethville).
— Scheerpeltz, 1933: 1421 (Glyphesthus; catalog).
— Scheerpeltz, 1975: 111 (Glyphesthus; subgenus Aleioglyphesthus; characters; Congo).
▴ Distribution: Congo].
evansi Bernhauer, 1931b: 85, 86 (Glyphesthus; Type locality: Belgisches Kongogebiet: 18 m. S. W von Elisabethville).
— Scheerpeltz, 1933: 1421 (Glyphesthus; catalog).
— Scheerpeltz, 1975: 109 (Glyphesthus; characters; Congo).
▴ Distribution: Congo.
hauseri Bernhauer, 1937: 305 (Glyphesthus, cited as Glyphestus; Type locality: Tanganyika: Uheheland, Kidugala).
— Scheerpeltz, 1975: 111 (Glyphesthus; subgenus Aleioglyphesthus; characters; Kenya; Urundi).
▴ Distribution: Kenya, Tanzania, Burundi.
leakeyi Bernhauer, 1937: 305 (Glyphesthus; cited as Glyphestus; Type locality: Kenya: Oldoway, Turrei).
modestus Bernhauer, 1929f: 123 (Glyphesthus; Type locality: Haut-Uele: Tuku; Moto).
— Bernhauer, 1931b: 85 (Glyphesthus; characters in key; Belgian Congo).
— Scheerpeltz, 1933: 1421 (Glyphesthus; catalog).
— Scheerpeltz, 1975: 108 (Glyphesthus; characters; Congo region).
▴ Distribution: Congo.
neavei Bernhauer, 1927: 57 (Glyphesthus; Type locality: Nyassaland: Mlanje).
— Bernhauer, 1931b: 85 (Glyphesthus; characters in key; Nyassaland).
— Scheerpeltz, 1933: 1421 (Glyphesthus; catalog).
— Scheerpeltz, 1975: 111 (Glyphesthus; subgenus Aleioglyphesthus; characters; Kenya).
▴ Distribution: Kenya, Malawi.
niger Kraatz, 1858: 365 (Glyphesthus; Type locality: Senegambia).
— Fauvel, 1899a: 32 (Glyphesthus; Senegambia).
— Bernhauer and Schubert, 1916: 411 (Glyphesthus; catalog).
— Bernhauer, 1931b: 85 (Glyphesthus; characters in key; Senegambien).
— Scheerpeltz, 1933: 1421 (Glyphesthus; catalog; Ethiopia).
— Scheerpeltz, 1975: 110 (Glyphesthus; characters; Guinea; Senegal; Niger; Ivory Coast).
— Levasseur, 1968a: 54 (Glyphesthus; Central African Republic).
▴ Distribution: “Senegambia”, Guinea, Ivory Coast, Niger, Central African Republic, Ethiopia.
nigerrimus Cameron, 1951c: 407 (Glyphesthus; Type locality: Kenya Colony: Kitui).
parcepunctatus Bernhauer, 1927g: 229 (Glyphesthus; Type locality: Harrar, Abessynien).
— Bernhauer, 1931b: 85 (Glyphesthus; characters in key; Abessinien).
— Scheerpeltz, 1933: 1421 (Glyphesthus; catalog).
— Scheerpeltz, 1975: 107 (Glyphesthus; characters; Abyssinia).
▴ Distribution: Ethiopia.
picipennis Roth, 1851: 118 (Acylophorus; Type locality: Abyssinien).
— Fauvel, 1899a: 32 (Glyphesthus; Senegambie portugaise; Abyssinia; Tigre).
— Bernhauer and Schubert, 1916: 411 (Glyphesthus; catalog).
— Bernhauer, 1931b: 85 (Glyphesthus; characters in key; Abessinia).
— Scheerpeltz, 1933: 1421 (Glyphesthus; catalog).
— Scheerpeltz, 1975: 109 (Glyphesthus; characters; Guinea; Senegal; Niger; Cameroon; Aequatoria; Abyssinia).
▴ Distribution: “Senegambia”, Senegal, Guinea, Niger, Cameroon, “Equatoria”, Ethiopia.
rufipennis Kraatz, 1858: 365 (Glyphesthus; Type locality: Senegambia).
rufipennis Kraatz, 1858, see: picipennis Roth, 1851.
sanguinipennis Bernhauer, 1931b: 85, 86 (Glyphesthus; Type locality: Mashonaland; Natal: Colenso; Delagoabai).
— Scheerpeltz, 1933: 1421 (Glyphesthus; catalog).
— Scheerpeltz, 1975: 108 (Glyphesthus; characters; South Africa).
▴ Distribution: South Africa, Zimbabwe.
zimmeri Scheerpeltz, 1975: 103, 112 (Glyphesthus; subgenus Aleioglyphesthus; Type locality: Umgebung von Ugano im Matengo-Hochlande, West-Kenyas).
Hesperomimus
[1 species; Oriental Region]
Hesperomimus Cameron, 1937: 17 (species included: abdominalis; [Note: For the generic description Cameron spelled the name Hesperominus and for the species, Hesperomimus. Blackwelder, 1952: 183, argued that Hesperomimus is the correct spelling]). Type species: Hesperomimus abdominalis Cameron, fixed by monotypy.
abdominalis Cameron, 1937: 17 (Hesperomimus; Type locality: Java: Batoerraden, G. Slamat).
Hesperopalpus
[3 species; Palaearctic Region]
Hesperopalpus Shibata, 1973: 23 (species included: venustus). Type species: Hesperopalpus venustus Shibata, fixed by original designation and monotypy.
— Shibata, 1986a: 124 (supplemental list of species from Taiwan).
— Schillhammer, 1999c: 303 (synopsis of genus; additional characters; notes; key to species).
hesperiformis Cameron, 1932: 153 (Philonthus; Type locality: Nilgiri Hills).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Schillhammer, 1996a: 83 (Hesperopalpus; characters; India).
— Schillhammer, 1999c: 307 (Hesperopalpus; characters; India).
▴ Distribution: India.
venustus Shibata, 1973: 25 (Hesperopalpus; Type locality: Taiwan: Koantauchi, Nantou Hsien).
vietnamensis Schillhammer, 1999c: 306 (Hesperopalpus; Type locality: Vietnam: Taj Nguen, Bud Loij, 65 km N. Ankche).
Hesperus
[201 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Oriental, Australian, and Oceanic Regions]
Hesperus Fauvel, 1874: 200 [= 1874a: 426] (species included: rufipennis, baltimorensis, apicialis [cited as apicalis]). Type species: Staphylinus rufipennis Gravenhorst, fixed by subsequent designation by R. Lucas, 1920: 324.
— Fauvel, 1875a: XXIX [= 1875b: 231] (catalog).
— Mulsant and Rey, 1876: 559 [= 1877: 415] (characters).
— Duvivier, 1883: 150 (catalog).
— Olliff, 1887: 508 (characters; Australian species).
— Heyden, Reitter, and Weise, 1891: 102 (list of species of Europe and Caucasus).
— Ganglbauer, 1895: 461 (characters).
— Fauvel, 1903d: 269 (key to New Caledonia species).
— Heyden, Reitter, and Weise, 1906: 163 (list of species of Europe and Caucasus).
— Reitter, 1909: 116, 122 (characters).
— Petri, 1912: 71 (list of species of Siebenbürgen).
— Bernhauer and Schubert, 1914: 363 (world catalog; 25 species).
— Leng, 1920: 108 (catalog of North American species).
— Cameron, 1921a: 354, 405 (characters; key to Singapore species; catalog).
— Gridelli, 1924 (characters; anatomy; revision of species; key to species).
— Winkler, 1925: 381 (catalog for Palaearctic region).
— Cameron, 1925: 64 (catalog of Indian species).
— Porta, 1926: 100 (characters; species of Italy).
— Bernhauer, 1928g: 116 (key to African species).
— Bruch, 1928a: 428 (catalog; Argentina).
— Roubal, 1930: 378 (catalog; Slovakia).
— Cameron, 1932: 156 (characters; key to species of British India).
— Scheerpeltz, 1933: 1371 (world catalog supplement).
— Scheerpeltz, 1936a: 488 (key to Nearctic and Neotropical species).
— Blackwelder, 1944: 138 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 184 (type species).
— Scheerpeltz, 1956b: 21 (key to African species).
— Moore, 1958 (characters; notes; key to North American species).
— Kocher, 1958: 152 (checklist of species; Morocco).
— Smetana, 1958: 258 (characters; habitus of one species; type species: rufipennis; Czechoslovakian species).
— Coiffait, 1963: 8, 18 (characters).
— Székessy, 1963: 56 (characters).
— Lohse, 1964: 174 (characters; key to central European species).
— Moore, 1965: 98, 101 (characters; notes).
— Scheerpeltz, 1968a: 79 (catalog of Austrian species).
— Scheerpeltz, 1971 (key to species).
— Last, 1973 (characters).
— Tikhomirova, 1973a: 185 (checklist of species of USSR).
— Coiffait, 1974: 320 (characters; species of western Palaearctic region).
— Moore and Legner, 1975: 16 (catalog; North America north of Mexico).
— Moore and Legner, 1979: 22 (characters; notes).
— Burakowski, Mroczkowski, and Stefańska, 1980: 33 (catalog; Poland).
— Shibata, 1983: 137 (checklist of species of Japan).
— Hammond, 1984: 205 (checklist of species of Borneo).
— Tóth, 1984: 41 (characters; habitus; species of Hungary).
— Shibata, 1986a: 123 (supplemental list of species from Taiwan).
— Shibata, 1990: 212 (key to species of Taiwan).
— Smetana, 1995e: 32, 486 (characters; notes; key to North American species; phylogenetic relationships).
— Ciceroni and Zanetti, 1995: 27 (list of species of Italy).
— Downie and Arnett, 1996: 383 (characters; key to species of Northeastern North America).
— Lundgren, 1998: 51 (list of species from Florida; records for other States listed for each species).
— Newton, Thayer, Ashe, and Chandler, 2000: 396 (6 Nearctic species; notes; characters in key).
— Hayashi, 2000: 23 (notes on empodial setae).
Hesperotropis Gridelli, 1924: 196 (subgenus of Hesperus; species included: perfoliatus; [Note: Following the description of the subgenus Hesperotropis Gridelli described Hesperus perfoliatus and that species was followed by Hesperus borneensis, H. bakeri, H. modestus, H. moultoni, H. ornatus, H. phaenomenalis, H. semicoeruleus, and H. terminalis but it is unclear that Gridelli intended them to be assigned to the new subgenus. In his key to species (page 182) Gridelli did not group H. perfoliatus in any special way with the other species and Scheerpeltz, 1933: 1371 and 1971: 193 included only H. perfoliatus in the subgenus]). Type species: Hesperus (Hesperotropis) perfoliatus Gridelli, fixed by monotypy [Note: If only one species was originally included then the type species is fixed by monotypy; if the other eight species were included then Blackwelder, 1952: 184, fixed the type species by subsequent designation].
— Cameron, 1932: 156 (subgenus of Hesperus).
— Scheerpeltz, 1933: 1371 (subgenus of Hesperus).
— Blackwelder, 1952: 184 (type species).
— Scheerpeltz, 1971: 193 (subgenus of Hesperus; characters).
abnormalis Cameron, 1952: 252 (Hesperus; Type locality: New Guinea: Snow Mountains: Moss Forest Camp, 2800 m).
addisadebaensis Scheerpeltz, 1971: 184 (Hesperus; Type locality: Aethiopien: Addisadeba).
aeneipennis Cameron, 1950c: 95 (Hesperus; Type locality: Belgian Congo: Mayidi).
aeneus Gridelli, 1924: 183, 191 (Hesperus; Type locality: Sumatra occid., Si Rambé).
— Scheerpeltz, 1933: 1371 (Hesperus; catalog).
— Scheerpeltz, 1971: 166 (Hesperus; characters).
▴ Distribution: Indonesia.
africanus Bernhauer, 1915d: 145 (Hesperus; Type locality: Erythraea: Ghinda).
— Bernhauer, 1928g: 117 (Hesperus; characters in key).
— Scheerpeltz, 1933: 1371 (Hesperus; catalog).
— Scheerpeltz, 1956b: 24 (Hesperus; characters; Erythrea).
— Scheerpeltz, 1971: 185 (Hesperus; characters).
— Scheerpeltz, 1974c: 24 (Hesperus; Sudan).
▴ Distribution: Eritrea, Sudan.
alboguttatus Cameron, 1946: 11 (Hesperus; Type locality: Philippines: Mindanao: Surigao).
amabilis Kraatz, 1859: 97 (Philonthus; Type locality: India borealis).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog).
— Cameron, 1932: 152 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1332 (Philonthus; catalog).
— Schillhammer, 1999a: 62 (Hesperus).
▴ Distribution: India.
amans Last, 1987: 33 (Hesperus; Type locality: Papua New Guinea: New Guinea NE: Wau, 1700 m).
anchora Fauvel, 1877: 258 (Cafius; Type locality: Nouvelle Calédonie).
— Fauvel, 1889b: 263 (Hesperus; New Caledonia).
— Fauvel, 1903d: 269 (Hesperus; New Caledonia).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Scheerpeltz, 1971: 178 (Hesperus; characters).
▴ Distribution: New Caledonia.
andrewesi Cameron, 1932: 157 (Hesperus; Type locality: Nilgiri Hills).
— Scheerpeltz, 1933: 1371 (Hesperus; catalog).
— Scheerpeltz, 1971: 167 (Hesperus; characters).
▴ Distribution: India.
apicialis Say, 1830: 37 [= 1834: 451] (Staphylinus; Type locality: Middle States [Note: See neotype designation by Smetana, 1995e]. Originally cited from the United States).
— Erichson, 1840: 931 (Philonthus; cited as apicalis; [Note: Bernhauer and Schubert, 1914: 364 cited apicalis Erichson as a synonym of apicialis; Erichson did not describe the species as new, he attributed it to Say but misspelled it]).
— Fauvel, 1874: 200 [= 1874a: 426] (Hesperus; cited as apicalis).
— Horn, 1884: 220 (Philonthus; characters; Canada to Georgia).
— Blatchley, 1910: 390 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Scheerpeltz, 1936a: 489 (Hesperus; characters in key).
— Moore, 1958: 315 (Hesperus; characters; Ohio, Missouri, Alabama).
— Scheerpeltz, 1971: 189 (Hesperus; characters).
— Frank, 1984: 478 (Hesperus; characters; Florida).
— Campbell and Davies, 1991: 119 (Hesperus; checklist of species; Canada).
— Smetana, 1995e: 498 (Hesperus; neotype designation: Middle States; characters; notes; Ontario; Quebec; Alabama; Connecticut; Illinois; Indiana; Iowa; Kansas; Kentucky; Maine; Maryland; Massachusetts; Michigan; Nebraska; New Hampshire; New Jersey; New York; North Carolina; Ohio; Pennsylvania; Rhode Island; Texas; Virginia; West Virginia).
— Downie and Arnett, 1996: 383 (Hesperus; characters; USA).
▴ Distribution: Canada, USA.
haematurus Erichson, 1840: 504 (Philonthus; Type locality: America septentrionalis; [Note: See lectotype designation by Smetana, 1995e]).
— Erichson, 1840: 931 (Philonthus; synonym of apicialis).
— LeConte, 1859: 566 (Philonthus; synonym of apicialis).
— Fauvel, 1874: 200 [= 1874a: 426] (Hesperus; synonym of apicialis).
— Bernhauer and Schubert, 1914: 364 (Hesperus; synonym of apicialis).
— Scheerpeltz, 1971: 189 (Hesperus; synonym of apicialis).
— Smetana, 1995e: 499 (Hesperus; lectotype designation; synonym of apicialis).
argentinus Scheerpeltz, 1971: 191 (Hesperus; Type locality: Argentinien: Santiago del Estero).
arizonicus Moore, 1958: 312 (Hesperus; Type locality: Patagonia, Arizona).
— Moore 1974: 22 (Hesperus; characters of female; Arizona).
— Smetana, 1995e: 504 (Hesperus; characters; notes; Arizona).
▴ Distribution: USA.
asperatus Last, 1987: 33 (Hesperus; Type locality: Papua New Guinea: New Guinea NE: Morobe Distr., Wau, 1200 m).
assecla Last, 1968: 350 (Philonthus; Type locality: New Guinea: Takai).
— Last, 1973: 425 (Hesperus).
— Last, 1987: 49 (Hesperus; Papua New Guinea).
▴ Distribution: New Guinea.
australis W. J. MacLeay, 1873: 139 (Philonthus; Type locality: Gayndah).
— Fauvel, 1877: 260 (Hesperus; characters; Australia).
— Fauvel, 1878g: 543 (Hesperus; characters; Australia).
— Olliff, 1887: 510 (Hesperus; characters; Australia).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Scheerpeltz, 1971: 176 (Hesperus; characters).
▴ Distribution: Australia.
babai Shibata, 1990: 209 (Hesperus; subgenus Hesperotropis; Type locality: Taiwan: Nantou Hsien: Sungkang, about 2200 m alt.).
bafutensis Levasseur, 1967a: 954 (Hesperus; Type locality: Cameroun: forêt de Bafut, N'guemba).
bakeri Bernhauer, 1919: 364 (Hesperus; Type locality: Baguio: Benguet).
— Gridelli, 1924: 199 (Hesperus; characters; Philippines).
— Scheerpeltz, 1933: 1371 (Hesperus; catalog).
— Scheerpeltz, 1971: 172 (Hesperus; characters).
▴ Distribution: Philippines.
obscuripennis Scheerpeltz, 1971: 172 (Hesperus; variety of bakeri; [Note: Nomen nudum (Article 15.2)]; cited from Philippinen: Luzon: Balbalan).
bakerianus Bernhauer, 1926b: 41 (Hesperus; Type locality: N. Viscaya, Imugin).
— Scheerpeltz, 1933: 1371 (Hesperus; catalog).
— Scheerpeltz, 1971: 170 (Hesperus; characters).
▴ Distribution: Philippines.
baltimorensis Gravenhorst, 1802: 163 (Staphylinus; Type locality: America septentr.; [Note: See lectotype designation by Smetana, 1995e]).
— Gravenhorst, 1806: 112 (Staphylinus).
— Nordmann, 1837: 109 (Philonthus; America septentrionalis).
— Erichson, 1840: 503 (Philonthus; characters; America septentrionalis).
— Fauvel, 1874:: 200 [= 1874a: 426] (Hesperus).
— Horn, 1884: 220 (Philonthus; characters; middle and southern States).
— Blatchley, 1910: 390 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Scheerpeltz, 1936a: 490 (Hesperus; characters in key).
— Moore, 1958: 314 (Hesperus; characters; Indiana, New York, Alabama, Missouri).
— Scheerpeltz, 1971: 189 (Hesperus; characters).
— Frank, 1984: 478 (Hesperus; characters; Florida).
— Smetana, 1995e: 492 (Hesperus; lectotype designation; characters; notes; Alabama; Arkansas; Connecticut; Delaware; Georgia; Illinois; Indiana; Iowa; Kansas; Kentucky; Louisiana; Maryland; Massachusetts; Michigan; Mississippi; Missouri; Montana; Nebraska; New Hampshire; New Jersey; New York; North Carolina; Ohio; Oklahoma; Pennsylvania; South Carolina; Virginia).
— Downie and Arnett, 1996: 383 (Hesperus; characters; USA).
▴ Distribution: Canada, USA.
barrioni Last, 1991: 1076 (Hesperus; Type locality: Philippines: Luzon Island: Siniloan, Laguna, Magsaysay village).
belonuchiformis Bernhauer, 1915d: 145 (Hesperus; Type locality: Abessinien: Boroda).
— Bernhauer, 1928g: 117 (Hesperus; characters in key).
— Scheerpeltz, 1933: 1371 (Hesperus; catalog).
— Scheerpeltz, 1956b: 23 (Hesperus; characters; Abyssinia).
— Scheerpeltz, 1971: 184 (Hesperus; characters).
▴ Distribution: Ethiopia.
bicolor Smetana, 1995e: 497 (Hesperus; Type locality: Florida: Coronado Bch.).
bifuscipennis Lea, 1931: 373 (Hesperus; Type locality: Papua: Mount Lamington).
— Scheerpeltz, 1933: 1371 (Hesperus; catalog).
— Scheerpeltz, 1971: 177 (Hesperus; characters).
▴ Distribution: New Guinea.
birmanus Cameron, 1932: 160 (Hesperus; Type locality: Burma: Karen Mts.; Assam: Sadiya).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1971: 167 (Hesperus; characters).
▴ Distribution: India, Myanmar.
borneensis Bernhauer, 1915f: 142 (Hesperus; Type locality: Sarawak: Mt. Matang, 2000 Fuss).
— Gridelli, 1924: 198 (Hesperus; characters; Sarawak).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1971: 167 (Hesperus; characters).
▴ Distribution: Borneo, Indonesia.
rufipennis Cameron, 1918b: 171 (Leucitus; [preoccupied]; Type locality: Borneo, Quop, W. Sarawak; also unnamed variety from Bedagei, Sumatra).
bruchi Bernhauer, 1921c: 169 (Hesperus; Type locality: Argentinien: Prov. Santiago d. Estero).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1936a: 490 (Hesperus; characters in key).
— Scheerpeltz, 1971: 191 (Hesperus; characters).
▴ Distribution: Argentina.
brunneipennis Scheerpeltz, 1971 [Note: Nomen nudum], see: rufipennis Gravenhorst, 1802.
buluensis Last, 1973: 432 (Hesperus; Type locality: New Guinea: Garaina, Bulu Valley).
nexilis Last, 1973: 432 (Hesperus; variety of buluensis; [Note: Nomen nudum (Article 15.2)]; cited from New Guinea: Upper Wario, 5000 ft.).
burgeoni Bernhauer, 1928g: 119 (Hesperus; Type locality: Belgisch-Kongo: Haut-Uele: Moto; Watsa).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1956b: 26 (Hesperus; characters; Belgian Congo).
— Scheerpeltz, 1971: 183 (Hesperus; characters).
▴ Distribution: Congo.
cafioides Fauvel, 1889c: 187 (Hesperus; Type locality: Liberia Africae).
— Fauvel, 1903e: 240 (Hesperus; Cameroon; Liberia; Assinie; Angola; Gabon; Congo; Sierra Leone).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Bernhauer, 1928g: 117 (Hesperus; characters in key).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Bernhauer, 1942c: 363 (Hesperus; Cameroon).
— Scheerpeltz, 1956b: 23 (Hesperus; characters; Liberia; Nigeria; Togo; Cameroon; French Congo; Belgian Congo).
— Scheerpeltz, 1971: 187 (Hesperus; characters).
— Scheerpeltz, 1971c: 170 (Hesperus; Cameroon).
▴ Distribution: Guinea, Sierra Leone, Liberia, Ivory Coast, Togo, Nigeria, Cameroon, Gabon, French Equatorial Africa, Congo, Angola.
laniger Fauvel, 1889b: 188 (Hesperus; Type locality: Stanley Pool, Congo).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Bernhauer, 1928g: 117 (Hesperus; variety of cafioides; characters in key).
— Scheerpeltz, 1933: 1372 (Hesperus; variety of cafioides).
— Scheerpeltz, 1956b: 23 (Hesperus; cited as valid species; characters; Belgian Congo).
— Levasseur, 1966a: 1214 (Hesperus; Cameroon; Guinea; Ivory Coast; Congo).
— Scheerpeltz, 1971: 187 (Hesperus; variety of cafioides; characters).
candidus Last, 1968: 353 (Philonthus; Type locality: New Guinea: Okapa).
— Last, 1973: 425 (Hesperus).
— Last, 1987: 49 (Hesperus; Papua New Guinea).
▴ Distribution: New Guinea.
caulis Last, 1981: 128 (Hesperus; Type locality: Goroka).
chinensis Cameron, 1940d: 252 (Hesperus; Type locality: China).
cingulatus Cameron, 1926, see: feae Fauvel, 1895.
coelestinus Bernhauer, 1926b: 41 (Hesperus; Type locality: Luzon: Balbalan).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1971: 174 (Hesperus; cited as coelestis; characters).
▴ Distribution: Philippines.
colmani Last, 1987: 33 (Hesperus; Type locality: Papua Guinea: New Guinea NE: Mount Missim, 2100 m).
coloratus Last, 1987: 34 (Hesperus; Type locality: Papua New Guinea: New Guinea NE: Torricelli Mts., Mobetei, 750 m).
conciliatus Last, 1981: 129 (Hesperus; Type locality: Daulo Pass, Asato-Chimbu Divide).
confusus Last, 1981: 128 (Hesperus; Type locality: Goroka, 8400 feet).
copiosus Last, 1968: 359 (Philonthus; Type locality: New Guinea: Okapa).
— Last, 1973: 425 (Hesperus).
— Last, 1987: 49 (Hesperus; Papua New Guinea).
▴ Distribution: New Guinea.
delectus Last, 1968: 365 (Philonthus; Type locality: New Guinea: Purosa).
depressus Bernhauer, 1908c: 109 (Hesperus; Type locality: Kamerun).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Bernhauer, 1928g: 119 (Hesperus; characters in key; Belgian Congo; Cameroon).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1971: 182 (Hesperus; characters).
▴ Distribution: Cameroon, Congo.
diabolicus Bernhauer, 1926b: 42 (Hesperus; Type locality: Luzon: Mt. Polis).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1971: 170 (Hesperus; characters).
▴ Distribution: Philippines.
distincticornis Cameron, 1937, see: roepkei Bernhauer, 1911.
divinus Bernhauer, 1915g: 190 (Hesperus; Type locality: Deutsch-Neuguinea: Lordberg).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1971: 180 (Hesperus; characters).
▴ Distribution: New Guinea.
dohertyi Cameron, 1942c: 845 (Hesperus; Type locality: S. Celebes: Tamba).
drescheri Cameron, 1937: 16 (Hesperus; Type locality: Java: Batoerraden, G. Slamat; G. Tangkoeban Prahoe, alt. 4000–5000 ft).
effectus Last, 1987: 34 (Hesperus; Type locality: Papua New Guinea: New Guinea NE: Wau, 1200 m).
eidmanni Scheerpeltz, 1936a: 484, 491 (Hesperus; Type locality: Brasilien: Umgebung von Mendes, E. d. Rio).
excellens Cameron, 1952: 250 (Hesperus; Type locality: Misool Island: Fakal).
excelsus Cameron, 1931, see: Philonthus.
feae Fauvel, 1895: 260 (Hesperus; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Carin Asciuii Ghécu, 1400–1500 m; Sikkim, Darjeeling).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Gridelli, 1924: 189 (Hesperus; characters; Burma; India).
— Cameron, 1932: 158 (Hesperus; characters; Burma; India).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1971: 155, 157 (Hesperus; characters).
— Schillhammer, 1999a: 62 (Hesperus; species may be synonymous with amabilis Kraatz but further study is needed).
▴ Distribution: Myanmar, India.
cingulatus Cameron, 1926b: 361 (Hesperus; Type locality: Mussorie District, Arni Gad. Chakrata District, Bangar. Simla Hills, Theog, 6000 to 7600 feet).
— Cameron, 1932: 158 (Hesperus; synonym of feae).
— Scheerpeltz, 1933: 1372 (Hesperus; variety of feae).
— Scheerpeltz, 1971: 155, 158 (Hesperus; aberration of feae; characters).
ferrugineus Bernhauer, 1925: 36 (Hesperus; Type locality: Argentinien: Alta Gracia La Granja, Sierra de Cordoba).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1936a: 490 (Hesperus; characters in key).
— Scheerpeltz, 1971: 192 (Hesperus; characters).
▴ Distribution: Argentina.
flavoris Last, 1987: 36 (Hesperus; Type locality: Irian: Vogelkop, Fak Fak, S. coast of Bomberai, 100–700 m).
fraternus Bernhauer, 1928g: 117 (Hesperus; Type locality: Natal: Frere; Zululand).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1956b: 24 (Hesperus; characters; Natal; Zululand).
— Scheerpeltz, 1971: 185 (Hesperus; characters).
▴ Distribution: South Africa.
fraudator Last, 1981: 133 (Hesperus; Type locality: Madang District, Finisterre Mts., Budemu, c. 4000 feet).
gagativentris Lea, 1931: 374 (Hesperus; Type locality: Papua: Mount Lamington).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1971: 179 (Hesperus; characters).
▴ Distribution: New Guinea.
gestroi Bernhauer, 1915k: 302 (Hesperus; Type locality: Insel Fernando Poo: Moka, 1300–1500 m).
— Bernhauer, 1915o: 363 (Hesperus; Fernando Poo).
— Bernhauer, 1928g: 116 (Hesperus; characters in key).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1956b: 21 (Hesperus; characters).
— Scheerpeltz, 1971: 189 (Hesperus; characters).
▴ Distribution: Equatorial Guinea.
giluensis Last, 1987: 36 (Hesperus; Type locality: Papua New Guinea: Mount Giluwe, 2800–3200 m).
glaucus Last, 1981: 133 (Hesperus; Type locality: New Guinea: Lufa).
gloriosus Lea, 1925: 227 (Hesperus; Type locality: Queensland: Coen River; Cairns District).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1971: 179 (Hesperus; characters).
▴ Distribution: Australia.
graciliventris Cameron, 1952: 254 (Hesperus; Type locality: New Guinea: Snow Mountains: Araucaria Camp, 800 m).
obscuritus Last, 1987: 48 (Hesperus; variety of graciliventris; [Note: Nomen nudum (Article 15.2)]; cited from Papua New Guinea: Wau, Big Wau Creek, 1300 m; Morobe District, Wau, 1300 m; Anga Gorge E of Mendi; Huon Pen., Pindiu. Irian: Vogelkop, bomberai, 700–900 m).
gratiosus Cameron, 1943a: 344 (Hesperus; Type locality: New Guinea: Finsch Haven).
gratus Cameron, 1943a: 342 (Philonthus; [preoccupied]; Type locality: New Guinea: Wareo, Finsch Haven).
— Last, 1987: 49 (Hesperus; Papua New Guinea).
— Herman, 2001: 45 (Hesperus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: New Guinea.
gressitti Last, 1987: 36 (Hesperus; Type locality: Papua New Guinea: New Guinea NE: Karimui, south of Goroka, 1000 m).
gridellianus Bernhauer, 1928g: 117 (Hesperus; Type locality: Portugiesisch-Guinea: Rio Cassine; Bahia de San Carlos).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1956b: 22 (Hesperus; characters).
— Scheerpeltz, 1971: 188 (Hesperus; characters).
▴ Distribution: Guinea-Bissau.
gridellii Cameron, 1926, see: Philonthus.
haematurus Erichson, 1840, see: apicialis Say, 1830.
haemorrhoidalis W. J. MacLeay, 1873: 140 (Philonthus; Type locality: Gayndah).
— Fauvel, 1877: 260 (Hesperus; characters; Australia).
— Olliff, 1887: 508 (Hesperus; characters; Australia).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Lea, 1925: 227 (Hesperus; characters; Australia).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1971: 178 (Hesperus; characters).
▴ Distribution: Australia.
mirabilis Fauvel, 1877: 260 (Hesperus; Type locality: Australie, Queensland, Wide Bay).
— Fauvel, 1878g: 543 (Hesperus; Australia).
— Olliff, 1887: 508 (Hesperus; synonym of haemorrhoidalis).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Bernhauer, 1920: 16 (Hesperus; Australia).
— Lea, 1925: 227 (Hesperus; synonym of haemorrhoidalis).
— Scheerpeltz, 1933: 1372 (Hesperus; synonym of haemorrhoidalis).
— Scheerpeltz, 1971: 178 (Hesperus; synonym of haemorrhoidalis).
hagenensis Last, 1987: 37 (Hesperus; Type locality: Papua New Guinea: New Guinea NE: Tomba, slopes of Mt. Hagen, 2450 m).
heinrichi Last, 1975a: 6 (Hesperus; Type locality: Papua: Madang).
heynei Wendeler, 1927: 7 (Hesperus; Type locality: Costa Rica: Turrialba, 800–900 m).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1936a: 489 (Hesperus; characters in key).
— Scheerpeltz, 1971: 190 (Hesperus; characters).
▴ Distribution: Costa Rica.
hilleri Last, 1987: 37 (Hesperus; Type locality: Papua New Guinea: New Guinea NE: Karimui, S of Goroka, 1000 m).
hirsutus Last, 1973: 431 (Hesperus; Type locality: New Guinea: Garaina).
hobbyi Bernhauer, 1936e: 214 (Hesperus; Type locality: Sarawak: foot of Mt. Dulit, Junction of rivers Tinjar and Lejok).
holzschuhi Schillhammer, 1991: 52 (Hesperus; Type locality: E-Nepal: Koshi, Basantapur, 2300 m).
honestus Last, 1981: 120 (Philonthus; Type locality: New Guinea: Mount Hagen).
hornabrooki Last, 1968: 354 (Philonthus; Type locality: New Guinea: Okapa).
— Last, 1973: 425 (Hesperus).
— Last, 1987: 49 (Hesperus; Papua New Guinea; Irian).
▴ Distribution: New Guinea.
huonensis Last, 1981: 129 (Hesperus; Type locality: Tep Tep, Finnisterre Range, Huon Pen).
inaequalis Fauvel, 1895: 258 (Hesperus; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Carin Ghécu, 1300–1400 m).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Gridelli, 1924: 193 (Hesperus; characters; Burma).
— Cameron, 1932: 160 (Hesperus; characters; Burma).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1965: 108 (Hesperus; Burma).
— Scheerpeltz, 1971: 162 (Hesperus; characters).
▴ Distribution: Myanmar.
indigaceus Fauvel, 1878g: 544 (Hesperus; Type locality: Nouvelle-Calédonie, Ile des Pins).
— Fauvel, 1889b: 263 (Hesperus; New Caledonia).
— Fauvel, 1903d: 269 (Hesperus; Iles des Pins).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1966e: 402 (Hesperus; characters; New Caledonia).
— Scheerpeltz, 1971: 181 (Hesperus; characters).
▴ Distribution: New Caledonia.
ignoratus Ito, 1994c: 81 (Hesperus; Type locality: Japan: Kyoto Pref.: Ogoshi).
interjectus Last, 1987: 37 (Hesperus; Type locality: Papua New Guinea NE: Karimui, S of Goroka, 1000 m. Also cited from Irian).
javanus Cameron, 1937: 15 (Hesperus; Type locality: Java: Batoerraden, G. Slamat).
junius Last, 1987: 38 (Hesperus; Type locality: Papua New Guinea: New Guinea NE: Daulo Pass, 2500 m, Asaro-Chimbu Div.).
justus Last, 1968: 348 (Philonthus; Type locality: New Guinea: Okapa).
— Last, 1973: 425 (Hesperus).
— Last, 1987: 50 (Hesperus; Papua New Guinea).
▴ Distribution: New Guinea.
kaindiensis Last, 1980: 155 (Hesperus; Type locality: New Guinea: Wau, Mt. Kaindi, 2000 m).
kokodanus Cameron, 1937a: 101 (Philonthus; Type locality: New Guinea: Kokoda, alt. 1200 ft.).
kambaitiensis Scheerpeltz, 1965: 242 (Hesperus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).
kraatzi Eppelsheim, 1895a: 129 (Hesperus; Type locality: Togoland).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Bernhauer, 1928g: 116 (Hesperus; characters in key).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1956a: 485 (Hesperus; Guinea; Liberia; Nigeria; Cameroon; French Congo; Belgian Congo; Uganda; Kenya; Tanganyika).
— Scheerpeltz, 1956b: 22 (Hesperus; characters).
— Scheerpeltz, 1971: 188 (Hesperus; characters).
▴ Distribution: Guinea, Liberia, Nigeria, Cameroon, French Equatorial Africa, Congo, Uganda, Kenya, Tanzania.
laevigatus Fauvel, 1895: 259 (Hesperus; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Pulo-Penang; Malacca; Singapore; Sumatra; Borneo; Java).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Cameron, 1921a: 378, 405 (Hesperus; characters; Singapore).
— Gridelli, 1924: 183 (Hesperus; characters; Burma; Sumatra).
— Bernhauer, 1928: 17 (Hesperus; Sumatra).
— Cameron, 1932: 157 (Hesperus; characters; Burma).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Cameron, 1933d: 390 (Hesperus; Sumatra).
— Scheerpeltz, 1965: 108 (Hesperus; Burma).
— Scheerpeltz, 1971: 159 (Hesperus; characters).
— Frank, 1982: 36 (Hesperus; list of parasites).
▴ Distribution: Myanmar, Malaysia, Singapore, Borneo, Indonesia.
sarawakensis Gridelli, 1924: 185 (Hesperus; variety of laevigatus; Type locality: Borneo: Sarawak).
— Scheerpeltz, 1933: 1373 (Hesperus; variety of laevigatus).
— Scheerpeltz, 1971: 159 (Hesperus; variety of laevigatus; characters).
laniger Fauvel, 1889, see: cafioides Fauvel, 1889.
levitus Last, 1987: 38 (Hesperus; Type locality: Papua New Guinea NE: Morobe Distr., Wau, 1200 m. Also cited from New Britain).
lifuanus Fauvel, 1903d: 269 (Hesperus; Type locality: Lifou).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Scheerpeltz, 1971: 179 (Hesperus; characters).
▴ Distribution: New Caledonia.
locus Last, 1987: 39 (Hesperus; Type locality: Papua New Guinea NE: Daulo Pass, Asaro-Chimbu Div., 2400 m).
longicornis Cameron, 1950e: 430 (Hesperus; Type locality: Cameroon: Ngauyanga).
longipennis Cameron, 1952: 253 (Hesperus; Type locality: New Guinea: Snow Mountains: Lower Mist Camp, 1700 m).
luluanus Scheerpeltz, 1956b: 23 (Hesperus; Type locality: Belgisch-Kongo).
luluanus Scheerpeltz, 1971: 184 (Hesperus; [preoccupied]; Type locality: Kongo Gebiet und Uganda: Kapanga).
luluanus Scheerpeltz, 1971, see: luluanus Scheerpeltz, 1956.
luzonicus Bernhauer, 1926b: 43 (Hesperus; Type locality: Luzon: Balbalan).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1971: 171 (Hesperus; characters).
▴ Distribution: Philippines.
maai Last, 1987: 39 (Hesperus; Type locality: Irian: Vogelkop, Bomberai, 700–900 m).
maculicollis Bernhauer, 1926b: 43 (Hesperus; Type locality: Mindanao: Surigao, Panaon).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1971: 161 (Hesperus; characters).
▴ Distribution: Philippines.
malaisei Scheerpeltz, 1965: 240 (Hesperus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).
malayanus Cameron, 1932a: 142 (Hesperus; Type locality: Selangor: Bukit Kutu).
mandatus Last, 1987: 39 (Hesperus; Type locality: Papua New Guinea: New Guinea NE: Morobe Distr., Wau, 1200 m).
maris Last, 1987: 40 (Hesperus; Type locality: Papua New Guinea: New Guinea NE: Torricelli Mts., Mobitei, 750 m).
meyeri Bernhauer, 1928g: 119 (Hesperus; Type locality: Uganda).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1956b: 26 (Hesperus; characters; Uganda).
— Scheerpeltz, 1971: 182 (Hesperus; characters).
▴ Distribution: Uganda.
mirabilis Fauvel, 1877, see: haemorrhoidalis W. J. MacLeay, 1873.
mirificus Herman, 2001: 45 (Hesperus; replacement name for mirus Last).
mirus Last, 1981: 132 (Hesperus; [preoccupied]; Type locality: Frigano, Okapa, Eastern Highlands).
mirus Bernhauer, 1915d: 146 (Hesperus; Type locality: Kamerun).
— Bernhauer, 1915o: 363 (Hesperus; Cameroon).
— Bernhauer, 1928g: 116 (Hesperus; characters in key).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1956b: 21 (Hesperus; characters; Cameroon).
— Levasseur, 1968a: 52 (Hesperus; Central African Republic).
— Scheerpeltz, 1971: 187 (Hesperus; characters).
▴ Distribution: Cameroon, Central African Republic.
mirus Last, 1981, see: mirificus Herman, 2001.
mixtus Last, 1968: 366 (Philonthus; Type locality: New Guinea: Karimu).
— Last, 1973: 425 (Hesperus).
— Last, 1981: 128 (Hesperus; New Guinea).
— Last, 1987: 50 (Hesperus; Papua New Guinea).
▴ Distribution: New Guinea.
modestus Bernhauer, 1915h: 229 (Hesperus; Type locality: Java: Preanger).
— Gridelli, 1924: 199 (Hesperus; characters; Java).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1971: 170 (Hesperus; characters).
▴ Distribution: Indonesia.
modulatus Last, 1968: 358 (Philonthus; Type locality: New Guinea: Buana, Sarawaged Range, 3000 ft., 20–30 miles north of Lae).
— Last, 1973: 425, 426 (Hesperus; characters).
— Last, 1987: 50 (Hesperus; Papua New Guinea).
▴ Distribution: New Guinea.
monstratus Last, 1987: 40 (Hesperus; Type locality: Papua New Guinea: Owen Stanley Range, Goilala: Loloipa).
montanellus Bernhauer, 1926b: 43 (Hesperus; Type locality: Luzon: Mt. Banahao).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1971: 171 (Hesperus; characters).
▴ Distribution: Philippines.
moresbyensis Last, 1987: 42 (Hesperus; Type locality: Papua New Guinea: Bisianumu, E of Port Moresby, 500 m).
moroensis Last, 1981: 132 (Hesperus; Type locality: Madang district, Finisterre, Mts., Moro, c. 5550 feet).
moultoni Bernhauer, 1915f: 143 (Hesperus; Type locality: Britisch-Nord-Borneo: Mt. Kinabalu).
— Gridelli, 1924: 199 (Hesperus; characters; Borneo).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1971: 172 (Hesperus; characters).
▴ Distribution: Borneo.
multipunctus Last, 1981: 130 (Hesperus; Type locality: New Hebrides: West Santo, Tartarii, 4000 ft.).
natalensis Scheerpeltz, 1956b: 22 (Hesperus; Type locality: Natal).
natalensis Scheerpeltz, 1971: 188 (Hesperus; [preoccupied]; Type locality: Süd-Afrika: Natal).
natalensis Scheerpeltz, 1971, see: natalensis Scheerpeltz, 1956.
nexilis Last, 1973 [Note: Nomen nudum], see: buluensis Last, 1973.
nigriventris Gridelli, 1924: 182, 186 (Hesperus; Type locality: Isola Nias, Sumatra).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1971: 169 (Hesperus; characters).
▴ Distribution: Indonesia.
nobilis Bernhauer, 1928g: 119 (Hesperus; Type locality: Belgisch-Kongo: Bas, Uele).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1956b: 27 (Hesperus; characters; Belgian Congo).
— Scheerpeltz, 1971: 183 (Hesperus; characters).
▴ Distribution: Congo.
novoteutonicus Wendeler, 1956a: 232 (Hesperus; Type locality: Brasilien: Nova Teutonia).
obesus Last, 1989: 878 (Hesperus; Type locality: Papua New Guinea: Onerunka, near Kainantu).
obscuricollis Cameron, 1941c: 379 (Hesperus; Type locality: Philippines: Mindanao: Surigao).
obscuricollis Scheerpeltz, 1971, see: septuosus Herman, 2001.
obscuripennis Scheerpeltz, 1971 [Note: Nomen nudum], see: bakeri Bernhauer, 1919.
obscurior Bernhauer, 1915d: 145 (Hesperus; variety of belonuchiformis; Type locality: Abessinien: Errer; Djerer-Tal).
— Bernhauer, 1928g: 118 (Hesperus; characters in key).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1956b: 24 (Hesperus; characters; Abyssinia).
— Scheerpeltz, 1971: 185 (Hesperus; characters).
▴ Distribution: Ethiopia.
obscuritus Last, 1987 [Note: Nomen nudum], see: graciliventris Cameron, 1952.
orientalis Cameron, 1950d: 21 (Hesperus; Type locality: Perak: Larut Hills, 3000–3700 feet).
ornaticornis Wendeler, 1927: 6 (Hesperus; Type locality: Costa Rica: Turrialba, 800 m).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1936a: 489 (Hesperus; characters in key).
— Scheerpeltz, 1971: 190 (Hesperus; characters).
▴ Distribution: Costa Rica.
ornatus Sharp, 1889: 43 (Hesperus; Type locality: Hitoyoshi; Yuyama; Nikko).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Gridelli, 1924: 199 (Hesperus; characters; Japon).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1971: 156 (Hesperus; characters).
▴ Distribution: Japan.
orroriensis Cameron, 1937a: 101 (Philonthus; Type locality: New Guinea: Orrori, alt. 3000 ft.).
— Last, 1968: 364 (Philonthus; New Guinea).
— Last, 1987: 48 (Hesperus; Papua New Guinea).
▴ Distribution: New Guinea.
overlaeti Bernhauer, 1939g: 229 (Hesperus; Type locality: Belgisches Kongo: Lulua: Kapanga).
— Scheerpeltz, 1956b: 26 (Hesperus; characters; Belgian Congo).
— Scheerpeltz, 1971: 182 (Hesperus; characters).
▴ Distribution: Congo.
pacificus Olliff, 1887: 509 (Hesperus; Type locality: Lord Howe Island).
— Olliff, 1889: 81 (Hesperus; Lord Howe Island).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Scheerpeltz, 1971: 179 (Hesperus; characters).
▴ Distribution: Australia.
pendleburyi Cameron, 1936b: 45 (Hesperus; Type locality: Pahang: Cameron Highlands, 4000–5000 feet).
perakensis Cameron, 1950d: 20 (Hesperus; Type locality: Perak).
perfoliatus Gridelli, 1924: 182, 196 (Hesperus; subgenus Hesperotropis; Type locality: Pangherang-Pisang, Sumatra occ.).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1971: 193 (Hesperus; subgenus Hesperotropis; characters).
▴ Distribution: Indonesia.
perraulti Coiffait, 1976: 227 (Hesperus; Type locality: Tahiti: Papenoo).
phaenomenalis Bernhauer, 1914c: 102 (Hesperus; Type locality: Philippinen: Los Banos).
— Gridelli, 1924: 200 (Hesperus; characters; Philippines).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1971: 174 (Hesperus; characters).
▴ Distribution: Philippines.
viridis Bernhauer, 1926b: 40 (Hesperus; variety of phaenomenalis; Type locality: Tayabas, Malinao).
— Scheerpeltz, 1933: 1373 (Hesperus; variety of phaenomenalis).
— Scheerpeltz, 1971: 174 (Hesperus; variety of phaenomenalis; characters).
picticollis Gridelli, 1924: 183, 194 (Hesperus; Type locality: Isole Mentawei; Sipor Sereinu).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1971: 164 (Hesperus; characters).
▴ Distribution: Indonesia.
picticornis Lea, 1925: 227 (Hesperus; Type locality: New South Wales: Tweed River).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1971: 177 (Hesperus; characters).
▴ Distribution: Australia.
plagiarius Last, 1987: 42 (Hesperus; Type locality: Papua New Guinea: New Guinea NE: Huon Peninsula, Pindiu).
plumbeus Last, 1981: 127 (Hesperus; Type locality: Morobe Dist., Edie CK, Bulldog Road, 9700 ft.).
postulator Last, 1987: 42 (Hesperus; Type locality: Papua New Guinea: New Guinea NE: Morobe Distr., Wau, 1300 m).
preangeranus Cameron, 1937: 16 (Hesperus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft).
primitus Last, 1987: 43 (Hesperus; Type locality: Irian: Waris S. of Jayapura, Hollandia, 450–500 m. Also cited from Papua New Guinea).
privus Last, 1973: 429 (Hesperus; Type locality: Okapa).
probatus Last, 1987: 43 (Hesperus; Type locality: Papua New Guinea: New Guinea NE: Aiyura, 1600 m).
pseudobirmanus Scheerpeltz, 1965: 238 (Hesperus; Type locality: N.E. Burma, Kambaiti, 2000 m).
pseudoviolaceus Last, 1981: 130 (Hesperus; Type locality: Wantout, Huon Peninsular).
pulleinei Olliff, 1887: 512 (Hesperus; [Note: Olliff's description was published before Blackburn's]; Type locality: Burnside, Adelaide, South Australia).
— Blackburn, 1888: 7 (Hesperus; [Note: Bernhauer and Schubert, 1914: 364 cited Blackburn as the author for this name, and although Olliff also cited Blackburn as the author, Olliff's description appeared first]; characters; Australia).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Scheerpeltz, 1971: 180 (Hesperus; characters).
▴ Distribution: Australia.
puncticeps Bernhauer, 1928g: 118 (Hesperus; Type locality: Uganda: Entebbe).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1956b: 25 (Hesperus; characters; Uganda).
— Scheerpeltz, 1971: 186 (Hesperus; characters).
▴ Distribution: Uganda.
punctipennis Cameron, 1941c: 380 (Hesperus; Type locality: Philippines: Tangcolan: Bukidnon).
purpuripennis Bernhauer, 1915g: 189 (Hesperus; Type locality: Ralums).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1971: 180 (Hesperus; characters).
— Last, 1987: 50 (Hesperus; Papua New Guinea).
▴ Distribution: New Guinea.
quinquecolor Lea, 1931: 372 (Hesperus; Type locality: New Guinea: Wareo).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1971: 180 (Hesperus; characters).
▴ Distribution: New Guinea.
rarus Last, 1989: 880 (Hesperus; Type locality: Papua New Guinea: Kainantu, Onerunka).
raynori Last, 1987: 44 (Hesperus; Type locality: Papua New Guinea: Owen Stanley Range, Goilala, Loloipa).
ricardi Last, 1973: 431 (Hesperus; Type locality: New Guinea: Upper Wario, 5000 ft.).
transformis Last, 1981: 132 (Hesperus; variety of ricardi, cited as richardi; [Note: Nomen nudum (Article 15.2)]; cited from Wantoat Valley, N. Guinea).
roepkei Bernhauer, 1911: 89 (Hesperus; Type locality: Java: Bandoeng; Borneo: Sarawak).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Gridelli, 1924: 185 (Hesperus; characters; Sumatra; Isole Mentawei).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1971: 160 (Hesperus; characters).
▴ Distribution: Borneo, Indonesia.
distincticornis Cameron, 1937: 15 (Hesperus; Type locality: Java: Batoerraden, G. Slamat; Bandoeng Dago; Tjilatjap; Patimoean, Zuid Preanger).
rubidus Last, 1987: 44 (Hesperus; Type locality: Papua New Guinea: New Guinea NE: Eliptamin Valley, 1200–1350 m).
ruficeps Cameron, 1941c: 380 (Hesperus; Type locality: Philippines: Tangcolan: Bukidnon).
rufipennis Gravenhorst, 1802: 40 (Staphylinus; [preoccupied]; Type locality: Not cited, but presumably Brunsvigae).
— Latreille, 1804: 327 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 104 (Staphylinus).
— Nordmann, 1837: 110 (Philonthus; Germany).
— Erichson, 1839a: 477 (Philonthus; characters; Germany).
— Erichson, 1840: 504 (Philonthus; characters; Germany; France; Sardinia).
— Heer, 1841: 583 (Philonthus; characters; Switzerland).
— Redtenbacher, 1849: 825 (Philonthus; characters; Austria).
— Küster, 1854: 17 (Philonthus; characters; Germany; France; Switzerland; Sardinia).
— Fairmaire and Laboulbène, 1856: 530 (Philonthus; characters; France).
— Kraatz, 1857d: 621 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 197 (Philonthus; characters; Austria).
— Fauvel, 1874: 200 [= 1874a: 426] (Hesperus; characters; France; Germany; Austria; Italy; Sardinia).
— Redtenbacher, 1874: 215 (Philonthus; characters; Austria).
— Mulsant and Rey, 1876: 562 [= 1877: 418] (Cafius; characters; France).
— Seidlitz, 1891: 390 (Philonthus; subgenus Hesperus; characters; Europe).
— Seidlitz, 1891a: 415 (Philonthus; subgenus Hesperus; characters; Europe).
— Ganglbauer, 1895: 462 (Hesperus; characters; middle Europe; Italy; Sardinia).
— Fauvel, 1902b: 107 (Hesperus; Morocco).
— Montandon, 1908: 79 (Hesperus; Romania).
— Reitter, 1909: 123 (Hesperus; characters; Germany).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Gridelli, 1924: 188 (Hesperus; characters; Italy; Morocco; Sardinia).
— Portevin, 1929: 355 (Hesperus; characters; France).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Smetana, 1958: 258 (Hesperus; characters; ecology; Czechoslovakia).
— Kocher, 1958: 152 (Hesperus; Morocco).
— Lohse, 1964: 173 (Hesperus; characters; central Europe).
— Smetana, 1964c: 168 (Hesperus; Czechoslovakia).
— Horion, 1965: 121 (Hesperus; France; Switzerland; Italy; Sardinia; Hercegovina; Serbia; Romania; Czech Republic; Slovakia; Poland; Germany; Austria).
— Scheerpeltz, 1968a: 79 (Hesperus; catalog; Austria).
— Scheerpeltz, 1971: 154 (Hesperus; characters).
— Coiffait, 1974: 321 (Hesperus; characters; Europe; Italy; Sardinia; Morocco).
— Burakowski, Mroczkowski, and Stefańska, 1980: 33 (Hesperus; catalog; Poland).
— Tóth, 1984: 41 (Hesperus; characters; Hungary).
— Lucht, 1987: 104 (Hesperus; checklist; Germany; Poland; Czechoslovakia; Austria; France).
— Schülke and Uhlig, 1988: 11 (Hesperus; notes; Germany).
— Gusarov, 1989: 11 (Hesperus; habitat notes; Ukraine).
— Drugmand and Coulon, 1993: 239 (Hesperus; collecting notes; Luxembourg).
— Smetana, 1993b: 48 (Hesperus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 27 (Hesperus; Italy).
— Herman, 2001: 45 (Hesperus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Europe, Morocco.
brunneipennis Scheerpeltz, 1971: 154 (Hesperus; aberration of rufipennis; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from nördlichere Teile des Verbreitungsgebietes).
rufipennis Cameron, 1918, see: borneensis Bernhauer, 1915.
sarawakensis Gridelli, 1924, see: laevigatus Fauvel, 1895.
saltus Last, 1968: 353 (Philonthus; Type locality: New Guinea: Okapa).
— Last, 1973: 425 (Hesperus).
— Last, 1987: 50 (Hesperus; Papua New Guinea).
▴ Distribution: New Guinea.
schedli Scheerpeltz, 1956b: 22, 27 (Hesperus; Type locality: Belgisch-Kongo: Hembe Bitale [Note: Type locality cited in Scheerpeltz, 1956a]).
scutellaris Bernhauer, 1926b: 40 (Hesperus; Type locality: Luzon: Mt. Makiling).
— Scheerpeltz, 1933: 1373 (Hesperus; catalog).
— Scheerpeltz, 1971: 175 (Hesperus; characters).
▴ Distribution: Philippines.
segmentarius Cameron, 1941c: 379 (Hesperus; Type locality: Not cited, presumably the Philippines).
semicoeruleus Bernhauer, 1915h: 229 (Hesperus; Type locality: Java: Preanger).
— Gridelli, 1924: 200 (Hesperus; characters; Java).
— Scheerpeltz, 1933: 1373 (Hesperus; cited as semicaeruleus; catalog).
— Scheerpeltz, 1935a: 634 (Hesperus; cited as semicaeruleus; Indonesia).
— Scheerpeltz, 1971: 173 (Hesperus; cited as semicaeruleus; characters).
▴ Distribution: Indonesia.
varicornis Cameron, 1937: 15 (Hesperus; Type locality: Java: G. Tangkoeban Prahoe, Preanger, alt. 4000–5000 ft).
semirufus Fauvel, 1878g: 543 (Hesperus; Type locality: Australie septentrionale).
— Olliff, 1887: 511 (Hesperus; characters; Australia).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Scheerpeltz, 1971: 176 (Hesperus; characters).
— Bellas, Brown, and Moore, 1974 (Hesperus; defensive secretions).
▴ Distribution: Australia.
septuosus Herman, 2001: 45 (Hesperus; replacement name for obscuricollis Scheerpeltz).
obscuricollis Scheerpeltz, 1971: 168 (Hesperus; [preoccupied]; Type locality: Philippinen: Luzon: Los Baños, Subuagra).
serpentinus Horn, 1884: 219 (Philonthus; Type locality: Kentucky).
— Bernhauer and Schubert, 1914: 355 (Philonthus; catalog).
— Smetana, 1995e: 490 (Hesperus; characters; notes; Alabama; Florida; Kentucky; North Carolina; Pennsylvania).
— Downie and Arnett, 1996: 381 (Philonthus; characters; USA).
▴ Distribution: USA.
setosus Cameron, 1952: 251 (Hesperus; Type locality: New Guinea: Snow Mountains: Moss Forest Camp, 2800 m).
similis Last, 1987: 45 (Hesperus; Type locality: Irian: Genjam, 40 km W of Jayapura, 100–200 m).
singularis Last, 1973: 428 (Philonthus; Type locality: New Guinea: Garaina, Bula Valley).
sociabilis Last, 1987: 45 (Hesperus; Type locality: Papua New Guinea: Madang).
solivagus Last, 1981: 133 (Hesperus; Type locality: Lufa, Mount Michael).
sparsiceps Bernhauer, 1928g: 116 (Hesperus; Type locality: Insel Fernando Po: Basile, 400–600 m).
— Scheerpeltz, 1933: 1374 (Hesperus; catalog).
— Bernhauer, 1942c: 363 (Hesperus; Cameroon).
— Scheerpeltz, 1956b: 21 (Hesperus; characters).
— Scheerpeltz, 1971: 188 (Hesperus; characters).
▴ Distribution: Equatorial Guinea.
sparsior Bernhauer, 1928g: 118 (Hesperus; Type locality: Tanganyika: Terr.: Konga).
— Scheerpeltz, 1933: 1374 (Hesperus; catalog).
— Scheerpeltz, 1956b: 25 (Hesperus; characters; Tanganyika).
— Scheerpeltz, 1971: 186 (Hesperus; characters).
— Scheerpeltz, 1974c: 24 (Hesperus; Ethiopia; Sudan).
▴ Distribution: Sudan, Ethiopia, Tanzania.
sparsus Last, 1987: 45 (Hesperus; Type locality: Papua New Guinea: New Guinea NE: Finisterre Range, Saidor, Kaimbaui Village).
spectabilis Bernhauer, 1928g: 118 (Hesperus; Type locality: Natal: Malvern).
— Scheerpeltz, 1933: 1374 (Hesperus; catalog).
— Scheerpeltz, 1956b: 25 (Hesperus; characters; Natal).
— Scheerpeltz, 1971: 186 (Hesperus; characters).
▴ Distribution: South Africa.
spinosus Last, 1981: 126 (Hesperus; Type locality: Kassem Pass).
splendoris Last, 1987: 46 (Hesperus; Type locality: Papua New Guinea: New Guinea NE: Mt. Amingwiwa, 3000–3400 m).
stehri Moore, 1958: 313 (Hesperus; Type locality: Ash Cave, Hocking County, Ohio).
— Smetana, 1995e: 502 (Hesperus; characters; notes; Arkansas; Georgia; Kansas; Ohio; Oklahoma; Tennessee).
— Downie and Arnett, 1996: 383 (Hesperus; characters; USA).
▴ Distribution: USA.
strigiventris Eppelsheim, 1895a: 127 (Philonthus; Type locality: Togoland).
— Fauvel, 1903e: 242 (Philonthus; Cameroon; Togo; Gabon).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Bernhauer, 1915o: 363 (Hesperus; Belgian Congo).
— Bernhauer, 1928g: 118 (Hesperus; characters in key).
— Scheerpeltz, 1933: 1374 (Hesperus; catalog).
— Scheerpeltz, 1956b: 26 (Hesperus; characters; Togo; Guinea).
— Tottenham, 1962: 144 (Hesperus; Uganda).
— Levasseur, 1966a: 1214 (Hesperus; Congo; Togo; Gabon; Zaire).
— Levasseur, 1968a: 52 (Hesperus; Central African Republic).
— Scheerpeltz, 1971: 183 (Hesperus; characters).
▴ Distribution: Guinea, Togo, Cameroon, Central African Republic, Gabon, Congo, Uganda.
subgrandis Last, 1987: 46 (Hesperus; Type locality: Irian: Sentani, 90 m).
subsimilis Cameron, 1933d: 390 (Hesperus; Type locality: Sumatra: Gunung Singgalang, 1800 m).
sumatrensis Cameron, 1933d: 390 (Hesperus; Type locality: Sumatra: Fort de Kock).
taiwanensis Shibata, 1973: 21 (Hesperus; Type locality: Taiwan: Koantauchi, Nantou Hsien).
tapiniensis Last, 1987: 46 (Hesperus; Type locality: Papua New Guinea: New Guinea SE: Tapini, 1000 m).
tectus Last, 1968: 364 (Philonthus; Type locality: New Guinea: Buana, Sarawaged Range, 3000 ft., 20–30 miles north of Lae).
temporatus Last, 1968: 348 (Philonthus; Type locality: New Guinea: Okapa).
— Last, 1973: 425, 426 (Hesperus; characters).
— Last, 1987: 50 (Hesperus; Papua New Guinea).
▴ Distribution: New Guinea.
tenuis Last, 1973: 430 (Hesperus; Type locality: Okapa).
terminalis Cameron, 1918a: 104 (Hesperus; Type locality: Java: Silabintanah).
— Gridelli, 1924: 200 (Hesperus; characters; Java).
— Scheerpeltz, 1933: 1374 (Hesperus; catalog).
— Scheerpeltz, 1971: 171 (Hesperus; characters).
▴ Distribution: Indonesia.
terminicornis Gridelli, 1924: 182, 192 (Hesperus; Type locality: Beccari a Sumatra, Monte Singalang).
— Scheerpeltz, 1933: 1374 (Hesperus; catalog).
— Scheerpeltz, 1971: 169 (Hesperus; characters).
▴ Distribution: Indonesia.
tiro Sharp, 1874: 49 (Philonthus; Type locality: Copper Temple, Nagasaki).
— Bernhauer and Schubert, 1914: 358 (Philonthus; catalog).
— Scheerpeltz, 1933: 1374 (Hesperus; catalog).
— Nakane, 1963a: 92 (Hesperus; characters; habitus photograph; Japan).
— Scheerpeltz, 1971: 156 (Hesperus; characters).
— J. Li, 1993: 40 (Hesperus; China).
— Watanabe, 1996a: 12 (Hesperus; mine collection; Japan).
▴ Distribution: Japan, China.
tokyanus R. Dvořák, 1957: 9 (Hesperus; Type locality: Mount Takar, Tokyo, Japon).
toxopei Cameron, 1952: 253 (Hesperus; Type locality: New Guinea: Snow Mountains: Rattan Camp, 1200 m).
transformis Last, 1981 [Note: Nomen nudum], see: ricardi Last, 1973.
tremolerasi Bernhauer, 1927e: 246 (Hesperus; Type locality: Uruguay: Montevideo).
— Scheerpeltz, 1933: 1374 (Hesperus; catalog).
— Scheerpeltz, 1936a: 490 (Hesperus; characters in key).
— Scheerpeltz, 1971: 191 (Hesperus; characters).
▴ Distribution: Uruguay.
tricolor Cameron, 1952: 251 (Hesperus; Type locality: New Guinea: Snow Mountains: Moss Forest Camp, 2800 m).
trimerus Fauvel, 1895: 261 (Hesperus; Type locality: Bírmanie, Carin Cheba, 900–1100 m).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Gridelli, 1924: 189 (Hesperus; characters; Burma).
— Cameron, 1932: 159 (Hesperus; characters; Burma; India).
— Scheerpeltz, 1933: 1374 (Hesperus; catalog).
— Scheerpeltz, 1965: 108 (Hesperus; Burma).
— Scheerpeltz, 1971: 158 (Hesperus; characters).
▴ Distribution: Myanmar, India.
uelensis Bernhauer, 1932b: 153 (Hesperus; Type locality: Haut-Uele: Moto).
— Scheerpeltz, 1933: 1374 (Hesperus; catalog).
— Scheerpeltz, 1956b: 25 (Hesperus; characters; Belgian Congo).
— Scheerpeltz, 1971: 181 (Hesperus; characters).
▴ Distribution: Congo.
uhligi Last, 1990: 1 (Hesperus; Type locality: Deutsch Neu Guinea: Mäanderberg, 470 m).
varicornis Cameron, 1937, see: semicaeruleus Bernhauer, 1915.
venator Last, 1987: 48 (Hesperus; Type locality: Irian: Vogelkop, Bomberai, 700–900 m).
venustulus Last, 1968: 351 (Philonthus; Type locality: New Guinea: Okapa).
venustus Last, 1968: 350 (Philonthus; Type locality: New Guinea: Agakomatuso).
vethi Bernhauer, 1915h: 230 (Hesperus; Type locality: Java: Preanger).
— Gridelli, 1924: 191 (Hesperus; characters; Java).
— Scheerpeltz, 1933: 1374 (Hesperus; catalog).
— Scheerpeltz, 1971: 165 (Hesperus; characters).
▴ Distribution: Indonesia.
viridiaureus Bernhauer, 1926b: 41 (Hesperus; variety of phaenomenalis; Type locality: Mindanao: Surigao; Luzon: Imugan).
— Scheerpeltz, 1933: 1374 (Hesperus; catalog).
— Scheerpeltz, 1971: 175 (Hesperus; characters).
▴ Distribution: Philippines.
viridipennis Cameron, 1952: 249 (Hesperus; Type locality: Misool Island).
viridis Bernhauer, 1926, see: phaenomenalis Bernhauer, 1914.
viridisplendens Scheerpeltz, 1971: 175 (Hesperus; Type locality: Philippinen: S. O. Luzon: Mt. Balusan).
vividus Cameron, 1931, see: Philonthus.
wasmanni Fauvel, 1895: 260 (Hesperus; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Sikkim, Darjeeling).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Gridelli, 1924: 187 (Hesperus; characters; Burma; India).
— Cameron, 1932: 158 (Hesperus; characters; Burma; India).
— Scheerpeltz, 1933: 1374 (Hesperus; catalog).
— Scheerpeltz, 1971: 155, 158 (Hesperus; characters).
▴ Distribution: Myanmar, India.
zischkai Scheerpeltz, 1971: 192 (Hesperus; Type locality: Süd-ost-Bolivien in der Sierra de Cochabamba).
Hybridolinus
[8 species; Palaearctic Region]
Hybridolinus Schillhammer, 1998a: 146 (species included: hesperoides, singularis, diabolicus, daliensis, similis, jizushanus, decipiens). Type species: Hybridolinus daliensis Schillhammer, fixed by original designation.
daliensis Schillhammer, 1998a: 149 (Hybridolinus; Type locality: China: Yunnan, Dali).
decipiens Schillhammer, 1998a: 150 (Hybridolinus; Type locality: China: Yunnan, Baishui).
diabolicus Schillhammer, 1998a: 154 (Hybridolinus; Type locality: Taiwan: Taichung Hsien, Anmashan, 2230 m).
flavocinctus Schillhammer, 2000b: 143 (Hybridolinus; Type locality: China: SE Sichuan: Jinfo Shan, 29°01N, 107°14E, 1750 m).
hesperoides Schillhammer, 1998a: 154 (Hybridolinus; Type locality: China: Kuatun, 2300 m).
jizushanus Schillhammer, 1998a: 152 (Hybridolinus; Type locality: China: Yunnan, Jizu-Shan).
similis Schillhammer, 1998a: 150 (Hybridolinus; Type locality: China: Sichuan, Daxue Shan, Gongga Shan Mt., Hailuogou glacier park, 102.04 E, 29.36 N, river valley ca. 1 km above Camp 1, 2100 m).
singularis Schillhammer, 1998a: 152 (Hybridolinus; Type locality: China: Yunnan, Jizu-Shan).
Jurecekia
[2 species; Palaearctic Region]
Jurecekia Rambousek, 1920: 16, 19 (species included: paradoxa). Type species: Jurecekia paradoxa Rambousek, fixed by monotypy.
— Winkler, 1925: 381 (catalog for Palaearctic region).
— Cameron, 1932: 62 (subgenus of Philonthus).
— Scheerpeltz, 1933: 1371 (world catalog supplement).
— Blackwelder, 1952: 206 (type species: paradoxa).
— Tikhomirova, 1973a: 182 (checklist of species of USSR).
— Coiffait, 1974: 316 (characters; key to species of western Palaearctic region).
— Smetana, 1995e: 50 (characters).
asphaltina Erichson, 1840: 436 (Philonthus; Type locality: Germania meridionalis [Note: Erroneous type locality, see Horion, 1965: 138]).
— Kraatz, 1857d: 576 (Philonthus; characters; Germany).
— Redtenbacher, 1874: 216 (Philonthus; characters).
— Ganglbauer, 1895: 445 (Philonthus; characters; southern Germany).
— Bernhauer, 1899a: 434 (Philonthus; characters; Austria).
— Reitter, 1909: 126 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 329 (Philonthus; catalog).
— Scheerpeltz, 1933: 1371 (Jurecekia; catalog).
— Horion, 1965: 138 (Jurecekia; species not found in central Europe).
— Coiffait, 1974: 317 (Jurecekia; characters; Russia; Transcaucasus; Germany).
▴ Distribution: Russia, Transcaucasus.
paradoxa Rambousek, 1920: 17, 19 (Jurecekia; Type locality: Russie aux environs de Totskoe, (Gouber. Samara) au bord de Samarka).
— Rambousek, 1923: 54 (Philonthus; synonym of asphaltinus).
— Scheerpeltz, 1933: 1371 (Jurecekia; synonym of asphaltina).
— Coiffait, 1974: 318 (Jurecekia; synonym of asphaltina).
paradoxa Rambousek, 1921, see: asphaltina Erichson, 1840.
rufipes Coiffait, 1969b: 39 (Jurecekia; Type locality: Atlosar région d'Akmolinsk, Kazakstan).
Laetulonthus
[2 species; Nearctic Region]
Laetulonthus Moore and Legner, 1973c: 213 (species included: laetulus). Type species: Staphylinus laetulus Say, fixed by original designation.
— Moore and Legner, 1975: 17 (catalog; North America north of Mexico).
— Moore and Legner, 1979: 23 (characters; notes).
— Frank, 1986a: 370 (checklist of species of Florida).
— Smetana, 1995e: 505 (characters; notes; phylogenetic relationships; key to species).
— Downie and Arnett, 1996: 383 (characters).
— Lundgren, 1998: 51 (list of species from Florida; records for other States listed for each species).
— Newton, Thayer, Ashe, and Chandler, 2000: 396 (2 Nearctic species; notes; characters in key).
laetulus Say, 1834: 449 (Staphylinus; [Note: This species was described in a note within a discussion of blandus Gravenhorst]; Type locality: Not cited [Note: See neotype designation by Smetana, 1995e]).
— Melsheimer, 1844: 36 (Philonthus; characters; Pennsylvania).
— LeConte, 1859: 564 (Philonthus).
— Horn, 1884: 184 (Philonthus; characters; Canada to Georgia west to the Mississippi).
— Blatchley, 1910: 381 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 343 (Philonthus; catalog).
— Moore and Legner, 1973c: 215 (Philonthus; characters; notes).
— Legner and Moore, 1977: 176 (Laetulonthus; found under bark).
— Frank, 1984: 478 (Laetulontus; notes; Florida).
— Campbell and Davies, 1991: 119 (Laetulonthus; checklist of species; Canada).
— Smetana, 1995e: 508 (Laetulonthus; neotype designation: [Note: Locality not cited; specimen selected from LeConte collection with “yellow round disc”]; characters; notes; Ontario; Quebec; Alabama; Arkansas; Connecticut; Florida; Georgia; Illinois; Indiana; Kentucky; Louisiana; Maine; Maryland; Massachusetts; Michigan; Mississippi; New Hampshire; New Jersey; New York; North Carolina; Ohio; Oklahoma; Pennsylvania; South Carolina; Texas; Vermont; Virginia; West Virginia; Wisconsin).
— Downie and Arnett, 1996: 383 (Laetulonthus; characters; USA).
▴ Distribution: Canada, USA.
nobilis Bernhauer, 1910b: 381 (Philonthus; Type locality: Neu Mexico: Porvenir).
Leptopeltus
[3 species; Neotropical Region]
Leptopeltus Bernhauer, 1906c: 337 (species included: flavipennis, netolitzkyi, montivagans). Type species: Philonthus flavipennis Erichson, fixed by subsequent designation by R. Lucas, 1920: 371.
— Bernhauer and Schubert, 1914: 365 (world catalog; 3 species).
— Bruch, 1928a: 428 (catalog; Argentina).
— Scheerpeltz, 1933: 1374 (world catalog supplement).
— Blackwelder, 1944: 138 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 217 (type species).
— Scheerpeltz, 1960c: 114 (key to species).
flavipennis Erichson, 1840: 460 (Philonthus; Type locality: Bogotae in Columbia).
— Solsky, 1872: 299 (Philonthus; Colombia).
— Bernhauer, 1906c: 338 (Leptopeltus).
— Bernhauer and Schubert, 1914: 365 (Leptopeltus; catalog).
— Scheerpeltz, 1933: 1374 (Leptopeltus; catalog; Bolivia, Peru, Ecuador, Venezuela).
— Scheerpeltz, 1960c: 115 (Leptopeltus; characters; Venezuela to Chile).
▴ Distribution: Venezuela, Colombia, Ecuador, Peru, Bolivia.
montivagans Bernhauer, 1906c: 339 (Leptopeltus; Type locality: Bolivia: Hochanden, 4000 m).
— Bernhauer and Schubert, 1914: 365 (Leptopeltus; catalog).
— Scheerpeltz, 1960c: 115 (Leptopeltus; characters; Bolivia).
▴ Distribution: Bolivia.
netolitzkyi Bernhauer, 1906c: 338 (Leptopeltus; Type locality: Bolivia: La Paz).
Leucitus
[12 species; Oriental, Australian, and Oceanic Regions]
Leucitus Fauvel, 1878f: 253 (species included: stenoides [with doubt] and argyreus). Type species: Leucitus argyreus Fauvel, fixed by virtual monotypy.
— Duvivier, 1883: 152 (catalog).
— Bernhauer and Schubert, 1914: 372 (world catalog; 8 species).
— Scheerpeltz, 1933: 1380 (world catalog supplement).
— Blackwelder, 1952: 219 (type species: argyreus).
— Last, 1975a: 1 (characters; type species designation).
— Hammond, 1984: 206 (checklist of species of Borneo).
amicus Bernhauer, 1915g: 193 (Leucitus; Type locality: Neu-Guinea: Sekar).
— Scheerpeltz, 1933: 1380 (Leucitus; catalog).
— Last, 1975a: 3 (Leucitus; characters).
▴ Distribution: New Guinea.
argyreus Fauvel, 1878f: 254 (Leucitus; Type locality: Amberbaki, New Guinea; [Note: See lectotype designation by Last, 1975a]. Also cited from Nouvelle-Guinée, Ramoi; Iles Arou, Wokan; Mysol).
— Fauvel, 1879: 96 (Leucitus; characters; notes; Dorei; Vaigheu).
— Heller, 1910: 10 (Leucitus; characters in key).
— Bernhauer and Schubert, 1914: 372 (Leucitus; catalog).
— Arrow, 1915: 500 (Leucitus; Dutch New Guinea).
— Cameron, 1924: 87 (Leucitus; New Guinea).
— Cameron, 1931: 363 (Leucitus; New Guinea).
— Last, 1975a: 3 (Leucitus; lectotype designation: Amberbaki, New Guinea; characters).
— Last, 1987: 55 (Leucitus; Papua New Guinea).
▴ Distribution: New Guinea, Indonesia.
benigseni Bernhauer, 1927, see: Philonthus.
bernhaueri Heller, 1910: 10 (Leucitus; Type locality: Nova Guinea britannica, Astrolabe montes).
— Bernhauer and Schubert, 1914: 373 (Leucitus; catalog).
— Last, 1975a: 4 (Leucitus; characters).
— Last, 1987: 55 (Leucitus; Papua New Guinea; Irian).
▴ Distribution: New Britain, New Guinea.
biroi Last, 1980: 155 (Leucitus; Type locality: New Guinea: Sattelberg, Huon Golf).
freudei Last, 1975a: 5 (Leucitus; Type locality: Papua/Wau, Keinde).
mandibularis Heller, 1910: 9 (Leucitus; Type locality: Papua New Guinea, 780 m).
— Bernhauer and Schubert, 1914: 373 (Leucitus; catalog).
— Last, 1975a: 4 (Leucitus; characters).
▴ Distribution: New Guinea.
paradiseus Fauvel, 1879: 96 (Leucitus; Type locality: Nouva Guinea, Fly River, Mt. Hautukeen; [Note: See lectotype designation by Last, 1975a]. Originally cited from Nouvelle Guinée, Fly River).
— Bernhauer and Schubert, 1914: 373 (Leucitus; catalog).
— Last, 1975a: 4 (Leucitus; lectotype designation: Nouva Guinea, Fly River, Mt. Hautukeen; characters).
▴ Distribution: New Guinea.
pulcher Wendeler, 1924, see: quadripunctatus Bernhauer, 1915.
quadripunctatus Bernhauer, 1915g: 193 (Leucitus; Type locality: Neu-Guinea: Sattelberg).
— Scheerpeltz, 1933: 1380 (Leucitus; catalog).
— Cameron, 1937a: 104 (Leucitus; New Guinea).
— Last, 1975a: 1 (Leucitus).
— Last, 1975a: 3 (Leucitus; characters).
— Last, 1987: 55 (Leucitus; Papua New Guinea; Irian).
▴ Distribution: New Guinea.
pulcher Wendeler, 1924: 340 (Leucitus; Type locality: Neu-Pommern, Süd-See).
— Scheerpeltz, 1933: 1380 (Leucitus; catalog).
— Last, 1975a: 1 (Leucitus; synonym of quadripunctatus).
rufipennis Cameron, 1918, see: Hesperus.
semichalceus Heller, 1910: 9 (Leucitus; Type locality: Papua New Guinea, 780–900 m).
— Bernhauer and Schubert, 1914: 373 (Leucitus; catalog).
— Last, 1975a: 5 (Leucitus; characters).
▴ Distribution: New Guinea.
somoruensis Last, 1968: 360 (Philonthus; Type locality: New Guinea: Mobutie Somoru).
stenoides Gravenhorst, 1806: 90 (Staphylinus; Type locality: Iauae).
— Erichson, 1840: 493, 930 (Philonthus; Java).
— Fauvel, 1878f: 253 (Leucitus; characters; Java).
— Bernhauer and Schubert, 1914: 373 (Leucitus; catalog).
— Last, 1975a: 1 (Leucitus).
— Schillhammer, 1999b: 95 (Leucitus; based on original description, species definitely belongs in Leucitus; type locality in Java probably erroneous; species probably from New Guinea; type specimen probably lost).
▴ Distribution: Indonesia.
superbus Arrow, 1915: 499 (Philonthus; Type locality: Utakwa, River, Dutch New Guinea).
Linoderus
[1 species; Neotropical Region]
Linoderus Sharp, 1885: 452 (species included: gracilipes). Type species: Linoderus gracilipes Sharp, fixed by monotypy.
— Bernhauer and Schubert, 1914: 367 (world catalog; 1 species).
— Blackwelder, 1944: 136 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 221 (type species: gracilipes).
gracilipes Sharp, 1885: 452 (Linoderus; Type locality: Panama, Volcan de Chiriqui, Bugaba).
Mentophilonthus
[8 species; Ethiopian Region]
Mentophilonthus Levasseur, 1966: 207 (species included: odzalaensis, triseriatus, descarpentriesi, mongendensis, dilutior, lampropterus). Type species: Philonthus triseriatus Bernhauer, fixed by original designation.
bangoranensis Levasseur, 1980: 354 (Mentophilonthus; Type locality: République Centrafricaine: Bangoran, SW de Ndélé).
centrafricanus Levasseur, 1980: 356 (Mentophilonthus; Type locality: République Centrafricaine: Bangoran, SW de Ndélé).
descarpentriesi Levasseur, 1966: 208, 209 (Mentophilonthus; Type locality: Congo ex français, Odzala. Also cited from Belgian Congo).
— Cameron, 1929: 61 (Philonthus; [Note: Misidentified as triseriatus according to Levasseur, 1966: 209]; characters; Belgian Congo).
— Tottenham, 1949b: 300 (Philonthus; [Note: Misidentified as triseriatus according to Levasseur, 1966: 209]; characters; Belgian Congo).
— Levasseur, 1966a: 1211 (Mentophilonthus; Belgian Congo; Congo).
▴ Distribution: Congo Republic, Congo.
dilutior Bernhauer, 1928g: 113 (Philonthus; Type locality: Belgisch-Kongo: Mongende).
— Scheerpeltz, 1933: 1340 (Philonthus; catalog).
— Levasseur, 1966: 208, 211 (Mentophilonthus; characters; Belgian Congo).
▴ Distribution: Congo.
seriatipennis Bernhauer, 1928g: 114 (Philonthus; Type locality: Belgisch-Kongo: Mongende).
— Cameron, 1929: 61 (Philonthus; characters; Belgian Congo).
— Scheerpeltz, 1933: 1360 (Philonthus; catalog).
— Tottenham, 1949b: 300 (Philonthus; characters).
— Levasseur, 1966: 211 (Mentophilonthus; synonym of dilutior).
lampropterus Bernhauer, 1934g: 239 (Philonthus; Type locality: Kivu: Kibumba).
— Cameron, 1950g: 35 (Philonthus; Belgian Congo).
— Levasseur, 1966: 213 (Mentophilonthus; characters; Belgian Congo).
▴ Distribution: Congo.
minor Bernhauer, 1928, see: triseriatus Bernhauer, 1928.
mongendensis Bernhauer, 1928g: 114 (Philonthus; Type locality: Belgisch-Kongo: Mongende).
— Scheerpeltz, 1933: 1351 (Philonthus; catalog).
— Levasseur, 1966: 208, 211 (Mentophilonthus; characters; Belgian Congo).
▴ Distribution: Congo.
odzalaensis Levasseur, 1966: 208, 211 (Mentophilonthus; Type locality: Odzala, Congo ex française. Also cited from Gabon).
— Levasseur, 1966a: 1211 (Mentophilonthus; Congo; Zaire; Gabon).
— Levasseur, 1968a: 53 (Mentophilonthus; Central African Republic).
▴ Distribution: Gabon, Congo Republic, Congo, Central African Republic.
parcior Bernhauer, 1928, see: triseriatus Bernhauer, 1928.
seriatipennis Bernhauer, 1928, see: dilutior Bernhauer, 1928.
triseriatus Bernhauer, 1928g: 112 (Philonthus; Type locality: Belgisch-Kongo: Mongende).
— Scheerpeltz, 1933: 1365 (Philonthus; catalog).
— Levasseur, 1966: 208 (Mentophilonthus; characters; Congo; Ruanda).
— Levasseur, 1966a: 1210 (Mentophilonthus; Congo; Zaire).
▴ Distribution: Congo Republic, Congo, Rwanda.
minor Bernhauer, 1928g: 113 (Philonthus; [preoccupied]; Type locality: Belgisch-Kongo: Mongende).
parcior Bernhauer, 1928g: 113 (Philonthus; Type locality: Belgisch-Kongo: Mongende).
Moeocerus
[13 species; Ethiopian Region]
Moeocerus Fauvel, 1899: 100 (replacement name for Homoeocerus Fauvel). Type species: Homoeocerus mimus Fauvel, fixed by subsequent designation Blackwelder, 1952: 250.
— Bernhauer and Schubert, 1916: 410 (world catalog; 7 species).
— Scheerpeltz, 1933: 1420 (world catalog supplement).
— Gridelli, 1939: 322 (characters; key to species).
Homoeocerus Fauvel, 1899a: 27 (species included: mimus, simplex, ciliatus, robustus, spinulosus, puncticollis; preoccupied by Burmeister, 1835, and Kolenati, 1859). Type species: Homoeocerus mimus Fauvel by objective synonym with Moeocerus for which Blackwelder fixed the type species. [Note: Blackwelder stated that R. Lucas, 1920: 421, designated the type species but the designation was ambiguous].
— Fauvel, 1907: 48 (synonym of Moeocerus).
— Bernhauer and Schubert, 1916: 410 (synonym of Moeocerus).
— Blackwelder, 1952: 190 (type species; synonym of Moeocerus).
Polystictomoeocerus Scheerpeltz, 1977: 145 (subgenus of Moeocerus; species included: puncticollis, griseosericans). Type species: Homoeocerus puncticollis Kraatz, fixed by original designation.
abessinus Bernhauer, 1917g: 46 (Moeocerus; Type locality: Abessinien: Harrar).
— Scheerpeltz, 1933: 1420 (Moeocerus; catalog).
— Gridelli, 1939: 326, 329 (Moeocerus; characters; Abyssinia).
— Scheerpeltz, 1977: 146 (Moeocerus; subgenus Moeocerus).
▴ Distribution: Ethiopia.
ciliatus Roth, 1851: 118 (Acylophorus; Type locality: Abyssinien).
— Fauvel, 1899a: 30 (Homoeocerus; characters; Tigre; Abyssinie).
— Bernhauer and Schubert, 1916: 411 (Moeocerus; catalog).
— Gridelli, 1939: 326 (Moeocerus; characters; Abyssinia).
— Scheerpeltz, 1977: 146 (Moeocerus; subgenus Moeocerus).
▴ Distribution: Ethiopia.
doriae Gridelli, 1939: 335 (Moeocerus; Type locality: Cheren, Eritrea; regione dell'Eritrea, Paese dei Bogos).
fauveli Gridelli, 1939: 337 (Moeocerus; Type locality: Maritzburg, Natal).
griseosericans Scheerpeltz, 1977: 143 (Moeocerus; subgenus Polystictomoeocerus; Type locality: Ed. Luja in der Umgebung von Kondué im Kongogebiet).
lineiventris Fauvel, 1907: 48 (Moeocerus; Type locality: Kilimandjaro: Kibosho, 1700 m).
— Bernhauer and Schubert, 1916: 411 (Moeocerus; catalog).
— Scheerpeltz, 1977: 146 (Moeocerus; subgenus Moeocerus).
▴ Distribution: Tanzania.
mimus Fauvel, 1899a: 29 (Homoeocerus; Type locality: Natal).
— Bernhauer and Schubert, 1916: 411 (Moeocerus; catalog).
— Gridelli, 1939: 326, 334 (Moeocerus; characters).
— Scheerpeltz, 1977: 146 (Moeocerus; subgenus Moeocerus).
▴ Distribution: South Africa.
overlaeti Bernhauer, 1935d: 104 (Moeocerus; Type locality: Lulua: Kapanga).
— Gridelli, 1939: 326 (Moeocerus; Abyssinia).
— Scheerpeltz, 1977: 146 (Moeocerus; subgenus Moeocerus).
▴ Distribution: Congo, Ethiopia.
puncticollis Kraatz, 1858: 363 (Homorocerus; Type locality: Senegambia).
— Fauvel, 1899a: 32 (Homoeocerus; Senegambie portugaise).
— Bernhauer and Schubert, 1916: 411 (Moeocerus; catalog).
— Gridelli, 1939: 325 (Moeocerus; Abyssinia).
— Tottenham, 1961: 205 (Moeocerus; Senegal).
— Scheerpeltz, 1977: 145 (Moeocerus; subgenus Polystictomoeocerus).
▴ Distribution: “Senegambia”, Senegal, Ethiopia.
robustus Gestro, 1881: 658 (Homorocerus; Type locality: Let Marefia).
— Fauvel, 1899a: 30 (Homoeocerus; characters; Abyssinie; Choa: Let Marefia; Giuba).
— Bernhauer and Schubert, 1916: 411 (Moeocerus; catalog).
— Gridelli, 1939: 330 (Moeocerus; characters; Abyssinia).
— Scheerpeltz, 1977: 146 (Moeocerus; subgenus Moeocerus).
▴ Distribution: Ethiopia.
simplex Fauvel, 1899a: 30 (Homoeocerus; Type locality: Angola).
— Bernhauer, 1915o: 363 (Moeocerus; Belgian Congo).
— Bernhauer and Schubert, 1916: 411 (Moeocerus; catalog).
— Scheerpeltz, 1977: 147 (Moeocerus; subgenus Moeocerus).
▴ Distribution: Angola, Congo.
spinulosus Solsky, 1864a: 375 (Homorocerus; Type locality: Insel Gorea an der Westküste von Afrika).
— Fauvel, 1899a: 31 (Homoeocerus; characters; Natal; Abyssinia).
— Bernhauer and Schubert, 1916: 411 (Moeocerus; catalog).
— Bernhauer and Chapman, 1939: 73 (Moeocerus; Kenya).
— Scheerpeltz, 1977: 147 (Moeocerus; subgenus Moeocerus).
▴ Distribution: Senegal, Ethiopia, Kenya, South Africa.
zavattarii Gridelli, 1939: 332 (Moeocerus; Type locality: Moyale; Arero).
Mysolius
[3 species; Oriental and Australian Regions]
Mysolius Fauvel, 1878f: 255 (species included: aurichalceus). Type species: Mysolius aurichlaceus Fauvel, fixed by monotypy.
— Duvivier, 1883: 152 (catalog).
— Olliff, 1887: 497 (characters; Australian species).
— Bernhauer and Schubert, 1914: 373 (world catalog; 2 species).
— Blackwelder, 1952: 256 (type species: aurichalceus).
aurichalceus Fauvel, 1878f: 256 (Mysolius; Type locality: Nouvelle-Guinée, Romoi; Iles Arou, Wokan; Mysol; Salvatti).
— Fauvel, 1879: 97 (Mysolius; characters; Fly River).
— Bernhauer and Schubert, 1914: 373 (Mysolius; catalog).
— Cameron, 1924: 87 (Philonthus; New Guinea).
— Cameron, 1931: 363 (Mysolius; New Guinea).
— Last, 1987: 55 (Mysolius; Irian).
▴ Distribution: New Guinea, Indonesia.
diversipes Fauvel, 1879: 98 (Mysolius; race of aurichalceus; Type locality: Iles Arou; Nouvelle Guinée; Fly River).
chalcopterus Olliff, 1887: 497 (Mysolius; Type locality: Mulgrave River, North Queensland).
— Bernhauer and Schubert, 1914: 373 (Mysolius; catalog).
— Bernhauer, 1920: 16 (Mysolius; Australia).
— Last, 1987: 55 (Mysolius; Papua New Guinea).
▴ Distribution: Australia, New Guinea.
cupreosplendens Cameron, 1943a: 344 (Actinus; Type locality: New Guinea: Milne Bay).
diversipes Fauvel, 1879, see: aurichalceus Fauvel, 1878.
Neobisnius
[72 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, and Australian Regions]
Neobisnius Ganglbauer, 1895: 464 (species included: villosulus; procerulus; prolixus). Type species: Gabrius villosulus Stephens, fixed by subsequent designation by R. Lucas, 1920: 434.
— Thomson, 1859: 25 (cited as Bisnius; [Note: Thomson's use of Bisnius was a misidentification of Neobisnius; Tottenham, 1949 cited it with a type species and as a synonym of Neobisnius as though it were an available name]; characters; type species: cephalotes).
— Thomson, 1860: 168 (cited as Bisnius; [Note: Thomson's use of Bisnius was a misidentification of Neobisnius; Tottenham, 1949 cited it with a type species and as a synonym of Neobisnius as though it were an available name]; characters).
— Heyden, 1880: 73 (cited as Actobius; list of species of Siberian region).
— Heyden, Reitter, and Weise, 1906: 163 (list of species of Europe and Caucasus).
— Reitter, 1909: 116, 123 (characters; key to species).
— Petri, 1912: 71 (list of species of Siebenbürgen).
— Bernhauer and Schubert, 1914: 322 (world catalog; 39 species).
— Leng, 1920: 106 (catalog of North American species).
— Winkler, 1925: 374 (catalog for Palaearctic region).
— Cameron, 1925: 54 (catalog of Indian species).
— Porta, 1926: 99 (characters; key to species of Italy).
— Bruch, 1928a: 427 (catalog; Argentina).
— Portevin, 1929: 356 (key to French species).
— Roubal, 1930: 366 (catalog; Slovakia).
— Cameron, 1932: 56 (characters; key to species of British India).
— Joy, 1932: 126 (key to British species).
— Scheerpeltz, 1933: 1326 (world catalog supplement).
— Normand, 1935: 370 (list of species of Tunisia).
— Blackwelder, 1943: 440 (synonym of Erichsonius; type species: cinerascens).
— Gridelli, 1943: 111 (characters; notes; key to a few species).
— Kloet and Hincks, 1945: 165 (list of British species).
— Last, 1948: 148 (key to European species).
— Tottenham, 1949: 371, 417 (type species: villosulus Stephens = elongatulus Erichson; checklist of British species).
— V. Hansen, 1952: 29 (characters; key to Danish species).
— Blackwelder, 1952: 258 (type species).
— Hatch, 1957: 173, 174 (characters; key to species of Pacific Northwest).
— Kocher, 1958: 142 (checklist of species; Morocco).
— Smetana, 1958: 134 (characters; habitus of one species; type species: villosulus; key to Czechoslovakian species).
— Smetana, 1959a: 209 (list of species collected in Albania).
— Ferreira, 1962a: 17 (list of Portuguese species).
— Coiffait, 1963: 8, 18 (characters; key to French species groups and included French species).
— Székessy, 1963: 53 (characters).
— Palm, 1963b: 58 (characters; key to species of Sweden).
— Lohse, 1964: 171 (characters; key to central European species).
— Moore, 1965: 98, 100 (characters; notes).
— Coiffait and Saiz, 1968: 348, 360 (characters; notes).
— Scheerpeltz, 1968a: 64 (catalog of Austrian species).
— Shibata, 1973c: 56 (checklist of species of Taiwan).
— Tikhomirova, 1973a: 182 (checklist of species of USSR).
— Coiffait, 1974: 322 (characters; key to species of western Palaearctic region).
— Moore and Legner, 1975: 18 (catalog; North America north of Mexico).
— Pope, 1977: 29 (list of British species).
— Topp, 1978: 312 (larval characters).
— Moore and Legner, 1979: 24 (characters; notes).
— Muona, 1979: 14 (list of Scandinavian species).
— Uhlig, Vogel, and Sieber, 1980: 246 (some species collected in Germany).
— Burakowski, Mroczkowski, and Stefańska, 1980: 29 (catalog; Poland).
— Frank, 1981: 2 (revision of and key to New World species; characters; checklist of New World species; species group classification).
— Uhlig and Vogler, 1981: 110 (list of some species collected in Germany; habitat and collecting notes).
— Shibata, 1983: 88 (checklist of species of Japan).
— Hammond, 1984: 206 (checklist of species of Borneo).
— Tóth, 1984: 34 (characters; habitus; key to species of Hungary).
— Frank, 1986a: 370 (checklist of species of Florida).
— Segers, 1986: 46 (checklist of species; Belgium).
— Drugmand 1987b (comments on distribution of Belgian species).
— Campbell and Davies, 1991: 117 (checklist of species; Canada).
— Silfverberg, 1992: 17 (list of species of Fennoscandia, Denmark, and the Baltic States).
— Smetana, 1993b: 47 (list of species of Czech Republic and Slovakia).
— Hernández, Outerelo, and Gamarra, 1994: 188 (checklist of species; Canary Islands).
— Smetana, 1995e: 50 (characters).
— Ciceroni and Zanetti, 1995: 27 (list of species of Italy).
— M. Hansen, 1996: 104 (list of Danish species).
— M. Hansen, Mahler, Palm, and Pedersen, 1996: 243 (additions to list of Danish species).
— Cho, 1996a: 159 (characters; key to species of Korea).
— Downie and Arnett, 1996: 374 (characters; key to species of Northeastern North America).
— Lundgren, 1998: 51 (list of species from Florida; records for other States listed for each species).
— Bain, 1998: 39 (17th century remains; Massachusetts).
— M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (list of species collected in 1997; Denmark).
— Newton, Thayer, Ashe, and Chandler, 2000: 396 (14 Nearctic species; notes; characters in key).
— Hayashi, 2000: 23 (notes on empodial setae).
adustus Sharp, 1887, see: terminalis LeConte, 1863.
afghanicus Coiffait, 1979d: 559 (Neobisnius; Type locality: entre Camdech et Kouchtous, 1400 à 2000 m).
— Tronquet, 1981: 71 (Neobisnius; Afghanistan).
— Coiffait, 1982b: 76 (Neobisnius; note).
▴ Distribution: Afghanistan.
agnatus Erichson, 1840, see: sobrinus Erichson, 1840.
alaiensis Bernhauer, 1914b: 66 (Neobisnius; Type locality: Nord-Alai: Fergana; [Note: See lectotype designation by Gusarov, 1995a]).
— Scheerpeltz, 1933: 1326 (Neobisnius; catalog).
— Coiffait, 1974: 329 (Neobisnius; characters; Transcaspian region; Afghanistan).
— Gusarov, 1995a: 92 [= 1996: 55] (Neobisnius; lectotype designation).
▴ Distribution: Uzbekistan, Afghanistan.
alternans Sharp, 1885, see: jocosus Horn, 1884.
angusticollis Kolenati, 1846, see: prolixus Erichson, 1840.
angustus Cho, 1996a: 162 (Neobisnius; Type locality: Korea: Chejudo, Kyoraeri).
arduus Sharp, 1885, see: sobrinus Erichson, 1840.
armuellensis Bierig, 1933: 56 (Neobisnius; Type locality: Panama: Puerti Armuelles).
— Frank, 1981: 28 (Neobisnius; characters; Costa Rica; Panama; Colombia).
▴ Distribution: Colombia, Panama, Costa Rica.
bakeri Bernhauer, 1916h: 424 (Neobisnius; Type locality: Luzon: Mt. Makiling).
bicolor Drugmand, 1990, see: Philonthus.
boops Fauvel, 1905b: 177 (Actobius; Type locality: forêt Tanala).
brasilianus Wendeler, 1956a: 230 (Neobisnius; Type locality: Brasilien: Nova Teutonia).
cameroni Wendeler, 1928b: 298 (Neobisnius; replacement name for fraternus Cameron).
— Cameron, 1932: 58 (Neobisnius; characters; India).
— Scheerpeltz, 1933: 1326 (Neobisnius; catalog).
▴ Distribution: India.
fraternus Cameron, 1926b: 349 (Neobisnius; [preoccupied]; Type locality: Dehra Dun).
— Wendeler, 1928b: 298 (Neobisnius; synonym of cameroni).
— Cameron, 1932: 58 (Neobisnius; synonym of cameroni).
— Scheerpeltz, 1933: 1326 (Neobisnius; synonym of cameroni).
carbonarius Bierig, 1933, see: nigrocoeruleus Cameron, 1922.
cavifrons Bierig, 1933, see: simplex Sharp, 1885.
cerrutii Gridelli, 1943, see: lathrobioides Baudi, 1848.
chengkouensis Zheng, 1994: 213, 214 (Neobisnius; Type locality: China: Sichuan: Chengkou).
cinctus Latreille, 1804, see: villosulus Stephens, 1833.
cinerascens Stephens, 1833, see: procerulus Gravenhorst, 1806.
concolor Sharp, 1885: 458 (Actobius; Type locality: Panama, Volcan de Chiriqui 4000 to 6000 feet, Peña Blanca 3000 to 4000 feet).
— Bernhauer and Schubert, 1914: 322 (Neobisnius; catalog).
— Frank, 1981: 38 (Neobisnius; characters; Panama).
▴ Distribution: Panama.
cooteri Last, 1987: 50 (Neobisnius; Type locality: Papua New Guinea: 20 km SE of Port Moresby).
decolor Coiffait, 1974, see: procerulus Gravenhorst, 1806.
deletus Sharp, 1876, see: ludicrus Erichson, 1840.
delicatulus Sharp, 1885, see: terminalis LeConte, 1863.
demmeli Bierig, 1933: 54 (Neobisnius; Type locality: Cuba, Prov. P. del Rio, Sierra del Rosario, cercanía de Cayajabos).
— Blackwelder, 1943: 441 (Erichsonius; synonym of humilis).
— Blackwelder, 1944: 131 (Erichsonius; synonym of humilis).
— Frank, 1981: 32 (Neobisnius; valid species; characters; Cuba).
▴ Distribution: West Indies.
dissimilis Baudi, 1870, see: prolixus Erichson, 1840.
edznai Frank, 1981: 44 (Neobisnius; Type locality: Mexico: Campeche, Edzná, marsh, 150 m. Also cited from Guatemala).
elegantulus Horn, 1884, see: terminalis LeConte, 1863.
elongatulus Erichson, 1839, see: villosulus Stephens, 1833.
facilis Sharp, 1885: 458 (Actobius; Type locality: Guatemala, San Geronimo).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Scheerpeltz, 1971b: 205 (Neobisnius; Guatemala; El Salvador; Nicaragua).
— Frank, 1981: 37 (Neobisnius; characters; Guatemala).
▴ Distribution: Guatemala, El Salvador, Nicaragua.
fatalis Tottenham, 1956: 259 (Neobisnius; Type locality: Urundi: Kitega, 1600–1700 m).
fauveli Smetana, 1963, see: ludicrus Erichson, 1840.
filiformis Wollaston, 1854, see: procerulus Gravenhorst, 1806.
filius Last, 1981, see: Philonthus.
flavicornis Fauvel, 1895: 261 (Actobius; Type locality: Bírmanie, Carin Asciuii Ghécu, 1400–1500 m).
— Fauvel, 1905e: 83 (Actobius; Java).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Cameron, 1932: 61 (Actobius; characters; Burma).
— Scheerpeltz, 1933: 1329 (Neobisnius; catalog).
▴ Distribution: Myanmar, Indonesia.
flavipes Bierig, 1933: 50 (Neobisnius; Type locality: Panama: Puerto Armuelles).
flavomaculatus Bernhauer, 1908d: 336 (Neobisnius; Type locality: Brasilien: S. Catharina; Grenada, W. J[I?]).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Frank, 1981: 15 (Neobisnius; characters; Uruguay; Paraguay; Brazil; Bolivia; Colombia; Grenada).
▴ Distribution: Uruguay, Paraguay, Brazil, Colombia, West Indies.
formosae Cameron, 1949: 175 (Neobisnius; Type locality: Formosa: Takesaki: near Mt. Ari).
formosus Fall, 1901, see: infimus Horn, 1884.
fortis Sharp, 1885: 457 (Actobius; Type locality: Panama, Volcan de Chiriqui 4000 to 6000 feet, Bugaba 800 to 1500 feet).
— Bernhauer, 1910b: 383 (Neobisnius).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Scheerpeltz, 1933: 1327 (Neobisnius; catalog; Argentina).
— Frank, 1981: 36 (Neobisnius; characters; Panama; Colombia).
▴ Distribution: Panama, Colombia, Argentina.
fraternus Bernhauer, 1921c: 105 (Neobisnius; Type locality: Argentinien: Prov. Jujuy).
— Scheerpeltz, 1933: 1327 (Neobisnius; catalog).
— Frank, 1981: 29 (Neobisnius; characters; Argentina).
▴ Distribution: Argentina.
fraternus Cameron, 1926, see: cameroni Wendeler, 1928.
fulvicornis Notman, 1920, see: villosulus Stephens, 1833.
funerulus Cameron, 1922: 119 (Neobisnius; Type locality: St. Vincent).
— Scheerpeltz, 1933: 1327 (Neobisnius; catalog).
— Blackwelder, 1943: 441 (Erichsonius; synonym of humilis).
— Blackwelder, 1944: 131 (Erichsonius; synonym of humilis).
— Frank, 1981: 34 (Neobisnius; valid species; characters; St. Lucia; St. Vincent).
▴ Distribution: West Indies.
gratus LeConte, 1863a: 38 (Philonthus; Type locality: Colorado River, near Fort Yuma, California; [Note: See lectotype designation by Frank, 1981]).
— Fauvel, 1874: 201 [= 1874a: 427] (Erichsonius).
— Horn, 1884: 231 (Actobius; characters; south of California; Arizona).
— Fall and Cockerell, 1907: 166 (Actobius; New Mexico).
— Bernhauer, 1912e: 176 (Neobisnius).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Scheerpeltz, 1933: 1327 (Neobisnius; catalog; Argentina).
— Frank, 1981: 16 (Neobisnius; lectotype designation; characters; Utah; Texas; New Mexico; Arizona; California; Sinaloa).
▴ Distribution: USA, Mexico, Argentina.
hauseri Bernhauer, 1941, see: prolixus Erichson, 1840.
humilis Erichson, 1840: 512 (Philonthus; Type locality: Puerto Rico; [Note: See lectotype designation by Smetana, 1963]).
— Chevrolat and Fauvel, 1863: 435 (Philonthus; characters; Cuba).
— Fauvel, 1874: 201 [= 1874a: 427] (Erichsonius).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Blackwelder, 1943: 441 (Erichsonius; characters; Cuba; Jamaica; Haiti; Dominican Republic; Puerto Rico; Virgin Islands; St. Croix; St. Lucia; Barbados; St. Vincent; Grenada).
— Smetana, 1963: 4 (Neobisnius; lectotype designation).
— Frank, 1981: 18 (Neobisnius; characters; Jamaica; Cuba; Haiti; Dominican Republic; U.S Virgin Islands).
▴ Distribution: West Indies.
margipallens Jacquelin du Val, 1857a: 41 (Lathrobium; [Note: This name is currently cited both in Lobrathium, where it is a valid species, and here in Neobisnius]; Type locality: Not cited, presumably from Cuba).
— Chevrolat and Fauvel, 1863: 435 (Philonthus; synonym of humilis).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; synonym of humilis).
— Blackwelder, 1943: 441 (Erichsonius; synonym of humilis).
— Blackwelder, 1944: 131 (Erichsonius; synonym of humilis).
— Frank, 1981: 18 (Neobisnius; synonym of humilis).
limbatus Cameron, 1922: 118 (Neobisnius; Type locality: Haiti).
— Scheerpeltz, 1933: 1327 (Neobisnius; catalog).
— Blackwelder, 1943: 441 (Erichsonius; synonym of humilis).
— Blackwelder, 1944: 131 (Erichsonius; synonym of humilis).
— Frank, 1981: 18 (Neobisnius; synonym of humilis).
imitator Scheerpeltz, 1974: 136 (Neobisnius; Type locality: S. W. Afrika: Kaokoveld, Otju, 45 miles WSW Ohopoho).
infimus Horn, 1884: 227 (Actobius; Type locality: San Jose, California; [Note: See lectotype designation by Frank, 1981]).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
— Frank, 1981: 52 (Neobisnius; lectotype designation; characters; California; Arizona).
▴ Distribution: USA.
formosus Fall, 1901: 221 (Actobius; Type locality: Pasadena; Pomona).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Frank, 1981: 52 (Neobisnius; synonym of infimus).
inornatus Sharp, 1889: 43 (Actobius; Type locality: Yokohama).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
— Cameron, 1933a: 170 (Neobisnius).
— Watanabe and Shibata, 1972: 63 (Neobisnius; Yaku-shima Island).
— J. Li, 1993: 33 (Neobisnius, cited as Noebinius; China).
— Cho, 1996a: 159 (Neobisnius; characters; Korea).
▴ Distribution: Japan, Korea, China.
iranicus Scheerpeltz, 1961, see: Philonthus.
jocosus Horn, 1884: 232 (Actobius; Type locality: Delaware; [Note: See lectotype designation by Frank, 1981]. Also cited from North Carolina).
— Blatchley, 1910: 394 (Actobius; characters; Indiana).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Frank, 1981: 27 (Neobisnius; lectotype designation: Delaware; characters; Massachusetts; New York; New Jersey; Maryland; Delaware; Virginia; Florida; Indiana; Kansas; Louisiana; Texas; Nuevo Leon; Veracruz; Tabasco; Chiapas; Guatemala; El Salvador; Honduras).
— Downie and Arnett, 1996: 375 (Neobisnius; characters; USA).
▴ Distribution: USA, Mexico, Guatemala, Honduras, El Salvador.
alternans Sharp, 1885: 461 (Actobius; Type locality: Paso Antonio, Escuintla, Guatemala; [Note: See lectotype designation by Frank, 1981]. Originally cited from Guatemala, San Geronimo, Torola, Paso Antonio).
— Bernhauer and Schubert, 1914: 322 (Neobisnius; catalog).
— Frank, 1981: 27 (Neobisnius; lectotype designation: Paso Antonio, Escuintla, Guatemala; synonym of jocosus).
jucundus Horn, 1884: 232 (Actobius; Type locality: South Carolina; [Note: See lectotype designation by Frank, 1981]).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Frank, 1981: 51 (Neobisnius; lectotype designation; characters; Ontario; Alberta; British Columbia; New Hampshire; Connecticut; Massachusetts; Rhode Island; New York; Pennsylvania; Michigan; Illinois; Iowa; Minnesota; Kansas; Nebraska; Colorado; New Jersey; Maryland; Virginia; South Carolina).
— Downie and Arnett, 1996: 375 (Neobisnius; characters; USA).
— Schmidt, 1999: 108 (Neobisnius; food for adults and larvae used for rearing species; rearing methods).
▴ Distribution: Canada, USA.
kaletiensis Last, 1964: 178 (Neobisnius; Type locality: Sarawak: Kaleti).
lathrobioides Baudi, 1848: 130 (Philonthus; Type locality: Pedemontio; [Note: See lectotype designation by Bordoni, 1975g]).
— Kraatz, 1857d: 621 (Philonthus).
— Fauvel, 1867c: 113 [= 1868b: 212] (Philonthus; synonym of procerulus).
— Ganglbauer, 1895: 465 (Neobisnius; synonym of procerulus).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; synonym of procerulus).
— Smetana, 1958: 137 (Neobisnius; synonym of procerulus).
— Bordoni 1975g: 82 (Neobisnius; lectotype designation; valid species; characters).
— Frank, 1981: 9 (Neobisnius; characters; Massachusetts; New York; Illinois; Nevada; California).
— Lohse and Lucht, 1989: 160 (Neobisnius; notes).
— Borges, 1990: Table II (Neobisnius; Azores).
— Maachi, 1991: 250 (Neobisnius; Morocco).
— Smetana, 1993b: 47 (Neobisnius; Czech Republic; Slovakia).
— Hodge and Jones, 1995: 38 (Neobisnius; characters; Britain).
— Ciceroni and Zanetti, 1995: 27 (Neobisnius; Italy).
— Downie and Arnett, 1996: 375 (Neobisnius; characters; USA).
▴ Distribution: Egypt, Tunisia, Algeria, Europe, Caucasus, Turkey, Canada, USA.
cerrutii Gridelli, 1943: 121 (Neobisnius; Type locality: Mekfi, in Basilicata).
— Last, 1948: 148 (Neobisnius; characters; Britain).
— V. Hansen, 1952: 30 (Neobisnius; characters; Denmark).
— Fagel, 1954 (Neobisnius; characters; separated from semipunctatus; Italy; Belgium; France; Britain; Germany; Austria; Algeria; Tunisia; Egypt; Caucasus; Canada: Quebec; USA: California; Nevada).
— Horion, 1954: 9 (Neobisnius; characters; Italy; England; Norway; Sweden; Denmark; Germany).
— Smetana, 1958: 138 (Neobisnius; characters; ecology; Czechoslovakia).
— Smetana, 1960b: 259 (Neobisnius; Czechoslovakia).
— Scheerpeltz, 1961d: 127 (Erichsonius; Italy).
— Palm, 1963b: 59 (Neobisnius; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 172 (Neobisnius; characters; central Europe).
— Scheerpeltz, 1964a: 299 (Neobisnius; Crete).
— Focarile, 1964: 57 (Neobisnius; Italy).
— Scheerpeltz, 1965b: 401 (Neobisnius; Sicily).
— Scheerpeltz, 1965c: 505 (Neobisnius; Sicily).
— Horion, 1965: 116 (Neobisnius; Italy; Spain; Holland; Belgium; France; Czech Republic; Slovakia; Britain; Denmark; Norway; Sweden; Finland; Germany; Austria).
— Smetana, 1967i: 551 (Neobisnius; Turkey).
— Szujecki, 1968a: 727 (Neobisnius; Poland).
— Coiffait, 1974: 327 (Neobisnius; characters; Europe; Portugal; Asia Minor).
— Bordoni 1975g: 82 (Neobisnius; synonym of lathrobioides).
— Pope, 1977: 29 (Neobisnius; Britain).
— Muona, 1979: 14 (Neobisnius; synonym of lathrobioides).
— Frank, 1981: 9 (Neobisnius; synonym of lathrobioides).
— Tóth, 1984: 37 (Neobisnius; [Note: I assume Tóth overlooked the synonymy of cerrutii]; characters; Hungary).
— Lucht, 1987: 104 (Neobisnius; checklist; Germany; Poland; Czechoslovakia; Austria; Benelux; Denmark; Sweden).
— Lohse and Lucht, 1989: 160 (Neobisnius; synonym of lathrobioides).
— Smetana, 1993b: 47 (Neobisnius; synonym of lathrobioides).
— Ciceroni and Zanetti, 1995: 27 (Neobisnius; synonym of lathrobioides).
rubripennis Gridelli, 1943: 122 (Neobisnius; variety of cerrutii; Type locality: presso Venezia, nei terreni di bonifica di Punta Sabbioni).
— Last, 1948: 148 (Neobisnius; variety of cerrutii; characters; Britain).
— Smetana, 1958: 138 (Neobisnius; aberration of cerrutii).
— Coiffait, 1974: 327 (Neobisnius; synonym of cerrutii).
— Pope, 1977: 29 (Neobisnius; synonym of cerrutii).
lepidulus LeConte, 1863a: 37 (Philonthus; [Note: Preoccupied as secondary homonym in Philonthus]; Type locality: Georgia, near the mountains).
— Horn, 1884: 233 (Actobius; characters; Georgia).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Frank, 1981: 23 (Neobisnius; characters; Georgia).
▴ Distribution: USA.
limbatus Cameron, 1922, see: humilis Erichson.
loebli Last, 1989: 881 (Neobisnius; Type locality: Papua New Guinea: Kokina).
longulus Kraatz, 1859, see: praelongus Gemminger and Harold, 1868.
ludicrus Erichson, 1840: 514 (Philonthus; Type locality: Guyana Venezuelensis; [Note: See lectotype designation by Frank, 1981]).
— Fauvel, 1874: 201 [= 1874a: 427] (Erichsonius).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Scheerpeltz, 1933: 1327 (Neobisnius; catalog; Antilles).
— Frank, 1981: 20 (Neobisnius; lectotype designation; characters; South Carolina; Georgia; Florida; Alabama; Louisiana; Texas; Nayarit; Guerrero; Veracruz; Oaxaca; Tabasco; Guatemala; Honduras; Costa Rica; Panama; Colombia; Bolivia; Venezuela; Trinidad and Tobago; St. Lucia; Barbados; St. Vincent; Grenada; Brazil).
▴ Distribution: Bolivia, Venezuela, Colombia, Panama, Costa Rica, Honduras, Guatemala, Mexico, West Indies, USA.
umbripennis LeConte, 1863a: 38 (Philonthus; Type locality: Middle States).
— Fauvel, 1874: 201 [= 1874a: 427] (Erichsonius).
— Horn, 1884: 233 (Actobius; characters; Pennsylvania to Florida).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; catalog).
— Notman, 1920b: 705 (Neobisnius; Florida).
— Frank, 1981: 20 (Neobisnius; synonym of ludicrus).
deletus Sharp, 1876: 167 (Philonthus; Type locality: Tapajos).
— Bernhauer and Schubert, 1914: 335 (Philonthus; catalog).
— Frank, 1981: 20 (Neobisnius; synonym of ludicrus).
fauveli Smetana, 1963: 2 (Neobisnius; Type locality: Louisiane).
maculicollis Scheerpeltz, 1972: 107 (Neobisnius; Type locality: Argentinien: Buenos Aires, Ezeiza).
madangensis Last, 1981, see: Philonthus.
margianus Bordoni, 1975, see: procerulus Gravenhorst, 1806.
margipallens Jacquelin du Val, 1857, see: humilis Erichson, 1840.
maximus Bernhauer, 1910b: 383 (Neobisnius; Type locality: Mexiko: Vera Cruz, Cordoba).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Frank, 1981: 37 (Neobisnius; characters; Veracruz).
▴ Distribution: Mexico.
mediopolitus Lea, 1929: 203 (Neobisnius; Type locality: Victoria: Melbourne).
mixtus Sharp, 1885: 460 (Actobius; Type locality: San Jerónimo, Baja Verapaz, Guatemala; [Note: See lectotype designation by Frank, 1981]. Originally cited from Mexico, Cordova, Vera Cruz, Tehuacan; Guatemala, San Geronimo).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Bierig, 1933: 56 (Neobisnius; characters; Panama).
— Frank, 1981: 24 (Neobisnius; lectotype designation: San Jerónimo, Baja Verapaz, Guatemala; characters; Puebla; Veracruz; San Luis Potosi; Guatemala).
▴ Distribution: Mexico, Guatemala, Panama.
moestus Sharp, 1887: 790 (Actobius; Type locality: Mexico, near the city).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Frank, 1981: 45 (Neobisnius; characters; Mexico).
▴ Distribution: Mexico.
mundus Last, 1981, see: Philonthus.
musonoiensis Levasseur, 1969: 881 (Neobisnius; Type locality: Katanga: Musonoie, région de Kolwezi).
nigripes Bernhauer, 1941a: 227 (Neobisnius; Type locality: Fukien: Kwangtseh; [Note: See lectotype designation by Gusarov, 1995a]).
— Coiffait, 1974: 331 (Neobisnius; characters; China).
— Gusarov, 1995a: 92 [= 1996: 55] (Neobisnius; lectotype designation; characters).
▴ Distribution: China.
nigritulus Cameron, 1922: [Note: Nomen nudum, Articles 12.1, 13.1.1; Scheerpeltz, 1933: 1327, and Blackwelder, 1944: 131, listed this species as described by Cameron, 1922: 119; I cannot find the species there or elsewhere.
nigrocoeruleus Cameron, 1922: 118 (Neobisnius; Type locality: Haiti).
— Scheerpeltz, 1933: 1327 (Neobisnius; cited as nigrocaeruleus; catalog).
— Blackwelder, 1943: 441 (Erichsonius; synonym of humilis).
— Blackwelder, 1944: 131 (Erichsonius; synonym of humilis).
— Frank, 1981: 26 (Neobisnius; valid species; characters; Jamaica; Cuba; Haiti; Dominican Republic).
▴ Distribution: West Indies.
carbonarius Bierig, 1933: 49 (Neobisnius; Type locality: Cuba: Prov. P. del Rio, Sierra del Rosario, Cayajabos).
— Blackwelder, 1943: 441 (Erichsonius; synonym of humilis).
— Blackwelder, 1944: 131 (Erichsonius; synonym of humilis).
— Frank, 1981: 26 (Neobisnius; synonym of nigrocoeruleus).
nitidulus Sharp, 1885: 460 (Actobius; Type locality: Guatemala City, Department of Guatemala, Guatemala; [Note: See lectotype designation by Frank, 1981]. Originally cited from Guatemala, near the city, San Geronimo).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Frank, 1981: 34 (Neobisnius; lectotype designation: Guatemala City, Department of Guatemala, Guatemala; characters; Veracruz; Guatemala; El Salvador).
▴ Distribution: El Salvador, Guatemala, Mexico.
politus Sharp, 1885: 461 (Actobius; Type locality: Mexico).
— Sharp, 1887: 790 (Actobius; Guatemala).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Frank, 1981: 34 (Neobisnius; synonym of nitidulus).
nothocreatus Frank, 1981: 11 (Neobisnius; Type locality: California: Calaveras Co.: Mokelumne Hill).
nouristanicus Coiffait, 1979, see: orbus Kiesenwetter, 1850.
novus Last, 1981: 126 (Neobisnius; Type locality: Goroka, Kami River).
occidentoides Frank, 1981: 47 (Neobisnius; Type locality: USA: Texas, San Patricio Co., Lake Corpus Christi St. Pk. Also cited from Manitoba; Alberta; Vermont; New York; Ohio; Virginia; North Carolina; Kentucky; Tennessee; Missouri; Alabama; Mississippi; Illinois; Minnesota; Kansas; Oklahoma; Arkansas; Louisiana; North Dakota; Montana; Idaho; Washington; South Dakota; Nebraska; Wyoming; Colorado; Utah; Nevada; Texas; New Mexico; Arizona; California; Mexico).
— Downie and Arnett, 1996: 375 (Neobisnius; characters; USA).
— Schmidt, 1999: 108 (Neobisnius; food for adults and larvae used for rearing species; rearing methods).
▴ Distribution: Canada, USA, Mexico.
ocreatus Horn, 1884, see: sobrinus Erichson, 1840.
oculatus Fauvel, 1905b: 178 (Actobius; Type locality: Diego-Suarez; forêts au nord de Fort Dauphin. Afrique orientale anglaise: Baie de Kavirondo, Victoria Nyanza. Congo: Chiloango; Boma. Gabon: Loango).
— Fauvel, 1907: 37 (Actobius; Kavirondo Bay, Victoria-Nyanza; Congo; Gabon; Madagascar).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Bernhauer and Chapman, 1939: 64 (Neobisnius; Kenya).
— Scheerpeltz, 1974: 134 (Neobisnius; South Africa; South West Africa).
— Lecoq, 1990: 203 (Neobisnius; Mascarene Islands).
▴ Distribution: Mascarene Islands, Madagascar, Kenya, Congo, Gabon, Namibia, South Africa.
omnirufus Frank, 1981: 31 (Neobisnius; Type locality: Argentina: Mendoza).
orbus Kiesenwetter, 1850: 220 (Philonthus; Type locality: Perpiniacum, Gerona).
— Kiesenwetter, 1851a: 418 (Philonthus; characters; France; Spain).
— Fairmaire and Laboulbène, 1856: 531 (Philonthus; characters; France).
— Kraatz, 1857d: 621 (Philonthus).
— Baudi, 1870: 387 (Philonthus; characters).
— Fauvel, 1874: 204 [= 1874a: 430] (Erichsonius; characters; France; Italy; Egypt; Madeira; Canary Islands).
— Fauvel, 1876a: 72 [= 1876: 257] (Actobius; Spain; Corsica; Syria).
— Mulsant and Rey, 1876: 592 [= 1877: 448] (Bisnius; characters; France).
— Fauvel, 1878: 121 [= 1878a: 41] (Actobius; Algeria; Egypt).
— Fauvel, 1886: 52 [= 1886a: 44] (Actobius; Morocco; Egypt; Algeria).
— Fauvel, 1902b: 107 (Actobius; Morocco; Algeria; Tunisia; Egypt; Canaries).
— Sainte-Claire Deville, 1906: 94 (Neobisnius; Corsica).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Portevin, 1929: 356 (Neobisnius; characters; France).
— Peyerimhoff, 1931: 31 (Neobisnius; Algeria).
— Scheerpeltz, 1933: 1327 (Neobisnius; catalog).
— Gridelli, 1943: 125 (Neobisnius; characters; Egypt; Morocco; Italy).
— Last, 1948: 149 (Neobisnius; characters in key and aedeagal illustration).
— Cameron, 1952d: 824 (Neobisnius; Mauritania).
— Jarrige, 1958: 92 (Neobisnius; Algeria).
— Coiffait, 1966b: 338 (Neobisnius; Mauritania; Hoggar).
— Smetana, 1967i: 551 (Neobisnius; Turkey).
— Coiffait, 1968a: 136 (Neobisnius; Chad).
— Coiffait, 1974: 328 (Neobisnius; characters; Mediterranean region; Spain; North Africa; Italy; Anatolia; Egypt; Mauritania; Canary Islands; Pyrenees; Corsica).
— Maachi, 1991: 250 (Neobisnius; Morocco).
— Ciceroni and Zanetti, 1995: 27 (Neobisnius; Italy).
— Gusarov, 1997: 287 (Neobisnius; Saudi Arabia).
— Assing and Wunderle, 2001: 36 (Neobisnius; Cyprus).
▴ Distribution: Afghanistan, Syria, Turkey, Italy, France, Spain, Madeira, Canary Islands, Cape Verde Islands, Mauritania, Morocco, Algeria, Tunisia, Egypt, Chad, Saudi Arabia, Madagascar, Réunion.
tenellus Wollaston, 1864: 576 (Philonthus; Type locality: Teneriffe: Barranco Santo near Santa Cruz).
— Wollaston, 1865: 495 (Philonthus; habitat; Canary Islands).
— Wollaston, 1867: 240 (Philonthus; characters; notes; Cape Verde Islands).
— Fauvel, 1874: 204 [= 1874a: 430] (Erichsoninus; synonym of orbus).
— Fauvel, 1902b: 107 (Actobius; synonym of orbus).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; synonym of orbus).
— Gridelli, 1943: 125 (Neobisnius; synonym of orbus).
— Coiffait, 1974: 329 (Neobisnius; synonym of orbus).
putiphar Saulcy, 1864: 640 (Philonthus; Type locality: Kafr-Zaïat, au bord du Nil).
— Baudi, 1870: 387 (Philonthus; characters).
— Fauvel, 1874: 204 [= 1874a: 430] (Erichsoninus; synonym of orbus).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; synonym of orbus).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; synonym of orbus).
— Gridelli, 1943: 125 (Neobisnius; synonym of orbus).
— Coiffait, 1974: 329 (Neobisnius; synonym of orbus).
nouristanicus Coiffait, 1979d: 559 (Neobisnius; subspecies of orbus; Type locality: gorges de Chikari, entre Djelalabad et Kaboul).
sanguinicollis Lecoq, 1990: 205 (Neobisnius; subspecies of orbus; Type locality: Madagascar Centre: Isalo. Also cited from Réunion Island).
orientalis Cameron, 1933d: 386 (Neobisnius; Type locality: Sumatra: Fort de Kock).
paederoides LeConte, 1863a: 38 (Philonthus; [Note: Preoccupied secondary homonym in Philonthus]; Type locality: “Central Valley”, Illinois, Missouri, etc., USA; [Note: See lectotype designation by Frank, 1981]. Originally cited from Western States; Lake Superior; Fort Yuma and San Diego, California).
— Fauvel, 1874: 201 [= 1874a: 427] (Erichsonius).
— Horn, 1884: 231 (Actobius; characters; Canada to Florida and westward to the Pacific coast).
— Fall and Cockerell, 1907: 166 (Actobius; New Mexico).
— Blatchley, 1910: 393 (Actobius; characters; Indiana).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Notman, 1920: 17 (Actobius; New York).
— Notman, 1920a: 180 (Actobius; New York).
— Hatch, 1957: 175 (Neobisnius; characters; British Columbia; Washington; Idaho).
— Frank, 1981: 17 (Neobisnius; lectotype designation: “Central Valley”, Illinois, Missouri, etc., USA; characters; New York; Maryland; Ohio; Wisconsin; Illinois; Kansas; North Carolina; South Carolina; Georgia; Florida; Alabama; Mississippi; Texas).
— Downie and Arnett, 1996: 374 (Neobisnius; characters; USA).
▴ Distribution: Canada, USA.
palmula Gravenhorst, 1802, see: villosulus Stephens, 1833.
parcepunctatus Bernhauer, 1912e: 176 (Neobisnius; Type locality: Argentinien: Chaunar Region; [Note: According to Frank, 1981: 12, 13 this locality is probably Chañar Region, La Rioja]).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Frank, 1981: 12 (Neobisnius; characters; Argentina).
▴ Distribution: Argentina.
piger Horn, 1884, see: senilis Horn, 1884.
planatus Gravenhorst, 1806, see: procerulus Gravenhorst, 1806.
politus Sharp, 1885, see: nitidulus Sharp, 1885.
praelongus Gemminger and Harold, 1868: 591 (Philonthus; replacement name for longulus Kraatz).
— Fauvel, 1874: 201 [= 1874a: 427] (Erichsonius).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Bernhauer, 1922a: 150 (Neobisnius; Cambodia).
— Cameron, 1932: 56 (Neobisnius; characters; Bihar; Assam; India).
— Scheerpeltz, 1933: 1327 (Neobisnius; catalog).
— Cameron, 1933d: 386 (Neobisnius; Sumatra).
— Cameron, 1937: 4 (Neobisnius; Java).
— Scheerpeltz, 1954: 113 (Neobisnius; India).
— Scheerpeltz, 1957c: 233 (Neobisnius; Sumba).
— Last, 1961: 306 (Neobisnius; Thailand).
— Scheerpeltz, 1965: 106 (Neobisnius; Burma).
— Watanabe and Shibata, 1976: 332 (Neobisnius; Malaysia).
— Coiffait, 1981f: 330 (Neobisnius; Andaman Islands).
— Barrion, Litsinger, and Last, 1991: 1078 (Neobisnius; associated with rice agroecosystem).
— Shibata, 1993a: 318 (Neobisnius; Taiwan).
— Ito, 1996a: 84 (Neobisnius; Japan; Thailand).
▴ Distribution: Sri Lanka, India, Andaman Islands, Myanmar, Thailand, Cambodia, Vietnam, Malaysia, Indonesia, Philippines, Taiwan, Japan.
longulus Kraatz, 1859: 99 (Philonthus; [Note: When this name was replaced it was con-generic with Philonthus longulus Motschulsky which had been described in Gabrius and now is in Bisnius]; Type locality: Ceylan; India orientalis).
— Fauvel, 1878f: 257 (Actobius).
— Fauvel, 1886b: 150 (Actobius; Philippines; Ceylon; East Indies; Annam; Sumatra; Celebes).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; synonym of praelongus).
— Cameron, 1932: 57 (Neobisnius; synonym of praelongus).
— Cameron, 1933d: 386 (Neobisnius; synonym of praelongus).
procerulus Gravenhorst, 1806: 95 (Staphylinus; Type locality: Brunsvigae).
— Mannerheim, 1830: 34 (Gyrohypnus; Finland; Sweden).
— Mannerheim, 1831: 448 (Gyrohypnus; Finland; Sweden).
— Lacordaire, 1835: 409 (Staphylinus; characters; France).
— Nordmann, 1837: 120 (Gyrohypnus; characters; Germany; Finland).
— Erichson, 1839a: 479 (Philonthus; characters; Germany).
— Heer, 1839: 272 (Philonthus; characters; Switzerland).
— Erichson, 1840: 541 (Philonthus; characters; Germany).
— Kolenati, 1846: 15 (Leptacinus; Elisabethopol).
— Hochhuth, 1849: 146 (Philonthus; characters).
— Redtenbacher, 1849: 707 (Philonthus; characters; Austria).
— Fairmaire and Laboulbène, 1856: 530 (Philonthus; characters; France).
— Kraatz, 1857d: 624 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 198 (Philonthus; characters; Austria).
— Thomson, 1860: 169 (Bisnius; characters; Scandinavia).
— Fauvel, 1874: 205 [= 1874a: 431] (Erichsonius; characters; France; Algeria; Caucasus; Bukara).
— Redtenbacher, 1874: 216 (Philonthus; characters; Austria).
— Solsky, 1874: 184 (Philonthus; characters; Turkestan).
— Seidlitz, 1875: 279 (Philonthus; characters; Baltic region).
— Fauvel, 1876a: 72 [= 1876: 257] (Actobius; Syria).
— Mulsant and Rey, 1876: 586 [= 1877: 442] (Bisnius; characters; France).
— Fauvel, 1878: 121 [= 1878a: 41] (Actobius; Algeria).
— Fauvel, 1886: 53 [= 1886a: 45] (Actobius; Tunisia; Algeria).
— Fowler, 1888: 285 (Actobius; characters; Britain).
— Ganglbauer, 1895: 465 (Neobisnius; characters; Europe; Mediterranean region; North America; Chile [Note: This Chilean record probably refers to semipunctatus]; Australia).
— Fauvel, 1902b: 107 (Actobius; Barbary; Egypt; Madeira; Canaries).
— Sainte-Claire Deville, 1906: 94 (Neobisnius; Corsica).
— Fauvel, 1907: 37 (Actobius; Nakuro; Nairobi; Voi; Naivasha; Egypt; Abyssinia; Madagascar; Mediterranean region; Caucasus; Europe; Boukharie; North America).
— Reitter, 1909: 123 (Neobisnius; characters; Germany).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; catalog).
— Johansen, 1914: 424 (Bisnius; characters; Denmark).
— Lea, 1925: 220 (Neobisnius; Australian introduction).
— Portevin, 1929: 356 (Neobisnius; characters; France).
— Bernhauer, 1931d: 583 (Neobisnius; Abyssinia).
— Scheerpeltz, 1933: 1327 (Neobisnius; catalog).
— Méquignon, 1942a: 20 (Neobisnius; Azores).
— Gridelli, 1943: 118 (Neobisnius; characters; Italy; Sardinia).
— Last, 1948: 148 (Neobisnius; characters in key and aedeagal illustration).
— Cameron, 1950g: 33 (Neobisnius; Belgian Congo).
— V. Hansen, 1952: 31 (Neobisnius; characters).
— Fagel, 1954 (Neobisnius; characters; separated from semipunctatus; Europe and North Africa).
— Hatch, 1957: 175 (Neobisnius; characters; Washington; Oregon; [Note: These records may be erroneous since Frank, 1981 does not report the species from North America]).
— Smetana, 1958: 136 (Neobisnius; characters; ecology; Czechoslovakia).
— Lohse, 1964: 172 (Neobisnius; characters; central Europe).
— Horion, 1965: 115 (Neobisnius; Turkey; Caucasus; Uzbekistan; Iberian Peninsula; Corsica; Sardinia; Sicily; Italy; Dalmatia; Albania; Macedonia; Bulgaria; Montenegro; Bosnia; Holland; Belgium; France; Switzerland; Czechoslovakia; Hungary; Poland; Russia; Germany; Austria).
— Smetana, 1967i: 551 (Neobisnius; Bulgaria; Turkey).
— Bolov, 1969a: 328 (Neobisnius; collecting notes; Russia).
— Coiffait, 1973a: 274 (Neobisnius; Morocco).
— Coiffait, 1974: 329 (Neobisnius; characters; Europe; England; Anatolia; Caucasus).
— Pope, 1977: 29 (Neobisnius; Britain).
— Mahler and Pritzl, 1980: 121 (Neobisnius; notes; Denmark).
— Outerelo, 1980a: 56 (Neobisnius; Spain).
— Tóth, 1984: 37 (Neobisnius; characters; Hungary).
— Ádám, 1987: 149 (Neobisnius; collecting notes; Hungary).
— Lucht, 1987: 104 (Neobisnius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark).
— Borges, 1990: Table II (Neobisnius; Azores).
— Maachi, 1991: 250 (Neobisnius; Morocco).
— Smetana, 1993b: 47 (Neobisnius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 27 (Neobisnius; Italy).
— Luff, 1996: 30 (Neobisnius; collecting notes; Isle of Man).
▴ Distribution: Madagascar, Congo, Kenya, Ethiopia, Egypt, Tunisia, Algeria, Morocco, Canary Islands, Madeira, Azores, Europe, Russia, Azerbaijan, Turkey, Syria, Afghanistan, Uzbekistan, Australia, USA (?).
planatus Gravenhorst, 1806: 108 (Staphylinus; Type locality: Borussia).
— Mannerheim, 1830: 34 (Gyrohypnus; Russia).
— Mannerheim, 1831: 448 (Gyrohypnus; Russia).
— Erichson, 1840: 511 (Philonthus; characters; Germany).
— Kraatz, 1857d: 624 (Philonthus; synonym of procerulus).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; synonym of procerulus).
— Smetana, 1958: 137 (Neobisnius; synonym of procerulus).
cinerascens Stephens, 1833: 252 (Gabrius; Type locality: Devonshire).
— Stephens, 1839: 401 (Gabrius; characters; England).
— Gemminger and Harold, 1868: 591 (Philonthus; synonym of procerulus).
— Bernhauer and Schubert, 1914: 323 (Neobisnius; synonym of procerulus).
— Smetana, 1958: 137 (Neobisnius; synonym of procerulus).
filiformis Wollaston, 1854: 585 (Philonthus; Type locality: Madeira: Santa Anna).
— Wollaston, 1857: 192 (Philonthus; habitat; Madeira Islands).
— Wollaston, 1865: 496 (Philonthus; habitat; notes; Madeira Islands).
— Wollaston, 1871: 304 (Philonthus; notes; characters; Madeira).
— Fauvel, 1874: 201 [= 1874a: 427] (Erichsonius).
— Fauvel, 1902b: 107 (Actobius; synonym of procerulus).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; variety of procerulus).
— Smetana, 1958: 137 (Neobisnius; synonym of procerulus).
xantholinoides Wollaston, 1864: 577 (Philonthus; Type locality: Teneriffe: Santa Cruz).
— Wollaston, 1865: 496 (Philonthus; habitat; Canary Islands).
— Fauvel, 1874: 206 [= 1874a: 432] (Erichsoninus; synonym of prolixus).
— Fauvel, 1902b: 107 (Actobius; synonym of procerulus).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; synonym of procerulus).
— Smetana, 1958: 137 (Neobisnius; synonym of procerulus).
decolor Coiffait, 1974: 325, 330 (Neobisnius; subspecies of procerulus; Type locality: Nord Afghanistan: Bala Murghab, 470 m; [Note: This subspecies and Neobisnius procerulus margianus have the same type locality]).
margianus Bordoni, 1975a: 151 (Neobisnius; subspecies of procerulus; Type locality: N-Afghanistan: Prov. Herat: Bala Murghab, 470 m).
prolixus Erichson, 1840: 510 (Philonthus; Type locality: Austria).
— Redtenbacher, 1849: 707 (Philonthus; characters; Austria).
— Kiesenwetter, 1851a: 418 (Philonthus; France; Spain; Dalmatia).
— Fairmaire and Laboulbène, 1856: 530 (Philonthus; characters; France).
— Kraatz, 1857d: 625 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 198 (Philonthus; characters; Austria).
— Hochhuth, 1862: 65 (Philonthus; Russia).
— Fauvel, 1865a: 297 [= 1865b: 47] (Philonthus; habitat; France).
— Fauvel, 1874: 206 [= 1874a: 432] (Erichsonius; characters; France; Britain; Algeria; Caucasus; Iran; Canary Islands).
— Redtenbacher, 1874: 216 (Philonthus; characters; Austria).
— Mulsant and Rey, 1876: 589 [= 1877: 445] (Bisnius; characters; France).
— Fauvel, 1878: 121 [= 1878a: 41] (Actobius; Algeria).
— Fauvel, 1886: 53 [= 1886a: 45] (Actobius; Algeria).
— Fowler, 1888: 286 (Actobius; characters; Britain).
— Ganglbauer, 1895: 465 (Neobisnius; characters; middle Europe; Mediterranean region).
— Fauvel, 1902b: 107 (Actobius; synonym of procerulus).
— Reitter, 1909: 123 (Neobisnius; characters; Germany).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; catalog).
— Johansen, 1914: 425 (Bisnius; characters; Denmark).
— Portevin, 1929: 356 (Neobisnius; characters; France).
— Scheerpeltz, 1933: 1327 (Neobisnius; catalog).
— Gridelli, 1943: 124 (Neobisnius; characters; Italy; Albania; Croatia; Carniola; Germany).
— Last, 1948: 149 (Neobisnius; characters in key and aedeagal illustration).
— V. Hansen, 1952: 31 (Neobisnius; characters).
— Smetana, 1958: 138 (Neobisnius; characters; ecology; Czechoslovakia).
— Lohse, 1964: 172 (Neobisnius; characters; central Europe).
— Horion, 1965: 117 (Neobisnius; Turkey; Caucasus; Britain; Holland; France; Switzerland; Portugal; Spain; Italy; Dalmatia; Bosnia Hercegovina; Serbia; Bulgaria; Greece; Romania; Hungary; Czechoslovakia; Poland; Germany; Austria).
— Smetana, 1967i: 552 (Neobisnius; Yugoslavia; Turkey).
— Coiffait, 1974: 328 (Neobisnius; characters; Europe; Balkans; Caucasus; Transcaucasus).
— Smetana 1975c: 168 (Neobisnius; Mongolia).
— Pope, 1977: 29 (Neobisnius; Britain).
— Tronquet, 1981: 71 (Neobisnius; Afghanistan).
— Tóth, 1984: 36 (Neobisnius; characters; Hungary).
— Lucht, 1987: 104 (Neobisnius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).
— Pilon and Zanetti, 1991: 55 (Neobisnius; Italy).
— Smetana, 1993b: 48 (Neobisnius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 27 (Neobisnius; Italy).
— Gusarov, 1995a: 92 [= 1996: 55] (Neobisnius; synonymic notes).
— Kashcheev, 1999a: 57 (Neobisnius; Kazakhstan).
▴ Distribution: Algeria, Canary Islands, Europe, Russia, Armenia, Azerbaijan, Turkey, Iran, Afghanistan, Uzbekistan, Kazakhstan, Mongolia.
angusticollis Kolenati, 1846: 16 (Leptacinus; Type locality: Armenia montuosa et Caucasus occidentalis).
— Hochhuth, 1849: 111 (Leptacinus; characters; Caucasus; Armenia).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; synonym of prolixus).
— Smetana, 1958: 138 (Neobisnius; synonym of prolixus).
— Coiffait, 1974: 328 (Neobisnius; synonym of prolixus).
pubipennis Kolenati, 1846: 16 (Leptacinus; Type locality: mons Ssarijal provinciae Elisa-bethopoleos).
— Hochhuth, 1849: 111 (Leptacinus; characters; Elisabethopol).
— Hochhuth, 1851: 10 (Leptacinus; notes).
— Fauvel, 1869: 489 (Philonthus; synonym of prolixus).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; synonym of prolixus).
— Smetana, 1958: 138 (Neobisnius; synonym of prolixus).
— Coiffait, 1974: 328 (Neobisnius; synonym of prolixus).
dissimilis Baudi, 1870: 387 (Philonthus; Type locality: Cypro).
— Fauvel, 1874: 206 [= 1874a: 432] (Erichsoninus; synonym of prolixus).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; synonym of prolixus).
— Gridelli, 1943: 125 (Neobisnius; probably a synonym of orbus).
— Smetana, 1958: 138 (Neobisnius; synonym of prolixus).
hauseri Bernhauer, 1941a: 227 (Neobisnius; Type locality: Buchara: Mts. Karateghin, Sary-pul, 1482 m).
— Coiffait, 1970d: 143 (Neobisnius; Uzbekistan).
— Gusarov, 1995a: 92 [= 1996: 55] (Neobisnius; synonym of prolixus).
pubipennis Kolenati, 1846, see: prolixus Erichson, 1840.
pumilus Sharp, 1874: 49 (Philonthus; Type locality: Osaka).
— Fauvel, 1874: 201 [= 1874a: 427] (Erichsonius).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; catalog).
— Cameron, 1949b: 467 (Neobisnius; Japan, Formosa).
— Watanabe, 1961: 353 (Leptacinus; Izu Islands).
— Nakane, 1963a: 90 (Neobisnius; characters; habitus photograph; Japan).
— Cho, 1996a: 162 (Neobisnius; characters; Korea).
▴ Distribution: Taiwan, Japan, Korea.
putiphar Saulcy, 1864, see: orbus Kiesenwetter, 1850.
richteri Bernhauer, 1927e: 245 (Neobisnius; Type locality: Argentinien: Prov. Buenos Aires).
— Scheerpeltz, 1933: 1327 (Neobisnius; catalog).
— Frank, 1981: 35 (Neobisnius; characters; Argentina; Brazil).
▴ Distribution: Argentina, Brazil.
rubripennis Gridelli, 1943, see: lathrobioides Baudi, 1848.
rufipennis Cameron, 1919a: 252 (Neobisnius; Type locality: Ceylon: Nuwera Eliya, alt. 6230–8000 feet).
— Cameron, 1932: 58 (Neobisnius; characters; Ceylon).
— Scheerpeltz, 1933: 1327 (Neobisnius; catalog).
— Scheerpeltz, 1954: 113 (Neobisnius; Ceylon).
▴ Distribution: Sri Lanka.
sanguinicollis Lecoq, 1990, see: orbus Kiesenwetter, 1850.
sanguinipennis Scheerpeltz, 1972: 110 (Neobisnius; Type locality: Argentinien: Rio Negro, El Bolson, 350 m).
scutellaris Bernhauer, 1908d: 336 (Neobisnius; Type locality: Paraguay).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; catalog).
— Scheerpeltz, 1933: 1328 (Neobisnius; catalog; Argentina).
— Scheerpeltz, 1972: 17 (Neobisnius; Argentina).
— Frank, 1981: 30 (Neobisnius; characters; Bolivia; Brazil; Paraguay; Argentina; Uruguay).
▴ Distribution: Brazil, Bolivia, Paraguay, Uruguay, Argentina.
semipunctatus Fairmaire and Germain, 1861: 434 (Othius; Type locality: Santiago).
— Fauvel, 1866a: 346 [= 1866b: 97] (Philonthus; characters; Chile).
— Fauvel, 1874: 205 [= 1874a: 431] (Erichsonius; characters; France; Spain; Algeria; Chile; Australia [Note: This species only occurs in Chile according to Fagel, 1954, so all but Chile of the preceding citations are erroneous]).
— Fauvel, 1876a: 72 [= 1876: 257] (Actobius; Ethiopia [Note: This species only occurs in Chile according to Fagel, 1954, so the preceeding citation is erroneous]).
— Mulsant and Rey, 1876: 842 [= 1877: 698] (Bisnius; characters; France [Note: This species only occurs in Chile according to Fagel, 1954, so the citation is erroneous]).
— J. Sahlberg, 1876: 23 (Bisnius; characters; Finland [Note: This species only occurs in Chile according to Fagel, 1954, so the citation is erroneous]).
— Fauvel, 1878: 121 [= 1878a: 41] (Actobius; Algeria [Note: This species only occurs in Chile according to Fagel, 1954, so the citation is erroneous]).
— Fauvel, 1878g: 545 (Actobius; characters; Australia [Note: This species only occurs in Chile according to Fagel, 1954, so the citation is erroneous]).
— Horn, 1884: 227 (Actobius; characters; Colorado; Nevada [Note: This species only occurs in Chile according to Fagel, 1954, so the citation is erroneous]).
— Fauvel, 1886: 53 [= 1886a: 45] (Actobius; Algeria [Note: This species only occurs in Chile according to Fagel, 1954, so the citation is erroneous]).
— Ganglbauer, 1895: 465 (Neobisnius; synonym of procerulus).
— Blackburn, 1896: 262 (Actobius; Australia [Note: This species only occurs in Chile according to Fagel, 1954, so the citation is erroneous]).
— Fauvel, 1902b: 107 (Actobius; synonym of procerulus).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; variety of procerulus).
— Portevin, 1929: 356 (Neobisnius; characters; France [Note: This species only occurs in Chile according to Fagel, 1954, so the citation is erroneous]).
— Scheerpeltz, 1933: 1327 (Neobisnius; variety of procerulus).
— Gridelli, 1943: 127 (Neobisnius; characters).
— Fagel, 1954 (Neobisnius; valid species; characters; separated from procerulus; all distributional records outside of Chile probably represent different species).
— Hatch, 1957: 175 (Neobisnius; variety of procerulus; characters; British Columbia; Washington; Oregon; [Note: This species only occurs in Chile according to Fagel, 1954, so the citation by Hatch is erroneous]).
— Coiffait and Saiz, 1968: 361 (Neobisnius; characters; Chile).
— Saiz, 1971b: 349 (Neobisnius; Chile).
— Frank, 1981: 10 (Neobisnius; characters; Chile).
▴ Distribution: Chile.
semirufus Bernhauer, 1921c: 105 (Neobisnius; Type locality: Argentinien: Prov. Salta).
— Scheerpeltz, 1933: 1328 (Neobisnius; catalog).
— Frank, 1981: 33 (Neobisnius; characters; Argentina; Bolivia).
▴ Distribution: Argentina, Bolivia.
senilis Horn, 1884: 226 (Actobius; Type locality: Garland, Costilla Co., Colorado; [Note: See lectotype designation by Frank, 1981]. Originally cited from Colorado; Nevada; California).
— Fall and Cockerell, 1907: 166 (Actobius; New Mexico).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
— Hatch, 1957: 175 (Neobisnius; characters; British Columbia; Washington; Oregon; Idaho).
— Frank, 1981: 45 (Neobisnius; lectotype designation: Garland, Costilla Co., Colorado; characters; British Columbia; Washington; Idaho; Oregon; Colorado; Nevada; New Mexico; California; Mexico).
▴ Distribution: Canada, USA, Mexico.
piger Horn, 1884: 219 (Philonthus; Type locality: Not cited).
simplex Sharp, 1885: 459 (Actobius; Type locality: Mexico, Mirador).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; catalog).
— Frank, 1981: 42 (Neobisnius; characters; Mexico; Guatemala; El Salvador; Costa Rica; Panama; Brazil).
▴ Distribution: Brazil, Panama, Costa Rica, El Salvador, Guatemala, Mexico.
cavifrons Bierig, 1933: 51 (Neobisnius; Type locality: Panama: Puerto Armuelles. Also listed from Costa Rica).
simulator Smetana, 1963, see: sobrinus Erichson, 1840.
sobrinus Erichson, 1840: 512 (Philonthus; Type locality: Pensylvania; [Note: See lectotype designation by Smetana, 1963]).
— Fauvel, 1874: 201 [= 1874a: 427] (Erichsonius).
— Horn, 1884: 228 (Actobius; characters; east of Rocky Mountains; California).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; catalog).
— Notman, 1920: 17 (Actobius; New York).
— Notman, 1920a: 180 (Actobius; New York).
— Smetana, 1963: 4, 5 (Neobisnius; characters; lectotype designation; Pennsylvania).
— Smetana, 1965: 10 (Neobisnius; Nova Scotia; Newfoundland).
— Frank, 1981: 39 (Neobisnius; characters; Newfoundland; New Brunswick; Quebec; Ontario; Maine; New Hampshire; Vermont; Massachusetts; Connecticut; New York; Pennsylvania; Illinois; Wisconsin; Iowa; Nebraska; Oregon; New Jersey; Kentucky; Missouri; Kansas; Colorado; Utah; Maryland; Virginia; West Virginia; North Carolina; Tennessee; South Carolina; Georgia; Florida; Louisiana; Texas; New Mexico; Arizona; California; Mexico; Guatemala; Costa Rica; Venezuela; Colombia; Brazil).
— Schmidt, 1994 (Neobisnius; egg, larval, and pupal characters; biological notes; Nebraska).
— Downie and Arnett, 1996: 375 (Neobisnius; characters; USA).
— Schmidt, 1999: 108 (Neobisnius; food for adults and larvae used for rearing species; rearing methods).
▴ Distribution: Canada, USA, Mexico, Guatemala, Costa Rica, Colombia, Venezuela, Brazil.
agnatus Erichson, 1840: 513 (Philonthus; Type locality: valle Araguensi Columbiae; [Note: See lectotype designation by Smetana, 1963]).
— Solsky, 1868a: 137 (Philonthus; Mexico).
— Fauvel, 1874: 201 [= 1874a: 427] (Erichsonius).
— Horn, 1884: 229 (Actobius; characters; Louisiana).
— Sharp, 1885: 459 (Actobius; notes; Mexico; Louisiana; Colombia).
— Bernhauer and Schubert, 1914: 322 (Neobisnius; catalog).
— Smetana, 1963: 4 (Neobisnius; characters; lectotype designation).
— Frank, 1981: 39 (Neobisnius; synonym of sobrinus).
ocreatus Horn, 1884: 228 (Actobius; Type locality: southern California; [Note: See lectotype designation by Frank, 1981]. Originally cited from California from the middle regions southward).
— Bernhauer and Schubert, 1914: 325 (Actobius; catalog).
— Frank, 1981: 39 (Neobisnius; lectotype designation: southern California; synonym of sobrinus).
arduus Sharp, 1885: 458 (Actobius; Type locality: Puebla, Mexico; [Note: See lectotype designation by Frank, 1981]. Originally cited from Mexico, Guanajuato, Cordova, Puebla, Mirador, Tehuacan, Jalapa; Guatemala, near the city, San Geronimo, Torola, Senahu, Balheu, Tactic, Panajachel 5000 feet, Cerro Zunil 4000 feet; Panama, San Miguel in the Pearl Islands).
— Bernhauer and Schubert, 1914: 322 (Neobisnius; catalog).
— Frank, 1981: 39 (Neobisnius; lectotype designation: Puebla, Mexico; synonym of sobrinus).
simulator Smetana, 1963: 3 (Neobisnius; Type locality: West-Newfoundland: Curling).
subaeneus Bernhauer, 1928: 15 (Neobisnius; Type locality: Sumatra: Medan).
— Scheerpeltz, 1933: 1328 (Neobisnius; catalog).
— Cameron, 1933d: 386 (Neobisnius; Sumatra).
▴ Distribution: Indonesia.
subprolixus Coiffait, 1982: 68 (Neobisnius; Type locality: Nepal: Ufer des Tila Khola, près Jumla).
sumbaensis Scheerpeltz, 1957c: 233, 316 (Neobisnius; Type locality: Sumba: Baing).
tenellus Wollaston, 1864, see: orbus Kiesenwetter, 1850.
terminalis LeConte, 1863a: 38 (Philonthus; Type locality: Maryland; [Note: See lectotype designation by Frank, 1981]).
— Fauvel, 1874: 201 [= 1874a: 427] (Erichsonius).
— Horn, 1884: 232 (Actobius; characters; Canada to middle States; Texas).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; catalog).
— Notman, 1919: 136 (Actobius; New York).
— Notman, 1920: 17 (Actobius; New York).
— Notman, 1920a: 180 (Actobius; New York).
— Bierig, 1933: 56 (Neobisnius).
— Smetana, 1965: 11 (Neobisnius; Nova Scotia).
— Frank, 1981: 13 (Neobisnius; lectotype designation; characters; Nova Scotia; Quebec; Maine; New Hampshire; New York; Pennsylvania; Michigan; Iowa; Colorado; Maryland; Virginia; Texas; New Mexico; Arizona; California; Baja California; Tamaulipas; Guerrero; Sinaloa; Puebla; Oaxaca; Guatemala; Costa Rica).
— Dearborn and Donahue, 1994: 14 (Neobisnius; Maine).
— Downie and Arnett, 1996: 374 (Neobisnius; characters; USA).
▴ Distribution: Canada, USA, Mexico, Guatemala, Costa Rica, Panama.
elegantulus Horn, 1884: 232 (Actobius; Type locality: Arizona; [Note: See lectotype designation by Frank, 1981]. Also cited from California).
— Bernhauer and Schubert, 1914: 322 (Neobisnius; catalog).
— Frank, 1981: 13 (Neobisnius; lectotype designation: Arizona; synonym of terminalis).
delicatulus Sharp, 1885: 461 (Actobius; Type locality: Guatemala, Pantaleon 1700 feet, Paso Antonio 400 feet).
— Bernhauer and Schubert, 1914: 322 (Neobisnius; catalog).
— Frank, 1981: 13 (Neobisnius; synonym of terminalis).
adustus Sharp, 1887: 790 (Actobius; Type locality: Mexico, La Noria in Sinaloa, Tehuantepec).
— Bernhauer and Schubert, 1914: 322 (Neobisnius; catalog).
— Bierig, 1933: 48 (Neobisnius; characters; Panama).
— Frank, 1981: 13 (Neobisnius; synonym of terminalis).
transvaalensis Scheerpeltz, 1974: 134 (Neobisnius; Type locality: Transvaal: Kruger National Park, Letaba Camp).
umbripennis LeConte, 1863, see: ludicrus Erichson, 1840.
vigii Frank, 1981: 43 (Neobisnius; Type locality: Colombia: Magdalena, Parque Tayrona, 21 mi. E of Santa Marta).
villosulus Stephens, 1833: 251 (Gabrius; Type locality: Suffolk).
— Stephens, 1839: 401 (Gabrius; characters; England).
— Fauvel, 1874: 204 [= 1874a: 430] (Erichsonius; characters; France; Caucasus).
— J. Sahlberg, 1876: 22 (Bisnius; Finland).
— Fowler, 1888: 285 (Actobius; characters; Britain).
— Ganglbauer, 1895: 464 (Neobisnius; characters; Europe).
— Reitter, 1909: 123 (Neobisnius; characters; Germany).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; catalog).
— Johansen, 1914: 424 (Bisnius; characters; Denmark).
— Portevin, 1929: 356 (Neobisnius; characters; France).
— Scheerpeltz, 1933: 1328 (Neobisnius; catalog).
— Tottenham, 1949: 417 (Neobisnius; checklist; Britain).
— V. Hansen, 1952: 30 (Neobisnius; characters; Denmark).
— Blackwelder, 1952: 258 (Neobisnius; type species of genus).
— Smetana, 1958: 135 (Neobisnius; characters; ecology; Czechoslovakia).
— Palm, 1963b: 58 (Neobisnius; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 171 (Neobisnius; characters; central Europe).
— Horion, 1965: 114 (Neobisnius; Russia; Finland; Sweden; Norway; Britain; Denmark; Holland; Belgium; France to Poland; Romania; Italy; Slovenia; Dalmatia; Bosnia; Germany; Austria).
— Szujecki, 1968a: 726 (Neobisnius; cited as villosus; Poland).
— Tikhomirova, 1973a: 182 (Neobisnius; checklist; Soviet Union).
— Coiffait, 1974: 327 (Neobisnius; characters; Europe).
— Moore and Legner, 1975: 18 (Neobisnius; type species of genus).
— Pope, 1977: 29 (Neobisnius; Britain).
— Muona, 1979: 14 (Neobisnius; Finland; Russia; Sweden; Norway; Denmark).
— Moore and Legner, 1979: 25 (Neobisnius; type species of genus).
— Burakowski, Mroczkowski, and Stefańska, 1980: 31 (Neobisnius; catalog; Poland).
— Frank, 1981: 8 (Neobisnius; characters; Quebec; Ontario; New York; Minnesota; Oregon).
— Uhlig and Vogler, 1981: 110 (Neobisnius; notes; Germany).
— Tóth, 1984: 35 (Neobisnius; characters; Hungary).
— Lucht, 1987: 104 (Neobisnius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Gusarov, 1989: 10 (Neobisnius; habitat notes; Ukraine).
— Lohse and Lucht, 1989: 160 (Neobisnius; notes).
— Zanetti, 1989: 112 (Neobisnius; Italy).
— Silfverberg, 1992: 17 (Neobisnius; Finland; Russia; Sweden; Norway; Denmark; Estonia; Lithuania).
— Smetana, 1993b: 48 (Neobisnius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 27 (Neobisnius; Italy).
— Downie and Arnett, 1996: 375 (Neobisnius; characters; USA).
— M. Hansen, 1996: 104 (Neobisnius; checklist; Denmark).
— Anderson, Nash, and O'Connor, 1997: 18 (Neobisnius; Ireland).
— Herman, 2001: 26 (Neobisnius; nomen protectum).
▴ Distribution: Europe, Canada, USA.
palmula Gravenhorst, 1802: 49 (Staphylinus; [Note: This name is older than villosulus]; Type locality: Not cited, but presumably Brunsvigae).
— Latreille, 1804: 333 (Staphylinus; synonym of cinctus Latreille).
— Gravenhorst, 1806: 107 (Staphylinus).
— Gyllenhal, 1810: 359 (Staphylinus; synonym of pilicornis Paykull).
— Zetterstedt, 1828: 81 (Xantholinus; synonym of pilicornis Paykull).
— Erichson, 1840: 514 (Philonthus; characters; Germany).
— Redtenbacher, 1849: 825 (Philonthus; characters; Austria).
— Kraatz, 1857d: 625 (Philonthus; notes).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; synonym of villosulus).
— Scheerpeltz, 1933: 1328 (Neobisnius; synonym of villosulus).
— Gridelli, 1943: 118 (Neobisnius; synonym of elongatulus).
— Smetana, 1958: 135 (Neobisnius; synonym of villosulus).
— Coiffait, 1974: 327 (Neobisnius; synonym of villosulus).
— Herman, 2001: 26 (Neobisnius; nomen oblitum).
cinctus Latreille, 1804: 333 (Staphylinus; [preoccupied]; Type locality: Brunswick).
— Erichson, 1840: 514 (Philonthus; synonym of palmula).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; synonym of villosulus).
— Smetana, 1958: 135 (Neobisnius; synonym of villosulus).
elongatulus Erichson, 1839a: 479 (Philonthus; Type locality: Mark Brandenburg).
— Heer, 1839: 272 (Philonthus; characters; Switzerland).
— Erichson, 1840: 511 (Philonthus; characters; Germany).
— Jacobsen, 1849: 706 (Philonthus; characters; Denmark).
— Hochhuth, 1849: 146 (Philonthus; characters; Mingrelien).
— Redtenbacher, 1849: 707 (Philonthus; characters; Austria).
— Fairmaire and Laboulbène, 1856: 531 (Philonthus; characters; France).
— Kraatz, 1857d: 623 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 197 (Philonthus; characters; Austria).
— Thomson, 1859: 25 (type species of Bisnius).
— Thomson, 1860: 168 (Bisnius; characters; Scandinavia).
— Fauvel, 1865a: 297 [= 1865b: 47] (Philonthus; habitat; France).
— Fauvel, 1874: 204 [= 1874a: 430] (Erichsonius; synonym of villosulus).
— Redtenbacher, 1874: 216 (Philonthus; characters; Austria).
— Seidlitz, 1875: 279 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 594 [= 1877: 450] (Bisnius; characters; France).
— Ganglbauer, 1895: 464 (Neobisnius; synonym of villosulus).
— Bernhauer and Schubert, 1914: 324 (Neobisnius; synonym of villosulus).
— Scheerpeltz, 1933: 1328 (Neobisnius; synonym of villosulus).
— Gridelli, 1943: 118 (Neobisnius; characters; Italy; Balkan peninsula).
— Last, 1948: 149 (Neobisnius; characters in key and aedeagal illustration).
— Smetana, 1958: 135 (Neobisnius; synonym of villosulus).
— Coiffait, 1974: 327 (Neobisnius; synonym of villosulus).
— Lohse and Lucht, 1989: 160 (Neobisnius; synonym of villosulus).
fulvicornis Notman, 1920: 27 (Actobius; Type locality: Schoharie, N.Y.).
— Scheerpeltz, 1933: 1329 (Neobisnius; catalog).
— Frank, 1981: 8 (Neobisnius; synonym of villosulus).
xantholinoides Wollaston, 1864, see: procerulus Gravenhorst, 1806.
Neoleucitus
[1 species; Oriental Region]
Neoleucitus Wendeler, 1924: 341 (species included: rugosus). Type species: Neoleucitus rugosus Wendeler, fixed by monotypy.
— Scheerpeltz, 1933: 1380 (world catalog supplement).
— Blackwelder, 1952: 259 (type species: rugosus).
rugosus Wendeler, 1924: 342 (Neoleucitus; Type locality: Luzon-Beenguet, Baguio, St. Thomas, 6500 ft.).
Odontolinus
[1 species; Neotropical Region]
Odontolinus Sharp, 1885: 454 (species included: fasciatus). Type species: Odontolinus fasciatus Sharp, fixed by monotypy.
— Bernhauer and Schubert, 1914: 372 (world catalog; 1 species).
— Blackwelder, 1944: 138 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 269 (type species: fasciatus).
fasciatus Sharp, 1885: 454 (Odontolinus; Type locality: Panama, Bugaba 800 to 1500 feet, Volcan de Chiriqui 2500 to 4000 feet).
Onthostygnus
[2 species; Neotropical Region]
Onthostygnus Sharp, 1884: 392 (species included: fasciatus; [Note: A second species, pollens, was included on the next page which was published in 1885]). Type species: Onthostygnus fasciatus Sharp, fixed by monotypy.
— Bernhauer and Schubert, 1914: 366 (world catalog; 2 species).
— Blackwelder, 1944: 136 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 275 (type species: fasciatus).
fasciatus Sharp, 1884: 392 (Onthostygnus; Type locality: Mexico, Jalapa).
pollens Sharp, 1885: 393 (Onthostygnus; Type locality: Mexico, Jalapa).
Ophionthus
[1 species; Neotropical Region]
Ophionthus Bernhauer, 1908d: 328 (species included: serpentinus). Type species: Ophionthus serpentinus Bernhauer, fixed by monotypy.
— Bernhauer and Schubert, 1914: 372 (world catalog; 1 species).
— Blackwelder, 1944: 138 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 276 (type species: serpentinus).
serpentinus Bernhauer, 1908d: 329 (Ophionthus; Type locality: Peruvia centralis).
Orthidus
[1 species; Palaearctic Region]
Orthidus Mulsant and Rey, 1876: 339 [= 1876: 195] (species included: cribratus). Type species: Philonthus cribratus Erichson, fixed by monotypy.
— Heyden, Reitter, and Weise, 1891: 102 (list of species of Europe and Caucasus).
— Ganglbauer, 1895: 436 (characters).
— Heyden, Reitter, and Weise, 1906: 168 (list of species of Europe and Caucasus).
— Reitter, 1909: 116, 122 (characters).
— Bernhauer and Schubert, 1914: 361 (world catalog; 1 species).
— Cameron, 1919b: 207 (synonym of Phucobius).
— Cameron, 1921a: 354, 405 (characters; key to Singapore species; catalog).
— Winkler, 1925: 381 (catalog for Palaearctic region).
— Porta, 1926: 99 (characters; species of Italy).
— Portevin, 1929: 354 (Orthidus; characters; France).
— Scheerpeltz, 1933: 1368 (world catalog supplement).
— Normand, 1935: 372 (list of species of Tunisia).
— Blackwelder, 1952: 278 (type species: cribratus).
— Coiffait, 1956e: 183, 187, 220 (characters; notes).
— Kocher, 1958: 152 (checklist of species; Morocco).
— Smetana, 1959a: 214 (list of species collected in Albania).
— Coiffait, 1963: 8, 20 (characters).
— Tikhomirova, 1973a: 186 (checklist of species of USSR).
— Coiffait, 1974: 350 (characters; species of western Palaearctic region).
— Moore and Legner, 1976: 541 (includes seashore species; notes).
— Smetana, 1995e: 49 (characters).
— Ciceroni and Zanetti, 1995: 27 (list of species of Italy).
atlanticus Coiffait, 1956, see: cribratus Erichson, 1840.
cribratus Erichson, 1840: 431 (Philonthus; Type locality: Istria; Gallia meridionali; Sardinia; Lusitania).
— Küster, 1853a: 45 (Philonthus; characters; France; Spain; Portugal; Sardinia; Istria; Dalmatia).
— Fairmaire and Laboulbène, 1856: 514 (Philonthus; characters; France).
— Kraatz, 1857d: 569 (Philonthus; characters).
— Redtenbacher, 1857: 189 (Philonthus; characters; Austria).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius; characters; France; Austria; Italy; Corsica; Sardinia; Portugal; Algeria).
— Redtenbacher, 1874: 208 (Philonthus; characters; Austria).
— Mulsant and Rey, 1876: 341 [= 1877: 197] (Anodus; characters; France).
— Fauvel, 1878: 120 [= 1878a: 40] (Cafius; Algeria; Tunisia).
— Fauvel, 1886: 52 [= 1886a: 44] (Cafius; Morocco; Tunisia; Algeria).
— Ganglbauer, 1895: 436 (Orthidus; characters; coast of western Mediterranean region; Atlantic coast of France).
— Fauvel, 1902b: 106 (Cafius; Morocco; Algeria; Tunisia).
— Sainte-Claire Deville, 1906: 95 (Orthidus; Corsica).
— Reitter, 1909: 122 (Orthidus).
— Bernhauer and Schubert, 1914: 361 (Orthidus; catalog).
— Scheerpeltz, 1933: 1368 (Orthidus; catalog).
— Coiffait, 1956e: 221 (Orthidus; characters).
— Focarile, 1964: 58 (Orthidus; Italy).
— Coiffait, 1974: 352 (Orthidus; characters; Mediterranean coast of Europe and North Africa).
— Bordoni and Carfi, 1978 (Orthidus; note on peg setae of aedeagus).
— Ciceroni and Zanetti, 1995: 27 (Orthidus; Italy).
▴ Distribution: Tunisia, Algeria, Morocco, Portugal, Spain, France, Italy, Croatia.
atlanticus Coiffait, 1956e: 221 (Orthidus; subspecies of cribratus; Type locality: côtes atlantiques de la Bretagne aux Pyrénées et probablement en Espagne).
cupreipennis Cameron, 1918, see: Phucobius.
Pachypelmopus
[1 species; Australian Region]
Pachypelmopus Scheerpeltz, 1966b: 204 (species included: buergersi). Type species: Diplostictus buergersi Bernhauer, fixed by original designation.
buergersi Bernhauer, 1915g: 192 (Diplostictus; cited as bürgersi; Type locality: Deutsch-Neuguinea: Hundsteinspitze).
Paederallus
[1 species; Neotropical Region]
Paederallus Sharp, 1885: 456 (species included: fragilis). Type species: Paederallus fragilis Sharp, fixed by monotypy.
— Bernhauer and Schubert, 1914: 326 (world catalog; 1 species).
— Blackwelder, 1944: 132 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 285 (type species: fragilis).
fragilis Sharp, 1885: 457 (Paederallus; Type locality: Panama, Peña Blanca 3000 to 4000 feet).
Paederomimus
[57 species; Neotropical Region]
Paederomimus Sharp, 1885: 438 (species included: laetus, lustralis, aeniceps, cognatus, traili, capitalis, lustrator, aeneicollis, palpalis, aberrans, conformis, propinquus, regillus, abactus, longipes, gentilis, mandibularis, chontalensis, salvini, capitatus, hilaris, nobilis, nigrinus, angularis, pectoralis, punctipennis, vicinus, simplex, mexicanus, maculipes, difformiceps, cephalax, nigripennis, insolitus, angusticollis, cribricollis, contractus, confertus, ornatellus, basipennis, distans, cribratus, perforatus, corrosus). Type species: Paederomimus difformiceps Sharp, fixed by original designation.
— Bernhauer and Schubert, 1914: 367 (world catalog; 50 species).
— Scheerpeltz, 1933: 1376 (world catalog supplement).
— Blackwelder, 1943: 395 (characters; type species: difformiceps; key to species of West Indies).
— Blackwelder, 1944: 136 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 286 (type species: difformiceps).
abactus Sharp, 1876: 177 (Philonthus; Type locality: Ega).
— Sharp, 1885: 438 (Paederomimus).
— Bernhauer and Schubert, 1914: 367 (Paederomimus; catalog).
▴ Distribution: Brazil.
aberrans Sharp, 1876: 174 (Philonthus; Type locality: Ega).
— Sharp, 1885: 438 (Paederomimus).
— Bernhauer and Schubert, 1914: 367 (Paederomimus; catalog).
▴ Distribution: Brazil.
aeneiceps Sharp, 1876: 169 (Philonthus; Type locality: Ega).
— Sharp, 1885: 438 (Paederomimus).
— Bernhauer and Schubert, 1914: 367 (Paederomimus; catalog).
▴ Distribution: Brazil.
aeneicollis Sharp, 1876: 172 (Philonthus; Type locality: Ega; St. Paulo).
— Sharp, 1885: 438 (Paederomimus).
— Bernhauer and Schubert, 1914: 367 (Paederomimus; catalog).
▴ Distribution: Brazil.
angularis Erichson, 1840: 477 (Philonthus; Type locality: Mexico).
— Sharp, 1885: 443 (Paederomimus; Mexico).
— Bernhauer and Schubert, 1914: 367 (Paederomimus; catalog).
— Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 62 (Paederomimus; necrophilous species; distribution in Mexico).
▴ Distribution: Mexico.
angusticollis Erichson, 1840: 487 (Philonthus; Type locality: Mexico).
— Sharp, 1885: 447 (Paederomimus; notes; Mexico; Guatemala).
— Bernhauer and Schubert, 1914: 367 (Paederomimus; catalog).
▴ Distribution: Mexico, Guatemala.
aterrimus Bernhauer, 1917d: 36 (Paederomimus; Type locality: West-Columbien: Umgebung von Cali am Rio Cauca, Cañon del Mte. Tolima, 1700 m).
basipennis Sharp, 1885: 450 (Paederomimus; Type locality: Panama, Chiriqui).
capitalis Sharp, 1876: 171 (Philonthus; Type locality: Ega).
— Sharp, 1885: 438 (Paederomimus).
— Bernhauer and Schubert, 1914: 367 (Paederomimus; catalog).
▴ Distribution: Brazil.
capitatus Sharp, 1885: 441 (Paederomimus; Type locality: Mexico, Jalapa, Playa Vicente).
cephalax Sharp, 1885: 446 (Paederomimus; Type locality: Panama, David).
chontalensis Sharp, 1885: 440 (Paederomimus; Type locality: Nicaragua, Chontales).
cognatus Sharp, 1876: 169 (Philonthus; [preoccupied]; Type locality: Ega).
— Sharp, 1885: 438 (Paederomimus).
— Bernhauer and Schubert, 1914: 367 (Paederomimus; catalog).
— Herman, 2001: 46 (Paederominus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Brazil.
confertus Sharp, 1885: 449 (Paederomimus; Type locality: Panama, David, Volcan de Chiriqui about 4000 feet).
conformis Sharp, 1876: 175 (Philonthus; [Note: Preoccupied as secondary homonym in Philonthus]; Type locality: Amazons).
— Sharp, 1885: 438 (Paederomimus).
— Bernhauer and Schubert, 1914: 367 (Paederomimus; catalog).
▴ Distribution: Brazil.
contractus Sharp, 1885: 448 (Paederomimus; Type locality: Guatemala, San Geronimo, Cubilguitz; Cahabon, Chacoj; Nicaragua).
corrosus Sharp, 1885: 451 (Paederomimus; Type locality: Panama, Bugaba 800 to 1500 feet).
cribratus Sharp, 1885: 450 (Paederomimus; Type locality: Panama, Volcan de Chiriqui 2000 to 3000 feet).
cribricollis Erichson, 1840: 487 (Philonthus; Type locality: Columbia; Mexico).
— Solsky, 1875: 15 (Philonthus; Peru).
— Sharp, 1885: 448 (Paederomimus; notes; Mexico; Colombia).
— Fauvel, 1891: 118 (Paederomimus; Venezuela).
— Fauvel, 1901: 87 (Paederomimus; Colombia; Venezuela; Antilles).
— Bernhauer and Schubert, 1914: 368 (Paederomimus; catalog).
— Blackwelder, 1943: 396 (Paederomimus; characters; Grenada; Trinidad).
— Marcuzzi, 1977: 12 (Paederomimus, cited as Paederimimus; Venezuela).
▴ Distribution: West Indies, Mexico, Venezuela, Colombia, Peru.
trinidadensis Bernhauer, 1915l: 296 (Paederomimus; Type locality: Trinidad: Caparo).
— Scheerpeltz, 1933: 1376 (Paederomimus; catalog).
— Blackwelder, 1943: 397 (Paederomimus; synonym of cribricollis).
cupreonitens Schubert, 1909: 294 (Paederomimus; Type locality: südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d'Orizaba).
densiceps Bernhauer, 1908d: 332 (Paederomimus; Type locality: Brasilien: Petropolis).
difformiceps Sharp, 1885: 445 (Paederomimus; Type locality: Guatemala, near the city, Capetillo).
distans Sharp, 1885: 450 (Paederomimus; Type locality: Guatemala, Zapote).
flavoguttatus Bernhauer, 1908d: 332 (Paederomimus; Type locality: Brasilien).
gentilis Sharp, 1885: 439 (Paederomimus; Type locality: Mexico, Cerro de Plumas, Jalapa).
— Bernhauer and Schubert, 1914: 368 (Paederomimus; catalog).
— Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 62 (Paederomimus; necrophilous species; distribution in Mexico).
▴ Distribution: Mexico.
hilaris Sharp, 1885, see: sharpi Bernhauer and Schubert, 1914.
hilaris Fauvel, 1874: 246 [= 1874a: 472] (Philonthus; replacement name for laetus Erichson).
laetus Erichson, 1840: 477 (Philonthus; [preoccupied]; Type locality: Columbia; Brasilia).
— Sharp, 1885: 438 (Paederomimus).
— Bernhauer and Schubert, 1914: 368 (Paederomimus; synonym of hilaris).
imitator Bierig, 1937: 198 (Paederomimus; Type locality: Panama: Volcan de Chiriqui, 1500 m).
insolitus Sharp, 1885: 447 (Paederomimus; Type locality: Panama, Bugaba 800 to 1500 feet).
insularis Bernhauer, 1908, see: lustralis Erichson, 1840.
interjectus Bernhauer, 1908d: 333 (Paederomimus; Type locality: St. Vincent: W. J [I?]. Windward side).
— Bernhauer and Schubert, 1914: 368 (Paederomimus; catalog).
— Blackwelder, 1943: 397 (Paederomimus; characters; St. Vincent).
▴ Distribution: West Indies.
smithi Cameron, 1922: 121 (Paederomimus; Type locality: St. Vincent).
— Scheerpeltz, 1933: 1376 (Paederomimus; catalog).
— Blackwelder, 1943: 397 (Paederomimus; synonym of interjectus).
klimschi Bernhauer, 1917d: 36 (Paederomimus; Type locality: Brasilien: S. Catharina).
laetus Erichson, 1840, see: hilaris Fauvel, 1874.
longipes Sharp, 1876: 178 (Philonthus; Type locality: Obydos).
— Sharp, 1885: 438 (Paederomimus).
— Bernhauer and Schubert, 1914: 368 (Paederomimus; catalog).
▴ Distribution: Brazil.
lustralis Erichson, 1840: 489 (Philonthus; Type locality: Puerto Rico).
— Sharp, 1885: 438 (Paederomimus).
— Bernhauer and Schubert, 1914: 368 (Paederomimus; catalog).
— Blackwelder, 1943: 398 (Paederomimus; characters; Puerto Rico; St. Thomas).
▴ Distribution: West Indies.
insularis Bernhauer, 1908d: 333 (Paederomimus; Type locality: Ins. St. Thomas).
— Bernhauer and Schubert, 1914: 368 (Paederomimus; catalog).
— Blackwelder, 1943: 398 (Paederomimus; synonym of lustralis).
lustrator Sharp, 1876: 172 (Philonthus; Type locality: Para).
— Sharp, 1885: 438 (Paederomimus).
— Bernhauer and Schubert, 1914: 368 (Paederomimus; catalog).
▴ Distribution: Brazil.
maculipes Sharp, 1885: 445 (Paederomimus; Type locality: Mexico, Cordova, Jalapa).
mandibularis Sharp, 1885: 439 (Paederomimus; Type locality: Guatemala, Capetillo).
mexicanus Sharp, 1885: 444 (Paederomimus; Type locality: Mexico, Jalapa).
nigrinus Sharp, 1885: 442 (Paederomimus; Type locality: Nicaragua, Chontales).
nigripennis Sharp, 1885: 446 (Paederomimus; Type locality: Panama, Bugaba 800 to 1500 feet, Volcan de Chiriqui from 2000 to 4000 feet).
nigrocyaneus Schubert, 1909: 294 (Paederomimus; Type locality: südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d'Orizaba).
nobilis Sharp, 1885: 442 (Paederomimus; Type locality: Panama, Volcan de Chiriqui 4000 to 6000 feet).
ornatellus Sharp, 1885: 449 (Paederomimus; Type locality: Panama, Bugaba 800 to 1500 feet).
pallidus Bierig, 1937: 197 (Paederomimus; Type locality: Panama: France Field, Zona del Canal).
palpalis Sharp, 1876: 173 (Philonthus; Type locality: Amazons).
— Sharp, 1885: 438 (Paederomimus).
— Bernhauer and Schubert, 1914: 368 (Paederomimus; catalog).
▴ Distribution: Brazil.
pectoralis Sharp, 1885: 443 (Paederomimus; Type locality: Guatemala, near the city, Capetillo, El Tumbador 2500 feet, Cerro Zunil 4000 to 5000 feet).
perforatus Sharp, 1885: 451 (Paederomimus; Type locality: Panama, Volcan de Chiriqui 2000 to 3000 feet).
praenobilis Bernhauer, 1910b: 379 (Paederomimus; Type locality: Mexiko: Vera Cruz, Cordoba).
propinquus Sharp, 1876: 176 (Philonthus; Type locality: Ega).
— Sharp, 1885: 438 (Paederomimus).
— Bernhauer and Schubert, 1914: 368 (Paederomimus; catalog).
▴ Distribution: Brazil.
punctipennis Sharp, 1885: 443 (Paederomimus; Type locality: Panama, Bugaba 800 to 1500 feet).
regillus Sharp, 1876: 176 (Philonthus; Type locality: St. Paulo).
— Sharp, 1885: 438 (Paederomimus; cited as regellus).
— Bernhauer and Schubert, 1914: 368 (Paederomimus; catalog).
▴ Distribution: Brazil.
salvini Sharp, 1885: 440 (Paederomimus; Type locality: Panama, Bugaba 800 to 1500 feet, Volcan de Chiriqui 2000 to 3000 feet).
sharpi Bernhauer and Schubert, 1914: 368 (Paederomimus; replacement name for hilaris Sharp).
hilaris Sharp, 1885: 441 (Paederomimus; [preoccupied]; Type locality: Panama, Volcan de Chiriqui 3000 to 4000 feet).
simplex Sharp, 1885: 444 (Paederomimus; Type locality: Guatemala, near the city, San Geronimo, San Joaquin in Vera Paz).
smithi Cameron, 1922, see: interjectus Bernhauer, 1908.
sulciceps Bernhauer, 1917d: 35 (Paederomimus; Type locality: Peru: Chanchamayo).
sydowi Bernhauer, 1917d: 37 (Paederomimus; Type locality: Brasilien: S. Paulo, Umgebung von Ribeirao Preto Fundao).
tetricus Blackwelder, 1943: 395 (Paederomimus; Type locality: Trinidad: Port of Spain).
traili Sharp, 1876: 170 (Philonthus; Type locality: Ananá).
— Sharp, 1885: 438 (Paederomimus).
— Bernhauer and Schubert, 1914: 368 (Paederomimus; catalog).
▴ Distribution: Brazil.
trinidadensis Bernhauer, 1915, see: cribricollis Erichson, 1840.
vicinus Sharp, 1885: 444 (Paederomimus; Type locality: Panama, Volcan de Chiriqui 2000 to 3000 feet).
Paracraspedomerus
[1 species; Oceanic Region]
Paracraspedomerus Moore, 1960: 99 (species included: speculifrons). Type species: Cafius speculifrons Fauvel, fixed by original designation and monotypy.
Xystridiogaster Scheerpeltz, 1966e: 402 (species included: speculifrons). Type species: Cafius speculifrons Fauvel, fixed by original designation (p. 405).
speculifrons Fauvel, 1877: 259 (Cafius; Type locality: Nouvelle Calédonie).
— Fauvel, 1889b: 263 (Cafius; notes; New Caledonia).
— Fauvel, 1903d: 269 (Hesperus; New Caledonia).
— Bernhauer and Schubert, 1914: 364 (Hesperus; catalog).
— Scheerpeltz, 1933: 1374 (Hesperus; catalog).
— Moore, 1960: 99 (Paracraspedomerus; New Caledonia).
— Scheerpeltz, 1966e: 402 (Xystridiogaster; characters; New Caledonia).
— Legner and Moore, 1977: 176 (Paracraspedomerus; found under bark).
▴ Distribution: New Caledonia.
Pescolinus
[2 species; Neotropical Region]
Pescolinus Sharp, 1885: 453 (species included: palmatus). Type species: Pescolinus palmatus Sharp, fixed by monotypy.
— Bernhauer and Schubert, 1914: 369 (world catalog; 1 species).
— Bierig, 1937: 191 (characters).
— Blackwelder, 1944: 136 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 299 (type species: palmatus).
palmatus Sharp, 1885: 453 (Pescolinus; Type locality: Panama, Volcan de Chiriqui 4000 to 6000 feet).
schmidti Bierig, 1937: 193 (Pescolinus; Type locality: Costa Rica: Vara Blanca, 2000 m).
Peucoglyphus
[1 species; Oriental Region]
Peucoglyphus Bernhauer, 1926f: 317 (species included: corporaali). Type species: Peucoglyphus corporaali Bernhauer, fixed by monotypy.
— Scheerpeltz, 1933: 1371 (world catalog supplement).
— Blackwelder, 1952: 299 (type species: corporaali).
corporaali Bernhauer, 1926f: 318 (Peucoglyphus; Type locality: Buru).
Phileciton
[2 species; Neotropical Region]
Phileciton Wasmann, 1894: 211 (species included: badariottii). Type species: Phileciton badariottii Wasmann, fixed by monotypy.
— Bernhauer and Schubert, 1914: 326 (world catalog; 2 species).
— Blackwelder, 1944: 132 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 300 (type species: badariottii).
— Seevers, 1965: 328 (doubtfully distinct from Erichsonius and Neobisnius; myrmecophilus species; list of species).
badariottii Wasmann, 1894: 212 (Phileciton; Type locality: Sales. bei Lorena, Prov. S. Paulo).
— Bernhauer and Schubert, 1914: 326 (Phileciton; catalog).
— Seevers, 1965: 328 (Phileciton; notes; Brazil).
▴ Distribution: Brazil.
wasmanni Bernhauer, 1908d: 334 (Phileciton; Type locality: Brasilien: S. Paulo).
Philonthus
[1255 species (19 doubtful); Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions]
Philonthus Stephens, 1829a: 23 (species included: laminatus, aeneus, splendens, politus, decorus, atratus, fimetarius, marginatus, concinnus, obscurus, varians, simplex, aterrimus, opacus, nitens, agilis, sanguinolentus, bimaculatus, cor[r]uscus, micans, and punctus along with 25 unavailable names; [Note: There is some dispute concerning the authorship of Philonthus. The name was used in 1827 and again in 1829. Westwood (1827: 58) described the larva of “Philonthus politus?” and presented fairly detailed characters and illustrations for the species. However, since he was unsure that the larva was identified accurately, we can reject as unavailable that use of Philonthus (Articles 12.2.5). In 1829 Philonthus was published in separate catalogs by both Curtis and Stephens. Tottenham (1962: 153) wrote that W. D. Hincks had informed him “. . . that he has seen definite evidence that the relevant [for date of Philonthus] part of Curtis' work was published in June . . .” . Earlier in the same paragraph Tottenham stated an August, 1829 publication date for Stephens' work. According to a letter from N. D. Riley, a former Keeper at the British Museum (Natural History), sent to J. G. Franclemont in 1949 there is evidence to support a June 1, 1829 publication of Stephens' “Nomenclature . . .” Unfortunately neither Tottenham nor Riley documented their opinions. Smetana (1995e: 51, 56) concluded no dates had been established and fixed the date of publication for both as December 31, 1829. According to evidence presented by D. Fletcher, 1979: xv, the date of publication was June 1, 1829 (see note attached to the citation for Stephens, 1829a in bibliography)]). Type species: Staphylinus splendens Fabricius, fixed by subsequent designation by Curtis, 1836: pl. 610.
— Curtis, 1829: 25 (catalog; Britain).
— Stephens, 1829: 279 (catalog; Britain).
— Stephens, 1832: 200, 226 (characters).
— Curtis, 1836: pl. 610 (characters; type species; Britain).
— Westwood, 1838a: 16 (characters; type species).
— Erichson, 1839a: 445 (characters).
— Stephens, 1839: 385 (characters).
— Heer, 1839: 257 (characters).
— Erichson, 1840: 426 (characters).
— Schaum, 1852: 26 (list of species; Europe).
— Lacordaire, 1854: 80 (characters; notes; list of species).
— Wollaston, 1854: 580 (characters; notes).
— Fairmaire and Laboulbène, 1856: 513 (characters).
— Jacquelin du Val, 1856: 35 (characters).
— Giebel, 1856: 72 (fossil species).
— Gistel, 1856: 388 (list of species).
— Kraatz, 1857d: 565 (characters).
— Redtenbacher, 1857: 188 (characters; key to Austrian species).
— Thomson, 1858: 30 (characters).
— G. Waterhouse, 1858: 24 (catalog of British species).
— Thomson, 1859: 24 (characters; type species: splendens).
— Schaum, 1859: 26 (catalog; European species).
— Thomson, 1860: 153 (characters).
— LeConte, 1861a: 65 (characters).
— Gredler, 1863: 104 (list of species from Tirol).
— LeConte, 1863: 24 (list of North American species).
— Fauvel, 1866a: 341 [= 1866b: 92] (characters).
— Wencker and Silbermann, 1866: 29 (list of species; collecting notes; France).
— Gemminger and Harold, 1868: 584 (catalog).
— Jacquelin du Val, 1868: 68 (catalog; European species).
— Sharp, 1871d: 11 (list of British species).
— Fauvel, 1874: 207 [= 1874a: 433] (characters; key to species; larval characters).
— Redtenbacher, 1874: 207 (characters; key to Austrian species).
— Oustalet, 1874: 167 (characters; description of fossil species).
— Fauvel, 1875a: XXIX [= 1875b: 231] (catalog).
— Mulsant and Rey, 1876: 344 [= 1877: 200] (characters; key to French species).
— Sharp, 1876: 165 (characters; notes).
— Fauvel, 1877: 262 (notes; key to species).
— Provancher, 1877: 248, 730 (characters; key to species).
— Fauvel, 1878f: 257 (characters; notes; key to New Guinea species).
— Broun, 1880: 110 (characters).
— Heyden, 1880: 73 (list of species of Siberian region).
— Duvivier, 1883: 144 (catalog).
— Lynch, 1884: 146 (characters; key for Argentine species).
— Horn, 1884: 179 (characters; key to North American species).
— Sharp, 1885: 395 (characters; notes; Mexican and Central American species).
— Fowler, 1888: 258 (notes; larval characters; key to British species).
— Seidlitz, 1891: 382 (key to some European species).
— Seidlitz, 1891a: 407 (key to some European species).
— Heyden, Reitter, and Weise, 1891: 103 (list of species of Europe and Caucasus).
— Heyden, 1893: 42 (list of species of Siberian region).
— Ganglbauer, 1895: 437 (adult and larval characters; key to middle European species).
— Heyden, 1896: 32 (list of species of Siberian region).
— Fauvel, 1897d: 299 (catalog of species of Barbary; Azores; Salvages; Madeira; Canaries).
— Germain, 1903: 414 (characters; notes).
— Fauvel, 1903d: 270 (key to New Caledonia species).
— Heyden, Reitter, and Weise, 1906: 161 (list of species of Europe and Caucasus).
— Reitter, 1909: 116 (characters).
— Blatchley, 1910: 379 (characters; key to Indiana species).
— Petri, 1912: 71 (list of species of Siebenbürgen).
— Bernhauer and Schubert, 1914: 326 (world catalog; 574 species).
— Johansen, 1914: 388 (characters; key to Danish species).
— Leng, 1920: 106 (catalog of North American species).
— Cameron, 1921a: 354, 405 (characters; key to Singapore species; catalog).
— Gridelli, 1923: 24 (key to Mediterranean region species).
— Winkler, 1925: 374 (catalog for Palaearctic region).
— Cameron, 1925: 54 (catalog of Indian species).
— Porta, 1926: 100 (characters; key to species of Italy).
— Bruch, 1928a: 427 (catalog; Argentina).
— Portevin, 1929: 356 (adult and larval characters; key to French species).
— Roubal, 1930: 367 (catalog; Slovakia).
— Cameron, 1932: 62 (characters; key to species of British India).
— Joy, 1932: 117 (key to British species).
— Scheerpeltz, 1933: 1329 (world catalog supplement).
— Normand, 1935: 370 (list of species of Tunisia).
— Tottenham, 1937a: 190 (discussion of some of Stephens' types specimens).
— Paulian, 1941: 223 (larval characters).
— Blackwelder, 1943: 398 (characters; type species: laminatus; key to species of West Indies).
— Blackwelder, 1944: 132 (checklist of species from Mexico, Central and South America, and the West Indies).
— Jeannel and Paulian, 1945: 79 (characters; notes; type species: politus).
— Kloet and Hincks, 1945: 165 (list of British species).
— Tottenham, 1949b (key to some African species).
— Jeannel and Jarrige, 1949: 374 (notes; cave species).
— Tottenham, 1949: 372, 417 (type species: splendens Fabricius; notes; checklist of British species).
— V. Hansen, 1952: 35 (characters; key to Danish species).
— Blackwelder, 1952: 301 (type species).
— Hatch, 1957: 173, 179, 205 (characters; key to species of Pacific Northwest; aedeagal key).
— Kocher, 1958: 144 (checklist of species; Morocco).
— Smetana, 1958: 143 (characters; habitus of one species; type species: splendens; key to Czechoslovakian species).
— Smetana, 1959 (key to species of central Europe).
— Smetana, 1959a: 209 (list of species collected in Albania).
— Tottenham, 1962: 153, 159 (author of genus is Curtis not Stephens because Curtis published his article in June and Stephens in August [Note: See my bibliographic citations for Curtis, 1829 and Stephens, 1829a]; key to various African species groups).
— Ferreira, 1962a: 17 (list of Portuguese species).
— Coiffait, 1963: 8, 14 (characters; key to French subgenera and species groups; list of French species included in species groups).
— Székessy, 1963: 56 (characters).
— Palm, 1963b: 61 (characters; key to species of Sweden).
— Lohse, 1964: 174 (characters; key to central European subgenera and species).
— Kerstens, 1964: 12 (notes on sordidus group).
— Moore, 1965: 97, 99 (characters; notes).
— Pototskaia, 1966: 152, 156 (larval characters; key to some species using larval characters).
— Coiffait, 1967a (key to subgenera, species groups, and species of western Palaearctic).
— Pototskaia, 1967: 52 (larval characters).
— Last, 1968: 373 (key to some species of New Guinea).
— Coiffait and Saiz, 1968: 348, 350 (characters; notes; key to Chilean species).
— Scheerpeltz, 1968a: 65 (catalog of Austrian species).
— Bolov, 1969a: 328 (collecting notes for some species; Russia).
— Kasule, 1970: 64, 65 (larval characters).
— Shibata, 1973c: 57 (checklist of species of Taiwan).
— Bordoni, 1973j: 703 (list of species from islands near Sicily).
— Tikhomirova, 1973a: 183 (checklist of species of USSR).
— Coiffait, 1974: 116 (adult and larval characters; key to subgenera, species groups, and species of western Palaearctic region).
— Moore and Legner, 1975: 22 (catalog; North America north of Mexico).
— Moore and Legner, 1976: 541 (includes seashore species; notes).
— Pope, 1977: 29 (list of British species).
— Moore and Miller, 1978: 38 (two unnamed species from Pleistocene asphalt deposits; California).
— Topp, 1978: 312 (larval characters).
— E. Kangas, 1979: 101 (key to species of varians group).
— Moore and Legner, 1979: 28 (characters; notes).
— Muona, 1979: 14 (list of Scandinavian species).
— Uhlig, 1979: 252 (some species collected in Germany).
— M. Dvořák, 1979: 115 (some species collected in Slovakia).
— Uhlig, Vogel, and Sieber, 1980: 246 (some species collected in Germany).
— Burakowski, Mroczkowski, and Stefańska, 1980: 38 (catalog; Poland).
— Uhlig and Vogler, 1981: 111 (list of some species collected in Germany; habitat and collecting notes).
— Spahr, 1981: 96 (references to specimens from amber and copal).
— Boller, 1983 (larval characters; larval key to some species).
— Dettner, 1983 (comparative anatomy of defensive glands and defensive secretions).
— Shibata, 1983: 90 (checklist of species of Japan).
— Hammond, 1984: 206 (checklist of species of Borneo).
— Tóth, 1984: 41 (characters; habitus; key to species of Hungary).
— Yuh, Paik, Kwon, and Lee, 1985: 240 (checklist of species of Korea).
— Frank, 1986a: 370 (checklist of species of Florida).
— Muona and Viramo, 1986: 13 (list of species of northeastern Finland).
— Segers, 1986: 43 (checklist of species; Belgium).
— Drugmand 1987b (comments on distribution of Belgian species).
— Ádám, 1987: 149 (list of 33 species with collecting notes; Hungary).
— Lohse and Lucht, 1989: 163 (supplement to key to species of Lohse, 1964).
— Lecoq, 1990: 202 (key to species of Mascarene Islands).
— Angelini, 1991: 192 (list of species from high elevations of Calabria, Italy).
— M. Hansen, Kristensen, Mahler, Pedersen, 1991: 107 (list of Danish species).
— Smetana, 1991a: 227 (key to species of furvus complex of Mexico and Central America).
— Campbell and Davies, 1991: 117 (checklist of species; Alaska and Canada).
— Silfverberg, 1992: 16 (list of species of Fennoscandia, Denmark, and the Baltic States).
— Navarrete-Heredia and Márquez-Luna, 1993 (cave collections; Mexico).
— Siitonen, 1993: 230 (list of Finnish species).
— Hayashi, 1993: 81 (key to species of prolatus group).
— Burakowski, 1993: 22, 23 (rearing techniques).
— Welch, 1993: 228 (ovariole structure and development).
— Smetana, 1993b: 48 (list of species of Czech Republic and Slovakia).
— Hayashi, 1994: 120, 126 (notes on classification of Philonthus and related genera; diagnostic characters; subgeneric characters).
— Hernández, Outerelo, and Gamarra, 1994: 188 (checklist of species; Canary Islands).
— Smetana, 1995e: 51 (characters; classification and revision of and key to species of USA and Canada; phylogeny of species groups; author of genus is Stephens).
— Hu and Frank, 1995b (surface structure of eggs of five species and key for identification of eggs [p.587]).
— Navarrete-Heredia, 1995: 96 (list of species of Mexico).
— M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 28 (additional records to Danish list of species).
— Ciceroni and Zanetti, 1995: 27 (list of species of Italy).
— M. Hansen, 1996: 104 (list of Danish species).
— M. Hansen, Mahler, Palm, and Pedersen, 1996: 243 (additions to list of Danish species).
— Lauterer and Opravil, 1996: 244 (subfossil; 16th century; Czech Republic).
— Downie and Arnett, 1996: 375 (characters; key to species of Northeastern North America).
— Lecoq, 1997: 280 (key to “yellow” species of Madagascar).
— Schillhammer, 1998 (revision of and key to species of cyanipennis group; notes).
— Lundgren, 1998: 51 (list of species from Florida; records for other States listed for each species).
— M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (list of species collected in 1997; Denmark).
— Assing, Frisch, Kahlen, et al., 1998: 137 (supplement to key to species in Lohse, 1964; notes).
— Schillhammer, 1999: 57 (key to species of spinipes and cinctulus species groups).
— Assing and Schülke, 1999: 14 (authorship of Philonthus; notes; synonymic discussion).
— Schillhammer, 2000: 113 (revision of and key to species of politus complex; species group listing; correction on page 159 of scale bars in Schillhammer, 1999).
— Newton, Thayer, Ashe, and Chandler, 2000: 396 (112 Nearctic species; notes; characters in key).
— Hayashi, 2000: 23 (notes on empodial setae).
— Naomi, Kuranishi, Saito, and Maruyama, 2000: 109 (collected from Kuril Islands and Kamchatka).
— Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 62 (necrophilous species in Mexico).
Laxobates Gistel, 1834: 8 (species included: coenosus, cyanipennis, splendens, laminatus, aeneus, metallicus, cephalotes, decorus, politus, atratus, carbonarius, ebeninus cited as ebeneninus, varians, varius, immundus, fuscus, sanguinolentus, tenuis, fimetarius, agilis, splendidulus along with 12 unavailable names). Type species: Staphylinus splendens Fabricius, fixed by subsequent designation by Blackwelder, 1952: 212.
Eccoptolonthus Bernhauer, 1912f: 206 (subgenus of Philonthus; species included: conradti). Type species: Philonthus (Eccoptolanthus) conradti Bernhauer, fixed by monotypy.
— Bernhauer and Schubert, 1914: 326 (synonym of Philonthus).
— Cameron, 1932: 62 (subgenus of Philonthus).
— Blackwelder, 1943: 399 (subgenus of Philonthus; type species: conradti).
— Blackwelder, 1952: 138 (subgenus of Philonthus; type species: conradti).
Pseudophilonthus Bernhauer, 1915k: 302 (subgenus of Philonthus; species included: bicoloripennis). Type species: Philonthus (Pseudophilonthus) bicoloripennis Bernhauer, fixed by monotypy.
— Bernhauer, 1932b: 147 (valid genus).
— Scheerpeltz, 1933: 1330 (subgenus of Philonthus).
— Blackwelder, 1943: 399 (subgenus of Philonthus; type species: bicoloripennis).
— Blackwelder, 1952: 329 (subgenus of Philonthus; type species: bicoloripennis).
Onychophilonthus Neresheimer and Wagner, 1924: 156 (subgenus of Philonthus; species included: marginatus). Type species: Staphylinus marginatus Ström, fixed by monotypy.
— Scheerpeltz, 1933: 1330 (subgenus of Philonthus).
— Blackwelder, 1943: 399 (subgenus of Philonthus; type species: marginatus).
— Tottenham, 1949: 372, 419 (type species: marginatus Ström; subgenus of Philonthus; checklist of British species).
— Blackwelder, 1952: 276 (subgenus of Philonthus; type species: marginatus).
— Smetana, 1958: 143, 226 (subgenus of Philonthus; characters; type species).
— Smetana, 1959: 170 (subgenus of Philonthus; characters).
— Smetana, 1963d: 402 (subgenus of Philonthus; characters; key to species).
— Coiffait, 1974: 116, 120, 190, 305 (subgenus of Philonthus; characters; key to species of western Palaearctic region).
Nephronthus Bernhauer, 1932b: 147 (subgenus of Pseudophilonthus; species included: grandis). Type species: Pseudophilonthus (Nephronthus) grandis Bernhauer, fixed by monotypy.
— Scheerpeltz, 1933: 1330 (subgenus of Philonthus).
— Blackwelder, 1943: 399 (subgenus of Philonthus; type species: grandis).
— Blackwelder, 1952: 261 (subgenus of Philonthus; type species: grandis).
Philonthopsis Cameron, 1932: 261 (species included: antennalis). Type species: Philonthopsis antennalis Cameron, 1932, fixed by monotypy.
— Scheerpeltz, 1933: 1418 (world catalog supplement).
— Blackwelder, 1952: 301 (type species: sabulosus).
— Scheerpeltz, 1965: 279 (characters; type species: antennalis).
— Schillhammer, 1994: 74 (synonym of Philonthus).
Cephalonthus Blackwelder, 1952: 96 (subgenus of Philonthus; for species included see Bernhauer, 1940b). Type species: Philonthus caffer Boheman, fixed by original designation (see Herman, 2001).
— Bernhauer, 1940b: 635 (subgenus of Philonthus; species included: lewisius, caffer, ustus, kochianus; characters; [Note: This name is unavailable because Bernhauer did not designate a type species]).
— Herman, 2001: 9 (name unavailable when proposed by Bernhauer, 1940, made available by Blackwelder, 1952; Blackwelder author of name, not Bernhauer).
Philonthoblerius Tottenham, 1949b: 300 (subgenus of Philonthus; species included: schoutedeni). Type species: Philonthus schoutedeni Bernhauer, fixed by original designation and monotypy.
Raucalius Tottenham, 1949b: 304 (subgenus of Philonthus; species included: rugulipennis, peripateticus, madianus). Type species: Philonthus (Raucalius) peripateticus Tottenham, fixed by original designation.
— Blackwelder, 1952: 337 (subgenus of Philonthus; type species: peripateticus).
— Levasseur, 1967b: 1624 (notes; characters).
Spatulonthus Tottenham, 1955: 154 (subgenus of Philonthus; species included: longicornis, kasaiensis, vestigator, ugandae, bestialis, bisignatus, deleterius, delusor, ridens, minutus, sanamus, nitidicollis, incognitus, peregrinus, igacus, labdanus, maskinius, ponderosus, tachyoryctides, limulus, bishanus). Type species: Philonthus longicornis Stephens, fixed by original designation.
— Tottenham, 1955: 155 (key to species).
— Coiffait, 1963: 8, 18 (characters; notes).
— Coiffait and Saiz, 1968: 348, 360 (characters; notes).
— Bordoni, 1973j: 707 (list of species from islands near Sicily).
— Coiffait, 1974: 308 (adult and larval characters; key to species of western Palaearctic region).
— Burakowski, Mroczkowski, and Stefańska, 1980: 36 (catalog; Poland).
— Segers, 1986: 46 (checklist of species; Belgium).
— Drugmand 1987b (comments on distribution of Belgian species).
— Ádám, 1987: 151 (list of three species with collecting notes; Hungary).
— Silfverberg, 1992: 17 (list of species of Fennoscandia, Denmark, and the Baltic States).
— Smetana, 1995e: 51 (synonym of Philonthus).
Kenonthus Coiffait, 1960: 361 (subgenus of Philonthus; species included: montivagus, aerosus, laevicollis, nimbicola). Type species: Philonthus montivagus Heer, fixed by original designation.
— Coiffait, 1974: 116, 120, 186, 298 (subgenus of Philonthus; characters; key to species of western Palaearctic region).
Trionthus Coiffait, 1963: 14 (subgenus of Philonthus; species included: lepidus). Type species: Staphylinus lepidus Gravenhorst, fixed by monotypy.
— Coiffait, 1974: 116, 120, 193 (subgenus of Philonthus; characters; key to species of western Palaearctic region).
— Ádám, 1987: 151 (cited as valid genus; list of one species with collecting notes; Hungary).
— Smetana, 1995e: 51 (synonym of Philonthus).
Paragabrius Coiffait, 1963: 12 (species included: micans, micanthoides, furcifer, palustris fulvipes, virgo, fagniezi, truquii, mimus). Type species: Staphylinus micans Gravenhorst, fixed by original designation.
— Coiffait, 1974: 84 (characters; key to species of western Palaearctic region).
— Pace and Zanetti, 1977: 301 (characters; notes; relationships).
— Burakowski, Mroczkowski, and Stefańska, 1980: 34 (catalog; Poland).
— Segers, 1986: 43 (checklist of species; Belgium).
— Drugmand 1987b (comments on distribution of Belgian species).
— Ádám, 1987: 151 (cited as valid genus; list of three species with collecting notes; Hungary).
— Angelini, 1991: 192 (list of species from high elevations of Calabria, Italy).
— Silfverberg, 1992: 16 (list of species of Fennoscandia, Denmark, and the Baltic States).
— Smetana, 1995e: 51 (synonym of Philonthus).
Kirschenblatia Bolov and Kryzhanovskij, 1969: 512 (species included: kabardensis). Type species: Kirschenblatia kabardensis Bolov and Kryzhanovskij, fixed by original designation.
— Bolov and Kryzhanovskij, 1969a: 327 (translation of Bolov and Kryzhanovskij, 1969).
— Schülke and Uhlig, 1989: 247 (synonym of Philonthus).
Palaeophilonthus Coiffait, 1972: 50 (proposed for Coiffait's [1965g: 616] misidentification of Sectophilonthus; species included: rossicus). Type species: Sectophilonthus rossicus Coiffait, fixed by monotypy.
— Coiffait, 1965g: 616 (characters; type species; [Note: Misidentified as Sectophilonthus according to Coiffait, 1972: 50]).
— Coiffait, 1974: 5 (characters; species of western Palaearctic region).
— Smetana, 1995e: 51 (synonym of Philonthus).
Metagabrius Coiffait, 1974: 84, 86 (subgenus of Paragabrius; species included: juvenilis, fuscatus). Type species: Philonthus juvenilis Peyerimhoff, fixed by original designation.
Parambigus Marcuzzi, 1977: 12 (subgenus of Philonthus; without description; [Note: Nomen nudum, Article 13.1.1]; species included: lucidus Fauvel).
Glaphyronthus Scheerpeltz, 1978: 220 (subgenus of Philonthus; species included: tenellicornis). Type species: Philonthus (Glaphyronthus) tenellicornis Scheerpeltz, fixed by original designation and monotypy.
Paralionthus Ádám, 1996: 236 (species included: punctus). Type species: Staphylinus punctus Gravenhorst, fixed by original designation and monotypy.
Piezarthrus Lecoq, 1998: 63 (subgenus of Philonthus; species included: pollux, steineri, pauliani, viettei, marojejy, humbloti, castor, rectus). Type species: Philonthus pollux Fauvel, fixed by original designation.
abactus Sharp, 1876, see: Paederomimus.
abauriae Gridelli, 1924, see: Rabigus.
abavus† Scudder, 1900: 57 (Philonthus; fossil; Type locality: Florissant, Colorado).
abdicans Tottenham, 1949b: 313 (Philonthus; Type locality: South Africa).
abdominalis Erichson, 1847: 89 (Philonthus; Type locality: Republica Peruana).
aberdarensis Bernhauer, 1939, see: Gabrius.
aberrans Sharp, 1876, see: Paederomimus.
aberrans Cameron, 1932: 111 (Philonthus; [preoccupied]; Type locality: Mussoorie district; Mossy Falls; Keyarkuli. Assam: Naga Hills, Laimatak).
— Scheerpeltz, 1933: 1330 (Philonthus; catalog).
— Herman, 2001: 46 (Philonthus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: India.
aberti Hatch, 1957: 191 (Philonthus; Type locality: Oregon: L. Abert).
abkhazensis Coiffait, 1963, see: rufipes Stephens, 1832.
abnormalis Sharp, 1885: 397 (Philonthus; Type locality: Mexico, Oaxaca).
— Bernhauer and Schubert, 1914: 326 (Philonthus; catalog).
— Navarrete-Heredia, 1995: 96 (Philonthus; Mexico; Guatemala).
▴ Distribution: Guatemala, Mexico.
abyssinus Fauvel, 1881a: 203 (Belonuchus; Type locality: Scioa, Mahal-Uonz; Let Marefia).
— Fauvel, 1907: 39 (Philonthus; Naivasha; Nairobi; Kilimandjaro; Kiboscho; Abyssinia; Gallas; Choa; Usambara; Mozambique; Mashunaland; Cape; Angola).
— Eichelbaum, 1913: 130 (Philonthus; Tanzania).
— Bernhauer and Schubert, 1914: 326 (Philonthus; catalog).
— Rambousek, 1922: 1008 (Philonthus; Ethiopia).
— Scheerpeltz, 1933: 1330 (Philonthus; catalog).
— Tottenham, 1949b: 315 (Philonthus).
— Cameron, 1950g: 38 (Philonthus; Belgian Congo).
— Tottenham, 1962: 164 (Philonthus; characters).
— Frank, 1982: 38 (Philonthus; list of parasites).
▴ Distribution: Ethiopia, Kenya, Congo, Tanzania, Mozambique, Zimbabwe, South Africa, Angola.
holomelas Eppelsheim, 1895: 201 (Philonthus; Type locality: Ganale Gudda, Arussi Galla).
— Eppelsheim, 1895: 213 (Philonthus; synonym of abyssinus).
— Fauvel, 1907: 39 (Philonthus; synonym of abyssinus).
— Bernhauer and Schubert, 1914: 327 (Philonthus; synonym of abyssinus).
acamas Smetana, 1995e: 281 (Philonthus; Type locality: Oregon: Austin House, Jct. Hwys. 26 & 7, 1005 m. Also cited from California, Colorado, Nevada, Oregon).
accedens Sharp, 1885: 403 (Philonthus; Type locality: Guatemala, Capetillo).
— Bernhauer and Schubert, 1914: 327 (Philonthus; catalog).
— Frank, 1982: 38 (Philonthus; list of parasites).
▴ Distribution: Guatemala.
acceptus Last, 1973: 429 (Philonthus; Type locality: N.E. New Guinea: Karkar Island).
aciculatus Stephens, 1832, see: sanguinolentus Gravenhorst, 1802.
acroleucus Kraatz, 1859: 91 (Philonthus; Type locality: Ceylan; India orientalis).
— Bernhauer, 1902b: 27 (Philonthus; Ceylon).
— Bernhauer and Schubert, 1914: 327 (Philonthus; catalog).
— Cameron, 1932: 115 (Philonthus; characters; Ceylon).
— Scheerpeltz, 1933: 1330 (Philonthus; catalog; Philippines).
— Cameron, 1933d: 387 (Philonthus; Sumatra).
— Cameron, 1937: 5 (Philonthus; Java).
— Frank, 1982: 38 (Philonthus; list of parasites).
▴ Distribution: Sri Lanka, “East Indies”, Indonesia, Philippines.
acromyrmecis Scheerpeltz, 1976b: 135 (Philonthus; subgenus Philonthus; Type locality: Kolonie Nova Teutonia, Brasilien).
actinus Bernhauer, 1917i: 103 (Philonthus; Type locality: Bolivien: Yuracaris).
activus Cameron, 1937: 8 (Philonthus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft).
actocharis Sharp, 1887: 788 (Philonthus; Type locality: Mexico, Tehuantepec).
— Bernhauer and Schubert, 1914: 327 (Philonthus; catalog).
— Navarrete-Heredia, 1995: 96 (Philonthus; Mexico).
▴ Distribution: Mexico.
aculeatus Coiffait, 1963: 28 (Philonthus; Type locality: Liban: environ de Beyrouth).
— Coiffait, 1967a: 415 (Philonthus; characters; Lebanon).
— Coiffait, 1974: 265 (Philonthus; characters; Lebanon).
— Raichev, 1979: 83 (Philonthus; Bulgaria).
▴ Distribution: Lebanon, Bulgaria.
acuminatus Kraatz, 1859: 93 (Philonthus; Type locality: Ceylan).
— Bernhauer and Schubert, 1914: 327 (Philonthus; catalog).
— Cameron, 1932: 142 (Philonthus; characters; Ceylon).
— Scheerpeltz, 1933: 1330 (Philonthus; catalog).
▴ Distribution: Sri Lanka.
addendus Sharp, 1867: 440 (Philonthus; Type locality: Corstorphine hills near Edinburgh).
— Rye, 1863: 82 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 327 cited temporalis Rye as a synonym of addendus; Rye did not describe the species, he attributed it to Mulsant]; characters).
— Fauvel, 1874: 217 [= 1874a: 443] (Philonthus; characters; France; Lake Baikal; Amur region).
— Mulsant and Rey, 1876: 376 [= 1877: 232] (Philonthus; characters; France).
— J. Sahlberg, 1876: 13 (Philonthus; characters; Finland).
— Sharp, 1889: 35 (Philonthus; notes; Japan).
— Fowler, 1888: 263 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 445 (Philonthus; characters; north and middle Europe; east Siberia).
— Poppius, 1905: 9 (Philonthus; Orkney Islands).
— Reitter, 1909: 125 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 327 (Philonthus; catalog).
— Johansen, 1914: 395 (Philonthus; characters; Denmark).
— Portevin, 1929: 358 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1330 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— V. Hansen, 1952: 51 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1955b: 228 (Philonthus; notes).
— Smetana, 1958: 165 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 147 (Philonthus; characters).
— Smetana, 1960b: 259 (Philonthus; Czechoslovakia).
— Palm, 1963b: 67 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 176 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 127 (Philonthus; Russia; Finland; Sweden; Norway; Britain; Denmark; Belgium; France; Switzerland; Italy; Bulgaria; Romania; Czech Republic; Slovakia; Poland; Austria; Germany).
— Coiffait, 1967a: 392 (Philonthus; characters).
— Raianu, 1968a: 462 (Philonthus; Romania).
— Szujecki, 1968a: 727 (Philonthus; Poland).
— Szujecki, 1969a: 255 (Philonthus; notes; Poland).
— Coiffait, 1974: 223 (Philonthus; characters; Europe; Britain; Siberia; Japan).
— Shilov, 1975a: 375 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Pope, 1977: 29 (Philonthus; Britain).
— Raichev, 1979: 82 (Philonthus; Bulgaria).
— Tóth, 1984: 48 (Philonthus; characters; Hungary).
— Yuh, Paik, Kwon, and Lee, 1985: 240 (Philonthus; Korea).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Schülke and Uhlig, 1988: 11 (Philonthus; notes; Germany).
— J. Li, 1993: 36 (Philonthus; China).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Hayashi, 1994: 124 (Philonthus; characters; Japan).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; Italy).
— Sterrenburg and Schülke, 1997: 18 (Philonthus; Netherlands).
— Owen, 1997a: 149 (Philonthus; collecting sites; Ireland).
— Schillhammer, 1999a: 61 (Philonthus).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
— Schillhammer, 2000: 134 (Philonthus; characters; Russia; Mongolia; North Korea; Japan; China).
▴ Distribution: Europe, Russia, Kazakhstan, Mongolia, Korea, China, Japan.
temporalisoides Drugmand, 1988b: 249 (Philonthus; subgenus Philonthus; Type locality: U.R.S.S.: lac Baikal, Sibérie).
— Schillhammer, 1999a: 61 (Philonthus; synonym of addendus).
— Schillhammer, 2000: 134 (Philonthus; synonym of addendus).
adjacens Cameron, 1937: 7 (Philonthus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft).
adscitus Märkel and Kiesenwetter, 1848, see: laevicollis Lacordaire, 1835.
adustus Casey, 1915, see: Gabrius.
adversarius Bernhauer, 1917g: 46 (Philonthus; replacement name for kamerunensis Bernhauer, 1915).
kamerunensis Bernhauer, 1915k: 301 (Philonthus; [preoccupied]; Type locality: Kamerun).
adversus Bernhauer and Schubert, 1914: 327 (Philonthus; replacement name for misellus Fauvel).
— Cameron, 1932: 100 (Philonthus; characters; Burma; Assam; Darjeeling).
— Scheerpeltz, 1933: 1330 (Philonthus; catalog).
▴ Distribution: Myanmar, India.
misellus Fauvel, 1895: 262 (Philonthus; [preoccupied]; Type locality: Bírmanie, Carin Cheba, 900–1100 m).
— Bernhauer and Schubert, 1914: 327 (Philonthus; synonym of adversus).
— Cameron, 1932: 100 (Philonthus; synonym of adversus).
adversus Cameron, 1925: 59 (Philonthus; [preoccupied]; replacement name for misellus Fauvel).
adversus Cameron, 1925, see: adversus Bernhauer and Schubert, 1914.
aeger Eppelsheim, 1895c: 59 (Philonthus; Type locality: Simla im Himalaya).
— Bernhauer and Schubert, 1914: 327 (Philonthus; catalog).
— Cameron, 1932: 83 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1330 (Philonthus; catalog).
— Coiffait, 1982a: 233 (Philonthus; Bhutan).
— Schillhammer, 2000: 140 (Philonthus; characters; India; Nepal; China).
▴ Distribution: China, Nepal, Bhutan, India.
aeolus Herman, 2001: 46 (Philonthus; replacement name for temporalis Coiffait).
temporalis Coiffait, 1977b: 221 (Paragabrius; [preoccupied]; Type locality: Bhutan: Dorjula, 2900 m).
— Smetana, 1995e: 51 (Philonthus; transferred from Paragabrius by reason of Smetana's synonymy of the genus with Philonthus).
— Herman, 2001: 46 (Philonthus; synonym of aeolus).
aemulus Tottenham, 1939d: 233 (Philonthus; replacement name for serraticornis Bernhauer).
— Tottenham, 1949b: 361 (Philonthus; characters).
— Tottenham, 1961: 205 (Philonthus; Senegal).
— Levasseur, 1966a: 1214 (Philonthus; Tanganyika; Senegal; Congo).
— Levasseur, 1968a: 54 (Philonthus; Central African Republic).
▴ Distribution: Namibia, Congo, Tanzania, Central African Republic, Senegal.
serraticornis Bernhauer, 1908c: 110 (Philonthus; [preoccupied]; Type locality: Deutsch-Südwestafrika: Otjosondu).
antennatus Bernhauer and Schubert, 1914: 328 (Philonthus; replacement name for serraticornis Bernhauer; [preoccupied]).
aeneiceps Sharp, 1876, see: Paederomimus.
aeneicollis Sharp, 1876, see: Paederomimus.
aeneipennis Boheman, 1858: 30 (Philonthus; Type locality: China).
— Fauvel, 1903: 159 (Philonthus; India).
— Fauvel, 1904a: 59 (Philonthus; Belgaum; Ceylan; Japan; Celebes; Maurice; China; New Guinea).
— Bernhauer, 1907: 386 (Philonthus; Japan).
— Bernhauer and Schubert, 1914: 327 (Philonthus; catalog).
— Bernhauer, 1922c: 232 (Philonthus; Formosa).
— Bernhauer, 1928: 16 (Philonthus; Sumatra).
— Cameron, 1932: 113 (Philonthus; characters; Ceylon; India).
— Scheerpeltz, 1933: 1330 (Philonthus; catalog).
— Cameron, 1933d: 386 (Philonthus; Sumatra).
— Cameron, 1937: 5 (Philonthus; Java).
— Cameron, 1949b: 467 (Philonthus; China).
— Scheerpeltz, 1957c: 233 (Philonthus; Sumba).
— Nakane and Sawada, 1960: A123 (Philonthus; Yakushima Island).
— Scheerpeltz, 1960: 5 (Philonthus; Pakistan; Iran; Afghanistan).
— Nakane, 1963a: 91 (Philonthus; characters; habitus photograph; Japan).
— Scheerpeltz, 1976a: 87 (Philonthus; Nepal).
— Watanabe and Shibata, 1972: 64 (Philonthus; Yaku-shima Island).
— Coiffait, 1977b: 206 (Philonthus; Bhutan).
— Coiffait, 1981f: 329 (Philonthus; Andaman Islands).
— Coiffait, 1982a: 232 (Philonthus; India).
— Frank, 1982: 38 (Philonthus; list of parasites).
— J. Li, 1993: 36 (Philonthus; China).
— Yadav, Chand, and Yadav, 1994: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
— Watanabe, 1998a: 140 (Philonthus; Japan).
— Watanabe, 1998b: 262 (Philonthus; Japan).
▴ Distribution: China, Japan, Taiwan, New Guinea, Indonesia, Andaman Islands, Bhutan, Nepal, India, Pakistan, Afghanistan, Iran, Yemen, Mauritius, Tanzania, Senegal.
cliens Eppelsheim, 1890b: 277 (Philonthus; Type locality: Nagpore, India centralis).
— Fauvel, 1903: 159 (Philonthus; variety of aeneipennis).
— Fauvel, 1904a: 59 (Philonthus; variety of aeneipennis; Khandesh; Kanara; Aden).
— Bernhauer and Schubert, 1914: 327 (Philonthus; variety of aeneipennis).
— Cameron, 1932: 113 (Philonthus; synonym of aeneipennis).
— Scheerpeltz, 1933: 1330 (Philonthus; variety of aeneipennis).
— Gridelli, 1936a: 151 (Philonthus; characters; valid species).
— Tottenham, 1961: 204 (Philonthus; Senegal).
— Tottenham, 1962: 139, 223, 226 (Philonthus; characters; Tanganyika).
— Scheerpeltz, 1976a: 87 (Philonthus; synonym of aeneipennis).
— Coiffait, 1982: 28 (Philonthus; subspecies of erythropus; Nepal).
— Coiffait, 1982a: 233 (Philonthus; variety of aeneipennis; Nepal).
— Coiffait, 1984e: 373 (Philonthus; variety of aeneipennis; Nepal).
punctatissimus Schubert, 1908: 619 (Philonthus; Type locality: Pathanka, Himalaya).
— Bernhauer and Schubert, 1914: 352 (Philonthus; catalog).
— Cameron, 1932: 113 (Philonthus; synonym of aeneipennis).
— Scheerpeltz, 1933: 1330 (Philonthus; synonym of aeneipennis).
lindemanni Scheerpeltz, 1960: 5 (Philonthus; variety of aeneipennis; Type locality: West Pakistan: Rawalpindi Umg.).
aeneipennis Cameron, 1932, see: distincticornis Cameron, 1932.
aeneoceps Broun, 1880: 111 (Philonthus; Type locality: Whangarei Harbour).
aenescens Solsky, 1870a: 262 (Philonthus; Type locality: Mexico).
— Sharp, 1885: 408 (Philonthus; Mexico).
— Bernhauer and Schubert, 1914: 327 (Philonthus; catalog).
▴ Distribution: Mexico.
aeneus De Geer, 1774, see: laminatus Creutzer, 1799.
aeneus Rossi, 1790, see: politus Linné, 1758.
aequalis Horn, 1884: 205 (Philonthus; Type locality: Canada, Lake Superior region; [Note: See lectotype designation by Smetana, 1995e]).
— Bernhauer and Schubert, 1914: 327 (Philonthus; catalog).
— Smetana, 1963a: 4 (Philonthus; characters; Montreal; Illinois; Iowa; Massachusetts; Kansas; Missouri; New Jersey).
— Frank, 1982: 38 (Philonthus; list of parasites).
— Smetana, 1995e: 368 (Philonthus; lectotype designation: Lake Superior; characters; notes; Manitoba; Ontario; Quebec; Arkansas; Connecticut; Georgia; Illinois; Indiana; Iowa; Kansas; Kentucky; Maine; Maryland; Massachusetts; Michigan; Missouri; Nebraska; New Hampshire; New Jersey; New York; North Carolina; Ohio; Oklahoma; Pennsylvania; Tennessee; Texas; Virginia; West Virginia; Wisconsin).
— Downie and Arnett, 1996: 380 (Philonthus; characters; USA).
▴ Distribution: Canada, USA.
viperinus Horn, 1884: 205 (Philonthus; cited as a Fauvel manuscript name without characters or locality; [Note: Nomen nudum, Article 12.1]).
— Scheerpeltz, 1933: 1367 (Philonthus; cited as a valid species; [Note: The name is still unavailable]).
— Smetana, 1963a: 3, 4 (Philonthus; [Note: Smetana cited “. . . viperinus Fauvel, in litt.” (p. 3) followed by a long discussion then listed it as a synonym of aequalis; it is still unavailable]).
aeratus Stephens, 1832, see: intermedius Lacordaire, 1835.
aeripennis Cameron, 1943: 130 (Philonthus; Type locality: Darjeeling: Ghum district: Tiger Hill, altitude 8500–10,000 feet).
aerosus Kiesenwetter, 1851a: 416 (Philonthus; Type locality: Alpes de la Styrie).
— Kraatz, 1857d: 587 (Philonthus; characters; Alps).
— Redtenbacher, 1857: 191 (Philonthus; characters; Austria).
— Fauvel, 1874: 232 [= 1874a: 458] (Philonthus; characters; Styria; Moravia; Switzerland; Tirol; Sicily).
— Redtenbacher, 1874: 210 (Philonthus; characters; Austria).
— Ganglbauer, 1895: 446 (Philonthus; characters; Alps).
— Reitter, 1909: 125 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 327 (Philonthus; catalog).
— Scheerpeltz, 1933: 1331 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Smetana, 1958: 171 (Philonthus; subgenus Philonthus; characters; ecology; presence in Czechoslovakia doubtful).
— Smetana, 1959: 147 (Philonthus; characters).
— Coiffait, 1960: 363, 364 (Philonthus; subgenus Kenonthus; characters; Mountains of central Europe especially the Alps).
— Lohse, 1964: 177 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 131 (Philonthus; France; Switzerland; Italy; Germany; Austria).
— Coiffait, 1967a: 435 (Philonthus; subgenus Kenonthus; characters; Europe).
— Coiffait, 1974: 298 (Philonthus; subgenus Kenonthus; characters; Alps).
— Shilov, 1975a: 375 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Tóth, 1984: 47 (Philonthus; characters).
— Lucht, 1987: 104 (Philonthus; checklist; Germany; Czechoslovakia; Austria; Switzerland).
— Pilon and Zanetti, 1991: 59 (Philonthus; Italy).
— Smetana, 1993b: 48 (Philonthus; subgenus Kenonthus; Czech Republic ?; Slovakia ?).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; Italy).
▴ Distribution: Germany, Austria, Switzerland, Italy, France.
aeruginosus Nordmann, 1837: 95 (Philonthus; Type locality: Brasilia meridionalis).
— Erichson, 1840: 469 (Philonthus; characters; Brazil; Colombia).
— Bernhauer and Schubert, 1914: 327 (Philonthus; catalog).
— Scheerpeltz, 1933: 1331 (Philonthus; catalog; Argentina).
— Frank, 1982: 38 (Philonthus; list of parasites).
▴ Distribution: Colombia, Brazil, Argentina.
aethiopicus Bernhauer, 1915d: 142 (Philonthus; Type locality: Abessinien: Umgebung von Harrar).
— Scheerpeltz, 1933: 1331 (Philonthus; catalog).
— Tottenham, 1949b: 350 (Philonthus; characters).
▴ Distribution: Ethiopia.
aethiops Bernhauer, 1915d: 137 (Philonthus; Type locality: Abessinien: Harrar).
— Scheerpeltz, 1933: 1331 (Philonthus; catalog).
— Scheerpeltz, 1933: 1331 (Philonthus; catalog).
— Bernhauer, 1942c: 361 (Philonthus; Cameroon).
— Tottenham, 1949b: 317 (Philonthus; characters).
— Cameron, 1950g: 36 (Philonthus; Belgian Congo).
— Tottenham, 1962: 164 (Philonthus; characters).
▴ Distribution: Ethiopia, Congo, Cameroon.
affinis Lynch, 1884, see: lynchi Bernhauer and Schubert, 1914.
affinis Roth, 1851: 118 (Philonthus; Type locality: Abyssinien).
— Fauvel, 1907: 38 (Philonthus; Naivasha; Khartoum; Abyssinia).
— Bernhauer and Schubert, 1914: 327 (Philonthus; catalog).
— Bernhauer and Chapman, 1939: 66 (Philonthus; Kenya).
— Tottenham, 1949b: 306 (Philonthus; characters).
— Tottenham, 1962: 160 (Philonthus; characters).
▴ Distribution: Sudan, Ethiopia, Kenya.
africanus Fauvel, 1881a: 202 (Philonthus; Type locality: Scioa, Mahal-Uonz).
— Bernhauer and Schubert, 1914: 327 (Philonthus; catalog).
— Scheerpeltz, 1933: 1331 (Philonthus; catalog).
▴ Distribution: “Central Africa”.
agilis Gravenhorst, 1806, see: parvicornis Gravenhorst, 1802.
agnatus Erichson, 1840, see: Neobisnius.
agyrtes Smetana, 1973, see: Bisnius.
alaiensis Fauvel, 1900, see: Bisnius.
alaskensis Smetana, 1965, see: Bisnius.
alberti Schillhammer, 2000: 135 (Philonthus; replacement name for picipes Fauvel; West Palaearctic as far east as the Caucasus).
picipes Fauvel, 1875a: XXXI [= 1875b: 233] (Philonthus; [preoccupied]; Type locality: Caucase).
— Bernhauer and Schubert, 1914: 350 (Philonthus; catalog).
— Scheerpeltz, 1933: 1355 (Philonthus; catalog).
— Scheerpeltz, 1944a: 144 (Philonthus; characters).
— Lohse, 1956: 87 (Philonthus; characters; notes).
— Smetana, 1958: 167 (Philonthus; subgenus Philonthus; characters; ecology).
— Smetana, 1959: 148 (Philonthus; characters).
— Lohse, 1964: 178 (Philonthus; subgenus Philonthus; characters; central Europe).
— Smetana, 1965f: 253 (Philonthus; notes).
— Horion, 1965: 139 (Philonthus; Turkey; Caucasus; Transcaucasus; Bulgaria; Hungary; Romania; Austria).
— Coiffait, 1967a: 395 (Philonthus; characters; Caucasus; Transcaucasus; Balkans; central Europe).
— Fagel, 1968a: 193 (Philonthus; Turkey).
— Raianu, 1968a: 462 (Philonthus; characters; Romania).
— Scheerpeltz, 1968a: 72 (Philonthus; catalog; Austria).
— Tikhomirova, 1973a: 183 (Philonthus; checklist; USSR).
— Coiffait, 1974: 229 (Philonthus; characters; Caucasus; Transcaucasus; Balkans; Europe).
— Smetana, 1977: 794 (Philonthus; Turkey).
— Coiffait, 1978d: 164 (Philonthus; Turkey).
— Raichev, 1979: 82 (Philonthus; Bulgaria).
— Tóth, 1984: 52 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; Austria).
— K. Koch, 1989: 298 (Philonthus; habitat).
— Schuh, Schillhammer, and Zettel, 1992: 220 (Philonthus; notes; Austria).
— Ádám, 1996a: 21 (Bisnius; Hungary).
— Solodovnikov, 1998: 340 (Philonthus; bionomic notes; distribution in northwest Caucasus; Russia).
— Herman, 2001: 47 (Philonthus; name will be replaced by Schillhammer).
— Schillhammer, 2000: 135 (Philonthus; synonym of alberti).
▴ Distribution: Europe, Russia, Caucasus, Turkey.
albertisi Fauvel, 1878f: 260 (Philonthus; Type locality: Nouvelle-Guinée; Andai).
— Fauvel, 1879: 99 (Philonthus; Dorei).
— Bernhauer and Schubert, 1914: 372 (Leucitus; catalog).
— Last, 1975a: 1 (Philonthus)
▴ Distribution: New Guinea.
albilabris Nordmann, 1837: 86 (Philonthus; Type locality: Japonia).
— Erichson, 1840: 454 (Philonthus; characters; Japan).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog; China).
— Scheerpeltz, 1933: 1331 (Philonthus; catalog).
— Schillhammer, 1999a: 61 (Philonthus).
▴ Distribution: Japan, China.
frater Bernhauer, 1907: 387 (Philonthus; Type locality: Kamakura; Takakijama).
— Bernhauer and Schubert, 1914: 339 (Philonthus; catalog).
— Scheerpeltz, 1933: 1343 (Philonthus; catalog).
— Ito, 1994c: 84 (Philonthus; characters; Japan).
— Schillhammer, 1999a: 61 (Philonthus; synonym of albilabris).
albionicus Mannerheim, 1843, see: Bisnius.
albipes Gravenhorst, 1802: 28 (Staphylinus; Type locality: Brunsvigae).
— Latreille, 1804: 316 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 76 (Staphylinus; characters).
— Gyllenhal, 1810: 327 (Staphylinus; characters; Sweden).
— Zetterstedt, 1828: 76 (Staphylinus; characters; Lapland).
— C. Sahlberg, 1830: 323 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 29 (Staphylinus; Finland).
— Mannerheim, 1831: 443 (Staphylinus; Finland).
— Lacordaire, 1835: 405 (Staphylinus; characters; France).
— Zetterstedt, 1838: 64 (Staphylinus; characters; habitat; Lapland).
— Erichson, 1839a: 457 (Philonthus; characters; Germany).
— Heer, 1839: 262 (Philonthus; characters; Switzerland).
— Erichson, 1840: 449 (Philonthus; characters; Germany; Sweden; France).
— Laporte, 1840: 178 (Staphylinus; characters; Germany; France).
— Jacobsen, 1849: 692 (Philonthus; characters; Denmark).
— Redtenbacher, 1849: 702 (Philonthus; characters; Austria).
— Hardy, 1851: 27 (Philonthus; notes; England).
— Fairmaire and Laboulbène, 1856: 520 (Philonthus; characters; France).
— Kraatz, 1857d: 585 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 192 (Philonthus; characters; Austria).
— Thomson, 1860: 163 (Philonthus; characters; Scandinavia).
— Hochhuth, 1862: 57 (Philonthus; Russia).
— Fauvel, 1865a: 296 [= 1865b: 46] (Philonthus; habitat; France).
— Fauvel, 1874: 256 [= 1874a: 482] (Philonthus; characters; France; Turkey; Italy; Caucasus; Lake Baikal; Amur region).
— Redtenbacher, 1874: 211 (Philonthus; characters; Austria).
— Seidlitz, 1875: 275 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 415 [= 1877: 271] (Philonthus; characters; France).
— J. Sahlberg, 1876: 20 (Philonthus; Finland).
— Fowler, 1888: 266 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 449 (Philonthus; characters; Palaearctic region).
— Fauvel, 1902b: 113 (Philonthus; Tunisia).
— Poppius, 1909: 26 (Philonthus; subgenus Gefyrobius; Lena Valley, Siberia).
— Reitter, 1909: 128 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog).
— Johansen, 1914: 400 (Philonthus; characters; Denmark).
— Portevin, 1929: 361 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1331 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— V. Hansen, 1952: 59 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1958: 198 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 152 (Philonthus; characters).
— Smetana, 1963c: 296 (Philonthus; subgenus Philonthus; Mongolia).
— Lohse, 1964: 180 (Philonthus; subgenus Philonthus; characters; central Europe).
— Smetana, 1965e: 174 (Philonthus; Mongolia).
— Horion, 1965: 146 (Philonthus; Siberia; Mongolia; Russia; Lapland; Scotland; widespread in central Europe; Serbia; Bosnia; Italy).
— Pototskaia, 1966: 156 (Philonthus; larval characters).
— Pototskaia, 1967: 57 (Philonthus; larval characters).
— Coiffait, 1967a: 411 (Philonthus; characters).
— Smetana, 1967d: 206 (Philonthus; Mongolia).
— Raianu, 1968a: 467 (Philonthus; Romania).
— Szujecki, 1968a: 730 (Philonthus; Poland).
— Coiffait, 1974: 257 (Philonthus; characters; Palaearctic region).
— Smetana 1975c: 171 (Philonthus; Mongolia).
— Shilov, 1975a: 376 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Shilov, 1976: 151 (Philonthus; Mongolia).
— Pope, 1977: 29 (Philonthus; Britain).
— Frank, 1982: 38 (Philonthus; list of parasites).
— Boller, 1983: 118 (Philonthus; larval characters).
— Tóth, 1984: 58 (Philonthus; characters; Hungary).
— Lucht, 1987: 104 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Pilon and Zanetti, 1991: 58 (Philonthus; Italy).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; Italy).
— Kashcheev, 1999a: 58 (Philonthus; Kazakhstan).
▴ Distribution: Tunisia, Europe, Turkey, Caucasus, Russia, Kazakhstan, Mongolia.
simplex Marsham, 1802: 505 (Staphylinus; Type locality: Britain).
— Gyllenhal, 1810: 345 (Staphylinus; synonym of virgo).
— Curtis, 1829: 25 (Philonthus; catalog; Britain).
— Stephens, 1832: 234 (Philonthus; characters; England).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Stephens, 1839: 396 (Philonthus; characters; England).
— Kraatz, 1857d: 602 (Philonthus; synonym of varians).
— G. Waterhouse, 1858: 24 (Philonthus; synonym of albipes).
— Gemminger and Harold, 1868: 585 (Philonthus; synonym of albipes).
— Ganglbauer, 1895: 449 (Philonthus; synonym of albipes).
— Bernhauer and Schubert, 1914: 328 (Philonthus; synonym of albipes).
— Smetana, 1958: 199 (Philonthus; synonym of albipes).
— Coiffait, 1974: 257 (Philonthus; synonym of albipes).
microcephalus Gravenhorst, 1806: 70 (Staphylinus; Type locality: Parisiae).
— Erichson, 1840: 459 (Philonthus; characters; France).
— Kraatz, 1857d: 575 (Philonthus).
— Fauvel, 1874: 256 [= 1874a: 482] (Philonthus; synonym of albipes).
— Redtenbacher, 1874: 211 (Philonthus; characters; Austria).
— Ganglbauer, 1895: 449 (Philonthus; synonym of albipes).
— Bernhauer and Schubert, 1914: 328 (Philonthus; synonym of albipes).
— Scheerpeltz, 1933: 1331 (Philonthus; synonym of albipes).
— Smetana, 1958: 199 (Philonthus; synonym of albipes).
— Coiffait, 1974: 257 (Philonthus; synonym of albipes).
hirtipennis Stephens, 1832: 221 (Quedius; Type locality: London; Suffolk).
— Stephens, 1839: 391 (Quedius; variety of picipes; characters; England).
— G. Waterhouse, 1858: 24 (Philonthus; synonym of albipes).
— Gemminger and Harold, 1868: 585 (Philonthus; synonym of albipes).
— Ganglbauer, 1895: 449 (Philonthus; synonym of albipes).
— Bernhauer and Schubert, 1914: 328 (Philonthus; synonym of albipes).
— Smetana, 1958: 199 (Philonthus; synonym of albipes).
— Coiffait, 1974: 257 (Philonthus; synonym of albipes).
nitescens Stephens, 1832: 222 (Quedius; Type locality: Norfolk: Swansea).
— Stephens, 1839: 392 (Quedius; characters; England).
— G. Waterhouse, 1858: 24 (Philonthus; synonym of albipes).
— Gemminger and Harold, 1868: 585 (Philonthus; synonym of albipes).
— Ganglbauer, 1895: 449 (Philonthus; synonym of albipes).
— Bernhauer and Schubert, 1914: 328 (Philonthus; synonym of albipes).
— Smetana, 1958: 199 (Philonthus; synonym of albipes).
— Coiffait, 1974: 257 (Philonthus; synonym of albipes).
alcyoneus Erichson, 1840: 476 (Philonthus; Type locality: Sardinia).
— Kraatz, 1857d: 596 (Philonthus).
— Fauvel, 1874: 223 [= 1874a: 449] (Philonthus; characters; Corsica; Sardinia; Sicily; Algeria).
— Mulsant and Rey, 1876: 490 [= 1877: 346] (Philonthus; characters; France).
— Fauvel, 1878: 122 [= 1878a: 42] (Philonthus; Algeria).
— Fauvel, 1886: 54 [= 1886a: 46] (Philonthus; notes; Morocco; Tunisia; Algeria).
— Fauvel, 1902b: 109 (Philonthus; Morocco; Algeria; Tunisia).
— Sainte-Claire Deville, 1906: 97 (Philonthus; Corsica).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog).
— Portevin, 1929: 363 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1331 (Philonthus; catalog).
— Scheerpeltz, 1965b: 403 (Philonthus; Sicily).
— Scheerpeltz, 1965c: 506 (Philonthus; Sicily).
— Coiffait, 1967a: 425 (Philonthus; characters; North Africa; Sicily; Sardinia; Corsica).
— Coiffait, 1974: 206 (Philonthus; characters; Corsica; Sardinia).
— Maachi, 1991: 251 (Philonthus; Morocco).
— Aranda and Outerelo, 1991: 121 (Philonthus; characters).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; Italy).
— Marchal and Chardonnet, 2000: 72 (Philonthus; Corsica).
▴ Distribution: Tunisia, Algeria, Morocco, Spain, France, Italy, Lebanon.
intermedius Ragusa, 1891: 223 (Philonthus; variety of alcyoneus; [preoccupied]; Type locality: Sardegna; Sicilia; Algeria).
— Bernhauer and Schubert, 1914: 328 (Philonthus; variety of alcyoneus).
— Peyerimhoff, 1931: 31 (Philonthus; subspecies of intermedius; notes; Algeria).
— Scheerpeltz, 1933: 1331 (Philonthus; synonym of ragusae).
— Jarrige, 1958: 92 (Philonthus; subspecies of alcyoneus; notes; Algeria).
— Coiffait, 1966b: 338 (Philonthus; subspecies of alcyoneus; Hoggar).
devillei Chapman, 1932: 36 (Philonthus; subgenus Gefyrobius; variety of alcyoneus; Type locality: Sicile: Messine).
— Coiffait, 1967a: 427 (Philonthus; variety of alcyoneus; characters).
— Coiffait, 1974: 206 (Philonthus; variety of alcyoneus; Sicily).
ragusai Scheerpeltz, 1933: 1331 (Philonthus; cited as ragusae; replacement name for intermedius Ragusa).
— Coiffait, 1967a: 425 (Philonthus; cited as ragusae; subspecies of alcyoneus; characters; Sicily; North Africa; Andalousia; Lebanon).
— Coiffait, 1974: 206 (Philonthus;cited as ragusae; subspecies of alcyoneus; Sicily; Spain; Tunisia; Algeria; Morocco; Spain).
— Pace and Zanetti, 1977: 299 (Philonthus; cited as ragusai [Note: Correction of erroneous original spelling of a patronym for Enrico Ragusa]; subspecies of alcyoneus; Italy).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; cited as ragusai; subspecies of alcyoneus; Italy).
wagneri Coiffait, 1963: 23 (Philonthus; aberration of alcyoneus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Corse: Guagno-les-bains).
aletes Smetana, 1967d: 200 (Philonthus; subgenus Philonthus; Type locality: Mongolei: Chentej aimak, zwischen Somon Zenchermandal und Somon Zargaltchan, 1400 m).
— Shilov, 1976: 148 (Philonthus; Mongolia).
— Schillhammer, 2000: 138 (Philonthus; characters; Mongolia).
▴ Distribution: Mongolia.
algarum Sharp, 1874, see: Cafius.
algiricus Motschulsky, 1858, see: longicornis Stephens, 1832.
alienus Eppelsheim, 1889, see: Rabigus.
aliicornis Bernhauer, 1939, see: Gabrius.
aliquatenus Schubert, 1908: 618 (Philonthus; Type locality: Kulu, Himalaya, ca. 2000 m).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog).
— Cameron, 1932: 107 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1332 (Philonthus; catalog).
— Coiffait, 1981e: 349 (Philonthus; Kashmir).
— Coiffait, 1982: 28 (Philonthus; Nepal).
— Coiffait, 1982a: 233 (Philonthus; Bhutan; Nepal; India).
— Coiffait, 1984e: 373 (Philonthus; Nepal).
▴ Distribution: India, Bhutan, Nepal.
alius Bernhauer and Schubert, 1914: 328 (Philonthus; replacement name for crassulus Sharp).
— Navarrete-Heredia, 1995: 96 (Philonthus; Mexico; Guatemala).
— Marquez-Luna and Navarrete Heredia, 1995: 35 (Philonthus; in detritus of Atta mexicana; Mexico).
▴ Distribution: Mexico, Guatemala.
crassulus Sharp, 1885: 396 (Philonthus; [preoccupied]; Type locality: Mexico, Las Vigas; Guanajuato).
allardi Levasseur, 1962: 239 (Philonthus; Type locality: Kolwezi [Lualaba], Haut Katanga, Congo belge).
allardianus Levasseur, 1962: 237 (Philonthus; Type locality: Kolwezi [Lualaba], Haut Katanga, Congo belge).
alluaudanus Jeannel and Paulian, 1945, see: Gabronthus.
almorensis Cameron, 1932, see: Gabrius.
alpestris Heer, 1839, see: Quedius.
alpigradus Mulsant and Rey, 1876, see: varians Paykull, 1789.
alpinus Eppelsheim, 1875a: 409 (Philonthus; variety of albipes; Type locality: Stilfser-Joch in Tyrol).
— Ganglbauer, 1895: 449 (Philonthus; synonym of albipes).
— Bernhauer and Schubert, 1914: 328 (Philonthus; variety of albipes).
— Donisthorpe, 1930: 96 (Philonthus; aberration of albipes; characters; Britain).
— Scheerpeltz, 1933: 1331 (Philonthus; variety of albipes).
— Lohse, 1956: 89 (Philonthus; characters; notes).
— Smetana, 1958: 199 (Philonthus; aberration of albipes).
— Smetana, 1959: 152 (Philonthus; characters).
— V. Hansen, 1962: 243 (Philonthus; Denmark).
— Palm, 1963b: 76 (Philonthus; characters; Sweden; Denmark).
— Lohse, 1964: 180 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 147 (Philonthus; Denmark; Sweden; Germany; Austria; Czechoslovakia; Poland; Romania; Italy; France; Austria; Germany).
— Coiffait, 1967a: 411 (Philonthus; characters; Europe).
— Coiffait, 1974: 257 (Philonthus; characters; Britain; mountain of Europe; Spain; France).
— Tóth, 1984: 58 (Philonthus; characters; Hungary).
— Lucht, 1987: 104 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Pilon and Zanetti, 1991: 58 (Philonthus; Italy).
— Vorst, 1992: 101 (Philonthus; collecting notes; Netherlands).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Hodge and Jones, 1995: 38 (Philonthus; characters; Britain).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; Italy).
▴ Distribution: Europe.
altaicus Bernhauer, 1923, see: kiautschauensis Bernhauer, 1915.
altaicus Coiffait, 1967, see: politus Linné, 1758.
alterius Cameron, 1951c: 402 (Philonthus; Type locality: Not cited).
— Tottenham, 1961: 204 (Philonthus; Senegal).
— Tottenham, 1962: 195 (Philonthus; notes).
— Lecoq, 1990: 200 (Philonthus; Mascarene Islands).
▴ Distribution: Mascarene Islands, Sudan, Ghana, Senegal.
daunus Tottenham, 1953: 147 (Philonthus; Type locality: Sudan: Khartoum, Zaphiro. Also cited from Gold Coast).
— Tottenham, 1962: 195 (Philonthus; synonym of alterius).
— Lecoq, 1990: 200 (Philonthus; synonym of alterius).
alternans Sharp, 1885: 418 (Philonthus; Type locality: Mexico, Jalapa, Oaxaca).
alticola Sharp, 1885: 400 (Philonthus; Type locality: Guatemala, Totonicapam, 8500 to 10,500 feet).
altissimus Bernhauer, 1939, see: Gabrius.
altivagans Fauvel, 1907: 42 (Philonthus; Type locality: Kilimandjaro: Kiboscho, 3200 m).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog).
— Bernhauer and Chapman, 1939: 72 (Philonthus; Kilimandjaro).
— Tottenham, 1962: 137, 174, 176 (Philonthus; characters; notes; Tanganyika).
▴ Distribution: Tanzania.
swynnertoni Tottenham, 1953b: 496 (Philonthus; Type locality: Kilimanjaro: Shira Plateau, Camp 2, 12,450 ft.).
— Tottenham, 1962: 137 (Philonthus; variety of altivagans; Tanganyika).
— Tottenham, 1962: 174, 177 (Philonthus; variety of altivagans).
discalis Tottenham, 1962: 174 (Philonthus; variety of altivagans; Type locality: Mt. Meru).
alumnus Erichson, 1840: 490 (Philonthus; Type locality: ins. Puerto Rico; [Note: See lectotype designation by Smetana, 1995e]).
— Fall and Cockerell, 1907: 166 (Philonthus; [Note: May be a misidentification of alumnus and may be rufulus according to Smetana, 1995e: 434]; New Mexico).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog).
— Notman, 1920b: 706 (Philonthus; [Note: May be a misidentification of alumnus and may be rufulus according to Smetana, 1995e: 434]; Florida).
— Scheerpeltz, 1933: 1332 (Philonthus; catalog).
— Moore and Legner, 1973a: 45 (Philonthus; notes; [Note: May be a misidentification of alumnus and may be rufulus according to Smetana, 1995e: 434]; California).
— Holcomb, 1977 (Philonthus; notes on aggression, feeding, mating, grooming, and cleaning).
— Smetana, 1994: 342 (Philonthus; characters; lectotype designation: Puerto Rico; Cayman Islands; Cuba; Haiti; Mona Island; Puerto Island; specimens determined as alumnus from North America are different species).
— Navarrete-Heredia, 1995: 96 (Philonthus; Mexico; Puerto Rico; USA).
— Downie and Arnett, 1996: 379 (Philonthus; characters; USA).
▴ Distribution: West Indies, Mexico.
alutaceus Horn, 1884: 183 (Philonthus; Type locality: southern regions of California; [Note: See lectotype designation by Smetana, 1995e]).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog).
— Smetana, 1991a: 235 (Philonthus; characters; Mexico).
— Smetana, 1995e: 106 (Philonthus; characters; lectotype designation: So. Cal. [southern regions of California]; Arizona; California; Mexico).
▴ Distribution: Mexico, USA.
aluticollis Bernhauer, 1921c: 107 (Philonthus; Type locality: Argentinien: Jujuy, Cueva Iturbe, 3700 m).
alutipennis Bernhauer, 1928g: 111 (Philonthus; Type locality: Belg.-Kongo: Mongende).
alveus Last, 1968: 369 (Philonthus; Type locality: New Guinea: Yentchan Sepik, Main River).
amabilis Kraatz, 1859, see: Hesperus.
amamiensis Hayashi, 1993: 78 (Philonthus; Type locality: Japan: Hatsuno, Amami Ohshima Is.).
amazonicus Sharp, 1876, see: figulus Erichson, 1840.
ambiguus Erichson, 1840: 441 (Philonthus; Type locality: Columbia).
amblyterus Olliff, 1887, see: politus Linné, 1758.
amicus Sharp, 1874: 45 (Philonthus; Type locality: Orakami Marsh, Nagasaki).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog).
— Scheerpeltz, 1933: 1332 (Philonthus; catalog).
— Watanabe and Shibata, 1972: 64 (Philonthus; Yaku-shima Island).
— Frank, 1982: 38 (Philonthus; list of parasites).
— Watanabe, 1998a: 140 (Philonthus; Japan).
— Watanabe, 1998b: 262 (Philonthus; Japan).
▴ Distribution: Japan.
aminius Tottenham, 1955: 173 (Philonthus; Type locality: Kenya: Muguga).
amnicola Gistel, 1857: 62 (Philonthus; [Note: Nomen dubium]; Type locality: Rhaetia).
amoenus Olivier, 1795, see: cyanipennis Fabricius, 1793.
amoenus Guérin-Méneville, 1844, see: Phanolinus.
analis Heer, 1839, see: Gabrius.
analis Erichson, 1840, see: Xenopygus.
analis Fauvel, 1907, see: trunculus Herman, 2001.
analokus Coiffait, 1982, see: obsoletus Eppelsheim, 1895.
ancora Reitter, 1909: 125 (Philonthus; Type locality: Transkaspien: Aulie-Ata).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog).
— Scheerpeltz, 1933: 1332 (Philonthus; catalog).
— Smetana, 1965f: 253 (Philonthus; characters).
— Coiffait, 1967a: 392 (Philonthus; characters; Transcaspian region).
— Coiffait, 1974: 277 (Philonthus; characters; Turkestan).
▴ Distribution: Kazakhstan.
andalusiacus Coiffait, 1966, see: quisquiliarius Gyllenhal, 1810.
andamanicus Coiffait, 1981f: 334 (Philonthus; Type locality: South Andaman, Port Blair, Chiriyatapu).
andrewesi Cameron, 1920a: 214 (Philonthus; Type locality: Nilgiri Hills).
— Cameron, 1932: 77 (Philonthus; aberration of cinctulus).
— Scheerpeltz, 1933: 1336 (Philonthus; aberration of cinctulus).
— Schillhammer, 1999: 63 (Philonthus; valid species).
▴ Distribution: India.
anepsius Cameron, 1926b: 353 (Philonthus; Type locality: Siwaliks; Mohan Rau).
— Cameron, 1932: 128 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1332 (Philonthus; catalog).
▴ Distribution: India.
angolensis Cameron, 1959: 116 (Philonthus; Type locality: Angola: Lunda N.E.: Riv. Tchikapa).
anguinus Fauvel, 1874: 238 [= 1874a: 464] (Philonthus; Type locality: Hautes Pyrénées, Arrens).
— Mulsant and Rey, 1876: 422 [= 1877: 278] (Philonthus; characters; France).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog).
— Portevin, 1929: 365 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1332 (Philonthus; catalog).
— Coiffait, 1967a: 408 (Philonthus; characters; Pyrenees; Maritime Alps).
— Coiffait, 1974: 285 (Philonthus; characters; Pyrenees; Alps).
▴ Distribution: France.
angularis Erichson, 1840, see: Paederomimus.
anguliceps Bernhauer, 1902c: 57 (Philonthus; Type locality: Turkestan, Aulie Ata).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog).
— Scheerpeltz, 1933: 1332 (Philonthus; catalog).
— Coiffait, 1967a: 393 (Philonthus; characters; Turkestan).
— Coiffait, 1974: 278 (Philonthus; characters; Turkestan).
▴ Distribution: Kazakhstan.
angulicollis Motschulsky, 1860, see: politus Linné, 1758.
angustatus Kraatz, 1859, see: Gabronthus.
angusticollis Erichson, 1840, see: Paederomimus.
angusticeps Bernhauer, 1933, see: Gabrius.
angusticeps Bernhauer, 1934, see: stenocephalus Scheerpeltz, 1972.
annandalei Cameron, 1924, see: maindroni Fauvel, 1903.
antennalis Cameron, 1932, see: distincticornis Cameron, 1932.
antennalis Cameron, 1937, see: antennarius Herman, 2001.
antennarius Herman, 2001: 46 (Philonthus; replacement name for antennalis).
antennalis Cameron, 1937: 7 (Philonthus; [preoccupied]; Type locality: Java: Bandoeng Dago).
antennatus Guérin-Méneville, 1844, see: Styngetus.
antennatus Motschulsky, 1858, see: distinctus Gemminger and Harold, 1868.
antennatus Bernhauer and Schubert, 1914, see: aemulus Tottenham, 1939.
anthracinus Lea, 1925: 222 (Philonthus; Type locality: Western Australia: Albany).
anthrax Gravenhorst, 1802, see: ventralis Gravenhorst, 1802.
antipodum Fauvel, 1877: 263 (Philonthus; Type locality: Australie, Victoria; Queensland, Gayndah).
— Fauvel, 1878g: 546 (Philonthus).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog).
▴ Distribution: Australia.
aokii R. Dvořák, 1958: 135 (Philonthus; Type locality: Japan: Nikko, Senjo-ga-hara).
apenninus Fiori 1894, see: gagates Mulsant and Rey, 1876.
apheles Solsky, 1868a: 132 (Philonthus; Type locality: Mexique).
— Sharp, 1885: 405 (Philonthus; characters; Mexico).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog).
▴ Distribution: Mexico.
apicalis Leinberg, 1900, see: caucasicus Nordmann, 1837.
apicetestaceus Scheerpeltz, 1960c: 106 (Philonthus; Type locality: Bolivien: Yungas de Arepucho, Chacisacha, 1500 m).
apicicornis Guérin-Méneville, 1830: 62 (Staphylinus; Type locality: Nouvelle-Guinée).
— Erichson, 1840: 931 (Philonthus; characters; New Guinea).
— Fauvel, 1878f: 260 (Philonthus; characters; Nouvelle-Guinée, Doreï).
— Fauvel, 1879: 100 (Philonthus; characters; Mysol; Fly River).
— Bernhauer and Schubert, 1914: 329 (Philonthus; catalog).
▴ Distribution: New Guinea.
apicipennis Cameron, 1932, see: terminipennis Tottenham, 1939.
apicipennis Lynch, 1884, see: flavolimbatus Erichson, 1840.
apicipunctus Lea, 1925: 223 (Philonthus; Type locality: Queensland: Cairns).
apiciventris Sharp, 1885, see: Belonuchus.
approximans Sharp, 1887: 428 (Philonthus; Type locality: Panama, Volcan de Chiriqui 4000 to 6000 feet).
approximatus Hatch, 1957, see: Gabrius.
aprilinus Gistel, 1857: 61 (Philonthus; [Note: Nomen dubium]; Type locality: Galliae Alpes).
arabicus Bernhauer, 1935a: 45 (Philonthus; Type locality: Arabien: Yemen).
buettikeri Coiffait, 1979a: 165 (Philonthus; Type locality: Saudi Arabia: Abha-Gizan, km 28 Wadi ad Dilla, 700 m).
arabicus Coiffait, 1981, see: variipennis Kraatz, 1859.
arabiensis Herman, 2001: 46 (Philonthus; replacement name for thoracicus Coiffait).
thoracicus Coiffait, 1979a: 165 (Paragabrius; [preoccupied]; Type locality: Saudi Arabia: Hofuf).
— Smetana, 1995e: 51 (Philonthus; [Note: Paragabrius is synonymy of Philonthus]).
— Herman, 2001: 46 (Philonthus; synonym of arabiensis).
arachosicus Coiffait, 1983e: 478 (Philonthus; Type locality: Bashgultal, 1100 m, Nuristan, Afghanistan).
archangelicus Poppius, 1908: 46 (Philonthus; Type locality: Nord-Russland, unterer Lauf des Flusses Petschora).
— Bernhauer and Schubert, 1914: 329 (Philonthus; catalog).
— Scheerpeltz, 1933: 1332 (Philonthus; catalog).
— Coiffait, 1967a: 413 (Philonthus; characters; Russia).
— Coiffait, 1974: 292 (Philonthus; characters; Russia).
— Shilov, 1975a: 376 [= 1975b: 99] (Philonthus; Komi region of Russia).
— Boháč, 1988: 551 [= 1989: 34] (Philonthus; characters; Russia).
▴ Distribution: Russia.
arctifrons Broun, 1880, see: Quedius.
ardoini Levasseur, 1966b: 1495 (Philonthus; Type locality: Katire-Equatoria, république du Sudan).
— Levasseur, 1968a: 54 (Philonthus; Central African Republic).
▴ Distribution: Sudan, Central African Republic.
arenarius Kirshenblat, 1950, see: Bisnius.
argentinus Bernhauer, 1912d: 43 (Philonthus; Type locality: Argentinien: Prov. Tucuman).
argus Herman, 2001: 46 (Philonthus; replacement name for tucumanensis Bernhauer, 1934).
tucumanensis Bernhauer, 1934a: 118 (Philonthus; [preoccupied]; Type locality: Argentinien: Tucuman: Parque Aconquija).
argutus Herman, 2001: 46 (Philonthus; replacement name for nitens Kraatz).
nitens Kraatz, 1859: 82 (Philonthus; [preoccupied]; Type locality: Ceylan).
— Bernhauer and Schubert, 1914: 348 (Philonthus; catalog).
— Cameron, 1932: 96 (Philonthus; characters; Ceylon).
— Scheerpeltz, 1933: 1353 (Philonthus; catalog).
— Herman, 2001: 46 (Philonthus; synonym of argutus).
arizonensis Horn, 1884: 203 (Philonthus; Type locality: Arizona; [Note: See lectotype designation by Smetana, 1995e]. No locality originally cited).
— Bernhauer and Schubert, 1914: 329 (Philonthus; catalog).
— Smetana, 1995e: 374 (Philonthus; lectotype designation: Ariz [Arizona]; characters; notes; Arizona; Mexico).
▴ Distribution: Mexico, USA.
armatipes Sharp, 1885: 422 (Philonthus; Type locality: Guatemala, near the city at an elevation of 5000 feet).
armeniacus Bernhauer, 1901, see: diversiceps Bernhauer, 1901.
armeniacus Hochhuth, 1851: 17 (Philonthus; Type locality: Armenien).
— Bernhauer and Schubert, 1914: 329 (Philonthus; catalog).
— Scheerpeltz, 1933: 1332 (Philonthus; catalog).
— Coiffait, 1967a: 400 (Philonthus; note; Armenia).
— Coiffait, 1974: 281 (Philonthus; characters; Caucasus; Transcaucasus).
▴ Distribution: Armenia.
arrowianus Bernhauer, 1931d: 585 (Philonthus; Type locality: Abyssinia: Mt. Zukwala, ca. 9,000 ft.; Mt. Chillalo, ca. 8,000 ft.).
asemus Kraatz, 1859, see: longicornis Stephens, 1832.
asiaticus Bernhauer, 1902b: 27 (Philonthus; Type locality: Kanthaley).
— Bernhauer and Schubert, 1914: 329 (Philonthus; catalog).
— Cameron, 1932: 143 (Philonthus; characters; Ceylon).
— Scheerpeltz, 1933: 1332 (Philonthus; catalog).
▴ Distribution: Sri Lanka.
asper Horn, 1884: 185 (Philonthus; Type locality: Pennsylvania; District of Columbia).
— Bernhauer and Schubert, 1914: 329 (Philonthus; catalog).
— Smetana, 1995e: 131 (Philonthus; characters; Ontario; Quebec; Georgia; Illinois; Indiana; Maryland; New Jersey; New York; North Carolina; Ohio; Pennsylvania; South Carolina; Tennessee; Virginia; West Virginia).
— Downie and Arnett, 1996: 377 (Philonthus; characters; USA).
▴ Distribution: Canada, USA.
asperulus Fauvel, 1875a: XXX [= 1875b: 232] (Philonthus; Type locality: Syrie, Jaffa, Jérusalem).
— Bernhauer and Schubert, 1914: 329 (Philonthus; catalog).
— Scheerpeltz, 1933: 1332 (Philonthus; catalog).
— Coiffait, 1967a: 413 (Philonthus; characters; Syria; Palestine; Turkey).
— Coiffait, 1974: 292 (Philonthus; characters; Syria; Palestine; Turkey).
▴ Distribution: Israel, Syria, Turkey.
asphaltinus Erichson, 1840, see: Jurecekia.
assamensis Cameron, 1932: 100 (Philonthus; Type locality: Assam: Manipur).
— Scheerpeltz, 1933: 1333 (Philonthus; catalog).
— Coiffait, 1982a: 232 (Philonthus; India).
▴ Distribution: India.
assecla Last, 1968, see: Hesperus.
assimilis Nordmann, 1837, see: Quedius.
astutoides A. Strand, 1946, see: Gabrius.
astutus Erichson, 1840, see: Gabrius.
ater Ragusa, 1892, see: sanguinolentus Gravenhorst, 1802.
ater Ziegler, 1845, see: Quedius.
aterrimus Marsham, 1802, see: varians Paykull, 1789.
atkinsoni Cameron, 1932: 136 (Philonthus; Type locality: Bengal: Berhampore).
atramentarius Erichson, 1840: 440 (Philonthus; Type locality: Columbia).
atratoides Coiffait, 1963: 24 (Philonthus; Type locality: Corse: environs de Corte, Vallée de la Raconica, 200 m).
— Coiffait, 1967a: 391 (Philonthus; characters; Corsica).
— Coiffait, 1974: 249 (Philonthus; characters; Corsica).
▴ Distribution: France.
atratus Gravenhorst, 1802: 21 (Staphylinus; Type locality: Brunsvigae).
— Latreille, 1804: 310 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 84 (Staphylinus; characters; Germany; France).
— Gyllenhal, 1810: 319 (Staphylinus; characters; Sweden).
— Curtis, 1829: 25 (Philonthus; catalog; Britain).
— C. Sahlberg, 1830: 321 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 28 (Staphylinus; Finland; Sweden; Lapland).
— Mannerheim, 1831: 442 (Staphylinus; Finland; Sweden; Lapland).
— Stephens, 1832: 232 (Philonthus; characters; England).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Nordmann, 1837: 82 (Philonthus; Europe).
— Zetterstedt, 1838: 63 (Staphylinus; characters; habitat; Lapland).
— Erichson, 1839a: 450 (Philonthus; characters; Germany).
— Heer, 1839: 259 (Philonthus; characters; Switzerland).
— Stephens, 1839: 395 (Philonthus; characters; England).
— Erichson, 1840: 439 (Philonthus; characters; Europe).
— Kolenati, 1846: 19 (Philonthus; Georgia Caucasica).
— Jacobsen, 1849: 689 (Philonthus; characters; Denmark).
— Ménétriés, 1849: 44 (Philonthus; Samarkand).
— Hochhuth, 1849: 134 (Philonthus; characters).
— Redtenbacher, 1849: 700 (Philonthus; characters; Austria).
— Küster, 1853a: 49 (Philonthus; characters; Europe).
— Rosenhauer, 1856: 71 (Philonthus; notes).
— Fairmaire and Laboulbène, 1856: 517 (Philonthus; characters; France).
— Kraatz, 1857d: 586 (Philonthus; characters; Germany).
— Kraatz, 1857d: 586 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 329 cited “coerulescens Kraatz” as a synonym of atratus; Kraatz did not describe the species as new, he misspelled the name and attributed it to Lacordaire]; synonym of atratus).
— Redtenbacher, 1857: 190 (Philonthus; characters; Austria).
— Thomson, 1860: 158 (Philonthus; characters; Scandinavia).
— Hochhuth, 1862: 57 (Philonthus; Russia).
— Fauvel, 1865a: 296 [= 1865b: 46] (Philonthus; habitat; France).
— Hochhuth, 1872: 137 (Philonthus; notes; Ukraine).
— Fauvel, 1874: 230 [= 1874a: 456] (Philonthus; characters; France; Caucasus; Iran; Siberia; central Asia).
— Redtenbacher, 1874: 209 (Philonthus; characters; Austria).
— Solsky, 1874: 176 (Philonthus; characters; Turkestan).
— Seidlitz, 1875: 274 (Philonthus; characters; Baltic region).
— Fauvel, 1876a: 74 [= 1876: 259] (Philonthus; Syria; Buchara).
— Mulsant and Rey, 1876: 385 [= 1877: 241] (Philonthus; characters; France).
— J. Sahlberg, 1876: 16 (Philonthus; Finland).
— Fowler, 1888: 263 (Philonthus; characters; Britain).
— Eppelsheim, 1892: 332 (Philonthus; Turkmenia; Taschkent).
— Ganglbauer, 1895: 446 (Philonthus; characters; Palaearctic region).
— Poppius, 1909: 23 (Philonthus; Lena Valley, Siberia).
— Reitter, 1909: 126 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 329 (Philonthus; catalog).
— Johansen, 1914: 396 (Philonthus; characters; Denmark).
— Gridelli, 1922a: 176 (Philonthus; Persia).
— Portevin, 1929: 358 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1333 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Suomalainen, 1941 (Philonthus; characters; Finland).
— V. Hansen, 1952: 52 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1953: 117 (Philonthus; Turkey).
— Smetana, 1958: 168 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 147 (Philonthus; characters).
— Scheerpeltz, 1961d: 128 (Philonthus; Italy).
— Palm, 1963b: 68 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 177 (Philonthus; subgenus Philonthus; characters; central Europe).
— Smetana, 1964d: 74 (Philonthus; subgenus Philonthus; Czechoslovakia).
— Horion, 1965: 129 (Philonthus; Russia; Finland; Sweden; Norway; Britain; Denmark; all central Europe to Mediterranean).
— Sawada, 1966: 258 (Philonthus; Afghanistan).
— Coiffait, 1967a: 393 (Philonthus; characters).
— Raianu, 1968a: 464 (Philonthus; Romania).
— Szujecki, 1968a: 727 (Philonthus; Poland).
— Szujecki, 1969a: 255 (Philonthus; notes; Poland).
— Coiffait, 1974: 248 (Philonthus; characters; Holarctic region).
— Shilov, 1975a: 375 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Pace and Zanetti, 1977: 290 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Coiffait, 1978d: 164 (Philonthus; Turkey).
— Tronquet, 1981: 71 (Philonthus; Afghanistan).
— Coiffait, 1983e: 477 (Philonthus; Afghanistan).
— Tóth, 1984: 48 (Philonthus; characters; Hungary).
— Lucht, 1987: 104 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Lohse and Lucht, 1989: 165 (Philonthus; characters).
— Whitehead, 1990: 242 (Philonthus; collecting notes; England).
— J. Li, 1993: 33 (Philonthus; China).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; Italy).
— Downie and Arnett, 1996: 376 (Philonthus; characters; USA [Note: Probably Misidentified]).
▴ Distribution: Morocco, Europe, Russia, Georgia, Azerbaijan, Turkey, Syria, Iran, Afghanistan, Turkmenistan, Uzbekistan, China.
metallicus Gravenhorst, 1802: 168 (Staphylinus; Type locality: probably from near Brunsvigae).
— Latreille, 1804: 323 (Staphylinus; characters; Germany; France).
— Gravenhorst, 1806: 86 (Staphylinus; characters; France).
— Erichson, 1839a: 450 (Philonthus; synonym of atratus).
— Heer, 1839: 259 (Philonthus; synonym of atratus).
— Erichson, 1840: 439 (Philonthus; synonym of atratus).
— Redtenbacher, 1849: 700 (Philonthus; synonym of atratus).
— Rosenhauer, 1856: 71 (Philonthus; synonym of atratus).
— Kraatz, 1857d: 586 (Philonthus; synonym of atratus).
— Ganglbauer, 1895: 446 (Philonthus; synonym of atratus).
— Bernhauer and Schubert, 1914: 329 (Philonthus; synonym of atratus).
— Smetana, 1958: 168 (Philonthus; synonym of atratus).
— Coiffait, 1974: 248 (Philonthus; synonym of atratus).
lucens Mannerheim, 1830: 28 (Staphylinus; Type locality: Petropoli; Scania).
— Mannerheim, 1831: 442 (Staphylinus; Russia; Sweden).
— Runde, 1835: 7 (Staphylinus; characters; Germany).
— Nordmann, 1837: 81 (Philonthus; characters; Sweden; Prussia).
— Heer, 1839: 260 (Philonthus; characters; Switzerland).
— Stephens, 1839: 395 (Philonthus; characters; England).
— Jacobsen, 1849: 690 (Philonthus; characters; Denmark).
— Hochhuth, 1849: 135 (Philonthus; characters).
— Redtenbacher, 1849: 701 (Philonthus; characters; Austria).
— Küster, 1853a: 51 (Philonthus; characters; Germany).
— Fairmaire and Laboulbène, 1856: 518 (Philonthus; characters; France).
— Motschulsky, 1858d: 68 (Philonthus; synonym of atratus).
— Kraatz, 1857d: 582 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 191 (Philonthus; characters; Austria).
— Thomson, 1870: 126 (Philonthus; characters; Sweden).
— Redtenbacher, 1874: 210 (Philonthus; characters; Austria).
— J. Sahlberg, 1876: 16 (Philonthus; synonym of atratus).
— Ganglbauer, 1895: 446 (Philonthus; synonym of atratus).
— Bernhauer and Schubert, 1914: 329 (Philonthus; synonym of atratus).
— Johansen, 1914: 397 (Philonthus; characters; Denmark).
— Smetana, 1958: 168 (Philonthus; synonym of atratus).
— Coiffait, 1974: 248 (Philonthus; synonym of atratus).
picipes Stephens, 1832: 221 (Quedius; [Note: Preoccupied secondary homonym in Quedius]; Type locality: London; Suffolk).
— Stephens, 1839: 391 (Quedius; characters; England).
— Gemminger and Harold, 1868: 585 (Philonthus; synonym of atratus).
— Ganglbauer, 1895: 446 (Philonthus; synonym of atratus).
— Bernhauer and Schubert, 1914: 329 (Philonthus; synonym of atratus).
— Smetana, 1958: 168 (Philonthus; synonym of atratus).
— Coiffait, 1974: 248 (Philonthus; synonym of atratus).
pilipes Stephens, 1832: 231 (Philonthus; Type locality: London; Suffolk; Devonshire).
— Stephens, 1839: 396 (Philonthus; characters; England).
— G. Waterhouse, 1858: 24 (Philonthus; synonym of atratus).
janthinipennis Kolenati, 1846: 19 (Philonthus; Type locality: Caucasus Lhesgicus).
— Hochhuth, 1849: 137 (Philonthus; characters; Iberia; Lenkoran).
— Kraatz, 1857d: 586 (Philonthus; synonym of atratus).
— Fauvel, 1869: 488 (Philonthus; synonym of atratus).
— Ganglbauer, 1895: 446 (Philonthus; synonym of atratus).
— Bernhauer and Schubert, 1914: 329 (Philonthus; synonym of atratus).
— Gusmann, 1925: 57 (Philonthus; synonym of caerulescens, cited as coerulescens).
— Smetana, 1958: 168 (Philonthus; aberration of atratus).
— Smetana, 1959: 147 (Philonthus; aberration of atratus; characters).
— Coiffait, 1967a: 393 (Philonthus; aberration of atratus; characters).
— Coiffait, 1974: 248 (Philonthus; variety of atratus).
pseudocoerulescens Gusmann, 1925: 57 (Philonthus; proposed as subspecific name for misidentified aberration atratus ab. coerulescens listed in L. Benick, 1921e: 82; Type locality: Schönberg).
— Scheerpeltz, 1933: 1333 (Philonthus; synonym of atratus).
— Smetana, 1958: 168 (Philonthus; synonym of atratus).
— Coiffait, 1974: 248 (Philonthus; synonym of atratus).
megatlasicus C. Koch, 1941: 61 (Philonthus; subspecies of atratus; Type locality: Maroc: Haut Atlas).
— Jarrige, 1952a: 132 (Philonthus; subspecies of atratus; characters; Morocco).
— Smetana, 1958: 168 (Philonthus; synonym of atratus).
— Coiffait, 1974: 248 (Philonthus; synonym of atratus).
atriceps Horn, 1884, see: rufulus Horn, 1884.
atriceps Sharp, 1885, see: peratriceps Scheerpeltz, 1933.
atricollis Bernhauer, 1936a: 21 (Pseudophilonthus; Type locality: Lulua: Kapanga).
atricolor Sharp, 1885: 426 (Philonthus; Type locality: Mexico, Capulalpam, Cerro de Plumas; Guatemala, Totonicapam 8500 to 10,000 feet).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Navarrete-Heredia, 1995: 96 (Philonthus; Mexico; Guatemala).
▴ Distribution: Mexico, Guatemala.
atricoxis Cameron, 1943: 131 (Philonthus; Type locality: Kashmir: Gulmarg, altitude 8000–10,000 feet).
atrolucens Casey, 1915, see: Bisnius.
audanti Blackwelder, 1943: 406 (Philonthus; Type locality: Haiti: 5 miles south of St. Marc).
auripennis Bernhauer, 1916c: 37 (Philonthus; Type locality: Bolivien).
auropilosus Luze, 1912, see: Rabigus.
auroscutatus Fauvel, 1878f: 261 (Philonthus; Type locality: Nouvelle-Guinée, Hatam).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Cameron, 1924: 87 (Philonthus; New Guinea).
▴ Distribution: New Guinea.
aurosetosus Scheerpeltz, 1965: 205 (Philonthus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).
aurulentus Horn, 1884: 222 (Philonthus; Type locality: Vancouver; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from Washington Territory, British Columbia, Colorado, Michigan and Kentucky).
— Blatchley, 1910: 390 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Hatch, 1957: 181 (Philonthus; characters; British Columbia; Idaho; Washington; Oregon).
— Smetana, 1965a: 8 (Philonthus; characters; Alaska).
— Smetana, 1995e: 445 (Philonthus; lectotype designation: Vanc [Vancouver]; characters; notes; Alberta; British Columbia; Manitoba; Northwest Territories; Saskatchewan; Arizona; California; Colorado; Idaho; Montana; Nebraska; Nevada; New Mexico; North Dakota; Oregon; Utah; Washington; Wyoming).
— Downie and Arnett, 1996: 382 (Philonthus; characters; USA).
— Schmidt, 1999: 108 (Philonthus; food for adults and larvae used for rearing species; rearing methods).
▴ Distribution: Canada, USA.
pansatus Casey, 1915: 442 (Philonthus; Type locality: Colorado: Boulder Co.; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1355 (Philonthus; catalog).
— Smetana, 1995e: 445 (Philonthus; lectotype designation: Boulder Co., Colorado; synonym of aurulentus).
ruber Hatch, 1957: 181 (Philonthus; aberration of aurulentus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Oregon: L. Abert).
austellus Herman, 2001: 46 (Philonthus; replacement name for rufipes Boheman).
rufipes Boheman, 1848: 284 (Philonthus; [preoccupied]; Type locality: Caffraria interior).
— Bernhauer and Schubert, 1914: 353 (Philonthus; catalog).
— Scheerpeltz, 1974: 143 (Philonthus; South Africa).
— Herman, 2001: 46 (Philonthus; synonym of austellus).
australicus Cameron, 1943, see: Gabronthus.
australis W. J. MacLeay, 1873, see: Hesperus.
australis Cameron, 1943a: 342 (Philonthus; [preoccupied]; Type locality: Australia: Port Darwin).
— Herman, 2001: 46 (Philonthus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Australia.
autumnalis Cameron, 1932, see: Gabrius.
avicola Bernhauer, 1942: 25 (Philonthus; Type locality: Kostarika: Reventzon, Ebene Limon, Hamburgfarm).
azabuensis R. Dvořák, 1958: 137 (Philonthus; Type locality: Japan: Tokyo, Azabu).
azuripennis Cameron, 1928c: 563 (Philonthus; Type locality: Sikkim: Tangu, 14,000 feet; [Note: See lectotype designation by Schillhammer, 2000]. Also cited originally from Tibet: Gantza, 13,000 feet; Jelap La, 12,000 feet. Sikkim: Dentang, 15,000 feet).
— Cameron, 1932: 81 (Philonthus; characters; Tibet; India).
— Scheerpeltz, 1933: 1333 (Philonthus; catalog).
— Scheerpeltz, 1976: 34 (Philonthus; Nepal).
— Scheerpeltz, 1976a: 87 (Philonthus; Nepal).
— Coiffait, 1982: 28 (Philonthus; Nepal).
— Schillhammer, 1999a: 61 (Philonthus).
— Schillhammer, 2000: 150 (Philonthus; lectotype designation; characters; China; Nepal; Bhutan).
▴ Distribution: China, Nepal, Bhutan, India.
stoetzneri Bernhauer, 1929d: 109 (Philonthus; Type locality: Szetschwan: Sunpanting).
— Scheerpeltz, 1933: 1362 (Philonthus; catalog).
— Coiffait, 1967a: 397 (Philonthus; characters; China).
— Coiffait, 1974: 279 (Philonthus; cited as stotzneri; characters; China).
— Schillhammer, 1999a: 61 (Philonthus; synonym of azuripennis).
— Schillhammer, 2000: 150 (Philonthus; synonym of azuripennis).
trisulensis Coiffait, 1982: 60 (Philonthus; Type locality: Nepal: Trisuli Tal, Gosainkund, 3200 m).
— Schillhammer, 1999a: 61 (Philonthus; synonym of azuripennis).
— Schillhammer, 2000: 150 (Philonthus; synonym of azuripennis).
bacchus Last, 1981: 121 (Philonthus; Type locality: Madang Dist. Finisterre Mts. Damanti, 3500 ft.).
badagshanensis Scheerpeltz, 1963: 11 [Note: Nomen nudum, published without description, Article 13.1.1.
badius Kiesenwetter, 1858, see: Gabrius.
badius Motschulsky, 1858, see: castaneus Gemminger and Harold, 1868.
bagdadensis Coiffait, 1967, see: irakoiranicus Scheerpeltz, 1961.
baicalensis Bernhauer, 1927, see: Bisnius.
baicalicus Bernhauer, 1935, see: coracion Peyerimhoff, 1902.
bakeri Bernhauer, 1920d: 183 (Philonthus; Type locality: Philippinen: Los Banos).
balbus Tottenham, 1955: 177 (Philonthus; Type locality: Uganda: Kawanda).
— Levasseur, 1968a: 54 (Philonthus; Central African Republic).
▴ Distribution: Uganda, Central African Republic.
balkanicus R. Dvořák and Havelka, 1952, see: frigidus Märkel and Kiesenwetter, 1848.
bambusae Cameron, 1937: 12 (Philonthus; Type locality: Java: Batoerraden; G. Slamat).
banalis Tottenham, 1962: 180 (Philonthus; Type locality: Cameroons: Bamenda).
bangoranus Levasseur, 1980: 360 (Philonthus; Type locality: République Centrafricaine: Bangonran, SW de Ndélé).
barbarae Hromádka, 1992: 98 (Philonthus; Type locality: Nepal, Tanje).
bashgulensis Scheerpeltz, 1963: 12 [Note: Nomen nudum, published without description, Article 13.1.1.
basalis Motschulsky, 1858, see: Erichsonius.
basalis Horn, 1884, see: duplicatus Bernhauer and Schubert, 1914.
basicornis Eppelsheim, 1895a: 124 (Philonthus; Type locality: Togoland).
— Eppelsheim, 1895: 200 (Philonthus; Arussi Galla; Togo).
— Fauvel, 1907: 38 (Philonthus; Kilimandjaro; Abyssinia; Gallas; Sierra Leone; Gabon; Togo).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Tottenham, 1949b: 309 (Philonthus; characters).
— Cameron, 1950g: 36 (Philonthus; Belgian Congo).
— Tottenham, 1956: 226 (Philonthus; Ruanda).
— Tottenham, 1962: 161 (Philonthus; characters).
— Levasseur, 1966a: 1211 (Philonthus; Congo; Tanzania; Kenya; Abyssinia; Sierra Leone; Gabon; Togo).
▴ Distribution: Ethiopia, Kenya, Tanzania, Ruanda, Congo, Gabon, Togo, Sierra Leone.
basilewskyi Tottenham, 1962, see: Afrorabigus.
basipennis Coiffait, 1967, see: biskrensis Fagel, 1957.
basipennis Tottenham, 1949b: 351 (Philonthus; Type locality: N.W. Rhodesia: Namwaca).
basipilosus Schubert, 1908, see: Rabigus.
basiventris Sharp, 1885, see: Belonuchus.
bassus Tottenham, 1953: 146 (Philonthus; Type locality: Batavia. Also cited from Java, New Guinea and Celebes).
batkhizensis Coiffait, 1967, see: Bisnius.
batotensis Cameron, 1932: 99 (Philonthus; Type locality: Kashmir: Udhampur division, Batote, alt. 5500 feet).
— Scheerpeltz, 1933: 1333 (Philonthus; catalog).
— Scheerpeltz, 1963: 13 (Philonthus; Afghanistan).
— Coiffait, 1983e: 478 (Philonthus; Afghanistan).
▴ Distribution: India, Afghanistan.
beccarii Fauvel, 1878f: 262 (Philonthus; Type locality: Iles Arou, Wokan).
— Fauvel, 1879: 101 (Philonthus; Dorei).
— Bernhauer and Schubert, 1914: 372 (Leucitus; catalog).
— Cameron, 1931: 363 (Philonthus; New Guinea).
— Cameron, 1931: 363 (Leucitus; New Guinea).
— Scheerpeltz, 1933: 1334 (Philonthus; catalog).
— Last, 1968: 358 (Philonthus; New Guinea).
— Last, 1975a: 1 (Philonthus).
— Last, 1987: 31 (Philonthus; Papua New Guinea).
▴ Distribution: Indonesia, New Guinea.
benigseni Bernhauer, 1927h: 293 (Leucitus; Type locality: Neu-Guinea: Kani Gebirge).
beckeri Bernhauer, 1933, see: oberti Eppelsheim, 1889.
beesoni Cameron, 1926b: 357 (Philonthus; Type locality: Simla District; Gahan, Matiana, Theog, Fagu, 7000 to 8000 feet. Chakrata District; Binal Gad, 7000 to 8000).
— Cameron, 1932: 126 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1334 (Philonthus; catalog).
— Coiffait, 1979d: 552 (Philonthus; Afghanistan).
▴ Distribution: India, Afghanistan.
belesis Tottenham, 1956: 278 (Philonthus; Type locality: Ruanda: Rutovu, forêt du Rugege, 2350 m).
belingaensis Levasseur, 1980: 365 (Philonthus; Type locality: Gabon: Belinga).
bellator Tottenham, 1956: 281 (Philonthus; Type locality: Ruanda: Biumba, 2300 m).
bellatulus Last, 1981: 119 (Philonthus; Type locality: Goroka).
bellus Kraatz, 1859, see: paederoides Motschulsky, 1858.
belonuchoides Cameron, 1918: 88 (Philonthus; Type locality: Singapore: Keppel Harbour).
— Cameron, 1921a: 377, 405 (Philonthus; characters; Singapore).
— Scheerpeltz, 1933: 1334 (Philonthus; catalog).
— Cameron, 1933d: 387 (Philonthus; Sumatra).
— Cameron, 1937: 5 (Philonthus; Java).
▴ Distribution: Singapore, Indonesia.
bengalensis Bernhauer, 1911: 91 (Philonthus; Type locality: W. Bengal: Paresnath, 4000′).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Cameron, 1932: 87 (Philonthus; characters; Bengal; India).
— Scheerpeltz, 1933: 1334 (Philonthus; catalog).
▴ Distribution: India.
benigseni Bernhauer, 1927, see: beccarii Fauvel, 1878.
bernardinensis Fall, 1930, see: Gabrius.
bernhaueri Csiki, 1901, see: rectangulus Sharp, 1874.
bernhaueri Roubal, 1909, see: japonicus Sharp, 1874.
berytensis Jarrige, 1951a: 10 (Philonthus; Type locality: Beyrouth, Syrie).
— Coiffait, 1967a: 429 (Philonthus; characters; Lebanon).
— Coiffait, 1974: 208 (Philonthus; characters; Lebanon; Palestine).
▴ Distribution: Lebanon, Israel.
bestialis Bernhauer, 1939, see: longicornis Stephens, 1832.
besucheti Smetana, 1977: 795 (Philonthus; Type locality: Turquie Adiyaman, Golbasi, 900 m).
bhutanicus Coiffait, 1977, see: obsoletus Eppelsheim, 1895.
bhutanensis Herman, 2001: 46 (Philonthus; replacement name for gabrioides).
gabrioides Coiffait, 1984e: 376 (Philonthus; [preoccupied]; Type locality: Bhutan: Punakka, 1560 m).
bicaudus Sharp, 1889: 36 (Philonthus; Type locality: Niigata).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Scheerpeltz, 1933: 1334 (Philonthus; catalog).
▴ Distribution: Japan.
bicolor Redtenbacher, 1849, see: Quedius.
bicolor Fauvel, 1903e: 240 (Philonthus; [preoccupied]; Type locality: Cameroun).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Tottenham, 1949b: 306 (Philonthus; characters).
— Tottenham, 1962: 160 (Philonthus; characters).
— Herman, 2001: 46 (Philonthus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Cameroon.
bicolor Drugmand, 1990, see: biskrensis Fagel, 1957.
bicoloratus Cameron, 1931: 361 (Philonthus; Type locality: Finsch Haven, New Guinea).
— Scheerpeltz, 1933: 1334 (Philonthus; catalog).
— Last, 1968: 367 (Philonthus; New Guinea).
— Last, 1987: 31 (Philonthus; Papua New Guinea).
▴ Distribution: New Guinea.
bicoloripennis Bernhauer, 1915k: 302 (Philonthus; subgenus Pseudophilonthus; Type locality: Portugiesisch-Guinea: Bolama).
— Scheerpeltz, 1933: 1334 (Philonthus; catalog).
— Tottenham, 1949b: 324 (Philonthus; characters).
▴ Distribution: Guinea-Bissau.
bidentatus Horn, 1884, see: Gabrius.
bidentiformis Tottenham, 1949, see: pollux Fauvel, 1905.
biguttulus Bernhauer, 1908, see: kamerunensis Bernhauer and Schubert, 1914.
biguttulus Fauvel, 1907: 44 (Philonthus; variety of imitator; Type locality: Abyssinie; Escarpment).
— Bernhauer and Schubert, 1914: 355 (Philonthus; variety of sequens).
— Rambousek, 1922: 1009 (Philonthus; variety of sequens; British East Africa).
— Cameron, 1950g: 36 (Philonthus; variety of sequens; Ruanda).
— Tottenham, 1962: 223, 226 (Philonthus; valid species; characters).
▴ Distribution: Ethiopia, Kenya, Rwanda.
bilineatus Erichson, 1840, see: Cafius.
bimaculatus Gravenhorst, 1802, see: nitidicollis Lacordaire, 1835.
bimaculatus Marsham, 1802, see: varians Paykull, 1789.
binderi Roubal, 1910, see: japonicus Sharp, 1874.
binotatus Gravenhorst, 1806: 73 (Staphylinus; [preoccupied]; Type locality: Borussia).
— Erichson, 1840: 506 (Philonthus; characters; Prussia).
— Redtenbacher, 1849: 825 (Philonthus; characters; Austria).
— Kraatz, 1857d: 619 (Philonthus; variety of punctus).
— Redtenbacher, 1857: 197 (Philonthus; characters; Austria).
— Redtenbacher, 1874: 216 (Philonthus; characters; Austria).
— Ganglbauer, 1895: 458 (Philonthus; aberration of punctus).
— Bernhauer and Schubert, 1914: 352 (Philonthus; variety of punctus).
— Scheerpeltz, 1933: 1357 (Philonthus; variety of punctus).
— V. Hansen, 1952: 79 (Philonthus; subgenus Philonthus; characters; Denmark).
— Palm, 1952: 117 (Philonthus; characters; notes).
— Horion, 1954: 10 (Philonthus; characters; Germany).
— Lohse, 1956: 91 (Philonthus; characters; notes).
— Smetana, 1958: 225 (Philonthus; subgenus Philonthus; characters; ecology).
— Smetana, 1959: 170 (Philonthus; characters).
— Palm, 1963b: 97 (Philonthus; characters; Sweden; Denmark; Finland).
— Lohse, 1964: 186 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 176 (Philonthus; Russia; Sweden; Denmark; Holland; Germany; Austria).
— Coiffait, 1967a: 432 (Philonthus; characters; Europe).
— Smetana, 1967d: 208 (Philonthus; form of binotatus cited from Mongolia).
— Scheerpeltz, 1968a: 67 (Philonthus; catalog; Austria).
— Tikhomirova, 1973a: 184 (Philonthus; in checklist for USSR).
— Coiffait, 1974: 214 (Philonthus; characters; Denmark; Scandinavia; Siberia; Mongolia).
— Smetana 1975c: 172 (Philonthus; form of binotatus cited from Mongolia).
— Shilov, 1976: 153 (Philonthus; Mongolia).
— Muona, 1979: 14 (Philonthus; Sweden; Denmark).
— Burakowski, Mroczkowski, and Stefańska, 1980: 41 (Philonthus; catalog; Poland).
— Bangsholt, 1981: 69 (Philonthus; Denmark).
— Lucht, 1987: 105 (Philonthus; Germany; Austria; Denmark).
— Mahler, 1987: 198 (Philonthus; Denmark).
— Schülke and Uhlig, 1988: 11 (Philonthus; notes; Germany).
— Silfverberg, 1992: 16 (Philonthus; Sweden; Denmark; Estonia).
— Smetana, 1995e: 9, 445 (Philonthus; notes).
— M. Hansen, 1996: 106 (Philonthus; checklist; Denmark).
— Hansen, Jørum, Palm, and Pedersen, 1997: 128 (Philonthus; checklist; Denmark).
— Hansen, Pedersen, and Pritzl, 1999: 81 (Philonthus; Denmark).
— Burakowski, Mroczkowski, and Stefańska, 2000: 50 (Philonthus; subgenus Philonthus; catalog; Poland).
— Herman, 2001: 46 (Philonthus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Europe, Russia, Mongolia.
hanseni Palm, 1952: 120 (Philonthus; aberration of binotatus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]).
— Smetana, 1958: 225 (Philonthus; synonym of binotatus).
— Smetana, 1959: 170 (Philonthus; aberration of binotatus; characters).
— Coiffait, 1967a: 434 (Philonthus; aberration of binotatus; characters).
— Smetana, 1967d: 208 (Philonthus; form of binotatus; Mongolia).
— Coiffait, 1974: 214 (Philonthus; variety of binotatus).
— Smetana 1975c: 172 (Philonthus; form of binotatus; Mongolia).
biparamerosus Tottenham, 1949, see: Erichsonius.
bipartitus Cameron, 1937a: 104 (Philonthus; Type locality: New Guinea: Kokoda, alt. 1200 ft.).
bipunctatus Bernhauer, 1911, see: industanus Fauvel, 1903.
birmanus Fauvel, 1895: 261 (Philonthus; Type locality: Bírmanie, Carin Cheba, 900–1100 m).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Cameron, 1932: 85 (Philonthus; characters; Burma; India).
— Scheerpeltz, 1933: 1334 (Philonthus; catalog).
— Cameron, 1933d: 386 (Philonthus; Sumatra).
— Jarrige, 1951: 133 (Philonthus; Tonkin).
— Yadav, Chand, and Yadav, 1995: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
▴ Distribution: Indonesia, Vietnam, Myanmar, India.
biseriatus Stierlin, 1875, see: Quedius.
bishanus Tottenham, 1955: 166 (Philonthus; subgenus Spatulonthus; Type locality: Transvaal: Johannesburg).
bisignatus Boheman, 1848: 282 (Philonthus; Type locality: Portus Natalensis).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Bernhauer, 1931d: 587 (Philonthus; Abyssinia).
— Scheerpeltz, 1933: 1335 (Philonthus; catalog).
— Bernhauer and Chapman, 1939: 66 (Philonthus; Kenya).
— Tottenham, 1949b: 345 (Philonthus; characters).
— Cameron, 1950g: 37 (Philonthus; Belgian Congo).
— Tottenham, 1955: 158 (Philonthus; subgenus Spatulonthus; notes; characters; southeastern Africa).
— Tottenham, 1961: 205 (Philonthus; subgenus Spatulonthus; Senegal).
— Tottenham, 1962: 139 (Philonthus; subgenus Spatulonthus; Tanganyika).
— Coiffait, 1968a: 136 (Philonthus; Chad).
— Scheerpeltz, 1974: 146 (Philonthus; South West Africa; South Africa).
▴ Distribution: Senegal, Chad, Ethiopia, Kenya, Tanzania, Congo, Namibia, South Africa.
piceicoxis Eppelsheim, 1895a: 127 (Philonthus; Type locality: Togoland).
— Fauvel, 1903e: 241 (Philonthus; synonym of peregrinus).
— Fauvel, 1904a: 60 (Philonthus; synonym of peregrinus).
— Bernhauer and Schubert, 1914: 350 (Philonthus; catalog).
— Cameron, 1932: 133 (Philonthus; synonym of peregrinus).
— Scheerpeltz, 1933: 1335 (Philonthus; variety of bisignatus).
pustulatus Bernhauer, 1915d: 140 (Philonthus; variety of peregrinus; Type locality: Deutsch-Ostafrika: Kaguru; Irangi; an Bachufern; Moshi; Shirati, Boma Gombe).
bisinuatus Eppelsheim, 1889a: 175 (Philonthus; Type locality: Sze-tschuan).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Scheerpeltz, 1933: 1335 (Philonthus; catalog).
— Coiffait, 1967a: 393 (Philonthus; characters; Mongolia; [Note: Erroneous record according to Schillhammer, 2000: 140]).
— Coiffait, 1974: 224 (Philonthus; characters; China; Mongolia; [Note: Mongolia is erroneous record according to Schillhammer, 2000: 140]).
— Shilov, 1976: 148 (Philonthus; Mongolia; [Note: Erroneous record according to Schillhammer, 2000: 140]).
— Schillhammer, 2000: 139 (Philonthus; characters; China; records in Mongolia are erroneous).
▴ Distribution: China.
biskrensis Fagel, 1957e: 332 (Philonthus; Type locality: Algérie: oasis de Biskra, route de Taoggourt).
— Coiffait, 1967a: 431 (Philonthus; characters; Algeria).
— Coiffait, 1974: 197 (Philonthus; characters; Algeria).
— Coiffait, 1974: 330 (Neobisnius; [Note: Misidentified as hauseri according to Gusarov, 1995a: 93 [= 1996: 55]; characters; Buchara; Kazakstan).
— Maachi, 1991: 251 (Philonthus; Morocco).
— Gusarov, 1995a: 93 [= 1996: 55] (Philonthus; characters; synonymic notes; Algeria; Iran; France; Turkmenia; Kazakhstan; Uzbekistan; Mongolia).
▴ Distribution: France, Morocco, Algeria, Iran, Turkmenistan, Uzbekistan, Kazakhstan, Mongolia.
iranicus Scheerpeltz, 1961: 22 (Neobisnius; Type locality: Iran: Khuzistan, Shush, [Susa]).
— Coiffait, 1974: 330 (Neobisnius; characters; Iran).
— Gusarov, 1995a: 93 [= 1996: 55] (Philonthus; lectytype designation; synonym of biskrensis).
basipennis Coiffait, 1967a: 439 (Philonthus; subgenus Philonthus; [preoccupied]; Type locality: environs de Maku dans le Nord-Ouest de l'Iran).
— Coiffait, 1974: 197 (Philonthus; characters; Iran).
— Gusarov, 1995a: 93 [= 1996: 55] (Philonthus; synonym of biskrensis).
bicolor Drugmand, 1990: 213 (Neobisnius; [preoccupied]; Type locality: Turkmenistan: Repetek).
bistriatus Erichson, 1840, see: Cafius.
bituminosus† Heyden and Heyden, 1866: 135 (Philonthus; Oligocene fossil; Type locality: Braunkohle des Siebengebirges).
— Scudder, 1891: 566 (Philonthus; catalog; Aquitanian; Prussia).
— Handlirsch, 1907: 726 (Philonthus; catalog; Oligocene; Rheinland).
▴ Distribution: Germany.
blossius Smetana, 1995e: 363 (Philonthus; Type locality: Quebec: Pte. Gatineau. Also cited from Ontario, Massachusetts, New Jersey, Wisconsin).
bodemeyeri Eppelsheim, 1881, see: caerulescens Lacordaire, 1835.
bojeri† Heer, 1856: 17 (Philonthus; Type locality: Aix, Provence; Fossil).
— Oustalet, 1874: 169 (Philonthus; characters; France).
— Scudder, 1891: 566 (Philonthus; catalog; Ligurian; France).
— Handlirsch, 1907: 725 (Philonthus; cited as boyeri; catalog; Oligocene; France).
▴ Distribution: France.
bolivianus Bernhauer, 1916c: 35 (Philonthus; Type locality: Bolivien: Coroico, 1800 m).
bonariensis Bernhauer, 1909c: 245 (Philonthus; Type locality: Argentinien: Buenos Aires).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Frank, 1982: 38 (Philonthus; list of parasites).
▴ Distribution: Argentina.
bonus Tottenham, 1962: 207 (Philonthus; Type locality: Nigeria: Potiskum).
borneensis Bernhauer, 1915i: 254 (Philonthus; Type locality: Sarawak).
— Bernhauer, 1928: 16 (Philonthus; Sumatra).
— Scheerpeltz, 1933: 1335 (Philonthus; catalog).
— Cameron, 1933d: 386 (Philonthus; Sumatra).
— Frank, 1982: 38 (Philonthus; list of parasites).
▴ Distribution: Malaysia, Indonesia.
boreas Smetana, 1995e: 450 (Philonthus; Type locality: Northwest Territories: Inuvik. Also cited from Alberta, British Columbia, Manitoba, Newfoundland, Yukon Territory, Alaska, Colorado, Nevada).
bos Tottenham, 1962: 177 (Philonthus; Type locality: Natal: Nqutu).
bothellensis Hatch, 1957, see: carbonarius Gravenhorst, 1802.
bottegoi Eppelsheim, 1895: 207 (Philonthus; Type locality: Ganale Gudda, Arussi Galla).
boucardi Solsky, 1868, see: fibularius Erichson, 1840.
brachypterus Solsky, 1872: 297 (Philonthus; Type locality: Cayenne).
brasilianus Bernhauer, 1916c: 40 (Philonthus; Type locality: Brasilien: S. Paulo, Ypiranga).
brevicornis Gravenhorst, 1802, see: ebeninus Gravenhorst, 1802.
brevipennis Horn, 1884, see: Gabrius.
brevis Melsheimer, 1844, see: sericans Gravenhorst, 1802.
brevithorax Bernhauer, 1934: 8 (Philonthus; Type locality: Szechuan: Nitou Tatsienlu).
— Schillhammer, 2000: 147 (Philonthus; characters; China; India; Nepal; Bhutan).
— Schillhammer and Smetana, 2000: 179 (Philonthus; characters; teratological features; China).
▴ Distribution: China, Nepal, Bhutan, India.
brincki Scheerpeltz, 1974: 143 (Philonthus; Type locality: Basutoland: Mokhotlong, 7200 ft.).
broumandi Boháč, 1981: 357 (Philonthus; Type locality: Khouzestan: Dezfoul, Haft-tapeh, 20 m).
brouni Duvivier, 1883, see: Bisnius.
brounianus Bernhauer and Schubert, 1914, see: novaezeelandiae Duvivier, 1883.
brunneipennis Gerhardt, 1899, see: micans Gravenhorst, 1802.
brunneipennis Quedenfeldt, 1882, see: quisquiliarius Gyllenhal, 1810.
brunneus Gravenhorst, 1802, see: sericans Gravenhorst, 1802.
brunnicollis Hochhuth, 1851, see: Rabigus.
buanaensis Last, 1968: 370 (Philonthus; Type locality: New Guinea: Buana, Sarawaged Range, 3000 ft., 20–30 miles north of Lae).
bucephalus Horn, 1884, see: perversus Horn, 1884.
bucharensis Coiffait, 1967, see: Bisnius.
buchari Boháč, 1977, see: spinipes Sharp, 1874.
buettikeri Coiffait, 1979, see: arabicus Bernhauer, 1935.
bulbifer Tottenham, 1956a: 242 (Philonthus; Type locality: Cape Colony: Simon's Town).
buphthalmus Scheerpeltz, 1957c: 234, 319 (Philonthus; Type locality: Sumba: Kodi).
burgeoni Bernhauer, 1928, see: Gabrius.
burgeonianus Bernhauer, 1934g: 237 (Pseudophilonthus; Type locality: Baudouinville).
— Levasseur, 1968a: 53 (Philonthus; Central African Republic).
▴ Distribution: Congo, Central African Republic.
buruensis Cameron, 1942c: 844 (Philonthus; Type locality: Boeroe: Ilat).
busiris Smetana, 1995e: 358 (Philonthus; Type locality: Ontario: 5 mi. NW South March. Also cited from Alberta, British Columbia, Manitoba, Ontario, Quebec, Saskatchewan, Colorado, Connecticut, Illinois, Indiana, Maine, Massachusetts, Michigan, Minnesota, Montana, Nebraska, New Hampshire, New Jersey, New York, North Dakota, Pennsylvania, Rhode Island, Washington, West Virginia, Wisconsin, Wyoming).
caedator Tottenham, 1949b: 337 (Philonthus; Type locality: Haute-Uele: Watsa).
caeruleipennis Mannerheim, 1830: 27 (Staphylinus; [Note: This name was proposed for cyanipennis from America characterized by Gravenhorst who attributed the species to Fabricius]; Type locality: America septentr.; [Note: See lectotype designation by Schillhammer, 1998]).
— Gravenhorst, 1802: 168 (Staphylinus; [Note: Misidentification: Cited as cyanipennis but according to work by Smetana, 1995e: 117 it is probably caeruleipennis]; characters; America septentr.).
— Say, 1830: 38 (Staphylinus; [Note: Misidentification: Cited as cyanipennis but according to work by Smetana, 1995e: 117 it is probably caeruleipennis]; characters).
— Mannerheim, 1831: 441 (Staphylinus; America septentrionalis).
— Say, 1834: 451 (Staphylinus; provisional identification; [Note: Misidentification: Cited as cyanipennis but according to work by Smetana, 1995e: 117 it is probably caeruleipennis]; United States).
— Nordmann, 1837: 80 (Philonthus; cited as coeruleipennis; characters; America borealis).
— Erichson, 1840: 433 (Philonthus; cited as coeruleipennis; synonym of cyanipennis).
— LeConte, 1850: 220 (Philonthus; [Note: Misidentification: Cited as cyanipennis but according to work by Smetana, 1995e: 117 it is probably caeruleipennis]; Lake Superior).
— Kraatz, 1857d: 574 (Philonthus; cited as coeruleipennis; synonym of cyanipennis).
— Provancher, 1877: 249 (Philonthus; characters; [Note: Misidentification: Cited as cyanipennis but according to work by Smetana, 1995e: 117 it is probably caeruleipennis]; Quebec).
— Horn, 1884: 208 (Philonthus; characters; [Note: Misidentification: Cited as cyanipennis but according to work by Smetana, 1995e: 117 it is probably caeruleipennis]; eastern United States).
— Sharp, 1889: 35 (Philonthus; [Note: Misidentification: Cited as cyanipennis but according to work by Smetana, 1995e: 117 it is probably caeruleipennis]; Japan).
— Ganglbauer, 1895: 444 (Philonthus; cited as coeruleipennis; synonym of cyanipennis).
— Blatchley, 1910: 387 (Philonthus; characters; [Note: Misidentification: Cited as cyanipennis but according to work by Smetana, 1995e: 117 it is probably caeruleipennis]; Indiana).
— Bernhauer and Schubert, 1914: 334 (Philonthus; cited as coeruleipennis; synonym of cyanipennis).
— Hatch, 1957: 195 (Philonthus; characters; [Note: Misidentification: Cited as cyanipennis but according to work by Smetana, 1995e: 117 it is probably caeruleipennis]; British Columbia).
— Smetana, 1958: 161 (Philonthus; cited as coeruleipennis; synonym of cyanipennis).
— Nakane, 1963: A239 (Philonthus; [Note: Misidentification: Cited as cyanipennis but according to work by Smetana, 1995e: 117 it is probably caeruleipennis]; Japan).
— Nakane, 1963a: 90 (Philonthus; characters; habitus photograph; [Note: Misidentification: Cited as cyanipennis but according to work by Smetana, 1995e: 117 it is probably caeruleipennis]; Japan).
— Watanabe and Shibata, 1965: 321 (Philonthus; subgenus Philonthus; [Note: Misidentification: Cited as cyanipennis but according to work by Smetana, 1995e: 117 it is probably caeruleipennis]; Japan).
— Coiffait, 1974: 225 (Philonthus; synonym of cyanipennis).
— Dearborn and Donahue, 1993: 15 (Philonthus; [Note: Misidentification: Cited as cyanipennis but according to work by Smetana, 1995e: 117 it is probably caeruleipennis]; Maine).
— Hayashi, 1994: 121 (Philonthus; characters; [Note: Misidentification: Cited as cyanipennis but according to work by Smetana, 1995e: 117 it is probably caeruleipennis]; Japan).
— Smetana, 1995e: 117 (Philonthus; [Note: Cited as cyanipennis but in a footnote Smetana reported that cyanipennis is restricted to Eurasia and caeruleipennis is a valid species from Japan and North America]; characters; Alabama; Arkansas; Connecticut; Georgia; Illinois; Indiana; Iowa; Kansas; Kentucky; Louisiana; Maine; Maryland; Massachusetts; Michigan; Minnesota; Mississippi; Missouri; Nebraska; New Hampshire; New Jersey; New York; North Carolina; Ohio; Oklahoma; Pennsylvania; South Carolina; South Dakota; Tennessee; Texas; Vermont; Virginia; West Virginia; Wisconsin).
— Levesque and Levesque, 1996: 292 (Philonthus; [Note: Cited as cyanipennis but according to Smetana, 1995e: 117 all the records in America are actually caeruleipennis]; seasonal abundance; Quebec).
— Downie and Arnett, 1996: 380 (Philonthus; [Note: Cited as cyanipennis but according to Smetana, 1995e: 117 all the records in America are actually caeruleipennis]; characters; USA).
— Schillhammer, 1998: 105 (Philonthus; valid species; lectotype designation; characters; Japan; Russia).
— Schmidt, 1999: 108 (Philonthus; food for adults and larvae used for rearing species; rearing methods).
▴ Distribution: eastern Russia, Japan, Canada, USA.
laterangulus Schillhammer, 1998: 106 (Philonthus; subspecies of caeruleipennis; Type locality: USSR: Primorskij Kraj, Arsenev env.).
caerulescens Lacordaire, 1835: 393 (Staphylinus; Type locality: Paris).
— Erichson, 1839a: 450 (Philonthus; synonym of atratus).
— Erichson, 1840: 439 (Philonthus; synonym of atratus).
— Fauvel, 1869: 488 (Philonthus; synonym of atratus).
— Eppelsheim, 1892: 332 (Philonthus; Turkestan).
— Ganglbauer, 1895: 446 (Philonthus; aberration of atratus).
— Reitter, 1909: 126 (Philonthus; cited as coerulescens; characters; Germany).
— Bernhauer and Schubert, 1914: 332 (Philonthus; cited as coerulescens; catalog).
— Gusmann, 1925: 57 (Philonthus; valid species).
— Scheerpeltz, 1933: 1337 (Philonthus; cited as coerulescens; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Suomalainen, 1941 (Philonthus; characters; Finland).
— Smetana, 1958: 169 (Philonthus; subgenus Philonthus; cited as coerulescens; characters; ecology; Czechoslovakia).
— Smetana, 1959: 147 (Philonthus; characters).
— Lohse, 1964: 177 (Philonthus; subgenus Philonthus; cited as coerulescens; characters; central Europe).
— Horion, 1965: 130 (Philonthus; cited as coerulescens; France; Spain; Switzerland; Italy; Slovenia; Bosnia Hercegovina; Montenegro; Romania; Czech Republic; Slovakia; Poland; Ukraine; Austria; Germany; Caucasus).
— Coiffait, 1967a: 393 (Philonthus; cited as coerulescens; characters; Europe).
— Raianu, 1968a: 464 (Philonthus; cited as coerulescens; Romania).
— Coiffait, 1974: 249 (Philonthus; cited as coerulescens; characters; Europe; Scandinavia; Britain).
— Bordoni and Carfi, 1978 (Philonthus; cited as coerulescens; note on peg setae of aedeagus).
— Tóth, 1984: 49 (Philonthus; cited as coerulescens; characters).
— Lucht, 1987: 105 (Philonthus; cited as coerulescens; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France).
— Lohse and Lucht, 1989: 165 (Philonthus; cited as coerulescens; characters).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; cited as coerulescens; Italy).
▴ Distribution: Europe, Caucasus.
bodemeyeri Eppelsheim, 1881a: 376 (Philonthus; Type locality: Mehadia in Süd-Ungarn).
— Ganglbauer, 1895: 446 (Philonthus; characters; Austria; Hungary; Caucasus; Spain).
— Bernhauer and Schubert, 1914: 332 (Philonthus; synonym of coerulescens).
— Gusmann, 1925: 57 (Philonthus; synonym of caerulescens, cited as coerulescens).
— Scheerpeltz, 1933: 1337 (Philonthus; synonym of caerulescens, cited as coerulescens).
— Smetana, 1958: 169 (Philonthus; synonym of caerulescens, cited as coerulescens).
— Coiffait, 1974: 249 (Philonthus; synonym of caerulescens).
caffer Boheman, 1848: 280 (Philonthus; Type locality: Caffraria orientalis).
— Fauvel, 1907: 45 (Philonthus; Nairobi; Orange; Transvaal; Cape).
— Bernhauer, 1908c: 110 (Philonthus; German Southwest Africa).
— Bernhauer and Schubert, 1914: 331 (Philonthus; catalog).
— Bernhauer, 1915n: 319 (Philonthus; German Southwest Africa).
— Bernhauer, 1940b: 635 (Philonthus; subgenus Cephalonthus).
— Bernhauer, 1942c: 361 (Philonthus; Cameroon).
— Tottenham, 1949b: 333 (Philonthus; characters).
— Tottenham, 1962: 223, 225 (Philonthus; characters).
— Scheerpeltz, 1974: 145 (Philonthus; South Africa; Basutoland; South West Africa).
▴ Distribution: Kenya, Cameroon, Namibia, South Africa, Lesotho.
caffer Sachse, 1852: 142 (Philonthus; [preoccupied]; Type locality: Vorgeb. d. g. Hoff-nung).
— Fauvel, 1907: 45 (Philonthus; synonym of caffer Boheman).
— Bernhauer and Schubert, 1914: 331 (Philonthus; synonym of caffer).
capensis Gemminger and Harold, 1868: 586 (Philonthus; replacement name for caffer Sachse).
— Fauvel, 1907: 45 (Philonthus; synonym of caffer Boheman).
— Bernhauer and Schubert, 1914: 331 (Philonthus; synonym of caffer).
caffer Sachse, 1852, see: caffer Boheman, 1848.
cailleuxi Levasseur, 1980: 363 (Philonthus; Type locality: République Centrafricaine: Bangonran, SW de Ndélé).
calidus Harold, 1881, see: natalensis Boheman, 1848.
caliensis Bernhauer, 1916c: 33 (Philonthus; Type locality: West-Columbien: Umgebung von Cali in Caucatale).
californicus Mannerheim, 1843, see: furvus Nordmann, 1837.
callosipennis Scheerpeltz, 1960c: 100 (Philonthus; Type locality: Bolivien: Cochabamba, in den Yungas de Palmar, 2000 m).
cambeforti Lecoq, 1989: 93 (Philonthus; Type locality: Lamto, Côte d'Ivoire).
cameroni Scheerpeltz, 1933: 1335 (Philonthus; replacement name for fraternus Cameron).
fraternus Cameron, 1926b: 354 (Philonthus; [preoccupied]; Type locality: Simla Hills; Kotgarh, 7000 feet).
— Cameron, 1932: 121 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1335 (Philonthus; synonym of cameroni).
cameroni Pajni and Kohli, 1977, see: malcolmi Herman, 2001.
cameronianus Scheerpeltz, 1933: 1335 (Philonthus; replacement name for rufiventris Cameron).
rufiventris Cameron, 1918b: 170 (Philonthus; [preoccupied]; Type locality: Ceylon, Kandy).
— Cameron, 1932: 122 (Philonthus; characters; Ceylon).
— Scheerpeltz, 1933: 1335 (Philonthus; synonym of cameronianus).
canariensis Fauvel, 1898, see: Gabrius.
candens Erichson, 1840, see: Belonuchus.
candidus Last, 1968, see: Hesperus.
canescens Mäklin, 1852, see: Cafius.
cantonneti Levasseur, 1962a: 54 (Philonthus; Type locality: Kolwezi, Katanga).
capeneri Last, 1953: 115 (Philonthus; Type locality: Natal: Mtunzini).
— Tottenham, 1962: 138, 183 (Philonthus; characters; notes; Uganda).
▴ Distribution: Uganda, South Africa.
capensis Gemminger and Harold, 1868, see: caffer Boheman, 1848.
capitalis Sharp, 1876, see: Paederomimus.
capito Cameron, 1952: 248 (Philonthus; Type locality: New Guinea: Snow Mountains: Mist Camp, 1800 m).
— Last, 1968: 372 (Philonthus; New Guinea).
— Last, 1987: 31 (Philonthus; Papua New Guinea).
▴ Distribution: New Guinea.
capizzii Hatch, 1957, see: Gabrius.
captivus Levasseur, 1966b: 1497 (Philonthus; Type locality: région de Kolwezi, Katanga, Musonoie).
carbonarius Gravenhorst, 1802: 23 (Staphylinus; Type locality: Germany; [Note: See lectotype designation by Smetana, 1995e]. Originally no locality cited).
— Latreille, 1804: 313 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 67 (Staphylinus).
— C. Sahlberg, 1830: 321 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 28 (Staphylinus; Finland).
— Mannerheim, 1831: 442 (Staphylinus; Finland).
— Stephens, 1832: 230 (Philonthus; characters; England).
— Lacordaire, 1835: 393 (Staphylinus; characters; France).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Nordmann, 1837: 82 (Philonthus; Sweden; Germany).
— Erichson, 1839a: 455 (Philonthus; synonym of varius).
— Stephens, 1839: 395 (Philonthus; characters; England).
— Erichson, 1840: 447 (Philonthus; synonym of varius).
— Jacobsen, 1849: 688 (Philonthus; characters; Denmark).
— Hochhuth, 1849: 133 (Philonthus; characters).
— Hochhuth, 1849: 136 (Philonthus; synonym of varius).
— Redtenbacher, 1849: 824 (Philonthus; characters; Austria).
— Fairmaire and Laboulbène, 1856: 516 (Philonthus; characters; France).
— Kraatz, 1857d: 584 (Philonthus; synonym of varius).
— Fauvel, 1869: 489 (Philonthus; synonym of varius).
— Fauvel, 1874: 217 [= 1874a: 443] (Philonthus; characters; France; Italy).
— Ganglbauer, 1895: 448 (Philonthus; synonym of varius).
— Montandon, 1908: 80 (Philonthus; Romania).
— Bernhauer and Schubert, 1914: 359 (Philonthus; synonym of varius).
— Cameron, 1928c: 563 (Philonthus; Sikkim).
— Portevin, 1929: 359 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1335 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Aleksandrov, 1934: 153 (Philonthus; China).
— Tottenham, 1939d: 231 (Philonthus; suggests abandoning name).
— Scheerpeltz, 1944a: 145, 145 (Philonthus; illustrations).
— V. Hansen, 1952: 52 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1958: 166 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 148 (Philonthus; characters).
— Scheerpeltz, 1961d: 128 (Philonthus; Italy).
— Smetana, 1962a: 131 (Philonthus; larval characters).
— Palm, 1963b: 68 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 178 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 139 (Philonthus; widespread in northern and central Europe to southern Europe; Portugal; Bosnia Hercegovina; Serbia).
— Pototskaia, 1966: 158 (Philonthus; larval characters).
— Pototskaia, 1967: 63 (Philonthus; larval characters).
— Coiffait, 1967a: 400 (Philonthus; characters).
— Raianu, 1968a: 462 (Philonthus; Romania).
— Szujecki, 1968a: 727 (Philonthus; Poland).
— Coiffait, 1974: 252 (Philonthus; characters; Palaearctic region).
— Osella and Zanetti, 1975: 109 (Philonthus; collected from nest of Talpa europaea; notes; Italy).
— Shilov, 1975a: 375 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Scheerpeltz, 1976a: 87 (Philonthus; Nepal).
— Pace and Zanetti, 1977: 293 (Philonthus; Italy).
— Topp, 1979: 6 (Philonthus; development; diapause).
— Grigo and Topp, 1980: 22 (Philonthus; influence of temperature and photoperiod on oxygen consumption in diapause and non-diapause).
— Boller, 1983: 120 (Philonthus; larval characters).
— Tóth, 1984: 52 (Philonthus; characters; Hungary).
— Segers, 1987a: 180, 182 (Philonthus; cephalic, ocular, and elytral aberrations; Belgium).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Lohse and Lucht, 1989: 165 (Philonthus; note).
— Pilon and Zanetti, 1991: 62 (Philonthus; Italy).
— J. Li, 1993: 37 (Philonthus; China).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; Italy).
— Smetana, 1995e: 137 (Philonthus; lectotype designation; characters; notes; British Columbia; Manitoba; New Brunswick; Newfoundland; Nova Scotia; Ontario; Prince Edward Island; Quebec; Saskatchewan; Yukon Territory; California; Colorado; Connecticut; Delaware; Idaho; Illinois; Indiana; Louisiana; Maine; Maryland; Massachusetts; Michigan; Minnesota; Missouri; Montana; Nebraska; New Hampshire; New Jersey; New York; Ohio; Oklahoma; Oregon; Pennsylvania; Rhode Island; Utah; Virginia; Washington; West Virginia; Wisconsin).
— Levesque and Levesque, 1996: 292 (Philonthus; seasonal abundance; Quebec).
— Ryabukhin, 1999: 62 (Philonthus; catalog; habitat; distribution; northeastern Russia).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
— Marchal and Chardonnet, 2000: 71 (Philonthus; Corsica).
— Assing, 2001: 75 (Philonthus; collected from xerothermous site; Germany).
▴ Distribution: Algeria, Canary Islands, Europe, Russia, Caucasus, Turkey, Iran, Kazakhstan, Uzbekistan, India, Nepal, China, Canada, USA.
dubius Gravenhorst, 1806: 74 (Staphylinus; Type locality: Parisiae ?).
— Nordmann, 1837: 90 (Philonthus; France).
— Erichson, 1840: 448 (Philonthus; synonym of bimaculatus).
— Kraatz, 1857d: 584 (Philonthus; synonym of varius).
— Fauvel, 1869: 489 (Philonthus; synonym of varius).
— Bernhauer and Schubert, 1914: 359 (Philonthus; synonym of varius).
— Smetana, 1958: 184 (Philonthus; synonym of varius).
— Coiffait, 1974: 252 (Philonthus; synonym of carbonarius).
varius Gyllenhal, 1810: 321 (Staphylinus; Type locality: Suecia).
— Gravenhorst, 1802: 20 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 359 cited varians Gravenhorst as a synonym of varius; Gravenhorst did not describe the species as new, he attributed it to Paykull]; characters).
— Zetterstedt, 1828: 74 (Staphylinus; characters; Lapland).
— C. Sahlberg, 1830: 321 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 28 (Staphylinus; Finland).
— Mannerheim, 1831: 442 (Staphylinus; Finland).
— Ménétriés, 1832: 146 (Staphylinus; Caucasus-Persia region).
— Runde, 1835: 7 (Staphylinus; characters; Germany).
— Lacordaire, 1835: 395 (Staphylinus; characters; France).
— Nordmann, 1837: 84 (Philonthus; characters; Europe).
— Zetterstedt, 1838: 65 (Staphylinus; characters; habitat; Lapland).
— Erichson, 1839a: 455 (Philonthus; characters; Germany).
— Heer, 1839: 261 (Philonthus; characters; Switzerland).
— Stephens, 1839: 395 (Philonthus; characters; England).
— Erichson, 1840: 447 (Philonthus; characters; Europe).
— Dufour, 1843: 35 (Staphylinus; characters; France).
— P. Lucas, 1846: 111 (Philonthus; notes; Algeria).
— Jacobsen, 1849: 692 (Philonthus; characters; Denmark).
— Hochhuth, 1849: 136 (Philonthus; characters).
— Redtenbacher, 1849: 703 (Philonthus; characters; Austria).
— Küster, 1853a: 55 (Philonthus; characters; Europe).
— Hardy, 1851: 27 (Philonthus; notes; England).
— Fairmaire and Laboulbène, 1856: 519 (Philonthus; characters; France).
— Kraatz, 1857d: 583 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 193 (Philonthus; characters; Austria).
— Thomson, 1860: 163 (Philonthus; characters; Scandinavia).
— Hochhuth, 1862: 55 (Philonthus; Russia).
— Fauvel, 1865a: 296 [= 1865b: 46] (Philonthus; habitat; France).
— Wollaston, 1865: 490 (Philonthus; notes; Canary Islands).
— Baudi, 1870: 386 (Philonthus; characters).
— Fauvel, 1874: 250 [= 1874a: 476] (Philonthus; characters; France; Algeria; Cyprus; Turkey; Caucasus; Iran; Buchara).
— Redtenbacher, 1874: 211 (Philonthus; characters; Austria).
— Solsky, 1874: 176 (Philonthus; characters; Turkestan).
— Seidlitz, 1875: 275 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 399 [= 1877: 255] (Philonthus; characters; larval characters; France).
— J. Sahlberg, 1876: 19 (Philonthus; Finland).
— Fauvel, 1878: 125 [= 1878a: 45] (Philonthus; Algeria).
— Fauvel, 1886: 55 [= 1886a: 48] (Philonthus; Algeria).
— Fowler, 1888: 265 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 448 (Philonthus; characters; Palaearctic region).
— Fauvel, 1902b: 112 (Philonthus; Algeria).
— Sainte-Claire Deville, 1906: 98 (Philonthus; Corsica).
— Poppius, 1909: 24 (Philonthus; subgenus Gefyrobius; Lena Valley, Siberia).
— Reitter, 1909: 127 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 359 (Philonthus; catalog).
— Johansen, 1914: 398 (Philonthus; characters; Denmark).
— Gridelli, 1922a: 176 (Philonthus; Persia).
— Portevin, 1929: 361 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1366 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Jeannel and Jarrige, 1949: 375 (Philonthus; cave species; France).
— V. Hansen, 1952: 55 (Philonthus; subgenus Philonthus; characters; Denmark).
— Hatch, 1957: 197 (Philonthus; [Note: Gravenhorst cited as author]; characters; British Columbia; Washington; Oregon; Idaho).
— Smetana, 1958: 184 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 150 (Philonthus; characters).
— Scheerpeltz, 1960d: 135 (Philonthus; Portugal).
— Scheerpeltz, 1961d: 128 (Philonthus; Italy).
— Scheerpeltz, 1963a: 420 (Philonthus; Greece).
— Palm, 1963b: 72 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 179 (Philonthus; subgenus Philonthus; characters; central Europe).
— Smetana, 1964d: 75 (Philonthus; subgenus Philonthus; Czechoslovakia).
— Raianu, 1964: 103 (Philonthus; Romania).
— Smetana, 1965: 15 (Philonthus; notes; Miquelon; St. Pierre; Nova Scotia; Newfoundland).
— Horion, 1965: 144 (Philonthus; all Europe).
— Pototskaia, 1966: 156 (Philonthus; larval characters).
— Pototskaia, 1967: 57 (Philonthus; larval characters).
— Raianu, 1968a: 466 (Philonthus; Romania).
— Szujecki, 1968a: 729 (Philonthus; Poland).
— Coiffait, 1974: 252 (Philonthus; synonym of carbonarius).
— Shilov, 1975a: 375 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Pope, 1977: 30 (Philonthus; Britain).
— Hinton, 1981: 660 (Philonthus; characters of egg).
— Frank, 1982: 44 (Philonthus; list of parasites).
— Boller, 1983: 119 (Philonthus; larval characters).
— Tóth, 1984: 55 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Lohse and Lucht, 1989: 165 (Philonthus; synonym of carbonarius).
— Welch, 1993: 228 (Philonthus; ovariole number and ovary structure).
— M. Morgan, 1993: 56 (Philonthus; congregation; Britain).
— Smetana, 1993b: 48 (Philonthus; synonym of carbonarius).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; synonym of carbonarius).
— Smetana, 1995e: 137 (Philonthus; synonym of carbonarius).
— Downie and Arnett, 1996: 377 (Philonthus; [Note: Gravenhorst cited as author]; characters; USA).
— Owen, 1997a: 149 (Philonthus; cited as valid species; collecting sites; Ireland).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
picatus Stephens, 1832: 220 (Quedius; Type locality: London; Suffolk; Devonshire; [Note: See lectotype designation by Smetana and Herman, 1999]).
— Stephens, 1839: 391 (Quedius; characters; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of fumigatus).
— Smetana and Herman, 1999: 298 (Philonthus; lectotype designation: type locality not stated; synonym of carbonarius).
chalcopterus Stephens, 1832: 231 (Philonthus; Type locality: Norfolk; Suffolk).
— Stephens, 1839: 395 (Philonthus; synonym of varius).
— Hardy, 1851: 27 (Philonthus; synonym of varius).
nigroaeneus Stephens, 1832: 232 (Philonthus; cited as a variety of chalcopterus as “variety β. Ph. nigroaeneus”; Type locality: Norfolk; Suffolk).
tanaicus Hochhuth, 1851: 14 (Philonthus; Type locality: Don-Gebiet; [Note: See lectotype designation by Gusarov, 1992a]).
— Ganglbauer, 1895: 452 (Philonthus; synonym of dimidiatus).
— Bernhauer and Schubert, 1914: 335 (Philonthus; synonym of dimidiatus).
— Coiffait, 1974: 258 (Philonthus; synonym of dimidiatus).
— Gusarov, 1992a: 785 [= 1993a: 74] (Philonthus; synonym of carbonarius; lectotype designation: M. Don).
— Smetana, 1995e: 137 (Philonthus; synonym of carbonarius).
oligostigma Leinberg, 1900: 79 (Philonthus; variety of varius; Type locality: Helsingfors).
— Bernhauer and Schubert, 1914: 359 (Philonthus; aberration of varius).
— Smetana, 1955: 206 (Philonthus; aberration of varius; notes).
— Hatch, 1957: 197 (Philonthus; aberration of varius; characters).
— Smetana, 1958: 185 (Philonthus; synonym of bimaculatus).
— Smetana, 1959: 151 (Philonthus; aberration of bimaculatus; characters).
— Coiffait, 1967a: 398 (Philonthus; variety of nitidicollis; characters).
— Coiffait, 1970e: 62 (Philonthus; variety of nitidicollis; Cyprus).
— Coiffait, 1974: 251 (Philonthus; variety of nitidicollis).
— Smetana, 1995e: 137 (Philonthus; synonym of carbonarius).
shetlandicus Poppius, 1905: 9 (Philonthus; variety of varius; Type locality: Shetland Islands: Lerwick and Tingwall).
— Bernhauer and Schubert, 1914: 359 (Philonthus; variety of varius).
— Donisthorpe, 1930: 95 (Philonthus; variety of varius; characters; Britain).
— Scheerpeltz, 1933: 1367 (Philonthus; variety of varius).
— Coiffait, 1974: 252 (Philonthus; variety of carbonarius).
— Pope, 1977: 30 (Philonthus; synonym of varius).
— Smetana, 1995e: 137 (Philonthus; synonym of carbonarius).
thessalicus Roubal, 1916a: 45 (Philonthus; aberration of varius; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Thessalia).
— Scheerpeltz, 1933: 1367 (Philonthus; aberration of varius).
— Smetana, 1958: 184 (Philonthus; aberration of varius).
— Smetana, 1959: 150 (Philonthus; aberration of varius; characters).
— Coiffait, 1967a: 400 (Philonthus; aberration of carbonarius; characters).
— Coiffait, 1974: 252 (Philonthus; variety of carbonarius).
— Tóth, 1984: 55 (Philonthus; aberration of varius; characters).
— Smetana, 1995e: 137 (Philonthus; synonym of carbonarius).
menetriesi Kirshenblat, 1933: 102 (Philonthus; variety of varius; proposed for misidentification of picimanus by Fauvel; Type locality: Not cited).
— Fauvel, 1874: 250 [= 1874a: 476] (Philonthus; synonym of varius; [Note: Cited as picimanus; Kirshenblat, 1933 regarded this citation to be a misidentification and renamed it]).
— Tottenham, 1939d: 232 (Philonthus; aberration of varius).
— Smetana, 1958: 184 (Philonthus; synonym of varius).
— Smetana, 1959: 150 (Philonthus; aberration of varius; characters).
— Coiffait, 1967a: 400 (Philonthus; aberration of carbonarius; characters).
— Coiffait, 1974: 252 (Philonthus; variety of carbonarius).
— Tóth, 1984: 55 (Philonthus; aberration of varius; characters).
— Smetana, 1995e: 137 (Philonthus; synonym of carbonarius).
vinohradensis R. Dvořák and Havelka, 1952: 41 (Philonthus; variety of varius; Type locality: ČSR: Bohemia, Praha-Vinohrady).
— Smetana, 1955b: 223 (Philonthus; variety of varius; characters).
— Smetana, 1958: 184 (Philonthus; aberration of varius).
— Coiffait, 1974: 252 (Philonthus; cited as vinoghradensis; synonym of carbonarius).
— Smetana, 1995e: 137 (Philonthus; synonym of carbonarius).
bothellensis Hatch, 1957: 198 (Philonthus; Type locality: Washington: Bothell).
pindeus Coiffait, 1976b: 86 (Philonthus; Type locality: Grèce, Katara, 1700 m).
— Schillhammer, 1992: 66 (Philonthus; synonym of carbonarius).
— Smetana, 1995e: 137 (Philonthus; synonym of carbonarius).
caribaeus Bierig, 1940a: 141 (Philonthus; Type locality: Cuba: Provincia de la Habana y de Pinar del Rio).
— Blackwelder, 1943: 415 (Philonthus; republication of original description; Cuba).
▴ Distribution: West Indies.
carinulatus Sharp, 1885: 402 (Philonthus; Type locality: Mexico, Jalapa; Oaxaca, Las Vigas).
caroli Last, 1973: 427 (Philonthus; Type locality: Madang).
carpenteri Bernhauer, 1937: 301 (Philonthus; Type locality: Uganda: Kigezi, 6800 ft.).
— Bernhauer, 1940: 136 (Philonthus; Uganda).
— Tottenham, 1949b: 309 (Philonthus; characters).
— Tottenham, 1956: 226, 278 (Philonthus; notes; Ruanda).
— Tottenham, 1962: 163 (Philonthus; characters).
▴ Distribution: Uganda, Rwanda.
castaneicollis Lea, 1931: 371 (Philonthus; Type locality: New Guinea: Finsch Haven).
castaneipennis Kraatz, 1859, see: Erichsonius.
castaneipennis Cameron, 1918: 87 (Philonthus; [preoccupied]; Type locality: Singapore: Mandai).
— Cameron, 1921a: 376, 405 (Philonthus; characters; Singapore).
— Scheerpeltz, 1933: 1335 (Philonthus; catalog).
▴ Distribution: Singapore.
castanopterus Stephens, 1832, see: see: rubripennis Stephens, 1832.
castaneus Gemminger and Harold, 1868: 586 (Philonthus; replacement name for badius Motschulsky).
— Bernhauer and Schubert, 1914: 331 (Philonthus; catalog).
— Cameron, 1932: 99 (Philonthus; characters; East Indies).
— Scheerpeltz, 1933: 1335 (Philonthus; catalog).
— Biswas, 1986: 127 (Philonthus; India).
▴ Distribution: “East Indies”.
badius Motschulsky, 1858: 661 (Gabrius; [Note: Preoccupied as secondary homonym]; Type locality: Indes orientales).
— Bernhauer and Schubert, 1914: 331 (Philonthus; synonym of castaneus).
— Cameron, 1932: 99 (Philonthus; synonym of castaneus).
— Cameron, 1934d: 79 (Philonthus; synonym of castaneus).
— Cameron, 1940: 89 (Philonthus; characters; synonym of castaneus).
castor Jarrige, 1970: 51 (Philonthus; Type locality: Madagascar: Mt. Tsaratanana, 1500 m).
— Jarrige, 1978: 289 (Philonthus; Madagascar).
— Lecoq, 1998: 70 (Philonthus; subgenus Piezarthrus; characters; Madagascar).
▴ Distribution: Madagascar.
catamarcanus Bernhauer, 1916c: 36 (Philonthus; Type locality: Argentinien: Prov. Catamarca).
densior Bernhauer, 1916c: 36 (Philonthus; variety of catamarcanus; Type locality: Provinz Tucuman).
caucasicus Nordmann, 1837: 99 (Philonthus; Type locality: provinciae Caucasicae).
— Erichson, 1840: 466 (Philonthus; characters; Asia occidentalis).
— Hochhuth, 1849: 143 (Philonthus; characters).
— Redtenbacher, 1849: 704 (Philonthus; characters; Austria).
— Kraatz, 1857d: 597 (Philonthus; synonym of ebeninus).
— Redtenbacher, 1857: 194 (Philonthus; characters; Austria).
— Hochhuth, 1862: 60 (Philonthus; characters; Russia).
— Gemminger and Harold, 1868: 587 (Philonthus; synonym of dimidiatus).
— Fauvel, 1874: 233 [= 1874a: 459] (Philonthus; synonym of dimidiatus).
— Redtenbacher, 1874: 213 (Philonthus; characters; Austria).
— J. Sahlberg, 1876: 17 (Philonthus; synonym of dimidiatus).
— Ganglbauer, 1895: 452 (Philonthus; synonym of dimidiatus).
— Bernhauer and Schubert, 1914: 335 (Philonthus; synonym of dimidiatus).
— Scheerpeltz, 1933: 1340 (Philonthus; synonym of dimidiatus).
— Tottenham, 1939d: 233 (Philonthus; replace preoccupied dimidiatus).
— Smetana, 1958: 176 (Philonthus; synonym of dimidiatus).
— Lohse and Lucht, 1989: 168 (Philonthus; note).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Smetana, 1995e: 211 (Philonthus; characters; notes; Alberta; British Columbia; Manitoba; Ontario; Quebec; Saskatchewan; California; Colorado; Connecticut; Idaho; Massachusetta; Minnesota; Montana; Nevada; New York; Oregon; Texas; Utah; Washington; Wisconsin).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; Italy ?).
▴ Distribution: Europe, Russia, Caucasus, Turkey, Iran, Afghanistan, Uzbekistan, Kazakhstan, “Karakorum region”, India, Mongolia, Canada, USA.
dimidiatus C. Sahlberg, 1830: 326 (Staphylinus; [Note: Preoccupied; this name was a secondary homonym of Paederus dimidiatus Panzer, 1795, when both names were in Philonthus. Tottenham (1939d: 233) recognized that fact and cited P. caucasicus, a junior synonym of P. dimidiatus, as the valid name of the species. Furthermore, the Sahlberg name evidently is a junior primary homonym of Staphylinus dimidiatus Say, 1830, which may have been published August 20, 1830 (Bousquet, 1993), whereas the month of publication for the Sahlberg name is December as no month has been specified for the work]; Type locality: Willnäs).
— Mannerheim, 1830: 30 (Staphylinus; Finland).
— Mannerheim, 1831: 444 (Staphylinus; Finland).
— Nordmann, 1837: 97 (Philonthus; characters; Finland).
— Erichson, 1840: 480 (Philonthus; characters; Finland).
— Kraatz, 1857d: 597 (Philonthus; synonym of ebeninus).
— Hochhuth, 1862: 61 (Philonthus; notes; Russia).
— Fauvel, 1874: 233 [= 1874a: 459] (Philonthus; characters; Finland; Prussia; Austria; Russia; Caucasus; Cyprus; Iran; Buchara; Siberia; western Asia).
— Seidlitz, 1875: 276 (Philonthus; variety of quisquiliarius; characters; Baltic region).
— Mulsant and Rey, 1876: 449 [= 1877: 305] (Philonthus; characters; France).
— J. Sahlberg, 1876: 17 (Philonthus; characters; Finland).
— Ganglbauer, 1895: 452 (Philonthus; characters; eastern Europe; eastern Mediterranean region; central Siberia).
— Reitter, 1909: 129 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 335 (Philonthus; catalog).
— Gridelli, 1922a: 176 (Philonthus; characters; Persia).
— Scheerpeltz, 1933: 1340 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Gridelli, 1934: 74 (Philonthus; Karakoram region).
— Tottenham, 1939d: 233 (Philonthus; synonym of caucasicus; preoccupied).
— Smetana, 1953: 118 (Philonthus; Turkey).
— Hatch, 1957: 187 (Philonthus; characters; Washington; Oregon).
— Smetana, 1958: 176 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 156 (Philonthus; characters).
— Palm, 1963b: 84 (Philonthus; characters; Finland).
— Lohse, 1964: 182 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 155 (Philonthus; Finland; Russia; Ukraine; Hungary; Czechoslovakia; Austria; Germany).
— Smetana, 1966: 330 (Philonthus; Czechoslovakia).
— Pototskaia, 1966: 156 (Philonthus; larval characters).
— Pototskaia, 1967: 61 (Philonthus; larval characters).
— Coiffait, 1967a: 415 (Philonthus; characters; Europe; North Africa; Caucasus; Siberia).
— Szujecki, 1968a: 728 (Philonthus; Poland).
— Szujecki, 1969a: 256 (Philonthus; notes; Poland).
— Jarrige, 1971: 495 (Philonthus; Iran).
— Coiffait, 1974: 258 (Philonthus; characters; Europe; Finland; Mediterranean region; Caucasus; Persia; Turkestan; Siberia).
— Smetana 1975c: 170 (Philonthus; Mongolia).
— Shilov, 1976: 149 (Philonthus; Mongolia).
— Coiffait, 1979d: 552 (Philonthus; Afghanistan).
— Coiffait, 1982: 29 (Philonthus; cited as valid species; India).
— Coiffait, 1983e: 478 (Philonthus; Afghanistan).
— Tóth, 1984: 67 (Philonthus; characters).
— Lucht, 1987: 104 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; France).
— Lohse and Lucht, 1989: 168 (Philonthus; synonym of caucasicus).
— Gusarov, 1992a: 785 [= 1993a: 74] (Philonthus; synonymic notes; cited as valid species).
— Smetana, 1993b: 48 (Philonthus; synonym of caucasicus).
— Smetana, 1995e: 211 (Philonthus; synonym of caucasicus).
— Downie and Arnett, 1996: 378 (Philonthus; characters; USA).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
rutilipennis Hochhuth, 1851: 13 (Philonthus; Type locality: Rossia borealis; [Note: See lectotype designation by Gusarov, 1992a]).
— Bernhauer and Schubert, 1914: 354 (Philonthus; catalog).
— Scheerpeltz, 1933: 1359 (Philonthus; catalog).
— Coiffait, 1967a: 437 (Philonthus; characters; Russia).
— Coiffait, 1974: 303 (Philonthus; subgenus Kenonthus?; characters; Russia; Siberia).
— Shilov, 1975a: 376 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Gusarov, 1992a: 786 [= 1993a: 75] (Philonthus; synonym of dimidiatus; lectotype designation: Ross. b.).
— Smetana, 1995e: 211 (Philonthus; synonym of caucasicus).
apicalis Leinberg, 1900a: 184 (Philonthus; variety of dimidiatus; Type locality: Helsingfors).
— Bernhauer and Schubert, 1914: 335 (Philonthus; aberration of dimidiatus).
— Smetana, 1958: 177 (Philonthus; aberration of dimidiatus).
— Smetana, 1959: 156 (Philonthus; aberration of dimidiatus; characters).
— Coiffait, 1967a: 415 (Philonthus; aberration of apicalis; characters).
— Coiffait, 1974: 258 (Philonthus; variety of dimdiatus).
— Tóth, 1984: 67 (Philonthus; variety of dimidiatus).
— Smetana, 1995e: 211 (Philonthus; synonym of caucasicus).
unicolor Leinberg, 1900a: 184 (Philonthus; [preoccupied]; Type locality: Helsingfors).
— Bernhauer and Schubert, 1914: 335 (Philonthus; aberration of dimidiatus).
— Scheerpeltz, 1933: 1340 (Philonthus; synonym of dimidiatus).
— Smetana, 1958: 177 (Philonthus; synonym of dimidiatus).
— Coiffait, 1974: 258 (Philonthus; synonym of dimdiatus).
— Smetana, 1995e: 211 (Philonthus; synonym of caucasicus).
heinemanni Bernhauer, 1913: 130 (Philonthus; variety of dimidiatus; Type locality: Braunschweig).
— Bernhauer and Schubert, 1914: 335 (Philonthus; aberration of dimidiatus).
— Scheerpeltz, 1933: 1340 (Philonthus; aberration of dimidiatus).
— Smetana, 1958: 177 (Philonthus; aberration of dimidiatus).
— Smetana, 1959: 156 (Philonthus; aberration of dimidiatus; characters).
— Coiffait, 1967a: 415 (Philonthus; aberration of dimidiatus; characters).
— Szujecki, 1968a: 728 (Philonthus; aberration of dimidiatus; Poland).
— Szujecki, 1969a: 256 (Philonthus; aberration of dimidiatus; notes; Poland).
— Coiffait, 1974: 258 (Philonthus; variety of dimdiatus).
— Tóth, 1984: 67 (Philonthus; aberration of dimidiatus).
— Smetana, 1995e: 211 (Philonthus; synonym of caucasicus).
— Kashcheev, 1999a: 57 (Philonthus; subspecies of dimidiatus; Kazakhstan).
caurinus Horn, 1884: 203 (Philonthus; Type locality: California; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from Vancouver to northern California).
— Bernhauer and Schubert, 1914: 331 (Philonthus; catalog).
— Hatch, 1957: 186 (Philonthus; characters; British Columbia; Idaho; Washington; Oregon).
— Smetana, 1995e: 364 (Philonthus; lectotype designation: Cal [California]; characters; notes; British Columbia; California; Idaho; Oregon; Washington).
▴ Distribution: Canada, USA.
cautus Erichson, 1840: 446 (Philonthus; Type locality: Carolina meridionalis; [Note: See lectotype designation by Smetana, 1995e]).
— Horn, 1884: 188 (Philonthus; characters; Carolina; Georgia; Florida).
— Bernhauer and Schubert, 1914: 331 (Philonthus; catalog).
— Woodruff, 1982: 27, 38 (Philonthus; in Gopher Tortoise burrows; notes; Florida).
— Smetana, 1995e: 181 (Philonthus; lectotype designation: Carolina meridionalis; characters; Alabama; Arkansas; Florida; Georgia; Louisiana; Maryland; Mississippi; North Carolina; South Carolina; Texas; Virginia).
▴ Distribution: USA.
cavifrons Sharp, 1885: 408 (Philonthus; Type locality: Panama, Bugaba 800 to 1500 feet).
celatus Sharp, 1885: 420 (Philonthus; Type locality: Mexico, Mirador, Oaxaca, Tehuacan, Yolos, Cuernavaca).
celebensis Cameron, 1942c: 843 (Philonthus; Type locality: S. Celebes: Lompa-Battau).
celer Gravenhorst, 1806, see: ventralis Gravenhorst, 1802.
centralis Sharp, 1885: 422 (Philonthus; Type locality: Mexico, Jalapa; Guatemala, San Geronimo; Panama, Volcan de Chiriqui).
— Bernhauer and Schubert, 1914: 331 (Philonthus; catalog).
— Frank, 1982: 38 (Philonthus; list of parasites).
▴ Distribution: Mexico, Guatemala, Panama.
cephalicus Casey, 1915, see: Bisnius.
cerambus Smetana, 1995e: 101 (Philonthus; Type locality: New Mexico: Linclon N. F., 1 mi. SE Cloudcroft, 8750 ft. Also cited from Alberta, Arizona, Colorado, Utah, Wyoming).
cervicalis Casey, 1915, see: longicornis Stephens, 1832.
cervicornis Schubert, 1911, see: Prionophilonthus.
cervus Tottenham, 1956: 289 (Philonthus; Type locality: Ruanda: terr. Kibuye, Yanina, 2300 m).
chalceipennis Fauvel, 1878, see: quisquiliarius Gyllenhal, 1810.
chalceus Stephens, 1832: 227 (Philonthus; [Note: Nomen dubium; most previous uses of chalceus refer to succicola]; Type locality: London).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Stephens, 1839: 394 (Philonthus; characters; England).
— Hardy, 1851: 25 (Philonthus; notes; England).
— Bernhauer, 1926g: 2 (Philonthus; [Note: Based on Tottenham, 1937a, probably a misidentification]; Kamchatka region).
— Cameron, 1932: 80 (Philonthus; [Note: Based on Tottenham, 1937a, probably a misidentification]; characters; India).
— Tottenham, 1937a: 190 (Philonthus; according to Stephens' description, this species refers either to laminatus or intermedius; type series includes four species namely, laminatus, intermedius, carbonarius and chalceus; [Note: No decision was reached concerning the disposition of chalceus Stephens]).
— Tottenham, 1939d: 232 (Philonthus; [Note: After discussing chalceus and removing the species that had been listed as synonyms nothing was decided about chalceus, the name was left in limbo; Ganglbauer's 1895 use of chalceus was regarded as referring to succicola, no specific mention was made of other usages of chalceus]).
— Smetana, 1967d: 202 (Philonthus; [Note: Based on Tottenham, 1937a, probably a misidentification]; Mongolia).
— Scheerpeltz, 1976a: 87 (Philonthus; [Note: Based on Tottenham, 1937a, probably a misidentification]; Nepal).
— Shilov, 1976: 148 (Philonthus; [Note: Based on Tottenham, 1937a, probably a misidentification]; Mongolia).
— Coiffait, 1982a: 233 (Philonthus; [Note: Based on Tottenham, 1937a, probably a misidentification]; Nepal).
▴ Distribution: Britain.
chalcopterus Stephens, 1832, see: carbonarius Gravenhorst, 1802.
chalybeipennis W. J. MacLeay, 1873, see: Quedius.
championi Sharp, 1885: 427 (Philonthus; Type locality: Panama, Peña Blanca 3000 to 4000 feet).
chandigarhensis Pajni and Kohli, 1977: 515 (Philonthus; Type locality: India: Chandigarh, University Campus).
chapmani Bernhauer, 1939, see: Gabrius.
chappuisi Bernhauer, 1939a: 84 (Philonthus; Type locality: Kenya: Mt. Aberdare, vers. Ouest, Mais. forest. Kinangop, 2600 m).
chatterjeei Cameron, 1926b: 356 (Philonthus; Type locality: Mussorie District; Arni Gad).
— Cameron, 1932: 115 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1336 (Philonthus; catalog).
▴ Distribution: India.
cheesmani Cameron, 1937a: 101 (Philonthus; Type locality: New Guinea: Diene, alt. 1200 ft.).
— Last, 1968: 347 (Philonthus; New Guinea).
— Last, 1987: 31 (Philonthus; Papua New Guinea).
▴ Distribution: New Guinea.
chehirensis Bernhauer, 1935a: 46 (Philonthus; Type locality: Klein-Asien: Ak-Chehir).
chilensis Bernhauer and Schubert, 1914, see: Loncovilius.
chilenus Bernhauer, 1939, see: Philonthellus.
chinensis Bernhauer, 1916e: 30 (Philonthus; Type locality: Kiautschau).
— Scheerpeltz, 1933: 1336 (Philonthus; catalog).
— Coiffait, 1967a: 413 (Philonthus; characters; China).
— Coiffait, 1974: 291 (Philonthus; characters; China).
▴ Distribution: China.
chiriquensis Sharp, 1885: 404 (Philonthus; Type locality: Panama, Volcan de Chiriqui between 4000 and 6000 feet).
chlorinus Tottenham, 1939, see: gracilior Casey, 1915.
chlorocephalus Solsky, 1872: 301 (Philonthus; Type locality: Cayenne).
chloropterus Bernhauer, 1939a: 82 (Philonthus; Type locality: Kenya: Camp III de l'Elgo, zone de Bruyeres, est, 3500 m).
chonrima Hayashi, 1993, see: prolatus Sharp, 1874.
chopardi Cameron, 1950b: 214 (Philonthus; Type locality: Agadez; Téouar, 800–900 m; Monts Tarraouaji, 900 m).
— Cameron, 1952d: 824 (Philonthus; Mauritania; Senegal; Niger; Dahomey).
— Cameron, 1952e: 832 (Philonthus; Dahomey).
— Lecoq, 1990: 199 (Philonthus; notes; Mascarene Islands).
▴ Distribution: Morocco, Mauritania, Senegal, Benin, Niger, Sudan, Congo, Kenya, Tanzania, Mozambique, Zambia, Zimbabwe, South Africa, Mascarene Islands, Yemen, Saudi Arabia.
misor Tottenham, 1953: 146 (Philonthus; Type locality: Cape Colony: Uitenhage. Also cited from Sudan, Nyassa, Yemen, Rhodesia; N.W. Rhodesia, Mashonaland, German East Africa).
— Tottenham, 1962: 138, 194 (Philonthus; notes; Tanganyika).
— Coiffait, 1966b: 338 (Philonthus; Mauritania; Senegal).
— Levasseur, 1966a: 1213 (Philonthus; Congo; Sudan; Nyassa; Tanganyika; Kenya; Arabia; Zaire; Rhodesia; Cape).
— Coiffait, 1967a: 424 (Philonthus; characters; Africa).
— Coiffait, 1974: 202 (Philonthus; characters; Morocco).
— Lecoq, 1990: 199 (Philonthus; synonym of chopardi).
christiei Bernhauer, 1920d: 181 (Philonthus; Type locality: Sikkim).
— Cameron, 1932: 139 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1336 (Philonthus; catalog).
▴ Distribution: India.
masuriensis Cameron, 1926b: 358 (Philonthus; Type locality: Mussorie District; Dhobi Ghat. Chakrata District; Bodyar. Simla Hills; Kotgarh, 5000 to 8000 feet).
— Cameron, 1932: 139 (Philonthus; synonym of christiei).
— Scheerpeltz, 1933: 1336 (Philonthus; synonym of christiei, cited as christei).
chrysoscutum Tikhomirova, 1973: 165 (Philonthus; Type locality: pos. Khasan).
chrysurus Nordmann, 1837, see: Xanthopygus.
chujoi R. Dvořák, 1958: 138 (Philonthus; Type locality: Japan: Honshu: Numata Gumma).
chungkingensis Bernhauer, 1943, see: Gabrius.
chyuluensis Cameron, 1942, see: Gabronthus.
cicatricosus Erichson, 1840, see: Cafius.
cinctipennis Fauvel, 1875a: XXX [= 1875b: 232 (Philonthus; Type locality: Egypte, Le Caire).
— Fauvel, 1878: 123 [= 1878a: 43] (Philonthus; Egypt).
— Fauvel, 1886b: 150 (Philonthus; notes; Philippines; Annam; Cochinchine; Egypt).
— Fauvel, 1902b: 110 (Philonthus; Egypt).
— Fauvel, 1904a: 59 (Philonthus; Khandesh; Bírmanie; Annam; Indochina; Philippines; Sumatra; Egypt; Ethiopia; Somaliland).
— Bernhauer and Schubert, 1914: 332 (Philonthus; catalog).
— Bernhauer, 1928: 16 (Philonthus; Sumatra).
— Cameron, 1932: 117 (Philonthus; characters; Burma; India).
— Scheerpeltz, 1933: 1336 (Philonthus; catalog).
— Cameron, 1933d: 386 (Philonthus; Sumatra).
— Scheerpeltz, 1963b: 428 (Philonthus; Egypt).
— Coiffait, 1967a: 422 (Philonthus; characters; Egypt).
— Coiffait, 1974: 204 (Philonthus; characters; Egypt; Iraq; intertropical region of Africa and Asia).
— Frank, 1982: 39 (Philonthus; list of parasites).
— Yadav, Chand, and Yadav, 1995: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
— Gusarov, 1997: 286 (Philonthus; Oman).
▴ Distribution: Egypt, Ethiopia, “Somaliland”, Oman, Iraq, India, Myanmar, Vietnam, Indonesia, Philippines.
cinctulus Gravenhorst, 1802: 167 (Staphylinus; Type locality: Bengala).
— Gravenhorst, 1806: 89 (Staphylinus).
— Erichson, 1840: 432 (Philonthus; characters; Bengal).
— Kraatz, 1859: 78 (Philonthus; characters; Ceylon; India borealis; Bengalia).
— Bernhauer, 1902b: 26 (Philonthus; Ceylon).
— Fauvel, 1903: 158 (Philonthus; Genji; Burma; Annam).
— Fauvel, 1904a: 58 (Philonthus; Khandesh; Ceylan; Bírmanie; Annam).
— Bernhauer and Schubert, 1914: 332 (Philonthus; catalog).
— Cameron, 1932: 77 (Philonthus; characters; Ceylon; India).
— Scheerpeltz, 1933: 1336 (Philonthus; catalog).
— Scheerpeltz, 1960: 4 (Philonthus; Pakistan).
— Yadav, Chand, and Yadav, 1995: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
— Schillhammer, 1999: 61 (Philonthus; characters; Pakistan; India; Nepal; Sri Lanka; Myanmar).
▴ Distribution: Pakistan, Nepal, India, Sri Lanka, Myanmar, Vietnam.
cinctus Fauvel, 1905a: 140 (Philonthus; Type locality: Abyssinie: Massouah, Moncullo, Archico; Bogos: Lebka. Senegal: Saine-Louis; Bissao).
— Bernhauer and Schubert, 1914: 332 (Philonthus; catalog).
— Tottenham, 1949b: 328 (Philonthus; characters).
— Tottenham, 1961: 203 (Philonthus; Senegal).
— Tottenham, 1962: 161 (Philonthus; characters).
— Scheerpeltz, 1974: 142 (Philonthus; South West Africa; South Africa).
▴ Distribution: Senegal, Ethiopia, Namibia, South Africa.
cinctutus Melsheimer, 1844, see: hepaticus Erichson, 1840.
cingulatus Fauvel, 1878f: 259 (Philonthus; Type locality: Nouvelle-Guinée, Andai).
circumcinctus Eppelsheim, 1895: 204 (Philonthus; Type locality: Ganale Gudda, Arussi Galla).
— Fauvel, 1907: 43 (Philonthus; Kilimandjaro; Nairobi; Zululand; Angola).
— Bernhauer and Schubert, 1914: 332 (Philonthus; catalog).
— Scheerpeltz, 1933: 1336 (Philonthus; catalog).
— Tottenham, 1949b: 345 (Philonthus; characters).
— Tottenham, 1962: 202, 203, 205 (Philonthus; characters; Ethiopia; Kenya).
— Coiffait, 1967a: 417 (Philonthus; characters; Sahara).
— Coiffait, 1974: 293 (Philonthus; characters; Abyssinia; East Africa; Angola; southern Sahara).
— Scheerpeltz, 1974c: 22 (Philonthus; Sudan).
▴ Distribution: Sudan, Ethiopia, Kenya, Tanzania, Angola, South Africa.
circumductus Fauvel, 1895: 263 (Philonthus; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Philippines, Antipolo).
— Bernhauer and Schubert, 1914: 332 (Philonthus; catalog).
— Cameron, 1921a: 377, 405 (Philonthus; characters; Singapore).
— Bernhauer, 1928: 16 (Philonthus; Sumatra).
— Cameron, 1932: 118 (Philonthus; characters; Burma).
— Cameron, 1933d: 387 (Philonthus; Sumatra).
— Frank, 1982: 39 (Philonthus; list of parasites).
— Yadav, Chand, and Yadav, 1995: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
▴ Distribution: Myanmar, Singapore, Indonesia, Philippines.
clambus Tottenham, 1949b: 325 (Philonthus; Type locality: S. Rhodesia).
claudus† Scudder, 1900a: 82 (Philonthus; pleistocene fossil; Type locality: Scarborough).
clementi Levasseur, 1962a: 60 (Philonthus; Type locality: Ditinn, Guinée. Also cited from Upper Volta).
cliens Eppelsheim, 1890, see: aeneipennis Boheman, 1858.
clunalis Horn, 1884, see: Gabrius.
cochleatus Scheerpeltz, 1937b: 211 (Philonthus; Type locality: Bulgaria: Umgebung von Varna).
— Gridelli, 1943: 129 (Philonthus; characters; Albania).
— Smetana, 1954b: 177 (Philonthus; characters; Germany; Austria; Turkey).
— Scheerpeltz, 1956c: 1100 (Philonthus; Cyprus).
— Lohse, 1956: 89 (Philonthus; characters; notes).
— Smetana, 1958: 192 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 158 (Philonthus; characters).
— Lohse, 1964: 182 (Philonthus; subgenus Philonthus; characters; central Europe).
— Focarile, 1964: 58 (Philonthus; Italy).
— Horion, 1965: 157 (Philonthus; Bulgaria; Albania; Italy; France; Corsica; Morocco; Turkey; Switzerland; Germany; Austria; Czechoslovakia).
— Raianu, 1967: 270 (Philonthus; cited as cohleatus; characters; ecological notes; Romania).
— Coiffait, 1974: 314 (Spatulonthus; characters; Balkans; Asia Minor; Armenia; Italy; central Europe; Corsica).
— Shilov, 1975a: 375 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Pace and Zanetti, 1977: 300 (Spatulonthus; Italy).
— Tóth, 1984: 68 (Philonthus; characters).
— Lucht, 1987: 106 (Philonthus; checklist; Germany; Czechoslovakia; Austria; Switzerland).
— Smetana, 1993b: 48 (Philonthus; subgenus Spatulonthus; Slovakia).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; Italy).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
▴ Distribution: Europe, Cyprus, Turkey, Armenia, Russia, Kazakhstan.
coelestis Bernhauer, 1933a: 40 (Philonthus; Type locality: Tatsienlu-Kiulung).
coelites Erichson, 1840, see: Chroaptomus.
coenicola Gistel, 1857: 23 (Philonthus; [Note: Nomen dubium]; Type locality: Bayern).
coenosus Gravenhorst, 1806, see: nitidus Fabricius, 1787.
coetus Last, 1981: 122 (Philonthus; Type locality: Musa).
cognatus Stephens, 1832: 229 (Philonthus; Type locality: London).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Hardy, 1851: 26 (Philonthus; synonym of scutatus).
— G. Waterhouse, 1858: 24 (Philonthus; synonym of politus).
— Ganglbauer, 1895: 447 (Philonthus; synonym of fuscipennis).
— Bernhauer and Schubert, 1914: 339 (Philonthus; synonym of fuscipennis).
— Smetana, 1958: 182 (Philonthus; synonym of fuscipennis).
— Coiffait, 1967a: 397 (Philonthus; characters).
— Jarrige, 1971: 495 (Philonthus; Iran).
— Coiffait, 1974: 231 (Philonthus; characters; Holarctic region).
— Bordoni, 1974b: 9 (Philonthus; subgenus Philonthus; Italy).
— Pace and Zanetti, 1977: 293 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Coiffait, 1978d: 164 (Philonthus; Turkey).
— Hinton, 1981: 661 (Philonthus; characters of egg).
— Segers, 1987a: 182 (Philonthus; elytral aberration; Belgium).
— Lohse and Lucht, 1989: 165 (Philonthus; note).
— Pilon and Zanetti, 1991: 63 (Philonthus; Italy).
— Welch, 1993: 228 (Philonthus; ovariole number and ovary structure).
— J. Li, 1993: 37 (Philonthus; China).
— M. Morgan, 1993: 56 (Philonthus; congregation; Britain).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Smetana, 1995e: 120 (Philonthus; characters; Alberta; British; Coumbia; New Brunswick; Newfoundland; Nova Scotia; Ontario; Prince Edward Island; Quebec; Arkansas; California; Colorado; Connecticut; Idaho; Illinois; Indiana; Iowa; Maine; Maryland; Massachusetts; Montana; New Hampshire; New Jersey; New York; North Carolina; North Dakota; Ohio; Oregon; Pennsylvania; Tennessee; Utah; Washington; Wisconsin).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; Italy).
— Lys, 1995 (Philonthus; predator in cereal field; Switzerland).
— Levesque and Levesque, 1996: 292 (Philonthus; seasonal abundance; Quebec).
— Staniec, 1996: 118 (Philonthus; pupal characters).
— Downie and Arnett, 1996: 377 (Philonthus; characters; USA).
— Owen, 1997a: 149 (Philonthus; collecting sites; Ireland).
— Smith and Webb, 1999 (Philonthus; aggregation behavior; Britain).
— Staniec, 1999a: 56 (Philonthus; characters of pupa; Poland).
— Schillhammer, 2000: 135 (Philonthus; characters; Russia; Tajikistan; India).
— Marchal and Chardonnet, 2000: 72 (Philonthus; Corsica).
— Assing, 2001: 75 (Philonthus; collected from xerothermous site; Germany).
▴ Distribution: Tunisia, Algeria, Europe, Turkey, Iran, Kazakhstan, Tajikistan, Russia, China, Canada, USA.
fuscipennis Mannerheim, 1830: 28 (Staphylinus; [preoccupied]; Type locality: Scania Sveciae).
— Mannerheim, 1831: 442 (Staphylinus; Sweden).
— Fabricius, 1775: 266 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 340 cited politus Fabricius as a synonym of fuscipennis; Fabricius did not describe the species as new, he attributed it to Linné]; characters).
— Nordmann, 1837: 82 (Philonthus; characters; Sweden).
— Motschulsky, 1858d: 68 (Philonthus; synonym of politus).
— J. Sahlberg, 1876: 19 (Philonthus; synonym of politus).
— Ganglbauer, 1895: 447 (Philonthus; characters; Palaearctic region; North America).
— Fauvel, 1902b: 112 (Philonthus; Algeria).
— Sainte-Claire Deville, 1906: 98 (Philonthus; Corsica).
— Reitter, 1909: 127 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 339 (Philonthus; catalog).
— Johansen, 1914: 397 (Philonthus; characters; Denmark).
— Scheerpeltz, 1933: 1344 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Kloet and Hincks, 1945: 166 (indicated as an invalid name).
— Normand, 1947a: 8 (Philonthus; Tunisia).
— V. Hansen, 1952: 54 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1953: 118 (Philonthus; Turkey).
— Hatch, 1957: 196 (Philonthus; characters; British Columbia; Washington; Oregon; Idaho).
— Smetana, 1958: 182 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 149 (Philonthus; characters).
— Scheerpeltz, 1961d: 128 (Philonthus; Italy).
— Fagel, 1963: 429 (Philonthus; Turkey).
— Palm, 1963b: 71 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 178 (Philonthus; subgenus Philonthus; characters; central Europe).
— Smetana, 1964d: 74 (Philonthus; subgenus Philonthus; Czechoslovakia).
— Scheerpeltz, 1965b: 401 (Philonthus; Sicily).
— Scheerpeltz, 1965c: 506 (Philonthus; Sicily).
— Smetana, 1965: 15 (Philonthus; notes; Miquelon; St. Pierre; Nova Scotia; Newfoundland).
— Szujecki, 1965 (Philonthus; notes on biology and development).
— Horion, 1965: 140 (Philonthus; widespread in Europe).
— Pototskaia, 1966: 158 (Philonthus; larval characters).
— Pototskaia, 1967: 63 (Philonthus; larval characters).
— Raianu, 1968a: 466 (Philonthus; Romania).
— Szujecki, 1968a: 729 (Philonthus; Poland).
— Coiffait, 1974: 231 (Philonthus; synonym of cognatus).
— Shilov, 1975a: 375 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Pope, 1977: 29 (Philonthus; synonym of cognatus).
— Frank, 1982: 41 (Philonthus; list of parasites).
— Boller, 1983: 120 (Philonthus; larval characters).
— Tóth, 1984: 52 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Lohse and Lucht, 1989: 165 (Philonthus; synonym of cognatus).
— Nowosad, 1990: 105 (Philonthus; collected from mole nest; notes; Poland).
— Dearborn and Donahue, 1993: 14 (Philonthus; Maine).
— Smetana, 1993b: 48 (Philonthus; synonym of cognatus).
— Smetana, 1995e: 120 (Philonthus; synonym of cognatus).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; synonym of cognatus).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
— Schillhammer, 2000: 135 (Philonthus; synonym of cognatus).
maculicornis Stephens, 1832: 229 (Philonthus; Type locality: London; Norfolk; Suffolk; Devonshire).
— Curtis, 1829: 25 (Philonthus; nomen nudum; catalog; Britain).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Holme, 1837: 63 (Philonthus; Scilly Islands).
— Hardy, 1851: 26 (Philonthus; synonym of politus).
— G. Waterhouse, 1858: 24 (Philonthus; synonym of politus).
— Ganglbauer, 1895: 447 (Philonthus; synonym of fuscipennis).
— Bernhauer and Schubert, 1914: 339 (Philonthus; synonym of fuscipennis).
— Coiffait, 1974: 231 (Philonthus; synonym of cognatus).
— Smetana, 1995e: 120 (Philonthus; synonym of cognatus).
— Schillhammer, 2000: 135 (Philonthus; synonym of cognatus).
melanopterus Stephens, 1832: 229 (Philonthus; Type locality: London; Norfolk).
— Curtis, 1829: 25 (Philonthus; nomen nudum; catalog; Britain).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Stephens, 1839: 395 (Philonthus; variety of politus; characters; England).
— Hardy, 1851: 26 (Philonthus; synonym of politus).
— G. Waterhouse, 1858: 24 (Philonthus; synonym of politus).
— Fauvel, 1869: 489 (Philonthus; synonym of politus).
— Ganglbauer, 1895: 447 (Philonthus; synonym of fuscipennis).
— Bernhauer and Schubert, 1914: 339 (Philonthus; synonym of fuscipennis).
— Smetana, 1958: 182 (Philonthus; synonym of fuscipennis).
— Coiffait, 1974: 231 (Philonthus; synonym of cognatus).
— Smetana, 1995e: 120 (Philonthus; synonym of cognatus).
— Schillhammer, 2000: 135 (Philonthus; synonym of cognatus).
microcephalus Stephens, 1832: 229 (Philonthus; [preoccupied]; Type locality: London).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Stephens, 1839: 395 (Philonthus; synonym of carbonarius).
— Hardy, 1851: 26 (Philonthus; synonym of carbonarius).
— Ganglbauer, 1895: 447 (Philonthus; synonym of fuscipennis).
— Bernhauer and Schubert, 1914: 339 (Philonthus; synonym of fuscipennis).
— Smetana, 1958: 182 (Philonthus; synonym of fuscipennis).
— Coiffait, 1974: 231 (Philonthus; synonym of cognatus).
— Smetana, 1995e: 120 (Philonthus; synonym of cognatus).
— Schillhammer, 2000: 135 (Philonthus; synonym of cognatus).
cognatus Sharp, 1876, see: Paederomimus.
coiffaiti Levasseur, 1962a: 56 (Philonthus; Type locality: Kolwezi, Katanga).
coiffaiti Hromádka, 1992, see: purpuripennis Reitter, 1887.
coiffaitianus Levasseur, 1980: 358 (Philonthus; Type locality: Gabon: Belinga).
collarti Cameron, 1932b: 133 (Philonthus; Type locality: Bondia, Likimi).
collartianus Cameron, 1935, see: Gabrius.
collis Last, 1987: 25 (Philonthus; Type locality: Papua New Guinea: New Guinea NE: Mount Missim, 2400–2800 m).
colon Sharp, 1885: 419 (Philonthus; Type locality: Mexico, Cordova).
coloratus Tottenham, 1939 [Note: Nomen nudum], see: debilis Gravenhorst, 1802.
coloripennis Bernhauer, 1910, see: Bisnius.
colubrinus Fauvel, 1868, see: Gabrius.
columbianus Baudi, 1848: 126 (Philonthus; Type locality: Novae Valenciae, Columbia).
— Fauvel, 1891: 111 (Philonthus; cited as columbinus; Venezuela).
— Bernhauer and Schubert, 1914: 332 (Philonthus; cited as columbinus; catalog).
— Scheerpeltz, 1933: 1337 (Philonthus; catalog).
▴ Distribution: Colombia, Venezuela.
combustus Fauvel, 1907: 44 (Philonthus; Type locality: steppe de Barta; Angola).
— Bernhauer and Schubert, 1914: 332 (Philonthus; catalog).
— Tottenham, 1949b: 333 (Philonthus; characters).
— Tottenham, 1962: 139, 223 (Philonthus; characters; Kenya).
▴ Distribution: Angola, Kenya.
commodatus Last, 1968: 356 (Philonthus; Type locality: New Guinea: Buana, Sairuwaget Range, 3000 ft., 20–30 miles north of Lae).
communis Motschulsky, 1860: 568 (Gabrius; Type locality: Daourie).
— Hochhuth, 1862: 57 (Philonthus; characters; Dauria near Lake Baikal).
— Solsky, 1871c: 349 (Philonthus; characters; Siberia).
— Bernhauer, 1913: 131 (Philonthus; characters; East Siberia).
— Bernhauer and Schubert, 1914: 332 (Philonthus; catalog).
— Scheerpeltz, 1933: 1337 (Philonthus; catalog).
— Wüsthoff, 1937a: 347 (Philonthus; illustration of aedeagus).
— Coiffait, 1967a: 411 (Philonthus; characters).
— Coiffait, 1974: 256 (Philonthus; characters; Siberia).
▴ Distribution: Russia.
comoranus Lecoq, 1996: 150 (Philonthus; Type locality: l'archipel des Comores: Grande-Comore, La Convalescence, 1750 m).
complicatus Tottenham, 1961: 209 (Philonthus; Type locality: Senegal: Parc National du Niokolo-Koba, Badi).
comptus Haldeman, 1852, see: furvus Nordmann, 1837.
concinnus Gravenhorst, 1802: 21 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae; [Note: See neotype designation by Smetana, 1995e]).
— Curtis, 1829: 25 (Philonthus; catalog; Britain).
— Stephens, 1832: 233 (Philonthus; characters; England).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Erichson, 1839a: 461 (Philonthus; synonym of varians).
— Stephens, 1839: 396 (Philonthus; variety of varius; characters; England).
— Erichson, 1840: 461 (Philonthus; synonym of ebeninus variety minor).
— Kraatz, 1857d: 597 (Philonthus; synonym of ebeninus).
— Thomson, 1860: 159 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 333 cited varians Thomson as a synonym of concinnus; Thomson did not describe the species as new, he attributed it to Gravenhorst who attributed it to Paykull]; characters; Scandinavia).
— Fauvel, 1874: 233 [= 1874a: 459] (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 33 cited ebeninus Fauvel ex parte as a synonym of concinnus; Fauvel did not describe the species as new, he attributed it to Kraatz]; characters; France).
— Mulsant and Rey, 1876: 449 [= 1877: 305] (Philonthus; characters; larval characters; France).
— Eppelsheim, 1892: 332 (Philonthus; Turkestan).
— Ganglbauer, 1895: 452 (Philonthus; characters; Palaearctic region).
— Fauvel, 1902b: 110 (Philonthus; synonym of ebeninus).
— Sainte-Claire Deville, 1906: 96 (Philonthus; Corsica).
— Montandon, 1908: 80 (Philonthus; Romania).
— Poppius, 1909: 24 (Philonthus; Lena Valley, Siberia).
— Reitter, 1909: 129 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 333 (Philonthus; catalog).
— Johansen, 1914: 405 (Philonthus; characters; Denmark).
— Gridelli, 1920a: 16 (Philonthus; characters; Sicily; Corsica; Sardinia; Italy).
— Gridelli, 1922a: 176 (Philonthus; characters; Persia).
— Portevin, 1929: 362 (Philonthus; characters; France).
— Gridelli, 1930: 72 (Philonthus; Libya).
— Jacobs, 1931: 146 (Philonthus; characters).
— Scheerpeltz, 1933: 1337 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— C. Koch, 1941: 63 (Philonthus; Morocco).
— Méquignon, 1942a: 20 (Philonthus; Azores).
— Jeannel and Jarrige, 1949: 374 (Philonthus; cave species; Algeria).
— V. Hansen, 1952: 64 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1953: 118 (Philonthus; Turkey).
— Coiffait, 1955e: 426 (Philonthus; Lebanon).
— Scheerpeltz, 1956c: 1100 (Philonthus; Cyprus).
— Hatch, 1957: 187 (Philonthus; characters; British Columbia; Washington; Oregon; Idaho).
— Smetana, 1958: 175 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 157 (Philonthus; characters).
— Scheerpeltz, 1960d: 135 (Philonthus; Portugal).
— Scheerpeltz, 1961d: 128 (Philonthus; Italy).
— Scheerpeltz, 1963: 11 (Philonthus; Afghanistan).
— Scheerpeltz, 1963a: 420 (Philonthus; subgenus Philonthus; Greece).
— Palm, 1963b: 82 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 182 (Philonthus; subgenus Philonthus; characters; central Europe).
— Scheerpeltz, 1964a: 299 (Philonthus; subgenus Philonthus; Crete).
— Focarile, 1964: 59 (Philonthus; Italy).
— Scheerpeltz, 1965b: 401 (Philonthus; Sicily).
— Scheerpeltz, 1965c: 505 (Philonthus; Sicily).
— Smetana, 1965: 14 (Philonthus; notes; Nova Scotia; Newfoundland).
— Horion, 1965: 154 (Philonthus; Madeira; Canaries; Azores; North Africa; Turkey; Afghanistan; Uzbekistan; all Europe).
— Sawada, 1966: 259 (Philonthus; Afghanistan).
— Coiffait, 1967a: 417 (Philonthus; preoccupied; synonym of ochropus).
— Smetana, 1967i: 554 (Philonthus; Turkey).
— Raianu, 1968a: 464 (Philonthus; Romania).
— Szujecki, 1968a: 728 (Philonthus; Poland).
— Coiffait, 1974: 261 (Philonthus; cited as concinus; synonym of ochropus).
— Smetana 1975c: 169 (Philonthus; Mongolia).
— Shilov, 1975a: 375 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Shilov, 1976: 149 (Philonthus; Mongolia).
— Pope, 1977: 29 (Philonthus; Britain).
— Smetana, 1978: 116 (Philonthus; collecting notes; Israel).
— Muona, 1979: 14 (Philonthus; synonym of ochropus; [Note: Philonthus concinnus Gravenhorst was thought to be preoccupied by Xylodromus concinnus Marsham but Gravenhorst's name is older]).
— Frank, 1982: 39 (Philonthus; list of parasites).
— H. Franz, 1984: 59 (Philonthus; Macedonia).
— Tóth, 1984: 66 (Philonthus; characters; Hungary).
— Lucht, 1987: 104 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Pilon and Zanetti, 1991: 58 (Philonthus; Italy).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Smetana, 1995e: 206 (Philonthus; neotype designation: Locality not cited; characters; notes; Alberta; British Columbia; Manitoba; New Brunswick; Newfoundland; Nova Scotia; Ontario Prince Edward Island; Quebec; Saskatchewan; Arizona; California; Colorado; Connecticut; Idaho; Illinois; Indiana; Iowa; Kansas; Maine; Maryland; Massachusetts; Michigan; Missouri; Montana; Nebraska; Nevada; New Hampshire; New Jersey; New York; North Carolina; North Dakota; Ohio; Oregon; Pennsylvania; South Dakota; Utah; Virginia; Washington; Wisconsin; Wyoming).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; Italy).
— Ádám, 1996: 245 (Bisnius; synonym of ochropus).
— Ryabukhin, 1999: 63 (Philonthus; catalog; habitat; distribution; northeastern Russia).
▴ Distribution: Libya, Algeria, Morocco, Europe, Russia, Turkey, Cyprus, Lebanon, Iran, Afghanistan, Uzbekistan, Mongolia, Canada, USA.
ochropus Gravenhorst, 1802: 39 (Staphylinus; Type locality: presumably Germany; [Note: See lectotype designation by Smetana, 1995e]. No locality originally cited). No locality originally cited).
— Latreille, 1804: 322 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 67 (Staphylinus).
— Gyllenhal, 1810: 336 (Staphylinus; characters; Sweden).
— C. Sahlberg, 1830: 325 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 29 (Staphylinus; Finland; Sweden).
— Mannerheim, 1831: 443 (Staphylinus; Finland; Sweden).
— Runde, 1835: 8 (Staphylinus; characters; Germany).
— Nordmann, 1837: 93 (Philonthus; Sweden; Prussia).
— Erichson, 1839a: 462 (Philonthus; synonym of varians).
— Erichson, 1840: 462 (Philonthus; synonym of ebeninus).
— Kraatz, 1857d: 597 (Philonthus; synonym of ebeninus).
— Fauvel, 1869: 488 (Philonthus; synonym of concinnus).
— Fauvel, 1874: 234 [= 1874a: 459] (Philonthus; synonym of ebeninus).
— Ganglbauer, 1895: 452 (Philonthus; aberration of concinnus).
— Bernhauer and Schubert, 1914: 333 (Philonthus; aberration of concinnus).
— Gridelli, 1920a: 16 (Philonthus; aberration of concinnus; characters).
— Scheerpeltz, 1933: 1337 (Philonthus; aberration of concinnus).
— Smetana, 1958: 175 (Philonthus; aberration of concinnus).
— Smetana, 1959: 157 (Philonthus; aberration of concinnus).
— Coiffait, 1967a: 417 (Philonthus; cited as valid species; characters).
— Coiffait, 1970d: 143 (Philonthus; cited as valid species; Uzbekistan).
— Coiffait, 1970e: 62 (Philonthus; Cyprus).
— Jarrige, 1971: 495 (Philonthus; Iran).
— Coiffait, 1974: 261 (Philonthus; cited as valid species; characters; Palaearctic region).
— Bordoni, 1975a: 152 (Philonthus; cited as valid species; Afghanistan).
— Pace and Zanetti, 1977: 291 (Philonthus; cited as valid species; Italy).
— Pope, 1977: 29 (Philonthus; synonym of concinnus).
— Muona, 1979: 14 (Philonthus; cited as valid name).
— Outerelo, 1980a: 56 (Philonthus; Spain).
— Hinton, 1981: 660 (Philonthus; characters of egg).
— Coiffait, 1983e: 478 (Philonthus; Afghanistan).
— Bordoni, 1984: 341 (Philonthus; Lebanon).
— Tóth, 1984: 66 (Philonthus; aberration of concinnus).
— Borges, 1990: Table II (Philonthus; cited as valid species; Azores).
— Smetana, 1993b: 48 (Philonthus; synonym of concinnus).
— Smetana, 1995e: 206 (Philonthus; lectotype designation: presumably Germany; synonym of concinnus).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; synonym of concinnus).
— Downie and Arnett, 1996: 378 (Philonthus; characters; USA).
— Ádám, 1996: 245 (Bisnius; valid name; Hungary).
irregularis Mannerheim, 1830: 30 (Staphylinus; Type locality: Petropoli).
— Mannerheim, 1831: 444 (Staphylinus; Russia).
— Nordmann, 1837: 93 (Philonthus; Russia).
— Erichson, 1840: 481 (Philonthus; characters; Petropoli).
— Motschulsky, 1858d: 68 (Philonthus; synonym of sordidus).
— Kraatz, 1857d: 596 (Philonthus).
— Fauvel, 1874: 234 [= 1874a: 459] (Philonthus; synonym of ebeninus).
— Ganglbauer, 1895: 452 (Philonthus; synonym of concinnus).
— Bernhauer and Schubert, 1914: 333 (Philonthus; synonym of concinnus).
— Smetana, 1958: 175 (Philonthus; synonym of concinnus).
— Coiffait, 1967a: 417 (Philonthus; variety of ochropus; characters).
— Coiffait, 1974: 261 (Philonthus; variety of ochropus).
— Smetana, 1995e: 206 (Philonthus; synonym of concinnus).
minor Erichson, 1840: 461 (Philonthus; variety of ebeninus; Type locality: Not cited).
— Ganglbauer, 1895: 452 (Philonthus; variety of concinnus).
— Bernhauer and Schubert, 1914: 333 (Philonthus; variety of concinnus).
— Smetana, 1958: 175 (Philonthus; synonym of concinnus).
— Coiffait, 1974: 261 (Philonthus; synonym of ochropus).
melanarius Mulsant and Rey, 1876: 403 [= 1877: 259] (Philonthus; Type locality: les Alpes et les montagnes du Lyonnais).
— Ganglbauer, 1895: 452 (Philonthus; synonym of concinnus).
— Bernhauer and Schubert, 1914: 333 (Philonthus; synonym of concinnus).
— Smetana, 1958: 175 (Philonthus; synonym of concinnus).
— Coiffait, 1974: 261 (Philonthus; synonym of ochropus).
— Smetana, 1995e: 206 (Philonthus; synonym of concinnus).
imperfectus J. Sahlberg, 1903a: 5 (Philonthus; variety of concinnus; Type locality: Not cited, presumably Italy).
— Bernhauer and Schubert, 1914: 333 (Philonthus; variety of concinnus).
— Gridelli, 1920a: 16 (Philonthus; aberration of concinnus; characters).
— Scheerpeltz, 1933: 1337 (Philonthus; variety of concinnus).
— Smetana, 1958: 175 (Philonthus; aberration of concinnus).
— Coiffait, 1974: 261 (Philonthus; variety of ochropus).
— Smetana, 1995e: 206 (Philonthus; synonym of concinnus).
ochripennis Gerhardt, 1910: 555 (Philonthus; aberration of concinnus; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]; cited from Liegnitz).
— Scheerpeltz, 1933: 1337 (Philonthus; aberration of concinnus).
— Smetana, 1953: 118 (Philonthus; aberration of concinnus; Turkey).
— Smetana, 1958: 175 (Philonthus; aberration of concinnus).
— Smetana, 1959: 157 (Philonthus; aberration of concinnus).
— Coiffait, 1967a: 417 (Philonthus; aberration of ochropus; characters).
— Coiffait, 1974: 261 (Philonthus; variety of ochropus).
— Tóth, 1984: 66 (Philonthus; aberration of concinnus).
— Smetana, 1995e: 206 (Philonthus; synonym of concinnus).
concolor Kraatz, 1859: 80 (Philonthus; Type locality: Ceylan).
— Bernhauer and Schubert, 1914: 333 (Philonthus; catalog).
— Cameron, 1932: 85 (Philonthus; characters; Ceylon).
— Scheerpeltz, 1933: 1337 (Philonthus; catalog).
▴ Distribution: Sri Lanka.
concolor Gridelli, 1920 [Note: Nomen nudum], see: fenestratus Fauvel, 1872.
confertoides Hatch, 1957, see: stictus Hausen, 1891.
confertus LeConte, 1863a: 40 (Philonthus; Type locality: Illinois; Nebraska; [Note: See lectotype designation by Smetana, 1995e]).
— Horn, 1884: 221 (Philonthus; characters; Canada; Iowa; Illinois; Kansas).
— Bernhauer and Schubert, 1914: 333 (Philonthus; catalog).
— Smetana, 1995e: 461 (Philonthus; lectotype designation [Note: Locality not cited]; characters; notes; Arkansas; Connecticut; Illinois; Indiana; Iowa; Kentucky; Massachusetts; Missouri; Nebraska; New York; Ohio; Pennsylvania; Tennessee; Wisconsin).
— Downie and Arnett, 1996: 382 (Philonthus; characters; USA).
▴ Distribution: USA.
confinis A. Strand, 1941: 35 (Philonthus; Type locality: Dänemark: Birkelse; Norwegen: Lalm Tolstadskog; Sibirien und Mongolei: Beikem, Kemtschick und Usti Abakansk).
— V. Hansen, 1952: 70 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1958: 198 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 161 (Philonthus; characters).
— Smetana, 1963c: 296 (Philonthus; subgenus Philonthus; Mongolia).
— Palm, 1963b: 88 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 184 (Philonthus; subgenus Philonthus; characters; central Europe).
— Smetana, 1965e: 173 (Philonthus; Mongolia).
— Smetana, 1965f: 255 (Philonthus; notes).
— Horion, 1965: 161 (Philonthus; Norway; Sweden; Finland; Denmark; France; Italy; Bosnia; Poland; Czechoslovakia; Germany).
— Coiffait, 1967a: 420 (Philonthus; characters; Europe).
— Smetana, 1967d: 204 (Philonthus; Mongolia).
— Raianu, 1968: 117 (Philonthus; characters; ecological notes; Romania).
— Raianu, 1968a: 467 (Philonthus; Romania).
— Szujecki, 1968a: 730 (Philonthus; Poland).
— Coiffait, 1974: 243 (Philonthus; characters; Europe; Siberia; Mongolia; France).
— Smetana 1975c: 171 (Philonthus; Mongolia).
— Shilov, 1976: 151 (Philonthus; Mongolia).
— Pace and Zanetti, 1977: 295 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Raichev, 1979: 83 (Philonthus; Bulgaria).
— Tóth, 1984: 71 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; France; Benelux; Denmark; Sweden).
— Schuh, Schillhammer, and Zettel, 1992: 220 (Philonthus; notes; Austria).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Slovakia).
— Hodge and Jones, 1995: 38 (Philonthus; characters; Britain).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Assing, Frisch, Kahlen, et al., 1998: 137 (Philonthus; Austria).
— Tronquet, 1999a: 214 (Philonthus; notes; France).
▴ Distribution: Europe, Russia, Mongolia.
isthmius E. Kangas, 1979: 100 (Philonthus; Type locality: Kivennapa [= Pervomajskoje], USSR).
conformis Lacordaire, 1835, see: discoideus Gravenhorst, 1802.
conformis Sharp, 1876, see: Paederomimus.
confusus Cameron, 1932, see: Gabrius.
congoensis Bernhauer, 1928g: 110 (Philonthus; Type locality: Belgisch-Kongo: Haut-Uele: Moto).
— Cameron, 1929: 64 (Philonthus; characters; Belgian Congo).
— Scheerpeltz, 1933: 1337 (Philonthus; catalog).
— Tottenham, 1949b: 335 (Philonthus; characters).
▴ Distribution: Congo.
congruens Cameron, 1932: 89 (Philonthus; Type locality: Mussoorie District: Arni Gad).
conicus Bernhauer, 1939, see: Gabrius.
conradti Bernhauer, 1912f: 206 (Philonthus; subgenus Eccoptolonthus; Type locality: Südost Kamerun: Lolodorf).
— Bernhauer and Schubert, 1914: 333 (Philonthus; catalog).
— Tottenham, 1949b: 356 (Philonthus; characters).
— Cameron, 1950g: 38 (Philonthus; subgenus Eccoptolonthus; Belgian Congo).
— Tottenham, 1961: 204 (Philonthus; Senegal).
— Tottenham, 1962: 230 (Philonthus; subgenus Eccoptolonthus; characters; Cameroon; Congo; S. Rhodesia; N. Rhodesia; Tanganyika).
▴ Distribution: Senegal, Cameroon, Congo, Tanzania, Zambia, Zimbabwe.
consors Cameron, 1932: 112 (Philonthus; Type locality: Ceylon).
consputus Baudi, 1848, see: frigidus Märkel and Kiesenwetter, 1848.
constricticeps Tottenham, 1949b: 347 (Philonthus; Type locality: Mashonaland: Salisbury).
contaminatus Gravenhorst, 1802, see: sanguinolentus Gravenhorst, 1802.
contiguus Motschulsky, 1860, see: lepidus Gravenhorst, 1802.
convalescens Eppelsheim, 1890b: 276 (Philonthus; Type locality: Simla, 7000′).
— Bernhauer and Schubert, 1914: 333 (Philonthus; catalog).
— Cameron, 1928c: 564 (Philonthus; Sikkim; Tibet).
— Cameron, 1932: 109 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1337 (Philonthus; catalog).
— Scheerpeltz, 1976: 35 (Philonthus; Nepal).
— Scheerpeltz, 1976a: 88 (Philonthus; Nepal).
— Coiffait, 1983d: 161 (Philonthus; Nepal).
— Yadav, Chand, and Yadav, 1994: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
▴ Distribution: India, Nepal, China.
convergens Casey, 1915, see: Rabigus.
convexicollis Lynch, 1884: 157 (Philonthus; Type locality: Chacabuco in Provincia Bonaerensi occidentali).
— Bernhauer and Schubert, 1914: 333 (Philonthus; catalog).
— Frank, 1982: 39 (Philonthus; list of parasites).
▴ Distribution: Argentina.
convexus Bernhauer, 1912g: 253 (Philonthus; Type locality: Luzon: Rizal, Montalban Gorge).
copiosus Last, 1968, see: Hesperus.
coprophilus Jarrige, 1949: 70 (Philonthus; replacement name for rubromaculatus Bernhauer).
— Tottenham, 1955: 158 (Philonthus; synonym of longicornis).
— Smetana, 1958: 192 (Philonthus; subgenus Philonthus; cited as valid species; characters; ecology; Czechoslovakia).
— Smetana, 1959: 159 (Philonthus; characters).
— Lohse, 1964: 183 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 158 (Philonthus; Italy; France; Belgium; Hungary; Slovakia; Czech Republic; Germany).
— Smetana, 1966: 330 (Philonthus; Czechoslovakia).
— Coiffait, 1974: 313 (Spatulonthus; characters; Europe; North Africa).
— Pace and Zanetti, 1977: 300 (Spatulonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Tóth, 1984: 69 (Philonthus; characters; Hungary).
— Last, 1987a: 221 (Philonthus; characters; Britain).
— Lucht, 1987: 106 (Philonthus; checklist; Germany; Czechoslovakia; Benelux).
— Schuh, Schillhammer, and Zettel, 1992: 221 (Philonthus; Austria).
— Smetana, 1993b: 48 (Philonthus; subgenus Spatulonthus; Czech Republic; Slovakia).
— Hodge and Jones, 1995: 39 (Philonthus; characters; Britain).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Gerend and Braunert, 1997: 197 (Philonthus; notes; Luxembourg).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
— Burakowski, Mroczkowski, and Stefańska, 2000: 49 (Spatulonthus; catalog; Poland).
▴ Distribution: Europe, North Africa, Kazakhstan.
rubromaculatus Bernhauer, 1914d: 9 (Philonthus; form of longicornis; [preoccupied]; Type locality: Süditalien, Prov. di Aquila, Castel di Sangro).
— Bernhauer, 1923d: 177 (Philonthus; variety of longicornis; characters; Italy).
— Scheerpeltz, 1933: 1349 (Philonthus; variety of longicornis).
— Blackwelder, 1943: 410 (Philonthus; synonym of longicornis).
— Tottenham, 1955: 158 (Philonthus; synonym of longicornis).
— Smetana, 1958: 192 (Philonthus; synonym of coprophilus).
— Smetana, 1959: 159 (Philonthus; synonym of coprophilus).
— Coiffait, 1974: 313 (Spatulonthus; synonym of coprophilus).
coracion Peyerimhoff, 1902: 246 (Philonthus; Type locality: au bord du lac de Bens, 2300 m, Alpes Maritimes).
— Bernhauer and Schubert, 1914: 333 (Philonthus; catalog).
— Scheerpeltz, 1933: 1337 (Philonthus; catalog).
— Coiffait, 1967a: 420 (Philonthus; characters; Alps).
— Jarrige, 1968b: 74 (Philonthus; notes; France).
— Coiffait, 1974: 204 (Philonthus; characters; Alps).
— Focarile, 1988: 181 (Philonthus; characters; Italy).
— Aranda and Outerelo, 1991: 119 (Philonthus; characters; comparison with species of quisquiliarius group; Spain).
— Wittwer, 1993: 250 (Philonthus; Switzerland).
— Schülke, 1993c (Philonthus; characters; notes; Swizerland; Italy).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Tronquet, 1998: 11 (Philonthus; notes; France).
— Assing, Frisch, Kahlen, et al., 1998: 138 (Philonthus; characters).
— Schillhammer, 1999a: 61 (Philonthus).
▴ Distribution: Spain, France, Switzerland, Italy, Russia.
baicalicus Bernhauer, 1935a: 45 (Philonthus; Type locality: Baikalgebiet).
corallicola Fairmaire, 1849, see: Remus.
corallipennis Sharp, 1876: 166 (Philonthus; Type locality: Para, Obydos, Ega, Tapajos).
cordicollis Gerstaecker, 1867, see: Diatrechus.
cordilleranus Bernhauer, 1916c: 34 (Philonthus; Type locality: West-Columbien: Ost-Cordilleren, La Pena bei Carmen, 3500 m).
corticalis Sharp, 1885: 413 (Philonthus; Type locality: Panama, Volcan de Chiriqui 2000 to 3000 feet).
coruscoides Scheerpeltz, 1963: 13 [Note: Nomen nudum, published without description, Article 13.1.1.
corruscus Gravenhorst, 1802: 33 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).
— Marsham, 1802: 511 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 333 cited nitidus Marsham, 1802 as a synonym of corruscus Gravenhorst; Marsham did not describe the species as new, he attributed it to Fabricius]; characters; Britain).
— Latreille, 1804: 319 (Staphylinus; cited as coruscus; characters; Germany).
— Gravenhorst, 1806: 51 (Staphylinus).
— Curtis, 1829: 26 (Philonthus; catalog; Britain).
— Stephens, 1832: 239 (Philonthus; characters; England).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Nordmann, 1837: 98 (Philonthus; cited as corrusus; Germany; France; Tauria).
— Holme, 1837: 63 (Philonthus; characters; notes; Scilly Islands).
— Erichson, 1839a: 463 (Philonthus; characters; Germany).
— Heer, 1839: 265 (Philonthus; characters; Switzerland).
— Stephens, 1839: 398 (Philonthus; characters; England).
— Erichson, 1840: 465 (Philonthus; characters; Europe).
— Kolenati, 1846: 20 (Philonthus; Transcaucasia).
— Motschulsky, 1849: 87 (Philonthus; Spain).
— Hochhuth, 1849: 143 (Philonthus; characters).
— Redtenbacher, 1849: 704 (Philonthus; characters; Austria).
— Küster, 1854: 7 (Philonthus; characters; Europe).
— Rosenhauer, 1856: 71 (Philonthus; Granada).
— Fairmaire and Laboulbène, 1856: 525 (Philonthus; characters; France).
— Kraatz, 1857d: 597 (Philonthus; synonym of ebeninus).
— Redtenbacher, 1857: 194 (Philonthus; characters; Austria).
— Peyron, 1858: 425 (Philonthus; Turkey).
— Fauvel, 1874: 234 [= 1874a: 459] (Philonthus; variety of ebeninus).
— Redtenbacher, 1874: 212 (Philonthus; characters; Austria).
— Mulsant and Rey, 1876: 447 [= 1877: 303] (Philonthus; characters; France).
— Eppelsheim, 1892: 332 (Philonthus; Turkestan).
— Ganglbauer, 1895: 452 (Philonthus; cited as coruscus; characters; Europe; Mediterranean region).
— Fauvel, 1902b: 110 (Philonthus; Algeria).
— Montandon, 1908: 80 (Philonthus; cited as coruscus; Romania).
— Reitter, 1909: 129 (Philonthus; cited as coruscus; characters; Germany).
— Bernhauer and Schubert, 1914: 333 (Philonthus; cited as coruscus; catalog).
— Johansen, 1914: 404 (Philonthus; cited as coruscus; characters; Denmark).
— Gridelli, 1920a: 15, 16 (Philonthus; characters; Italy).
— Portevin, 1929: 362 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1338 (Philonthus; cited as coruscus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; cited as coruscus; illustration of aedeagus).
— V. Hansen, 1952: 64 (Philonthus; subgenus Philonthus; cited as coruscus; characters; Denmark).
— Smetana, 1953: 118 (Philonthus; Turkey).
— Smetana, 1958: 173 (Philonthus; subgenus Philonthus; cited as coruscus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 155 (Philonthus; cited as coruscus; characters).
— Lohse, 1964: 181 (Philonthus; subgenus Philonthus; cited as coruscus; characters; central Europe).
— Horion, 1965: 152 (Philonthus; cited as coruscus; Iberian Peninsula to Greece and Turkey; France; Switzerland; Belgium; Holland; Poland; Russia; Ireland; Britain; Denmark; Norway; Sweden; Finland; Germany; Austria).
— Coiffait, 1967a: 415 (Philonthus; characters; Turkestan).
— Raianu, 1968a: 464 (Philonthus; cited as coruscus; Romania).
— Szujecki, 1968a: 728 (Philonthus; cited as coruscus; Poland).
— Coiffait, 1974: 259 (Philonthus; characters; Europe; Britain; Denmark; Scandinavia; Anatolia; Lebanon; Caucasus; Transcaucasus; Turkestan).
— Osella and Zanetti, 1975: 108 (Philonthus; cited as coruscus; collected from nest of Talpa europaea; notes; Italy).
— Bordoni, 1975a: 152 (Philonthus; Afghanistan).
— Pace and Zanetti, 1977: 290 (Philonthus; cited as coruscus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Smetana, 1978: 116 (Philonthus; collecting notes; Israel).
— Coiffait, 1983e: 478 (Philonthus; Afghanistan).
— Tóth, 1984: 63 (Philonthus; cited as coruscus; characters; Hungary).
— Lucht, 1987: 104 (Philonthus; cited as coruscus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Welch, 1993: 228 (Philonthus; ovariole number and ovary structure).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Assing, 2001: 75 (Philonthus; collected from xerothermous site; Germany).
▴ Distribution: Algeria, Europe, Russia, Georgia, Armenia, Turkey, Lebanon, Israel, Afghanistan, Turkmenistan, Uzbekistan.
planus Lacordaire, 1835: 401 (Staphylinus; [preoccupied]; Type locality: Paris).
— Erichson, 1839a: 464 (Philonthus; synonym of corruscus).
— Erichson, 1840: 465 (Philonthus; synonym of corruscus).
— Küster, 1854: 7 (Philonthus; synonym of corruscus).
— Fairmaire and Laboulbène, 1856: 525 (Philonthus; synonym of corruscus).
— Kraatz, 1857d: 597 (Philonthus; synonym of ebeninus).
— Ganglbauer, 1895: 452 (Philonthus; synonym of corruscus).
— Bernhauer and Schubert, 1914: 333 (Philonthus; synonym of corruscus).
— Smetana, 1958: 173 (Philonthus; synonym of corruscus).
— Coiffait, 1974: 259 (Philonthus; synonym of corruscus).
pragensis Roubal, 1948: 123 (Philonthus; aberration of varius; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Bohemia: Praha-Sarka).
— Smetana, 1958: 173 (Philonthus; synonym of corruscus).
— Coiffait, 1974: 259 (Philonthus; synonym of corruscus).
corvinus Erichson, 1839a: 462 (Philonthus; Type locality: Mark Brandenburg).
— Erichson, 1840: 462 (Philonthus; characters; Germany).
— Kolenati, 1846: 20 (Philonthus; Caucasus; Transcaucasus).
— Jacobsen, 1849: 697 (Philonthus; characters; Denmark).
— Hochhuth, 1849: 139 (Philonthus; characters; Lenkoran).
— Redtenbacher, 1849: 704 (Philonthus; characters; Austria).
— Rosenhauer, 1856: 71 (Philonthus; Andalusia; Xerez; Cadiz; Algeciras; Malaga; Granada).
— Fairmaire and Laboulbène, 1856: 524 (Philonthus; characters; France).
— Kraatz, 1857d: 599 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 195 (Philonthus; characters; Austria).
— Thomson, 1860: 160 (Philonthus; characters; Scandinavia).
— Hochhuth, 1862: 58 (Philonthus; Russia).
— Fauvel, 1874: 236 [= 1874a: 462] (Philonthus; characters; Scandinavia; Britain; Germany; Switzerland; Austria).
— Redtenbacher, 1874: 213 (Philonthus; characters; Austria).
— J. Sahlberg, 1876: 18 (Philonthus; Finland).
— Fowler, 1888: 271 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 452 (Philonthus; characters; Scandinavia; England; Germany; Austria; Switzerland).
— Reitter, 1909: 129 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 333 (Philonthus; catalog).
— Johansen, 1914: 405 (Philonthus; characters; Denmark).
— Scheerpeltz, 1933: 1338 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; cited as cervinus; illustration of aedeagus).
— V. Hansen, 1952: 73 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1958: 210 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 161 (Philonthus; characters).
— Palm, 1963b: 90 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 184 (Philonthus; subgenus Philonthus; characters; central Europe).
— Smetana, 1964d: 75 (Philonthus; subgenus Philonthus; Czechoslovakia).
— Horion, 1965: 162 (Philonthus; Russia; Finland Sweden; Norway; Britain; Ireland; Denmark; Ukraine; Poland; Czech Republic; Slovakia; Hungary; Romania; Italy; Switzerland; France; Holland; Austria; Germany).
— Coiffait, 1967a: 422 (Philonthus; characters; Europe).
— Coiffait, 1974: 203 (Philonthus; characters; Europe; Britain).
— Shilov, 1975a: 376 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Pope, 1977: 29 (Philonthus; Britain).
— Raichev, 1979: 81 (Philonthus; Bulgaria).
— Tóth, 1984: 73 (Philonthus; characters).
— Drugmand, 1987a: 311 (Philonthus; Belgium).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; Benelux; Denmark; Sweden).
— Schülke and Uhlig, 1988: 11 (Philonthus; Germany).
— Aranda and Outerelo, 1991: 121 (Philonthus; characters).
— Wittwer, 1993: 250 (Philonthus; Switzerland).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Allen, 1994a: 30 (Philonthus; Britain).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
▴ Distribution: Europe, Russia, Azerbaijan, Kazakhstan.
coryndoni Cameron, 1942, see: Gabrius.
costatus Baudi, 1848, see: varians Paykull, 1789.
couleensis Hatch, 1957: 185 (Philonthus; Type locality: Washington: Grand Coulee, Dry Falls).
— Smetana, 1965: 25 (Philonthus; subgenus Philonthus; characters; Miquelon; Nova Scotia; Newfoundland).
— Smetana, 1995e: 327 (Philonthus; characters; notes; Alberta; British Columbia; Manitoba; New Brunswick; Newfoundland; Northwest Territories; Nova Scotia; Ontario; Quebec; Saskatchewan; Idaho; Illinois; Indiana; Massachusetts; Michigan; New Jersey; New York; Washington; Wisconsin).
▴ Distribution: Canada, USA.
couloni Drugmand, 1987: 272 (Philonthus; subgenus Philonthus; Type locality: Belgique: province de Brabant, Forest).
coxalus Hochhuth 1872, see: Gabrius.
coxatus Curtis, 1836, see: laminatus Creutzer, 1799.
coxatus Bernhauer, 1914, see: Gabrius.
crassicornis Fauvel, 1895: 264 (Philonthus; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Tenasserim, Malewoon; Tonkin; Singapore; Java; Ceylan).
— Fauvel, 1904a: 59 (Philonthus; Nilghiris; Ceylan; Bírmanie; Tenasserim; Tonkin; Singapore; Sumatra; Java; Lombock; Maurice; Réunion).
— Fauvel, 1905e: 83 (Philonthus; Java).
— Bernhauer and Schubert, 1914: 333 (Philonthus; catalog).
— Cameron, 1921a: 377, 405 (Philonthus; characters; Singapore).
— Bernhauer, 1928: 16 (Philonthus; Sumatra).
— Cameron, 1932: 114 (Philonthus; characters; Ceylon; India).
— Scheerpeltz, 1933: 1338 (Philonthus; catalog).
— Cameron, 1933d: 387 (Philonthus; Sumatra).
— Jarrige, 1957: 109 (Philonthus; Réunion; also listed from Madagascar and Mauritius).
— Hayashi, 1996: 110 (Philonthus; Taiwan).
— Lecoq, 1990: 199 (Philonthus; Mascarene Islands).
▴ Distribution: Madagascar, Mascarene Islands, Sri Lanka, India, Myanmar, Vietnam, Singapore, Indonesia, Taiwan.
crassulus Horn, 1884, see: Gabrius.
crassulus Sharp, 1885, see: alius Bernhauer and Schubert, 1914.
crebrior Bernhauer, 1931, see: donckieri Bernhauer, 1915.
crestonensis Hatch, 1957, see: Bisnius.
cribellatus Solsky, 1868, see: Styngetus.
cribratus Erichson, 1840, see: Orthidus.
cribrellus Fauvel, 1891: 112 (Philonthus; Type locality: Caracas; Nouvelle Grenade).
cribricephalicus Tottenham, 1949b: 323 (Philonthus; Type locality: Mashonaland: Salisbury. Also cited from Natal).
cribriceps Bernhauer, 1921c: 105 (Philonthus; Type locality: Brasilien: Petropolis).
cribricollis Erichson, 1840, see: Paederomimus.
cribripennis Germain, 1903, see: Loncovilius.
cribripennis Sharp, 1885: 403 (Philonthus; Type locality: Guatemala; Panama, Volcan de Chiriqui 2000 to 3000 feet).
cribriventris Bernhauer, 1912d: 43 (Philonthus; Type locality: Argentinien: Prov. Tucuman; Chaunar Region [Note: In a discussion of Neobisnius parcepunctatus, Frank, 1981: 12, 13 thought this locality probably referred to Chañar Region, La Rioja]).
crinitus Schubert, 1909: 293 (Philonthus; Type locality: südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d'Orizaba).
crotchi Horn, 1884: 203 (Philonthus; Type locality: Vancouver; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from Vancouver to Arizona; near Lake Tahoe).
— Bernhauer and Schubert, 1914: 334 (Philonthus; catalog).
— Hatch, 1957: 186 (Philonthus; subspecies of lomatus; characters; British Columbia; Washington; Oregon).
— Smetana, 1963a: 5 (Philonthus; aedeagus).
— Smetana, 1995e: 416 (Philonthus; lectotype designation: Van [Vancouver]; characters; notes; Alberta; British Columbia; Alaska; California; Montana; Nevada; Oregon; Washington).
▴ Distribution: Canada, USA.
nigritoides Smetana, 1964: 35 (Philonthus; subgenus Philonthus; Type locality: Alaska: Kodiak Island).
— Smetana, 1965a: 7 (Philonthus; characters; Alaska).
— Smetana, 1995e: 416 (Philonthus; synonym of crotchi).
cruentatus Gmelin, 1790: 2035 (Staphylinus; Type locality: Europa).
— Zschach, 1788: 44 (Staphylinus; species number 975; characters stated but not named and referred to as cruentatus by Gmelin).
— Panzer, 1795: 355 (Staphylinus; [Misidentification: Bernhauer and Schubert, 1914: 334 cited bipustulatus Panzer as a synonym of cruentatus Gmelin but Panzer did not describe bipustulatus as new, he attributed it to Fabricius who attributed it to Linné]; characters; Germany).
— Panzer, 1796c: 10 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 334 cited bipustulatus Panzer as a synonym of cruentatus; Panzer did not describe bipustulatus as new, he attributed it to Linné]; illustration; characters).
— Turton, 1802: 519 (Staphylinus; characters; Europe).
— Latreille, 1804: 321 (Staphylinus; cited as bipustulatus Panzer; characters; Germany; France).
— Panzer, 1808: 8 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 334 cited bipustulatus Panzer, 1795: 10 as a synonym of cruentatus Gmelin but Panzer did not describe bipustulatus as new, he attributed it to Fabricius who attributed it to Linné]; characters; illustration).
— Kraatz, 1857d: 601 (Philonthus; [Note: Cited as bipustulatus Panzer]; characters; Germany).
— Thomson, 1860: 164 (Philonthus; [Note: Cited as bipustulatus Panzer]; characters; Scandinavia).
— Fauvel, 1874: 253 [= 1874a: 479] (Philonthus; characters; France; Caucasus; Transcaucasia; Buchara; Lake Baikal).
— Seidlitz, 1875: 277 (Philonthus; cited as bipustulatus Panzer; characters; Baltic region).
— Mulsant and Rey, 1876: 468 [= 1877: 325] (Philonthus; characters; France).
— J. Sahlberg, 1876: 20 (Philonthus; Finland).
— Fowler, 1888: 272 (Philonthus; characters; Britain).
— Eppelsheim, 1892: 334 (Philonthus; Turkestan).
— Ganglbauer, 1895: 454 (Philonthus; characters; Europe; Transcaucasian region; Siberia).
— Sainte-Claire Deville, 1906: 99 (Philonthus; Corsica).
— Reitter, 1909: 130 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 334 (Philonthus; catalog).
— Johansen, 1914: 408 (Philonthus; characters; Denmark).
— Gridelli, 1920a: 18, 20 (Philonthus; characters; Italy).
— Portevin, 1929: 363 (Philonthus; characters; France).
— Cameron, 1932: 130 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1338 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Voris, 1936 (Philonthus; rapid spread in North America; habitat; New York; Michigan; Indiana; Kentucky; Tennessee; Illinois; Wisconsin; Minnesota; North Dakota; South Dakota; Nebraska; Missouri; Kansas; Colorado; Wyoming; Utah; Nevada; California; Idaho; Oregon; Washington; Ontario; Manitoba).
— Uyttenboogaart, 1937: 82 (Philonthus; notes; Canary Islands).
— C. Koch, 1941: 63 (Philonthus; Morocco).
— Hatch, 1957: 189 (Philonthus; characters; British Columbia; Idaho; Washington; Oregon).
— Smetana, 1958: 194 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 157, 158 (Philonthus; characters).
— Scheerpeltz, 1963: 13 (Philonthus; Afghanistan).
— Palm, 1963b: 88 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 183 (Philonthus; subgenus Philonthus; characters; central Europe).
— Focarile, 1964: 59 (Philonthus; Italy).
— Smetana, 1965: 16 (Philonthus; notes; Miquelon; St. Pierre; Newfoundland).
— Horion, 1965: 158 (Philonthus; Ireland; Britain; Holland; Belgium; France to Portugal and Spain; Corsica; Italy; Istria; Dalmatia; Bosnia; Serbia; Macedonia; Greece; France to Poland and Russia; Germany; Austria; Denmark; Sweden; Norway; Finland).
— Coiffait, 1967a: 419 (Philonthus; characters; Europe; Siberia).
— Raianu, 1968a: 466 (Philonthus; Romania).
— Szujecki, 1968a: 730 (Philonthus; Poland).
— Coiffait, 1970d: 143 (Philonthus; Uzbekistan).
— Coiffait, 1974: 241 (Philonthus; characters; Europe; Britain; Scandinavia; Siberia; North Africa; Lebanon; Caucasus; India).
— Shilov, 1975a: 375 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Pace and Zanetti, 1977: 295 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Hinton, 1981: 661 (Philonthus; characters of egg).
— Frank, 1982: 39 (Philonthus; list of parasites).
— Coiffait, 1983e: 477 (Philonthus; Afghanistan).
— Bordoni, 1984: 342 (Philonthus; Lebanon).
— Tóth, 1984: 70 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Nowosad, 1990: 105 (Philonthus; collected from mole nest; notes; Poland).
— Pilon and Zanetti, 1991: 59 (Philonthus; Italy).
— Hunter, Kincher, Bay, Beerwinkle, 1991: 168 (Philonthus; seasonal distribution and diel flight activity; Texas).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Smetana, 1995e: 232 (Philonthus; characters; notes; Alberta; British Columbia; Manitoba; New Brunswick; Newfoundland; Nova Scotia; Ontario; Prince Edward Island; Quebec; Saskatchewan; Alabama; Arizona; Arkansas; California; Colorado; Connecticut; Idaho; Illinois; Indiana; Iowa; Kansas; Kentucky; Maryland; Massachusetts; Michigan; Missouri; Montana; Nebraska; Nevada; New Hampshire; New Jersey; New Mexico; New York; North Carolina; North Dakota; Ohio; Oklahoma; Oregon; Pennsylvania; South Carolina; South Dakota; Tennessee; Texas; Vermont; Virginia; Washington; Wisconsin; Wyoming; Mexico).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Downie and Arnett, 1996: 378 (Philonthus; characters; USA).
— Schmidt, 1999: 108 (Philonthus; food for adults and larvae used for rearing species; rearing methods).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
▴ Distribution: Morocco, Europe, Russia, Georgia, Armenia, Lebanon, Kazakhstan, Afghanistan, Uzbekistan, India, Canada, USA, Mexico.
extinctus Bernhauer, 1900b: 55 (Philonthus; variety of cruentatus; Type locality: Not cited).
— Bernhauer and Schubert, 1914: 334 (Philonthus; variety of cruentatus).
— Gridelli, 1920a: 18 (Philonthus; aberration of cruentatus; characters).
— Cameron, 1932: 130 (Philonthus; synonym of cruentatus).
— Scheerpeltz, 1933: 1338 (Philonthus; variety of cruentatus).
— Smetana, 1958: 194 (Philonthus; aberration of cruentatus).
— Smetana, 1959: 157 (Philonthus; aberration of cruentatus).
— Coiffait, 1967a: 419 (Philonthus; aberration of cruentatus; characters).
— Szujecki, 1968a: 730 (Philonthus; aberration of cruentatus; Poland).
— Coiffait, 1974: 241 (Philonthus; variety of cruentatus).
— Tronquet, 1981: 70 (Philonthus; variety of cruentatus; Afghanistan).
— Smetana, 1995e: 232 (Philonthus; synonym of cruentatus).
immaculatus Gusmann, 1924: 249 (Philonthus; aberration of cruentatus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Ln. Strandweide; Sohl. Travestrand; Dmd.-Strd.; Schwn.; Hhm. Teschower Travestrand).
— Scheerpeltz, 1933: 1338 (Philonthus; synonym of cruentatus).
— Smetana, 1958: 194 (Philonthus; synonym of cruentatus).
— Coiffait, 1974: 242 (Philonthus; synonym of cruentatus).
— Smetana, 1995e: 232 (Philonthus; synonym of cruentatus).
cruenticollis Lea, 1925: 225 (Philonthus; Type locality: Queensland: Townsville).
cruentus Bernhauer, 1915k: 300 (Philonthus; Type locality: Insel S. Thomé: Agua-Ize, 200–700 m).
csikii Bernhauer, 1917g: 46 (Philonthus; Type locality: Africa or.: Kibosho; Arusha-Ju).
— Scheerpeltz, 1933: 1338 (Philonthus; catalog).
— Tottenham, 1962: 136, 179, 181 (Philonthus; characters; notes; Tanganyika).
▴ Distribution: Tanzania.
cumaeus Tottenham, 1949b: 333 (Philonthus; Type locality: S. Rhodesia: Mt. Selinda).
cunctans Horn, 1884: 205 (Philonthus; Type locality: District of Columbia; [Note: See lectotype designation by Smetana, 1995e]).
— Bernhauer and Schubert, 1914: 334 (Philonthus; catalog).
— Frank, 1982: 39 (Philonthus; list of parasites).
— Smetana, 1995e: 394 (Philonthus; lectotype designation: D.C. [= District of Columbia]; characters; notes; Ontario; Georgia; Indiana; Kentucky; Maryland; Massachusetts; Michigan; New Hampshire; New Jersey; New York; North Carolina; Ohio; Pennsylvania; Tennessee; Vermont; Virginia; West Virginia).
— Downie and Arnett, 1996: 380 (Philonthus; characters; USA).
▴ Distribution: Canada, USA.
cunctator Sharp, 1889: 41 (Philonthus; Type locality: Yokohama).
— Bernhauer and Schubert, 1914: 334 (Philonthus; catalog).
— Scheerpeltz, 1933: 1338 (Philonthus; catalog).
— Nakane, 1963a: 91 (Philonthus; characters; habitus photograph; Japan).
— Yuh, Paik, Kwon, and Lee, 1985: 254 (Philonthus; Korea).
— Kanehisa, Tsumuki, and Kawazu, 1994 (Philonthus; cited as cuncutator; chemical secretions).
▴ Distribution: Japan, Korea.
cunicularius Gistel, 1857: 80 (Philonthus; [Note: Nomen dubium]; Type locality: Bavaria).
cupreipennis Cameron, 1926, see: tractatus Eppelsheim, 1895.
cupreonitens Eppelsheim, 1895a: 125 (Philonthus; Type locality: Gabon).
— Fauvel, 1903e: 241 (Philonthus; Cameroon; Gabon; Togo).
— Bernhauer and Schubert, 1914: 334 (Philonthus; catalog).
— Tottenham, 1949b: 332 (Philonthus; characters).
— Cameron, 1950g: 36 (Philonthus; Belgian Congo).
— Scheerpeltz, 1971c: 170 (Philonthus; Cameroon).
▴ Distribution: Togo, Cameroon, Gabon, Congo.
cupreotinctus Lea, 1925: 222 (Philonthus; Type locality: Queensland: Cairns).
cupreus Guérin-Méneville, 1844, see: Phanolinus.
cupripennis Guérin-Méneville, 1844, see: Styngetus.
curiosus Tottenham, 1949b: 298 (Philonthus; Type locality: N.W. Rhodesia: Kashitu, N. of Broken Hill).
curtipennis Bernhauer, 1902, see: tardus Kraatz, 1859.
curvabilis Herman, 2001: 47 (Philonthus; replacement name for sinuatus Tottenham).
sinuatus Tottenham, 1949b: 341 (Philonthus; [preoccupied]; Type locality: N.W. Rhodesia: Kasitu, N. of Broken Hill).
cushmani Hatch, 1957, see: Gabrius.
cyanelytrius Kraatz, 1859: 80 (Philonthus; Type locality: India borealis).
— Bernhauer and Schubert, 1914: 334 (Philonthus; catalog).
— Cameron, 1932: 86 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1338 (Philonthus; catalog).
— Schillhammer, 1998: 108 (Philonthus; characters; India; Nepal).
▴ Distribution: India, Nepal.
gridellii Cameron, 1926b: 360 (Hesperus; Type locality: Almora, Kali Valley, 9000 feet).
— Cameron, 1932: 161 (Hesperus; characters; India).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Scheerpeltz, 1971: 172 (Hesperus; characters).
— Schillhammer, 1994: 74 (Philonthus; synonym of cyanelytrius).
— Schillhammer, 1998: 108 (Philonthus; synonym of cyanelytrius).
cyaneoaureus Bernhauer, 1935c: 90 (Philonthus; Type locality: Venezuela).
cyaneonitens Schillhammer, 1991, see: distincticornis Cameron, 1932.
cyaneoviolaceus Bernhauer, 1915h: 227 (Philonthus; Type locality: Java: Goenoeng Gedeh).
— Bernhauer, 1928: 16 (Philonthus; Sumatra).
— Scheerpeltz, 1933: 1339 (Philonthus; catalog).
— Cameron, 1933b: 344 (Philonthus; characters; British North Borneo).
— Cameron, 1933d: 386 (Philonthus; Sumatra).
— Scheerpeltz, 1935a: 634 (Philonthus; Indonesia).
— Cameron, 1937: 5 (Philonthus; Java).
▴ Distribution: Borneo, Indonesia.
cyanescens Guérin-Méneville, 1844, see: Phanolinus.
cyanicornis Mannerheim, 1830, see: politus Linné, 1758.
cyanipennis Fabricius, 1793: 525 (Staphylinus; Type locality: Parisiae).
— Fabricius, 1801: 597 (Staphylinus; characters; France).
— Turton, 1802: 512 (Staphylinus; characters; France).
— Latreille, 1804: 308 (Staphylinus; synonym of amoenus Olivier).
— Gravenhorst, 1806: 86 (Staphylinus; characters).
— Curtis, 1829: 25 (Quedius; catalog; Britain).
— Stephens, 1832: 230 (Philonthus; characters; England).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Lacordaire, 1835: 387 (Staphylinus; characters; France).
— Heer, 1839: 260 (Philonthus; characters; Switzerland).
— Stephens, 1839: 395 (Philonthus; characters; England).
— Erichson, 1840: 433 (Philonthus; characters; France; Austria; Chersonesus Taurica; Pennsylvania [Note: Based on work by Smetana, 1995e: 117 the last locality is misidentification of caeruleipennis]).
— Laporte, 1840: 177 (Staphylinus; characters; France).
— Dufour, 1843: 33 (Staphylinus; characters; France).
— Redtenbacher, 1849: 700 (Philonthus; characters; Austria).
— Fairmaire and Laboulbène, 1856: 518 (Philonthus; characters; France).
— Kraatz, 1857d: 573 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 190 (Philonthus; characters; Austria).
— Fauvel, 1874: 215 [= 1874a: 441] (Philonthus; cited as cyaneipennis; characters; France; Germany; Switzerland; Austria; Sardinia; Russia; Dauria; North America [Note: Based on work by Smetana, 1995e: 117 the last locality is misidentification of caeruleipennis]).
— Redtenbacher, 1874: 209 (Philonthus; characters; Austria).
— Mulsant and Rey, 1876: 366 [= 1877: 223] (Philonthus; characters; France).
— Ganglbauer, 1895: 444 (Philonthus; characters; middle Europe; Siberia; Siberia; Japan; North America [Note: Based on work by Smetana, 1995e: 117 the last two localities are misidentifications of caeruleipennis]).
— Reitter, 1909: 126 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 334 (Philonthus; catalog).
— Šulc, 1922 (Philonthus; osmeterium; anatomy; histology; physiology).
— Portevin, 1929: 358 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1339 (Philonthus; catalog).
— Aleksandrov, 1934: 152 (Philonthus; China).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Scheerpeltz and Höfler, 1948: 156 (Philonthus; characters; fungus inhabitant).
— Smetana, 1955b: 228 (Philonthus; notes).
— Smetana, 1958: 161 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 149 (Philonthus; characters).
— Lohse, 1964: 178 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 133 (Philonthus; Ukraine; Poland; Czech Republic; Slovakia; Romania; Bosnia; Croatia; Slovakia; Italy; Sardinia; Switzerland; France; Belgium; Germany; Austria).
— Coiffait, 1967a: 393 (Philonthus; characters).
— Raianu, 1968a: 462 (Philonthus; Romania).
— Coiffait, 1974a: 198 (Philonthus; Siberia).
— Coiffait, 1974: 225 (Philonthus; characters; Palaearctic region).
— Tóth, 1984: 50 (Philonthus; characters; Hungary).
— Yuh, Paik, Kwon, and Lee, 1985: 240 (Philonthus; Korea).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).
— J. Li, 1993: 36 (Philonthus; China).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Kanehisa, Tsumuki, and Kawazu, 1994 (Philonthus; cited as cyanippenis; chemical secretions).
— Smetana, 1995e: 117 (Philonthus; in a footnote Smetana reported Schillhammer's discovery that cyanipennis is restricted to temperate continental Eurasia; the North American and Japanese records refer to caeruleipennis; [Note: According to Smetana's interpretation, all citations of cyanipennis in North America and Japan have been moved to caeruleipennis]).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Schillhammer, 1998: 104 (Philonthus; characters; Russia, Kazakhstan; China; Korea).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
▴ Distribution: Europe, Russia, China, Korea.
amoenus Olivier, 1795:(42): 26 (Staphylinus; Type locality: Fontainebleau).
— Fabricius, 1801: 597 (Staphylinus; synonym of cyanipennis Fabricius).
— Latreille, 1804: 308 (Staphylinus; characters).
— Gravenhorst, 1806: 86 (Staphylinus; synonym of cyanipennis Fabricius).
— Stephens, 1839: 395 (Philonthus; synonym of cyanipennis).
— Erichson, 1840: 433 (Philonthus; synonym of cyanipennis).
— Redtenbacher, 1849: 700 (Philonthus; synonym of cyanipennis).
— Kraatz, 1857d: 574 (Philonthus; synonym of cyanipennis).
— Ganglbauer, 1895: 444 (Philonthus; synonym of cyanipennis).
— Bernhauer and Schubert, 1914: 334 (Philonthus; synonym of cyanipennis).
— Smetana, 1958: 161 (Philonthus; synonym of cyanipennis).
— Coiffait, 1974: 225 (Philonthus; synonym of cyanipennis).
— Schillhammer, 1998: 104 (Philonthus; synonym of cyanipennis).
cyanopterus Cameron, 1945c: 715 (Philonthus; Type locality: South Africa: Simon's Town).
cyclopius Cameron, 1939b: 146 (Philonthus; Type locality: New Guinea: Cyclops Mts.: Mt. Lina, altitude 3500–4500 feet).
cylindricollis Cameron, 1934e: 373 (Philonthus; Type locality: Mayumbe: Tshiobo N'Goy).
— Cameron, 1952e: 831 (Philonthus; Dahomey).
— Tottenham, 1962: 187, 188 (Philonthus; characters).
▴ Distribution: Congo, Benin.
daimio Sharp, 1889: 42 (Philonthus; Type locality: Otaru; Hakodate).
— Bernhauer and Schubert, 1914: 334 (Philonthus; catalog).
— Scheerpeltz, 1933: 1339 (Philonthus; catalog).
▴ Distribution: Japan.
dampfi Bernhauer, 1929h: 194 (Philonthus; Type locality: Mexiko: Desierto de los Leones).
danicus Coiffait, 1963: 23 (Philonthus; Type locality: Israël: Jaffa. Also cited from Lebanon).
— Coiffait, 1967a: 420 (Philonthus; characters; Palestine; Lebanon).
— Coiffait, 1974: 246 (Philonthus; characters; Lebanon; Israel).
— Drugmand, 1987: 275 (Philonthus; characters).
▴ Distribution: Israel, Lebanon.
dauloensis Last, 1987: 25 (Philonthus; Type locality: Papua New Guinea: New Guinea NE: Asaru-Chimbu Div., Daulo Pass, 2400 m).
daunus Tottenham, 1953, see: alterius Cameron, 1951.
davus Smetana, 1995e: 97 (Philonthus; Type locality: California: S. Bern. Mts., Hwy. 38, 3 mi SW Onyx Summit, 7700 ft. Also cited from Alberta, British Columbia, Arizona, Colorado, Idaho, Kansas, Montana, Nevada, Oregon, Utah, Washington).
debiliformis Cameron, 1951c: 402 (Philonthus; Type locality: Kenya Colony: Magadi).
— Tottenham, 1961: 204 (Philonthus; Senegal).
— Tottenham, 1962: 138, 194 (Philonthus; characters; Tanganyika).
— Herman, 2001: 28 (Philonthus; synonymy).
▴ Distribution: Senegal, Kenya, Tanzania.
obscuratus Cameron, 1951c: 402 (Philonthus; [preoccupied]; Type locality: Kenya Colony, Maktau).
— Tottenham, 1962: 195 (Philonthus; this species is probably identical to debiliformis).
— Herman, 2001: 28 (Philonthus; synonym of debiliformis).
debilis Gravenhorst, 1802: 35 (Staphylinus; Type locality: Europa; [Note: See lectotype designation by Smetana, 1995e]. No locality originally cited).
— Latreille, 1804: 320 (Staphylinus; characters; Germany).
— Lacordaire, 1835: 407 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 334 cited agilis Boisduval and Lacordaire as a synonym of debilis Gravenhorst; Lacordaire did not describe the species as new, he attributed it to Gravenhorst]; characters; France).
— Erichson, 1839a: 467 (Philonthus; characters; Germany).
— Heer, 1839: 266 (Philonthus; characters; Switzerland).
— Heer, 1839: 266 (Philonthus; [Note: Misidentified as celer according to Smetana, 1958: 180) characters; Switzerland).
— Erichson, 1840: 472 (Philonthus; characters; Germany; France; Egypt; United States).
— Hochhuth, 1849: 142 (Philonthus; characters).
— Redtenbacher, 1849: 705 (Philonthus; characters; Austria).
— LeConte, 1850: 220 (Philonthus; Lake Superior).
— Küster, 1854: 10 (Philonthus; characters; Germany; Switzerland; France; Egypt).
— Rosenhauer, 1856: 71 (Philonthus; Granada).
— Fairmaire and Laboulbène, 1856: 524 (Philonthus; characters; France).
— Kraatz, 1857d: 604 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 195 (Philonthus; characters; Austria).
— Peyron, 1858: 425 (Philonthus; Turkey).
— Hochhuth, 1862: 61 (Philonthus; Russia).
— Thomson, 1867: 149 (Philonthus; characters; Scandinavia).
— Baudi, 1870: 386 (Philonthus; characters).
— Fauvel, 1874: 228 [= 1874a: 454] (Philonthus; characters; France; Algeria; Egypt; Turkey; Caucasus; Transcaucasia; Iran; Madeira; North America).
— Redtenbacher, 1874: 213 (Philonthus; characters; Austria).
— Seidlitz, 1875: 276 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 461 [= 1877: 317] (Philonthus; characters; France).
— J. Sahlberg, 1876: 18 (Philonthus; Finland).
— Fauvel, 1878: 123 [= 1878a: 43] (Philonthus; Algeria).
— Horn, 1884: 194 (Philonthus; characters; eastern Atlantic region west to Kansas and Nebraska).
— Fauvel, 1886: 54 [= 1886a: 46] (Philonthus; Morocco; Egypt; Algeria).
— Sharp, 1889: 40 (Philonthus; Japan).
— Fowler, 1888: 271 (Philonthus; characters; Britain).
— Eppelsheim, 1892: 332 (Philonthus; Turkestan).
— Ganglbauer, 1895: 453 (Philonthus; characters; Palaearctic region; North America).
— Fauvel, 1902b: 110 (Philonthus; Morocco; Algeria; Tunisia; Egypt).
— Montandon, 1908: 80 (Philonthus; Romania).
— Fall and Cockerell, 1907: 166 (Philonthus; New Mexico).
— Sainte-Claire Deville, 1906: 96 (Philonthus; Corsica).
— Reitter, 1909: 128 (Philonthus; characters; Germany).
— Blatchley, 1910: 383 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 334 (Philonthus; catalog).
— Johansen, 1914: 406 (Philonthus; characters; Denmark).
— Gridelli, 1922a: 176 (Philonthus; Persia).
— Portevin, 1929: 361 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1339 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— C. Koch, 1941: 63 (Philonthus; Morocco).
— V. Hansen, 1952: 66 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1953: 118 (Philonthus; Turkey).
— Hatch, 1957: 187 (Philonthus; characters; British Columbia; Idaho; Washington; Oregon).
— Smetana, 1958: 180 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 154 (Philonthus; characters).
— Scheerpeltz, 1961d: 128 (Philonthus; Italy).
— Palm, 1963b: 83 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 182 (Philonthus; subgenus Philonthus; characters; central Europe).
— Scheerpeltz, 1965b: 401 (Philonthus; Sicily).
— Scheerpeltz, 1965c: 506 (Philonthus; Sicily).
— Smetana, 1965: 14 (Philonthus; Newfoundland).
— Horion, 1965: 154 (Philonthus; all Europe).
— Sawada, 1966: 259 (Philonthus; Afghanistan).
— Pototskaia, 1966: 157 (Philonthus; larval characters).
— Pototskaia, 1967: 61 (Philonthus; larval characters).
— Coiffait, 1967a: 413 (Philonthus; characters).
— Szujecki, 1968a: 729 (Philonthus; Poland).
— Coiffait, 1974: 265 (Philonthus; characters; Holarctic region).
— Osella and Zanetti, 1975: 109 (Philonthus; collected from nest of Talpa europaea; notes; Italy).
— Smetana 1975c: 170 (Philonthus; Mongolia).
— Shilov, 1976: 149 (Philonthus; Mongolia).
— Pace and Zanetti, 1977: 292 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Smetana, 1978: 117 (Philonthus; collecting notes; Israel).
— Frank, 1982: 39 (Philonthus; list of parasites).
— Tóth, 1984: 65 (Philonthus; characters; Hungary).
— Lucht, 1987: 104 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Pilon and Zanetti, 1991: 58 (Philonthus; Italy).
— J. Li, 1993: 37 (Philonthus; China).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Smetana, 1995e: 195 (Philonthus; lectotype designation: Europa; characters; notes; Alberta; British Columbia; Manitoba; Now Brunswick; Newfoundland; Nova Scotia; Ontario; Prince Edward Island; Quebec; Saskatchewan; Arizona; California; Colorado; Connecticut; Florida; Idaho; Illinois; Indiana; Iowa; Kansas; Maine; Maryland; Massachusetts; Michigan; Minnesota; Montana; New Hampshire; New Jersey; New York; Ohio; Oregon; Pennsylvania; Rhode Island; South Dakota; Utah; Virginia; Washington; West Virginia; Wisconsin; Wyoming).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Downie and Arnett, 1996: 378 (Philonthus; characters; USA).
▴ Distribution: Egypt, Tunisia, Algeria, Morocco, Europe, Russia, Caucasus, Turkey, Israel, Iran, Afghanistan, Turkmenistan, Uzbekistan, Mongolia, China, Japan, Canada, USA.
lucidus Gravenhorst, 1802: 21 (Staphylinus; Type locality: Europa; [Note: See lectotype designation by Smetana, 1995e]. No locality originally cited).
— Latreille, 1804: 310 (Staphylinus; characters; notes; France; Germany).
— C. Sahlberg, 1830: 322 (Staphylinus; synonym of fimetarius Gravenhorst).
— Stephens, 1832: 232 (Philonthus; characters; England).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Erichson, 1839a: 467 (Philonthus; synonym of debilis).
— Stephens, 1839: 396 (Philonthus; characters; England).
— Erichson, 1840: 472 (Philonthus; synonym of debilis).
— Küster, 1854: 10 (Philonthus; synonym of debilis).
— Kraatz, 1857d: 604 (Philonthus; synonym of debilis).
— Fauvel, 1869: 489 (Philonthus; synonym of varius).
— Ganglbauer, 1895: 453 (Philonthus; synonym of debilis).
— Bernhauer and Schubert, 1914: 334, 359 (Philonthus; synonym of debilis [334] and varius [359]).
— Smetana, 1958: 180, 184 (Philonthus; synonym of debilis [180] and varius [184]).
— Coiffait, 1974: 265 (Philonthus; synonym of debilis).
— Smetana, 1995e: 195 (Philonthus; lectotype designation: Europa; synonym of debilis).
melanocephalus Heer, 1839: 269 (Philonthus; Type locality: Zürich am Horn).
— Kraatz, 1857d: 596 (Philonthus).
— Fauvel, 1874: 228 [= 1874a: 454] (Philonthus; synonym of melanocephalus).
— Ganglbauer, 1895: 453 (Philonthus; synonym of debilis).
— Bernhauer and Schubert, 1914: 334 (Philonthus; synonym of debilis).
— Smetana, 1958: 180 (Philonthus; synonym of debilis).
— Coiffait, 1974: 265 (Philonthus; synonym of debilis).
— Smetana, 1995e: 195 (Philonthus; synonym of debilis).
coloratus Tottenham, 1939a: 202 (Philonthus; aberration of debilis; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from England: Chichester; Jugoslavia: Mostarsko-blato).
— Smetana, 1959: 155 (Philonthus; aberration of debilis; characters).
— Smetana, 1958: 180 (Philonthus; aberration of debilis).
— Coiffait, 1967a: 413 (Philonthus; characters).
— Coiffait, 1974: 265 (Philonthus; variety of debilis).
— Tóth, 1984: 67 (Philonthus; aberration of debilis).
— Smetana, 1995e: 195 (Philonthus; synonym of debilis).
— Hodge and Jones, 1995: 39 (Philonthus; aberration of debilis; characters; Britain).
deceptivus Cameron, 1943, see: Gabrius.
deceptor Cameron, 1932, see: Gabrius.
decipiens LeConte, 1863, see: Cafius.
decipiens Horn, 1884, see: Gabrius.
decoloratus Kirshenblat, 1933: 101 (Philonthus; Type locality: Transbaikalien: Umgebung von Tshita).
— Coiffait, 1967a: 389 (Philonthus; characters; Transbaikal).
— Coiffait, 1974: 304 (Philonthus; subgenus Kenonthus?; characters; Transbaikal).
— Boháč, 1988: 552 [= 1989: 35] (Philonthus; subgenus Philonthus; characters; Russia).
— Schillhammer, 2000: 128 (Philonthus; characters; Russia; Mongolia; Korea; Japan; China).
▴ Distribution: Russia, Mongolia, China, Korea, Japan.
decorus Gravenhorst, 1802: 19 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).
— Gravenhorst, 1806: 85 (Staphylinus; characters).
— Gyllenhal, 1810: 316 (Staphylinus; characters; Sweden).
— Curtis, 1829: 25 (Philonthus; catalog; Britain).
— C. Sahlberg, 1830: 320 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 27 (Staphylinus; Sweden; Runsala Island).
— Mannerheim, 1831: 441 (Staphylinus; Sweden; Runsala Island).
— Stephens, 1832: 230 (Philonthus; characters; England).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Runde, 1835: 7 (Staphylinus; characters; Germany).
— Lacordaire, 1835: 391 (Staphylinus; characters; France).
— Nordmann, 1837: 82 (Philonthus; Sweden; Germany).
— Erichson, 1839a: 451 (Philonthus; characters; Germany).
— Heer, 1839: 260 (Philonthus; characters; Switzerland).
— Stephens, 1839: 395 (Philonthus; characters; England).
— Erichson, 1840: 442 (Philonthus; characters; Europe).
— Jacobsen, 1849: 689 (Philonthus; characters; Denmark).
— Redtenbacher, 1849: 701 (Philonthus; characters; Austria).
— Küster, 1853a: 50 (Philonthus; characters; Europe).
— Hardy, 1851: 26 (Philonthus; notes; England).
— Fairmaire and Laboulbène, 1856: 517 (Philonthus; characters; France).
— Kraatz, 1857d: 580 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 191 (Philonthus; characters; Austria).
— Thomson, 1860: 163 (Philonthus; characters; Scandinavia).
— Fauvel, 1874: 246 [= 1874a: 472] (Philonthus; characters; France; Italy).
— Redtenbacher, 1874: 210 (Philonthus; characters; Austria).
— Seidlitz, 1875: 275 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 391 [= 1877: 247] (Philonthus; characters; France).
— J. Sahlberg, 1876: 19 (Philonthus; Finland).
— Fowler, 1888: 264 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 447 (Philonthus; characters; north and middle Europe).
— Poppius, 1905: 9 (Philonthus; Shetland Islands).
— Reitter, 1909: 127 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 334 (Philonthus; catalog).
— Johansen, 1914: 396 (Philonthus; characters; Denmark).
— Šulc, 1922 (Philonthus; osmeterium; anatomy; histology; physiology).
— Portevin, 1929: 360 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1339 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— V. Hansen, 1952: 54 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1958: 181 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 149 (Philonthus; characters).
— Palm, 1963b: 71 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 177 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 131 (Philonthus; northern and central Europe; France; Italy; Bosnia Hercegovina).
— Evans, 1965 (Philonthus; feeding behavior; Britain).
— Pototskaia, 1966: 158 (Philonthus; larval characters).
— Pototskaia, 1967: 63 (Philonthus; larval characters).
— Coiffait, 1967a: 397 (Philonthus; characters; Europe).
— Raianu, 1968a: 466 (Philonthus; Romania).
— Frank, 1968 (Philonthus; biological notes).
— Szujecki, 1968a: 729 (Philonthus; Poland).
— Kasule, 1970: 68 (Philonthus; larval characters).
— Coiffait, 1974: 230 (Philonthus; characters; Europe; Britain; Russia; Italy; Balkans).
— Pace and Zanetti, 1977: 292 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Raichev, 1979: 82 (Philonthus; Bulgaria).
— Hinton, 1981: 661 (Philonthus; characters of egg).
— Frank, 1982: 40 (Philonthus; list of parasites).
— Boller, 1983: 119 (Philonthus; larval characters).
— Tóth, 1984: 47 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Nowosad, 1990: 103 (Philonthus; collected from mole nest; notes; Poland).
— Pilon and Zanetti, 1991: 60 (Philonthus; Italy).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Staniec, 1996: 118 (Philonthus; pupal characters).
— K. Bland, 1997: 204 (Philonthus; eaten by toad; Scotland).
— Staniec, 1999a: 56 (Philonthus; characters of pupa; Poland).
— Schillhammer, 2000: 153 (Philonthus; characters; Russia).
— Owen, 2000: 251 (Philonthus; pitfall trap collections from underground at roots of trees; Britain).
— Assing, 2001: 75 (Philonthus; collected from xerothermous site; Germany).
▴ Distribution: Europe, Russia.
rossicus Coiffait, 1965g: 616 (Sectophilonthus; Type locality: Vologolskaia obl., Evropeiskaia chast' SSSR).
— Coiffait, 1967a: 397 (Philonthus; notes).
— Coiffait, 1974: 6 (Palaeophilonthus; characters; Russia).
— Abdullah, 1975 (Sectophilonthus; aedeagal characters).
— Smetana, 1995e: 51 (Philonthus; transferred from Palaeophilonthus by reason of Smetana's synonym of the genus with Philonthus).
— Schillhammer, 2000: 153 (Philonthus; synonym of decorus).
definitus Cameron, 1950g: 35 (Philonthus; Type locality: Belgian Congo: vers Rweru, volc. Mikeno, alt. 2400 m).
dejectus Cameron, 1932: 108 (Philonthus; Type locality: Chakrata district; Bangar, alt. 6000 feet).
— Scheerpeltz, 1933: 1339 (Philonthus; catalog).
— Coiffait, 1982: 30 (Philonthus; Nepal).
▴ Distribution: India, Nepal.
delectus Last, 1968, see: Hesperus.
deleterius Tottenham, 1955: 160 (Philonthus; subgenus Spatulonthus; Type locality: Kenya: Muguga).
— Tottenham, 1962: 139 (Philonthus; subgenus Spatulonthus; Tanganyika).
▴ Distribution: Kenya, Tanzania.
deletus Sharp, 1876, see: Neobisnius.
delicatulus Boheman, 1858: 29 (Philonthus; Type locality: China).
— Fauvel, 1904a: 58 (Philonthus; Belgaum; Nilghiris; Ceylan; Bírmanie; Indochina; Annam; China; Sumatra; Java).
— Bernhauer and Schubert, 1914: 335 (Philonthus; catalog).
— Cameron, 1921a: 376, 405 (Philonthus; characters; Singapore).
— Cameron, 1932: 98 (Philonthus; characters; Ceylon).
— Scheerpeltz, 1933: 1339 (Philonthus; catalog).
— Cameron, 1933d: 386 (Philonthus; Sumatra).
— Scheerpeltz, 1954: 113 (Philonthus, cited as Phylonthus; Ceylon).
— Yadav, Chand, and Yadav, 1995: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
▴ Distribution: China, Indonesia, Singapore, Vietnam, Myanmar, India, Sri Lanka.
lativentris Motschulsky, 1858: 663 (Philonthus; Type locality: Indes orientales).
— Fauvel, 1904a: 58 (Philonthus; synonym of delicatulus).
— Bernhauer and Schubert, 1914: 335 (Philonthus; synonym of delicatulus).
— Cameron, 1932: 98 (Philonthus; synonym of delicatulus).
subirideus Kraatz, 1859: 81 (Philonthus; Type locality: Ceylan).
— Fauvel, 1904a: 58 (Philonthus; synonym of delicatulus).
— Bernhauer and Schubert, 1914: 335 (Philonthus; synonym of delicatulus).
— Cameron, 1932: 98 (Philonthus; synonym of delicatulus).
delusor Tottenham, 1955: 159 (Philonthus; subgenus Spatulonthus; Type locality: Abyssinia: Djem-Djem Forest).
densecaudatus Bernhauer, 1928g: 109 (Philonthus; Type locality: Belgisch-Kongo: Elisabethville, Pweto).
— Scheerpeltz, 1933: 1339 (Philonthus; catalog).
— Tottenham, 1949b: 345 (Philonthus; characters).
— Tottenham, 1962: 202, 203, 205 (Philonthus; characters; notes).
▴ Distribution: Congo, Ethiopia.
katompeanus Bernhauer, 1932b: 150 (Philonthus; Type locality: Katanga: Katompe).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
— Bernhauer and Chapman, 1939: 70 (Philonthus; Ethiopia).
— Tottenham, 1962: 205 (Philonthus; synonym of densecaudatus).
densicauda Bernhauer, 1943b: 290 (Philonthus; Type locality: Kongo: Elisabethville).
— Tottenham, 1955: 172 (Philonthus; characters).
— Tottenham, 1962: 202, 203, 207 (Philonthus; characters; Sudan).
▴ Distribution: Congo, Sudan.
densior Bernhauer, 1916, see: catamarcanus Bernhauer, 1916.
densipennis Bernhauer, 1908c: 110 (Philonthus; Type locality: Kamerun).
densiventris Bernhauer, 1915a: 24 (Philonthus; Type locality: Philippinen: Luzon: Mt. Makiling).
densus Cameron, 1926b: 355 (Philonthus; Type locality: Chakrata District; Chulli Khud, Sainj Khud, Korawa Khud, 6500 to 8000 feet).
— Cameron, 1932: 125 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1339 (Philonthus; catalog).
— Yadav, Chand, and Yadav, 1995: 6 (Philonthus; cited as denus; morphology of mitotic and meiotic chromosomes).
▴ Distribution: India.
dentiphallus Schillhammer, 1999: 60 (Philonthus; Type locality: China: Guizhou prov., Anshun Prefecture, Pingba, ca. 50 km SW Guiyang City).
dentipes Eppelsheim, 1889, see: Gabrius.
deplanatus Sharp, 1885: 423 (Philonthus; Type locality: Guatemala, San Geronimo).
depravus Cameron, 1932a: 141 (Philonthus; Type locality: Selangor: Bukit Kutu).
depressipennis Sharp, 1889: 37 (Philonthus; Type locality: Ishiuchi; Konose).
— Bernhauer and Schubert, 1914: 335 (Philonthus; catalog).
— Scheerpeltz, 1933: 1339 (Philonthus; catalog).
— Kanehisa, Tsumuki, and Kawazu, 1994 (Philonthus; chemical secretions).
▴ Distribution: Japan.
depressus Steinheil, 1869, see: pallipes Blanchard, 1842.
derennei Drugmand, 1987: 274 (Philonthus; subgenus Philonthus; Type locality: Belgique: province de Brabant, Boitsfort).
derivatus Cameron, 1937: 9 (Philonthus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft).
deuvei Coiffait, 1982a: 253 (Philonthus; Type locality: Inde: Pir Panjal, Bhadarwak, Samum, 3200 m).
devillei Chapman, 1932, see: alcyoneus Erichson, 1840.
diabolicus Cameron, 1942b: 327 (Philonthus; Type locality: Chyulu Hills, altitude 5600 feet).
diallus Tottenham, 1953, see: eidmannianus Scheerpeltz, 1929.
diamantinus Bernhauer, 1917i: 102 (Philonthus; Type locality: Bolivien: Yuracaris).
diatrechoides Tottenham, 1962: 232 (Philonthus; replacement name for nigrolucens Cameron; characters).
nigrolucens Cameron, 1959: 116 (Diatrechus; [preoccupied]; Type locality: Angola: Lunda N.E.: Riv. Tchikapa).
dictator Fagel, 1972: 101 (Philonthus; Type locality: Sainte-Hélène: Basse Fisher's Valley, 1000 ft.).
— Wollaston, 1877: 30 (Philonthus; [Note: Misidentified as turbidus according to Fagel, 1972: 101]; characters; St. Helena).
▴ Distribution: Saint Helena.
differens Cameron, 1932: 141 (Philonthus; Type locality: Nilgiri Hills).
diffusiventris Bernhauer, 1933, see: oberti Eppelsheim, 1889.
dignus Last, 1968: 347 (Philonthus; Type locality: New Guinea: Okapa).
uniformis Last, 1981: 120 (Philonthus; variety of dignus; [Note: Nomen nudum (Article 15.2); preoccupied]; cited from New Guinea: Garaina, Bulu Valley).
dilucidus Tottenham, 1962: 154 (Philonthus; Type locality: Nigeria: Ibadan).
dilutior Bernhauer, 1928, see: Mentophilonthus.
dilutipes Fauvel, 1898a: 117 (Philonthus; Type locality: La Digue; Iles Maurice et La Réunion. Madagascar: Diego Suarez; baie d'Antongil; Tananarive; Java oriental: Monts Tengger).
— H. Kolbe, 1910: 20 (Philonthus; Seychelles).
— Bernhauer and Schubert, 1914: 335 (Philonthus; catalog).
— Tottenham, 1949b: 339 (Philonthus; characters).
— Lecoq, 1990: 199 (Philonthus; Mascarene Islands).
▴ Distribution: Mauritius, Réunion, Madagascar, Seychelles, Indonesia.
dimidiaticornis Fauvel, 1905d: 197 (Philonthus; Type locality: Kioboscho, Kilimandjaro).
— Bernhauer and Schubert, 1914: 335 (Philonthus; catalog).
— Tottenham, 1949b: 309 (Philonthus).
— Tottenham, 1956: 225 (Philonthus; Ruanda).
— Tottenham, 1962: 135, 161, 165 (Philonthus; characters; Tanganyika; Nigeria; Natal).
▴ Distribution: Nigeria, Tanzania, Rwanda, South Africa.
dimidiatipennis Erichson, 1840: 500 (Philonthus; Type locality: Sicilia; Mesopotamia).
— Hochhuth, 1849: 145 (Philonthus; characters; Armenia).
— Kiesenwetter, 1851a: 417 (Philonthus; France).
— Fairmaire and Laboulbène, 1856: 530 (Philonthus; characters; France).
— Kraatz, 1857d: 621 (Philonthus).
— Hochhuth, 1862: 65 (Philonthus; Russia).
— Fauvel, 1874: 238 [= 1874a: 464] (Philonthus; characters; France; Sicily; Corsica; Sardinia; Algeria; Armenia; Mesopotamia; Russia).
— Solsky, 1874: 183 (Philonthus; characters; Turkestan).
— Fauvel, 1876a: 74 [= 1876: 259] (Philonthus; Spain; Turkestan; Buchara).
— Mulsant and Rey, 1876: 521 [= 1877: 377] (Philonthus; characters; France).
— Fauvel, 1878: 123 [= 1878a: 43] (Philonthus; Algeria).
— Fauvel, 1886: 55 [= 1886a: 47] (Philonthus; Algeria).
— Eppelsheim, 1892: 334 (Philonthus; Turkestan).
— Ganglbauer, 1895: 457 (Philonthus; characters; Mediterranean region; southern Russia).
— Fauvel, 1902b: 107 (Actobius; Morocco; Algeria).
— Sainte-Claire Deville, 1906: 97 (Philonthus; Corsica).
— Bernhauer and Schubert, 1914: 335 (Philonthus; catalog).
— Portevin, 1929: 364 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1340 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Bernhauer, 1936: 5 (Philonthus; Mongolia).
— Smetana, 1953: 119 (Philonthus; Turkey).
— Lohse, 1956: 91 (Philonthus; notes).
— Scheerpeltz, 1958a: 22 (Philonthus; Turkey).
— Scheerpeltz, 1961: 7 (Philonthus; Iran).
— Scheerpeltz, 1961h: 253 (Philonthus; Afghanistan).
— Raianu, 1963: 83 (Philonthus; Romania).
— Lohse, 1964: 185 (Philonthus; subgenus Philonthus; characters; central Europe).
— Focarile, 1964: 59 (Philonthus; Italy).
— Horion, 1965: 171 (Philonthus; North Africa; Turkey; Russia; Transcaucasus; Turkestan; Turkmenistan; northern China; Greece; France; Holland).
— Coiffait, 1967a: 431 (Philonthus; characters).
— Smetana, 1967d: 208 (Philonthus; Mongolia).
— Smetana, 1967i: 555 (Philonthus; notes; Turkey).
— Coiffait, 1974: 196 (Philonthus; characters; Europe; Anatolia; Mongolia; France; Corsica).
— Smetana 1975c: 172 (Philonthus; Mongolia).
— Zerche, 1975a (Philonthus; notes; Germany).
— Bordoni, 1975a: 153 (Philonthus; Afghanistan).
— Shilov, 1976: 152 (Philonthus; Mongolia).
— Pope, 1977: 29 (Philonthus; Britain).
— Coiffait, 1983e: 477 (Philonthus; Afghanistan).
— Tóth, 1984: 77 (Philonthus; characters).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Benelux).
— Lohse and Lucht, 1989: 168 (Philonthus; note).
— Hodge and Jones, 1995: 39 (Philonthus; characters; Britain).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
▴ Distribution: Algeria, Morocco, Spain, France, Italy, Greece, Holland, Romania, Russia, Armenia, Turkey, Iraq, Iran, Afghanistan, Uzbekistan, Mongolia.
rubellus Solsky, 1874: 179 (Philonthus; Type locality: Maracandam).
— Eppelsheim, 1892: 334 (Philonthus; variety of dimidiatipennis; Turkestan).
— Bernhauer and Schubert, 1914: 335 (Philonthus; variety of dimidiatipennis).
dimidiatus Say, 1830, see: sericans Gravenhorst, 1802.
dimidiatus C. Sahlberg, 1830, see: caucasicus Nordmann, 1837.
dimidiatus Lacordaire, 1835, see: quisquiliarius Gyllenhal, 1810.
dimorphus Bernhauer, 1903a: 29 (Philonthus; Type locality: Sumatra).
— Bernhauer and Schubert, 1914: 335 (Philonthus; catalog).
— Cameron, 1933d: 387 (Philonthus; Sumatra).
▴ Distribution: Indonesia.
dionysiae Levasseur, 1962a: 52 (Philonthus; Type locality: Kolwezi, Katanga).
discalis Tottenham, 1962, see: altivagans Fauvel, 1907.
discedens Schubert, 1909: 291 (Philonthus; Type locality: südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d'Orizaba).
discipennis Fauvel, 1879: 100 (Philonthus; Type locality: Nouvelle Guinée; Fly River).
— Bernhauer and Schubert, 1914: 335 (Philonthus; catalog).
— Cameron, 1937a: 100 (Philonthus; New Guinea).
— Last, 1987: 31 (Philonthus; Papua New Guinea).
▴ Distribution: New Guinea.
discoideisimilis Scheerpeltz, 1974: 149 (Philonthus; Type locality: Cape Prov.: Near Areb Mnt., ab 22 miles ENE Springbok. Also cited from South West Africa).
discoideus Gravenhorst, 1802: 38 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae; [Note: See lectotype designation by Smetana, 1995e]).
— Latreille, 1804: 322 (Staphylinus; characters; Germany; France).
— Gravenhorst, 1806: 58 (Staphylinus; characters; Germany; France).
— Gravenhorst, 1806: 60 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 336 cited testaceus Gravenhorst as a synonym of discoideus but Gravenhorst did not describe the species as new, he attributed it to Paykull]; characters; Germany; France).
— Gyllenhal, 1810: 331 (Staphylinus; characters; Sweden).
— Germar, 1827: 4 (Staphylinus; characters).
— C. Sahlberg, 1830: 324 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 29 (Staphylinus; Finland; Sweden).
— Mannerheim, 1831: 443 (Staphylinus; Finland; Sweden).
— Runde, 1835: 8 (Staphylinus; characters; Germany).
— Lacordaire, 1835: 403 (Staphylinus; characters; France).
— Nordmann, 1837: 97 (Philonthus; Sweden Germany; England).
— Erichson, 1839a: 469 (Philonthus; characters; Germany).
— Heer, 1839: 268 (Philonthus; characters; Switzerland).
— Stephens, 1839: 392 (Quedius; characters; England).
— Erichson, 1840: 474 (Philonthus; characters; Europe; Cuba).
— Jacobsen, 1849: 700 (Philonthus; characters; Denmark).
— Redtenbacher, 1849: 703 (Philonthus; characters; Austria).
— Küster, 1854: 12 (Philonthus; characters; Europe; Cuba).
— Hardy, 1851: 30 (Philonthus; notes; England).
— Rosenhauer, 1856: 71 (Philonthus; Cadiz).
— Fairmaire and Laboulbène, 1856: 526 (Philonthus; characters; France).
— Guérin-Méneville, 1857: 40 (Philonthus; characters).
— Wollaston, 1857: 190 (Philonthus; characters; habitat; Madeira Islands).
— Kraatz, 1857d: 605 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 193 (Philonthus; characters; Austria).
— Peyron, 1858: 426 (Philonthus; Turkey).
— Thomson, 1860: 160 (Philonthus; characters; Scandinavia).
— Chevrolat and Fauvel, 1863: 435 (Philonthus; characters; Cuba).
— Wollaston, 1864: 573 (Philonthus; notes; Canary Islands).
— Wollaston, 1865: 493 (Philonthus; habitat; notes; Madeira Islands; Canary Islands).
— Wollaston, 1867: 238 (Philonthus; characters; notes; Cape Verde Islands).
— Fauvel, 1874: 228 [= 1874a: 454] (Philonthus; characters; France; Barbary; Cyprus; Turkey; Buchara; Madeira, Canary, Cape Verde Islands; North America; Cuba).
— Redtenbacher, 1874: 212 (Philonthus; characters; Austria).
— Solsky, 1874: 179 (Philonthus; characters; Turkestan).
— Seidlitz, 1875: 276 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 485 [= 1877: 341] (Philonthus; characters; France).
— J. Sahlberg, 1876: 18 (Philonthus; Finland).
— Wollaston, 1877: 28 (Philonthus; characters; St. Helena).
— Fauvel, 1878: 123 [= 1878a: 43] (Philonthus; Barbary).
— Fauvel, 1878g: 547 (Philonthus; characters; Australia).
— Horn, 1884: 196 (Philonthus; characters; Atlantic region to Nevada and Arizona).
— Sharp, 1885: 409 (Philonthus; Mexico; Guatemala; Cuba; Australia).
— Blackburn and Sharp, 1885: 222 (Philonthus; habitat; Hawaii).
— Fauvel, 1886: 54 [= 1886a: 46] (Philonthus; Barbary).
— Fowler, 1888: 275 (Philonthus; characters; Britain).
— Fauvel, 1891: 115 (Philonthus; Venezuela).
— Ganglbauer, 1895: 455 (Philonthus; characters; Europe).
— Fauvel, 1902b: 110 (Philonthus; Barbary; Egypt; Madeira; Canaries).
— Sainte-Claire Deville, 1906: 96 (Philonthus; Corsica).
— Sharp, 1908: 548 (Philonthus; Hawaii).
— Reitter, 1909: 129 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 335 (Philonthus; catalog).
— Johansen, 1914: 410 (Philonthus; characters; Denmark).
— Bernhauer, 1920: 16 (Philonthus; Australia).
— Lea, 1925: 220 (Philonthus; Australian introduction).
— Portevin, 1929: 362 (Philonthus; characters; France).
— Gridelli, 1930: 74 (Philonthus; Libya).
— Cameron, 1932: 116 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1340 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Scheerpeltz, 1935a: 654 (Philonthus; Australia).
— Cameron, 1937: 5 (Philonthus; Java).
— Bernhauer, 1940c: 4 (Philonthus; Azores; Madeira).
— C. Koch, 1941: 63 (Philonthus; Morocco).
— Méquignon, 1942a: 21 (Philonthus; Azores).
— Blackwelder, 1943: 407 (Philonthus; characters; Cuba; Jamaica; Haiti; Puerto Rico; St. Croix; St. Kitts; Antigua; Montserrat; Guadeloupe; Grenada).
— V. Hansen, 1952: 73 (Philonthus; subgenus Philonthus; characters; Denmark).
— Cameron, 1951: 282 (Philonthus; Egypt).
— Cameron, 1952d: 824 (Philonthus; Mauritania).
— Scheerpeltz, 1956a: 484 (Philonthus; Belgian Congo).
— Smetana, 1958: 212 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 161 (Philonthus; characters).
— Tottenham, 1961: 204 (Philonthus; Senegal).
— Scheerpeltz, 1961: 6 (Philonthus; Iran).
— Tottenham, 1962: 216, 217, 221 (Philonthus; characters; Cape Verde; Senegal; Nigeria; Cameroon; Tanganyika; Rhodesia).
— Scheerpeltz, 1963b: 429 (Philonthus; Egypt).
— Palm, 1963b: 90 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 184 (Philonthus; subgenus Philonthus; characters; central Europe).
— Scheerpeltz, 1964a: 300 (Philonthus; subgenus Philonthus; Crete).
— Horion, 1965: 161 (Philonthus; widespread in southern and central Europe, sparse in northern Europe).
— Coiffait, 1967a: 422 (Philonthus; characters).
— Raianu, 1968a: 467 (Philonthus; Romania).
— Szujecki, 1968a: 731 (Philonthus; Poland).
— Fagel, 1972: 101 (Philonthus; notes; Sainte Helene).
— Moore and Legner, 1974a: 39 (Philonthus; spread by commerce).
— Coiffait, 1974: 195 (Philonthus; characters; Europe; Britain; Scandinavia; Anatolia; France).
— Scheerpeltz, 1974c: 22 (Philonthus; Sudan).
— Moore, 1975c: 106 (Philonthus; Hawaii).
— Coiffait, 1976: 217 (Philonthus; Tahiti).
— Pace and Zanetti, 1977: 297 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Smetana, 1978: 118 (Philonthus; collecting notes; Israel).
— Coiffait, 1979a: 162 (Philonthus; Saudi Arabia).
— Frank, 1982: 40 (Philonthus; list of parasites).
— Frank, 1984: 479 (Philonthus; characters; notes; Costa Rica).
— Tóth, 1984: 71 (Philonthus; characters; Hungary).
— Nekulisianu, 1985 (Philonthus; notes on biology; larval characters; Moldavia).
— Coiffait, 1986: 394 (Philonthus; Saudi Arabia).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Borges, 1990: Table II (Philonthus; Azores).
— Lecoq, 1990: 199 (Philonthus; Mascarene Islands).
— Watanabe, 1993b (Philonthus; Izu Islands).
— J. Li, 1993: 36 (Philonthus; China).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Nishida, 1994: 73 (Philonthus; Hawaii).
— Smetana, 1995e: 184 (Philonthus; lectotype designation; introduced species; characters; Ontario; Quebec; Arizona; California; Connecticut; Colorado; Florida; Georgia; Hawaii; Idaho; Illinois; Maine; Maryland; Massachusetts; Michigan; Nevada; New Hampshire; New Jersey; New York; Pennsylvania; Texas; Utah; Mexico).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Downie and Arnett, 1996: 378 (Philonthus; characters; USA).
— Gusarov, 1997: 287 (Philonthus; Saudi Arabia).
— Nishida, 1997: 62 (Philonthus; Hawaii).
— Watanabe, 1998a: 140 (Philonthus; Japan).
— Watanabe, 1998b: 262 (Philonthus; Japan).
— Lepeyre and Dauphin, 1999: 25 (Philonthus; teratological specimen; France).
— Assing and Wunderle, 2001: 37 (Philonthus; Cyprus).
▴ Distribution: Saint Helena, Cape Verde Islands, Canary Islands, Madeira, Azores, Zimbabwe, Mascarene Islands, Tanzania, Congo, Cameroon, Nigeria, Senegal, Mauritania, Morocco, Libya, Sudan, Egypt, Saudi Arabia, Israel, Europe, Russia, Turkey, Iran, Uzbekistan, China, India, Indonesia, Australia, Society Islands, Hawaii, Canada, USA, West Indies, Guatemala, Costa Rica, Venezuela.
suturalis Marsham, 1802: 509 (Staphylinus; Type locality: Britain).
— Gyllenhal, 1810: 331 (Staphylinus; synonym of discoideus).
— Germar, 1827: 4 (Staphylinus; synonym of discoideus).
— Blackwelder, 1943: 407 (Philonthus; synonym of discoideus).
— Smetana, 1958: 213 (Philonthus; synonym of discoideus).
— Coiffait, 1974: 195 (Philonthus; synonym of discoideus).
— Smetana, 1995e: 184 (Philonthus; synonym of discoideus).
lepidulus Stephens, 1832: 223 (Quedius; Type locality: London; Suffolk; Devonshire).
— Stephens, 1839: 392 (Quedius; characters; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of discoideus).
— Gemminger and Harold, 1868: 587 (Philonthus; synonym of discoideus).
— Bernhauer and Schubert, 1914: 336 (Philonthus; synonym of discoideus).
— Cameron, 1932: 116 (Philonthus; synonym of discoideus).
— Blackwelder, 1943: 407 (Philonthus; synonym of discoideus).
— Smetana, 1958: 213 (Philonthus; synonym of discoideus).
— Coiffait, 1974: 195 (Philonthus; synonym of discoideus).
— Smetana, 1995e: 185 (Philonthus; synonym of discoideus).
— Herman, 2001: 49 (Philonthus; nomen oblitum).
suturalis Stephens, 1832: 224 (Quedius; [preoccupied]; Type locality: London; Norfolk; Suffolk).
— Stephens, 1839: 392 (Quedius; synonym of discoideus).
— Hardy, 1851: 30 (Philonthus; synonym of discoideus).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of discoideus).
— Bernhauer and Schubert, 1914: 336 (Philonthus; synonym of discoideus).
— Cameron, 1932: 116 (Philonthus; synonym of discoideus).
— Blackwelder, 1943: 407 (Philonthus; synonym of discoideus).
conformis Lacordaire, 1835: 398 (Staphylinus; Type locality: Paris).
— Erichson, 1839a: 469 (Philonthus; synonym of discoideus).
— Erichson, 1840: 474 (Philonthus; synonym of discoideus).
— Fairmaire and Laboulbène, 1856: 526 (Philonthus; synonym of discoideus).
— Guérin-Méneville, 1857: 40 (Philonthus; synonym of discoideus).
— Kraatz, 1857d: 605 (Philonthus; synonym of discoideus).
— Ganglbauer, 1895: 455 (Philonthus; synonym of discoideus).
— Bernhauer and Schubert, 1914: 336 (Philonthus; synonym of discoideus).
— Cameron, 1932: 116 (Philonthus; synonym of discoideus).
— Blackwelder, 1943: 407 (Philonthus; synonym of discoideus).
— Smetana, 1958: 213 (Philonthus; synonym of discoideus).
— Coiffait, 1974: 195 (Philonthus; synonym of discoideus).
— Smetana, 1995e: 185 (Philonthus; synonym of discoideus).
ruficornis Melsheimer, 1844: 38 (Philonthus; Type locality: Pennsylvania; [Note: See lectotype designation by Smetana, 1995e]).
— Fauvel, 1891: 115 (Philonthus; synonym of discoideus).
— Ganglbauer, 1895: 455 (Philonthus; synonym of discoideus).
— Bernhauer and Schubert, 1914: 336 (Philonthus; synonym of discoideus).
— Cameron, 1932: 116 (Philonthus; synonym of discoideus).
— Blackwelder, 1943: 407 (Philonthus; synonym of discoideus).
— Smetana, 1958: 213 (Philonthus; synonym of discoideus).
— Coiffait, 1974: 195 (Philonthus; synonym of discoideus).
— Smetana, 1995e: 185 (Philonthus; lectotype designation: Pennsylvania; synonym of discoideus).
rufipennis Gerhardt, 1910: 555 (Philonthus; aberration of discoideus; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]; cited from Liegnitz).
— Cameron, 1932: 116 (Philonthus; aberration of discoideus).
— Scheerpeltz, 1933: 1340 (Philonthus; synonym of gerhardtianus).
— Blackwelder, 1943: 407 (Philonthus; synonym of discoideus).
— Smetana, 1959: 161 (Philonthus; synonym of discoideus).
— Smetana, 1995e: 185 (Philonthus; synonym of discoideus).
— Coiffait, 1974: 195 (Philonthus; synonym of discoideus).
gerhardtianus Scheerpeltz, 1933: 1340 (Philonthus; variety of discoideus replacement name for rufipennis Gerhardt).
— Smetana, 1958: 213 (Philonthus; aberration of discoideus).
— Smetana, 1959: 161 (Philonthus; aberration of discoideus; characters).
— Coiffait, 1967a: 422 (Philonthus; aberration of discoideus; characters).
— Coiffait, 1974: 195 (Philonthus; variety of discoideus).
— Tóth, 1984: 72 (Philonthus; aberration of discoideus).
— Smetana, 1995e: 185 (Philonthus; synonym of discoideus).
discrepens Sharp, 1889: 38 (Philonthus; Type locality: Nikko).
— Bernhauer and Schubert, 1914: 336 (Philonthus; catalog).
— Scheerpeltz, 1933: 1340 (Philonthus; catalog).
▴ Distribution: Japan.
discretus Sharp, 1876b: 428 (Philonthus; Type locality: Mexico City).
— Sharp, 1885: 427 (Philonthus; notes; Mexico).
— Bernhauer and Schubert, 1914: 336 (Philonthus; catalog).
▴ Distribution: Mexico.
disjunctus Bernhauer and Schubert, 1914, see: Gabrius.
disjunctus Cameron, 1925, see: Gabrius.
dispersepunctatus Scheerpeltz, 1965: 209 (Philonthus; Type locality: N.E. Burma, Kambaiti, 2000 m).
dispersus Fauvel, 1891: 111 (Philonthus; Type locality: Colonie Tovar; Lagonite; Nouvelle Grenade).
— Bernhauer and Schubert, 1914: 336 (Philonthus; catalog).
— Marcuzzi, 1977: 11 (Philonthus; Venezuela).
▴ Distribution: Venezuela, Colombia.
dispersus Fauvel, 1895, see: fauvelianus Bernhauer and Schubert, 1914.
dissentaneus Tottenham, 1961: 210 (Philonthus; Type locality: Senegal: Parc National du Niokolo-Koba, Badi).
dissimilis Baudi, 1870, see: Neobisnius.
distans Fauvel, 1895, see: Gabrius.
distans Horn, 1884, see: Gabrius.
distincticolle Cameron, 1932, see: vagus Fauvel, 1895.
distincticornis Cameron, 1932: 87 (Philonthus; Type locality: Sikkim: Tonglo).
— Scheerpeltz, 1933: 1340 (Philonthus; catalog).
— Coiffait, 1983d: 161 (Philonthus; Nepal).
— Schillhammer, 1998: 107 (Philonthus; characters; Nepal, India).
▴ Distribution: India, Nepal.
antennalis Cameron, 1932: 262 (Philonthopsis; Type locality: Darjeeling Dist.: Ghum, alt. 8000 feet).
— Scheerpeltz, 1933: 1418 (Philonthopsis; catalog).
— Schillhammer, 1994: 74 (Philonthus; synonym of distincticornis).
— Schillhammer, 1998: 107 (Philonthus; synonym of distincticornis).
aeneipennis Cameron, 1932: 262 (Philonthopsis; variety of antennalis; [preoccupied]; Type locality: Tonglu, 10,074 feet, on the Nepal-Sikkim frontier).
— Scheerpeltz, 1933: 1418 (Philonthopsis; variety of antennalis).
— Schillhammer, 1998: 107 (Philonthus; synonym of distincticornis).
cyaneonitens Schillhammer, 1991: 51 (Philonthus; Type locality: Nepal: Parbat Distr.: Ghoropani Pass, n slope, 2800 m).
— Schillhammer, 1994: 74 (Philonthus; synonym of distincticornis).
— Schillhammer, 1998: 107 (Philonthus; synonym of distincticornis).
distinctus Gemminger and Harold, 1868: 587 (Philonthus; replacement name for antennatus Motschulsky).
— Bernhauer and Schubert, 1914: 336 (Philonthus; catalog).
— Cameron, 1932: 133 (Philonthus; characters; East Indies).
— Scheerpeltz, 1933: 1340 (Philonthus; catalog).
▴ Distribution: “East Indies”.
antennatus Motschulsky, 1858: 664 (Philonthus; [preoccupied]; Type locality: Indes orientales).
— Bernhauer and Schubert, 1914: 336 (Philonthus; synonym of distinctus).
— Cameron, 1932: 133 (Philonthus; synonym of distinctus).
distinguendus Cameron, 1950, see: Gabrius.
distinguendus Cameron, 1951, see: secus Herman, 2001.
divergens Cameron, 1937: 6 (Philonthus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft).
diversiceps Bernhauer, 1901b: 653 (Philonthus; replacement name for armeniacus Bernhauer).
— Bernhauer and Schubert, 1914: 336 (Philonthus; catalog).
— Gridelli, 1920a: 20 (Philonthus; characters; Italy).
— Scheerpeltz, 1933: 1340 (Philonthus; catalog; Bulgaria; Sardinia).
— Gridelli, 1943: 130 (Philonthus; Macedonia; Sardinia; Italy; Dalmatia).
— Smetana, 1955: 207 (Philonthus; notes).
— Smetana, 1958: 214 (Philonthus; subgenus Philonthus; characters; ecology).
— Smetana, 1959: 162 (Philonthus; characters).
— Lohse, 1964: 184 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 167 (Philonthus; Macedonia; Sardinia; Bulgaria; Albania; Hungary; Austria; Slovakia).
— Coiffait, 1967a: 425 (Philonthus; characters; Caucasus; Asia Minor; Balkans; central Europe).
— Coiffait, 1974: 202 (Philonthus; characters; Transcaucasus; Caucasus; Russia; Europe).
— Pace and Zanetti, 1977: 297 (Philonthus; Italy).
— Tóth, 1984: 74 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Austria).
— Schuh, Schillhammer, and Zettel, 1992: 221 (Philonthus; Austria).
— Schillhammer, 1993: 326 (Philonthus; Austria).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Schillhammer, 1995: 231 (Philonthus; Austria).
— Assing and Wunderle, 2001: 37 (Philonthus; Cyprus).
▴ Distribution: Turkey, Armenia, Russia, Europe.
armeniacus Bernhauer, 1901: 538 (Philonthus; [preoccupied]; Type locality: Kaukasus, armenisches Gebirge).
— Bernhauer and Schubert, 1914: 336 (Philonthus; synonym of diversiceps).
— Smetana, 1958: 214 (Philonthus; synonym of diversiceps).
— Smetana, 1959: 162 (Philonthus; synonym of diversiceps).
— Coiffait, 1974: 202 (Philonthus; synonym of diversiceps).
diversipennis Bernhauer, 1899, see: Bisnius.
diversus Schubert, 1906a: 382 (Philonthus; Type locality: Kashmir, Sintan).
— Bernhauer and Schubert, 1914: 336 (Philonthus; catalog).
— Cameron, 1932: 95 (Philonthus; characters; Kashmir).
— Scheerpeltz, 1933: 1340 (Philonthus; catalog).
— Shilov, 1976: 151 (Philonthus; Mongolia).
▴ Distribution: India, Mongolia.
dives Erichson, 1847, see: Phanolinus.
divisifrons Cameron, 1950, see: Gabrius.
divisus Sharp, 1891: 41 (Philonthus; Type locality: Machachi, 9–10,000 feet).
dohertyi Cameron, 1932: 102 (Philonthus; Type locality: Burma: Tavoy).
dolichoderes Lea, 1925: 221 (Philonthus; Type locality: Lord Howe Island).
dollmani Bernhauer, 1934g: 236 (Pseudophilonthus; Type locality: N.W. Rhodesia: Kashitu, N. of Broken Hill; Baudouinville im belgischen Kongo).
— Tottenham, 1949b: 362 (Philonthus; characters).
— Levasseur, 1962: 242 (Philonthus; characters; Zaire).
▴ Distribution: Congo, Zambia.
donckieri Bernhauer, 1915i: 254 (Philonthus; Type locality: Südliches Brit. Indien: Madura, Chambaganur).
— Cameron, 1932: 78 (Philonthus; characters; Ceylon; India).
— Scheerpeltz, 1933: 1340 (Philonthus; catalog).
— Scheerpeltz, 1976a: 87 (Philonthus; Nepal).
— Coiffait, 1981d: 323 (Philonthus; Nepal).
— Coiffait, 1982: 29 (Philonthus; cited as “dockkieri Bernhauer, 1911: 254”; characters; Nepal).
— Coiffait, 1982a: 233 (Philonthus; India).
— Schillhammer, 1999a: 61 (Philonthus).
— Schillhammer, 2000: 126 (Philonthus; characters; China; India; Nepal; Laos; Vietnam).
▴ Distribution: China, Nepal, India, Sri Lanka, Laos, Veitnam.
tripunctatus Cameron, 1919a: 253 (Philonthus; Type locality: Ceylon: Dikoya, alt. 3800–4200; [Note: See lectotype designation by Schillhammer, 2000]).
— Cameron, 1932: 78 (Philonthus; synonym of donckieri).
— Scheerpeltz, 1933: 1341 (Philonthus; synonym of donckieri).
— Schillhammer, 2000: 126 (Philonthus; lectotype designation; synonym of donckieri).
crebrior Bernhauer, 1931c: 129 (Philonthus; Type locality: Hongkong).
— Scheerpeltz, 1933: 1338 (Philonthus; catalog).
— Schillhammer, 1999a: 61 (Philonthus; synonym of donckieri).
— Schillhammer, 2000: 126 (Philonthus; synonym of donckieri).
donisthorpei Dollmann, 1910, see: intermedius Lacordaire, 1835.
doriae Fauvel, 1878f: 262 (Philonthus; Type locality: Nouvelle-Guinée, Hatam).
— Fauvel, 1879: 101 (Philonthus; Amberbaki; Mysol).
— Bernhauer and Schubert, 1914: 336 (Philonthus; catalog).
▴ Distribution: New Guinea.
dorjulensis Coiffait, 1977, see: Gabrius.
dorylophilus Cameron, 1933: 42 (Philonthus; Type locality: Belgian Congo: Nizi: Blukwa).
downiei Hatch, 1957, see: Gabrius.
drescheri Cameron, 1937: 8 (Philonthus; Type locality: Java: Batoerraden. G. Slamat).
dryas Smetana, 1995e: 435 (Philonthus; Type locality: Arizona: Wickenburg, Maricopa Co.).
dubiosus Bernhauer, 1936h: 324 (Philonthus; Type locality: Kenya Colony: Kaimosi).
dubius Heer, 1839, see: Quedius.
dubius LeConte, 1863, see: Cafius.
duodecimpunctatus Bernhauer, 1932b: 149 (Philonthus; Type locality: Haut-Uele: Moto).
kapanganus Bernhauer, 1936a: 22 (Philonthus; subspecies of duodecimpunctatus; Type locality: Lulua-Gebiet: Kapanga).
duplex Bernhauer, 1920d: 182 (Philonthus; Type locality: Ostindien).
— Cameron, 1932: 105 (Philonthus; characters; “East Indies”).
— Scheerpeltz, 1933: 1341 (Philonthus; catalog).
▴ Distribution: “East Indies”.
duplicatus Bernhauer and Schubert, 1914: 336 (Philonthus; replacement name for basalis Horn).
— Hatch, 1957: 197 (Philonthus; characters; Washington; Oregon; Idaho).
— Smetana, 1995e: 142 (Philonthus; characters; notes; Alberta; British Columbia; Manitoba; Northwest Territories; Saskatchewan; Yukon Territory; Alasaka; California; Colorado; Idaho; Kansas; Oregon; Utah; Washington; Wyoming).
▴ Distribution: Canada, USA.
basalis Horn, 1884: 187 (Philonthus; [preoccupied]; Type locality: Alma, Colorado, 10,000 feet; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from Swift Current, British America; Wyoming).
— Fall, 1926: 149 (Philonthus; Yukon Territory).
— Smetana, 1965a: 6 (Philonthus; characters; British Columbia; Alaska).
— Smetana, 1995e: 142 (Philonthus; lectotype designation: Alma, Colorado, 10,000 feet; synonym of duplicatus).
duplicatus Tikhomirova, 1973, see: wuesthoffi Bernhauer, 1938.
duplopunctatus Motschulsky, 1860, see: rotundicollis Ménétriés, 1832.
duploseriatus Solsky, 1868a: 136 (Philonthus; Type locality: Mexique).
— Sharp, 1885: 427 (Philonthus; Mexico).
— Bernhauer and Schubert, 1914: 336 (Philonthus; catalog).
▴ Distribution: Mexico.
ebeninus Gravenhorst, 1802: 170 (Staphylinus; Type locality: Brunsvigae).
— Latreille, 1804: 323 (Staphylinus; characters; Germany).
— Gyllenhal, 1810: 337 (Staphylinus; characters; illustration).
— C. Sahlberg, 1830: 325 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 30 (Staphylinus; Finland; Sweden; Germany).
— Mannerheim, 1831: 444 (Staphylinus; Finland; Sweden; Germany).
— Lacordaire, 1835: 394 (Staphylinus; characters; France).
— Nordmann, 1837: 93 (Philonthus; Sweden; Germany; France).
— Erichson, 1839a: 461 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 336 cited varians Erichson as a synonym of ebeninus Gravenhorst; Erichson did not describe the species as new, he attributed it to Fabricius who attributed it to Paykull]; characters; Germany).
— Erichson, 1839a: 462 (Philonthus; synonym of varians).
— Heer, 1839: 264 (Philonthus; synonym of varians).
— Erichson, 1840: 461 (Philonthus; characters; Europe; Siberia; Asia minor; Syria; Egypt).
— Laporte, 1840: 178 (Staphylinus; characters; France).
— Grimm, 1845: 135 (Philonthus; with Formica rufa; Germany).
— P. Lucas, 1846: 112 (Philonthus; notes; Algeria).
— Jacobsen, 1849: 696 (Philonthus; characters; Denmark).
— Ménétriés, 1849: 44 (Philonthus; Turcomanie).
— Hochhuth, 1849: 138 (Philonthus; characters; Caucasus; Imeretia).
— Redtenbacher, 1849: 703 (Philonthus; characters; Austria).
— Kiesenwetter, 1851a: 417 (Philonthus; France; Spain).
— Rosenhauer, 1856: 71 (Philonthus; Malaga).
— Fairmaire and Laboulbène, 1856: 523 (Philonthus; characters; France).
— Kraatz, 1857d: 596 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 193 (Philonthus; characters; Austria).
— Kraatz, 1858a: 61 (Philonthus; Crete).
— Peyron, 1858: 425 (Philonthus; Turkey).
— Thomson, 1860: 159 (Philonthus; characters; Scandinavia).
— Hochhuth, 1862: 58 (Philonthus; Russia).
— Fauvel, 1869: 488 (Philonthus; synonym of brevicornis).
— Redtenbacher, 1874: 212 (Philonthus; characters; Austria).
— Solsky, 1874: 177 (Philonthus; characters; Turkestan).
— Seidlitz, 1875: 276 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 445 [= 1877: 301] (Philonthus; characters; France).
— J. Sahlberg, 1876: 17 (Philonthus; Finland).
— Fauvel, 1878: 123 [= 1878a: 43] (Philonthus; Barbary).
— Fauvel, 1886: 55 [= 1886a: 47] (Philonthus; Barbary).
— Fowler, 1888: 270 (Philonthus; characters; Britain).
— Eppelsheim, 1892: 332 (Philonthus; Turkestan).
— Ganglbauer, 1895: 451 (Philonthus; characters; Palaearctic region).
— Fauvel, 1902b: 110 (Philonthus; Barbary; Madeira; Canaries).
— Sainte-Claire Deville, 1906: 96 (Philonthus; Corsica).
— Montandon, 1908: 80 (Philonthus; Romania).
— Reitter, 1909: 129 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 336 (Philonthus; catalog).
— Johansen, 1914: 404 (Philonthus; characters; Denmark).
— Gridelli, 1920a: 15 (Philonthus; characters; Sicily; Corsica; Sardinia; Italy).
— Gridelli, 1922a: 176 (Philonthus; Persia).
— Scheerpeltz, 1933: 1341 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; cited as ebenius; illustration of aedeagus).
— Jeannel and Jarrige, 1949: 374 (Philonthus; cave species; Hungary).
— V. Hansen, 1952: 63 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1953: 117 (Philonthus; Turkey).
— Scheerpeltz, 1956c: 1100 (Philonthus; Cyprus).
— Smetana, 1958: 172 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 157 (Philonthus; characters).
— Palm, 1963b: 82 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 182 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 157 (Philonthus; widespread in southern and central Europe, sparse in northern Europe).
— Coiffait, 1967a: 415 (Philonthus; characters).
— Szujecki, 1968a: 728 (Philonthus; Poland).
— Jarrige, 1971: 495 (Philonthus; Iran).
— Coiffait, 1974: 260 (Philonthus; characters; Palaearctic region).
— Shilov, 1976: 149 (Philonthus; Mongolia).
— Pace and Zanetti, 1977: 290 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Frank, 1982: 40 (Philonthus; list of parasites).
— Tóth, 1984: 67 (Philonthus; characters; Hungary).
— Lucht, 1987: 104 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Gusarov, 1991a: 8 (Philonthus; notes).
— Wittwer, 1993: 250 (Philonthus; Switzerland).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Sterrenburg and Schülke, 1997: 18 (Philonthus; Netherlands).
▴ Distribution: Egypt, Algeria, Canary Islands, Madeira, Europe, Russia, Georgia, Turkey, Syria, Iran, Turkmenistan, Mongolia.
brevicornis Gravenhorst, 1802: 22 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).
— Latreille, 1804: 312 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 69 (Staphylinus).
— Nordmann, 1837: 94 (Philonthus; Germany).
— Erichson, 1839a: 462 (Philonthus; synonym of varians).
— Erichson, 1840: 461 (Philonthus; synonym of ebeninus).
— P. Lucas, 1846: 112 (Philonthus; synonym of ebeninus).
— Kraatz, 1857d: 596 (Philonthus; synonym of ebeninus).
— Ganglbauer, 1895: 451 (Philonthus; synonym of ebeninus).
— Bernhauer and Schubert, 1914: 336 (Philonthus; synonym of ebeninus).
— Scheerpeltz, 1933: 1341 (Philonthus; synonym of ebeninus).
— Smetana, 1958: 172 (Philonthus; synonym of ebeninus).
— Coiffait, 1974: 260 (Philonthus; synonym of ebeninus).
longicornis Waltl, 1835: 57 (Staphylinus; variety of quisquilarius; [preoccupied]; Type locality: Andalusien).
— Erichson, 1840: 929 (Philonthus; [Note: Cited as quisquiliarius variety longicornis]; synonym of ebeninus).
lhesgicus Kolenati, 1846: 19 (Philonthus; Type locality: mons Ssalwat Caucasi Lhesgici).
— Hochhuth, 1849: 136 (Philonthus; characters).
— Fauvel, 1874: 234 [= 1874a: 459] (Philonthus; synonym of ebeninus).
— Ganglbauer, 1895: 451 (Philonthus; synonym of ebeninus).
— Bernhauer and Schubert, 1914: 336 (Philonthus; synonym of ebeninus).
— Smetana, 1958: 172 (Philonthus; synonym of ebeninus).
— Coiffait, 1974: 260 (Philonthus; synonym of ebeninus).
incompletus Hochhuth, 1849: 153 (Quedius; Type locality: Tiflis).
— Bernhauer and Schubert, 1914: 359 (Philonthus; synonym of varians).
— Cameron, 1932: 131 (Philonthus; synonym of varians).
— Scheerpeltz, 1933: 1366 (Philonthus; synonym of varians).
— Blackwelder, 1943: 412 (Philonthus; synonym of varians).
— Smetana, 1958: 195 (Philonthus; synonym of varians).
— Gusarov, 1991a: 8 (Philonthus; synonym of ebeninus).
— Smetana, 1995e: 235 (Philonthus; synonym of varians [Note: I assume Smetana overlooked Gusarov, 1991]).
setosus J. Sahlberg, 1876: 14 (Philonthus; Type locality: Kontschosero i Ryska Karelen).
— Bernhauer and Schubert, 1914: 355 (Philonthus; catalog).
— Scheerpeltz, 1933: 1360 (Philonthus; catalog).
— Smetana, 1958: 172 (Philonthus; synonym of ebeninus).
— Coiffait, 1974: 260 (Philonthus; synonym of ebeninus).
— Shilov, 1975a: 376 [= 1975b: 98] (Philonthus; Komi region of Russia).
quadripunctatus Fiori, 1894: 91 (Philonthus; Type locality: Monte Paderno, nelle colline Bolognesi; Catanzaro).
— Bernhauer and Schubert, 1914: 336 (Philonthus; synonym of ebeninus).
— Smetana, 1958: 172 (Philonthus; synonym of ebeninus).
— Coiffait, 1974: 260 (Philonthus; synonym of ebeninus).
eccoptomus Tottenham, 1962: 231 (Philonthus; subgenus Eccoptolonthus; Type locality: Natal).
egelegus Last, 1981: 121 (Philonthus; Type locality: Romad).
egens Sharp, 1874, see: Gabrius.
egregius Erichson, 1840, see: Styngetus.
eidmanni Scheerpeltz, 1929, see: Bisnius.
eidmannianus Scheerpeltz, 1929a: 121 (Philonthus; Type locality: Umgebung von Hang-Tschou in der Provinz Tschekiang in China).
— Scheerpeltz, 1933: 1341 (Philonthus; catalog).
— Coiffait, 1967a: 424 (Philonthus; characters; China).
— Coiffait, 1974: 295 (Philonthus; characters; China).
— J. Li, 1993: 35 (Philonthus; China).
▴ Distribution: China, Japan.
diallus Tottenham, 1953: 145 (Philonthus; Type locality: China: Jentschoufu, Schant.).
— Coiffait, 1967a: 424 (Philonthus; characters; China).
— Coiffait, 1974: 200 (Philonthus; characters; China).
— Hayashi, 1994b: 140 (Philonthus; Japan).
— Schillhammer, 1996a: 82 (Philonthus; synonym of eidmannianus).
elatus Erichson, 1840, see: Diatrechus.
electus Bernhauer and Schubert, 1914: 336 (Philonthus; replacement name for elegantulus Schubert).
elegantulus Schubert, 1906: 373 (Philonthus; [preoccupied]; Type locality: Kwai, Usambara).
elegans Sharp, 1885: 428 (Philonthus; Type locality: Panama, Peña Blanca 3000 to 4000 feet).
elegantissimus Cameron, 1925a: 186 (Philonthus; Type locality: Tjibodas, 1400 m).
elegantulus Luze, 1904: 97 (Philonthus; Type locality: Kisil-kum mer.: Kalma-tai).
— Bernhauer and Schubert, 1914: 337 (Philonthus; catalog).
— Scheerpeltz, 1933: 1341 (Philonthus; catalog).
— Pototskaia, 1966c: 940 (Philonthus; larval characters).
— Coiffait, 1967a: 410 (Philonthus; characters; Turkestan).
— Coiffait, 1974: 288 (Philonthus; characters; Turkestan; Transcaspian region).
▴ Distribution: Turkmenistan.
elegantulus Schubert, 1906, see: electus Bernhauer and Schubert, 1914.
elgonensis Tottenham, 1940d: 149 (Philonthus; Type locality: Uganda: Mt. Elgon, 10,000 ft.).
— Tottenham, 1949b: 315 (Philonthus; characters).
— Tottenham, 1956: 226 (Philonthus; Ruanda).
— Tottenham, 1962: 164 (Philonthus; characters).
▴ Distribution: Uganda, Rwanda.
elongatulus Erichson, 1839, see: Neobisnius.
elypticus Bernhauer, 1932b: 150 (Philonthus; Type locality: Haut-Uele: Moto).
elytratus Coiffait, 1963 [Note: Nomen nudum], see: frigidoides Coiffait, 1963.
emas Schillhammer, 1998: 111 (Philonthus; Type locality: E-Malaysia: Sabah Crocker Range, Gg. Emas, 1500–1700 m).
emdeni Bernhauer, 1931: 2 (Philonthus; Type locality: Szetschwan: Sunpanting).
— Scheerpeltz, 1933: 1341 (Philonthus; catalog).
— Coiffait, 1967a: 398 (Philonthus; cited as emdemi; characters; China).
— Coiffait, 1974: 280 (Philonthus; cited as endemi; characters; China).
— Schillhammer, 1999a: 61 (Philonthus).
— Schillhammer, 2000: 155 (Philonthus; characters; China).
▴ Distribution: China.
kiulungensis Bernhauer, 1933a: 42 (Philonthus; Type locality: Tatsienlu-Kiulung).
— Schillhammer, 1999a: 61 (Philonthus; synonym of emdeni).
— Schillhammer, 2000: 155 (Philonthus; synonym of emdeni).
emelinae Coiffait and Saiz, 1968: 354 (Philonthus; subgenus Philonthus; Type locality: Chile: Romeral, Curicó).
enodis Broun, 1880: 111 (Philonthus; Type locality: Parua).
ephippium Nordmann, 1837: 111 (Philonthus; Type locality: Grusinia).
— Erichson, 1840: 507 (Philonthus; characters; Grusia).
— Kraatz, 1857d: 619 (Philonthus; variety of punctus).
— Solsky, 1870: 460 (Philonthus; notes; characters; Astrakhan).
— Bernhauer and Schubert, 1914: 337 (Philonthus; catalog).
— Scheerpeltz, 1933: 1341 (Philonthus; catalog).
— Coiffait, 1967a: 434 (Philonthus; characters; Russia; Turkestan).
— Smetana, 1967d: 208 (Philonthus; Mongolia).
— Coiffait, 1974: 269 (Philonthus; characters; Russia; Transcaucasus; Transcaspian region; Mongolia).
— Smetana 1975c: 172 (Philonthus; Mongolia).
— Shilov, 1976: 152 (Philonthus; Mongolia).
▴ Distribution: Georgia, Russia, Kyrgyzstan, Mongolia.
scutellatus Motschulsky, 1860: 565 (Philonthus; Type locality: bords du fl. Oural et les steppes des Kirguises).
— Hochhuth, 1862: 64 (Philonthus; characters; Ural River; Kirghiz Steppes).
— Solsky, 1870: 460 (Philonthus; synonym of ephippium).
— Fauvel, 1875a: XXX [= 1875b: 232] (Philonthus; synonym of ephippium).
— Bernhauer and Schubert, 1914: 337 (Philonthus; synonym of ephippium).
— Smetana, 1967d: 208 (Philonthus; synonym of ephippium).
uralensis Reitter, 1909: 131 (Philonthus; Type locality: Südrussland: Uralsk).
— Bernhauer and Schubert, 1914: 358 (Philonthus; catalog).
— Scheerpeltz, 1933: 1365 (Philonthus; catalog).
— Smetana, 1965f: 257 (Philonthus; synonym of ephippium).
— Smetana, 1967d: 208 (Philonthus; synonym of ephippium).
— Coiffait, 1974: 269 (Philonthus; synonym of ephippium).
eppelsheimi Coiffait, 1967, see: indubius Luze, 1904.
eremus Gistel, 1857: 79 (Philonthus; [Note: Nomen dubium]; Type locality: Helvetia).
eriwanensis Bernhauer and Schubert, 1914: 337 (Philonthus; replacement name for sanguinipennis Reitter).
— Scheerpeltz, 1933: 1341 (Philonthus; catalog).
— Coiffait, 1967a: 395 (Philonthus; cited as erevanensis; characters; Transcaucasus).
— Coiffait, 1974: 226 (Philonthus; characters; Transcaucasus; Armenia).
▴ Distribution: Azerbaijan, Armenia.
sanguinipennis Reitter, 1898: 115 (Philonthus; [preoccupied]; Type locality: Araxesthal bei Ordubad).
erraticus Nordmann, 1837: 90 (Philonthus; Type locality: Rossia meridionalis).
erythropterus Eichelbaum, 1910, see: rudipennis Fauvel, 1907.
erythropterus Fauvel, 1872, see: turbatus Erichson, 1840.
erythropus Kraatz, 1859: 88 (Philonthus; Type locality: Ceylan; India australis; India borealis).
— Fauvel, 1879: 104 (Philonthus; characters; notes; New Guinea; Celebes; Philippines; China; India; Ceylon).
— Fauvel, 1886b: 150 (Philonthus; Philippines; Ceylon; East Indies; China; Annam; Sumatra; Java; New Guinea).
— Sharp, 1889: 38 (Philonthus; Japan).
— Fauvel, 1903: 159 (Philonthus; synonym of aeneipennis).
— Bernhauer and Schubert, 1914: 327 (Philonthus; synonym of aeneipennis).
— Cameron, 1932: 113 (Philonthus; synonym of aeneipennis).
— Gridelli, 1936a: 150 (Philonthus; characters; valid species).
— Scheerpeltz, 1957c: 234 (Philonthus; valid species; Sumba).
— Coiffait, 1967a: 424 (Philonthus; characters; China; Southeast Asia).
— Coiffait, 1974: 207 (Philonthus; characters).
— Coiffait, 1981b: 236 (Philonthus; Saudi Arabia).
— Coiffait, 1982: 28 (Philonthus; Nepal).
— Coiffait, 1986: 394 (Philonthus; Saudi Arabia).
▴ Distribution: Saudi Arabia, Sri Lanka, India, Nepal, Vietnam, Indonesia, New Guinea, Philippines, China.
kuluensis Schubert, 1908: 617 (Philonthus; Type locality: Kulu, Himalaya, ca. 1500 m).
— Bernhauer and Schubert, 1914: 343 (Philonthus; catalog).
— Cameron, 1932: 108 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
— Scheerpeltz, 1976a: 88 (Philonthus; Nepal).
— Schillhammer, 1991: 54 (Philonthus; synonym of erythropus).
erythrostictus Kraatz, 1859, see: rubricollis Motschulsky, 1858.
erythrostomus Hochhuth, 1851, see: Gabrius.
eryx Smetana, 1995e: 306 (Philonthus; Type locality: California: Ventura Co.: Fillmore, Santa Clara River. Also cited from Arizona, Mexico).
escurialensis Perez, 1865, see: Rabigus.
euphorbiarum Peyerimhoff, 1949: 251 (Philonthus; subgenus Gefyrobius; Type locality: Maroc atlantique: Agadir et bords de l'Ocean, entre Agadir et Mogador).
— Coiffait, 1967a: 417 (Philonthus; characters; Morocco).
— Coiffait, 1974: 247 (Philonthus; characters; Morocco).
▴ Distribution: Morocco.
eustilboides Cameron, 1933d: 390 (Philonthus; Type locality: Sumatra: Gunung Singgalang).
eustilbus Kraatz, 1859: 98 (Philonthus; Type locality: Ceylan).
— Bernhauer and Schubert, 1914: 337 (Philonthus; catalog).
— Cameron, 1932: 150 (Philonthus; characters; Ceylon; India).
— Scheerpeltz, 1933: 1341 (Philonthus; catalog).
— Cameron, 1933d: 387 (Philonthus; Sumatra).
— Scheerpeltz, 1954: 114 (Philonthus, cited as Phylonthus; Ceylon).
— Coiffait, 1982a: 232 (Philonthus; India).
▴ Distribution: India, Sri Lanka, Indonesia.
versicolor Cameron, 1919a: 254 (Philonthus; Type locality: Ceylon).
— Cameron, 1932: 150 (Philonthus; synonym of eustilbus).
— Scheerpeltz, 1933: 1341 (Philonthus; synonym of eustilbus).
evitendus Tottenham, 1939, see: Philonthellus.
exactus Schubert, 1911: 27 (Philonthus; Type locality: Usambara, Nguelo).
excellens Bernhauer, 1916c: 37 (Philonthus; Type locality: West-Columbien: Umgebung von Cali).
excelsior Bernhauer, 1939a: 83 (Philonthus; Type locality: Kenya: Marakwet, Elgeyo Escarpment, 2500 m).
excelsus Cameron, 1931: 361 (Hesperus; [Note: Cameron, 1937a: 101 stated that, in the original description, because of a printer's error, this species was erroneously cited in Hesperus rather than Philonthus]; Type locality: Mount Lamington, New Guinea).
— Scheerpeltz, 1933: 1372 (Hesperus; catalog).
— Cameron, 1937a: 101 (Philonthus; New Guinea).
▴ Distribution: New Guinea.
excelsus Cameron, 1932, see: Gabrius.
excelsus Bernhauer, 1941, see: supernus Herman, 2001.
excisicauda Bernhauer, 1939, see: Gabrius.
exemplaris Wendeler, 1956a: 231 (Philonthus; Type locality: note cited but presumably Brazil).
exiguus Nordmann, 1837, see: Gabrius.
exilis Kraatz, 1851, see: Gabronthus.
explanipes Tottenham, 1939, see: simpliciventris Bernhauer, 1933.
explorator Cameron, 1932: 104 (Philonthus; Type locality: Nilgiri Hills).
— Scheerpeltz, 1933: 1341 (Philonthus; catalog).
— Coiffait, 1982: 31 (Philonthus; Nepal).
— Hayashi, 1997b: 38 (Philonthus; Taiwan; Thailand).
▴ Distribution: India, Nepal, Thailand; Taiwan.
exsectus Tottenham, 1956: 285 (Philonthus; Type locality: Ruanda: Rutovu, forêt du Rugege, 2350 m).
extensus Korge, 1971 [Note: Nomen nudum], see: velatipennis Solsky, 1870.
extinctus Bernhauer, 1900, see: cruentatus Gmelin, 1790.
extremus Sharp, 1885: 415 (Philonthus; Type locality: Guatemala, San Geronimo).
ezoensis Hayashi, 1993, see: prolatus Sharp, 1874.
fascialis Tottenham, 1962: 164 (Philonthus; Type locality: Not cited, presumably Africa).
fageli Levasseur, 1962: 234 (Philonthus; Type locality: Zilo [Lualaba], Haut Katanga, Congo belge).
fagelianus Tikhomirova, 1973, see: wuesthoffi Bernhauer, 1938.
fagniezi Jarrige, 1951a: 8 (Philonthus; Type locality: Bou-Berak, Kabylie).
— Coiffait, 1963: 12 (Paragabrius).
— Smetana, 1965f: 257 (Philonthus; Algeria).
— Coiffait, 1974: 95 (Paragabrius; characters; Algeria).
▴ Distribution: Algeria.
falcozi Coiffait, 1987, see: variipennis Kraatz, 1859.
fallaciosus Horn, 1884, see: Gabrius.
falsesordidus Coiffait, 1982a: 254 (Philonthus; Type locality: Pakistan: Lalazar, 3000 m).
fasciatus Nordmann, 1837, see: Oligotergus.
fasciiventris Jarrige, 1948a: 19 (Philonthus; Type locality: Madagascar: Montagne d'Ambre).
fassli Bernhauer, 1916c: 29 (Philonthus; Type locality: Peru: Huancabamba).
fatalis Tottenham, 1956: 286 (Philonthus; Type locality: Urundi: Bururi, 1800–2000 m).
fauvelii Coquerel, 1866, see: Gabrius.
fauvelianus Bernhauer and Schubert, 1914: 337 (Philonthus; replacement name for dispersus Fauvel).
— Cameron, 1932: 119 (Philonthus; characters; Burma).
— Scheerpeltz, 1933: 1342 (Philonthus; catalog).
▴ Distribution: Myanmar.
dispersus Fauvel, 1895: 264 (Philonthus; [preoccupied]; Type locality: Bírmanie, Carin Asciuii Ghécu, 1400–1500 m).
fauvelianus Cameron, 1925: 59 (Philonthus; [preoccupied]; replacement name for disperus Fauvel).
fauvelianus Cameron, 1925, see: fauvelianus Bernhauer and Schubert, 1914.
femineus Hatch, 1957, see: Gabrius.
femoralis Hochhuth, 1851, see: Gabrius.
femoralis Mäklin, 1853, see: Cafius.
fenderi Hatch, 1957, see: Gabrius.
fenestratus Fauvel, 1872b: 35 (Philonthus; Type locality: Bône; Ténériffe).
— Wollaston, 1854: 583 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 337 cited bipustulatus Wollaston as a synonym of fenestratus; Wollaston did not describe the species as new, he attributed it to Panzer who attributed it to Linné]; characters; notes; Madeira Islands).
— Wollaston, 1864: 570 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 337 cited bipustulatus Wollaston as a synonym of fenestratus; Wollaston did not describe the species as new, he attributed it to Panzer who attributed it ot Linné]; characters; Canary Islands).
— Fauvel, 1874: 253 [= 1874a: 479] (Philonthus; characters; France; Sicily; Algeria; Madeira; Canary Islands).
— Fauvel, 1876a: 75 [= 1876: 260] (Philonthus; Spain; Tunisia).
— Mulsant and Rey, 1876: 469 [= 1877: 325] (Philonthus; characters; France).
— Fauvel, 1878: 125 [= 1878a: 45] (Philonthus; Algeria; Tunisia).
— Fauvel, 1886: 56 [= 1886a: 48] (Philonthus; Morocco; Tunisia; Algeria).
— Fauvel, 1902b: 112 (Philonthus; Barbary; Egypt; Canaries).
— Sainte-Claire Deville, 1906: 99 (Philonthus; Corsica).
— Bernhauer and Schubert, 1914: 337 (Philonthus; catalog).
— Gridelli, 1920a: 18, 20 (Philonthus; characters; France; Spain; Caucasus; Algeria; Canary Islands; Italy).
— Portevin, 1929: 363 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1342 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Smetana, 1953: 118 (Philonthus; Turkey).
— Scheerpeltz, 1956c: 1100 (Philonthus; Cyprus).
— Smetana, 1963b: 36 (Philonthus; subgenus Philonthus; Madeira).
— Smetana, 1965f: 255 (Philonthus; notes).
— Coiffait, 1967a: 419 (Philonthus; characters).
— Coiffait, 1974: 242 (Philonthus; characters; North Africa; Sicily; Sardinia; Spain; Madeira; Canary Islands; France; Britain).
— Pace and Zanetti, 1977: 294 (Philonthus; Italy).
— Tóth, 1984: 70 (Philonthus; characters).
— Borges, 1990: Table II (Philonthus; Azores).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
▴ Distribution: Egypt, Tunisia, Algeria, Morocco, Canary Islands, Madeira, Azores, Britain, Spain, France, Italy, Cyprus, Turkey, Caucasus.
concolor Gridelli, 1920a: 18, 19, 20 (Philonthus; aberration of fenestratus; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]; cited from Isola di Unie, Quarnero, Istria).
— Scheerpeltz, 1933: 1342 (Philonthus; aberration of fenestratus).
— Normand, 1947a: 8 (Philonthus; aberration of fenestratus; Tunisia).
— Smetana, 1963b: 36 (Philonthus; subgenus Philonthus; aberration of fenestratus; Madeira).
— Coiffait, 1974: 243 (Philonthus; variety of fenestratus).
— Tóth, 1984: 70 (Philonthus; variety of fenestratus).
fenyesi Bernhauer, 1910b: 381 (Philonthus; Type locality: Mexiko: Vera Cruz, Cordoba).
feralis Erichson, 1840: 469 (Philonthus; Type locality: Columbia; [Note: See lectotype designation by Smetana, 1994]).
— Guérin-Méneville, 1855: 583 (Philonthus; Brazil).
— Solsky, 1868a: 135 (Philonthus; Mexico).
— Sharp, 1885: 416 (Philonthus; Mexico; Guatemala; Costa Rica; Panama; Venezuela).
— Fauvel, 1891: 115 (Philonthus; synonym of longicornis).
— Fauvel, 1903d: 271 (Philonthus; synonym of longicornis).
— Bernhauer and Schubert, 1914: 337 (Philonthus; catalog).
— Scheerpeltz, 1933: 1349 (Philonthus; synonym of longicornis).
— Blackwelder, 1943: 410 (Philonthus; synonym of longicornis).
— Smetana, 1958: 189 (Philonthus; synonym of longicornis).
— Smetana, 1994: 346 (Philonthus; characters; lectotype designation: Colombia; valid species; Colombia).
▴ Distribution: Brazil, Venezuela, Colombia, Panama, Costa Rica, Guatemala, Mexico.
ferreipennis Horn, 1884: 202 (Philonthus; Type locality: Washington Territory; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from Washington Territory to the southern part of California).
— Bernhauer and Schubert, 1914: 338 (Philonthus; catalog).
— Hatch, 1957: 184 (Philonthus; characters; British Columbia; Idaho; Washington; Oregon).
— Smetana, 1963a: 5 (Philonthus; aedeagus).
— Smetana, 1995e: 348 (Philonthus; lectotype designation: W.T [Washington Territory]; characters; notes; British Columbia; California; Idaho; Nevada; Oregon; Washington).
▴ Distribution: Canada, USA, Mexico.
stigmatias Smetana, 1995e: 350 (Philonthus; subspecies of ferreipennis; Type locality: California: San Diego Co.: Mt. Laguna. Also cited from Arizona, Colorado, New Mexico, Utah and Mexico).
ferreirai Scheerpeltz, 1978: 209 (Philonthus; Type locality: Dadim-Bere, Timor).
ferreiranus Scheerpeltz, 1978: 215 (Philonthus; Type locality: Dadim-Bere, Timor).
ferrugatus Erichson, 1840, see: Belonuchus.
fervidus Erichson, 1840, see: Eugastus.
fibularius Erichson, 1840: 442 (Philonthus; Type locality: Mexico).
— Sharp, 1885: 402 (Philonthus; characters; Mexico).
— Bernhauer and Schubert, 1914: 338 (Philonthus; catalog).
▴ Distribution: Mexico.
boucardi Solsky, 1868a: 127 (Philonthus; Type locality: Mexique).
— Sharp, 1885: 404 (Philonthus; notes; Mexico).
— Bernhauer and Schubert, 1914: 338 (Philonthus; synonym of fibularius).
figulus Erichson, 1840: 464 (Philonthus; Type locality: Brasilia).
— R. Sahlberg, 1847: 790 (Philonthus; Brazil).
— Chevrolat and Fauvel, 1863: 434 (Philonthus; characters; Brazil, Cayenne, Cuba).
— Fauvel, 1877a: xxviii (Philonthus; Brazil).
— Sharp, 1885: 415 (Philonthus; notes; Mexico; Guatemala; Nicaragua; Costa Rica; Panama).
— Fauvel, 1891: 115 (Philonthus; Venezuela).
— Fauvel, 1901: 86 (Philonthus; Central America; Cuba; Saint Vincent; Venezuela; Guyana; Colombia; Bolivia; Brazil; Paraguay).
— Bernhauer and Schubert, 1914: 338 (Philonthus; catalog).
— Scheerpeltz, 1933: 1342 (Philonthus; catalog; Argentina).
— Blackwelder, 1943: 409 (Philonthus; characters; Cuba; Puerto Rico; St. Vincent; Trinidad; Guatemala).
▴ Distribution: Argentina, Paraguay, Bolivia, Brazil, French Guiana, Guyana, Venezuela, Colombia, Panama, Costa Rica, Nicaragua, Guatemala, Mexico, West Indies.
amazonicus Sharp, 1876: 165 (Philonthus; Type locality: Para, Ega).
— Sharp, 1885: 415 (Philonthus; synonym of figulus).
— Fauvel, 1891: 115 (Philonthus; synonym of figulus).
— Bernhauer and Schubert, 1914: 338 (Philonthus; synonym of figulus).
— Blackwelder, 1943: 409 (Philonthus; synonym of figulus).
fijiensis Cameron, 1943b: 565 (Philonthus; Type locality: Fiji: Waidina River).
filator Tottenham, 1953: 147 (Philonthus; Type locality: Turkestan: Aulie Ata, Syr Daria. Also cited from Transcaspia and East Siberia).
— Coiffait, 1967a: 425 (Philonthus; characters; Turkestan; Siberia).
— Coiffait, 1974: 201 (Philonthus; characters; Turkestan; Transcaspian region; Siberia).
▴ Distribution: Russia, Uzbekistan, Tajikistan.
rubrofilator Coiffait, 1967a: 441 (Philonthus; subgenus Philonthus; variety of filator; Type locality: Djiliktchl, Tougail, Tadjikistan).
filicornis Horn, 1884, see: Bisnius.
filiformis Wollaston, 1854, see: Neobisnius.
filius Last, 1981: 124 (Neobisnius; Type locality: New Guinea: Musa).
filum Kiesenwetter, 1849, see: Remus.
fimbriolatus Erichson, 1840, see: Gabrius.
fimetarioides Scheerpeltz, 1976, see: Gabrius.
finitimus Casey, 1915, see: stictus Hausen, 1891.
finitoris Last, 1981: 122 (Philonthus; Type locality: Musa).
finmarkicus Munster, 1921, see: Bisnius.
fischeri Bernhauer, 1903a: 26 (Philonthus; Type locality: Sumatra).
— Bernhauer and Schubert, 1914: 338 (Philonthus; catalog).
— Cameron, 1933d: 387 (Philonthus; Sumatra).
▴ Distribution: Indonesia.
fissatus Tottenham, 1962: 210 (Philonthus; Type locality: Congo: Haut-Uele, Watsa. Also cited from Uganda and Natal).
fissilis Sharp, 1885: 399 (Philonthus; Type locality: Guatemala, Totonicapam, 8000 to 10,500 feet; [Note: See lectotype designation by Smetana, 1991a]).
— Smetana, 1991a: 234 (Philonthus; lectotype designation: Guatemala, Totonicapam, 8000 to 10,500 feet; characters; Guatemala).
▴ Distribution: Guatemala.
flagrans Erichson, 1840, see: Chroaptomus.
flandrianus Cameron, 1952: 329 (Philonthus; Type locality: Belgian Congo: Tshuapa: Flandria).
flavibasis Casey, 1915: 437 (Philonthus; Type locality: Montana: Helena; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1342 (Philonthus; catalog).
— Hatch, 1957: 186 (Philonthus; characters; British Columbia).
— Smetana, 1995e: 385 (Philonthus; lectotype designation: Helena, Montana; characters; notes; Alberta; Manitoba; Ontario; Quebec; Saskatchewan; Connecticut; Indiana; Maryland; Massachusetts; Michigan; Montana; New York; Pennsylvania; Wisconsin).
▴ Distribution: Canada, USA.
flavicauda Bernhauer, 1936h: 325 (Philonthus; Type locality: Kenya Colony: Kaimosi; Kakamega, Yala River).
— Tottenham, 1949b: 306 (Philonthus; characters; Belgian Congo).
— Cameron, 1950g: 37 (Philonthus; Belgian Congo).
— Tottenham, 1962: 160 (Philonthus; characters).
▴ Distribution: Kenya, Congo.
flavicornis Fairmaire, 1892, see: infirmus Erichson, 1840.
flavicoxis Bernhauer, 1912d: 43 (Philonthus; Type locality: Argentinien: Prov. Tucuman).
flavimanus Gemminger and Harold, 1868, see: Gabrius.
flavipennis Erichson, 1840, see: Leptopeltus.
flavipes Kraatz, 1859: 88 (Philonthus; Type locality: Ceylan).
— Bernhauer, 1902b: 27 (Philonthus; Ceylon).
— Fauvel, 1904a: 58 (Philonthus; Belgaum; Kanara; Ceylan; Bírmanie; Siam; Tonkin; Indochina; Sumatra).
— Bernhauer and Schubert, 1914: 338 (Philonthus; catalog).
— Bernhauer, 1922a: 151 (Philonthus; Samoa).
— Bernhauer, 1928: 16 (Philonthus; Sumatra).
— Cameron, 1932: 110 (Philonthus; characters; Ceylon).
— Scheerpeltz, 1933: 1342 (Philonthus; catalog; Mauritius; Réunion).
— Cameron, 1933d: 386 (Philonthus; Sumatra).
— Last, 1964: 177 (Philonthus; Brunei).
— Lecoq, 1990: 199 (Philonthus; Mascarene Islands).
— Hayashi, 1994c: 4 (Philonthus; Japan).
▴ Distribution: Mascarene Islands, Sri Lanka, India, Myanmar, Thailand, Vietnam, Indonesia, Brunei, Samoa, Japan.
flaviventris Jarrige, 1948a: 21 (Philonthus; Type locality: Madagascar: Montagne d'Ambre).
flavocinctus Motschulsky, 1858: 663 (Philonthus; Type locality: Indes orientales).
— Fauvel, 1904a: 58 (Philonthus; Belgaum; Ceylan; Bírmanie; Annam; Maurice; Réunion).
— Bernhauer and Schubert, 1914: 338 (Philonthus; catalog).
— Cameron, 1921a: 377, 405 (Philonthus; characters; Singapore).
— Bernhauer, 1922c: 232 (Philonthus; Formosa).
— Cameron and Scott, 1928: 272 (Philonthus; Rodriguez Island).
— Cameron, 1932: 116 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1342 (Philonthus; catalog).
— Cameron, 1933d: 387 (Philonthus; Sumatra).
— Cameron, 1934d: 79 (Philonthus; type examined; two specimens representing leucopygus and minutus).
— Cameron, 1950b: 214 (Philonthus; Sudan).
— Jarrige, 1957: 110 (Philonthus; Réunion).
— Scheerpeltz, 1966e: 402 (Philonthus; New Caledonia).
— Legner and Moore, 1977: 176 (Philonthus; found under bark).
— Frank, 1982: 40 (Philonthus; list of parasites).
— Lecoq, 1990: 197 (Philonthus; Mascarene Islands).
— Shibata, 1992: 188 (Philonthus; Ryukyu Archipelago).
▴ Distribution: Sudan, Mascarene Islands, Sri Lanka, India, Myanmar, Vietnam, Singapore, Indonesia, New Caledonia, Taiwan, Japan.
rufomarginatus Kraatz, 1859: 90 (Philonthus; Type locality: Ceylan).
— Fauvel, 1904a: 58 (Philonthus; synonym of flavocinctus).
— Bernhauer and Schubert, 1914: 338 (Philonthus; synonym of flavocinctus).
— Cameron, 1932: 116 (Philonthus; synonym of flavocinctus).
— Jarrige, 1957: 110 (Philonthus; synonym of flavocinctus).
flavolimbatus Erichson, 1840: 471 (Philonthus; Type locality: Columbia; [Note: See lectotype designation by Smetana, 1995e]).
— Erichson, 1840: 929 (Philonthus; characters; Carolina meridionalis).
— Horn, 1884: 191 (Philonthus; characters; Gulf States and Arizona, extending to South America).
— Sharp, 1885: 416 (Philonthus; Mexico; Guatemala; Nicaragua; Panama; Colombia; Brazil; South Carolina).
— Fauvel, 1891: 116 (Philonthus; Venezuela).
— Fauvel, 1901: 86 (Philonthus; southern United States; Central America; Martinique; Venezuela; Peru; Brazil; Argentina).
— Fall and Cockerell, 1907: 166 (Philonthus; New Mexico).
— Bernhauer and Schubert, 1914: 338 (Philonthus; catalog).
— Notman, 1920b: 706 (Philonthus; Florida).
— Blackwelder, 1943: 414 (Philonthus; characters; Martinique; Granadines; Texas; Arizona).
— Scheerpeltz, 1975a: 243 (Philonthus; collecting notes; Brazil).
— Marcuzzi, 1977: 12 (Philonthus; Venezuela).
— Frank, 1982: 40 (Philonthus; list of parasites).
— Hunter, Kincher, Bay, Beerwinkle, 1991: 168 (Philonthus; seasonal distribution and diel flight activity; Texas).
— Smetana, 1995e: 200 (Philonthus; lectotype designation: Colombia; characters; notes; Arizona; California; Colorado; Florida; Georgia; Kansas; Mississippi; Missouri; New Mexico; New York; North Carolina; South Carolina; South Dakota; Tennessee; Texas; Utah; Mexico).
— Hu and Frank, 1995b: 584 (Philonthus; surface structure of eggs).
— Hu and Frank, 1997 (Philonthus; predation of eggs and larvae of Haematobia irritans; Florida).
▴ Distribution: Canada, USA, West Indies, Mexico, Guatemala, Nicaragua, Panama, Colombia, Venezuela, Brazil, Peru, Argentina.
apicipennis Lynch, 1884: 155 (Philonthus; Type locality: Chacabuco; América Meridional, Setentrional).
— Fauvel, 1891: 116 (Philonthus; synonym of flavolimbatus).
— Bernhauer and Schubert, 1914: 338 (Philonthus; synonym of flavolimbatus).
— Blackwelder, 1943: 412 (Philonthus; synonym of flavolimbatus).
— Smetana, 1995e: 200 (Philonthus; synonym of flavolimbatus).
wacoensis Casey, 1915: 430 (Philonthus; Type locality: Texas: Waco; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1368 (Philonthus; catalog).
— Smetana, 1995e: 200 (Philonthus; lectotype designation: Waco, Texas; synonym of flavolimbatus).
flavoscutellatus Bernhauer, 1928g: 110 (Philonthus; Type locality: Belg.-Kongo: Mongende).
flavoterminatus Wollaston, 1877, see: peliomerus Kraatz, 1859.
fletcheri Cameron, 1943: 129 (Philonthus; Type locality: Kashmir: Gulmarg).
flohri Sharp, 1876b: 429 (Philonthus; Type locality: Mexico City; [Note: See lectotype designation by Smetana, 1991a]).
— Sharp, 1885: 398 (Philonthus; variety of furvus; Mexico).
— Bernhauer and Schubert, 1914: 339 (Philonthus; variety of furvus).
— Scheerpeltz, 1971b: 205 (Philonthus; Guatemala; Mexico; El Salvador; Nicaragua).
— Smetana, 1991a: 231 (Philonthus; valid species; lectotype designation: Mexico City; characters; Mexico).
▴ Distribution: Mexico, Guatemala, El Salvador, Nicaragua.
flumineus Casey, 1915: 439 (Philonthus; Type locality: Missouri: St. Louis; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1342 (Philonthus; catalog).
— Smetana, 1965: 23 (Philonthus; subgenus Philonthus; [Note: Misidentified as flumineus according to Smetana, 1995e: 405]; characters; Nova Scotia; Newfoundland).
— Smetana, 1995e: 405 (Philonthus; lectotype designation: St. Louis, Missouri; characters; notes; Manitoba; New Brunswick; Newfoundland; Nova Scotia; Ontario; Quebec; Alabama; Arkansas; Connecticut; Georgia; Illinois; Indiana; Iowa; Kansas; Kentucky; Louisiana; Maine; Maryland; Massachusetts; Michigan; Minnesota; Mississippi; Missouri; Montana; New Hampshire; New Jersey; New York; North Carolina; Ohio; Oklahoma; Pennsylvania; Rhode Island; Tennessee; Texas; Vermont; Virginia; West Virginia; Wisconsin).
▴ Distribution: Canada, USA.
fluviatilis Cameron, 1932, see: Gabrius.
foetidus Cameron, 1932: 129 (Philonthus; Type locality: Chakrata district: Kanasar, Chulli Khud, Bodyar, alt. 7500–8300 feet. Simla Hills, alt. 7600 feet).
— Scheerpeltz, 1933: 1342 (Philonthus; catalog).
— Scheerpeltz, 1965: 107 (Philonthus; Burma).
— Coiffait, 1982: 31 (Philonthus; Nepal).
— Yadav, Chand, and Yadav, 1994: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
— Hayashi, 1997b: 38 (Philonthus; Thailand; Taiwan).
▴ Distribution: India, Nepal, Myanmar, Thailand, Taiwan.
fonticola Erichson, 1840, see: Cafius.
forcipatus Hatch, 1957, see: Gabrius.
formaneki Roubal, 1911: 6 (Philonthus; Type locality: Kr. Poljana).
— Bernhauer and Schubert, 1914: 353 (Philonthus; synonym of rotundicollis).
— Cameron, 1932: 84 (Philonthus; synonym of rotundicollis).
— Gridelli, 1943: 128 (Philonthus; valid species; characters).
— Smetana, 1965f: 254 (Philonthus; notes).
— Jarrige, 1971: 494 (Philonthus; Iran).
▴ Distribution: Iran, Russian Caucasus.
incompletus Roubal, 1911: 7 (Philonthus; variety of formaneki; [preoccupied]; Type locality: Kr. Poljana).
— Bernhauer and Schubert, 1914: 353 (Philonthus; synonym of rotundicollis).
— Cameron, 1932: 84 (Philonthus; aberration of rotundicollis).
— Gridelli, 1943: 128 (Philonthus; variety of formaneki).
formicarius Bernhauer, 1908d: 334 (Philonthus; Type locality: Brasilien: Mar d'Hespanha).
formosae Bernhauer, 1922c: 232 (Philonthus; Type locality: Kanku, Pilam, Alikang).
formosae Cameron, 1949, see: taiwanensis Shibata, 1993.
forsteri Scheerpeltz, 1960c: 104 (Philonthus; Type locality: Bolivien: Songotal, Cuticucho, 3700 m).
forsterianus Scheerpeltz, 1960c: 110 (Philonthus; Type locality: Bolivien: Yungas de Corami, 2500 m).
forticornis Cameron, 1937: 12 (Philonthus; Type locality: Java: Batoerraden. G. Slamat).
fortunatus Wollaston, 1865, see: ventralis Gravenhorst, 1802.
foveifrons Cameron, 1932, see: Gabronthus.
foveifrons J. Sahlberg, 1913 [Note: Nomen nudum (Articles 1.3.4, 45.5)], see: Bisnius.
franzi Schillhammer, 1998: 111 (Philonthus; Type locality: Medan, - O. Küste v. Sumatra).
fraseri Cameron, 1950d: 19 (Philonthus; Type locality: Pahang: Frazer's Hill, 4200 feet).
frater Bernhauer, 1907, see: albilabris Nordmann, 1837.
fraternus Cameron, 1926, see: cameroni Scheerpeltz, 1933.
fraternus Fall, 1926, see: quadricollis Horn, 1884.
fraxinatus Tottenham, 1949b: 341 (Philonthus; Type locality: Upper Senegal: Badoumbe).
freyi Bernhauer, 1939k: 591 (Philonthus; Type locality: China: Tienmuschan).
— Coiffait, 1967a: 422 (Philonthus; characters; China).
— Coiffait, 1974: 295 (Philonthus; characters; China).
— Hayashi, 1994b: 140 (Philonthus; Japan).
▴ Distribution: China, Japan.
friebi Bernhauer, 1927, see: Bisnius.
frigidoides Coiffait, 1963: 25 (Philonthus; Type locality: Liban: Col de Dahr el Baïdar. Also cited from Anatolia).
— Coiffait, 1967a: 400 (Philonthus; characters; Lebanon; Asia Minor; Transcaucasus).
— Coiffait, 1974: 255 (Philonthus; characters; Lebanon; Anatolia).
— Coiffait, 1978d: 164 (Philonthus; Turkey).
▴ Distribution: Lebanon, Turkey, Caucasus.
elytratus Coiffait, 1963: 27 (Philonthus; aberration of frigidoides; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Liban: Col de Dahr el Baïdar. Also cited from Anatolia).
— Coiffait, 1967a: 400 (Philonthus; aberration of frigidoides; characters).
— Coiffait, 1974: 255 (Philonthus; variety of frigidoides).
frigidus Märkel and Kiesenwetter, 1848a: 324 (Philonthus; Type locality: Kärnthner Alpen).
— Hochhuth, 1851: 17 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 339 cited janthinipennis Hochhuth as a synonym of frigidus; Hochhuth did not describe the species as new, he attributed it to Kolenati]; characters; notes).
— Kraatz, 1857d: 590 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 192 (Philonthus; characters; Austria).
— Fauvel, 1874: 235 [= 1874a: 461] (Philonthus; characters; France; Germany; Switzerland; Austria; Italy; Spain; Caucasus; Turkey).
— Fauvel, 1876a: 74 [= 1876: 259] (Philonthus; Italy).
— Redtenbacher, 1874: 211 (Philonthus; characters; Austria).
— Mulsant and Rey, 1876: 408 [= 1877: 264] (Philonthus; characters; France).
— Ganglbauer, 1895: 448 (Philonthus; characters; middle Europe; Spain; Asia Minor; Caucasus).
— Reitter, 1909: 128 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 339 (Philonthus; catalog).
— Portevin, 1929: 361 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1343 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Aleksandrov, 1934: 153 (Philonthus; China).
— C. Koch, 1941: 63 (Philonthus; Morocco).
— Smetana, 1958: 186 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 150 (Philonthus; characters).
— Coiffait, 1963: 24 (Philonthus; characters; notes; Alps).
— Lohse, 1964: 179 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 144 (Philonthus; Caucasus; Turkey; Lebanon; France; Switzerland; Italy; Germany; Austria; Bosnia Hercegovina; Serbia; Macedonia; Bulgaria; Romania; Urals).
— Coiffait, 1967a: 400 (Philonthus; characters; Alps; Massif Central).
— Raianu, 1968a: 466 (Philonthus; Romania).
— Coiffait, 1974: 253 (Philonthus; characters; Alps; Bulgaria; Greece; Caucasus).
— Pace and Zanetti, 1977: 294 (Philonthus; Italy).
— Tóth, 1984: 56 (Philonthus; characters; Hungary).
— Lucht, 1987: 104 (Philonthus; checklist; Germany; Austria).
— Lohse and Lucht, 1989: 165 (Philonthus; characters).
— Pilon and Zanetti, 1991: 57 (Philonthus; Italy).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Slovakia ?).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Assing, Frisch, Kahlen, et al., 1998: 137 (Philonthus; note).
▴ Distribution: Europe, Georgia, Turkey.
consputus Baudi, 1848: 127 (Philonthus; Type locality: valle Augustana).
— Kraatz, 1857b: 176 (Philonthus; synonym of frigidus).
— Kraatz, 1857d: 574 (Philonthus).
— Kraatz, 1857d: 1060 (Philonthus; synonym of frigidus).
— Ganglbauer, 1895: 448 (Philonthus; synonym of frigidus).
— Bernhauer and Schubert, 1914: 339 (Philonthus; synonym of frigidus).
— Smetana, 1958: 186 (Philonthus; synonym of frigidus).
— Coiffait, 1974: 253 (Philonthus; synonym of frigidus).
balkanicus R. Dvořák and Havelka, 1952: 41 (Philonthus; variety of albipes; Type locality: Yougoslavie: Serbia: Kozuf Planina).
— R. Dvořák, 1954a: 9 (Philonthus; subspecies of frigidus; [Note: Cited as a new subspecies but also stated to be the same as the 1954 use by Dvořák and Havelka]; Balkan region).
— Smetana, 1955b: 223 (Philonthus; variety of frigidus; characters).
vitochensis Coiffait, 1974: 143, 253, 254 (Philonthus; subspecies of frigidus; Type locality: Bulgarie: Aleko, Mont Vitocha près de Sofia vers 2000 m. Also cited from Greece).
svanetiensis Coiffait, 1974: 143, 253, 254 (Philonthus; subspecies of frigidus; Type locality: Caucase: Mestia Massif Svanetia, vers 2400 m).
frontalis Nordmann, 1837, see: Quedius.
frontalis Schubert, 1908, see: obsoletus Eppelsheim, 1895.
fuentei Roubal, 1909, see: Bisnius.
fugax Faldermann, 1835: 125 (Staphylinus; Type locality: [area] Persico-Armeniaca).
— Hochhuth, 1849: 134 (Philonthus; characters; Caucasus).
— Bernhauer and Schubert, 1914: 339 (Philonthus; catalog).
— Scheerpeltz, 1933: 1343 (Philonthus; catalog).
— Coiffait, 1967a: 392 (Philonthus; characters; Transcaucasian).
— Coiffait, 1974: 277 (Philonthus; characters; Caucasus; Transcaucasus).
— Schillhammer, 2000: 159 (Philonthus; species incertae sedis).
▴ Distribution: Caucasus.
fulcinius Smetana, 1995e: 85 (Philonthus; Type locality: Washington: Olympic Nat. Pk., Hoh Ranger Stn. 600 ft. Also cited from British Columbia).
fulgipennis Sharp, 1885: 410 (Philonthus; Type locality: Panama: Chiriqui; Volcan de Chiriqui below 4000 feet; Boquete 3500 feet).
fulvipes Fabricius, 1793, see: rubripennis Stephens, 1832.
fulvipes Stephens, 1833, see: ventralis Gravenhorst, 1802.
fulvitarsis Motschulsky, 1860a: 80 (Philonthus; Type locality: montagnes Noura-Ellia à Ceylan).
fumarius Gravenhorst, 1806: 67 (Staphylinus; Type locality: Brunsvigae).
— Nordmann, 1837: 100 (Philonthus; characters; Germany).
— Erichson, 1839a: 472 (Philonthus; characters; Germany).
— Heer, 1839: 270 (Philonthus; characters; Switzerland).
— Erichson, 1840: 482 (Philonthus; characters; Germany).
— Kiesenwetter, 1848a: 20 (Philonthus; characters; Germany).
— Jacobsen, 1849: 702 (Philonthus; characters; Denmark).
— Redtenbacher, 1849: 706 (Philonthus; characters; Austria).
— Fairmaire and Laboulbène, 1856: 527 (Philonthus; characters; France).
— Kraatz, 1857d: 610 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 196 (Philonthus; characters; Austria).
— Thomson, 1867a: 42 (Philonthus; characters; Scandinavia).
— Fauvel, 1874: 257 [= 1874a: 483] (Philonthus; characters; France; Austria).
— Redtenbacher, 1874: 214 (Philonthus; characters; Austria).
— Mulsant and Rey, 1876: 498 [= 1877: 354] (Philonthus; characters; France).
— Fowler, 1888: 277 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 456 (Philonthus; characters; north and middle Europe).
— Reitter, 1909: 130 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 339 (Philonthus; catalog).
— Johansen, 1914: 411 (Philonthus; characters; Denmark).
— Portevin, 1929: 364 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1343 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— V. Hansen, 1952: 75 (Philonthus; subgenus Philonthus; characters; Denmark).
— Jarrige, 1951a: 7 (Philonthus; characters).
— Smetana, 1958: 218 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 168 (Philonthus; characters).
— Palm, 1963b: 92 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 185 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 171 (Philonthus; Ireland Britain; Denmark; Norway; Sweden; Finland; Holland; Belgium; France; Switzerland; Italy; Hercegovina; Serbia; Turkey; Romania; Hungary; Czechoslovakia; Poland; Ukraine; Germany; Austria).
— Coiffait, 1967a: 429 (Philonthus; characters; Europe).
— Raianu, 1968a: 468 (Philonthus; Romania).
— Coiffait, 1974: 271 (Philonthus; characters; Europe; Britain; Scandinavia).
— Pope, 1977: 29 (Philonthus; Britain).
— Hinton, 1981: 660 (Philonthus; characters of egg).
— Frank, 1982: 41 (Philonthus; list of parasites).
— Tóth, 1984: 75 (Philonthus; characters; Hungary).
— Lucht, 1987: 104 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Pilon and Zanetti, 1991: 64 (Philonthus; Italy).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Owen, 1997a: 149 (Philonthus; collecting sites; Ireland).
— Kashcheev, 1999a: 58 (Philonthus; Kazakhstan).
▴ Distribution: Europe, Kazakhstan.
fumigatus Erichson, 1839, see: rufipes Stephens, 1832.
fumosus Solsky, 1868, see: longicornis Stephens, 1832.
funebris Last, 1981 [Note: Nomen nudum], see: specialis Cameron, 1937.
funebris Last, 1987 [Note: Nomen nudum], see: specialis Cameron, 1937.
funeralis Cameron, 1943: 130 (Philonthus; Type locality: Darjeeling: Ghum district: Tiger Hill, altitude 8500–10,000 feet).
furcifer Renkonen, 1937: 73 (Philonthus; Type locality: Finnland: Lohja; Tvärminne; Kauniainen; Tuusula; Pudasjärvi; Kuolajärvi; Muonio; Lapponia).
— Wüsthoff, 1937a: 347 (Philonthus; illustration of aedeagus).
— Tottenham, 1939a: 201 (Philonthus; characters; Britain).
— V. Hansen, 1952: 77 (Philonthus; subgenus Philonthus; characters; Denmark).
— Lohse, 1956: 90 (Philonthus; characters; notes).
— Smetana, 1958: 219 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 168 (Philonthus; characters).
— Coiffait, 1963: 12 (Paragabrius).
— Palm, 1963b: 94 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 185 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 168 (Philonthus; Russia; Finland; Sweden; Norway; Britain; Denmark; France; Germany; Austria; Switzerland; Romania).
— Coiffait, 1974: 93 (Paragabrius; characters; Finland; Scandinavia; Britain; Ireland; France).
— Pope, 1977: 29 (Philonthus; Britain).
— Tóth, 1984: 76 (Philonthus; characters).
— Lucht, 1987: 106 (Philonthus; checklist; Germany; Poland; Austria; Denmark; Sweden).
— Smetana, 1993b: 48 (Philonthus; subgenus Paragabrius; Czech Republic ?).
— Hodge and Jones, 1995: 39 (Philonthus; characters; Britain).
— Owen, 1996: 64 (Philonthus; characters; Ireland).
— Owen, 1997a: 149 (Philonthus; collecting sites; Ireland).
▴ Distribution: Europe.
furtivus Cameron, 1932, see: Gabrius.
furvus Nordmann, 1837: 88 (Philonthus; Type locality: Mexico).
— Erichson, 1840: 434 (Philonthus; characters; Mexico).
— Solsky, 1868a: 127 (Philonthus; characters; Mexico).
— Horn, 1884: 181 (Philonthus; characters; Newfoundland to Vancouver south to Arizona and Mexico; California).
— Sharp, 1885: 398 (Philonthus; characters; notes; Mexico; Guatemala; Costa Rica).
— Fall and Cockerell, 1907: 166 (Philonthus; New Mexico).
— Bernhauer and Schubert, 1914: 339 (Philonthus; catalog).
— Hatch, 1957: 195 (Philonthus; characters; British Columbia; Washington; Oregon; Idaho).
— Smetana, 1965: 13 (Philonthus; characters; Miquelon; Labrador; Nova Scotia; Newfoundland).
— Smetana, 1965a: 3 (Philonthus; British Columbia).
— Frank, 1982: 41 (Philonthus; list of parasites).
— Smetana, 1991a: 229 (Philonthus; characters; Mexico).
— Smetana, 1995e: 90 (Philonthus; characters; Alberta; British Columbia; Manitoba; New Brunswick; Newfoundland; Northwest Territories; Nova Scotia; Ontario; Quebec; Saskatchewan; Yukon Territory; Arizona; California; Colorado; Maine; Montana; Nebraska; Nevada; New Mexico; New York; Ohio; Oregon; Utah; Washington; Wisconsin; Wyoming; Mexico).
— Downie and Arnett, 1996: 377 (Philonthus; characters; USA).
▴ Distribution: Canada, USA, Mexico.
californicus Mannerheim, 1843b: 230 (Philonthus; Type locality: California).
— Horn, 1884: 182 (Philonthus; variety of furvus).
— Bernhauer and Schubert, 1914: 339 (Philonthus; synonym of furvus).
— Hatch, 1957: 195 (Philonthus; aberration of furvus; characters).
— Smetana, 1995e: 90 (Philonthus; synonym of furvus).
comptus Haldeman, 1852: 375 (Philonthus; Type locality: Not cited).
— Bernhauer and Schubert, 1914: 339 (Philonthus; synonym of furvus).
— Smetana, 1991a: 230 (Philonthus; synonym of furvus).
— Smetana, 1995e: 90 (Philonthus; synonym of furvus).
fuscatus Kraatz, 1859: 95 (Philonthus; Type locality: Ceylan).
— Bernhauer and Schubert, 1914: 339 (Philonthus; catalog).
— Cameron, 1932: 141 (Philonthus; characters; Ceylon).
— Scheerpeltz, 1933: 1344 (Philonthus; catalog).
— Coiffait, 1967a: 429 (Philonthus; characters; Mesopotamia; India; China).
— Coiffait, 1974: 98 (Paragabrius; subgenus Metagabrius; characters; Ceylon; India; China; Mesopotamia).
▴ Distribution: Iraq, India, Sri Lanka, China.
fuscicornis Stephens, 1833, see: ventralis Gravenhorst, 1802.
fuscicornis Nordmann, 1837, see: longicornis Stephens, 1832.
fuscicoxis Scheerpeltz, 1933 [Note: Nomen nudum], see: varians Paykull, 1789.
fuscipennis Mannerheim, 1830, see: cognatus Stephens, 1832.
fuscus Last, 1989: 880 (Gabrius; Type locality: Papua New Guinea: Kainantu, Onerunka).
fusiformis Melsheimer, 1844: 38 (Philonthus; Type locality: Pennsylvania; [Note: See lectotype designation by Smetana, 1995e]).
— Horn, 1884: 200 (Philonthus; characters; middle States and Canada, west to Colorado).
— Blatchley, 1910: 385 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 340 (Philonthus; catalog).
— Notman, 1920: 16 (Philonthus; New York).
— Smetana, 1995e: 376 (Philonthus; lectotype designation: Pennsylvania; characters; notes; New Brunswick; Ontario; Quebec; Arkansas; Connecticut; Illinois; Indiana; Iowa; Kansas; Massachusetts; Michigan; New Hampshire; New Jersey; New York; Ohio; Oklahoma; Pennsylvania; Rhode Island; South Dakota; Texas; Wisconsin).
— Downie and Arnett, 1996: 379 (Philonthus; characters; USA).
▴ Distribution: Canada, USA.
gabonensis Levasseur, 1966a: 1211 (Philonthus; Type locality: Belinga, Mission biologique au Gabon. Also cited from Congo).
— Levasseur, 1968a: 53 (Philonthus; Central African Republic).
▴ Distribution: Gabon, Congo, Central African Republic.
gabrioides Bernhauer, 1913, see: Gabrius.
gabrioides Coiffait, 1984, see: bhutanensis Herman, 2001.
gaerdesi Scheerpeltz, 1966: 65 (Philonthus; subgenus Philonthus; Type locality: Südwest-Afrika: Okahandja in der Umgebung von Okahandja).
gagates Mulsant and Rey, 1876: 454 [= 1877: 310] (Philonthus; Type locality: Provence et dans le Languedoc).
— Sainte-Claire Deville, 1906: 97 (Philonthus; Corsica).
— Bernhauer and Schubert, 1914: 340 (Philonthus; catalog).
— Scheerpeltz, 1933: 1344 (Philonthus; catalog).
— Coiffait, 1967a: 413 (Philonthus; characters; Europe; France; Corsica; Italy).
— Coiffait, 1974: 264 (Philonthus; characters; Mediterranean region).
— Pace and Zanetti, 1977: 292 (Philonthus; Italy).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
▴ Distribution: France, Italy.
apenninus Fiori, 1894: 92 (Philonthus; Type locality: Toano, Cusna e Riarbero nell'appennino Reggiano; Sila de di Serra S. Bruno nell'appennino Calabrese; Adriano sull'Argentaro in Toscana).
— Bernhauer and Schubert, 1914: 340 (Philonthus; synonym of gagates).
— Fiori, 1915: 15 (Philonthus; Italy).
— Scheerpeltz, 1933: 1344 (Philonthus; synonym of gagates).
— Coiffait, 1974: 264 (Philonthus; synonym of gagates).
ganeshensis Coiffait, 1983d: 164 (Philonthus; Type locality: Nepal: Ganesh Himal, entre Salmi et Rupchet, 1500 à 3600 m).
ganglbaueri Bernhauer, 1898, see: juvenilis Peyeron, 1858.
gardneri Cameron, 1932: 101 (Philonthus; Type locality: Bengal: Darjeeling, Rangirum, alt. 6000 feet).
gastoni Tikhomirova, 1973, see: kiautschauensis Bernhauer, 1916.
gastralis Sharp, 1874: 48 (Philonthus; Type locality: Nagasaki).
— Bernhauer and Schubert, 1914: 340 (Philonthus; catalog).
— Scheerpeltz, 1933: 1344 (Philonthus; catalog).
— Nakane, 1963a: 91 (Philonthus; characters; habitus photograph; Japan).
— Watanabe and Shibata, 1965: 321 (Philonthus; subgenus Bisnius; Japan).
— Frank, 1982: 41 (Philonthus; list of parasites).
— Watanabe, 1995d: 256 (Philonthus; Okushiri-tô Island).
▴ Distribution: Japan.
gaudens Tottenham, 1939d: 234 (Philonthus; replacement name for nitescens).
— Smetana, 1995e: 87 (Philonthus; characters; British Columbia; California; Oregon; Washington).
▴ Distribution: Canada, USA.
nitescens Horn, 1884: 182 (Philonthus; [preoccupied]; Type locality: southern California; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from Vancouver, Wash. Territory and the south of California).
— Bernhauer and Schubert, 1914: 348 (Philonthus; catalog).
— Hatch, 1957: 196 (Philonthus; characters; British Columbia; Washington; Oregon).
— Smetana, 1995e: 87 (Philonthus; lectotype designation: So. Cal [southern California]; synonym of gaudens).
gaudichaudi Blanchard, 1842: 81 (Philonthus; Type locality: Rio-Janeiro).
gavius Smetana, 1995e: 291 (Philonthus; Type locality: Louisiana: Concordia Pa.: 5 mi. W. Ferriday. Also cited from Arkansas, Florida, Illinois, Louisiana, Okalahoma, Tennessee, Texas).
gavoyi Bernhauer, 1928, see: Gabrius.
gawensis Last, 1981: 117 (Philonthus; Type locality: Gawe, near Morawaka, Eastern Highlands).
gedyeanus Bernhauer, 1940: 137 (Philonthus; Type locality: Kenya: Mt. Kenya, 8000 ft.; West Aberdare Mts., 10–11,000 ft.).
gedyei Cameron, 1951, see: Gabrius.
gemelloides Coiffait, 1983e: 479 (Philonthus; Type locality: Bashgultal, 1150 m, Nuristan, Afghanistan).
gemellus Kraatz, 1859: 91 (Philonthus; Type locality: Ceylan).
— Fauvel, 1904a: 58 (Philonthus; Belgaum; Nilghiris; Ceylan; Java).
— Fauvel, 1905e: 83 (Philonthus; Java).
— Bernhauer and Schubert, 1914: 340 (Philonthus; catalog).
— Cameron, 1921a: 377, 405 (Philonthus; characters; Singapore).
— Cameron, 1932: 124 (Philonthus; characters; Ceylon; India).
— Scheerpeltz, 1933: 1344 (Philonthus; catalog).
— Cameron, 1937: 5 (Philonthus; Java).
— Frank, 1982: 41 (Philonthus; list of parasites).
— Lecoq, 1990: 199 (Philonthus; Mascarene Islands).
— Shibata, 1992: 188 (Philonthus; Ryukyu Archipelago).
— Yadav, Chand, and Yadav, 1995: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
▴ Distribution: Mascarene Islands, India, Sri Lanka, Singapore, Indonesia, Japan.
geminus Kraatz, 1859: 87 (Philonthus; Type locality: Ceylan).
— Fauvel, 1903: 159 (Philonthus; Ceylan; India; China; Burma; Japan; Tonkin; Pulo-Penang; Sumatra; Mentawei Islands; Celebes).
— Fauvel, 1904a: 58 (Philonthus; Nilghiris; China; Japan; Bírmanie; Tonkin; Pulo-Penang; Sumatra; Java; Mentawei Island; Celebes).
— Bernhauer and Schubert, 1914: 340 (Philonthus; catalog).
— Cameron, 1921a: 378, 405 (Philonthus; characters; Singapore).
— Cameron, 1932: 103 (Philonthus; characters; Ceylon; India).
— Scheerpeltz, 1933: 1344 (Philonthus; catalog).
— Cameron, 1937: 5 (Philonthus; Java).
— Yadav, Chand, and Yadav, 1995: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
▴ Distribution: Sri Lanka, India, Myanmar, Malaysia, Vietnam, Singapore, Indonesia.
gentilicius Cameron, 1926b: 353 (Philonthus; Type locality: Chakrata District; Manjgaon, Bangar, Mohna. Mussorie District; Arni Gad. Dehra Dun; Siwaliks, 2500 to 6000 feet).
— Cameron, 1932: 120 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1344 (Philonthus; catalog).
▴ Distribution: India.
gentilis Horn, 1884: 185 (Philonthus; Type locality: Arizona; [Note: See lectotype designation by Smetana, 1995e]).
— Bernhauer and Schubert, 1914: 340 (Philonthus; catalog).
— Smetana, 1995e: 160 (Philonthus; lectotype designation: Ariz. [Arizona]; characters; Arizona; Texas; Mexico).
▴ Distribution: Mexico; USA.
georgianus Sachse, 1852, see: lomatus Erichson, 1840.
gerardi Bernhauer, 1928g: 108 (Philonthus; Type locality: Belgisch-Kongo: Manyema, Sibatwa Kilengwe).
— Scheerpeltz, 1933: 1344 (Philonthus; catalog).
— Tottenham, 1949b: 353 (Philonthus; characters).
▴ Distribution: Congo.
gerhardti Jakobson, 1909, see: micans Gravenhorst, 1802.
gerhardtianus Scheerpeltz, 1933, see: discoideus Gravenhorst, 1802.
germaini Scheerpeltz, 1933, see: Loncovilius.
germanus Sharp, 1874, see: Bisnius.
gestroi Fauvel, 1878f: 263 (Philonthus; Type locality: Nouvelle-Guinée, Andai).
— Fauvel, 1879: 102 (Philonthus; Dorei; Amberbaki).
— Bernhauer and Schubert, 1914: 340 (Philonthus; catalog).
▴ Distribution: New Guinea.
geus Tottenham, 1953: 146 (Philonthus; Type locality: Rarotonga).
— Barrion, Litsinger, and Last, 1991: 1078 (Philonthus; cited subspecies of variipennis; associated with rice agroecosystem).
▴ Distribution: Cook Islands.
ghilianii Kraatz, 1857, see: laetus Heer, 1839.
ghilarovi Tikhomirova, 1973: 167 (Philonthus; Type locality: Suputinskii zapovednik).
gigas Bernhauer, 1915d: 136 (Philonthus; Type locality: Abessinien: Boroda).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Tottenham, 1962: 160 (Philonthus; characters).
▴ Distribution: Ethiopia.
gilleti Levasseur, 1980: 361 (Philonthus; Type locality: Tchad: Parc Zakouma, Mas Dhour).
gilvipes Erichson, 1839, see: lepidus Gravenhorst, 1802.
gilvisuturatus Coiffait, 1983e: 481 (Philonthus; Type locality: Bashgultal, 1150 m, Nuristan, Afghanistan).
girardi Lecoq, 1989: 92 (Philonthus; Type locality: Mont Nimba, République Populaire de Guinée).
glaberrimus Scheerpeltz, 1965: 224 (Philonthus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).
glenelgi Blackburn, 1903: 94 (Philonthus; Type locality: Victoria, Glenelg R. District).
— Lea, 1925: 220 (Philonthus; characters; Australia).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
▴ Distribution: Australia.
glenoides Schubert, 1908, see: Craspedomerus.
godmani Sharp, 1885: 413 (Philonthus; Type locality: Mexico, Orizaba; Guatemala, Calderas 1200 feet, Panajachel 5000 feet, San Geronimo 3000 feet; Panama, Boquete 3500 feet).
— Bernhauer and Schubert, 1914: 340 (Philonthus; catalog).
▴ Distribution: Mexico, Guatemala, Panama.
gongulus Tottenham, 1949b: 356 (Philonthus; Type locality: Kivu: N'Gama).
gopheri Hubbard, 1894a: 308 (Philonthus; Type locality: Crescent City, Florida).
— Hubbard, 1896: 300, 301 (Philonthus; notes).
— Notman, 1920b: 706 (Philonthus; Florida).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Young and Goff, 1939: 59 (Philonthus; in Gopher Tortoise burrows; notes; Florida).
— Woodruff, 1982: 27 (Philonthus; in Gopher Tortoise burrows; notes; Florida).
— Smetana, 1995e: 260 (Philonthus; characters; Florida).
▴ Distribution: USA.
gorokaensis Last, 1981: 123 (Philonthus; Type locality: Goroka).
gracilentus Cameron, 1925a: 186 (Philonthus; Type locality: Tjibodas, 1400 m).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Cameron, 1937: 5 (Philonthus; Java).
▴ Distribution: Indonesia.
gracilicornis Heer, 1839, see: Quedius.
gracilicornis Horn, 1884, see: incertus Solsky, 1868.
gracilior Casey, 1915: 444 (Philonthus; Type locality: California: Anderson Valley, Mendocino Co.; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Smetana, 1995e: 310 (Philonthus; lectotype designation: Anderson Valley, Mendocino Co., Califormia; characters; British Columbia; New Brunswick; Nova Scotia; Ontario; Quebec; Arkansas; California; Idaho; Illinois; Indiana; Iowa; Kansas; Maryland; Maine; Michigan; Missouri; Montana; New Hampshire; New Jersey; New York; Ohio; Oregon; Pennsylvania; Virginia; Washington; West Virginia).
▴ Distribution: Canada, USA.
viridanus Horn, 1884: 221 (Philonthus; [preoccupied]; Type locality: New York; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from Middle States westward to Missouri).
— Bernhauer and Schubert, 1914: 361 (Philonthus; catalog).
— Notman, 1920: 16 (Philonthus; New York).
— Tottenham, 1939d: 233 (Philonthus; synonym of chlorinus).
— Hatch, 1957: 182 (Philonthus; characters; British Columbia; Washington; Idaho).
— Smetana, 1965: 28 (Philonthus; synonym of chlorinus).
— Dearborn and Donahue, 1994: 15 (Philonthus; Maine).
— Smetana, 1995e: 310 (Philonthus; lectotype designation: N.Y. [New York]; synonym of gracilior).
strigicollis Notman, 1919: 136 (Philonthus; Type locality: Mooers, Clinton Co., New York; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1362 (Philonthus; catalog).
— Smetana, 1995e: 310 (Philonthus; lectotype designation: Mooers, Clinton Co., New York; synonym of gracilior).
— Downie and Arnett, 1996: 381 (Philonthus; characters; USA).
chlorinus Tottenham, 1939d: 233 (Philonthus; replacement name for viridanus Horn).
— Smetana, 1965: 28 (Philonthus; subgenus Philonthus; characters; Nova Scotia).
— Smetana, 1995e: 310 (Philonthus; synonym of gracilior).
— Downie and Arnett, 1996: 382 (Philonthus; characters; USA).
gracilis Letzner, 1846, see: Rabigus.
gracillimus Sharp, 1876: 168 (Philonthus; Type locality: Ega).
grandiceps Kraatz, 1859, see: Belonuchus.
grandiceps Poppius, 1909, see: poppiusi Bernhauer and Schubert, 1914.
grandiceps Bernhauer, 1916, see: ridiculus Blackwelder, 1944.
grandicollis Horn, 1884: 193 (Philonthus; Type locality: California, and as far north as Vancouver).
— Bernhauer and Schubert, 1914: 340 (Philonthus; catalog).
— Hatch, 1957: 191 (Philonthus; characters; British Columbia; Washington; Oregon; Idaho).
— Smetana, 1995e: 300 (Philonthus; characters; notes; British Columbia; Arizona; California; Idaho; Montana; Nevada; Oregon; Utah; Washington; Wyoming; Mexico).
▴ Distribution: Canada, USA, Mexico.
grandis Bernhauer, 1932b: 147 (Pseudophilonthus; subgenus Nephronthus; Type locality: Haut-Uele: Abimva; Moto; Yebo Moto, Tora).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Tottenham, 1949b: 361 (Philonthus; characters).
▴ Distribution: Congo.
grassei Levasseur, 1980: 363 (Philonthus; Type locality: Gabon: Belinga).
gratellus Cameron, 1932, see: Gabrius.
gratiosus Erichson, 1840: 461 (Philonthus; Type locality: Bogotae in Columbia).
gratiosus Brisout, 1866, see: suavus Brisout, 1867.
gratus LeConte, 1863, see: Neobisnius.
gratus Cameron, 1943, see: Hesperus.
greipeli W. Kolbe, 1931 [Note: Nomen nudum], see: intermedius Lacordaire, 1835.
gridellii Cameron, 1926, see: cyanelytrius Kraatz, 1859.
griseolus Sharp, 1885: 399 (Philonthus; Type locality: Costa Rica, Irazu, 6000 to 7000 feet).
— Bernhauer and Schubert, 1914: 340 (Philonthus; catalog).
— Smetana, 1991a: 233 (Philonthus; characters; Costa Rica).
▴ Distribution: Costa Rica.
grossepunctatus Bernhauer, 1939, see: nudus Sharp, 1874.
guatemalenus Sharp, 1885: 403 (Philonthus; Type locality: Guatemala, San Geronimo).
guerreroensis Bernhauer, 1910b: 380 (Philonthus; Type locality: Mexiko: Omilteme, Guerrero, 8000 F.).
gulmargensis Coiffait, 1982: 64 (Philonthus; Type locality: Inde: Cachemire, Gulmarg, Pir Panjal Gebirge, 2800 m).
gyllenhali Gistel, 1857: 74 (Philonthus; [Note: Nomen dubium]; Type locality: Germania).
haematodes Bernhauer, 1915d: 138 (Philonthus; Type locality: Abessinien: Kunhe, Boroda).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Tottenham, 1949b: 308 (Philonthus; characters).
— Tottenham, 1962: 162 (Philonthus; characters).
▴ Distribution: Ethiopia.
haematurus Erichson, 1840, see: Hesperus.
haemorrhoidalis W. J. MacLeay, 1873, see: Hesperus.
haemorrhoidalis Brancsik, 1893, see: Diatrechus.
hanae Vávra and Schillhammer, 1993, see: Bisnius.
hanseni Palm, 1952 [Note: Nomen nudum], see: binotatus Gravenhorst, 1806.
hargreavesi Tottenham, 1949b: 354 (Philonthus; Type locality: Sierra Leone: Njala).
harrisii Melsheimer, 1844, see: politus Linné, 1758.
havaniensis Laporte, 1835: 116 (Staphylinus; Type locality: Ile de Cuba).
— Bernhauer and Schubert, 1914: 340 (Philonthus; catalog).
— Blackwelder, 1943: 418 (Philonthus; characters; Cuba; Jamaica; Haiti; Puerto Rico; St. John; St. Croix; Antigua).
— Ramos, 1946: 33 (Philonthus; Mona Island, West Indies).
— Blackwelder, 1947: 123 (Philonthus; Cayman Islands).
▴ Distribution: West Indies.
obscurus Fauvel, 1863: 434 (Philonthus; [preoccupied]; Type locality: Not cited, presumably Cuba).
— Fauvel, 1869: 489 (Philonthus; synonym of havaniensis).
— Fauvel, 1891: 115 (Philonthus; synonym of suspectus).
— Bernhauer and Schubert, 1914: 340 (Philonthus; synonym of havaniensis).
— Blackwelder, 1943: 418 (Philonthus; synonym of havaniensis).
havelkai R. Dvořák, 1958: 140 (Philonthus; Type locality: Japan: Osaka, Yodokawa).
hebridensis Fauvel, 1878g: 549 (Philonthus; Type locality: Nouvelle-Hébrides).
heidenreichi Hubenthal, 1911, see: Bisnius.
heilongjiangensis J. Li, 1993: 60 (Philonthus; cited as heilougjiangensis; Type locality: Mount Laoye).
heinemanni Bernhauer, 1913, see: caucasicus Nordmann, 1837.
heinlii Gistel, 1857: 23 (Philonthus; [Note: Nomen dubium]; Type locality: Bayern).
helveticus Mulsant and Rey, 1876, see: micans Gravenhorst, 1802.
helvolus Erichson, 1840: 494 (Philonthus; Type locality: Madagascar).
hepaticus Erichson, 1840: 451 (Philonthus; Type locality: America septentrionalis; [Note: See lectotype designation by Smetana, 1995e]).
— Fauvel, 1878g: 546 (Philonthus; characters; Australia).
— Lynch, 1884: 153 (Philonthus; characters; Argentina).
— Horn, 1884: 188 (Philonthus; characters; California; Nevada; Arizona).
— Fauvel, 1891: 112 (Philonthus; notes; Venezuela).
— Fauvel, 1901: 85 (Philonthus; Colombia; northern United States to Argentina; southern Australia).
— Fall and Cockerell, 1907: 166 (Philonthus; New Mexico).
— Blatchley, 1910: 381 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 341 (Philonthus; catalog).
— Notman, 1920b: 705 (Philonthus; Florida).
— Lea, 1925: 220 (Philonthus; Australian introduction).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Blackwelder, 1943: 401 (Philonthus; characters; Bahamas; Cuba; Jamaica; Haiti; Dominican Republic; Puerto Rico; St. Thomas; St. Croix; St. Kitts; Antigua; Montserrat; Guadeloupe; Les Saintes; Dominica; Barbados; St. Vincent; Grenadines; Tobago; Trinidad).
— Coiffait and Saiz, 1968: 355 (Philonthus; characters; Chile).
— Saiz, 1971b: 342 (Philonthus; Argentina; Chile).
— Frank, 1982: 41 (Philonthus; list of parasites).
— Hunter, Kincher, Bay, Beerwinkle, 1991: 168 (Philonthus; seasonal distribution and diel flight activity; Texas).
— Smetana, 1995e: 175 (Philonthus; lectotype designation: America septentrionalis; characters; British Columbia; Alberta; Arizona; Arkansas; California; Colorado; Delaware; Florida; Georgia; Illinois; Indiana; Iowa; Kansas Kentucky; Louisiana Maryland; Massachusetts; Michigan; Mississippi; Missouri; Montana; Nebraska; New Jersey; New Mexico; New York; North Carolina; Ohio; Oklahoma; Pennsylvania; South Carolina; Tennessee; Texas; Utah; Virginia; West Virginia; Wyoming; Mexico).
— Hu and Frank, 1995b: 586 (Philonthus; surface structure of eggs).
— Hu and Frank, 1997 (Philonthus; predation of eggs and larvae of Haematobia irritans; Florida).
— Downie and Arnett, 1996: 377 (Philonthus; characters; USA).
▴ Distribution: Canada, USA, West Indies, Mexico, Guatemala, Nicaragua, Panama, Colombia, Venezuela, Brazil, Argentina, Chile, Australia, New Zealand.
vilis Erichson, 1840: 451 (Philonthus; Type locality: Americae meridionalis ins. St. Thoma; [Note: See lectotype designation by Smetana, 1995e]).
— Guérin-Méneville, 1857: 39 (Philonthus; characters).
— Chevrolat and Fauvel, 1863: 433 (Philonthus; characters; St. Thomas, Cuba).
— Fleutiaux and Sallé, 1889: 379 (Philonthus; Guadeloupe Island).
— Sharp, 1885: 406 (Philonthus; notes; Mexico; Guatemala; Nicaragua; Antilles).
— Fauvel, 1891: 113 (Philonthus; Venezuela).
— Fauvel, 1901: 85 (Philonthus; Colombia; Central America; Antilles; Venezuela; Brazil).
— Bernhauer and Schubert, 1914: 360 (Philonthus; catalog).
— Blackwelder, 1943: 401 (Philonthus; synonym of hepaticus).
— Blackwelder, 1944: 133 (Philonthus; synonym of hepaticus).
— Scheerpeltz, 1971b: 205 (Philonthus; Guatemala).
— Smetana, 1995e: 175 (Philonthus; lectotype designation; synonym of hepaticus).
nanus Melsheimer, 1844: 36 (Philonthus; Type locality: Pennsylvania; [Note: See lectotype designation by Smetana, 1995e]).
— Gemminger and Harold, 1868: 589 (Philonthus; synonym of hepaticus).
— Lynch, 1884: 153 (Philonthus; synonym of hepaticus).
— Horn, 1884: 233 (Philonthus; synonymy with hepaticus may be erroneous).
— Fauvel, 1891: 112 (Philonthus; synonym of hepaticus).
— Bernhauer and Schubert, 1914: 341 (Philonthus; synonym of hepaticus).
— Scheerpeltz, 1933: 1345 (Philonthus; synonym of hepaticus).
— Blackwelder, 1943: 401 (Philonthus; synonym of hepaticus).
— Smetana, 1995e: 175 (Philonthus; lectotype designation: Pennsylvania; synonym of hepaticus).
cinctutus Melsheimer, 1844: 37 (Philonthus; [Note: This name is often cited as cinctulus; Melsheimer spelled it cinctutus when describing it (1844) and later when cataloging it (1853: 157)]; Type locality: Pennsylvania; [Note: See lectotype designation by Smetana, 1995e]).
— Horn, 1884: 223 (Philonthus; may be synonym of hepaticus).
— Fauvel, 1891: 112 (Philonthus; synonym of hepaticus).
— Bernhauer and Schubert, 1914: 341 (Philonthus; cited as cinctulus; synonym of hepaticus).
— Blackwelder, 1943: 401 (Philonthus; synonym of hepaticus).
— Smetana, 1995e: 175 (Philonthus; lectotype designation: Pennsylvania; synonym of hepaticus).
palleolus Melsheimer, 1844: 37 (Philonthus; variety of nanus; Type locality: Not cited).
— Gemminger and Harold, 1868: 589 (Philonthus; synonym of hepaticus).
— Lynch, 1884: 153 (Philonthus; synonym of hepaticus).
— Fauvel, 1891: 112 (Philonthus; synonym of hepaticus).
— Bernhauer and Schubert, 1914: 341 (Philonthus; synonym of hepaticus).
— Blackwelder, 1943: 401 (Philonthus; synonym of hepaticus).
— Smetana, 1995e: 175 (Philonthus; synonym of hepaticus).
rufipennis Solier, 1849: 317 (Staphylinus; [preoccupied]; Type locality: Chile: Santa Rosa; Illapel).
— Kraatz, 1859a: 5 (Philonthus; notes).
— Fairmaire and Germain, 1861: 434 (Philonthus; synonym of pyropterus).
— Fauvel, 1864a: 125 (Philonthus; synonym of pyropterus).
— Fauvel, 1866a: 343 [= 1866b: 94] (Philonthus; synonym of pyropterus).
— Fauvel, 1891: 112 (Philonthus; synonym of hepaticus).
— Bernhauer and Schubert, 1914: 341 (Philonthus; synonym of hepaticus).
— Blackwelder, 1943: 401 (Philonthus; synonym of hepaticus).
— Coiffait and Saiz, 1968: 355 (Philonthus; synonym of hepaticus).
— Saiz, 1969a: 9 (Philonthus; synonym of hepaticus).
varicolor Boheman, 1858: 29 (Philonthus; Type locality: California).
— Fauvel, 1891: 112 (Philonthus; synonym of hepaticus).
— Bernhauer and Schubert, 1914: 341 (Philonthus; synonym of hepaticus).
— Blackwelder, 1943: 401 (Philonthus; synonym of hepaticus).
— Smetana, 1995e: 175 (Philonthus; synonym of hepaticus).
pyropterus Kraatz, 1859a: 12, note (Philonthus; replacement name for rufipennis Solier).
— Fairmaire and Germain, 1861: 431 (Philonthus; characters; Chile).
— Fauvel, 1864a: 125 (Philonthus; notes; Chile).
— Fauvel, 1866a: 343 [= 1866b: 94] (Philonthus; characters; Chile).
— Fauvel, 1878g: 546 (Philonthus; synonym of hepaticus).
— Fauvel, 1891: 112 (Philonthus; synonym of hepaticus).
— Bernhauer and Schubert, 1914: 341 (Philonthus; synonym of hepaticus).
— Blackwelder, 1943: 401 (Philonthus; synonym of hepaticus).
— Coiffait and Saiz, 1968: 355 (Philonthus; synonym of hepaticus).
— Kuschel, 1990: 26 (Philonthus; cited as valid species; introduced, South American species; winged; habitat; New Zealand).
parvimanus Sharp, 1885: 406 (Philonthus; Type locality: Mexico: Cordova; Guatemala: near the City, Paso Antonio, Camperico, San Joaquin in Vera Paz; Nicaragua: Chontales; Panama: David).
— Fauvel, 1891: 112 (Philonthus; synonym of hepaticus).
— Bernhauer and Schubert, 1914: 341 (Philonthus; synonym of hepaticus).
— Blackwelder, 1943: 401 (Philonthus; synonym of hepaticus).
oklahomensis Hatch and Ortenburger, 1930: 12 (Philonthus; Type locality: Okla., Murray Co., Dougherty).
— Scheerpeltz, 1933: 1354 (Philonthus; catalog).
— Smetana, 1995e: 175 (Philonthus; synonym of hepaticus).
herilis Erichson, 1840, see: Gastrisus.
hesperiformis Cameron, 1932, see: Hesperopalpus.
hesperius Fauvel, 1878, see: Gabrius.
heterocerus Cameron, 1950, see: Gabrius.
heterodoxus Mulsant and Rey, 1876: 419 [= 1877: 275] (Philonthus; Type locality: Biguglia en Corse).
— Sainte-Claire Deville, 1906: 96 (Philonthus; Corsica).
— Bernhauer and Schubert, 1914: 341 (Philonthus; catalog).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Coiffait, 1967a: 420 (Philonthus; characters; Corsica).
— Coiffait, 1973: 112 (Philonthus; characters; Sicily; Italy).
— Coiffait, 1974: 274 (Philonthus; characters; Corsica; Sicily; Italy; Corfu).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
▴ Distribution: Corsica, Sicily, Corfu.
heterogaster Cameron, 1942c: 845 (Philonthus; Type locality: N. Celebes: Toli-Toli).
heteropus Sharp, 1887: 788 (Philonthus; Type locality: Mexico, near the city).
hilaris Fauvel, 1874, see: Paederomimus.
himalayicus Bernhauer, 1915, see: Gabrius.
hindostanus Schubert, 1911, see: Rabigus.
hirsutus Tottenham, 1949b: 312 (Philonthus; Type locality: Lulua: Kapanga).
hirtipennis Stephens, 1832, see: albipes Gravenhorst, 1802.
hirtiventris Sharp, 1885: 410 (Philonthus; Type locality: Mexico, Oaxaca, Mirador).
histrio Sharp, 1874, see: Cafius.
hoegei Sharp, 1885: 400 (Philonthus; Type locality: Mexico, Las Vigas; Milpas).
holomelas Eppelsheim, 1895, see: abyssinus Fauvel, 1881.
honestus Last, 1981, see: Hesperus.
hongkongensis Bernhauer, 1931c: 130 (Philonthus; Type locality: Hongkong).
hornabrooki Last, 1968, see: Hesperus.
hornaditanus Rambousek, 1925: 75 (Philonthus; Type locality: Jujuy: Hornadita 3400–3600 m).
horni† Scudder, 1900: 56 (Philonthus; fossil; Type locality: Florissant, Colorado).
horni Bernhauer and Schubert, 1914, see: Gabrius.
horridulus Last, 1968: 370 (Philonthus; Type locality: New Guinea: Karimu).
hosmanni Bernhauer, 1912d: 41 (Philonthus; Type locality: Argentinien: Prov. Buenos Aires).
hospes Erichson, 1843: 221 (Philonthus; Type locality: Angola).
— Boheman, 1848: 279 (Philonthus; characters; South Africa).
— Bernhauer and Schubert, 1914: 341 (Philonthus; catalog).
— Rambousek, 1922: 1009 (Philonthus; Ethiopia).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Tottenham, 1949b: 331 (Philonthus; characters).
— Tottenham, 1956: 226 (Philonthus; Ruanda).
— Tottenham, 1961: 204 (Philonthus; Senegal).
— Levasseur, 1966a: 1211 (Philonthus; Congo; Senegal; Gabon; Kenya; Zaire; Madagascar).
— Levasseur, 1968a: 53 (Philonthus; Central African Republic).
— Scheerpeltz, 1974c: 21 (Philonthus; Sudan; Somalia).
▴ Distribution: Senegal, Central African Republic, Sudan, Ethiopia, Somalia, Kenya, Tanzania, Rwanda, Congo, Gabon, Angola, South Africa, Madagascar.
inaequalis Eppelsheim, 1895: 200 (Philonthus; Type locality: Boran Galla; Madagascar).
— Bernhauer and Schubert, 1914: 341 (Philonthus; synonym of hospes).
— Gridelli, 1939: 316 (Philonthus; characters; Abyssinia).
— Scheerpeltz, 1974c: 21 (Philonthus; synonym of hospes).
necopinus Schubert, 1911: 23 (Philonthus; Type locality: Afr. or. Germ., Kilimandjaro).
— Bernhauer and Schubert, 1914: 347 (Philonthus; catalog).
— Scheerpeltz, 1933: 1345 (Philonthus; synonym of hospes).
— Scheerpeltz, 1974c: 21 (Philonthus; synonym of hospes).
hova Fauvel, 1905b: 180 (Philonthus; Type locality: Tananarive).
hudsonicus Horn, 1884: 201 (Philonthus; Type locality: Canada; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from Canada and the Hudson's Bay region).
— Bernhauer and Schubert, 1914: 341 (Philonthus; catalog).
— Smetana, 1965: 22 (Philonthus; subgenus Philonthus; characters; notes; Newfoundland).
— Smetana, 1995e: 411 (Philonthus; lectotype designation: Canada; characters; notes; Alberta; New Brunswick; Newfoundland; Northwest Territories; Nova Scotia; Ontario; Quebec).
— Downie and Arnett, 1996: 379 (Philonthus; characters; Canada).
▴ Distribution: Canada.
hulunbeierensis J. Li, 1993, see: spinipes Sharp, 1874.
humbloti Lecoq, 1998: 68 (Philonthus; subgenus Piezarthrus; Type locality: Iles Comores).
humeralis Fauvel, 1878f: 267 (Philonthus; Type locality: Nouvelle-Guinée, Andai; Iles Arou, Wokan).
— Fauvel, 1879: 105 (Philonthus; Dorei).
— Bernhauer and Schubert, 1914: 341 (Philonthus; catalog).
— Cameron, 1931: 363 (Philonthus; New Guinea).
— Last, 1968: 368 (Philonthus; New Guinea).
▴ Distribution: New Guinea, Indonesia.
humidulus Cameron, 1932, see: Gabrius.
humidus Last, 1968: 368 (Philonthus; Type locality: New Guinea: Kanu River).
humilis Erichson, 1840, see: Neobisnius.
humilis Cameron, 1932: 106 (Philonthus; [preoccupied]; Type locality: Tehri Garhwal State: Aglar River).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Herman, 2001: 47 (Philonthus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: India.
hybridus Erichson, 1840, see: Quedius.
hybridus Cameron, 1930b: 163 (Philonthus; [preoccupied]; Type locality: North Borneo: Bettotan).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Herman, 2001: 47 (Philonthus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Borneo.
hylaeus Smetana, 1995e: 399 (Philonthus; Type locality: Louisiana: Concordia Pa.: 5 mi W. Ferriday. Also cited from Alabama; Arkansas; Florida; Georgia; Illinois; Mississippi; Oklahoma; Texas).
hyperboreus J. Sahlberg, 1880, see: Bisnius.
ichor Tottenham, 1961: 204, 209 (Philonthus; [Note: This name is validated in the commentary on sanguineus]; Type locality: Senegal: Badi; Wassadou).
idiocerus Kraatz, 1859: 85 (Philonthus; Type locality: Ceylan).
— Bernhauer, 1902b: 26 (Philonthus; Ceylon).
— Fauvel, 1905e: 83 (Philonthus; Java; Celebes; Sumatra; Indochina; Burma; Annam; India; Ceylon).
— Bernhauer and Schubert, 1914: 341 (Philonthus; catalog).
— Cameron, 1932: 101 (Philonthus; characters; Ceylon; India; Burma).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Cameron, 1937: 5 (Philonthus; Java).
— Yadav, Chand, and Yadav, 1995: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
▴ Distribution: Sri Lanka, India, Myanmar, Vietnam, Indonesia.
idoneus Sharp, 1885: 411 (Philonthus; Type locality: Guatemala, El Tumbador 2500 feet).
igacus Tottenham, 1955: 164 (Philonthus; subgenus Spatulonthus; Type locality: Cape Colony: Uitenhage).
— Tottenham, 1956: 227 (Philonthus; subgenus Spatulonthus; Ruanda).
— Tottenham, 1962: 140 (Philonthus; subgenus Spatulonthus; Tanganyika).
▴ Distribution: South Africa, Tanzania, Rwanda.
ignipennis Tottenham, 1962: 162 (Philonthus; Type locality: Not cited, presumably Africa).
ignobilis Cameron, 1943, see: Gabrius.
ilus Tottenham, 1953, see: variipennis Kraatz, 1859.
imitator Fauvel, 1895, see: Gabrius.
imitator Fauvel, 1907, see: sequens Bernhauer and Schubert, 1914.
immaculatus Gusmann, 1924 [Note: Nomen nudum], see: cruentatus Gmelin, 1790.
immundus Gravenhorst, 1806, see: ventralis Gravenhorst, 1802.
immundus: Gyllenhal, 1810 [Note: Nomen nudum], see: rufipes Stephens, 1832.
immutatus Motschulsky, 1860, see: latiusculus Hochhuth, 1851.
imperfectus J. Sahlberg, 1903, see: concinnus Gravenhorst, 1802.
imperialis Bernhauer, 1916c: 28 (Philonthus; Type locality: West-Columbien: Umgebung von Cali, St. Antonio, 2000 m).
impressicollis Stephens, 1835, see: Bisnius.
impressifrons Broun, 1880, see: Bisnius.
impuncticollis Bernhauer, 1932b: 153 (Philonthus; Type locality: Haut-Uele: Abimva).
— Scheerpeltz, 1933: 1346 (Philonthus; catalog).
— Tottenham, 1949b: 296 (Philonthus; characters).
▴ Distribution: Congo.
inaequalis Eppelsheim, 1895, see: hospes Erichson, 1843.
inachus Smetana, 1995e: 103 (Philonthus; Type locality: Arizona: Huachuca Mts., Ramsey Cn., 6300 ft. Also cited from California, Colorado, Nebraska, New Mexico, Texas, Utah, Wyoming).
incertus Solsky, 1868a: 135 (Philonthus; Type locality: Mexique).
— Sharp, 1885: 416 (Philonthus; notes; Mexico).
— Fall and Cockerell, 1907: 166 (Philonthus; New Mexico).
— Bernhauer and Schubert, 1914: 342 (Philonthus; catalog).
— Frank, 1982: 41 (Philonthus; list of parasites).
— Smetana, 1995e: 221 (Philonthus; characters; Arizona; Colorado; New Mexico; Utah).
▴ Distribution: Mexico, USA.
gracilicornis Horn, 1884: 190 (Philonthus; [preoccupied]; Type locality: Arizona; [Note: See lectotype designation by Smetana, 1995e]).
— Sharp, 1887: 788 (Philonthus; synonym of incertus).
— Bernhauer and Schubert, 1914: 342 (Philonthus; synonym of incertus).
— Smetana, 1995e: 221 (Philonthus; lectotype designation: Ariz. [Arizona]; synonym of incertus).
incisiventris Lea, 1925: 223 (Philonthus; Type locality: Queensland: Cairns; Mulgrave River).
incisus Schillhammer, 2000: 128 (Philonthus; Type locality: Kamtschatka, Sevo-See, Milkovo).
inclitus Fauvel, 1878f: 264 (Philonthus; Type locality: Iles Arou, Wokan).
incognitus Bernhauer, 1931d: 585 (Philonthus; Type locality: Abyssinia: Mt. Chillalo, 8,000 ft., 12,000–13,000 ft.; Wachacha Ravine, near Addis Ababa, ca. 8,000 ft.; between Jem-Jem and Wouramboulchi, nearly 9,000 ft.).
— Scheerpeltz, 1933: 1346 (Philonthus; catalog).
— Bernhauer and Chapman, 1939: 69 (Philonthus; Kenya).
— Cameron, 1950g: 35 (Philonthus; Belgian Congo).
— Tottenham, 1955: 162 (Philonthus; subgenus Spatulonthus; notes).
— Tottenham, 1962: 228 (Philonthus; characters).
▴ Distribution: Ethiopia, Kenya, Congo.
kristenseni Bernhauer, 1931d: 585 (Philonthus; subspecies of longicornis; [preoccupied]; Type locality: Abyssinia, Dire Dawa).
kristensenianus Scheerpeltz, 1933: 1346 (Philonthus; subspecies of incognitus; replacement name for kristenseni Bernhauer).
incompletus Hochhuth, 1849, see: ebeninus Gravenhorst, 1802.
incompletus Roubal, 1911, see: formaneki, Roubal, 1911.
inconspicuus Lea, 1929: 204 (Philonthus; Type locality: Queensland: Coen River; Mulgrave River).
inconstans Sharp, 1889, see: Rabigus.
incultus Cameron, 1932: 98 (Philonthus; Type locality: Chakrata district: Bodyar, alt. 8300 feet).
indicus Cameron, 1920a: 216 (Philonthus; Type locality: Nilgiri Hills).
— Cameron, 1932: 111 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1346 (Philonthus; catalog).
▴ Distribution: India.
indigaceus Fauvel, 1891: 110 (Philonthus; Type locality: Caracas; Nouvelle Grenade).
indubius Luze, 1904: 95 (Philonthus; Type locality: Seravschan: Postigau; See Margusan).
— Bernhauer and Schubert, 1914: 342 (Philonthus; catalog).
— Scheerpeltz, 1933: 1346 (Philonthus; catalog; Transcaspian region).
— Coiffait, 1967a: 410 (Philonthus; characters; Turkestan).
— Coiffait, 1974: 287 (Philonthus; characters; Turkestan; Transcaspian region).
— Boháč, 1988: 551 [= 1989: 34] (Philonthus; characters; synonym of umbratilis).
— Schillhammer, 1992: 66 (Philonthus; distinct from umbratilis).
▴ Distribution: Tajikistan, Uzbekistan, Afghanistan.
eppelsheimi Coiffait, 1967a: 445 (Philonthus; subgenus Philonthus; Type locality: Taschkent, S.S.R. d'Ouzbekistan).
— Coiffait, 1974: 273 (Philonthus; characters; Turkestan; Transcaspian region).
— Coiffait, 1983e: 478 (Philonthus; Afghanistan).
— Schillhammer, 1992: 66 (Philonthus; synonym of indubius).
industanus Fauvel, 1903: 158 (Philonthus; Type locality: Nilghiris; [Note: See lectotype designation by Schillhammer, 1999]. Originally cited from Coonoor; also: Bengale occidental, Kurseong and Sikkim: Darjeeling).
— Bernhauer and Schubert, 1914: 342 (Philonthus; catalog).
— Cameron, 1932: 78 (Philonthus; characters; India).
— Cameron, 1932: 81 (Philonthus; [Note: Misidentification: Cameron's use of carbonarius was a misidentification according to Schillhammer, 1999: 59]; characters; India).
— Scheerpeltz, 1933: 1346 (Philonthus; catalog).
— Yadav, Chand, and Yadav, 1995: 6 (Philonthus; cited as industaneus; morphology of mitotic and meiotic chromosomes).
— Schillhammer, 1999: 59 (Philonthus; lectotype designation: Nilghiris; characters; India; Nepal Sri Lanka; Vietnam; China; Malaysia; Indonesia).
▴ Distribution: China, India, Nepal, Sri Lanka, Vietnam, Malaysia, Indonesia.
bipunctatus Bernhauer, 1911: 90 (Philonthus; Type locality: Little Nepal valley: Chitlong).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Cameron, 1932: 74 (Philonthus; characters; Nepal; India).
— Scheerpeltz, 1933: 1334 (Philonthus; catalog).
— Paulian, 1941: 229 (Philonthus; larval characters).
— Yadav, Chand, and Yadav, 1995: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
— Schillhammer, 1999: 59 (Philonthus; synonym of industanus).
pubipennis Cameron, 1919a: 253 (Philonthus; Type locality: Ceylon: Kitulgalle, alt. 1700 feet).
— Cameron, 1932: 75 (Philonthus; characters; Ceylon).
— Scheerpeltz, 1933: 1356 (Philonthus; catalog).
— Schillhammer, 1999: 59 (Philonthus; synonym of industanus).
infantilis Cameron, 1947: 2 (Philonthus; subgenus Gabrius; Type locality: Nizi: Blukwa).
infimus Sharp, 1885: 414 (Philonthus; Type locality: Guatemala, Las Mercedes 3000 feet, Chiacam; Panama, Volcan de Chiriqui).
infirmus Erichson, 1840: 472 (Philonthus; Type locality: Arabia felice).
— Bernhauer and Schubert, 1914: 342 (Philonthus; catalog).
— Scheerpeltz, 1933: 1346 (Philonthus; catalog; Mesopotamia).
— Jarrige, 1952a: 133 (Philonthus; Mauritania; Egypt).
— Cameron, 1952d: 824 (Philonthus; Mauritania).
— Scheerpeltz, 1963b: 428 (Philonthus; Egypt).
— Coiffait, 1966b: 338 (Philonthus; Mauritania; Hoggar).
— Coiffait, 1967a: 413 (Philonthus; characters; Mesopotamia; Arabia; Sahara; Mauritania).
— Coiffait, 1968a: 136 (Philonthus; notes; Chad).
— Coiffait, 1974: 243 (Philonthus; characters; Mauritania; Sahara; Arabia; Mesopotamia).
— Gusarov, 1997: 287 (Philonthus; Oman).
▴ Distribution: Mauritania, Algeria, Chad, Egypt, Djibouti, Oman, Saudi Arabia, Iraq.
flavicornis Fairmaire, 1892: 89 (Philonthus; Type locality: Obock).
— Bernhauer and Schubert, 1914: 338 (Philonthus; catalog).
— Peyerimhoff, 1931: 31 (Philonthus; Algeria).
— Scheerpeltz, 1933: 1342 (Philonthus; catalog).
— Jarrige, 1952a: 133 (Philonthus; synonym of infirmus).
— Coiffait, 1974: 243 (Philonthus; synonym of infirmus).
ingratulus Cameron, 1932: 133 (Philonthus; Type locality: Mussoorie district: Arni Gad).
ingratus Fauvel, 1905, see: Gabrius.
inipendezius Tottenham, 1962: 190 (Philonthus; Type locality: Tanganyika: E. Usambara, Amani, 3300 ft.).
innocuus Horn, 1884: 199 (Philonthus; Type locality: Arizona; [Note: See lectotype designation by Smetana, 1995e]).
— Fall and Cockerell, 1907: 166 (Philonthus; New Mexico).
— Bernhauer and Schubert, 1914: 342 (Philonthus; catalog).
— Smetana, 1995e: 438 (Philonthus; lectotype designation: Ariz. [Arizona]; characters; notes; Arizona; Colorado; New Mexico; Texas; Utah).
▴ Distribution: USA.
inopinatus Smetana, 1963, see: rotundicollis Ménétriés, 1832.
inornatus Cameron, 1920, see: tardus Kraatz, 1859.
inquietus Erichson, 1840, see: Bisnius.
inquinatus Stephens, 1832, see: quisquiliarius Gyllenhal, 1810.
insignis Luze, 1904, see: Gabrius.
insignitus Fauvel, 1875a: XXIX [= 1875b: 231] (Philonthus; Type locality: Sibérie, région Baïkal).
— Bernhauer and Schubert, 1914: 342 (Philonthus; catalog).
— Scheerpeltz, 1933: 1346 (Philonthus; catalog).
— Coiffait, 1967a: 432 (Philonthus; characters; Siberia).
— Coiffait, 1974: 297 (Philonthus; characters; central Siberia; East Asia).
— Shilov, 1976: 153 (Philonthus; Mongolia).
▴ Distribution: Russia, Mongolia.
instabilis Horn, 1884, see: Bisnius.
instigma Coiffait, 1970 [Note: Nomen nudum], see: nitidicollis Lacordaire, 1835.
insularis Bernhauer and Schubert, 1916: 425 (Quedius; replacement name for puncticollis Broun; [Note: Preoccupied secondary homonym in Philonthus]).
— Newton, 1996: 160 (Philonthus; [Note: Cited with only Bernhauer as author of species]).
▴ Distribution: New Zealand.
puncticollis Broun, 1894: 422 (Quedius; [preoccupied]; Type locality: West Plains, Invercargill).
insularis Gautier, 1868, see: Gabrius.
intaminatus Stephens, 1832, see: varians Paykull, 1789.
interantennalis Lea, 1931: 370 (Philonthus; Type locality: New Guinea: Finsch Haven).
intermedius Lacordaire, 1835: 388 (Staphylinus; Type locality: Paris).
— Nordmann, 1837: 74 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 342 cited laminatus Nordmann as a synonym of intermedius Lacordaire; Nordmann did not describe the species as new, he attributed it to Creutzer]; characters; Russia; Sweden; Germany; France).
— Erichson, 1839a: 447 (Philonthus; characters; Germany).
— Heer, 1839: 257 (Philonthus; characters; Switzerland).
— Erichson, 1840: 429 (Philonthus; characters; Germany; France; Italy; Greece; Mesopotamia).
— P. Lucas, 1846: 111 (Philonthus; notes; Algeria).
— Hochhuth, 1849: 132 (Philonthus; characters; Lenkoran; Tiflis).
— Redtenbacher, 1849: 700 (Philonthus; characters; Austria).
— Küster, 1853a: 43 (Philonthus; characters; Germany; Switzerland; France).
— Hardy, 1851: 25 (Philonthus; synonym of chalceus).
— Fairmaire and Laboulbène, 1856: 513 (Philonthus; characters; France).
— Kraatz, 1857d: 570 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 189 (Philonthus; characters; Austria).
— Kraatz, 1858a: 61 (Philonthus; Greece, Crete).
— Peyron, 1858: 425 (Philonthus; Turkey).
— Thomson, 1860: 155 (Philonthus; characters; Scandinavia).
— Fauvel, 1874: 213 [= 1874a: 439] (Philonthus; characters; France; Europe; Barbary; Cyprus; Turkey; Caucasus; Transcaucasia; Mesopotamia; Iran).
— Redtenbacher, 1874: 208 (Philonthus; characters; Austria).
— Seidlitz, 1875: 272 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 350 [= 1877: 206] (Philonthus; characters; France).
— Fauvel, 1878: 121 [= 1878a: 41] (Philonthus; Morocco; Algeria).
— Fauvel, 1886: 53 [= 1886a: 45] (Philonthus; Algeria).
— Fowler, 1888: 260 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 442 (Philonthus; characters; Europe; Mediterranean region).
— Fauvel, 1902b: 108 (Philonthus; Morocco; Algeria; Tunisia).
— Sainte-Claire Deville, 1906: 95 (Philonthus; Corsica).
— Montandon, 1908: 80 (Philonthus; Romania).
— Reitter, 1909: 124 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 342 (Philonthus; catalog).
— Johansen, 1914: 392 (Philonthus; characters; Denmark).
— Gridelli, 1922a: 175 (Philonthus; Persia).
— Portevin, 1929: 357 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1346 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Tottenham, 1949: 417 (Philonthus; checklist; Britain).
— V. Hansen, 1952: 48 (Philonthus; subgenus Philonthus; characters; Denmark).
— Scheerpeltz, 1956c: 1100 (Philonthus; Cyprus).
— Smetana, 1958: 156 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Kocher, 1958: 144 (Philonthus; Morocco).
— Smetana, 1959: 145 (Philonthus; characters).
— Smetana, 1959a: 209 (Philonthus; Albania).
— Scheerpeltz, 1961d: 127 (Philonthus; Italy).
— Ferreira, 1962: 17 (Philonthus; Portugal).
— Scheerpeltz, 1963a: 419 (Philonthus; subgenus Philonthus; Greece).
— Palm, 1963b: 64 (Philonthus; characters; Sweden; Denmark).
— Lohse, 1964: 175 (Philonthus; subgenus Philonthus; characters; central Europe).
— Scheerpeltz, 1964a: 299 (Philonthus; subgenus Philonthus; Crete).
— Raianu, 1964: 103 (Philonthus; Romania).
— Scheerpeltz, 1965b: 401 (Philonthus; Sicily).
— Horion, 1965: 123 (Philonthus; all southern and central Europe; east Europe in Poland; Czechoslovakia, Romania, and Balkans; northern Europe in Britain, Denmark and Sweden).
— Coiffait, 1967a: 390 (Philonthus; characters; Europe; North Africa).
— Raianu, 1968a: 462 (Philonthus; Romania).
— Szujecki, 1968a: 727 (Philonthus; Poland).
— Bolov, 1969: 514 (Philonthus; collecting notes; Russia).
— Bolov, 1969a: 328 (Philonthus; collecting notes; Russia).
— Coiffait, 1970e: 62 (Philonthus; Cyprus).
— Coiffait, 1974: 220 (Philonthus; characters; Europe; Britain; Scandinavia; North Africa).
— Pace and Zanetti, 1977: 288 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Bordoni and Carfi, 1978 (Philonthus; note on peg setae of aedeagus).
— Raichev, 1979: 82 (Philonthus; Bulgaria).
— Muona, 1979: 14 (Philonthus; Sweden; Denmark; Baltic States).
— Burakowski, Mroczkowski, and Stefańska, 1980: 56 (Philonthus; catalog; Poland).
— Hinton, 1981: 661 (Philonthus; characters of egg).
— Frank, 1982: 41 (Philonthus; list of parasites).
— Bordoni, 1984: 342 (Philonthus; Lebanon).
— Tóth, 1984: 44 (Philonthus; characters; Hungary).
— Ádám, 1987: 150 (Philonthus; collecting notes; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Gusarov, 1989: 10 (Philonthus; habitat notes; Ukraine).
— Pilon and Zanetti, 1991: 57 (Philonthus; Italy).
— Angelini, 1991: 192 (Philonthus; Italy).
— Silfverberg, 1992: 16 (Philonthus; Sweden; Denmark; Estonia; Latvia; Lithuania).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— M. Hansen, 1996: 104 (Philonthus; checklist; Denmark).
— Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 37 (Philonthus; Latvia).
— Anderson, Nash, and O'Connor, 1997: 18 (Philonthus; Ireland).
— Assing and Wunderle, 2001: 37 (Philonthus; Cyprus).
— Herman, 2001: 27 (Philonthus; nomen protectum).
▴ Distribution: Tunisia, Algeria, Morocco, Europe, Russia, Georgia, Azerbaijan, Turkey, Lebanon, Iraq, Iran.
aeratus Stephens, 1832: 228 (Philonthus; [Note: This name is older than intermedius]; Type locality: Suffolk; London).
— Curtis, 1829: 25 (Philonthus; nomen nudum; catalog; Britain).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Holme, 1837: 63 (Philonthus; Scilly Islands).
— Stephens, 1839: 394 (Philonthus; characters; England).
— Hardy, 1851: 25 (Philonthus; synonym of chalceus).
— Gemminger and Harold, 1868: 589 (Philonthus; synonym of intermedius).
— Ganglbauer, 1895: 443 (Philonthus; synonym of intermedius).
— Bernhauer and Schubert, 1914: 342 (Philonthus; synonym of intermedius).
— Smetana, 1958: 156 (Philonthus; synonym of intermedius).
— Coiffait, 1974: 220 (Philonthus; synonym of intermedius).
— Herman, 2001: 27 (Philonthus; nomen oblitum).
donisthorpei Dollmann, 1910: 295 (Philonthus; variety of intermedius; Type locality: Ditchling, Sussex).
— Bernhauer and Schubert, 1914: 342 (Philonthus; variety of intermedius).
— Scheerpeltz, 1933: 1347 (Philonthus; variety of intermedius).
— Smetana, 1959: 145 (Philonthus; aberration of intermedius; characters).
— Smetana, 1958: 156 (Philonthus; variety of intermedius).
— Coiffait, 1967a: 390 (Philonthus; aberration of intermedius; characters).
— Szujecki, 1968a: 727 (Philonthus; variety of intermedius; Poland).
— Coiffait, 1974: 220 (Philonthus; variety of intermedius).
— Pope, 1977: 29 (Philonthus; synonym of intermedius).
greipeli W. Kolbe, 1931: 11 (Philonthus; aberration of intermedius; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Altvatergeb.).
— Scheerpeltz, 1933: 1346 (Philonthus; aberration of intermedius).
— Smetana, 1958: 156 (Philonthus; variety of intermedius).
— Smetana, 1959: 145 (Philonthus; aberration of intermedius; characters).
— Coiffait, 1967a: 390 (Philonthus; aberration of intermedius; characters).
— Coiffait, 1974: 220 (Philonthus; variety of intermedius).
intermedius Ragusa, 1891, see: alcyoneus Erichson, 1840.
intermistus Last, 1968: 354 (Philonthus; Type locality: New Guinea: Okapa).
— Last 1981: 120 (Philonthus; New Guinea).
— Last, 1987: 31 (Philonthus; Papua New Guinea).
▴ Distribution: New Guinea.
interocularis Bernhauer, 1915d: 143 (Philonthus; Type locality: Abessinien: Boroda).
interpositus Bernhauer, 1915h: 228 (Philonthus; Type locality: Java: Preanger).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
— Cameron, 1937: 5 (Philonthus; Java).
▴ Distribution: Indonesia.
interpunctatus Hochhuth, 1860, see: Bisnius.
intrudens Tottenham, 1949, see: Bisnius.
invelatus† Scudder, 1900: 55 (Philonthus; fossil; Type locality: Florissant, Colorado).
inversus Horn, 1884, see: Bisnius.
iopterus Erichson, 1840: 928 (Philonthus; Type locality: Bogotae, Columbia).
— Fauvel, 1891: 114 (Philonthus; Venezuela).
— Bernhauer and Schubert, 1914: 342 (Philonthus; catalog).
▴ Distribution: Colombia, Venezuela.
irakoiranicus Scheerpeltz, 1961: 27 (Philonthus; Type locality: Iran: Khuzistan, Shush, [Susa]).
bagdadensis Coiffait, 1967a: 441 (Philonthus; subgenus Philonthus; Type locality: Bagdad).
iranicus Scheerpeltz, 1961, see: biskrensis Fagel, 1957.
iridescens Tottenham, 1949b: 326 (Philonthus; Type locality: Zululand: Eshowe).
— Tottenham, 1955: 173 (Philonthus; characters).
— Tottenham, 1962: 181 (Philonthus; characters).
▴ Distribution: South Africa.
iridicollis Bernhauer, 1932b: 152 (Philonthus; Type locality: Kiambi; Mulongo, Niunzu).
iridiventris Solsky, 1868, see: piceatus Nordmann, 1837.
irinae Ryvkin, 1987: 128 [= 1987a: 98] (Philonthus; subgenus Philonthus; Type locality: Khr. Zhamar-Doban, r. Solzan).
irinus Horn, 1884: 186 (Philonthus; Type locality: Columbus, Texas; [Note: See lectotype designation by Smetana, 1995e]).
— Bernhauer and Schubert, 1914: 342 (Philonthus; catalog).
— Smetana, 1995e: 163 (Philonthus; lectotype designation: Columbus, Texas; characters; Texas).
▴ Distribution: USA.
iris Sharp, 1885: 401 (Philonthus; Type locality: Mexico; Cuernavaca, Capulalpam).
irregularis Mannerheim, 1830, see: concinnus Gravenhorst, 1802.
irritans Tottenham, 1953a: 139 (Philonthus; Type locality: Rhodesia: Mashonaland, Salisbury).
ischnocerus Cameron, 1935a: 374 (Philonthus; Type locality: Nizi: Blukwa).
islamicus Tronquet, 1981: 79 (Philonthus; Type locality: Afghanistan: massif du Kakrak, 2600 m, Koh-I-Baba).
isthmius E. Kangas, 1979, see: confinis Stand, 1941.
iteratus Sharp, 1887: 788 (Philonthus; Type locality: Mexico, Villa Lerdo in Durango).
jacksoni Cameron, 1951, see: Gabrius.
jacobsoni Cameron, 1933d: 388 (Philonthus; Type locality: Sumatra: Gunung Singgalang).
jalapensis Bernhauer, 1910b: 382 (Philonthus; Type locality: Mexiko: Jalapa).
janthinipennis Kolenati, 1846, see: atratus Gravenhorst, 1802.
janus Smetana, 1995e: 324 (Philonthus; Type locality: Quebec: Parc Gatineau, Lac Bourgeois. Also cited from Manitoba, Ontario, Quebec, Saskatchewan, Maryland, Massachusetts, New Hampshire, New Jersey, New York).
japonicus Sharp, 1874: 40 (Philonthus; Type locality: Japan; [Note: See lectotype designation by Schillhammer, 2000]. Originally cited from Simabara).
— Matsumura, 1911: 114 (Philonthus; Japan).
— Bernhauer and Schubert, 1914: 341 (Philonthus; catalog).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog; East Siberia).
— Aleksandrov, 1934: 153 (Philonthus; China).
— Nakane and Sawada, 1960: A123 (Philonthus; Yakushima Island).
— Nakane, 1963a: 90 (Philonthus; characters; habitus photograph; Japan).
— Watanabe and Shibata, 1965: 321 (Philonthus; subgenus Philonthus; Japan).
— Watanabe and Shibata, 1972: 63 (Philonthus; Yaku-shima Island).
— Yuh, Paik, Kwon, and Lee, 1985: 240 (Philonthus; Korea).
— J. Li, 1993: 36 (Philonthus; China).
— Kanehisa, Tsumuki, and Kawazu, 1994 (Philonthus; chemical secretions).
— Watanabe, 1998: 313 (Philonthus; Japan).
— Schillhammer, 1999a: 61 (Philonthus).
— Schillhammer, 2000: 124 (Philonthus; lectotype designation; characters; Russia; Korea; China; Japan).
▴ Distribution: Japan, Korea, China, Russia.
bernhaueri Roubal, 1909a: 374 (Philonthus; variety of japonicus; [preoccupied]; Type locality: Kurilen, Iturup).
binderi Roubal, 1910a: 263 (Philonthus; replacement name for bernhaueri Roubal; valid species; characters).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Schillhammer, 1999a: 61 (Philonthus; synonym of japonicus).
— Schillhammer, 2000: 124 (Philonthus; synonym of japonicus).
seishinensis Bernhauer, 1936g: 308 (Philonthus; Type locality: Mandschurei: Chiau-tou. Also cited from Korea).
— J. Li, 1993: 35 (Philonthus; cited as seighinensis; China).
— Schillhammer, 1999a: 61 (Philonthus; synonym of japonicus).
— Schillhammer, 2000: 124 (Philonthus; synonym of japonicus).
jarrigei Levasseur, 1962: 240 (Philonthus; Type locality: Ruwe [Lualaba], Haut Katanga, Congo belge).
javanus Bernhauer, 1920d: 183 (Philonthus; Type locality: Java: Preanger, Tjigembong).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
— Cameron, 1937: 5 (Philonthus; Java).
▴ Distribution: Indonesia.
jaynesi Bernhauer, 1934: 117 (Philonthus; Type locality: Argentinien: Tucuman: Parque Aconquija).
jeanneli Bernhauer, 1939a: 82 (Philonthus; Type locality: Kenya: Mt. Aberdare, Westhang, mais. forest Kinangop, 2600 m; Marakwet, Chip Cherangani, 3500 m).
— Tottenham, 1949b: 311 (Philonthus; characters; Kenya).
— Tottenham, 1962: 161 (Philonthus; characters).
▴ Distribution: Kenya.
jeannelianus Bernhauer, 1939, see: Gabrius.
jeholensis Kamiya, 1936: 2, 5 (Philonthus; Type locality: Jehol: Weng-niu-te).
jelskii Solsky, 1872, see: Belonuchus.
jenseni Bernhauer, 1912d: 42 (Philonthus; Type locality: Argentinien: Chaunar-Region [Note: In a discussion of Neobisnius parcepunctatus, Frank, 1981: 12, 13 thought this locality probably referred to Chañar Region, La Rioja]).
jenseni A. Strand, 1941, see: strandi Smetana, 1959.
jilinensis J. Li, 1993: 49 (Philonthus; Type locality: Han Cong Ling, Dunhua, Jilin Province).
jocquei Lecoq, 1996: 148 (Philonthus; Type locality: l'archipel des Comores: Grande-Comore, Ivembeni).
johnsoni Fall, 1916, see: nudus Sharp, 1874.
jonenensis Sawada, 1965: 12 (Philonthus; subgenus Philonthus; Type locality: Japan: Pref. Nagano: Mt. Jônen).
jucundus Cameron, 1937a: 103 (Philonthus; Type locality: New Guinea: Kokoda, alt. 1200 ft.).
jujuyensis Bernhauer, 1921c: 107 (Philonthus; Type locality: Argentinien: Jujuy, Cueva Iturbe).
junius Last, 1981: 118 (Philonthus; Type locality: Blackwater River, Sepik).
jureceki Boháč and Hromádka, 1977, see: jurecekianus Boháč and Hromádka, 1980.
jureceki Rambousek, 1913, see: nimbicola Fauvel, 1874.
jurecekianus Boháč and Hromádka, 1980: 430 (Philonthus; replacement name for jureceki Boháč and Hromádka).
jureceki Boháč and Hromádka, 1977: 283 (Philonthus; [preoccupied]; Type locality: Bulgaria: Knazevo, nr. Sofia).
jurgans Tottenham, 1937: 176 (Philonthus; Type locality: England: Kennington; Wicken Fen; Bow St.; Castle Eden; Newick; Arundel; Leds; East Ardsley; Skipwith Common; Thorpe Bay; Richmond; Rudyard; Wixford; Timperley. Also from Germany, Switzerland, and Austria).
— Wüsthoff, 1934: 61 (Philonthus; aberration of varians; [Note: Cited as unicolor, a misidentification of jurgans according to Smetana, 1995e: 241]; illustration of aedeagus).
— V. Hansen, 1952: 71 (Philonthus; subgenus Philonthus; characters; Denmark).
— Palm, 1952: 121 (Philonthus; characters; notes).
— Smetana, 1958: 197 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 157, 160 (Philonthus; characters).
— Smetana, 1960b: 259 (Philonthus; Czechoslovakia).
— Palm, 1963b: 88 (Philonthus; characters; Sweden; Denmark).
— Lohse, 1964: 184 (Philonthus; subgenus Philonthus; characters; central Europe).
— Focarile, 1964: 59 (Philonthus; Italy).
— Scheerpeltz, 1965b: 402 (Philonthus; Sicily).
— Horion, 1965: 160 (Philonthus; Britain; Denmark; Sweden; France; Switzerland; Italy; Albania; Czech Republic; Germany; Austria).
— Coiffait, 1967a: 419 (Philonthus; characters).
— Raianu, 1968: 117 (Philonthus; characters; ecological notes; Romania).
— Szujecki, 1969a: 256 (Philonthus; notes; Poland).
— Coiffait, 1974: 246 (Philonthus; characters; England; Europe; Belgium; Italy; France).
— Pace and Zanetti, 1977: 296 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Frank, 1982: 41 (Philonthus; list of parasites).
— Tóth, 1984: 71 (Philonthus; characters).
— Drugmand, 1987: 275 (Philonthus; characters).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; Benelux; Denmark; Sweden).
— Pilon and Zanetti, 1991: 64 (Philonthus; Italy).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Smetana, 1995e: 241 (Philonthus; characters; British Columbia; Labrador; New Brunswick; Newfoundland; Ontario; Quebec; California; Idaho; Illinois; Maine; Massachusetts; Michigan; New Hampshire; New York; Oregon; Pennsylvania; Vermont; Washington).
— Hodge and Jones, 1995: 39 (Philonthus; characters; Britain).
▴ Distribution: Europe, Canada, USA.
pinguis Tottenham, 1949c: 41 (Philonthus; variety of jurgans; Type locality: Christchurch, Hants.).
— Smetana, 1958: 197 (Philonthus; synonym of jurgans).
— Smetana, 1959: 157 (Philonthus; aberration of jurganus; characters).
— Coiffait, 1967a: 419 (Philonthus; aberration of jurgans; characters).
— Coiffait, 1974: 246 (Philonthus; variety of jurgans).
— Pope, 1977: 29 (Philonthus; synonym of jurgans).
— Smetana, 1995e: 241 (Philonthus; synonym of jurgans).
justus Last, 1968, see: Hesperus.
juvenilis Peyron, 1858: 426 (Philonthus; Type locality: Cilicie, near Tarsous).
— Baudi, 1870: 386 (Philonthus; characters; Cyprus).
— Bernhauer and Schubert, 1914: 342 (Philonthus; catalog).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
— Gridelli, 1943: 130 (Philonthus; characters; Bosnia; Dalmatia; Albania; Asia Minor).
— Smetana, 1953: 119 (Philonthus; Turkey).
— Jarrige, 1951a: 7 (Philonthus; characters).
— Coiffait, 1974: 97 (Paragabrius; subgenus Metagabrius; characters; Caucasus; Asia Minor; Corfu; Balkans; Hungary).
— Tóth, 1984: 76 (Philonthus; characters).
— Lohse and Lucht, 1989: 168 (Philonthus; characters).
▴ Distribution: Romania, Hungary, Greece, Bosnia Hercegovina, Croatia, Albania, Greece, Cyprus, Turkey, Caucasus.
tibialis Eppelsheim, 1876: 431 (Philonthus; Type locality: Nauplia, Griechenland).
— Bernhauer and Schubert, 1914: 342 (Philonthus; synonym of juvenilis).
— Gridelli, 1943: 130 (Philonthus; synonym of juvenilis).
ganglbaueri Bernhauer, 1898: 341 (Philonthus; Type locality: Herkulesbad).
— Bernhauer, 1900: 46 (Philonthus; notes).
— Bernhauer and Schubert, 1914: 342 (Philonthus; synonym of juvenilis).
— Gridelli, 1943: 130 (Philonthus; synonym of juvenilis).
thaseus Coiffait, 1976b: 89 (Philonthus; Type locality: Ile de Thasos, Teològus).
kabardensis Bolov and Kryzhanovskij, 1969, see: spinipes Sharp, 1874.
kainpitensis Last, 1989: 881 (Gabrius; Type locality: Papua New Guinea: Morobe, Kainpit).
kamatembeanus Cameron, 1950g: 38 (Philonthus; subgenus Gabrius; Type locality: Belgian Congo: forest near mont Kamatembe, alt. 2200 m).
kamerunensis Bernhauer and Schubert, 1914: 342 (Philonthus; replacement name for biguttulus Bernhauer).
biguttulus Bernhauer, 1908: 109 (Philonthus; [preoccupied]; Type locality: Kamerun).
kamerunensis Bernhauer, 1915, see: adversarius Bernhauer, 1917.
kamimurai Sawada, 1965: 15 (Paragabrius; Type locality: Japan: Pref. Nagano: Mt. Jônen).
— Smetana, 1995e: 51 (Philonthus; transferred from Paragabrius by reason of Smetana's synonymy of Paragabrius and Philonthus).
▴ Distribution: Japan.
kanalensis Fauvel, 1889b: 264 (Philonthus; Type locality: Kanala).
— Fauvel, 1903d: 270 (Philonthus; New Caledonia).
— Bernhauer and Schubert, 1914: 343 (Philonthus; catalog).
▴ Distribution: New Caledonia.
kapanganus Bernhauer, 1936, see: duodecimpunctatus Bernhauer, 1932.
karimuensis Last, 1968: 371 (Philonthus; Type locality: New Guinea: Karimu).
kasaiensis Bernhauer, 1928g: 109 (Philonthus; Type locality: Belgisch-Kongo: Kasai: Nyombe).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
— Bernhauer, 1940: 136 (Philonthus; Uganda).
— Tottenham, 1955: 160 (Philonthus; subgenus Spatulonthus; characters; notes; Uganda).
— Levasseur, 1971: 299 (Spatulonthus; Dahomey; Ivory Coast; Sierra Leone).
▴ Distribution: Uganda, Congo, Benin, Ivory Coast, Sierra Leone.
kashituensis Tottenham, 1949b: 324 (Philonthus; Type locality: N.W. Rhodesia: Kashitu).
kashmiranus Bernhauer, 1915b: 52 (Philonthus; Type locality: West-Himalaya: Kashmir, Kishtewer Panjal).
— Cameron, 1932: 126 (Philonthus; characters; Kashmir).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
▴ Distribution: India.
kashmiricus Cameron, 1932: 94 (Philonthus; Type locality: Kashmir: Sindh Valley, Haran Plantation).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
— Coiffait, 1982: 30 (Philonthus; Nepal).
▴ Distribution: India, Nepal.
kaszabi Smetana, 1967d: 204 (Philonthus; subgenus Philonthus; Type locality: Mongolei: Central aimak, Ulan-Baator, Nucht im Bogdo ul, 1600 m).
— Smetana 1975c: 171 (Philonthus; Mongolia).
— Shilov, 1976: 150 (Philonthus; Mongolia).
— Smetana, 1995e: 229 (Philonthus; characters; notes; British Columbia; Manitoba; Northwest Territories; Yukon Territory; Alaska; Arizona; California; Colorado; New Mexico; Oregon; Washington).
▴ Distribution: Mongolia, Canada, USA.
katangaensis Levasseur, 1967b: 1625 (Philonthus; subgenus Raucalius; Type locality: Kolwezi, Katanga).
katompeanus Bernhauer, 1932, see: densecaudatus Bernhauer, 1928.
katonae Bernhauer, 1915, see: Gabrius.
kempi Cameron, 1924a: 118 (Philonthus; Type locality: Garo Hills, Assam, Siju Cave).
— Cameron, 1932: 89 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
— Jeannel and Jarrige, 1949: 372 (Philonthus; cave species; Assam).
— Scheerpeltz, 1965: 106 (Philonthus; Burma).
— Coiffait, 1981e: 349 (Philonthus; Baltistan).
▴ Distribution: Myanmar, India, Pakistan.
kenyanus Bernhauer, 1939a: 86 (Philonthus; Type locality: Kenya: Lokitang, Turkana Nord, 750 m).
kerinciensis Schillhammer, 1998: 110 (Philonthus; Type locality: Sumatra: Jambi, Mt. Kerinci, 1900 m).
kessleri Blatchley, 1936, see: Gabrius.
khnzoriani Coiffait, 1967, see: kiautschauensis Bernhauer, 1915.
khouzestanicus Boháč, 1981: 356 (Philonthus; cited as knouzestanicus; Type locality: SE Iran, Tis.).
kiangsiensis Bernhauer, 1933: 30 (Philonthus; Type locality: Kiang-Si: Chang-Tsin-cheng, Südostchina).
kiautschauensis Bernhauer, 1916e: 30 (Philonthus; Type locality: Kiautschau).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
— Schillhammer, 1999a: 61 (Philonthus).
▴ Distribution: China, Russia.
altaicus Bernhauer, 1923a: 124 (Philonthus; Type locality: Semenowsk im Altai).
— Scheerpeltz, 1933: 1332 (Philonthus; catalog).
— Schillhammer, 1999a: 61 (Philonthus; synonym of kiautschauensis).
khnzoriani Coiffait, 1967d: 352 (Philonthus; subgenus Philonthus; Type locality: Lac de Teletsk, Altai).
— Coiffait, 1974a: 198 (Philonthus; Siberia).
— Coiffait, 1974: 276 (Philonthus; characters; central and east Asia).
— Schillhammer, 1999a: 61 (Philonthus; synonym of kiautschauensis).
septempunctatus Tikhomirova, 1973: 169 (Philonthus; [preoccupied]; Type locality: Suputinskii zapovednik).
— Coiffait, 1974a: 198 (Philonthus; synonym of khnzoriani).
— Coiffait, 1974: 276 (Philonthus; synonym of khnzoriani).
gastoni Tikhomirova, 1973a: 184 (Philonthus; replacement name for septempunctatus Tikhomirova).
kigenius Tottenham, 1962: 200 (Philonthus; Type locality: Cape Colony: Simon's Town).
kisirensis Coiffait, 1966c: 509 (Paragabrius; Type locality: Turquie d'Asie, Kisir Dag, 2600 m, villayet de Kars).
— Coiffait, 1974: 92 (Paragabrius; characters; Turkey).
— Smetana, 1995e: 51 (Philonthus; transferred from Paragabrius by reason of Smetana's synonymy of Paragabrius and Philonthus).
▴ Distribution: Turkey.
kiulungensis Bernhauer, 1933, see: emdeni Bernhauer, 1931.
kivuensis Bernhauer, 1934g: 238 (Philonthus; Type locality: Kivu: Kibumba).
kiyoyamai Hayashi, 1993: 80 (Philonthus; Type locality: Taiwan: Pinglin, Taipei Hsien).
klimschi Bernhauer, 1916c: 38 (Philonthus; Type locality: Brasilien: S. Catharina).
klugi Bordoni, 1984, see: sanamus Tottenham, 1955.
kneri† Giebel, 1856: 72 (Philonthus; Fossil; Type locality: Aix, Provence).
kobensis Sharp, 1874: 46 (Philonthus; Type locality: Kobé).
— Duvivier, 1883: 146 (Philonthus; cited as kobeensis; [Note: Misspelling; Bernhauer and Schubert, 1914: 343 cited kobeensis Duvivier as a synonym of kobensis Sharp; Duvivier did not propose an emendation or replacement name, he misspelled kobensis]; catalog).
— Bernhauer and Schubert, 1914: 343 (Philonthus; catalog).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
— Nakane, 1963a: 91 (Philonthus; characters; habitus photograph; Japan).
▴ Distribution: Japan.
kochianus Bernhauer, 1940b: 635 (Philonthus; subgenus Cephalonthus; Type locality: Atlas, Asni).
— Coiffait, 1967a: 411 (Philonthus; characters; Morocco).
— Coiffait, 1974: 289 (Philonthus; characters; Morocco).
▴ Distribution: Morocco.
kokodanus Cameron, 1937, see: Hesperus.
kotgarhensis Cameron, 1932: 109 (Philonthus; Type locality: Simla Hills: Kotgarh, alt. 7000 feet. Dehra Dun).
kraatzi Bernhauer, 1908c: 110 (Philonthus; Type locality: Kamerun).
— Bernhauer and Schubert, 1914: 343 (Philonthus; catalog).
— Tottenham, 1949b: 347 (Philonthus; characters).
▴ Distribution: Cameroon.
krali Hromádka, 1987a: 147 (Philonthus; Type locality: USSR: Uzbekistan: Aktas, 1100–1200, Taskent env.).
kralianus Hromádka, 1987, see: liopterus Sharp, 1889.
kremeni Lecoq, 1997: 284 (Philonthus; Type locality: Madagascar Est, parc de Ranomafana, 800 m).
kristenseni Bernhauer, 1915d: 138 (Philonthus; Type locality: Abessinien: Boroda, Kunhe).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
— Tottenham, 1949b: 308 (Philonthus; notes).
— Tottenham, 1962: 161, 165 (Philonthus; characters).
▴ Distribution: Ethiopia.
kristenseni Bernhauer, 1931, see: incognitus Bernhauer, 1931.
kristensenianus Scheerpeltz, 1933, see: incognitus Bernhauer, 1931.
kryzhanovskii Tikhomirova, 1973, see: liopterus Sharp, 1889.
kuanshanensis Bernhauer, 1914, see: Gabrius.
kuliabensis Bernhauer, 1915, see: Gabrius.
kuluensis Schubert, 1908, see: erythropus Kraatz, 1859.
labastiei Levasseur, 1980: 358 (Philonthus; Type locality: République Centrafricaine: Bangonran, SW de Ndélé).
labdanus Tottenham, 1955: 164 (Philonthus; subgenus Spatulonthus; Type locality: Angola).
labilis Erichson, 1840: 500 (Philonthus; Type locality: Guyana Venezuelensis).
lacustrinus Scheerpeltz, 1933, see: Rabigus.
lacustris Bernhauer, 1915d: 141 (Philonthus; Type locality: Abessinien: See Shalo).
— Bernhauer, 1922b: 176 (Philonthus; Aldabra).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
— Coiffait, 1968a: 136 (Philonthus; Chad).
— Scheerpeltz, 1974c: 22 (Philonthus; Sudan).
▴ Distribution: Chad, Sudan, Ethiopia, Seychelles.
lacustris Casey, 1915, see: Rabigus.
laetabilis Olliff, 1887, see: longicornis Stephens, 1832.
laeticollis Bernhauer, 1933, see: Rabigus.
laetipennis Sharp, 1885: 409 (Philonthus; Type locality: Mexico, Misantla).
laetus Heer, 1839: 259 (Philonthus; Type locality: Walliseralpen).
— Kraatz, 1857d: 574 (Philonthus; characters).
— Gemminger and Harold, 1868: 588 (Philonthus; synonym of ghilianii).
— Fauvel, 1874: 246 [= 1874a: 472] (Philonthus; characters; France; Italy; Sardinia).
— Ganglbauer, 1895: 447 (Philonthus; characters; western Alps; Sardinia).
— Bernhauer and Schubert, 1914: 343 (Philonthus; catalog).
— Portevin, 1929: 360 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
— Coiffait, 1967a: 397 (Philonthus; characters).
— Coiffait, 1974: 230 (Philonthus; characters; Alps).
— Pilon and Zanetti, 1991: 60 (Philonthus; notes; Italy).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Schillhammer, 2000: 154 (Philonthus; characters; France; Italy).
▴ Distribution: France, Italy.
ghilianii Kraatz, 1857d: 574 (Philonthus; Type locality: Piemonteser Alpen).
— Fauvel, 1874: 246 [= 1874a: 472] (Philonthus; synonym of laetus).
— Mulsant and Rey, 1876: 388 [= 1877: 244] (Philonthus; characters; France).
— Ganglbauer, 1895: 447 (Philonthus; synonym of laetus).
— Bernhauer and Schubert, 1914: 343 (Philonthus; synonym of laetus).
— Coiffait, 1974: 231 (Philonthus; synonym of laetus).
— Schillhammer, 2000: 154 (Philonthus; synonym of laetus).
laetus Erichson, 1840, see: Paederomimus.
laeus Olliff, 1887, see: politus Linné, 1758.
laevicollis Lacordaire, 1835: 397 (Staphylinus; [preoccupied by levicollis Brullé according to Article 58.1]; Type locality: Paris).
— Redtenbacher, 1849: 699 (Philonthus; characters; Austria).
— Kiesenwetter, 1851a: 414 (Philonthus; Pyrenees).
— Küster, 1853a: 46 (Philonthus; characters; Europe).
— Fairmaire and Laboulbène, 1856: 514 (Philonthus; characters; France).
— Kraatz, 1857d: 571 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 189, 985 (Philonthus; characters; Austria).
— Kraatz, 1858a: 61 (Philonthus).
— Fauvel, 1874: 232 [= 1874a: 458] (Philonthus; characters; France; Germany; Switzerland; Austria; Russia).
— Redtenbacher, 1874: 208, 216 (Philonthus; characters; Austria).
— Seidlitz, 1875: 272 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 356 [= 1877: 212] (Philonthus; characters; France).
— Ganglbauer, 1895: 443 (Philonthus; characters; Germany; Alps; Carpathians).
— Reitter, 1909: 125 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 343 (Philonthus; catalog).
— Portevin, 1929: 357 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1347 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Smetana, 1958: 158 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 145 (Philonthus; characters).
— Coiffait, 1960: 363, 365 (Philonthus; subgenus Kenonthus; characters; Alps; Carpathians; Massif Central).
— Lohse, 1964: 176 (Philonthus; subgenus Philonthus; characters; central Europe).
— Raianu, 1964: 103 (Philonthus; Romania).
— Horion, 1965: 124 (Philonthus; Germany; France; Switzerland; Austria; Italy; Czech Republic; Slovakia; Romania; Poland).
— Coiffait, 1967a: 435 (Philonthus; subgenus Kenonthus; characters; mountains of central Europe).
— Raianu, 1968a: 462 (Philonthus; Romania).
— Szujecki, 1969a: 255 (Philonthus; notes; Poland).
— Coiffait, 1974: 299 (Philonthus; subgenus Kenonthus; characters; Europe).
— Tóth, 1984: 46 (Philonthus; characters).
— Lucht, 1987: 104 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France).
— Smetana, 1993b: 48 (Philonthus; subgenus Kenonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy ?).
▴ Distribution: Europe, Russia.
adscitus Märkel and Kiesenwetter, 1848a: 322 (Philonthus; Type locality: Kärnthner Alpen).
— Fairmaire and Laboulbène, 1856: 514 (Philonthus; characters; France).
— Kraatz, 1857d: 571 (Philonthus; synonym of laevicollis).
— Bernhauer and Schubert, 1914: 343 (Philonthus; synonym of laevicollis).
— Smetana, 1958: 158 (Philonthus; synonym of laevicollis).
— Coiffait, 1974: 299 (Philonthus; synonym of laevicollis).
laevis Cameron, 1942a: 136 (Philonthus; Type locality: Borneo: Martapura).
laeviventris Schubert, 1909: 291 (Philonthus; Type locality: südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d'Orizaba).
laminatus Creutzer, 1799: 128 (Staphylinus; Type locality: Neuwaldegg).
— Panzer, 1799e: 20 (Staphylinus; characters; illustrations).
— Gravenhorst, 1802: 16 (Staphylinus; characters).
— Marsham, 1802: 504 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 343 cited politus Marsham as a synonym of laminatus; Marsham did not describe the species as new, he attributed it to Linné]; characters; Britain).
— Latreille, 1804: 307 (Staphylinus; characters; Sweden; Germany; France).
— Gravenhorst, 1806: 86 (Staphylinus; characters; France; Prussia).
— Gyllenhal, 1810: 298 (Staphylinus; characters; Sweden).
— Zetterstedt, 1828: 70 (Staphylinus; characters; Lapland).
— Curtis, 1829: 25 (Philonthus; catalog; Britain).
— C. Sahlberg, 1830: 313 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 25 (Staphylinus; Finland; Sweden).
— Mannerheim, 1831: 439 (Staphylinus; Finland; Sweden).
— Stephens, 1832: 226 (Philonthus; characters; England).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Runde, 1835: 5 (Staphylinus; characters; Germany).
— Lacordaire, 1835: 389 (Staphylinus; characters; France).
— Zetterstedt, 1838: 60 (Staphylinus; characters; habitat; Lapland).
— Erichson, 1839a: 446 (Philonthus; characters; Germany).
— Heer, 1839: 257 (Philonthus; characters; Switzerland).
— Stephens, 1839: 394 (Philonthus; characters; England).
— Erichson, 1840: 430 (Philonthus; characters; Europe).
— Laporte, 1840: 177 (Staphylinus; characters; France; Germany).
— Kolenati, 1846: 19 (Philonthus; Transcaucasia).
— Jacobsen, 1849: 686 (Philonthus; characters; Denmark).
— Hochhuth, 1849: 132 (Philonthus; characters; Tiflis; Helenendorf).
— Redtenbacher, 1849: 700 (Philonthus; characters; Austria).
— Kiesenwetter, 1851a: 414 (Philonthus; France).
— Küster, 1853a: 44 (Philonthus; characters; Europe).
— Hardy, 1851: 25 (Philonthus; notes; England).
— Rosenhauer, 1856: 70 (Philonthus; Cadiz).
— Fairmaire and Laboulbène, 1856: 514 (Philonthus; characters; France).
— Kraatz, 1857d: 570 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 189 (Philonthus; characters; Austria).
— Thomson, 1860: 155 (Philonthus; characters; Scandinavia).
— Fauvel, 1865a: 295 [= 1865b: 45] (Philonthus; habitat; France).
— Fauvel, 1874: 229 [= 1874a: 455] (Philonthus; characters; France; Turkey; Caucasus; Transcaucasia).
— Redtenbacher, 1874: 208 (Philonthus; characters; Austria).
— Seidlitz, 1875: 272 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 353 [= 1877: 209] (Philonthus; characters; France).
— J. Sahlberg, 1876: 13 (Philonthus; Finland).
— Fowler, 1888: 260 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 443 (Philonthus; characters; Europe; Mediterranean region).
— Xambeu, 1901: 49 (Philonthus; larval characters).
— Poppius, 1905: 9 (Philonthus; Orkney Islands).
— Reitter, 1909: 125 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 343 (Philonthus; catalog).
— Johansen, 1914: 393 (Philonthus; characters; Denmark).
— Portevin, 1929: 357 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1348 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Paulian, 1941: 227 (Philonthus; larval characters).
— Tottenham, 1949: 417 (Philonthus; checklist; Britain).
— V. Hansen, 1952: 49 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1953: 117 (Philonthus; Turkey).
— Smetana, 1958: 157 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 145 (Philonthus; characters).
— Scheerpeltz, 1961d: 127 (Philonthus; Italy).
— Raianu, 1963: 82 (Philonthus; Romania).
— Palm, 1963b: 65 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 175 (Philonthus; subgenus Philonthus; characters; central Europe).
— Smetana, 1964d: 74 (Philonthus; subgenus Philonthus; Czechoslovakia).
— Horion, 1965: 124 (Philonthus; most of Europe).
— Pototskaia, 1966: 156 (Philonthus; larval characters).
— Coiffait, 1967a: 389 (Philonthus; characters; Europe; Caucasus; Asia Minor).
— Pototskaia, 1967: 57 (Philonthus; larval characters).
— Szujecki, 1968a: 727 (Philonthus; Poland).
— Scheerpeltz, 1968a: 71 (Philonthus; catalog; Austria).
— Jarrige, 1971: 494 (Philonthus; Iran).
— Coiffait, 1974: 221 (Philonthus; characters; Europe; Britain; Scandinavia; Turkey; Caucasus; Transcaucasus).
— Shilov, 1975a: 375 [= 1975b: 97] (Philonthus; Komi region of Russia).
— Coiffait, 1976b: 81 (Philonthus; Greece).
— Pace and Zanetti, 1977: 289 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Coiffait, 1978d: 164 (Philonthus; Turkey).
— Muona, 1979: 14 (Philonthus; Finland; Russia; Sweden; Norway; Denmark; Baltic States).
— Burakowski, Mroczkowski, and Stefańska, 1980: 57 (Philonthus; catalog; Poland).
— Waterston, Owen, Welch, Bacchus and Hammond, 1981: 274 (Philonthus; Outer Hebrides).
— Tóth, 1984: 43 (Philonthus; characters).
— Muona and Viramo, 1986: 13 (Philonthus; Finland).
— Ádám, 1987: 150 (Philonthus; collecting notes; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Pilon and Zanetti, 1991: 56 (Philonthus; Italy).
— Silfverberg, 1992: 16 (Philonthus; Finland; Russia; Sweden; Norway; Denmark; Estonia; Latvia; Lithuania).
— Welch, 1993: 228 (Philonthus; ovariole number and ovary structure).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Franzen, 1996: 206 (Philonthus; collecting notes; Germany).
— Tagliapietra and Zanetti, 1996: 129 (Philonthus; ecological notes; Italy).
— M. Hansen, 1996: 104 (Philonthus; checklist; Denmark).
— Smoleński, 1997: 79 (Philonthus; collecting notes; Poland).
— T. Wagner, 1997: 234 (Philonthus; collecting notes; Germany).
— Wenzel, 1997: 346 (Philonthus; collecting notes; Germany).
— Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 37 (Philonthus; Latvia).
— Anderson, Nash, and O'Connor, 1997: 18 (Philonthus; Ireland).
— Herman, 2001: 27 (Philonthus; nomen protectum).
▴ Distribution: Europe, Russia, Georgia, Turkey, Iran.
aeneus De Geer, 1774: 23 (Staphylinus; [Note: This name is older than laminatus]; Type locality: Not cited).
— O. Müller, 1776: 98 (Staphylinus; characters).
— Fourcroy, 1785: 168 (Staphylinus; characters; France).
— Villers, 1789: 422, 423 (Staphylinus; characters; France).
— Gmelin, 1790: 2034 (Staphylinus; characters).
— Olivier, 1795:(42): 25 (Staphylinus; synonym of politus Linné).
— Turton, 1802: 518 (Staphylinus; characters; Europe).
— Gravenhorst, 1802: 17 (Staphylinus; synonym of laminatus Creutzer).
— Marsham, 1802: 511 (Staphylinus; [Note: This name is cited by Bernhauer and Schubert, 1914: 343 as, synonym of laminatus Creutzer with Marsham as the author but Marsham attributed the species to De Geer]; characters; Britain).
— Gyllenhal, 1810: 298 (Staphylinus; synonym of laminatus Creutzer).
— Stephens, 1832: 227 (Philonthus; characters; England).
— Curtis, 1829: 25 (Philonthus; catalog; Britain).
— Erichson, 1839a: 449 (Philonthus; characters; Germany).
— Stephens, 1839: 394 (Philonthus; variety of laminatus; characters; England).
— Jacobsen, 1849: 687 (Philonthus; characters; Denmark).
— Gemminger and Harold, 1868: 589 (Philonthus; [Note: Marsham, 1802, cited as author but Marsham attributed the species to De Geer]; synonym of laminatus).
— Ganglbauer, 1895: 443 (Philonthus; [Note: Marsham, 1802, cited as author but Marsham attributed the species to De Geer]; synonym of laminatus).
— Bernhauer and Schubert, 1914: 343 (Philonthus; synonym of laminatus).
— Smetana, 1958: 157 (Philonthus; synonym of laminatus).
— Herman, 2001: 27 (Philonthus; nomen oblitum).
matutinalis Faldermann, 1835: 124 (Staphylinus; Type locality: [area] Persico-Armeniaca).
— Gemminger and Harold, 1868: 589 (Philonthus; synonym of laminatus).
— Ganglbauer, 1895: 443 (Philonthus; synonym of laminatus).
— Bernhauer and Schubert, 1914: 343 (Philonthus; synonym of laminatus).
— Smetana, 1958: 157 (Philonthus; synonym of laminatus).
— Coiffait, 1974: 221 (Philonthus; synonym of laminatus).
coxatus Curtis, 1836: pl. 610 (Philonthus; Type locality: Britain).
viridanus Nordmann, 1837: 74 (Philonthus; Type locality: Germania; Livonia).
— Erichson, 1839a: 446 (Philonthus; synonym of laminatus).
— Heer, 1839: 257 (Philonthus; synonym of laminatus).
— Erichson, 1840: 430 (Philonthus; synonym of laminatus).
— Kraatz, 1857d: 570 (Philonthus; synonym of laminatus).
— Ganglbauer, 1895: 443 (Philonthus; synonym of laminatus).
— Bernhauer and Schubert, 1914: 343 (Philonthus; synonym of laminatus).
— Smetana, 1958: 157 (Philonthus; synonym of laminatus).
pseudonitidus Everts, 1923: 211 (Philonthus; aberration of laminatus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Zeeburg, Amsterdam).
— Scheerpeltz, 1933: 1348 (Philonthus; aberration of laminatus).
— Smetana, 1959: 145 (Philonthus; aberration of laminatus; characters).
— Smetana, 1958: 157 (Philonthus; aberration of laminatus).
— Scheerpeltz, 1961d: 128 (Philonthus; aberration of laminatus; Italy).
— Coiffait, 1967a: 389 (Philonthus; variety of laminatus; characters).
— Coiffait, 1974: 221 (Philonthus; variety of laminatus).
lampropterus Bernhauer, 1934, see: Mentophilonthus.
lamtoensis Levasseur, 1967b: 1626 (Philonthus; subgenus Raucalius; Type locality: Côte d'Ivoire, Lamto, Toumodi).
lan Schillhammer, 1998: 107 (Philonthus; Type locality: China: Yunnan, Heishui, 35 km N Lijiang, 27°13′N, 100°19′E).
lapidicola Cameron, 1932, see: Gabrius.
lassallei Coiffait, 1981d: 327 (Philonthus; Type locality: Nohabkarat, env. Danan, Nepal central, 2400 m).
lasti Tottenham, 1955: 176 (Philonthus; Type locality: Belgian Congo).
latecinctus Sharp, 1885: 415 (Philonthus; Type locality: Guatemala, Cerro Zunil 4000 to 5000 feet, San Geronimo).
laterangulus Schillhammer, 1998, see: caeruleipennis Mannerheim, 1830.
lathrobioides Baudi, 1848, see: Neobisnius.
laticeps Zetterstedt, 1828, see: politus Linné, 1758.
laticollis Fauvel, 1874, see: Gabrius.
laticornis Bernhauer, 1906a: 190 (Philonthus; Type locality: Madagaskar).
— Bernhauer and Schubert, 1914: 343 (Philonthus; catalog).
— Tottenham, 1949b: 362 (Philonthus; characters).
— Lecoq, 1997: 285 (Philonthus; characters; Madagascar).
▴ Distribution: Madagascar.
latiusculus Hochhuth, 1851: 15 (Philonthus; Type locality: Dahurien).
— Hochhuth, 1862: 53 (Philonthus; Russia).
— Poppius, 1909: 23 (Philonthus; Lena Valley, Siberia).
— Bernhauer and Schubert, 1914: 343 (Philonthus; catalog).
— Scheerpeltz, 1933: 1348 (Philonthus; catalog).
— Smetana, 1963c: 295 (Philonthus; subgenus Philonthus; Mongolia).
— Smetana, 1965e: 173 (Philonthus; Mongolia).
— Coiffait, 1967a: 397 (Philonthus; characters; central Asia).
— Smetana, 1967d: 203 (Philonthus; Mongolia).
— Coiffait, 1974: 267 (Philonthus; characters; Siberia; Mongolia).
— Smetana 1975c: 170 (Philonthus; Mongolia).
— Shilov, 1976: 149 (Philonthus; Mongolia).
— Ryabukhin, 1999: 64 (Philonthus; catalog; habitat; distribution; northeastern Russia).
— Schillhammer, 2000: 154 (Philonthus; characters; Russia; Mongolia; China).
▴ Distribution: Russia, Mongolia, China.
immutatus Motschulsky, 1860: 569 (Gabrius; Type locality: Mongolie près de Kiahta).
— Hochhuth, 1862: 54 (Philonthus; characters; Siberia; Kiachta).
— Fauvel, 1875a: XXXI [= 1875b: 233] (Philonthus; synonym of latiusculus).
— Bernhauer and Schubert, 1914: 343 (Philonthus; synonym of latiusculus).
— Coiffait, 1974: 267 (Philonthus; synonym of latiusculus).
— Schillhammer, 2000: 154 (Philonthus; synonym of latiusculus).
lativentris Motschulsky, 1858, see: delicatulus Boheman, 1858.
lautus Casey, 1915, see: Bisnius.
laxatus Fauvel, 1875a: XXXI [= 1875b: 233] (Philonthus; Type locality: Caucase).
— Hochhuth, 1862: 55 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 343 cited lucens Hochhuth as a synonym of laxatus; Hochhuth neither described the species as new nor provided characters for it, he attributed it to Mannerheim]; Armenia).
— Bernhauer and Schubert, 1914: 343 (Philonthus; catalog).
— Scheerpeltz, 1933: 1348 (Philonthus; catalog).
— Gridelli, 1943: 129 (Philonthus; characters; Caucasus).
— Smetana, 1963d: 405 (Philonthus; subgenus Onychophilonthus; characters; Caucasus).
— Coiffait, 1967a: 437 (Philonthus; subgenus Onychophilonthus; characters; Caucasus).
— Korge, 1971a: 7 (Philonthus; subgenus Onychophilonthus; Turkey).
— Coiffait, 1974: 306 (Philonthus; subgenus Onychophilonthus; characters; Russia; Caucasus; Transcaucasus).
— Solodovnikov, 1998a: 15 (Philonthus; notes; Russia, Krasnodar territory).
▴ Distribution: Russia, Caucasus, Armenia, Turkey.
laxellus Casey, 1915, see: Rabigus.
leai Scheerpeltz, 1933: 1348 (Philonthus; replacement name for pauper Lea).
pauper Lea, 1931: 371 (Philonthus; [preoccupied]; Type locality: Papua: Mount Lamington).
lebedevi Bernhauer, 1910, see: Gabrius.
lebkanus Fauvel, 1907: 39 (Philonthus; Type locality: Nairobi; Kilimandjaro; riviere Himo; Abyssinie: Lebka; Bogos, 1500 m).
— Bernhauer and Schubert, 1914: 343 (Philonthus; catalog).
— Tottenham, 1962: 161 (Philonthus; characters).
▴ Distribution: Kenya, Ethiopia.
lecontei Horn, 1884: 222 (Philonthus; Type locality: California; [Note: See lectotype designation by Smetana, 1995e]. Also cited from Colorado).
— Bernhauer and Schubert, 1914: 344 (Philonthus; catalog).
— Hatch, 1957: 181 (Philonthus; characters; British Columbia; Washington).
— Smetana, 1995e: 467 (Philonthus; lectotype designation: Cal [California]; characters; notes; British Columbia; Arizona; California; Colorado; Nevada; New Mexico; Oregon; Utah; Washington).
▴ Distribution: Canada, USA.
lederi Eppelsheim, 1893: 49 (Philonthus; Type locality: Ost-Sibirien, in der Nähe von Irkutsk, von Tunka bis Munku-Sardik im Sajan'schen Gebirge).
— Bernhauer and Schubert, 1914: 344 (Philonthus; catalog).
— Scheerpeltz, 1933: 1348 (Philonthus; catalog).
— Jansson, 1943 (Philonthus; characters; Finland).
— Nakane, 1963: A239 (Philonthus; [Note: Nakane stated some doubt about identification of these specimens]; Japan).
— Smetana, 1963d: 408 (Philonthus; subgenus Onychophilonthus; characters; Siberia; Mongolia; Kamtschatka).
— Palm, 1963b: 86 (Philonthus; characters; Sweden).
— Coiffait, 1967a: 437 (Philonthus; subgenus Onychophilonthus; characters; Scandinavia; Asia).
— Coiffait, 1974: 307 (Philonthus; subgenus Onychophilonthus; characters; Mongolia; Baikal region).
— Shibata, 1974: 34 (Philonthus; Japan).
— Shilov, 1976: 153 (Philonthus; Mongolia).
— Siitonen, 1993: 227 (Philonthus; notes; Finland).
— J. Li, 1993: 37 (Philonthus; China).
▴ Distribution: Finland, Russia, Mongolia, China, Japan.
ledouxi Coiffait, 1979, see: variipennis Kraatz, 1859.
leechensis Hatch, 1957: 190 (Philonthus; Type locality: British Columbia: Aspen Grove. Also cited from Washington).
— Smetana, 1965: 21 (Philonthus; subgenus Philonthus; Labrador; Newfoundland).
— Smetana, 1965a: 7 (Philonthus; British Columbia; Alaska).
— Smetana, 1995e: 277 (Philonthus; characters; Alberta; British Columbia; Labrador; Manitoba; Newfoundland; Northwest Territories; Quebec; Saskatchewan; Yukon Territory; Alaska; Arizona; California; Colorado; Idaho; Minnesota; Montana; Oregon; Washington; Wisconsin).
▴ Distribution: Canada, USA.
leonensis Cameron, 1934e: 372 (Philonthus; Type locality: Sierra Leone: Freetown).
lepidulus Stephens, 1832, see: discoideus Gravenhorst, 1802.
lepidulus LeConte, 1863, see: Neobisnius.
lepidus Gravenhorst, 1802: 31 (Staphylinus; Type locality: Not cited).
— Latreille, 1804: 318 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 74 (Staphylinus; characters).
— Gravenhorst, 1806: 74 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 344 cited flavopterus Gravenhorst as a synonym of lepidus but Gravenhorst did not describe the species as new, he attributed it to Fourcroy in whose work Geoffroy's new species are published]; characters; France).
— Gyllenhal, 1810: 328 (Staphylinus; characters; Sweden).
— Curtis, 1829: 28 (Bolitobius; catalog; Britain).
— C. Sahlberg, 1830: 323 (Staphylinus; characters; Finland).
— Runde, 1835: 10 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 344 cited fulvipes Runde as a synonym of lepidus Gravenhorst; Runde did not describe the species as new, he attributed it to Gyllenhal who attributed it to Panzer who attributed it to Fabricius]; characters; Germany).
— Lacordaire, 1835: 401 (Staphylinus; characters; France).
— Erichson, 1839a: 457 (Philonthus; characters; Germany).
— Heer, 1839: 262 (Philonthus; characters; Switzerland).
— Erichson, 1840: 450 (Philonthus; characters; Germany; Sweden; France).
— Jacobsen, 1849: 693 (Philonthus; characters; Denmark).
— Redtenbacher, 1849: 701 (Philonthus; characters; Austria).
— Küster, 1853a: 57 (Philonthus; characters; Sweden; Finland; Germany; France; Switzerland).
— Fairmaire and Laboulbène, 1856: 515 (Philonthus; characters; France).
— Kraatz, 1857d: 588 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 191 (Philonthus; characters; Austria).
— Thomson, 1860: 163 (Philonthus; characters; Scandinavia).
— Hochhuth, 1862: 57 (Philonthus; Russia).
— Fauvel, 1874: 248 [= 1874a: 474] (Philonthus; characters; France; Austria; Lake Baikal; Amur region).
— Redtenbacher, 1874: 210 (Philonthus; characters; Austria).
— Seidlitz, 1875: 275 (Philonthus; characters; Baltic region).
— Fauvel, 1876a: 75 [= 1876: 260] (Philonthus; Dauria).
— Mulsant and Rey, 1876: 413 [= 1877: 269] (Philonthus; characters; France).
— J. Sahlberg, 1876: 20 (Philonthus; Finland).
— Fowler, 1888: 266 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 449 (Philonthus; characters; north and middle Europe; east Siberia).
— Poppius, 1909: 25 (Philonthus; subgenus Gefyrobius; Lena Valley, Siberia).
— Reitter, 1909: 128 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 344 (Philonthus; catalog).
— Johansen, 1914: 399 (Philonthus; characters; Denmark).
— Portevin, 1929: 361 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1348 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— V. Hansen, 1952: 57 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1958: 187 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 149 (Philonthus; characters).
— Palm, 1963b: 73 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 179 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 143 (Philonthus; Irkut and Vladivostok regions; Finland; Sweden; Norway; Denmark; Britain; Russia; Ukraine; Baltics; Poland; Hungary; Romania; Serbia; Bosnia; Croatia; Italy; Switzerland; France; Belgium; Holland; Germany; Austria).
— Coiffait, 1967a: 434 (Philonthus; subgenus Trionthus; characters; Europe).
— Raianu, 1968a: 466 (Philonthus; Romania).
— Szujecki, 1968a: 730 (Philonthus; Poland).
— Coiffait, 1974: 193 (Philonthus; subgenus Trionthus; characters; Europe; Siberia; France).
— Shilov, 1976: 150 (Philonthus; Mongolia).
— Pope, 1977: 29 (Philonthus; Britain).
— Tóth, 1984: 54 (Philonthus; characters; Hungary).
— Ádám, 1987: 151 (Trionthus; collecting notes; Hungary).
— Lucht, 1987: 106 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Pilon and Zanetti, 1991: 57 (Philonthus; Italy).
— J. Li, 1993: 37 (Philonthus; cited as lepoidus; China).
— Smetana, 1993b: 48 (Philonthus; subgenus Trionthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
▴ Distribution: Europe, Russia, Kazakhstan, Mongolia, China.
gilvipes Erichson, 1839a: 456 (Philonthus; Type locality: Berlin).
— Erichson, 1840: 448 (Philonthus; characters; Germany).
— Redtenbacher, 1849: 702 (Philonthus; characters; Austria).
— Kraatz, 1857d: 588 (Philonthus; synonym of lepidus).
— Redtenbacher, 1857: 192 (Philonthus; characters; Austria).
— Gemminger and Harold, 1868: 589 (Philonthus; synonym of lepidus).
— Fauvel, 1874: 248 [= 1874a: 474] (Philonthus; synonym of lepidus).
— Redtenbacher, 1874: 211 (Philonthus; characters; Austria).
— Seidlitz, 1875: 275 (Philonthus; variety of lepidus; characters; Baltic region).
— Ganglbauer, 1895: 449 (Philonthus; synonym of lepidus).
— Reitter, 1909: 128 (Philonthus; variety of gilvipes; characters; Germany).
— Bernhauer and Schubert, 1914: 344 (Philonthus; variety of lepidus).
— Scheerpeltz, 1933: 1348 (Philonthus; variety of lepidus).
— Smetana, 1958: 187 (Philonthus; aberration of lepidus).
— Smetana, 1959: 149 (Philonthus; aberration of lepidus; characters).
— Coiffait, 1974: 193 (Philonthus; variety of lepidus).
contiguus Motschulsky, 1860: 568 (Gabrius; Type locality: environs de Verkhnie-Oudinsk au delà du lac Baical).
— Hochhuth, 1862: 55 (Philonthus; characters; Lake Baikal).
— Bernhauer and Schubert, 1914: 344 (Philonthus; synonym of lepidus).
— Smetana, 1958: 187 (Philonthus; synonym of lepidus).
— Coiffait, 1974: 193 (Philonthus; synonym of lepidus).
lepineyi Scheerpeltz, 1934a: 98 (Philonthus; subgenus Gefyrobius; Type locality: Soudan, Nema).
leptocerus Fauvel, 1895: 266 (Philonthus; Type locality: Bírmanie, Rangoon; Teinzo; Cochinchine, Long-Xuyen).
— Bernhauer and Schubert, 1914: 344 (Philonthus; catalog).
— Cameron, 1932: 150 (Philonthus; characters; Burma).
— Scheerpeltz, 1933: 1348 (Philonthus; catalog).
— Schillhammer, 1999a: 61 (Philonthus).
▴ Distribution: Myanmar, Cambodia, Vietnam, Indonesia.
pseudooculatus Bernhauer, 1928: 16 (Philonthus; Type locality: Ostindien: Kambodja; also listed from Sumatra).
— Scheerpeltz, 1933: 1356 (Philonthus; catalog).
— Cameron, 1933d: 387 (Philonthus; cited as pseudoculatus; Sumatra).
— Schillhammer, 1999a: 61 (Philonthus; synonym of leptocerus).
leucopygus Kraatz, 1859: 90 (Philonthus; Type locality: India orientalis).
— Bernhauer and Schubert, 1914: 344 (Philonthus; catalog).
— Cameron, 1932: 118 (Philonthus; characters; East Indies).
— Scheerpeltz, 1933: 1348 (Philonthus; catalog; Malay Peninsula; Sumatra).
▴ Distribution: “East Indies”, Malaysia, Indonesia.
leucotus Erichson, 1840: 507 (Philonthus; Type locality: Nepal).
— Kraatz, 1859: 98 (Philonthus; characters; Nepal).
— Bernhauer and Schubert, 1914: 344 (Philonthus; catalog).
— Cameron, 1932: 152 (Philonthus; characters; Nepal).
— Scheerpeltz, 1933: 1349 (Philonthus; catalog).
▴ Distribution: Nepal.
levatis Last, 1973: 426 (Philonthus; Type locality: New Guinea: Wau, 4000 ft.).
leveri Cameron, 1943b: 565 (Philonthus; Type locality: Fiji: Vanua Levu).
lewisius Sharp, 1874: 42 (Philonthus; Type locality: Nagasaki).
— Fauvel, 1903: 159 (Philonthus; synonym of aeneipennis).
— Fauvel, 1904a: 59 (Philonthus; synonym of aeneipennis).
— Bernhauer and Schubert, 1914: 327 (Philonthus; synonym of aeneipennis).
— Bernhauer, 1922c: 232 (Philonthus; Formosa).
— Cameron, 1932: 113 (Philonthus; synonym of aeneipennis).
— Gridelli, 1936a: 147 (Philonthus; characters; valid species).
— Bernhauer, 1940b: 635 (Philonthus; subgenus Cephalonthus).
— Cameron, 1949b: 468 (Philonthus; cited as valid species; Japan, Formosa).
— Hasegawa and Kanie, 1992: 20 (Philonthus; seashore collection; Japan).
— Hayashi, 1994: 125 (Philonthus; valid species; characters; Japan).
— Shibata, 1994a: 106 (Philonthus; characters; Taiwan).
▴ Distribution: Japan, Taiwan.
leyei Levasseur, 1980: 356 (Philonthus; Type locality: Sénégal: M'boro).
lhesgicus Kolenati, 1846, see: ebeninus Gravenhorst, 1802.
libanicus Saulcy, 1864, see: Gabronthus.
lidarensis Cameron, 1932: 94 (Philonthus; Type locality: Kashmir: Lidar Valley, Liderwat, alt. 9000 feet).
— Scheerpeltz, 1933: 1349 (Philonthus; catalog).
— Cameron, 1934: 19 (Philonthus; Indian Tibet).
— Coiffait, 1982a: 232 (Philonthus; Kashmir).
▴ Distribution: India.
limaensis Solsky, 1872: 300 (Philonthus; Type locality: Lima).
limbatus Fauvel, 1900, see: Gabronthus.
limbipennis Fauvel, 1905b: 179 (Philonthus; Type locality: Diego-Suarez; forêt Audroy; Ambovombe).
— Bernhauer and Schubert, 1914: 344 (Philonthus; catalog).
— Tottenham, 1962: 216 (Philonthus; characters).
▴ Distribution: Madagascar.
limbukus Tottenham, 1955: 168 (Philonthus; Type locality: Kenya: Muguga).
limulus Tottenham, 1955: 166 (Philonthus; subgenus Spatulonthus; Type locality: Transvaal: Pretoria).
lindbergi Scheerpeltz, 1958a: 17 (Philonthus; Type locality: Van, Ostufer des Van-Sees in Armenien).
— Coiffait, 1967a: 403 (Philonthus; characters; Asia Minor).
— Coiffait, 1974: 282 (Philonthus; characters; Armenia).
▴ Distribution: Turkey.
lindemanni Scheerpeltz, 1960, see: aeneipennis Boheman, 1858.
lindrothi Smetana, 1965: 18 (Philonthus; subgenus Philonthus; Type locality: W. Newfoundland, Deer Lake. Also cited from Nova Scotia).
— Smetana, 1995e: 271 (Philonthus; characters; notes; Alberta; British Columbia; Manitoba; New Brunswick; Newfoundland; Northwest Territories; Nova Scotia; Ontario; Quebec; Saskatchewan; Alaska; Arizona; California Colorado; Idaho; Illinois, Indiana, Iowa; Kansas; Maine; Massachusetts; Michigan; Minnesota; Missouri; Montana; Nebraska; Nevada; New Hampshire; New Jersey; New York; North Carolina; North Dakota; Ohio; Oregon; Pennsylvania; Rhode Island; South Dakota; Vermont; Washington; Wisconsin).
▴ Distribution: Canada, USA.
linearis Casey, 1915, see: Gabrius.
linearis Hochhuth, 1849, see: quisquiliarius Gyllenhal, 1810.
lineifrons Cameron, 1926, see: Gabronthus.
lineiventris Bernhauer, 1937b: 611 (Philonthus; Type locality: N. W. Uluguru).
lineolatus Tottenham, 1962: 214, 216 (Philonthus; Type locality: Mauritius).
linkei Bernhauer, 1908, see: longicornis Stephens, 1832.
linki Solsky, 1867: 81 (Philonthus; Type locality: près de Samara).
— Bernhauer and Schubert, 1914: 344 (Philonthus; catalog).
— Scheerpeltz, 1933: 1349 (Philonthus; catalog).
— Lohse, 1956: 91 (Philonthus; characters; notes).
— Coiffait, 1974: 270 (Philonthus; subgenus Philonthus; characters; Russia; western Siberia).
— J. Li, 1993: 33 (Paragabrius; cited as linkei; China).
▴ Distribution: Russia, China.
liopterus Sharp, 1889: 36 (Philonthus; Type locality: Yezo).
— Bernhauer and Schubert, 1914: 344 (Philonthus; catalog).
— Scheerpeltz, 1933: 1349 (Philonthus; catalog).
— Nakane, 1963a: 91 (Philonthus; characters; habitus photograph; Japan).
— Kanehisa, Tsumuki, and Kawazu, 1994 (Philonthus; cited as lioterus; chemical secretions).
— Schillhammer, 1999a: 61 (Philonthus).
▴ Distribution: Japan, Russia, Tajikistan.
kryzhanovskii Tikhomirova, 1973: 166 (Philonthus; Type locality: Suputinskii zapovednik).
kralianus Hromádka, 1987a: 149 (Philonthus; Type locality: USSR: Tadzikistan, Ziddy, Hissar. chr. 2100 m).
lissonurus Tottenham, 1949b: 341 (Philonthus; Type locality: Equateur: Boende).
— Levasseur, 1966a: 1213 (Philonthus; Congo; Zaire; Cameroon; Ivory Coast).
— Levasseur, 1968a: 54 (Philonthus; Central African Republic).
▴ Distribution: Congo Republic, Congo, Central African Republic, Ivory Coast.
lithocharinus LeConte, 1863, see: Cafius.
littorinus Gistel, 1857: 75 (Philonthus; [Note: Nomen dubium]; Type locality: Alpes Helvetiae).
lituratus Stephens, 1832, see: varians Paykull, 1789.
lividipes Baudi, 1848, see: Gabrius.
lomatus Erichson, 1840: 482 (Philonthus; Type locality: Amer. b.; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from America septentrionalis).
— LeConte, 1850: 220 (Philonthus; Lake Superior).
— Provancher, 1877: 250 (Philonthus; characters; Quebec).
— Horn, 1884: 204 (Philonthus; characters; North America).
— Blatchley, 1910: 386 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 344 (Philonthus; catalog).
— Notman, 1919: 135 (Philonthus; New York).
— Notman, 1919a: 95 (Philonthus; Ontario).
— Notman, 1920a: 180 (Philonthus; New York).
— Notman, 1920b: 706 (Philonthus; Florida).
— Scheerpeltz, 1933: 1349 (Philonthus; catalog).
— Hatch, 1957: 186 (Philonthus; characters; British Columbia; Washington).
— Smetana, 1965: 22 (Philonthus; subgenus Philonthus; notes; Nova Scotia; Newfoundland).
— Smetana, 1965a: 7 (Philonthus; British Columbia).
— Frank, 1984: 479 (Philonthus; notes; characters; Haiti).
— Dearborn and Donahue, 1994: 15 (Philonthus; Maine).
— Smetana, 1995e: 390 (Philonthus; lectotype designation: Amer. b.; characters; notes; Alberta, British Columbia, Manitoba, New Brunswick, Newfoundland, Northwest Territories, Nova Scotia, Ontario, Quebec, Saskatchewan, Alabama, Arkansas, Connecticut, Georgia, Idaho, Illinois, Indiana, Iowa, Kansas, Kentucky, Louisiana, Maine, Maryland, Massachusetts, Michigan, Minnesota, Missouri, Montana, Nebraska, New Hampshire, New Jersey, New York, North Carolina, North Dakota, Ohio, Oklahoma, Oregon, Pennsylvania, Rhode Island, South Dakota, Texas, Vermont, Virginia, Washington, West Virginia, Wisconsin, Wyoming).
— Downie and Arnett, 1996: 380 (Philonthus; characters; USA).
▴ Distribution: Canada, USA, Haiti.
georgianus Sachse, 1852: 143 (Philonthus; Type locality: Georgia).
— Fauvel, 1869: 488 (Philonthus; synonym of sobrinus).
— Bernhauer and Schubert, 1914: 344 (Philonthus; synonym of lomatus).
— Scheerpeltz, 1933: 1349 (Philonthus; aberration of lomatus).
— Smetana, 1995e: 390 (Philonthus; synonym of lomatus).
londianus Cameron, 1951, see: Gabrius.
longiceps Fauvel, 1879, see: minutus Boheman, 1848.
longicollis Bernhauer, 1908, see: Bisnius.
longicornis Stephens, 1832: 237 (Philonthus; Type locality: London).
— Curtis, 1829: 26 (Philonthus; nomen nudum; catalog; Britain).
— Erichson, 1839a: 466 (Philonthus; [Note: Bernhauer and Schubert, 1914: 345 cited varians variety b as a synonym of longicornis; Erichson included it under opacus, but the variety is unavailable]; characters).
— Stephens, 1839: 397 (Philonthus; characters; England).
— Fairmaire, 1849: 290 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 345 cited varians Fairmaire as a synonym of longicornis; Fairmaire did not describe the species as new, he attributed it to Paykull]; notes; Tahiti).
— Hardy, 1851: 29 (Philonthus; synonym of varians).
— Fauvel, 1874: 254 [= 1874a: 480] (Philonthus; characters; France; Mediterranean region; Siberia; Japan; Ceylon; Azores; Canary Islands; Madeira; Cape Verde Islands; Ascension Island; South Africa; Madagascar; Tahiti; Australia; North and South America).
— Solsky, 1874: 178 (Philonthus; characters; Turkestan).
— J. Sahlberg, 1876: 20 (Philonthus; Finland).
— Wollaston, 1877: 28 (Philonthus; characters; St. Helena).
— Fauvel, 1877: 264 (Philonthus; characters; Australia; Tahiti).
— Fauvel, 1878: 125 [= 1878a: 45] (Philonthus; North Africa).
— Fauvel, 1878g: 548 (Philonthus; New Caledonia).
— Horn, 1884: 195 (Philonthus; characters; North America).
— Sharp, 1885: 416 (Philonthus; Mexico).
— Fauvel, 1886: 56 [= 1886a: 48] (Philonthus; North Africa).
— Fowler, 1888: 273 (Philonthus; characters; Britain).
— Fauvel, 1889b: 266 (Philonthus; New Caledonia).
— Fauvel, 1891: 115 (Philonthus; Venezuela).
— Ganglbauer, 1895: 454 (Philonthus; characters; Europe).
— Fauvel, 1902b: 112 (Philonthus; Barbary; Egypt; Azores; Madeira; Canaries).
— Bernhauer, 1902b: 27 (Philonthus; Ceylon).
— Fauvel, 1903: 159 (Philonthus; India).
— Fauvel, 1903d: 271 (Philonthus; New Caledonia; Iles des Pins).
— Fauvel, 1904a: 59 (Philonthus; Nilghiris).
— Fall and Cockerell, 1907: 166 (Philonthus; New Mexico).
— Bernhauer, 1908: 19 (Philonthus; Australia).
— Bernhauer, 1908c: 110 (Philonthus; German Southwest Africa).
— Reitter, 1909: 130 (Philonthus; characters; Germany).
— Blatchley, 1910: 383 (Philonthus; characters; Indiana).
— Eichelbaum, 1913: 130 (Philonthus; Tanzania).
— Bernhauer and Schubert, 1914: 344 (Philonthus; catalog).
— Johansen, 1914: 408 (Philonthus; characters; Denmark).
— Bernhauer, 1920: 16 (Philonthus; Australia).
— Gridelli, 1920a: 18, 19 (Philonthus; characters; Italy).
— Bernhauer, 1921a: 42 (Philonthus; Easter Island).
— Bernhauer, 1922c: 232 (Philonthus; Formosa).
— Cameron, 1924: 87 (Philonthus; New Guinea).
— Dodero, 1925: 5 (Philonthus; Libya).
— Lea, 1925: 220 (Philonthus; Australian introduction).
— Portevin, 1929: 363 (Philonthus; characters; France).
— Gridelli, 1930: 74 (Philonthus; Libya).
— Bernhauer, 1931d: 587 (Philonthus; Abyssinia).
— Cameron, 1932: 127 (Philonthus; characters; Ceylon; India).
— Scheerpeltz, 1933: 1349 (Philonthus; catalog).
— Cameron, 1933d: 386 (Philonthus; Sumatra).
— Cameron, 1933e: 82 (Philonthus; Society Islands).
— Cameron, 1933f: 80 (Philonthus; Marquesas Islands).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Scheerpeltz, 1935a: 649 (Philonthus; Australia).
— Cameron, 1936: 6 (Philonthus; Austral Islands).
— C. Koch, 1936a: 75 (Philonthus; Libya).
— Wüsthoff, 1937a: 347 (Philonthus; illustration of aedeagus).
— Tottenham, 1937a: 190 (Philonthus; specimens of longicornis in Stephens' collection with type labels are not longicornis).
— Gridelli, 1939: 317 (Philonthus; Abyssinia).
— Bernhauer, 1940c: 4 (Philonthus; Azores).
— Méquignon, 1942a: 20 (Philonthus; Azores).
— Blackwelder, 1943: 410 (Philonthus; characters; Haiti; Puerto Rico; St. Kitts; Antigua; Montserrat; Grenada; Argentina; Chile).
— Cameron, 1949b: 468 (Philonthus; Japan).
— Cameron, 1950b: 214 (Philonthus; Sudan).
— V. Hansen, 1952: 67 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1953: 118 (Philonthus; Turkey).
— Coiffait, 1954: 165 (Philonthus; Canary Islands).
— Tottenham, 1955: 163 (Philonthus; subgenus Spatulonthus; notes; examined no African specimens).
— Scheerpeltz, 1956a: 484 (Philonthus; Belgian Congo).
— Scheerpeltz, 1956c: 1100 (Philonthus; Cyprus).
— Scheerpeltz, 1957: 154 (Philonthus; Tanganyika).
— Smetana, 1958: 189 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 159 (Philonthus; characters).
— Nakane, 1963a: 91 (Philonthus; characters; habitus photograph; Japan).
— Smetana, 1963b: 36 (Philonthus; subgenus Philonthus; Madeira).
— Palm, 1963b: 86 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 183 (Philonthus; subgenus Philonthus; characters; central Europe).
— Focarile, 1964: 59 (Philonthus; Italy).
— Horion, 1965: 158 (Philonthus; all Europe).
— Raianu, 1967: 268 (Philonthus; characters; ecological notes; Romania).
— Szujecki, 1968a: 730 (Philonthus; Poland).
— Smetana, 1970: 59 (Philonthus; distributional summary; Azores).
— Scheerpeltz, 1971c: 170 (Philonthus; Cameroon).
— Fagel, 1972: 103 (Spatulonthus; notes; Sainte Helene).
— Moore and Legner, 1974a: 39 (Philonthus; spread by commerce).
— Coiffait, 1974: 311 (Spatulonthus; characters; Europe; Britain; Scandinavia; Anatolia).
— Scheerpeltz, 1974: 146 (Philonthus; Basutoland; South Africa).
— Scheerpeltz, 1974c: 22 (Philonthus; Ethiopia; Sudan).
— Moore, 1975c: 106 (Philonthus; Hawaii).
— Coiffait, 1976: 217 (Spatulonthus; Tahiti).
— Pace and Zanetti, 1977: 299 (Spatulonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Marcuzzi, 1977: 12 (Philonthus; Venezuela).
— Frank, 1982: 41 (Philonthus; list of parasites).
— Coiffait, 1984e: 374 (Spatulonthus; Nepal).
— Tóth, 1984: 68 (Philonthus; characters; Hungary).
— Yuh, Paik, Kwon, and Lee, 1985: 241 (Philonthus; Korea).
— Coiffait, 1986: 394 (Spatulonthus; Saudi Arabia).
— Last, 1987: 31 (Philonthus; Papua New Guinea).
— Lucht, 1987: 106 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Borges, 1990: Table II (Spatulonthus; Azores).
— Kuschel, 1990: 26 (Philonthus; introduced, Palaearctic species; winged; habitat; New Zealand).
— Lecoq, 1990: 203 (Spatulonthus; Mascarene Islands).
— Barrion, Litsinger, and Last, 1991: 1078 (Philonthus; associated with rice agroecosystem).
— Pilon and Zanetti, 1991: 64 (Philonthus; Italy).
— J. Li, 1993: 37 (Philonthus; China).
— Smetana, 1993b: 48 (Philonthus; subgenus Spatulonthus; Czech Republic; Slovakia).
— Nishida, 1994: 73 (Philonthus; Hawaii).
— Hernández, Outerelo, and Gamarra, 1994: 188 (Spatulonthus; checklist of species; Canary Islands).
— Yadav, Chand, and Yadav, 1995: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Smetana, 1995e: 244 (Philonthus; characters; Manitoba; Nova Scotia; Ontario; Quebec; Alabama; Alaska; Arizona; Arkansas; California; Colorado; Florida; Georgia; Illinois; Indiana; Kansas; Maine; Maryland; Massachusetts; Michigan; Missouri; Nevada; New Jersey; New Mexico; New York; North Carolina; Oregon; Pennsylvania; South Carolina; Texas; Utah; Vermont; Virginia; West Virginia; Washington; Wisconsin; Mexico).
— Hu and Frank, 1995b: 584 (Philonthus; surface structure of eggs).
— Desender and Baert, 1996: 43 (Philonthus; Easter Island).
— Hu and Frank, 1997 (Philonthus; predation of eggs and larvae of Haematobia irritans; Florida).
— Nishida, 1997: 62 (Philonthus; Hawaii).
— Smithers, 1998: 16 (Philonthus; references; Norfolk Island).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
— Staniec, 1999a: 56 (Philonthus; characters of pupa; Poland).
▴ Distribution: Cosmopolitan.
scybalarius Nordmann, 1837: 94 (Philonthus; Type locality: Berolinum).
— Erichson, 1839a: 465 (Philonthus; synonym of opacus).
— P. Lucas, 1846: 112 (Philonthus; synonym of varians).
— Wollaston, 1857: 189 (Philonthus; habitat; Madeira Islands).
— Guérin-Méneville, 1857: 40 (Philonthus; synonym of varians).
— Kraatz, 1857d: 601 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 985 (Philonthus; characters; Austria).
— Kraatz, 1859: 86 (Philonthus; characters; Ceylon).
— Thomson, 1860: 164 (Philonthus; characters; Scandinavia).
— Wollaston, 1864: 571 (Philonthus; notes; Canary Islands).
— Wollaston, 1865: 492 (Philonthus; habitat; notes; Madeira Islands; Canary Islands).
— Crotch, 1867a: 383 (Philonthus; Azores).
— Wollaston, 1867: 237 (Philonthus; characters; notes; Cape Verde Islands).
— Gemminger and Harold, 1868: 590 (Philonthus; synonym of longicornis).
— Wollaston, 1871: 302 (Philonthus; synonym of longicornis).
— Fauvel, 1874: 254 [= 1874a: 480] (Philonthus; synonym of longicornis).
— Sharp, 1874: 44 (Philonthus; notes; Japan).
— Redtenbacher, 1874: 213 (Philonthus; characters; Austria).
— Seidlitz, 1875: 277 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 472 [= 1877: 328] (Philonthus; characters; France).
— Fauvel, 1877: 264 (Philonthus; synonym of longicornis).
— Sharp, 1885: 416 (Philonthus; synonym of longicornis).
— Blackburn and Sharp, 1885: 222 (Philonthus; habitat; Hawaii).
— Ganglbauer, 1895: 454 (Philonthus; synonym of longicornis).
— Fauvel, 1902b: 112 (Philonthus; synonym of longicornis).
— Fauvel, 1903: 159 (Philonthus; synonym of longicornis).
— Fauvel, 1904a: 59 (Philonthus; synonym of longicornis).
— Sharp, 1908: 547 (Philonthus; Hawaii).
— Bernhauer and Schubert, 1914: 345 (Philonthus; synonym of longicornis).
— Cameron, 1932: 127 (Philonthus; synonym of longicornis).
— Scheerpeltz, 1933: 1349 (Philonthus; synonym of longicornis).
— Blackwelder, 1943: 410 (Philonthus; synonym of longicornis).
— Smetana, 1958: 189 (Philonthus; synonym of longicornis).
— Fagel, 1972: 103 (Philonthus; synonym of longicornis).
— Coiffait, 1974: 313 (Spatulonthus; synonym of longicornis).
— Nishida, 1994: 73 (Philonthus; Hawaii).
— Smetana, 1995e: 244 (Philonthus; synonym of longicornis).
fuscicornis Nordmann, 1837: 96 (Philonthus; [preoccupied]; Type locality: Promontorium bonae spei).
— Kraatz, 1857d: 601 (Philonthus; synonym of scybalarius).
— Gemminger and Harold, 1868: 590 (Philonthus; synonym of longicornis).
— Wollaston, 1871: 302 (Philonthus; synonym of longicornis).
— Ganglbauer, 1895: 454 (Philonthus; synonym of longicornis).
— Bernhauer and Schubert, 1914: 344 (Philonthus; synonym of longicornis).
— Cameron, 1932: 127 (Philonthus; synonym of longicornis).
— Blackwelder, 1943: 410 (Philonthus; synonym of longicornis).
— Smetana, 1958: 189 (Philonthus; synonym of longicornis).
— Smetana, 1995e: 244 (Philonthus; synonym of longicornis).
promtus Erichson, 1840: 929 (Philonthus; Type locality: Pensylvania; [Note: See lectotype designation by Smetana, 1995e]).
— LeConte, 1850: 220 (Philonthus; Lake Superior).
— Fauvel, 1874: 254 [= 1874a: 480] (Philonthus; synonym of longicornis).
— Provancher, 1877: 250 (Philonthus; characters; Quebec).
— Fauvel, 1891: 115 (Philonthus; synonym of longicornis).
— Ganglbauer, 1895: 454 (Philonthus; synonym of longicornis).
— Bernhauer and Schubert, 1914: 345 (Philonthus; cited as promptus; synonym of longicornis).
— Cameron, 1932: 127 (Philonthus; cited as promptus; synonym of longicornis).
— Blackwelder, 1943: 410 (Philonthus; synonym of longicornis).
— Smetana, 1958: 189 (Philonthus; synonym of longicornis).
— Smetana, 1995e: 244 (Philonthus; lectotype designation: Pennsylvania; synonym of longicornis).
algiricus Motschulsky, 1858: 663 (Philonthus; Type locality: Algérie).
— Fauvel, 1878: 122 [= 1878a: 42] (Philonthus; Algeria).
— Fauvel, 1886: 53 [= 1886a: 45] (Philonthus; Algeria).
— Fauvel, 1891: 115 (Philonthus; synonym of longicornis).
— Fauvel, 1902b: 112 (Philonthus; synonym of longicornis).
— Fauvel, 1904a: 59 (Philonthus; synonym of longicornis).
— Bernhauer and Schubert, 1914: 344 (Philonthus; synonym of longicornis).
— Cameron, 1932: 127 (Philonthus; synonym of longicornis).
— Blackwelder, 1943: 410 (Philonthus; synonym of longicornis).
— Smetana, 1958: 189 (Philonthus; synonym of longicornis).
— Smetana, 1995e: 244 (Philonthus; synonym of longicornis).
asemus Kraatz, 1859: 86 (Philonthus; Type locality: India orientalis).
— Bernhauer and Schubert, 1914: 329 (Philonthus; catalog).
— Cameron, 1932: 127 (Philonthus; synonym of longicornis).
— Scheerpeltz, 1933: 1349 (Philonthus; synonym of longicornis).
— Blackwelder, 1943: 410 (Philonthus; synonym of longicornis).
— Smetana, 1958: 189 (Philonthus; synonym of longicornis).
— Smetana, 1995e: 244 (Philonthus; synonym of longicornis).
pedestris Walker, 1859: 51 (Philonthus; Type locality: Ceylon).
— Fauvel, 1903: 159 (Philonthus; synonym of longicornis).
— Fauvel, 1903d: 271 (Philonthus; synonym of longicornis).
— Fauvel, 1904a: 59 (Philonthus; synonym of longicornis).
— Bernhauer and Schubert, 1914: 345 (Philonthus; synonym of longicornis).
— Cameron, 1932: 127 (Philonthus; synonym of longicornis).
— Blackwelder, 1943: 410 (Philonthus; synonym of longicornis).
— Smetana, 1958: 189 (Philonthus; synonym of longicornis).
— Smetana, 1995e: 244 (Philonthus; synonym of longicornis).
fumosus Solsky, 1868a: 134 (Philonthus; Type locality: Mexique).
— Sharp, 1885: 416 (Philonthus; synonym of longicornis).
— Fauvel, 1891: 115 (Philonthus; synonym of longicornis).
— Fauvel, 1903d: 271 (Philonthus; synonym of longicornis).
— Fauvel, 1904a: 59 (Philonthus; synonym of longicornis).
— Bernhauer and Schubert, 1914: 344 (Philonthus; synonym of longicornis).
— Cameron, 1932: 127 (Philonthus; synonym of longicornis).
— Blackwelder, 1943: 410 (Philonthus; synonym of longicornis).
— Smetana, 1958: 189 (Philonthus; synonym of longicornis).
— Smetana, 1995e: 244 (Philonthus; synonym of longicornis).
laetabilis Olliff, 1887: 501 (Cafius; Type locality: Port Lincoln, South Australia; Hobart, Tasmania).
— Lea, 1899: 540 (Cafius; notes).
— Fauvel, 1903: 159 (Philonthus; synonym of longicornis).
— Fauvel, 1903d: 271 (Philonthus; synonym of longicornis).
— Fauvel, 1904a: 59 (Philonthus; synonym of longicornis).
— Bernhauer and Schubert, 1914: 345 (Philonthus; synonym of longicornis).
— Cameron, 1932: 127 (Philonthus; synonym of longicornis).
— Blackwelder, 1943: 410 (Philonthus; synonym of longicornis).
— Smetana, 1958: 189 (Philonthus; synonym of longicornis).
— Smetana, 1995e: 244 (Philonthus; synonym of longicornis).
linkei Bernhauer, 1908b: 34 (Philonthus; variety of longicornis; Type locality: Mölkau bei Leipzig).
— Bernhauer and Schubert, 1914: 345 (Philonthus; variety of longicornis).
— Scheerpeltz, 1933: 1349 (Philonthus; variety of longicornis).
— Blackwelder, 1943: 410 (Philonthus; synonym of longicornis).
— Smetana, 1958: 189 (Philonthus; aberration of longicornis).
— Smetana, 1959: 159 (Philonthus; aberration of longicornis; characters).
— Coiffait, 1967a: 432 (Philonthus; cited as valid species; characters; Russia; Siberia).
— Coiffait, 1974: 270 (Philonthus; cited as valid species; characters; Russia; Siberia).
— Coiffait, 1974: 313 (Spatulonthus; variety of longicornis).
— Smetana, 1995e: 244 (Philonthus; synonym of longicornis).
scutellatus Casey, 1915: 434 (Philonthus; [preoccupied]; Type locality: California: Paraiso Hot Springs, Monterey Co.).
— Scheerpeltz, 1933: 1360 (Philonthus; catalog).
— Smetana, 1995e: 244 (Philonthus; synonym of longicornis).
cervicalis Casey, 1915: 437 (Philonthus; Type locality: New York: Ithaca).
— Scheerpeltz, 1933: 1336 (Philonthus; catalog).
— Smetana, 1995e: 244 (Philonthus; synonym of longicornis).
— Downie and Arnett, 1996: 378 (Philonthus; characters; USA).
piceicornis Gridelli, 1920a: 18, 19, 20 (Philonthus; aberration of longicornis; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied by picicornis Horn according to Article 58.2]; cited from Mte. Rima, Giudicarie, Trentino; Liguria, Genova; Salonicco, Macedonia).
— Donisthorpe, 1930: 96 (Philonthus; aberration of longicornis; characters; Britain).
— Scheerpeltz, 1933: 1349 (Philonthus; aberration of longicornis).
— Blackwelder, 1943: 410 (Philonthus; synonym of longicornis).
— Smetana, 1958: 189 (Philonthus; aberration of longicornis).
— Smetana, 1959: 159 (Philonthus; aberration of longicornis; characters).
— Coiffait, 1974: 313 (Spatulonthus; variety of longicornis).
— Pope, 1977: 29 (Philonthus; synonym of longicornis).
— Smetana, 1995e: 244 (Philonthus; synonym of longicornis).
bestialis Bernhauer, 1939a: 86 (Philonthus; variety of longicornis; Type locality: British Ostafrika: M. Kenya, Westhang, Waldzone, Podocarpus, 2400 m).
— Tottenham, 1955: 158 (Philonthus; subgenus Spatulonthus; notes; Kenya).
— Tottenham, 1962: 139 (Philonthus; subgenus Spatulonthus; Tanganyika).
longicornis Waltl, 1835, see: ebeninus Gravenhorst, 1802.
longipalpis Kashcheev, 1999a: 58 (Philonthus; Type locality: Altai, r. Bukhtarma, okr. pos. Archaty).
longipennis Provancher, 1877, see: Bisnius.
longipes Sharp, 1876, see: Paederomimus.
longiventris Casey, 1915, see: Rabigus.
longulus Kraatz, 1859, see: Neobisnius.
lopezi Hatch, 1957, see: Gabrius.
lualabaensis Levasseur, 1966b: 1494 (Philonthus; Type locality: Kolwezi, Katanga).
lubomiri Herman, 2001: 47 (Philonthus; replacement name for simonae Hromádka).
simonae Hromádka, 1992: 99 (Philonthus; [preoccupied]; Type locality: Nepal, Manana).
lucanicornis Bernhauer, 1908c: 111 (Philonthus; Type locality: Gabun).
— Bernhauer and Schubert, 1914: 345 (Philonthus; catalog).
— Tottenham, 1949b: 361 (Philonthus; characters).
▴ Distribution: Gabon.
lucens Mannerheim, 1830, see: atratus Gravenhorst, 1802.
lucidanus Blackwelder, 1944: 134 (Philonthus; replacement name for lucidus Fauvel).
lucidus Fauvel, 1891: 116 (Philonthus; [preoccupied]; Type locality: Colonie Tovar; Nouvelle Grenada).
— Bernhauer and Schubert, 1914: 345 (Philonthus; catalog).
— Blackwelder, 1944: 134 (Philonthus; synonym of lucidanus).
lucidus Fauvel, 1891, see: lucidanus Blackwelder, 1944.
lucidus Gravenhorst, 1802, see: debilis Gravenhorst, 1802.
lucilius Sharp, 1885: 415 (Philonthus; Type locality: Panama, Volcan de Chiriqui 2000 to 3000 feet).
ludicrus Erichson, 1840, see: Neobisnius.
luenaensis Levasseur, 1962a: 58 (Philonthus; Type locality: Luena, Haut Lomami, Katanga).
lulengae Bernhauer, 1932b: 151 (Philonthus; Type locality: Kivu: Lulenga).
— Scheerpeltz, 1933: 1349 (Philonthus; catalog).
— Bernhauer and Chapman, 1939: 70 (Philonthus; cited as lulangae; Kenya).
▴ Distribution: Congo.
luluanus Bernhauer, 1935d: 102 (Philonthus; Type locality: Lulua: Kapanga).
luridus Runde, 1835, see: micans Gravenhorst, 1802.
lustralis Erichson, 1840, see: Paederomimus.
lustrator Sharp, 1876, see: Paederomimus.
luteocinctus Scheerpeltz, 1965: 222 (Philonthus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).
luxurians Erichson, 1840: 491 (Philonthus; Type locality: Sardinia).
— Küster, 1854: 15 (Philonthus; characters; Sardinia).
— Kraatz, 1857d: 610 (Philonthus).
— Mulsant and Rey, 1876: 514 [= 1877: 370] (Philonthus; characters; France).
— Sainte-Claire Deville, 1906: 98 (Philonthus; Corsica).
— Bernhauer and Schubert, 1914: 345 (Philonthus; catalog).
— Scheerpeltz, 1933: 1349 (Philonthus; catalog).
— Jarrige, 1951a: 7 (Philonthus; characters).
— Coiffait, 1967a: 427 (Philonthus; characters; Corsica).
— Coiffait, 1974: 271 (Philonthus; characters; Corsica; Sardinia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Marchal and Chardonnet, 2000: 72 (Philonthus; Corsica).
▴ Distribution: France, Italy.
lynchi Bernhauer and Schubert, 1914: 345 (Philonthus; replacement name for affinis Lynch).
affinis Lynch, 1884: 160 (Philonthus; [preoccupied]; Type locality: Baradero).
macellus Fauvel, 1877: 264 (Philonthus; Type locality: Australie, Victoria).
— Fauvel, 1878g: 547 (Philonthus; Australia).
— Blackburn, 1896: 262 (Philonthus; Australia).
— Bernhauer and Schubert, 1914: 345 (Philonthus; catalog).
— Last, 1987: 31 (Philonthus; Papua New Guinea).
▴ Distribution: Australia, New Guinea.
macies Sharp, 1874, see: Bisnius.
macilentus Sharp, 1885: 423 (Philonthus; Type locality: Mexico, Cordova; also with a variety from Mexico: Las Vigas, Jalapa, Cerro de Plumas).
mcnamarae Cameron, 1931: 362 (Philonthus; Type locality: New Guinea: Mount Lamington; Finsch Haven).
— Scheerpeltz, 1933: 1350 (Philonthus; catalog).
— Last, 1968: 372 (Philonthus; cited as macnamarae; New Guinea).
— Last 1981: 124 (Philonthus; cited as macnamarae; New Guinea).
— Last, 1987: 32 (Philonthus; cited as macmarae; Papua New Guinea; New Britain).
▴ Distribution: New Guinea, New Britain.
macrocephalus Sharp, 1889: 42 (Philonthus; Type locality: Ichiuchi; Yuyama).
— Bernhauer and Schubert, 1914: 345 (Philonthus; catalog).
— Scheerpeltz, 1933: 1350 (Philonthus; catalog).
▴ Distribution: Japan.
maculatus Cameron, 1920a: 217 (Philonthus; Type locality: Mysore, alt. 4400 feet).
— Cameron, 1932: 130 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1350 (Philonthus; catalog).
▴ Distribution: India.
maculicollis Cameron, 1937: 10 (Philonthus; Type locality: Java: G. Tangkoeban Prahoe).
maculicornis Stephens, 1832, see: cognatus Stephens, 1832.
maculipennis Fauvel, 1903e: 242 (Philonthus; Type locality: Cameroun; Gabon: Loango).
— Bernhauer and Schubert, 1914: 345 (Philonthus; catalog).
— Bernhauer, 1942c: 361 (Philonthus; Cameroon).
— Tottenham, 1949b: 348 (Philonthus; characters).
— Levasseur, 1968a: 53 (Philonthus; Central African Republic).
— Levasseur, 1971: 299 (Philonthus; Sierra Leone; Liberia; Central African Republic).
▴ Distribution: Sierra Leone, Liberia, Cameroon, Central African Republic, Gabon.
madangensis Last, 1981: 124 (Neobisnius; Type locality: New Guinea: Musa).
madianus Bernhauer, 1936f: 263 (Philonthus; Type locality: Angola: Ebanga. Uganda: Madi).
— Tottenham, 1949b: 304 (Philonthus; subgenus Raucalius; similar to rugulipennis).
▴ Distribution: Angola, Uganda.
madurensis Bernhauer, 1915i: 255 (Philonthus; Type locality: Südliches Indien: Madura, Chambaganoor).
— Bernhauer, 1922c: 232 (Philonthus; Formosa).
— Cameron, 1932: 121 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1350 (Philonthus; catalog).
▴ Distribution: India, Taiwan.
mafuluanus Cameron, 1937a: 102 (Philonthus; Type locality: New Guinea: Mafulu, alt. 400 ft.).
magniceps Cameron, 1948, see: Gabrius.
magnicollis Bernhauer, 1915, see: Gabrius.
magnificus Bernhauer, 1933, see: purpuripennis Reitter, 1887.
mahagianus Cameron, 1933: 42 (Philonthus; Type locality: Belgian Congo: Mahagi-Port).
maindroni Fauvel, 1903: 158 (Philonthus; Type locality: Coonoor).
— Bernhauer and Schubert, 1914: 345 (Philonthus; catalog).
— Cameron, 1932: 88 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1350 (Philonthus; catalog).
— Scheerpeltz, 1965: 106 (Philonthus; Burma).
— Hayashi, 1997b: 38 (Philonthus; Taiwan).
▴ Distribution: India, Myanmar, Taiwan.
annandalei Cameron, 1924a: 118 (Philonthus; Type locality: Dehra Dun).
— Cameron, 1932: 88 (Philonthus; synonym of maindroni).
— Scheerpeltz, 1933: 1350 (Philonthus; synonym of maindroni).
— Jeannel and Jarrige, 1949: 374 (Philonthus; cave species; Assam).
malaisei Scheerpeltz, 1965: 203 (Philonthus; Type locality: N.E. Burma, Kambaiti, 7000 ft).
malcolmi Herman, 2001: 47 (Philonthus; replacement name for cameroni Pajni and Kohli).
cameroni Pajni and Kohli, 1977: 513 (Philonthus; [preoccupied]; Type locality: India: Chandigarh, Sector 35).
malheurensis Hatch, 1957, see: Erichsonius.
malkini Hatch, 1957, see: Gabrius.
malleus Tottenham, 1962: 169 (Philonthus; Type locality: Uganda: Ruwenzori Range, Namwamba Valley).
manaiaensis Broun, 1910, see: novaezeelandiae Duvivier, 1883.
mandibularis Kirby, 1837, see: politus Linné, 1758.
mandschuricus Bernhauer, 1914, see: Gabrius.
mannerheimi Fauvel, 1869: 488 (Philonthus; proposed for the lucens misidentified by Erichson; Type locality: Not cited).
— Erichson, 1839a: 452 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 345 cited lucens Erichson as a synonym of mannerheimi Fauvel; Erichson did not describe the species as new, he attributed it to Mannerheim]; characters; Germany).
— Erichson, 1840: 443 (Philonthus; [Note: See preceding citation; cited as lucens, but Erichson's use of the name was a misidentification]; characters; Germany).
— Fauvel, 1874: 247 [= 1874a: 473] (Philonthus; characters; France; Sicily; Sardinia).
— Fauvel, 1876a: 74 [= 1876: 259] (Philonthus; Spain).
— Mulsant and Rey, 1876: 396 [= 1877: 252] (Philonthus; [Note: Cited as lucens Erichson, who misidentified the species, and with mannerheimi Fauvel as a synonym]; characters; France).
— Fowler, 1888: 265 (Philonthus; cited as lucens Erichson; characters; Britain).
— Ganglbauer, 1895: 447 (Philonthus; characters; Europe).
— Reitter, 1909: 127 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 345 (Philonthus; catalog).
— Portevin, 1929: 360 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1350 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Aleksandrov, 1934: 153 (Philonthus; China).
— V. Hansen, 1952: 55 (Philonthus; subgenus Philonthus; characters; Denmark).
— Palm, 1953: 12 (Philonthus; notes; Sweden).
— Smetana, 1958: 183 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 150 (Philonthus; characters).
— Palm, 1963b: 72 (Philonthus; characters; Sweden; Denmark).
— Lohse, 1964: 178 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 140 (Philonthus; Ireland; Britain; Denmark; Sweden; France; Czechoslovakia; Poland; Hungary; Romania; Ukraine; Spain; Italy; Bosnia; Macedonia; Germany; Austria).
— Coiffait, 1967a: 398 (Philonthus; characters; Europe).
— Raianu, 1968a: 466 (Philonthus; Romania).
— Szujecki, 1968a: 729 (Philonthus; Poland).
— Last, 1974 (Philonthus; comparisons to related species; characters; England).
— Coiffait, 1974: 232 (Philonthus; characters; Europe; Britain; Sweden).
— Shilov, 1975a: 375 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Pope, 1977: 29 (Philonthus; Britain).
— Raichev, 1979: 82 (Philonthus; Bulgaria).
— Frank, 1982: 42 (Philonthus; list of parasites).
— Tóth, 1984: 53 (Philonthus; characters; Hungary).
— Schülke and Uhlig, 1988: 11 (Philonthus; Germany).
— Pilon and Zanetti, 1991: 62 (Philonthus; Italy).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Luff, 1996: 30 (Philonthus; collecting notes; Isle of Man).
— Schillhammer, 2000: 136 (Philonthus; characters; West Palaearctic).
▴ Distribution: Europe, China.
manyemae Bernhauer, 1932b: 148 (Philonthus; subgenus Gabrius; Type locality: Manyema, r. Kasa-Niemba; Katanga: Katompe; Mulongo, Niunzu).
— Scheerpeltz, 1933: 1350 (Philonthus; catalog).
— Tottenham, 1949b: 344 (Philonthus; characters; should not be in Gabrius).
▴ Distribution: Congo.
marakwetensis Bernhauer, 1939, see: Gabrius.
marcelli† Heer, 1856: 17 (Philonthus; Type locality: Aix, Provence; Fossil).
— Oustalet, 1874: 170 (Philonthus; characters; France).
— Scudder, 1891: 566 (Philonthus; catalog; Ligurian; France).
— Handlirsch, 1907: 725 (Philonthus; catalog; Oligocene; France).
▴ Distribution: France.
marcescens Tottenham, 1949b: 342 (Philonthus; Type locality: Jebel Haina, Sudan).
sudanensis Cameron, 1952d: 824 (Philonthus; Type locality: Mauritanie: Atar).
marcidulus† Scudder, 1900: 54 (Philonthus; fossil; Type locality: Florissant, Colorado).
— Handlirsch, 1907: 725 (Philonthus; catalog; Miocene; Colorado).
— Wickham, 1913: 288 (Philonthus; fossil; doubtful identification).
▴ Distribution: USA.
marcidus Wollaston, 1864: 571 (Philonthus; Type locality: Lanzarote; Fuerteventura; Teneriffe; Hierro; Gomera; Palma).
— Wollaston, 1865: 492 (Philonthus; habitat; notes; Canary Islands).
— Wollaston, 1867: 279 (Philonthus; synonym of concinnus).
— Fauvel, 1869: 488 (Philonthus; synonym of concinnus).
— Wollaston, 1871: 303 (Philonthus; synonym of concinnus).
— Fauvel, 1874: 234 [= 1874a: 459] (Philonthus; synonym of ebeninus).
— Ganglbauer, 1895: 452 (Philonthus; synonym of concinnus).
— Fauvel, 1902b: 110 (Philonthus; synonym of ebeninus).
— Bernhauer and Schubert, 1914: 333 (Philonthus; synonym of concinnus).
— Coiffait, 1954: 165 (Philonthus; characters; valid species; Canary Islands).
— Smetana, 1958: 175 (Philonthus; synonym of concinnus).
— Coiffait, 1967a: 417 (Philonthus; characters; Canaries).
— Coiffait, 1974: 260 (Philonthus; characters; Canary Islands).
▴ Distribution: Canary Islands.
mareki Coiffait, 1967a: 449 (Philonthus; subgenus Kenonthus; Type locality: Belan Tatry, Slovaquie septentrionale).
— Coiffait, 1974: 302 (Philonthus; subgenus Kenonthus; characters; Slovakia).
— Tóth, 1984: 46 (Philonthus; characters).
— Lohse, 1988: 42 (Philonthus; characters).
— Lohse and Lucht, 1989: 163 (Philonthus; characters).
— Smetana, 1993b: 48 (Philonthus; subgenus Kenonthus; Slovakia).
— Burakowski, Mroczkowski, and Stefańska, 2000: 51 (Philonthus; subgenus Kenonthus; catalog; Poland).
▴ Distribution: Slovakia.
marginatus O. Müller, 1764: 23 (Staphylinus; [Note: This name was described as new in Staphylinus separately by O. Müller, 1764, Ström, 1768, and Fabricius, 1775. Goeze, 1777, listed the three citations as referring to the same species. Fabricius, 1781, cited his earlier listing and that of Ström as referring to the same species. Erichson, 1840 cited Fabricius as the author and the use by O. Müller and Ström as subsequent citations. From Gravenhorst in 1802 until 1895 most authors cited Fabricius as the author of marginatus; from Ganglbauer, 1895, to present most authors cited Ström as author]; Type locality: Not cited).
— O. Müller, 1776: 97 (Staphylinus; characters).
— Goeze, 1777: 726 (Staphylinus; [Note: Cited references to marginatus by Ström, 1768, and Fabricius, 1775, as referring to marginatus O. Müller]; characters).
— Villers, 1789: 419 (Staphylinus; [Note: Both the O. Müller and Ström references to marginatus cited as referring to the same species]; characters; France).
— Fauvel, 1874: 252 [= 1874a: 478] (Philonthus; attributed to Müller; characters; France; Italy).
— J. Sahlberg, 1876: 20 (Philonthus; attributed to Müller; Finland).
— Johansen, 1914: 398 (Philonthus; attributed to Müller; characters; Denmark).
— Herman, 2001: 27 (Philonthus; valid name for species).
▴ Distribution: Europe, Russia, Kazakhstan, Turkmenistan, China.
marginatus Ström, 1768: 332 [= 1770: 281] (Staphylinus; [preoccupied]; Type locality: Norske).
— Gmelin, 1790: 2028 (Staphylinus; cited marginatus of Fabricius and Ström as referring to the same species; characters).
— Fabricius, 1793: 526 (Staphylinus; species attributed to Ström; characters; England, Norway).
— Olivier, 1795:(42): 26 (Staphylinus; species attributed to Ström; characters; France, Sweden, England).
— Fabricius, 1801: 597 (Staphylinus; attributed to Ström; characters).
— Jacobsen, 1849: 691 (Philonthus; characters; Denmark).
— Ganglbauer, 1895: 448 (Philonthus; attributed to Ström; characters; north and middle Europe).
— Poppius, 1905: 10 (Philonthus; Orkney Islands; Shetland Islands).
— Reitter, 1909: 127 (Philonthus; attributed to Ström; characters; Germany).
— Poppius, 1913: 10 (Philonthus; attributed to Linné; Faroes).
— Bernhauer and Schubert, 1914: 345 (Philonthus; attributed to Ström; catalog).
— Portevin, 1929: 360 (Philonthus; attributed to Ström; characters; France).
— Scheerpeltz, 1933: 1350 (Philonthus; attributed to Ström; catalog).
— Wüsthoff, 1934: 61 (Philonthus; attributed to Ström; illustration of aedeagus).
— West, 1937: 39 (Philonthus; attributed to Ström; Faroe Islands).
— V. Hansen, 1952: 56 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1958: 226 (Philonthus; subgenus Onychophilonthus; attributed to Ström; characters; ecology; Czechoslovakia).
— Smetana, 1959: 172 (Philonthus; subgenus Onychophilonthus; characters).
— Smetana, 1963d: 404 (Philonthus; subgenus Onychophilonthus; characters; Europe).
— Palm, 1963b: 73 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 186 (Philonthus; subgenus Onychophilonthus; characters; central Europe).
— Horion, 1965: 177 (Philonthus; subgenus Onychophilonthus; Russia; Lapland; Northern Cape; Scotland; France; Italy; Romania; Hungary; Czechoslovakia; Poland; Ukraine; Germany; Austria).
— Coiffait, 1967a: 437 (Philonthus; subgenus Onychophilonthus; characters; Europe).
— Raianu, 1968a: 468 (Philonthus; Romania).
— Szujecki, 1968a: 731 (Philonthus; Poland).
— Kasule, 1970: 65 (Philonthus; larval characters).
— Coiffait, 1974: 305 (Philonthus; subgenus Onychophilonthus; attributed to Ström; characters; Europe; Britain; Scandinavia; Siberia).
— Shilov, 1975a: 376 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Pope, 1977: 29 (Philonthus; attributed to Ström; Britain).
— Raichev, 1979: 83 (Philonthus; Bulgaria).
— Hinton, 1981: 660 (Philonthus; characters of egg).
— Waterston, Owen, Welch, Bacchus and Hammond, 1981: 274 (Philonthus; Outer Hebrides).
— Frank, 1982: 42 (Philonthus; list of parasites).
— Boller, 1983: 120 (Philonthus; larval characters).
— Tóth, 1984: 43 (Philonthus; characters; Hungary).
— Lucht, 1987: 106 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Pilon and Zanetti, 1991: 60 (Philonthus; Italy).
— Welch, 1993: 228 (Philonthus; ovariole number and ovary structure).
— J. Li, 1993: 36 (Philonthus; China).
— Smetana, 1993b: 48 (Philonthus; subgenus Onychophilonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Kashcheev, 1999a: 58 (Philonthus; Kazakhstan).
— Herman, 2001: 27 (Philonthus; synonym of marginatus O. Müller).
marginatus Fabricius, 1775: 266 (Staphylinus; [preoccupied]; Type locality: Anglia).
— Fabricius, 1781: 336 (Staphylinus; Fabricius cited both his 1775 reference and Ström's article as referring to the same species; characters; England, Norway).
— Fabricius, 1787: 221 (Staphylinus; cited without attribution]; characters).
— Paykull, 1789: 44 (Staphylinus; attributed Fabricius, 1787]; redescription; Sweden).
— Panzer, 1795: 355 (Staphylinus; species attributed to Fabricius, 1793]; characters; Germany).
— Paykull, 1800: 392 (Staphylinus; attributed to Fabricius; characters; Sweden).
— Turton, 1802: 512 (Staphylinus; characters; Europe).
— Gravenhorst, 1802: 24 (Staphylinus; attributed to Fabricius; characters).
— Marsham, 1802: 512 (Staphylinus; attributed to Fabricius with an article by O. Müller as a subsequent citation]; characters; Britain).
— Latreille, 1804: 313 (Staphylinus; attributed to Fabricius; characters; France; England; Sweden).
— Gravenhorst, 1806: 71 (Staphylinus; attributed to Fabricius; France).
— Gyllenhal, 1810: 322 (Staphylinus; attributed to Fabricius; characters; Sweden).
— Zetterstedt, 1828: 75 (Staphylinus; attributed to Fabricius; characters; Lapland).
— Curtis, 1829: 25 (Philonthus; attributed to Fabricius; catalog; Britain).
— C. Sahlberg, 1830: 322 (Staphylinus; attributed to Paykull who attributed it to Fabricius; characters; Finland).
— Mannerheim, 1830: 28 (Staphylinus; attributed to Paykull who attributed it to Fabricius; Finland; Sweden; Lapland).
— Mannerheim, 1831: 442 (Staphylinus; attributed to Paykull who attributed it to Fabricius; Finland; Sweden; Lapland).
— Stephens, 1832: 233 (Philonthus; attributed to Fabricius; characters; England).
— Runde, 1835: 7 (Staphylinus; attributed to Paykull who attributed it to Fabricius; characters; Germany).
— Lacordaire, 1835: 397 (Staphylinus; attributed to Fabricius; characters; France).
— Curtis, 1836: pl. 610 (Philonthus; attributed to Fabricius; characters; Britain).
— Nordmann, 1837: 84 (Philonthus; attributed to Fabricius; Sweden; Germany; France).
— Zetterstedt, 1838: 64 (Staphylinus; attributed to Fabricius; characters; habitat; Lapland).
— Erichson, 1839a: 453 (Philonthus; attributed to Fabricius; characters; Germany).
— Heer, 1839: 261 (Philonthus; attributed to Fabricius; characters; Switzerland).
— Stephens, 1839: 396 (Philonthus; attributed to Fabricius; characters; England).
— Erichson, 1840: 444 (Philonthus; attributed to Fabricius and cited descriptions by O. Müller and Ström as subsequent references; characters; Europe).
— Laporte, 1840: 178 (Staphylinus; attributed to Fabricius; characters; England; France).
— Ménétriés, 1849: 44 (Philonthus; attributed to Fabricius; Turcomanie).
— Redtenbacher, 1849: 702 (Philonthus; attributed to Fabricius; characters; Austria).
— Küster, 1853a: 53 (Philonthus; attributed to Fabricius; characters; Europe).
— Hardy, 1851: 26 (Philonthus; attributed to Fabricius; notes; England).
— Fairmaire and Laboulbène, 1856: 515 (Philonthus; attributed to Fabricius; characters; France).
— Kraatz, 1857d: 587 (Philonthus; attributed to Fabricius; characters; Germany).
— Redtenbacher, 1857: 192 (Philonthus; attributed to Fabricius; characters; Austria).
— Thomson, 1860: 165 (Philonthus; attributed to Fabricius; characters; Scandinavia).
— Redtenbacher, 1874: 211 (Philonthus; attributed to Fabricius; characters; Austria).
— Seidlitz, 1875: 275 (Philonthus; attributed to Fabricius; characters; Baltic region).
— Mulsant and Rey, 1876: 410 [= 1877: 266] (Philonthus; attributed to Fabricius; characters; France).
— Fowler, 1888: 266 (Philonthus; attributed to Fabricius; characters; Britain).
— Herman, 2001: 27 (Philonthus; synonym of marginatus O. Müller).
rubromarginatus Gabriel, 1909: 416 (Philonthus; aberration of marginatus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]).
— Bernhauer and Schubert, 1914: 345 (Philonthus; synonym of marginatus).
— Smetana, 1958: 226 (Philonthus; synonym of marginatus).
— Smetana, 1959: 172 (Philonthus; subgenus Onychophilonthus; aberration of marginatus; characters).
— Smetana, 1963d: 404 (Philonthus; synonym of marginatus).
— Coiffait, 1974: 305 (Philonthus; variety of marginatus; characters).
marginatus Ström, 1768, see: marginatus O. Müller, 1764.
marginatus Fabricius, 1775, see: marginatus O. Müller, 1764.
marginellus Schubert, 1902, see: xanthoraphis Eppelsheim, 1895.
marginipennis Wollaston, 1867: 236 (Philonthus; Type locality: Cape Verde Islands: S. Vicente).
— Bernhauer and Schubert, 1914: 345 (Philonthus; catalog).
— Tottenham, 1949b: 328 (Philonthus; characters).
— Cameron, 1950b: 214 (Philonthus; Sudan).
— Cameron, 1952e: 831 (Philonthus; Dahomey; Senegal).
— Coiffait, 1967a: 401 (Philonthus; characters; Cape Verde Islands; intertropical Africa).
— Coiffait, 1974: 215 (Philonthus; characters; Cape Verde Islands; western Africa; Sahara).
— Coiffait, 1979a: 162 (Philonthus; Saudi Arabia).
▴ Distribution: Cape Verde Islands, Senegal, Benin, Sudan, Saudi Arabia.
maridadus Tottenham, 1962: 223 (Philonthus; Type locality: Kenya: Nairobi. Also cited from Tanganyika).
marojejy Lecoq, 1998: 67 (Philonthus; subgenus Piezarthrus; Type locality: Madagascar Est: massif du Marojejy).
marphensis Coiffait, 1982: 68 (Philonthus; Type locality: Nepal: Thakkhola, Alt Marpha, 3200 m).
marshalli Cameron, 1951, see: Gabrius.
marshalli Tottenham, 1949b: 335 (Philonthus; Type locality: Natal: Frere).
martialis Cameron, 1932, see: Gabrius.
maryensis Last, 1987: 26 (Philonthus; Type locality: Papua New Guinea: New Guinea NE: Mount St. Mary, 1900–3000 m).
maskinius Tottenham, 1955: 164 (Philonthus; subgenus Spatulonthus; Type locality: Abyssinia: Mt. Chillalo).
masuriensis Cameron, 1926, see: christiei Bernhauer, 1920.
mateui Levasseur, 1962: 233 (Philonthus; Type locality: Rgueibat Temba, Mauritanie).
matutinalis Faldermann, 1835, see: laminatus Creutzer, 1799.
mauretanicus Peyerimhoff, 1937, see: Gabrius.
mediocris Cameron, 1943, see: Gabrius.
megacephalus Heer, 1839, see: Bisnius.
megatlasicus C. Koch, 1941, see: atratus Gravenhorst, 1802.
meges Tottenham, 1949b: 332 (Philonthus; Type locality: Abyssinia: Mt. Zuquala, circa 9000 feet).
megophthalmus Cameron, 1948, see: Gabrius.
malaccanus Cameron, 1950d: 19 (Philonthus; Type locality: Singapore).
melampus Smetana, 1991a: 236 (Philonthus; Type locality: Mexico: Chiapas: Mt. Tzontehuitz, Chis., 9500 ft.).
melanarius Mulsant and Rey, 1876, see: concinnus Gravenhorst, 1802.
melancholicus Sharp, 1885: 425 (Philonthus; Type locality: Mexico, Capulalpam).
melanocephalus Heer, 1839, see: debilis Gravenhorst, 1802.
melanocephalus Hochhuth, 1851, see: Erichsonius.
melanopterus Stephens, 1832, see: cognatus Stephens, 1832.
melanopus Sharp, 1885: 405 (Philonthus; Type locality: Mexico: Jalapa; Costa Rica: Volcan de Irazu 6000 to 7000 feet; Panama: Volcan de Chiriqui, between 2500 and 4000 feet).
menai Last, 1987: 26 (Philonthus; Type locality: Papua New Guinea: New Guinea NE: Mt. St. Mary, 1900–3000 m).
mendax Runde, 1835, see: rubripennis Stephens, 1832.
mendosus Bernhauer, 1928, see: Gabrius.
menetriesi Kirshenblat, 1933, see: carbonarius Gravenhorst, 1802.
mequignoni Jarrige, 1938, see: rectangulus Sharp, 1874.
mercurii Tikhomirova, 1973: 169 (Philonthus; Type locality: Suputinskii zapovednik).
meridioafricanus Scheerpeltz, 1974: 158 (Philonthus; Type locality: South West Africa: Kaokoveld, Anabib, Orupembe, 100 miles W Ohopoho).
meridionalis Cameron, 1945, see: Gabronthus.
merops Smetana, 1963d: 407 (Philonthus; subgenus Onychophilonthus; Type locality: Mongolia: See Kulun).
— Coiffait, 1967a: 439 (Philonthus; subgenus Onychophilonthus; characters; Mongolia).
— Coiffait, 1974: 308 (Philonthus; subgenus Onychophilonthus; characters; Mongolia).
— Shilov, 1976: 153 (Philonthus; Mongolia).
— Shibata, 1991b: 256 (Philonthus; characters; Taiwan).
▴ Distribution: Mongolia, Taiwan.
merorientis Smetana, 1995e: 408 (Philonthus; Type locality: Florida: L. Istokpoga n. L. Placid. Also cited from Alabama, Arkansas, Florida, Georgia, Louisiana, Mississippi, South Carolina, Texas).
metallicus Gravenhorst, 1802, see: atratus Gravenhorst, 1802.
metallifer Bernhauer, 1916c: 31 (Philonthus; Type locality: Bolivien und Peru).
methneri Bernhauer, 1915d: 139 (Philonthus; Type locality: Deutsch-Ostafrika: West-Kilimandscharo, 1500–1700 m).
— Scheerpeltz, 1933: 1350 (Philonthus; catalog).
— Tottenham, 1949b: 308 (Philonthus; notes).
— Tottenham, 1962: 135, 162, 163, 166 (Philonthus; characters; Tanganyika; Kenya).
▴ Distribution: Kenya, Tanzania.
mexicanus Sharp, 1876, see: piceatus Nordmann, 1837.
micans Gravenhorst, 1802: 25 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).
— Latreille, 1804: 314 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 76 (Staphylinus; characters).
— Gyllenhal, 1810: 344 (Staphylinus; characters; Sweden).
— Zetterstedt, 1828: 78 (Staphylinus; characters; Lapland).
— Curtis, 1829: 26 (Philonthus; catalog; Britain).
— Mannerheim, 1830: 31 (Staphylinus; Finland; Sweden).
— Mannerheim, 1831: 445 (Staphylinus; Finland; Sweden).
— Stephens, 1832: 240 (Philonthus; characters; England).
— C. Sahlberg, 1830: 329 (Staphylinus; characters; Finland).
— Lacordaire, 1835: 406 (Staphylinus; characters; France).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Nordmann, 1837: 102 (Philonthus; Europe).
— Zetterstedt, 1838: 65 (Staphylinus; characters; habitat; Lapland).
— Erichson, 1839a: 473 (Philonthus; characters; Germany).
— Heer, 1839: 270 (Philonthus; characters; Switzerland).
— Stephens, 1839: 398 (Philonthus; characters; England).
— Erichson, 1840: 484 (Philonthus; characters; Europe).
— Laporte, 1840: 178 (Staphylinus; characters; France; Sweden).
— Kiesenwetter, 1848a: 21 (Philonthus; characters; Europe).
— Jacobsen, 1849: 703 (Philonthus; characters; Denmark).
— Redtenbacher, 1849: 706 (Philonthus; characters; Austria).
— Hardy, 1851: 30 (Philonthus; notes; England).
— Fairmaire and Laboulbène, 1856: 528 (Philonthus; characters; France).
— Kraatz, 1857d: 612 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 196 (Philonthus; characters; Austria).
— Peyron, 1858: 426 (Philonthus; Turkey).
— Thomson, 1860: 165 (Philonthus; characters; Scandinavia).
— Redtenbacher, 1867: 29 (Philonthus; Java [Note: This may be a misidentification (Schillhammer, personal communication)]).
— Fauvel, 1874: 258 [= 1874a: 484] (Philonthus; characters; France; Turkey; North America).
— Redtenbacher, 1874: 215 (Philonthus; characters; Austria).
— Solsky, 1874: 180 (Philonthus; characters; Turkestan).
— Seidlitz, 1875: 278 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 503 [= 1877: 359] (Philonthus; characters; France).
— J. Sahlberg, 1876: 21 (Philonthus; Finland).
— Horn, 1884: 204 (Philonthus; characters; eastern United States from Massachusetts to Michigan).
— Fowler, 1888: 277 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 457 (Philonthus; characters; Palaearctic region; North America).
— Fauvel, 1902b: 113 (Philonthus; Morocco; Algeria).
— Montandon, 1908: 80 (Philonthus; Romania).
— Reitter, 1909: 130 (Philonthus; characters; Germany).
— Blatchley, 1910: 385 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 346 (Philonthus; catalog).
— Johansen, 1914: 412 (Philonthus; characters; Denmark).
— Portevin, 1929: 364 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1350 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Wüsthoff, 1937a: 347 (Philonthus; illustration of aedeagus).
— Jarrige, 1951a: 7, 9 (Philonthus; characters).
— V. Hansen, 1952: 76 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1958: 220 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 166 (Philonthus; characters).
— Coiffait, 1963: 12 (Paragabrius).
— Palm, 1963b: 94 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 185 (Philonthus; subgenus Philonthus; characters; central Europe).
— Focarile, 1964: 59 (Philonthus; Italy).
— Horion, 1965: 169 (Philonthus; all Europe).
— Raianu, 1968a: 468 (Philonthus; Romania).
— Szujecki, 1968a: 731 (Philonthus; Poland).
— Coiffait, 1974: 94 (Paragabrius; characters; Holarctic).
— Shilov, 1975a: 376 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Pace and Zanetti, 1977: 303 (Paragabrius; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Frank, 1982: 42 (Philonthus; list of parasites).
— Tóth, 1984: 77 (Philonthus; characters; Hungary).
— Lucht, 1987: 106 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Smetana, 1993b: 48 (Philonthus; subgenus Paragabrius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Downie and Arnett, 1996: 380 (Philonthus; characters; USA).
— Owen, 1997a: 149 (Philonthus; collecting sites; Ireland).
— Kashcheev, 1999a: 58 (Philonthus; Kazakhstan).
▴ Distribution: Algeria, Morocco, Europe, Turkey, Kazakhstan.
obscuripennis Stephens, 1832: 236 (Philonthus; Type locality: London; Suffolk; Norfolk; Devonshire).
— Curtis, 1829: 25 (Philonthus; nomen nudum; catalog; Britain).
— Stephens, 1839: 397 (Philonthus; characters; England).
— Hardy, 1851: 29 (Philonthus; synonym of varians).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of micans).
— Bernhauer and Schubert, 1914: 346 (Philonthus; synonym of micans).
— Jarrige, 1951a: 9 (Philonthus; synonym of micans).
— Smetana, 1958: 220 (Philonthus; synonym of micans).
— Coiffait, 1974: 94 (Paragabrius; synonym of micans).
luridus Runde, 1835: 9 (Staphylinus; Type locality: Halae).
— Erichson, 1839a: 473 (Philonthus; synonym of micans).
— Erichson, 1840: 484 (Philonthus; synonym of micans).
— Kraatz, 1857d: 612 (Philonthus; cited as lucidus; synonym of micans).
— Ganglbauer, 1895: 457 (Philonthus; synonym of micans).
— Bernhauer and Schubert, 1914: 346 (Philonthus; synonym of micans).
— Coiffait, 1974: 94 (Paragabrius; synonym of micans).
pullus Runde, 1835: 9 (Staphylinus; Type locality: Halae).
— Erichson, 1839a: 473 (Philonthus; synonym of micans).
— Erichson, 1840: 484 (Philonthus; synonym of micans).
— Kraatz, 1857d: 612 (Philonthus; synonym of micans).
— Ganglbauer, 1895: 457 (Philonthus; synonym of micans).
— Bernhauer and Schubert, 1914: 346 (Philonthus; synonym of micans).
— Jarrige, 1951a: 9 (Philonthus; synonym of micans).
— Smetana, 1958: 220 (Philonthus; synonym of micans).
— Coiffait, 1974: 94 (Paragabrius; synonym of micans).
helveticus Mulsant and Rey, 1876: 506 [= 1877: 362] (Philonthus; Type locality: Suisse).
— Duvivier, 1883: 147 (Philonthus; catalog; synonym of micans).
— Ganglbauer, 1895: 457 (Philonthus; synonym of micans).
— Bernhauer and Schubert, 1914: 346 (Philonthus; synonym of micans).
— Jarrige, 1951a: 9 (Philonthus; synonym of micans).
— Smetana, 1958: 220 (Philonthus; synonym of micans).
— Coiffait, 1974: 94 (Paragabrius; synonym of micans).
toenianus Mulsant and Rey, 1876: 506 [= 1877: 362] (Philonthus; Type locality: environs de Lyon).
— Duvivier, 1883: 147 (Philonthus; catalog; synonym of micans).
— Ganglbauer, 1895: 457 (Philonthus; synonym of micans).
— Bernhauer and Schubert, 1914: 346 (Philonthus; synonym of micans).
— Jarrige, 1951a: 9 (Philonthus; synonym of micans).
— Smetana, 1958: 220 (Philonthus; synonym of micans).
— Coiffait, 1974: 94 (Paragabrius; synonym of micans).
brunneipennis Gerhardt, 1899: 218 (Philonthus; variety of micans; [preoccupied]; Type locality: Not cited).
— Bernhauer and Schubert, 1914: 346 (Philonthus; synonym of micans).
— Scheerpeltz, 1933: 1331 (Philonthus; aberration of agilis).
— Scheerpeltz, 1933: 1351 (Philonthus; synonym of gerhardti).
— Jarrige, 1951a: 9 (Philonthus; cited as brunncipennis; synonym of micans).
— Smetana, 1958: 193 (Philonthus; aberration of agilis).
— Smetana, 1959: 158 (Philonthus; aberration of agilis; characters).
— Smetana, 1959: 168 (Philonthus; synonym of micans).
— Coiffait, 1974: 94 (Paragabrius; synonym of micans).
gerhardti Jakobson, 1909: 508 (Philonthus; replacement name for brunneipennis Gerhardt).
— Scheerpeltz, 1933: 1351 (Philonthus; variety of micans).
— Smetana, 1958: 220 (Philonthus; aberration of micans).
— Smetana, 1959: 168 (Philonthus; aberration of micans; characters).
— Coiffait, 1974: 94 (Paragabrius; variety of micans).
micanticollis Sharp, 1889: 36 (Philonthus; Type locality: Nagasaki).
— Bernhauer and Schubert, 1914: 346 (Philonthus; catalog).
— Scheerpeltz, 1933: 1351 (Philonthus; catalog).
— Nakane, 1963a: 91 (Philonthus; characters; habitus photograph; Japan).
— Frank, 1982: 42 (Philonthus; list of parasites).
— Yuh, Paik, Kwon, and Lee, 1985: 241 (Philonthus; Korea).
— J. Li, 1993: 36 (Philonthus; China).
▴ Distribution: Japan, Korea, China.
micantoides G. Benick and Lohse, 1956: 90 (Philonthus; Type locality: Tesperhude, Umg. Hamburg).
— Smetana, 1959: 166 (Philonthus; characters).
— V. Hansen, 1962: 243 (Philonthus; may not be separate from micans; Denmark).
— Coiffait, 1963: 12 (Paragabrius).
— Lohse, 1964: 185 (Philonthus; subgenus Philonthus; characters; central Europe).
— Smetana, 1964d: 77 (Philonthus; subgenus Philonthus; Czechoslovakia).
— Coiffait, 1974: 94 (Paragabrius; characters; France).
— Pope, 1977: 29 (Philonthus; Britain).
— Tóth, 1984: 77 (Philonthus; characters).
— Lucht, 1987: 106 (Philonthus; checklist; Germany; Czechoslovakia; Austria; Benelux).
— Hodge and Jones, 1995: 39 (Philonthus; characters; Britain).
— Luff, 1996: 30 (Philonthus; collecting notes; Isle of Man).
— Burakowski, Mroczkowski, and Stefańska, 2000: 49 (Paragabrius; catalog; Poland).
▴ Distribution: Britain, Denmark, France, Germany, Czechoslovakia.
microcephalus Gravenhorst, 1806, see: albipes Gravenhorst, 1802.
microcephalus Stephens, 1832, see: cognatus Stephens, 1832.
microphthalmus Horn, 1884, see: Gabrius.
microtophilus Coiffait, 1967, see: Bisnius.
mifanus Tottenham, 1956: 283 (Philonthus; Type locality: Ruanda: Rutovu, forêt du Rugege, 2350 m).
mimeticus Tottenham, 1962: 228 (Philonthus; subgenus Spatulonthus; Type locality: Gaboon).
mimulus Rottenberg, 1870, see: Gabronthus.
mimulus Sharp, 1874, see: Cafius.
mimus Smetana, 1959a: 211 (Philonthus; subgenus Philonthus; Type locality: Albanien: Butrint).
— Coiffait, 1963: 12 (Paragabrius).
— Coiffait, 1970e: 62 (Paragabrius; Corfu).
— Coiffait, 1974: 93 (Paragabrius; characters; Albania; Turkey).
— Smetana, 1977: 795 (Philonthus; Turkey).
— Coiffait, 1978d: 164 (Paragabrius; Turkey).
— Smetana, 1995e: 51 (Philonthus; transferred from Paragabrius by reason of Smetana's synonymy of Paragabrius and Philonthus).
▴ Distribution: Greece, Albania, Turkey.
minax Stephens, 1833, see: Bisnius.
minimus Cameron, 1933a: 170 (Philonthus; subgenus Gabrius; Type locality: Japan: Chuzenji).
minor Bernhauer, 1928, see: Mentophilonthus.
minor Erichson, 1840, see: concinnus Gravenhorst, 1802.
minutissimus Bernhauer, 1915, see: Gabronthus.
minutus Boheman, 1848: 283 (Philonthus; Type locality: Portus Natalensis).
— Fauvel, 1902b: 112 (Philonthus; Morocco; Algeria; Tunisia).
— Fauvel, 1904a: 60 (Philonthus; Kanara; Nilghiris; Ceylan; Bírmanie; China; Japan; Annam; Malacca; Borneo; Sumatra; Java; Celebes; New Guinea; Arabia; Egypt; Abyssinia; Senegal; Mozambique; Natal; Cape; Barbary; Spain).
— Fauvel, 1907: 45 (Philonthus; Nairobi; near Nairobi).
— Bernhauer, 1907: 386 (Philonthus; Japan).
— Bernhauer and Schubert, 1914: 346 (Philonthus; catalog).
— Bernhauer, 1920: 16 (Philonthus; Australia).
— Bernhauer, 1922c: 232 (Philonthus; Formosa).
— Lea, 1925: 220, 221 (Philonthus; characters; Australian introduction).
— Cameron, 1931: 363 (Philonthus; New Guinea).
— Cameron, 1932: 132 (Philonthus; characters; Ceylon; India).
— Scheerpeltz, 1933: 1351 (Philonthus; catalog).
— Scheerpeltz, 1935a: 654 (Philonthus; Australia).
— Adachi, 1937: 58 (Philonthus; Izu-Oshima).
— Bernhauer and Chapman, 1939: 71 (Philonthus; Kenya).
— Gridelli, 1939: 317 (Philonthus; Abyssinia).
— Tottenham, 1949b: 345 (Philonthus; characters).
— Cameron, 1950g: 36 (Philonthus; Belgian Congo).
— Tottenham, 1955: 167 (Philonthus; subgenus Spatulonthus; characters; notes).
— Scheerpeltz, 1960: 5 (Philonthus; Pakistan).
— Last, 1968: 370 (Philonthus; New Guinea).
— Coiffait, 1974: 315 (Spatulonthus; characters; Mediterranean region; Egypt; North Africa; Spain).
— Frank, 1982: 42 (Philonthus; list of parasites).
— Yuh, Paik, Kwon, and Lee, 1985: 241 (Philonthus; Korea).
— Last, 1987: 32 (Philonthus; Papua New Guinea).
— Lohse and Lucht, 1989: 168 (Philonthus; characters).
— Maachi, 1991: 251 (Spatulonthus; Morocco).
— Barrion, Litsinger, and Last, 1991: 1078 (Philonthus; associated with rice agroecosystem).
— J. Li, 1993: 37 (Philonthus; China).
— Yadav, Chand, and Yadav, 1995: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
▴ Distribution: Senegal, Morocco, Spain, Algeria, Tunisia, Egypt, Saudi Arabia, Ethiopia, Kenya, Congo, Mozambique, South Africa, Pakistan, India, Sri Lanka, Myanmar, Vietnam, Malaysia, Singapore, Borneo, Indonesia, New Guinea, China, Korea, Japan, Taiwan, Australia.
parviceps Kraatz, 1859: 86 (Philonthus; Type locality: Ceylan).
— Bernhauer, 1902b: 27 (Philonthus; Ceylon).
— Fauvel, 1904a: 60 (Philonthus; synonym of minutus).
— Bernhauer and Schubert, 1914: 346 (Philonthus; synonym of minutus).
— Cameron, 1932: 132 (Philonthus; synonym of minutus).
— Coiffait, 1974: 315 (Spatulonthus; synonym of minutus).
mutans Sharp, 1874: 43 (Philonthus; Type locality: northern China; Japan).
— Fauvel, 1904a: 60 (Philonthus; synonym of minutus).
— Bernhauer and Schubert, 1914: 346 (Philonthus; synonym of minutus).
— Cameron, 1932: 132 (Philonthus; synonym of minutus).
— Coiffait, 1974: 315 (Spatulonthus; synonym of minutus).
rufocinctus Fauvel, 1878: 126 [= 1878a: 46] (Philonthus; Type locality: Bône; Espagne: Algésiras).
— Fauvel, 1886: 57 [= 1886a: 49] (Philonthus; Morocco; Algeria).
— Fauvel, 1902b: 112 (Philonthus; synonym of minutus).
— Fauvel, 1904a: 60 (Philonthus; cited as rafocinctus; synonym of minutus).
— Bernhauer and Schubert, 1914: 346 (Philonthus; synonym of minutus).
— Cameron, 1932: 132 (Philonthus; synonym of minutus).
— Coiffait, 1974: 315 (Spatulonthus; synonym of minutus).
longiceps Fauvel, 1879: 104 (Philonthus; Type locality: Nouvelle Guinée, Fly River; Célèbes; Borneo; Malacca).
— Fauvel, 1904a: 60 (Philonthus; synonym of minutus).
— Bernhauer and Schubert, 1914: 346 (Philonthus; synonym of minutus).
— Cameron, 1921a: 377, 405 (Philonthus; characters; Singapore).
— Cameron, 1924: 87 (Philonthus; New Guinea).
— Bernhauer, 1926f: 317 (Philonthus; variety of minutus; Buru).
— Cameron, 1928f: 410 (Philonthus; Borneo).
— Cameron, 1932: 132 (Philonthus; synonym of minutus).
— Scheerpeltz, 1933: 1351 (Philonthus; synonym of minutus).
— Coiffait, 1974: 315 (Spatulonthus; synonym of minutus).
mirabilis Erichson, 1840, see: Xanthopygus.
mirei Levasseur, 1967a: 958 (Philonthus; Type locality: forêt de Bafut, N'guemba, 1900 m).
misellus Fauvel, 1895, see: adversus Bernhauer and Schubert, 1914.
misellus Sharp, 1885: 421 (Philonthus; Type locality: Guatemala near the city, Quiche Mountains 7000 to 9000 feet).
misor Tottenham, 1953, see: chopardi Cameron, 1950.
mivutanus Tottenham, 1962: 184 (Philonthus; Type locality: Kenya: Kanzeko).
— Coiffait, 1968a: 136 (Philonthus; Chad).
— Schillhammer, 1996a: 82 (Philonthus; Oman; Cape Verde Islands).
▴ Distribution: Oman, Kenya, Chad, Cape Verde Islands.
mixtus Last, 1968, see: Hesperus.
mjobergi Cameron, 1928g: 445 (Philonthus; cited as mjöbergi; [Note: Since this name is based on the Swedish name Mjöberg, the umlaut should be dropped and without further modification and the name should be emended to mjobergi]; Type locality: Borneo: Baram district).
— Scheerpeltz, 1933: 1351 (Philonthus; cited as mjöbergi; catalog).
— Hammond, 1984: 206 (Philonthus; cited as mjoebergi; Borneo).
— Herman, 2001: 12 (Philonthus; spelling corrected).
▴ Distribution: Borneo.
mlogosiensis Bernhauer, 1934f: 207 (Philonthus; Type locality: Kenya: Mlogosi).
— Tottenham, 1962: 202, 203, 209 (Philonthus; characters; Kenya; Cameroon).
▴ Distribution: Kenya, Cameroon.
mnemon Smetana, 1995e: 157 (Philonthus; Type locality: Arizona: Chiric. Mts., Ash Spg., 7 mi. SW Portal, 6000 ft. Also cited from New Mexico and Mexico).
modestus Cameron, 1932: 102 (Philonthus; Type locality: India).
— Scheerpeltz, 1933: 1351 (Philonthus; catalog).
— Scheerpeltz, 1954: 113 (Philonthus, cited as Phylonthus; Ceylon).
▴ Distribution: India, Sri Lanka.
modulatus Last, 1968, see: Hesperus.
modus Last, 1987: 26 (Philonthus; Type locality: Papua New Guinea: New Guinea NE: Karimi, 1800 m).
moestus Gravenhorst, 1806, see: umbrinus Gravenhorst, 1802.
moldavicus Wendeler, 1924: 343 (Philonthus; Type locality: Rumänien: Moldau).
— Scheerpeltz, 1933: 1351 (Philonthus; catalog).
— Coiffait, 1967a: 390 (Philonthus; characters; Romania).
— Coiffait, 1974: 298 (Philonthus; characters).
▴ Distribution: Romania.
molliculus Casey, 1915, see: Gabrius.
mollis Sharp, 1885: 408 (Philonthus; Type locality: Panama, San Feliz).
monachus Bernhauer, 1908d: 335 (Philonthus; Type locality: Paraguay).
mongendensis Bernhauer, 1928, see: Mentophilonthus.
mongolicus Csiki, 1901: 104 (Philonthus; variety of lepidus; Type locality: Mongolia: Chara-gol).
— Bernhauer and Schubert, 1914: 344 (Philonthus; variety of lepidus).
— Smetana, 1963c: 295 (Philonthus; subgenus Philonthus; valid species; characters; Mongolia).
— Smetana, 1965e: 173 (Philonthus; Mongolia).
— Coiffait, 1967a: 434 (Philonthus; subgenus Trionthus; characters; Mongolia).
— Smetana, 1967d: 203 (Philonthus; Mongolia).
— Coiffait, 1974: 194 (Philonthus; characters; Mongolia).
— Smetana 1975c: 170 (Philonthus; Mongolia).
— Shilov, 1976: 150 (Philonthus; Mongolia).
▴ Distribution: Mongolia.
montanellus Bernhauer, 1934g: 238 (Philonthus; Type locality: Kivu: Tshibinda; Lubero).
montanus Heer, 1839, see: Quedius.
montanus Bernhauer, 1934, see: Gabrius.
monticola Bernhauer, 1931, see: Gabrius.
montivagans Cameron, 1943: 129 (Philonthus; Type locality: Darjeeling: Ghum district).
montivagoides Coiffait, 1963: 28 (Philonthus; subgenus Kenonthus; Type locality: Alpes Maritimes: environs de Beuil).
— Coiffait, 1967a: 437 (Philonthus; subgenus Kenonthus; characters; Alps).
— Coiffait, 1974: 301 (Philonthus; subgenus Kenonthus; characters; Alps).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
▴ Distribution: France.
montivagus Heer, 1839: 275 (Philonthus; Type locality: Matt; Mühlebachalp; Krauchthalalp; Bergli; Pilatus; Engadin; Beverserthal; Camogaskerthal; Val Emet; Nufenen; Valser-berg; Luckmanier; Urserenthal; beim St. Annagletscher; Gotthard; Furca; Gemmi; Engelbergerjoch; Finnelalp; Ollen).
— Heer, 1834: 75 (Staphylinus; subgenus Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 346 and others cited nigrita Heer as a synonym of montivagus; Heer did not describe the species as new, he attributed it to Gravenhorst and in 1839 cited it as a synonym of montivagus]; elevational listing).
— Erichson, 1840: 432 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 433 cited laevicollis Erichson as a synonym of montivagus; Erichson did not describe the species as new, he attributed it to Boisduval and Lacordaire]; characters; Carinthia).
— Fairmaire and Laboulbène, 1856: 514 (Philonthus; characters; France).
— Kraatz, 1857d: 572 (Philonthus; characters; Germany).
— Kraatz, 1857d: 572 (Philonthus; [Note: Cited as nigrita Heer, a misidentification]; synonym of montivagus).
— Fauvel, 1865a: 295 [= 1865b: 45] (Philonthus; habitat; France).
— Fauvel, 1874: 231 [= 1874a: 457] (Philonthus; characters; France; Germany; Switzerland; Austria).
— Redtenbacher, 1874: 216 (Philonthus; characters).
— Mulsant and Rey, 1876: 359 [= 1877: 215] (Philonthus; characters; France).
— Ganglbauer, 1895: 443 (Philonthus; characters; Germany; Alps; Carpathians).
— Ganglbauer, 1895: 443 (Philonthus; [Note: Cited as nigrita Heer, a misidentification]; synonym of montivagans).
— Xambeu, 1900: 29 (Philonthus; larval characters).
— Reitter, 1909: 125 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 346 (Philonthus; catalog).
— Bernhauer and Schubert, 1914: 346 (Philonthus; [Note: Cited as nigrita Heer, a misidentification]; synonym of montivagus).
— Portevin, 1929: 357 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1351 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Smetana, 1958: 159 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 145 (Philonthus; characters).
— Coiffait, 1960: 363, 365 (Philonthus; subgenus Kenonthus; characters; Alps; Carpathians; Massif Central; Pyrenees).
— Lohse, 1964: 176 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 126 (Philonthus; France; Switzerland; Italy; Germany; Austria; Czech Republic; Slovakia; Poland; Romania).
— Coiffait, 1967a: 437 (Philonthus; subgenus Kenonthus; characters; Europe).
— Raianu, 1968a: 462 (Philonthus; Romania).
— Coiffait, 1974: 300 (Philonthus; subgenus Kenonthus; characters; Europe).
— Coiffait, 1974: 300 (Philonthus; [Note: Cited as nigrita Heer, a misidentification]; synonym of montivagus).
— Tóth, 1984: 46 (Philonthus; characters).
— Lohse, 1988: 42 (Philonthus; characters).
— Lucht, 1987: 104 (Philonthus; checklist; Poland; Czechoslovakia; France).
— Lohse and Lucht, 1989: 163 (Philonthus; characters).
— Pilon and Zanetti, 1991: 57 (Philonthus; Italy).
— Smetana, 1993b: 48 (Philonthus; subgenus Kenonthus; Czech Republic; Slovakia ?).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
▴ Distribution: Europe.
tripunctatus Coiffait, 1987: 497 (Philonthus; subspecies of montivagus; [Note: This name was originally cited by Coiffait, 1960, as an aberration of montivagus and was used subsequently by him as an aberration or, in 1974, a variety; the name was unavailable for all those uses; Coiffait made the name available in 1987 when he cited it as a subspecies of montivagus; characters had been presented in earlier articles when the name was still unavailable; preoccupied]; Type locality: Pyrénées: Luchon, cirque de la Glère).
— Coiffait, 1960: 364, 365 (Philonthus; subgenus Kenonthus; aberration of montivagus; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]; characters; cited from Pyrénées: Luchon, cirque de la Glère).
— Coiffait, 1967a: 435 (Philonthus; subgenus Kenonthus; aberration of montivagus; [Note: Nomen nudum (Articles 1.3.4, 45.5); characters).
— Coiffait, 1974: 301 (Philonthus; subgenus Kenonthus; variety of montivagus; [Note: Nomen nudum (Articles 1.3.4, 45.5); characters).
seguyi Coiffait, 1987: 497 (Philonthus; subspecies of montivagus; replacement name for tripunctatus Coiffait).
mocquerysi Fauvel, 1903e: 241 (Philonthus; Type locality: Cameroun; Gabon).
— Bernhauer and Schubert, 1914: 347 (Philonthus; catalog).
— Bernhauer, 1915o: 362 (Philonthus; cited as moquerisi; Cameroon).
— Tottenham, 1949b: 347 (Philonthus; characters).
— Frank, 1982: 42 (Philonthus; cited as moquerysi; list of parasites).
▴ Distribution: Cameroon, Gabon.
monaeses Smetana, 1995e: 402 (Philonthus; Type locality: Louisiana: Concordia Pa.: 5 mi. W. Ferriday. Also cited from Ontario, Arkansas, Connecticut, Illinois, Kentucky, Maine, Maryland, Massachusetts, New Hampshire, New Jersey, New York, North Carolina, Ohio, Oklahoma, and Tennessee).
morio Boheman, 1848: 279 (Philonthus; Type locality: Caffraria orientalis).
— Fauvel, 1907: 45 (Philonthus; Kibosho; Abyssinia; Cormoso; Angola; Fernando Po; Natal).
— Bernhauer and Schubert, 1914: 347 (Philonthus; catalog).
— Scheerpeltz, 1933: 1352 (Philonthus; catalog).
— Bernhauer, 1942c: 362 (Philonthus; Cameroon).
— Tottenham, 1949b: 333 (Philonthus; characters).
— Tottenham, 1956: 226 (Philonthus; Ruanda).
— Tottenham, 1961: 204 (Philonthus; Senegal).
— Tottenham, 1962: 138 (Philonthus; Tanganyika; Kenya).
— Levasseur, 1966a: 1213 (Philonthus; Congo; Senegal to the Cape).
— Levasseur, 1968a: 53 (Philonthus; Central African Republic).
— Scheerpeltz, 1974: 142 (Philonthus; South Africa).
— Scheerpeltz, 1974c: 21 (Philonthus; Sudan; Ethiopia).
▴ Distribution: South Africa, Angola, Congo, Rwanda, Tanzania, Comoros, Kenya, Ethiopia, Sudan, Central African Republic, Cameroon, Bioko, Senegal.
sericeiventris Eppelsheim, 1895: 203 (Philonthus; Type locality: Territoire des Cormoso).
— Bernhauer and Schubert, 1914: 347 (Philonthus; synonym of morio).
— Gridelli, 1939: 318 (Philonthus; Abyssinia).
— Scheerpeltz, 1974c: 21 (Philonthus; synonym of morio).
temporasinus Schubert, 1911: 24 (Philonthus; Type locality: Zoutpansberg, 800 m, Transvaal).
— Bernhauer and Schubert, 1914: 357 (Philonthus; catalog).
— Scheerpeltz, 1933: 1352 (Philonthus; synonym of morio).
— Scheerpeltz, 1974c: 21 (Philonthus; synonym of morio).
morosus Casey, 1915: 441 (Philonthus; Type locality: California: San Francisco Co.; [Note: See lectotype designation by Smetana, 1995e]. Also cited from California: Lake Co.).
— Scheerpeltz, 1933: 1352 (Philonthus; catalog).
— Smetana, 1995e: 452 (Philonthus; lectotype designation: S. Francisco Co., Cal.; characters; notes; Alberta; British Columbia; Saskatchewan; California; Idaho; Montana; Nevada; Oregon; Utah; Washington; Mexico).
▴ Distribution: Canada, USA, Mexico.
mortuorum Cameron, 1951c: 401 (Philonthus; Type locality: Kenya Colony: Nairobi).
motoensis Cameron, 1929: 62 (Philonthus; Type locality: Haut Uele: Moto).
— Scheerpeltz, 1933: 1352 (Philonthus; catalog).
— Bernhauer, 1942c: 361 (Philonthus; Cameroon).
— Tottenham, 1962: 188, 189 (Philonthus; characters).
▴ Distribution: Congo, Cameroon.
moucheti Levasseur, 1962: 236 (Philonthus; Type locality: Bertoua, Cameroun).
mubilibanus Cameron, 1950, see: Gabrius.
multicolor Lea, 1925: 224 (Philonthus; Type locality: North-western Australia: Behn River; Queensland: Coen River).
multiformis Last, 1968, see: specialis Cameron, 1937.
multipunctatus Blatchley, 1910, see: Gabrius.
multipunctatus Mannerheim, 1830, see: punctus Gravenhorst, 1802.
mundus Last, 1981: 124 (Neobisnius; Type locality: New Guinea: Zokoyoi River, Goroka).
— Last, 1987: 51 (Neobisnius; Papua New Guinea).
— Schillhammer, 1996a: 82 (Philonthus).
▴ Distribution: New Guinea.
murex Last, 1993: 149 (Philonthus; Type locality: New Guinea).
musonoiensis Levasseur, 1966b: 1499 (Philonthus; Type locality: région de Kolwezi, Katanga, Musonoie).
musurus Tottenham, 1956a: 241 (Philonthus; Type locality: Uganda: Jinja).
mutans Sharp, 1874, see: minutus Boheman, 1848.
mutatus Gemminger and Harold, 1868, see: Cafius.
muticus Sharp, 1876: 167 (Philonthus; Type locality: Ega, St. Paulo).
nairobianus Cameron, 1951c: 400 (Philonthus; Type locality: Kenya Colony: Nairobi, 5500 feet).
nairobiensis Fauvel, 1907: 41 (Philonthus; Type locality: Nairobi; steppe Masai pres Nairobi).
nakanei Sawada, 1965: 14 (Philonthus; subgenus Onychophilonthus; Type locality: Japan: Pref. Nagano: Mt. Jônen).
nanellus Casey, 1915, see: Gabrius.
nanus Melsheimer, 1844, see: hepaticus Erichson, 1840.
natalensis Boheman, 1848: 278 (Philonthus; Type locality: Portus Natalensis).
— Bernhauer and Schubert, 1914: 347 (Philonthus; catalog).
— Tottenham, 1949b: 313 (Philonthus; characters).
— Tottenham, 1962: 163 (Philonthus; characters).
— Scheerpeltz, 1974: 142 (Philonthus; South Africa).
▴ Distribution: South Africa, Tanzania, Kenya, Ethiopia.
vittatus Roth, 1851: 118 (Philonthus; Type locality: Abyssinien).
— Fauvel, 1907: 39 (Philonthus; variety of natalensis; Naivasha; Kilimandjaro; Abyssinia; Zanguebar; South Africa).
— Bernhauer and Schubert, 1914: 347 (Philonthus; synonym of natalensis).
— Rambousek, 1922: 1008 (Philonthus; variety of natalensis; Ethiopia).
— Tottenham, 1962: 161 (Philonthus; variety of natalensis; characters).
calidus Harold, 1881: 262 (Philonthus; Type locality: Not cited presumably East Africa).
— Fauvel, 1907: 39 (Philonthus; synonym of natalensis variety vittatus).
— Bernhauer and Schubert, 1914: 347 (Philonthus; synonym of natalensis).
nauticus Fairmaire, 1849, see: Cafius.
necopinus Schubert, 1911, see: hospes Erichson, 1843.
neglectus Cameron, 1933d: 387 (Philonthus; Type locality: Sumatra: Fort de Kock).
negligens Tottenham, 1962: 218 (Philonthus; Type locality: D. Ostafrika: Ussure).
negligens Smetana, 1966, see: Bisnius.
nematocerus Casey, 1915, see: Beeria.
nemorosus Gistel, 1857: 89 (Philonthus; [Note: Nomen dubium]; Type locality: Bavaria).
neobisniformis Bernhauer, 1915, see: Gabrius.
neonatus Smetana, 1965: 24 (Philonthus; subgenus Philonthus; Type locality: Springfield. Also cited from Nova Scotia).
— Horn, 1884: 200 (Philonthus; [Note: Misidentified as fulvipes according to Smetana, 1995e: 356]; characters; Canada; Massachusetts; Michigan; New York).
— Smetana, 1995e: 356 (Philonthus; characters; notes; Nova Scotia; Ontario; Quebec; Arkansas; Indiana; Iowa; Kansas; Kentucky; Maine; Maryland; Massachusetts; Michigan; Mississippi; Missouri; New Hampshire; New Jersey; New York; Pennsylvania).
— Downie and Arnett, 1996: 379 (Philonthus; characters; USA).
▴ Distribution: Canada, USA.
nepalicus Coiffait, 1977, see: Craspedomerus.
nepos Smetana, 1965f: 256 (Philonthus; subgenus Philonthus; Type locality: Turkestan: Buchara).
neptunus Smetana, 1995e: 436 (Philonthus; Type locality: Florida: Monroe Co.: Middle Torch Key, Sec. 17, Lazelle Place. Also cited from Bahama Islands).
neuter Tottenham, 1962: 219 (Philonthus; Type locality: Capeland: Willowmore. Also cited from Southwest Africa).
nguembaensis Levasseur, 1968: 1384 (Philonthus; Type locality: Cameroun: forêt de Bafut, Nguemba).
nichinaiensis Coiffait, 1982: 59 (Philonthus; Type locality: Inde: Cachemire, Sonamarg, Himalaya-Hauptkette, Nichina Tal, 3100–3200 m).
nidicola Bernhauer, 1921c: 108 (Philonthus; Type locality: Provinz Buenos Aires, Monte Veloz).
niger Melsheimer, 1844, see: Bisnius.
niger O. Müller, 1764, see: splendens Fabricius, 1793.
nigerrimus Erichson, 1840: 434 (Philonthus; Type locality: Mexico; [Note: See lectotype designation by Smetana, 1991a]).
— Sharp, 1885: 398 (Philonthus; notes; Mexico).
— Bernhauer and Schubert, 1914: 347 (Philonthus; catalog).
— Smetana, 1991a: 234 (Philonthus; lectotype designation: Mexico; characters; Mexico).
▴ Distribution: Mexico.
nigrus Coiffait, 1974 [Note: Nomen nudum], see: oberti Eppelsheim, 1889.
nigriceps Gemminger and Harold, 1868, see: Erichsonius.
nigriceps Eppelsheim, 1885: 112 (Philonthus; subgenus Rabigus; [preoccupied]; Type locality: Goldküste: Adda).
— Bernhauer and Schubert, 1914: 347 (Philonthus; catalog).
— Tottenham, 1949b: 353 (Philonthus; characters).
— Cameron, 1952e: 831 (Philonthus; Dahomey).
— Tottenham, 1961: 204 (Philonthus; Senegal).
— Levasseur, 1966a: 1214 (Philonthus; Senegal; Upper Volta; Ivory Coast; Congo).
— Herman, 2001: 47 (Philonthus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Senegal, Burkina Faso, Ivory Coast, Ghana, Benin, Congo.
nigricollis Gernhardt, 1908, see: Rabigus.
nigricolor Cameron, 1942b: 328 (Philonthus; Type locality: Chyulu Hills, altitude 5600 feet).
nigricoxis Cameron, 1928c: 564 (Philonthus; Type locality: Sikkim: Karponang, 10,000 feet. Tibet: Tuna, 14,500 feet).
— Cameron, 1932: 129 (Philonthus; characters; Tibet; India).
— Scheerpeltz, 1933: 1352 (Philonthus; catalog).
— Coiffait, 1982: 30 (Philonthus; Nepal).
— Coiffait, 1983: 273 (Philonthus; Kashmir).
▴ Distribution: India, Nepal, China.
nigrinus Boheman, 1858: 30 (Philonthus; Type locality: Promontorium Bonae Spei).
— Bernhauer and Schubert, 1914: 347 (Philonthus; catalog).
— Scheerpeltz, 1974: 149 (Philonthus; Basutoland; South West Africa; South Africa).
▴ Distribution: Lesotho, South Africa, Namibia.
nigripennis Outerelo, 1976 [Note: Nomen nudum], see: rubripennis Stephens, 1832.
nigripes Cameron, 1932a: 141 (Philonthus; Type locality: Selangor: The Gap).
— Cameron, 1933b: 345 (Philonthus; British North Borneo).
— Cameron, 1937: 5 (Philonthus; notes; Java).
▴ Distribution: Borneo, Indonesia.
nigrita Gravenhorst, 1806: 68 (Staphylinus; Type locality: Not cited).
— Gyllenhal, 1810: 345 (Staphylinus; synonym of virgo Gravenhorst).
— Nordmann, 1837: 101 (Philonthus; Germany).
— Erichson, 1839a: 471 (Philonthus; characters; Germany).
— Heer, 1839: 269 (Philonthus; characters; Switzerland).
— Stephens, 1839: 398 (Philonthus; characters; England).
— Erichson, 1840: 481 (Philonthus; characters; Germany; Sweden).
— Kiesenwetter, 1848a: 19 (Philonthus; characters; Sweden; Germany; France; Switzerland).
— Redtenbacher, 1849: 825 (Philonthus; characters).
— Kiesenwetter, 1851a: 417 (Philonthus; France).
— Fairmaire and Laboulbène, 1856: 526 (Philonthus; characters; France).
— Kraatz, 1857d: 611 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 198 (Philonthus; characters; Germany).
— Thomson, 1860: 165 (Philonthus; characters; Scandinavia).
— Fauvel, 1874: 258 [= 1874a: 484] (Philonthus; characters; France).
— Redtenbacher, 1874: 217 (Philonthus; characters; Germany).
— Seidlitz, 1875: 277 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 496 [= 1877: 352] (Philonthus; characters; France).
— J. Sahlberg, 1876: 21 (Philonthus; Finland).
— Fowler, 1888: 277 (Philonthus; characters; Britain).
— Eppelsheim, 1892: 334 (Philonthus; Turkmenistan; Turkestan).
— Ganglbauer, 1895: 456 (Philonthus; characters; Europe).
— Sainte-Claire Deville, 1906: 99 (Philonthus; Corsica).
— Reitter, 1909: 130 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 347 (Philonthus; catalog).
— Johansen, 1914: 412 (Philonthus; characters; Denmark).
— Fiori, 1915: 15 (Philonthus; Italy).
— Portevin, 1929: 364 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1352 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Jarrige, 1951a: 7 (Philonthus; characters).
— V. Hansen, 1952: 75 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1958: 219 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 168 (Philonthus; characters).
— Palm, 1963b: 94 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 185 (Philonthus; subgenus Philonthus; characters; central Europe).
— Smetana, 1964d: 76 (Philonthus; subgenus Philonthus; notes; Czechoslovakia).
— Horion, 1965: 172 (Philonthus; Scotland; North Cape; Lapland; Holland; Belgium; France; Switzerland to Czechoslovakia; Poland and Russia; Portugal; Spain; Balearics; Corsica; Sicily; Italy; Austria; Germany).
— Coiffait, 1967a: 431 (Philonthus; characters; Europe; Caucasus; Siberia).
— Szujecki, 1968a: 731 (Philonthus; Poland).
— Korge, 1971a: 7 (Philonthus; Turkey).
— Coiffait, 1974: 274 (Philonthus; characters; Europe; Britain; Scandinavia; Caucasus; Siberia).
— Pope, 1977: 29 (Philonthus; Britain).
— Frank, 1982: 42 (Philonthus; list of parasites).
— Hinton, 1981: 661 (Philonthus; characters of egg).
— Tóth, 1984: 76 (Philonthus; characters; Hungary).
— Lucht, 1987: 104 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Pilon and Zanetti, 1991: 60 (Philonthus; Italy).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Sterrenburg and Schülke, 1997: 18 (Philonthus; Netherlands).
— Owen, 1997a: 149 (Philonthus; collecting sites; Ireland).
▴ Distribution: Europe, Caucasus, Russia, Turkey, Turkmenistan, Uzbekistan.
nigritoides Smetana, 1964, see: crotchi Horn, 1884.
nigrituloides Cameron, 1932, see: Gabrius.
nigritus Runde, 1835, see: succicola Thomson, 1860.
nigriventris Thomson, 1867, see: Bisnius.
nigroaeneus Stephens, 1832, see: carbonarius Gravenhorst, 1802.
nigrocinctus Bernhauer, 1915d: 142 (Philonthus; Type locality: Abessinien: Kunhe).
nigrolineatus Bernhauer, 1913, see: virgatus Sharp, 1889.
nigrolucens Cameron, 1937: 9 (Philonthus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft).
nigrolucens Cameron, 1959, see: diatrechoides Tottenham, 1962.
nigropolitus Bernhauer, 1916, see: rotundicollis Ménétriés, 1832.
nigroscutellatus Lecoq, 1997: 280 (Philonthus; Type locality: Madagascar Est. parc de Ranomafana, 900 m).
nilgiriensis Cameron, 1920, see: quediiformis Cameron, 1932.
nilgiriensis Cameron, 1920, see: remotus Fauvel, 1895.
nimbicola Fauvel, 1874: 231 [= 1874a: 457] (Philonthus; Type locality: Mont Rosa, Macugnaga).
— Mulsant and Rey, 1876: 361 [= 1877: 217] (Philonthus; characters; France).
— Ganglbauer, 1895: 443 (Philonthus; characters; Europe).
— Bernhauer and Schubert, 1914: 348 (Philonthus; catalog).
— Scheerpeltz, 1933: 1351 (Philonthus; variety of montivagus).
— Wüsthoff, 1937: 32 (Philonthus; illustration of aedeagus).
— Lohse, 1956: 87 (Philonthus; cited as valid species; characters; notes).
— Smetana, 1959: 146 (Philonthus; variety of montivagus; characters).
— Coiffait, 1960: 363, 365 (Philonthus; subgenus Kenonthus; characters; Alps).
— Coiffait, 1967a: 435 (Philonthus; subgenus Kenonthus; characters; Alps).
— Coiffait, 1974: 298 (Philonthus; subgenus Kenonthus; characters; Alps).
— Lohse, 1988: 42 (Philonthus; characters).
— Lucht, 1987: 104 (Philonthus; subspecies of montivagus; checklist; Germany; Austria; Switzerland).
— Lohse and Lucht, 1989: 163 (Philonthus; characters).
— Pilon and Zanetti, 1991: 59 (Philonthus; notes; Italy).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
▴ Distribution: France, Italy.
jureceki Rambousek, 1913: 116, 118 (Quedius; subgenus Ediquus; Type locality: Folgario; Mont Cenis).
— Gridelli, 1924d: 69 (Quedius; characters; Italy).
— Scheerpeltz, 1933: 1351 (Philonthus; synonym of montivagus).
— Coiffait, 1974: 298 (Philonthus; synonym of nimbicola).
nimboides Tottenham, 1949b: 315 (Philonthus; Type locality: S. Rhodesia: Vumbu Mts., alt. 5700 feet; Cloudlands, Vumbu Mts., alt. 6000 feet).
nimeaglius Tottenham, 1962: 172 (Philonthus; Type locality: Kenya: Molo, Mau Escarpment, 2150–2200 m).
nimeikopus Tottenham, 1962: 199 (Philonthus; Type locality: Congo: Blukwa).
nitens Gravenhorst, 1802, see: varians Paykull, 1789.
nitens Kraatz, 1859, see: argutus Herman, 2001.
nitescens Horn, 1884, see: gaudens Tottenham, 1939.
nitescens Stephens, 1832, see: albipes Gravenhorst, 1802.
nitidicollis Lacordaire, 1835: 401 (Staphylinus; Type locality: Paris).
— Erichson, 1839a: 454 (Philonthus; synonym of bimaculatus).
— Erichson, 1840: 448 (Philonthus; synonym of bimaculatus).
— Kraatz, 1857d: 584 (Philonthus; synonym of varius).
— Fauvel, 1869: 489 (Philonthus; synonym of varius).
— Ganglbauer, 1895: 448 (Philonthus; variety of varius).
— Reitter, 1909: 128 (Philonthus; aberration of varius; characters; Germany).
— Bernhauer and Schubert, 1914: 359 (Philonthus; aberration of varius).
— Donisthorpe, 1930: 95 (Philonthus; aberration of bimaculatus; characters; Britain).
— Scheerpeltz, 1933: 1334 (Philonthus; synonym of bimaculatus).
— Scheerpeltz, 1933: 1353 (Philonthus; catalog).
— Cameron, 1950g: 36 (Philonthus; Belgian Congo).
— Smetana, 1953: 118 (Philonthus; aberration of bimaculatus; Turkey).
— Smetana, 1958: 185 (Philonthus; aberration of bimaculatus).
— Smetana, 1959: 151 (Philonthus; aberration of bimaculatus; characters).
— Scheerpeltz, 1961d: 129 (Philonthus; aberration of bimaculatus; Italy).
— Scheerpeltz, 1963: 12 (Philonthus; aberration of bimaculatus; Afghanistan).
— Scheerpeltz, 1965b: 402 (Philonthus; aberration of bimaculatus; Sicily).
— Coiffait, 1967a: 398 (Philonthus; characters; Europe; North Africa).
— Coiffait, 1970e: 62 (Philonthus; Laconia).
— Coiffait, 1974: 250 (Philonthus; characters; Europe; Britain; Sweden; North Africa; Anatolia; Lebanon; Caucasus; Persia; Transcaspian region).
— Bordoni, 1975a: 152 (Philonthus; Afghanistan).
— Pace and Zanetti, 1977: 294 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Raichev, 1979: 83 (Philonthus; Bulgaria).
— Tronquet, 1981: 70 (Philonthus; Afghanistan).
— Coiffait, 1983e: 478 (Philonthus; Afghanistan).
— Bordoni, 1984: 342 (Philonthus; Lebanon).
— Tóth, 1984: 55 (Philonthus; characters; Hungary).
— Lohse and Lucht, 1989: 165 (Philonthus; note).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Assing and Wunderle, 2001: 37 (Philonthus; Cyprus).
▴ Distribution: Libya, Tunisia, Algeria, Morocco, Europe, Russia, Caucasus, Turkey, Lebanon, Israel, Iran, “Transcaspia”, Afghanistan.
bimaculatus Gravenhorst, 1802: 38 (Staphylinus; [preoccupied]; Type locality: Not cited, but presumably Brunsvigae).
— Latreille, 1804: 321 (Staphylinus; characters; Germany; France).
— Gravenhorst, 1806: 63 (Staphylinus).
— Curtis, 1829: 26 (Philonthus; catalog; Britain).
— Stephens, 1832: 239 (Philonthus; characters; England).
— Runde, 1835: 9 (Staphylinus; characters; Germany).
— Lacordaire, 1835: 400 (Staphylinus; characters; France).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Nordmann, 1837: 92 (Philonthus; characters; Germany).
— Erichson, 1839a: 454 (Philonthus; characters; Germany).
— Heer, 1839: 261 (Philonthus; characters; Switzerland).
— Stephens, 1839: 398 (Philonthus; characters; England).
— Erichson, 1840: 448 (Philonthus; characters; Prussia; Germany; France; Italy).
— Jacobsen, 1849: 692 (Philonthus; characters; Denmark).
— Redtenbacher, 1849: 700 (Philonthus; characters; Austria).
— Küster, 1853a: 56 (Philonthus; characters; Prussia; Germany; Switzerland; France; Italy).
— Rosenhauer, 1856: 71 (Philonthus; Cadiz; Algeciras).
— Fairmaire and Laboulbène, 1856: 515 (Philonthus; characters; France).
— Kraatz, 1857d: 584 (Philonthus; synonym of varius).
— Redtenbacher, 1857: 190 (Philonthus; characters; Austria).
— Fauvel, 1874: 250 [= 1874a: 476] (Philonthus; variety of varius).
— Redtenbacher, 1874: 209 (Philonthus; characters; Austria).
— Fauvel, 1878: 125 [= 1878a: 45] (Philonthus; variety of varius; Algeria).
— Fauvel, 1886: 56 [= 1886a: 48] (Philonthus; Algeria).
— Fauvel, 1902b: 112 (Philonthus; Morocco; Algeria; Tunisia).
— Reitter, 1909: 128 (Philonthus; aberration of varius; characters; Germany).
— Bernhauer and Schubert, 1914: 359 (Philonthus; aberration of varius).
— Gridelli, 1924e: 177 (Philonthus; Persia).
— Gridelli, 1930: 72 (Philonthus; Libya).
— Scheerpeltz, 1933: 1334 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Kloet and Hincks, 1945: 166 (Philonthus; cited as an invalid name; Britain).
— V. Hansen, 1952: 56 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1953: 118 (Philonthus; Turkey).
— Smetana, 1958: 185 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 151 (Philonthus; characters).
— Scheerpeltz, 1961h: 252 (Philonthus; Afghanistan).
— Scheerpeltz, 1963: 11 (Philonthus; Afghanistan).
— Palm, 1963b: 72 (Philonthus; characters; Sweden; Denmark).
— Lohse, 1964: 179 (Philonthus; subgenus Philonthus; characters; central Europe).
— Scheerpeltz, 1964a: 300 (Philonthus; subgenus Philonthus; Crete).
— Horion, 1965: 144 (Philonthus; Turkey; Caucasus; Siberia; Mesopotamia; Afghanistan; Uzbekistan; Iberian Peninsula to Italy; Dalmatia; Corfu; Montenegro; Greece; France to Poland and Russia; Denmark; Sweden; Germany; Austria).
— Raianu, 1968a: 466 (Philonthus; Romania).
— Szujecki, 1968a: 729 (Philonthus; Poland).
— Coiffait, 1974: 250 (Philonthus; synonym of nitidicollis).
— Pope, 1977: 29 (Philonthus; synonym of nitidicollis).
— Smetana, 1978: 117 (Philonthus; collecting notes; Israel).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Lohse and Lucht, 1989: 165 (Philonthus; synonym of nitidicollis).
— Smetana, 1993b: 48 (Philonthus; synonym of nitidicollis).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; synonym of nitidicollis).
rosarius Tottenham, 1945: 71 (Philonthus; variety of nitidicollis; replacement name for bimaculatus Gravenhorst; [Note: Unnecessary replacement name; nitidicollis predates rosarius]).
— Coiffait, 1967a: 398 (Philonthus; variety of nitidicollis; characters).
— Coiffait, 1970e: 62 (Philonthus; variety of nitidicollis; Cyprus).
— Coiffait, 1974: 250 (Philonthus; variety of nitidicollis).
— Pope, 1977: 29 (Philonthus; synonym of nitidicollis).
instigma Coiffait, 1970e: 64 (Philonthus; variety of nitidicollis; [Note: Nomen nudum (Article 15.2)]; cited from Val Cédri, 1500 m, Chypre).
nitidicollis Klug, 1855, see: sanamus Tottenham, 1955.
nitidus Fabricius, 1787: 220 (Staphylinus; Type locality: Kiliae).
— Paykull, 1789: 40 (Staphylinus; characters; Sweden).
— Gmelin, 1790: 2028 (Staphylinus; characters).
— Fabricius, 1793: 524 (Staphylinus; characters).
— Schrank, 1798: 640 (Staphylinus; characters; notes).
— Cederhjelm, 1798: 118 (Staphylinus; characters).
— Paykull, 1800: 390 (Staphylinus; characters; Sweden).
— Fabricius, 1801: 596 (Staphylinus; characters; Germany).
— Turton, 1802: 511 (Staphylinus; characters; Kiel).
— Latreille, 1804: 318 (Staphylinus; characters).
— Gravenhorst, 1806: 42 (Staphylinus; characters).
— Gyllenhal, 1810: 315 (Staphylinus; characters; Sweden).
— Germar, 1827: 2 (Staphylinus; characters; Germany; Sweden; Italy).
— Zetterstedt, 1828: 74 (Staphylinus; characters; Lapland).
— Curtis, 1829: 25 (Quedius; catalog; Britain).
— C. Sahlberg, 1830: 320 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 27 (Staphylinus; Finland; Germany; Russia).
— Mannerheim, 1831: 441 (Staphylinus; Finland; Germany; Russia).
— Stephens, 1832: 225 (Quedius; characters; England).
— Lacordaire, 1835: 377 (Emus; characters; France).
— Nordmann, 1837: 90 (Philonthus; Sweden; Germany).
— Zetterstedt, 1838: 63 (Staphylinus; characters; habitat; Lapland).
— Stephens, 1839: 393 (Quedius; characters; England).
— Erichson, 1839a: 448 (Philonthus; characters; Germany).
— Erichson, 1840: 435 (Philonthus; characters; Europe).
— Jacobsen, 1849: 687 (Philonthus; characters; Denmark).
— Redtenbacher, 1849: 700 (Philonthus; characters; Austria).
— Kiesenwetter, 1851a: 416 (Philonthus; France).
— Küster, 1853a: 47 (Philonthus; characters; Europe).
— Fairmaire and Laboulbène, 1856: 515 (Philonthus; characters; France).
— Kraatz, 1857d: 576 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 190 (Philonthus; characters; Austria).
— Thomson, 1860: 156 (Philonthus; characters; Scandinavia).
— Hochhuth, 1862: 53 (Philonthus; Russia).
— Fauvel, 1865a: 295 [= 1865b: 45] (Philonthus; habitat; France).
— Fauvel, 1874: 214 [= 1874a: 440] (Philonthus; characters; France; Scandinavia; Germany; Switzerland; Austria; Italy; Russia).
— Redtenbacher, 1874: 209 (Philonthus; characters; Austria).
— Seidlitz, 1875: 273 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 364 [= 1877: 220] (Philonthus; characters; France).
— J. Sahlberg, 1876: 13 (Philonthus; Finland).
— Ganglbauer, 1895: 443 (Philonthus; characters; north and middle Europe).
— Bordas, 1899: 230 (Philonthus; defensive glands).
— Reitter, 1909: 126 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 348 (Philonthus; catalog).
— Johansen, 1914: 393 (Philonthus; characters; Denmark).
— Šulc, 1922 (Philonthus; osmeterium; anatomy; histology; physiology).
— Portevin, 1929: 358 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1353 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— V. Hansen, 1952: 50 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1958: 160 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 149 (Philonthus; characters).
— Palm, 1963b: 67 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 177 (Philonthus; subgenus Philonthus; characters; central Europe).
— Scheerpeltz, 1964: 28 (Philonthus; subgenus Philonthus; Mongolia).
— Raianu, 1964: 103 (Philonthus; Romania).
— Horion, 1965: 132 (Philonthus; Russia; Finland; Sweden; Norway; Denmark; Baltics; Ukraine; Poland; Czechoslovakia; Hungary; Romania; Serbia; Italy; Switzerland; France; Holland; Germany; Austria).
— Smetana, 1966: 330 (Philonthus; Czechoslovakia).
— Pototskaia, 1966: 158 (Philonthus; larval characters).
— Pototskaia, 1967: 63 (Philonthus; larval characters).
— Coiffait, 1967a: 393 (Philonthus; characters; Europe; Siberia).
— Raianu, 1968a: 462 (Philonthus; Romania).
— Szujecki, 1968a: 727 (Philonthus; Poland).
— Coiffait, 1974: 225 (Philonthus; characters; Europe; Siberia; Britain).
— Shilov, 1975a: 375 [= 1975b: 97] (Philonthus; Komi region of Russia).
— Shilov, 1976: 147 (Philonthus; Mongolia).
— Tóth, 1984: 49 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Schülke and Uhlig, 1989: 243 (Philonthus; characters).
— Pilon and Zanetti, 1991: 61 (Philonthus; Italy).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
— Schillhammer, 2000: 137 (Philonthus; characters; Russia; Mongolia).
▴ Distribution: Europe, Russia, Kazakhstan, Mongolia.
saxatilis Schrank, 1781: 234 (Staphylinus; [Note: Nomen dubium]; Type locality: Austria).
— Herbst, 1784: 150 (Staphylinus; Germany).
— Rossi, 1790: 250 (Staphylinus; characters; Italy).
— Gmelin, 1790: 2031 (Staphylinus; characters; Germany, France).
— Turton, 1802: 516 (Staphylinus; characters; Germany).
— Gravenhorst, 1802: 33 (Staphylinus; synonym of nitidus Fabricius).
coenosus Gravenhorst, 1806: 51 (Staphylinus; Type locality: Norimbergae; Borussia).
— Gyllenhal, 1810: 315 (Staphylinus; synonym of nitidus Fabricius).
— C. Sahlberg, 1830: 320 (Staphylinus; synonym of nitidus Fabricius).
— Lacordaire, 1835: 386 (Staphylinus; characters; France).
— Zetterstedt, 1838: 63 (Staphylinus; synonym of nitidus Fabricius).
— Erichson, 1839a: 448 (Philonthus; synonym of nitidus).
— Erichson, 1840: 436 (Philonthus; synonym of nitidus).
— Redtenbacher, 1849: 700 (Philonthus; synonym of nitidus).
— Ganglbauer, 1895: 443 (Philonthus; synonym of nitidus).
— Bernhauer and Schubert, 1914: 348 (Philonthus; synonym of nitidus).
— Smetana, 1958: 160 (Philonthus; synonym of nitidus).
— Coiffait, 1974: 225 (Philonthus; synonym of nitidus).
— Schillhammer, 2000: 137 (Philonthus; synonym of nitidus).
niziensis Cameron, 1933, see: xanthoraphis Eppelsheim, 1895.
nobilis Bernhauer, 1910, see: Laetulonthus.
notabilis Kraatz, 1859: 79 (Philonthus; Type locality: Ceylan; India orientalis).
— Fauvel, 1886b: 150 (Philonthus; Philippines; Ceylon; East Indies; Annam).
— Bernhauer and Schubert, 1914: 348 (Philonthus; catalog; Philippines).
— Cameron, 1921a: 376, 405 (Philonthus; characters; Singapore).
— Cameron, 1932: 76 (Philonthus; characters; Ceylon; India).
— Scheerpeltz, 1933: 1353 (Philonthus; catalog).
— Scheerpeltz, 1954: 113 (Philonthus, cited as Phylonthus; India).
— Scheerpeltz, 1960: 5 (Philonthus; Pakistan).
— Nakane, 1963a: 91 (Philonthus; characters; habitus photograph; Japan).
— Scheerpeltz, 1976a: 87 (Philonthus; Nepal).
— Coiffait, 1981f: 329 (Philonthus; Andaman Islands).
— Ikudome and Hayakawa, 1994: 187 (Philonthus; cattle dung collections; Ryukyu Islands).
— Yadav, Chand, and Yadav, 1995: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
▴ Distribution: “East Indies”, Pakistan, India, Nepal, Sri Lanka, Andaman Islands, Vietnam, Singapore, Philippines, Japan.
taprobanus Cameron, 1918b: 171 (Quedius; subgenus Raphirus; Type locality: Ceylon: Kandy).
noualhieri Fauvel, 1898, see: Bisnius.
novaeguineae Cameron, 1937a: 102 (Philonthus; Type locality: New Guinea: Kokoda, alt. 1200 ft.).
novaezeelandiae Duvivier, 1883: 147 (Philonthus; replacement name for ruficornis Broun).
— Bernhauer and Schubert, 1914: 348 (Philonthus; catalog).
— Herman, 2001: 27 (Philonthus; synonymy).
▴ Distribution: New Zealand.
ruficornis Broun, 1880: 111 (Philonthus; [preoccupied]; Type locality: Whangarei Harbour).
— Bernhauer and Schubert, 1914: 348 (Philonthus; synonym of novaezeelandiae, cited as novaezealandiae).
manaiaensis Broun, 1910: 15 (Quedius; replacement name for ruficornis Broun; [Note: Unnecessary replacement name]).
brounianus Bernhauer and Schubert, 1914: 330 (Philonthus; replacement name for ruficornis Broun; [Note: Unnecessary replacement name]).
novellus Last, 1952: 90 (Philonthus; Type locality: Belgian Congo: Jadotville).
novus Cameron, 1935, see: Gabrius.
nudus Sharp, 1874: 36 (Philonthus; subgenus Cafius; Type locality: Amakusa and Iwosima).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Kirshenblat, 1938: 531 (Cafius; Russia).
— Watanabe, 1961: 354 (Cafius; Izu Islands).
— Watanabe and Shibata, 1965: 321 (Cafius; subgenus Cafius; Japan).
— Moore, 1965: 99, 101 (Philonthus; characters; notes).
— Smetana, 1965a: 9 (Cafius; subgenus Cafius; British Columbia).
— Coiffait, 1974a: 198 (Cafius; Siberia).
— Moore, 1977b: 407 (Philonthus; larval characters; Washington).
— Yuh, Paik, Kwon, and Lee, 1985: 243 (Cafius; Korea).
— J. Li, 1993: 40 (Cafius; China).
— Kanehisa, Tsumuki, and Kawazu, 1994 (Cafius; chemical secretions).
— Smetana, 1995e: 257 (Philonthus; characters; British Columbia; Washington).
— Watanabe, 1998: 314 (Philonthus; Japan).
— Schillhammer, 1999a: 62 (Philonthus).
— Ryabukhin, 1999: 60 (Cafius; catalog; habitat; distribution; northeastern Russia).
— Naomi, Kuranishi, Saito, and Maruyama, 2000: 109 (Philonthus; collected from North Kuril Islands).
▴ Distribution: Russia, China, Korea, Japan, Canada, USA.
johnsoni Fall, 1916: 13 (Cafius; Type locality: Seattle, Washington; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1369 (Cafius; catalog).
— Hatch, 1957: 211 (Cafius; subspecies of nudus; characters; British Columbia; Washington).
— Moore, 1965: 99 (Philonthus; synonym of nudus).
— Smetana, 1995e: 257 (Philonthus; lectotype designation: Seattle; synonym of nudus).
sharpi C. Koch, 1936b: 182 (Cafius; aberration of nudus; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]; cited from Japan).
grossepunctatus Bernhauer, 1939e: 154 (Philonthus; Type locality: Japan: Oita, Beppu, N.O. Kiushiu).
nugax Sharp, 1885: 414 (Philonthus; Type locality: Mexico, Mirador).
numata R. Dvořák, 1958: 139 (Philonthus; Type locality: Japan: Honshu: Numata Gumma).
— Watanabe, 1993c (Philonthus; Sadogashima Island).
— J. Li, 1993: 35 (Philonthus; China).
▴ Distribution: Japan, China.
nuristanicus Scheerpeltz, 1961b: 48 (Philonthus; subgenus Gefyrobius; Type locality: Afghanistan: Gusalek).
oberti Eppelsheim, 1889: 174 (Philonthus; Type locality: Daurien; [Note: See lectotype designation by Schillhammer, 2000]).
— Bernhauer and Schubert, 1914: 348 (Philonthus; catalog).
— Scheerpeltz, 1933: 1353 (Philonthus; catalog).
— Coiffait, 1967a: 397 (Philonthus; characters; Siberia; eastern Asia).
— Coiffait, 1974: 232 (Philonthus; characters; Siberia; East Asia).
— Schillhammer, 1999a: 62 (Philonthus; synonyms).
— Schillhammer, 2000: 131 (Philonthus; lectotype designation; characters; Russia; Mongolia; North Korea; South Korea; Japan; China).
▴ Distribution: Russia, Mongolia, China, North Korea, South Korea, Japan.
beckeri Bernhauer, 1933: 30 (Philonthus; Type locality: Szetschwan: Kinfushan).
— Coiffait, 1967a: 397 (Philonthus; characters; China).
— Coiffait, 1974: 279 (Philonthus; characters; China).
— Schillhammer, 1999a: 62 (Philonthus; synonym of oberti).
— Schillhammer, 2000: 131 (Philonthus; synonym of oberti).
diffusiventris Bernhauer, 1933a: 41 (Philonthus; Type locality: Tatsienlu-Kiulung).
— Coiffait, 1967a: 397 (Philonthus; characters; China).
— Coiffait, 1974: 278 (Philonthus; characters; China).
— Schillhammer, 1999a: 62 (Philonthus; synonym of oberti).
— Schillhammer, 2000: 131 (Philonthus; synonym of oberti).
pseudojaponicus Bernhauer, 1936g: 307 (Philonthus; Type locality: North Alp. Japan: Kamikochi; Jesso: Nemoro).
— Nakane, 1963: A239 (Philonthus; Japan).
— Sawada, 1965: 15 (Philonthus; subgenus Philonthus; characters; Japan).
— J. Li, 1993: 36 (Philonthus; China).
— Schillhammer, 1999a: 62 (Philonthus; synonym of oberti).
— Schillhammer, 2000: 131 (Philonthus; synonym of oberti).
reflexiventris Tikhomirova, 1973: 164 (Philonthus; Type locality: Suputinskii zapovednik; Kedrovaia Pad).
— Schillhammer, 1999a: 62 (Philonthus; synonym of oberti).
— Schillhammer, 2000: 131 (Philonthus; synonym of oberti).
nigrus Coiffait, 1974a: 202 (Philonthus; variety of oberti; [Note: Nomen nudum (Article 15.2)]; cited from Khassan, Réserve Kedrovaia-Pad).
obex Tottenham, 1962: 182 (Philonthus; Type locality: Tanganyika Terr.: Bunduki, Uluguru Mts., 1500 m., gorge de la Mungula).
obirensis Lohse, 1988: 42 (Philonthus; Type locality: Obir, Kärnten).
— Lohse, 1964: 176 (Philonthus; subgenus Philonthus; subspecies of montivagus; cited as nimbicola; [Note: Misidentification of obirensis according to Lohse and Lucht, 1989]; characters; Alps).
— Lohse and Lucht, 1989: 163 (Philonthus; characters; Alps).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
▴ Distribution: Austria.
oblitus Jarrige, 1951a: 9 (Philonthus; Type locality: Hyères, Var).
— Smetana, 1965f: 257 (Philonthus; Dalmatia; Haifa; France; Italy; Albania; Greece).
— Coiffait, 1967a: 429 (Philonthus; characters; France; Pyrenees).
— Coiffait, 1974: 198 (Philonthus; characters; France).
— Maachi, 1991: 251 (Philonthus; Morocco).
— Frank, 1982: 42 (Philonthus; list of parasites).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
▴ Distribution: Morocco, France, Italy, Greece, Albania, Israel.
obliviosus Levasseur, 1968: 1387 (Philonthus; Type locality: mont Cameroun).
obscuratus Cameron, 1950, see: Gabrius.
obscuratus Cameron, 1951, see: debiliformis Cameron, 1951.
obscuricollis Bernhauer, 1922, see: paederoides Motschulsky, 1858.
obscuricollis Bernhauer, 1935, see: rubricollis Motschulsky, 1858.
obscuripennis Stephens, 1832, see: micans Gravenhorst, 1802.
obscuripes Brisout, 1863, see: rubripennis Stephens, 1832.
obscurus Fauvel, 1863, see: havaniensis Laporte, 1835.
obscurus Gravenhorst, 1802: 174 (Staphylinus; [Note: Nomen dubium; preoccupied]; Type locality: Brunsvigae).
— Latreille, 1804: 324 (Staphylinus; characters; Germany).
— Curtis, 1829: 25 (Philonthus; catalog; Britain).
— Stephens, 1832: 234 (Philonthus; characters; England).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Stephens, 1839: 396 (Philonthus; characters; England).
— Herman, 2001: 47 (Philonthus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Britain, Germany.
obsoletipennis Bernhauer, 1942c: 362 (Philonthus; Type locality: Mont Cameroun, 1800–2000 m).
obsoletus Eppelsheim, 1895c: 61 (Philonthus; Type locality: Not cited).
— Bernhauer and Schubert, 1914: 348 (Philonthus; catalog).
— Cameron, 1928c: 564 (Philonthus; Sikkim; Tibet).
— Cameron, 1932: 106 (Philonthus; characters; Sikkim; Tibet).
— Scheerpeltz, 1933: 1353 (Philonthus; catalog).
— Scheerpeltz, 1976: 35 (Philonthus; Nepal).
— Scheerpeltz, 1976a: 88 (Philonthus; Nepal).
— Schillhammer, 1996a: 82 (Philonthus; synonyms).
▴ Distribution: China, Nepal, India, Bhutan, Myanmar.
frontalis Schubert, 1908: 616 (Philonthus; [preoccupied]; Type locality: Kulu, Himalaya, ca. 2000 m).
— Bernhauer and Schubert, 1914: 339 (Philonthus; catalog).
— Cameron, 1932: 107 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1343 (Philonthus; catalog).
— Scheerpeltz, 1965: 107 (Philonthus; Burma).
— Schillhammer, 1991: 54 (Philonthus; synonym of obsoletus).
— Schillhammer, 1996a: 82 (Philonthus; synonym of obsoletus).
bhutanicus Coiffait, 1977b: 224 (Philonthus; Type locality: Bhutan: Gogona, 3100 m).
analokus Coiffait, 1982: 63 (Philonthus; Type locality: Nepal: Dampelek, près de Jumla).
— Schillhammer, 1994: 74 (Philonthus; synonym of obsoletus).
— Schillhammer, 1996a: 82 (Philonthus; synonym of obsoletus).
ocaleoides J. Sahlberg, 1908, see: Rabigus.
occidentalis Horn, 1884: 200 (Philonthus; Type locality: Wyoming; [Note: See lectotype designation by Smetana, 1995e]. Also cited from Colorado; Utah; Indian Territory;).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Hatch, 1957: 184 (Philonthus; characters; British Columbia; Washington; Oregon; Idaho).
— Smetana, 1995e: 344 (Philonthus; lectotype designation: Wy. [Wyoming]; characters; notes; Alberta; British Columbia; Manitoba; Northwest Territories; Ontario; Saskatchewan; Arizona; California; Colorado; Idaho; Montana; Nevada; New Mexico; North Dakota; Oregon; South Dakota; Utah; Washington; Wyoming).
▴ Distribution: Canada, USA.
occipitalis Fauvel, 1879: 103 (Philonthus; Type locality: Nouvelle Guinée, Fly River).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Last, 1968: 364 (Philonthus; New Guinea).
— Last, 1987: 32 (Philonthus; Papua New Guinea).
▴ Distribution: New Guinea.
occultus Sharp, 1885: 407 (Philonthus; Type locality: Guatemala).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Frank, 1982: 42 (Philonthus; list of parasites).
▴ Distribution: Guatemala.
oceanus Hatch, 1957, see: Gabrius.
ochricornis Tottenham, 1962: 215, 216, 221 (Philonthus; Type locality: Not cited, presumably Africa).
ochrigonalis Tottenham, 1962: 156 (Philonthus; Type locality: S. Sudan: 18 m. E. of Yirol).
ochripennis Cameron, 1929: 63 (Philonthus; Type locality: Haut Uele: Moto).
— Scheerpeltz, 1933: 1354 (Philonthus; catalog).
— Tottenham, 1949b: 332 (Philonthus; characters).
▴ Distribution: Congo.
ochripennis Gerhardt, 1910 [Note: Nomen nudum], see: concinnus Gravenhorst, 1802.
ochromerus Sharp, 1885: 406 (Philonthus; Type locality: Mexico: Cordova; Guatemala: near the city, San Geronimo, Zapote, Capetillo, Calderas, Cerro Zunil, Lanquin, Senahu and San Joaquin in Vera Paz; Panama, Bugaba, Volcan de Chiriqui 2000 to 4000 feet).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Frank, 1982: 42 (Philonthus; list of parasites).
▴ Distribution: Mexico, Guatemala, Panama.
ochropus Gravenhorst, 1802, see: concinnus Gravenhorst, 1802.
octopunctus Bernhauer, 1928g: 112 (Philonthus; Type locality: Belgisch-Kongo: Magumbe Tshela; Tengo Katanta, Manyema).
— Scheerpeltz, 1933: 1354 (Philonthus; cited as octopunctatus; catalog).
— Cameron, 1950g: 36 (Philonthus; cited as octopunctatus; Belgian Congo).
▴ Distribution: Congo.
ocularis Bernhauer, 1928g: 107 (Philonthus; Type locality: Belg.-Kongo: Haut-Uele: Watsa, Moto).
oculatus Bernhauer, 1902b: 28 (Philonthus; Type locality: Negombo).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Bernhauer, 1922a: 151 (Philonthus; Indochina; Cambodia).
— Cameron, 1932: 143 (Philonthus; characters; Ceylon).
— Scheerpeltz, 1933: 1354 (Philonthus; catalog).
▴ Distribution: Sri Lanka, Cambodia, Vietnam.
ocypoides Schubert, 1911: 22 (Philonthus; Type locality: Nguelo, Usambara).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Tottenham, 1949b: 361 (Philonthus; characters).
— Tottenham, 1953a: 141 (Philonthus; characters in key).
— Tottenham, 1961: 204 (Philonthus; Senegal).
▴ Distribution: Tanzania, Senegal.
oebalus Tottenham, 1953: 147 (Philonthus; Type locality: Caucasus: Armenien Geb.).
— Smetana, 1955: 207 (Philonthus; characters; notes; Hungary).
— Smetana, 1958: 215 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 164 (Philonthus; characters).
— Lohse, 1964: 184 (Philonthus; subgenus Philonthus; characters; Slovakia).
— Coiffait, 1967a: 425 (Philonthus; synonym of viridipennis).
— Coiffait, 1974: 202 (Philonthus; synonym of viridipennis).
— Lucht, 1987: 105 (Philonthus; checklist; Czechoslovakia).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; cited as valid species; Slovakia).
▴ Distribution: Armenia, Hungary, Slovakia.
oenotrus Smetana, 1991a: 228 (Philonthus; Type locality: Mexico: Chiapas, Mt. Tzontehuitz, Chis. 9500 ft.).
ohizumi R. Dvořák, 1958: 136 (Philonthus; Type locality: Japan: Tokyo, Ohizumi).
— Nakane and Sawada, 1960: A124 (Philonthus; cited as ohizumii; Yakushima Island).
— Watanabe and Shibata, 1972: 64 (Philonthus; cited as ohizumii; Yaku-shima Island).
▴ Distribution: Japan.
oklahomensis Hatch and Ortenburger, 1930, see: hepaticus Erichson, 1840.
oligostigma Leinberg, 1900, see: carbonarius Gravenhorst, 1802.
olfactorius Gistel, 1857: 19 (Philonthus; [Note: Nomen dubium]; Type locality: Styria).
onthomanes Gistel, 1857: 84 (Philonthus; [Note: Nomen dubium]; Type locality: Bavaria).
opacigaster Bierig, 1940a: 143 (Philonthus; subgenus Gabrius; Type locality: Panamá: Volcán de Chiriqui, 1500 m).
opacinus Scheerpeltz, 1933, see: quisquiliarius Gyllenhal, 1810.
opacipennis Notman, 1919: 135 (Philonthus; Type locality: Mooers, Clinton Co., New York).
— Scheerpeltz, 1933: 1354 (Philonthus; catalog).
— Smetana, 1965: 26 (Philonthus; subgenus Philonthus; characters; Nova Scotia; Newfoundland).
— Smetana, 1995e: 332 (Philonthus; characters; notes; Alberta; British Columbia; Manitoba; New Brunswick; Newfoundland; Northwest Territories; Nova Scotia; Ontario; Quebec; Saskatchewan; Alaska; Connecticut; Illinois; Iowa; Maine; Massachusetts; Michigan; New Hampshire; New Jersey; New York; Ohio; Pennsylvania; West Virginia; Wisconsin; Wyoming).
— Downie and Arnett, 1996: 380 (Philonthus; characters; USA).
▴ Distribution: Canada, USA.
opaciventris Fauvel, 1907, see: xanthoraphis Eppelsheim, 1895.
opacus Gravenhorst, 1802, see: varians Paykull, 1789.
opacus LeConte, 1863, see: Cafius.
opacus Gerhardt, 1910, see: quisquiliarius Gyllenhal, 1810.
ophthalmicus Bernhauer, 1941c: 288 [= 1951: 278] (Philonthus; Type locality: Süd Peru: Ica).
optatus Sharp, 1885: 427 (Philonthus; Type locality: Mexico, Guanajuato).
opulentus Erichson, 1840, see: Gastrisus.
orbus Kiesenwetter, 1850, see: Neobisnius.
orcinus Tottenham, 1949b: 353 (Philonthus; Type locality: Kivu: Lulenga).
oreophilus Fauvel, 1877: 265 (Philonthus; Type locality: Australie, Cap York, Somerset).
— Fauvel, 1878g: 548 (Philonthus).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Bernhauer, 1920: 16 (Philonthus; Australia).
— Cameron, 1931: 363 (Philonthus; New Guinea).
— Scheerpeltz, 1933: 1354 (Philonthus; catalog).
— Frank, 1982: 42 (Philonthus; list of parasites).
▴ Distribution: Australia, New Guinea.
oribates Jeannel and Paulian, 1945, see: Gabronthus.
orientalis Cameron, 1937: 8 (Philonthus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft).
orientis Cameron, 1946a: 685 (Philonthus; Type locality: Siam: Renong).
ornatipennis Reitter, 1901, see: ustulatus Fauvel, 1875.
ornatus Blackburn, 1888b: 47 (Philonthus; Type locality: South Australia).
— Blackburn, 1896: 262 (Philonthus; Australia).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Lea, 1925: 220 (Philonthus; characters; Australia).
▴ Distribution: Australia.
orphanulus Erichson, 1840, see: Erichsonius.
orphanus Erichson, 1840: 452 (Philonthus; Type locality: Caracas Columbiae [= Venezuela]; [Note: See lectotype designation by Smetana, 1992d]).
— Fauvel, 1891: 113 (Philonthus; synonym of vilis).
— Bernhauer and Schubert, 1914: 360 (Philonthus; variety of vilis).
— Blackwelder, 1943: 401 (Philonthus; synonym of hepaticus).
— Smetana, 1992d: 135 (Philonthus; characters; lectotype designation: Caracas; valid species; Venezuela).
▴ Distribution: Venezuela.
orphanus Erichson, 1840, see: Erichsonius.
orroriensis Cameron, 1937, see: Hesperus.
osseticus Kolenati, 1846, see: Gabrius.
ottawensis Casey, 1915, see: Gabrius.
ottoi Herman, 2001: 47 (Philonthus; replacement name for trapeziceps Scheerpeltz, 1965).
trapeziceps Scheerpeltz, 1965: 213 (Philonthus; [preoccupied]; Type locality: N.E. Burma, Kambaiti, 2000 m).
ovaliceps Fall, 1930, see: Gabrius.
ovaticeps Sharp, 1885: 421 (Philonthus; Type locality: Mexico, Las Vigas).
ovatus Bernhauer, 1942c: 363 (Philonthus; Type locality: Monts Bambouto, 2300–2500 m).
overlaeti Bernhauer, 1935d: 103 (Philonthus; Type locality: Lulua: Lunene; Kasai).
overlaetianus Bernhauer, 1936a: 22 (Philonthus; Type locality: Lulua: Kapanga).
oviceps Lea, 1925: 222 (Philonthus; Type locality: Queensland: Townsville).
ovicollis Fauvel, 1905, see: ovithorax Bernhauer and Schubert, 1914.
ovithorax Bernhauer and Schubert, 1914: 349 (Philonthus; replacement name for ovicollis Fauvel).
ovicollis Fauvel, 1905b: 179 (Philonthus; [Note: Preoccupied at the time it was replaced by Bernhauer and Schubert by a Broun name described in Staphylinus, moved to Philonthus and now in Gabrius]; Type locality: Madagascar sud: Andrahomana).
oxyporinus Sharp, 1885, see: Belonuchus.
pachycephalus Nordmann, 1837, see: Bisnius.
pacificus Erichson, 1840, see: Cafius.
paederinus Sachse, 1852, see: Bisnius.
paederoides Motschulsky, 1858: 662 (Gabrius; Type locality: Indes orientales).
— Fauvel, 1886b: 150 (Philonthus; Philippines; Ceylon; East Indies; Siam; Cochinchine).
— Bernhauer, 1902b: 26 (Philonthus; Ceylon).
— Fauvel, 1903: 159 (Philonthus; Ceylon; India; Sikkim; Burma; Siam; Indochina; Cambodia; Tonkin; Philippines; Sumatra).
— Fauvel, 1904a: 59 (Philonthus; Belgaum; Tudia; Nilghiris; Sikkim; Bírmanie; Siam; Indochina; Cambodia; Tonkin; Philippines; Sumatra).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Bernhauer, 1928: 16 (Philonthus; Sumatra).
— Cameron, 1932: 114 (Philonthus; characters; Ceylon; India).
— Scheerpeltz, 1933: 1354 (Philonthus; catalog).
— Cameron, 1933d: 387 (Philonthus; Sumatra).
— Cameron, 1937: 5 (Philonthus; Java).
— Scheerpeltz, 1954: 114 (Philonthus, cited as Phylonthus; Ceylon).
— Scheerpeltz, 1961f: 135 (Philonthus; Ceylon).
— Last, 1961: 306 (Philonthus; Thailand; Cambodia).
— Scheerpeltz, 1976a: 88 (Philonthus; Nepal).
— Coiffait, 1979d: 552 (Philonthus; Afghanistan).
— Coiffait, 1982: 28 (Philonthus; Nepal).
— Coiffait, 1982a: 232 (Philonthus; India).
— Barrion, Litsinger, and Last, 1991: 1078 (Philonthus; associated with rice agroecosystem).
▴ Distribution: Afghanistan, India, Sri Lanka, Nepal, Myanmar, Thailand, Cambodia, Vietnam, Indonesia, Philippines.
bellus Kraatz, 1859: 83 (Philonthus; Type locality: Ceylan).
— Fauvel, 1903: 159 (Philonthus; synonym of paederoides).
— Fauvel, 1904a: 59 (Philonthus; synonym of paederoides).
— Bernhauer and Schubert, 1914: 349 (Philonthus; synonym of paederoides).
— Cameron, 1932: 114 (Philonthus; synonym of paederoides).
— Scheerpeltz, 1976a: 88 (Philonthus; synonym of paederoides).
obscuricollis Bernhauer, 1922a: 151 (Philonthus; variety of paederoides; Type locality: Indochina: Me-Kong).
paederoides LeConte, 1863, see: Neobisnius.
paederomimus Eppelsheim, 1895: 206 (Philonthus; Type locality: Ganale Gudda, Arussi Galla).
— Fauvel, 1907: 42 (Philonthus; Voi; Nairobi; Abyssinia; Gallas).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Rambousek, 1922: 1010 (Philonthus; Ethiopia; British East Africa).
— Scheerpeltz, 1933: 1354 (Philonthus; catalog).
— Tottenham, 1949b: 321 (Philonthus; characters).
— Tottenham, 1962: 187 (Philonthus; characters).
▴ Distribution: Ethiopia, Kenya.
paganettii Linke, 1915: 111 (Philonthus; Type locality: Calabrien Sta. Enfemia).
— Gridelli, 1923: 24, 26 (Philonthus; characters; Calabria).
— Scheerpeltz, 1933: 1343 (Philonthus; synonym of fulvipes).
— Coiffait, 1974: 97 (Paragabrius; characters; Italy).
— Pace and Zanetti, 1977: 301 (Paragabrius; notes; Italy).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
▴ Distribution: Italy.
pagmanensis Coiffait, 1983e: 482 (Philonthus; Type locality: Pagmangebirge, 2300 m).
pahangensis Cameron, 1950d: 17 (Philonthus; Type locality: Pahang: Bentong; Indo-China: Saigon).
pakanus Tottenham, 1962: 169 (Philonthus; Type locality: Natal).
palcoensis Bernhauer, 1941c: 288 [= 1951: 277] (Philonthus; Type locality: Süd Peru: Palco, 2600 m; Tambo).
palleolus Melsheimer, 1844, see: hepaticus Erichson, 1840.
palliatus Gravenhorst, 1806: 53 (Staphylinus; Type locality: America septentrionalis; [Note: See lectotype designation by Smetana, 1995e]).
— Erichson, 1840: 474 (Philonthus; characters; America septentrionalis).
— Horn, 1884: 192 (Philonthus; characters; Canada, the middle States, west to Colorado).
— Blatchley, 1910: 383 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Notman, 1919: 135 (Philonthus; New York).
— Smetana, 1965: 18 (Philonthus; Nova Scotia).
— Smetana, 1995e: 265 (Philonthus; lectotype designation: Am. spt. [America septentrionlis]; characters; Alberta; New Brunswick; Nova Scotia; Ontario; Prince Edward Island; Quebec; Colorado; Connecticut; Illinois; Indiana; Iowa; Kansas; Maine; Maryland; Massachusetts; Michigan; Montana; New Hampshire; New Jersey; New York; Ohio; Pennsylvania; Wisconsin; Wyoming).
— Downie and Arnett, 1996: 378 (Philonthus; characters; USA).
▴ Distribution: Canada, USA.
pallidicauda Bernhauer, 1939g: 228 (Philonthus; subgenus Pseudophilonthus; Type locality: Belgisches Kongo: Lulua: Kapanga; Sandoa).
pallidipennis Fauvel, 1905d: 197 (Philonthus; Type locality: Gabon: Libreville; Monts de Cristal; Loango).
pallidipes Lynch, 1884, see: pallipes Blanchard, 1842.
pallipes Blanchard, 1842: 82 (Philonthus; Type locality: îles du Parana, en face de Corrientes).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Scheerpeltz, 1960c: 114 (Philonthus; Bolivia; Uruguay; Argentina; Paraguay; Brazil; Ecuador).
▴ Distribution: Uruguay, Argentina, Paraguay, Bolivia, Brazil, Ecuador.
depressus Steinheil, 1869: 253 (Philonthus; Type locality: Sierra de Mendoza).
pallidipes Lynch, 1884: 152 (Philonthus; [Note: At the heading for this species, Lynch cited pallidipes Blanchard and listed pallipes Blanchard as a synonym, but it is unclear whether the former name is a misspelling or an emendation]; Type locality: îles du Parana, en face de Corrientes).
palmatus Sharp, 1885: 397 (Philonthus; Type locality: Mexico, Jalapa, Oaxaca; La Parada).
palmi Smetana, 1955, see: Bisnius.
palousi Hatch, 1957, see: Bisnius.
palpalis Sharp, 1876, see: Paederomimus.
palustris Brisout, 1859: ccxxxii (Philonthus; Type locality: Collioure).
— Fauvel, 1869: 489 (Philonthus; synonym of virgo).
— Fauvel, 1874: 260 [= 1874a: 486] (Philonthus; synonym of virgo).
— Bernhauer and Schubert, 1914: 360 (Philonthus; synonym of virgo).
— Jarrige, 1951a: 7, 8 (Philonthus; valid species; characters; France; Sardinia; Sicily; Morocco).
— Coiffait, 1963: 12 (Paragabrius).
— Coiffait, 1974: 95 (Paragabrius; characters; France).
▴ Distribution: Morocco, France, Italy.
pamirensis Sharp, 1878, see: Rabigus.
panelii Scheerpeltz, 1974c: 22 (Philonthus; Type locality: Sudan: Attara).
papuanus Cameron, 1931: 362 (Philonthus; Type locality: Mount Lamington, New Guinea).
— Scheerpeltz, 1933: 1354 (Philonthus; catalog).
— Last, 1968: 370 (Philonthus; New Guinea).
— Last, 1987: 32 (Philonthus; Papua New Guinea; British North Borneo).
▴ Distribution: New Guinea, Borneo.
paradisianus Heer, 1839, see: Quedius.
paradoxus Cameron, 1932: 151 (Philonthus; Type locality: Dehra Dun).
parafrigidus Coiffait, 1963: 27 (Philonthus; Type locality: Alpes Maritimes: Col de la Cayolle).
— Coiffait, 1967a: 400 (Philonthus; characters; Maritime Alps; France).
— Jarrige, 1968b: 74 (Philonthus; France).
— Coiffait, 1974: 255 (Philonthus; characters; Alps).
▴ Distribution: France.
pseudopyrenaeus Coiffait, 1974: 145, 255 (Philonthus; variety of parafrigidus; [Note: Nomen nudum (Article 15.2)]; cited from Haute Savoie, Lac Blanc, 2300 m).
parajaponicus J. Li, 1993: 61 (Philonthus; Type locality: Mount Laoye, Shangzhi, Heilongjiang).
parallelus Kraatz, 1859, see: Cafius.
paramerus Coiffait, 1982a: 255 (Spatulonthus; Type locality: Nepal: Thodung via Those, 3100 m).
parasanguineus Levasseur, 1962: 242 (Philonthus; Type locality: Thies, Sénégal).
parcior Bernhauer, 1928, see: Mentophilonthus.
parcipennis Bernhauer, 1915d: 141 (Philonthus; Type locality: Ostafrika: Kilimandscharo).
parciventris Bernhauer, 1936h: 325 (Philonthus; Type locality: Kenya Colony: Katamayu R.).
— Cameron, 1950g: 36 (Philonthus; Belgian Congo).
— Tottenham, 1962: 179 (Philonthus; characters).
▴ Distribution: Kenya, Congo.
parcus Sharp, 1874, see: Bisnius.
parehmontanus Eichelbaum, 1912: 174 (Philonthus; Type locality: Parehberge 2000 m).
— Eichelbaum, 1913: 130 (Philonthus; Tanzania).
— Scheerpeltz, 1933: 1354 (Philonthus; catalog).
▴ Distribution: Tanzania.
parkeri Cameron, 1932, see: Gabrius.
particeps Tottenham, 1962: 175 (Philonthus; Type locality: Tanganyika Terr.: Kilimanjaro, Bismarck Hügel, 3100 m).
parumpunctatus Erichson, 1840, see: Bisnius.
parustus Tottenham, 1962: 193 (Philonthus; Type locality: S. Africa).
parviceps Kraatz, 1859, see: minutus Boheman, 1848.
parvicornis Gravenhorst, 1802: 23 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).
— Latreille, 1804: 313 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 69 (Staphylinus).
— Nordmann, 1837: 82 (Philonthus; Germany).
— Erichson, 1839a: 466 (Philonthus; synonym of opacus).
— Guérin-Méneville, 1857: 40 (Philonthus; synonym of varians).
— Kraatz, 1857d: 603 (Philonthus; synonym of agilis).
— Ganglbauer, 1895: 454 (Philonthus; synonym of agilis).
— Bernhauer and Schubert, 1914: 328 (Philonthus; synonym of agilis).
— Smetana, 1958: 193 (Philonthus; synonym of agilis).
— Coiffait, 1974: 314 (Spatulonthus; characters; Holarctic region).
— Pace and Zanetti, 1977: 300 (Spatulonthus; Italy).
— Coiffait, 1983: 274 (Philonthus; Kashmir).
— Coiffait, 1983e: 478 (Philonthus; Afghanistan).
— Tóth, 1984: 68 (Philonthus; characters; Hungary).
— Ádám, 1996: 245 (Spatulonthus; valid name; Hungary).
— Assing, Frisch, Kahlen, et al., 1998: 137 (Philonthus; note).
— Marchal and Chardonnet, 2000: 72 (Philonthus; Corsica).
▴ Distribution: Ethiopia, Tunisia, Algeria, Morocco, Canary Islands, Europe, Russia, Turkey, Iran, Turkmenistan, Kazakhstan, Uzbekistan, Afghanistan, “Karakorum region”, Mongolia, China, India, New Guinea, Taiwan, Japan.
agilis Gravenhorst, 1806: 77 (Staphylinus; Type locality: Not cited).
— Gyllenhal, 1810: 341 (Staphylinus; characters; Sweden).
— Zetterstedt, 1828: 77 (Staphylinus; characters; Lapland).
— Curtis, 1829: 26 (Philonthus; catalog; Britain).
— C. Sahlberg, 1830: 328 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 30 (Staphylinus; Finland; Sweden).
— Mannerheim, 1831: 444 (Staphylinus; Finland; Sweden).
— Stephens, 1832: 237 (Philonthus; characters; England).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Nordmann, 1837: 98 (Philonthus; Europe).
— Zetterstedt, 1838: 64 (Staphylinus; characters; habitat; Lapland).
— Erichson, 1839a: 466 (Philonthus; synonym of opacus).
— Erichson, 1839a: 466 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 328 cited opacus variety d Erichson as a synonym of agilis; the variety is unavailable];
— Stephens, 1839: 397 (Philonthus; characters; England).
— Erichson, 1840: 470 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 328 cited varians Erichson as a synonym of agilis; Erichson did not describe the species as new, he attributed it to Paykull]; characters; Europe; South Africa; West Indies).
— Hardy, 1851: 29 (Philonthus; synonym of varians).
— Kraatz, 1857d: 603 (Philonthus; characters; Germany).
— Guérin-Méneville, 1857: 40 (Philonthus; synonym of varians).
— Redtenbacher, 1857: 985 (Philonthus; characters; Austria).
— Kraatz, 1858a: 61 (Philonthus; Crete).
— Thomson, 1860: 165 (Philonthus; characters; Scandinavia).
— Rye, 1864a: 56 (Philonthus; characters; notes; Britain).
— Fauvel, 1869: 488 (Philonthus; synonym of parvicornis).
— Fauvel, 1874: 256 [= 1874a: 482] (Philonthus; race of varians; characters; Barbary; Cyprus; Turkey; Buchara; Japan; Ethiopia; Canary Islands).
— Sharp, 1874: 44 (Philonthus; characters; Japan).
— Redtenbacher, 1874: 217 (Philonthus; characters).
— Seidlitz, 1875: 277 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 478 [= 1877: 334] (Philonthus; characters; France).
— J. Sahlberg, 1876: 21 (Philonthus; Finland).
— Fauvel, 1878: 126 [= 1878a: 46] (Philonthus; variety of varians; Algeria).
— Fauvel, 1886: 56 [= 1886a: 48] (Philonthus; variety of varians; Morocco; Tunisia; Algeria).
— Fowler, 1888: 273 (Philonthus; characters; Britain).
— Eppelsheim, 1892: 334 (Philonthus; Turkestan).
— Ganglbauer, 1895: 454 (Philonthus; characters; Palaearctic region; North America; Africa).
— Fauvel, 1902b: 112 (Philonthus; Barbary; Madeira).
— Poppius, 1909: 25 (Philonthus; subgenus Gefyrobius; Lena Valley, Siberia).
— Reitter, 1909: 130 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 327 (Philonthus; catalog).
— Johansen, 1914: 409 (Philonthus; characters; Denmark).
— Gridelli, 1920a: 17, 20 (Philonthus; characters; Italy).
— Bernhauer, 1922c: 232 (Philonthus; Formosa).
— Bernhauer, 1926g: 2 (Philonthus; Kamchatka region).
— Scheerpeltz, 1933: 1331 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Gridelli, 1934: 74 (Philonthus; Karakoram region).
— Wüsthoff, 1937a: 347 (Philonthus; illustration of aedeagus).
— Bernhauer and Chapman, 1939: 71 (Philonthus; Kenya).
— C. Koch, 1941: 63 (Philonthus; Morocco).
— Cameron, 1949b: 468 (Philonthus; Japan).
— V. Hansen, 1952: 68 (Philonthus; subgenus Philonthus; characters; Denmark).
— Hatch, 1957: 189 (Philonthus; characters; British Columbia; Washington [Note: According to Smetana, 1995e: 229 agilis does not occur in North America]).
— Smetana, 1958: 193 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 158 (Philonthus; characters).
— Scheerpeltz, 1960b: 68 (Philonthus; subgenus Gefyrobius; Afghanistan).
— Smetana, 1963c: 296 (Philonthus; subgenus Philonthus; Mongolia).
— Palm, 1963b: 87 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 183 (Philonthus; subgenus Philonthus; characters; central Europe).
— Scheerpeltz, 1964: 28 (Philonthus; subgenus Spatulonthus; Mongolia).
— Focarile, 1964: 58 (Philonthus; Italy).
— Smetana, 1965e: 174 (Philonthus; Mongolia).
— Horion, 1965: 158 (Philonthus; all Europe).
— Pototskaia, 1966: 158 (Philonthus; larval characters).
— Pototskaia, 1967: 63 (Philonthus; larval characters).
— Smetana, 1967d: 204 (Philonthus; Mongolia).
— Raianu, 1967: 271 (Philonthus; characters; ecological notes; Romania).
— Raianu, 1968a: 466 (Philonthus; Romania).
— Szujecki, 1968a: 730 (Philonthus; Poland).
— Jarrige, 1971: 496 (Spatulonthus; Iran).
— Coiffait, 1974: 314 (Philonthus; synonym of parvicornis).
— Smetana 1975c: 171 (Philonthus; Mongolia).
— Shilov, 1976: 150 (Philonthus; Mongolia).
— Pope, 1977: 29 (Philonthus; Britain).
— Frank, 1982: 38 (Philonthus; list of parasites).
— Last, 1987: 31 (Philonthus; Papua New Guinea).
— Lucht, 1987: 106 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Pilon and Zanetti, 1991: 64 (Philonthus; Italy).
— J. Li, 1993: 37 (Philonthus, cited as Philothus; China).
— Smetana, 1993b: 48 (Philonthus; subgenus Spatulonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 27 (Philonthus; Italy).
— M. Hansen, 1996: 105 (Philonthus; synonym of parvicornis).
— Downie and Arnett, 1996: 378 (Philonthus; [Note: This is a misidentification; according to Smetana, 1995e: 229, agilis does not occur in North America]; characters; USA).
— Ádám, 1996: 245 (Spatulonthus; synonym of agilis).
— Gerend and Braunert, 1997: 198 (Philonthus; notes; Luxembourg; synonym of parvicornis).
— Assing, Frisch, Kahlen, et al., 1998: 137 (Philonthus; synonym of parvicornis).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
parvicornis Fauvel, 1907, see: serenus Herman, 2001.
parvimanus Sharp, 1885, see: hepaticus Erichson, 1840.
parvus Horn, 1884, see: Gabrius.
pauliani Jarrige, 1970: 51 (Philonthus; Type locality: Madagascar: Mt. Tsaratanana, 1500 m).
— Lecoq, 1998: 66 (Philonthus; subgenus Piezarthrus; characters; Madagascar).
▴ Distribution: Madagascar.
pauper Bernhauer, 1916c: 32 (Philonthus; Type locality: West-Columbien: Umgebung von Cali in Caucatale, Alto de las cruces, 2200 m).
pauper Lea, 1931, see: leai Scheerpeltz, 1933.
pauxillus Solsky, 1868a: 133 (Philonthus; Type locality: Mexique).
— Lynch, 1884: 153 (Philonthus; synonym of hepaticus).
— Fauvel, 1891: 113 (Philonthus; synonym of vilis).
— Bernhauer and Schubert, 1914: 341 (Philonthus; synonym of hepaticus).
— Bernhauer and Schubert, 1914: 360 (Philonthus; synonym of vilis).
— Blackwelder, 1943: 401 (Philonthus; synonym of hepaticus).
— Blackwelder, 1944: 133 (Philonthus; synonym of hepaticus).
— Smetana, 1995e: 179 (Philonthus; valid species; characters; notes; Alabama; Florida; North Carolina; Texas; Mexico).
▴ Distribution: Mexico, USA.
pavidus Erichson, 1847: 89 (Philonthus; Type locality: Republica Peruana).
peculiaris Last, 1974, see: rectangulus Sharp, 1874.
pedarius Cameron, 1950g: 37 (Philonthus; variety of affinis; Type locality: Ruanda: Ninda, alt. 2150 m).
— Tottenham, 1956: 225, 276 (Philonthus; valid species; characters; Ruanda).
— Tottenham, 1962: 160 (Philonthus; characters).
▴ Distribution: Rwanda.
pedestris Walker, 1859, see: longicornis Stephens, 1832.
pediculus Nordmann, 1837, see: Quedius.
peliomeroides Cameron, 1934e: 372 (Philonthus; Type locality: Mayumbe: Tshiobo N'Goy).
— Cameron, 1950g: 38 (Philonthus; Belgian Congo).
— Cameron, 1952e: 832 (Philonthus; Dahomey).
— Tottenham, 1962: 188, 189 (Philonthus; characters).
▴ Distribution: Congo, Benin.
peliomerus Kraatz, 1859: 82 (Philonthus; Type locality: Ceylan).
— Fauvel, 1898a: 118 (Philonthus; Seychelles).
— Bernhauer, 1902b: 26 (Philonthus; Ceylon).
— Fauvel, 1904a: 58 (Philonthus; Khandesh; Ceylan; Bírmanie; Java; Seychelles; Saint Helena).
— Bernhauer, 1907: 387 (Philonthus; Japan).
— H. Kolbe, 1910: 20 (Philonthus; Seychelles).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Bernhauer, 1915o: 362 (Philonthus; French Congo).
— Cameron, 1932: 87 (Philonthus; characters; Ceylon; India).
— Scheerpeltz, 1933: 1354 (Philonthus; catalog).
— Cameron, 1933d: 386 (Philonthus; Sumatra).
— Tottenham, 1962: 188 (Philonthus; characters).
— Fagel, 1972: 100 (Philonthus; notes; Sainte Helene).
— Lecoq, 1990: 197 (Philonthus; notes; Mascarene Islands).
— Schillhammer, 1999a: 62 (Philonthus; may be endemic to Sri Lanka and perhaps southern India, other records may be misidentifications but further study is needed).
▴ Distribution: Sainte Helena, French Equatorial Africa, Seychelles, Mascarene Islands, Sri Lanka, India, Myanmar, Indonesia.
flavoterminatus Wollaston, 1877: 26 (Philonthus; Type locality: St. Helena).
— Fauvel, 1904a: 58 (Philonthus; synonym of peliomerus).
— Bernhauer and Schubert, 1914: 350 (Philonthus; synonym of peliomerus).
— Cameron, 1932: 87 (Philonthus; synonym of peliomerus).
— Fagel, 1972: 100 (Philonthus; synonym of peliomerus).
peliomixtus Tottenham, 1962: 190 (Philonthus; Type locality: Natal: Mouth of Umkomaas).
pellax Tottenham, 1955: 170 (Philonthus; Type locality: Kenya: Muguga).
pelzelmayeri Bernhauer, 1939, see: Gabrius.
pendleburyi Cameron, 1930a: 155 (Philonthus; Type locality: Malay Peninsula. Negri Sembilan; Port Dickson).
pansatus Casey, 1915, see: aurulentus Horn, 1884.
pentheri Ganglbauer, 1905, see: Bisnius.
peratriceps Scheerpeltz, 1933: 1355 (Philonthus; replacement name for atriceps Sharp).
atriceps Sharp, 1885: 404 (Philonthus; [preoccupied]; Type locality: Mexico, Jalapa, Las Vigas).
— Bernhauer and Schubert, 1914: 329 (Philonthus; catalog).
— Scheerpeltz, 1933: 1355 (Philonthus; synonym of peratriceps).
— Frank, 1982: 38 (Philonthus; list of parasites).
peregrinoides Scheerpeltz, 1974: 146 (Philonthus; Type locality: Cape Prov.: Upington).
peregrinus Fauvel, 1866e: 315 (Philonthus; Type locality: La Réunion: Bourbon; Mayotte, Sainte-Marie-de-Madagascar).
— Fauvel, 1867b: 65 [= 1868a: 69] (Philonthus; characters; Réunion Islands; Madagascar).
— Fauvel, 1874: 255 [= 1874a: 481] (Philonthus; synonym of longicornis).
— Ganglbauer, 1895: 454 (Philonthus; synonym of longicornis).
— Fauvel, 1898a: 118 (Philonthus; Seychelles).
— Fauvel, 1903e: 241 (Philonthus; Cameroon; tropical and southern Africa; Madagascar; Comoro Islands; Mauritia; Réunion; Seychelles; Sikkim; Kashmir).
— Fauvel, 1904a: 60 (Philonthus; Kashmir; Sikkim; Seychelles; Maurice; Réunion; Madagascar; Comoro Islands; tropical Africa; Cape).
— H. Kolbe, 1910: 19 (Philonthus; Seychelles).
— Eichelbaum, 1913: 130 (Philonthus; Tanzania).
— Bernhauer and Schubert, 1914: 350 (Philonthus; catalog).
— Bernhauer, 1922b: 176 (Philonthus; variety of bisignatus; Seychelles).
— Rambousek, 1922: 1008 (Philonthus; Ethiopia).
— Cameron, 1932: 133 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1355 (Philonthus; catalog).
— Wüsthoff, 1937a: 347 (Philonthus; illustration of aedeagus).
— Bernhauer and Chapman, 1939: 69 (Philonthus; Kenya).
— Bernhauer, 1940: 136 (Philonthus; Uganda).
— Cerruti, 1941 (Philonthus; larval characters).
— Bernhauer, 1942c: 362 (Philonthus; Cameroon; Ivory Coast).
— Tottenham, 1949b: 345 (Philonthus; characters).
— Cameron, 1950g: 33 (Philonthus; Belgian Congo).
— Cameron, 1952e: 832 (Philonthus; Dahomey).
— Tottenham, 1955: 165 (Philonthus; subgenus Spatulonthus; characters; notes; Africa; Seychelles; India).
— Tottenham, 1956: 227 (Philonthus; subgenus Spatulonthus; Ruanda; Urundi).
— Tottenham, 1961: 205 (Philonthus; subgenus Spatulonthus; Senegal).
— Tottenham, 1962: 140 (Philonthus; subgenus Spatulonthus; Tanganyika; Uganda).
— Levasseur, 1966a: 1214 (Spatulonthus; Congo).
— Levasseur, 1968a: 53 (Spatulonthus; Central African Republic).
— Levasseur, 1971: 299 (Spatulonthus; Sierra Leone).
— Scheerpeltz, 1974: 146 (Philonthus; South West Africa; South Africa).
— Scheerpeltz, 1976a: 89 (Philonthus; Nepal).
— Jarrige, 1978: 291 (Spatulonthus; Madagascar).
— Coiffait, 1982: 29 (Philonthus; Nepal).
— Coiffait, 1983: 274 (Philonthus; Kashmir).
— Lecoq, 1990: 203 (Spatulonthus; Mascarene Islands).
▴ Distribution: Senegal, Sierra Leone, Ivory Coast, Benin, Cameroon, Central African Republic, Ethiopia, Uganda, Rwanda, Burundi, Kenya, Tanzania, Congo, Namibia, South Africa, Comoros, Madagascar, Mascarene Islands, Seychelles, India, Nepal.
perexcelsus Tottenham, 1939, see: Gabrius.
perfidiosus Last, 1968: 357 (Philonthus; Type locality: New Guinea: Karimu).
perforatus Fauvel, 1889, see: subvirescens Thomson, 1884.
perfusorius Last, 1968: 356 (Philonthus; Type locality: New Guinea: Wanitabe).
perillustris Last, 1968: 358 (Philonthus; Type locality: New Guinea: Buana, Sarawaged Range, 3000 ft., 20–30 miles north of Lae).
— Last, 1987: 32 (Philonthus; Papua New Guinea; New Ireland; New Britain).
▴ Distribution: New Guinea, New Ireland, New Britain.
peringueyi Tottenham, 1962: 162 (Philonthus; Type locality: Not cited, presumably Africa).
peripateticus Tottenham, 1949b: 303 (Philonthus; subgenus Raucalius; Type locality: Nyassa).
perniger Bernhauer, 1920d: 181 (Philonthus; Type locality: Südliches Ostindien: Madura: Chambaganoor).
— Cameron, 1932: 142 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1355 (Philonthus; catalog).
▴ Distribution: India.
perplexus Fairmaire and Germain, 1861: 431 (Philonthus; Type locality: Santiago).
— Fauvel, 1866a: 345 [= 1866b: 96] (Philonthus; [Note: Misidentified as varians according to Coiffait and Saiz, 1968: 360]; characters; Chile).
— Fauvel, 1869: 489 (Philonthus; synonym of longicornis).
— Fauvel, 1874: 255 [= 1874a: 481] (Philonthus; synonym of longicornis).
— Ganglbauer, 1895: 454 (Philonthus; synonym of longicornis).
— Bernhauer and Schubert, 1914: 345 (Philonthus; synonym of longicornis).
— Cameron, 1932: 127 (Philonthus; synonym of longicornis).
— Blackwelder, 1943: 410 (Philonthus; synonym of longicornis).
— Smetana, 1958: 189 (Philonthus; synonym of longicornis).
— Coiffait and Saiz, 1968: 360 (Spatulonthus; valid species; characters; Chile).
— Saiz, 1971b: 348 (Spatulonthus; notes; Chile).
— Desender and Baert, 1996: 36 (Spatulonthus; Easter Island).
▴ Distribution: Chile, Easter Island.
perroti Jarrige, 1951: 133 (Philonthus; Type locality: Tam-Dao, Tonkin).
persimilis Cameron, 1926b: 358 (Philonthus; Type locality: Siwaliks; Kheri Rau, Song River. Mussorie District; Aglar River, Arni Gad, Keyarkuli, 6000 feet. Simla District; Fagu, 8000 feet).
— Cameron, 1932: 138 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1355 (Philonthus; catalog).
— Coiffait, 1983e: 478 (Philonthus; Afghanistan).
▴ Distribution: India, Afghanistan.
perthenus Bernhauer, 1916a: 5 (Philonthus; Type locality: Westaustralien: Perth).
peruvianus Bernhauer, 1916c: 28 (Philonthus; Type locality: Peru: Chanchamayo).
pervagus Cameron, 1932b: 134 (Philonthus; Type locality: Tshiobo N'Goy, Mayumbe).
— Scheerpeltz, 1933: 1355 (Philonthus; catalog).
— Tottenham, 1962: 230 (Philonthus; characters).
▴ Distribution: Congo.
perversus Horn, 1884: 183 (Philonthus; Type locality: Vancouver).
— Bernhauer and Schubert, 1914: 350 (Philonthus; catalog).
— Hatch, 1957: 196 (Philonthus; characters).
— Smetana, 1995e: 153 (Philonthus; characters; British Columbia; California; Oregon; Washington).
▴ Distribution: Canada, USA.
bucephalus Horn, 1884: 184 (Philonthus; Type locality: California; Vancouver).
— Bernhauer and Schubert, 1914: 331 (Philonthus; catalog).
— Hatch, 1957: 196 (Philonthus; characters; British Columbia; Oregon).
— Smetana, 1995e: 153 (Philonthus; synonym of perversus).
pettiti Horn, 1884: 201 (Philonthus; Type locality: California; [Note: See lectotype designation by Smetana, 1995e]. Also cited from Washington Territory).
— Bernhauer and Schubert, 1914: 350 (Philonthus; catalog).
— Hatch, 1957: 185 (Philonthus; characters; British Columbia; Idaho; Washington; Oregon).
— Smetana, 1995e: 425 (Philonthus; lectotype designation: Cal [California]; characters; notes; British Columbia; California; Nevada; Oregon; Washington).
▴ Distribution: Canada, USA.
phaeopus Stephens, 1832, see: quisquiliarius Gyllenhal, 1810.
phaon Smetana, 1995e: 167 (Philonthus; Type locality: Arizona: Graham Co., Pinaleno Mts., Wet Cn., 6100 ft.).
pharao Saulcy, 1864, see: turbidus Erichson, 1840.
pheres Smetana, 1963, see: renominatus Cameron, 1937.
philippinus Cameron, 1941b: 447 (Philonthus; Type locality: Philippines: Luzon: Balbalan).
phoculus Tottenham, 1949b: 309 (Philonthus; Type locality: Abyssinia).
pholeophilus Kirshenblat, 1950, see: Bisnius.
picatus Stephens, 1832, see: carbonarius Gravenhorst, 1802.
picatus Kirby, 1837, see: sericans Gravenhorst, 1802.
piceatus Nordmann, 1837: 85 (Philonthus; Type locality: Mexico).
— Erichson, 1840: 446 (Philonthus; characters; Mexico).
— Solsky, 1868a: 130 (Philonthus; Mexico).
— Sharp, 1885: 400 (Philonthus; characters; notes; Mexico; Guatemala; Costa Rica; Panama; Texas).
— Bernhauer and Schubert, 1914: 350 (Philonthus; catalog).
— Smetana, 1995e: 135 (Philonthus; characters; Arizona; Kansas; Texas; Mexico).
▴ Distribution: USA, Mexico, Guatemala, Costa Rica, Panama.
iridiventris Solsky, 1868a: 128 (Philonthus; Type locality: Mexique).
— Sharp, 1885: 400 (Philonthus; synonym of piceatus).
— Bernhauer and Schubert, 1914: 350 (Philonthus; synonym of piceatus).
— Smetana, 1995e: 135 (Philonthus; synonym of piceatus).
mexicanus Sharp, 1876b: 430 (Philonthus; Type locality: Mexico City).
— Sharp, 1885: 400 (Philonthus; synonym of piceatus).
— Bernhauer and Schubert, 1914: 350 (Philonthus; synonym of piceatus).
— Smetana, 1995e: 135 (Philonthus; synonym of piceatus).
piceicornis Gridelli, 1920 [Note: Nomen nudum], see: longicornis Stephens, 1832.
piceicoxis Eppelsheim, 1895, see: bisignatus Boheman, 1848.
piceicoxis Gridelli, 1920 [Note: Nomen nudum], see: varians Paykull, 1789.
picicollis Stephens, 1832, see: ventralis Gravenhorst, 1802.
picicornis Horn, 1884, see: Bisnius.
picimanus Ménétriés, 1832, see: rotundicollis Ménétriés, 1832.
picipennis Heer, 1839, see: Quedius.
picipennis Mäklin, 1852, see: Gabrius.
picipes Stephens, 1832, see: atratus Gravenhorst, 1802.
picipes Fauvel, 1875, see: alberti Schillhammer, 2000.
picnocara Gistel, 1857: 61 (Philonthus; [Note: Nomen dubium]; Type locality: Helvetia).
picticollis Fauvel, 1879: 102 (Philonthus; Type locality: Nouvelle Guinée, Amberbaki).
piger Horn, 1884, see: Neobisnius.
pilicornis Chapman, 1939: 69 (Philonthus; subgenus Gefyrobius; Type locality: Kenya Colony: Chaîne de l'Aberdare: maison forestière du Kinangop, versant ouest, alt. 2600 m).
pilipennis W. J. MacLeay, 1873: 140 (Philonthus; Type locality: Gayndah).
— Fauvel, 1877: 264 (Philonthus; characters; Australia).
— Bernhauer and Schubert, 1914: 350 (Philonthus; catalog).
— Bernhauer, 1920: 16 (Philonthus; Australia).
— Cameron, 1931: 363 (Philonthus; New Guinea).
— Scheerpeltz, 1933: 1355 (Philonthus; catalog).
▴ Distribution: Australia, New Guinea.
pilipes Stephens, 1832, see: atratus Gravenhorst, 1802.
pilipes Bierig, 1940, see: Gabrius.
pilosus Last, 1981: 118 (Philonthus; Type locality: Daullo Pass).
pindeus Coiffait, 1976, see: carbonarius Gravenhorst, 1802.
pinguis Tottenham, 1949, see: jurgans Tottenham, 1937.
piochardi Fauvel, 1875a: XXIX [= 1875b: 231] (Philonthus; Type locality: Syrie, Naplouse).
— Bernhauer and Schubert, 1914: 350 (Philonthus; catalog).
— Scheerpeltz, 1933: 1355 (Philonthus; catalog).
— Coiffait, 1967a: 395 (Philonthus; characters).
— Coiffait, 1974: 267 (Philonthus; characters; Palestine; Syria; Asia Minor; Persia).
▴ Distribution: Turkey, Syria, Israel, Iran.
pisciformis Fauvel, 1874, see: Gabrius.
placatus Tottenham, 1955, see: sanamus Tottenham, 1955.
placidus Erichson, 1848, see: Bisnius.
plagiatus Fauvel, 1874, see: Bisnius.
planulatus Sharp, 1885: 407 (Philonthus; Type locality: Panama: Volcan de Chiriqui 3000 to 4000 feet, Peña Blanca 3000 to 4000 feet).
planus Lacordaire, 1835, see: corruscus Gravenhorst, 1802.
planus Last, 1987, see: vertumnus Herman, 2001.
plasoni Bernhauer, 1902: 166 (Philonthus; Type locality: Port Daufin, Madagaskar).
platypterus Sharp, 1885: 412 (Philonthus; Type locality: Guatemala, Cerro Zunil 4000 to 5000 feet).
pluripunctus Cameron, 1937: 14 (Philonthus; Type locality: Java: Bandoeng Dago. Also cited from Lombok).
politulus W. J. MacLeay, 1873, see: Quedius.
politus Linné, 1758: 422 (Staphylinus; Type locality: Europa).
— Linné, 1760: 231 (Staphylinus; characters; Sweden).
— Scopoli, 1763: 100 (Staphylinus; characters; Carniola).
— O. Müller, 1764: 23 (Staphylinus; characters).
— Pontoppidans, 1765: 211 (Staphylinus; Denmark).
— Linné, 1767: 683 (Staphylinus; characters; Europe).
— Ström, 1765: 404 (Staphylinus; characters; Denmark) [= 1767: 363].
— Füessly, 1775: 21 (Staphylinus).
— O. Müller, 1776: 98 (Staphylinus; characters).
— Goeze, 1777: 717 (Staphylinus; characters).
— Schrank, 1781: 231 (Staphylinus; characters; Austria).
— Fabricius, 1781: 335 (Staphylinus; characters; Europe).
— Herbst, 1784: 149 (Staphylinus; Germany).
— Fabricius, 1787: 220 (Staphylinus; characters).
— Paykull, 1789: 42 (Staphylinus; redescription; Sweden).
— Rossi, 1790: 245 (Staphylinus; characters; Italy).
— Villers, 1789: 412 (Staphylinus; characters; France).
— Gmelin, 1790: 2028 (Staphylinus; characters).
— Fabricius, 1793: 524 (Staphylinus; characters).
— Olivier, 1795:(42): 25 (Staphylinus; characters).
— Panzer, 1795: 354 (Staphylinus; [Note: The use of politus here by Panzer may be a misidentification of punctus; see the citations for punctus]; characters; Germany).
— Schrank, 1798: 641 (Staphylinus; characters; notes).
— Cederhjelm, 1798: 118 (Staphylinus; characters).
— Paykull, 1800: 391 (Staphylinus; characters; Sweden).
— Fabricius, 1801: 594 (Staphylinus; characters).
— Turton, 1802: 511 (Staphylinus; characters; Europe).
— Gravenhorst, 1802: 17 (Staphylinus; characters).
— Marsham, 1802: 497 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 351 cited similis Marsham as a synonym of politus; Marsham did not describe the species as new, he attributed it to Paykull]; characters).
— Latreille, 1804: 308 (Staphylinus; characters).
— Gravenhorst, 1806: 85 (Staphylinus).
— Latreille, 1806: 287 (Staphylinus; characters; France).
— Gyllenhal, 1810: 317 (Staphylinus; characters; Sweden).
— Leach, 1815: 91 (Staphylinus; characters).
— Westwood, 1827: 58 (Philonthus; larval description).
— Zetterstedt, 1828: 74 (Staphylinus; characters; Lapland).
— Curtis, 1829: 25 (Philonthus; catalog; Britain).
— C. Sahlberg, 1830: 320 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 28 (Staphylinus; Finland; Sweden; Lapland; Germany).
— Mannerheim, 1831: 442 (Staphylinus; Finland; Sweden; Lapland; Germany).
— Stephens, 1832: 228 (Philonthus; characters; England).
— Ménétriés, 1832: 145 (Staphylinus; Caucasus-Persia region).
— Runde, 1835: 7 (Staphylinus; characters; Germany).
— Lacordaire, 1835: 390, 392 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 351 cited metallicus Boisduval and Lacordaire and atratus Boisduval and Lacordaire as synonyms of politus Linné; Lacordaire did not describe the species as new, he attributed them to Gravenhorst]; characters; France).
— Lacordaire, 1835: 392 (Staphylinus; characters; France).
— Curtis, 1836: pl. 610 (Philonthus; characters; reference to possible larval illustration; Britain).
— Nordmann, 1837: 81 (Philonthus; Europe).
— Holme, 1837: 63 (Philonthus; Scilly Islands).
— Kirby, 1837: 91 (Philonthus; characters; notes; Nova Scotia).
— Westwood, 1838a: 16 (Philonthus; type species).
— Zetterstedt, 1838: 63 (Staphylinus; characters; habitat; Lapland).
— Brullé, 1839: 60 (Staphylinus; Canary Islands).
— Erichson, 1839a: 452 (Philonthus; characters; Germany).
— Heer, 1839: 260 (Philonthus; characters; Switzerland).
— Stephens, 1839: 395 (Philonthus; characters; England).
— Erichson, 1840: 443 (Philonthus; characters; Europe).
— Laporte, 1840: 178 (Staphylinus; characters; France).
— Heer, 1841: 582 (Philonthus; notes).
— Westwood, 1845: 47 (Philonthus; Linné confused several species; characters; notes).
— Kolenati, 1846: 19, 20 (Philonthus; Caucasus; Transcaucasia; Iberia).
— Jacobsen, 1849: 690 (Philonthus; characters; Denmark).
— Ménétriés, 1849: 44 (Philonthus; Turcomanie).
— Hochhuth, 1849: 133 (Philonthus; characters).
— Redtenbacher, 1849: 701 (Philonthus; characters; Austria).
— Küster, 1853a: 52 (Philonthus; characters; Europe).
— Hardy, 1851: 26 (Philonthus; notes; England).
— Fairmaire and Laboulbène, 1856: 518 (Philonthus; characters; France).
— Kraatz, 1857d: 581 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 191 (Philonthus; characters; Austria).
— Kraatz, 1858a: 61 (Philonthus; Greece).
— Peyron, 1858: 425 (Philonthus; Turkey).
— Thomson, 1860: 162 (Philonthus; characters; Scandinavia).
— Hochhuth, 1862: 54 (Philonthus; Russia).
— Fauvel, 1874: 247 [= 1874a: 473] (Philonthus; characters; France; Algeria; Turkey; Caucasus; Transcaucasus; North America).
— Redtenbacher, 1874: 210 (Philonthus; characters; Austria).
— Seidlitz, 1875: 273 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 394 [= 1877: 250] (Philonthus; characters; France).
— J. Sahlberg, 1876: 19 (Philonthus; Finland).
— Fauvel, 1878: 125 [= 1878a: 45] (Philonthus; Algeria).
— Horn, 1884: 186 (Philonthus; characters; North Carolina).
— Fauvel, 1886: 56 [= 1886a: 48] (Philonthus; Algeria).
— Fowler, 1888: 265 (Philonthus; characters; Britain).
— Fauvel, 1902b: 108 (Philonthus; Morocco; Algeria; Tunisia; Azores; Madeira; Salvages).
— Blatchley, 1910: 380 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 350, 356 (Philonthus; [Note: Cited as a valid species and as a synonym of splendens]; catalog).
— Gridelli, 1922a: 175 (Philonthus; Persia).
— Lea, 1925: 220 (Philonthus; Australian introduction).
— Bernhauer, 1926g: 2 (Philonthus; Kamchatka region).
— Portevin, 1929: 360 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1355 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Bernhauer, 1940c: 4 (Philonthus; Azores).
— Paulian, 1941: 224 (Philonthus; larval characters).
— Méquignon, 1942a: 20 (Philonthus; Azores).
— Hinton, 1945: 75 (Philonthus; characters).
— V. Hansen, 1952: 50 (Philonthus; subgenus Philonthus; characters; Denmark).
— Hatch, 1957: 198 (Philonthus; characters; British Columbia; Idaho; Washington; Oregon).
— Scheerpeltz, 1958a: 16 (Philonthus; Turkey).
— Smetana, 1958: 163 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 146 (Philonthus; characters).
— Smetana, 1958b (Philonthus; larval characters).
— Larsson and Gígja, 1959: 83 (Philonthus; biological notes; Iceland).
— Smetana, 1963b: 36 (Philonthus; subgenus Philonthus; Madeira).
— Palm, 1963b: 67 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 177 (Philonthus; subgenus Philonthus; characters; central Europe).
— Scheerpeltz, 1964: 28 (Philonthus; subgenus Philonthus; Mongolia).
— Smetana, 1964d: 74 (Philonthus; subgenus Philonthus; Czechoslovakia).
— Smetana, 1965: 12 (Philonthus; Newfoundland).
— Horion, 1965: 129 (Philonthus; all Europe).
— Pototskaia, 1966: 158 (Philonthus; larval characters).
— Pototskaia, 1967: 63 (Philonthus; larval characters).
— Coiffait, 1967a: 392 (Philonthus; characters).
— Smetana, 1967d: 202 (Philonthus; Mongolia).
— Coiffait and Saiz, 1968: 351, 352, 353 (Philonthus; characters; probably an introduced species; Chile).
— Raianu, 1968a: 462 (Philonthus; Romania).
— Szujecki, 1968a: 727 (Philonthus; Poland).
— Saiz, 1971b: 341 (Philonthus; Chile).
— Scheerpeltz, 1972: 17 (Philonthus; Argentina).
— Bacchus and Hammond, 1972: 63 (Philonthus; lion dung; Britain).
— Bellas, Brown, and Moore, 1974 (Philonthus; defensive secretions).
— Coiffait, 1974: 221 (Philonthus; characters; Holarctic region; Australia; New Zealand).
— Shilov, 1975a: 375 [= 1975b: 97] (Philonthus; Komi region of Russia).
— Smetana 1975c: 169 (Philonthus; Mongolia).
— Shilov, 1976: 148 (Philonthus; Mongolia).
— Pace and Zanetti, 1977: 289 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Bordoni, 1978a: 59 (Philonthus; cave collection; Turkey).
— Frank, 1982: 42 (Philonthus; list of parasites).
— Boller, 1983: 119 (Philonthus; larval characters).
— Tóth, 1984: 48 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Israelson, 1990: 2 (Philonthus; Azores).
— Borges, 1990: Table II (Philonthus; Azores).
— Kuschel, 1990: 26 (Philonthus; introduced, Palaearctic species; winged; habitat; New Zealand).
— Pilon and Zanetti, 1991: 63 (Philonthus; Italy).
— Welch, 1993: 228 (Philonthus; ovariole number and ovary structure).
— J. Li, 1993: 37 (Philonthus; China).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Smetana, 1995e: 73 (Philonthus; characters; Alberta; British Columbia, Manitoba; New Brunswick; Newfoundland; Northwest Territories, Nova Scotia; Ontario; Quebec; Saskatchewan; Arizona; California; Colorado; Connecticut; Georgia; Idaho; Illinois; Indiana; Iowa; Kansas; Kentucky; Maine; Maryland; Massachusetts; Michigan; Minnesota; Nebraska; Nevada; New Hampshire; New Jersey; New Mexico; New York; North Carolina; North Dakota; Ohio; Oregon; Pennsylvania; Rhode Island; South Carolina; South Dakota; Utah; Vermont; Virginia; Washington; West Virginia; Wisconsin; Wyoming).
— Sadler and Dugmore, 1995: 144 (Philonthus; habitat notes; Iceland).
— Hodge and Jones, 1995: 40 (Philonthus; characters; Britain).
— Downie and Arnett, 1996: 376 (Philonthus; characters; USA).
— Bain, 1998: 39, 42, 43 (Philonthus; 17th century remains; introduced; Massachusetts).
— Emberson, 1998: 36 (Philonthus; collecting notes; Chatham Islands).
— Outerelo, Gamarra, and Salgado, 1998: 129 (Philonthus; subgenus Philonthus; collected from cave; Spain; Portugal).
— Schmidt, 1999: 108 (Philonthus; food for adults and larvae used for rearing species; rearing methods).
— Ryabukhin, 1999: 60 (Philonthus; catalog; habitat; distribution; northeastern Russia).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
— Schillhammer, 2000: 122 (Philonthus; characters; Kazakhstan; Mongolia; China; India).
▴ Distribution: Tunisia, Algeria, Morocco, Canary Islands, Madeira, Azores, Iceland, Europe, Russia, Caucasus, Turkey, Iran, Turkmenistan, Kazakhstan, Mongolia, China, India, Canada, USA, Argentina, Chile, New Zealand, Chatham Islands, Australia.
aeneus Rossi, 1790: 249 (Staphylinus; [preoccupied]; Type locality: Etruria).
— Olivier, 1795:(42): 25 (Staphylinus; synonym of politus Linné).
— Gravenhorst, 1802: 17 (Staphylinus; characters).
— Latreille, 1804: 307 (Staphylinus; characters; France; Italy).
— Gravenhorst, 1806: 86 (Staphylinus; characters).
— Gyllenhal, 1810: 314 (Staphylinus; characters; Sweden).
— Zetterstedt, 1828: 73 (Staphylinus; characters; Lapland).
— C. Sahlberg, 1830: 319 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 27 (Staphylinus; Finland; Russia).
— Mannerheim, 1831: 441 (Staphylinus; Finland; Russia).
— Runde, 1835: 6 (Staphylinus; characters; Germany).
— Lacordaire, 1835: 389 (Staphylinus; characters; France).
— Nordmann, 1837: 81 (Philonthus; Sweden; Germany; France; Russia).
— Zetterstedt, 1838: 62 (Staphylinus; characters; habitat; Lapland).
— Heer, 1839: 258 (Philonthus; characters; Switzerland).
— Stephens, 1839: 395 (Philonthus; characters; England).
— Erichson, 1840: 437 (Philonthus; characters; Eurasia; America septentrionalis).
— Erichson, 1840: 444 (Philonthus; synonym of politus).
— Laporte, 1840: 178 (Staphylinus; characters; France).
— Hochhuth, 1849: 133 (Philonthus; characters; Imeretia).
— Redtenbacher, 1849: 701 (Philonthus; characters; Austria).
— LeConte, 1850: 220 (Philonthus; Lake Superior).
— Küster, 1853a: 48 (Philonthus; characters; Europe).
— Hardy, 1851: 25 (Philonthus; notes; England).
— Wollaston, 1854: 580 (Philonthus; characters; notes; Madeira Islands).
— Scriba, 1855: 300 (Philonthus; Venezuela).
— Rosenhauer, 1856: 70 (Philonthus).
— Fairmaire and Laboulbène, 1856: 516 (Philonthus; characters; France).
— Staudinger, 1857: 286 (Tachinus; notes; Iceland).
— Wollaston, 1857: 188 (Philonthus; habitat; Madeira Islands).
— Kraatz, 1857d: 578 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 191 (Philonthus; characters; Austria).
— Thomson, 1860: 157 (Philonthus; characters; Scandinavia).
— Fauvel, 1865a: 296 [= 1865b: 46] (Philonthus; habitat; France).
— Wollaston, 1865: 490 (Philonthus; notes; Madeira Islands; Salvage Islands).
— Crotch, 1867a: 383 (Philonthus; Azores).
— Solsky, 1871c: 348 (Philonthus; notes; Siberia).
— Fauvel, 1874: 216 [= 1874a: 442] (Philonthus; characters; France; Europe; Algeria; Turkey; Caucasus; Siberia; Amur region; Madeira; North America; Tasmania).
— Redtenbacher, 1874: 210 (Philonthus; characters; Austria).
— Seidlitz, 1875: 273 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 370 [= 1877: 226] (Philonthus; characters; larval characters; France).
— J. Sahlberg, 1876: 13 (Philonthus; Finland).
— Fauvel, 1877: 262 (Philonthus; characters; Australia).
— Provancher, 1877: 249 (Philonthus; characters; Quebec).
— Fauvel, 1878: 121 [= 1878a: 41] (Philonthus; Algeria).
— Fauvel, 1878g: 546 (Philonthus; Australia).
— Horn, 1884: 181 (Philonthus; characters).
— Fauvel, 1886: 53 [= 1886a: 45] (Philonthus; Algeria).
— Fowler, 1888: 262 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 444 (Philonthus; characters; Palaearctic and Nearctic regions; New Zealand; Tasmania).
— Fauvel, 1902b: 108 (Philonthus; synonym of politus).
— Sainte-Claire Deville, 1906: 95 (Philonthus; Corsica).
— Fall and Cockerell, 1907: 166 (Philonthus; New Mexico).
— Montandon, 1908: 80 (Philonthus; Romania).
— Poppius, 1909: 23 (Philonthus; Lena Valley, Siberia).
— Reitter, 1909: 125 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 350 (Philonthus; synonym of politus).
— Johansen, 1914: 394 (Philonthus; characters; Denmark).
— Lea, 1925: 220 (Philonthus; synonym of politus).
— Portevin, 1929: 358 (Philonthus; characters; France).
— Aleksandrov, 1934: 152 (Philonthus; China).
— West, 1937: 39 (Philonthus; Faroe Islands).
— Smetana, 1958: 163 (Philonthus; synonym of politus).
— Pototskaia, 1967: 57 (Philonthus; larval characters).
— Smetana, 1967d: 202 (Philonthus; synonym of politus).
— Coiffait and Saiz, 1968: 353 (Philonthus; synonym of politus).
— Coiffait, 1974: 221 (Philonthus; synonym of politus).
— Pope, 1977: 29 (Philonthus; synonym of politus).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; synonym of politus).
— Smetana, 1995e: 73 (Philonthus; synonym of politus).
laticeps Zetterstedt, 1828: 73 (Staphylinus; Type locality: Juckasjervi Lapponiae Tornensis).
— Zetterstedt, 1838: 62 (Staphylinus; characters; habitat; Lapland).
— Erichson, 1840: 437 (Philonthus; synonym of aeneus).
— Kraatz, 1857d: 579 (Philonthus; synonym of aeneus).
— Ganglbauer, 1895: 444 (Philonthus; synonym of aeneus).
— Bernhauer and Schubert, 1914: 351 (Philonthus; synonym of politus).
— Smetana, 1958: 163 (Philonthus; synonym of politus).
— Coiffait, 1974: 221 (Philonthus; synonym of politus).
— Smetana, 1995e: 73 (Philonthus; synonym of politus).
— Schillhammer, 2000: 122 (Philonthus; synonym of politus).
cyanicornis Mannerheim, 1830: 27 (Staphylinus; Type locality: America borealis).
— Mannerheim, 1831: 441 (Staphylinus; Finland).
— Nordmann, 1837: 88 (Philonthus; America septentrionalis).
— Kraatz, 1857d: 579 (Philonthus; synonym of aeneus).
— Ganglbauer, 1895: 444 (Philonthus; synonym of aeneus).
— Bernhauer and Schubert, 1914: 351 (Philonthus; synonym of politus).
— Smetana, 1958: 163 (Philonthus; synonym of politus).
— Coiffait, 1974: 221 (Philonthus; synonym of politus).
— Smetana, 1995e: 73 (Philonthus; synonym of politus).
— Schillhammer, 2000: 122 (Philonthus; synonym of politus).
puncticollis Stephens, 1835: 439 (Bisnius; [preoccupied]; Type locality: Oxton Bogs).
— Curtis, 1829: 25 (Philonthus; nomen nudum; catalog; Britain).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Kirby, 1837: 91 (Philonthus; synonym of politus Linné).
— Hardy, 1851: 31 (Philonthus; synonym of minax).
— G. Waterhouse, 1858: 24 (Philonthus; synonym of aeneus).
— Gemminger and Harold, 1868: 585 (Philonthus; synonym of aeneus).
— Ganglbauer, 1895: 444 (Philonthus; synonym of aeneus).
— Bernhauer and Schubert, 1914: 351 (Philonthus; synonym of politus).
— Smetana, 1958: 163 (Philonthus; synonym of politus).
— Coiffait, 1974: 221 (Philonthus; synonym of politus).
— Smetana, 1995e: 73 (Philonthus; synonym of politus).
— Schillhammer, 2000: 122 (Philonthus; synonym of politus).
mandibularis Kirby, 1837: 92 (Philonthus; Type locality: Not cited).
— Erichson, 1840: 928 (Philonthus; synonym of aeneus).
— Kraatz, 1857d: 579 (Philonthus; synonym of aeneus).
— Horn, 1876: 151 (Philonthus; synonym of aeneus).
— Ganglbauer, 1895: 444 (Philonthus; synonym of aeneus).
— Bernhauer and Schubert, 1914: 351 (Philonthus; synonym of politus).
— Smetana, 1958: 163 (Philonthus; synonym of politus).
— Smetana, 1995e: 73 (Philonthus; synonym of politus).
— Schillhammer, 2000: 122 (Philonthus; synonym of politus).
harrisii Melsheimer, 1844: 35 (Philonthus; Type locality: Pennsylvania).
— LeConte, 1850: 220 (Philonthus; synonym of aeneus).
— Fauvel, 1869: 488 (Philonthus; synonym of aeneus).
— Fauvel, 1874: 216 [= 1874a: 442] (Philonthus; synonym of aeneus).
— Ganglbauer, 1895: 444 (Philonthus; synonym of aeneus).
— Bernhauer and Schubert, 1914: 351 (Philonthus; synonym of politus).
— Smetana, 1958: 163 (Philonthus; synonym of politus).
— Smetana, 1995e: 73 (Philonthus; synonym of politus).
— Schillhammer, 2000: 122 (Philonthus; synonym of politus).
angulicollis Motschulsky, 1860b: 121 (Philonthus; Type locality: Daourie et des bords du fl. Amour).
— Motschulsky, 1859: 491 (Philonthus; Amur River).
— Fauvel, 1874: 216 [= 1874a: 442] (Philonthus; synonym of aeneus).
— Ganglbauer, 1895: 444 (Philonthus; synonym of aeneus).
— Bernhauer and Schubert, 1914: 351 (Philonthus; synonym of politus).
— Smetana, 1958: 163 (Philonthus; synonym of politus).
— Coiffait, 1974: 221 (Philonthus; synonym of politus).
— Smetana, 1995e: 73 (Philonthus; synonym of politus).
— Schillhammer, 2000: 122 (Philonthus; synonym of politus).
amblyterus Olliff, 1887: 502 (Cafius; Type locality: Launceston, Hobart, Tasmania).
— Bernhauer and Schubert, 1914: 361 (Cafius; catalog).
— Lea, 1925: 220 (Philonthus; synonym of politus).
— Scheerpeltz, 1933: 1355 (Philonthus; synonym of politus).
— Smetana, 1958: 163 (Philonthus; synonym of politus).
— Coiffait, 1974: 221 (Philonthus; synonym of politus).
— Smetana, 1995e: 73 (Philonthus; synonym of politus).
— Schillhammer, 2000: 122 (Philonthus; synonym of politus).
laeus Olliff, 1887: 503 (Cafius; Type locality: Sydney, New South Wales; Adelaide, South Australia; Launceston, Tasmania).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Lea, 1925: 220 (Philonthus; synonym of politus).
— Scheerpeltz, 1933: 1355 (Philonthus; synonym of politus).
— Smetana, 1958: 163 (Philonthus; synonym of politus).
— Coiffait, 1974: 221 (Philonthus; synonym of politus).
— Smetana, 1995e: 73 (Philonthus; synonym of politus).
— Schillhammer, 2000: 122 (Philonthus; synonym of politus).
altaicus Coiffait, 1967d: 352 (Philonthus; subgenus Philonthus; subspecies of politus; [preoccupied]; Type locality: Tchouia, km. 511, Mejtoueryk, Altai).
— Coiffait, 1974: 221 (Philonthus; subspecies of politus).
— Schillhammer, 2000: 122 (Philonthus; synonym of politus).
temperei Coiffait, 1987: 497 (Philonthus; subspecies of politus; replacement name for altaicus Coiffait).
pollux Fauvel, 1905b: 178 (Philonthus; Type locality: Diego-Suarez; forêts au nord de Fort Dauphin; [Note: See lectotype designation by Lecoq, 1998]. Also cited from baie d'Antongil; forêt de Moramanga; Tananarive).
— Bernhauer and Schubert, 1914: 351 (Philonthus; catalog).
— Lecoq, 1998: 64 (Philonthus; subgenus Piezarthrus; lectotype designation: Madagascar nord, Diego Suarez; characters; Madagascar).
▴ Distribution: Madagascar.
bidentiformis Tottenham, 1949b: 362 (Philonthus; Type locality: Madagascar).
ponderosus Tottenham, 1955: 165 (Philonthus; subgenus Spatulonthus; Type locality: Natal: Frere. Also cited from Rhodesia).
— Tottenham, 1962: 140 (Philonthus; subgenus Spatulonthus; Uganda).
▴ Distribution: South Africa, Zimbabwe, Uganda.
poephagus Cameron, 1928, see: purpuripennis Reitter, 1887.
poppiusi Bernhauer and Schubert, 1914: 351 (Philonthus; replacement name for grandiceps Poppius).
— Scheerpeltz, 1933: 1356 (Philonthus; catalog).
— Coiffait, 1967a: 415 (Philonthus; characters; Siberia).
— Coiffait, 1974: 293 (Philonthus; characters; Siberia).
▴ Distribution: Russia.
grandiceps Poppius, 1909: 24 (Philonthus; [preoccupied]; Type locality: Tscheljabinsk, W. Sibirien).
populus Last, 1987: 27 (Philonthus; Type locality: Papua New Guinea: New Guinea NE: S. Highlands, Dimifa, S.E. of Mount Giluwe, 2200 m).
porus Tottenham, 1955: 171 (Philonthus; Type locality: Nigeria: Ibadan).
potakus Tottenham, 1956: 290 (Philonthus; subgenus Spatulonthus; Type locality: Ruanda: Contrefort Est Muhavura, 2100 m).
potanini Eppelsheim, 1889, see: Bisnius.
praelongus Gemminger and Harold, 1868, see: Neobisnius.
praetor Tottenham, 1949b: 319 (Philonthus; Type locality: S. Rhodesia: Bulawayo).
upotovus Tottenham, 1962: 173 (Philonthus; variety of praetor; [Note: Nomen nudum (Article 15.2)]; cited from Natal: Nqutu).
pragensis Roubal, 1948 [Note: Nomen nudum], see: corruscus Gravenhorst, 1802.
preangeranus Cameron, 1937: 6 (Philonthus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft).
prescottianus Sharp, 1887: 787 (Philonthus; Type locality: Mexico, Amecameca in Morelos, Salazar).
pressus Last, 1968: 365 (Philonthus; Type locality: New Guinea: Okasa, 4500 ft.).
pretiosus Erichson, 1840, see: Phanolinus.
primarius Last, 1987: 28 (Philonthus; Type locality: Papua New Guinea: New Guinea NE: Mount Piora, 2800 m).
princeps Bernhauer, 1902, see: Rabigus.
prismalis Fauvel, 1891: 114 (Philonthus; Type locality: Colonie Tovar).
proselytus Herman, 2001: 47 (Philonthus; replacement name for tamulus Tottenham).
tamulus Tottenham, 1949b: 358 (Philonthus; [preoccupied]; Type locality: Kivu: Tshibinda).
productus Kraatz, 1859: 89 (Philonthus; Type locality: Ceylan).
— Bernhauer, 1902b: 27 (Philonthus; Ceylon).
— Bernhauer and Schubert, 1914: 351 (Philonthus; catalog).
— Cameron, 1932: 105 (Philonthus; characters; Ceylon; India; Malay Peninsula; S. China).
— Scheerpeltz, 1933: 1356 (Philonthus; catalog).
— Schillhammer, 1999a: 62 (Philonthus).
▴ Distribution: Sri Lanka, India, “Malay Peninsula”, China.
sulciceps Bernhauer, 1931c: 130 (Philonthus; Type locality: Hongkong).
— Scheerpeltz, 1933: 1363 (Philonthus; catalog).
— Schillhammer, 1999a: 62 (Philonthus; synonym of productus).
prolatus Sharp, 1874: 46 (Philonthus; Type locality: Nagasaki).
— Matsumura, 1911: 114 (Philonthus; Japan).
— Bernhauer and Schubert, 1914: 351 (Philonthus; catalog).
— Scheerpeltz, 1933: 1356 (Philonthus; catalog).
— Hayashi, 1993: 71 (Philonthus; characters; Japan).
— Hayashi, 1994: 126 (Philonthus; characters; Japan).
— Nishida, 1994: 73 (Philonthus; Hawaii).
— Nishida, 1997: 62 (Philonthus; Hawaii).
▴ Distribution: Korea, Japan, Hawaii.
chonrima Hayashi, 1993: 74 (Philonthus; subspecies of prolatus; Type locality: Korea: Kwangneung).
ezoensis Hayashi, 1993: 75 (Philonthus; subspecies of prolatus; Type locality: Japan: Hokkaido: Yoneoka, Okushiri Is.).
prolixicornis Sharp, 1885: 424 (Philonthus; Type locality: Guatemala, Torola).
prolixus Erichson, 1840, see: Neobisnius.
promtus Erichson, 1840, see: longicornis Stephens, 1832.
propinquus Sharp, 1876, see: Paederomimus.
propinquus Cameron, 1933, see: Gabrius.
propinquus Kirshenblat, 1950, see: Bisnius.
proportionalis Schubert, 1909: 293 (Philonthus; Type locality: südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d'Orizaba).
protenus Schubert, 1906a: 382 (Philonthus; Type locality: Kashmir, Songam).
— Bernhauer and Schubert, 1914: 351 (Philonthus; catalog).
— Cameron, 1932: 95 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1356 (Philonthus; catalog).
— Scheerpeltz, 1976: 35 (Philonthus; Nepal).
— Coiffait, 1982: 29 (Philonthus; Nepal).
— Schillhammer, 1991: 54 (Philonthus; characters; notes).
▴ Distribution: India, Nepal, China.
signifrons Schubert, 1908: 618 (Philonthus; Type locality: Kulu, Himalaya, ca. 2000 m).
— Bernhauer and Schubert, 1914: 355 (Philonthus; catalog).
— Cameron, 1932: 106 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1361 (Philonthus; catalog).
— Schillhammer, 1991: 54 (Philonthus; synonym of protenus).
simlaensis Bernhauer, 1911: 90 (Philonthus; Type locality: Simla, 7000′; Simla hills: Phagu, 8700′).
— Bernhauer and Schubert, 1914: 355 (Philonthus; catalog).
— Cameron, 1932: 95 (Philonthus; synonym of protenus).
— Scheerpeltz, 1933: 1356 (Philonthus; synonym of protenus).
yatungensis Cameron, 1928c: 564 (Philonthus; Type locality: Tibet: Yatung, 10,000 feet).
— Cameron, 1932: 95 (Philonthus; synonym of protenus).
— Scheerpeltz, 1933: 1356 (Philonthus; synonym of protenus).
protervus Casey, 1915, see: stictus Hausen, 1891.
proximatus Schubert, 1908, see: tractatus Eppelsheim, 1895.
proximus Wollaston, 1857, see: ventralis Gravenhorst, 1802.
proximus Kraatz, 1859, see: simpliciventris Bernhauer, 1933.
pruinosus Erichson, 1840, see: Remus.
pseudabyssinus Tottenham, 1940d: 148 (Philonthus; Type locality: S. Africa: Delagoa Bay; also listed from E. Africa; Mashonaland; Angola; Kenya; Belgian Congo).
— Tottenham, 1949b: 305, 315 (Philonthus).
— Tottenham, 1956: 226 (Philonthus; Ruanda).
— Tottenham, 1962: 164 (Philonthus; characters).
▴ Distribution: South Africa, Zimbabwe, Angola, Rwanda, Kenya.
pseudeustilbus Scheerpeltz, 1965: 226 (Philonthus; Type locality: N.E. Burma, Kambaiti, 2000 m).
pseudocoerulescens Gusmann, 1925, see: atratus Gravenhorst, 1802.
pseudoconvalescens Scheerpeltz, 1965: 215 (Philonthus; Type locality: N.E. Burma, Kambaiti, 2000 m).
pseudogeminus Cameron, 1950d: 19 (Philonthus; Type locality: Kuala Lumpur).
pseudogracilis Reitter, 1909, see: Rabigus.
pseudojaponicus Bernhauer, 1936g: 307 (Philonthus; Type locality: North Alp. Japan: Kamikochi; Jesso: Nemoro).
— Nakane, 1963: A239 (Philonthus; Japan).
— Sawada, 1965: 15 (Philonthus; subgenus Philonthus; characters; Japan).
— J. Li, 1993: 36 (Philonthus; China).
▴ Distribution: Japan, China.
pseudolodes Smetana, 1996b: 220 (Philonthus; replacement name for pseudolus Smetana).
pseudolus Smetana, 1995e: 283 (Philonthus; [preoccupied]; Type locality: Manitoba: Riding Mtn. N. P., Clear Lake, north end. Also cited from Alberta, British Columbia, Ontario, Arizona, California, Colorado).
pseudolus Last, 1989: 878 (Philonthus; Type locality: Papua New Guinea: Kainantu).
pseudolus Smetana, 1995, see: pseudolodes Smetana, 1996.
pseudonigriceps Tottenham, 1962: 229 (Philonthus; Type locality: S. Sudan: Prov. Bahr el Ghazal, 8°30′N, 28°30′E).
pseudonitidus Everts, 1923 [Note: Nomen nudum], see: laminatus Creutzer, 1799.
pseudooculatus Bernhauer, 1928, see: leptocerus Fauvel, 1895.
pseudoparcus Brunne, 1976, see: Bisnius.
pseudopennatus Cameron, 1945, see: Gabrius.
pseudopyrenaeus Coiffait, 1974 [Note: Nomen nudum], see: parafrigidus Coiffait, 1963.
pseudosulcatus Cameron, 1942a: 136 (Philonthus; Type locality: Borneo: Sandakan. Also at Singapore).
pseudotardus Cameron, 1950d: 17 (Philonthus; Type locality: Perak: Larut Hills, 3700 feet).
pseudovarians A. Strand, 1941: 34 (Philonthus; Type locality: Bygdøy bei Oslo; Ullern in V. Aker; Lysaker; Lillestrøm; Kongsberg).
— V. Hansen, 1952: 70 (Philonthus; subgenus Philonthus; characters; Denmark).
— Horion, 1954: 9 (Philonthus; characters; Germany).
— Smetana, 1958: 197 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 160 (Philonthus; characters).
— Palm, 1963b: 88 (Philonthus; characters; Denmark; Norway).
— Lohse, 1964: 183 (Philonthus; subgenus Philonthus; characters; central Europe).
— Smetana, 1964d: 75 (Philonthus; subgenus Philonthus; Czechoslovakia).
— Smetana, 1965f: 255 (Philonthus; notes).
— Horion, 1965: 160 (Philonthus; Norway; Denmark; France; Spain; Italy; Czech Republic; Slovakia; Austria; Germany).
— Coiffait, 1967a: 420 (Philonthus; characters; Europe).
— Raianu, 1968: 115 (Philonthus; characters; ecological notes; Romania).
— Raianu, 1968a: 467 (Philonthus; characters; Romania).
— Szujecki, 1969a: 256 (Philonthus; notes; Poland).
— Coiffait, 1974: 245 (Philonthus; characters; Norway; Europe; Corfu).
— Tóth, 1984: 70 (Philonthus; characters; Hungary).
— Jarrige, 1968b: 74 (Philonthus; France).
— Drugmand, 1987a: 311 (Philonthus; characters; Belgium).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria).
— Pilon and Zanetti, 1991: 64 (Philonthus; Italy).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
▴ Distribution: Europe.
pseudovarius Bernhauer, 1923a: 125 (Philonthus; Type locality: Amurgebiet).
— Scheerpeltz, 1933: 1356 (Philonthus; catalog).
— Shilov, 1976: 150 (Philonthus; Mongolia).
▴ Distribution: Mongolia, “Amur region”.
pseudovittiger Bernhauer, 1939, see: vittiger Fauvel, 1907.
psyllophagus Kirshenblat, 1933, see: Bisnius.
puberulus Horn, 1884: 193 (Philonthus; Type locality: California; [Note: See lectotype designation by Smetana, 1995e]. Also cited from Arizona).
— Bernhauer and Schubert, 1914: 351 (Philonthus; catalog).
— Hatch, 1957: 189 (Philonthus; characters; British Columbia; Idaho; Washington; Oregon).
— Smetana, 1995e: 296 (Philonthus; lectotype designation: Cal [California]; characters; British Columbia; California; Nevada; Oklahoma; Oregon; Utah; Washington).
▴ Distribution: Canada, USA.
pubes Horn, 1884: 192 (Philonthus; Type locality: Thurston Co., Washington Territory; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from Washington Territory and Nevada).
— Bernhauer and Schubert, 1914: 351 (Philonthus; catalog).
— Hatch, 1957: 190 (Philonthus; characters; British Columbia; Washington; Oregon; Idaho).
— Smetana, 1995e: 254 (Philonthus; lectotype designation: Thurston Co., W.T. [Washington Territory]; characters; British Columbai; California; Idaho; Montana; Nevada; Oregon; Washington; Wyoming).
▴ Distribution: Canada, USA.
pubipennis Cameron, 1919, see: industanus Fauvel, 1903.
puella Nordmann, 1837, see: Bisnius.
pugetensis Hatch, 1957, see: Bisnius.
pulchellus Kraatz, 1859, see: Gabronthus.
pulchellus Melsheimer, 1844, see: Bisnius.
pulcher Bernhauer, 1908d: 335 (Philonthus; Type locality: Peru).
pulcherrimus Bernhauer, 1915d: 139 (Philonthus; Type locality: Abessinien: Dire Daoua).
pulchricolor Bernhauer, 1923c: 63 (Philonthus; replacement name for quadricolor Bernhauer).
quadricolor Bernhauer, 1916c: 30 (Philonthus; [preoccupied]; Type locality: West-Columbien: Umgebung von Cali im Caucatal, Rio Aquatal, 1800 m).
pullus Nordmann, 1837, see: Rabigus.
pullus Runde, 1835, see: micans Gravenhorst, 1802.
pumilio Casey, 1915, see: Rabigus.
pumilus Sharp, 1874, see: Neobisnius.
punctatellus Heer, 1839, see: Quedius.
punctatellus Horn, 1884, see: Gabrius.
punctatissimus Schubert, 1908, see: aeneipennis Boheman, 1858.
punctativentris Bernhauer, 1933a: 41 (Philonthus; Type locality: Tatsienlu-Kiulung).
punctatus Fauvel, 1874, see: punctus Gravenhorst, 1802.
puncticeps Cameron, 1926b: 352 (Philonthus; Type locality: Simla Hills; Gahan, 7000 feet).
— Cameron, 1932: 90 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1357 (Philonthus; catalog).
▴ Distribution: India.
puncticollis Stephens, 1832: 228 (Philonthus; [Note: Nomen dubium]; Type locality: London; Norfolk; Suffolk; Devonshire).
puncticollis Stephens, 1835, see: politus Linné, 1758.
puncticollis Boheman, 1858, see: Cafius.
puncticollis Broun, 1894, see: insularis Bernhauer and Schubert, 1916.
punctifer Nordmann, 1837: 107 (Philonthus; Type locality: Mexico).
— Erichson, 1840: 498 (Philonthus; characters; Mexico).
— Sharp, 1885: 427 (Philonthus; notes; Mexico; Costa Rica).
— Bernhauer and Schubert, 1914: 352 (Philonthus; catalog).
▴ Distribution: Mexico, Costa Rica.
punctifrons Cameron, 1926b: 352 (Philonthus; Type locality: Mussorie District; Keyarkuli. Chakrata District; Chulli Khud, 8000 feet).
— Cameron, 1932: 89 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1357 (Philonthus; catalog).
▴ Distribution: India.
punctipennis Wollaston, 1857, see: wollastoni Scheerpeltz, 1933.
punctiventris Stephens, 1832, see: varians Paykull, 1789.
punctiventris Kraatz, 1857, see: temporalis Mulsant and Rey, 1853.
punctus Gravenhorst, 1802: 20 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).
— Panzer, 1796c: 7 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 352 cited politus Panzer as a synonym of punctus Gravenhorst; Panzer did not describe the species as new, he attributed it to Linné]; illustration).
— Panzer, 1808: 7 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 352 cited politus Panzer as a synonym of punctus Gravenhorst; Panzer did not describe the species as new, he attributed it to Linné]; characters; illustrations).
— Gravenhorst, 1806: 85 (Staphylinus).
— Gyllenhal, 1810: 346 (Staphylinus; characters; Sweden).
— Gyllenhal, 1810: 346 (Staphylinus; [Note: Cited as politus Panzer which is a misidentification of punctus Gravenhorst]).
— Curtis, 1829: 26 (Philonthus; catalog; Britain).
— C. Sahlberg, 1830: 330 (Staphylinus; characters; Finland).
— C. Sahlberg, 1830: 330 (Staphylinus; [Note: Cited as politus Panzer which is a misidentification of punctus Gravenhorst]).
— Mannerheim, 1830: 31 (Staphylinus; Finland).
— Mannerheim, 1831: 445 (Staphylinus; Finland).
— Stephens, 1832: 240 (Philonthus; characters; England).
— Runde, 1835: 10 (Staphylinus; characters; Germany).
— Lacordaire, 1835: 393 (Staphylinus; characters; France).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Erichson, 1839a: 477 (Philonthus; characters; Germany).
— Erichson, 1839a: 477 (Philonthus; [Note: Cited as politus Panzer which is a misidentification of punctus Gravenhorst]).
— Heer, 1839: 271 (Philonthus; characters; Switzerland).
— Stephens, 1839: 398 (Philonthus; characters; England).
— Erichson, 1840: 498 (Philonthus; characters; Germany; Sardinia).
— Erichson, 1840: 498 (Philonthus; [Note: Cited as politus Panzer which is a misidentification of punctus Gravenhorst]).
— Laporte, 1840: 178 (Staphylinus; characters; France).
— Jacobsen, 1849: 704 (Philonthus; characters; Denmark).
— Hochhuth, 1849: 145 (Philonthus; characters).
— Redtenbacher, 1849: 707 (Philonthus; characters; Austria).
— Redtenbacher, 1849: 707 (Philonthus; [Note: Cited as politus Panzer which is a misidentification of punctus Gravenhorst]).
— Fairmaire and Laboulbène, 1856: 529 (Philonthus; characters; France).
— Kraatz, 1857d: 618 (Philonthus; characters; Germany).
— Kraatz, 1857d: 618 (Philonthus; [Note: Cited as politus Panzer which is a misidentification of punctus Gravenhorst]).
— Redtenbacher, 1857: 197 (Philonthus; characters; Austria).
— Peyron, 1858: 427 (Philonthus; Turkey).
— Thomson, 1860: 161 (Philonthus; characters; Scandinavia).
— Hochhuth, 1862: 64 (Philonthus; Russia).
— Hochhuth, 1872: 144 (Philonthus; notes; Ukraine).
— Redtenbacher, 1874: 215 (Philonthus; characters; Austria).
— Seidlitz, 1875: 278 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 515 [= 1877: 371] (Philonthus; characters; France).
— Fauvel, 1886: 53 [= 1886a: 45] (Philonthus; Algeria).
— Fowler, 1888: 280 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 458 (Philonthus; characters; Europe; Mediterranean region).
— Fauvel, 1902b: 108 (Philonthus; Morocco; Algeria; Tunisia).
— Sainte-Claire Deville, 1906: 99 (Philonthus; Corsica).
— Poppius, 1909: 27 (Philonthus; subgenus Gefyrobius; Lena Valley, Siberia).
— Reitter, 1909: 131 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 352 (Philonthus; catalog).
— Bernhauer and Schubert, 1914: 352 (Philonthus; [Note: Cited as politus Panzer which is a misidentification of punctus Gravenhorst]).
— Johansen, 1914: 415 (Philonthus; characters; Denmark).
— Portevin, 1929: 365 (Philonthus; cited as punctatus; characters; France).
— Scheerpeltz, 1933: 1357 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— V. Hansen, 1952: 78 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1953: 119 (Philonthus; Turkey).
— Smetana, 1958: 224 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1958: 224 (Philonthus; [Note: Cited as politus Panzer which is a misidentification of punctus Gravenhorst]).
— Smetana, 1959: 170 (Philonthus; characters).
— Scheerpeltz, 1960d: 136 (Philonthus; Portugal).
— Palm, 1963b: 97 (Philonthus; characters; Sweden; Denmark; Finland).
— Lohse, 1964: 186 (Philonthus; subgenus Philonthus; characters; central Europe).
— Focarile, 1964: 59 (Philonthus; Italy).
— Raianu, 1964: 104 (Philonthus; Romania).
— Horion, 1965: 175 (Philonthus; Russia; Finland; Sweden; Britain; Denmark; Holland; France; Switzerland; Spain; Corsica; Sardinia; Sicily; Italy; Dalmatia; Hercegovina; Serbia; Albania; Macedonia; Bulgaria; Romania; Hungary; Czechoslovakia; Poland; Germany; Austria).
— Coiffait, 1967a: 432 (Philonthus; characters; Siberia; Turkestan).
— Raianu, 1968a: 468 (Philonthus; Romania).
— Coiffait, 1974: 268 (Philonthus; characters; Europe; Britain; Scandinavia; North Africa; Caucasus; Turkestan; Siberia).
— Coiffait, 1974: 268 (Philonthus; [Note: Cited as politus Panzer which is a misidentification of punctus Gravenhorst]).
— Shilov, 1975a: 376 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Pace and Zanetti, 1977: 299 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Frank, 1982: 43 (Philonthus; list of parasites).
— Tóth, 1984: 78 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Bordoni, 1995: 88 (Philonthus; ecological notes; Italy).
— Ádám, 1996: 236 (Paralionthus).
— Sterrenburg and Schülke, 1997: 18 (Philonthus; Netherlands).
— Owen, 1997a: 149 (Philonthus; collecting sites; Ireland).
▴ Distribution: Tunisia, Algeria, Morocco, Europe, Russia, Caucasus, Turkey, Iran, “Turkestan”.
multipunctatus Mannerheim, 1830: 31 (Staphylinus; [preoccupied]; Type locality: Scania Sveciae: Esperöd).
— Mannerheim, 1831: 445 (Staphylinus; Sweden).
— Ménétriés, 1832: 146 (Staphylinus; Caucasus-Persia region).
— Nordmann, 1837: 106 (Philonthus; characters; Germany; Sweden).
— Faldermann, 1835: 131 (Staphylinus; characters; Persia-Armenia).
— Erichson, 1839a: 477 (Philonthus; synonym of punctus).
— Heer, 1839: 272 (Philonthus; synonym of punctus).
— Erichson, 1840: 498 (Philonthus; synonym of punctus).
— Hochhuth, 1849: 145 (Philonthus; synonym of punctus).
— Redtenbacher, 1849: 707 (Philonthus; synonym of punctus).
— Motschulsky, 1858d: 69 (Philonthus; synonym of punctus).
— Kraatz, 1857d: 618 (Philonthus; synonym of punctus).
— Fauvel, 1874: 219 [= 1874a: 445] (Philonthus; synonym of punctus).
— Ganglbauer, 1895: 458 (Philonthus; synonym of punctus).
— Bernhauer and Schubert, 1914: 352 (Philonthus; synonym of punctus).
— Smetana, 1958: 224 (Philonthus; synonym of punctus).
— Coiffait, 1974: 268 (Philonthus; synonym of punctus).
punctatus Fauvel, 1874: 219 [= 1874a: 445] (Philonthus; emendation of punctus).
— Mulsant and Rey, 1876: 516 [= 1877: 372] (Philonthus; synonym of punctus).
— Fauvel, 1878: 122 [= 1878a: 42] (Philonthus; Algeria).
— Ganglbauer, 1895: 458 (Philonthus; synonym of punctus).
— Bernhauer and Schubert, 1914: 352 (Philonthus; synonym of punctus).
— Scheerpeltz, 1933: 1357 (Philonthus; synonym of punctus).
— Smetana, 1958: 224 (Philonthus; synonym of punctus).
— Coiffait, 1974: 268 (Philonthus; synonym of punctus).
rapillyi Jarrige, 1971: 487 (Philonthus; subspecies of punctus; Type locality: Sud Iran, Dacht Argen).
purkynei Smetana, 1963c: 297 (Philonthus; subgenus Rabigus; Type locality: Nord-Mongolei: Karakorum).
— Smetana, 1967d: 209 (Philonthus; subgenus Rabigus; Mongolia).
— Coiffait, 1974: 104 (Rabigus; characters; Mongilia).
— Smetana 1975c: 172 (Philonthus; Mongolia).
— Shilov, 1976: 153 (Philonthus; Mongolia).
▴ Distribution: Mongolia.
purpuripennis Reitter, 1887: 214 (Philonthus; Type locality: Höhe von 13–14,000 Fuss zwischen dem Fluss Bytchu und dem Blauen Fluss; Blauer Fluss, 12,500–13,000 Fuss; Gebirge an den Flüssen Kong-tchum-tchu, Tchum-tcha-uma und By-djun in der Höhe von 13 bis 14,000 Fuss; Thal des Bytchu, 13,000 Fuss; Thal des Tala-tchu).
— Bernhauer and Schubert, 1914: 352 (Philonthus; catalog).
— Scheerpeltz, 1933: 1357 (Philonthus; catalog).
— Coiffait, 1967a: 390 (Philonthus; characters; central Asia; China).
— Coiffait, 1974: 269 (Philonthus; characters; central Asia).
— Schillhammer, 1999a: 62 (Philonthus).
— Schillhammer, 2000: 142 (Philonthus; characters; Nepal; India; China).
▴ Distribution: China, Nepal, India.
poephagus Cameron, 1928c: 562 (Philonthus; Type locality: Tibet: Rongshar Valley, 15,000 feet; [Note: See lectotype designation by Schillhammer, 2000]. Also originally cited from Ling-ka, 14,000 feet; Kampa Dzong, 14,500 feet).
— Cameron, 1932: 79 (Philonthus; characters; Tibet).
— Scheerpeltz, 1933: 1355 (Philonthus; catalog).
— Coiffait, 1982: 30 (Philonthus; Nepal).
— Biswas and Biswas, 1985: 145 (Philonthus; India).
— Schillhammer, 1999a: 62 (Philonthus; synonym of purpuripennis).
— Schillhammer, 2000: 142 (Philonthus; lectotype designation: Tibet, Rongshar Valley; synonym of purpuripennis).
magnificus Bernhauer, 1933a: 39 (Philonthus; Type locality: Tatsienlu-Kiulung).
— Smetana, 1963e: 72 (Philonthus; synonym of purpuripennis).
— Coiffait, 1974: 269 (Philonthus; synonym of purpuripennis).
— Schillhammer, 2000: 142 (Philonthus; synonym of purpuripennis).
coiffaiti Hromádka, 1992: 97 (Philonthus; [preoccupied]; Type locality: Nepal, Managn [Note: Correct spelling is Manang]).
— Schillhammer, 1999a: 62 (Philonthus; synonym of purpuripennis).
— Schillhammer, 2000: 142 (Philonthus; synonym of purpuripennis).
pusillus Heer, 1839, see: Gabrius.
pustulatus Bernhauer, 1915, see: bisignatus Boheman, 1848.
putiphar Saulcy, 1864, see: Neobisnius.
putridarius Gistel, 1857: 19 (Philonthus; [Note: Nomen dubium]; Type locality: Styriae).
pygmaeus Kraatz, 1859, see: Gabronthus.
pyrenaeus Kiesenwetter, 1850: 219 (Philonthus; Type locality: Pyrenaei).
— Kiesenwetter, 1851a: 414 (Philonthus; characters; Pyrenees).
— Fairmaire and Laboulbène, 1856: 519 (Philonthus; characters; France).
— Kraatz, 1857d: 574 (Philonthus; characters).
— Fauvel, 1865a: 296 [= 1865b: 46] (Philonthus; habitat; France).
— Fauvel, 1874: 236 [= 1874a: 462] (Philonthus; synonym of frigidus).
— Mulsant and Rey, 1876: 405 [= 1877: 261] (Philonthus; characters; France).
— Ganglbauer, 1895: 448 (Philonthus; synonym of frigidus).
— Bernhauer and Schubert, 1914: 339 (Philonthus; variety of frigidus).
— Smetana, 1958: 186 (Philonthus; synonym of frigidus).
— Smetana, 1959: 150 (Philonthus; aberration of frigidus; characters).
— Coiffait, 1963: 25 (Philonthus; valid species; characters; Pyrenees).
— Coiffait, 1967a: 400 (Philonthus; characters; Pyrenees).
— Coiffait, 1974: 254 (Philonthus; characters; Pyrenees).
— Outerelo, 1976: 266 (Philonthus; Spain).
▴ Distribution: France, Spain.
pyropterus Kraatz, 1859, see: hepaticus Erichson, 1840.
quadraticeps Boheman, 1858: 28 (Philonthus; Type locality: Buenos Ayres).
— Lynch, 1884: 150 (Philonthus; characters; Argentina).
— Bernhauer and Schubert, 1914: 352 (Philonthus; catalog).
— Frank, 1982: 43 (Philonthus; list of parasites).
▴ Distribution: Argentina.
quadraticollis Sharp, 1885, see: suspectus Erichson, 1840.
quadratus Kraatz, 1859, see: Belonuchus.
quadriceps Cameron, 1950, see: Gabronthus.
quadricollis Horn, 1884: 194 (Philonthus; Type locality: Illinois; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from New York to Missouri and Michigan).
— Fauvel, 1889b: 265 (Philonthus; synonym of quisquiliarius).
— Ganglbauer, 1895: 455 (Philonthus; synonym of quisquiliarius).
— Fauvel, 1903d: 271 (Philonthus; synonym of quisquiliarius).
— Blatchley, 1910: 383 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 352 (Philonthus; catalog).
— Notman, 1919: 135 (Philonthus; New York).
— Notman, 1920a: 180 (Philonthus; New York).
— Hatch, 1957: 190 (Philonthus; characters; British Columbia; Washington).
— Smetana, 1995e: 287 (Philonthus; lectotype designation: Illinois; characters; notes; Alberta; British Columbia; Manitoba; New Brunswick; Nova Scotia; Ontario; Quebec; Saskatchewan; Arkansas; California; Colorado; Connecticut; Delaware; Idaho; Illinois; Mississippi; Missouri; Montana; Nebraska; Nevada; New Hampshire; New Jersey; New York; North Carolina; North Dakota; Ohio; Oregon; Pennsylvania; South Dakota; Tennessee; Utah; Vermont; Virginia; Washington; Wisconsin; Wyoming).
— Downie and Arnett, 1996: 378 (Philonthus; characters; USA).
▴ Distribution: Canada, USA.
fraternus Fall, 1926: 149 (Philonthus; Type locality: Dawson, Yukon Territory).
— Scheerpeltz, 1933: 1343 (Philonthus; catalog).
— Smetana, 1995e: 287 (Philonthus; synonym of quadricollis).
quadricolor Bernhauer, 1916, see: pulchricolor Bernhauer, 1923.
quadricolor Fauvel, 1879: 99 (Philonthus; Type locality: Nouvelle Guinée, Amberbaki).
quadripunctatus Fiori, 1894, see: ebeninus Gravenhorst, 1802.
quadripunctulus Scheerpeltz, 1974: 156 (Philonthus; Type locality: S. W. Africa: Kaokoveld, Anabib, Orupembe, 100 miles W Ohopoho. Also cited from South Africa).
quadrulus Horn, 1884: 221 (Philonthus; Type locality: Arizona; [Note: See lectotype designation by Smetana, 1995e]. Also cited from California).
— Fall and Cockerell, 1907: 166 (Philonthus; New Mexico).
— Bernhauer and Schubert, 1914: 352 (Philonthus; catalog).
— Smetana, 1995e: 314 (Philonthus; lectotype designation: Arizona; characters; notes; British Columbia; Arizona; California; New Mexico; Oregon; Texas).
▴ Distribution: Canada, USA.
sagax Casey, 1915: 445 (Philonthus; Type locality: Arizona: Nogales; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1359 (Philonthus; catalog).
— Smetana, 1995e: 314 (Philonthus; lectotype designation: Nogales, Ari [Arizona]; synonym of quadrulus).
quediiformis Cameron, 1932: 76 (Philonthus; replacment name for nilgiriensis).
nilgiriensis Cameron, 1920a: 219 (Quedius; subgenus Quedionuchus; [preoccupied]; Type locality: Nilgiri Hills).
— Cameron, 1932: 76 (Philonthus; synonym of quediiformis).
— Scheerpeltz, 1933: 1357 (Philonthus; synonym of quediifornis).
quediinus Horn, 1884, see: Bisnius.
quediodes Sharp, 1874: 38 (Philonthus; Type locality: Nagasaki).
— Bernhauer and Schubert, 1914: 352 (Philonthus; catalog).
— Scheerpeltz, 1933: 1357 (Philonthus; catalog; East Asia).
▴ Distribution: Japan, “East Asia”.
quediomimus Bernhauer, 1939a: 85 (Philonthus; Type locality: Kenya: Mt. Elgon, Osthang, Suam fishing hut, 2400 m).
quirinus Smetana, 1995e: 336 (Philonthus; Type locality: Louisiana).
quisquiliariformis Scheerpeltz, 1958a: 18 (Philonthus; Type locality: Kotoum).
— Coiffait, 1967a: 425 (Philonthus; characters; Turkey).
— Coiffait, 1974: 295 (Philonthus; characters; Turkey).
▴ Distribution: Turkey.
quisquiliarius Gyllenhal, 1810: 335 (Staphylinus; Type locality: Suecia).
— C. Sahlberg, 1830: 325 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 29 (Staphylinus; Finland; Sweden).
— Mannerheim, 1831: 443 (Staphylinus; Finland; Sweden).
— Runde, 1835: 8 (Staphylinus; characters; Germany).
— Lacordaire, 1835: 406 (Staphylinus; characters; France).
— Nordmann, 1837: 93 (Philonthus; Sweden; Germany; Russia).
— Erichson, 1839a: 469 (Philonthus; characters; Germany).
— Heer, 1839: 267 (Philonthus; characters; Switzerland).
— Erichson, 1840: 475 (Philonthus; characters; Europe; Syria; Egypt).
— Jacobsen, 1849: 700 (Philonthus; characters; Denmark).
— Hochhuth, 1849: 141 (Philonthus; characters; Iberia).
— Redtenbacher, 1849: 703 (Philonthus; characters; Austria).
— Kiesenwetter, 1851a: 416 (Philonthus; France).
— Rosenhauer, 1856: 71 (Philonthus; Xerez; Cartama).
— Fairmaire and Laboulbène, 1856: 523 (Philonthus; characters; France).
— Kraatz, 1857d: 607 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 193 (Philonthus; characters; Austria).
— Kraatz, 1858a: 61 (Philonthus; Greece).
— Peyron, 1858: 426 (Philonthus; Turkey).
— Thomson, 1860: 160 (Philonthus; characters; Scandinavia).
— Fauvel, 1865a: 297 [= 1865b: 47] (Philonthus; habitat; France).
— Fauvel, 1874: 237 [= 1874a: 463] (Philonthus; characters; France; Barbary; Egypt; Turkey; Caucasus; Transcaucasia; Japan; USA).
— Sharp, 1874: 44 (Philonthus; cited as quisquiliarus; Japan [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Redtenbacher, 1874: 212 (Philonthus; characters; Austria).
— Seidlitz, 1875: 276 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 492 [= 1877: 348] (Philonthus; characters; France).
— J. Sahlberg, 1876: 18 (Philonthus; Finland).
— Fauvel, 1878: 123 [= 1878a: 43] (Philonthus; Algeria).
— Fauvel, 1886: 55 [= 1886a: 47] (Philonthus; Morocco; Egypt; Algeria).
— Fowler, 1888: 275 (Philonthus; characters; Britain).
— Fauvel, 1889b: 265 (Philonthus; New Caledonia [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Eppelsheim, 1892: 334 (Philonthus; Uzbekistan).
— Ganglbauer, 1895: 455 (Philonthus; characters; Palaearctic and Nearctic regions; Africa; China; Siam; New Caledonia [Note: According to Tottenham, 1953: 143 specimens from the latter locality are probably misidentified]).
— Fauvel, 1902b: 110 (Philonthus; Barbary; Egypt).
— Fauvel, 1903d: 271 (Philonthus; New Caledonia [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Sainte-Claire Deville, 1906: 97 (Philonthus; Corsica).
— Montandon, 1908: 80 (Philonthus; Romania).
— Poppius, 1909: 26 (Philonthus; subgenus Gefyrobius; Lena Valley, Siberia).
— Reitter, 1909: 129 (Philonthus; characters; Germany).
— Eichelbaum, 1913: 130 (Philonthus; Tanzania [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Bernhauer and Schubert, 1914: 352 (Philonthus; catalog).
— Johansen, 1914: 411 (Philonthus; characters; Denmark).
— Bernhauer, 1915n: 319 (Philonthus; German Southwest Africa [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Bernhauer, 1922c: 232 (Philonthus; Formosa [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Rambousek, 1922: 1009 (Philonthus; Ethiopia; British East Africa [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Lea, 1925: 220, 221 (Philonthus; Australian introduction [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Bernhauer, 1926g: 2 (Philonthus; Kamchatka region).
— Bernhauer, 1928: 16 (Philonthus; Sumatra [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Portevin, 1929: 362 (Philonthus; characters; France).
— Cameron, 1932: 123 (Philonthus; characters; Ceylon; India [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Scheerpeltz, 1933: 1357 (Philonthus; catalog).
— Cameron, 1933d: 387 (Philonthus; Sumatra [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Bernhauer, 1936: 4 (Philonthus; Mongolia).
— Gridelli, 1934: 74 (Philonthus; Karakoram region).
— Cameron, 1937: 5 (Philonthus; Java [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Bernhauer and Chapman, 1939: 71 (Philonthus; Kenya [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Hafez, 1939 (Philonthus; life history; egg, larval, and pupal characters).
— Méquignon, 1942a: 21 (Philonthus; Azores).
— Cameron, 1949b: 468 (Philonthus; Japan [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— V. Hansen, 1952: 74 (Philonthus; subgenus Philonthus; characters; Denmark).
— Tottenham, 1953: 143 (Philonthus; limited to Europe and parts of Asia and North Africa, not in North America or the Oriental and Australasian regions).
— Smetana, 1953: 119 (Philonthus; Turkey).
— Scheerpeltz, 1954: 114 (Philonthus, cited as Phylonthus; India [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Scheerpeltz, 1956a: 484 (Philonthus; Belgian Congo).
— Smetana, 1958: 215 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 164 (Philonthus; cited as quisquiliarus; characters).
— Scheerpeltz, 1960d: 135 (Philonthus; Portugal).
— Scheerpeltz, 1961g: 238 (Philonthus; Madagascar [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Scheerpeltz, 1961h: 253 (Philonthus; Afghanistan).
— Scheerpeltz, 1963: 13 (Philonthus; Afghanistan).
— Scheerpeltz, 1963b: 429 (Philonthus; Egypt).
— Raianu, 1963: 82 (Philonthus; Romania).
— Palm, 1963b: 91 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 184 (Philonthus; subgenus Philonthus; characters; central Europe).
— Smetana, 1964d: 76 (Philonthus; subgenus Philonthus; Czechoslovakia).
— Focarile, 1964: 59 (Philonthus; Italy).
— Scheerpeltz, 1965: 107 (Philonthus; Burma [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Scheerpeltz, 1965b: 402 (Philonthus; Sicily).
— Scheerpeltz, 1965c: 506 (Philonthus; Sicily).
— Horion, 1965: 166 (Philonthus; widespread in central and southern Europe).
— Coiffait, 1967a: 424 (Philonthus; characters; Europe).
— Smetana, 1967d: 208 (Philonthus; Mongolia).
— Smetana, 1967i: 555 (Philonthus; Turkey).
— Raianu, 1968a: 467 (Philonthus; Romania).
— Szujecki, 1968a: 731 (Philonthus; Poland).
— Coiffait, 1970d: 143 (Philonthus; Uzbekistan).
— Coiffait, 1973a: 274 (Philonthus; Morocco).
— Moore and Legner, 1974a: 39 (Philonthus; spread by commerce).
— Coiffait, 1974: 199 (Philonthus; characters; Europe; Britain; Scandinavia; Asia Minor; Lebanon; North Africa).
— Scheerpeltz, 1974: 151 (Philonthus; South Africa [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Scheerpeltz, 1974c: 22 (Philonthus; Sudan; Ethiopia [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Bordoni, 1975a: 153 (Philonthus; Afghanistan).
— Osella and Zanetti, 1975: 109 (Philonthus; collected from nest of Talpa europaea; notes; Italy).
— Watanabe and Shibata, 1976: 332 (Philonthus; Malaysia [Note: According to Tottenham, 1953: 143 this is probably a misidentification]).
— Scheerpeltz, 1976a: 89 (Philonthus; Nepal).
— Shilov, 1976: 152 (Philonthus; Mongolia).
— Pace and Zanetti, 1977: 298 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Hinton, 1981: 661 (Philonthus; characters of egg).
— Tronquet, 1981: 71 (Philonthus; cited as quisquilarius; Afghanistan).
— Coiffait, 1982a: 232 (Philonthus; Nepal).
— Frank, 1982: 43 (Philonthus; list of parasites).
— Boller, 1983: 118 (Philonthus; larval characters).
— Tóth, 1984: 74 (Philonthus; cited as quisquilarius; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Nekulisianu, Ostafichuk and Tsygankova, 1988 (Philonthus; larval characters; biological notes).
— Borges, 1990: Table II (Philonthus; Azores).
— Aranda and Outerelo, 1991: 121 (Philonthus; characters).
— Pilon and Zanetti, 1991: 64 (Philonthus; Italy).
— J. Li, 1993: 37 (Philonthus; China).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Hernández, Outerelo, and Gamarra, 1994: 192 (Philonthus; Canary Islands).
— Yadav, Chand, and Yadav, 1995: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Smetana, 1995e: 291 (Philonthus; does not occur in North America).
— Mikat, Fremuth, and Prouza, 1997: 111 (Philonthus; Czech Republic).
— Owen, 1997a: 149 (Philonthus; collecting sites; Ireland).
— Duverger, 1998 (Philonthus; host to three species of Laboulbeniaceae).
— Assing, Frisch, Kahlen, et al., 1998: 138 (Philonthus; characters).
▴ Distribution: Egypt, Algeria, Morocco, Azores, Europe, Russia, Caucasus, Turkey, Syria, Afghanistan, Uzbekistan, Nepal, Mongolia, China, “Karakorum region”. Note: According to Schillhammer (personal communication) the records for the Oriental region cited above refer to P. variipennis and P. bassus.
inquinatus Stephens, 1832: 223 (Quedius; Type locality: Suffolk).
— Curtis, 1837: pl. 638 (Quedius; Type locality: Britain).
— Stephens, 1839: 392 (Quedius; characters; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of quisquiliarius).
— Fauvel, 1874: 237 [= 1874a: 463] (Philonthus; variety of quisquiliarius).
— Ganglbauer, 1895: 455 (Philonthus; variety of quisquiliarius).
— Fauvel, 1903: 159 (Philonthus; variety of quisquiliarius; India).
— Fauvel, 1904a: 59 (Philonthus; variety of quisquiliarius; Belgaum; Persia; Europe; India; Japan; Malaysia; Celebes).
— Bernhauer and Schubert, 1914: 352 (Philonthus; variety of punctus).
— Cameron, 1932: 123 (Philonthus; synonym of quisquiliarius).
— Scheerpeltz, 1933: 1358 (Philonthus; aberration of quisquiliarius).
— Smetana, 1953: 119 (Philonthus; variety of quisquiliarius; Turkey).
— Scheerpeltz, 1954: 114 (Philonthus, cited as Phylonthus; variety of quisquiliarius; Ceylon).
— Smetana, 1958: 215 (Philonthus; aberration of quisquiliarius).
— Smetana, 1959: 164 (Philonthus; aberration of quisquiliarius; characters).
— Scheerpeltz, 1961h: 253 (Philonthus; aberration of quisquiliarius; Afghanistan).
— Smetana, 1964d: 76 (Philonthus; subgenus Philonthus; aberration of quisquiliarius; Czechoslovakia).
— Scheerpeltz, 1965: 107 (Philonthus; variety of quisquiliarius; Burma).
— Coiffait, 1967a: 424 (Philonthus; aberration of quisquiliarius; characters).
— Smetana, 1967i: 555 (Philonthus; aberration of quisquiliarius; Turkey).
— Coiffait, 1974: 199 (Philonthus; characters; variety of quisquiliarius).
— Pope, 1977: 29 (Philonthus; synonym of quisquiliarius).
— Tronquet, 1981: 71 (Philonthus; variety of quisquiliarius; Afghanistan).
— Tóth, 1984: 74 (Philonthus; variety of quisquiliarius; Hungary).
phaeopus Stephens, 1832: 236 (Philonthus; Type locality: London; Suffolk).
— Curtis, 1829: 25 (Philonthus; nomen nudum; catalog; Britain).
— Stephens, 1839: 397 (Philonthus; characters; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of quisquiliarius).
— Fauvel, 1874: 237 [= 1874a: 463] (Philonthus; synonym of quisquiliarius).
— Ganglbauer, 1895: 455 (Philonthus; synonym of quisquiliarius).
— Bernhauer and Schubert, 1914: 352 (Philonthus; synonym of punctus).
— Cameron, 1932: 123 (Philonthus; synonym of quisquiliarius).
— Smetana, 1958: 215 (Philonthus; synonym of quisquiliarius).
— Coiffait, 1974: 199 (Philonthus; synonym of quisquiliarius).
dimidiatus Lacordaire, 1835: 402 (Staphylinus; [preoccupied]; Type locality: Paris).
— Erichson, 1839a: 467 (Philonthus; characters; Germany).
— Heer, 1839: 266 (Philonthus; characters; Switzerland).
— Erichson, 1840: 475 (Philonthus; synonym of rubidus).
— Jacobsen, 1849: 700 (Philonthus; synonym of rubidus).
— Fairmaire and Laboulbène, 1856: 522 (Philonthus; synonym of rubidus).
— Kraatz, 1857d: 607 (Philonthus; synonym of quisquiliarius).
— Redtenbacher, 1857: 194 (Philonthus; characters; Austria).
— Ganglbauer, 1895: 455 (Philonthus; synonym of quisquiliarius).
— Bernhauer and Schubert, 1914: 352 (Philonthus; synonym of punctus).
— Cameron, 1932: 123 (Philonthus; synonym of quisquiliarius).
— Scheerpeltz, 1933: 1358 (Philonthus; synonym of quisquiliarius).
— Smetana, 1958: 215 (Philonthus; synonym of quisquiliarius).
— Coiffait, 1974: 199 (Philonthus; synonym of quisquiliarius).
— Pope, 1977: 29 (Philonthus; synonym of quisquiliarius).
rubidus Erichson, 1840: 475 (Philonthus; Type locality: Germania; Gallia).
— Heer, 1841: 583 (Philonthus; synonym of dimidiatus Lacordaire).
— Jacobsen, 1849: 700 (Philonthus; characters; Denmark).
— Redtenbacher, 1849: 704 (Philonthus; characters; Austria).
— Kiesenwetter, 1851a: 417 (Philonthus; France).
— Fairmaire and Laboulbène, 1856: 522 (Philonthus; characters; France).
— Kraatz, 1857d: 607 (Philonthus; synonym of quisquiliarius).
— Peyron, 1858: 426 (Philonthus; Turkey).
— Thomson, 1860: 161 (Philonthus; synonym of quisquiliarius).
— Fauvel, 1874: 237 [= 1874a: 463] (Philonthus; synonym of quisquiliarius).
— Redtenbacher, 1874: 213 (Philonthus; characters; Austria).
— Ganglbauer, 1895: 455 (Philonthus; synonym of quisquiliarius).
— Bernhauer and Schubert, 1914: 352 (Philonthus; synonym of punctus).
— Cameron, 1932: 123 (Philonthus; synonym of quisquiliarius).
— Scheerpeltz, 1933: 1358 (Philonthus; synonym of quisquiliarius).
linearis Hochhuth, 1849: 140 (Philonthus; Type locality: Kaukasien).
— Hochhuth, 1862: 63 (Philonthus; Russia).
— Fauvel, 1874: 237 [= 1874a: 463] (Philonthus; synonym of quisquiliarius).
— Ganglbauer, 1895: 455 (Philonthus; synonym of quisquiliarius).
— Bernhauer and Schubert, 1914: 352 (Philonthus; synonym of punctus).
— Cameron, 1932: 123 (Philonthus; synonym of quisquiliarius).
— Smetana, 1958: 215 (Philonthus; synonym of quisquiliarius).
— Coiffait, 1974: 199 (Philonthus; synonym of quisquiliarius).
— Gusarov, 1992a: 786 [= 1993a: 75] (Philonthus; synonym of quisquiliarius).
sinuatus Wollaston, 1867: 239 (Philonthus; Type locality: Cape Verde Islands: S. Antao).
— Fauvel, 1902b: 110 (Philonthus; synonym of quisquiliarius).
— Bernhauer and Schubert, 1914: 352 (Philonthus; synonym of punctus).
— Cameron, 1932: 123 (Philonthus; synonym of quisquiliarius).
— Smetana, 1958: 215 (Philonthus; synonym of quisquiliarius).
— Coiffait, 1974: 199 (Philonthus; synonym of quisquiliarius).
chalceipennis Fauvel, 1878g: 547 (Philonthus; Type locality: Nouvelle-Calédonie, Kanala).
— Fauvel, 1889b: 265 (Philonthus; synonym of quisquiliarius).
— Bernhauer and Schubert, 1914: 352 (Philonthus; synonym of punctus).
— Cameron, 1932: 123 (Philonthus; synonym of quisquiliarius).
— Tottenham, 1953: 148 (Philonthus; notes; probably a valid species and may be same as geus).
— Smetana, 1958: 215 (Philonthus; synonym of quisquiliarius).
stoliczkae Sharp, 1878: 170 (Philonthus; Type locality: Yarkand).
— Sharp, 1890: 40 (Philonthus; [Note: Cited by Bernhauer and Schubert, 1914: 353 as the original description]; characters; Yarkand).
— Fauvel, 1903: 159 (Philonthus; synonym of quisquiliarius variety inquinatus).
— Fauvel, 1904a: 59 (Philonthus; synonym of quisquiliarius variety inquinatus).
— Bernhauer and Schubert, 1914: 353 (Philonthus; synonym of punctus).
— Tottenham, 1953: 148 (Philonthus; notes; probably a valid species).
— Smetana, 1958: 215 (Philonthus; synonym of quisquiliarius).
brunneipennis Quedenfeldt, 1882: 182 (Philonthus; Type locality: Marocco: Tanger).
— Quedenfeldt, 1884: 374 (Philonthus; characters; Morocco).
— Fauvel, 1886: 55 [= 1886a: 47] (Philonthus; Morocco).
— Fauvel, 1903d: 271 (Philonthus; synonym of quisquiliarius).
— Bernhauer and Schubert, 1914: 352 (Philonthus; synonym of quisquiliarius).
— Cameron, 1932: 123 (Philonthus; synonym of quisquiliarius).
— Smetana, 1958: 215 (Philonthus; synonym of quisquiliarius).
— Coiffait, 1974: 199 (Philonthus; synonym of quisquiliarius).
rubrosuturatus Bernhauer, 1902: 166 (Philonthus; variety of quisquiliarius; Type locality: Port Daufin, Madagaskar).
opacus Gerhardt, 1910: 555 (Philonthus; [preoccupied]; Type locality: Liegnitz).
— Scheerpeltz, 1933: 1358 (Philonthus; synonym of quisquiliarius).
— Smetana, 1958: 215 (Philonthus; synonym of quisquiliarius).
— Coiffait, 1974: 199 (Philonthus; synonym of quisquiliarius).
samoensis Bernhauer, 1922a: 150 (Philonthus; variety of quisquiliarius; Type locality: Samoa: Apia).
— Cameron, 1932: 123 (Philonthus; synonym of quisquiliarius).
— Scheerpeltz, 1933: 1358 (Philonthus; variety of quisquiliarius).
— Tottenham, 1953: 148 (Philonthus; notes; probably a valid species).
opacinus Scheerpeltz, 1933: 1358 (Philonthus; variety of quisquiliarius; replacement name for opacus Gerhardt).
— Smetana, 1959: 164 (Philonthus; aberration of quisquiliarius; characters).
— Smetana, 1958: 215 (Philonthus; synonym of quisquiliarius).
— Coiffait, 1967a: 424 (Philonthus; aberration of quisquiliarius; characters).
— Coiffait, 1974: 199 (Philonthus; characters; variety of quisquiliarius).
— Tronquet, 1981: 71 (Philonthus; variety of quisquiliarius; Afghanistan).
andalusiacus Coiffait, 1966c: 510 (Philonthus; subspecies of quisquiliarius; Type locality: Andalousei, San Pedro Alcantara, Marbella).
quisquiliaroides Scheerpeltz, 1974: 152 (Philonthus; Type locality: Cape Prov.: Steenkamp Puts, 37 miles NW Upington).
rabidus Tottenham, 1962: 167 (Philonthus; Type locality: Tanganyika Terr.: Ngorongoro, 2500–2600 m. Also cited from Kenya).
ragusai Scheerpeltz, 1933, see: alcyoneus Erichson, 1840.
rainieri Hatch, 1957, see: Bisnius.
ralumensis Bernhauer, 1915g: 188 (Philonthus; Type locality: Ralums).
— Cameron, 1931: 363 (Philonthus; New Guinea).
— Scheerpeltz, 1933: 1358 (Philonthus; catalog).
▴ Distribution: New Guinea.
ramosus Levasseur, 1968: 1385 (Philonthus; Type locality: mont Cameroun).
rapaciosus Tottenham, 1949b: 350 (Philonthus; Type locality: Lulua: Kapanga).
rapillyi Jarrige, 1971, see: punctus Gravenhorst, 1802.
rauli Scheerpeltz, 1978: 212 (Philonthus; Type locality: Dadim-Bere, Timor).
rectangulus Sharp, 1874: 42 (Philonthus; Type locality: Nagasaki).
— Bernhauer and Schubert, 1914: 353 (Philonthus; catalog).
— Bernhauer, 1922c: 232 (Philonthus; Formosa).
— Scheerpeltz, 1933: 1358 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Gridelli, 1936a: 154 (Philonthus; notes on distribution).
— Walles, 1938: 69, 70 (Philonthus; Poland).
— Adachi, 1937: 57 (Philonthus; characters; Izu-Oshima).
— Tottenham, 1935 (Philonthus; characters; Britain; France).
— Cameron, 1949b: 468 (Philonthus; Japan).
— V. Hansen, 1952: 63 (Philonthus; subgenus Philonthus; characters; Denmark).
— Coiffait, 1953: 78 (Philonthus; notes on distribution).
— Smetana, 1953: 118 (Philonthus; Turkey).
— Coiffait, 1954: 165 (Philonthus; notes; Canary Islands).
— Lohse, 1956: 88 (Philonthus; notes).
— Hatch, 1957: 192 (Philonthus; characters; British Columbia; Washington; Oregon; Idaho).
— Smetana, 1958: 174 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 156 (Philonthus; characters).
— Scheerpeltz, 1961d: 128 (Philonthus; Italy).
— Nakane, 1963a: 91 (Philonthus; characters; habitus photograph; Japan).
— Scheerpeltz, 1963: 13 (Philonthus; Afghanistan).
— Lohse, 1964: 181 (Philonthus; subgenus Philonthus; characters; central Europe).
— Last, 1961: 306 (Philonthus; Thailand).
— Palm, 1963b: 81 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Horion, 1965: 153 (Philonthus; Japan; Siberia; China; Afghanistan; widespread in Europe).
— Coiffait, 1967a: 415 (Philonthus; characters; Holarctic region; Chile).
— Last, 1968: 369 (Philonthus; New Guinea).
— Coiffait and Saiz, 1968: 351, 353, 354 (Philonthus; characters; introduced species; Chile).
— Raianu, 1969 (Philonthus; characters; ecological notes; Romania).
— Saiz, 1971b: 342 (Philonthus; notes; Chile).
— Watanabe and Shibata, 1972: 64 (Philonthus; Yaku-shima Island).
— Bacchus and Hammond, 1972: 63 (Philonthus; donkey dung; Britain).
— Coiffait, 1974: 194 (Philonthus; characters; Japan; China; Manchuria; Formosa; Palaearctic region; Madeira; Chile).
— Moore, 1975c: 106 (Philonthus; Hawaii).
— Coiffait, 1976: 217 (Philonthus; Tahiti).
— Coiffait, 1977b: 206 (Philonthus; Bhutan).
— Pace and Zanetti, 1977: 291 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Scheerpeltz, 1978: 219 (Philonthus; Timor).
— Raichev, 1979: 82 (Philonthus; Bulgaria).
— Frank, 1982: 43 (Philonthus; list of parasites).
— Boller, 1983: 117 (Philonthus; larval characters).
— Coiffait, 1984e: 373 (Philonthus; Nepal).
— Tóth, 1984: 63 (Philonthus; characters; Hungary).
— Yuh, Paik, Kwon, and Lee, 1985: 242 (Philonthus; cited as rectangules; Korea).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Israelson, 1990: 2 (Philonthus; Azores).
— Borges, 1990: Table II (Philonthus; Azores).
— Kuschel, 1990: 26 (Philonthus; introduced, Palaearctic species; winged; habitat; New Zealand).
— Pilon and Zanetti, 1991: 58 (Philonthus; Italy).
— Hunter, Kincher, Bay, Beerwinkle, 1991: 168 (Philonthus; seasonal distribution and diel flight activity; Texas).
— J. Li, 1993: 35 (Philonthus; China).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Dearborn and Donahue, 1994: 15 (Philonthus; Maine).
— Kanehisa, Tsumuki, and Kawazu, 1994 (Philonthus; chemical secretions).
— Nishida, 1994: 73 (Philonthus; Hawaii).
— Hayashi, 1994: 125 (Philonthus; characters; Japan).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; introduced in Italy).
— Smetana, 1995e: 216 (Philonthus; characters; Alberta; British Columbia; Manitoba; New Brunswick; Newfoundland; ontario; Quebec; Saskatchewan; Alaska; Arkansas; Arizona; California; Colorado; Connecticut; Florida; Georgia; Idaho; Illinois; Indiana; Iowa; Kansas; Kentucky; Louisiana; Maine; Maryland; Massachusetts; Michigan; Minnesota; Mississippi; Missouri; Montana; Nebraska; New Hampshire; New Jersey; New Mexico; New York; North Carolina; North Dakota; Ohio; Oklahoma; Oregon; Pennsylvania; South Carolina; South Dakota; Tennessee; Texas; Utah; Virginia; Washington; Wisconsin).
— Hodge and Jones, 1995: 40 (Philonthus; characters; Britain).
— Downie and Arnett, 1996: 378 (Philonthus; characters; USA).
— Gerend and Braunert, 1997: 197 (Philonthus; notes; Luxembourg).
— Sterrenburg and Schülke, 1997: 18 (Philonthus; Netherlands).
— Nishida, 1997: 62 (Philonthus; Hawaii).
— Nishida, 1997: 62 (Philonthus; Hawaii).
— Schmidt, 1999: 108 (Philonthus; food for adults and larvae used for rearing species; rearing methods).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
— Ødegaard, 1999: 71 (Philonthus; invasive species associated with compost heaps in Nordic countries).
▴ Distribution: Canary Islands, Madeira, Azores, Europe, Turkey, Kazakhstan, Afghanistan, Nepal, Bhutan, Thailand, Indonesia, New Guinea, Society Islands, China, Korea, Japan, Taiwan, Hawaii, New Zealand, Canada, USA, Chile.
bernhaueri Csiki, 1901: 104 (Philonthus; Type locality: China: Peking).
— Bernhauer and Schubert, 1914: 353 (Philonthus; synonym of rectangulus).
— Tottenham, 1941 (Philonthus; synonym of rectangulus).
— Smetana, 1958: 174 (Philonthus; synonym of rectangulus).
— Smetana, 1959: 156 (Philonthus; synonym of rectangulus).
— Coiffait, 1974: 194 (Philonthus; synonym of rectangulus).
— Scheerpeltz, 1978: 219 (Philonthus; synonym of rectangulus).
— Smetana, 1995e: 216 (Philonthus; synonym of rectangulus).
tetragonocephalus Notman, 1924: 271 (Philonthus; Type locality: Cranberry Lake, New York).
— Scheerpeltz, 1933: 1364 (Philonthus; catalog).
— Tottenham, 1941 (Philonthus; synonym of rectangulus).
— Hatch, 1957: 192 (Philonthus; synonym of rectangulus).
— Smetana, 1958: 174 (Philonthus; synonym of rectangulus).
— Smetana, 1959: 156 (Philonthus; synonym of rectangulus).
— Coiffait and Saiz, 1968: 354 (Philonthus; synonym of rectangulus).
— Coiffait, 1974: 194 (Philonthus; synonym of rectangulus).
— Smetana, 1995e: 216 (Philonthus; synonym of rectangulus).
rufipennis Wüsthoff, 1936: 236 (Philonthus; form of rectangulus; [preoccupied]; Type locality: Not cited but presumably Rheinprovinz).
— Smetana, 1958: 174 (Philonthus; aberration of rectangulus).
— Smetana, 1959: 156 (Philonthus; aberration of rectangulus; characters).
— Coiffait, 1967a: 415 (Philonthus; aberration of rectangulus; characters).
— Coiffait, 1974: 194 (Philonthus; variety of rectangulus).
— Smetana, 1995e: 216 (Philonthus; synonym of rectangulus).
mequignoni Jarrige, 1938: 206 (Philonthus; Type locality: France: Jouy-sous-Thelle, Oise).
— Tottenham, 1941 (Philonthus; synonym of rectangulus).
— Paulian, 1941: 228 (Philonthus; larval characters).
— Gridelli, 1943: 128 (Philonthus; notes).
— Smetana, 1958: 174 (Philonthus; synonym of rectangulus).
— Smetana, 1959: 156 (Philonthus; synonym of rectangulus).
— Coiffait and Saiz, 1968: 354 (Philonthus; synonym of rectangulus).
— Coiffait, 1974: 194 (Philonthus; synonym of rectangulus).
— Smetana, 1995e: 216 (Philonthus; synonym of rectangulus).
peculiaris Last, 1974b: 192 (Philonthus; subspecies of rectangulus; Type locality:
New Guinea: Okapa).
recticollis Cameron, 1947: 2 (Philonthus; Type locality: Nizi: Blukwa).
rectilaterus Sharp, 1885: 425 (Philonthus; Type locality: Mexico, Las Vigas; Guatemala, Dueñas).
— Bernhauer and Schubert, 1914: 353 (Philonthus; catalog).
— Scheerpeltz, 1971b: 205 (Philonthus; Guatemala; Mexico; El Salvador; Nicaragua; Costa Rica).
▴ Distribution: Mexico, Guatemala, El Salvador, Nicaragua, Costa Rica.
rectus Tottenham, 1962: 197 (Philonthus; Type locality: Madagascar mer. or: Midongy).
— Lecoq, 1998: 71 (Philonthus; subgenus Piezarthrus; characters; Madagascar).
▴ Distribution: Madagascar.
reflexiventris Tikhomirova, 1973, see: oberti Eppelsheim, 1889.
regillus Sharp, 1876, see: Paederomimus.
reinecki Schubert, 1902: 405 (Philonthus; Type locality: Pietermaritzburg).
— Fauvel, 1907: 38 (Philonthus; Nairobi; Natal).
— Bernhauer and Schubert, 1914: 353 (Philonthus; catalog).
— Tottenham, 1955: 168 (Philonthus; characters).
— Tottenham, 1962: 135, 153 (Philonthus; notes; characters; Tanganyika).
▴ Distribution: South Africa, Tanzania, Kenya.
reitteri Eppelsheim, 1889, see: Bisnius.
reitterianus Smetana, 1955: 208 (Philonthus; Type locality: Buchara).
— Coiffait, 1967a: 429 (Philonthus; characters; Turkestan).
— Coiffait, 1974: 275 (Philonthus; characters; Uzbekistan).
▴ Distribution: Uzbekistan.
remotus Fauvel, 1895: 264 (Philonthus; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Carin Asciuii Ghécu, 1400–1500 m).
— Bernhauer and Schubert, 1914: 353 (Philonthus; catalog).
— Cameron, 1932: 110 (Philonthus; characters; Burma; India).
— Scheerpeltz, 1933: 1358 (Philonthus; catalog).
— Cameron, 1937: 5 (Philonthus; Java).
▴ Distribution: India, Myanmar, Indonesia.
nilgiriensis Cameron, 1920a: 215 (Philonthus; Type locality: Nilgiri Hills).
— Cameron, 1932: 110 (Philonthus; synonym of remotus).
— Scheerpeltz, 1933: 1358 (Philonthus; synonym of remotus).
renominatus Cameron, 1937: 5 (Philonthus; replacement name for vicinus Cameron, 1933).
— Cameron, 1937: 5 (Philonthus; Java).
— Herman, 2001: 28 (Philonthus; synonymy).
▴ Distribution: Indonesia, Malaysia.
vicinus Cameron, 1933d: 389 (Philonthus; [preoccupied]; Type locality: Sumatra: Fort de Kock; Gunung Singgalang; Malay Peninsula).
pheres Smetana, 1963e: 74 (Philonthus; replacement name for vicinus Cameron, 1933; [Note: Unnecessary replacement name]).
renominatus Cameron, 1951, see: repetitus Herman, 2001.
repetitus Herman, 2001: 48 (Philonthus; replacement name for renominatus Cameron, 1951).
renominatus Cameron, 1951c: 402 (Philonthus; [preoccupied]; Type locality: Kenya Colony: Nairobi).
reskai R. Dvořák, 1954, see: Rabigus.
rhodesiae Tottenham, 1949b: 319 (Philonthus; Type locality: N.W. Rhodesia: Kashitu, N. of Broken Hill).
rhodesianus Tottenham, 1949b: 348 (Philonthus; Type locality: N.W. Rhodesia: Kashitu, N. of Broken Hill. Belgian Congo: 10 miles S.W. of Elizabethville).
richteri Bernhauer, 1912, see: Chroaptomus.
ridens Tottenham, 1955: 167 (Philonthus; subgenus Spatulonthus; Type locality: S. Rhodesia: Umtali; also listed from N.W. Rhodesia and Transvaal).
— Tottenham, 1956: 227 (Philonthus; subgenus Spatulonthus; Ruanda).
▴ Distribution: Rwanda, Zambia, Zimbabwe, South Africa.
ridiculus Blackwelder, 1944: 134 (Philonthus; replacement name for grandiceps Bernhauer).
grandiceps Bernhauer, 1916c: 33 (Philonthus; [preoccupied]; Type locality: West-Columbien: Umgebung von Cali in Caucatale).
riedeli Bernhauer, 1916c: 39 (Philonthus; Type locality: Brasilien: S. Paulo, Umgebung von Ribeirao Preto Fundao).
riftensis Fauvel, 1907: 38 (Philonthus; Type locality: Kajabe, Rift Valley).
— Bernhauer and Schubert, 1914: 353 (Philonthus; catalog).
— Tottenham, 1962: 161 (Philonthus; characters).
▴ Distribution: Kenya.
riparius Cameron, 1926b: 351 (Philonthus; Type locality: Mussorie District; Mossy Falls, Kolhu Khet. Chakrata District; Mohna, Binal Gadh, Manjgaon, Chulli Khud. Simla Hills, Kotgarh, Gahan, 5500 to 8000 feet).
— Cameron, 1928c: 563 (Philonthus; Sikkim; Tibet).
— Cameron, 1932: 91 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1358 (Philonthus; catalog).
— Coiffait, 1977b: 209 (Philonthus; Bhutan).
— Coiffait, 1979d: 552 (Philonthus; Afghanistan).
— Tronquet, 1981: 70 (Philonthus; Afghanistan).
▴ Distribution: China, Bhutan, India, Afghanistan.
rivularis Kiesenwetter, 1858, see: Erichsonius.
rivularis Cameron, 1932: 138 (Philonthus; [preoccupied]; Type locality: Dehra Dun: Song River. Mussoorie: Arni Gad, Aglar River. Simla Hills: Fagu. Sikkim: Namsoo).
— Scheerpeltz, 1933: 1358 (Philonthus; catalog).
— Coiffait, 1983: 274 (Philonthus; Kashmir).
— Herman, 2001: 48 (Philonthus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: India.
robinsoni Lecoq, 1997: 285 (Philonthus; Type locality: Madagascar Sambirano, Nosy Be, forêt de Lokobe).
robusticornis Bernhauer, 1915, see: Afrorabigus.
roeri Last, 1992: 543 (Philonthus; Type locality: South West Africa (Namibia): Nyangana/Okavango).
rollei Bernhauer, 1943b: 290 (Philonthus; Type locality: Abessynien: Harrar).
rosarius Tottenham, 1945, see: nitidicollis Lacordaire, 1835.
roscius Smetana, 1995e: 78 (Philonthus; Type locality: Arizona: Chiricahua Mts., Rustler Park, 8400 ft.).
rossicus Coiffait, 1965, see: decorus Gravenhorst, 1802.
rotundiceps Stephens, 1833, see: ventralis Gravenhorst, 1802.
rotundicollis Ménétriés, 1832: 145 (Staphylinus; Type locality: Caucase).
— Nordmann, 1837: 88 (Philonthus; characters; Caucasus).
— Faldermann, 1835: 126 (Staphylinus; characters; Persia-Armenia).
— Ménétriés, 1849: 44 (Philonthus; Turcomanie).
— Hochhuth, 1849: 135 (Philonthus; synonym of scutatus).
— Fauvel, 1874: 230 [= 1874a: 456] (Philonthus; characters; France; Austria; Sardinia; Caucasus; Iran; Buchara; central Asia; Siberia; Dauria; Amur region).
— Solsky, 1874: 175 (Philonthus; characters; Turkestan).
— Fauvel, 1876a: 73 [= 1876: 258] (Philonthus; Turkestan).
— J. Sahlberg, 1876: 13 (Philonthus; Finland).
— Eppelsheim, 1892: 332 (Philonthus; Turkestan).
— Ganglbauer, 1895: 446 (Philonthus; characters; Palaearctic region).
— Poppius, 1909: 24 (Philonthus; Lena Valley, Siberia).
— Reitter, 1909: 126 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 353 (Philonthus; catalog).
— Bernhauer, 1926g: 2 (Philonthus; Kamchatka region).
— Portevin, 1929: 359 (Philonthus; characters; France).
— Cameron, 1932: 84 (Philonthus; characters; Pamirs).
— Scheerpeltz, 1933: 1358 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Aleksandrov, 1934: 153 (Philonthus; China).
— Smetana, 1958: 170 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 148 (Philonthus; characters).
— Scheerpeltz, 1960b: 68 (Philonthus; subgenus Philonthus; Afghanistan).
— Scheerpeltz, 1961b: 34 (Philonthus; Afghanistan).
— Scheerpeltz, 1963: 11 (Philonthus; Afghanistan).
— Palm, 1963b: 69 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 178 (Philonthus; subgenus Philonthus; characters; central Europe).
— Smetana, 1965e: 173 (Philonthus; Mongolia).
— Horion, 1965: 136 (Philonthus; Pamirs; Afghanistan; Transbaikal to Vladivostok; Mongolia; Russia; Finland; Sweden; Britain; Holland; Belgium; France; Spain; Sardinia; Italy; Bosnia; Macedonia; Romania; Czechoslovakia; Poland; Russia; Germany; Austria).
— Sawada, 1966: 259 (Philonthus; Afghanistan).
— Coiffait, 1967a: 395 (Philonthus; characters; Europe; Caucasus).
— Raianu, 1968a: 464 (Philonthus; Romania).
— Szujecki, 1968a: 728 (Philonthus; Poland).
— Szujecki, 1969a: 255 (Philonthus; notes; Poland).
— Coiffait, 1974: 227 (Philonthus; characters; Europe; Britain; central Asia).
— Shilov, 1975a: 375 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Bordoni, 1975a: 152 (Philonthus; Afghanistan).
— Shilov, 1976: 148 (Philonthus; Mongolia).
— Pope, 1977: 29 (Philonthus; Britain).
— Coiffait, 1979d: 552 (Philonthus; Afghanistan).
— Tronquet, 1981: 71 (Philonthus; Afghanistan).
— Coiffait, 1982a: 233 (Philonthus; Pakistan).
— Coiffait, 1983e: 477 (Philonthus; Afghanistan).
— Tóth, 1984: 50 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).
— Schülke and Uhlig, 1988: 11 (Philonthus; notes; Germany).
— Nowosad, 1990: 107 (Philonthus; collected from mole nest; notes; Poland).
— Pilon and Zanetti, 1991: 62 (Philonthus; Italy).
— J. Li, 1993: 36 (Philonthus; China).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Hayashi, 1994: 122 (Philonthus; characters; Japan).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Sterrenburg and Schülke, 1997: 18 (Philonthus; Netherlands).
— Ryabukhin, 1999: 62 (Philonthus; catalog; habitat; distribution; northeastern Russia).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
▴ Distribution: Europe, Russia, Caucasus, Iran, Turkmenistan, Kazakhstan, Uzbekistan, “Pamir region”, Afghanistan, Pakistan, Mongolia, China, Japan.
picimanus Ménétriés, 1832: 146 (Staphylinus; Type locality: Caucase).
— Faldermann, 1835: 132 (Staphylinus; characters; Persia-Armenia).
— Hochhuth, 1849: 155 (Quedius; characters).
— Ganglbauer, 1895: 448 (Philonthus; synonym of varius).
— Bernhauer and Schubert, 1914: 359 (Philonthus; aberration of varius).
— Scheerpeltz, 1933: 1366 (Philonthus; aberration of varius).
— Kirshenblat, 1933: 101 (Philonthus; characters; Caucasus).
— Tottenham, 1939d: 232 (Philonthus; valid species).
— Coiffait, 1967a: 434 (Philonthus; subgenus Kenonthus; characters).
— Coiffait, 1974: 303 (Philonthus; subgenus Kenonthus?; characters; Caucasus).
— Boháč, 1988: 551, 552 [= 1989: 35] (Philonthus; characters; synonym of rotundicollis).
— Hodge and Jones, 1995: 40 (Philonthus; aberration of varius; characters; Britain).
scutatus Erichson, 1840: 438 (Philonthus; Type locality: Germania meridionalis; Hungaria; Sardinia; Persia).
— Ménétriés, 1849: 44 (Philonthus; synonym of rotundicollis).
— Hochhuth, 1849: 134 (Philonthus; characters; Helenendorf).
— Redtenbacher, 1849: 701 (Philonthus; characters; Austria).
— Hardy, 1851: 26 (Philonthus; notes; England).
— Fairmaire and Laboulbène, 1856: 517 (Philonthus; characters; France).
— Kraatz, 1857d: 580 (Philonthus; characters; Austria, Germany).
— Redtenbacher, 1857: 191 (Philonthus; characters; Austria).
— Thomson, 1860: 156 (Philonthus; characters; Scandinavia).
— Hochhuth, 1862: 53 (Philonthus; Russia).
— Fauvel, 1874: 230 [= 1874a: 456] (Philonthus; synonym of rotundicollis).
— Redtenbacher, 1874: 210 (Philonthus; characters; Austria).
— Seidlitz, 1875: 273 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 389 [= 1877: 245] (Philonthus; characters; France).
— J. Sahlberg, 1876: 13 (Philonthus; synonym of rotundicollis).
— Fowler, 1888: 264 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 446 (Philonthus; synonym of rotundicollis).
— Bernhauer and Schubert, 1914: 353 (Philonthus; synonym of rotundicollis).
— Gridelli, 1920a: 16 (Philonthus; aberration of varius; characters; Italy).
— Cameron, 1932: 84 (Philonthus; synonym of rotundicollis).
— Scheerpeltz, 1933: 1358 (Philonthus; synonym of rotundicollis).
— Smetana, 1958: 170 (Philonthus; synonym of rotundicollis).
— Coiffait, 1974: 227 (Philonthus; synonym of rotundicollis).
— Scheerpeltz, 1974c: 22 (Philonthus; variety of peregrinus; Sudan; Somalia).
— Pope, 1977: 29 (Philonthus; synonym of rotundicollis).
— Boháč, 1988: 552 [= 1989: 35] (Philonthus; synonym of rotundicollis).
duplopunctatus Motschulsky, 1860: 567 (Philonthus; Type locality: lac Baical).
— Hochhuth, 1862: 53 (Philonthus; characters; Transbaikal).
— Fauvel, 1874: 230 [= 1874a: 456] (Philonthus; synonym of rotundicollis).
— Ganglbauer, 1895: 446 (Philonthus; synonym of rotundicollis).
— Bernhauer and Schubert, 1914: 353 (Philonthus; synonym of rotundicollis).
— Cameron, 1932: 84 (Philonthus; synonym of rotundicollis).
— Smetana, 1958: 170 (Philonthus; synonym of rotundicollis).
— Coiffait, 1974: 227 (Philonthus; synonym of rotundicollis).
— Boháč, 1988: 552 [= 1989: 35] (Philonthus; synonym of rotundicollis).
sinuatocollis Motschulsky, 1860b: 121 (Philonthus; Type locality: Nord de la Daourie jusqu'au Kamtschatka).
— Motschulsky, 1862a: 11 (Philonthus; Japan).
— Fauvel, 1874: 230 [= 1874a: 456] (Philonthus; synonym of rotundicollis).
— Ganglbauer, 1895: 446 (Philonthus; synonym of rotundicollis).
— Bernhauer and Schubert, 1914: 353 (Philonthus; synonym of rotundicollis).
— Cameron, 1932: 84 (Philonthus; synonym of rotundicollis).
— Smetana, 1958: 170 (Philonthus; synonym of rotundicollis).
— Coiffait, 1974: 227 (Philonthus; synonym of rotundicollis).
— Boháč, 1988: 552 [= 1989: 35] (Philonthus; cited as sinuaticollis; synonym of rotundicollis).
nigropolitus Bernhauer, 1916d: 26 (Philonthus; subspecies of rotundicollis; Type locality: Turkestan: Aulie Ata, Alai; Tien-schan).
— Scheerpeltz, 1933: 1359 (Philonthus; variety of rotundicollis).
— Coiffait, 1967a: 395 (Philonthus; characters; Turkestan; central Asia).
— Coiffait, 1974: 227 (Philonthus; subspecies of rotundicollis).
inopinatus Smetana, 1963c: 294 (Philonthus; subgenus Philonthus; subspecies of rotundicollis; Type locality: Nord-Mongolei, Karakorum).
— Smetana, 1967d: 203 (Philonthus; subspecies of rotundicollis; Mongolia).
— Coiffait, 1974: 227 (Philonthus; subspecies of rotundicollis).
— Smetana 1975c: 169 (Philonthus; subspecies of rotundicollis; Mongolia).
ruandae Cameron, 1950, see: vanhoofi Bernhauer, 1935.
rubellus Solsky, 1874, see: dimidiatipennis Erichson, 1840.
ruber Hatch, 1957 [Note: Nomen nudum], see: aurulentus Horn, 1884.
rubidus Erichson, 1840, see: quisquiliarius Gyllenhal, 1810.
rubiginosus Solsky, 1867, see: turbidus Erichson, 1840.
rubricatus Tottenham, 1962: 212 (Philonthus; Type locality: Nigeria: Jos).
rubricollis Motschulsky, 1858: 665 (Philonthus; Type locality: Indes orientales).
— Bernhauer and Schubert, 1914: 353 (Philonthus; catalog).
— Cameron, 1932: 141 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1359 (Philonthus; catalog).
▴ Distribution: “East Indies”, India, China.
erythrostictus Kraatz, 1859: 94 (Philonthus; Type locality: India orientalis).
— Bernhauer and Schubert, 1914: 353 (Philonthus; synonym of rubricollis).
— Cameron, 1932: 141 (Philonthus; synonym of rubricollis).
obscuricollis Bernhauer, 1935: 7 (Philonthus; variety of rubricollis; [preoccupied]; Type locality: China: Amoy).
rubripennis Kiesenwetter, 1844, see: Gabrius.
rubripennis Stephens, 1832: 239 (Philonthus; Type locality: Norwich; on the coast of Norfolk).
— Curtis, 1829: 26 (Philonthus; nomen nudum; catalog; Britain).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Stephens, 1839: 398 (Philonthus; characters; England).
— Hardy, 1851: 30 (Philonthus; synonym of rubripennis).
— Rye, 1864a: 56 (Philonthus; characters; notes; Britain).
— Fauvel, 1874: 259 [= 1874a: 485] (Philonthus; synonym of fulvipes).
— Ganglbauer, 1895: 457 (Philonthus; synonym of fulvipes).
— Bernhauer and Schubert, 1914: 339 (Philonthus; synonym of fulvipes).
— Tottenham, 1949: 419 (Philonthus; valid species; Britain).
— Smetana, 1958: 221 (Philonthus; synonym of fulvipes).
— Coiffait, 1974: 92 (Paragabrius; synonym of fulvipes).
— Pope, 1977: 29 (Philonthus; valid species; Britain; [Note: Pope considered fulvipes as used by British authors to be a misidentification of rubripennis]).
— Lohse and Lucht, 1989: 168 (Philonthus; note).
— Pilon and Zanetti, 1991: 64 (Philonthus; Italy).
— Smetana, 1993b: 48 (Philonthus; subgenus Paragabrius; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Smetana, 1995e: 358 (Philonthus; North American records are erroneous).
— M. Hansen, 1996: 106 (Philonthus; valid species; Denmark).
— Assing, Frisch, Kahlen, et al., 1998: 138 (Philonthus; valid name).
▴ Distribution: Algeria, Europe, Turkey, Azerbaijan.
fulvipes Fabricius, 1793: 526 (Staphylinus; [preoccupied]; Type locality: Germania).
— Panzer, 1795: 355 (Staphylinus; characters; Germany).
— Panzer, 1796c: 9 (Staphylinus; characters; illustration).
— Fabricius, 1801: 597 (Staphylinus; characters).
— Turton, 1802: 512 (Staphylinus; characters; Germany).
— Gravenhorst, 1802: 24 (Staphylinus; characters).
— Latreille, 1804: 314 (Staphylinus; characters).
— Gravenhorst, 1806: 75 (Staphylinus; characters).
— Gyllenhal, 1810: 343 (Staphylinus; characters; Sweden).
— Zetterstedt, 1828: 71 (Staphylinus; synonym of laevigatus Gyllenhal).
— Curtis, 1829: 24 (Quedius; catalog; Britain).
— Mannerheim, 1830: 30 (Staphylinus; Finland; Germany).
— Mannerheim, 1831: 444 (Staphylinus; Finland; Germany).
— C. Sahlberg, 1830: 329 (Staphylinus; characters; Finland).
— Lacordaire, 1835: 402 (Staphylinus; characters; France).
— Nordmann, 1837: 102 (Philonthus; Europe).
— Kirby, 1837: 93 (Philonthus; characters; notes; boreal America).
— Erichson, 1839a: 473 (Philonthus; characters; Germany).
— Heer, 1839: 270 (Philonthus; characters; Switzerland).
— Erichson, 1840: 485 (Philonthus; characters; Europe; America septentrionalis).
— Laporte, 1840: 178 (Staphylinus; characters; France; Sweden; Germany).
— Kiesenwetter, 1848a: 22 (Philonthus; characters; Europe).
— Jacobsen, 1849: 703 (Philonthus; characters; Denmark).
— Hochhuth, 1849: 143 (Philonthus; characters; Lenkoran).
— Redtenbacher, 1849: 706 (Philonthus; characters; Austria).
— Kiesenwetter, 1851a: 417 (Philonthus; France).
— Hardy, 1851: 30 (Philonthus; notes; England).
— Fairmaire and Laboulbène, 1856: 528 (Philonthus; characters; France).
— Kraatz, 1857d: 614 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 196 (Philonthus; characters; Austria).
— Peyron, 1858: 427 (Philonthus; Turkey).
— Thomson, 1860: 166 (Philonthus; characters; Scandinavia).
— Fauvel, 1874: 259 [= 1874a: 485] (Philonthus; characters; France; Turkey; Caucasus; Transcaucasia).
— Redtenbacher, 1874: 214 (Philonthus; characters; Austria).
— Seidlitz, 1875: 278 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 506 [= 1877: 362] (Philonthus; characters; France).
— J. Sahlberg, 1876: 21 (Philonthus; Finland).
— Provancher, 1877: 730 (Philonthus; characters; Quebec).
— Fauvel, 1886: 57 [= 1886a: 49] (Philonthus; Algeria).
— Fowler, 1888: 279 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 457 (Philonthus; characters; Palaearctic region; North America).
— Fauvel, 1902b: 113 (Philonthus; Algeria).
— Reitter, 1909: 130 (Philonthus; characters; Germany).
— Blatchley, 1910: 385 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 339 (Philonthus; catalog).
— Johansen, 1914: 413 (Philonthus; characters; Denmark).
— Notman, 1919: 135 (Philonthus; New York).
— Notman, 1920b: 706 (Philonthus; Florida).
— Gridelli, 1924e: 177 (Philonthus; Caucasus).
— Portevin, 1929: 364 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1343 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Tottenham, 1949: 419 (Philonthus; synonym of rubripennis).
— V. Hansen, 1952: 77 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1958: 221 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Coiffait, 1963: 12 (Paragabrius).
— Palm, 1963b: 95 (Philonthus; characters; Sweden; Denmark; Finland).
— Lohse, 1964: 185 (Philonthus; subgenus Philonthus; characters; central Europe).
— Smetana, 1964d: 77 (Philonthus; subgenus Philonthus; Czechoslovakia).
— Horion, 1965: 167 (Philonthus; widespread in southern and central Europe; Russia; Finland; Sweden; Britain; Denmark).
— Raianu, 1968a: 468 (Philonthus; Romania).
— Szujecki, 1968a: 731 (Philonthus; Poland).
— Coiffait, 1974: 92 (Paragabrius; characters; Holarctic region).
— Shilov, 1975a: 376 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Pace and Zanetti, 1977: 301 (Paragabrius; Italy).
— Muona, 1979: 14 (Philonthus; synonym of rubripennis).
— Outerelo, 1980a: 56 (Paragabrius; Spain).
— Frank, 1982: 40 (Philonthus; list of parasites).
— Boller, 1983: 117 (Philonthus; larval characters).
— Tóth, 1984: 75 (Philonthus; characters; Hungary).
— Lucht, 1987: 106 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Lohse and Lucht, 1989: 168 (Philonthus; synonym of rubripennis).
— Smetana, 1993b: 48 (Philonthus; synonym of rubripennis).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; synonym of rubripennis).
— M. Hansen, 1996: 106 (Philonthus; synonym of rubripennis).
— Assing, Frisch, Kahlen, et al., 1998: 138 (Philonthus; synonym of rubripennis).
castanopterus Stephens, 1832: 225 (Quedius; Type locality: London).
— Stephens, 1839: 392 (Quedius; characters; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of rubripennis).
mendax Runde, 1835: 10 (Staphylinus; Type locality: Halae).
— Erichson, 1839a: 474 (Philonthus; synonym of fulvipes).
— Erichson, 1840: 485 (Philonthus; synonym of fulvipes).
— Kraatz, 1857d: 614 (Philonthus; synonym of fulvipes).
— Ganglbauer, 1895: 457 (Philonthus; synonym of fulvipes).
— Bernhauer and Schubert, 1914: 339 (Philonthus; synonym of fulvipes).
— Smetana, 1958: 221 (Philonthus; synonym of fulvipes).
— Coiffait, 1974: 92 (Paragabrius; synonym of fulvipes).
obscuripes Brisout, 1863: 34 (Philonthus; Type locality: le midi du la France, les Pyrénées).
— Fauvel, 1867c: 113 [= 1868b: 211] (Philonthus; synonym of fulvipes).
— Fauvel, 1874: 259 [= 1874a: 485] (Philonthus; synonym of fulvipes).
— Ganglbauer, 1895: 457 (Philonthus; synonym of fulvipes).
— Bernhauer and Schubert, 1914: 339 (Philonthus; synonym of fulvipes).
— Coiffait, 1974: 92 (Paragabrius; synonym of fulvipes).
nigripennis Outerelo, 1976a: 258 (Paragabrius; mutation of fulvipes; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Sierra de Guadarrama).
rubrithorax Kirshenblat, 1950, see: Bisnius.
rubrofilator Coiffait, 1967, see: filator Tottenham, 1953.
rubromaculatus Bernhauer, 1914, see: coprophilus Jarrige, 1949.
rubromaculatus Blanchard, 1842: 82 (Philonthus; Type locality: Uruguay: Montevideo).
— Lynch, 1884: 151 (Philonthus; characters; Argentina).
— Bernhauer and Schubert, 1914: 353 (Philonthus; catalog).
— Scheerpeltz, 1933: 1359 (Philonthus; catalog).
▴ Distribution: Uruguay, Argentina.
rubromarginatus Gerhardt, 1909 [Note: Nomen nudum], see: marginatus Ström, 1768.
rubrosuturatus Bernhauer, 1902, see: quisquiliarius Gyllenhal, 1810.
rubrovittatus Tottenham, 1949b: 320 (Philonthus; Type locality: Kivu: Lulenga).
rudebecki Scheerpeltz, 1974: 153 (Philonthus; Type locality: Natal: Royal Natal National Park, The Cascades, 6000 ft.).
rudipennis Fauvel, 1907: 41 (Philonthus; Type locality: Kilimandjaro: Kibonoto).
— Eichelbaum, 1913: 130 (Philonthus; Tanzania).
— Bernhauer and Schubert, 1914: 353 (Philonthus; catalog).
— Tottenham, 1949b: 326 (Philonthus; characters).
— Cameron, 1950g: 36 (Philonthus; Belgian Congo).
— Tottenham, 1956: 226 (Philonthus; Urundi).
— Tottenham, 1962: 181 (Philonthus; characters).
▴ Distribution: Tanzania, Burundi, Congo.
erythropterus Eichelbaum, 1910: 88 (Philonthus; variety of rudipennis; [preoccupied]; Type locality: Meru, 3500 m).
— Eichelbaum, 1913: 130 (Philonthus; variety of rudipennis; Tanzania).
— Bernhauer and Schubert, 1914: 353 (Philonthus; variety of rudipennis).
rudiventris Bernhauer, 1937b: 613 (Philonthus; subgenus Rabigus; Type locality: Kahama).
rufescens Bernhauer, 1915n: 319 (Philonthus; Type locality: Kapland, Umgebung von Kapstadt).
— Scheerpeltz, 1933: 1359 (Philonthus; catalog).
— Tottenham, 1949b: 322 (Philonthus; characters).
▴ Distribution: South Africa.
ruficapillus Reitter, 1887, see: Rabigus.
ruficauda Bernhauer, 1934a: 119 (Philonthus; Type locality: Argentinien: Tucuman).
ruficollis Schubert, 1908, see: Tolmerinus.
ruficornis Melsheimer, 1844, see: discoideus Gravenhorst, 1802.
ruficornis Hochhuth, 1860, see: rufipes Stephens, 1832.
ruficornis Broun, 1880, see: novaezeelandiae Duvivier, 1883.
rufimanus Heer, 1839: 266 (Philonthus; Type locality: Basel; Bern; Genf).
— Redtenbacher, 1849: 703 (Philonthus; [Note: Cited descriptions by Heer and Erichson as referring to the same species]; characters; Austria).
— Kraatz, 1857d: 609 (Philonthus; [Note: Cited descriptions by Heer and Erichson as referring to the same species]).
— Redtenbacher, 1857: 193 (Philonthus; [Note: Cited descriptions by Heer and Erichson as referring to the same species]; characters; Austria).
— Fauvel, 1874: 223 [= 1874a: 449] (Philonthus; characters; France; Germany; Switzerland; Austria; Italy; Russia; Crete; Syria; Caucasus).
— Redtenbacher, 1874: 212 (Philonthus; characters; Austria).
— Herman, 2001: 28 (Philonthus; synonymy).
▴ Distribution: Europe, Russia, Caucasus, Azerbaijan, Turkey, Lebanon, Israel, Syria, Afghanistan, Uzbekistan.
rufimanus Erichson, 1840: 476 (Philonthus; [preoccupied]; Type locality: Carinthia).
— Erichson, 1840: 930 (Philonthus; characters; Switzerland; Sardinia).
— Kolenati, 1846: 21 (Philonthus; Iberia; provincia Elisabethopol).
— Hochhuth, 1849: 140 (Philonthus; characters; Lenkoran).
— Kiesenwetter, 1851a: 417 (Philonthus; France).
— Fairmaire and Laboulbène, 1856: 526 (Philonthus; characters; France).
— Kraatz, 1857d: 609 (Philonthus; characters; Germany).
— Kraatz, 1858a: 61 (Philonthus; Crete).
— Hochhuth, 1862: 63 (Philonthus; Russia).
— Fauvel, 1865a: 297 [= 1865b: 47] (Philonthus; habitat; France).
— Baudi, 1870: 386 (Philonthus; characters; Lebanon).
— Fauvel, 1876a: 73 [= 1876: 258] (Philonthus; Spain).
— Mulsant and Rey, 1876: 488 [= 1877: 344] (Philonthus; characters; France).
— Eppelsheim, 1892: 334 (Philonthus; Turkestan).
— Ganglbauer, 1895: 456 (Philonthus; characters; middle Europe; Mediterranean region).
— Reitter, 1909: 130 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 353 (Philonthus; catalog).
— Portevin, 1929: 362 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1359 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Smetana, 1953: 119 (Philonthus; Turkey).
— Smetana, 1955b: 229 (Philonthus; notes).
— Smetana, 1958: 213 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 162 (Philonthus; characters).
— Scheerpeltz, 1961d: 129 (Philonthus; Italy).
— Lohse, 1964: 184 (Philonthus; subgenus Philonthus; characters; central Europe).
— Scheerpeltz, 1964a: 300 (Philonthus; subgenus Philonthus; Crete).
— Scheerpeltz, 1965b: 402 (Philonthus; Sicily).
— Horion, 1965: 163 (Philonthus; Turkey; Spain; France; Switzerland; Sardinia; Sicily; Italy; Greece; Crete; Cyprus; Macedonia; Bulgaria; Albania; Bosnia Hercegovina; Serbia; Dalmatia; Slovenia; Romania; Slovakia; Austria; Germany).
— Coiffait, 1967a: 425 (Philonthus; characters; Europe).
— Smetana, 1967i: 555 (Philonthus; Turkey).
— Coiffait, 1974: 205 (Philonthus; characters; central Europe; Spain; Italy; Corfu; Turkey; Syria; Caucasus).
— Pace and Zanetti, 1977: 298 (Philonthus; Italy).
— Smetana, 1978: 118 (Philonthus; collecting notes; Israel).
— Tronquet, 1981: 70 (Philonthus; Afghanistan).
— Tóth, 1984: 73 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Czechoslovakia; Austria; Switzerland; France).
— Aranda and Outerelo, 1991: 121 (Philonthus; characters).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic ?; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Herman, 2001: 28 (Philonthus; synonym of rufimanus Heer).
rufimanus Erichson, 1840, see: rufimanus Heer, 1839.
rufimargo Reitter, 1909: 128 (Philonthus; Type locality: Armenisches Gebirge: Gouvernement Erivan).
— Bernhauer and Schubert, 1914: 353 (Philonthus; catalog).
— Scheerpeltz, 1933: 1359 (Philonthus; catalog).
— Smetana, 1965f: 254 (Philonthus; notes).
— Coiffait, 1967a: 398 (Philonthus; characters; Caucasus).
— Coiffait, 1974: 251 (Philonthus; characters; Armenia).
▴ Distribution: Armenia.
rufipennis Solier, 1849, see: hepaticus Erichson, 1840.
rufipennis Gerhardt, 1910 [Note: Nomen nudum], see: discoideus Gravenhorst, 1802.
rufipennis Delahon, 1914 [Note: Nomen nudum], see: Bisnius.
rufipennis Wüsthoff, 1936, see: rectangulus Sharp, 1874.
rufipes Stephens, 1832: 222 (Quedius; [Note: Preoccupied secondary homonym in Quedius]; Type locality: London; [Note: See lectotype designation by Smetana and Herman, 1999]).
— Stephens, 1839: 391 (Quedius; characters; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of fumigatus).
— Smetana and Herman, 1999: 298 (Philonthus; valid species; lectotype designation).
▴ Distribution: Azores, Europe, Russia, Georgia, Turkey.
immundus: Gyllenhal, 1810: 337 (Staphylinus; [Note: Nomen nudum. This name has a long history as the valid name for the species but it is a misidentification. Gyllenhal did not describe this species, he attributed it to Gravenhorst, 1806: 66; however since immundus Gravenhorst is a synonym of ventralis Gravenhorst the species referred to as immundus Gyllenhal needs a new name]; characters; Sweden).
— C. Sahlberg, 1830: 326 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 30 (Staphylinus; Finland; Sweden).
— Mannerheim, 1831: 444 (Staphylinus; Finland; Sweden).
— Nordmann, 1837: 93 (Philonthus; Sweden; Germany; France).
— Heer, 1839: 264 (Philonthus; characters; Switzerland).
— Erichson, 1840: 479 (Philonthus; characters; Sweden).
— Jacobsen, 1849: 701 (Philonthus; characters; Denmark).
— Hochhuth, 1849: 138 (Philonthus; characters).
— Kraatz, 1857d: 596 (Philonthus).
— Thomson, 1860: 164 (Philonthus; synonym of scybalarius).
— Fauvel, 1874: 226 [= 1874a: 452] (Philonthus; characters; France; Caramania).
— Ganglbauer, 1895: 453 (Philonthus; characters; Europe; Asia Minor).
— Bernhauer, 1899a: 435 (Philonthus; [Note: Misidentification: Scheerpeltz, 1933: 1346 cited apenninus Bernhauer as a synonym of immundus; Bernhauer did not describe the species as new, he attributed it to Fiori]; characters; notes).
— Montandon, 1908: 80 (Philonthus; Romania).
— Reitter, 1909: 128 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 341 (Philonthus; catalog).
— Johansen, 1914: 406 (Philonthus; characters; Denmark).
— Portevin, 1929: 361 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1346 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Bernhauer, 1940c: 4 (Philonthus; Azores).
— V. Hansen, 1952: 65 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1958: 179 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 155 (Philonthus; characters).
— Palm, 1963b: 82 (Philonthus; characters; Sweden; Denmark).
— Lohse, 1964: 182 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 153 (Philonthus; Turkey; Azores; Russia; Poland; Czechoslovakia; Hungary; Romania; Bosnia Hercegovina; Dalmatia; Italy; Switzerland; France; Belgium; Holland; Britain; Ireland; Denmark; Sweden; Germany; Austria).
— Coiffait, 1967a: 413 (Philonthus; characters; Europe; Asia Minor).
— Smetana, 1967i: 554 (Philonthus; Turkey).
— Raianu, 1968a: 466 (Philonthus; cited as imundus; Romania).
— Szujecki, 1968a: 729 (Philonthus; Poland).
— Coiffait, 1974: 263 (Philonthus; characters; all Europe; Britain; Scandinavia; Asia Minor; Mediterranean region).
— Pope, 1977: 29 (Philonthus; Britain).
— Hinton, 1981: 660 (Philonthus; characters of egg).
— Tóth, 1984: 65 (Philonthus; characters; Hungary).
— Lucht, 1987: 104 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Borges, 1990: Table II (Philonthus; Azores).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy ?).
— Luff, 1996: 30 (Philonthus; collecting notes; Isle of Man).
— Smetana and Herman, 1999: 297 (Philonthus; misidentification and unavailable name; valid species name is rufipes Stephens).
fumigatus Erichson, 1839a: 463 (Philonthus; Type locality: Mark Brandenburg).
— Erichson, 1840: 463 (Philonthus; characters; Germany; France).
— Heer, 1841: 582 (Philonthus; characters; Switzerland).
— Jacobsen, 1849: 697 (Philonthus; characters; Denmark).
— Motschulsky, 1849: 87 (Philonthus; Spain).
— Redtenbacher, 1849: 703 (Philonthus; characters; Austria).
— Fairmaire and Laboulbène, 1856: 524 (Philonthus; characters; France).
— Kraatz, 1857d: 599 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 194 (Philonthus; characters; Austria).
— Peyron, 1858: 425 (Philonthus; Turkey).
— Thomson, 1860: 161 (Philonthus; characters; Scandinavia).
— Hochhuth, 1862: 60 (Philonthus; Russia).
— Fauvel, 1874: 226 [= 1874a: 452] (Philonthus; synonym of immundus).
— Redtenbacher, 1874: 212 (Philonthus; characters; Austria).
— Seidlitz, 1875: 276 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 457 [= 1877: 313] (Philonthus; characters; larval characters; France).
— Fowler, 1888: 271 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 453 (Philonthus; synonym of immundus).
— Bernhauer and Schubert, 1914: 342 (Philonthus; synonym of immundus).
— Scheerpeltz, 1933: 1346 (Philonthus; synonym of immundus).
— Smetana, 1958: 179 (Philonthus; synonym of immundus).
— Coiffait, 1974: 263 (Philonthus; synonym of immundus).
— Pope, 1977: 29 (Philonthus; synonym of immundus).
— Smetana and Herman, 1999: 297 (Philonthus; synonym of rufipes).
ruficornis Hochhuth, 1860: 566 (Philonthus; [preoccupied]; Type locality: Russie méri-donale aux environs de la ville de Kharkov; [Note: See lectotype designation by Gusarov, 1993]).
— Hochhuth, 1862: 58 (Philonthus; characters; southern Russia).
— Bernhauer and Schubert, 1914: 353 (Philonthus; catalog).
— Scheerpeltz, 1933: 1359 (Philonthus; catalog).
— Coiffait, 1967a: 413 (Philonthus; characters; Russia; Crimea; Georgia).
— Coiffait, 1974: 291 (Philonthus; characters; Russia; Crimea; Georgia).
— Gusarov, 1993: 70 (Philonthus; lectotype designation: Rus. mer.; synonym of immundus).
undae Penecke, 1901: 13 (Philonthus; Type locality: Mur bei Graz).
— Bernhauer and Schubert, 1914: 342 (Philonthus; aberration of immundus).
— Scheerpeltz, 1933: 1346 (Philonthus; aberration of immundus).
— Smetana, 1958: 179 (Philonthus; synonym of immundus).
— Coiffait, 1974: 263 (Philonthus; synonym of immundus).
abkhazensis Coiffait, 1963: 27 (Philonthus; subspecies of immundus; Type locality: Otkapa, Abkhazie).
— Coiffait, 1967a: 415 (Philonthus; subspecies of immundus; characters).
— Coiffait, 1974: 263, 264 (Philonthus; subspecies of immundus; characters; Caucasus).
rufipes Boheman, 1848, see: austellus Herman, 2001.
rufithorax Fauvel, 1878f: 264 (Philonthus; Type locality: Nouvelle-Guinée).
— Bernhauer and Schubert, 1914: 353 (Philonthus; catalog).
— Cameron, 1931: 363 (Philonthus; New Guinea).
— Cameron, 1937a: 100 (Philonthus; New Guinea).
— Last, 1968: 360 (Philonthus; New Guinea).
— Last, 1987: 32 (Philonthus; Papua New Guinea).
▴ Distribution: New Guinea.
rufiventris Cameron, 1918, see: cameronianus Scheerpeltz, 1933.
rufiventris Sharp, 1887: 787 (Philonthus; Type locality: Mexico, Amecameca in Morelos).
rufocaudus Sharp, 1885: 412 (Philonthus; Type locality: Panama, Bugaba, Volcan de Chiriqui).
rufocinctus Fauvel, 1878, see: minutus Boheman, 1848.
rufoflavus Cameron, 1936, see: Gabronthus.
rufomarginatus Kraatz, 1859, see: flavocinctus Motschulsky, 1858.
rufoniger Cameron, 1937: 13 (Philonthus; Type locality: Java: Kopeng).
rufoplagiatus Solsky, 1868a: 130 (Philonthus; Type locality: Mexique).
— Sharp, 1885: 400 (Philonthus; synonym of piceatus).
— Bernhauer and Schubert, 1914: 354 (Philonthus; catalog).
▴ Distribution: Mexico.
rufopygus Sharp, 1885: 411 (Philonthus; Type locality: Panama, Bugaba).
rufulus Horn, 1884: 199 (Philonthus; variety of alumnus; Type locality: Not cited).
— Horn, 1884: 198 (Philonthus; [Note: Misidentified as alumnus according to Smetana, 1995e: 430] characters; Florida to Arizona, north to New York).
— Scheerpeltz, 1933: 1332 (Philonthus; variety of alumnus).
— Smetana, 1995e: 432 (Philonthus; characters; notes; British Columbia; Nova Scotia; Ontario; Alabama; Arizona; Arkansas; California; Colorado; Delaware; Florida; Georgia; Illinois; Indiana; Iowa; Kansas; Kentucky; Louisiana; Maine; Maryland; Massachusetts, Michigan; Minnesota; Mississippi; Missouri; Nebraska; New Jersey; New Mexico; New York; North Carolina; Ohio; Oklahoma; Oregon; Pennsylvania; South Carolina; South Dakota; Tennessee; Texas; Utah; Virginia; Washington; Wisconsin; Mexico; Bahama Islands; Cuba).
▴ Distribution: Canada, USA, Mexico, West Indies.
atriceps Horn, 1884: 199 (Philonthus; variety of alumnus; Type locality: New Jersey; [Note: See lectotype designation by Smetana, 1995e]. No locality originally cited).
— Scheerpeltz, 1933: 1332 (Philonthus; variety of alumnus).
— Smetana, 1995e: 430 (Philonthus; lectotype designation: N. J. [New Jersey]; synonym of rufulus).
rufus Fauvel, 1898c: 184 (Philonthus; Type locality: Madagascar Est, Antanambe, baie d'Antongil; [Note: See lectotype designation by Lecoq, 1997]. Also cited from Suberbieville; Diego-Suarez).
— Bernhauer and Schubert, 1914: 354 (Philonthus; catalog).
— Scheerpeltz, 1961g: 238 (Philonthus; Madagascar).
— Lecoq, 1997: 282 (Philonthus; lectotype designation: Madagascar Est, Antanambe, baie d'Antongil; characters; Madagascar).
▴ Distribution: Madagascar.
rugosipennis Chapman, 1939: 68 (Philonthus; Type locality: Kenya Colony: Massif du Marakwet, alt. 2500 m).
rugulipennis Tottenham, 1949b: 301 (Philonthus; subgenus Raucalius; Type locality: Lulua: Kapanga).
rupicola Cameron, 1943, see: Gabrius.
rusticus Sharp, 1885: 421 (Philonthus; Type locality: Mexico, Las Vigas, Cordova).
— Bernhauer and Schubert, 1914: 354 (Philonthus; catalog).
— Scheerpeltz, 1971b: 205 (Philonthus; Guatemala; Mexico; El Salvador).
▴ Distribution: Mexico, Guatemala, El Salvador.
rutilipennis Hochhuth, 1851, see: caucasicus Nordmann, 1837.
rutiliventris Sharp, 1874: 47 (Philonthus; Type locality: Nagasaki; eastern Siberia, Lake Baikal; China).
— Matsumura, 1911: 114 (Philonthus; Japan).
— Bernhauer and Schubert, 1914: 354 (Philonthus; catalog).
— Scheerpeltz, 1933: 1359 (Philonthus; catalog).
— Adachi, 1937: 56 (Philonthus; Izu-Oshima).
— Nakane, 1963a: 91 (Philonthus; characters; habitus photograph; Japan).
— Watanabe and Shibata, 1972: 64 (Philonthus; Yaku-shima Island).
— Yuh, Paik, Kwon, and Lee, 1985: 242 (Philonthus; Korea).
— J. Li, 1993: 35 (Philonthus; China).
▴ Distribution: Russia, China, Korea, Japan.
ruweanus Levasseur, 1962a: 55 (Philonthus; Type locality: Ruwe, région de Kolwezi, Katanga).
sadales Smetana, 1995e: 355 (Philonthus; Type locality: Oregon: Mt. Hood, Timberline Rd., 4500–5000 ft.).
sagax Casey, 1915, see: quadrulus Horn, 1884.
sagax Smetana, 1963, see: sessor Smetana, 1965.
sahlbergorum Renkonen, 1937, see: truquii Peyron, 1858.
salinus Kiesenwetter, 1844: 347, note (Philonthus; Type locality: Leipzig).
— Kiesenwetter, 1848a: 23 (Philonthus; characters; Germany).
— Redtenbacher, 1849: 825 (Philonthus; characters; Austria).
— Kiesenwetter, 1851a: 417 (Philonthus; France).
— Fairmaire and Laboulbène, 1856: 529 (Philonthus; characters; France).
— Kraatz, 1857d: 613 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 196 (Philonthus; characters; Austria).
— Kraatz, 1858a: 61 (Philonthus; Greece).
— Fauvel, 1874: 259 [= 1874a: 485] (Philonthus; characters; France; Germany; Hungary; Greece; Caucasus).
— Solsky, 1874: 181 (Philonthus; characters; Turkestan).
— Fauvel, 1876a: 75 [= 1876: 260] (Philonthus; Turkestan).
— Redtenbacher, 1874: 215 (Philonthus; characters; Austria).
— Mulsant and Rey, 1876: 509 [= 1877: 365] (Philonthus; characters; France).
— Ganglbauer, 1895: 457 (Philonthus; characters; middle Europe; Mediterranean region).
— Montandon, 1908: 80 (Philonthus; Romania).
— Reitter, 1909: 131 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 354 (Philonthus; catalog).
— Johansen, 1914: 414 (Philonthus; characters; Denmark).
— Portevin, 1929: 364 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1359 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— V. Hansen, 1952: 77 (Philonthus; subgenus Philonthus; characters; Denmark).
— Scheerpeltz, 1958a: 21 (Philonthus; Armenia).
— Smetana, 1958: 222 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 166 (Philonthus; characters).
— Scheerpeltz, 1961d: 129 (Philonthus; Italy).
— Scheerpeltz, 1963a: 420 (Philonthus; Greece).
— Raianu, 1963: 83 (Philonthus; Romania).
— Palm, 1963b: 95 (Philonthus; characters; Sweden; Denmark).
— Lohse, 1964: 185 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 169 (Philonthus; Kazakhstan; Uzbekistan; Turkmenistan; Russia; Turkey; Romania; Greece; Italy; France; Germany; Denmark; Sweden; Austria).
— Smetana, 1966: 331 (Philonthus; notes; Czechoslovakia).
— Coiffait, 1967a: 427 (Philonthus; characters; European coasts).
— Raianu, 1968a: 468 (Philonthus; Romania).
— Jarrige, 1971: 494 (Philonthus; Iran).
— Coiffait, 1974: 198 (Philonthus; characters; Europe; Denmark; Sweden; Asia Minor; France).
— Tóth, 1984: 75 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Czechoslovakia; Austria; Denmark; Sweden).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
▴ Distribution: Europe, Armenia, Turkey, Iran, “Turkestan”.
sallaei Sharp, 1885: 398 (Philonthus; Type locality: Mexico, La Parada; [Note: See lectotype designation by Smetana, 1995e]).
— Bernhauer and Schubert, 1914: 354 (Philonthus; catalog).
— Smetana, 1991a: 232 (Philonthus; characters; lectotype designation; Mexico).
▴ Distribution: Mexico.
salti Tottenham, 1953b: 497 (Philonthus; Type locality: Uganda: Kawanda, near Kampala).
— Tottenham, 1953a: 141 (Philonthus; characters in key).
— Levasseur, 1966a: 1214 (Philonthus; Congo; Zaire; Cameroon).
— Levasseur, 1968a: 54 (Philonthus; Central African Republic).
▴ Distribution: Uganda, Congo, Central African Republic, Cameroon.
saltus Last, 1968, see: Hesperus.
samoensis Bernhauer, 1922, see: quisquiliarius Gyllenhal, 1810.
sanamus Tottenham, 1955: 163 (Philonthus; subgenus Spatulonthus; replacement name for nitidicollis Klug. Also cited from South Africa and Rhodesia).
— Tottenham, 1956: 227 (Philonthus; subgenus Spatulonthus; Ruanda).
— Byrne, 1993 (Philonthus; egg, larval, pupal, and adult characters; South Africa).
— Herman, 2001: 27 (Philonthus; synonymy).
▴ Distribution: South Africa, Zimbabwe, Mozambique, Rwanda.
nitidicollis Klug, 1855: 644 (Philonthus; [preoccupied]; Type locality: Mossambique: Sena).
— Fauvel, 1891: 115 (Philonthus; synonym of longicornis).
— Bernhauer and Schubert, 1914: 348 (Philonthus; catalog).
placatus Tottenham, 1955: 163 (Philonthus; subgenus Spatulonthus; form of sanamus; Type locality: Not cited).
klugi Bordoni, 1984: 344 (Philonthus; replacement name for nitidicollis Klug).
sanguineus Fauvel, 1907: 46 (Philonthus; Type locality: Meru; Ikutha; Natal; Senegal; Sierra Leone; Congo).
— Eichelbaum, 1913: 130 (Philonthus; Tanzania).
— Bernhauer and Schubert, 1914: 354 (Philonthus; catalog).
— Bernhauer, 1915n: 319 (Philonthus; German Southwest Africa).
— Tottenham, 1949b: 347 (Philonthus; characters).
— Cameron, 1952e: 832 (Philonthus; Dahomey).
— Tottenham, 1961: 204, 209 (Philonthus; notes; Senegal).
— Tottenham, 1962: 139 (Philonthus; Tanganyika).
— Scheerpeltz, 1974c: 24 (Philonthus; Sudan).
▴ Distribution: Senegal, Sierra Leone, Benin, Congo, Sudan, Tanzania.
sanguinicollis Fauvel, 1877, see: subcingulatus W. J. MacLeay, 1873.
sanguinipennis Quedenfeldt, 1882: 182 (Philonthus; Type locality: Marocco: flumen Tenfift prope urbem Marocco; Urica in monte Atlante).
— Quedenfeldt, 1884: 373 (Philonthus; characters; Morocco).
— Fauvel, 1886: 55 [= 1886a: 47] (Philonthus; notes; Morocco).
— Fauvel, 1902b: 110 (Philonthus; Morocco; Algeria).
— Bernhauer and Schubert, 1914: 354 (Philonthus; catalog).
— Scheerpeltz, 1933: 1359 (Philonthus; catalog).
— Coiffait, 1967a: 417 (Philonthus; characters; Morocco; Algeria).
— Coiffait, 1974: 216 (Philonthus; characters; Algeria; Morocco).
— Maachi, 1991: 250 (Philonthus; Morocco).
▴ Distribution: Morocco, Algeria.
sanguinipennis Reitter, 1898, see: eriwanensis Bernhauer and Schubert, 1914.
sanguinithorax Scheerpeltz, 1965: 211 (Philonthus; Type locality: Burma: Rangoon).
sanguinolentus Gravenhorst, 1802: 36 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae; [Note: See lectotype designation by Smetana, 1995e]).
— Latreille, 1804: 320 (Staphylinus; characters; France; Germany).
— Gravenhorst, 1806: 64 (Staphylinus; characters; Germany; France).
— Gyllenhal, 1810: 338 (Staphylinus; characters; Sweden).
— Curtis, 1829: 26 (Philonthus; catalog; Britain).
— C. Sahlberg, 1830: 326 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 30 (Staphylinus; Finland; Sweden).
— Mannerheim, 1831: 444 (Staphylinus; Finland; Sweden).
— Stephens, 1832: 238 (Philonthus; characters; England).
— Runde, 1835: 8 (Staphylinus; characters; Germany).
— Lacordaire, 1835: 399 (Staphylinus; characters; France).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Nordmann, 1837: 97 (Philonthus; Europe).
— Holme, 1837: 63 (Philonthus; Scilly Islands).
— Erichson, 1839a: 464 (Philonthus; characters; Germany).
— Heer, 1839: 265 (Philonthus; characters; Switzerland).
— Stephens, 1839: 398 (Philonthus; characters; England).
— Erichson, 1840: 467 (Philonthus; characters; Europe).
— Laporte, 1840: 178 (Staphylinus; characters; France).
— Jacobsen, 1849: 698 (Philonthus; characters; Denmark).
— Redtenbacher, 1849: 704 (Philonthus; characters; Austria).
— Küster, 1854: 8 (Philonthus; characters; Europe).
— Hardy, 1851: 29 (Philonthus; notes; England).
— Fairmaire and Laboulbène, 1856: 525 (Philonthus; characters; France).
— Kraatz, 1857d: 600 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 194 (Philonthus; characters; Austria).
— Thomson, 1860: 161 (Philonthus; characters; Scandinavia).
— Fauvel, 1874: 225 [= 1874a: 451] (Philonthus; characters; France; Algeria).
— Redtenbacher, 1874: 213 (Philonthus; characters; Austria).
— Seidlitz, 1875: 276 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 464 [= 1877: 320] (Philonthus; characters; France).
— J. Sahlberg, 1876: 19 (Philonthus; Finland).
— Fauvel, 1878: 123 [= 1878a: 43] (Philonthus; Algeria).
— Fauvel, 1886: 54 [= 1886a: 46] (Philonthus; Algeria).
— Fowler, 1888: 272 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 452 (Philonthus; characters; Europe; Mediterranean region).
— Fauvel, 1902b: 109 (Philonthus; Algeria).
— Reitter, 1909: 129 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 354 (Philonthus; catalog).
— Johansen, 1914: 407 (Philonthus; characters; Denmark).
— Portevin, 1929: 362 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1359 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Paulian, 1941: 227 (Philonthus; larval characters).
— V. Hansen, 1952: 66 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1958: 177 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 155 (Philonthus; characters).
— Palm, 1963b: 83 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 181 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 153 (Philonthus; Algeria; Siberia; Scotland; Lapland; Russia; Spain; Sardinia; Italy; Bosnia; Croatia; Germany; Austria).
— Pototskaia, 1966: 156 (Philonthus; larval characters).
— Pototskaia, 1967: 57 (Philonthus; larval characters).
— Coiffait, 1967a: 417 (Philonthus; characters; Siberia).
— Raianu, 1968a: 464 (Philonthus; Romania).
— Szujecki, 1968a: 729 (Philonthus; Poland).
— Coiffait, 1974: 208 (Philonthus; characters; Europe; Britain; Scandinavia; Siberia; Armenia; north Africa).
— Pace and Zanetti, 1977: 292 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Hinton, 1981: 661 (Philonthus; characters of egg).
— Frank, 1982: 43 (Philonthus; list of parasites).
— Tóth, 1984: 65 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
— Smetana, 1995e: 251 (Philonthus; lectotype designation; characters; British Columbia; Idaho; Oregon; Washington).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
▴ Distribution: Algeria, Europe, Russia, Armenia, Kazakhstan, Canada, USA.
contaminatus Gravenhorst, 1802: 174 (Staphylinus; Type locality: Brunsvigae; [Note: See lectotype designation by Smetana, 1995e]).
— Latreille, 1804: 325 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 64 (Staphylinus; variety of sanguinolentus Gravenhorst).
— Runde, 1835: 9 (Staphylinus; characters; Germany).
— Erichson, 1840: 467 (Philonthus; synonym of sanguinolentus).
— Redtenbacher, 1849: 704 (Philonthus; synonym of sanguinolentus).
— Küster, 1854: 8 (Philonthus; synonym of sanguinolentus).
— Kraatz, 1857d: 600 (Philonthus; synonym of sanguinolentus).
— Fauvel, 1869: 489 (Philonthus; synonym of sanguinolentus).
— Fauvel, 1874: 226 [= 1874a: 452] (Philonthus; synonym of sanguinolentus).
— Ganglbauer, 1895: 452 (Philonthus; aberration of sanguinolentus).
— Bernhauer and Schubert, 1914: 354 (Philonthus; synonym of sanguinolentus).
— Scheerpeltz, 1933: 1359 (Philonthus; synonym of sanguinolentus).
— Smetana, 1958: 178 (Philonthus; synonym of sanguinolentus).
— Coiffait, 1974: 209 (Philonthus; synonym of sanguinolentus).
— Smetana, 1995e: 251 (Philonthus; lectotype designation; synonym of sanguinolentus).
aciculatus Stephens, 1832: 238 (Philonthus; Type locality: London).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Stephens, 1839: 397 (Philonthus; characters; England).
— Hardy, 1851: 29 (Philonthus; synonym of varians).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of sanguinolentus).
— Bernhauer and Schubert, 1914: 354 (Philonthus; aberration of sanguinolentus).
— Tottenham, 1937a: 190 (Philonthus; according to the description by Stephens the species may have been varians but the specimens labelled as types in the collection are sanguinolentus).
— Smetana, 1958: 178 (Philonthus; aberration of sanguinolentus).
— Smetana, 1959: 155 (Philonthus; aberration of sanguinolentus; characters).
— Coiffait, 1967a: 417 (Philonthus; aberration of sanguinolentus; characters).
— Coiffait, 1974: 209 (Philonthus; variety of sanguinolentus).
— Pope, 1977: 29 (Philonthus; synonym of sanguinolentus).
— Smetana, 1995e: 251 (Philonthus; synonym of sanguinolentus).
— Hodge and Jones, 1995: 40 (Philonthus; aberration of sanguinolentus; characters; Britain).
unicolor Seidlitz, 1875: 276 (Philonthus; variety of sanguinolentus; [preoccupied]; Type locality: Not cited).
— Bernhauer and Schubert, 1914: 354 (Philonthus; aberration of sanguinolentus).
— Scheerpeltz, 1933: 1359 (Philonthus; aberration of sanguinolentus).
— Smetana, 1958: 178 (Philonthus; aberration of sanguinolentus).
— Smetana, 1959: 155 (Philonthus; aberration of sanguinolentus; characters).
— Coiffait, 1967a: 417 (Philonthus; aberration of sanguinolentus; characters).
— Coiffait, 1974: 209 (Philonthus; variety of sanguinolentus).
— Tóth, 1984: 65 (Philonthus; aberration of sanguinolentus).
— Smetana, 1995e: 251 (Philonthus; synonym of sanguinolentus).
ater Ragusa, 1892: 167 (Philonthus; variety of sanguinolentus; Type locality: Sicilia: Palermo).
— Bernhauer and Schubert, 1914: 354 (Philonthus; synonym of sanguinolentus).
— Smetana, 1958: 178 (Philonthus; synonym of sanguinolentus).
— Coiffait, 1974: 209 (Philonthus; synonym of sanguinolentus).
— Smetana, 1995e: 252 (Philonthus; synonym of sanguinolentus). suturamonus Tottenham, 1939a: 201 (Philonthus; aberration of sanguinolentus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from England: Bagley Wood).
— Smetana, 1959: 155 (Philonthus; aberration of sanguinolentus; characters).
— Smetana, 1958: 178 (Philonthus; aberration of sanguinolentus).
— Coiffait, 1967a: 417 (Philonthus; aberration of sanguinolentus; characters).
— Coiffait, 1974: 209 (Philonthus; variety of sanguinolentus).
— Pope, 1977: 29 (Philonthus; synonym of sanguinolentus).
— Smetana, 1995e: 252 (Philonthus; synonym of sanguinolentus).
— Hodge and Jones, 1995: 40 (Philonthus; aberration of sanguinolentus; characters; Britain).
sanguinosus Fauvel, 1889b: 265 (Philonthus; Type locality: Marais de l'Anse Vata).
— Fauvel, 1903d: 271 (Philonthus; synonym of thermarum).
— Bernhauer and Schubert, 1914: 354 (Philonthus; catalog).
▴ Distribution: New Caledonia.
saphyreus Schillhammer, 2000: 149 (Philonthus; Type locality: China: Sichuan: Daxue Shan, Hailuogou Glacier Park [Gongga Shan] Camp, 2100 m, 29.38N, 102.04E).
sarawakensis Bernhauer, 1915f: 141 (Philonthus; Type locality: Sarawak: Mt. Matang, 1000 Fuss).
— Cameron, 1930b: 162 (Philonthus; British North Borneo).
— Scheerpeltz, 1933: 1360 (Philonthus; catalog).
— Cameron, 1937: 5 (Philonthus; Java).
▴ Distribution: Borneo, Indonesia.
satanus Cameron, 1942b: 327 (Philonthus; Type locality: Chyulu Hills, altitude 5200 feet).
saxatilis Schrank, 1781, see: nitidus Fabricius, 1787.
scansor Gistel, 1857: 70 (Philonthus; [Note: Nomen dubium]; Type locality: Tyrolis).
schaeuffelei Scheerpeltz, 1961: 25 (Philonthus; Type locality: Iran: Khuzistan, Shadegan).
— Coiffait, 1974: 296 (Philonthus; cited as schaeuffeli; characters; Iran).
— Coiffait, 1981b: 236 (Philonthus; cited as schaeuffeli; Saudi Arabia).
▴ Distribution: Iran, Saudi Arabia.
schawalleri Coiffait, 1982: 62 (Philonthus; Type locality: Inde: Cachemire, Sonamarg, Himalaya-Hauptkette, 3000 m).
schoutedeni Bernhauer, 1928g: 115 (Philonthus; Type locality: Belgisch-Kongo: Mongende).
— Scheerpeltz, 1933: 1360 (Philonthus; catalog).
— Tottenham, 1949b: 300 (Philonthus; subgenus Philonthoblerius; characters; Belgian Congo).
▴ Distribution: Congo.
schroederi Eichelbaum, 1912: 174 (Philonthus; Type locality: Parehberge, 1100 m).
schwarzi Horn, 1884: 201 (Philonthus; Type locality: Cambridge, Massachusetts; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from Massachusetts; Detroit).
— Blatchley, 1910: 385 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 354 (Philonthus; catalog).
— Smetana, 1995e: 421 (Philonthus; lectotype designation: Cambr. [presumably Cambridge, Massachusetts]; characters; notes; Manitoba; New Brunswick; Nova Scotia; Ontario; Quebec; Colorado; Connecticut; Illinois; Indiana; Maine; Massachusetts; Michigan; Minnesota; Montana; New Hampshire; New Jersey; New York; Pennsylvania; Vermont; Wisconsin). — Downie and Arnett, 1996: 379 (Philonthus; characters; USA).
▴ Distribution: Canada, USA.
scintillans Solsky, 1868, see: scintillatus Blackwelder, 1944.
scintillatus Blackwelder, 1944: 134 (Philonthus; replacement name for scintillans Solsky).
scintillans Solsky, 1868a: 131 (Philonthus; [preoccupied; secondary homonym of Quedius scintillans (Gravenhorst) in Philonthus where the Gravenhorst name was cited by Redtenbacher (1849: 825)]; Type locality: Mexique).
— Sharp, 1885: 405 (Philonthus; Mexico).
— Bernhauer and Schubert, 1914: 354 (Philonthus; catalog).
— Blackwelder, 1944f: 134 (Philonthus; synonym of scintillatus).
scoticus Joy and Tomlin, 1913, see: Bisnius.
scotti Bernhauer, 1931d: 586 (Philonthus; Type locality: Abyssinia: Mt. Chillalo, 9000 ft. and 10,000 ft.).
— Scheerpeltz, 1933: 1360 (Philonthus; catalog).
— Tottenham, 1949b: 308 (Philonthus; characters).
— Tottenham, 1962: 162, 165 (Philonthus; characters).
▴ Distribution: Ethiopia.
scribae Fauvel, 1867, see: Bisnius.
scutatus Eppelsheim, 1895, see: varians Paykull, 1789.
scutatus Erichson, 1840, see: rotundicollis Ménétriés, 1832.
scutellatus Casey, 1915, see: longicornis Stephens, 1832.
scutellatus Motschulsky, 1860, see: ephippium Nordmann, 1837.
scybalarius Nordmann, 1837, see: longicornis Stephens, 1832.
seattlensis Hatch, 1957, see: Gabrius.
secus Herman, 2001: 48 (Philonthus; replacement name for distinguendus Cameron, 1951).
distinguendus Cameron, 1951c: 400 (Philonthus; [preoccupied]; Type locality: Kenya Colony: Nairobi).
segregatus Cameron, 1933b: 345 (Philonthus; Type locality: British North Borneo: Lumu Lumu, 5500 feet).
seguyi Coiffait, 1987, see: montivagus Heer, 1839.
seishinensis Bernhauer, 1936, see: japonicus Sharp, 1874.
selangorensis Cameron, 1932a: 141 (Philonthus; Type locality: Selangor: Bukit Kutu).
semiaenescens Bernhauer, 1920d: 182 (Philonthus; Type locality: West-Himalaya: Murree, 7500′).
— Cameron, 1932: 94 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1360 (Philonthus; catalog).
— Scheerpeltz, 1965: 106 (Philonthus; Burma).
▴ Distribution: India, Myanmar.
semialutaceus Bernhauer, 1937b: 612 (Philonthus; Type locality: Utzungwe Berge, 1200–1600 m).
semicupreus Fauvel, 1891: 114 (Philonthus; Type locality: Caracas).
semicyaneus Schubert, 1909: 290 (Philonthus; Type locality: südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d'Orizaba).
semiruber Horn, 1884: 187 (Philonthus; Type locality: California; Arizona).
— Fall and Cockerell, 1907: 166 (Philonthus; New Mexico).
— Bernhauer and Schubert, 1914: 355 (Philonthus; catalog).
— Smetana, 1995e: 109 (Philonthus; characters; Alberta; Arizona; California; Colorado; Kansas; Nebraska; New Mexico; Oklahoma; Oklahoma; South Dakota; Texas; Utah).
▴ Distribution: Canada, USA.
semunciarius Tottenham, 1940a: 20 (Philonthus; Type locality: Costa Rica: El Congo, 750 m).
senegalensis Bernhauer, 1937a: 300 (Philonthus; Type locality: Haut Senegal: Bodoumbé).
senium Smetana, 1995e: 413 (Philonthus; Type locality: Massachusetts: Tyngs. Also cited from Maine, Massachusetts, New Hampshire).
separatus Cameron, 1950g: 38 (Philonthus; subgenus Gabrius; Type locality: Belgian Congo: lac N'Gando, pied volc. Karisimbi, alt. 2400 m).
sepilibilis Tottenham, 1949b: 339 (Philonthus; Type locality: Congo Region).
septempunctatus Cameron, 1930, see: Gabrius.
septempunctatus Tikhomirova, 1973, see: kiautschauensis Bernhauer, 1916.
septentrionis Fall, 1926, see: subvirescens Thomson, 1884.
septentrionum Eppelsheim, 1893: 50 (Philonthus; Type locality: Ost-Sibirien, in der Nähe von Irkutsk, von Tunka bis Munku-Sardik im Sajan'schen Gebirge).
— Bernhauer and Schubert, 1914: 355 (Philonthus; catalog).
— Scheerpeltz, 1933: 1360 (Philonthus; catalog).
— Smetana, 1963c: 296 (Philonthus; subgenus Philonthus; Mongolia).
— Smetana, 1965e: 174 (Philonthus; Mongolia).
— Coiffait, 1967a: 398 (Philonthus; characters; central Asia).
— Smetana, 1967d: 206 (Philonthus; Mongolia).
— Coiffait, 1974: 280 (Philonthus; characters; central Asia; Baikal region; Mongolia).
— Smetana 1975c: 171 (Philonthus; Mongolia).
— Shilov, 1976: 151 (Philonthus; Mongolia).
▴ Distribution: Russia, Mongolia.
sequens Bernhauer and Schubert, 1914: 355 (Philonthus; replacement name for imitator Fauvel).
— Scheerpeltz, 1933: 1360 (Philonthus; catalog).
— Bernhauer and Chapman, 1939: 67 (Philonthus; Kenya).
— Tottenham, 1949b: 333 (Philonthus; characters).
— Tottenham, 1962: 222, 225 (Philonthus; specimens of the syntypic series from the Cape are a differenct species, probably biguttulus; characters).
▴ Distribution: Ethiopia, Kenya, South Africa?.
imitator Fauvel, 1907: 44 (Philonthus; [preoccupied]; Type locality: Abyssinie; Cap).
— Scheerpeltz, 1933: 1360 (Philonthus; synonym of sequens).
— Tottenham, 1962: 225 (Philonthus; synonym of sequens).
serenus Herman, 2001: 48 (Philonthus; replacement name for parvicornis Fauvel).
parvicornis Fauvel, 1907: 40 (Philonthus; [preoccupied]; Type locality: Victoria Nyanza: ile de Lusinga; Transvaal, Hamman'skraal).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Tottenham, 1949b: 328 (Philonthus; characters).
— Herman, 2001: 48 (Philonthus; synonym of serenus).
seriatipennis Bernhauer, 1928, see: Mentophilonthus.
sericans Gravenhorst, 1802: 171 (Staphylinus; Type locality: America septentr.).
— Gravenhorst, 1806: 59 (Staphylinus; characters).
— Erichson, 1840: 486 (Philonthus; synonym of brunneus).
— Bernhauer and Schubert, 1914: 331 (Philonthus; synonym of brunneus).
— Smetana, 1995e: 319 (Philonthus; characters; notes; New Brunswick; Ontario; Quebec; Alabama; Arizona; Arkansas; California; Colorado; Connecticut; Florida; Georgia; Illinois; Indiana; Iowa; Kansas; Kentucky; Louisiana; Maine; Maryland; Massachusetts; Michigan; Minnesota; Mississippi; Missouri; Nebraska; New Hampshire; New Jersey; New York; North Carolina; Ohio; Oklahoma; Pennsylvania; Rhode Island; South Carolina; South Dakota; Tennessee; Texas; Vermont; Virginia; West Virginia; Wisconsin; Mexico).
— Hu and Frank, 1995b: 585 (Philonthus; surface structure of eggs).
— Hu and Frank, 1997 (Philonthus; predation of eggs and larvae of Haematobia irritans; Florida).
▴ Distribution: Canada, USA, Mexico, Argentina.
brunneus Gravenhorst, 1802: 172 (Staphylinus; [preoccupied]; Type locality: America septentr.).
— Gravenhorst, 1806: 57 (Staphylinus).
— Erichson, 1840: 486 (Philonthus; characters; America septentrionalis).
— LeConte, 1850: 220 (Philonthus; Lake Superior).
— Horn, 1884: 206 (Philonthus; characters; Canada to Texas, west to Nebraska).
— Blatchley, 1910: 386 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 331 (Philonthus; catalog).
— Scheerpeltz, 1933: 1335 (Philonthus; catalog; Argentina).
— Muona, 1977: 16 (Philonthus; [Note: The name brunneus Gravenhorst is cited as being preoccupied by brunneus Fabricius, 1798. Gravenhorst's name is preoccupied by brunneus Paykull the name used by Fabricius, 1798 was a subsequent usage of Paykull's name]; synonym of sericans).
— Hunter, Kincher, Bay, Beerwinkle, 1991: 168 (Philonthus; cited as valid species; seasonal distribution and diel flight activity; Texas).
— Smetana, 1995e: 319 (Philonthus; synonym of sericans).
— Downie and Arnett, 1996: 379 (Philonthus; characters; USA).
dimidiatus Say, 1830: 37 [= 1834: 450] (Staphylinus; [preoccupied]; Type locality: Pennsylvania and Missouri).
— Erichson, 1840: 929 (Philonthus; synonym of ventralis).
— LeConte, 1859: 565 (Philonthus; probably synonym of ventralis).
— Gemminger and Harold, 1868: 586 (Philonthus; synonym of brevis).
— Bernhauer and Schubert, 1914: 331 (Philonthus; synonym of brunneus).
— Smetana, 1995e: 319 (Philonthus; synonym of sericans).
picatus Kirby, 1837: 93 (Philonthus; [preoccupied]; Type locality: America borealis: lat. 54°).
— Gemminger and Harold, 1868: 586 (Philonthus; synonym of brunneus).
— Horn, 1876: 151 (Philonthus; synonym of brunneus).
— Bernhauer and Schubert, 1914: 331 (Philonthus; synonym of brunneus).
— Smetana, 1995e: 319 (Philonthus; synonym of sericans).
brevis Melsheimer, 1844: 37 (Philonthus; replacement name for dimidiatus Say).
— Bernhauer and Schubert, 1914: 331 (Philonthus; synonym of brunneus).
— Smetana, 1995e: 319 (Philonthus; synonym of sericans).
solitarius Sharp, 1885: 424 (Philonthus; Type locality: Mexico, Jalapa).
— Bernhauer and Schubert, 1914: 355 (Philonthus; catalog).
— Smetana, 1995e: 2 (Philonthus; synonym of sericans [Note: Synonymy cited in Abstract, page 2, but not under species account on page 319]).
sericans Sharp, 1874, see: sublucanus Herman, 2001.
sericatus Cameron, 1930b: 162 (Philonthus; Type locality: North Borneo: Bettotan near Sandakan).
sericeicollis Fauvel, 1878f: 266 (Philonthus; Type locality: Nouvelle-Guinée, Andai; Iles Arou, Wokan; Borneo).
— Fauvel, 1879: 103 (Philonthus; Mysol; Ternate).
— Duvivier, 1883: 148 (Philonthus; cited as sericcicollis; catalog).
— Bernhauer and Schubert, 1914: 355 (Philonthus; catalog).
— Bernhauer, 1926f: 317 (Philonthus; Buru).
— Cameron, 1931: 363 (Philonthus; New Guinea).
— Last, 1987: 32 (Philonthus; New Ireland; New Britain).
▴ Distribution: Borneo, Indonesia, New Guinea, New Ireland, New Britain.
sericeiventris Eppelsheim, 1895, see: morio Boheman, 1848.
sericeus Stephens, 1832: 231 (Philonthus; Type locality: London).
sericinus Horn, 1884: 182 (Philonthus; Type locality: Canada; Pennsylvania; Texas).
— Bernhauer and Schubert, 1914: 355 (Philonthus; catalog).
— Smetana, 1995e: 126 (Philonthus; characters; Ontario; Quebec; Arkansas; Connecticut; Illinois; Indiana; Iowa; Maine; Maryland; Massachusetts; Minnesota; New Hampshire; New Jersey; New York; North Carolina; Ohio; Pennsylvania; Vermont; Virginia; Wisconsin).
— Downie and Arnett, 1996: 376 (Philonthus; characters; USA).
▴ Distribution: Canada, USA.
sericoilius Cameron, 1920, see: Quediosoma.
serpens Sharp, 1885: 422 (Philonthus; Type locality: Guatemala, San Geronimo, Torola; Panama, Volcan de Chiriqui 4000 to 6000 feet; Peña Blanca 3000 to 4000 feet).
serpentinus Horn, 1884, see: Hesperus.
serraticornis Bernhauer, 1908, see: aemulus Tottenham, 1939.
serraticornis Sharp, 1876: 179 (Philonthus; Type locality: Ega).
servilis Cameron, 1932: 431 (Philonthus; Type locality: Waziristan).
sessor Smetana, 1965: 58 (Philonthus; replacement name for sagax Smetana).
— Smetana, 1995e: 361 (Philonthus; characters; notes; New Brunswick; Ontario; Quebec; Iowa; Massachusetts; New Hampshire; New Jersey; New York; Pennsylvania; Vermont; Wisconsin).
▴ Distribution: Canada, USA.
sagax Smetana, 1963a: 1 (Philonthus; [preoccupied]; Type locality: New Jersey. Also cited from Massachusetts, Illinois, and Iowa).
seticornis Sharp, 1885: 410 (Philonthus; Type locality: Guatemala, Cerro Zunil 4000 to 5000 feet).
setosus J. Sahlberg, 1876, see: ebeninus Gravenhorst, 1802.
sexpunctatus Last, 1968: 372 (Philonthus; Type locality: New Guinea: Yentchan, Sepik).
sextus Schaeffer, 1766 [Note: Nomen nudum], see: splendens Fabricius, 1793.
shangzhiensis J. Li, 1993: 52 (Philonthus; Type locality: Mount Laoye, Shangzhi, Heilongjiang Province).
sharpi Fauvel, 1879: 101 (Philonthus; Type locality: Nouvelle Guinée, Dorei).
— Bernhauer and Schubert, 1914: 355 (Philonthus; catalog).
— Last, 1987: 32 (Philonthus; Papua New Guinea).
▴ Distribution: New Guinea.
sharpi C. Koch, 1936 [Note: Nomen nudum], see: nudus Sharp, 1874.
sharpianus Cameron, 1930, see: Gabrius.
shetlandicus Poppius, 1905, see: carbonarius Gravenhorst, 1802.
shulli Hatch, 1957, see: Gabrius.
siculus Gridelli, 1923: 23 (Philonthus; Type locality: Sicilia, Pachino).
— Sainte-Claire Deville, 1926: 120 (Philonthus; subgenus Gabrius; notes; Corsica).
— Scheerpeltz, 1933: 1361 (Philonthus; catalog).
— Coiffait, 1974: 96 (Paragabrius; characters; Corsica; Sardinia; Sicily; Algeria; Morocco).
— Ciceroni and Zanetti, 1995: 28 (Philonthus; Italy).
▴ Distribution: Algeria, Morocco, France, Italy.
sideropterus Kolenati, 1846, see: splendens Fabricius, 1793.
siegwaldii Mannerheim, 1843, see: Bisnius.
signaticornis Mulsant and Rey, 1853, see: Erichsonius.
signifrons Schubert, 1908, see: protenus Schubert, 1906.
silvaticus Cameron, 1922: 119 (Philonthus; Type locality: Jamaica).
— Scheerpeltz, 1933: 1361 (Philonthus; catalog).
— Blackwelder, 1943: 416 (Philonthus; characters; Jamaica).
▴ Distribution: West Indies.
silvestris Bernhauer, 1929h: 194 (Philonthus; Type locality: Mexiko: Desierto de los Leones, 3000 m).
silvicola Bernhauer, 1912h: 479 (Philonthus; Type locality: Zentralafrika: Urwald hinter den Randbergen des Nordwestufers des Tanganjika-Sees).
simlaensis Bernhauer, 1911, see: protenus Schubert, 1906.
simonae Levasseur, 1962: 238 (Philonthus; Type locality: Kolwezi [Lualaba], Haut Katanga, Congo belge).
simonae Hromádka, 1992, see: lubomiri Herman, 2001.
simplex Marsham, 1802, see: albipes Gravenhorst, 1802.
simplex Stephens, 1833, see: ventralis Gravenhorst, 1802.
simpliciventris Bernhauer, 1933a: 39 (Philonthus; Type locality: Szetschwan: Giufu-Shan).
— Schillhammer, 1999a: 62 (Philonthus).
— Schillhammer, 2000: 125 (Philonthus; characters; China; Taiwan; Nepal; India).
▴ Distribution: Taiwan, China, Nepal, India, Europe.
proximus Kraatz, 1859: 80 (Philonthus; [preoccupied]; Type locality: India borealis).
— Fauvel, 1874: 215 [= 1874a: 441] (Philonthus; characters; France; Europe; Siberia; India).
— J. Sahlberg, 1876: 13 (Philonthus; Finland).
— Fowler, 1888: 262 (Philonthus; characters; Britain).
— Bernhauer and Schubert, 1914: 332 (Philonthus; synonym of chalceus).
— Portevin, 1929: 358 (Philonthus; characters; France).
— Cameron, 1932: 80 (Philonthus; synonym of chalceus).
— Scheerpeltz, 1933: 1336 (Philonthus; synonym of chalceus).
— Tottenham, 1939b: 218 (Philonthus; valid species but name is preoccupied).
— Smetana, 1958: 164 (Philonthus; synonym of chalceus).
— Coiffait, 1974: 223 (Philonthus; synonym of succicola).
— Schillhammer, 2000: 125 (Philonthus; synonym of simpliciventris).
explanipes Tottenham, 1939b: 218 (Philonthus; replacement name for proximus Kraatz; valid species).
— Schillhammer, 1999a: 62 (Philonthus; synonym of simpliciventris; [Note: Although Schillhammer did not list proximus Kraatz as a junior synonym of simpliciventris, it must be regarded as such because explanipes is a replacement name for it]).
— Schillhammer, 2000: 125 (Philonthus; synonym of simpliciventris).
simulans Wollaston, 1857, see: Gabrius.
sinayotus Tottenham, 1962: 226 (Philonthus; subgenus Spatulonthus; Type locality: Tanganyika Terr.: Bunduki, Uluguru Mt., moy. Mgeta, 1300 m. Also cited from Kenya).
sinetuber Coiffait, 1977, see: Craspedomerus.
singgalangensis Cameron, 1933d: 388 (Philonthus; Type locality: Sumatra: Gunung Singgalang).
singhalensis Cameron, 1932: 96 (Philonthus; Type locality: Ceylon: Dikoya, alt. 3800–4200 feet. Bogawantalawa, alt. 4900–5200 feet).
singularis Last, 1973, see: Hesperus.
sinuatocollis Motschulsky, 1860, see: rotundicollis Ménétriés, 1832.
sinuatus Wollaston, 1867, see: quisquiliarius Gyllenhal, 1810.
sinuatus Tottenham, 1949, see: curvabilis Herman, 2001.
sinus Last, 1968: 363 (Philonthus; Type locality: New Guinea: Okapa).
sithanus Tottenham, 1962: 215, 216, 221 (Philonthus; Type locality: Not cited, presumably Africa).
siwalikensis Cameron, 1926b: 357 (Philonthus; Type locality: Siwaliks; Kheri Rau, Mohan Rau, Lachiwala, Song River).
— Cameron, 1932: 137 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1361 (Philonthus; catalog).
▴ Distribution: India.
smaragdinus Tottenham, 1949b: 348 (Philonthus; Type locality: Ituri: Blukwa).
sobrinus Erichson, 1840, see: Neobisnius.
socius Cameron, 1937: 10 (Philonthus; Type locality: Java: Djeroeklegi, Zuid Banjoemas; Tjilatjap).
sodalis Cameron, 1943, see: Gabrius.
sogdianus Kirshenblat, 1950, see: Bisnius.
solidus Sharp, 1874: 43 (Philonthus; Type locality: Nagasaki; northern China).
— Fauvel, 1903: 159 (Philonthus; synonym of geminus).
— Fauvel, 1904a: 58 (Philonthus; synonym of geminus).
— Bernhauer and Schubert, 1914: 355 (Philonthus; catalog).
— Scheerpeltz, 1933: 1363 (Philonthus; synonym of tardus).
— Cameron, 1949b: 468 (Philonthus; cited as valid species; Japan).
— Shibata, 1983: 120 (Philonthus; synonym of tardus).
— Kanehisa, Tsumuki, and Kawazu, 1994 (Philonthus; chemical secretions).
— Watanabe, 1998: 313 (Philonthus; cited as valid species; Japan).
▴ Distribution: Japan.
solitarius Sharp, 1885, see: sericans Gravenhorst, 1802.
somaliensis Eppelsheim, 1895: 202 (Philonthus; Type locality: Ganale Gudda, Arussi Galla).
— Fauvel, 1907: 40 (Philonthus; Kilimandjaro; Abyssinia; Gallas).
— Bernhauer and Schubert, 1914: 355 (Philonthus; catalog; Somaliland; British East Africa).
▴ Distribution: “Somaliland”, Ethiopia, Kenya, Tanzania.
somoruensis Last, 1968, see: Leucitus.
sordidioides Scheerpeltz, 1962d: 329 (Philonthus; Type locality: Umgebung von Rustenburg, etwa 100 km NW von Johannesburg in Transvaal).
sosis Tottenham, 1953: 147 (Philonthus; Type locality: Sudan).
— Tottenham, 1962: 138, 194 (Philonthus; synonym of debiliformis).
— Coiffait, 1966b: 338 (Philonthus; cited as valid species; Mauritania; Senegal).
— Coiffait, 1968a: 136 (Philonthus; notes; Chad).
▴ Distribution: Senegal, Mauritania, Chad, Sudan.
sospitalis Gistel, 1857: 37 (Philonthus; [Note: Nomen dubium]; Type locality: Bayern).
spadiceus Sharp, 1889: 41 (Philonthus; Type locality: Kawasaki near Yokohama; Awomori; Sapporo).
— Bernhauer and Schubert, 1914: 356 (Philonthus; catalog).
— Scheerpeltz, 1933: 1361 (Philonthus; catalog).
— Nakane, 1963a: 91 (Philonthus; characters; habitus photograph; Japan).
— Hayashi, 1993: 76 (Philonthus; characters; Japan).
— Kanehisa, Tsumuki, and Kawazu, 1994 (Philonthus; chemical secretions).
▴ Distribution: Japan.
sparsipennis Cameron, 1933b: 344 (Philonthus; Type locality: British North Borneo: Lumu Lumu, 5500 feet).
sparsus P. Lucas, 1846, see: Bisnius.
specialis Cameron, 1937a: 103 (Philonthus; Type locality: Kokoda, alt. 1300 ft.).
— Last, 1968: 360 (Philonthus; New Guinea).
— Last, 1987: 30 (Philonthus; Papua New Guinea).
▴ Distribution: New Guinea.
multiformis Last, 1968: 362 (Philonthus; Type locality: New Guinea: Buana, Sarawaged Range, 3000 ft., 20–30 miles north of Lae).
— Last, 1973: 425 (Hesperus).
— Last, 1987: 30 (Philonthus; synonym of specialis aberration funebris).
funebris Last, 1981: 120 (Philonthus; variety of specialis; [Note: Nomen nudum (Article 15.2)]; cited from Managalase Plateau North District, Papua).
funebris Last, 1987: 30 (Philonthus; aberration of specialis; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from many localities in Papua New Guinea and Irian Jaya).
speciosus Cameron, 1926b: 355 (Philonthus; Type locality: Dehra Dun).
— Cameron, 1932: 122 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1361 (Philonthus; catalog).
— Schillhammer, 1996a: 82 (Philonthus; Turkey; Oman).
▴ Distribution: India, Oman, Turkey.
specularis Fauvel, 1878f: 267 (Philonthus; Type locality: Nouvelle-Guinée, Hatam).
speculicollis Schubert, 1909: 292 (Philonthus; Type locality: südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d'Orizaba).
speculipennis Bernhauer, 1921b: 75 (Philonthus; Type locality: West-Kolumbien: Umgebung von Cali am Cauca, Canon del Mte. Tolima, 1700 m).
speculum Lokay, 1919: 24 (Philonthus; Type locality: Transsylvania, Sibin.).
— Scheerpeltz, 1933: 1361 (Philonthus; catalog).
— Coiffait, 1967a: 389 (Philonthus; characters; Transylvania).
— Coiffait, 1974: 304 (Philonthus; subgenus Kenonthus?; characters; Romania).
— Tóth, 1984: 45 (Philonthus; characters).
▴ Distribution: Romania.
spermophili Ganglbauer, 1897, see: Bisnius.
sphagnicola Sjöberg, 1950, see: Gabrius.
sphagnorum Smetana, 1995e: 388 (Philonthus; Type locality: New Brunswick: Kouchi-bouguac N.P. Also cited from Ontario and Quebec).
spiniformis Hatch, 1957: 188 (Philonthus; Type locality: Oregon: West Woodburn. Also cited from Washington and British Columbia).
— Smetana, 1995e: 224 (Philonthus; characters; notes; Alberta; British Columbia; Manitoba; Nova Scotia; Ontario; Saskatchewan; Arizona; California; Colorado; Idaho; Iowa; Maine; Montana; Nevada; New Hampshire; New Mexico; New York; North Carolina; North Dakota; Oregon; Pennsylvania; Utah; Washington; Wisconsin; Wyoming).
▴ Distribution: Canada, USA.
woodburnensis Hatch, 1957: 187 (Philonthus; Type locality: Oregon: West Woodburn).
spinipes Sharp, 1874: 39 (Philonthus; Type locality: Japan; [Note: See lectotype designation by Schülke and Uhlig, 1989]. Originally cited from Hiogo; Nagasaki; Mantchuria).
— Bernhauer and Schubert, 1914: 356 (Philonthus; catalog).
— Scheerpeltz, 1933: 1361 (Philonthus; catalog).
— Aleksandrov, 1934: 152 (Philonthus; listed with three characterized but unnamed varieties; notes; China).
— Adachi, 1937: 56 (Philonthus; Izu-Oshima).
— Nakane, 1963a: 90 (Philonthus; characters; habitus photograph; Japan).
— Watanabe and Shibata, 1972: 63 (Philonthus; Yaku-shima Island).
— Frank, 1982: 43 (Philonthus; list of parasites).
— Yuh, Paik, Kwon, and Lee, 1985: 242 (Philonthus; Korea).
— Lundberg, 1986: 148 (Philonthus; Sweden).
— Schülke and Uhlig, 1989: 243, 247 (Philonthus; characters; lectotype designation: Japan; Korea; China; Japan; Caucasus; Bulgaria; Romania; Czechoslovakia; Poland; Austria).
— Schülke and Uhlig, 1989b (Philonthus; notes; Bulgaria; Armenia; Yugoslavia; Austria; Czech Republic; Slovakia; Germany).
— Lohse and Lucht, 1989: 163 (Philonthus; characters).
— Pilon and Zanetti, 1991: 61 (Philonthus; notes; Italy).
— Shibata, 1991b: 256 (Philonthus; Taiwan).
— Callot, 1993: 13 (Philonthus; characters; notes; France).
— Bruge, 1993: 29 (Philonthus; subgenus Kenonthus; Belgium).
— Pedersen, 1993 (Philonthus; characters; Denmark).
— J. Li, 1993: 33 (Philonthus; characters; China).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; introduced species; Czech Republic; Slovakia).
— Kanehisa, Tsumuki, and Kawazu, 1994 (Philonthus; chemical secretions).
— Bruge, 1994a (Philonthus; subgenus Kenonthus; Belgium; Italy).
— Ciceroni and Zanetti, 1995: 29 (Philonthus; introduced in Italy).
— Allen and Owen, 1997 (Philonthus; characters; collecting notes; Britain).
— Tronquet, 1998: 11 (Philonthus; notes; France).
— Schillhammer, 1999: 58 (Philonthus; characters; Japan; China: South Korea; Russia).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
— Tronquet, 1999a: 214 (Philonthus; notes; France).
— Schillhammer, 2000: 159 (Philonthus; note on subspecies).
— Burakowski, Mroczkowski, and Stefańska, 2000: 51 (Philonthus; subgenus Philonthus; catalog; Poland).
▴ Distribution: Europe, Armenia, Kazakhstan, China, Korea, Japan, Taiwan.
kabardensis Bolov and Kryzhanovskij, 1969: 515 (Kirschenblatia; Type locality: Kabardino-Balkarskaia ASSR, Argudan v 35 km iugo-vostochnee Nal'chika).
— Bolov and Kryzhanovskij, 1969a: 329 (Kirschenblatia; characters; Kabardino-Balkaria, Argudan, 35 mi. southeast of Nal'chik).
— Boháč, 1977: 22 (Kirschenblatia; characters).
— Schülke and Uhlig, 1989: 247 (Philonthus; synonym of spinipes).
— Schillhammer, 1999: 58 (Philonthus; synonym of spinipes).
— Schillhammer, 2000: 159 (Philonthus; note).
buchari Boháč, 1977: 20 (Kirschenblatia; Type locality: Caucasus c., 10 km north of Tyrnyauz, 1500 m).
— Schülke and Uhlig, 1989: 247 (Philonthus; synonym of spinipes).
— Schillhammer, 1999: 58 (Philonthus; synonym of spinipes).
hulunbeierensis J. Li, 1993: 35 (Philonthus; subspecies of spinipes cited as spinipts; Type locality: Hairaerbayinhushe, N.E. China).
splendens Fabricius, 1793: 523 (Staphylinus; Type locality: Germania).
— Panzer, 1795: 353 (Staphylinus; characters; Germany).
— Cederhjelm, 1798: 118 (Staphylinus; characters).
— Paykull, 1800: 373 (Staphylinus; characters; Sweden).
— Fabricius, 1801: 594 (Staphylinus; characters; Germany).
— Turton, 1802: 511 (Staphylinus; characters; Germany).
— Gravenhorst, 1802: 16 (Staphylinus; characters).
— Latreille, 1804: 306 (Staphylinus; characters; Sweden; Germany).
— Gravenhorst, 1806: 87 (Staphylinus; characters; Prussia).
— Gyllenhal, 1810: 297 (Staphylinus; characters; Sweden).
— Zetterstedt, 1828: 69 (Staphylinus; characters; Lapland).
— Curtis, 1829: 25 (Philonthus; catalog; Britain).
— C. Sahlberg, 1830: 313 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 25 (Staphylinus; Finland; Sweden).
— Mannerheim, 1831: 439 (Staphylinus; Finland; Sweden).
— Stephens, 1832: 227 (Philonthus; characters; England).
— Ménétriés, 1832: 145 (Staphylinus; Caucasus).
— Runde, 1835: 5 (Staphylinus; characters; Germany).
— Lacordaire, 1835: 387 (Staphylinus; characters; France).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Nordmann, 1837: 73 (Philonthus; characters; Sweden; Russia; Germany; France).
— Holme, 1837: 63 (Philonthus; Scilly Islands).
— Zetterstedt, 1838: 60 (Staphylinus; characters; habitat; Lapland).
— Erichson, 1839a: 447 (Philonthus; characters; Germany).
— Heer, 1839: 258 (Philonthus; characters; Switzerland).
— Stephens, 1839: 394 (Philonthus; characters; England).
— Erichson, 1840: 429 (Philonthus; characters; Europe).
— Laporte, 1840: 177 (Staphylinus; characters; France).
— Kolenati, 1846: 18 (Philonthus; Transcaucasia).
— Jacobsen, 1849: 687 (Philonthus; characters; Denmark).
— Ménétriés, 1849: 44 (Philonthus; Turcomanie).
— Hochhuth, 1849: 131 (Philonthus; characters; Caucasus; Iberia).
— Redtenbacher, 1849: 699 (Philonthus; characters; Austria).
— Kiesenwetter, 1851a: 414 (Philonthus; France).
— Küster, 1853a: 42 (Philonthus; characters; Europe).
— Hardy, 1851: 25 (Philonthus; notes; England).
— Fairmaire and Laboulbène, 1856: 513 (Philonthus; characters; France).
— Kraatz, 1857d: 569 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 189 (Philonthus; characters; Austria).
— Kraatz, 1858a: 61 (Philonthus; Crete).
— Thomson, 1860: 155 (Philonthus; characters; Scandinavia).
— Hochhuth, 1872: 136 (Philonthus; notes; Ukraine).
— Fauvel, 1874: 213 [= 1874a: 439] (Philonthus; characters; France; Europe; Algeria; Caucasus; Transcaucasia; Lake Baikal).
— Redtenbacher, 1874: 208 (Philonthus; characters; Austria).
— Seidlitz, 1875: 272 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 347 [= 1877: 203] (Philonthus; characters; France).
— J. Sahlberg, 1876: 12 (Philonthus; Finland).
— Fauvel, 1878: 121 [= 1878a: 41] (Philonthus; Algeria).
— Fauvel, 1886: 53 [= 1886a: 45] (Philonthus; Algeria).
— Fowler, 1888: 260 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 442 (Philonthus; characters; Europe; Mediterranean region; Siberia).
— Fauvel, 1902b: 108 (Philonthus; Algeria).
— Reitter, 1909: 124 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 356 (Philonthus; [Note: The name politus Linné is cited on page 356 as a synonym of splendens and on page 350 as a valid species]; catalog).
— Johansen, 1914: 392 (Philonthus; characters; Denmark).
— Šulc, 1922 (Philonthus; osmeterium; anatomy; histology; physiology).
— Portevin, 1929: 357 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1361 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Paulian, 1941: 229 (Philonthus; larval characters).
— Tottenham, 1949: 417 (Philonthus; checklist; Britain).
— V. Hansen, 1952: 48 (Philonthus; subgenus Philonthus; characters; Denmark).
— Smetana, 1958: 155 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Kocher, 1958: 144 (Philonthus; Morocco).
— Smetana, 1959: 144 (Philonthus; characters).
— Palm, 1963b: 64 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 176 (Philonthus; subgenus Philonthus; characters; central Europe).
— Raianu, 1964: 103 (Philonthus; Romania).
— Horion, 1965: 123 (Philonthus; Mediterranean region; Germany; Austria).
— Pototskaia, 1966: 158 (Philonthus; larval characters).
— Pototskaia, 1967: 63 (Philonthus; larval characters).
— Coiffait, 1967a: 389 (Philonthus; characters; Europe; Siberia; North Africa).
— Raianu, 1968a: 462 (Philonthus; Romania).
— Szujecki, 1968a: 727 (Philonthus; Poland).
— Scheerpeltz, 1968a: 74 (Philonthus; catalog; Austria).
— Kasule, 1970: 68 (Philonthus; larval characters).
— Coiffait, 1974: 266 (Philonthus; characters; Europe; Britain; Scandinavia; North Africa; Anatolia; Caucasus).
— Shilov, 1975a: 375 [= 1975b: 97] (Philonthus; Komi region of Russia).
— Pope, 1977: 29 (Philonthus; Britain).
— Muona, 1979: 14 (Philonthus; Finland; Russia; Sweden; Norway; Denmark; Baltic States).
— Burakowski, Mroczkowski, and Stefańska, 1980: 67 (Philonthus; catalog; Poland).
— Hinton, 1981: 661 (Philonthus; characters of egg).
— Waterston, Owen, Welch, Bacchus and Hammond, 1981: 274 (Philonthus; Outer Hebrides).
— Boller, 1983: 119 (Philonthus; larval characters).
— Tóth, 1984: 45 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Pilon and Zanetti, 1991: 56 (Philonthus; Italy).
— Silfverberg, 1992: 17 (Philonthus; Finland; Russia; Sweden; Norway; Denmark; Estonia; Latvia; Lithuania).
— Welch, 1993: 228 (Philonthus; ovariole number and ovary structure).
— J. Li, 1993: 35 (Philonthus; China [Note: This may be a misidentification (Schillhammer, personal communication)]).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Hayashi, 1994: 115 (Philonthus; characters; England).
— Ciceroni and Zanetti, 1995: 29 (Philonthus; Italy).
— Ádám, 1996: 244 (Philonthus; synonym of niger).
— Smoleński, 1997: 79 (Philonthus; collecting notes; Poland).
— M. Hansen, 1996: 104 (Philonthus; checklist; Denmark).
— Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 37 (Philonthus; Latvia).
— Anderson, Nash, and O'Connor, 1997: 19 (Philonthus; Ireland).
— Owen, 1997a: 149 (Philonthus; collecting sites; Ireland).
— Kashcheev, 1999a: 57 (Philonthus; Kazakhstan).
— Schillhammer, 2000: 141 (Philonthus; characters; Russia; Korea).
— Herman, 2001: 28 (Philonthus; prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.3).
▴ Distribution: Algeria, Europe, Russia, Georgia, Turkey, Turkmenistan, Kazakhstan, China ?, Korea.
niger O. Müller, 1764: 23 (Staphylinus; [Note: This name is older than splendens]; Type locality: Not cited).
— O. Müller, 1776: 98 (Staphylinus; characters).
— Goeze, 1777: 725 (Staphylinus; characters).
— Villers, 1789: 424 (Staphylinus; characters).
— Gmelin, 1790: 2032 (Staphylinus; page misnumbered as 1032; characters; Denmark).
— Erichson, 1840: 429 (Philonthus; synonym of splendens).
— Kraatz, 1857d: 569 (Philonthus; synonym of splendens).
— Bernhauer and Schubert, 1914: 356 (Philonthus; synonym of splendens).
— Smetana, 1958: 155 (Philonthus; synonym of splendens).
— Ádám, 1996: 244 (Philonthus; valid name; Hungary).
— Schillhammer, 2000: 141 (Philonthus; synonym of splendens).
— Herman, 2001: 28 (Philonthus; synonym of splendens).
sextus Schaeffer, 1766: xxxix, 12 (Staphylinus; [Note: Nomen nudum, Article 11.4]; Type locality: Ratisbon).
— Turton, 1802: 511 (Staphylinus; characters; cited as equivalent to politus).
— Erichson, 1840: 429 (Philonthus; synonym of splendens).
— Kraatz, 1857d: 569 (Philonthus; synonym of splendens).
— Schillhammer, 2000: 141 (Philonthus; synonym of splendens).
sideropterus Kolenati, 1846: 18 (Philonthus; variety of splendens; Type locality: Transcaucasia).
— Hochhuth, 1849: 129 (Philonthus; characters; Tiflis; Helenendorf).
— Kraatz, 1857d: 569 (Philonthus; synonym of splendens).
— Ganglbauer, 1895: 442 (Philonthus; synonym of splendens).
— Bernhauer and Schubert, 1914: 356 (Philonthus; synonym of splendens).
— Scheerpeltz, 1933: 1362 (Philonthus; aberration of splendens).
— Smetana, 1955: 205 (Philonthus; aberration of splendens; notes).
— Smetana, 1958: 155 (Philonthus; subspecies of splendens).
— Smetana, 1959: 144 (Philonthus; subspecies of splendens; characters).
— Coiffait, 1967a: 389 (Philonthus; subspecies of splendens; characters; Caucasus).
— Coiffait, 1974: 266 (Philonthus; synonym of splendens).
— Schillhammer, 2000: 142 (Philonthus; subspecies of sideropterus; characters; Turkey; Caucasus).
squalidus Fauvel, 1879: 105 (Philonthus; Type locality: Nouvelle Guinée).
stacesmithi Hatch, 1957, see: Bisnius.
staudingeri Bernhauer, 1928g: 110 (Philonthus; Type locality: Kongostaat).
— Scheerpeltz, 1933: 1362 (Philonthus; catalog).
— Tottenham, 1962: 202, 203, 211 (Philonthus; characters).
▴ Distribution: Congo.
steineri Lecoq, 1998: 66 (Philonthus; subgenus Piezarthrus; Type locality: Madagascar Est: Prov. Fianarantsoa, 7 km. W Ranomafana, 900 m).
stenocephalus Scheerpeltz, 1972: 18 (Philonthus; replacement name for angusticeps Bernhauer).
angusticeps Bernhauer, 1934a: 118 (Philonthus; [preoccupied]; Type locality: Argentinien: Prov. Tucuman).
stenoderus Reiche, 1861, see: Bisnius.
stercorosus Tottenham, 1962: 186 (Philonthus; Type locality: S. Rhodesia: Wankie Game Reserve).
stictus Hausen, 1891: 321 (Philonthus; Type locality: Lachine, Montreal Island).
— Bernhauer and Schubert, 1914: 356 (Philonthus; catalog).
— Casey, 1915: 450 (Philonthus; republication of characters of original description).
— Smetana, 1995e: 463 (Philonthus; characters; notes; Alberta; British Columbia; Manitoba; New Brunswick; Ontario; Quebec; Saskatchewan; Colorado; Illinois; Iowa; Minnesota; Montana; Nebraska; North Dakota; Utah; Wisconsin).
▴ Distribution: Canada, USA.
protervus Casey, 1915: 443 (Philonthus; Type locality: northern Illinois; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1356 (Philonthus; catalog).
— Dearborn and Donahue, 1994: 15 (Philonthus; Maine).
— Smetana, 1995e: 463 (Philonthus; lectotype designation; synonym of stictus).
— Downie and Arnett, 1996: 382 (Philonthus; characters; USA).
finitimus Casey, 1915: 443 (Philonthus; Type locality: Canada: Hull, Province of Quebec; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1342 (Philonthus; catalog).
— Smetana, 1995e: 463 (Philonthus; lectotype designation; synonym of stictus).
— Downie and Arnett, 1996: 382 (Philonthus; cited as valid species; characters; Canada).
confertoides Hatch, 1957: 182 (Philonthus; Type locality: British Columbia: Enderby).
stigmatias Smetana, 1995, see: ferreipennis Horn, 1884.
stiphrogaster Cameron, 1937: 13 (Philonthus; Type locality: Java: G. Goentoer).
stoetzneri Bernhauer, 1929, see: azuripennis Cameron, 1928.
stoliczkae Sharp, 1878, see: quisquiliarius Gyllenhal, 1810.
stolzmanni Bernhauer, 1916f: 187 (Philonthus; Type locality: Bolivien).
straatmani Last, 1987: 28 (Philonthus; Type locality: Papua New Guinea: Solomon Islands: Bougainville, Mount Balki, 2000–2400 m).
stragulatus Erichson, 1840: 468 (Philonthus; Type locality: Massiliae).
— Fairmaire and Laboulbène, 1856: 525 (Philonthus; characters; France).
— Kraatz, 1857d: 596 (Philonthus).
— Peyron, 1858: 425 (Philonthus; Turkey).
— Fauvel, 1874: 254 [= 1874a: 480] (Philonthus; characters; France; Italy; Sicily; Algeria; Syria; Caramania).
— Mulsant and Rey, 1876: 466 [= 1877: 322] (Philonthus; characters; France).
— Fauvel, 1878: 125 [= 1878a: 45] (Philonthus; Algeria).
— Fauvel, 1886: 56 [= 1886a: 48] (Philonthus; Algeria).
— Ganglbauer, 1895: 453 (Philonthus; characters; Mediterranean region).
— Fauvel, 1902b: 112 (Philonthus; Algeria; Tunisia).
— Bernhauer and Schubert, 1914: 356 (Philonthus; catalog).
— Gridelli, 1920a: 17 (Philonthus; characters; Dalmatia; Croatia).
— Portevin, 1929: 363 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1362 (Philonthus; catalog).
— Smetana, 1965f: 255 (Philonthus; notes).
— Coiffait, 1974: 311 (Spatulonthus; characters; Mediterranean region; North Africa; Lebanon).
— Tóth, 1984: 68 (Philonthus; characters).
— Ciceroni and Zanetti, 1995: 29 (Philonthus; Italy ?).
▴ Distribution: Tunisia, Algeria, France, Italy, Croatia, Turkey, Syria, Lebanon.
strandi Smetana, 1959: 161 (Philonthus; replacement name for jenseni A. Strand).
— Smetana, 1967d: 204 (Philonthus; Mongolia).
— Coiffait, 1974: 294 (Philonthus; characters; Siberia; Mongolia).
— Shilov, 1976: 151 (Philonthus; Mongolia).
▴ Distribution: Russia, Mongolia.
jenseni A. Strand, 1941: 36 (Philonthus; [preoccupied]; Type locality: Sibirien und Mongolei: Kalna, Beikem, Sistikem und Usti Abakansk).
— Smetana, 1959: 161 (Philonthus; synonym of strandi).
— Coiffait, 1967a: 420 (Philonthus; characters; Siberia; Mongolia).
— Smetana, 1967d: 204 (Philonthus; synonym of strandi).
strigicollis Notman, 1919, see: gracilior Casey, 1915.
strigiventris Eppelsheim, 1895, see: Hesperus.
stygialis Sharp, 1885: 405 (Philonthus; Type locality: Guatemala, Balheu in Vera Paz; Panama: Peña Blanca 3000 to 4000 feet).
stysi Hromádka, 1987: 21 (Philonthus; subgenus Philonthus; Type locality: Ethiopia: Bahr-Dar).
suavus Brisout, 1867: 116 (Philonthus; replacement name for gratiosus Brisout).
— Fauvel, 1874: 223 [= 1874a: 449] (Philonthus; cited as suavis; characters; Spain; Portugal).
— Mulsant and Rey, 1876: 491 [= 1877: 347] (Philonthus; cited as suavis; characters; France).
— Bernhauer and Schubert, 1914: 356 (Philonthus; cited as suavis; catalog).
— Scheerpeltz, 1933: 1362 (Philonthus; cited as suavis; catalog).
— Scheerpeltz, 1960d: 135 (Philonthus; Portugal).
— Coiffait, 1967a: 425 (Philonthus; cited as suavis; characters; Iberian Peninsula).
— Coiffait, 1974: 206 (Philonthus; cited as suavis; characters; Spain).
— Aranda and Outerelo, 1991: 121 (Philonthus; cited as suavis; characters).
▴ Distribution: Portugal, Spain, France.
gratiosus Brisout, 1866: 359 (Philonthus; [preoccupied]; Type locality: Espagne: bords du Mançanarès et environs de Madrid).
— Fauvel, 1874: 223 [= 1874a: 449] (Philonthus; synonym of suavus, cited as suavis).
— Bernhauer and Schubert, 1914: 356 (Philonthus; synonym of suavus, cited as suavis).
— Coiffait, 1974: 206 (Philonthus; synonym of suavus, cited as suavis).
subaeneicollis Bernhauer, 1931d: 585 (Philonthus; Type locality: Abyssinia: Mt. Zukwala, ca. 9,000 ft.; Jem-Jem Forest, 8,000–9,000 ft.; Mt. Chillalo, ca. 9,000 ft.).
— Scheerpeltz, 1933: 1362 (Philonthus; catalog).
— Tottenham, 1955: 172 (Philonthus; characters).
— Tottenham, 1962: 181 (Philonthus; characters).
▴ Distribution: Ethiopia.
subaeneipennis Bernhauer, 1916c: 32 (Philonthus; Type locality: Columbien: Muzo, Rio Cantinere, 500 m).
subaereipennis Bernhauer, 1939, see: Bisnius.
subcingulatus W. J. MacLeay, 1873: 141 (Philonthus; Type locality: Gayndah).
— Fauvel, 1877: 270 (Quedius; characters; Australia).
— Blackburn, 1896: 262 (Philonthus; Australia).
— Bernhauer and Schubert, 1916: 435 (Philonthus with the following statement: “Under Philonthus should be inserted the following, which Fauvel erroneously placed in Quedius:”).
— Bernhauer, 1920: 16 (Philonthus; Australia).
— Lea, 1925: 221 (Philonthus; from nest of Pheidole sp.; Australia).
— Scheerpeltz, 1933: 1362 (Philonthus; catalog).
▴ Distribution: Australia, New Caledonia.
sanguinicollis Fauvel, 1877: 265 (Philonthus; Type locality: Australie, Victoria; Rivière Paroo).
— Fauvel, 1878g: 548 (Philonthus; Australia).
— Fauvel, 1889b: 266 (Philonthus; notes; New Caledonia).
— Fauvel, 1903d: 271 (Philonthus; New Caledonia; Australia).
— Bernhauer and Schubert, 1916: 435 (Philonthus; synonym of subcingulatus).
— Bernhauer, 1916a: 5 (Philonthus; Australia).
— Lea, 1925: 220 (Philonthus; synonym of subcingulatus).
— Scheerpeltz, 1933: 1362 (Philonthus; synonym of subcingulatus).
subdecorus Cameron, 1950d: 18 (Philonthus; Type locality: Negri Sembilan: Bukit Tangga).
subdepressus Bernhauer, 1939, see: Gabrius.
subfoetidus Coiffait, 1983d: 166 (Philonthus; Type locality: Nepal: Ganesh Himal, entre Salmi et Rupchet, 1500 à 3600 m).
subfuscus Gyllenhal, 1810, see: umbratilis Gravenhorst, 1802.
subirideus Kraatz, 1859, see: delicatulus Boheman, 1858.
subjectus Cameron, 1932: 108 (Philonthus; Type locality: Sikkim: Tonglo).
sublaevipennis Bernhauer, 1903a: 28 (Philonthus; Type locality: Sumatra).
— Bernhauer and Schubert, 1914: 356 (Philonthus; catalog).
— Cameron, 1933d: 387 (Philonthus; Sumatra).
— Cameron, 1937: 5 (Philonthus; Java).
▴ Distribution: Indonesia.
sublaevis Bernhauer, 1915a: 24 (Philonthus; Type locality: Philippinen: Luzon: Mt. Makiling).
sublucanus Herman, 2001: 48 (Philonthus; replacement name for sericans Sharp).
sericans Sharp, 1874: 45 (Philonthus; [preoccupied]; Type locality: Nagasaki).
— Bernhauer and Schubert, 1914: 355 (Philonthus; catalog).
— Scheerpeltz, 1933: 1360 (Philonthus; catalog).
— Nakane, 1963a: 91 (Philonthus; characters; habitus photograph; Japan).
— Nakane and Sawada, 1960: A123 (Philonthus; subgenus Bisnius?; Yakushima Island).
— Watanabe and Shibata, 1972: 64 (Philonthus; Yaku-shima Island).
— Yuh, Paik, Kwon, and Lee, 1985: 254 (Philonthus; Korea).
— J. Li, 1993: 35 (Philonthus; China).
— Kanehisa, Tsumuki, and Kawazu, 1994 (Philonthus; chemical secretions).
— Herman, 2001: 48 (Philonthus; synonym of sublucanus).
submetallicus Cameron, 1933, see: Gabrius.
submicans Last, 1961: 308 (Philonthus; Type locality: Thailand: Chieng Mai).
subnigrituloides Scheerpeltz, 1933, see: Gabrius.
subnigritulus Reitter, 1909 [Note: Nomen nudum], see: Gabrius.
subnitens Poppius, 1909: 26 (Philonthus; Type locality: Lena-Ufer auf Inseln bei Önkyr-yrjä).
— Bernhauer and Schubert, 1914: 356 (Philonthus; catalog).
— Scheerpeltz, 1933: 1363 (Philonthus; catalog).
— Coiffait, 1967a: 429 (Philonthus; characters; Siberia).
— Coiffait, 1974: 297 (Philonthus; characters; Siberia).
▴ Distribution: Russia.
subnivalis Gistel, 1857: 65 (Philonthus; [Note: Nomen dubium]; Type locality: Alpes Helvetiae).
subopacus Hochhuth, 1851, see: Erichsonius.
subrufus Last, 1968: 367 (Philonthus; Type locality: New Guinea: Sarawaged Range, 20–30 miles north of Lae, 3000 ft.).
subrugosus Mulsant and Rey, 1876, see: Bisnius.
subtilepunctatus Scheerpeltz, 1965: 216 (Philonthus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).
subtilicornis Bernhauer, 1932b: 148 (Pseudophilonthus; Type locality: Haut-Uele: Moto Yebo).
— Scheerpeltz, 1933: 1363 (Philonthus; catalog).
— Tottenham, 1962: 232 (Philonthus; characters; Congo; Uganda).
▴ Distribution: Congo, Uganda.
subvarians Sawada, 1965: 13 (Philonthus; subgenus Philonthus; Type locality: Japan: Hokkaido: Sôunkyô).
subvirescens Thomson, 1884: 1032 (Philonthus; Type locality: Lappland).
— Horn, 1884: 187 (Philonthus; [Note: Misidentified as atratus according to Smetana, 1995e: 148]; characters; Hudson Bay).
— Ganglbauer, 1895: 446 (Philonthus; synonym of atratus).
— Poppius, 1909: 23 (Philonthus; Lena Valley, Siberia).
— Bernhauer and Schubert, 1914: 329 (Philonthus; synonym of atratus).
— Scheerpeltz, 1933: 1363 (Philonthus; valid species; catalog).
— Suomalainen, 1941 (Philonthus; subspecies of atratus; characters; Finland).
— Hatch, 1957: 197 (Philonthus; [Note: Misidentified as atratus according to Smetana, 1995e: 148]; characters; British Columbia; Washington; Oregon).
— Smetana, 1963c: 294 (Philonthus; subgenus Philonthus; Mongolia).
— Palm, 1963b: 68 (Philonthus; characters; Sweden; Norway; Finland).
— Smetana, 1965a: 4 (Philonthus; characters; notes; cited as valid species; British Columbia; Alaska).
— Coiffait, 1967a: 392 (Philonthus; characters; Scandinavia).
— Coiffait, 1974: 250 (Philonthus; characters; Scandinavia).
— Smetana 1975c: 169 (Philonthus; Mongolia).
— Shilov, 1976: 148 (Philonthus; Mongolia).
— Andersen, 1982: 66 (Philonthus; natural history; Norway).
— Lohse and Lucht, 1989: 165 (Philonthus; characters).
— Smetana, 1995e: 148 (Philonthus; characters; notes; Alberta; British Columbia; Northwest Territories; Yukon Territory; Alaska; California; Colorado; Idaho; Montana; New Mexico; Oregon; South Dakota; Utah; Washington; Wyoming).
— Ryabukhin, 1999: 64 (Philonthus; catalog; habitat; distribution; northeastern Russia).
▴ Distribution: Europe, Russia, Mongolia, Canada; Alaska.
perforatus Fauvel, 1889a: 114 (Philonthus; named to replace atratus as misidentified by Hamilton, 1884: 111, Transactions of the American Entomological Society; [Note: Fauvel cited 1884 as the date for the Hamilton article but it should probably be 1889, volume 16]; Type locality: California: Cisco).
— Bernhauer and Schubert, 1914: 350 (Philonthus; catalog).
— Smetana, 1995e: 148 (Philonthus; synonym of subvirescens).
septentrionis Fall, 1926: 149 (Philonthus; Type locality: Dawson, Yukon Territory).
— Scheerpeltz, 1933: 1360 (Philonthus; catalog).
— Smetana, 1995e: 148 (Philonthus; synonym of subvirescens).
succicola Thomson, 1860: 157 (Philonthus; Type locality: Skåne).
— Erichson, 1839a: 448 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 332 cited carbonarius Erichson as a synonym of chalceus Stephens; Erichson did not describe the species as new, he attributed it to Gyllenhal who attributed it to Gravenhorst; P. chalceus was misidentified by most authors according to Tottenham, 1937a]; characters).
— Heer, 1839: 259 (Philonthus; [Note: Heer cited Erichson as the author but Erichson did not describe the species; he attributed it to Gyllenhal who cited Gravenhorst as author; P. chalceus was misidentified by most authors according to Tottenham, 1937a]; characters; Switzerland).
— Erichson, 1840: 437 (Philonthus; cited as carbonarius; [Note: Bernhauer and Schubert, 1914: 332 cited carbonarius Erichson, 1839a: 448 as a synonym of chalceus but it was a misidentification and so presumably was Erichson's 1840 citation of carbonarius; P. chalceus was misidentified by most authors according to Tottenham, 1937a]; characters; Europe).
— Kraatz, 1857d: 577 (Philonthus; [Note: Cited as carbonarius Erichson but Erichson did not describe the species and his use of the name is treated as a misidentification of chalceus; P. chalceus was misidentified by most authors according to Tottenham, 1937a]; characters; Germany).
— Redtenbacher, 1857: 190 (Philonthus; [Note: Cited as carbonarius Erichson but Erichson did not describe the species and his use is treated as a misidentification of chalceus; P. chalceus was misidentified by most authors according to Tottenham, 1937a]; characters; Austria).
— G. Waterhouse, 1858: 24 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; synonym of carbonarius).
— Rye, 1864a: 54 (Philonthus; characters; notes; Britain).
— Fauvel, 1869: 489 (Philonthus; synonym of chalceus).
— Fauvel, 1869: 488 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; valid species).
— Fauvel, 1874: 215 [= 1874a: 441] (Philonthus; synonym of proximus).
— Redtenbacher, 1874: 216 (Philonthus; characters).
— Seidlitz, 1875: 273 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 374 [= 1877: 229] (Philonthus; [Note: Cited as carbonarius Erichson but Erichson did not describe the species and his use is treated as a misidentification of chalceus; P. chalceus was misidentified by most authors according to Tottenham, 1937a]; characters; France).
— Ganglbauer, 1895: 444 (Philonthus; [Note: According to Tottenham, 1939d: 232 Ganglbauer's use of chalceus was a misidentification of succicola]; characters; Europe; Siberia; North India).
— Ganglbauer, 1895: 444 (Philonthus; synonym of chalceus).
— Poppius, 1905: 9 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; Shetland Islands).
— Reitter, 1909: 125 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; characters; Germany).
— Bernhauer and Schubert, 1914: 332 (Philonthus; synonym of chalceus).
— Bernhauer and Schubert, 1914: 331 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; catalog).
— Johansen, 1914: 394 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; characters; Denmark).
— Cameron, 1932: 80 (Philonthus; synonym of chalceus).
— Scheerpeltz, 1933: 1336 (Philonthus; synonym of chalceus).
— Scheerpeltz, 1933: 1336 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; catalog).
— Wüsthoff, 1934: 60 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; illustration of aedeagus).
— Aleksandrov, 1934: 153 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; China).
— West, 1937: 39 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; Faroe Islands).
— Tottenham, 1939d: 232 (Philonthus; valid species).
— Hinton, 1945: 76 (Philonthus; characters).
— Normand, 1947a: 8 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; Tunisia).
— V. Hansen, 1952: 51 (Philonthus; subgenus Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; characters; Denmark).
— Smetana, 1958: 164 (Philonthus; synonym of chalceus).
— Smetana, 1958: 164 (Philonthus; subgenus Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; characters; ecology; Czechoslovakia).
— Smetana, 1959: 146 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; characters).
— Scheerpeltz, 1961d: 128 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; Italy).
— Lohse, 1964: 177 (Philonthus; subgenus Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; characters; central Europe).
— Smetana, 1964d: 74 (Philonthus; subgenus Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; Czechoslovakia).
— Horion, 1965: 129 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; all Europe).
— Pototskaia, 1966: 146, 158 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; larval characters).
— Pototskaia, 1967: 63 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; larval characters).
— Coiffait, 1967a: 392 (Philonthus; characters).
— Scheerpeltz, 1968a: 67 (Philonthus; synonym of chalceus).
— Raianu, 1968a: 462 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; Romania).
— Szujecki, 1968a: 727 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; Poland).
— Szujecki, 1969a: 255 (Philonthus; notes; Poland).
— Jarrige, 1971: 494 (Philonthus; Iran).
— Bacchus and Hammond, 1972: 63 (Philonthus; lion dung; Britain).
— Coiffait, 1974a: 198 (Philonthus; Siberia).
— Coiffait, 1974: 222 (Philonthus; characters; Europe; Scandinavia; Britain; Mongolia).
— Coiffait, 1974: 223 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; synonym of succicola).
— Shilov, 1975a: 375 [= 1975b: 98] (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; Komi region of Russia).
— Legner and Moore, 1977: 176 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; found under bark).
— Pace and Zanetti, 1977: 289 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Muona, 1979: 14 (Philonthus; Finland; Russia; Sweden; Norway; Denmark; Baltic states).
— Tóth, 1984: 48 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; characters; Hungary).
— Segers, 1986: 45 (Philonthus; Belgium).
— Muona and Viramo, 1986: 13 (Philonthus; Finland).
— Drugmand, 1987b: 14, 47 (Philonthus; habitat; distribution in Belgium).
— Lucht, 1987: 105 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Lohse and Lucht, 1989: 163 (Philonthus; notes).
— Lohse and Lucht, 1989: 163 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; synonym of succicola).
— Pilon and Zanetti, 1991: 63 (Philonthus; Italy).
— Silfverberg, 1992: 16 (Philonthus; Finland; Russia; Sweden; Norway; Denmark; Latvia; Lithuania).
— J. Li, 1993: 37 (Philonthus; China; [Note: May be misidentification of P. decoloratus according to Schillhammer, 2000: 128]).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Philonthus; Italy).
— Muona and Viramo, 1995: 9 (Philonthus; collected from dung of Ursus arctos; Finland).
— Smetana, 1995e: 19, 72, 84 (Philonthus; comparative characters; reference to data on embryo-gensis).
— Hansen, 1996: 105 (Philonthus; distribution in Denmark).
— Ádám, 1996: 245 (Bisnius; Hungary).
— Ádám, 1996a: 21 (Bisnius; Hungary).
— Ádám, 1996b: 57 (Bisnius; Hungary).
— Meer, 1999: 9 (Philonthus; France).
— Staniec, 1999 (Philonthus; characters of pupa; Poland).
— Kashcheev, 1999a: 57 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola]; Kazakhstan).
— Jørum, 2000: 39 (Philonthus; Denmark).
— Schillhammer, 2000: 127 (Philonthus; characters; Russia; records from China and Mongolia are probably misidentifications of Philonthus decoloratus).
— Herman, 2001: 28 (Philonthus; nomen protectum).
▴ Distribution: Tunisia, Europe, Russia, Iran, Kazakhstan.
nigritus Runde, 1835: 7 (Staphylinus; Type locality: Halae).
— Erichson, 1840: 437 (Philonthus; synonym of carbonarius).
— Kraatz, 1857d: 577 (Philonthus; synonym of carbonarius Erichson).
— Ganglbauer, 1895: 444 (Philonthus; synonym of chalceus).
— Bernhauer and Schubert, 1914: 332 (Philonthus; synonym of chalceus).
— Cameron, 1932: 80 (Philonthus; synonym of chalceus).
— Tottenham, 1939d: 232 (Philonthus; synonym of succicola).
— Smetana, 1958: 164 (Philonthus; synonym of chalceus).
— Coiffait, 1974: 223 (Philonthus; synonym of succicola).
— Schillhammer, 2000: 127 (Philonthus; synonym of succicola).
— Herman, 2001: 28 (Philonthus; nomen oblitum).
wohlgrothi R. Dvořák, 1952: 79 (Philonthus; aberration of chalceus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Brunstatt, Haut-rhin, Carrière).
— Smetana, 1955b: 224 (Philonthus; aberration of chalceus; notes).
— Smetana, 1958: 164 (Philonthus; synonym of chalceus).
— Coiffait, 1974: 223 (Philonthus; synonym of succicola).
— Schillhammer, 2000: 127 (Philonthus; listed as unavailable name; synonym of succicola).
succinctus Guérin-Méneville, 1844: 13 (Philonthus; Type locality: Nouvelle Granade).
— Solsky, 1875: 15 (Philonthus; Peru).
— Bernhauer and Schubert, 1914: 356 (Philonthus; catalog).
▴ Distribution: Colombia, Peru.
sudanensis Cameron, 1952, see: marcescens Tottenham, 1949.
sulcatus Cameron, 1918: 87 (Philonthus; Type locality: Singapore: Keppel Harbour).
— Cameron, 1921a: 377, 405 (Philonthus; characters; Singapore).
— Scheerpeltz, 1933: 1363 (Philonthus; catalog).
— Yadav, Chand, and Yadav, 1995: 6 (Philonthus; morphology of mitotic and meiotic chromosomes).
▴ Distribution: Singapore.
sulciceps Bernhauer, 1931, see: productus Kraatz, 1859.
sulcicollis LeConte, 1863, see: Cafius.
sulcifrons Sharp, 1889, see: Gabronthus.
sumatrensis Cameron, 1933d: 388 (Philonthus; Type locality: Sumatra: Fort de Kock).
sumbaensis Scheerpeltz, 1957c: 234, 324 (Philonthus; Type locality: Sumba: Mau Marru).
superbus Arrow, 1915, see: Leucitus.
supernus Herman, 2001: 48 (Philonthus; replacement name for excelsus Bernhauer).
excelsus Bernhauer, 1941c: 287 [= 1951: 277] (Philonthus; [preoccupied]; Type locality: Süd-Peru: Ocaña, 3100 m; Oberh. Palco, 3240 m; Oberh. Totora, 4000 m).
suspectus Erichson, 1840: 465 (Philonthus; Type locality: Caracas Columbiae).
— Fauvel, 1891: 115 (Philonthus; Venezuela).
— Bernhauer and Schubert, 1914: 356 (Philonthus; catalog; Uruguay).
▴ Distribution: Mexico, Guatemala, Venezuela, Brazil, Uruguay.
quadraticollis Sharp, 1885: 409 (Philonthus; Type locality: Mexico, Cordova, Jalapa; Guatemala, San Geronimo; South America, Rio de Janeiro).
suspiciosus Cameron, 1932: 129 (Philonthus; Type locality: Burma: Ruby Mines; Tenasserim).
suspiritus Last, 1987: 28 (Philonthus; Type locality: Irian Jaya: New Guinea: S. Highlands, Dimifa, SE of Mt. Giluwe, 2200 m).
sutteri Scheerpeltz, 1957c: 234, 321 (Philonthus; Type locality: Sumba: Baing).
suturalis Marsham, 1802, see: discoideus Gravenhorst, 1802.
suturalis Stephens, 1832, see: discoideus Gravenhorst, 1802.
suturalis Nordmann, 1837, see: transbaicalia Hochhuth, 1851.
suturamonus Tottenham, 1939 [Note: Nomen nudum], see: sanguinolentus Gravenhorst, 1802.
suturorubineous Pajni and Kohli, 1977: 517 (Philonthus; Type locality: India: Chandigarh, Labour Colony).
svanetiensis Coiffait, 1974, see: frigidus Märkel and Kiesenwetter, 1848.
swynnertoni Tottenham, 1953, see: altivagans Fauvel, 1907.
sylvicola Cameron, 1949a: 320 (Philonthus; Type locality: Côte d'Ivoire: Mt. Tonkoui, 900–1200 m).
synonymicus Roubal, 1915, see: Bisnius.
syriacus Gridelli, 1924, see: Rabigus.
syrovatkai Roubal, 1913 [Note: Nomen nudum], see: Bisnius.
tabius Tottenham, 1955: 176 (Philonthus; Type locality: Cape Colony: Queenstown).
tachiniformis Say, 1830: 37 [= 1834: 450] (Staphylinus; Type locality: Mexico).
— Erichson, 1840: 930 (Philonthus; characters; Mexico).
— Sharp, 1885: 425 (Philonthus; notes; Mexico).
— Bernhauer and Schubert, 1914: 357 (Philonthus; catalog).
▴ Distribution: Mexico.
tachyoryctidis Jeannel and Paulian, 1945: 82 (Philonthus; Type locality: camp II de l'Elgon, alt. 2470 m. Also larval description included).
— Tottenham, 1955: 166 (Philonthus; subgenus Spatulonthus; characters).
— Tottenham, 1962: 140 (Philonthus; subgenus Spatulonthus; Tanganyika).
▴ Distribution: Kenya, Tanzania.
tahitiensis Coiffait, 1977a: 172 (Philonthus; Type locality: Tahiti, bords du Lac Vaihiria).
taiwanensis Shibata, 1993b: 54 (Philonthus; replacement name for formosae Cameron).
formosae Cameron, 1949: 176 (Philonthus; [preoccupied]; Type locality: Formosa: Toroon: near Mt. Ari).
— Shibata, 1993b: 54 (Philonthus; synonym of taiwanensis).
— Schillhammer, 1998: 109 (Philonthus; synonym of taiwanensis).
takaiensis Last, 1968: 351 (Philonthus; Type locality: New Guinea: Takai).
tamulus Cameron, 1932: 91 (Philonthus; Type locality: Nilgiri Hills).
tamulus Tottenham, 1949, see: proselytus Herman, 2001.
tanaicus Hochhuth, 1851, see: carbonarius Gravenhorst, 1802.
tandalensis Bernhauer, 1939a: 84 (Philonthus; Type locality: Ost-Afrika: Tandala).
tangamanus Tottenham, 1962: 171 (Philonthus; Type locality: Rhodesia: Mashonaland, Salisbury).
tantulus Luze, 1904, see: Bisnius.
taprobanus Cameron, 1918, see: notabilis Kraatz, 1859.
tardus Kraatz, 1859: 84 (Philonthus; Type locality: Ceylan).
— Bernhauer, 1902b: 26 (Philonthus; Ceylon).
— Bernhauer and Schubert, 1914: 357 (Philonthus; catalog).
— Bernhauer, 1922c: 232 (Philonthus; Formosa).
— Cameron, 1930b: 162 (Philonthus; British North Borneo).
— Cameron, 1932: 103 (Philonthus; characters; Ceylon; India).
— Scheerpeltz, 1933: 1363 (Philonthus; catalog).
— Cameron, 1933d: 386 (Philonthus; Sumatra).
— Cameron, 1937: 5 (Philonthus; Java).
— Bernhauer and Chapman, 1939: 71 (Philonthus; Kenya).
— Scheerpeltz, 1954: 113 (Philonthus, cited as Phylonthus; Ceylon).
— Nakane and Sawada, 1960: A123 (Philonthus; Yakushima Island).
— Watanabe, 1961: 354 (Philonthus; Izu Islands).
— Scheerpeltz, 1961f: 135 (Philonthus; Ceylon).
— Watanabe and Shibata, 1972: 64 (Philonthus; Yaku-shima Island).
▴ Distribution: Kenya, Sri Lanka, India, Borneo, Indonesia, Taiwan, Japan, China.
curtipennis Bernhauer, 1902b: 26 (Philonthus; Type locality: Bentota).
— Bernhauer and Schubert, 1914: 334 (Philonthus; catalog).
— Cameron, 1932: 103 (Philonthus; synonym of tardus).
— Scheerpeltz, 1933: 1363 (Philonthus; synonym of tardus).
inornatus Cameron, 1920a: 215 (Philonthus; variety of geminus; Type locality: Nilgiri Hills).
— Cameron, 1932: 103 (Philonthus; synonym of tardus).
— Scheerpeltz, 1933: 1363 (Philonthus; variety of tardus).
tarsalis Smetana, 1963d: 412 (Philonthus; subgenus Onychophilonthus; Type locality: Ost-Asien: UdSSR, Primorskij kraj, Küsten-Provinz, Naturschutzgebiet, “Kedrovaja pad”).
— Coiffait, 1967a: 439 (Philonthus; subgenus Onychophilonthus; characters; eastern Asia).
— Coiffait, 1974: 306 (Philonthus; subgenus Onychophilonthus; characters; East Asia).
— Hayashi, 1994: 124 (Philonthus; subgenus Onychophilonthus; characters; Japan).
▴ Distribution: Russia, Japan.
taterae Scheerpeltz, 1951b: 27 (Philonthus; Type locality: South Africa: Vredefort, Orange Free State; Zuurbekom, Transvaal, Roodepoort district).
taunggyiensis Scheerpeltz, 1965: 218 (Philonthus; Type locality: S. Shan-States, Burma, 1500 m, Taunggyi).
tauricus Nordmann, 1837, see: Quedius.
tectipennis Coiffait, 1967 [Note: Nomen nudum], see: velatipennis Solsky, 1870.
tectus Last, 1968, see: Hesperus.
temperei Coiffait, 1987, see: politus Linné, 1758.
temporalis Mulsant and Rey, 1853: 61 [= 1853a: 74] (Philonthus; Type locality: Suisse).
— Fairmaire and Laboulbène, 1856: 516 (Philonthus; characters; France).
— Kraatz, 1855: 23 (Xantholinus; notes).
— Kraatz, 1857d: 573, note (Philonthus; characters).
— Rye, 1864a: 55 (Philonthus; characters; notes; Britain).
— Fauvel, 1874: 218 [= 1874a: 444] (Philonthus; characters; France; Germany; Switzerland; Tirol; Italy).
— Redtenbacher, 1874: 216 (Philonthus; characters).
— Mulsant and Rey, 1876: 379 [= 1877: 235] (Philonthus; characters; France).
— Ganglbauer, 1895: 445 (Philonthus; characters; Alps; Jura; Rheinland).
— Reitter, 1909: 126 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 357 (Philonthus; catalog).
— Portevin, 1929: 359 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1363 (Philonthus; catalog).
— Wüsthoff, 1934: 60 (Philonthus; illustration of aedeagus).
— Wüsthoff, 1937a: 347 (Philonthus; illustration of aedeagus).
— Smetana, 1958: 168 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 148 (Philonthus; characters).
— Lohse, 1964: 178 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 138 (Philonthus; Holland; Switzerland; France; Italy; Bosnia; Romania; Poland; Austria; Germany).
— Coiffait, 1967a: 395 (Philonthus; characters; Europe).
— Coiffait, 1974: 228 (Philonthus; characters; Europe).
— Tóth, 1984: 51 (Philonthus; characters; Hungary).
— Drugmand, 1987a: 311 (Philonthus; Belgium).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Austria; Switzerland; France; Benelux).
— Pilon and Zanetti, 1991: 63 (Philonthus; Italy).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czechoslovakia ?).
— Ciceroni and Zanetti, 1995: 29 (Philonthus; Italy).
— Schillhammer, 2000: 137 (Philonthus; characters; West Palaearctic).
▴ Distribution: Europe.
punctiventris Kraatz, 1857d: 578 (Philonthus; [preoccupied]; Type locality: Baiern).
— Redtenbacher, 1857: 985 (Philonthus; characters; Austria).
— Fauvel, 1869: 489 (Philonthus; synonym of temporalis).
— Rye, 1870a: 82 (Philonthus; notes).
— Fauvel, 1874: 218 [= 1874a: 444] (Philonthus; synonym of temporalis).
— Redtenbacher, 1874: 216 (Philonthus; characters).
— Ganglbauer, 1895: 445 (Philonthus; synonym of temporalis).
— Bernhauer and Schubert, 1914: 357 (Philonthus; catalog).
— Scheerpeltz, 1933: 1363 (Philonthus; synonym of temporalis).
— Smetana, 1958: 168 (Philonthus; synonym of temporalis).
— Coiffait, 1974: 228 (Philonthus; synonym of temporalis).
— Schillhammer, 2000: 137 (Philonthus; synonym of temporalis).
temporalis Coiffait, 1977, see: aeolus Herman, 2001.
temporalisoides Drugmand, 1988, see: addendus Sharp, 1867.
temporasinus Schubert, 1911, see: morio Boheman, 1848.
temporatus Last, 1968, see: Hesperus.
tenebrosus Boheman, 1858: 28 (Philonthus; Type locality: Buenos Ayres).
— Lynch, 1884: 149 (Philonthus; characters; Argentina).
— Bernhauer and Schubert, 1914: 357 (Philonthus; catalog).
— Scheerpeltz, 1933: 1363 (Philonthus; catalog).
▴ Distribution: Argentina.
tenellicornis Scheerpeltz, 1978: 220 (Philonthus; subgenus Glaphyronthus; Type locality: Turisca, Timor).
tenellus Wollaston, 1864, see: Neobisnius.
tenuemicans Scheerpeltz, 1965: 207 (Philonthus; Type locality: N.E. Burma, Kambaiti, 2000 m).
tenuepunctatus Scheerpeltz, 1965: 220 (Philonthus; Type locality: N.E. Burma, Kambaiti, 2000 m).
tenuicornis Mulsant and Rey, 1853: 58 [= 1853a: 71] (Philonthus; Type locality: Lyonnais, Bourgogne).
— Gyllenhal, 1810: 319 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 331 cited carbonarius Gyllenhal as a valid species; Gyllenhal did not describe the species as new, he attributed it to Gravenhorst]; characters; Sweden).
— Fairmaire and Laboulbène, 1856: 516 (Philonthus; characters; France).
— Kraatz, 1855: 23 (Xantholinus; notes; France).
— Kraatz, 1857d: 574, note (Philonthus; characters).
— Rye, 1863: 82 (Philontus; [Note: Misidentification: Bernhauer and Schubert, 1914: 339 cited punctiventris Janson as a synonym of carbonarius Gyllenhal; Janson did not describe the species that was cited in Rye's article. Janson reported the species from Britain based on one specimen; Rye reported Janson's finding as punctiventris Kraatz]; notes; characters; Britain).
— Fauvel, 1874: 217 [= 1874a: 443] (Philonthus; synonym of carbonarius).
— Redtenbacher, 1874: 216 (Philonthus; characters).
— Seidlitz, 1875: 273 (Philonthus; cited as carbonarius Gyllenhal; characters; Baltic region).
— Mulsant and Rey, 1876: 381 [= 1877: 237] (Philonthus; characters; larval characters; France).
— J. Sahlberg, 1876: 14 (Philonthus; cited as carbonarius Gyllenhal; Finland).
— Fowler, 1888: 263 (Philonthus; cited as carbonarius Gyllenhal; characters; Britain).
— Ganglbauer, 1895: 445 (Philonthus; cited as carbonarius Gyllenhal and with tenuicornis as synonym; characters; Europe).
— Reitter, 1909: 126 (Philonthus; cited as carbonarius Gyllenhal; characters; Germany).
— Bernhauer and Schubert, 1914: 331 (Philonthus; synonym of carbonarius Gyllenhal; [Note: The carbonarius of Gyllenhal is an unavailable, misidentified species]).
— Johansen, 1914: 395 (Philonthus; [Note: Gyllenhal cited as author]; characters; Denmark).
— Cameron, 1932: 81 (Philonthus; synonym of carbonarius).
— Scheerpeltz, 1933: 1335 (Philonthus; synonym of carbonarius).
— Tottenham, 1939d: 231 (Philonthus; cited with carbonarius Gyllenhal as a junior synonym; [Note: Gyllenhal did not describe carbonarius]).
— Smetana, 1958: 166 (Philonthus; subgenus Philonthus; cited with carbonarius Gyllenhal as the senior name and tenuicornis as a junior synonym; [Note: carbonarius was described by Gravenhorst; the carbonarius Gyllenhal referred to by Gyllenhal has been associated with tenuicornis]; characters; ecology; Czechoslovakia).
— Coiffait, 1967a: 395 (Philonthus; characters; Europe).
— Jarrige, 1971: 495 (Philonthus; Iran).
— Coiffait, 1974: 228 (Philonthus; characters; Europe; Scandinavia; Britain).
— Pace and Zanetti, 1977: 289 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Raichev, 1979: 82 (Philonthus; Bulgaria).
— Hinton, 1981: 661 (Philonthus; characters of egg).
— Frank, 1982: 43 (Philonthus; list of parasites).
— Lohse and Lucht, 1989: 165 (Philonthus; note).
— Pilon and Zanetti, 1991: 62 (Philonthus; Italy).
— Welch, 1993: 228 (Philonthus; ovariole number and ovary structure).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Philonthus; Italy).
— Smetana, 1995e: 123 (Philonthus; characters; introduced species; British Columbia).
— Luff, 1996: 30 (Philonthus; collecting notes; Isle of Man).
— Schillhammer, 2000: 130 (Philonthus; characters; West Palaearctic region as far east as Caucasus and Iran; introduced in Nearctic).
▴ Distribution: Europe, Iran, Canada.
tenuimarginatus Scheerpeltz, 1960c: 112 (Philonthus; Type locality: Bolivien: Cochabamba, 2600 m).
tenuissimus Tottenham, 1949, see: Gabronthus.
tergomaculatus Lecoq, 1997: 282 (Philonthus; Type locality: Madagascar Est, Ambodivoangy).
terminalis LeConte, 1863, see: Neobisnius.
terminipennis Tottenham, 1939d: 233 (Philonthus; replacement name for apicipennis).
apicipennis Cameron, 1932: 117 (Philonthus; [preoccupied]; Type locality: Dehra Dun. Siwaliks: Mussoorie. Haldwani).
— Scheerpeltz, 1933: 1332 (Philonthus; catalog).
— Tottenham, 1939d: 233 (Philonthus; synonym of terminipennis).
— Coiffait, 1984e: 374 (Philonthus; cited as valid species; Nepal).
testaceipennis Erichson, 1840: 435 (Philonthus; Type locality: Mexico).
— Solsky, 1868a: 127 (Philonthus; Mexico).
— Sharp, 1885: 400 (Philonthus; characters; Mexico).
— Bernhauer and Schubert, 1914: 357 (Philonthus; catalog).
▴ Distribution: Mexico.
testaceipennis Roubal, 1948 [Note: Nomen nudum], see: Bisnius.
testaceocinctus Bernhauer, 1915d: 140 (Philonthus; Type locality: Deutsch-Ostafrika: Unjanjembe).
— Scheerpeltz, 1933: 1364 (Philonthus; catalog).
— Tottenham, 1962: 139, 215, 217, 218 (Philonthus; characters; Uganda).
▴ Distribution: Tanzania, Uganda.
testudo Smetana, 1995e: 263 (Philonthus; Type locality: Georgia: Tift Co.: Tifton. Also cited from Florida, Mississippi).
tetragonocephalus Notman, 1924, see: rectangulus Sharp, 1874.
tetramerus Fauvel, 1878f: 265 (Philonthus; Type locality: Nouvelle-Guinée, Hatam).
tetricus Bernhauer, 1933a: 42 (Philonthus; Type locality: Tatsienlu-Kiulung).
thailandicus Cameron, 1946a: 684 (Philonthus; Type locality: Siam: Renong).
thaseus Coiffait, 1976, see: juvenilis Peyron, 1858.
thermarum Aubé, 1850, see: Gabronthus.
thermicola Cameron, 1947, see: Gabrius.
thessalicus Roubal, 1916 [Note: Nomen nudum], see: carbonarius Gravenhorst, 1802.
thessalonicus Coiffait [Note: Nomen nudum], 1963, see: Bisnius.
theveneti Horn, 1884: 194 (Philonthus; Type locality: California; [Note: See lectotype designation by Smetana, 1995e]. Also cited from Nevada).
— Bernhauer and Schubert, 1914: 358 (Philonthus; catalog).
— Hatch, 1957: 190 (Philonthus; characters; British Columbia; Washington; Oregon; Idaho).
— J. B. Campbell and Hermanussen, 1974: 356 (Philonthus; natural history and predatory habits).
— Smetana, 1995e: 274 (Philonthus; lectotype designation: Cal [California]; characters; California; Nevada; Oregon).
▴ Distribution: Canada, USA.
thoracicus Gravenhorst, 1802: 170 (Staphylinus; [preoccupied]; Type locality: America septentr.; [Note: See lectotype designation by Smetana, 1995e]).
— Gravenhorst, 1806: 71 (Staphylinus; characters).
— Erichson, 1840: 481 (Philonthus; characters; America septentrionalis).
— Horn, 1884: 199 (Philonthus; characters; Gulf States; Michigan; Kansas).
— Blatchley, 1910: 384 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 358 (Philonthus; catalog).
— Smetana, 1995e: 380 (Philonthus; lectotype designation; characters; notes; Alberta; Manitoba; New Brunswick; Ontario; Quebec; Saskatchewan; Arkansas; Colorado; Illinois; Indiana; Iowa; Kansas; Kentucky; Maine; Maryland; Massachusetts; Minnesota; Missouri; Nebraska; New Hampshire; New Jersey; New York; North Carolina; North Dakota; Ohio; Oklahoma; Pennsylvania; South Dakota; Texas; Utah; Virginia; Wisconsin).
— Downie and Arnett, 1996: 379 (Philonthus; characters; USA).
— Herman, 2001: 47 (Philonthus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
▴ Distribution: Canada, USA.
thoracicus Melsheimer, 1844, see: Bisnius.
thoracicus Coiffait, 1979, see: arabiensis Herman, 2001.
tibialis Eppelsheim, 1876, see: juvenilis Peyron, 1858.
tienmuschanensis Bernhauer, 1939k: 591 (Philonthus; Type locality: N. W. China: Tienmuschan).
— Coiffait, 1967a: 411 (Philonthus; cited as tiemuschanensis; characters; China).
— Coiffait, 1974: 290 (Philonthus; cited as tiemuschanensis; characters; China).
▴ Distribution: China.
tilakholensis Coiffait, 1982: 65 (Philonthus; Type locality: Nepal: Ufer des Tila Khola, bei Jumla).
tillius Smetana, 1995e: 428 (Philonthus; Type locality: Manitoba: Winnipeg. Also cited from Alberta, Colorado, New Mexico, Wyoming).
timorensis Scheerpeltz, 1978: 207 (Philonthus; Type locality: La Queco, Timor).
tiro Sharp, 1874, see: Hesperus.
tirolensis Luze, 1903, see: Gabrius.
titschacki Bernhauer, 1941c: 286 [= 1951: 276] (Philonthus; Type locality: Süd-Peru: Hac. Huayuri).
toenianus Mulsant and Rey, 1876, see: micans Gravenhorst, 1802.
tomicus Tottenham, 1962: 225 (Philonthus; Type locality: Cameroon: Bamenda).
tottenhami Last, 1953: 114 (Philonthus; Type locality: Natal: Qudeni Forest).
toxopeanus Cameron, 1952: 248 (Philonthus; Type locality: New Guinea: Snow Mountains: Rattan Camp, 1150 m).
toxopei Cameron, 1937: 14 (Philonthus; Type locality: Java: Tjibodas, alt. 1400 m).
tractatoides Schillhammer, 2000: 148 (Philonthus; Type locality: Bhutan: Thimpu District Taba, 2800 m. Also cited from Nepal).
tractatus Eppelsheim, 1895c: 61 (Philonthus; Type locality: Simla im Himalaya).
— Bernhauer and Schubert, 1914: 358 (Philonthus; catalog).
— Cameron, 1928c: 563 (Philonthus; Sikkim; Tibet).
— Cameron, 1932: 82 (Philonthus; characters; Tibet; India).
— Scheerpeltz, 1933: 1364 (Philonthus; catalog).
— Coiffait, 1982: 30 (Philonthus; Nepal).
— Coiffait, 1984e: 373 (Philonthus; Nepal).
— Schillhammer, 1999a: 62 (Philonthus).
— Schillhammer, 2000: 146 (Philonthus; characters; China; Nepal).
▴ Distribution: Bhutan, India, Nepal, China.
proximatus Schubert, 1908: 616 (Philonthus; Type locality: Kulu, Himalaya).
— Bernhauer and Schubert, 1914: 351 (Philonthus; catalog).
— Cameron, 1928c: 563 (Philonthus; Sikkim; Tibet).
— Cameron, 1932: 82 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1356 (Philonthus; catalog).
— Coiffait, 1977b: 208 (Philonthus; Bhutan).
— Schillhammer, 1999a: 62 (Philonthus; synonym of tractatus).
— Schillhammer, 2000: 146 (Philonthus; synonym of tractatus).
cupreipennis Cameron, 1926b: 350 (Philonthus; Type locality: West Almora; Dhauli Ganga, 9520 feet).
— Cameron, 1932: 82 (Philonthus; synonym of tractatus).
— Scheerpeltz, 1933: 1364 (Philonthus; synonym of tractatus).
— Schillhammer, 2000: 146 (Philonthus; synonym of tractatus).
traili Sharp, 1876, see: Paederomimus.
transbaicalia Hochhuth, 1851: 10 (Philonthus; Type locality: Dahurien; [Note: See lectotype designation by Gusarov, 1992a]).
— Motschulsky, 1860b: 121 (Philonthus; cited as transbaicalicus; eastern Siberia).
— Hochhuth, 1862: 53 (Philonthus; characters; Russia).
— Solsky, 1871c: 348 (Philonthus; cited as transbaicalicus; characters; notes; Siberia).
— Fauvel, 1875a: XXIX [= 1875b: 231] (Philonthus; cited as transbaicalicus; synonym of suturalis).
— Bernhauer and Schubert, 1914: 357 (Philonthus; cited as transbaicalicus; synonym of suturalis).
— Smetana, 1967d: 202 (Philonthus; cited as transbaicalicus; synonym of suturalis).
— Coiffait, 1974: 302 (Philonthus; cited as transbaicalus; synonym of suturalis).
— Gusarov, 1992a: 785 [= 1993a: 74] (Philonthus; lectotype designation).
— Schillhammer, 2000: 157 (Philonthus; characters; Russia; Mongolia; China).
— Herman, 2001: 48 (Philonthus; valid name for species).
▴ Distribution: Russia, Mongolia.
suturalis Nordmann, 1837: 91 (Philonthus; [preoccupied]; Type locality: Sibiria; Rossia orientalis).
— Erichson, 1840: 459 (Philonthus; characters; Siberia; Russia orientalis).
— Poppius, 1909: 23 (Philonthus; Lena Valley, Siberia).
— Bernhauer and Schubert, 1914: 357 (Philonthus; catalog).
— Scheerpeltz, 1933: 1363 (Philonthus; catalog).
— Smetana, 1963c: 293 (Philonthus; subgenus Philonthus; Mongolia).
— Scheerpeltz, 1964: 28 (Philonthus; subgenus Philonthus; Mongolia).
— Smetana, 1965e: 171 (Philonthus; Mongolia).
— Coiffait, 1967a: 437 (Philonthus; subgenus Kenonthus; characters; central Asia; Lake Baikal).
— Smetana, 1967d: 202 (Philonthus; Mongolia).
— Coiffait, 1974: 302 (Philonthus; subgenus Kenonthus; characters; Siberia; Mongolia; Russia).
— Smetana 1975c: 169 (Philonthus; Mongolia).
— Shilov, 1976: 147 (Philonthus; Mongolia).
— Schillhammer, 2000: 157 (Philonthus; synonym of transbaicalia).
— Herman, 2001: 48 (Philonthus; synonym of transbaicalia).
trapeziceps Scheerpeltz, 1960c: 107 (Philonthus; Type locality: Bolivien: Yungas de Palmar, 1250 m).
trapeziceps Scheerpeltz, 1965, see: ottoi Herman, 2001.
trepidus Erichson, 1840: 489 (Philonthus; Type locality: Ins. St. John Portorico; [Note: See lectotype designation by Smetana, 1994]. Originally cited from Americae ins. St. Johannis; Puerto Rico).
— Gemminger and Harold, 1868: 589 (Philonthus; synonym of havaniensis).
— Fauvel, 1891: 115 (Philonthus; synonym of suspectus).
— Bernhauer and Schubert, 1914: 340 (Philonthus; synonym of havaniensis).
— Blackwelder, 1943: 418 (Philonthus; synonym of havaniensis).
— Smetana, 1994: 345 (Philonthus; characters; lectotype designation: Ins. St. John Portorico; listed as valid species; Puerto Rico; Virgin Islands; Antigua).
▴ Distribution: West Indies.
triangulum Horn, 1884: 202 (Philonthus; Type locality: California; [Note: See lectotype designation by Smetana, 1995e]. Also cited from Washington Territory; Nevada).
— Fall and Cockerell, 1907: 166 (Philonthus; New Mexico).
— Bernhauer and Schubert, 1914: 358 (Philonthus; catalog).
— Hatch, 1957: 185 (Philonthus; characters; British Columbia; Washington; Oregon; Idaho).
— Smetana, 1995e: 338 (Philonthus; characters; lectotype designation: Cal [California]; British Columbia; Arizona; California; Colorado; Idaho; Missouri; Nevada; Oregon; Utah; Washington; Mexico).
▴ Distribution: Canada, USA, Mexico.
tricoloricornis Coiffait, 1977, see: Craspedomerus.
tricoloris Schubert, 1908: 620 (Philonthus; Type locality: Palumbus, Himalaya).
— Bernhauer and Schubert, 1914: 358 (Philonthus; catalog).
— Cameron, 1932: 113 (Philonthus; synonym of aeneipennis).
— Scheerpeltz, 1933: 1331 (Philonthus; synonym of aeneipennis).
— Gridelli, 1934: 74 (Philonthus; characters; Karakoram region).
— Gridelli, 1936a: 150 (Philonthus; characters; valid species).
— Cerruti, 1951: 16 (Philonthus; India).
— Scheerpeltz, 1965: 107 (Philonthus; valid species; Burma).
— Scheerpeltz, 1976: 34 (Philonthus; Nepal).
— Scheerpeltz, 1976a: 87 (Philonthus; synonym of aeneipennis).
▴ Distribution: “Karakorum region”, India, Nepal, Myanmar.
trilobatus Tottenham, 1949b: 335 (Philonthus; Type locality: Lubero).
trinitatis Blackwelder, 1943: 418 (Philonthus; Type locality: Trinidad: St. Augustine).
tripunctatus Cameron, 1919, see: donckieri Bernhauer, 1915.
tripunctatus Coiffait, 1960 [Note: Nomen nudum], see: montivagus Heer, 1839.
triseriatus Bernhauer, 1928, see: Mentophilonthus.
tristichus Cameron, 1929: 62 (Philonthus; Type locality: Haut Uele: Mauda; Dika).
— Scheerpeltz, 1933: 1365 (Philonthus; catalog).
— Tottenham, 1962: 158 (Philonthus; aedeagus).
▴ Distribution: Congo.
tristipennis Bernhauer, 1941c: 289 [= 1951: 278] (Philonthus; Type locality: Süd Peru: Querobamba, 3520 m; Pampa de Cangallo, 3450 m).
trisulensis Coiffait, 1982, see: azuripennis Cameron, 1928.
trochanterinus Sharp, 1885: 417 (Philonthus; Type locality: Guatemala).
trochilus Solsky, 1872: 299 (Philonthus; Type locality: Venezuela).
— Fauvel, 1891: 114 (Philonthus; Venezuela).
— Bernhauer and Schubert, 1914: 358 (Philonthus; catalog).
▴ Distribution: Venezuela.
trossuloides Cameron, 1933, see: Gabrius.
trossulus Nordmann, 1837, see: Gabrius.
trunculus Herman, 2001: 48 (Philonthus; replacement name for analis Fauvel).
analis Fauvel, 1907: 46 (Philonthus; [preoccupied]; Type locality: Nairobi).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog).
— Herman, 2001: 48 (Philonthus; synonym of trunculus).
truquii Peyron, 1858: 427 (Philonthus; Type locality: Not cited, presumably Tarsous).
— Baudi, 1870: 386 (Philonthus; characters).
— Fauvel, 1874: 258 [= 1874a: 484] (Philonthus; synonym of micans).
— Ganglbauer, 1895: 457 (Philonthus; synonym of micans).
— Bernhauer and Schubert, 1914: 346 (Philonthus; synonym of micans).
— Jarrige, 1951a: 7, 9 (Philonthus; valid species; characters).
— Smetana, 1954b: 178 (Philonthus; valid species; Turkey).
— Coiffait, 1963: 12 (Paragabrius).
— Coiffait, 1974: 97 (Paragabrius; characters; Turkey).
▴ Distribution: Turkey, Syria.
sahlbergorum Renkonen, 1937: 76 (Philonthus; Type locality: Syrien: Sanamein).
tuberculatus Coiffait, 1974, see: wuesthoffi Bernhauer, 1939.
tucumanensis Bernhauer, 1927e: 245 (Philonthus; Type locality: Argentinien: Prov. Tucuman, San Pedro Colaloo).
tucumanensis Bernhauer, 1934, see: argus Herman, 2001.
tukchensis Coiffait, 1982: 67 (Philonthus; Type locality: Nepal: Thakkhola, Tukche, Thaksang, 3100–3300).
tumulinus Tottenham, 1955: 172 (Philonthus; Type locality: Transvaal: Pretoria).
— Tottenham, 1956a: 243 (Philonthus; characters; Transvaal; Cape Colony).
— Tottenham, 1961: 204 (Philonthus; Senegal).
— Lecoq, 1990: 202 (Philonthus; Mascarene Islands).
▴ Distribution: Senegal, South Africa, Mascarene Islands.
turbatus Erichson, 1840: 466 (Philonthus; Type locality: Sardinia).
— Kraatz, 1857d: 596 (Philonthus).
— Fauvel, 1874: 234 [= 1874a: 459] (Philonthus; synonym of corruscus).
— Fauvel, 1886: 54 [= 1886a: 46] (Philonthus; Algeria).
— Fauvel, 1902b: 109 (Philonthus; Algeria; Tunisia; Egypt).
— Bernhauer and Schubert, 1914: 358 (Philonthus; catalog).
— Scheerpeltz, 1933: 1365 (Philonthus; catalog).
— Coiffait, 1967a: 422 (Philonthus; characters; North Africa; Sardinia; Egypt).
— Coiffait, 1974: 262 (Philonthus; characters; Sardinia; North Africa).
— Ciceroni and Zanetti, 1995: 29 (Philonthus; Italy).
▴ Distribution: Egypt, Tunisia, Algeria, Italy.
erythropterus Fauvel, 1872b: 35 (Philonthus; Type locality: Oran).
— Fauvel, 1874: 225 [= 1874a: 451] (Philonthus; characters; Algeria).
— Fauvel, 1878: 122 [= 1878a: 42] (Philonthus; Algeria).
— Fauvel, 1886: 54 [= 1886a: 46] (Philonthus; synonym of turbatus).
— Fauvel, 1902b: 109 (Philonthus; synonym of turbatus).
— Bernhauer and Schubert, 1914: 358 (Philonthus; synonym of turbatus).
— Coiffait, 1974: 262 (Philonthus; synonym of turbatus).
turbidus Erichson, 1840: 484 (Philonthus; Type locality: Madagascar; [Note: See lectotype designation by Smetana, 1995e]).
— Cocquerel, 1866: 316 (Philonthus; characters; Réunion; Madagascar).
— Fauvel, 1867b: 66 [= 1868a: 70] (Philonthus; characters; Réunion; Madagascar).
— Wollaston, 1867: 240 (Philonthus; characters; notes; Cape Verde Islands).
— Fauvel, 1878: 124 [= 1878a: 44] (Philonthus; Egypt).
— Blackburn and Sharp, 1885: 222 (Philonthus; Hawaii).
— Fauvel, 1886: 55 [= 1886a: 47] (Philonthus; Morocco; Egypt).
— Fauvel, 1902b: 110 (Philonthus; Morocco; Egypt; Madeira; Canaries).
— Peyerimhoff, 1907: 14 (Philonthus; Sinai).
— Sharp, 1908: 548 (Philonthus; Hawaii).
— Bernhauer and Schubert, 1914: 358 (Philonthus; catalog).
— Bernhauer, 1915n: 319 (Philonthus; German Southwest Africa).
— Gridelli, 1930a: 88 (Philonthus; characters; Eritrea, Cape Verde).
— Cameron, 1932: 141 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1365 (Philonthus; catalog).
— Bernhauer and Chapman, 1939: 71 (Philonthus; Ethiopia).
— Tottenham, 1949b: 354 (Philonthus; characters).
— Tottenham, 1956a: 238 (Philonthus; characters; Madagascar; Cape Verde Archipelago; Canary Islands; Madeira; Gaboon; Anglo-Egyptian Sudan; British and Italian Somaliland; Tanganyika; Mozambique; Natal; Cape Colony; Hawaii).
— Tottenham, 1961: 204 (Philonthus; Senegal).
— Scheerpeltz, 1961g: 238 (Philonthus; Madagascar).
— Outerelo, Aranda, and Gamarra, 1992 (Philonthus; male and female genitalia; general distribution; Spain).
— Scheerpeltz, 1963b: 429 (Philonthus; Egypt).
— Coiffait, 1966b: 338 (Philonthus; Hoggar).
— Coiffait, 1967a: 431 (Philonthus; characters; Russia; Syria; Egypt; Erythrea; Hoggar; Cape Verde Islands).
— Coiffait, 1974: 213 (Philonthus; characters; Madagascar; Egypt; Sahara; Morocco; Cape Verde Islands; Syria; Russia).
— Scheerpeltz, 1974: 143 (Philonthus; South West Africa; South Africa).
— Scheerpeltz, 1974c: 21 (Philonthus; characters; Sudan; Ethiopia; Somalia).
— Moore, 1975c: 106 (Philonthus; Hawaii).
— Jarrige, 1978: 289 (Philonthus; Madagascar).
— Hoebeke, 1978: 508 (Philonthus; parasitized by proctotrupid).
— Coiffait, 1979a: 162 (Philonthus; Saudi Arabia).
— Coiffait, 1981b: 236 (Philonthus; Saudi Arabia).
— Frank, 1982: 44 (Philonthus; list of parasites).
— Coiffait, 1986: 393 (Philonthus; Saudi Arabia).
— Lecoq, 1990: 200 (Philonthus; notes; Mascarene Islands).
— Maachi, 1991: 251 (Philonthus; Morocco).
— Nishida, 1994: 73 (Philonthus; Hawaii).
— Ciceroni and Zanetti, 1995: 29 (Philonthus; Italy).
— Smetana, 1995e: 442 (Philonthus; lectotype designation; characters; notes; California).
— Gusarov, 1997: 286 (Philonthus; Saudi Arabia).
— Nishida, 1997: 62 (Philonthus; Hawaii).
▴ Distribution: Réunion, Madagascar, Mozambique, South Africa, Namibia, Gabon, Tanzania, Somalia, Ethiopia, Eritrea, Sudan, Saudi Arabia, Egypt, Morocco, Senegal, Cape Verde Islands, Canary Islands, Madeira, Spain, Russia, Syria, India, Hawaii, USA.
pharao Saulcy, 1864: 639 (Philonthus; Type locality: Le Caire).
— Fauvel, 1875a: XXXI [= 1875b: 233] (Philonthus; synonym of turbidus).
— Fauvel, 1902b: 110 (Philonthus; synonym of turbidus).
— Bernhauer and Schubert, 1914: 358 (Philonthus; synonym of turbidus).
— Cameron, 1932: 141 (Philonthus; synonym of turbidus).
— Tottenham, 1956a: 237 (Philonthus; probably synonym of turbidus).
— Coiffait, 1974: 213 (Philonthus; synonym of turbidus).
— Scheerpeltz, 1974c: 21 (Philonthus; synonym of turbidus).
— Smetana, 1995e: 442 (Philonthus; synonym of turbidus).
rubiginosus Solsky, 1867: 30 (Philonthus; Type locality: Egypte).
— Fauvel, 1869: 489 (Philonthus; synonym of turbidus).
— Fauvel, 1875a: XXXI [= 1875b: 233] (Philonthus; synonym of turbidus).
— Fauvel, 1878: 124 [= 1878a: 44] (Philonthus; synonym of turbidus).
— Fauvel, 1902b: 110 (Philonthus; synonym of turbidus).
— Bernhauer and Schubert, 1914: 358 (Philonthus; synonym of turbidus).
— Cameron, 1932: 141 (Philonthus; synonym of turbidus).
— Tottenham, 1956a: 237 (Philonthus; probably synonym of turbidus).
— Coiffait, 1974: 213 (Philonthus; synonym of turbidus).
— Scheerpeltz, 1974c: 21 (Philonthus; synonym of turbidus).
— Smetana, 1995e: 442 (Philonthus; synonym of turbidus).
turbo Smetana, 1995e: 276 (Philonthus; Type locality: Alberta: Upper Waterfowl Lk., Banff N. P., 5700 ft.).
turcmenus Fauvel, 1900, see: Gabrius.
turneri Cameron, 1951, see: Gabrius.
turneri Tottenham, 1949, see: Gabronthus.
ubadilius Tottenham, 1956: 280 (Philonthus; Type locality: Urundi: Bururi, 2100 m).
uelensis Bernhauer, 1928g: 112 (Philonthus; Type locality: Belgisch-Kongo: Stanleyville a Kilo).
— Scheerpeltz, 1933: 1365 (Philonthus; catalog).
— Tottenham, 1949b: 321 (Philonthus; characters).
▴ Distribution: Congo.
ugandae Bernhauer, 1937: 302 (Philonthus; Type locality: Uganda: Nakasongola, Buruli).
— Tottenham, 1955: 157 (Philonthus; subgenus Spatulonthus; notes; N.W. Rhodesia).
▴ Distribution: Uganda, Zambia.
ullrichi Last, 1989: 877 (Philonthus; Type locality: Papua New Guinea: Kainantu, Onerunka).
umboiensis Last, 1987: 30 (Philonthus; Type locality: Papua New Guinea: New Guinea NE: Umboi, Semoto Awelcom, 470–600 m).
umbratilis Gravenhorst, 1802: 170 (Staphylinus; Type locality: America septentr.).
— Gravenhorst, 1806: 69 (Staphylinus).
— Say, 1830: 38 (Staphylinus; characters).
— Say, 1834: 451 (Staphylinus; characters).
— Nordmann, 1837: 88 (Philonthus; characters; America septentrionalis).
— Erichson, 1839a: 454 (Philonthus; characters; Germany).
— Heer, 1839: 261 (Philonthus; characters; Switzerland).
— Erichson, 1840: 445 (Philonthus; characters; Europe; America septentrionalis).
— Kolenati, 1846: 20 (Philonthus; Caucasi declivitas septentrionalis).
— Jacobsen, 1849: 691 (Philonthus; characters; Denmark).
— Hochhuth, 1849: 136 (Philonthus; cited as umbratitus; characters).
— Redtenbacher, 1849: 824 (Philonthus; characters; Austria).
— Küster, 1853a: 54 (Philonthus; characters; Europe).
— Hardy, 1851: 27 (Philonthus; notes; England).
— Wollaston, 1854: 581 (Philonthus; characters; notes; Madeira Islands).
— Fairmaire and Laboulbène, 1856: 518 (Philonthus; characters; France).
— Wollaston, 1857: 189 (Philonthus; habitat; Madeira Islands).
— Kraatz, 1857d: 583 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 193 (Philonthus; characters; Austria).
— Thomson, 1860: 158 (Philonthus; characters; Scandinavia).
— Hochhuth, 1862: 55 (Philonthus; Russia).
— Wollaston, 1864: 569 (Philonthus; notes; Canary Islands).
— Fauvel, 1865a: 296 [= 1865b: 46] (Philonthus; habitat; France).
— Crotch, 1867a: 383 (Philonthus; Azores).
— Fauvel, 1874: 221 [= 1874a: 447] (Philonthus; characters; France; Syria; Caucasus; Madeira; North America).
— Redtenbacher, 1874: 212 (Philonthus; characters; Austria).
— Seidlitz, 1875: 274 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 420 [= 1877: 276] (Philonthus; characters; France).
— Horn, 1884: 184 (Philonthus; characters; Massachusetts; New Jersey; Lake Superior).
— Fowler, 1888: 267 (Philonthus; characters; Britain).
— Ganglbauer, 1895: 450 (Philonthus; characters; Europe; Mediterranean region; North America).
— Xambeu, 1900: 29 (Philonthus; larval characters).
— Fauvel, 1902b: 108 (Philonthus; Tunisia; Azores; Madeira; Canaries).
— Poppius, 1905: 10 (Philonthus; Orkney Islands; Shetland Islands).
— Sainte-Claire Deville, 1906: 96 (Philonthus; Corsica).
— Reitter, 1909: 126 (Philonthus; characters; Germany).
— Blatchley, 1910: 381 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 358 (Philonthus; catalog).
— Johansen, 1914: 401 (Philonthus; characters; Denmark).
— Portevin, 1929: 359 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1365 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Bernhauer, 1940c: 4 (Philonthus; Azores).
— Méquignon, 1942a: 21 (Philonthus; Azores).
— V. Hansen, 1952: 60 (Philonthus; subgenus Philonthus; characters; Denmark).
— Hatch, 1957: 195 (Philonthus; characters; British Columbia; Washington; Oregon).
— Smetana, 1958: 209 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 151 (Philonthus; characters).
— Scheerpeltz, 1961d: 129 (Philonthus; Italy).
— Palm, 1963b: 77 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 179 (Philonthus; subgenus Philonthus; characters; central Europe).
— Scheerpeltz, 1965b: 402 (Philonthus; Sicily).
— Smetana, 1965: 18 (Philonthus; notes; Nova Scotia).
— Horion, 1965: 146 (Philonthus; all Europe).
— Coiffait, 1967a: 403 (Philonthus; characters; Europe; Siberia; Madeira; Canary Islands).
— Raianu, 1968a: 467 (Philonthus; Romania).
— Szujecki, 1968a: 731 (Philonthus; Poland).
— Coiffait, 1974: 272 (Philonthus; characters; Holarctic).
— Pace and Zanetti, 1977: 297 (Philonthus; Italy).
— Pope, 1977: 29 (Philonthus; Britain).
— Frank, 1982: 44 (Philonthus; list of parasites).
— Boller, 1983: 118 (Philonthus; larval characters).
— Tóth, 1984: 57 (Philonthus; characters; Hungary).
— Lucht, 1987: 104 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Borges, 1990: Table II (Philonthus; Azores).
— Pilon and Zanetti, 1991: 57 (Philonthus; Italy).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Philonthus; Italy).
— Smetana, 1995e: 169 (Philonthus; characters; notes; Alberta; British Columbia; Manitoba; New Brunswick; Nova Scotia; Ontario; Prince Edward Island; Quebec; Idaho; Illinois; Indiana; Kentucky; Maine; Massachusetts; Michigan; New Hampshire; New Jersey; New York; Ohio; Oregon; Pennsylvania; Utah; Washington; Wisconsin).
— Downie and Arnett, 1996: 376 (Philonthus; characters; USA).
— Owen, 1997a: 149 (Philonthus; collecting sites; Ireland).
▴ Distribution: Tunisia, Canary Islands, Madeira, Azores, Europe, Russia, Caucasus, Canada, USA.
subfuscus Gyllenhal, 1810: 326 (Staphylinus; Type locality: Suecia).
— C. Sahlberg, 1830: 323 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 29 (Staphylinus; Finland).
— Mannerheim, 1831: 443 (Staphylinus; Finland).
— Stephens, 1832: 233 (Philonthus; characters; England).
— Runde, 1835: 8 (Staphylinus; characters; Germany).
— Lacordaire, 1835: 395 (Staphylinus; characters; France).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Nordmann, 1837: 84 (Philonthus; Germany; Sweden; France).
— Erichson, 1839a: 454 (Philonthus; synonym of umbratilis).
— Heer, 1839: 261 (Philonthus; synonym of umbratilis).
— Stephens, 1839: 396 (Philonthus; characters; England).
— Erichson, 1840: 445 (Philonthus; synonym of umbratilis).
— Kolenati, 1846: 20 (Philonthus; synonym of umbratilis).
— Redtenbacher, 1849: 824 (Philonthus; synonym of umbratilis).
— Hardy, 1851: 27 (Philonthus; synonym of umbratilis).
— Wollaston, 1854: 581 (Philonthus; synonym of umbratilis).
— Fairmaire and Laboulbène, 1856: 519 (Philonthus; synonym of umbratilis).
— Kraatz, 1857d: 583 (Philonthus; synonymn of umbratilis).
— Ganglbauer, 1895: 450 (Philonthus; synonym of umbratilis).
— Bernhauer and Schubert, 1914: 358 (Philonthus; synonym of umbratilis).
— Smetana, 1958: 209 (Philonthus; synonym of umbratilis).
— Coiffait, 1974: 273 (Philonthus; synonym of umbratilis).
— Boháč, 1988: 551 [= 1989: 34] (Philonthus; synonym of umbratilis).
— Smetana, 1995e: 169 (Philonthus; synonym of umbratilis).
umbratus Sharp, 1885: 424 (Philonthus; Type locality: Mexico, Las Vigas; Guatemala, Paso Antonio; Panama, Bugaba, Volcan de Chiriqui 4000 to 6000 feet, Peña Blanca 3000 to 4000 feet).
— Bernhauer and Schubert, 1914: 358 (Philonthus; catalog).
▴ Distribution: Mexico, Guatemala, Panama.
umbrinoides Smetana, 1995e: 268 (Philonthus; Type locality: Quebec: Parc Gatineau, Lac Borugeois. Also cited from Alberta, Manitoba, Nova Scotia, Ontario, Maine, Massachusetts, Michigan, Minnesota, New York, West Virginia, Wisconsin).
umbrinus Gravenhorst, 1802: 169 (Staphylinus; Type locality: America septentr.).
— Gravenhorst, 1806: 69 (Staphylinus).
— Erichson, 1840: 464 (Philonthus; characters; America septentrionalis).
— Horn, 1884: 191 (Philonthus; characters; New York to South Carolina).
— Blatchley, 1910: 382 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 358 (Philonthus; catalog).
— Smetana, 1965: 18 (Philonthus; characters; Nova Scotia).
— Smetana, 1995e: 113 (Philonthus; characters; Ontario; Alabama; Arkansas; Georgia; Illinois; Indiana; Kansas; Kentucky; Louisiana; Maryland; Massachusetts; Mississippi; Missouri; New Jersey; New York; North Carolina; Oklahoma; Pennsylvania; South Carolina; Tennessee; West Virginia; Virginia).
— Downie and Arnett, 1996: 378 (Philonthus; characters; USA).
▴ Distribution: Canada; USA.
moestus Gravenhorst, 1806: 47 (Staphylinus; Type locality: America septentr.).
— Nordmann, 1837: 97 (Philonthus; America borealis).
— Erichson, 1840: 464 (Philonthus; synonym of umbrinus).
— Bernhauer and Schubert, 1914: 358 (Philonthus; synonym of umbrinus).
— Smetana, 1995e: 113 (Philonthus; synonym of umbrinus).
umbripennis LeConte, 1863, see: Neobisnius.
undae Penecke, 1901, see: rufipes Stephens, 1832.
unguicularis Tottenham, 1949b: 323 (Philonthus; Type locality: Equateur: Flandria).
unicolor Stephens, 1832, see: varians Paykull, 1789.
unicolor Seidlitz, 1875, see: sanguinolentus Gravenhorst, 1802.
unicolor Leinberg, 1902, see: caucasicus Nordmann, 1837.
uniformis Cameron, 1937: 11 (Philonthus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft).
uniformis Last, 1981 [Note: Nomen nudum], see: dignus Last, 1968.
unzenensis Bernhauer, 1939, see: Gabrius.
upotovus Tottenham, 1962 [Note: Nomen nudum], see: praetor Tottenham, 1949.
upuzius Tottenham, 1962: 215, 216, 221, 222, 226 (Philonthus; Type locality: Not cited, presumably Africa).
uralensis Reitter, 1909, see: ephippium Nordmann, 1837.
usagarae Bernhauer, 1937b: 613 (Philonthus; Type locality: N. W. Usagara, 1700–1900 m).
usambaricus Bernhauer, 1937b: 612 (Philonthus; Type locality: W. Usambara).
ussuriensis Coiffait, 1974, see: virgatus Sharp, 1889.
ustipennis Fauvel, 1907, see: ustus Fauvel, 1907.
ustulatus Fauvel, 1875a: XXX [= 1875b: 232] (Philonthus; Type locality: Sibérie, région du Baïkal; Daourie).
— Bernhauer and Schubert, 1914: 358 (Philonthus; catalog).
— Scheerpeltz, 1933: 1365 (Philonthus; catalog).
— Smetana, 1963c: 295 (Philonthus; subgenus Philonthus; Mongolia).
— Smetana, 1965e: 173 (Philonthus; Mongolia).
— Coiffait, 1967a: 422 (Philonthus; characters; central Asia).
— Smetana, 1967d: 203 (Philonthus; Mongolia).
— Coiffait, 1974: 263 (Philonthus; characters; Siberia; Mongolia).
— Smetana 1975c: 170 (Philonthus; Mongolia).
— Shilov, 1976: 149 (Philonthus; Mongolia).
▴ Distribution: Russia, Mongolia.
ornatipennis Reitter, 1901b: 158 (Philonthus; Type locality: Quellgebiet des Irkut).
— Bernhauer and Schubert, 1914: 358 (Philonthus; synonym of ustulatus).
— Smetana, 1967d: 203 (Philonthus; synonym of ustulatus).
— Coiffait, 1974: 263 (Philonthus; synonym of ustulatus).
ustus Fauvel, 1907: 43 (Philonthus; Type locality: Mombasa).
— Bernhauer and Schubert, 1914: 358 (Philonthus; catalog).
— Scheerpeltz, 1933: 1365 (Philonthus; catalog).
— Bernhauer and Chapman, 1939: 67 (Philonthus; cited as uxtus; Kenya).
— Bernhauer, 1940b: 635 (Philonthus; subgenus Cephalonthus).
— Cameron, 1950g: 38 (Philonthus; Belgian Congo).
▴ Distribution: Congo, Kenya, Ethiopia.
ustipennis Fauvel, 1907: 43 (Philonthus; variety of ustus; Type locality: Ethiopie: Addis Abeba; Cap).
— Bernhauer and Schubert, 1914: 358 (Philonthus; variety of ustus).
— Rambousek, 1922: 1010 (Philonthus; Ethiopia).
— Scheerpeltz, 1933: 1366 (Philonthus; variety of ustus).
— Tottenham, 1962: 136 (Philonthus; Kenya).
vacans Tottenham, 1953, see: variipennis Kraatz, 1859.
vadoni Lecoq, 1997: 284 (Philonthus; Type locality: Madagascar Nord, montagne d'Ambre).
vagepunctatus Schubert, 1909: 292 (Philonthus; Type locality: südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d'Orizaba).
vagus Fauvel, 1895: 263 (Philonthus; Type locality: Bírmanie, Carin Asciuii Ghécu, 1400–1500 m; Tenasserim, Thagata; Malacca; Sumatra).
— Bernhauer and Schubert, 1914: 358 (Philonthus; catalog).
— Cameron, 1932: 119 (Philonthus; characters; Burma).
— Scheerpeltz, 1933: 1366 (Philonthus; catalog).
▴ Distribution: Myanmar, Malaysia, Indonesia.
distincticollis Cameron, 1932a: 141 (Philonthus; variety of vagus; Type locality: Selangor: The Gap, Bukit Kutu; Perak: Gunong Kledang).
validus Casey, 1915: 435 (Philonthus; Type locality: New York; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from New York: near the city and at “Cairo”; Pennsylvania: Harrisburg).
— Scheerpeltz, 1933: 1366 (Philonthus; catalog).
— Smetana, 1995e: 80 (Philonthus; lectotype designation: N.Y. [New York]; characters; Alberta; British Columbia; Manitoba; New Brunswick; Newfoundland; Nova Scotia; Ontario; Quebec; Alaska; Arkansas; Connecticut; Georgia; Illinois; Indiana; Iowa; Kansas; Kentucky; Maine; Maryland; Massachusetts; Michigan; Montana; Nebraska; New Hampshire; New Jersey; New York; North Carolina; North Dakota; Ohio; Oklahoma; Pennsylvania; South Carolina; Virginia; West Virginia; Wisconsin).
— Downie and Arnett, 1996: 380 (Philonthus; characters; USA).
— Downie and Arnett, 1996: 376 (Philonthus; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232); North America reports may be P. validus based on comments by Smetana, 1995a: 84-85]; characters; USA).
— Bain, 1998: 39 (Philonthus; 17th century remains; Massachusetts).
▴ Distribution: Canada, USA.
vanhoofi Bernhauer, 1935d: 102 (Philonthus; Type locality: W. Ruwenzori: Kalonge).
— Tottenham, 1956: 226, 278 (Philonthus; characters; Ruanda).
— Tottenham, 1962: 163 (Philonthus; characters).
▴ Distribution: Congo, Rwanda.
ruandae Cameron, 1950g: 37 (Philonthus; Type locality: Ruanda: Ninda, alt. 2150 m).
variabilis Eppelsheim, 1892: 333 (Philonthus; Type locality: Taschkent).
— Bernhauer and Schubert, 1914: 359 (Philonthus; catalog).
— Scheerpeltz, 1933: 1366 (Philonthus; catalog).
— Coiffait, 1967a: 401 (Philonthus; characters; Turkestan; Transcaspian region).
— Coiffait, 1974: 256 (Philonthus; characters; Uzbekistan; Turkestan).
▴ Distribution: Uzbekistan.
varians Paykull, 1789: 45 (Staphylinus; Type locality: Suecia).
— Fabricius, 1793: 524 (Staphylinus; characters).
— Olivier, 1795:(42): 27 (Staphylinus; characters).
— Panzer, 1795: 354 (Staphylinus; characters; Germany).
— Paykull, 1800: 393 (Staphylinus; characters; Sweden).
— Fabricius, 1801: 594 (Staphylinus; characters).
— Turton, 1802: 511 (Staphylinus; characters; Zealand).
— Gravenhorst, 1802: 37 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 359 cited bipustulatus Gravenhorst as a synonym of varians but Gravenhorst did not describe the species as new, he attributed it to Panzer who attributed it to Linné]; characters; varieties).
— Latreille, 1804: 309 (Staphylinus; characters).
— Gravenhorst, 1806: 82 (Staphylinus; characters; Germany).
— Gyllenhal, 1810: 342 (Staphylinus; characters; Sweden).
— Zetterstedt, 1828: 77 (Staphylinus; characters; Lapland).
— Curtis, 1829: 25 (Philonthus; catalog; Britain).
— C. Sahlberg, 1830: 329 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 30 (Staphylinus; Finland; Sweden).
— Mannerheim, 1831: 444 (Staphylinus; Finland; Sweden).
— Stephens, 1832: 234 (Philonthus; characters; England).
— Ménétriés, 1832: 146 (Staphylinus; Caucasus-Persia region).
— Runde, 1835: 9 (Staphylinus; characters; Germany).
— Lacordaire, 1835: 394 (Staphylinus; characters; France).
— Nordmann, 1837: 93 (Philonthus; Europe).
— Zetterstedt, 1838: 64 (Staphylinus; characters; habitat; Lapland).
— Heer, 1839: 264 (Philonthus; characters; Switzerland).
— Heer, 1841: 582 (Philonthus; notes).
— P. Lucas, 1846: 112 (Philonthus; notes; Algeria).
— Jacobsen, 1849: 699 (Philonthus; characters; Denmark).
— Redtenbacher, 1849: 705 (Philonthus; characters; Austria).
— Hardy, 1851: 29 (Philonthus; notes; England).
— Wollaston, 1854: 583 (Philonthus; characters; notes; Madeira Islands).
— Scriba, 1855: 300 (Philonthus; Venezuela).
— Fairmaire and Laboulbène, 1856: 524 (Philonthus; characters; France).
— Guérin-Méneville, 1857: 39 (Philonthus; characters; St. John; St. Vincent; Cuba).
— Kraatz, 1857d: 602 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 195 (Philonthus; characters; Austria).
— Peyron, 1858: 425 (Philonthus; Turkey).
— Chevrolat and Fauvel, 1863: 435 (Philonthus; characters; Cuba).
— Fauvel, 1874: 255 [= 1874a: 481] (Philonthus; characters; France; Cyprus; Syria; Turkey; Caucasus; Siberia).
— Redtenbacher, 1874: 213 (Philonthus; characters; Austria).
— Mulsant and Rey, 1876: 475 [= 1877: 331] (Philonthus; characters; France).
— Fauvel, 1878: 125 [= 1878a: 45] (Philonthus; Morocco; Algeria).
— Horn, 1884: 195 (Philonthus; characters; northern portions of Atlantic region west to Washington Territory and California).
— Fauvel, 1886: 56 [= 1886a: 48] (Philonthus; Morocco; Algeria).
— Fowler, 1888: 273 (Philonthus; characters; Britain).
— Eppelsheim, 1892: 334 (Philonthus; Turkestan).
— Ganglbauer, 1895: 454 (Philonthus; characters; Palaearctic region).
— Fauvel, 1902b: 112 (Philonthus; Morocco; Algeria).
— Poppius, 1905: 10 (Philonthus; Orkney Islands).
— Fauvel, 1907: 45 (Philonthus; Kijabe; Naivasha; Nairobi; Escarpment; Abyssinia; Usambara; Natal; Transvaal; Cape).
— Fall and Cockerell, 1907: 166 (Philonthus; New Mexico).
— Montandon, 1908: 80 (Philonthus; Romania).
— Sainte-Claire Deville, 1906: 99 (Philonthus; Corsica).
— Poppius, 1909: 25 (Philonthus; subgenus Gefyrobius; Lena Valley, Siberia).
— Reitter, 1909: 130 (Philonthus; characters; Germany).
— Eichelbaum, 1913: 130 (Philonthus; Tanzania).
— Bernhauer and Schubert, 1914: 359 (Philonthus; catalog).
— Johansen, 1914: 409 (Philonthus; characters; Denmark).
— Eichelbaum, 1915a: 92, 94 (Philonthus; abdominal segmentation; apical segements).
— Gridelli, 1920a: 18, 20 (Philonthus; characters; Italy).
— Rambousek, 1922: 1009 (Philonthus; Ethiopia).
— Šulc, 1922 (Philonthus; osmeterium; anatomy; histology; physiology).
— Portevin, 1929: 363 (Philonthus; characters; France).
— Cameron, 1932: 131 (Philonthus; characters; Burma).
— Scheerpeltz, 1933: 1366 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Paulian, 1941: 225 (Philonthus; larval characters).
— Blackwelder, 1943: 412 (Philonthus; characters; Cuba; St. John; St. Vincent).
— V. Hansen, 1952: 69 (Philonthus; subgenus Philonthus; characters; Denmark).
— Scheerpeltz, 1956c: 1101 (Philonthus; Cyprus).
— Hatch, 1957: 188 (Philonthus; characters; British Columbia; Washington; Oregon).
— Smetana, 1958: 195 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 160 (Philonthus; characters).
— Palm, 1963b: 87 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 184 (Philonthus; subgenus Philonthus; characters; central Europe).
— Smetana, 1965: 17 (Philonthus; Newfoundland).
— Horion, 1965: 160 (Philonthus; all Europe).
— Pototskaia, 1966: 157 (Philonthus; larval characters).
— Pototskaia, 1967: 61 (Philonthus; larval characters).
— Coiffait, 1967a: 419 (Philonthus; characters).
— Raianu, 1968: 114 (Philonthus; characters; ecological notes; Romania).
— Raianu, 1968a: 466 (Philonthus; Romania).
— Szujecki, 1968a: 730 (Philonthus; Poland).
— Coiffait, 1970e: 62 (Philonthus; Cyprus).
— Jarrige, 1971: 495 (Philonthus; Iran).
— Scheerpeltz, 1972: 19 (Philonthus; Argentina).
— Bacchus and Hammond, 1972: 63 (Philonthus; donkey dung; Britain).
— Coiffait, 1974: 244 (Philonthus; characters; Europe; Britain; Scandinavia).
— Shilov, 1975a: 375 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Coiffait, 1976: 217 (Philonthus; Tahiti).
— Shilov, 1976: 150 (Philonthus; Mongolia).
— Pace and Zanetti, 1977: 295 (Philonthus; Italy).
— Pope, 1977: 30 (Philonthus; Britain).
— Hinton, 1981: 661 (Philonthus; characters of egg).
— Coiffait, 1982a: 232 (Philonthus; India).
— Frank, 1982: 44 (Philonthus; list of parasites).
— Tóth, 1984: 71 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Pilon and Zanetti, 1991: 63 (Philonthus; notes; Italy).
— Maachi, 1991: 250 (Philonthus; Morocco).
— Welch, 1993: 228 (Philonthus; ovariole number and ovary structure).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Dearborn and Donahue, 1994: 15 (Philonthus; Maine).
— Ciceroni and Zanetti, 1995: 29 (Philonthus; Italy).
— Smetana, 1995e: 235 (Philonthus; characters; notes; Alberta; British Columbia; Manitoba; New Brunswick; Newfoundland; Nova Scotia; Ontario; Prince Edward Island; Quebec; Saskatchewan; California; Connecticut; Delaware; Idaho; Illinois; Indiana; Iowa; Kansas; Kentucky; Maine; Massachusetts; Michigan; Minnesota; New Hampshire; New York; North Carolina; Ohio; Oregon; Pennsylvania; Vermont; Washington; Wisconsina).
— Levesque and Levesque, 1996: 292 (Philonthus; seasonal abundance; Quebec).
— Downie and Arnett, 1996: 378 (Philonthus; characters; USA).
— Owen, 1997a: 149 (Philonthus; collecting sites; Ireland).
— Kashcheev, 1999a: 58 (Philonthus; Kazakhstan).
— Marchal and Chardonnet, 2000: 72 (Philonthus; Corsica).
▴ Distribution: South Africa, Gabon, Congo, Tanzania, Madagascar, Kenya, Ethiopia, Algeria, Morocco, Madeira, Europe, Russia, Caucasus, Turkey, Syria, Uzbekistan, Kazakhstan, Mongolia, India, Myanmar, Society Islands, Canada, USA, West Indies, Venezuela.
nitens Gravenhorst, 1802: 26 (Staphylinus; [preoccupied]; Type locality: Not cited, but presumably Brunsvigae).
— Latreille, 1804: 315 (Staphylinus; characters).
— Gravenhorst, 1806: 78 (Staphylinus; characters).
— Curtis, 1829: 25 (Philonthus; catalog; Britain).
— Stephens, 1832: 236 (Philonthus; characters; England).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Stephens, 1839: 397 (Philonthus; characters; England).
— Bernhauer and Schubert, 1914: 359 (Philonthus; synonym of varians).
— Cameron, 1932: 131 (Philonthus; synonym of varians).
— Blackwelder, 1943: 412 (Philonthus; synonym of varians).
— Smetana, 1958: 195 (Philonthus; synonym of varians).
— Smetana, 1995e: 235 (Philonthus; synonym of varians).
opacus Gravenhorst, 1802: 26 (Staphylinus; [preoccupied]; [Note: Bernhauer and Schubert, 1914: 359 cited Gyllenhal, 1810: 340 as the author but Gyllenhal attributed the species to Gravenhorst]; Type locality: Not cited, but presumably Brunsvigae).
— Latreille, 1804: 315 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 64 (Staphylinus; characters; Germany).
— Gyllenhal, 1810: 340 (Staphylinus; characters; Sweden).
— Zetterstedt, 1828: 77 (Staphylinus; characters; Lapland).
— Curtis, 1829: 25 (Philonthus; catalog; Britain).
— C. Sahlberg, 1830: 328 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 30 (Staphylinus; Finland; Sweden).
— Mannerheim, 1831: 444 (Staphylinus; Finland; Sweden).
— Stephens, 1832: 236 (Philonthus; characters; England).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Lacordaire, 1835: 404 (Staphylinus; characters; France).
— Nordmann, 1837: 98 (Philonthus; Sweden; Germany; England).
— Zetterstedt, 1838: 64 (Staphylinus; characters; habitat; Lapland).
— Erichson, 1839a: 465 (Philonthus; characters; Germany).
— Heer, 1839: 265 (Philonthus; characters; Switzerland).
— Heer, 1841: 583 (Philonthus; synonym of varians).
— P. Lucas, 1846: 112 (Philonthus; synonym of varians).
— Hardy, 1851: 29 (Philonthus; synonym of varians).
— Wollaston, 1854: 583 (Philonthus; synonym of varians).
— Fairmaire and Laboulbène, 1856: 524 (Philonthus; characters; France).
— Guérin-Méneville, 1857: 39 (Philonthus; synonym of varians).
— Kraatz, 1857d: 602 (Philonthus; synonym of varians).
— Thomson, 1860: 165 (Philonthus; characters; Scandinavia).
— Chevrolat and Fauvel, 1863: 435 (Philonthus; synonym of varians).
— Fauvel, 1866a: 345 [= 1866b: 96] (Philonthus; synonym of varians).
— Fauvel, 1869: 489 (Philonthus; synonym of varians).
— Seidlitz, 1875: 277 (Philonthus; characters; Baltic region).
— J. Sahlberg, 1876: 20 (Philonthus; Finland).
— Ganglbauer, 1895: 454 (Philonthus; synonym of varians).
— Bernhauer and Schubert, 1914: 359 (Philonthus; synonym of varians).
— Cameron, 1932: 131 (Philonthus; synonym of varians).
— Blackwelder, 1943: 412 (Philonthus; synonym of varians).
— Smetana, 1958: 195 (Philonthus; synonym of varians).
— Smetana, 1995e: 235 (Philonthus; synonym of varians).
aterrimus Marsham, 1802: 513 (Staphylinus; [preoccupied]; Type locality: Britain).
— Gyllenhal, 1810: 341 (Staphylinus; synonym of opacus Gravenhorst).
— Curtis, 1829: 25 (Philonthus; catalog; Britain).
— C. Sahlberg, 1830: 328 (Staphylinus; synonym of opacus Gravenhorst).
— Stephens, 1832: 235 (Philonthus; characters; England).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Stephens, 1839: 397 (Philonthus; characters; England).
— P. Lucas, 1846: 112 (Philonthus; synonym of varians).
— Hardy, 1851: 29 (Philonthus; synonym of varians).
— Guérin-Méneville, 1857: 40 (Philonthus; synonym of varians).
— Kraatz, 1857d: 602 (Philonthus; synonym of varians).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of varians).
— Chevrolat and Fauvel, 1863: 435 (Philonthus; synonym of varians).
— Fauvel, 1866a: 345 [= 1866b: 96] (Philonthus; synonym of varians).
— Fauvel, 1869: 489 (Philonthus; synonym of varians).
— Ganglbauer, 1895: 454 (Philonthus; synonym of varians).
— Bernhauer and Schubert, 1914: 359 (Philonthus; synonym of varians).
— Cameron, 1932: 131 (Philonthus; synonym of varians).
— Blackwelder, 1943: 412 (Philonthus; synonym of varians).
— Smetana, 1958: 195 (Philonthus; synonym of varians).
— Smetana, 1995e: 235 (Philonthus; synonym of varians).
bimaculatus Marsham, 1802: 525 (Staphylinus; [preoccupied]; Type locality: Britain).
— Holme, 1837: 63 (Philonthus; Scilly Islands).
— Guérin-Méneville, 1857: 39 (Philonthus; synonym of varians).
— Kraatz, 1857d: 602 (Philonthus; synonym of varians).
— Chevrolat and Fauvel, 1863: 435 (Philonthus; synonym of varians).
— Bernhauer and Schubert, 1914: 359 (Philonthus; synonym of varians).
— Cameron, 1932: 131 (Philonthus; synonym of varians).
— Blackwelder, 1943: 412 (Philonthus; synonym of varians).
— Smetana, 1958: 195 (Philonthus; synonym of varians).
— Smetana, 1995e: 235 (Philonthus; synonym of varians).
unicolor Stephens, 1832: 224 (Quedius; Type locality: London; Norfolk; Suffolk; Devonshire).
— Stephens, 1839: 392 (Quedius; characters; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of varians).
— Fauvel, 1869: 489 (Philonthus; synonym of varians).
— Bernhauer and Schubert, 1914: 359 (Philonthus; synonym of varians).
— Scheerpeltz, 1933: 1366 (Philonthus; aberration of varians).
— Blackwelder, 1943: 412 (Philonthus; synonym of varians).
— Hatch, 1957: 188 (Philonthus; cited as valid species; characters; British Columbia; Washington; Oregon).
— Smetana, 1958: 195 (Philonthus; synonym of varians).
— Smetana, 1959: 160 (Philonthus; aberration of varians; characters).
— Coiffait, 1967a: 419 (Philonthus; aberration of varians; characters).
— Coiffait, 1974: 244 (Philonthus; variety of varians).
— Pope, 1977: 30 (Philonthus; synonym of varians).
— Moore and Legner, 1975: 34 (Philonthus; cited as valid; catalog).
— Smetana, 1995e: 235 (Philonthus; synonym of varians).
intaminatus Stephens, 1832: 235 (Philonthus; Type locality: London; Suffolk; Devonshire).
— Curtis, 1829: 25 (Philonthus; nomen nudum; catalog; Britain).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Stephens, 1839: 397 (Philonthus; characters; England).
— Hardy, 1851: 29 (Philonthus; synonym of varians).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of varians).
— Bernhauer and Schubert, 1914: 359 (Philonthus; synonym of varians).
— Cameron, 1932: 131 (Philonthus; synonym of varians).
— Blackwelder, 1943: 412 (Philonthus; synonym of varians).
— Smetana, 1958: 195 (Philonthus; synonym of varians).
— Smetana, 1995e: 235 (Philonthus; synonym of varians).
punctiventris Stephens, 1832: 235 (Philonthus; Type locality: Suffolk).
— Curtis, 1829: 25 (Philonthus; nomen nudum; catalog; Britain).
— Stephens, 1839: 396 (Philonthus; characters; England).
— Hardy, 1851: 29 (Philonthus; synonym of varians).
— Rye, 1870a: 82 (Philonthus; synonym of varians).
— Fauvel, 1874: 256 [= 1874a: 482] (Philonthus; synonym of varians).
— Ganglbauer, 1895: 454 (Philonthus; synonym of varians).
— Bernhauer and Schubert, 1914: 359 (Philonthus; synonym of varians).
— Cameron, 1932: 131 (Philonthus; synonym of varians).
— Blackwelder, 1943: 412 (Philonthus; synonym of varians).
— Smetana, 1958: 195 (Philonthus; synonym of varians).
— Smetana, 1995e: 235 (Philonthus; synonym of varians).
lituratus Stephens, 1832: 238 (Philonthus; Type locality: London; Devonshire; Suffolk; near Edinburgh).
— Curtis, 1829: 26 (Philonthus; nomen nudum; catalog; Britain).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Stephens, 1839: 397 (Philonthus; characters; England; Scotland).
— Hardy, 1851: 29 (Philonthus; synonym of varians).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of varians).
— Bernhauer and Schubert, 1914: 359 (Philonthus; synonym of varians).
— Cameron, 1932: 131 (Philonthus; synonym of varians).
— Blackwelder, 1943: 412 (Philonthus; synonym of varians).
— Smetana, 1958: 195 (Philonthus; synonym of varians).
— Smetana, 1995e: 235 (Philonthus; synonym of varians).
costatus Baudi, 1848: 28 (Philonthus; Type locality: Sardinia).
— Kraatz, 1857b: 176 (Philonthus; synonym of varians).
— Kraatz, 1857d: 574 (Philonthus).
— Kraatz, 1857d: 1060 (Philonthus; synonym of varians).
— Bernhauer and Schubert, 1914: 359 (Philonthus; synonym of varians).
— Cameron, 1932: 131 (Philonthus; synonym of varians).
— Blackwelder, 1943: 412 (Philonthus; synonym of varians).
— Smetana, 1958: 195 (Philonthus; synonym of varians).
— Smetana, 1995e: 235 (Philonthus; synonym of varians).
alpigradus Mulsant and Rey, 1876: 481 [= 1877: 337] (Philonthus; Type locality: la Grande-Chartreuse et mont Dore, Auvergne).
— Ganglbauer, 1895: 454 (Philonthus; synonym of varians).
— Bernhauer and Schubert, 1914: 359 (Philonthus; synonym of varians).
— Cameron, 1932: 131 (Philonthus; synonym of varians).
— Blackwelder, 1943: 412 (Philonthus; synonym of varians).
— Smetana, 1958: 195 (Philonthus; synonym of varians).
— Smetana, 1995e: 235 (Philonthus; synonym of varians).
scutatus Eppelsheim, 1895a: 127 (Philonthus; variety of varians; [preoccupied]; Type locality: Gabon).
— Fauvel, 1907: 45 (Philonthus; variety of peregrinus; Kilimandjaro; Obock; Abyssinia; Togo; Madagascar).
— Bernhauer and Schubert, 1914: 350 (Philonthus; variety of peregrinus).
— Rambousek, 1922: 1009 (Philonthus; variety of peregrinus; Ethiopia).
— Blackwelder, 1943: 412 (Philonthus; synonym of varians).
— Cameron, 1950g: 34 (Philonthus; variety of peregrinus; Belgian Congo).
— Smetana, 1958: 195 (Philonthus; synonym of varians).
— Jarrige, 1978: 291 (Spatulonthus; Madagascar).
— Smetana, 1995e: 235 (Philonthus; synonym of varians).
piceicoxis Gridelli, 1920: 18, 19, 20 (Philonthus; aberration of varians; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]; cited from Mte. Rima, Giudicarie, Trentino).
— Cameron, 1932: 131 (Philonthus; aberration of varians).
— Scheerpeltz, 1933: 1366 (Philonthus; synonym of piceicoxis).
— Blackwelder, 1943: 412 (Philonthus; synonym of varians).
— Smetana, 1958: 195 (Philonthus; synonym of varians).
— Smetana, 1959: 160 (Philonthus; synonym of varians).
— Coiffait, 1974: 244 (Philonthus; synonym of varians).
— Smetana, 1995e: 235 (Philonthus; synonym of varians).
fuscicoxis Scheerpeltz, 1933: 1366 (Philonthus; [Note: Nomen nudum since it is a replacement for an aberration (Articles 1.3.4, 45.5)]; replacement name for piceicoxis Gridelli).
— Blackwelder, 1943: 412 (Philonthus; synonym of varians).
— Smetana, 1958: 196 (Philonthus; synonym of varians).
— Smetana, 1959: 160 (Philonthus; aberration of varians; characters).
— Coiffait, 1967a: 419 (Philonthus; aberration of varians; characters).
— Coiffait, 1974: 244 (Philonthus; variety of varians).
— Tóth, 1984: 71 (Philonthus; cited as fuscicornis; aberration of varians).
— Smetana, 1995e: 235 (Philonthus; synonym of varians).
varicolor Boheman, 1858, see: hepaticus Erichson, 1840.
variegatus Erichson, 1840, see: Cafius.
variicolor Nordmann, 1837, see: Quedius.
variipennis Kraatz, 1859: 85 (Philonthus; Type locality: Ceylan; India orientalis).
— Fauvel, 1886b: 150 (Philonthus; variety of quisquiliarius; Philippines; Ceylon; East Indies; Cochinchine; Annam; Sumatra; Java; Celebes).
— Bernhauer, 1902b: 27 (Philonthus; Ceylon).
— Fauvel, 1903: 159 (Philonthus; synonym of quisquiliarius variety inquinatus).
— Fauvel, 1904a: 59 (Philonthus; synonym of quisquiliarius variety inquinatus).
— Bernhauer and Schubert, 1914: 353 (Philonthus; synonym of punctus).
— Tottenham, 1953: 143 (Philonthus; characters; distinct from quisquiliarius).
— Last, 1961: 306 (Philonthus; Thailand).
— Coiffait, 1967a: 424 (Philonthus; characters; Southeast Asia; India; China).
— Coiffait, 1974: 201 (Philonthus; characters; India; Cambodia).
— Watanabe, 1989a: 56 (Philonthus; Japan).
— Barrion, Litsinger, and Last, 1991: 1078 (Philonthus; associated with rice agroecosystem).
— Shibata, 1994a: 106 (Philonthus; characters; Taiwan).
▴ Distribution: Saudi Arabia, Afghanistan, Sri Lanka, India, Thailand, Cambodia, Vietnam, Indonesia, Philippines, China, Taiwan, Japan.
vacans Tottenham, 1953: 145 (Philonthus; variety of variipennis; Type locality: Penang. Also cited from Siam).
— Last, 1961: 306 (Philonthus; variety of variipennis; Thailand; Cambodia).
— Last, 1964: 177 (Philonthus; variety of variipennis; Thailand).
— Coiffait, 1974: 201 (Philonthus; variety of variipennis; characters).
— Barrion, Litsinger, and Last, 1991: 1078 (Philonthus; subspecies of variipennis; associated with rice agroecosystem).
ilus Tottenham, 1953: 145 (Philonthus; variety of variipennis; Type locality: Philippine Islands: Manila).
— Last, 1961: 306 (Philonthus; variety of variipennis; Thailand).
— Coiffait, 1974: 201 (Philonthus; variety of variipennis; characters).
ledouxi Coiffait, 1979d: 558 (Philonthus; Type locality: Nouristan: route de Djelalabad, à Kaboul, gorges du Chikari).
— Coiffait, 1983e: 477 (Philonthus; Afghanistan).
— Schillhammer, 1996a: 82 (Philonthus; synonym of variipennis).
arabicus Coiffait, 1981b: 238 (Philonthus; subspecies of ledouxi; [preoccupied]; Type locality: Arabie Saoudite: Hofuf, 85 m).
falcozi Coiffait, 1987: 497 (Philonthus; replacement name for arabicus Coiffait).
varipennis Scriba, 1864, see: Bisnius.
variolosus Sharp, 1871, see: Cafius.
varipes Mulsant and Rey, 1861: 142 [= 1862: 126] (Philonthus; Type locality: Montagnes du Bourbonnais, de l'Auvergne et de la Provence).
— Fauvel, 1869: 489 (Philonthus; synonym of fulvipes).
— Fauvel, 1874: 259 [= 1874a: 485] (Philonthus; synonym of fulvipes).
— Ganglbauer, 1895: 457 (Philonthus; synonym of fulvipes).
— Bernhauer and Schubert, 1914: 339 (Philonthus; variety of fulvipes).
— Scheerpeltz, 1933: 1343 (Philonthus; variety of fulvipes).
— Smetana, 1955: 210 (Philonthus; variety of fulvipes; notes).
— Smetana, 1966b: 47 (Philonthus; cited as valid species; characters; England).
— Coiffait, 1974: 92 (Paragabrius; variety of fulvipes).
— Pope, 1977: 30 (Philonthus; Britain).
— Hodge and Jones, 1995: 40 (Philonthus; characters; Britain).
▴ Distribution: France, England.
varius Gyllenhal, 1810, see: carbonarius Gravenhorst, 1802.
varro Smetana, 1995e: 293 (Philonthus; Type locality: Quebec: Hwy. 148, 15 km E. Lachute. Also cited from Manitoba, New Brunswick, Connecticut, Maine, Massachusetts, New Hampshire, New York).
vegetus Last, 1968: 363 (Philonthus; Type locality: New Guinea: Okapa).
— Last 1981: 120 (Philonthus; characters; New Guinea).
— Last, 1987: 32 (Philonthus; Papua New Guinea).
▴ Distribution: New Guinea.
velatipennis Solsky, 1870: 461 (Philonthus; Type locality: environs de la ville d'Astrakhan).
— Solsky, 1874: 183 (Philonthus; characters; Turkestan).
— Bernhauer and Schubert, 1914: 359 (Philonthus; catalog).
— Scheerpeltz, 1933: 1367 (Philonthus; catalog).
— Coiffait, 1967a: 432 (Philonthus; characters; Russia; Caucasus; Turkestan).
— Jarrige, 1971: 494 (Philonthus; Iran).
— Coiffait, 1974: 210 (Philonthus; characters; Russia; Turkestan; Transcaspian region; Persia).
— Petrenko, 1980: 82 (Philonthus; Caucasus).
▴ Distribution: Russia, Turkey, Iran, “Turkestan”.
tectipennis Coiffait, 1967a: 432 (Philonthus; subgenus Philonthus; variety of velatipennis; [Note: Nomen nudum (Article 15.2)]).
— Coiffait, 1974: 210 (Philonthus; variety of velatipennis; [Note: Cited as a new variety but is still a nomen nudum]; characters; Type locality cited as Turkestan: Tschimagad).
extensus Korge, 1971a: 7 (Philonthus; aberration of velatipennis; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Kleinasien: Igdir nördlich des Ararat).
ventralis Gravenhorst, 1802: 174 (Staphylinus; Type locality: Lusitania; [Note: See lectotype designation by Smetana, 1995e]).
— Latreille, 1804: 324 (Staphylinus; characters; Portugal).
— Gravenhorst, 1806: 60 (Staphylinus).
— Gyllenhal, 1810: 334 (Staphylinus; characters; Sweden).
— C. Sahlberg, 1830: 325 (Staphylinus; characters; Finland).
— Mannerheim, 1830: 29 (Staphylinus; Finland; Sweden).
— Mannerheim, 1831: 443 (Staphylinus; Finland; Sweden).
— Stephens, 1832: 219 (Quedius; [Note: Misidentification: Bernhauer and Schubert, 1914: 360 cited fulvipes Stephens (1832: 219) as a synonym of ventralis Gravenhorst; Stephens did not describe the species as new, he attributed it to Fabricius]; characters; England).
— Stephens, 1832: 237 (Philonthus; characters; England).
— Ménétriés, 1832: 146 (Staphylinus; Caucasus-Persia region).
— Lacordaire, 1835: 407 (Staphylinus; characters; France).
— Curtis, 1836: pl. 610 (Philonthus; characters; Britain).
— Nordmann, 1837: 93 (Philonthus; Sweden; Germany; Russia).
— Erichson, 1839a: 468 (Philonthus; characters; Germany).
— Heer, 1839: 267 (Philonthus; characters; Switzerland).
— Stephens, 1839: 397 (Philonthus; characters; England).
— Erichson, 1840: 473 (Philonthus; characters; Europe; America septentrionalis; St. Vincent).
— Kolenati, 1846: 20 (Philonthus; Tanain; trans Caucasum).
— Jacobsen, 1849: 699 (Philonthus; characters; Denmark).
— Hochhuth, 1849: 142 (Philonthus; characters; Tiflis).
— Redtenbacher, 1849: 705 (Philonthus; characters; Austria).
— Küster, 1854: 11 (Philonthus; characters; Europe).
— Hardy, 1851: 30 (Philonthus; notes; England).
— Fairmaire and Laboulbène, 1856: 523 (Philonthus; characters; France).
— Kraatz, 1857d: 605 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 195 (Philonthus; characters; Austria).
— Kraatz, 1858a: 61 (Philonthus; Greece).
— Peyron, 1858: 425 (Philonthus; Turkey).
— Thomson, 1860: 160 (Philonthus; characters; Scandinavia).
— Hochhuth, 1862: 61 (Philonthus; Russia).
— Wollaston, 1867: 238 (Philonthus; characters; notes; Cape Verde Islands).
— Solsky, 1872: 303 (Philonthus; Cayenne).
— Fauvel, 1874: 227 [= 1874a: 453] (Philonthus; characters; France; Algeria; Cyprus; Turkey; Caucasus; Transcausia; Siberia; Madeira, Canary, and Cape Verde Islands; North America; Antilles; Guyana).
— Redtenbacher, 1874: 214 (Philonthus; characters; Austria).
— Seidlitz, 1875: 276 (Philonthus; characters; Baltic region).
— Fauvel, 1876a: 73 [= 1876: 258] (Philonthus; Tunisia; Azores; Cuba).
— Mulsant and Rey, 1876: 481 [= 1877: 337] (Philonthus; characters; larval characters; France).
— J. Sahlberg, 1876: 18 (Philonthus; Finland).
— Provancher, 1877: 250 (Philonthus; characters; Quebec).
— Fauvel, 1878: 123 [= 1878a: 43] (Philonthus; Algeria; Tunisia).
— Horn, 1884: 211 (Philonthus; characters; Michigan; Missouri; Nevada).
— Fauvel, 1886: 54 [= 1886a: 46] (Philonthus; Morocco; Tunisia; Algeria).
— Blackburn, 1888b: 48 (Philonthus; Australia).
— Fowler, 1888: 274 (Philonthus; characters; Britain).
— Fleutiaux and Sallé, 1889: 380 (Philonthus; Guadeloupe Island).
— Fauvel, 1889b: 265 (Philonthus; New Caledonia).
— Ganglbauer, 1895: 455 (Philonthus; characters; Palaearctic region; North and South America).
— Fauvel, 1902b: 109 (Philonthus; Barbary; Egypt; Azores; Madeira; Canaries).
— Fauvel, 1903d: 270 (Philonthus; New Caledonia).
— Fauvel, 1903e: 241 (Philonthus; Cameroon; Subcosmopolitan).
— Sainte-Claire Deville, 1906: 96 (Philonthus; Corsica).
— Ramos, 1946: 33 (Philonthus; Mona Island, West Indies).
— Reitter, 1909: 129 (Philonthus; characters; Germany).
— Blatchley, 1910: 388 (Philonthus; characters; Indiana).
— Bernhauer and Schubert, 1914: 360 (Philonthus; catalog).
— Johansen, 1914: 410 (Philonthus; characters; Denmark).
— Lea, 1925: 220 (Philonthus; Australian introduction).
— Rambousek, 1922: 1010 (Philonthus; Ethiopia).
— Bernhauer, 1928: 16 (Philonthus; Sumatra).
— Portevin, 1929: 362 (Philonthus; characters; France).
— Cameron, 1932: 124 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1367 (Philonthus; catalog).
— Cameron, 1933d: 387 (Philonthus; Sumatra).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Bernhauer, 1942c: 361 (Philonthus; Cameroon).
— Blackwelder, 1943: 404 (Philonthus; characters; Cuba; Jamaica; Dominican Republic; Puerto Rico; Culebra Island; St. Thomas; St. Croix; St. Kitts; Antigua; Montserrat; Guadeloupe; Dominica; St. Lucia; Barbados; St. Vincent; Grenada; Tobago; Trinidad).
— Blackwelder, 1947: 123 (Philonthus; Cayman Islands).
— V. Hansen, 1952: 74 (Philonthus; subgenus Philonthus; characters; Denmark).
— Tottenham, 1956: 226 (Philonthus; Ruanda).
— Scheerpeltz, 1956c: 1101 (Philonthus; Cyprus).
— Smetana, 1958: 211 (Philonthus; subgenus Philonthus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 161, 166 (Philonthus; characters).
— Tottenham, 1961: 204 (Philonthus; Senegal).
— Smetana, 1963b: 36 (Philonthus; subgenus Philonthus; Madeira).
— Palm, 1963b: 91 (Philonthus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 184 (Philonthus; subgenus Philonthus; characters; central Europe).
— Horion, 1965: 166 (Philonthus; widespread in southern and central Europe, sporadic in northern Europe).
— Levasseur, 1966a: 1213 (Philonthus; Congo).
— Coiffait, 1967a: 422 (Philonthus; characters).
— Levasseur, 1968a: 53 (Philonthus; Central African Republic).
— Szujecki, 1969a: 256 (Philonthus; notes; Poland).
— Scheerpeltz, 1971c: 170 (Philonthus; Cameroon).
— Moore and Legner, 1974: 39 (Philonthus; spread by commerce).
— Coiffait, 1974: 217 (Philonthus; characters; Europe; Britain; Scandinavia; north Africa).
— Pope, 1977: 30 (Philonthus; Britain).
— Smetana, 1978: 117 (Philonthus; collecting notes; Israel).
— Frank, 1982: 44 (Philonthus; list of parasites).
— Frank, 1984: 479 (Philonthus; notes; characters; Haiti; Costa Rica; Florida).
— Tóth, 1984: 73 (Philonthus; characters; Hungary).
— Lucht, 1987: 105 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Welch, 1993: 228 (Philonthus; ovariole number and ovary structure).
— Smetana, 1993b: 48 (Philonthus; subgenus Philonthus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Philonthus; Italy).
— Smetana, 1995e: 190 (Philonthus; lectotype designation; characters; Ontario; Quebec; Arizona; California; Florida; Illinois; Iowa; Kansas; Maryland; Massachusetts; Nevada; New Hampshire; New Jersey; New York; North Carolina; Pennsylvania; South Carolina; Tennessee; Texas; Washington; West Virginia).
— Hu and Frank, 1995b: 585 (Philonthus; surface structure of eggs).
— Downie and Arnett, 1996: 381 (Philonthus; characters; USA).
— Hu and Frank, 1997 (Philonthus; predation of eggs and larvae of Haematobia irritans; Florida).
▴ Distribution: Cosmopolitan.
anthrax Gravenhorst, 1802: 176 (Staphylinus; Type locality: America septentr.; [Note: See lectotype designation by Smetana, 1995e]).
— Gravenhorst, 1806: 67 (Staphylinus).
— Nordmann, 1837: 97 (Philonthus; America septentrionalis).
— Erichson, 1840: 473 (Philonthus; synonym of ventralis).
— Kolenati, 1846: 20 (Philonthus; synonym of ventralis).
— Küster, 1854: 11 (Philonthus; synonym of ventralis).
— Kraatz, 1857d: 605 (Philonthus; synonym of ventralis).
— Fleutiaux and Sallé, 1889: 380 (Philonthus; synonym of ventralis).
— Ganglbauer, 1895: 455 (Philonthus; synonym of ventralis).
— Bernhauer and Schubert, 1914: 360 (Philonthus; synonym of ventralis).
— Cameron, 1932: 124 (Philonthus; synonym of ventralis).
— Blackwelder, 1943: 404 (Philonthus; synonym of ventralis).
— Smetana, 1958: 211 (Philonthus; synonym of ventralis).
— Smetana, 1995e: 190 (Philonthus; lectotype designation; synonym of ventralis).
immundus Gravenhorst, 1806: 66 (Staphylinus; Type locality: Parisiae).
— Erichson, 1839a: 468 (Philonthus; synonym of ventralis).
— Erichson, 1840: 473 (Philonthus; synonym of ventralis).
— Kolenati, 1846: 20 (Philonthus; synonym of ventralis).
— Küster, 1854: 11 (Philonthus; synonym of ventralis).
— Kraatz, 1857d: 605 (Philonthus; synonym of ventralis).
— J. Sahlberg, 1876: 18 (Philonthus; synonym of ventralis).
— Fleutiaux and Sallé, 1889: 380 (Philonthus; synonym of ventralis).
— Ganglbauer, 1895: 455 (Philonthus; synonym of ventralis).
— Bernhauer and Schubert, 1914: 360 (Philonthus; synonym of ventralis).
— Cameron, 1932: 124 (Philonthus; synonym of ventralis).
— Blackwelder, 1943: 404 (Philonthus; synonym of ventralis).
— Smetana, 1958: 211 (Philonthus; synonym of ventralis).
— Coiffait, 1974: 217 (Philonthus; synonym of ventralis).
— Smetana, 1995e: 190 (Philonthus; synonym of ventralis).
celer Gravenhorst, 1806: 66 (Staphylinus; Type locality: Parisiae).
— Heer, 1839: 266 (Staphylinus; [Note: Bernhauer and Schubert, 1914: 334 attributed celer to Heer as a synonym of debilis but Heer did not describe the species as new, he attributed it to Gravenhorst]; characters).
— Nordmann, 1837: 94 (Philonthus; France; Germany).
— Erichson, 1840: 480 (Philonthus; characters; France).
— Kraatz, 1857d: 596 (Philonthus).
— Fauvel, 1874: 227 [= 1874a: 453] (Philonthus; synonym of ventralis).
— Ganglbauer, 1895: 453 (Philonthus; synonym of debilis).
— Bernhauer and Schubert, 1914: 334 (Philonthus; synonym of debilis).
— Smetana, 1958: 211 (Philonthus; synonym of ventralis).
— Coiffait, 1974: 217 (Philonthus; synonym of ventralis).
— Smetana, 1995e: 190 (Philonthus; synonym of ventralis).
picicollis Stephens, 1832: 224 (Quedius; Type locality: Norfolk; Suffolk; Devonshire).
— Stephens, 1839: 392 (Quedius; characters; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of ventralis).
— Fleutiaux and Sallé, 1889: 380 (Philonthus; synonym of ventralis).
— Bernhauer and Schubert, 1914: 360 (Philonthus; synonym of ventralis).
— Cameron, 1932: 124 (Philonthus; synonym of ventralis).
— Blackwelder, 1943: 404 (Philonthus; synonym of ventralis).
— Smetana, 1958: 212 (Philonthus; synonym of ventralis).
— Coiffait, 1974: 217 (Philonthus; synonym of ventralis).
— Smetana, 1995e: 190 (Philonthus; synonym of ventralis).
simplex Stephens, 1833: 248 (Bisnius; [preoccupied]; Type locality: Devonshire).
— Stephens, 1839: 399 (Bisnius; characters; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of ventralis).
fuscicornis Stephens, 1833: 248 (Bisnius; Type locality: Suffolk).
— Stephens, 1839: 399 (Bisnius; characters; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of ventralis).
fulvipes Stephens, 1833: 249 (Bisnius; [preoccupied]; Type locality: London).
— Stephens, 1839: 399 (Bisnius; characters; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of ventralis).
— Fleutiaux and Sallé, 1889: 380 (Philonthus; synonym of ventralis).
— Cameron, 1932: 124 (Philonthus; synonym of ventralis).
— Blackwelder, 1943: 404 (Philonthus; synonym of ventralis).
— Smetana, 1958: 212 (Philonthus; synonym of ventralis).
— Coiffait, 1974: 217 (Philonthus; synonym of ventralis).
— Smetana, 1995e: 190 (Philonthus; synonym of ventralis).
rotundiceps Stephens, 1833: 248 (Bisnius; Type locality: Norfolk; Suffolk).
— Stephens, 1839: 399 (Bisnius; characters; England).
— Hardy, 1851: 28 (Philonthus; notes; England).
— G. Waterhouse, 1858: 25 (Philonthus; synonym of ventralis).
— Fleutiaux and Sallé, 1889: 380 (Philonthus; synonym of ventralis).
— Bernhauer and Schubert, 1914: 360 (Philonthus; synonym of ventralis).
— Cameron, 1932: 124 (Philonthus; synonym of ventralis).
— Blackwelder, 1943: 404 (Philonthus; synonym of ventralis).
— Smetana, 1958: 212 (Philonthus; synonym of ventralis).
— Coiffait, 1974: 217 (Philonthus; synonym of ventralis).
— Smetana, 1995e: 190 (Philonthus; synonym of ventralis).
proximus Wollaston, 1857: 189 (Philonthus; Type locality: Madeira: near Funchal; Gorgulho; Praya Formoza. Porto Santo).
— Wollaston, 1864: 573 (Philonthus; characters; Canary Islands).
— Wollaston, 1865: 493 (Philonthus; habitat; notes; Madeira Islands; Canary Islands).
— Crotch, 1867a: 383 (Philonthus; Azores).
— Wollaston, 1867: 238 (Philonthus; synonym of ventralis).
— Wollaston, 1867: 279 (Philonthus; synonym of ventralis).
— Fauvel, 1869: 489 (Philonthus; synonym of ventralis).
— Fauvel, 1869a: 151 (Philonthus; synonym of ventralis).
— Wollaston, 1871: 303 (Philonthus; synonym of ventralis).
— Fauvel, 1902b: 109 (Philonthus; synonym of ventralis).
— Bernhauer and Schubert, 1914: 360 (Philonthus; variety of ventralis).
— Bernhauer, 1940c: 4 (Philonthus; variety of ventralis; Azores).
— Méquignon, 1942a: 21 (Philonthus; subspecies of ventralis; Azores).
— Blackwelder, 1943: 404 (Philonthus; synonym of ventralis).
— Borges, 1990: Table II (Philonthus; subspecies of ventralis; Azores).
fortunatus Wollaston, 1865: 493 (Philonthus; variety of proximus; Type locality: Teneriffe).
— Fauvel, 1902b: 110 (Philonthus; synonym of debilis).
— Bernhauer and Schubert, 1914: 360 (Philonthus; variety of ventralis).
— Blackwelder, 1943: 404 (Philonthus; synonym of ventralis).
venustulus Last, 1968, see: Hesperus.
venustus Last, 1968, see: Hesperus.
veronikae Hromádka, 1992: 101 (Philonthus; Type locality: Nepal, Khudi).
versicolor Cameron, 1919, see: eustilbus Kraatz, 1859.
versutus Horn, 1884, see: Bisnius.
verticalis Tottenham, 1956: 287 (Philonthus; Type locality: Ruanda: Rutovu, forêt du Rugege, 2350 m).
vertumnus Herman, 2001: 48 (Philonthus; replacement name for planus Last).
planus Last, 1987: 27 (Philonthus; [preoccupied]; Type locality: Papua New Guinea: Gazelle Peninsula, Upper Warangoi, 1250 m).
vestigator Tottenham, 1955: 162 (Philonthus; subgenus Spatulonthus; Type locality: Kenya: Ngonga Hills).
— Tottenham, 1962: 140 (Philonthus; subgenus Spatulonthus; Tanganyika).
▴ Distribution: Kenya, Tanzania.
vestitus Sharp, 1874, see: Cafius.
vesubiensis Coiffait, 1967a: 447 (Philonthus; subgenus Kenonthus; Type locality: Balme de la Frema, 2400 m, Alpes Maritimes).
— Jarrige, 1968b: 74 (Philonthus; France).
— Coiffait, 1974: 300 (Philonthus; subgenus Kenonthus; characters; Alps).
▴ Distribution: France.
veteratorius Broun, 1880: 112 (Philonthus; Type locality: Whangarei Harbour).
vicinus Cameron, 1932, see: Gabrius.
vicinus Cameron, 1933, see: renominatus Cameron, 1937.
victoriensis Lea, 1925: 224 (Philonthus; Type locality: Victoria: Melbourne).
viduus Erichson, 1840, see: Styngetus.
viduus Cameron, 1933, see: Gabrius.
viettei Jarrige, 1970: 50 (Philonthus; Type locality: Madagascar: Mt. Tsaratanana, matsabory en dessous de l'Andohanisambirano, 2050 m).
— Lecoq, 1998: 67 (Philonthus; subgenus Piezarthrus; characters; Madagascar).
▴ Distribution: Madagascar.
vilis Erichson, 1840, see: hepaticus Erichson, 1840.
villiersi Bernhauer, 1942c: 362 (Philonthus; Type locality: Mont Cameroun, 1800–2000 m).
vinohradensis R. Dvořák and Havelka, 1952, see: carbonarius Gravenhorst, 1802.
vinsoni Cameron, 1947, see: Erichsonius.
violaceus Fauvel, 1878f: 265 (Philonthus; Type locality: Ternate, Acquiconora).
— Fauvel, 1879: 103 (Philonthus; Batcian).
— Bernhauer and Schubert, 1914: 360 (Philonthus; catalog).
— Cameron, 1937a: 100 (Philonthus; New Guinea).
— Last, 1987: 32 (Philonthus; Papua New Guinea).
▴ Distribution: New Guinea, Indonesia.
viperinus Horn, 1884, see: aequalis Horn, 1884.
virgatus Sharp, 1889: 38 (Philonthus; Type locality: Nikko; Chiuzenji).
— Bernhauer and Schubert, 1914: 360 (Philonthus; catalog).
— Scheerpeltz, 1933: 1367 (Philonthus; catalog).
— J. Li, 1993: 33 (Philonthus; China).
— Kanehisa, Tsumuki, and Kawazu, 1994 (Philonthus; cited as virugatu; chemical secretions).
— Schillhammer, 1999a: 62 (Philonthus).
▴ Distribution: Japan, China, Russia.
nigrolineatus Bernhauer, 1913: 130 (Philonthus; Type locality: Ostsibirien: Schipka-Gora, Sotka-Gora).
— Bernhauer and Schubert, 1914: 348 (Philonthus; catalog).
— Scheerpeltz, 1933: 1353 (Philonthus; catalog).
— Coiffait, 1974: 316 (Spatulonthus; characters; Siberia).
— Boháč, 1988: 552 [= 1989: 35] (Spatulonthus; characters; Russia).
— Schillhammer, 1999a: 62 (Philonthus; synonym of virgatus).
ussuriensis Coiffait, 1974: 202 (Philonthus; Type locality: Shkotovo, Peishoula).
virgo Gravenhorst, 1802: 169 (Staphylinus; Type locality: Lusitania).
— Latreille, 1804: 323 (Staphylinus; characters; Portugal).
— Gravenhorst, 1806: 69 (Staphylinus).
— Gyllenhal, 1810: 345 (Staphylinus; characters; illustration).
— Zetterstedt, 1828: 78 (Staphylinus; characters; Lapland).
— Mannerheim, 1830: 31 (Staphylinus; Finland).
— Mannerheim, 1831: 445 (Staphylinus; Finland).
— C. Sahlberg, 1830: 330 (Staphylinus; characters; Finland).
— Nordmann, 1837: 101 (Philonthus; Germany; Sweden; Russia).
— Zetterstedt, 1838: 65 (Staphylinus; characters; habitat; Lapland).
— Heer, 1839: 269 (Philonthus; characters; Switzerland).
— Erichson, 1840: 483 (Philonthus; characters; Portugal; Sardinia; Sweden).
— Laporte, 1840: 178 (Staphylinus; characters; France).
— Jacobsen, 1849: 702 (Philonthus; characters; Denmark).
— Fairmaire and Laboulbène, 1856: 526 (Philonthus; characters; France).
— Kraatz, 1857d: 611 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 198 (Philonthus; characters; Germany).
— Thomson, 1860: 165 (Philonthus; characters; Scandinavia).
— Fauvel, 1874: 260 [= 1874a: 486] (Philonthus; characters; France; Lapland; Scandinavia; Germany; Switzerland; Portugal; Corsica; Sardinia; Algeria).
— Redtenbacher, 1874: 217 (Philonthus; characters; Germany).
— Mulsant and Rey, 1876: 501 [= 1877: 357] (Philonthus; characters; France).
— J. Sahlberg, 1876: 21 (Philonthus; Finland).
— Fauvel, 1878: 126 [= 1878a: 46] (Philonthus; Algeria).
— Fauvel, 1886: 57 [= 1886a: 49] (Philonthus; Algeria).
— Ganglbauer, 1895: 456 (Philonthus; characters; Europe; Algeria).
— Fauvel, 1902b: 113 (Philonthus; Morocco; Algeria; Tunisia).
— Sainte-Claire Deville, 1906: 99 (Philonthus; Corsica).
— Poppius, 1909: 26 (Philonthus; subgenus Gefyrobius; Lena Valley, Siberia).
— Reitter, 1909: 130 (Philonthus; characters; Germany).
— Bernhauer and Schubert, 1914: 360 (Philonthus; catalog).
— Johansen, 1914: 412 (Philonthus; characters; Denmark).
— Portevin, 1929: 364 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1367 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— Jarrige, 1951a: 7 (Philonthus; characters).
— Coiffait, 1963: 12 (Paragabrius).
— Focarile, 1964: 59 (Philonthus; Italy).
— Coiffait, 1973a: 274 (Paragabrius; Morocco).
— Coiffait, 1974: 95 (Paragabrius; characters; Portugal; Morocco).
— Shilov, 1975a: 376 [= 1975b: 98] (Philonthus; Komi region of Russia).
— Tóth, 1984: 76 (Philonthus; characters).
— Maachi, 1991: 251 (Paragabrius; Morocco).
— Outerelo, Gamarra, and Salgado, 1998:129 (Paragabrius; subgenus Paragabrius; collected from cave; Spain; Portugal).
▴ Distribution: Tunisia, Algeria, Morocco, Europe.
viridanus Horn, 1884, see: gracilior Casey, 1915.
viridanus Nordmann, 1837, see: laminatus Creutzer, 1799.
viridiniger Bernhauer, 1916c: 30 (Philonthus; Type locality: West-Columbien: Umgebung von Cali im Caucatale).
viridipennis Fauvel, 1875a: XXX [= 1875b: 232] (Philonthus; Type locality: Syrie, Tibériade, Bahr-el-Houleh).
— Bernhauer and Schubert, 1914: 361 (Philonthus; catalog).
— Scheerpeltz, 1933: 1368 (Philonthus; catalog).
— Coiffait, 1967a: 425 (Philonthus; characters; Syria; Lebanon; Palestine; Caucasus; central Europe; [Note: The last two regions cited referred to P. oebalus Tottenham which Coiffait listed as a junior synonym of P. viridipennis]).
— Coiffait, 1974: 202 (Philonthus; characters; Palestine; Syria; Caucasus; Europe; [Note: The last two regions cited referred to P. oebalus Tottenham which Coiffait listed as a junior synonym of P. viridipennis]).
— Tóth, 1984: 74 (Philonthus; characters; Hungary).
▴ Distribution: Syria, Lebanon, Israel.
viridovix Smetana, 1995e: 397 (Philonthus; Type locality: Arkansas: Newton Co.: Buffalo NR, Leatherwood Cr. Also cited from Illinois).
virilis Horn, 1884, see: Gabrius.
visokeanus Cameron, 1950, see: Gabrius.
vitochensis Coiffait, 1974, see: frigidus Märkel and Kiesenwetter, 1848.
vittatus Roth, 1851, see: natalensis Boheman, 1848.
vittiger Fauvel, 1907: 40 (Philonthus; Type locality: Escarpment).
— Bernhauer and Schubert, 1914: 361 (Philonthus; catalog).
— Tottenham, 1962: 136 (Philonthus; Tanganyika).
▴ Distribution: Kenya, Tanzania.
pseudovittiger Bernhauer, 1939a: 85 (Philonthus; variety of vittiger; Type locality: Kenya:
Mt. Elgon, Osthang, Suam fishing hut, 2400 m).
vividus Broun, 1880, see: Quedius.
vividus Cameron, 1931: 360 (Hesperus; [Note: The name is a secondary homonym in Philonthus; Cameron, 1937a: 101 stated that in his original description, because of a printer's error, this species was erroneously cited in Hesperus rather than Philonthus]; Type locality: Mount Lamington, New Guinea).
— Scheerpeltz, 1933: 1374 (Hesperus; catalog).
— Cameron, 1937a: 101 (Philonthus; New Guinea).
▴ Distribution: New Guinea.
volvulus Bernhauer, 1932b: 151 (Philonthus; Type locality: Haut-Uele: Yebo Moto).
— Scheerpeltz, 1933: 1368 (Philonthus; catalog).
— Bernhauer, 1947: 158 (Philonthus; Deutsch-Ostafrika).
— Tottenham, 1949b: 321 (Philonthus; characters).
— Scheerpeltz, 1974c: 22 (Philonthus; Sudan).
▴ Distribution: Sudan, Congo, Tanzania.
vulcanus Cameron, 1950, see: Gabrius.
vulgatus Casey, 1915: 442 (Philonthus; Type locality: Minnesota: Duluth; [Note: See lectotype designation by Smetana, 1995e]. Also cited from Canada: Ottawa; New York: Lake Champlain and Ithaca;).
— Scheerpeltz, 1933: 1368 (Philonthus; catalog).
— Smetana, 1965: 28 (Philonthus; subgenus Philonthus; characters; Nova Scotia).
— Smetana, 1965a: 8 (Philonthus; Alaska).
— Dearborn and Donahue, 1994: 15 (Philonthus; Maine).
— Smetana, 1995e: 456 (Philonthus; lectotype designation: Minnesota, Duluth; characters; notes; Alberta; British Columbia; Manitoba; Nova Scotia; Ontario; Quebec; Saskatchewan; Alaska; Colorado; Idaho; Illinois; Indiana; Iowa; Kansas; Maine; Massachusetts; Michigan; Minnesota; Montana; Nebraska; New Hampshire; New York; North Dakota; South Dakota; Utah; Washington; Wisconsin).
— Downie and Arnett, 1996: 382 (Philonthus; characters; USA).
▴ Distribution: Canada; USA.
wacoensis Casey, 1915, see: flavolimbatus Erichson, 1840.
wagneri Coiffait, 1963 [Note: Nomen nudum], see: alcyoneus Erichson, 1840.
waterhousei Cameron, 1922: 120 (Philonthus; Type locality: St. Vincent; Grenada).
— Scheerpeltz, 1933: 1368 (Philonthus; catalog).
— Blackwelder, 1943: 417 (Philonthus; characters; St. Vincent; Grenada).
▴ Distribution: West Indies.
watsoni Stephens, 1832, see: Bisnius.
wawawai Hatch, 1957, see: Gabrius.
weiseri Bernhauer, 1921c: 169 (Philonthus; Type locality: Argentinien: bei Jujuy, Cueva Iturbe, 3700 m; Rio Colorado).
weiserianus Bernhauer, 1921c: 106 (Philonthus; Type locality: Argentinien: Prov. Jujuy, Hornadita, 3400 m).
whymperi Sharp, 1891: 40 (Philonthus; Type locality: Ecuador: Hacienda of Gauchala, 9217 feet; Antisanilla to Piñantura, 11,000 feet; Machachi, 9–10,000 feet).
witteanus Cameron, 1950g: 34 (Philonthus; Type locality: Belgian Congo: Rutshuru, alt. 1285 m).
wittei Bernhauer, 1932b: 149 (Philonthus; Type locality: Katanga: Kansenia).
wittmeri Coiffait, 1977b: 225 (Philonthus; Type locality: Bhutan: Kotoka-Gogona, 2600–3400 m).
wittmeri Coiffait, 1982, see: Anisolinus.
wohlgrothi R. Dvořák, 1952 [Note: Nomen nudum], see: succicola Thomson, 1860.
wollastoni Scheerpeltz, 1933: 1368 (Philonthus; replacement name for punctipennis Wollaston).
— Tottenham, 1956a: 241 (Philonthus; synonym of turbidus).
— Levasseur, 1966a: 1214 (Philonthus; subspecies of turbidus; Congo; Cape Verde; Sudan; Somalia; Nigeria; Madagascar).
— Coiffait, 1967a: 431 (Philonthus; cited as valid species; characters; Madeira; Canary Islands; Cape Verde Islands).
— Coiffait, 1973a: 274 (Philonthus; Morocco).
— Coiffait, 1974: 207 (Philonthus; characters; Cape Verde Islands; Canary Islands; Madeira; intertropical Africa).
— Scheerpeltz, 1974c: 21 (Philonthus; characters; Sudan).
▴ Distribution: Madeira, Canary Islands, Cape Verde Islands, Morocco, Algeria, Nigeria, Sudan, Congo, Somalia, Madagascar.
punctipennis Wollaston, 1857: 192 (Philonthus; [Note: Preoccupied secondary homonym when punctipennis Solier was in Philonthus]; Type locality: Madeira: Santa Cruz).
— Wollaston, 1864: 575 (Philonthus; characters; Canary Islands).
— Wollaston, 1865: 495 (Philonthus; habitat; Madeira Islands; Canary Islands).
— Wollaston, 1867: 240 (Philonthus; synonym of turbidus).
— Wollaston, 1871: 303 (Philonthus; notes; synonym of turbidus).
— Fauvel, 1875a: XXXI [= 1875b: 233] (Philonthus; synonym of turbidus).
— Fauvel, 1902b: 110 (Philonthus; synonym of turbidus).
— Bernhauer and Schubert, 1914: 358 (Philonthus; synonym of turbidus).
— Gridelli, 1930a: 92 (Philonthus; [Note: According to Tottenham (1956a: 243) Gridelli's identification of punctipennis may have been erroneous and it may have been tumulinus]; characters; Cape Verde).
— Peyerimhoff, 1931: 32 (Philonthus; Algeria).
— Cameron, 1932: 141 (Philonthus; synonym of turbidus).
— Tottenham, 1956a: 241 (Philonthus; synonym of turbidus).
— Coiffait, 1974: 207 (Philonthus; synonym of wollastoni).
— Scheerpeltz, 1974c: 21 (Philonthus; synonym of wollastoni).
woodburnensis Hatch, 1957, see: spiniformis Hatch, 1957.
wuesthoffi Bernhauer, 1939d: 98 (Philonthus; Type locality: Japan: Yokkaichi, Meeresstrand. Also cited from Korea and China).
— Wüsthoff, 1937a: 347 (Philonthus; illustration of aedeagus; Manchuria).
— Watanabe, 1992 (Philonthus; Okushiri-tô Island).
— Watanabe and Shibata, 1972: 64 (Philonthus; Yaku-shima Island).
— Frank, 1982: 44 (Philonthus; list of parasites).
— Yuh, Paik, Kwon, and Lee, 1985: 242 (Philonthus; Korea).
— J. Li, 1993: 33 (Philonthus; China).
— Watanabe, 1998: 313 (Philonthus; Japan).
— Schillhammer, 1999a: 62 (Philonthus).
▴ Distribution: Japan, Russia, China, Korea.
duplicatus Tikhomirova, 1973: 167 (Philonthus; [preoccupied]; Type locality: Khasanskii raion).
fagelianus Tikhomirova, 1973a: 184 (Philonthus; replacement name for duplicatus Tikhomirova).
tuberculatus Coiffait, 1974a: 203 (Philonthus; Type locality: Shkotovo, Peishoula).
xantholinoides Wollaston, 1864, see: Neobisnius.
xantholinoides W. J. MacLeay, 1873, see: Heterothops.
xanthomerus Kraatz, 1859, see: Bisnius.
xanthopterus Nordmann, 1837, see: Belonuchus.
xanthoraphis Eppelsheim, 1895: 205 (Philonthus; Type locality: Ganale Gudda, Arussi Galla).
— Bernhauer and Schubert, 1914: 361 (Philonthus; catalog).
— Scheerpeltz, 1933: 1368 (Philonthus; catalog).
— Tottenham, 1962: 215 (Philonthus; characters).
— Tottenham, 1962: 216, 217 (Philonthus; cited as xanthorhaphis; characters).
— Scheerpeltz, 1974c: 22 (Philonthus; Somalia).
▴ Distribution: Ethiopia, Somalia, Kenya, Congo, South Africa.
marginellus Schubert, 1902: 406 (Philonthus; Type locality: Natal).
— Bernhauer and Schubert, 1914: 345 (Philonthus; catalog).
— Scheerpeltz, 1933: 1368 (Philonthus; synonym of xanthoraphis).
opaciventris Fauvel, 1907: 42 (Philonthus; Type locality: steppe Masai pres Nairobi).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Bernhauer and Chapman, 1939: 70 (Philonthus; Kenya).
— Tottenham, 1962: 217 (Philonthus; synonym of xanthoraphis).
niziensis Cameron, 1933: 42 (Philonthus; Type locality: Belgian Congo: Nizi: Blukwa).
— Cameron, 1950g: 36 (Philonthus; Belgian Congo).
— Tottenham, 1962: 217 (Philonthus; synonym of xanthoraphis).
yakimae Hatch, 1957: 191 (Philonthus; Type locality: Washington: Yakima R. Canyon).
yatungensis Cameron, 1928, see: protenus Schubert, 1906.
yemenicus Coiffait, 1981: 16 (Philonthus; Type locality: Yemen: Wadi Zabid).
yokohamus R. Dvořák, 1958: 138 (Philonthus; Type locality: Japan: Yokohama).
zaidius Tottenham, 1962: 158 (Philonthus; Type locality: Kenya: Timboroa, 2800 m. Also cited from Tanganyika).
zhangi J. Li, 1993: 47 (Philonthus; Type locality: Jiamusi, Heilongjiang).
zhuk Gusarov, 1995a: 93 [= 1996: 57] (Philonthus; Type locality: Ukraina: Krymskaia Obl.: nad Massandroi).
ziloanus Levasseur, 1962a: 59 (Philonthus; Type locality: Zilo, région de Kolwezi, Katanga).
zunilensis Sharp, 1885: 418 (Philonthus; Type locality: Guatemala, Cerro Zunil 4000 to 5000 feet).
Phucobius
[8 species; Palaearctic, Ethiopian, Oriental, and Oceanic Regions]
Phucobius Sharp, 1874: 35 (species included: simulator). Type species: Phucobius simulator Sharp, fixed by monotypy.
— Duvivier, 1883: 144 (catalog).
— Sharp, 1889: 108 (notes; valid genus).
— Bernhauer and Schubert, 1914: 373 (world catalog; 3 species).
— Winkler, 1925: 381 (catalog for Palaearctic region).
— Cameron, 1925: 65 (catalog of Indian species).
— Cameron, 1932: 178 (characters).
— Scheerpeltz, 1933: 1380 (world catalog supplement).
— Blackwelder, 1952: 307 (type species: simulator).
— Tikhomirova, 1973a: 186 (checklist of species of USSR).
— Shibata, 1984: 80 (checklist of species of Japan).
— Moore and Legner, 1976: 541 (includes seashore species; notes).
— Shibata, 1986a: 124 (supplemental list of species from Taiwan).
— Kanehisa, Tsumuki, and Kawazu, 1994 (chemical secretions).
— Smetana, 1995e: 50 (characters).
— Hayashi, 2000: 23 (notes on empodial setae).
africanus Bernhauer, 1937b: 617 (Phucobius; Type locality: W. Usambara: Kwarukussu, 1600 m).
congruus Walker, 1858: 205 (Ocypus; Type locality: Ceylon).
— Bernhauer and Schubert, 1914: 373 (Phucobius; catalog).
— Cameron, 1919b: 207 (Phucobius; notes; Ceylon).
— Cameron, 1932: 178 (Phucobius; characters; Ceylon).
— Scheerpeltz, 1933: 1380 (Phucobius; catalog).
▴ Distribution: Sri Lanka.
punctilinea Walker, 1858: 205 (Ocypus; Type locality: Ceylon).
— Fauvel, 1902a: 41 (Phucobius; synonym of congruus).
— Bernhauer and Schubert, 1914: 373 (Phucobius; synonym of congruus).
— Cameron, 1932: 178 (Phucobius; synonym of congruus).
horni Bernhauer, 1902b: 28 (Cafius; Type locality: Weligama).
— Fauvel, 1902a: 41 (Phucobius; synonym of congruus).
— Bernhauer and Schubert, 1914: 373 (Phucobius; synonym of congruus).
— Cameron, 1932: 178 (Phucobius; synonym of congruus).
cupreipennis Cameron, 1918: 89 (Orthidus; Type locality: Singapore: Pasir Panjang).
— Cameron, 1921a: 378, 405 (Orthidus; characters; Singapore).
— Scheerpeltz, 1933: 1368 (Orthidus; catalog).
— Watanabe and Shibata, 1976: 333 (Phucobius; Malaysia).
▴ Distribution: Singapore, Malaysia.
densipennis Bernhauer, 1931c: 131 (Phucobius; Type locality: Riu-Kiu-Inseln: Kagoshima).
— Scheerpeltz, 1933: 1380 (Phucobius; catalog).
— Ikudome and Hayakawa, 1994: 187 (Phucobius; cattle dung collections; Ryukyu Islands).
— Watanabe, 1998a: 140 (Phucobius; Japan).
▴ Distribution: Japan.
horni Bernhauer, 1902, see: congruus Walker, 1859.
pectoralis Boheman, 1858: 31 (Quedius; Type locality: China).
— Bernhauer and Schubert, 1914: 373 (Phucobius; catalog).
— Bernhauer, 1932c: 235 (Phucobius; notes).
— Scheerpeltz, 1933: 1380 (Phucobius; catalog).
▴ Distribution: China.
punctilinea Walker, 1858, see: congruus Walker, 1858.
semiaereus Cameron, 1934a: 22 (Phucobius; Type locality: New Hebrides: N.E. Malekula).
simulator Sharp, 1874: 35 (Phucobius; Type locality: Amakusa and Iwosima).
— Bernhauer and Schubert, 1914: 373 (Phucobius; catalog).
— Kirshenblat, 1938: 531 (Phucobius; Russia).
— Watanabe, 1961: 354 (Phucobius; Izu Islands).
— Nakane, 1963a: 92 (Phucobius; characters; habitus photograph; Japan).
— Watanabe and Shibata, 1972: 65 (Phucobius; Yaku-shima Island).
— Coiffait, 1974a: 199 (Phucobius; Siberia).
— Frank, 1982: 45 (Phucobius; list of parasites).
— Yuh, Paik, Kwon, and Lee, 1985: 243 (Phucobius; Korea).
— J. Li, 1993: 41 (Phucobius; China).
— Kanehisa, Tsumuki, and Kawazu, 1994 (Phucobius; chemical secretions).
▴ Distribution: Japan, Korea, China.
tricolor Bernhauer, 1917j: 125 (Phucobius; Type locality: Hongkong).
Platyschema
[1 species; Oriental Region]
Platyschema Cameron, 1937: 19 (species included: javana). Type species: Platyschema javana Cameron, fixed by monotypy.
javana Cameron, 1937: 19 (Platyschema; Type locality: Java: G. Tangkoeban Prahoe).
Proxenobius
[1 species; Neotropical Region]
Proxenobius Seevers, 1965: 326 (species included: borgmeieri). Type species: Proxenobius borgmeieri Seevers, fixed by original designation and monotypy.
borgmeieri Seevers, 1965: 327 (Proxenobius; Type locality: Mexico: Oaxaca: Modelo Forest, upper Coatzocoalcos River region. Also cited from Panama and Costa Rica).
Pseudocraspedomerus
[1 species; Australian Region]
Pseudocraspedomerus Bernhauer, 1927h: 294 (species included: alutaceus). Type species: Pseudocraspedomerus alutaceus Bernhauer, fixed by monotypy.
— Scheerpeltz, 1933: 1410 (world catalog supplement).
— Blackwelder, 1952: 325 (type species: alutaceus).
— Moore, 1960: 101 (characters in key).
alutaceus Bernhauer, 1927h: 294 (Pseudocraspedomerus; Type locality: Neu-Guinea: Kloofbivak).
Pseudomoeocerus
[1 species; Oriental Region]
Pseudomoeocerus Cameron, 1950d: 23 (species included: malayanus). Type species: Pseudomoeocerus malayanus Cameron, fixed by monotypy.
malayanus Cameron, 1950d: 23 (Pseudomoeocerus; Type locality: Pahang: Selangor, 2000 feet).
Pterygolaetus
[1 species; Neotropical Region]
Pterygolaetus Bierig, 1937: 194 (species included: williamsi). Type species: Pterygolaetus williamsi Bierig, fixed by original designation and monotypy.
— Blackwelder, 1944: 136 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 332 (type species: williamsi).
williamsi Bierig, 1937: 196 (Pterygolaetus; Type locality: Ecuador: Banos Or., 2700–3300 m).
Quediosoma
[1 species; Oriental Region]
Quediosoma Cameron, 1926b: 366 (species included: termitophaga). Type species: Quediosoma termitophaga Cameron, fixed by original designation and monotypy.
— Cameron, 1932: 256 (characters).
— Scheerpeltz, 1933: 1411 (world catalog supplement).
— Blackwelder, 1952: 335 (type species: termitophaga).
— Scheerpeltz, 1965: 278 (characters; type species: sericoilium = termitophagum).
sericoilium Cameron, 1920: 216 (Philonthus; Type locality: Nilgiri Hills).
— Cameron, 1932: 257 (Quediosoma; characters; India).
— Scheerpeltz, 1933: 1411 (Quediosoma; catalog).
— Seevers, 1957: 318 (Quediosoma; doubtfully termitophilous).
▴ Distribution: India.
termitophaga Cameron, 1926b: 367 (Quediosoma; Type locality: Dehra Dun).
— Cameron, 1932: 257 (Quediosoma; synonym of sericoilius).
— Scheerpeltz, 1933: 1411 (Quediosoma; synonym of sericoilium).
termitophaga Cameron, 1926, see: sericoilium Cameron, 1920.
Rabigus
[19 species; Nearctic, Palaearctic and Oriental Regions]
Rabigus Mulsant and Rey, 1876: 523 [= 1877: 379] (species included: tenuis, pullus). Type species: Philonthus pullus Nordmann, fixed by subsequent designation by Tottenham, 1939d: 229.
— Seidlitz, 1891: 389 (key to some European species).
— Seidlitz, 1891a: 414 (key to some European species).
— Heyden, Reitter, and Weise, 1891: 104 (list of species of Europe and Caucasus).
— Heyden, 1893: 43 (subgenus of Philonthus; list of species of Siberian region).
— Heyden, 1896: 33 (subgenus of Philonthus; list of species of Siberian region).
— Heyden, Reitter, and Weise, 1906: 168 (subgenus of Philonthus; list of species of Europe and Caucasus).
— Reitter, 1909: 124 (subgenus of Philonthus; characters).
— Bernhauer and Schubert, 1914: 326 (subgenus of Philonthus).
— Cameron, 1932: 62 (subgenus of Philonthus).
— Scheerpeltz, 1933: 1330 (subgenus of Philonthus).
— Tottenham, 1939d: 229 (type species designation: pullus).
— Blackwelder, 1943: 399 (subgenus of Philonthus; type species: pullus).
— Tottenham, 1949: 372,419 (type species: pullus Nordmann; subgenus of Philonthus; checklist of British species).
— Blackwelder, 1952: 336 (type species).
— Smetana, 1958: 143, 227 (subgenus of Philonthus; characters; type species).
— Smetana, 1959: 172 (subgenus of Philonthus; characters).
— Coiffait, 1963: 7, 18 (characters).
— Lohse, 1964: 186 (subgenus of Philonthus; characters; key to central European species).
— Coiffait, 1974: 99 (characters; key to species of western Palaearctic region).
— Shibata, 1983: 126 (checklist of species of Japan).
— Tóth, 1984: 79 (characters; habitus; key to species of Hungary).
— Segers, 1986: 43 (checklist of species; Belgium).
— Drugmand 1987b (comments on distribution of Belgian species).
— Lohse and Lucht, 1989: 168 (valid genus).
— Silfverberg, 1992: 16 (list of species of Fennoscandia, Denmark, and the Baltic States).
— Smetana, 1993b: 48 (list of species of Czech Republic and Slovakia).
— Smetana, 1995e: 32, 471 (characters; North American species; notes).
— Ciceroni and Zanetti, 1995: 29 (list of species of Italy).
— Newton, Thayer, Ashe, and Chandler, 2000: 397 (1 Nearctic species; notes; characters in key).
abauriae Gridelli, 1924e: 177, 180 (Philonthus; subgenus Rabigus; Type locality: Persia settentrionale).
— Scheerpeltz, 1933: 1330 (Philonthus; catalog).
— Smetana, 1965f: 257 (Philonthus; subgenus Rabigus; Caucasus; Persia).
— Coiffait, 1974: 107 (Rabigus; characters; Persia; Armenia).
— Raitschev, 1997a: 100 (Rabigus; notes; Bulgaria).
▴ Distribution: Iran, Armenia.
alienus Eppelsheim, 1889a: 178 (Philonthus; subgenus Rabigus; Type locality: Kan-ssu).
— Bernhauer and Schubert, 1914: 328 (Philonthus; catalog).
— Gridelli, 1924e: 181 (Philonthus; subgenus Rabigus).
— Scheerpeltz, 1933: 1331 (Philonthus; catalog).
— Coiffait, 1974: 109 (Rabigus; characters; Mongolia; China).
▴ Distribution: Mongolia, China.
auropilosus Luze, 1912, see: princeps Bernhauer, 1902.
basipilosus Schubert, 1908: 621 (Philonthus; Type locality: Kulu, Himalaya, ca. 2000 m).
— Bernhauer and Schubert, 1914: 330 (Philonthus; catalog).
— Cameron, 1932: 140 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1333 (Philonthus; catalog).
— Gridelli, 1934: 76 (Philonthus; subgenus Rabigus; characters; Karakoram region).
— Coiffait, 1979d: 552 (Rabigus; Afghanistan).
— Coiffait, 1981e: 349 (Rabigus; Baltistan).
— Coiffait, 1983e: 477 (Rabigus; Afghanistan).
▴ Distribution: India, Pakistan, Afghanistan.
brunnicollis Hochhuth, 1851, see: tenuis Fabricius, 1793.
convergens Casey, 1915, see: laxellus Casey, 1915.
dimidiatus Panzer, 1795, see: tenuis Fabricius, 1793.
escurialensis Perez, 1865: 12 (Philonthus; Type locality: los montes del Escorial antes de llegar á la cumbre).
— Bernhauer and Schubert, 1914: 337 (Philonthus; catalog).
— Scheerpeltz, 1933: 1341 (Philonthus; catalog).
— Gridelli, 1920a: 17 (Philonthus; characters; Portugal; Spain).
— Coiffait, 1974: 102 (Rabigus; characters; Spain; Portugal).
▴ Distribution: Spain, Portugal.
formosus Motschulsky, 1860: 567 (Gabrius; Type locality: Daghestan Caucasien).
— Hochhuth, 1862: 63 (Philonthus; characters; Daghestan).
— Solsky, 1871: 160 (Philonthus; characters; Uzbekistan; Djisak).
— Solsky, 1874: 182 (Philonthus; characters; Turkestan).
— Eppelsheim, 1892: 334 (Philonthus; Uzbekistan).
— Bernhauer and Schubert, 1914: 338 (Philonthus; catalog).
— Gridelli, 1924e: 179 (Philonthus; subgenus Rabigus; characters).
— Scheerpeltz, 1933: 1343 (Philonthus; catalog).
— Scheerpeltz, 1963: 13 (Philonthus; Afghanistan).
— Coiffait, 1974: 108 (Rabigus; characters; Daghestan; Caucasus).
▴ Distribution: Russia, Caucasus, Afghanistan, Uzbekistan.
gracilis Letzner, 1846, see: tenuis Fabricius, 1793.
hindostanus Schubert, 1911: 23 (Philonthus; Type locality: Schembagamur, Madura, Hindostan).
— Bernhauer and Schubert, 1914: 341 (Philonthus; catalog).
— Cameron, 1932: 97 (Philonthus; characters; India).
— Scheerpeltz, 1933: 1345 (Philonthus; catalog).
— Schillhammer, 1996a: 82 (Rabigus; notes).
▴ Distribution: India.
inconstans Sharp, 1889, see: tenuis Fabricius, 1793.
jumlensis Coiffait, 1982: 58 (Rabigus; Type locality: Nepal occidental: Dampelek, près de Jumla).
karakorus Coiffait, 1981e: 351 (Rabigus; Type locality: Karakorum, Satpora Lake, 2700 m, Skardu).
ladakhensis Coiffait, 1982: 58 (Rabigus; Type locality: Inde: Ladakh, Kargil, 2950 m).
lacustrinus Scheerpeltz, 1933, see: laxellus Casey, 1915.
lacustris Casey, 1915, see: laxellus Casey, 1915.
laeticollis Bernhauer, 1933, see: pullus Nordmann, 1837.
laxellus Casey, 1915: 431 (Philonthus; Type locality: New Mexico: Cloudcroft; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1348 (Philonthus; catalog).
— Smetana, 1995e: 473 (Rabigus; lectotype designation; characters; notes; Alberta; Manitoba; Northwest Territories; Ontario; Quebec; Saskatchewan; Arizona; Colorado; Idaho; Iowa;; Montana; Nebraska; New Mexico; New Hampshire; New York; North Dakota; South Dakota; Vermont; Wisconsin; Wyoming).
▴ Distribution: Canada, USA.
pumilio Casey, 1915: 431 (Philonthus; Type locality: Manitoba: Aweme; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1357 (Philonthus; catalog).
— Smetana, 1995e: 473 (Rabigus; lectotype designation; synonym of laxellus).
longiventris Casey, 1915: 432 (Philonthus; Type locality: Montana: Mullan; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1349 (Philonthus; catalog).
— Smetana, 1995e: 473 (Rabigus; lectotype designation; synonym of laxellus).
convergens Casey, 1915: 432 (Philonthus; Type locality: Montana: Helena; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1337 (Philonthus; catalog).
— Smetana, 1995e: 473 (Rabigus; lectotype designation; synonym of laxellus).
lacustris Casey, 1915: 432 (Philonthus; [preoccupied]; Type locality: Minnesota: Duluth; [Note: See lectotype designation by Smetana, 1995e]).
— Scheerpeltz, 1933: 1347 (Philonthus; synonym of lacustrinus).
— Smetana, 1995e: 473 (Rabigus; lectotype designation; synonym of laxellus).
lacustrinus Scheerpeltz, 1933: 1347 (Philonthus; replacement name for lacustris Casey).
longiventris Casey, 1915, see: laxellus Casey, 1915.
nigricollis Gerhardt, 1908, see: tenuis Fabricius, 1793.
ocaleoides J. Sahlberg, 1908: 35 (Philonthus; subgenus Rabigus; Type locality: Tarsi Anatoliae).
— Bernhauer and Schubert, 1914: 348 (Philonthus; catalog).
— Gridelli, 1924e: 181 (Philonthus; subgenus Rabigus).
— Scheerpeltz, 1933: 1353 (Philonthus; catalog).
— Smetana, 1959a: 212 (Philonthus; subgenus Rabigus; characters; Albania).
— Coiffait, 1974: 104 (Rabigus; characters; Anatolia; Balkans; Yugoslavia; Albania).
— Smetana, 1977: 794 (Rabigus; Turkey).
▴ Distribution: Turkey, “Balkan region”.
pamirensis Sharp, 1878: 170 (Philonthus; Type locality: on the road across the Pamir from Sarikol to Panjah).
— Sharp, 1890: 41 (Philonthus; [Note: Cited by Bernhauer and Schubert, 1914: 349, as the original description]; characters; Pamirs).
— Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).
— Gridelli, 1924e: 181 (Philonthus; subgenus Rabigus).
— Cameron, 1932: 127 (Philonthus; characters; Pamirs).
— Scheerpeltz, 1933: 1354 (Philonthus; catalog).
— Coiffait, 1974: 110 (Rabigus; characters; Pamirs).
— Boháč, 1988: 549 [= 1989: 32] (Rabigus; characters; Pamirs).
▴ Distribution: “Pamir region”.
princeps Bernhauer, 1902c: 56 (Philonthus; Type locality: Turkestan, Aulie Ata).
— Bernhauer and Schubert, 1914: 351 (Philonthus; catalog).
— Gridelli, 1924e: 180 (Philonthus; subgenus Rabigus; characters).
— Scheerpeltz, 1933: 1356 (Philonthus; catalog).
— Coiffait, 1974: 107 (Rabigus; characters; Turkestan; Turkey).
— Tronquet, 1981: 70 (Rabigus; Afghanistan).
▴ Distribution: Kazakhstan, Afghanistan, Turkey.
auropilosus Luze, 1912: 460 (Philonthus; subgenus Rabigus; Type locality: Türkei).
— Bernhauer and Schubert, 1914: 351 (Philonthus; synonym of princeps).
— Gridelli, 1924e: 181 (Philonthus; subgenus Rabigus).
— Scheerpeltz, 1933: 1356 (Philonthus; synonym of princeps).
— Coiffait, 1974: 106 (Rabigus; synonym of princeps).
pseudogracilis Reitter, 1909, see: pullus Nordmann, 1837.
pulloides Coiffait, 1982: 56 (Rabigus; Type locality: Inde: Cachemire, Sonamarg, Himalaya-Hauptkette, 2800 m).
— Coiffait, 1982a: 232 (Rabigus; Pakistan).
— Coiffait, 1983: 274 (Rabigus; Kashmir).
▴ Distribution: Pakistan, India.
pullus Nordmann, 1837: 104 (Philonthus; [Note: Preoccupied secondary homonym in Philonthus]; Type locality: Berolinum).
— Erichson, 1839a: 475 (Philonthus; characters; Germany).
— Erichson, 1840: 488 (Philonthus; characters; Germany; France).
— Heer, 1841: 583 (Philonthus; characters; Switzerland).
— Redtenbacher, 1849: 705 (Philonthus; characters; Austria).
— Fairmaire and Laboulbène, 1856: 527 (Philonthus; characters; France).
— Kraatz, 1857d: 617 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 195 (Philonthus; characters; Austria).
— Hochhuth, 1862: 63 (Philonthus; Russia).
— Fauvel, 1874: 251 [= 1874a: 477] (Philonthus; characters; France; Britain; Germany; Switzerland; Italy; Austria; Russia; Lake Baikal; Amur region).
— Redtenbacher, 1874: 214 (Philonthus; characters; Austria).
— Seidlitz, 1875: 278 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 528 [= 1877: 384] (Rabigus; characters; France).
— Fowler, 1888: 279 (Philonthus; characters; Britain).
— Eppelsheim, 1892: 334 (Philonthus; Turkmenistan; Uzbekistan).
— Ganglbauer, 1895: 459 (Philonthus; characters; middle Europe; Siberia).
— Reitter, 1909: 131 (Philonthus; subgenus Rabigus; characters; Germany).
— Bernhauer and Schubert, 1914: 351 (Philonthus; catalog).
— Gridelli, 1924e: 180 (Philonthus; subgenus Rabigus; characters).
— Portevin, 1929: 367 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1357 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— V. Hansen, 1952: 81 (Philonthus; subgenus Philonthus; characters).
— Smetana, 1958: 228 (Philonthus; subgenus Rabigus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 172 (Philonthus; subgenus Rabigus; characters).
— Lohse, 1964: 186 (Philonthus; subgenus Rabigus; characters; central Europe).
— Horion, 1965: 180 (Philonthus; subgenus Rabigus; Britain; Holland; Belgium; Switzerland; France; Spain; Italy; Dalmatia; Croatia; Bosnia Hercegovina; Serbia; Albania; Greece; Romania; Czech Republic; Slovakia; Poland; Russia; Germany; Austria).
— Raianu, 1968a: 468 (Philonthus; Romania).
— Szujecki, 1968a: 732 (Philonthus; Poland).
— Coiffait, 1974: 105 (Rabigus; characters; Europe; Britain; Asia Minor; Armenia; Siberia).
— Pope, 1977: 29 (Philonthus; Britain).
— Tóth, 1984: 80 (Rabigus; characters; Hungary).
— Ádám, 1987: 151 (Rabigus; collecting notes; Hungary).
— Lucht, 1987: 106 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).
— Lohse and Lucht, 1989: 168 (Rabigus).
— J. Li, 1993: 40 (Rabigus; China).
— Smetana, 1993b: 48 (Rabigus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Rabigus; Italy).
— Sterrenburg and Schülke, 1997: 20 (Rabigus; Netherlands).
▴ Distribution: Europe, Russia, Armenia, Turkey, Turkmenistan, Kazakhstan, Uzbekistan, Pakistan, China.
pseudogracilis Reitter, 1909: 131 (Philonthus; subgenus Rabigus; variety of pullus; Type locality: Not cited).
— Scheerpeltz, 1933: 1357 (Philonthus; aberration of pullus).
— Smetana, 1958: 228 (Philonthus; synonym of pullus).
— Smetana, 1959: 172 (Philonthus; subgenus Rabigus; aberration of pullus; characters).
— Coiffait, 1974: 105 (Rabigus; variety of pullus; characters; Europe; Kazakhstan).
— Tóth, 1984: 80 (Rabigus; aberration of pullus; characters; Hungary).
laeticollis Bernhauer, 1933: 30 (Philonthus; variety of pullus; Type locality: Thibet).
— Gridelli, 1934: 77 (Philonthus; subspecies of basipilosus; characters; Karakoram region).
— Coiffait, 1974: 105 (Rabigus; subspecies of pullus; characters; Tibet).
pumilio Casey, 1915, see: laxellus Casey, 1915.
queinneci Coiffait, 1983d: 164 (Rabigus; Type locality: Nepal: Ganesh Himal, entre Salmi et Rupchet, 1500 à 3500 m).
reskai R. Dvořák, 1954, see: tenuis Fabricius, 1793.
ruficapillus Reitter, 1887: 216 (Philonthus; Type locality: Keria 7,500 bis 8,000 F. [Xinjiang]).
— Bernhauer and Schubert, 1914: 353 (Philonthus; catalog).
— Gridelli, 1924e: 181 (Philonthus; subgenus Rabigus).
— Scheerpeltz, 1933: 1359 (Philonthus; catalog).
— Coiffait, 1974: 108 (Rabigus; characters; central Asia).
▴ Distribution: China.
syriacus Gridelli, 1924e: 178, 179 (Philonthus; subgenus Rabigus; Type locality: Tripoli de Siria).
— Scheerpeltz, 1933: 1363 (Philonthus; catalog).
— Coiffait, 1974: 109 (Rabigus; characters; Lebanon).
▴ Distribution: Syria, Lebanon.
tenuis Fabricius, 1793: 528 (Staphylinus; Type locality: Germania).
— Panzer, 1795: 357 (Staphylinus; characters; Germany).
— Fabricius, 1801: 599 (Staphylinus; characters; Germany).
— Turton, 1802: 513 (Staphylinus; characters; Germany).
— Gravenhorst, 1802: 39 (Staphylinus; characters).
— Latreille, 1804: 322 (Staphylinus; characters; Germany).
— Gravenhorst, 1806: 72 (Staphylinus; characters).
— Panzer, 1808: 11 (Staphylinus; characters; illustrations).
— Curtis, 1829: 28 (Bolitobius; catalog; Britain).
— Runde, 1835: 10 (Staphylinus; characters; Germany).
— Lacordaire, 1835: 403 (Staphylinus; characters; France).
— Nordmann, 1837: 103 (Philonthus; characters; Germany).
— Erichson, 1839a: 474 (Philonthus; characters; Germany).
— Heer, 1839: 270 (Philonthus; characters; Switzerland).
— Erichson, 1840: 488 (Philonthus; characters; Germany).
— Kolenati, 1846: 21 (Philonthus; Transcaucasia).
— Hochhuth, 1849: 144 (Philonthus; characters).
— Redtenbacher, 1849: 706 (Philonthus; characters; Austria).
— Fairmaire and Laboulbène, 1856: 528 (Philonthus; characters; France).
— Kraatz, 1857d: 617 (Philonthus; characters; Germany).
— Redtenbacher, 1857: 196 (Philonthus; characters; Austria).
— Hochhuth, 1862: 63 (Philonthus; Russia).
— Fauvel, 1874: 251 [= 1874a: 477] (Philonthus; characters; France; Finland; Germany; Russia; Austria; Switzerland; Italy; Sardinia; Transcaucasia; Amur region).
— Fauvel, 1876a: 75 [= 1876: 260] (Philonthus; Portugal).
— Redtenbacher, 1874: 214 (Philonthus; characters; Austria).
— Seidlitz, 1875: 278 (Philonthus; characters; Baltic region).
— Mulsant and Rey, 1876: 526 [= 1877: 382] (Rabigus; characters; France).
— J. Sahlberg, 1876: 22 (Philonthus; characters; Finland).
— Eppelsheim, 1892: 334 (Philonthus; Uzbekistan).
— Ganglbauer, 1895: 458 (Philonthus; subgenus Rabigus; characters; Palaearctic region).
— Poppius, 1909: 27 (Philonthus; subgenus Rabigus; Lena Valley, Siberia).
— Reitter, 1909: 131 (Philonthus; subgenus Rabigus; characters; Germany).
— Bernhauer and Schubert, 1914: 357 (Philonthus; catalog).
— Gridelli, 1924e: 177 (Philonthus; subgenus Rabigus; characters; Persia).
— Portevin, 1929: 367 (Philonthus; characters; France).
— Scheerpeltz, 1933: 1363 (Philonthus; catalog).
— Wüsthoff, 1934: 61 (Philonthus; illustration of aedeagus).
— V. Hansen, 1952: 81 (Philonthus; subgenus Philonthus; characters).
— Smetana, 1958: 227 (Philonthus; subgenus Rabigus; characters; ecology; Czechoslovakia).
— Smetana, 1959: 172 (Philonthus; subgenus Rabigus; characters).
— Palm, 1963b: 98 (Philonthus; subgenus Rabigus; characters; Sweden; Finland).
— Lohse, 1964: 186 (Philonthus; subgenus Rabigus; characters; central Europe).
— Horion, 1965: 178 (Philonthus; subgenus Rabigus; Russia; Baltics; Poland; Czechoslovakia; Romania; Serbia; Bosnia; Croatia; Italy; Spain; Portugal; France; Switzerland; Belgium; Holland; Germany; Austria; Finland; Sweden).
— Raianu, 1968a: 468 (Philonthus; Romania).
— Szujecki, 1968a: 731 (Philonthus; Poland).
— Korge, 1971a: 7 (Philonthus; Turkey).
— Coiffait, 1974: 106 (Rabigus; characters; Europe; Siberia; Britain; Scandinavia; Sweden; France).
— Shilov, 1976: 153 (Philonthus; Mongolia).
— Tóth, 1984: 79 (Rabigus; characters; Hungary).
— Lucht, 1987: 106 (Philonthus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).
— Lohse and Lucht, 1989: 168 (Rabigus).
— Nowosad, 1990: 113 (Philonthus; collected from mole nest; notes; Poland).
— Pilon and Zanetti, 1991: 64 (Rabigus; Italy).
— J. Li, 1993: 40 (Rabigus; China).
— Smetana, 1993b: 48 (Rabigus; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 29 (Rabigus; Italy).
— Schülke, 1998g: 90 (Rabigus; Germany).
▴ Distribution: Europe, Russia, Caucasus, Turkey, Iran, Uzbekistan, Mongolia.
dimidiatus Panzer, 1795: 362 (Paederus; Type locality: Germania).
— Panzer, 1796c: 24 (Paederus; characters; illustration).
— Panzer, 1808: 11 (Staphylinus; synonym of tenuis Fabricius).
— Gravenhorst, 1802: 40 (Staphylinus; synonym of tenuis Fabricius).
— Erichson, 1839a: 474 (Philonthus; synonym of tenuis).
— Heer, 1839: 270 (Philonthus; synonym of tenuis).
— Erichson, 1840: 488 (Philonthus; synonym of tenuis).
— Redtenbacher, 1849: 706 (Philonthus; synonym of tenuis).
— Kraatz, 1857d: 618 (Philonthus; synonym of tenuis).
— Ganglbauer, 1895: 458 (Philonthus; synonym of tenuis).
— Bernhauer and Schubert, 1914: 357 (Philonthus; synonym of tenuis).
— Smetana, 1958: 227 (Philonthus; synonym of tenuis).
— Coiffait, 1974: 106 (Rabigus; synonym of tenuis).
gracilis Letzner, 1846: 79 (Philonthus; Type locality: Not cited).
— Redtenbacher, 1857: 198 (Philonthus; characters; Germany).
— Kraatz, 1857d: 618 (Philonthus; synonym of tenuis).
— Redtenbacher, 1874: 217 (Philonthus; characters; Germany).
— Mulsant and Rey, 1876: 526 [= 1877: 382] (Rabigus; synonym of tenuis).
— Ganglbauer, 1895: 458 (Philonthus; aberration of tenuis).
— Bernhauer and Schubert, 1914: 357 (Philonthus; aberration of tenuis).
— Scheerpeltz, 1933: 1364 (Philonthus; aberration of tenuis).
— Smetana, 1959: 172 (Philonthus; subgenus Rabigus; aberration of tenuis; characters).
— Smetana, 1958: 227 (Philonthus; aberration of tenuis).
— Coiffait, 1974: 106 (Rabigus; variety of tenuis).
— Tóth, 1984: 80 (Rabigus; aberration of tenuis).
brunnicollis Hochhuth, 1851: 20 (Philonthus; Type locality: Kasan; [Note: See lectotype designation by Gusarov, 1992a]).
— Bernhauer and Schubert, 1914: 331 (Philonthus; catalog).
— Gridelli, 1924e: 180 (Philonthus; subgenus Rabigus; characters).
— Scheerpeltz, 1933: 1335 (Philonthus; catalog).
— Nakane, 1963a: 91 (Philonthus; characters; habitus photograph; Japan).
— Coiffait, 1974: 110 (Rabigus; characters; Russia).
— Shilov, 1976: 153 (Philonthus; Mongolia).
— Yuh, Paik, Kwon, and Lee, 1985: 243 (Rabigus; Korea).
— Gusarov, 1992a: 786 [= 1993a: 75] (Rabigus; synonym of tenuis; lectotype designation).
— J. Li, 1993: 40 (Rabigus; cited as valid species; China).
inconstans Sharp, 1889: 40 (Philonthus; Type locality: Osaka; Niigata; Yokohama; Oyama; Hakodate; Junsai).
— Bernhauer and Schubert, 1914: 331 (Philonthus; synonym of brunnicollis).
— Coiffait, 1974: 110 (Rabigus; synonym of brunnicollis).
nigricollis Gerhardt, 1908: 209 (Philonthus; variety of tenuis; Type locality: Ohlau: Rodeland).
— Bernhauer and Schubert, 1914: 357 (Philonthus; aberration of tenuis).
— Scheerpeltz, 1933: 1364 (Philonthus; aberration of tenuis).
— Smetana, 1958: 227 (Philonthus; aberration of tenuis).
— Smetana, 1959: 172 (Philonthus; subgenus Rabigus; aberration of tenuis; characters).
— Coiffait, 1974: 106 (Rabigus; variety of tenuis).
— Tóth, 1984: 80 (Rabigus; aberration of tenuis).
reskai R. Dvořák, 1954c: 36 (Philonthus; subgenus Rabigus; variety of tenuis; Type locality: Komorany, Bohême centr., vers Prague).
Remus
[4 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions]
Remus Holme, 1837: 64 (species included: sericeus). Type species: Remus sericeus Holme, fixed by monotypy.
— Westwood, 1838a: 16 (characters; type species).
— Stephens, 1839: 385 (characters).
— Erichson, 1840: 426 (synonym of Philonthus).
— Lacordaire, 1854: 80 (synonym of Philonthus).
— Kraatz, 1857d: 565 (synonym of Philonthus).
— Thomson, 1859: 25 (type species: cinerascens).
— Gemminger and Harold, 1868: 584 (synonym of Philonthus).
— Fauvel, 1874: 196 [= 1874a: 421] (synonym of Cafius).
— Ganglbauer, 1895: 463 (subgenus of Cafius).
— Reitter, 1909: 116 (characters).
— Bernhauer and Schubert, 1914: 361 (subgenus of Cafius).
— Johansen, 1914: 421 (characters).
— Porta, 1926: 98 (characters; key to species of Italy).
— Cameron, 1932: 154 (subgenus of Cafius).
— Joy, 1932: 126 (key to British species).
— Scheerpeltz, 1933: 1369 (Philonthus; subgenus of Cafius).
— Blackwelder, 1943: 435 (subgenus of Cafius; type species: sericeus).
— Kloet and Hincks, 1945: 166 (list of British species).
— Tottenham, 1949: 373, 420 (type species: sericeus Holme; subgenus of Cafius; checklist of British species).
— Blackwelder, 1952: 338 (subgenus of Cafius; type species: sericeus).
— V. Hansen, 1952: 33 (characters; Danish species).
— Hatch, 1957: 210 (subgenus of Cafius; characters; species of Pacific Northwest).
— Coiffait, 1963: 8, 20 (cited as valid genus; characters; list of French species).
— Palm, 1963b: 59 (characters; species of Sweden).
— Lohse, 1964: 173 (cited as valid genus; characters; key to central European species).
— Pototskaia, 1966: 152 (cited as valid genus; larval characters).
— Pototskaia, 1967: 53 (cited as valid genus; larval characters).
— Kasule, 1970: 64, 65 (cited as valid genus; larval characters).
— Bordoni, 1973j: 702 (cited as valid genus; list of species from islands near Sicily).
— Tikhomirova, 1973a: 182 (cited as valid genus; checklist of species of USSR).
— Coiffait, 1974: 332 (cited as valid genus; adult and larval characters; key to species of western Palaearctic region).
— Pope, 1977: 30 (subgenus of Cafius).
— Topp, 1978: 312 (larval characters).
— Muona, 1979: 14 (cited as valid genus; list of Scandinavian species).
— Shibata, 1983: 133 (subgenus of Cafius; checklist of species of Japan).
— Silfverberg, 1992: 17 (cited as valid genus; list of species of Fennoscandia, Denmark, and the Baltic States).
— Hernández, Outerelo, and Gamarra, 1994: 188 (checklist of species; Canary Islands).
— Ciceroni and Zanetti, 1995: 27 (list of species of Italy).
— M. Hansen, 1996: 104 (cited as valid genus; list of Danish species).
Menapius Holme, 1842: 128 (species included: sericeus; [Note: Nomen nudum. In a defense of his recognition of Remus as a genus distinct from Cafius, Holme mentioned that some specimens of sericeus had been placed in the British Museum under the name Menapius grisescens. Holme wrote, “I find that I was wrong in supposing that Mr. Rudd had taken the insects which I characterized . . . as Remus sericeus, on the coast of Yorkshire; his specimens were from the Isle of Wight, . . . Mr. Rudd had distinguished the specimens [from the Isle of Wight] which he placed in the British Museum by the MS. name of Menapius grisescens.” Blackwelder, 1952: 237, concluded that this citation constituted validation of the generic name Menapius by objective synonymy with Remus and that the type species would be sericeus by monotypy. Since Menapius was first published as a synonym of Remus, it is unavailable (Article 11.6)]).
Pseudidus Mulsant and Rey, 1876: 574 [= 1877: 430] (species included: sericeus, pruinosus, filum). Type species: Remus sericeus Holme, fixed by subsequent designation by Tottenham, 1939d: 229.
— Bernhauer and Schubert, 1914: 361 (synonym of Cafius).
— Cameron, 1932: 154 (synonym of Cafius).
— Tottenham, 1939d: 229 (type species designation: sericeus).
— Blackwelder, 1943: 435 (synonym of Remus; type species: sericeus).
— Tottenham, 1949: 373 (synonym of Remus; type species: sericeus Holme).
— Blackwelder, 1952: 324 (type species).
— Coiffait, 1974: 332 (synonym of Remus).
aegyptiacus Motschulsky, 1858, see: sericeus Holme, 1837.
corallicola Fairmaire, 1849: 289 (Philonthus; Type locality: Taïti).
— Fauvel, 1874: 196 [= 1874a: 422] (Cafius).
— Fauvel, 1877: 256 (Cafius; characters; Tahiti).
— Fauvel, 1889b: 262 (Cafius; New Caledonia).
— Fauvel, 1898a: 117 (Cafius; Seychelles).
— Fauvel, 1903d: 268 (Cafius; New Caledonia; Australia; Tahiti; Seychelles; Mauritius; Madagascar; Perim; Djibouti).
— H. Kolbe, 1910: 19 (Cafius; Seychelles).
— Bernhauer and Schubert, 1914: 362 (Cafius; catalog).
— Cameron, 1921a: 378, 405 (Cafius; characters; Singapore).
— Bernhauer, 1922b: 177 (Cafius; Seychelles).
— Lea, 1925: 225, 226 (Cafius; characters; Australian introduction).
— Cameron, 1927c: 264 (Cafius; Fiji; New Caledonia).
— Scheerpeltz, 1933: 1369 (Cafius; catalog).
— C. Koch, 1936b: 186 (Cafius; characters in key; Tahiti; New Caledonia; Australia; Mauritius; Singapore; Ceylon; Hong Kong).
— Cameron, 1937: 15 (Cafius; Java).
— Coiffait, 1974: 337 (Remus; characters).
— Watanabe and Shibata, 1976: 333 (Cafius; subgenus Bryonomus; Malaysia).
— Coiffait, 1976: 217 (Cafius; cited as corallicolla; Tahiti).
— Coiffait, 1977a: 171 (Cafius; Austral Islands).
— Lecoq, 1990: 205 (Remus; Mascarene Islands).
▴ Distribution: Austral Islands, Society Islands, Fiji, New Caledonia, Australia, China, Indonesia, Singapore, Malaysia, Seychelles, Mauritius, Madagascar, Djibouti, Yemen.
occidentalis Olliff, 1887: 508 (Cafius; [Note: Olliff's description was published before Blackburn's]; Type locality: West Australia).
— Blackburn, 1888b: 48 (Cafius; [Note: Bernhauer and Schubert, 1914: 363, cited Blackburn as the author for this name, and although Olliff also cited Blackburn as the author, Olliff's description appeared first]; characters; Australia).
— Bernhauer and Schubert, 1914: 363 (Cafius; catalog).
— Lea, 1925: 226 (Cafius; synonym of corallicola).
— Scheerpeltz, 1933: 1369 (Cafius; synonym of corallicola).
— Coiffait, 1974: 337 (Remus; synonym of corallicola, cited as corallicolla).
filum Kiesenwetter, 1849: 19 (Philonthus; Type locality: Kahr, Dalmatien).
— Kiesenwetter, 1851a: 417 (Philonthus; France).
— Fairmaire and Laboulbène, 1856: 532 (Philonthus; characters; France).
— Kraatz, 1857d: 621 (Philonthus).
— Fauvel, 1874: 199 [= 1874a: 425] (Staphylinus; synonym of sericeus).
— Mulsant and Rey, 1876: 580 [= 1877: 437] (Pseudidus; characters; France).
— Ganglbauer, 1895: 463 (Cafius; variety of sericeus).
— Fauvel, 1902b: 106 (Cafius; synonym of sericeus).
— Reitter, 1909: 123 (Remus; variety of sericeus; characters; Germany).
— Bernhauer and Schubert, 1914: 363 (Cafius; variety of sericeus).
— Gridelli, 1931: 125 (Cafius; characters in key).
— Scheerpeltz, 1933: 1369 (Cafius; catalog).
— C. Koch, 1936b: 192 (Cafius; characters).
— Coiffait, 1974: 337 (Remus; characters; Atlantic coast to the south of Ausdierne Bay; western Mediterranean coast; Dalmatia; Tripolitania; Sinai; Black Sea coast).
— Ciceroni and Zanetti, 1995: 27 (Remus; Italy).
— Assing and Wunderle, 2001: 37 (Remus; Cyprus).
▴ Distribution: France, Germany, Croatia, Bulgaria, Romania, Turkey, Egypt, Libya.
obscuricornis C. Koch, 1936, see: sericeus Holme, 1837.
occidentalis Olliff, 1887, see: corallicola Fairmaire, 1849.
pruinosus Erichson, 1840: 510 (Philonthus; Type locality: Sardinia).
— Kiesenwetter, 1851a: 417 (Philonthus; France).
— Fairmaire and Laboulbène, 1856: 532 (Philonthus; characters; France).
— Kraatz, 1857d: 621 (Philonthus).
— Peyron, 1858: 427 (Philonthus; Turkey).
— Hochhuth, 1862: 65 (Philonthus; Russia).
— Fauvel, 1874: 199 [= 1874a: 425] (Staphylinus; synonym of sericeus).
— Mulsant and Rey, 1876: 579 [= 1877: 435] (Pseudidus; characters; France).
— Ganglbauer, 1895: 463 (Cafius; synonym of sericeus).
— Fauvel, 1902b: 106 (Cafius; variety of sericeus).
— Bernhauer and Schubert, 1914: 363 (Cafius; variety of sericeus).
— Scheerpeltz, 1933: 1370 (Cafius; variety of sericeus).
— Bierig, 1934a: 67 (Cafius; characters; Cuba).
— C. Koch, 1936b: 192 (Cafius; characters).
— Coiffait, 1974: 336 (Remus; characters; Atlantic coast from Holland to Portugal; Mediterranean coast; Sardinia; Canary Islands).
— Israelson, 1990: 2 (Remus; cited as valid species; Azores).
— Borges, 1990: Table II (Remus; cited as valid species; Azores).
— Ciceroni and Zanetti, 1995: 27 (Remus; Italy).
▴ Distribution: Cuba, Canary Islands, Azores, Netherlands, Portugal, France, Turkey, Russia.
sericeus Holme, 1837: 64 (Remus; Type locality: Isle of Wight according to correction by Holm, 1842: 128; originally cited as Scilly Islands: St. Mary's).
— Westwood, 1838a: 16 (Remus; type species).
— Stephens, 1839: 401 (Remus; characters; England).
— Erichson, 1840: 509 (Philonthus; characters; France; England).
— Jacobsen, 1849: 706 (Philonthus; characters; Denmark).
— Kiesenwetter, 1849: 18 (Philonthus; characters; England).
— Kiesenwetter, 1851a: 417 (Philonthus; France).
— Fairmaire and Laboulbène, 1856: 532 (Philonthus; characters; France).
— Kraatz, 1857d: 621 (Philonthus).
— Thomson, 1860: 153 (Cafius; characters; Scandinavia).
— Wollaston, 1864: 576 (Philonthus; characters; Canary Islands).
— Wollaston, 1865: 495 (Philonthus; habitat; Canary Islands).
— Baudi, 1870: 386 (Philonthus; characters).
— Fauvel, 1874: 199 [= 1874a: 425] (Cafius; characters; France; Scandinavia; Britain; Mediterranean region; Madeira).
— Seidlitz, 1875: 279 (Philonthus; characters; Baltic region).
— Fauvel, 1876a: 72 [= 1876: 257] (Cafius; Morocco).
— Mulsant and Rey, 1876: 577 [= 1877: 433] (Pseudidus; characters; France).
— Fauvel, 1878: 120 [= 1878a: 40] (Cafius; Morocco; Algeria).
— Fauvel, 1878g: 542 (Cafius; characters; notes; Australia).
— Horn, 1884: 238 (Cafius; characters; Coney Island, near New York).
— Fauvel, 1886: 52 [= 1886a: 44] (Cafius; Morocco; Egypt; Algeria).
— Olliff, 1887: 507 (Cafius; characters; Australia).
— Fowler, 1888: 283 (Cafius; characters; Britain).
— Ganglbauer, 1895: 463 (Cafius; characters; north and middle Europe; Mediterranean region; North America).
— Fauvel, 1902b: 106 (Cafius; Barbary; Egypt; Canaries).
— Sainte-Claire Deville, 1906: 95 (Cafius; Corsica).
— Reitter, 1909: 123 (Remus; characters; Germany).
— Bernhauer and Schubert, 1914: 363 (Cafius; catalog).
— Johansen, 1914: 421 (Remus; characters; Denmark).
— Portevin, 1929: 355 (Cafius; characters; France).
— Gridelli, 1930: 72 (Cafius; Libya).
— Gridelli, 1931: 124 (Cafius; characters in key).
— Scheerpeltz, 1933: 1370 (Cafius; catalog).
— C. Koch, 1936b: 170 (Cafius; characters; Egypt).
— Paulian, 1941: 233, 327 (Remus; larval and pupal characters).
— V. Hansen, 1952: 33 (Remus; characters; Denmark).
— Scheerpeltz, 1960d: 136 (Cafius; subgenus Remus; Portugal).
— Palm, 1963b: 60 (Remus; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 173 (Remus; characters; central Europe).
— Horion, 1965: 121 (Remus; sea coast of Europe).
— Pototskaia, 1967: 53 (Remus; larval characters).
— James, Moore, and Legner, 1971: 282 (Cafius; larval characters).
— Coiffait, 1974: 335 (Remus; characters; Europe; England; Scandinavia; Bulgarian coast of Black Sea).
— Pope, 1977: 30 (Cafius; Britain).
— Frank, 1982: 34 (Cafius; list of parasites).
— Israelson, 1984: 145 (Remus; Azores).
— Lucht, 1987: 104 (Remus; checklist; Germany; Benelux; Denmark; Sweden).
— Lecoq, 1990: 203 (Remus; Mascarene Islands).
— Maachi, 1991: 251 (Cafius; Morocco).
— Ciceroni and Zanetti, 1995: 27 (Remus; Italy).
— Assing and Wunderle, 2001: 36 (Remus; Cyprus).
▴ Distribution: Norway, Sweden, Denmark, Germany, Britain, France, Portugal, Azores, Madeira, Canary Islands, Morocco, Algeria, Libya, Egypt, Turkey, Bulgaria, Mascarene Islands, Australia, USA.
aegyptiacus Motschulsky, 1858: 665 (Cafius; Type locality: Alexandrie en Egypte).
— Fauvel, 1874: 199 [= 1874a: 425] (Staphylinus; synonym of sericeus).
— Fauvel, 1902b: 106 (Cafius; synonym of sericeus).
— Bernhauer and Schubert, 1914: 363 (Cafius; synonym of sericeus).
obscuricornis C. Koch, 1936b: 170, 183 (Cafius; variety of sericeus; Type locality: östliches Mediterrangebiet).
— Scheerpeltz, 1958a: 22 (Cafius; variety of sericeus; Turkey).
— Coiffait, 1968c: 505 (Remus; variety of sericeus; Corsica).
— Coiffait, 1974: 336 (Remus; variety of sericeus; characters).
subniger Mulsant and Rey, 1876, see: Erichsonius.
Scelotrichus
[1 species; Australian Region]
Scelotrichus Bernhauer, 1915g: 190 (species included: elegans). Type species: Scelotrichus elegans Bernhauer, fixed by monotypy.
— Scheerpeltz, 1933: 1374 (world catalog supplement).
— Blackwelder, 1952: 345 (type species: elegans).
elegans Bernhauer, 1915g: 191 (Scelotrichus; Type locality: Deutsch-Neuguinea: Schrader-berg, 2100 m).
Sternotoxus
[2 species; Australian Region]
Sternotoxus Bernhauer, 1916a: 6 (species included: flavicornis). Type species: Sternotoxus flavicornis Bernhauer, fixed by monotypy.
— Scheerpeltz, 1933: 1326 (world catalog supplement).
— Blackwelder, 1952: 360 (type species: flavicornis).
dohrni Fauvel, 1877: 266 (Belonuchus; Type locality: Australie septentrionale, Cap York).
— Bernhauer and Schubert, 1914: 370 (Belonuchus; catalog).
— Lea, 1925: 228 (Belonuchus; Australia).
— Scheerpeltz, 1933: 1326 (Sternotoxus; catalog).
▴ Distribution: Australia.
flavicornis Bernhauer, 1916a: 6 (Sternotoxus; Type locality: Nordost-Australien).
Taxiplagus
[1 species; Oriental Region]
Taxiplagus Bernhauer, 1915h: 236 (species included: abnormalis). Type species: Taxiplagus abnormalis Bernhauer, fixed by monotypy.
— Scheerpeltz, 1933: 1414 (world catalog supplement).
— Blackwelder, 1952: 375 (type species: abnormalis).
abnormalis Bernhauer, 1915h: 237 (Taxiplagus; Type locality: Java: Preanger).
Thinocafius
[1 species; Australian Region]
Thinocafius Steel, 1949c: 309 (species included: insularis). Type species: Thinocafius insularis Steel, fixed by original designation.
— Blackwelder, 1952: 386 (type species: insularis).
— Moore and Legner, 1976: 540 (includes seashore species; notes).
insularis Steel, 1949c: 309 (Thinocafius; Type locality: Chatham Islands: Pitt Island).
Trachyphilonthus
[1 species; Oriental Region]
Trachyphilonthus Scheerpeltz, 1965: 234 (species included: malaisei). Type species: Trachyphilonthus malaisei Scheerpeltz, fixed by original designation and monotypy.
malaisei Scheerpeltz, 1965: 236 (Trachyphilonthus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).
Tropiopterius
[1 species; Australian Region]
Tropiopterius Bernhauer, 1915g: 194 (species included: purpuripennis). Type species: Tropiopterius purpuripennis Bernhauer, fixed by monotypy.
— Scheerpeltz, 1933: 1380 (world catalog supplement).
— Blackwelder, 1952: 399 (type species: purpuripennis).
purpuripennis Bernhauer, 1915g: 194 (Tropiopterius; Type locality: Neu-Guinea: Kapaur).
Xanthodermus
[1 species; Neotropical Region]
Xanthodermus Bernhauer, 1912d: 40 (species included: vestitus). Type species: Belonuchus vestitus R. Sahlberg, fixed by monotypy.
— Bernhauer and Schubert, 1914: 372 (world catalog; 1 species).
— Blackwelder, 1944: 138 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 404 (type species: vestitus).
vestitus R. Sahlberg, 1847: 788 (Belonuchus; Type locality: Rio Janeiro).
Xenobius
[1 species; Neotropical Region]
Xenobius Borgmeier, 1931: 358 (species included: rotundiceps). Type species: Xenobius rotundiceps Borgmeier, fixed by original designation and monotypy.
— Scheerpeltz, 1933: 1320 (world catalog supplement).
— Blackwelder, 1944: 149 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 406 (type species: rotundiceps).
— Seevers, 1965: 324 (characters; myrmecophilus species; list of species).
rotundiceps Borgmeier, 1931: 360 (Xenobius; Type locality: Campinas, Goyaz).
V. Staphylinine Group (Part 2)
Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina)," Bulletin of the American Museum of Natural History 2001(265), 2441-3020, (18 July 2019). https://doi.org/10.1206/0003-0090.265.1.5
