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18 July 2001 Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium.
VI. Staphylinine Group (Part 3)
Staphylininae: Staphylinini (Quediina, Staphylinina, Tanygnathinina, Xanthopygina), Xantholinini
Staphylinidae Incertae Sedis
Fossils, Protactinae
Lee H. Herman
Author Affiliations +
Abstract

This catalog (published in seven parts, all released on the same day) is based on only the published literature for the Staphylinidae. Of the 32 subfamilies, the following 28 are included herein: Apateticinae, Dasycerinae, Empelinae, Euaesthetinae, Glypholomatinae, Habrocerinae, Leptotyphlinae, Megalopsidiinae, Micropeplinae, Microsilphinae, Neophoninae, Olisthaerinae, Omaliinae, Osoriinae, Oxyporinae, Oxytelinae, Phloeocharinae, Piestinae, Protactinae†, Proteininae, Protopselaphinae, Pseudopsinae, Solieriinae, Staphylininae, Steninae, Tachyporinae, Trichophyinae, and Trigonurinae. The Aleocharinae, Paederinae, Pselphinae, and Scaphidiinae are excluded from this edition of the catalog. References to the original citation or description are given for available family-group, genus-group, and species-group names of both extant and extinct forms. The type genus is cited for each family-group name, the type species for each genus-group name, and the type locality for each species-group name. Where appropriate, all subgenera, subspecies, or synonyms are listed for each valid name. Annotated subsequent references are presented for all names. Distributional summaries are given for each valid taxon. Full bibliographic citations are in Part VII. A short historical review, coauthored with Aleš Smetana, follows the Introduction (Part I), with the main focus on biographical sketches that include many photographs. The goal of this catalog is to summarize the current state of knowledge of the family and to stimulate worldwide monographic studies.

Subtribe Quediina

Quediina Kraatz, 1857.

  • — Jakobson, 1908: 447 (characters).

  • — Jakobson, 1909: 514 (key to genera; list of species of Russia and western Europe).

  • — Johansen, 1914: 341 (characters; key to Danish genera).

  • — Hatch, 1957: 172, 214 (characters; key to species of Pacific Northwest).

  • — Arnett, 1963: 279 (characters; list of North American genera).

  • — Newton and Thayer, 1992: 65 (subtribe of Staphylinini).

  • — Smetana, 1995e: 48 (characters).

  • — Klimaszewski, Newton, and Thayer, 1996: 151 (list of New Zealand genera; number of New Zealand species).

  • — Smetana and Davies, 2000: 12, 14 (characters).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 331 (key to Nearctic genera).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 331, 392 (key to Nearctic genera; characters; notes).

Platycnemidiformes Nordmann, 1837: 6 (genera included: Platycnemus). Type genus: Platycnemus (see Newton and Thayer, 1992: 65).

  • — Newton and Thayer, 1992: 65 (synonym of Quediina).

  • — Newton, 1995: 48, 49, 51 (name emended to Platycnemini).

Platycnemini Nordmann, 1837.

  • — Newton, 1995: 51 (name emended from Platycnemidiformes; requested that the International Commission of Zoological Nomenclature Platycnemini not be given priority over junior family-group names in current general usage in the Staphylininae and that it be placed on the Official List of Family-Group Names in Zoology).

  • — ICZN, 1996: 215, 216 (Platycnemini placed on Official List of Family Group Names in Zoolgy; it and other family group names based on Platycnemus not given priority over junior family-group names in general current usage in the Staphylininae).

Quediiformes Kraatz, 1857d: 473 (genera included: Tanygnathus, Acylophorus, Euryporus, Heterothops, Quedius, Astrapaeus, Cordylaspis, Homorocerus, Haematodes, Scariphaeus, and with question Caranistes and Palaestrinus). Type genus: Quedius (see Newton and Thayer, 1992: 65).

Quediides Thomson, 1859: 25 (genera included: Velleius, Quedius, Microsaurus, Raphirus, Euryporus, Heterothops, Acylophorus, Tanygnathus). Type genus: Quedius (see Newton and Thayer, 1992: 65).

  • — Newton and Thayer, 1992: 65 (synonym of Quediina).

Quedii Kraatz, 1857.

  • — Scheerpeltz, 1933: 1420 (world catalog supplement).

  • — Blackwelder, 1944: 143 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Seevers, 1965: 329 (myrmecophilus species).

  • — Coiffait, 1978: 7,9 (characters; cited as new subtribe).

Quediinae Kraatz, 1857.

  • — Casey, 1915: 396 (characters; discussion).

  • — Moore, 1964a: 89 (characters; notes).

  • — Moore and Legner, 1974g: 559 (key to genera of America north of Mexico).

  • — Moore and Legner, 1974h: 472 (catalog of the taxonomy, biology, and ecology of developmental stages of species of America north of Mexico).

  • — Moore and Legner, 1975: 48 (catalog; North America north of Mexico).

  • — Moore and Legner, 1979: 39 (characters; tabular key to genera of North America north of Mexico).

  • — E. Matthews, 1982: 7, 24 (note; key to genera of Australia).

  • — Lawrence and Newton, 1995: 830 (included in Staphylininae).

  • — Downie and Arnett, 1996: 387 (characters; key to genera of Northeastern North America).

Quediini Kraatz, 1857.

  • — Porta, 1907: 85 (characters; key to genera).

  • — Reitter, 1908: 101, 105, 106 (characters; key to genera).

  • — Eichelbaum, 1909: 184 (list of genera with references; generic synonyms; number of genera; number of species in each genus; general distribution of genera).

  • — Blatchley, 1910: 367 (characters).

  • — Casey, 1915: 397 (key to North American genera).

  • — Bernhauer and Schubert, 1916: 409 (world catalog).

  • — Porta, 1926: 108 (characters; species of Italy).

  • — Portevin, 1929: 334 (characters; key to French genera).

  • — Cameron, 1932: 2, 264 (characters; key to genera of British India).

  • — Joy, 1932: 106 (key to British genera).

  • — Scheerpeltz, 1933: 1418 (world catalog supplement).

  • — Scheerpeltz, 1940: 48 (key to Palaearctic genera).

  • — V. Hansen, 1952: 112 (characters; key to Danish genera).

  • — Smetana, 1958: 312 (characters; key to Czechoslovakian genera).

  • — Scheerpeltz, 1962b: 263 (key to Palaearctic genera).

  • — Palm, 1963b: 123 (characters; key to genera and species of Sweden).

  • — Lohse, 1964: 202 (characters; key to central European genera and species).

  • — Pototskaia, 1967: 75 (larval characters).

  • — Coiffait and Saiz, 1968: 365 (list of Chilean species).

  • — Scheerpeltz, 1968a: 86 (catalog of Austrian species).

  • — Kasule, 1970: 56 (larval characters; larval key to some genera).

  • — Saiz, 1971 (notes on Chilean taxa).

  • — Smetana, 1971 (revision of species and genera for America north of Mexico; key to genera and species; characters; collecting techniques).

  • — Smetana, 1977b: 178 (characters; notes on classification; key to genera of Holarctic region).

  • — Coiffait, 1978: 7 (characters; key to subtribes and genera of western Palaearctic region).

  • — Muona, 1979: 15 (list of Scandinavian species).

  • — Smetana, 1981a: 354 (key to genera of Chile).

  • — Newton, 1985: 208 (notes on austral distribution and taxonomic notes).

  • — Outerelo and Gamarra, 1985: 61 (key to genera of Iberian Peninsula).

  • — Dettner, 1987: 30 (defensive glands).

  • — Smetana, 1988: 169, 178, 179 (characters of genital segments; characters; key to genera).

  • — Newton, 1995: 51 (requested that the International Commission of Zoological Nomenclature place Gyrohypnini on the Official List of Family-Group Names in Zoology and that it and other family-group names based on Quedius be given precedence over those based on Platycnemus).

  • — Dettner, 1993a (summary and discussion of defensive secretions and exocrine glands and their bearing on phylogeny).

  • — Smetana, 1995: 14, 23 (characters; revision of species of Taiwan; key to genera and species of Taiwan; cladistic analysis).

  • — Newton, 1995: 51 (requested that the International Commission of Zoological Nomenclature Quediini Kraatz be given precedence over Platycnemini Nordmann).

  • — Cho, 1996b: 108 (key to genera of Korea).

  • — ICZN, 1996: 216 (Quediini placed on Official List of Family Group Names in Zoology; it and other family group names based on Quedius have precedence over Platycnemini and other family group names based on Platycnemus).

Acylophorini Scheerpeltz, 1968a: 97 (catalog of Austrian species; genera included: Atangynathus). Type genus: Acylophorus (see Newton and Thayer, 1992: 65).

  • — Outerelo and Gamarra, 1985: 67 (genera of Iberian Peninsula).

  • — Newton and Thayer, 1992: 65 (synonym of Quediina).

Heterothopsi Coiffait, 1978: 7, 300 (genera included: Heterothops, Atanygnathus). Type genus: Heterothops (see Newton and Thayer, 1992: 65).

  • — Smetana, 1984c: 278 (characters; discussion; probably not valid).

  • — Newton and Thayer, 1992: 65 (synonym of Quediina).

Acylophorini Outerelo and Gamarra, 1985: 48, 67 (genera included: Acylophorus). Type genus: Acylophorus (see Newton and Thayer, 1992: 65).

  • — Newton and Thayer, 1992: 65 (synonym of Quediina).

Acylohsellus
[1 species; Oriental Region]

Acylohsellus Smetana, 1995d: 114 (species included: longiceps). Type species: Acylophorus longiceps Cameron, fixed by original designation and monotypy.

longiceps Cameron, 1918b: 172 (Acylophorus; Type locality: Borneo, Mt. Merinjak, Sarawak).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • — Smetana, 1995d: 116 (Acylohsellus; characters; Borneo; Sumatra).

  • Distribution: Borneo, Indonesia.

Acylophorus [128 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, and Australian Regions]

Acylophorus Nordmann, 1837: 127 (species included: ahrensii, femoralis). Type species: Acylophorus ahrensii Nordmann, fixed by subsequent designation by Blackwelder, 1943: 466.

  • — Erichson, 1839a: 482 (characters).

  • — Erichson, 1840: 519 (characters).

  • — Heer, 1841: 584 (characters).

  • — Lacordaire, 1854: 84 (characters; notes; list of species).

  • — Fairmaire and Laboulbène, 1856: 534 (characters).

  • — Jacquelin du Val, 1856: 38 (characters).

  • — Kraatz, 1857d: 479 (characters).

  • — Redtenbacher, 1857: 199 (characters; key to Austrian species).

  • — Thomson, 1858: 30 (characters).

  • — Thomson, 1859: 26 (characters; type species: glabricollis).

  • — Schaum, 1859: 25 (catalog; European species).

  • — Thomson, 1860: 181 (characters).

  • — LeConte, 1861a: 64 (characters).

  • — LeConte, 1863: 23 (list of North American species).

  • — Wencker and Silbermann, 1866: 27 (list of species; collecting notes; France).

  • — Gemminger and Harold, 1868: 566 (catalog).

  • — Jacquelin du Val, 1868: 71 (catalog; European species).

  • — Sharp, 1871d: 11 (list of British species).

  • — Fauvel, 1874: 316 [= 1874a: 542] (characters).

  • — Redtenbacher, 1874: 195 (characters).

  • — Fauvel, 1875a: XXXIV [= 1875b: 236] (catalog).

  • — Mulsant and Rey, 1876: 831 [= 1877: 687] (characters; France).

  • — Sharp, 1876: 97 (notes).

  • — LeConte, 1878a: 388 (key to North American species).

  • — Horn, 1878: 150 (characters; key to North American species).

  • — Duvivier, 1883: 136 (catalog).

  • — Sharp, 1884: 324 (notes; Mexican and Central American species).

  • — Sharp, 1888: 222 (characters).

  • — Seidlitz, 1891: 400 (key to some European species).

  • — Seidlitz, 1891a: 427 (key to some European species).

  • — Heyden, Reitter, and Weise, 1891: 98 (list of species of Europe and Caucasus).

  • — Ganglbauer, 1895: 380 (characters).

  • — Heyden, Reitter, and Weise, 1906: 177 (list of species of Europe and Caucasus).

  • — Porta, 1907: 88 (key to Italian species).

  • — Reitter, 1908: 106 (characters).

  • — Blatchley, 1910: 368 (characters).

  • — Petri, 1912: 80 (list of species of Siebenbürgen).

  • — Johansen, 1914: 342 (characters; key to species).

  • — Casey, 1915: 426 (characters; key to species).

  • — Bernhauer and Schubert, 1916: 437 (world catalog; 37 species).

  • — Leng, 1920: 109 (catalog of North American species).

  • — Cameron, 1921a: 355, 405 (characters; key to Singapore species; catalog).

  • — Munster, 1923: 203 (key to Scandinavian species).

  • — Winkler, 1925: 393 (catalog for Palaearctic region).

  • — Cameron, 1925: 73 (catalog of Indian species).

  • — Porta, 1926: 119 (characters; species of Italy).

  • — Bruch, 1928a: 429 (catalog; Argentina).

  • — Portevin, 1929: 335 (characters; key to French species).

  • — Roubal, 1930: 397 (catalog; Slovakia).

  • — Cameron, 1932: 302 (characters; key to species of British India).

  • — Joy, 1932: 106 (key to British species).

  • — Scheerpeltz, 1933: 1466 (world catalog supplement).

  • — Normand, 1935: 373 (list of species of Tunisia).

  • — Bierig, 1938: 119–123 (characters; notes; illustration of variation of fourth segment of maxillary palpus and mandibles; type species).

  • — Blackwelder, 1943: 466 (characters; type species: ahrensii; key to species of West Indies).

  • — Blackwelder, 1944: 145 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Kloet and Hincks, 1945: 168 (list of British species).

  • — Tottenham, 1949: 377, 423 (type species: glaberrimus Herbst = ahrensii Nordmann; checklist of British species).

  • — Blackwelder, 1952: 38 (type species: ahrensii Nordmann).

  • — V. Hansen, 1952: 161 (characters; Danish species).

  • — Hatch, 1957: 231 (characters; species of Pacific Northwest).

  • — Kocher, 1958: 161 (checklist of species; Morocco).

  • — Smetana, 1958: 398 (characters; habitus of one species; type species: glaberrimus; key to Czechoslovakian species).

  • — Smetana, 1959a: 216 (list of species collected in Albania).

  • — Scheerpeltz, 1962b: 267 (characters).

  • — Székessy, 1963: 62 (characters).

  • — Palm, 1963b: 158 (characters; key to species of Sweden).

  • — Lohse, 1964: 220 (characters; key to central European species).

  • — Scheerpeltz, 1968a: 97 (catalog of Austrian species).

  • — Smetana, 1971: 246 (revision of and key to species of Canada and United States; characters).

  • — Shibata, 1973c: 64 (checklist of species of Taiwan).

  • — Tikhomirova, 1973a: 190 (checklist of species of USSR).

  • — Moore and Legner, 1975: 48 (catalog; North America north of Mexico).

  • — Smetana, 1977b: 181 (characters).

  • — Pope, 1977: 32 (list of British species).

  • — Coiffait, 1978: 296 (characters; key to species of western Palaearctic region).

  • — Moore and Legner, 1979: 41 (characters; notes).

  • — Muona, 1979: 16 (list of Scandinavian species).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 154 (catalog; Poland).

  • — Uhlig and Vogler, 1981: 126 (list of some species collected in Germany; habitat and collecting notes).

  • — Hammond, 1984: 204 (checklist of species of Borneo).

  • — Tóth, 1984: 93 (characters; habitus; key to species of Hungary).

  • — Shibata, 1984: 139 (subgenus of Acylophorus; checklist of species of Japan).

  • — Frank, 1986a: 371 (checklist of species of Florida).

  • — Smetana, 1988: 337 (characters; key to species of Himalayan region).

  • — Campbell and Davies, 1991: 122 (checklist of species; Alaska and Canada).

  • — Silfverberg, 1992: 18 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Smetana, 1993b: 50 (list of species of Czech Republic and Slovakia).

  • — Smetana, 1995: 125 (notes; revision of species of Taiwan; key to subgenera and species of Taiwan; cladistic analysis).

  • — Ciceroni and Zanetti, 1995: 33 (list of species of Italy).

  • — M. Hansen, 1996: 108 (list of Danish species).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 245 (additions to list of Danish species).

  • — Downie and Arnett, 1996: 394 (characters; key to species of Northeastern North America).

  • — Lundgren, 1998: 50 (list of species from Florida; records for other States listed for each species).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 392 (18 Nearctic species; notes; characters in key).

Rhygmacera Motschulsky, 1845a: 40 (species included: nitida). Type species: Rhygmacera nitida Motschulsky, fixed by original designation and monotypy.

  • — Motschulsky, 1858: 656 (synonym of Acylophorus).

  • — Fauvel, 1874: 316 [= 1874a: 542] (synonym of Acylophorus).

  • — Ganglbauer, 1895: 380 (synonym of Acylophorus).

  • — Bernhauer and Schubert, 1916: 438 (synonym of Acylophorus).

  • — Cameron, 1932: 302 (synonym of Acylophorus).

  • — Blackwelder, 1943: 466 (synonym of Acylophorus; type species: nitida).

  • — Blackwelder, 1952: 341 (type species: nitida).

  • — Smetana, 1958: 398 (synonym of Acylophorus).

  • — Smetana, 1971: 246 (synonym of Acylophorus).

  • — Coiffait, 1978: 296 (synonym of Acylophorus).

Paracylophorus Bierig, 1938: 123 (subgenus of Acylophorus; species included: schmidti). Type species: Acylophorus (Paracylophorus) schmidti Bierig, fixed by original designation and monotypy.

  • — Blackwelder, 1952: 289 (subgenus of Acylophorus; type species: schmidti).

Neoacylophorus Bierig, 1938: 123 (subgenus of Acylophorus; species included: oculifer). Type species: Acylophorus (Neoacylophorus) oculifer Bierig, fixed by original designation and monotypy.

  • — Blackwelder, 1952: 258 (subgenus of Acylophorus; type species: oculifer).

Indoacylophorus Bierig, 1938: 123 (subgenus of Acylophorus; species included: microcephalus). Type species: Acylophorus (Indoacylophorus) microcephalus Cameron, by original designation and monotypy.

  • — Blackwelder, 1952: 199 (subgenus of Acylophorus; type species: microcephalus).

Amacylophorus Smetana, 1971: 247 (subgenus of Acylophorus; species included: pratensis). Type species: Acylophorus (Amacylophorus) pratensis LeConte, fixed by original designation and monotypy.

Palpacylophorus Smetana, 1971: 247, 268 (subgenus of Acylophorus; species included: leechi). Type species: Acylophorus (Palpacylophorus) leechi Smetana, fixed by original designation and monotypy.

acuminatus Sharp, 1876: 99 (Acylophorus; Type locality: Ega).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Brazil.

aenescens Blackwelder, 1943: 468 (Acylophorus; Type locality: Dominican Republic: Sanchez).

  • Distribution: West Indies.

agilis Smetana, 1971: 262 (Acylophorus; subgenus Acylophorus; Type locality: Indiana: Parke Co., Inc. Also cited from Maryland, Massachusetts, Michigan, New York, North Carolina, Pennsylvania, Rhode Island, and Missouri).

  • — Smetana, 1971b: 4 (Acylophorus; Michigan).

  • — Smetana, 1973: 1434 (Acylophorus; notes; North Carolina).

  • — Smetana, 1978a: 836 (Acylophorus; collecting notes; Illinois; Kentucky).

  • — Smetana, 1990d: 103 (Acylophorus; Arkansas; New Jersey).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • Distribution: USA.

ahrensii Nordmann, 1837, see: glaberrimus Herbst, 1784.

allardi Levasseur, 1968: 1400 (Acylophorus; Type locality: Katanga: Luena, Haut-Lomami).

  • Distribution: Congo.

alternans Smetana, 1971: 256 (Acylophorus; subgenus Acylophorus; Type locality: New York: Staten Is. Also cited from Missouri, New Jersey, and Ontario).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • Distribution: USA.

anguliceps Bierig, 1938: 133 (Acylophorus; Type locality: Costa Rica: La Caja; Vara Blanca).

  • Distribution: Costa Rica.

angusticeps Sharp, 1876: 99 (Acylophorus; Type locality: Tapajos).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Brazil.

antennalis Cameron, 1932b: 135 (Acylophorus; Type locality: Lolo Damvu, Mayumbe).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • Distribution: Congo.

asperatus Fauvel, 1878g: 561 (Acylophorus; Type locality: Australie méridionale, Victoria).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Bernhauer, 1920: 18 (Acylophorus; Australia).

  • Distribution: Australia.

asperipennis Smetana, 1967i: 561 (Acylophorus; Type locality: Türkei: Gaziantep).

  • — Korge, 1971a: 32 (Acylophorus; synonym of glaberrimus).

  • — Coiffait, 1978: 299 (Acylophorus; cited as valid species; characters; Anatolia; Lebanon).

  • Distribution: Lebanon, Turkey.

bafutensis Levasseur, 1968: 1401 (Acylophorus; Type locality: Cameroun: forêt de Bafut, Nguemba).

  • Distribution: Cameroon.

balchhi Smetana, 1988: 350 (Acylophorus; Type locality: India: Meghalaya, Khasi Hills, 1000 m, Mawsynram-Balat).

  • Distribution: India.

beesoni Cameron, 1926b: 371 (Acylophorus; Type locality: Kaligad, Dehra Dun; [Note: See lectotype designation by Smetana, 1988]).

  • — Cameron, 1932: 304 (Acylophorus; characters; India).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • — Smetana, 1988: 341 (Acylophorus; lectotype designation: Kaligad, Dehra Dun; characters; Uttar Pradesh; Nepal).

  • Distribution: India, Nepal.

ventralis Coiffait, 1983d: 168 (Acylophorus; Type locality: Nepal: Bandha, vallée de Katmandou).

  • — Smetana, 1988: 342 (Acylophorus; synonym of beesoni).

bipunctatus Cameron, 1920a: 219 (Acylophorus; Type locality: Nilgiri Hills).

  • — Cameron, 1932: 308 (Anchocerus; characters; India).

  • — Scheerpeltz, 1933: 1466, 1467 (Acylophorus and Anchocerus; [Note: Name cited in two genera]; catalog).

  • — Smetana, 1988: 359 (Acylophorus; characters; India).

  • Distribution: India.

borneensis Cameron, 1933b: 348 (Acylophorus; Type locality: British North Borneo: Kamborangah, 7200 feet).

  • Distribution: Borneo.

breviceps Cameron, 1939b: 147 (Acylophorus; Type locality: New Guinea: Cyclops Mts.: Sabron, altitude 930 feet).

  • Distribution: New Guinea.

bruchi Bernhauer, 1912e: 178 (Acylophorus; Type locality: Argentinien: Prov. Buenos Aires).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Argentina.

bumbunae Bordoni, 1994a: 311 (Acylophorus; Type locality: Sierra Leone: Northern Province: presso Bumbuna).

  • Distribution: Sierra Leone.

capensis Cameron, 1945c: 716 (Acylophorus; Type locality: South Africa: Simon's Town).

  • Distribution: South Africa.

caseyi Leng, 1920: 109 (Acylophorus; replacement name for longicornis Casey).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • — Smetana, 1965: 49 (Acylophorus; Nova Scotia).

  • — Smetana, 1971: 253 (Acylophorus; subgenus Acylophorus; characters; Nova Scotia; Ontario; Alabama; Illinois; Massachusetts; New Jersey; New York; Pennsylvania; Wisconsin).

  • — Smetana, 1973: 1432 (Acylophorus; notes; Quebec).

  • — Smetana, 1976c: 183 (Acylophorus; Ontario; Quebec).

  • — Smetana, 1978a: 833 (Acylophorus; collecting notes; Louisiana; Wisconsin; Quebec).

  • — Smetana, 1990d: 102 (Acylophorus; Maine; West Virginia; Wisconsin).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • Distribution: Canada, USA.

longicornis Casey, 1915: 427 (Acylophorus; [preoccupied]; Type locality: New York: Peekskill).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; synonym of caseyi).

  • — Smetana, 1965: 49 (Quedius; synonym of caseyi).

  • — Smetana, 1971: 253 (Acylophorus; synonym of caseyi).

cephalotes Bierig, 1938: 124 (Acylophorus; Type locality: Panama: France Field, Zona del Canal).

  • Distribution: Panama.

charaa Smetana, 1988: 356 (Acylophorus; Type locality: Nepal: Lalitpur Distr., 2 km S Godavari, 1700 m).

  • Distribution: Nepal.

chillo Smetana, 1988: 343 (Acylophorus; Type locality: India: Assam, Manas, 200 m).

  • Distribution: India.

chontalenus Sharp, 1884: 326 (Acylophorus; Type locality: Nicaragua).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Nicaragua.

ciliatus Roth, 1851, see: Moeocerus.

collarti Cameron, 1935a: 375 (Acylophorus; Type locality: Nizi: Blukwa).

  • Distribution: Congo.

congener Bierig, 1938: 127 (Acylophorus; Type locality: Panama: France Field, Zona; Puerto Armuelles, Costa pacifica).

  • Distribution: Panama.

congoensis Cameron, 1932b: 134 (Acylophorus; Type locality: Blukwa, Nizi).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • — Bernhauer, 1942c: 364 (Acylophorus; Cameroon).

  • — Cameron, 1950g: 43 (Acylophorus; Belgian Congo).

  • Distribution: Congo.

daai Smetana, 1988: 348 (Acylophorus; Type locality: Nepal: Khandbari District, below Sheduwa, 2100–2500 m. Also cited from India: Uttar Pradesh).

  • Distribution: Nepal, India.

dampfi Bierig, 1938: 134 (Acylophorus; Type locality: Mexico: Tenancingo, Desierto del Convento, 2400 m).

  • Distribution: Mexico.

dankalensis Bordoni, 1994a: 311 (Acylophorus; Type locality: Sierra Leone: North Province: Monti Loma, Dankale, 800 m).

  • Distribution: Sierra Leone.

darlingtoni Blackwelder, 1943: 470 (Acylophorus; Type locality: Trinidad: Cedros).

  • — Frank, 1982: 33 (Acylophorus; list of parasites).

  • Distribution: West Indies.

delphinus Fauvel, 1905b: 181 (Acylophorus; Type locality: Pays Androy: Ambovombe).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Jarrige, 1965: 141 (Acylophorus; characters; Madagascar).

  • Distribution: Madagascar.

densipennis Bernhauer, 1929f: 125 (Acylophorus; Type locality: Haut-Uele: Moto; Ituri: La Moto: Madyu).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • — Cameron, 1950g: 43 (Acylophorus; Belgian Congo).

  • Distribution: Congo.

densus LeConte, 1878a: 387 (Acylophorus; Type locality: Enterprise).

  • — Horn, 1878: 152 (Acylophorus; characters; Florida; Maryland).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Smetana, 1971: 254 (Acylophorus; subgenus Acylophorus; characters; Florida; Louisiana).

  • — Smetana, 1978a: 833 (Acylophorus; collecting notes; Florida; Illinois; Louisiana; Texas).

  • — Smetana, 1990d: 102 (Acylophorus; Georgia).

  • Distribution: USA.

didymus Sharp, 1884: 325 (Acylophorus; Type locality: Guatemala, Dueñas).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Guatemala.

dissimilis Smetana, 1971: 259 (Acylophorus; subgenus Acylophorus; Type locality: California: Westwood).

  • — Smetana, 1973: 1432 (Acylophorus; notes; California).

  • — Smetana, 1978a: 833 (Acylophorus; collecting notes; California).

  • — Smetana, 1990d: 103 (Acylophorus; Colorado).

  • Distribution: USA.

fallax Smetana, 1971: 254 (Acylophorus; subgenus Acylophorus; Type locality: Florida: Orange Co.).

  • — Smetana, 1990d: 102 (Acylophorus; Florida; Georgia).

  • Distribution: USA.

femoralis Nordmann, 1837: 128 (Acylophorus; Type locality: Rio-Janeiro).

  • — Erichson, 1840: 520 (Acylophorus; characters; Brazil).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Brazil.

filius Smetana, 1971: 266 (Acylophorus; subgenus Acylophorus; Type locality: Pennsylvania: Pittsburg. Also cited from Illinois, Iowa, Massachusetts, New York, and Pennsylvania).

  • — Smetana, 1971b: 4 (Acylophorus; Michigan).

  • — Smetana, 1973: 1434 (Acylophorus; notes; Michigan).

  • — Puthz, 1974: 12 (Acylophorus, cited as Acylophorys; Illinois).

  • — Smetana, 1978a: 838 (Acylophorus; collecting notes; New Jersey).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • Distribution: USA.

flavicollis Sachse, 1852: 143 (Acylophorus; Type locality: Georgia).

  • — Horn, 1878: 151 (Acylophorus; characters; Pennsylvania to Kansas and Texas).

  • — Blatchley, 1910: 368 (Acylophorus; characters; Indiana).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Smetana, 1971: 267 (Acylophorus; subgenus Acylophorus; characters; Arkansas; Illinois; Indiana; Iowa; Kansas; KenÏtucky; Michigan; Maryland; Missouri; New Mexico; New York; Ohio; Oklahoma; Pennsylvania; South Dakota; Tennessee; Texas; Virginia).

  • — Smetana, 1971b: 4 (Acylophorus; Michigan).

  • — Smetana, 1978a: 838 (Acylophorus; collecting notes; Illinois; Kentucky; Texas).

  • — Smetana, 1981b: 643 (Acylophorus; Arkansas; Ontario).

  • — Frank, 1982: 33 (Acylophorus; list of parasites).

  • — Smetana, 1990d: 103 (Acylophorus; Mississippi).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • — Schmidt, 1999: 108 (Acylophorus; food for adults and larvae used for rearing species; rearing methods).

  • Distribution: USA.

flavicornis Cameron, 1918c: 214 (Acylophorus; Type locality: Borneo, Mt. Matang, W. Sarawak).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • Distribution: Borneo.

flavipes LeConte, 1878, see: lecontei Duvivier, 1883.

flavipes Motschulsky, 1858: 657 (Acylophorus; Type locality: Indes orientales; [Note: See lectotype designation by Smetana, 1988]).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Cameron, 1932: 304 (Acylophorus; characters; East Indies).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • — Cameron, 1934d: 79 (Acylophorus; type examined).

  • — Cameron, 1940: 89 (Rhygmacera; characters).

  • — Coiffait, 1982a: 234 (Acylophorus; Nepal).

  • — Smetana, 1988: 356 (Acylophorus; lectotype designation; characters; species may not occur in the Himalaya).

  • Distribution: “East Indies”.

fordi Last, 1987: 53 (Acylophorus; Type locality: Papua New Guinea: New Guinea NE: Sepalakambang, Salawaket Range, 1920 m).

  • Distribution: New Guinea.

franckei Wendeler, 1956b: 264 (Acylophorus; Type locality: Brasilien: Nova Teutonia).

  • Distribution: Brazil.

furcatus Motschulsky, 1858: 657 (Acylophorus; Type locality: Indes orientales; [Note: See lectotype designation by Smetana, 1988]).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Bernhauer, 1922c: 233 (Acylophorus; Formosa).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog; Philippines).

  • — Cameron, 1934d: 79 (Acylophorus; type examined).

  • — Smetana, 1988: 345 (Acylophorus; lectotype designation; characters; Assam; Meghalaya; West Bengal).

  • — Smetana, 1995: 125 (Acylophorus; characters; bionomics; Taiwan).

  • Distribution: “East Indies”, India, Philippines, Taiwan.

gilensis LeConte, 1863a: 34 (Acylophorus; Type locality: Gila River below Pima villages; [Note: See lectotype designation by Smetana, 1971]).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; synonym of pronus).

  • — Smetana, 1971: 257 (Acylophorus; subgenus Acylophorus; lectotype designation; characters; Arizona; Arkansas; California; New Mexico; Texas; Utah).

  • — Smetana, 1971a: 1848 (Acylophorus; collecting notes; Arizona).

  • Distribution: USA.

glaberrimus Herbst, 1784: 151 (Staphylinus; Type locality: Berlin; [Note: See lectotype designation by Gusarov, 1993]).

  • — Gmelin, 1790: 2032 (Staphylinus; page misnumbered as 1032; characters).

  • — Turton, 1802: 516 (Staphylinus; characters; Germany).

  • — Fauvel, 1886: 61 [= 1886a: 53] (Acylophorus; Algeria).

  • — Fauvel, 1889a: 112 (Acylophorus; notes; North America and Europe).

  • — Ganglbauer, 1895: 381 (Acylophorus; characters; Europe; Mediterranean region; North America; Mexico).

  • — Fauvel, 1897d: 311 (Acylophorus; Morocco).

  • — Fauvel, 1902b: 119 (Acylophorus; Morocco; Algeria).

  • — Porta, 1907: 99 (Acylophorus; characters; Italy).

  • — Reitter, 1909: 107 (Acylophorus; characters; Germany).

  • — Johansen, 1914: 342 (Acylophorus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Munster, 1923: 203 (Acylophorus; Fennoscandia).

  • — Lea, 1925: 252, 253 (Acylophorus; Australia, introduction).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • — Bierig, 1938: 123 (Acylophorus; type species of Acylophorus).

  • — Normand, 1947a: 9 (Acylophorus; Tunisia).

  • — V. Hansen, 1952: 162 (Acylophorus; characters; Denmark).

  • — Smetana, 1958: 399 (Acylophorus; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1960d: 136 (Acylophorus; Portugal).

  • — Palm, 1963b: 158 (Acylophorus; characters; Sweden; Denmark).

  • — Lohse, 1964: 220 (Acylophorus; characters; central Europe).

  • — M. Dvořák, 1965: 89 (Acylophorus; Slovakia).

  • — Horion, 1965: 308 (Acylophorus; Spain; France; Switzerland; Holland; Britain; Italy; Albania; Bosnia Hercegovina; Serbia; Bulgaria; Greece; Romania; Hungary; Germany; Austria; Slovakia; Czech Republic; Poland; Ukraine; Denmark; Sweden).

  • — Smetana, 1967i: 563 (Acylophorus; characters).

  • — Smetana, 1971: 247 (Acylophorus; species does not occur in North America, previous citations referred to pratensis and perhaps gilensis; [Note: One name, luctuosus Solsky, described from Mexico, is still listed as a synonym of glaberrimus as it was not studied with by Smetana]).

  • — Korge, 1971a: 32 (Acylophorus; characters; Turkey).

  • — Pope, 1977: 32 (Acylophorus; Britain).

  • — Coiffait, 1978: 298 (Acylophorus; characters; Europe; Britain; Scandinavia; North Africa).

  • — Tóth, 1984: 94 (Acylophorus; characters; Hungary).

  • — Ádám, 1987: 153 (Acylophorus; collecting notes; Hungary).

  • — Lucht, 1987: 111 (Acylophorus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Gusarov, 1993: 72 (Acylophorus; lectotype designation; synonyms; Georgia; Ukraine).

  • — Wittwer, 1993: 251 (Acylophorus; Switzerland).

  • — Smetana, 1993b: 50 (Acylophorus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 33 (Acylophorus; Italy).

  • — Schülke, 1998g: 90 (Acylophorus; Germany).

  • Distribution: Tunisia, Algeria, Morocco, Europe, Georgia, Turkey, Australia.

glabricollis Lacordaire, 1835: 396 (Staphylinus; Type locality: Paris).

  • — Erichson, 1839a: 482 (Acylophorus; characters; Germany).

  • — Erichson, 1840: 519 (Acylophorus; characters; Europe).

  • — Heer, 1841: 584 (Acylophorus; characters; Switzerland).

  • — Redtenbacher, 1849: 826 (Acylophorus; characters; Austria).

  • — Kiesenwetter, 1851a: 418 (Acylophorus; Spain).

  • — Fairmaire and Laboulbène, 1856: 534 (Acylophorus; characters; France).

  • — Kraatz, 1857d: 480 (Acylophorus; characters; Germany).

  • — Redtenbacher, 1857: 199 (Acylophorus; characters; Austria).

  • — Peyron, 1858: 428 (Acylophorus; Turkey).

  • — Thomson, 1859: 26 (type species of Acylophorus).

  • — Fauvel, 1874: 316 [= 1874a: 542] (Acylophorus; characters; France; Algeria; Cyprus; Caramania; Caucasus; USA; Mexico).

  • — Redtenbacher, 1874: 195 (Acylophorus; characters; Austria).

  • — Seidlitz, 1875: 263 (Acylophorus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 834 [= 1877: 690] (Acylophorus; characters; France).

  • — Fauvel, 1878: 131 [= 1878a: 51] (Acylophorus; Algeria).

  • — Fowler, 1888: 223 (Acylophorus; characters; Britain).

  • — Ganglbauer, 1895: 381 (Acylophorus; synonym of glaberrimus).

  • — Fauvel, 1902b: 119 (Acylophorus; synonym of glaberrimus).

  • — Porta, 1907: 99 (Acylophorus; synonym of glaberrimus).

  • — Montandon, 1908: 80 (Acylophorus; Romania).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; synonym of glaberrimus).

  • — Portevin, 1929: 335 (Acylophorus; characters; France).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; synonym of glaberrimus).

  • — Pope, 1977: 32 (Acylophorus; synonym of glaberrimus).

  • — Coiffait, 1978: 298 (Acylophorus; synonym of glaberrimus).

ahrensii Nordmann, 1837: 127 (Acylophorus; Type locality: Grunewald prope Berolinum).

  • — Erichson, 1839a: 482 (Acylophorus; synonym of glabricollis).

  • — Erichson, 1840: 519 (Acylophorus; synonym of glabricollis).

  • — Kraatz, 1857d: 480 (Acylophorus; synonym of glabricollis).

  • — Ganglbauer, 1895: 381 (Acylophorus; synonym of glaberrimus).

  • — Porta, 1907: 99 (Acylophorus; synonym of glaberrimus).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; synonym of glaberrimus).

  • — Smetana, 1958: 399 (Acylophorus; synonym of glaberrimus).

  • — Coiffait, 1978: 298 (Acylophorus; synonym of glaberrimus).

rufilabris Zetterstedt, 1838: 65 (Staphylinus; Type locality: Lapponia Umensis).

  • — Erichson, 1840: 519 (Acylophorus; synonym of glabricollis).

  • — Kraatz, 1857d: 480 (Acylophorus; synonym of glabricollis).

  • — Ganglbauer, 1895: 381 (Acylophorus; synonym of glaberrimus).

  • — Porta, 1907: 99 (Acylophorus; synonym of glaberrimus).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; synonym of glaberrimus).

  • — Smetana, 1958: 399 (Acylophorus; synonym of glaberrimus).

  • — Coiffait, 1978: 298 (Acylophorus; synonym of glaberrimus).

nitidus Motschulsky, 1845a: 40 (Rhygmacera; Type locality: Géorgie; [Note: See lectotype designation by Gusarov, 1993]).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • — Coiffait, 1978: 300 (Acylophorus; characters).

  • — Gusarov, 1993: 72 (Acylophorus; lectotype designation; synonym of glaberrimus).

luctuosus Solsky, 1868a: 122 (Acylophorus; Type locality: Mexique).

  • — Fauvel, 1874: 316 [= 1874a: 542] (Acylophorus; synonym of glabricollis).

  • — Sharp, 1884: 325 (Acylophorus; notes; Mexico).

  • — Ganglbauer, 1895: 381 (Acylophorus; synonym of glaberrimus).

  • — Porta, 1907: 99 (Acylophorus; synonym of glaberrimus).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; synonym of glaberrimus).

  • — Smetana, 1958: 399 (Acylophorus; synonym of glaberrimus).

  • — Coiffait, 1978: 298 (Acylophorus; synonym of glaberrimus).

pulcher Scriba, 1870: 81 (Acylophorus; Type locality: Santiago de Compostella Galaeciae).

  • — Fauvel, 1874: 316 [= 1874a: 542] (Acylophorus; synonym of glabricollis).

  • — Ganglbauer, 1895: 381 (Acylophorus; synonym of glaberrimus).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; synonym of glaberrimus).

  • — Smetana, 1958: 399 (Acylophorus; synonym of glaberrimus).

  • — Coiffait, 1978: 298 (Acylophorus; synonym of glaberrimus).

glabricollis Lacordaire, 1835, see: glaberrimus Herbst, 1784.

grandis Bernhauer, 1929f: 124 (Acylophorus; Type locality: Manyema: Niemba-Kalembe).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • Distribution: Congo.

hatuey Bierig, 1938: 126 (Acylophorus; Type locality: Cuba: Sierra del Rosario, El Rangel; Laguna del Somorrostro; Aspiro).

  • — Blackwelder, 1943: 469 (Acylophorus; characters; Cuba).

  • Distribution: West Indies.

hayashii Smetana, 1995: 129 (Acylophorus; Type locality: Chipon, Formosa).

  • Distribution: Taiwan.

hwang Smetana, 1995: 127 (Acylophorus; Type locality: Taiwan: Koantauchi, Nantou Hsien).

  • Distribution: Taiwan.

honshuensis R. Dvořák, 1957: 10 (Acylophorus; Type locality: Nagoya-Honshu, Japon).

  • Distribution: Japan.

hubrichi Bernhauer, 1927e: 249 (Acylophorus; Type locality: Argentinien: Prov. Santa Fe, Rosario).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • Distribution: Argentina.

immotus† Scudder, 1900: 47 (Acylophorus; fossil; Type locality: Florissant, Colorado).

  • — Handlirsch, 1907: 723 (Acylophorus; catalog; Miocene; Colorado).

  • Distribution: USA.

indignus Blackburn, 1888: 4 (Acylophorus; Type locality: near Adelaide, banks of the Torrens).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Australia.

insulanus Bierig, 1938: 125 (Acylophorus; Type locality: Cuba: El Mariel; Laguna La Canoa, Portugues; Isla de Pinos).

  • — Blackwelder, 1943: 468 (Acylophorus; characters; Cuba; Isle of Pines).

  • Distribution: USA.

iridescens Sharp, 1876: 100 (Acylophorus; Type locality: Tapajos).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Brazil.

iridiventris Bernhauer, 1927e: 248 (Acylophorus; Type locality: Brasilien: S. Catharina).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • Distribution: Brazil.

jalapanus Schubert, 1909: 289 (Acylophorus; Type locality: südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d'Orizaba).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Mexico.

javanus Cameron, 1937: 30 (Acylophorus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft).

  • Distribution: Indonesia.

khairo Smetana, 1988: 344 (Acylophorus; Type locality: India: Meghalaya, Garo Hills, Songsak, 400 m).

  • Distribution: India.

kraatzi Bernhauer, 1908d: 336 (Acylophorus; Type locality: Paraguay).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Paraguay.

kumbuiensis Bordoni, 1994a: 312 (Acylophorus; Type locality: Sierra Leone: Eastern Province: Kambui Hills Kenema).

  • Distribution: Sierra Leone.

leechi Smetana, 1971: 268 (Acylophorus; subgenus Palpacylophorus; Type locality: Missouri: Butler Co., stream at Hwy. 60, 12 mi. SE Ellsinore. Also cited from Texas).

  • Distribution: USA.

lecontei Duvivier, 1883: 136 (Acylophorus; replacement name for flavipes LeConte).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Smetana, 1971: 260 (Acylophorus; subgenus Acylophorus; characters; Arkansas; Florida; Georgia; Maryland Mississippi; New Jersey; Pennsylvania; South Carolina).

  • — Smetana, 1978a: 834 (Acylophorus; collecting notes; Alabama; Kentucky; Louisiana; Mississippi; North Carolina; Texas).

  • — Smetana, 1981b: 643 (Acylophorus; collecting notes; Florida).

  • — Frank, 1982: 33 (Acylophorus; list of parasites).

  • — Smetana, 1990d: 103 (Acylophorus; Arkansas; West Virginia).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • Distribution: USA.

flavipes LeConte, 1878a: 387 (Acylophorus; [preoccupied]; Type locality: Capron).

  • — Horn, 1878: 152 (Acylophorus; characters; Florida).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; synonym of lecontei).

  • — Smetana, 1971: 260 (Acylophorus; synonym of lecontei).

leonensis Bordoni, 1994a: 312 (Acylophorus; Type locality: Sierra Leone: Western Area: Sugar Loaf Mt.).

  • Distribution: Sierra Leone.

lindbergi Scheerpeltz, 1958a: 26 (Acylophorus; Type locality: Van, Südost-Ufer des Van-Sees in Armenien).

  • — Smetana, 1967i: 563 (Acylophorus; characters).

  • — Coiffait, 1978: 299 (Acylophorus; characters; Turkey).

  • Distribution: Turkey.

lomaensis Levasseur, 1971: 303 (Acylophorus; Type locality: monts Loma, Sierra Leone, 1600 m).

  • Distribution: Sierra Leone.

longiceps Cameron, 1918, see: Acylohsellus.

longicornis Casey, 1915, see: caseyi Leng, 1920.

longicornis Erichson, 1840: 520 (Acylophorus; Type locality: Cayenna).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: French Guiana.

longistylus Casey, 1915: 426 (Acylophorus; Type locality: Florida).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; synonym of pronus).

  • — Smetana, 1971: 261 (Acylophorus; subgenus Acylophorus; characters; Alabama; Florida; Louisiana; New Jersey; New York; Pennsylvania).

  • — Smetana, 1978a: 835 (Acylophorus; collecting notes; Florida; Illinois; Louisiana; Mississippi; Texas).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • Distribution: USA.

lualabaensis Levasseur, 1968: 1404 (Acylophorus; Type locality: Katanga: région de Kolwezi, Zilo, Lualaba).

  • Distribution: Congo.

luctuosus Solsky, 1868, see: glaberrimus Herbst, 1784.

maculicornis Cameron, 1918c: 214 (Acylophorus; Type locality: Borneo, Mt. Matang, W. Sarawak).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • Distribution: Borneo.

malaisei Scheerpeltz, 1965: 297 (Acylophorus; Type locality: N.E. Burma, Kambaiti, 2000 m).

  • Distribution: Myanmar.

mareei Bernhauer, 1943b: 290 (Acylophorus; Type locality: Kongo: Lokandu).

  • Distribution: Congo.

marginalis Cameron, 1948: 42 (Acylophorus; Type locality: Bassin du Chari, Rivière Gribingui).

  • — Tottenham, 1961: 205 (Acylophorus; Senegal).

  • Distribution: Chad, Senegal.

metallicus Blackwelder, 1943: 469 (Acylophorus; Type locality: Dominican Republic: Sanchez).

  • Distribution: West Indies.

methneri Bernhauer, 1917g: 48 (Acylophorus; Type locality: Deutsch-Ostafrika: Kaguru).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — Bernhauer and Chapman, 1939: 74 (Acylophorus; Kenya).

  • — Scheerpeltz, 1974c: 25 (Acylophorus; Sudan).

  • Distribution: Sudan, Kenya, Tanzania.

mexicanus Sharp, 1884: 326 (Acylophorus; Type locality: Mexico, Oaxaca, Las Vigas).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Mexico.

microcephalus Cameron, 1932: 305 (Acylophorus; Type locality: Mussoorie district: Arni Gad and Kali Gad; [Note: See lectotype designation by Smetana, 1988]. Also cited from Mussoorie district: Kali Gad).

  • — Cameron, 1932: 304 (Acylophorus; [Note: Misidentified as furcatus according to Smetana, 1988: 352]; characters; India).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — Bierig, 1938: 123 (Acylophorus; type species of Indoacylophorus).

  • — Smetana, 1988: 352 (Acylophorus; lectotype designation: Arni Gad, Mussoorie; characters; Uttar Pradesh; West Bengal).

  • Distribution: India.

microcerus Fauvel, 1895: 276 (Acylophorus; Type locality: Bírmanie, Bhamo).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • — Cameron, 1932: 305 (Acylophorus; characters; Burma).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — Scheerpeltz, 1976: 44 (Acylophorus; Nepal).

  • Distribution: Myanmar.

minutus Jarrige, 1965: 142 (Acylophorus; Type locality: Perinet, Madagascar Est).

  • Distribution: Madagascar.

muscorum Jarrige, 1965: 140 (Acylophorus; Type locality: massif du Tsaratanana, 1500 m).

  • Distribution: Madagascar.

nepa Bierig, 1938: 131 (Acylophorus; Type locality: Panama: Puerto Armuelles).

  • Distribution: Panama.

nepalicus Coiffait, 1981, see: Erichsonius.

nigeriae Tottenham, 1961: 205 (Acylophorus; [Note: Nomen nudum, cited from Senegal with Cameron as author but I find no description of the species by either Cameron or Tottenham (Article 13.1.1)]).

nitidus Motschulsky, 1845, see: glaberrimus Herbst, 1784.

obscurus Cameron, 1942a: 139 (Acylophorus; Type locality: Brunei).

  • Distribution: Brunei.

oculifer Bierig, 1938: 137 (Acylophorus; subgenus Neoacylophorus; Type locality: Panama: France Field, Zona).

  • Distribution: Panama.

orientalis Fauvel, 1907: 48 (Acylophorus; Type locality: Voi; Kisoumou [Victoria Nyanza]; baie de Kavirondo; steppe Masai pres Nairobi).

  • — Eichelbaum, 1913: 129 (Acylophorus; Tanzania).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • — Bernhauer, 1931d: 588 (Acylophorus; Abyssinia).

  • — Scheerpeltz, 1974c: 25 (Acylophorus; Sudan).

  • Distribution: Sudan, Ethiopia, Kenya, Tanzania.

pacificus Bierig, 1938: 128 (Acylophorus; Type locality: Panama: France Field, Zona; Puerto Armuelles, Costa pacifica).

  • Distribution: Panama.

palpalis Sharp, 1884: 327 (Acylophorus; Type locality: Guatemala, San Geronimo).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • — Bierig, 1938: 135 (Acylophorus; Panama).

  • Distribution: Guatemala, Panama.

parviceps Sharp, 1884: 326 (Acylophorus; Type locality: Guatemala, near the city).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Guatemala.

persimilis Cameron, 1950d: 24 (Acylophorus; Type locality: Malacca: Perak).

  • Distribution: Indonesia.

picipennis Roth, 1851, see: Glyphesthus.

picipennis Bernhauer, 1929f: 124 (Acylophorus; [preoccupied]; Type locality: Luebo).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • Distribution: Congo.

picipes Erichson, 1840: 521 (Acylophorus; Type locality: Columbia).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Colombia.

planatus Sharp, 1884: 327 (Acylophorus; Type locality: Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Nicaragua.

plaumanni Wendeler, 1956b: 263 (Acylophorus; Type locality: Brasilien: Nova Teutonia).

  • Distribution: Brazil.

pratensis LeConte, 1863a: 34 (Acylophorus; Type locality: Kansas).

  • — Fauvel, 1874: 316 [= 1874a: 542] (Acylophorus; synonym of glabricollis).

  • — Horn, 1878: 151 (Acylophorus; characters; Kansas).

  • — Ganglbauer, 1895: 381 (Acylophorus; synonym of glaberrimus).

  • — Porta, 1907: 99 (Acylophorus; synonym of glaberrimus).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; synonym of glaberrimus).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; synonym of glaberrimus; valid species?).

  • — Smetana, 1965: 50 (Acylophorus; characters; Newfoundland).

  • — Smetana, 1971: 248 (Acylophorus; subgenus Amacylophorus; characters; Newfoundland; Quebec; Massachusetts; Minnesota).

  • — Smetana, 1973: 1432 (Acylophorus; notes; Quebec).

  • — Smetana, 1981b: 643 (Acylophorus; collecting notes; Ontario).

  • — Smetana, 1990d: 102 (Acylophorus; New Hampshire).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • Distribution: Canada, USA.

princeps Smetana, 1971: 255 (Acylophorus; subgenus Acylophorus; Type locality: Florida: Homestead, Dade Co.).

  • — Smetana, 1981b: 643 (Acylophorus; Florida).

  • Distribution: USA.

pronus Erichson, 1840: 521 (Acylophorus; Type locality: America septentrionalis).

  • — Horn, 1878: 151 (Acylophorus; characters; USA).

  • — Blatchley, 1910: 368 (Acylophorus; characters; Indiana).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • — Notman, 1919: 135 (Acylophorus; New York).

  • — Notman, 1920a: 180 (Acylophorus; New York).

  • — Notman, 1920b: 706 (Acylophorus; Florida).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — Hatch, 1957: 231 (Acylophorus; characters; British Columbia; Washington).

  • — Smetana, 1965: 48 (Acylophorus; characters; Nova Scotia; Newfoundland).

  • — Smetana, 1965a: 17 (Acylophorus; Alaska).

  • — Smetana, 1971: 264 (Acylophorus; subgenus Acylophorus; characters; British Columbia; Manitoba; New Brunswick; Newfoundland; Northwest Territories; Nova Scotia; Ontario; Quebec; Alaska; Connecticut; Illinois; Massachusetts; Maine; Michigan; Minnesota; New Hampshire; New Jersey; New York; Oregon; Pennsylvania; Rhode Island; Washington; Wisconsin).

  • — Smetana, 1971b: 4 (Acylophorus; Michigan).

  • — Smetana, 1976c: 184 (Acylophorus; California; Washington).

  • — Frank, 1982: 33 (Acylophorus; list of parasites).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • Distribution: Canada, USA.

proximus Sharp, 1884: 325 (Acylophorus; Type locality: Guatemala, Dueñas, Paso Antonio).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Guatemala.

pulcher Scriba, 1870, see: glaberrimus Herbst, 1784.

puncticeps Fauvel, 1895: 275 (Acylophorus; Type locality: Bírmanie, Carin Ghécu, 1300–1400 m).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • — Cameron, 1932: 303 (Acylophorus; characters; Burma).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — Smetana, 1988: 359 (Acylophorus; characters; Burma).

  • Distribution: Myanmar.

punctiventris Sharp, 1876: 98 (Acylophorus; Type locality: Ega; Tapajos).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Brazil.

punctulatus Bierig, 1938: 130 (Acylophorus; Type locality: Costa Rica: San José, La Caja).

  • Distribution: Costa Rica.

pusillus Smetana, 1973: 1432 (Acylophorus; Type locality: U.S.A.: Florida: Lake Istokpoga, near Lake Placid).

  • Distribution: USA.

raato Smetana, 1988: 351 (Acylophorus; Type locality: Nepal: Khandbari District, Arun River at Num, 1500–1600 m. Also cited from India: Meghalaya; West Bengal).

  • — Smetana, 1992b: 10 (Acylophorus; Nepal).

  • Distribution: Nepal, India.

richteri Bernhauer, 1912e: 177 (Acylophorus; Type locality: Argentinien: Prov. Buenos).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Argentina.

rossii Bordoni, 1994a: 309 (Acylophorus; Type locality: Sierra Leone: Western Area: Sugar Loaf Mt.).

  • Distribution: Sierra Leone.

rotundicollis Cameron, 1918: 90 (Acylophorus; Type locality: Singapore: Bukit Timah).

  • — Cameron, 1921a: 379, 405 (Acylophorus; characters; Singapore).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • Distribution: Singapore.

ruficeps Cameron, 1941c: 383 (Acylophorus; Type locality: Philippines: Mindanao: Surigao).

  • Distribution: Philippines.

ruficollis Kraatz, 1859, see: ruficollis Motschulsky, 1858.

ruficollis Motschulsky, 1858: 657 (Acylophorus; Type locality: Indes orientales; [Note: See lectotype designation by Smetana, 1988]).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • — Bernhauer, 1920: 18 (Acylophorus; Australia).

  • — Wendeler, 1924a: 344 (Acylophorus; East Indies; Ceylon; Australia).

  • — Bernhauer, 1928: 18 (Acylophorus; Sumatra).

  • — Cameron, 1932: 303 (Acylophorus; characters; Ceylon; Penang).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — Cameron, 1933d: 393 (Acylophorus; Sumatra).

  • — Last, 1964: 177 (Acylophorus; Thailand).

  • — Smetana, 1988: 349 (Acylophorus; lectotype designation; characters; Assam; Nepal).

  • — Barrion, Litsinger, and Last, 1991: 1078 (Acylophorus; associated with rice agroecosystem).

  • Distribution: “East Indies”, Nepal, Sri Lanka, Malaysia, Thailand, Indonesia, Australia.

ruficollis Kraatz, 1859: 65 (Acylophorus; [preoccupied]; Type locality: Ceylan).

  • — Lea, 1925: 252, 253 (Acylophorus; characters; Australia, introduction).

  • — Cameron, 1932: 303 (Acylophorus; synonym of ruficollis).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; synonym of ruficollis).

  • — Smetana, 1988: 349 (Acylophorus; synonym of ruficollis).

rufilabris Zetterstedt 1838, see: glaberrimus Herbst, 1784.

rufipennis Cameron, 1959: 117 (Acylophorus; Type locality: Angola: Dundo, River Luachimo).

  • Distribution: Angola.

sandersoni Smetana, 1971: 258 (Acylophorus; subgenus Acylophorus; Type locality: Arkansas: Washington Co.).

  • Distribution: USA.

schmidti Bierig, 1938: 135 (Acylophorus; subgenus Paracylophorus; Type locality: Costa Rica: San José, La Caja).

  • Distribution: Costa Rica.

scutellaris Bernhauer, 1916b: 21 (Acylophorus; Type locality: Borneo).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — Cameron, 1933d: 393 (Acylophorus; Sumatra).

  • — Cameron, 1937: 30 (Acylophorus; Java).

  • Distribution: Borneo, Indonesia.

similis Wendeler, 1956b: 264 (Acylophorus; Type locality: Brasilien: Nova Teutonia).

  • Distribution: Brazil.

simplex Sharp, 1884: 325 (Acylophorus; Type locality: Mexico, Oaxaca, Mirador, Cordova).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Mexico.

siyo Smetana, 1988: 344 (Acylophorus; Type locality: Nepal: Khandbari Distr., 2 km E Mansingma, 1900 m).

  • — Smetana, 1992b: 10 (Acylophorus; Nepal).

  • Distribution: Nepal.

sogai Jarrige, 1965: 142 (Acylophorus; Type locality: Mahanilazy, Ambatondrazaka, Madagascar Est).

  • Distribution: Madagascar.

tenuiceps Bernhauer, 1932b: 157 (Acylophorus; Type locality: Haut-Uele: Yebo; Moto).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • Distribution: Congo.

tenuipes Lea, 1929: 207 (Acylophorus; Type locality: Queensland: Brisbane).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • Distribution: Australia.

tibialis Cameron, 1932: 305 (Acylophorus; Type locality: Burma: Ruby Mines; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — Scheerpeltz, 1965: 112 (Acylophorus; Burma).

  • — Smetana, 1988: 357 (Acylophorus; lectotype designation; characters; Burma).

  • Distribution: Myanmar.

trigonocephalus Cameron, 1949a: 322 (Acylophorus; Type locality: Côte d'Ivoire: Mt. Tonkoui, alt. 900–1200 m).

  • Distribution: Ivory Coast.

trinitatis Blackwelder, 1943: 467 (Acylophorus; Type locality: Trinidad: foothills north of Tunapuna).

  • Distribution: West Indies.

tristis Jarrige, 1965: 140 (Acylophorus; Type locality: Tananarive).

  • Distribution: Madagascar.

usambarae Bernhauer, 1917g: 48 (Acylophorus; Type locality: West-Usambara).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • Distribution: Tanzania.

ventralis Coiffait, 1983, see: beesoni Cameron, 1926.

versicolor Sharp, 1887: 783 (Acylophorus; Type locality: Mexico, Tehuantepec).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Mexico.

wagenschieberi Kiesenwetter, 1850: 220 (Acylophorus; Type locality: Berolini).

  • — Kiesenwetter, 1851a: 419 (Acylophorus; characters; Germany).

  • — Redtenbacher, 1857: 199 (Acylophorus; characters; Germany).

  • — Kraatz, 1857d: 481 (Acylophorus; characters; Germany).

  • — Thomson, 1860: 182 (Acylophorus; characters; Scandinavia).

  • — Fauvel, 1874: 316 [= 1874a: 542] (Acylophorus; characters; Scandinavia; Germany; Greece).

  • — Redtenbacher, 1874: 195 (Acylophorus; characters).

  • — Seidlitz, 1875: 263 (Acylophorus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 836 [= 1877: 692] (Acylophorus; characters; France).

  • — Ganglbauer, 1895: 382 (Acylophorus; characters; Germany; Greece; Sweden).

  • — Reitter, 1909: 107 (Acylophorus; characters; Germany).

  • — Johansen, 1914: 344 (Acylophorus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • — Munster, 1923: 203 (Acylophorus; Fennoscandia).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — V. Hansen, 1952: 162 (Acylophorus; characters; Denmark).

  • — Smetana, 1958: 401 (Acylophorus; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 158 (Acylophorus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 220 (Acylophorus; characters; central Europe).

  • — Horion, 1965: 311 (Acylophorus; Russia; Finland; Sweden; Norway; Denmark; Poland; Czechoslovakia; Holland; Austria; Germany).

  • — Smetana, 1967i: 563 (Acylophorus; characters).

  • — Coiffait, 1978: 300 (Acylophorus; characters; Europe; Greece).

  • — Tóth, 1984: 94 (Acylophorus; characters).

  • — Segers, 1987: 60 (Acylophorus; Belgium).

  • — Lucht, 1987: 111 (Acylophorus; checklist; Germany; Poland; Czechoslovakia; Austria; Benelux; Denmark; Sweden).

  • — Smetana, 1993b: 50 (Acylophorus; Czech Republic).

  • — Sterrenburg and Schülke, 1997: 20 (Acylophorus; Netherlands).

  • — Staneic, 1998a: 5 (Acylophorus; habitat; life cycle; Poland).

  • Distribution: Russia, Finland, Sweden, Norway, Denmark, Holland, Germany, Poland, Czechoslovakia, Austria.

williamsi Bierig, 1938: 129 (Acylophorus; Type locality: Brasil: Belen, Para).

  • Distribution: Brazil.

zdenae Smetana, 1978a: 836 (Acylophorus; Type locality: Texas: Trinity Co.: 12 mi. SW Lufkin. Also cited from Illinois and Louisiana).

  • — Smetana, 1990d: 103 (Acylophorus; Arkansas).

  • Distribution: USA.

ziloensis Levasseur, 1968: 1403 (Acylophorus; Type locality: Katanga: région de Kolwezi, Zilo, Lualaba).

  • Distribution: Congo.

Anaquedius [1 species; Nearctic Region]

Anaquedius Casey, 1915: 397, 400 (species included: vernix). Type species: Quedius vernix LeConte, fixed by original designation and monotypy.

  • — Casey, 1915: 397 (type species: Quedius vernix).

  • — Scheerpeltz, 1933: 1427 (subgenus of Quedius).

  • — Blackwelder, 1952: 50 (type species: vernix).

  • — Smetana, 1971: 242 (valid genus; characters).

  • — Moore and Legner, 1975: 51 (catalog; North America north of Mexico).

  • — Smetana, 1977b: 181 (characters).

  • — Moore and Legner, 1979: 42 (characters; notes).

  • — Frank, 1986a: 371 (checklist of species of Florida).

  • — Downie and Arnett, 1996: 394 (characters).

  • — Lundgren, 1998: 50 (list of species from Florida; records for other States listed for each species).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 392 (1 Nearctic species; notes; characters in key).

vernix LeConte, 1878a: 389 (Quedius; Type locality: Cambridge, Massachusetts; [Note: See lectotype designation by Smetana, 1971]. Originally cited from Massachusetts; Michigan; Canada).

  • — Horn, 1878: 167 (Quedius; characters; Massachusetts, Canada, Illinois, Michigan).

  • — Blatchley, 1910: 373 (Quedius; characters; Indiana).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1465 (Quedius; catalog).

  • — Smetana, 1965: 35 (Quedius; subgenus Anaquedius; Nova Scotia).

  • — Smetana, 1971: 242 (Anaquedius; lectotype designation: Cambr. [Cambridge, Massachusetts]; characters; New Brunswick; Nova Scotia; Ontario; Quebec; Connecticut; Illinois; Indiana; Louisiana; Maine; Maryland; Massachusetts; Michigan; Minnesota; New Hamshire; New Jersey; New York; Ohio; Pennsylvania; Rhode Island; Virginia; Wisconsin).

  • — Smetana, 1973: 1432 (Anaquedius; notes; North Carolina).

  • — Smetana, 1978a: 831 (Anaquedius; collecting notes; Florida; Louisiana; Texas).

  • — Frank, 1982: 33 (Anaquedius; list of parasites).

  • — Smetana, 1990d: 102 (Anaquedius; Arkansas).

  • — Campbell and Davies, 1991: 122 (Anaquedius; checklist of species; Canada).

  • — Downie and Arnett, 1996: 394 (Anaquedius; characters; USA).

  • Distribution: Canada, USA.

Anchocerus [10 species; Palaearctic, Oriental, and Australian Regions]

Anchocerus Fauvel, 1905a: 141 (species included: birmanus). Type species: Anchocerus birmanus Fauvel, fixed by original designation and monotypy.

  • — Bernhauer and Schubert, 1916: 439 (world catalog; 1 species).

  • — Cameron, 1925: 73 (catalog of Indian species).

  • — Cameron, 1932: 306 (characters; key to species of British India).

  • — Scheerpeltz, 1933: 1467 (world catalog supplement).

  • — Blackwelder, 1952: 51(type species: birmanus).

  • — Scheerpeltz, 1962b: 268 (characters).

  • — Last, 1975: 441 (characters).

  • — Smetana, 1988: 361 (characters; key to species of Himalayan region).

  • — Smetana, 1995: 132 (notes; species of Taiwan).

birmanus Fauvel, 1905a: 141 (Anchocerus; Type locality: Tenasserim, Kawkareet, Bírmanie; [Note: See lectotype designation by Smetana, 1977a]. Originally cited from Bírmanie. Tenasserim. Java: Buitenzorg; Malang).

  • — Fauvel, 1905e: 84 (Anchocerus; Java).

  • — Bernhauer and Schubert, 1916: 439 (Anchocerus; catalog).

  • — Bernhauer, 1922a: 151 (Anchocerus; Indochina).

  • — Cameron, 1932: 306 (Anchocerus; characters; Burma: Tenasserim).

  • — Scheerpeltz, 1933: 1467 (Anchocerus; catalog).

  • — Cameron, 1937: 30 (Anchocerus; Java).

  • — Smetana, 1977a: 249 (Anchocerus; lectotype designation: Tenasserim, Kawkareet, Bírmanie; characters).

  • — Smetana, 1988: 364 (Anchocerus; characters; Burma; Uttar Pradesh).

  • Distribution: India, Myanmar, Indonesia.

monticola Cameron, 1926b: 371 (Anchocerus; Type locality: Kaligad, Dehra Dun; [Note: See lectotype designation by Smetana, 1988]. Originally cited from Mussoorie district; Kaligad).

  • — Cameron, 1932: 307 (Anchocerus; characters; India).

  • — Scheerpeltz, 1933: 1467 (Anchocerus; catalog).

  • — Smetana, 1988: 363 (Anchocerus; lectotype designation: Kaligad, Dehra Dun; characters; India).

  • Distribution: India.

nepalicus Smetana, 1988: 366 (Anchocerus; Type locality: Nepal: Khandbari District: For. above Ahale, 2300 m. Also cited from India).

  • — Smetana, 1992b: 11 (Anchocerus; Nepal).

  • Distribution: India, Nepal.

novaguinensis Last, 1975: 442 (Anchocerus; Type locality: Okapa).

  • Distribution: New Guinea.

okasaensis Last, 1975: 441 (Anchocerus; Type locality: Okasa, 4000 ft.).

  • Distribution: New Guinea.

punctatissimus Smetana, 1977a: 248 (Anchocerus; Type locality: Bhutan: Changra, 18 km S. Tongsa).

  • — Smetana, 1988: 365 (Anchocerus; characters; Bhutan; Nepal).

  • Distribution: Bhutan, Nepal.

punctus Last, 1980: 151 (Anchocerus; Type locality: New Guinea: Wau, Mount Kumbak).

  • Distribution: New Guinea.

shibatai Smetana, 1995: 132 (Anchocerus; Type locality: Taiwan: near Lienhuachih, Nantou Hsien, 700 m).

  • Distribution: Taiwan.

similis Last, 1980: 150 (Anchocerus; Type locality: New Guinea: Friedrich Wilhelmshafen).

  • Distribution: New Guinea.

wilhelmensis Last, 1980: 150 (Anchocerus; Type locality: New Guinea: Mount Wilhelm, Field Stat., 3200 m).

  • Distribution: New Guinea.

Arrowinus [1 species; Ethiopian Region]

Arrowinus Bernhauer, 1935f: 215 (species included: phaenomenalis). Type species: Arrowinus phaenomenalis Bernhauer, fixed by original designation and monotypy.

  • — Blackwelder, 1952: 63 (type species: phaenomenalis).

  • — Newton, 1985: 208 (comments on austral distribution and taxonomic notes).

phaenomenalis Bernhauer, 1935f: 216 (Arrowinus; Type locality: Natal).

  • Distribution: South Africa.

Astrapaeus [1 species; Palaearctic Region]

Astrapaeus Gravenhorst, 1802: 199 (species included: ulmi). Type species: Staphylinus ulmi Rossi, fixed by monotypy.

  • — Latreille, 1804: 284, 288 (characters).

  • — Latreille, 1802: 130 (characters).

  • — Latreille, 1806: 284 (characters).

  • — Latreille, 1810: 179, 427 (characters; type species).

  • — Leach, 1815: 91 (characters).

  • — Billberg, 1820: 13 (characters).

  • — Dejean, 1821: 23 (species in collection).

  • — Latreille, 1825: 244 (cited as Astrapée; characters).

  • — Berthold, 1827: 331 (characters).

  • — Le Peletier and Serville, 1828: 478 (characters; notes).

  • — Curtis, 1829: 24 (catalog; Britain).

  • — Stephens, 1829a: 22 (catalog; Britain).

  • — Stephens, 1829: 276 (catalog; Britain).

  • — Latreille, 1829: 433 (character).

  • — Mannerheim, 1830: 7, 19 (characters).

  • — Mannerheim, 1831: 421, 433 (characters).

  • — Stephens, 1832: 200, 213 (characters).

  • — Griffith and Pidgeon, 1832: 290 (characters).

  • — Dejean, 1833: 61 (in list of species).

  • — Gistel, 1834: 8 (in list of species).

  • — Lacordaire, 1835: 355 (characters).

  • — Nordmann, 1837: 16 (characters).

  • — Westwood, 1838a: 16 (characters; type species).

  • — Heer, 1839: 282 (characters).

  • — Stephens, 1839: 384 (characters).

  • — Erichson, 1840: 552 (characters).

  • — Laporte, 1840: 173 (characters).

  • — Redtenbacher, 1849: 712 (characters).

  • — Schaum, 1852: 27 (in list of species; Europe).

  • — Lacordaire, 1854: 86 (characters; notes; in list of species).

  • — Fairmaire and Laboulbène, 1856: 545 (characters).

  • — Jacquelin du Val, 1856: 40 (characters).

  • — Kraatz, 1857d: 521 (characters).

  • — Redtenbacher, 1857: 205 (characters; key to Austrian species).

  • — G. Waterhouse, 1858: 23 (catalog of British species).

  • — Schaum, 1859: 26 (catalog; European species).

  • — Gredler, 1863: 101 (in list of species from Tirol).

  • — Wencker and Silbermann, 1866: 28 (in list of species; collecting notes; France).

  • — Gemminger and Harold, 1868: 573 (catalog).

  • — Jacquelin du Val, 1868: 71 (catalog; European species).

  • — Fauvel, 1874: 313 [= 1874a: 539] (characters).

  • — Redtenbacher, 1874: 202 (characters).

  • — Fauvel, 1875a: XXXIV [= 1875b: 236] (catalog).

  • — Mulsant and Rey, 1876: 798 [= 1877: 654] (characters).

  • — Heyden, Reitter, and Weise, 1891: 99 (in list of species of Europe and Caucasus).

  • — Ganglbauer, 1895: 383 (characters).

  • — Heyden, Reitter, and Weise, 1906: 176 (species of Europe and Caucasus).

  • — Porta, 1907: 89 (key to Italian species).

  • — Reitter, 1908: 106 (characters).

  • — Petri, 1912: 80 (in list of species of Siebenbürgen).

  • — Bernhauer and Schubert, 1916: 414 (world catalog; 1 species).

  • — Winkler, 1925: 388 (catalog for Palaearctic region).

  • — Porta, 1926: 118 (characters).

  • — Portevin, 1929: 336 (characters; key to French species).

  • — Roubal, 1930: 386 (catalog; Slovakia).

  • — Scheerpeltz, 1933: 1425 (world catalog supplement).

  • — Blackwelder, 1952: 66 (type species: ulmi).

  • — Smetana, 1958: 322 (characters; habitus; type species: ulmi).

  • — Smetana, 1959a: 215 (species collected in Albania).

  • — Scheerpeltz, 1962b: 264 (characters; type species: ulmi).

  • — Székessy, 1963: 62 (characters).

  • — Lohse, 1964: 205 (characters; central European species).

  • — Scheerpeltz, 1968a: 87 (catalog of Austrian species).

  • — Bordoni, 1973j: 713 (in list of species from islands near Sicily).

  • — Tikhomirova, 1973a: 188 (in checklist of species of USSR).

  • — Smetana, 1977b: 181 (characters).

  • — Coiffait, 1978: 294 (characters; cited as Astrapeus).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 118 (catalog; Poland).

  • — Tóth, 1984: 97 (characters; habitus; species of Hungary).

  • — Drugmand 1987b (comments on distribution of Belgian species).

  • — Smetana, 1993b: 49 (in list of species of Czech Republic; cited as Astrapeus).

  • — Ciceroni and Zanetti, 1995: 31 (species of Italy).

  • — Sparacio, 1995: 164 (characters; notes; Sicily).

Systolastes Gistel, 1856: 388 (species included: ulmi). Type species: Staphylinus ulmi Rossi, fixed by monotypy as “Systolastes ulmi Pan.”

  • — Blackwelder, 1952: 368 (type species; synonym of Astrapaeus).

rufipennis Stephens, 1832, see: ulmi Rossi, 1790.

rufipes Latreille, 1806, see: Tasgius.

ulmi Rossi, 1790: 248 (Staphylinus; Type locality: Etruria).

  • — Olivier, 1795:(42): 17 (Staphylinus; characters; Italy).

  • — Gravenhorst, 1802: 199 (Astrapaeus; characters).

  • — Panzer, 1805: 4 (Astrapaeus; characters; illustrations).

  • — Gravenhorst, 1806: 47 (Staphylinus).

  • — Latreille, 1806: 284 (Astrapaeus; characters).

  • — Leach, 1815: 91 (Astrapaeus; characters; Italy; France).

  • — Le Peletier and Serville, 1828: 479 (notes).

  • — Mannerheim, 1830: 19 (Astrapaeus; southern Europe).

  • — Mannerheim, 1831: 433 (Astrapaeus; southern Europe).

  • — Griffith and Pidgeon, 1832: 290 (Astrapaeus).

  • — Nordmann, 1837: 16 (Astrapaeus; southern Europe).

  • — Westwood, 1838a: 16 (Astrapaeus; characters; type species).

  • — Heer, 1839: 282 (Astrapaeus; characters; Switzerland).

  • — Erichson, 1840: 553 (Astrapaeus; characters; France; Italy; Dalmatia; Hungary).

  • — Küster, 1847: 47 (Astrapaeus; characters; Germany; France; Italy; Dalmatia; Hungary).

  • — Hochhuth, 1849: 155 (Astrapaeus; characters; Helenendorf).

  • — Redtenbacher, 1849: 712 (Astrapaeus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 545 (Astrapaeus; characters; France).

  • — Kraatz, 1857d: 523 (Astrapaeus; characters; Germany).

  • — Redtenbacher, 1857: 205 (Astrapaeus; characters; Austria).

  • — Kraatz, 1858a: 56 (Astrapaeus; Greece).

  • — Fauvel, 1874: 314 [= 1874a: 540] (Astrapaeus; characters; France; Germany; Switzerland; Austria; Italy; Sardinia; Greece; Russia; Caucasus).

  • — Redtenbacher, 1874: 202 (Astrapaeus; characters; Austria).

  • — Mulsant and Rey, 1876: 801 [= 1877: 657] (Astrapaeus; characters; France).

  • — Seidlitz, 1891a: 420 (Astrapaeus; characters; Europe).

  • — Ganglbauer, 1895: 384 (Astrapaeus; characters; southern middle Europe; Mediterranean region).

  • — Sainte-Claire Deville, 1906: 104 (Astrapaeus; Corsica).

  • — Porta, 1907: 101 (Astrapaeus; characters; Italy).

  • — Montandon, 1908: 80 (Astrapaeus; Romania).

  • — Reitter, 1909: 107 (Astrapaeus; characters; Germany).

  • — Bernhauer and Schubert, 1916: 415 (Astrapaeus; catalog).

  • — Portevin, 1929: 336 (Astrapaeus; characters; France).

  • — Scheerpeltz, 1933: 1425 (Astrapaeus; catalog).

  • — Smetana, 1958: 322 (Astrapaeus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 205 (Astrapaeus; characters; central Europe).

  • — Raianu, 1964: 102 (Astrapaeus; Romania).

  • — Scheerpeltz, 1965b: 406 (Astrapaeus; Sicily).

  • — Scheerpeltz, 1965c: 511 (Astrapaeus; Sicily).

  • — Horion, 1965: 251 (Astrapaeus, cited as Astrapeus; Portugal; Spain; Corsica; Italy; Croatia; Bosnia Hercegovina; Montenegro; Albania; Greece; Corfu; Albania; Turkey; Slovenia; Serbia; Hungary; Czech Republic; Slovakia; Poland ?; Ukraine; Switzerland; France; Belgium; Austria; Germany ?).

  • — Bordoni, 1974b: 14 (Astrapaeus; Italy).

  • — Coiffait, 1978: 294 (Astrapaeus; characters; Europe; Britain; Scandinavia; North Africa).

  • — Tóth, 1984: 98 (Astrapaeus; characters; Hungary).

  • — Segers, 1986: 52 (Astrapaeus; checklist of species; Belgium).

  • — Lucht, 1987: 109 (Astrapaeus; checklist; Germany; Poland; Czechoslovakia; Austria; France; Benelux).

  • — Smetana, 1993b: 49 (Astrapaeus, cited as Astrapeus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Astrapaeus; Italy).

  • Distribution: Europe, England, Russia, Caucasus.

ulmineus Fabricius, 1801: 595 (Staphylinus; Type locality: Italia).

  • — Panzer, 1805: 4 (Astrapaeus; synonym of ulmi).

  • — Gravenhorst, 1806: 47 (Staphylinus; synonym of ulmi Rossi).

  • — Guérin-Méneville, 1829: pl. 9, fig. 2 (Astrapaeus; illustration).

  • — Lacordaire, 1835: 356 (Astrapaeus; characters; France).

  • — Erichson, 1840: 553 (Astrapaeus; synonym of ulmi).

  • — Laporte, 1840: 173 (Astrapaeus; characters; Europe and northern Asia).

  • — Guérin-Méneville, 1844a: 30 (Astrapaeus; France).

  • — Fairmaire and Laboulbène, 1856: 545 (Astrapaeus; synonym of ulmi).

  • — Kraatz, 1857d: 523 (Astrapaeus; synonym of ulmi).

  • — Ganglbauer, 1895: 384 (Astrapaeus; synonym of ulmi).

  • — Porta, 1907: 101 (Astrapaeus; synonym of ulmi).

  • — Bernhauer and Schubert, 1916: 415 (Astrapaeus; synonym of ulmi).

  • — Smetana, 1958: 323 (Astrapaeus; synonym of ulmi).

  • — Coiffait, 1978: 295 (Astrapaeus; synonym of ulmi).

rufipennis Stephens, 1832: 214 (Astrapaeus; Type locality: Coombe-wood; Devonshire).

  • — Curtis, 1829: 24 (Astrapaeus; nomen nudum; catalog; Britain).

  • — Stephens, 1839: 389 (Ocypus; characters; England).

  • — G. Waterhouse, 1858: 23 (Astrapaeus; synonym of ulmi).

  • — Gemminger and Harold, 1868: 573 (Astrapaeus; synonym of ulmi).

  • — Ganglbauer, 1895: 384 (Astrapaeus; synonym of ulmi).

  • — Porta, 1907: 101 (Astrapaeus; synonym of ulmi).

  • — Bernhauer and Schubert, 1916: 415 (Astrapaeus; synonym of ulmi).

  • — Smetana, 1958: 323 (Astrapaeus; synonym of ulmi).

  • — Coiffait, 1978: 295 (Astrapaeus; synonym of ulmi).

ulmineus Fabricius, 1801, see: ulmi Rossi, 1790.

Beeria [1 species; Nearctic Region]

Beeria Hatch, 1957: 214, 231 (species included: punctata). Type species: Beeria punctata Hatch, fixed by original designation and monotypy.

  • — Korge, 1971a: 36 (characters).

  • — Moore and Legner, 1975: 8 (catalog; North America north of Mexico).

  • — Smetana, 1977b: 181 (characters).

  • — Smetana, 1978a: 816 (characters).

  • — Smetana, 1995j: 131 (related to Strouhalium and Heinzia).

  • — Moore and Legner, 1979: 44 (characters; notes).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 392 (1 Nearctic species; notes; characters in key).

nematocera Casey, 1915: 436 (Philonthus; Type locality: British Columbia: Metlakatla).

  • — Scheerpeltz, 1933: 1352 (Philonthus; catalog).

  • — Hatch, 1957: 198 (Philonthus; characters; British Columbia).

  • — Smetana, 1977b: 184 (Beeria; characters; ecological notes; British Columbia; Oregon; Washington).

  • — Smetana, 1978a: 815 (Beeria; notes; collecting notes; British Columbia; Oregon; Washington).

  • — Smetana, 1981b: 631 (Beeria; Alaska).

  • — Smetana, 1990d: 95 (Beeria; British Columbia; Washington).

  • — Campbell and Davies, 1991: 120 (Beeria; checklist of species; Canada).

  • Distribution: Canada, USA.

punctata Hatch, 1957: 231 (Beeria; Type locality: Oregon: Benton Co.: Mary's Peak).

  • — Smetana, 1977b: 184 (Beeria; synonym of nematocera).

  • — Smetana, 1978a: 815 (Beeria; synonym of nematocera).

punctata Hatch, 1957, see: nematocera Casey, 1915.

Bolitogyrus [29 species; Neotropical, Palaearctic, and Oriental Regions]

Bolitogyrus Chevrolat, 1842: 641 (species included: buphthalmus). Type species: Quedius buphthalmus Erichson, fixed by monotypy (see Herman, 2001).

  • — Blackwelder, 1952: 82 (type species: cribripennis).

  • — Hammond, 1984: 205 (checklist of species of Borneo).

  • — Smetana, 1988: 314 (characters; species of Himalayan region).

  • — Smetana, 1995: 118 (revision of species of Taiwan; key to species of Taiwan).

  • — Smetana, 2000b: 329 (key to species of mainland China).

  • — Herman, 2001: 8 (discussion of availability of and type species for Bolitogyrus).

Cyrtothorax Kraatz, 1858: 366 (species included: sallei, nigricoxis, erythrurus). Type species: Cyrtothorax sallei Kraatz, fixed by subsequent designation by Scheerpeltz, 1974a: 175, 180 [Note: Blackwelder, 1952: 115 accepted C. buphthalmus Erichson as the type species designation by R. Lucas, 1920: 222, but that name was not among the originally included nominal species].

  • — Gemminger and Harold, 1868: 567 (catalog).

  • — Fauvel, 1878b: 163 [= 1878c: 83] (notes; key to species).

  • — Duvivier, 1883: 137 (catalog).

  • — Sharp, 1884: 340 (notes; Mexican and Central American species).

  • — Bernhauer and Schubert, 1916: 415 (world catalog; 9 species).

  • — Cameron, 1932: 277 (characters; key to species of British India).

  • — Scheerpeltz, 1933: 1426 (world catalog supplement).

  • — Blackwelder, 1944: 144 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 115 (type species).

  • — Scheerpeltz, 1962b: 265 (characters).

  • — Scheerpeltz, 1974a: 175, 176 (type species designation; notes; key to species).

  • — Shibata, 1986a: 125 (supplemental list of species from Taiwan).

  • — Smetana, 1988: 314 (synonym of Bolitogyrus).

  • — Smetana, 1995: 118 (synonym of Bolitogyrus).

bechyneorum Scheerpeltz, 1974a: 181, 189 (Cyrtothorax; Type locality: El Salvador: Gebiet des Vulcan San Vincente, Finca la Paz, Ahuachapan, Apaneca).

  • Distribution: El Salvador.

borneensis Cameron, 1942a: 138 (Cyrtothorax; Type locality: Martapura).

  • Distribution: Borneo.

bullatus Sharp, 1884: 340 (Cyrtothorax; Type locality: Panama, Boquete).

  • — Bernhauer and Schubert, 1916: 415 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 182 (Cyrtothorax; characters; Panama).

  • Distribution: Panama.

buphthalmus Erichson, 1840: 534 (Quedius; Type locality: Mexico).

  • — Fauvel, 1878b: 164 [= 1878c: 84] (Cyrtothorax; characters; Mexico).

  • — Sharp, 1884: 340 (Cyrtothorax; characters; notes; Mexico; Guatemala).

  • — Bernhauer and Schubert, 1916: 415 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 180 (Cyrtothorax; characters; Mexico; Guatemala; El Salvador).

  • Distribution: Mexico, Guatemala, El Salvador.

sallei Kraatz, 1858: 367 (Cyrtothorax; Type locality: Mexico).

  • — Gemminger and Harold, 1868: 567 (Cyrtothorax; synonym of buphthalmus).

  • — Bernhauer and Schubert, 1916: 415 (Cyrtothorax; synonym of buphthalmus).

  • — Scheerpeltz, 1974a: 180 (Cyrtothorax; synonym of buphthalmus).

nigricoxis Kraatz, 1858: 367 (Cyrtothorax; variety? of sallei; Type locality: Not cited).

caesareus Bernhauer, 1915f: 146 (Cyrtothorax; Type locality: Sarawak: Mts. Matang).

  • — Bernhauer and Schubert, 1916: 415 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 179 (Cyrtothorax; characters; Borneo).

  • Distribution: Borneo.

carnifex Fauvel, 1878b: 165 [= 1878c: 85] (Cyrtothorax; Type locality: Cambodge).

  • — Bernhauer and Schubert, 1916: 415 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 180 (Cyrtothorax; characters; Cambodia).

  • Distribution: Cambodia.

costaricensis Wendeler, 1927: 8 (Cyrtothorax; Type locality: Costa Rica: Turrialba).

  • — Scheerpeltz, 1933: 1426 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 181 (Cyrtothorax; characters; Costa Rica).

  • Distribution: Costa Rica.

cyanescens Sharp, 1884: 341 (Cyrtothorax; Type locality: Guatemala, Capetillo, Cerro Zunil 4000 to 5000 feet).

  • — Bernhauer and Schubert, 1916: 416 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 181 (Cyrtothorax; characters; Guatemala).

  • Distribution: Guatemala.

cyanipennis Zheng, 1988: 306, 308 (Cyrtothorax; Type locality: China: Sichuan, Wolong, alt. 1830 m).

  • — Smetana, 2000b: 329 (Bolitogyrus; characters in key).

  • Distribution: China.

doesburgi Scheerpeltz, 1974a: 178, 187 (Cyrtothorax; Type locality: Java: Djawa; Gg. Moerie Tjolo).

  • Distribution: Indonesia.

electus Smetana and Zheng, 2001: 465 (Bolitogyrus; replacement name for elegans Smetana and Zheng).

  • Distribution: China.

elegans Smetana and Zheng, 2000: 59 (Bolitogyrus; [Preoccupied]; Type locality: China: N. Yunnan, Yulongshan mts., 2500–2800 m, Ganhaizi/Lijiang road).

  • — Smetana, 2000b: 329 (Bolitogyrus; characters in key).

  • — Smetana and Zheng, 2001: 465 (Bolitogyrus; synonym of electus).

elegans Cameron, 1937: 28 (Cyrtothorax; Type locality: Java: Batoerraden, G. Slamat).

  • — Scheerpeltz, 1974a: 179 (Cyrtothorax; characters; Java).

  • Distribution: Indonesia.

elegans Smetana and Zheng, 2000, see: electus Smetana and Zheng, 2001.

erythrurus Kraatz, 1858: 368 (Cyrtothorax; Type locality: Nova Granada).

  • — Fauvel, 1878b: 164 [= 1878c: 84] (Cyrtothorax; New Granada).

  • — Bernhauer and Schubert, 1916: 416 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 182 (Cyrtothorax; characters; Colombia).

  • Distribution: Colombia.

fukienensis Scheerpeltz, 1974a: 177, 183 (Cyrtothorax; Type locality: Provinz Fukien in Südostchina, in Kuatun, Tschang-Sen).

  • — Smetana and Zheng, 2000: 55 (Bolitogyrus; characters; notes; China).

  • — Smetana, 2000b: 330 (Bolitogyrus; characters in key).

  • Distribution: China.

fulgidus Sharp, 1884: 341 (Cyrtothorax; Type locality: Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1916: 416 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 182 (Cyrtothorax; characters; Nicaragua).

  • Distribution: Nicaragua.

kitawakii Smetana and Zheng, 2000: 56 (Bolitogyrus; Type locality: China: NE Sichuan, Chengkou Xian Daba Shan abv. Bashan, 1600–1900 m).

  • — Smetana, 2000b: 330 (Bolitogyrus; characters in key).

  • Distribution: China.

nevermanni Scheerpeltz, 1974a: 181, 190 (Cyrtothorax; Type locality: Costarica: Carpintera).

  • Distribution: Costa Rica.

nigricoxis Kraatz, 1858, see: sallei Kraatz, 1858.

nigropolitus Smetana, 2000b: 327 (Bolitogyrus; Type locality: China: W. Sichuan: Ya'an Prefecture: Tianquan Co., Jiain Shan, Tal oberh. Labahe N. R. St., 57 km W Ya'an, 30°06N, 102°25E, 1800 m).

  • Distribution: China.

octomaculatus Cameron, 1937: 28 (Cyrtothorax; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft.).

  • — Scheerpeltz, 1974a: 177 (Cyrtothorax; characters; Java).

  • Distribution: Indonesia.

pictus Smetana and Zheng, 2000: 62 (Bolitogyrus; Type locality: China: Yun'an. okr. Puzenya, 900 m).

  • — Smetana, 2000b: 330 (Bolitogyrus; characters in key).

  • Distribution: China.

proximus Cameron, 1942a: 138 (Cyrtothorax; Type locality: Martapura).

  • Distribution: Borneo.

rufipennis Cameron, 1937: 27 (Cyrtothorax; Type locality: Java: Batoerraden, G. Slamat; Pengalengan, alt. 4000 ft.).

  • — Scheerpeltz, 1974a: 179 (Cyrtothorax; characters; Java).

  • Distribution: Indonesia.

rufomaculatus Shibata, 1979: 26 (Cyrtothorax; Type locality: near Lushan, about 1200 m alt., Nantou Hsien).

  • — Smetana, 1995: 118 (Bolitogyrus; characters; bionomics; Taiwan).

  • Distribution: Taiwan.

sallei Kraatz, 1858, see: buphthalmus Erichson, 1840.

salvini Sharp, 1884: 341 (Cyrtothorax; Type locality: Guatemala, Zapotes).

  • — Bernhauer and Schubert, 1916: 416 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 183 (Cyrtothorax; characters; Guatemala).

  • Distribution: Guatemala.

signatus Cameron, 1932: 278 (Cyrtothorax; Type locality: Ceylon: Dikoya, alt 3800–4200 feet).

  • — Scheerpeltz, 1933: 1426 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 178 (Cyrtothorax; characters; Ceylon).

  • Distribution: Sri Lanka.

strigifrons Wendeler, 1928a: 34 (Cyrtothorax; Type locality: Mexico: Jalapa).

  • — Scheerpeltz, 1933: 1426 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 183 (Cyrtothorax; characters; Mexico).

  • Distribution: Mexico.

taiwanensis Hayashi, 1991a: 45 (Cyrtothorax; Type locality: Taiwan: Taitung Hsien: Taitung).

  • — Smetana, 1995: 121 (Bolitogyrus; characters; bionomics; Taiwan).

  • — Smetana, 1996g: 27 (Bolitogyrus; notes; Taiwan).

  • Distribution: Taiwan.

vietnamensis Scheerpeltz, 1974a: 178, 185 (Cyrtothorax; Type locality: Nordvietnam, Bao Lac, nördlich von Hanoi).

  • Distribution: Vietnam.

vulneratus Fauvel, 1878b: 165 [= 1878c: 85] (Cyrtothorax; Type locality: Cochinchine).

  • — Bernhauer and Schubert, 1916: 416 (Cyrtothorax; catalog).

  • — Cameron, 1932: 277 (Cyrtothorax; characters; Burma: Mt. Mooleyit. Cochin China).

  • — Scheerpeltz, 1933: 1426 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 177 (Cyrtothorax; characters; Burma; Indochina).

  • — Biswas, 1986: 128 (Cyrtothorax; India).

  • — Smetana, 1988: 315 (Bolitogyrus; characters; Meghalaya; West Bengal; Nepal; Vietnam).

  • Distribution: Vietnam, Myanmar, Nepal, India.

Cheilocolpus [16 species; Neotropical Region]

Cheilocolpus Solier, 1849: 320 (species included: pyrostomus, angustatus, parvus). Type species: Staphylinus pyrostomus Solier, fixed by subsequent designation by Blackwelder, 1943: 99.

  • — Fauvel, 1874: 207 [= 1874: 433] (synonym of Philonthus).

  • — Bernhauer and Schubert, 1914: 326 (synonym of Philonthus).

  • — Cameron, 1932: 62 (synonym of Philonthus).

  • — Blackwelder, 1943: 399 (synonym of Philonthus; type species: pyrostomus).

  • — Blackwelder, 1952: 99 (type species).

  • — Smetana, 1958: 143 (synonym of Philonthus).

  • — Coiffait and Saiz, 1966: 386 (characters; key to Chilean species).

  • — Coiffait and Saiz, 1968: 365 (list of Chilean species).

  • — Saiz, 1971: 365, 387, 389 (characters; key to species of Chile; notes on biogeography; notes on phenetic relationships).

angustatus Solier, 1849: 320 (Staphylinus; [preoccupied]; Type locality: Chile).

  • — Kraatz, 1859a: 5 (Staphylinus; subgenus Cheilocolpus is a Philonthus; notes).

  • — Fairmaire and Germain, 1861: 433 (Staphylinus; characters; Chile).

  • — Fauvel, 1864a: 126 (Othius; characters; Chile).

  • — Fauvel, 1866a: 348 [= 1866b: 99] (Othius; characters; Chile).

  • — Germain, 1903: 417 (Heterothops; characters; Chile).

  • — Bernhauer and Schubert, 1916: 411 (Heterothops; catalog).

  • — Coiffait and Saiz, 1966: 397 (Cheilocolpus; characters; Chile).

  • — Saiz, 1969a: 9 (Cheilocolpus; notes).

  • — Saiz, 1971: 365, 370, 377 (Cheilocolpus; characters; Chile).

  • — Herman, 2001: 43 (Cheilocolpus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: Chile.

cryptobioides Fairmaire and Germain, 1861: 439 (Lithocharis; Type locality: Aculco).

  • — Bernhauer and Schubert, 1916: 411 (Heterothops; synonym of angustatus).

  • — Coiffait and Saiz, 1966: 398 (Cheilocolpus; synonym of angustatus).

brevipennis Germain, 1903: 424 (Heterothops; Type locality: selvas subandinas de Chillan).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Coiffait and Saiz, 1966: 392 (Cheilocolpus; characters; Chile).

  • — Saiz, 1971: 367 (Cheilocolpus; characters; Chile).

  • Distribution: Chile.

centralis Saiz, 1971: 375 (Cheilocolpus; Type locality: Chile: El Arrayán, Santiago).

  • Distribution: Chile.

consors Fauvel, 1866a: 340 [= 1866b: 91] (Quedius; Type locality: Cordilliere de Santiago).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Coiffait and Saiz, 1966: 393 (Cheilocolpus; characters; Chile).

  • — Saiz, 1971: 367, 370, 373 (Cheilocolpus; characters; Chile).

  • Distribution: Chile.

cryptobioides Fairmaire and Germain, 1861, see: angustatus Solier, 1849.

denticulatus Saiz, 1971: 379 (Cheilocolpus; Type locality: Chile: Bosque Relicto de Quinter, Valparaíso).

  • Distribution: Chile.

fulvicollis Fairmaire and Germain, 1861: 435 (Baptolinus; Type locality: Concepcion).

  • — Germain, 1903: 417 (Heterothops; characters; notes; Chile).

  • — Bernhauer and Schubert, 1916: 411 (Heterothops; synonym of angustatus).

  • — Coiffait and Saiz, 1966: 400 (Cheilocolpus; characters; Chile).

  • — Saiz, 1971: 365, 370 (Cheilocolpus; characters; Chile).

  • Distribution: Chile.

impressifrons Solier, 1849: 316 (Staphylinus; Type locality: Chile: provincia de Santiago).

  • — Kraatz, 1859a: 5 (Philonthus; notes).

  • — Fairmaire and Germain, 1861: 430 (Philonthus; characters; Chile).

  • — Fauvel, 1864a: 125 (Philonthus; notes; Chile).

  • — Fauvel, 1866a: 343 [= 1866b: 94] (Philonthus; characters; Chile).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Coiffait and Saiz, 1966: 396 (Cheilocolpus; characters; Chile).

  • — Coiffait and Saiz, 1968: 365 (Cheilocolpus; synonym of pyrostoma).

  • — Saiz, 1969a: 9 (Cheilocolpus; notes).

  • — Saiz, 1971: 367, 370, 374 (Cheilocolpus; characters; Chile).

  • — Frank, 1982: 34 (Cheilocolpus; list of parasites).

  • Distribution: Chile.

levigatus Saiz, 1971: 371 (Cheilocolpus; Type locality: Malacohuello, Sud du Chili).

  • Distribution: Chile.

magallanicus Saiz, 1971: 374 (Cheilocolpus; Type locality: Chile: Seno Otway, Rio El Ganso, Magallanes).

  • Distribution: Chile.

nanus Saiz, 1971: 377 (Cheilocolpus; Type locality: Chile: Nahuelbuta, Mallaco).

  • Distribution: Chile.

punctatus Saiz, 1971: 381 (Cheilocolpus; Type locality: Chile: Nahuelbuta, Malleco).

  • Distribution: Chile.

pygmaeus Saiz, 1971: 379 (Cheilocolpus; Type locality: Chile: Curacaví, Los Cerrillos, Santiago).

  • Distribution: Chile.

pyrostomus Solier, 1849: 320 (Staphylinus; Type locality: Chile: Illapel).

  • — Kraatz, 1859a: 5 (Staphylinus; subgenus Cheilocolpus is a Philonthus; notes).

  • — Fairmaire and Germain, 1861: 429 (Quedius; characters; Chile).

  • — Fauvel, 1864a: 125 (Quedius; notes; Chile).

  • — Fauvel, 1866a: 341 [= 1866b: 92] (Quedius; characters; Chile).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • — Coiffait and Saiz, 1966: 389 (Cheilocolpus; characters; Chile).

  • — Saiz, 1969: 3, 4 (Cheilocolpus; characters; collecting notes; Chile).

  • — Saiz, 1969a: 9 (Cheilocolpus; notes).

  • — Saiz, 1971: 367, 370, 371 (Cheilocolpus; characters; Chile).

  • — Saiz, 1971a (Cheilocolpus; ecological notes).

  • — Saiz, Solervicens, and Ojeda, 1989: 52 (Cheilocolpus; collecting notes; habitus; Chile).

  • Distribution: Chile.

tenuis Saiz, 1971: 375 (Cheilocolpus; Type locality: Chile: Misituni, Tarapacá, 4000 m).

  • Distribution: Chile.

valdiviensis Coiffait and Saiz, 1966: 394 (Cheilocolpus; Type locality: Chili: Valdivia, 180 m, 40°05′S, 73°13′W).

  • — Saiz, 1971: 367, 370, 373 (Cheilocolpus; characters; Chile).

  • Distribution: Chile.

vicunianus Germain, 1903: 426 (Heterothops; Type locality: Requínoa).

  • — Bernhauer and Schubert, 1916: 414 (Heterothops; catalog).

  • — Coiffait and Saiz, 1966: 392 (Cheilocolpus; characters; Chile).

  • — Saiz, 1971: 367 (Cheilocolpus; characters; Chile).

  • — Scheerpeltz, 1972: 23 (Heterothops; [Note: I assume Scheerpeltz overlooked transfer of this name to Cheilocolpus]; Argentina).

  • Distribution: Argentina, Chile.

Ctenandropus [2 species; Palaearctic, Oriental, Australian, and Oceanic Regions]

Ctenandropus Cameron, 1926b: 348 (species included: nigriceps). Type species: Ctenandropus nigriceps Cameron, fixed by original designation and monotypy.

  • — Cameron, 1932: 268 (characters).

  • — Scheerpeltz, 1933: 1425 (world catalog supplement).

  • — Blackwelder, 1952: 113 (type species: nigriceps).

  • — Scheerpeltz, 1962b: 264 (characters; type species: nigriceps).

  • — Smetana, 1988: 334 (characters; species of Himalayan region).

nigriceps Cameron, 1926b: 349 (Ctenandropus; Type locality: Asarori, Siwaliks; [Note: See lectotype designation by Smetana, 1988]. Originally cited from Siwaliks; Asarori, Timli. Andaman Islands, Port Blair).

  • — Cameron, 1927c: 264 (Ctenandropus; Fiji; Sumatra).

  • — Cameron, 1932: 268 (Ctenandropus; characters; India; Andaman Islands; Malay Peninsula; Sumatra; Philippines).

  • — Scheerpeltz, 1933: 1425 (Ctenandropus; catalog).

  • — Cameron, 1933d: 392 (Ctenandropus; Sumatra).

  • — Cameron, 1933e: 83 (Ctenandropus; Society Islands).

  • — Cameron, 1936: 6 (Ctenandropus; Society Islands; Rapa).

  • — Coiffait, 1976: 218 (Ctenandropus; Tahiti).

  • — Legner and Moore, 1977: 175 (Ctenandropus; cited as nicriceps; found under bark).

  • — Smetana, 1988: 334 (Ctenandropus; lectotype designation: Asarori, Siwaliks; characters; Uttar Pradesh).

  • — Lecoq, 1990: 207 (Ctenandropus; Mascarene Islands).

  • Distribution: India, Andaman Islands, Malaysia, Indonesia, Philippines, Fiji, Society Islands, Rapa Islands.

magniceps Bernhauer, 1920: 17 (Heterothops; Type locality: Herberton).

  • — Lea, 1925: 231 (Heterothops; Australia).

  • — Bernhauer, 1928: 18 (Heterothops; Sumatra).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Cameron, 1933d: 392 (Heterothops; Sumatra).

  • — Smetana, 1988: 334 (Ctenandropus).

  • Distribution: Australia, Indonesia.

Euristus [1 species; Ethiopian Region]

Euristus Fauvel, 1899: 23 (species included: globus). Type species: Euristus globus Fauvel, fixed by original designation and monotypy.

  • — Bernhauer and Schubert, 1916: 409 (world catalog; 1 species).

  • — Blackwelder, 1952: 158 (type species: globus).

globus Fauvel, 1899: 24 (Euristus; Type locality: Gabon: Loango).

  • — Bernhauer and Schubert, 1916: 409 (Euristus; catalog).

  • Distribution: Gabon.

Euryporus [6 species; Palaearctic Oriental, and Australian Regions]

Euryporus Erichson, 1839a: 496 (replacement name for Pelecyphorus Nordmann). Type species: Oxyporus picipes Paykull, fixed by objective synonymy with Pelecyphorus.

  • — Heer, 1839: 282 (characters).

  • — Stephens, 1839: 384 (characters).

  • — Erichson, 1840: 553 (characters).

  • — Redtenbacher, 1849: 712 (characters).

  • — Schaum, 1852: 27 (in list of species; Europe).

  • — Lacordaire, 1854: 86 (characters; notes; in list of species).

  • — Fairmaire and Laboulbène, 1856: 545 (characters).

  • — Jacquelin du Val, 1856: 39 (characters).

  • — Kraatz, 1857d: 481 (characters).

  • — Redtenbacher, 1857: 205 (characters; species of Austria).

  • — Thomson, 1858: 30 (characters).

  • — G. Waterhouse, 1858: 22 (catalog of British species).

  • — Thomson, 1859: 26 (characters; type species: picipes).

  • — Schaum, 1859: 25 (catalog; European species).

  • — Thomson, 1860: 179 (characters).

  • — LeConte, 1861a: 64 (characters).

  • — Wencker and Silbermann, 1866: 27 (in list of species; collecting notes; France).

  • — Gemminger and Harold, 1868: 566 (catalog).

  • — Jacquelin du Val, 1868: 71 (catalog; European species).

  • — Sharp, 1871d: 11 (in list of British species).

  • — Fauvel, 1874: 315 [= 1874a: 541] (characters).

  • — Redtenbacher, 1874: 196 (characters).

  • — Fauvel, 1875a: XXXIV [= 1875b: 236] (catalog).

  • — Mulsant and Rey, 1876: 803 [= 1877: 659] (characters).

  • — Duvivier, 1883: 136 (catalog).

  • — Sharp, 1888: 223 (characters).

  • — Ganglbauer, 1895: 382 (character).

  • — Heyden, Reitter, and Weise, 1906: 176 (in list of species of Europe and Caucasus).

  • — Porta, 1907: 88 (key to Italian species).

  • — Reitter, 1908: 106 (characters).

  • — Petri, 1912: 80 (in list of species of Siebenbürgen).

  • — Johansen, 1914: 344 (characters).

  • — Bernhauer and Schubert, 1916: 414 (world catalog; 5 species).

  • — Winkler, 1925: 388 (catalog for Palaearctic region).

  • — Porta, 1926: 118 (characters; key to species of Italy).

  • — Portevin, 1929: 335 (characters; key to French species).

  • — Roubal, 1930: 386 (catalog; Slovakia).

  • — Joy, 1932: 107 (key to British species).

  • — Scheerpeltz, 1933: 1424 (world catalog supplement).

  • — Normand, 1935: 372 (cited as Euriporus; in list of species of Tunisia).

  • — Kloet and Hincks, 1945: 167 (in list of British species).

  • — Tottenham, 1949: 376, 421 (type species: picipes Paykull; in checklist of British species).

  • — Blackwelder, 1952: 160 (type species: picipes).

  • — V. Hansen, 1952: 160 (characters; Danish species).

  • — Blackwelder, 1952: 160 (type species).

  • — Kocher, 1958: 156 (in checklist of species; Morocco).

  • — Smetana, 1958: 320 (characters; habitus of one species; type species: picipes; Czechoslovakian species).

  • — Scheerpeltz, 1962b: 264 (characters; type species: picipes).

  • — Székessy, 1963: 61 (characters).

  • — Palm, 1963b: 157 (characters; species of Sweden).

  • — Lohse, 1964: 204 (characters; central European species).

  • — Scheerpeltz, 1968a: 87 (catalog of Austrian species).

  • — Kasule, 1970: 57 (larval characters).

  • — Bordoni, 1973j: 713 (in list of species from islands near Sicily).

  • — Tikhomirova, 1973a: 188 (in checklist of species of USSR).

  • — Smetana, 1977b: 181 (characters).

  • — Pope, 1977: 31 (in list of British species).

  • — Coiffait, 1978: 290 (characters; key to species of western Palaearctic region).

  • — Muona, 1979: 15 (in list of Scandinavian species).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 117 (catalog; Poland).

  • — Uhlig and Vogler, 1981: 122 (in list of some species collected in Germany; collecting notes).

  • — Tóth, 1984: 94 (characters; habitus).

  • — Drugmand 1987b (comments on distribution of Belgian species).

  • — Silfverberg, 1992: 17 (in list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Siitonen, 1993: 230 (in list of Finnish species).

  • — Smetana, 1993b: 49 (in list of species of Czech Republic and Slovakia).

  • — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 30 (additional records to Danish list of species).

  • — Ciceroni and Zanetti, 1995: 31 (in list of species of Italy).

  • — M. Hansen, 1996: 108 (in list of Danish species).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 245 (additions to list of Danish species).

Pelecyphorus Nordmann, 1837: 13 (species included: picipes; preoccupied by Dejean, 1834). Type species: Oxyporus picipes Paykull, fixed by monotypy.

  • — Erichson, 1839a: 496 (synonym of Euryporus).

  • — Lacordaire, 1854: 86 (synonym of Euryporus).

  • — Jacquelin du Val, 1856: 39 (synonym of Euryporus).

  • — Fauvel, 1874: 315 [= 1874a: 541] (synonym of Euryporus).

  • — Ganglbauer, 1895: 382 (synonym of Euryporus).

  • — Bernhauer and Schubert, 1916: 414 (synonym of Euryporus).

  • — Tottenham, 1949: 376 (synonym of Euryporus; type species: picipes).

  • — Blackwelder, 1952: 296 (synonym of Euryporus; type species: picipes).

  • — Smetana, 1958: 320 (synonym of Euryporus).

aeneiventris P. Lucas, 1846: 115 (Euryporus; Type locality: Algeria: environs du moulin pres Lacalle; Sicile).

  • — Fauvel, 1878: 130 [= 1878a: 50] (Euryporus; Egypt; Algeria).

  • — Fauvel, 1886: 61 [= 1886a: 53] (Euryporus; Morocco; Algeria).

  • — Fauvel, 1897d: 310 (Euryporus; cited as Eurporus; Barbary).

  • — Fauvel, 1902b: 118 (Euryporus; Morocco; Algeria; Tunisia).

  • — Sainte-Claire Deville, 1906: 104 (Euryporus; Corsica).

  • — Porta, 1907: 100 (Euryporus; characters; Italy).

  • — Bernhauer and Schubert, 1916: 414 (Euryporus; catalog).

  • — Normand, 1947a: 9 (Euryporus, cited as Euriporus; Tunisia).

  • — Kocher, 1958: 156 (Euryporus; Morocco).

  • — Scheerpeltz, 1961d: 132 (Euryporus; Italy).

  • — Scheerpeltz, 1965b: 406 (Euryporus; Sicily).

  • — Scheerpeltz, 1965c: 511 (Euryporus; Sicily).

  • — Coiffait, 1968c: 505 (Euryporus; Corsica).

  • — Coiffait, 1978: 293 (Euryporus; characters; Spain; Italy; Corsica; North Africa).

  • — Sparacio, 1995: 163 (Euryporus; characters; summary of distribution; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 31 (Euryporus; Italy).

  • Distribution: Egypt, Tunisia, Algeria, Morocco, Spain, Italy.

meridionalis Baudi, 1848: 133 (Euryporus; Type locality: Sardinia).

  • — Gemminger and Harold, 1868: 566 (Euryporus; synonym of aeneiventris).

  • — Fauvel, 1902b: 118 (Euryporus; synonym of aeneiventris).

  • — Porta, 1907: 100 (Euryporus; synonym of aeneiventris).

  • — Bernhauer and Schubert, 1916: 414 (Euryporus; synonym of aeneiventris).

  • — Coiffait, 1978: 293 (Euryporus; synonym of aeneiventris).

argentatus Fauvel, 1881: 164 (Euryporus; Type locality: Moeara Laboe, Sumatra).

  • — Bernhauer and Schubert, 1916: 414 (Euryporus; catalog).

  • Distribution: Indonesia.

flavipes Fauvel, 1884, see: Pammegus.

meridionalis Baudi, 1848, see: aeneiventris P. Lucas, 1846.

multicavus Last, 1980: 154 (Euryporus; Type locality: New Guinea: Kiunga).

  • Distribution: New Guinea.

picipes Paykull, 1800: 426 (Oxyporus; Type locality: Suecia).

  • — Gyllenhal, 1810: 443 (Oxyporus; characters; Sweden).

  • — Olivier, 1811: 608 (Oxyporus; characters; Sweden).

  • — Nordmann, 1837: 14 (Pelecyphorus; Germany; Sweden).

  • — Erichson, 1839a: 496 (Euryporus; characters; Germany).

  • — Heer, 1839: 283 (Heterothops; characters; Switzerland).

  • — Stephens, 1839: 389 (Euryporus; characters; England).

  • — Erichson, 1840: 554 (Euryporus; characters; Sweden; Germany).

  • — Heer, 1841: 584 (Euryporus; Switzerland).

  • — Redtenbacher, 1849: 712 (Euryporus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 545 (Euryporus; characters; France).

  • — Kraatz, 1857d: 482 (Euryporus; characters; Germany).

  • — Redtenbacher, 1857: 205 (Euryporus; characters; Austria).

  • — Thomson, 1859: 26 (type species of Euryporus).

  • — Thomson, 1860: 180 (Euryporus; characters; Scandinavia).

  • — Fauvel, 1874: 315 [= 1874a: 541] (Euryporus; characters; France; Lapland; Scandinavia; Russia; Germany; Britain; Switzerland).

  • — Redtenbacher, 1874: 196 (Euryporus; characters; Austria).

  • — Seidlitz, 1875: 268 (Euryporus; characters; Baltic region).

  • — Fauvel, 1876a: 79 [= 1876: 264] (Euryporus; Spain).

  • — Mulsant and Rey, 1876: 806 [= 1877: 662] (Euryporus; characters; France).

  • — J. Sahlberg, 1876: 30 (Euryporus; Finland).

  • — Fowler, 1888: 223 (Euryporus; characters; Britain).

  • — Seidlitz, 1891: 395 (Euryporus; characters; Europe).

  • — Seidlitz, 1891a: 420 (Euryporus; characters; Europe).

  • — Ganglbauer, 1895: 383 (Euryporus; characters; north and middle Europe).

  • — Porta, 1907: 100 (Euryporus; characters; Italy).

  • — Reitter, 1909: 107 (Euryporus; characters; Germany).

  • — Johansen, 1914: 345 (Euryporus; characters; Denmark).

  • — Munster, 1923: 185 (Euryporus; Fennoscandia).

  • — Portevin, 1929: 336 (Euryporus; characters; France).

  • — Scheerpeltz, 1933: 1425 (Euryporus; catalog).

  • — V. Hansen, 1952: 160 (Euryporus; characters; Denmark).

  • — Smetana, 1958: 320 (Euryporus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 204 (Euryporus; characters; central Europe).

  • — Horion, 1965: 250 (Euryporus; Russia; Finland; Sweden; Norway; Denmark; Baltics; Poland; Czechoslovakia; Hungary; Romania; Italy; Switzerland; France; Belgium; Holland; Germany; Austria).

  • — Kasule, 1970: 57 (Euryporus?; larval characters).

  • — Pope, 1977: 31 (Euryporus; Britain).

  • — Coiffait, 1978: 291 (Euryporus; characters; Europe; Scandinavia; Britain).

  • — Tóth, 1984: 95 (Euryporus; characters).

  • — Segers, 1986: 52 (Euryporus; checklist of species; Belgium).

  • — Lucht, 1987: 108 (Euryporus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Smetana, 1993b: 49 (Euryporus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Euryporus; Italy).

  • — Sterrenburg and Schülke, 1997: 20 (Euryporus; Netherlands).

  • — Owen, 1999: 10 (Euryporus; collecting notes; Britain).

  • — Eyre, Luff, and Lott, 2000: 33 (Euryporus; Britain).

  • Distribution: Europe, Russia.

princeps Wollaston, 1864: 561 (Euryporus; Type locality: Grand Canary: Teror).

  • — Wollaston, 1865: 485 (Euryporus; habitat; Canary Islands).

  • — Fauvel, 1897d: 310 (Euryporus; cited as Eurporus; Canaries).

  • — Fauvel, 1902b: 118 (Euryporus; Canaries).

  • — Bernhauer and Schubert, 1916: 414 (Euryporus; catalog).

  • — Coiffait, 1978: 294 (Euryporus; characters; Canary Islands).

  • — Hernández, Outerelo, and Gamarra, 1994: 187 (Euryporus; checklist of species; Canary Islands).

  • Distribution: Canary Islands.

puncticollis Erichson, 1840, see: Tympanophorus.

warisensis Last, 1987: 51 (Euryporus; Type locality: Irian: Warris, S. of Jayapura, 450–500 m).

  • Distribution: New Guinea.

Haematodes [2 species; Neotropical Region]

Haematodes Laporte, 1835: 112 (species included: bicolor). Type species: Haematodes bicolor Laporte, fixed by monotypy.

  • — Erichson, 1839b: 340 (characters).

  • — Laporte, 1840: 175 (cited as Hematodes; characters).

  • — Lacordaire, 1854: 72 (characters; notes; list of species).

  • — Gemminger and Harold, 1868: 573 (catalog).

  • — Lynch, 1884: 115 (characters).

  • — Bernhauer and Schubert, 1916: 410 (world catalog; 2 species).

  • — Scheerpeltz, 1933: 1420 (world catalog supplement).

  • — Blackwelder, 1944: 144 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 179 (type species: bicolor).

Platycnemus Nordmann, 1837: 135 (species included: lateritius). Type species: Platycnemus lateritius Nordmann, fixed by monotypy.

  • — Lacordaire, 1854: 72 (synonym of Haematodes).

  • — Lynch, 1884: 115 (synonym of Haematodes).

  • — Bernhauer and Schubert, 1916: 410 (synonym of Haematodes).

  • — Blackwelder, 1952: 312 (type species: lateritius).

  • — Newton, 1995: 51 (requested that the International Commission of Zoological Nomenclature place Platycnemus on the Official List of Generic Names in Zoology).

  • — ICZN, 1996: 215 (placed on Official List of Generic Names in Zoology).

Platytoma Chevrolat, 1847: 263 ([Note: Nomen nudum (Article 11.6, 12.1), published without description or species and as a synonym of Haematodes. Blackwelder, 1952: 314 cited it as an isogenotypic synonym of Haematodes]).

  • — Lacordaire, 1854: 72 (synonym of Haematodes).

  • — Lynch, 1884: 115 (synonym of Haematodes).

bicolor Laporte, 1835: 113 (Haematodes; Type locality: Buenos Ayres).

  • — Blanchard, 1842: 76 (Haematodes; Uruguay; Brazil).

  • — Erichson, 1839b: 340 (Haematodes; characters; Brazil).

  • — Laporte, 1840: 175 (Haematodes; characters; Argentina).

  • — Lynch, 1884: 116 (Haematodes; characters; Argentina; Brazil; Uruguay).

  • — Bernhauer and Schubert, 1916: 410 (Haematodes; catalog).

  • — Scheerpeltz, 1933: 1420 (Haematodes; catalog).

  • Distribution: Brazil, Uruguay, Argentina.

lateritius Nordmann, 1837: 135 (Platycnemus; Type locality: Brasilia meridionalis).

  • — Blanchard, 1842: 76 (Haematodes; synonym of bicolor).

  • — Erichson, 1839b: 340 (Haematodes; synonym of bicolor).

  • — Lynch, 1884: 116 (Haematodes; synonym of bicolor).

  • — Bernhauer and Schubert, 1916: 410 (Haematodes; synonym of bicolor).

  • — Newton, 1995: 51 (requested that the International Commission of Zoological Nomenclature place lateritius on the Official List of Specific Names in Zoology).

  • — ICZN, 1996: 215 (placed on Official List of specific Names in Zoology).

lateritius Nordmann, 1837, see: bicolor Laporte, 1835.

tenuipes Kraatz, 1858: 363 (Haematodes; Type locality: Brasilia).

  • — Bernhauer and Schubert, 1916: 410 (Haematodes; catalog).

  • Distribution: Brazil.

Heinzia [1 species; Palaearctic Region]

Heinzia Korge, 1971a: 33, 36 (species included; variabilis). Type species: Heinzia variabilis Korge, fixed by original designation and monotypy.

  • — Smetana, 1977b: 181 (characters).

  • — Coiffait, 1978: 284 (characters).

variabilis Korge, 1971a: 33, 37 (Heinzia; Type locality: Anatolia bor.: Kaçkar-Daglari bei Iliça, Tal von Ardesen, 1000–1600 m).

  • — Coiffait, 1978: 284 (Heinzia; characters; Anatolia).

  • — Solodovnikov, 1998a: 16 (Heinzia; notes; Russia, Krasnodar territory).

  • Distribution: Turkey, Russia.

Hemiquedius [1 species; Nearctic Region]

Hemiquedius Casey, 1915: 397, 399 (species included: ferox). Type species: Quedius ferox LeConte, fixed by original designation and monotypy.

  • — Casey, 1915: 397 (type species: Quedius ferox).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 182 (type species: ferox).

  • — Smetana, 1971: 238 (valid genus; characters).

  • — Moore and Legner, 1975: 53 (catalog; North America north of Mexico).

  • — Smetana, 1977b: 181 (characters).

  • — Moore and Legner, 1979: 45 (characters; notes).

  • — Frank, 1986a: 371 (checklist of species of Florida).

  • — Downie and Arnett, 1996: 394 (characters).

  • — Lundgren, 1998: 51 (list of species from Florida; records for other States listed for each species).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 392 (1 Nearctic species; notes; characters in key).

ferox LeConte, 1878a: 388 (Quedius; Type locality: Enterprise, Florida; [Note: See lectotype designation by Smetana, 1971]. Also cited from Louisiana; Canada; Massachusetts).

  • — Horn, 1878: 166 (Quedius; characters; Massachusetts to Canada, Florida, Louisiana).

  • — Blatchley, 1910: 372 (Quedius; characters; Indiana).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Smetana, 1965: 35 (Quedius; subgenus Hemiquedius; Nova Scotia).

  • — Smetana, 1971: 239 (Hemiquedius; lectotype designation: Enterprise, Florida; characters; Ontario; Nova Scotia; Quebec; Alabama; Florida; Georgia; Illinois; Indiana; Iowa; Louisiana; Maine; Maryland; massachusetts; Michigan; Missouri; New Jersey; New York; North Carolina; Pennsylvania; Wisconsin).

  • — Smetana, 1971a: 1848 (Hemiquedius; notes).

  • — Smetana, 1978a: 831 (Hemiquedius; collecting notes; Florida; Illinois; Texas).

  • — Frank, 1982: 36 (Hemiquedius; list of parasites).

  • — Smetana, 1990d: 101 (Hemiquedius; Arkansas; Georgia; Louisiana; Mississippi; New Hampshire; New York; Tennessee).

  • — Campbell and Davies, 1991: 122 (Hemiquedius; checklist of species; Canada).

  • — Downie and Arnett, 1996: 394 (Hemiquedius; characters; USA).

  • Distribution: Canada, USA.

Heterothops [149 species; Neotropical, Nearctic, Palaearctic, Oriental, and Australian Regions]

Heterothops Stephens, 1829a: 23 (species included: binotatus and two unavailable names). Type species: Staphylinus binotatus Gravenhorst, fixed by monotypy.

  • — Stephens, 1829: 284 (catalog; Britain).

  • — Stephens, 1832: 200 (characters).

  • — Stephens, 1833: 256 (characters).

  • — Westwood, 1838a: 16 (characters; type species).

  • — Erichson, 1839a: 480 (characters).

  • — Heer, 1839: 280 (characters).

  • — Stephens, 1839: 385 (characters).

  • — Erichson, 1840: 515 (characters).

  • — Redtenbacher, 1849: 711 (characters).

  • — Schaum, 1852: 26 (list of species; Europe).

  • — Lacordaire, 1854: 83 (characters; notes; list of species).

  • — Fairmaire and Laboulbène, 1856: 532 (characters).

  • — Jacquelin du Val, 1856: 38 (characters).

  • — Kraatz, 1857d: 483 (characters).

  • — Redtenbacher, 1857: 198 (characters; key to Austrian species).

  • — Thomson, 1858: 30 (characters).

  • — G. Waterhouse, 1858: 22 (catalog of British species).

  • — Thomson, 1859: 26 (characters; type species: dissimilis).

  • — Schaum, 1859: 25 (catalog; European species).

  • — Thomson, 1860: 180 (characters).

  • — LeConte, 1861a: 64 (characters).

  • — LeConte, 1863: 23 (list of North American species).

  • — Fauvel, 1866a: 337 [= 1866b: 88] (characters).

  • — Wencker and Silbermann, 1866: 27 (cited as Heterotops; list of species; collecting notes; France).

  • — Gemminger and Harold, 1868: 567 (catalog).

  • — Jacquelin du Val, 1868: 71 (catalog; European species).

  • — Sharp, 1871d: 11 (list of British species).

  • — Fauvel, 1874: 309 [= 1874a: 535] (characters; key to species).

  • — Redtenbacher, 1874: 196 (characters; key to Austrian species).

  • — Fauvel, 1875a: XXXIV [= 1875b: 236] (catalog).

  • — Mulsant and Rey, 1876: 808 [= 1877: 664] (characters; key to species).

  • — Horn, 1878: 152 (characters; key for North American species).

  • — Heyden, 1880: 70 (list of species of Siberian region).

  • — Duvivier, 1883: 136 (catalog).

  • — Sharp, 1884: 323 (notes; Mexican and Central American species).

  • — Lynch, 1884: 122 (characters).

  • — Sharp, 1888: 224 (characters; key to British species).

  • — Seidlitz, 1891: 400 (key to some European species).

  • — Seidlitz, 1891a: 426 (key to some European species).

  • — Heyden, Reitter, and Weise, 1891: 99 (list of species of Europe and Caucasus).

  • — Heyden, 1893: 40 (list of species of Siberian region).

  • — Ganglbauer, 1895: 384 (adult and larval characters; key to middle European species).

  • — Heyden, 1896: 31 (list of species of Siberian region).

  • — Fauvel, 1897d: 310 (catalog of species of Barbary; Madeira; Canaries).

  • — Germain, 1903: 414 (characters; notes).

  • — Heyden, Reitter, and Weise, 1906: 176 (list of species of Europe and Caucasus).

  • — Porta, 1907: 89 (key to Italian species).

  • — Reitter, 1908: 106, 107 (characters; key to species).

  • — Blatchley, 1910: 368 (characters).

  • — Petri, 1912: 80 (list of species of Siebenbürgen).

  • — Johansen, 1914: 345 (characters; key to species).

  • — Bernhauer and Schubert, 1916: 411 (world catalog; 48 species).

  • — Leng, 1920: 109 (catalog of North American species).

  • — Munster, 1923: 184 (characters; key to Scandinavian species).

  • — Winkler, 1925: 387 (catalog for Palaearctic region).

  • — Cameron, 1925: 71 (catalog of Indian species).

  • — Porta, 1926: 118 (characters; key to species of Italy).

  • — Bruch, 1928a: 429 (catalog; Argentina).

  • — Portevin, 1929: 336 (key to French species).

  • — Roubal, 1930: 385 (catalog; Slovakia).

  • — Cameron, 1932: 265 (characters; key to species of British India).

  • — Joy, 1932: 107 (key to British species).

  • — Scheerpeltz, 1933: 1421 (world catalog supplement).

  • — Normand, 1935: 372 (list of species of Tunisia).

  • — Paulian, 1941: 254 (larval characters).

  • — Blackwelder, 1943: 464 (characters; type species: binotatus; key to species of West Indies).

  • — Blackwelder, 1944: 144 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Kloet and Hincks, 1945: 167 (list of British species).

  • — Tottenham, 1949: 376, 421 (type species: binotatus Gravenhorst; notes; checklist of British species).

  • — Blackwelder, 1952: 186 (type species: binotatus).

  • — V. Hansen, 1952: 158 (characters; key to Danish species).

  • — V. Hansen, 1952: 158 (characters; key to Danish species).

  • — Hatch, 1957: 214, 230 (characters; key to species of Pacific Northwest).

  • — Smetana, 1958: 314 (characters; habitus of one species; type species: binotatus; key to Czechoslovakian species).

  • — Kocher, 1958: 156 (checklist of species; Morocco).

  • — Smetana, 1959a: 215 (list of species collected in Albania).

  • — Scheerpeltz, 1962b: 263 (characters; type species: binotatus).

  • — Ferreira, 1962a: 18 (list of Portuguese species).

  • — Székessy, 1963: 59 (characters).

  • — Palm, 1963b: 154 (characters; key to species of Sweden).

  • — Lohse, 1964: 203 (characters; key to central European species).

  • — Pototskaia, 1967: 79 (larval characters).

  • — Smetana, 1967g: 466, 470 (natural history notes; key to central European species).

  • — Scheerpeltz, 1968a: 86 (catalog of Austrian species).

  • — Kasule, 1970: 57 (larval characters).

  • — Smetana, 1971: 14 (characters; key to species of America north of Mexico; species group classification).

  • — Smetana, 1971a: 1838 (additions to key to species).

  • — Tikhomirova, 1973a: 188 (checklist of species of USSR).

  • — Last, 1975: 425 (characters; species of New Guinea).

  • — Moore and Legner, 1975: 54 (catalog; North America north of Mexico).

  • — Moore and Legner, 1976: 539 (includes seashore species; notes).

  • — Smetana, 1977b: 181 (characters).

  • — Pope, 1977: 30 (list of British species).

  • — Coiffait, 1978: 300 (characters; key to species of western Palaearctic region).

  • — Smetana, 1978a: 821 (key to North American species).

  • — Topp, 1978: 312 (larval characters).

  • — Israelson, 1979a: 261 (discussion of characters of punctation, chaetotaxy, and aedeagus).

  • — Moore and Legner, 1979: 46 (characters; notes).

  • — Uhlig, 1979: 255 (some species collected in Germany).

  • — Uhlig, Vogel, and Sieber, 1980: 248 (some species collected in Germany).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 113 (catalog; Poland).

  • — Uhlig and Vogler, 1981: 121 (list of some species collected in Germany; habitat and collecting notes).

  • — Tóth, 1984: 98 (characters; habitus; key to species of Hungary).

  • — Shibata, 1984: 115 (checklist of species of Japan).

  • — Frank, 1986a: 371 (checklist of species of Florida).

  • — Muona and Viramo, 1986: 13 (list of species of northeastern Finland).

  • — Segers, 1986: 52 (checklist of species; Belgium).

  • — Drugmand 1987b (comments on distribution of Belgian species).

  • — Ádám, 1987: 152 (list of two species with collecting notes; Hungary).

  • — Smetana, 1988: 318 (characters; key to species of Himalayan region).

  • — Lohse and Lucht, 1989: 169 (supplement to key to species of Lohse, 1964).

  • — Angelini, 1991: 191 (list of species from high elevations of Calabria, Italy).

  • — M. Hansen, Kristensen, Mahler, Pedersen, 1991: 107 (list of Danish species).

  • — Campbell and Davies, 1991: 121 (checklist of species; Alaska and Canada).

  • — Silfverberg, 1992: 17 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Navarrete-Heredia and Márquez-Luna, 1993 (cave collections; Mexico).

  • — Welch, 1993: 229 (ovariole structure and development).

  • — Smetana, 1993b: 49 (list of species of Czech Republic and Slovakia).

  • — Ito, 1994a: 179 (key to Japanese species).

  • — Smetana, 1995: 121 (revision of species of Taiwan; key to species of Taiwan).

  • — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 30 (additional records to Danish list of species).

  • — Ciceroni and Zanetti, 1995: 31 (list of species of Italy).

  • — M. Hansen, 1996: 108 (list of Danish species).

  • — Cho, 1996b: 108 (key to species of Korea).

  • — Downie and Arnett, 1996: 388 (characters; key to species of Northeastern North America).

  • — Lundgren, 1998: 51 (list of species from Florida; records for other States listed for each species).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 74 (list of species collected in 1997; Denmark).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 393 (18 Nearctic species; notes; characters in key).

Trichopygus Nordmann, 1837: 137 (species included: subuliformis, dissimilis, nitens, pumilio). Type species: Tachyporus dissimilis Gravenhorst, fixed by subsequent designation by Tottenham, 1939d: 229.

  • — Erichson, 1839a: 480 (synonym of Heterothops).

  • — Lacordaire, 1854: 83 (synonym of Heterothops).

  • — Jacquelin du Val, 1856: 38 (synonym of Heterothops).

  • — Fauvel, 1874: 309 [= 1874a: 535] (synonym of Heterothops).

  • — Lynch, 1884: 122 (synonym of Heterothops).

  • — Ganglbauer, 1895: 386 (synonym of Heterothops).

  • — Bernhauer and Schubert, 1916: 411 (synonym of Heterothops).

  • — Cameron, 1932: 265 (synonym of Heterothops).

  • — Tottenham, 1939d: 229 (type species designation: dissimilis).

  • — Blackwelder, 1943: 464 (synonym of Heterothops; type species: dissimilis).

  • — Tottenham, 1949: 376 (synonym of Heterothops; type species: dissimilis).

  • — Blackwelder, 1952: 396 (type species).

  • — Smetana, 1958: 314 (synonym of Heterothops).

  • — Smetana, 1971: 14 (synonym of Heterothops).

  • — Coiffait, 1978: 301 (synonym of Heterothops).

acuminatus Fairmaire, 1860, see: Quedius.

aeneiventris Smetana, 1971a: 1836 (Heterothops; Type locality: Arizona: Cochise Co.: Huachuca Mts., Ramsey Canyon, 5200 ft.).

  • — Smetana, 1978a: 818 (Heterothops; collecting notes; Arizona).

  • Distribution: USA.

amabilis Last, 1975: 428 (Heterothops; Type locality: Okapa, 5800 ft.).

  • Distribution: New Guinea.

angusticeps Reitter, 1891: 18 (Heterothops; Type locality: Kasalinsk).

  • — Eppelsheim, 1892: 329 (Heterothops; Turkestan).

  • — Bernhauer and Schubert, 1916: 411 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1421 (Heterothops; catalog; Turkestan; Transcaspian region).

  • — Coiffait, 1978: 317 (Heterothops; characters; Transcaspian region).

  • Distribution: Kazakhstan, Uzbekistan.

apicicornis Bernhauer, 1917h: 86 (Heterothops; Type locality: West-Kolumbien: Umgebung von Cali am Rio Cauca, St. Antonio, 2000 m).

  • — Scheerpeltz, 1933: 1421 (Heterothops; catalog).

  • Distribution: Colombia.

apterus Lea, 1925: 232 (Heterothops; Type locality: Tasmania: Cradle Mountain).

  • — Scheerpeltz, 1933: 1421 (Heterothops; catalog).

  • Distribution: Australia.

aquitanicus Coiffait, 1977d: 295 (Heterothops; Type locality: Arcachon, Plage du Pilat).

  • — Coiffait, 1978: 315 (Heterothops; characters; France).

  • Distribution: France.

argus Wendeler, 1956b: 263 (Heterothops; Type locality: Brasilien: Nova Teutonia).

  • Distribution: Brazil.

arizonicus Smetana, 1971a: 1834 (Heterothops; Type locality: Arizona: Cochise Co.: Chiricahua Mts., Portal, 5000 ft.).

  • — Smetana, 1978a: 817 (Heterothops; collecting notes; Arizona).

  • Distribution: USA.

armeniacus Coiffait, 1977c: 142 (Heterothops; Type locality: Erevan, Kamaker, Arménie soviétique).

  • — Coiffait, 1978: 312 (Heterothops; characters; Armenia).

  • — Tronquet, 1981: 71 (Heterothops; Afghanistan).

  • Distribution: Armenia, Afghanistan.

asperatus Smetana, 1971: 34 (Heterothops; Type locality: California: Samoa, Humboldt Co. Also cited from British Columbia).

  • — Smetana, 1971b: 1 (Heterothops; Washington).

  • — Orth, Moore, and Fisher, 1978: 188 (Heterothops; characters; collecting notes; California).

  • Distribution: Canada, USA.

baicalensis Fauvel, 1900d: 233 (Heterothops; Type locality: Riviere Sludianka, Baical; Daourie).

  • — Bernhauer and Schubert, 1916: 411 (Heterothops; catalog).

  • — Smetana, 1967d: 213 (Heterothops; Mongolia).

  • — Smetana 1975c: 176 (Heterothops; Mongolia).

  • — Coiffait, 1978: 318 (Heterothops; characters; central Asia).

  • Distribution: Russia, Mongolia.

balthasari Smetana, 1967g: 468 (Heterothops; Type locality: Böhmen, Böhmisches Mittelgebirge, Berg Raná).

  • — Coiffait, 1978: 312 (Heterothops; cited as balthazari; characters; central Europe).

  • — Tóth, 1984: 100 (Heterothops; cited as balthazari; characters).

  • — Lohse and Lucht, 1989: 171 (Heterothops; characters).

  • — Smetana, 1993b: 49 (Heterothops; Czech Republic).

  • Distribution: Czech Republic.

basicornis Bernhauer, 1917h: 85 (Heterothops; Type locality: Bolivien: Yuracares).

  • — Scheerpeltz, 1933: 1421 (Heterothops; catalog).

  • Distribution: Bolivia.

bicolor Smetana, 1975c: 173 (Heterothops; Type locality: Mongolie: Bajanchongor aimak: Oase Echin gol, cca 90 km NO vom Grenzposten Caganbulag, 950 m).

  • Distribution: Mongolia.

bimaculatus Fauvel, 1878g: 557 (Heterothops; Type locality: Nouvelle-Galles du Sud, Sydney).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1421 (Heterothops; catalog).

  • Distribution: Australia.

binotatus Gravenhorst, 1802: 28 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).

  • — Latreille, 1804: 316 (Staphylinus; characters; Germany).

  • — Gyllenhal, 1810: 312 (Staphylinus; synonym of subuliformis Gravenhorst).

  • — C. Sahlberg, 1830: 319 (Staphylinus; synonym of subuliformis Gravenhorst).

  • — Stephens, 1833: 257 (Heterothops; characters; England).

  • — Westwood, 1838a: 16 (Heterothops; type species).

  • — Stephens, 1839: 401 (Heterothops; characters; England).

  • — Kraatz, 1857d: 485 (Heterothops; characters; Germany).

  • — Redtenbacher, 1857: 985 (Heterothops; characters; Austria).

  • — Fauvel, 1874: 310 [= 1874a: 536] (Heterothops; characters; France; Russia; Germany; Britain; Italy; Corsica; Sardinia; Cyprus; Caramania).

  • — Redtenbacher, 1874: 197 (Heterothops; characters).

  • — Fauvel, 1876a: 78 [= 1876: 263] (Heterothops; Algeria; Tunisia).

  • — Fauvel, 1878: 130 [= 1878a: 50] (Heterothops; Algeria; Tunisia).

  • — Fauvel, 1886: 60 [= 1886a: 52] (Heterothops; Algeria).

  • — Ganglbauer, 1895: 387 (Heterothops; characters; coast of North Sea and Baltic Sea; Mediterranean coast).

  • — Fauvel, 1902b: 118 (Heterothops; Algeria; Tunisia).

  • — Porta, 1907: 101 (Heterothops; characters; Italy).

  • — Sainte-Claire Deville, 1906: 103 (Heterothops; Corsica).

  • — Reitter, 1909: 108 (Heterothops; characters; Germany).

  • — Johansen, 1914: 346 (Heterothops; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Munster, 1923: 184 (Heterothops; Fennoscandia).

  • — Portevin, 1929: 337 (Heterothops; characters; France).

  • — Scheerpeltz, 1933: 1421 (Heterothops; catalog).

  • — V. Hansen, 1952: 159 (Heterothops; characters; Denmark).

  • — Scheerpeltz, 1960d: 136 (Heterothops; Portugal).

  • — Palm, 1963b: 156 (Heterothops; characters; Sweden; Denmark; Norway).

  • — Lohse, 1964: 204 (Heterothops; characters; central Europe).

  • — Horion, 1965: 245 (Heterothops; Syria; Cyprus; Turkey; Greece; Dalmatia; Italy; France; Spain; Tunisia; Algeria; Spain; Portugal; Ireland; Britain; Belgium; Holland; Norway; Denmark; Sweden; Germany).

  • — Smetana, 1967g: 470 (Heterothops; characters).

  • — Szujecki, 1969a: 258 (Heterothops; notes; Poland).

  • — Pope, 1977: 30 (Heterothops; Britain).

  • — Coiffait, 1978: 308 (Heterothops; characters; Europe; Britain; Scandinavia; North Africa).

  • — Israelson, 1979a: 265, 266 (Heterothops; characters).

  • — Frank, 1982: 37 (Heterothops; list of parasites).

  • — Tóth, 1984: 100 (Heterothops; characters).

  • — Lucht, 1987: 108 (Heterothops; checklist; Germany; Poland; Benelux; Denmark; Sweden).

  • — Lohse and Lucht, 1989: 169 (Heterothops; characters).

  • — Smetana, 1993b: 49 (Heterothops; Czech Republic).

  • — Ciceroni and Zanetti, 1995: 31 (Heterothops; Italy).

  • — Owen, 1997a: 149 (Heterothops; collecting sites; Ireland).

  • — Herman, 2001: 25 (Heterothops; synonymy).

  • Distribution: Tunisia, Algeria, Europe, Turkey, Russia.

sericans Stephens, 1832: 221 (Quedius; Type locality: Suffolk).

  • — Stephens, 1839: 391 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 22 (Heterothops; synonym of binotatus).

holmensis Stephens, 1833: 257 (Heterothops; Type locality: on the sea-shore near Holm, in Norfolk).

  • — Stephens, 1839: 401 (Heterothops; variety of binotatus; characters; England).

  • — G. Waterhouse, 1858: 22 (Heterothops; synonym of binotatus).

  • — Gemminger and Harold, 1868: 566 (Heterothops; synonym of binotatus).

  • — Ganglbauer, 1895: 387 (Heterothops; synonym of binotatus).

  • — Porta, 1907: 101 (Heterothops; synonym of binotatus).

  • — Bernhauer and Schubert, 1916: 411 (Heterothops; synonym of binotatus).

  • — Coiffait, 1978: 309 (Heterothops; synonym of binotatus).

kirbiella Stephens, 1833: 257 (Heterothops; Type locality: Sussex).

  • — Stephens, 1839: 401 (Heterothops; variety of binotatus; characters; England).

  • — G. Waterhouse, 1858: 22 (Heterothops; synonym of binotatus).

  • — Gemminger and Harold, 1868: 567 (Heterothops; cited as kirbyella; synonym of binotatus).

  • — Ganglbauer, 1895: 387 (Heterothops; cited as kirbyella; synonym of binotatus).

  • — Porta, 1907: 101 (Heterothops; cited as kirbyella; synonym of binotatus).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; cited as kirbyella; synonym of binotatus).

  • — Coiffait, 1978: 309 (Heterothops; cited as kirbyellus; synonym of binotatus).

binotatus Erichson, 1840: 516 (Heterothops; [preoccupied]; [Note: Bernhauer and Schubert, 1916: 412 listed Erichson's reference for binotatus as a subsequent citation to binotatus Gravenhorst. Erichson attributed binotatus to no one so he evidently described it as new but it may be the same as binotatus Gravenhorst]; Type locality: Anglia).

  • — Fairmaire and Laboulbène, 1856: 533 (Heterothops; characters; France).

  • — Peyron, 1858: 428 (Heterothops; Turkey).

  • — Seidlitz, 1875: 263 (Heterothops; characters; Baltic region).

  • — Mulsant and Rey, 1876: 811 [= 1877: 667] (Heterothops; characters; France).

  • — Fowler, 1888: 224 (Heterothops; characters; Britain).

  • — Herman, 2001: 25 (Heterothops; synonym of binotatus).

binotatus Erichson, 1840, see: binotatus Gravenhorst, 1802.

bonariensis Lynch, 1884: 123 (Heterothops; Type locality: Chacabuco).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • Distribution: Argentina.

boops Bernhauer, 1910b: 383 (Heterothops; Type locality: Mexiko: Cuernavaca).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • Distribution: Mexico.

brevipennis Germain, 1903, see: Cheilocolpus.

brunnipennis Kiesenwetter, 1858: 56 (Heterothops; Type locality: bei Nauplia [bei Tirynth]; [Note: See lectotype designation by Israelson, 1979a]).

  • — Baudi, 1870: 383 (Heterothops; cited as brunneipennis; characters; Sardinia).

  • — Fauvel, 1874: 312 [= 1874a: 538] (Heterothops; cited as brunneipennis; synonym of dissimilis).

  • — Fowler, 1888: 225 (Heterothops; synonym of dissimilis).

  • — Ganglbauer, 1895: 388 (Heterothops; cited as brunneipennis; synonym of dissimilis).

  • — Porta, 1907: 103 (Heterothops; cited as brunneipennis; aberration of dissimilis).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; cited as brunneipennis; variety of dissimilis).

  • — Scheerpeltz, 1933: 1422 (Heterothops; cited as brunneipennis; aberration of dissimilis).

  • — Gridelli, 1934: 82 (Heterothops; cited as brunneipennis; variety of dissimilis; Karakoram region).

  • — C. Koch, 1936a: 76 (Heterothops; cited as brunneipennis; Libya).

  • — Normand, 1947a: 9 (Heterothops; cited as brunneipennis; subspecies of dissimilis; Tunisia).

  • — Cameron, 1951: 283 (Heterothops; cited as brunneipennis; Egypt).

  • — Smetana, 1958: 318 (Heterothops; cited as brunneipennis; aberration of dissimilis).

  • — Scheerpeltz, 1963: 14 (Heterothops; cited as brunneipennis; Afghanistan).

  • — Focarile, 1964: 57 (Heterothops; cited as valid species; Italy).

  • — Coiffait, 1973a: 275 (Heterothops; cited as brunneipennis; variety of dissimilis; Morocco).

  • — Coiffait, 1978: 314 (Heterothops; cited as brunneipennis; variety of dissimilis; characters).

  • — Muona, 1979: 15 (Heterothops; cited as brunneipennis; synonym of dissimilis).

  • — Israelson, 1979a: 263, 265 (Heterothops; lectotype designation; valid species distinct from dissimilis; Swedish record of brunnipennis are minutus; notes; discussion; characters).

  • Distribution: Egypt, Libya, Europe, Afghanistan, “Karakorum region”.

brunneus Smetana, 1971: 32 (Heterothops; Type locality: California: Antelope Spgs., Inyo Co.).

  • — Smetana, 1978a: 821 (Heterothops; collecting notes; California).

  • Distribution: USA.

buettikeri Coiffait, 1981b: 238 (Heterothops; Type locality: Arabie Saoudite: Hofuf).

  • — Gusarov, 1997: 288 (Heterothops; Saudi Arabia).

  • Distribution: Saudi Arabia.

buluensis Last, 1975: 438 (Heterothops; Type locality: New Guinea: Garaina, Bulu Valley).

  • Distribution: New Guinea.

burdurensis Štourač, 2000: 67 (Heterothops; Type locality: Turquie: Burdur, Çerçin lac Burdur, 870 m).

  • Distribution: Turkey.

californicus LeConte, 1863, see: fusculus LeConte, 1863.

campbelli Smetana, 1971: 22 (Heterothops; Type locality: Illinois: Pine Hills Field Sta., Union Co.).

  • — Smetana, 1978a: 817 (Heterothops; collecting notes; North Carolina; Oklahoma; Texas).

  • — Smetana, 1981b: 632 (Heterothops; collecting notes; Florida; Illinois; Kentucky; Michigan).

  • — Smetana, 1990d: 96 (Heterothops; Arkansas; Louisiana; Mississippi).

  • — Downie and Arnett, 1996: 388 (Heterothops; characters; USA).

  • Distribution: USA.

canariensis Israelson, 1979a: 267 (Heterothops; Type locality: Los Tiles, Gran Canaria, Canary Islands).

  • — Wollaston, 1864: 562 (Heterothops; [Note: Misidentified as minutus according to Israelson, 1979a: 267]; notes; habitat; Canary Islands).

  • — Wollaston, 1865: 485 (Heterothops; [Note: Misidentified as minutus according to Israelson, 1979a: 267]; habitat; notes; Canary Islands).

  • — Coiffait, 1954: 167 (Heterothops; [Note: Misidentified as minutus according to Israelson, 1979a: 267]; Canary Islands).

  • — Hernández, Outerelo, and Gamarra, 1994: 188 (Heterothops; checklist of species; Canary Islands).

  • — Assing, 1999a: 346 (Heterothops; Canary Islands).

  • — Assing and Wunderle, 1999: 437 (Heterothops; Canary Islands).

  • Distribution: Canary Islands.

carbonatus Fall, 1901: 220 (Heterothops; Type locality: maritime southern California).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Hatch, 1957: 231 (Heterothops; characters; British Columbia; Washington; Oregon; Idaho).

  • — Smetana, 1971: 23 (Heterothops; characters; British Columbia; Arizona; California; Idaho; Oregon; Washington).

  • — Smetana, 1971a: 1833 (Heterothops; collecting notes; Arizona).

  • — Smetana, 1971b: 1 (Heterothops; British Columbia).

  • Distribution: Canada, USA.

castaneus Lea, 1925: 238 (Heterothops; Type locality: Lord Howe Island).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • Distribution: Lord Howe Island.

cephalota Coiffait, 1977d: 297 (Heterothops; Type locality: Darvaz, Noulvantch, Tadjikistan).

  • — Coiffait, 1978: 313 (Heterothops; characters; Transcaucasia).

  • Distribution: Tajikistan, “Transcaucasus”.

clarki Lea, 1925: 237 (Heterothops; Type locality: Western Australia: Swan River).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • Distribution: Australia.

cognatus Sharp, 1874: 20 (Heterothops; Type locality: Japan).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • — Watanabe, 1961: 354 (Heterothops; Izu Islands).

  • — Nakane, 1963a: 94 (Heterothops; characters; habitus photograph; Japan).

  • — Watanabe and Shibata, 1972: 65 (Heterothops; Yaku-shima Island).

  • — Yuh, Paik, Kwon, and Lee, 1985: 249 (Heterothops; Korea).

  • — Watanabe, 1993d (Heterothops; Sarushima Island).

  • — J. Li, 1993: 38 (Heterothops; China).

  • — Cho, 1996b: 108 (Heterothops; characters; Korea; also listed from China).

  • — Watanabe, 1998: 314 (Heterothops; Japan).

  • Distribution: Japan, Korea, China.

confertus Smetana, 1971: 25 (Heterothops; Type locality: Arizona: Madera Cn. S. Rita Mts., 6200 ft., S. Cruz Co.).

  • — Smetana, 1971a: 1834 (Heterothops; collecting notes; Arizona).

  • — Smetana, 1978a: 818 (Heterothops; collecting notes; Texas).

  • Distribution: USA.

conformis Smetana, 1971: 30 (Heterothops; Type locality: British Columbia: 8 mi. W. Creston. Also cited from Arizona, California, Colorado, Idaho, Nevada, New Mexico, Oregon, Washington).

  • — Smetana, 1971a: 1834 (Heterothops; collecting notes; California).

  • — Smetana, 1971b: 1 (Heterothops; British Columbia).

  • — Smetana, 1973: 1422 (Heterothops; California; Idaho; Oregon).

  • — Smetana, 1978a: 819 (Heterothops; collecting notes; Arizona; California).

  • Distribution: Canada, USA.

conticens† Scudder, 1900: 48 (Heterothops; Type locality: Florissant, Colorado).

  • — Handlirsch, 1907: 723 (Heterothops; cited as conticeus; catalog; Miocene; Colorado).

  • Distribution: USA.

conviva Smetana, 1967d: 212 (Heterothops; Type locality: Mongolei: Chentej aimak, 20 km SW von Somon Batnorov, 1000 m).

  • — Smetana 1975c: 176 (Heterothops; Mongolia).

  • Distribution: Mongolia.

crisbus Last, 1975: 436 (Heterothops; Type locality: Okasa, North Fore census division, Okapa district, 3000 ft.).

  • Distribution: New Guinea.

discoideus Fairmaire and Germain, 1861, see: Loncovilius.

dissimilis Gravenhorst, 1802: 125 (Tachyporus; Type locality: Brunsvigae).

  • — Gravenhorst, 1806: 23 (Tachinus; characters; Germany).

  • — Erichson, 1839a: 480 (Heterothops; characters; Germany).

  • — Heer, 1839: 281 (Heterothops; characters; Switzerland).

  • — Erichson, 1840: 517 (Heterothops; characters; Germany; Sweden).

  • — Kolenati, 1846: 21 (Heterothops; Caucasus; Iberia).

  • — P. Lucas, 1846: 113 (Heterothops; notes; Algeria).

  • — Jacobsen, 1849: 707 (Heterothops; characters; Denmark).

  • — Hochhuth, 1849: 146 (Heterothops; characters; Lenkoran).

  • — Redtenbacher, 1849: 712 (Heterothops; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 533 (Heterothops; characters; France).

  • — Jacquelin du Val, 1857: pl. 15, fig. 74 (Heterothops; [Note: Misidentification: Bernhauer and Schubert, 1916: 412 cited binotatus Jacquelin du Val as a synonym of dissimilis; Jacquelin du Val did not describe the species as new, he attributed it to Erichson]; illustration).

  • — Kraatz, 1857d: 485 (Heterothops; characters; Germany).

  • — Redtenbacher, 1857: 199 (Heterothops; characters; Austria).

  • — Thomson, 1859: 26 (type species of Heterothops).

  • — Thomson, 1860: 180 (Heterothops; [Note: Misidentification: Bernhauer and Schubert, 1916: 412 cited praevius Thomson as a synonym of dissimilis; Thomson did not describe the species as new, he attributed it to Erichson]; characters; Scandinavia).

  • — Thomson, 1860: 181 (Heterothops; characters; Scandinavia).

  • — Hochhuth, 1862: 43 (Heterothops; characters; Caucasus; Altai).

  • — Gredler, 1863: 99 (Heterothops; Austria).

  • — Wollaston, 1871: 298 (Heterothops; Madeira and Canary Islands).

  • — Fauvel, 1874: 312 [= 1874a: 538] (Heterothops; characters; France; Madeira and Canary Islands; Barbary; Caucasus; Transcaucasus; Daghestan; Altai region; Siberia).

  • — Redtenbacher, 1874: 196 (Heterothops; characters; Austria).

  • — Seidlitz, 1875: 263 (Heterothops; characters; Baltic region).

  • — Mulsant and Rey, 1876: 823 [= 1877: 679] (Heterothops; characters; France).

  • — J. Sahlberg, 1876: 30 (Heterothops; Finland).

  • — Fauvel, 1878: 130 [= 1878a: 50] (Heterothops; Morocco; Algeria; Tunisia).

  • — Fauvel, 1886: 60 [= 1886a: 52] (Heterothops; North Africa).

  • — Fowler, 1888: 225 (Heterothops; characters; Britain).

  • — Ganglbauer, 1895: 387 (Heterothops; characters; Palaearctic region).

  • — Fauvel, 1902b: 118 (Heterothops; Barbary; Egypt; Madeira; Canaries).

  • — Sainte-Claire Deville, 1906: 104 (Heterothops; Corsica).

  • — Porta, 1907: 103 (Heterothops; characters; Italy).

  • — Peyerimhoff, 1907: 14 (Heterothops; Sinai).

  • — Montandon, 1908: 80 (Heterothops; Romania).

  • — Reitter, 1909: 108 (Heterothops; characters; Germany).

  • — Johansen, 1914: 347 (Heterothops; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Munster, 1923: 185 (Heterothops; Fennoscandia).

  • — Portevin, 1929: 337 (Heterothops; characters; France).

  • — Gridelli, 1930: 78 (Heterothops; Libya).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • — Uyttenboogaart, 1937: 82 (Heterothops; characters; notes).

  • — V. Hansen, 1952: 159 (Heterothops; characters; Denmark).

  • — Smetana, 1958: 318 (Heterothops; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1961: 7 (Heterothops; Iran).

  • — Palm, 1963b: 156 (Heterothops; characters; Sweden; Denmark; Finland).

  • — Lohse, 1964: 204 (Heterothops; characters; central Europe).

  • — Scheerpeltz, 1965c: 510 (Heterothops; Sicily).

  • — Horion, 1965: 247 (Heterothops; widespread in southern and central Europe; Denmark; Britain; Sweden; Finland).

  • — Smetana, 1967g: 471 (Heterothops; characters).

  • — Smetana, 1967i: 557 (Heterothops; Turkey).

  • — Szujecki, 1968a: 733 (Heterothops; Poland).

  • — Coiffait, 1973a: 275 (Heterothops; Morocco).

  • — Osella and Zanetti, 1975: 113 (Heterothops; collected from nest of Talpa europaea; notes; Italy).

  • — Pope, 1977: 30 (Heterothops; Britain).

  • — Coiffait, 1978: 314 (Heterothops; characters; Europe; Britain; Scandinavia; Madeira; Canaries; North Africa).

  • — Israelson, 1979a: 265, 266 (Heterothops; characters).

  • — Frank, 1982: 37 (Heterothops; list of parasites).

  • — Tóth, 1984: 102 (Heterothops; characters; Hungary).

  • — Lucht, 1987: 108 (Heterothops; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Lohse and Lucht, 1989: 171 (Heterothops; characters).

  • — Nowosad, 1990: 124 (Heterothops; collected from mole nest; notes; Poland).

  • — Maachi, 1991: 251 (Heterothops; Morocco).

  • — Janák and Vysoky, 1992: 137 (Heterothops; collected with Formica rufa and truncorum; Bohemia).

  • — Smetana, 1993b: 49 (Heterothops; Czech Republic; Slovakia).

  • — Outerelo, Palmer and Pons, 1995: 80 (Heterothops; Mallorca).

  • — Ciceroni and Zanetti, 1995: 31 (Heterothops; probably a species complex; Italy).

  • — Ryabukhin, 1999: 69 (Heterothops; catalog; habitat; distribution; northeastern Russia).

  • — Kashcheev, 1999a: 58 (Heterothops; Kazakhstan).

  • Distribution: Libya, Tunisia, Algeria, Morocco, Canary Islands, Maderia, Europe, Russia, Armenia, Azerbaijan, Turkey, Iran, Kazakhstan.

flavolimbatus Motschulsky, 1860: 571 (Heterothops; Type locality: Armenie; Daghestan).

  • — Hochhuth, 1862: 42 (Heterothops; characters; Armenia; Daghestan).

  • — Fauvel, 1874: 312 [= 1874a: 538] (Heterothops; synonym of dissimilis).

  • — Mulsant and Rey, 1876: 826 [= 1877: 682] (Heterothops; characters; France).

  • — Fowler, 1888: 225 (Heterothops; synonym of dissimilis).

  • — Ganglbauer, 1895: 388 (Heterothops; synonym of dissimilis).

  • — Porta, 1907: 103 (Heterothops; synonym of dissimilis).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; synonym of dissimilis).

  • — Smetana, 1958: 318 (Heterothops; synonym of dissimilis).

  • — Coiffait, 1978: 314 (Heterothops; synonym of dissimilis).

minutus Motschulsky, 1860: 572 (Heterothops; [preoccupied]; Type locality: fl. Irtysch dans la Sibérie occidentale).

  • — Hochhuth, 1862: 43 (Heterothops; notes; Siberia).

  • — Fauvel, 1874: 313 [= 1874a: 539] (Heterothops; synonym of dissimilis).

  • — Ganglbauer, 1895: 388 (Heterothops; synonym of dissimilis).

  • — Porta, 1907: 103 (Heterothops; synonym of dissimilis).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; synonym of dissimilis).

  • — Smetana, 1958: 318 (Heterothops; synonym of dissimilis).

pygmaeus Gemminger and Harold, 1868: 567 (Heterothops; replacement name for minutus Motschulsky).

  • — Fauvel, 1874: 313 [= 1874a: 539] (Heterothops; synonym of dissimilis).

  • — Ganglbauer, 1895: 388 (Heterothops; synonym of dissimilis).

  • — Porta, 1907: 103 (Heterothops; synonym of dissimilis).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; synonym of dissimilis).

  • — Smetana, 1958: 318 (Heterothops; synonym of dissimilis).

distinguendus Hochhuth, 1872: 130 (Heterothops; Type locality: Not cited, but presumably from Gouvernements Kiew and/or Volhynien).

  • — Fauvel, 1874: 311 [= 1874a: 537] (Heterothops; synonym of praevius).

  • — Ganglbauer, 1895: 387 (Heterothops; synonym of praevius).

  • — Porta, 1907: 102 (Heterothops; synonym of praevius).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; synonym of praevius).

  • — Smetana, 1958: 316 (Heterothops; synonym of praevius).

  • — Coiffait, 1978: 310 (Heterothops; synonym of praevius).

  • — Gusarov, 1992a: 786 [= 1993a: 76] (Heterothops; synonym of dissimilis).

parvicornis Mulsant and Rey, 1876: 826 [= 1877: 682 (Heterothops; Type locality: Lille).

  • — Ganglbauer, 1895: 388 (Heterothops; synonym of dissimilis).

  • — Porta, 1907: 103 (Heterothops; synonym of dissimilis).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; synonym of dissimilis).

  • — Smetana, 1958: 318 (Heterothops; synonym of dissimilis).

  • — Coiffait, 1978: 314 (Heterothops; variety of dissimilis; characters).

sericans Mulsant and Rey, 1876: 820 [= 1877: 676] (Heterothops; [preoccupied]; Type locality: Provence).

  • — Ganglbauer, 1895: 387 (Heterothops; characters; Europe).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog; western Mediterranean region).

  • — Lohse, 1964: 204 (Heterothops; characters; central Europe).

  • — Coiffait, 1978: 314 (Heterothops; synonym of dissimilis).

  • — Tóth, 1984: 102 (Heterothops; variety of dissimilis; characters).

paradoxus Mulsant and Rey, 1876: 823 [= 1877: 679] (Heterothops; Type locality: Not cited).

  • — Ganglbauer, 1895: 387 (Heterothops; synonym of sericans).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; synonym of sericans).

fraudulentus Coiffait, 1977c: 143 (Heterothops; variety of dissimilis; [Note: Nomen nudum (Article 15.2]; cited from Portugal, environs de Faro).

  • — Coiffait, 1978: 314 (Heterothops; variety of dissimilis; characters).

distinguendus Hochhuth, 1872, see: dissimilis Gravenhorst, 1802.

dolichocephalus Lea, 1925: 231 (Heterothops; Type locality: Tasmania; Waratah).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • Distribution: Australia.

eidmanni Scheerpeltz, 1936a: 502 (Heterothops; Type locality: Brasilien: Mendes, E. d. Rio).

  • Distribution: Brazil.

exilis Casey, 1886, see: occidentis Casey, 1886.

exilis Erichson, 1840: 518 (Heterothops; Type locality: Columbia).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • Distribution: Colombia.

fallax Lynch, 1884: 124 (Heterothops; Type locality: Chacabuco).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • Distribution: Argentina.

fauveli Bernhauer and Schubert, 1916: 412 (Heterothops; replacement name for flavicollis Fauvel).

  • — Bernhauer, 1920: 17 (Heterothops; Australia).

  • — Lea, 1925: 230 (Heterothops; synonym of xantholinoides).

  • — Scheerpeltz, 1933: 1424 (Heterothops; synonym of xantholinoides).

  • — Herman, 2001: 26 (Heterothops; synonymy).

  • Distribution: Australia.

xantholinoides W. J. MacLeay, 1873: 141 (Philonthus; [preoccupied by xantholinoides Wollaston]; Type locality: Gayndah).

  • — Fauvel, 1877: 275 (Heterothops; characters; Australia).

  • — Bernhauer and Schubert, 1916: 437 (Quedius; catalog).

  • — Bernhauer, 1920: 17 (Heterothops; Australia).

  • — Lea, 1925: 230 (Heterothops; characters; Australia).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • — Herman, 2001: 26 (Heterothops; synonym of fauveli).

flavicollis Fauvel, 1878g: 559 (Heterothops; [preoccupied]; Type locality: Australie méridionale: Nouvelle-Galles du Sud, Sydney).

  • — Lea, 1925: 230 (Heterothops; synonym of xantholinoides).

  • — Scheerpeltz, 1933: 1424 (Heterothops; synonym of xantholinoides).

flavicollis Fauvel, 1878, see: fauveli Bernhauer and Schubert, 1916.

flavicollis Motschulsky, 1858, see: Gabronthus.

flavicornis Bernhauer, 1917h: 85 (Heterothops; Type locality: Bolivien: Yuracares).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • Distribution: Bolivia.

flavolimbatus Motschulsky, 1860, see: dissimilis Gravenhorst, 1802.

flavus Scheerpeltz, 1972: 122 (Heterothops; Type locality: Argentinien: Rio Negro, El Bolson, Mt. Piltriquitron, NW Tal, 460 m).

  • Distribution: Argentina.

formicetorum Bernhauer, 1912d: 46 (Heterothops; Type locality: Argentinien: Buenos Aires).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Frank, 1982: 37 (Heterothops; list of parasites).

  • Distribution: Argentina.

forsteri Scheerpeltz, 1960c: 128 (Heterothops; Type locality: Bolivien: Yungas de Arepucho, Sihuencas, 2200–2500 m).

  • Distribution: Bolivia.

franzi Coiffait, 1982: 82 (Heterothops; Type locality: Nepal: environs de Pina, region de Rara-See).

  • — Smetana, 1988: 328 (Heterothops; characters; Uttar Pradesh; Nepal).

  • Distribution: Nepal, India.

frater Last, 1975: 433 (Heterothops; Type locality: Okasa, 3000 ft., North Fore census division, Okapa subdistrict).

  • Distribution: New Guinea.

fraternus Smetana, 1971: 35 (Heterothops; Type locality: Washington: Paradise Park, alt. 6000 ft., Mt. Rainier. Also cited from Oregon and Washington).

  • — Smetana, 1971b: 1 (Heterothops; Washington).

  • Distribution: USA.

fraudulentus Coiffait, 1977 [Note: Nomen nudum], see: dissimilis Gravenhorst, 1802.

fulvipennis Bernhauer, 1921, see: laxus Blackwelder, 1944.

fulvipennis Motschulsky, 1860, see: praevius Erichson, 1839.

fulvus Scheerpeltz, 1972: 119 (Heterothops; Type locality: Argentinien: Rio Negro, El Bolson, Mt. Piltriquitron, 1170 m).

  • Distribution: Argentina.

fumigatus LeConte, 1863, see: fusculus LeConte, 1863.

fundatus Last, 1980: 151 (Heterothops; Type locality: New Guinea: Mt. Wilhelm, 3900 m).

  • Distribution: New Guinea.

funeralis Last, 1975: 432 (Heterothops; Type locality: Karimui, 3500 ft, S.W. Goroka, 6°32′[S],144°47′[E]).

  • — Last, 1987: 55 (Heterothops; Papua New Guinea; Irian).

  • Distribution: New Guinea.

fusculus LeConte, 1863a: 35 (Heterothops; Type locality: Cambridge, Massachusetts; [Note: See lectotype designation by Smetana, 1971]).

  • — Horn, 1878: 153 (Heterothops; variety of fumigatus; characters).

  • — Blatchley, 1910: 369 (Heterothops; characters; Indiana).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1422 (Heterothops; synonym of fumigatus).

  • — Smetana, 1971: 26 (Heterothops; cited as valid species; lectotype designation; characters; Alberta; British Columbia; Manitoba; Nova Scotia; Ontario; Quebec; Saskatchewan; Arizona; California; Colorado; Idaho; Illinois; Iowa; Maine; Massachusetts; Michigan; Minnesota; Montana; Nevada; New Hampshire; New Jersey; New Mexico; New York; North Carolina; North Dakota; Oregon; Pennsylvania; Washington; Wisconsin).

  • — Smetana, 1971b: 1 (Heterothops; Michigan).

  • — Levesque and Levesque, 1996: 292 (Heterothops; seasonal abundance; Quebec).

  • — Downie and Arnett, 1996: 388 (Heterothops; characters; USA).

  • Distribution: Canada, USA.

californicus LeConte, 1863a: 35 (Heterothops; Type locality: San Francisco, California; [Note: See lectotype designation by Smetana, 1971]).

  • — Horn, 1878: 153 (Heterothops; variety of fumigatus; characters).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1422 (Heterothops; synonym of fumigatus).

  • — Hatch, 1957: 231 (Heterothops; synonym of fumigatus).

  • — Smetana, 1971: 26 (Heterothops; lectotype designation; synonym of fusculus).

fumigatus LeConte, 1863a: 35 (Heterothops; Type locality: Detroit; [Note: See lectotype designation by Smetana, 1971]).

  • — Horn, 1878: 153 (Heterothops; characters; Michigan; Texas; California; middle States; Canada).

  • — Fall and Cockerell, 1907: 166 (Heterothops; New Mexico).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • — Hatch, 1957: 231 (Heterothops; characters; British Columbia; Washington; Oregon; Idaho).

  • — Smetana, 1965: 34 (Heterothops; Newfoundland).

  • — Smetana, 1971: 26 (Heterothops; lectotype designation; synonym of fusculus).

gemellus Smetana, 1971: 19 (Heterothops; Type locality: California: Pasadena).

  • — Smetana, 1978a: 817 (Heterothops; collecting notes; California).

  • Distribution: USA.

gestroi Gridelli, 1922: 25 (Heterothops; Type locality: Persia settentrionalis).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • — Coiffait, 1978: 309 (Heterothops; characters; Iran; Lebanon; Uzbekistan).

  • Distribution: Lebanon, Iran, Uzbekistan.

gijiensis Last, 1987: 54 (Heterothops; Type locality: Irian: Vogelkop, Lake Anggi Giji, 2000–2100 m).

  • Distribution: New Guinea.

gnarus Last, 1975: 435 (Heterothops; Type locality: Daulo Pass, 8000 ft., Osaro Climbu Divide).

  • Distribution: New Guinea.

grandiceps Bernhauer, 1917h: 84 (Heterothops; Type locality: Brasilien: Blumenau).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • Distribution: Brazil.

gressitti Last, 1987: 53 (Heterothops; Type locality: Papua New Guinea: New Guinea NE: Asaro-Chimbu Div., Daulo Pass, 2800 m).

  • Distribution: New Guinea.

giulianii Smetana, 1978a: 819 (Heterothops; Type locality: California: Inyo Co.: Rock Creek, 10,000 ft.).

  • Distribution: USA.

hindustanus Cameron, 1932: 267 (Heterothops; Type locality: Chakrata district: Korawa Khud, alt. 9000 feet; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • — Scheerpeltz, 1965: 111 (Heterothops; Burma).

  • — Coiffait, 1982: 33 (Heterothops; Nepal).

  • — Smetana, 1988: 325 (Heterothops; lectotype designation; characters; Uttar Pradesh; Nepal).

  • Distribution: India, Nepal, Myanmar.

holmensis Stephens, 1833, see: binotatus Gravenhorst, 1802.

hornabrooki Last, 1975: 430 (Heterothops; Type locality: N.E. New Guinea: Daulo Pass, North of Goroka, Eastern Highlands, 8200 ft.).

  • Distribution: New Guinea.

impressifrons Germain, 1903: 422 (Heterothops; Type locality: cordilleras de Chillan, cerca de las Termas).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • Distribution: Chile.

indicus Cameron, 1926b: 365 (Heterothops; Type locality: Mossy Falls, Mussoorie; characters; Himachal Pradesh; Uttar Pradesh; [Note: See lectotype designation by Smetana, 1988]. Originally cited from Mussoorie district; Arni Gad. Chakrata and Simla Hills).

  • — Scheerpeltz, 1933: 1423 (Heterothops; synonym of oculatus).

  • — Smetana, 1988: 322 (Heterothops; valid species; lectotype designation: Mossy Falls, Mussoorie; characters; Himachal Pradesh; Uttar Pradesh).

  • Distribution: India.

insolitus Sharp, 1884: 323 (Heterothops; Type locality: Guatemala, Cerro Zunil 4000 to 5000 feet).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • Distribution: Guatemala.

iranicus Štourač, 2000: 68 (Heterothops; Type locality: SE Iran: 40 km SW Zábolí).

  • Distribution: Iran.

ishiharai Ito, 1994a: 177 (Heterothops; Type locality: Japan: Yamanashi Pref.: Gozaishi Spa, alt. 1400 m).

  • Distribution: Japan.

jureceki Štourač, 2000: 70 (Heterothops; Type locality: Tockoe, Ros. or., Gub Samara).

  • Distribution: Russia.

kentiae Lea, 1925: 238 (Heterothops; Type locality: Lord Howe Island, summit of Mount Gower).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • Distribution: Lord Howe Island.

khairo Smetana, 1988: 326 (Heterothops; Type locality: India: Dehra Dun).

  • Distribution: India.

kirbiella Stephens, 1833, see: binotatus Gravenhorst, 1802.

laeticolor Reitter, 1891: 17 (Heterothops; Type locality: Kasalinsk).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • — Coiffait, 1978: 318 (Heterothops; characters; Transcaspian region; Transcaucasus).

  • Distribution: Kazakhstan, “Transcaucasus”.

laticeps Fauvel, 1878g: 558 (Heterothops; Type locality: Australie orientale: Queensland, Wide Bay).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Lea, 1925: 230 (Heterothops; valid species; characters; Australia).

  • Distribution: Australia.

laxus Blackwelder, 1944: 144 (Heterothops; replacement name for fulvipennis Bernhauer).

  • Distribution: Argentina.

fulvipennis Bernhauer, 1921c: 172 (Heterothops; [preoccupied]; Type locality: Argentinien: Prov. Jujuy, bei Hornadita, 3400 und bei Cueva Iturbe, 3700 m).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • — Blackwelder, 1944: 144 (Heterothops; synonym of laxus).

limbatus Heer, 1839, see: Quedius.

limosus Motschulsky, 1858, see: praevius Erichson, 1839.

longiceps Wendeler, 1956b: 262 (Heterothops; Type locality: Brasilien: Nova Teutonia).

  • Distribution: Brazil.

luctuosus Fauvel, 1877: 275 (Heterothops; Type locality: Australie).

  • — Fauvel, 1878g: 558 (Heterothops; Australia).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Australia.

macrops Coiffait, 1977c: 142 (Heterothops; Type locality: Arménie, Dvine, 15 km au Sud de Erevan, Arménie soviétique).

  • — Coiffait, 1978: 313 (Heterothops; characters; Armenia).

  • Distribution: Armenia.

magniceps Bernhauer, 1920, see: Ctenandropus.

malaisei Scheerpeltz, 1965: 280 (Heterothops; Type locality: N.E. Burma, Kambaiti, 7000 ft.).

  • Distribution: Myanmar.

marmotae Smetana, 1971a: 1837 (Heterothops; Type locality: Ontario; Ottawa, Kanata).

  • — Smetana, 1973: 1421 (Heterothops; Ontario).

  • — Downie and Arnett, 1996: 388 (Heterothops; characters; Canada).

  • Distribution: Canada.

marocanus Coiffait, 1977d: 296 (Heterothops; Type locality: Maroc: Aguelmane Sidi Ali).

  • — Coiffait, 1978: 316 (Heterothops; characters; Morocco).

  • Distribution: Morocco.

mediocris Fall, 1907, see: occidentis Casey, 1886.

mediofuscus Lea, 1925: 233 (Heterothops; Type locality: Tasmania: Waratah).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Australia.

megalops Cameron, 1959: 117 (Heterothops; Type locality: Angola: Vila Luso).

  • Distribution: Angola.

melanocerus Solsky, 1871: 156 (Heterothops; Type locality: Samarkand).

  • — Fauvel, 1874: 311 [= 1874a: 537] (Heterothops; synonym of praevius).

  • — Solsky, 1874: 171 (Heterothops; characters; Turkestan).

  • — Eppelsheim, 1892: 329 (Heterothops; valid species; Turkestan).

  • — Ganglbauer, 1895: 387 (Heterothops; synonym of praevius).

  • — Porta, 1907: 102 (Heterothops; synonym of praevius).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Coiffait, 1978: 314 (Heterothops; characters; Transcaucasus).

  • — Tronquet, 1981: 71 (Heterothops; Afghanistan).

  • Distribution: Uzbekistan, Afghanistan.

meurguesae Tronquet, 1981: 80 (Heterothops; Type locality: Afghanistan: Kenjan, 2000 m).

  • Distribution: Afghanistan.

mexicanus Schubert, 1909: 290 (Heterothops; Type locality: südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d'Orizaba).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • Distribution: Mexico.

microtophilus Coiffait, 1977c: 141 (Heterothops; Type locality: Goukassian Zouigakhpiour).

  • — Coiffait, 1978: 310 (Heterothops; characters; Armenia).

  • Distribution: Armenia.

minor Smetana, 1971: 36 (Heterothops; Type locality: Ontario: Ottawa. Also cited from Alberta, British Columbia, Manitoba, Nova Scotia, Maine, New Hampshire, and Washington).

  • — Smetana, 1973: 1422 (Heterothops; Quebec).

  • — Smetana, 1976c: 169 (Heterothops; Alberta; Newfoundland; Northwest Territories).

  • — Smetana, 1981b: 633 (Heterothops; Newfoundland).

  • — Downie and Arnett, 1996: 388 (Heterothops; characters; Canada; USA).

  • Distribution: Canada, USA.

minutus Motschulsky, 1860, see: dissimilis Gravenhorst, 1802.

minutus Wollaston, 1860: 53 (Heterothops; Type locality: Palmeira above Funchal; [Note: See lectotype designation by Israelson, 1979a]).

  • — Fauvel, 1869: 486 (Heterothops; synonym of praevius).

  • — Fauvel, 1869a: 151 (Heterothops; synonym of praevius).

  • — Wollaston, 1871: 298l, 299 (Heterothops; notes; synonym of dissimilis).

  • — Fauvel, 1874: 312 [= 1874a: 538] (Heterothops; synonym of dissimilis).

  • — Ganglbauer, 1895: 388 (Heterothops; synonym of dissimilis).

  • — Porta, 1907: 103 (Heterothops; synonym of dissimilis).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; synonym of dissimilis).

  • — Uyttenboogaart, 1937: 81 (Heterothops; characters; notes; Canary Islands).

  • — Smetana, 1958: 318 (Heterothops; synonym of dissimilis).

  • — Smetana, 1963b: 39 (Heterothops; Madeira).

  • — Coiffait, 1978: 314 (Heterothops; synonym of dissimilis).

  • — Israelson, 1979a: 264, 266 (Heterothops; lectotype designation; valid species; Swedish specimens identified as brunnipennis identical to minutus; characters; species may be introduced into Madeira; specimens from Canary and Cape Verde Islands identified as minutus are different species; Madeira; Sweden).

  • — Lohse and Lucht, 1989: 171 (Heterothops; characters; all central Europe).

  • — Borges, 1990: Table II (Heterothops; cited as valid species; Azores).

  • — Hodge and Jones, 1995: 41 (Heterothops; characters; Britain).

  • — Štourač, 1998b: 135 (Heterothops; notes; Czech Republic).

  • — Burakowski, Mroczkowski, and Stefańska, 2000: 53 (Heterothops; catalog; Poland).

  • Distribution: Azores, Madeira, Britain, Sweden, central Europe.

mirabilis Last, 1975: 426 (Heterothops; Type locality: New Guinea: Okapa).

  • — Last, 1987: 55 (Heterothops; Papua New Guinea).

  • Distribution: New Guinea.

mirus Lea, 1925: 234 (Heterothops; Type locality: Lord Howe Island).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Lord Howe Island.

montanus Iablokov-Khnzorian, 1966: 174 (Heterothops; Type locality: ArmSSR: Argats, u ozera Karilich).

  • — Coiffait, 1978: 311 (Heterothops; characters; Armenia).

  • Distribution: Armenia.

montanus Last, 1975, see: tumulus Herman, 2001.

myrmeciae Lea, 1929: 205 (Heterothops; Type locality: Victoria: Northern Gippsland; Fern Tree Gully).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Australia.

nidicola Thomson, 1870, see: niger Kraatz, 1868.

nidorum Smetana, 1967d: 211 (Heterothops; Type locality: Mongolei: Cojbalsan aimak, 80 km NV von Cojbalsan, 700 m).

  • — Smetana 1975c: 175 (Heterothops; Mongolia).

  • Distribution: Mongolia.

niger Kraatz, 1868: 352 (Heterothops; Type locality: Thüringen).

  • — Fauvel, 1874: 311 [= 1874a: 537] (Heterothops; cited as nigra; synonym of praevius).

  • — Redtenbacher, 1874: 197 (Heterothops; characters).

  • — Mulsant and Rey, 1876: 818 [= 1877: 674] (Heterothops; characters; France).

  • — Fowler, 1888: 225 (Heterothops; synonym of praevius).

  • — Ganglbauer, 1895: 387 (Heterothops; synonym of praevius).

  • — Joy, 1906a: 199 (Heterothops; notes; mole's nest; Britain).

  • — Porta, 1907: 102 (Heterothops; synonym of praevius).

  • — Montandon, 1908: 80 (Heterothops; Romania).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; synonym of praevius).

  • — Scheerpeltz, 1933: 1423 (Heterothops; variety of praevius).

  • — Paulian, 1941: 256 (Heterothops; larval characters).

  • — Smetana, 1958: 317 (Heterothops; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 204 (Heterothops; characters; central Europe).

  • — Smetana, 1964d: 78 (Heterothops; Czechoslovakia).

  • — Focarile, 1964: 57 (Heterothops; Italy).

  • — Horion, 1965: 247 (Heterothops; Denmark; Sweden; Britain; Holland; Belgium; France; Switzerland; Italy; Czechoslovakia; Poland; Germany; Austria).

  • — Pototskaia, 1967: 79 (Heterothops; larval characters).

  • — Smetana, 1967g: 471 (Heterothops; characters).

  • — Bolov, 1969a: 329 (Heterothops; collecting notes; Russia).

  • — Osella and Zanetti, 1975: 112 (Heterothops; collected from nest of Talpa europaea; notes; Italy).

  • — Pope, 1977: 30 (Heterothops; Britain).

  • — Coiffait, 1978: 312 (Heterothops; characters; Europe; Britain; Scandinavia).

  • — Israelson, 1979a: 263, 265 (Heterothops; subspecies of praevius; discussion of status; characters; collecting notes).

  • — Tóth, 1984: 100 (Heterothops; characters; Hungary).

  • — Lucht, 1987: 108 (Heterothops; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Drugmand, 1988a: 80 (Heterothops; characters).

  • — Lohse and Lucht, 1989: 171 (Heterothops; characters).

  • — Nowosad, 1990: 120 (Heterothops; collected from mole nest; notes; Poland).

  • — Janák and Vysoky, 1992: 137 (Heterothops; collected with Formica pratensis; Bohemia).

  • — Welch, 1993: 229 (Heterothops; ovariole number and ovary structure).

  • — Smetana, 1993b: 49 (Heterothops; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Heterothops; Italy).

  • Distribution: Britain, Sweden, Denmark, Holland; Belgium, France, Germany, Poland, Czechoslovakia, Switzerland, Italy, Austria, Hungary, Romania, Russia.

nidicola Thomson, 1870: 127 (Heterothops; Type locality: Bastad in nordvestra Skåne).

  • — Fauvel, 1874: 311 [= 1874a: 537] (Heterothops; synonym of praevius).

  • — Fowler, 1888: 225 (Heterothops; synonym of praevius).

  • — Ganglbauer, 1895: 387 (Heterothops; synonym of praevius).

  • — Porta, 1907: 102 (Heterothops; synonym of praevius).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; synonym of praevius).

  • — Smetana, 1958: 317 (Heterothops; synonym of niger).

  • — Coiffait, 1978: 312 (Heterothops; synonym of niger).

nigerrimus Bernhauer, 1914b: 67 (Heterothops; Type locality: Ghissar-Gebirge in Turkestan).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Scheerpeltz, 1963: 14 (Heterothops; Afghanistan).

  • — Coiffait, 1978: 316 (Heterothops; characters; Transcaspian region).

  • Distribution: Tajikistan, Afghanistan.

nigritulus R. Sahlberg, 1847: 790 (Quedius; Type locality: Rio Janeiro).

  • — Fauvel, 1891: 120 (Heterothops).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • Distribution: Brazil.

nigrofrater Lea, 1925: 236 (Heterothops; Type locality: Victoria: Warburton, Belgrave, Killara; Melbourne. Tasmania: Mount Wellington).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Australia.

nitens Nordmann, 1837: 138 (Trichopygus; Type locality: Taurus-Chersonesus).

  • — Erichson, 1840: 518 (Heterothops; characters; Chersonesus Tauricus).

  • — Redtenbacher, 1857: 199 (Heterothops; characters; Germany).

  • — Redtenbacher, 1874: 196 (Heterothops; characters; Germany).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Russia.

obscuripennis Lea, 1925: 237 (Heterothops; Type locality: South Australia: Kangaroo Island).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Australia.

occidentis Casey, 1886: 263 (Heterothops; replacement name for exilis Casey).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Smetana, 1971: 17 (Heterothops; characters; California; New Mexico).

  • — Legner and Moore, 1977: 175 (Heterothops; cited as occidentalis; found under bark).

  • Distribution: USA.

exilis Casey, 1886: 208 (Heterothops; [preoccupied]; Type locality: California: Monterey Co.).

  • — Scheerpeltz, 1933: 1423 (Heterothops; synonym of occidentis).

  • — Smetana, 1971: 17 (Heterothops; synonym of occidentis).

mediocris Fall, 1907: 220 (Heterothops; Type locality: Las Vegas Hot Springs, New Mexico; Riverside, California).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Smetana, 1971: 17 (Heterothops; synonym of occidentis).

ocularis Blackwelder, 1943: 465 (Heterothops; Type locality: Trinidad: mouth of cave in Aripo Valley, elevation 2600 feet).

  • Distribution: West Indies.

oculatus Fauvel, 1895: 275 (Heterothops; Type locality: Tenasserim, mont Mooleyit, 1800–1900 m; [Note: See lectotype designation by Smetana, 1988]).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Cameron, 1932: 267 (Heterothops; characters; Burma; India).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Scheerpeltz, 1965: 111 (Heterothops; Burma).

  • — Coiffait, 1982a: 234 (Heterothops; Pakistan).

  • — Smetana, 1988: 320 (Heterothops; lectotype designation; characters; Burma; Nepal).

  • — Smetana, 1992b: 10 (Heterothops; Nepal).

  • — Smetana, 1995: 122 (Heterothops; characters; bionomics; Taiwan).

  • Distribution: Pakistan, India, Nepal, Myanmar, Taiwan.

okapaensis Last, 1975: 431 (Heterothops; Type locality: Okapa, 5800 ft.).

  • Distribution: New Guinea.

orbicularis Sharp, 1884: 324 (Heterothops; Type locality: Guatemala, San Juan in Vera Paz; Panama, Volcan de Chiriqui 3000 to 4000 feet, Boquete 3500 feet).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • Distribution: Guatemala, Panama.

papuanus Cameron, 1937a: 104 (Quedius; subgenus Sauridus; Type locality: New Guinea: Mt. Tafa, alt. 8500 ft.).

  • — Last, 1975: 425, 438 (Heterothops; New Guinea).

  • Distribution: New Guinea.

paradoxus Mulsant and Rey, 1876, see: dissimilis Gravenhorst, 1802.

paralleliceps Bernhauer, 1917h: 87 (Heterothops; Type locality: West-Kolumbien: Quindici Pass bei Linia, 3600 m).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Scheerpeltz, 1960c: 128 (Heterothops; Bolivia).

  • Distribution: Colombia, Bolivia.

parvicornis Mulsant and Rey, 1876, see: dissimilis Gravenhorst, 1802.

parvipennis Scheerpeltz, 1972: 116 (Heterothops; Type locality: Argentinien: Rio Negro, El Bolson, Mt. Piltriquitron, 1000 m).

  • Distribution: Argentina.

persimilis Cameron, 1932: 267 (Heterothops; Type locality: Simla Hills: Kotgarh, alt. 7000 feet; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Scheerpeltz, 1965: 111 (Heterothops; Burma).

  • — Smetana, 1988: 326 (Heterothops; lectotype designation; characters; Himachal Pradesh; Uttar Pradesh; Nepal).

  • Distribution: India, Nepal, Myanmar.

picipennis Fauvel, 1877: 276 (Heterothops; Type locality: Australie, Victoria).

  • — Fauvel, 1878g: 558 (Heterothops; notes; Australia).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Lea, 1925: 231 (Heterothops; from nest of Myrmecia sp.; Australia).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Scheerpeltz, 1935a: 655 (Heterothops; Australia).

  • Distribution: Australia.

picticollis Lea, 1925: 232 (Heterothops; Type locality: Victoria: Dividing Range; Ferntree Gully; Killara).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Australia.

pictus Lea, 1925: 239 (Heterothops; Type locality: Victoria: Victorian Alps, Dividing Range, Belgrave, Ferntree Gully, Warburton, Bright, Wandin, Emerald. Tasmania, Zeehan, Western Australia: Donnybrook).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Australia.

praevius Erichson, 1839a: 480 (Heterothops; Type locality: Mark Brandenburg).

  • — Lacordaire, 1835: 385 (Emus; [Note: Misidentification: Bernhauer and Schubert, 1916: 413 cited subuliformis Boisduval and Lacordaire as a synonym of praevius Erichson; Lacordaire did not describe the species as new, he attributed it to Gravenhorst]; characters; France).

  • — Nordmann, 1837: 137 (Trichopygus; [Note: Misidentification: Bernhauer and Schubert, 1916: 413 cited dissimilis Nordmann as a synonym of praevius Erichson; Nordmann did not describe the species as new, he attributed it to Gravenhorst]; characters; Germany; Russia).

  • — Heer, 1839: 281 (Heterothops; characters; Switzerland).

  • — Erichson, 1840: 516 (Heterothops; characters; Germany).

  • — Märkel, 1841: 217 (Heterothops; in ant nest).

  • — Redtenbacher, 1849: 711 (Heterothops; characters; Austria).

  • — Rosenhauer, 1856: 71 (Philonthus; Granada).

  • — Fairmaire and Laboulbène, 1856: 533 (Heterothops; characters; France).

  • — Kraatz, 1857d: 484 (Heterothops; characters; Germany).

  • — Redtenbacher, 1857: 199 (Heterothops; characters; Austria).

  • — Fauvel, 1874: 311 [= 1874a: 537] (Heterothops; characters; France; Europe; Caspian Sea; Iran; Buchara).

  • — Redtenbacher, 1874: 196 (Heterothops; characters; Austria).

  • — Seidlitz, 1875: 263 (Heterothops; characters; Baltic region).

  • — Mulsant and Rey, 1876: 814 [= 1877: 670] (Heterothops; characters; larval characters; France).

  • — J. Sahlberg, 1876: 30 (Heterothops; Finland).

  • — Fowler, 1888: 225 (Heterothops; characters; Britain).

  • — Ragusa, 1892: 167 (Heterothops; Sicily).

  • — Ganglbauer, 1895: 387 (Heterothops; characters; Europe; eastern Mediterranean region).

  • — Fauvel, 1902b: 118 (Heterothops; Algeria).

  • — Porta, 1907: 102 (Heterothops; characters; Italy).

  • — Reitter, 1909: 108 (Heterothops; characters; Germany).

  • — Johansen, 1914: 346 (Heterothops; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Šulc, 1922 (Heterothops; osmeterium; anatomy; histology; physiology).

  • — Portevin, 1929: 336 (Heterothops; characters; France).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Bernhauer, 1936: 5 (Heterothops; Mongolia).

  • — Paulian, 1941: 256 (Heterothops; larval characters).

  • — Hinton, 1945: 69 (Heterothops; characters).

  • — Jeannel and Jarrige, 1949: 377 (Heterothops; cave species; Spain; Algeria).

  • — V. Hansen, 1952: 159 (Heterothops; characters; Denmark).

  • — Scheerpeltz, 1958a: 23 (Heterothops; Turkey).

  • — Smetana, 1958: 316 (Heterothops; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1963: 14 (Heterothops; Afghanistan).

  • — Palm, 1963b: 156 (Heterothops; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 204 (Heterothops; characters; central Europe).

  • — Scheerpeltz, 1965c: 510 (Heterothops; Sicily).

  • — Horion, 1965: 246 (Heterothops; Caucasus; Transbaikal; Afghanistan; Russia; Finland; Sweden; Norway; Britain; Denmark; France; Belgium; Holland; Germany; Austria; Czechoslovakia; Poland; Romania; Portugal; Balaearics; Italy).

  • — Pototskaia, 1967: 79 (Heterothops; larval characters).

  • — Smetana, 1967g: 470 (Heterothops; characters).

  • — Szujecki, 1968a: 733 (Heterothops; Poland).

  • — Osella and Zanetti, 1975: 110 (Heterothops; collected from nest of Talpa europaea; notes; Italy).

  • — Pope, 1977: 30 (Heterothops; Britain).

  • — Coiffait, 1978: 310 (Heterothops; characters; Europe; Britain; Scandinavia; North Africa).

  • — Israelson, 1979a: 265, 266 (Heterothops; characters).

  • — Frank, 1982: 37 (Heterothops; list of parasites).

  • — Tóth, 1984: 101 (Heterothops; characters; Hungary).

  • — Lucht, 1987: 108 (Heterothops; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Drugmand, 1988a: 80 (Heterothops; characters).

  • — Lohse and Lucht, 1989: 169 (Heterothops; characters).

  • — Nowosad, 1990: 117 (Heterothops; collected from mole nest; notes; Poland).

  • — Smetana, 1993b: 49 (Heterothops; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Heterothops; Italy).

  • — Ryabukhin, 1999: 70 (Heterothops; catalog; habitat; distribution; northeastern Russia).

  • Distribution: Algeria, Europe, Russia, Iran, Afghanistan, Uzbekistan, Mongolia.

limosus Motschulsky, 1858: 660 (Heterothops; cited as Heteropos; Type locality: Styrie).

  • — Fauvel, 1874: 311 [= 1874a: 537] (Heterothops; synonym of praevius).

  • — Ganglbauer, 1895: 387 (Heterothops; synonym of praevius).

  • — Porta, 1907: 102 (Heterothops; synonym of praevius).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; synonym of praevius).

  • — Smetana, 1958: 316 (Heterothops; synonym of praevius).

fulvipennis Motschulsky, 1860: 572 (Heterothops; Type locality: environs de St. Pétersbourg sur les bords de la Néva).

  • — Fauvel, 1874: 311 [= 1874a: 537] (Heterothops; synonym of praevius).

  • — Ganglbauer, 1895: 387 (Heterothops; synonym of praevius).

  • — Porta, 1907: 102 (Heterothops; synonym of praevius).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; synonym of praevius).

  • — Smetana, 1958: 316 (Heterothops; synonym of praevius).

  • — Coiffait, 1978: 310 (Heterothops; synonym of praevius).

primarius Last, 1975: 426 (Heterothops; Type locality: N.E. New Guinea: Mount Michael, 8–1000 ft.).

  • Distribution: New Guinea.

pumilio Nordmann, 1837, see: quadripunctulus Gravenhorst, 1806.

pusillus Coiffait, 1982: 83 (Heterothops; Type locality: Nepal: environs de Pina, region de Rara-See).

  • — Smetana, 1988: 323 (Heterothops; characters; Himachal Pradesh; Uttar Pradesh; West Bengal; Nepal).

  • — Smetana, 1992b: 10 (Heterothops; Nepal).

  • Distribution: Nepal, India.

pusio LeConte, 1863a: 35 (Heterothops; Type locality: New York).

  • — Horn, 1878: 153 (Heterothops; characters; New York; Pennsylvania; Ohio; Arizona).

  • — Blatchley, 1910: 369 (Heterothops; characters; Indiana).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Smetana, 1971: 20 (Heterothops; characters; British Columbia; Ontario; Quebec; Arizona; Arkansas; California; Illinois; Indiana; Iowa; Kentucky; Maine; Massachusetts; Michigan; Missouri; New Jersey; Ohio; Oregon; Pennsylvania; Wisconsin).

  • — Smetana, 1973: 1421 (Heterothops; Quebec).

  • — Legner and Moore, 1977: 176 (Heterothops; found under bark).

  • — Smetana, 1978a: 817 (Heterothops; collecting notes; Arizona; Arkansas; California; Oklahoma; Texas).

  • — Smetana, 1981b: 631 (Heterothops; Ontario).

  • — Smetana, 1990d: 96 (Heterothops; Saskatchewan).

  • — Downie and Arnett, 1996: 388 (Heterothops; characters; USA).

  • — Schmidt, 1999: 108 (Heterothops; food for adults and larvae used for rearing species; rearing methods).

  • Distribution: Canada, USA.

pygmaeus Gemminger and Harold, 1868, see: dissimilis Gravenhorst, 1802.

quadriceps Bernhauer, 1917h: 87 (Heterothops; Type locality: Argentinien: Prov. Tucuman).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • Distribution: Argentina.

quadripunctulus Gravenhorst, 1806: 24 (Tachinus; Type locality: Brunsvigae).

  • — Zetterstedt, 1828: 72 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1916: 413 cited subuliformis Zetterstedt as a synonym of quadripunctulus; Zetterstedt did not describe the species as new, he attributed it to Gravenhorst]; characters; Lapland).

  • — Erichson, 1839a: 481 (Heterothops; characters; Germany).

  • — Heer, 1839: 281 (Heterothops; characters; Switzerland).

  • — Erichson, 1840: 517 (Heterothops; characters; Germany).

  • — Jacobsen, 1849: 707 (Heterothops; characters; Denmark).

  • — Redtenbacher, 1849: 712 (Heterothops; cited as quadripunctatus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 533 (Heterothops; characters; France).

  • — Kraatz, 1857d: 486 (Heterothops; characters; Germany).

  • — Redtenbacher, 1857: 199 (Heterothops; characters; Austria).

  • — Thomson, 1860: 181 (Heterothops; characters; Scandinavia).

  • — Gredler, 1863: 99 (Heterothops; Austria).

  • — Fauvel, 1874: 312 [= 1874a: 538] (Heterothops; characters; France; Lapland; Scandinavia; Britain; Germany; Switzerland; Austria; Tirol; Russia).

  • — Redtenbacher, 1874: 196 (Heterothops; characters; Austria).

  • — Seidlitz, 1875: 263 (Heterothops; characters; Baltic region).

  • — Mulsant and Rey, 1876: 829 [= 1877: 685] (Heterothops; characters; France).

  • — J. Sahlberg, 1876: 30 (Heterothops; Finland).

  • — Fowler, 1888: 225 (Heterothops; characters; Britain).

  • — Ganglbauer, 1895: 388 (Heterothops; characters; north and middle Europe).

  • — Porta, 1907: 104 (Heterothops; characters; Italy).

  • — Reitter, 1909: 108 (Heterothops; characters; Germany).

  • — Johansen, 1914: 347 (Heterothops; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Portevin, 1929: 337 (Heterothops; characters; France).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • — V. Hansen, 1952: 160 (Heterothops; characters; Denmark).

  • — Smetana, 1958: 319 (Heterothops; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 157 (Heterothops; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 204 (Heterothops; characters; central Europe).

  • — Smetana, 1964d: 78 (Heterothops; Czechoslovakia).

  • — Horion, 1965: 249 (Heterothops; Russia; Finland; Sweden; Norway; Denmark; Britain; Poland; Czechoslovakia; Hungary; Hercegovina; Italy; Switzerland; Austria; Germany; France; Holland).

  • — Smetana, 1967g: 471 (Heterothops; characters).

  • — Szujecki, 1968a: 733 (Heterothops; Poland).

  • — Coiffait, 1978: 316 (Heterothops; characters; Britain; France).

  • — Israelson, 1979a: 265, 266 (Heterothops; characters).

  • — Tóth, 1984: 101 (Heterothops; characters; Hungary).

  • — Lucht, 1987: 108 (Heterothops; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Lohse and Lucht, 1989: 171 (Heterothops; characters).

  • — Smetana, 1993b: 49 (Heterothops; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Heterothops; Italy ?).

  • Distribution: Europe, Russia.

pumilio Nordmann, 1837: 138 (Trichopygus; Type locality: Berolini).

  • — Erichson, 1839a: 481 (Heterothops; synonym of quadripunctulus).

  • — Erichson, 1840: 517 (Heterothops; synonym of quadripunctulus).

  • — Redtenbacher, 1849: 712 (Heterothops; synonym of quadripunctulus).

  • — Kraatz, 1857d: 486 (Heterothops; synonym of quadripunctulus).

  • — Ganglbauer, 1895: 388 (Heterothops; synonym of quadripunctula).

  • — Porta, 1907: 104 (Heterothops; synonym of quadripunctula).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; synonym of quadripunctula).

  • — Smetana, 1958: 319 (Heterothops; synonym of quadripunctulus).

  • — Coiffait, 1978: 316 (Heterothops; synonym of quadripunctula).

rambouseki Blackwelder, 1943: 465 (Heterothops; Type locality: Cuba: district of Jamaica, province of Habana. Also cited from Jamaica).

  • Distribution: West Indies.

remotus Sharp, 1884: 323 (Heterothops; Type locality: Mexico, Mirador; Guatemala, Cerro Zunil 4000 to 5000 feet, San Geronimo; Panama, Volcan de Chiriqui 2500 to 4000 feet, Pena Blanca 3000 to 4000 feet).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • Distribution: Panama, Guatemala, Mexico.

riparius Peyerimhoff, 1929: 56 (Heterothops; Type locality: Hoggar de 1,700 a 2,400 m: Adrar Amezzeroui, Aguelmane Imerrera, Tihentekert, Arrem de Tinozel, Tezeit. Tifedest, vers 1150 m: Oued Aguelil, oued Ahetes. Haut Igharghar a Timenain. Tassili occidental, 600–800 m: Tim Tahart et Tin Eselmaken, Amguid).

  • — Peyerimhoff, 1931: 32 (Heterothops; characters; Algeria).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • — Jarrige, 1958: 92 (Heterothops; Algeria).

  • — Coiffait, 1966b: 338 (Heterothops; Mauritania).

  • — Coiffait, 1978: 311 (Heterothops; characters; Sahara).

  • Distribution: Algeria, Mauritania.

rotundiceps Sharp, 1889: 35 (Heterothops; Type locality: Nikko; Kiga).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Watanabe and Shibata, 1972: 65 (Heterothops; Yaku-shima Island).

  • — Cho, 1996b: 109 (Heterothops; characters; Korea).

  • Distribution: Japan, Korea.

rubicundulus Last, 1980: 152 (Heterothops; Type locality: New Guinea: Mt. Wilhelm, Keglsugl).

  • Distribution: New Guinea.

rufosuturalis Lea, 1925: 234 (Heterothops; Type locality: Queensland: Goodna; Brisbane; New South Wales: Tweed River).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • Distribution: Australia.

rufulus Last, 1980: 152 (Heterothops; Type locality: New Guinea: Mount Wilhelm, Field Station).

  • Distribution: New Guinea.

rufus Last, 1987: 53 (Heterothops; Type locality: Irian: Vogelkop, Fak Fak, S coast of Bomberai, 100–700 m).

  • Distribution: New Guinea.

saano Smetana, 1988: 329 (Heterothops; Type locality: India: Kashmir, Aru).

  • Distribution: India.

saltus Last, 1975: 427 (Heterothops; Type locality: Okapa, 5800 ft.).

  • Distribution: New Guinea.

saphaa Smetana, 1988: 330 (Heterothops; Type locality: Nepal: 148 Manang Dist., Marayandi, 3800–4100 m, oberhalb Manang Richtung Thorung).

  • Distribution: Nepal.

semicupreus Fauvel, 1878g: 557 (Heterothops; Type locality: Australie orientale: Queensland, Wide Bay).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Lea, 1929: 205 (Heterothops; characters).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • Distribution: Australia.

sericans Stephens, 1832, see: binotatus Gravenhorst, 1802.

sericans Mulsant and Rey, 1876, see: dissimilis Gravenhorst, 1802.

simulator Last, 1980: 153 (Heterothops; Type locality: New Guinea: Mount Wilhelm).

  • Distribution: New Guinea.

simulatrix Smetana, 1971: 37 (Heterothops; Type locality: California: Mokell. Hill, Calaveras Co.).

  • Distribution: USA.

sordidus Smetana, 1971: 32 (Heterothops; Type locality: British Columbia: 8 mi. W. Creston).

  • — Smetana, 1973: 1422 (Heterothops; Northwest Territories).

  • — Smetana, 1981b: 632 (Heterothops; collecting notes; Alaska).

  • Distribution: Canada, USA.

stiglundbergi Israelson, 1979: 259 (Heterothops; Type locality: Sweden: Lomma).

  • — Lohse and Lucht, 1989: 170 (Heterothops; characters).

  • — Silfverberg, 1992: 17 (Heterothops; Finland; Russia; Sweden; Denmark; Lithuania).

  • — Smetana, 1993b: 49 (Heterothops; Czech Republic).

  • — Assing, Frisch, Kahlen, et al., 1998: 139 (Heterothops; all central Europe).

  • — Ødegaard, 1999: 72 (Heterothops; invasive species associated with compost heaps in Sweden, Demark, Finland, and Norway).

  • — Burakowski, Mroczkowski, and Stefańska, 2000: 53 (Heterothops; catalog; Poland).

  • Distribution: Europe.

subterraneus Bernhauer, 1921c: 173 (Heterothops; Type locality: Argentinien: Prov. Buenos Aires, Monte Veloz).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • Distribution: Argentina.

tantillus Lea, 1925: 235 (Heterothops; Type locality: Queensland: Cairns district).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • Distribution: Australia.

tantulus Last, 1975: 429 (Heterothops; Type locality: Okapa, 5800 ft.).

  • Distribution: New Guinea.

tanygnathoides Reitter, 1891: 17 (Heterothops; Type locality: Kasalinsk, am Aralsee).

  • — Eppelsheim, 1892: 329 (Heterothops; Turkestan).

  • — Bernhauer and Schubert, 1916: 414 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • — Coiffait, 1978: 317 (Heterothops; cited as tanygnatoides; characters; Transcaspian region).

  • Distribution: Kazakhstan, Uzbekistan.

taurus Blackburn, 1888: 4 (Heterothops; Type locality: about 30 miles north of Port Lincoln).

  • — Blackburn, 1891: 69 (Quedius; characters; notes).

  • — Bernhauer and Schubert, 1916: 414 (Heterothops; catalog).

  • — Lea, 1925: 240 (Quedius; synonym of hybridus).

  • Distribution: Australia.

tenuicornis Sharp, 1884: 324 (Heterothops; Type locality: Guatemala, near the city).

  • — Bernhauer and Schubert, 1916: 414 (Heterothops; catalog).

  • Distribution: Guatemala.

thaxteri Bernhauer, 1917h: 86 (Heterothops; Type locality: Argentinien).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • — Frank, 1982: 37 (Heterothops; list of parasites).

  • Distribution: Argentina.

tibialis Fauvel, 1878g: 559 (Heterothops; Type locality: Australie méridionale: Victoria).

  • — Bernhauer and Schubert, 1916: 414 (Heterothops; catalog).

  • — Lea, 1925: 231 (Heterothops; characters; Australia).

  • Distribution: Australia.

titschacki Bernhauer, 1941c: 289 [= 1951: 279] (Heterothops; Type locality: Süd Peru: Palco, 2600 m).

  • Distribution: Peru.

topali Scheerpeltz, 1972: 112 (Heterothops; Type locality: Argentinien: Rio Negro, El Bolson, Mt. Piltriquitron, 1500 m).

  • Distribution: Argentina.

tovarensis Fauvel, 1891: 120 (Heterothops; Type locality: Colonie Tovar).

  • — Bernhauer and Schubert, 1916: 414 (Heterothops; catalog).

  • Distribution: Venezuela.

tumulus Herman, 2001: 45 (Heterothops; replacement name for montanus Last).

  • Distribution: New Guinea.

montanus Last, 1975: 434 (Heterothops; [preoccupied] Type locality: N.E. New Guinea: Mount Hagen, 9000 ft., Western Highlands District).

  • — Last, 1987: 55 (Heterothops; Papua New Guinea).

  • — Herman, 2001: 45 (Heterothops; synonym of tumulus).

tzaw Smetana, 1995: 124 (Heterothops; Type locality: Taiwan: near Tungpu, Nantou-Hsien, 2600 m).

  • Distribution: Taiwan.

ubiquitosus Lea, 1925: 236 (Heterothops; Type locality: Tasmania: Mount Wellington, Hobart, Huon River, Latrobe, Stanley, Ulverstone, Launceston, Frankford, Burnie. Victoria: Melbourne, Beaconfield, Emerald, Belgrave, Ferntree Gully. South Australia: Myponga, Lucindale).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • Distribution: Australia.

velatus Wendeler, 1956b: 262 (Heterothops; Type locality: Brasilien: Nova Teutonia).

  • Distribution: Brazil.

vicunianus Germain, 1903, see: Cheilocolpus.

wouei Drugmand, 1988a: 74 (Heterothops; Type locality: Vierves-sur-Viroin, Province de Namur, Belgique; also included larval characters).

  • Distribution: Belgium.

willbergi Reitter, 1891: 18 (Heterothops; Type locality: Kasalinsk).

  • — Bernhauer and Schubert, 1916: 414 (Heterothops; catalog).

  • — Coiffait, 1978: 318 (Heterothops; cited as wilbergi; characters; Transcaucasus).

  • Distribution: Kazakhstan.

wittei Bernhauer, 1927, see: Rolla.

wollastoni Israelson, 1979a: 266 (Heterothops; Type locality: S. Antao, Cape Verde Islands).

  • — Wollaston, 1867: 235 (Heterothops; [Note: Misidentified as minutus according to Israelson, 1979a: 266] characters; notes; Cape Verde Islands).

  • Distribution: Cape Verde Islands.

xantholinoides W. J. MacLeay, 1873, see: fauveli Bernhauer and Schubert, 1916.

Indoquedius [14 species; Palaearctic and Oriental Regions]

Indoquedius Blackwelder, 1952: 199 (subgenus of Quedius; name made available by Blackwelder's designation of type species [Articles 13.3, 50.1]; species included: see Cameron, 1932: 281, 300). Type species: Quedius oculatus Fauvel, fixed by original designation by Blackwelder, 1952: 199, and selected from among species included by Cameron, 1932 (see Herman, 2001).

  • — Cameron, 1932: 281, 300 (subgenus of Quedius; species included: oculatus, filicornis, bipunctatus; characters; [Note: Cameron (1932) did not designate a type species for Indoquedius so the name was unavailable (Article 13.3). Blackwelder (1952) made the name available when he designated the type species]).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Korge, 1963a: 85 (subgenus of Quedius; characters; notes).

  • — Shibata, 1973c: 63 (subgenus of Quedius; checklist of species of Taiwan).

  • — Shibata, 1984: 137 (subgenus of Quedius; checklist of species of Japan).

  • — Smetana, 1988: 300 (valid genus; characters; key to species of Himalayan region).

  • — Smetana, 1995: 112 (characters; key to and revision of species of Taiwan).

  • — Herman, 2001: 9 (type species; author of name is Blackwelder, not Cameron).

aberrans Cameron, 1932: 299 (Quedius; subgenus Raphirus; Type locality: Sikkim: Rungbong Valley).

  • — Smetana, 1988: 311 (Indoquedius; characters; India).

  • Distribution: India.

akalita Smetana, 1977, see: sikkimensis Cameron, 1932.

baliyo Smetana, 1988: 304 (Indoquedius; Type locality: Nepal: Prov. Bagmati: Malemchi, 2800 m. Also cited from India).

  • — Smetana, 1992b: 10 (Indoquedius; Nepal).

  • Distribution: Nepal, India.

bing Smetana, 1995: 113 (Indoquedius; Type locality: Taiwan: Nantou Hsien: Hwy. 14, Fengnan, 700 m).

  • Distribution: Taiwan.

bipunctatus Eppelsheim, 1895b: 390 (Quedius; Type locality: Parasnath-Gebirge; [Note: See lectotype designation by Smetana, 1988]).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Cameron, 1932: 301 (Quedius; subgenus Indoquedius; characters; Bihar).

  • — Scheerpeltz, 1933: 1442 (Quedius; synonym of grandiceps).

  • — Smetana, 1988: 305 (Indoquedius; cited as valid species; lectotype designation; characters; Himachal Pradesh; Uttar Pradesh).

  • Distribution: India.

conicus Champion, 1922, see: filicornis Eppelsheim, 1895.

cyanipennis Coiffait, 1982, see: sikkimensis Cameron, 1932.

daai Smetana, 1988: 307 (Indoquedius; Type locality: Inde: Kumaon, UP, Bhim Tal env. 1500 m. Also from Himachal Pradesh).

  • Distribution: India.

filicornis Eppelsheim, 1895c: 55 (Quedius; Type locality: Sikkim im Himalaya).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Cameron, 1932: 301 (Quedius; subgenus Indoquedius; characters; India).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Korge, 1963a: 87 (Quedius; subgenus Indoquedius).

  • — Smetana, 1988: 309 (Indoquedius; characters; Sikkim; Uttar Pradesh; Nepal).

  • — Smetana, 1992b: 10 (Indoquedius; Nepal).

  • Distribution: India, Nepal.

conicus Champion, 1922: 33 (Quedius; subgenus Raphirus; Type locality: Ranikhet Division of Kumaon).

  • — Cameron, 1932: 301 (Quedius; synonym of filicornis).

  • — Scheerpeltz, 1933: 1440 (Quedius; synonym of filicornis).

  • — Smetana, 1988: 309 (Indoquedius; synonym of filicornis).

formosae Cameron, 1949: 176 (Quedius; subgenus Indoquedius; Type locality: Formosa: Tamari, Taito).

  • — Korge, 1963a: 87 (Quedius; subgenus Indoquedius).

  • — Smetana, 1995: 114 (Indoquedius; characters; Taiwan).

  • Distribution: Taiwan.

grandiceps Kraatz, 1879: 122 (Quedius; Type locality: Amur).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1442 (Quedius; catalog).

  • — Korge, 1963a: 87 (Quedius; subgenus Indoquedius).

  • — Herman, 2001: 83 (Indoquedius).

  • Distribution: Russia.

juno Sharp, 1874: 24 (Quedius; Type locality: Yamato).

  • — Sharp, 1889: 29 (Quedius; Japan).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1445 (Quedius; catalog).

  • — Korge, 1963a: 87 (Quedius; subgenus Indoquedius).

  • — Nakane, 1963a: 94 (Quedius; characters; habitus photograph; Japan).

  • — Smetana, 1988: 300 (Indoquedius).

  • Distribution: Japan.

oculatus Fauvel, 1895: 274 (Quedius; Type locality: Bírmanie, Carin Asciuii Ghécu, 1400–1500 m; [Note: See lectotype designation by Smetana, 1988]. Also cited from Bírmanie: Carin Cheba, 900–1100 m).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; catalog).

  • — Cameron, 1932: 300 (Quedius; subgenus Indoquedius; characters; Burma; Tonkin).

  • — Scheerpeltz, 1933: 1454 (Quedius; catalog).

  • — Korge, 1963a: 87 (Quedius; subgenus Indoquedius).

  • — Smetana, 1988: 306 (Indoquedius; lectotype designation: Carin Asciuii Ghécu, 1400–1500 m; characters; Burma).

  • Distribution: Myanmar, Vietnam.

praeditus Sharp, 1889: 29 (Quedius; Type locality: Miyanoshita; Junsai).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1458 (Quedius; catalog).

  • — Korge, 1963a: 87 (Quedius; subgenus Indoquedius).

  • — Yuh, Paik, Kwon, and Lee, 1985: 249 (Quedius; subgenus Indoquedius; Korea).

  • — Smetana, 1988: 300 (Indoquedius).

  • — J. Li, 1993: 41 (Quedius; China).

  • — Cho, 1996b: 113 (Quedius; subgenus Indoquedius; characters; Korea).

  • — Watanabe, 1996a: 15 (Indoquedius; cave collection; Japan).

  • Distribution: Japan, Korea, China.

saathi Smetana, 1988: 310 (Indoquedius; Type locality: India: Meghalaya, Khasi Hills, Shillong, 1850–1950 m).

  • Distribution: India.

shibatai Smetana, 1995: 115 (Indoquedius; Type locality: Taiwan: near Fenchifu, Chiai).

  • Distribution: Taiwan.

sikkimensis Cameron, 1932: 294 (Quedius; subgenus Raphirus; Type locality: Sikkim: Lachung, alt. 9000–10,000 feet; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • — Smetana, 1988: 302 (Indoquedius; lectotype designation; characters; Bhutan; Arunachal Pradesh).

  • Distribution: India, Bhutan.

akalita Smetana, 1977a: 246 (Quedius; subgenus Indoquedius; Type locality: Bhutan: Thimphu).

  • — Smetana, 1988: 302 (Indoquedius; synonym of sikkimensis).

cyanipennis Coiffait, 1982a: 273 (Algon; Type locality: Bhoutan: Batbalithan, Bhumtang, 2600 m).

  • — Smetana, 1988: 302 (Indoquedius; synonym of sikkimensis).

Ioma [1 species; Ethiopian Region]

Ioma Blackwelder, 1952: 199 (replacement name for Tachinopsis Fauvel). Type species: Tachinopsis setigera Fauvel, fixed by objective synonymy with Tachinopsis.

Tachinopsis Fauvel, 1899: 22 (species included: setigera; preoccupied by Coquillett, 1897). Type species: Tachinopsis setigera Fauvel, fixed by monotypy.

  • — Bernhauer and Schubert, 1916: 440 (world catalog; 1 species).

  • — Blackwelder, 1952: 369 (synonym of Ioma; type species: setigera).

setigera Fauvel, 1899: 22 (Tachinopsis; Type locality: Abyssinie: Bogos, Sciotel; Haut-Sénégal: Kayes).

  • — Bernhauer and Schubert, 1916: 440 (Tachinopsis; synonym of setigera).

  • — Coiffait, 1968a: 137 (Tachinopsis; Chad).

  • — Scheerpeltz, 1974c: 25 (Tachinopsis; Sudan).

  • Distribution: Ethiopia, Sudan, Chad, Mali.

Loncovilius [7 species; Neotropical Region]

Loncovilius Germain, 1903: 439 (species included; semiflavus, lividipennis). Type species: Quedius semiflavus Fairmaire and Germain, fixed by subsequent designation by Blackwelder, 1952: 226.

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 226 (type species).

  • — Coiffait and Saiz, 1966: 401 (characters; key to Chilean species).

  • — Coiffait and Saiz, 1968: 365 (list of Chilean species).

  • — Saiz, 1971: 387, 389 (notes on biogeography; notes on phenetic relationships).

  • — Herman, 2001: 24 (synonymy).

Lienturius Coiffait and Saiz, 1966: 402 (subgenus of Loncovilius; species included: germaini, discoideus, leiocephalus, heeri). Type species: Staphylinus leiocephalus Solier, fixed by original designation.

  • — Saiz, 1971: 382 (subgenus of Loncovilius; key to species of Chile).

Apoquedius Scheerpeltz, 1972: 24, 25 (subgenus of Quedius; species included: aeneipennis, piciformis). Type species: Quedius aeneipennis Fairmaire and Germain, fixed by original designation.

  • — Herman, 2001: 24 (synonym of Loncovilius).

aeneipennis Fairmaire and Germain, 1861: 428 (Quedius; [Note: Preoccupied secondary homonym in Quedius]; Type locality: Aculco).

  • — Fauvel, 1866a: 339 [= 1866b: 90] (Quedius; characters; Chile).

  • — Germain, 1903: 435 (Quedius; characters; Chile).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Coiffait and Saiz, 1966: 404 (Loncovilius; synonym of heeri).

  • — Coiffait and Saiz, 1968: 365 (Loncovilius; synonym of heeri).

  • — Scheerpeltz, 1972: 24 (Quedius; subgenus Apoquedius; type species of subgenus; cited as valid species; Argentina).

  • — Herman, 2001: 26 (Loncovilius; valid name).

  • Distribution: Chile, Argentina.

heeri Blackwelder, 1944: 144 (Quedius; replacement name for aeneipennis Fairmaire and Germain; [Note: Unnecessary replacement name; the senior name in Quedius, aeneipennis Heer, was originally published without a description, so is an unavailable name]).

  • — Coiffait and Saiz, 1966: 404 (Loncovilius; subgenus Lienturius; characters; Chile).

  • — Saiz, 1971: 382, 383 (Loncovilius; subgenus Lienturius; characters; Chile).

  • — Herman, 2001: 26 (Loncovilius; synonym of aeneipennis).

brevis Saiz, 1971: 384 (Loncovilius; subgenus Lienturius; Type locality: Chile: Chiloé).

  • Distribution: Chile.

chilensis Bernhauer and Schubert, 1914: 332 (Philonthus; replacement name for cribripennis Germain; [Note: This name is older than germaini. It is preoccupied as secondary homonym by chilensis Solier, a name described in Staphylinus and formerly in Philonthus but now in Bisnius]).

  • — Scheerpeltz, 1933: 1344 (Philonthus; synonym of germaini).

  • — Coiffait and Saiz, 1966: 406 (Loncovilius; synonym of germaini).

  • — Coiffait and Saiz, 1968: 365 (Loncovilius; synonym of germaini).

  • — Herman, 2001: 26 (Loncovilius; valid name).

  • Distribution: Chile.

cribripennis Germain, 1903: 412 (Philonthus; [preoccupied]; Type locality: los arenales maritimos de las provincias centrales, Chile).

  • — Bernhauer and Schubert, 1914: 332 (Philonthus; synonym of chilensis).

  • — Scheerpeltz, 1933: 1344 (Philonthus; synonym of germaini).

  • — Coiffait and Saiz, 1966: 406 (Loncovilius; synonym of germaini).

  • — Coiffait and Saiz, 1968: 365 (Loncovilius; synonym of germaini).

germaini Scheerpeltz, 1933: 1344 (Philonthus; replacement name for cribripennis Germain).

  • — Coiffait and Saiz, 1966: 406 (Loncovilius; subgenus Lienturius; characters; Chile).

  • — Saiz, 1971: 382 (Loncovilius; subgenus Lienturius; characters; Chile).

  • — Herman, 2001: 26 (Loncovilius; synonym of chilensis).

cribripennis Germain, 1903, see: chilensis Bernhauer and Schubert, 1914.

discoideus Fairmaire and Germain, 1861: 427 (Heterothops; Type locality: Santiago; Quillota).

  • — Fauvel, 1866a: 337 [= 1866b: 88] (Heterothops; characters; Chile).

  • — Germain, 1903: 409, 444 (Bolitobius; cited as a new species as “discoideus mihi” with discoideus Fairmaire and Germain as a synonym; characters).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Coiffait and Saiz, 1966: 407 (Loncovilius; subgenus Lienturius; characters; Chile).

  • — Saiz, 1969: 3, 4 (Loncovilius; characters; collecting notes; Chile).

  • — Saiz, 1971: 382, 383 (Loncovilius; subgenus Lienturius; characters; Chile).

  • — Saiz, 1971a (Loncovilius; ecological notes).

  • — Saiz, Solervicens, and Ojeda, 1989: 50 (Loncovilius; collecting notes; habitus; Chile).

  • Distribution: Chile.

germaini Scheerpeltz, 1933, see: chilensis Bernhauer and Schubert, 1914.

heeri Blackwelder, 1944, see: aeneipennis Fairmaire and Germain, 1861.

leiocephalus Solier, 1849: 318 (Staphylinus; Type locality: Chile).

  • — Kraatz, 1859a: 5 (Quedius; notes).

  • — Fairmaire and Germain, 1861: 429 (Quedius; characters; Chile).

  • — Fauvel, 1864a: 124 (Quedius; notes; Chile).

  • — Fauvel, 1866a: 338 [= 1866b: 89] (Quedius; characters; Chile).

  • — Germain, 1903: 428 (Quedius; characters; notes; Chile).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; catalog).

  • — Coiffait and Saiz, 1966: 403 (Loncovilius; subgenus Lienturius; characters; Chile).

  • — Saiz, 1969a: 9 (Loncovilius; subgenus Lienturius; notes).

  • — Saiz, 1971: 382, 383 (Loncovilius; subgenus Lienturius; characters; Chile).

  • Distribution: Chile.

lividipennis Fairmaire and Germain, 1861: 430 (Quedius; Type locality: Chillan).

  • — Fauvel, 1866a: 338 [= 1866b: 89] (Quedius; characters; Chile).

  • — Germain, 1903: 443 (Loncovilius; characters).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; catalog).

  • — Coiffait and Saiz, 1966: 409 (Loncovilius; subgenus Loncovilius; characters; Chile).

  • — Saiz, 1971: 385 (Loncovilius; subgenus Loncovilius; Chile).

  • — Scheerpeltz, 1972: 23 (Quedius; subgenus Loncovilius; Argentina).

  • Distribution: Chile, Argentina.

semiflavus Fairmaire and Germain, 1861: 428 (Quedius; Type locality: Concepcion).

  • — Gemminger and Harold, 1868: 570 (Quedius; synonym of lividipennis).

  • — Germain, 1903: 443 (Loncovilius; characters).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; catalog).

  • — Coiffait and Saiz, 1966: 411 (Loncovilius; subgenus Loncovilius; characters; Chile).

  • — Saiz, 1971: 385 (Loncovilius; subgenus Loncovilius; Chile).

  • Distribution: Chile.

Lonia [1 species; Australian Region]

Lonia E. Strand, 1943: 96 (replacement name for Colonia Olliff). Type species: Colonia regalis Olliff, fixed by objective synonymy with Colonia.

  • — Blackwelder, 1952: 227 (type species).

Colonia Olliff, 1887: 493 (species included: regalis; preoccupied by Gray, 1829, and Schaufuss, 1850). Type species: Colonia regalis Olliff, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 397 (world catalog; 1 species).

  • — Blackwelder, 1952: 102 (type species: regalis).

regalis Olliff, 1887: 494 (Colonia; Type locality: Lismore, Richmond River, New South Wales).

  • — Bernhauer and Schubert, 1914: 397 (Colonia; catalog).

  • — Lea, 1925: 229 (Colonia; Australia).

  • Distribution: Australia.

Mimosticus [1 species; Neotropical Region]

Mimosticus Sharp, 1884: 327 (species included: viridipennis). Type species: Mimosticus viridipennis Sharp, fixed by monotypy.

  • — Bernhauer and Schubert, 1916: 437 (world catalog; 1 species).

  • — Blackwelder, 1944: 145 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 249 (type species: viridipennis).

viridipennis Sharp, 1884: 328 (Mimosticus; Type locality: Panama, Volcan de Chiriqui 2000 to 3000 feet).

  • — Bernhauer and Schubert, 1916: 437 (Mimosticus; catalog).

  • Distribution: Panama.

Paratolmerus [2 species; Palaearctic and Oriental Regions]

Paratolmerus Cameron, 1932: 169 (species included: pilosiventris). Type species: Paratolmerus pilosiventris Cameron, fixed by monotypy.

  • — Scheerpeltz, 1933: 1376 (world catalog supplement).

  • — Blackwelder, 1952: 294 (type species: pilosiventris).

  • — Smetana, 1988: 335 (characters).

pilosiventris Cameron, 1932: 170 (Paratolmerus; Type locality: Sikkim: Gopaldhara, Rungbong Valley; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1376 (Paratolmerus; catalog).

  • — Smetana, 1988: 336 (Paratolmerus; lectotype designation; characters; Sikkim; Nepal).

  • Distribution: India, Nepal.

siamensis Rougemont, 1991: 221 (Paratolmerus; Type locality: Thailand: Kanchanaburi Prov.: river Kwai Noi at Ban Sai Yok).

  • Distribution: Thailand.

Parisanopus [1 species; Neotropical Region]

Parisanopus Brèthes, 1900: 216 (species included: castaneipennis). Type species: Parisanopus castaneipennis Brèthes, fixed by monotypy.

  • — Bernhauer and Schubert, 1916: 415 (world catalog; 1 species).

  • — Scheerpeltz, 1933: 1426 (world catalog supplement).

  • — Blackwelder, 1944: 144 (in checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 294 (type species: castaneipennis).

castaneipennis Brèthes, 1900: 216 (Parisanopus; Type locality: Palermo, Buenos Aires).

  • — Bernhauer and Schubert, 1916: 415 (Parisanopus; catalog).

  • — Scheerpeltz, 1933: 1426 (Parisanopus; catalog; Argentina; Paraguay).

  • Distribution: Argentina, Paraguay.

Philonthellus [1 species; Neotropical Region]

Philonthellus Bernhauer, 1939: 13 (subgenus of Philonthus; species included: chilenus). Type species: Philonthus (Philonthellus) chilenus Bernhauer, by monotypy.

  • — Blackwelder, 1952: 301 (type species: chilenus).

  • — Coiffait and Saiz, 1968: 347, 349 (valid genus; characters).

chilenus Bernhauer, 1939: 13 (Philonthus; subgenus Philonthellus; Type locality: Isla Calbuco, Süd-Chile).

  • — Coiffait and Saiz, 1968: 349 (Philonthellus; characters; Chile).

  • — Saiz, 1971b: 341 (Philonthellus; Chile).

  • — Scheerpeltz, 1972: 18 (Philonthus; Argentina).

  • Distribution: Chile, Argentina.

evitendus Tottenham, 1939d: 234 (Philonthus; replacement name for chilensis Bernhauer; [Note: Unnecessary replacement name, Bernhauer used chilenus]).

  • — Coiffait and Saiz, 1968: 349 (Philonthellus; synonym of chilenus).

evitendus Tottenham, 1939, see: chilenus Bernhauer, 1939.

Pseudorientis [3 species; Palaearctic Region]

Pseudorientis Watanbe, 1970: 70 (species included: shinobuae). Type species: Pseudorientis shinobuae Watanabe, fixed by original designation and monotypy.

  • — Tikhomirova, 1973a: 188 (checklist of species of USSR).

  • — Smetana, 1977b: 181 (characters).

  • — Shibata, 1984: 115 (checklist of species of Japan).

  • — Smetana, 1995c: 345 (key to species).

gongga Smetana, 1995c: 344 (Pseudorientis; Type locality: China: Sichuan, Gongga Shan above Camp 3, 3050 m).

  • — Smetana, 1998d: 98 (Pseudorientis; characters of female; China).

  • Distribution: China.

shinobuae Watanabe, 1970: 72 (Pseudorientis; Type locality: Japan: Honshu: Gumma Pref.: near Lake Marunuma, Oku-Nikko).

  • Distribution: Japan.

uenoi Smetana, 1995c: 341 (Pseudorientis; Type locality: China: Yunnan, Heishui, 35 km N Lijiang, 27°13′N, 100°19′E).

  • Distribution: China.

Quediocafus [3 species; Australian Region]

Quediocafus Cameron, 1945d: 791 (species included: hudsoni). Type species: Quediocafus hudsoni Cameron, fixed by original designation and monotypy.

  • — Blackwelder, 1952: 334 (type species: hudsoni).

  • — Klimaszewski, Newton, and Thayer, 1996: 151 (endemic to New Zealand).

hudsoni Cameron, 1945d: 791 (Quediocafus; Type locality: New Zealand: Hokitika).

  • Distribution: New Zealand.

insolitus Sharp, 1886a: 379 (Quedius; Type locality: New Zealand; Dunedin).

  • — Broun, 1893: 1029 (Quedius; characters; New Zealand).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • — Newton, 1996: 160 (Quediocafus).

  • — Marris, 2000: 175 (Quediocafus; habitat; Bollons Island [Antipodes]; also known from New Zealand).

  • Distribution: New Zealand, Antipodes Islands.

taieriensis Broun, 1894: 424 (Quedius; Type locality: Taieri, Otago).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • — Newton, 1996: 160 (Quediocafus).

  • Distribution: New Zealand.

Quediomacrus [2 species; Neotropical Region]

Quediomacrus Sharp, 1884: 339 (species included: puniceipennis, pollens). Type species: Quedius puniceipennis Solsky, fixed by subsequent designation by R. Lucas, 1920: 563.

  • — Bernhauer and Schubert, 1916: 410 (world catalog; 2 species).

  • — Blackwelder, 1944: 144 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 334 (type species).

pollens Sharp, 1884: 339 (Quediomacrus; Type locality: Guatemala, near the city and at Purula).

  • — Bernhauer and Schubert, 1916: 410 (Quediomacrus; catalog).

  • Distribution: Guatemala.

puniceipennis Solsky, 1868a: 123 (Quedius; Type locality: Mexique).

  • — Sharp, 1884: 339 (Quediomacrus; Mexico).

  • — Bernhauer and Schubert, 1916: 410 (Quediomacrus; catalog).

  • — Smetana, 1971b: 4 (Quediomacrus; Mexico).

  • — Frank, 1982: 45 (Quediomacrus; list of parasites).

  • Distribution: Mexico.

Quediomimus [1 species; Australian Region]

Quediomimus Cameron, 1948c: 724 (species included: brookesi). Type species: Quediomimus brookesi Cameron, fixed by original designation and monotypy.

brookesi Cameron, 1948c: 725 (Quediomimus; Type locality: New Zealand: Katikati: Bay of Plenty).

  • — Klimaszewski, Newton, and Thayer, 1996: 151 (Quediomimus; endemic to New Zealand).

  • Distribution: New Zealand.

Quediopsis [2 species; Australian Region]

Quediopsis Fauvel, 1878g: 560 (species included: lugubris, abdominalis). Type species: Quediopsis lugubris Fauvel, fixed by subsequent designation by Blackwelder, 1952: 335.

  • — Duvivier, 1883: 136 (catalog).

  • — Bernhauer and Schubert, 1916: 437 (world catalog; 2 species).

abdominalis Fauvel, 1878g: 561 (Quediopsis; Type locality: Australie méridionale, Victoria).

  • — Bernhauer and Schubert, 1916: 437 (Quediopsis; catalog).

  • Distribution: Australia.

lugubris Fauvel, 1878g: 560 (Quediopsis; Type locality: Australie méridionale, Victoria).

  • — Bernhauer and Schubert, 1916: 437 (Quediopsis; catalog).

  • Distribution: Australia.

Quedius [789 species (3 doubtful); Neotropical, Nearctic, Ethiopian, Palaearctic, Oriental, and Australian Regions]

Quedius Stephens, 1829a: 22 (species included: tristis, variabilis, fulvipes, impressus, rufitarsis, nitidus and cyanipennis along with 32 unavailable names; [Note: The same questions and controversy concerning the authorship and date of publication for Philonthus apply to Quedius. The issue is discussed in my “note” attached to the citation of the original description for Philonthus]). Type species: Staphylinus levicollis Brullé, fixed by subsequent designation in Opinion 1851 (ICZN, 1996: 215) [Note: Curtis, (1837: pl. 638) fixed Staphylinus tristis Gravenhorst as the type species Quedius. Gravenhorst, who misidentified the species, attributed the name to Fabricius so his use of the name is unavailable. Blackwelder (1952: 335) fixed Quedius fulgidus Fabricius, a species that was in Staphylinus, Ocypus, or Pseudocypus since its description. The problem is discussed by Newton, 1995: 49 and action by the Commission resolved the problem].

  • — Curtis, 1829: 24 (catalog; Britain).

  • — Stephens, 1829: 277 (catalog; Britain).

  • — Stephens, 1832: 200, 214 (characters).

  • — Curtis, 1837: pl. 638 (characters; type species; habitat; larval characters).

  • — Westwood, 1838a: 16 (characters; type species).

  • — Erichson, 1839a: 483 (characters).

  • — Stephens, 1839: 385 (characters).

  • — Erichson, 1840: 523 (characters).

  • — Schaum, 1852: 26 (list of species; Europe).

  • — Lacordaire, 1854: 84 (characters; notes; list of species).

  • — Fairmaire and Laboulbène, 1856: 535 (characters).

  • — Jacquelin du Val, 1856: 37 (characters).

  • — Giebel, 1856: 72 (fossil species).

  • — Kraatz, 1857d: 486 (characters).

  • — Redtenbacher, 1857: 199 (characters; key to Austrian species).

  • — Thomson, 1858: 30 (characters).

  • — G. Waterhouse, 1858: 22 (catalog of British species).

  • — Thomson, 1859: 25 (characters; type species: molochinus).

  • — Schaum, 1859: 25 (catalog; European species).

  • — Thomson, 1860: 173 (characters).

  • — LeConte, 1861a: 64 (characters).

  • — LeConte, 1863: 23 (list of North American species).

  • — Gredler, 1863: 99 (list of species from Tirol).

  • — Fauvel, 1866a: 337 [= 1866b: 88] (characters).

  • — Wencker and Silbermann, 1866: 27 (list of species; collecting notes; France).

  • — Gemminger and Harold, 1868: 568 (catalog).

  • — Jacquelin du Val, 1868: 69 (catalog; European species).

  • — Sharp, 1871d: 11 (list of British species).

  • — Fauvel, 1874: 266 [= 1874a: 492] (characters; larval characters; key to species).

  • — Redtenbacher, 1874: 197 (characters; key to Austrian species).

  • — Oustalet, 1874: 171 (characters; description of fossil species).

  • — Fauvel, 1875a: XXXII [= 1875b: 234] (catalog).

  • — Mulsant and Rey, 1876: 613 [= 1877: 469] (characters; key to French species).

  • — Sharp, 1876: 100 (notes).

  • — Fauvel, 1877: 267 (notes; key to species).

  • — Provancher, 1877: 244, 728 (characters; key to species).

  • — Fauvel, 1878f: 272 (characters; notes; key to New Guinea species).

  • — Horn, 1878: 154 (characters; notes; key to North American species).

  • — Heyden, 1880: 71 (list of species of Siberian region).

  • — Duvivier, 1883: 137 (catalog).

  • — Sharp, 1884: 328 (notes; Mexican and Central American species).

  • — Lynch, 1884: 119 (characters).

  • — Sharp, 1888: 226 (adult and larval characters; key to British species).

  • — Seidlitz, 1891: 395 (key to some European species).

  • — Seidlitz, 1891a: 421 (key to some European species).

  • — Heyden, Reitter, and Weise, 1891: 99 (list of species of Europe and Caucasus).

  • — Heyden, 1893: 41 (list of species of Siberian region).

  • — Ganglbauer, 1895: 391 (characters; key to middle European species).

  • — Heyden, 1896: 32 (list of species of Siberian region).

  • — Fauvel, 1897d: 305 (catalog of species of Barbary; Canaries).

  • — Germain, 1903: 414 (characters; notes).

  • — Heyden, Reitter, and Weise, 1906: 172 (list of species of Europe and Caucasus).

  • — Porta, 1907: 90 (key to Italian species).

  • — Reitter, 1908: 106, 108 (characters; key to subgenera).

  • — Blatchley, 1910: 369 (characters; key to Indiana species).

  • — Petri, 1912: 76 (list of species of Siebenbürgen).

  • — Johansen, 1914: 349 (characters; key to species).

  • — Casey, 1915: 398 (type species: Quedius molochinus).

  • — Bernhauer and Schubert, 1916: 417 (world catalog; 278 species).

  • — Leng, 1920: 109 (catalog of North American species).

  • — Munster, 1923: 186 (characters; key to subgenera and Scandinavian species).

  • — Gridelli, 1924d (anatomy; characters; key to Palaearctic subgenera and species; revision of species).

  • — Winkler, 1925: 388 (catalog for Palaearctic region).

  • — Cameron, 1925: 72 (catalog of Indian species).

  • — Porta, 1926: 109 (characters; key to species of Italy).

  • — Bruch, 1928a: 429 (catalog; Argentina).

  • — Portevin, 1929: 337 (characters; key to French species).

  • — Roubal, 1930: 387 (catalog; Slovakia).

  • — Cameron, 1932: 280 (characters; key to subgenera and species of British India).

  • — Joy, 1932: 108 (key to British species).

  • — Scheerpeltz, 1933: 1427 (world catalog supplement).

  • — Normand, 1935: 372 (list of species of Tunisia).

  • — Voris, 1939 (egg, larval, and pupal characters).

  • — Paulian, 1941: 259 (larval characters).

  • — Blackwelder, 1944: 144 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Kloet and Hincks, 1945: 167 (list of British species).

  • — Jeannel and Jarrige, 1949: 377 (cave species; notes).

  • — Tottenham, 1949: 376, 421 (type species: levicollis Brullé = tristis Gravenhorst; checklist of British species).

  • — V. Hansen, 1952: 115 (characters; key to Danish species).

  • — Hatch, 1957: 215 (characters; key to subgenera and species of Pacific Northwest).

  • — Kocher, 1958: 157 (checklist of species; Morocco).

  • — Smetana, 1958: 327 (characters; type species: tristis Gravenhorst; key to Czechoslovakian species).

  • — Smetana, 1959a: 215 (list of species collected in Albania).

  • — Smetana, 1960e: 150 (key to European species and subgenus).

  • — Coiffait, 1961a: 58 (key to and list of species of molochinus complex).

  • — Scheerpeltz, 1962b: 267 (characters).

  • — Ferreira, 1962a: 18 (list of Portuguese species).

  • — Coiffait, 1963a: 372 (key to subgenera).

  • — Székessy, 1963: 61 (characters).

  • — Palm, 1963b: 124 (characters; key to subgenera and species of Sweden).

  • — Lohse, 1964: 205 (characters; key to central European subgenera and species).

  • — Smetana, 1965: 38 (key to Nearctic species of subgenus Quedius).

  • — Coiffait and Saiz, 1966: 411 (Chilean species).

  • — Pototskaia, 1967: 79 (larval characters).

  • — Coiffait and Saiz, 1968: 365 (list of Chilean species).

  • — Scheerpeltz, 1968a: 87 (catalog of Austrian species).

  • — Bolov, 1969a: 328 (collecting notes for some species; Russia).

  • — Kasule, 1970: 57, 59 (larval characters; notes).

  • — Smetana, 1971 (revision of and key to subgenus and species of America north of Mexico; characters).

  • — Korge, 1971a: 37 (characters).

  • — Shibata, 1973c: 63 (checklist of species of Taiwan).

  • — Bordoni, 1973b: 41 (key to subgenera).

  • — Bordoni, 1973j: 710 (list of species from islands near Sicily).

  • — Tikhomirova, 1973a: 188 (checklist of species of USSR).

  • — Moore and Legner, 1975: 56 (catalog; North America north of Mexico).

  • — Bordoni, 1976b: 88 (revision of and key to Italian species).

  • — Pototskaia, 1976: 171 (larval characters).

  • — Smetana, 1977b: 181 (characters).

  • — Pope, 1977: 31 (list of British species).

  • — Coiffait, 1978: 9 (adult and larval characters; key to subgenera and species of western Palaearctic region).

  • — Smetana, 1978b: 87 (key to species of Pacific coastal area of Siberia, from Kamchatka Peninsula northwards).

  • — Topp, 1978: 312 (larval characters).

  • — Moore and Legner, 1979: 47 (characters; notes).

  • — Muona, 1979: 15 (list of Scandinavian species).

  • — Uhlig, 1979: 255 (some species collected in Germany).

  • — M. Dvořák, 1979: 116 (some species collected in Slovakia).

  • — Uhlig, Vogel, and Sieber, 1980: 248 (some species collected in Germany).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 119 (catalog; Poland).

  • — Spahr, 1981: 96 (references to specimens from amber and copal).

  • — Uhlig and Vogler, 1981: 122 (list of some species collected in Germany; habitat and collecting notes).

  • — Hammond, 1984: 207 (checklist of species of Borneo).

  • — Tóth, 1984: 102 (characters; habitus; key to species of Hungary).

  • — Shibata, 1984: 119 (checklist of species of Japan).

  • — Frank, 1986a: 371 (checklist of species of Florida).

  • — Shibata, 1986a: 125 (supplemental list of species from Taiwan).

  • — Muona and Viramo, 1986: 13 (list of species of northeastern Finland).

  • — Segers, 1986: 49 (checklist of species; Belgium).

  • — Dettner, 1987: 30 (no defensive glands).

  • — Ádám, 1987: 152 (list of six species with collecting notes; Hungary).

  • — Smetana, 1988: 181 (characters; notes; key to genera of Himalayan region; discussion of difficulty of defining subgenera).

  • — Lohse and Lucht, 1989: 171 (notes; supplement to key to species of Lohse, 1964).

  • — Angelini, 1991: 193 (list of species from high elevations of Calabria, Italy).

  • — M. Hansen, Kristensen, Mahler, Pedersen, 1991: 107 (list of Danish species).

  • — Campbell and Davies, 1991: 121 (checklist of species; Alaska and Canada).

  • — Silfverberg, 1992: 17 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Siitonen, 1993: 230 (list of Finnish species).

  • — Welch, 1993: 229 (ovariole structure and development).

  • — Smetana, 1993b: 49 (list of species of Czech Republic and Slovakia).

  • — Hernández, Outerelo, and Gamarra, 1994: 188 (checklist of species; Canary Islands).

  • — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 29 (additional records to Danish list of species).

  • — Smetana, 1995: 14, 24 (revision of species of Taiwan; key to subgenera and species of Taiwan; cladistic analysis).

  • — Newton, 1995: 51 (requested that the International Commission on Zoological Nomenclature set aside all previous designations of type species for Quedius and designate Staphylinus levicollis Brullé as type species).

  • — M. Hansen, 1996: 107 (list of Danish species).

  • — Ciceroni and Zanetti, 1995: 31 (list of species of Italy).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 244 (additions to list of Danish species).

  • — Cho, 1996b: 110 (key to species of Korea).

  • — ICZN, 1996: 215 (type species designated as Staphylinus levicollis Brullé; placed on Official List of Generic Names in Zoology).

  • — Downie and Arnett, 1996: 388 (characters; key to subgenera and species of Northeastern North America).

  • — Lundgren, 1998: 52 (list of species from Florida; records for other States listed for each species).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (list of species collected in 1997; Denmark).

  • — Staniec, 1999a: 55 (key to known pupae of Poland).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 393 (91 Nearctic species; notes; characters in key).

  • — Naomi, Kuranishi, Saito, and Maruyama, 2000: 110 (collected from Kuril Islands and Kamchatka).

  • — Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 60 (necrophilous species in Mexico).

Raphirus Stephens, 1829a: 23 (species included: semiobscurus, attenuatus, praecox along with six unavailable names). Type species: Staphylinus attenuatus Gravenhorst, fixed by subsequent designation by Westwood, 1838a: 16.

  • — Stephens, 1829: 282 (catalog; Britain).

  • — Stephens, 1832: 200 (characters).

  • — Stephens, 1833: 241 (characters).

  • — Westwood, 1838a: 16 (characters; type species).

  • — Erichson, 1839a: 483 (synonym of Quedius).

  • — Stephens, 1839: 385 (characters).

  • — Lacordaire, 1854: 84 (synonym of Quedius).

  • — Kraatz, 1857d: 486 (synonym of Quedius).

  • — G. Waterhouse, 1858: 22 (synonym of Quedius).

  • — Thomson, 1859: 26 (characters; type species: attenuatus).

  • — Thomson, 1860: 177 (characters).

  • — Mulsant and Rey, 1876: 759 [= 1877: 615] (subgenus of Quedius; characters; key to species).

  • — Ganglbauer, 1895: 391 (characters; subgenus of Quedius).

  • — Reitter, 1909: 109 (subgenus of Quedius; characters; key to species).

  • — Casey, 1915: 398, 416 (characters; type species: fulvicollis; notes; key to species).

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Gridelli, 1924d: 87 (subgenus of Quedius; characters; key to species).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Scheerpeltz, 1933: 1429 (synonym of Quedius).

  • — Tottenham, 1945: 70 (Genus dubium because attenuatus, the species designated as type species, was misidentified by Stephens).

  • — Blackwelder, 1952: 336 (type species).

  • — Hatch, 1957: 215, 228 (subgenus of Quedius; characters; key to species of Pacific Northwest).

  • — Smetana, 1958: 327, 366 (synonym of Quedius).

  • — Lohse, 1964: 212 (subgenus of Quedius; characters; key to central European species).

  • — Smetana, 1971: 39, 183 (subgenus of Quedius; characters; notes; key to North American species).

  • — Coiffait, 1978: 9, 13, 125 (subgenus of Quedius; characters; key to species of western Palaearctic region).

  • — Smetana, 1988: 231 (notes; key to species of Himalayan region).

  • — Shibata, 1984: 134 (subgenus of Quedius; checklist of species of Japan).

  • — Ádám, 1987: 152 (list of two species with collecting notes; Hungary).

  • — Lohse and Lucht, 1989: 171 (subgenus of Quedius; notes).

  • — Smetana, 1995: 24, 25, 65 (subgenus of Quedius; characters; key to species of Taiwan).

Microsaurus Dejean, 1833: 61 (species included: lateralis, fuliginosus, molochinus, nitidus, scitus, laevigatus, ochripennis, praecox, attenuatus, scintillans, boops). Type species: Staphylinus lateralis Gravenhorst, fixed by subsequent designation by Westwood, 1838a: 16.

  • — Stephens, 1835: 435 (characters).

  • — Westwood, 1838a: 16 (characters; type species).

  • — Stephens, 1839: 384 (characters).

  • — Lacordaire, 1854: 84 (synonym of Quedius).

  • — Kraatz, 1857d: 486 (synonym of Quedius).

  • — Thomson, 1859: 25 (characters; type species: lateralis).

  • — Thomson, 1860: 174 (characters).

  • — Mulsant and Rey, 1876: 685 [= 1877: 541] (subgenus of Quedius; characters; key to species).

  • — Ganglbauer, 1895: 391 (characters; subgenus of Quedius).

  • — Reitter, 1909: 108, 109 (subgenus of Quedius; characters; key to species).

  • — Casey, 1915: 398, 408 (characters; type species: Quedius fulgidus; key to species).

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Gridelli, 1924d: 18 (subgenus of Quedius; revision of concept; characters; key to species).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Tottenham, 1949: 376, 421 (type species: lateralis Gravenhorst; subgenus of Quedius; checklist of British species).

  • — Blackwelder, 1952: 247 (type species).

  • — Hatch, 1957: 215, 219 (subgenus of Quedius; characters; key to species of Pacific Northwest).

  • — Smetana, 1958: 327 (subgenus of Quedius).

  • — Lohse, 1964: 206 (subgenus of Quedius; characters; key to central European species).

  • — Smetana, 1971: 38, 49 (subgenus of Quedius; characters; key to species of Canada and USA).

  • — Bordoni, 1973g: 124 (subgenus of Quedius; characters; key to species groups).

  • — Bordoni, 1974: 1 (characters; notes; anatomy; key to species and species groups of Italy).

  • — Smetana, 1976: 113 (subgenus of Quedius; key to species of Mexico).

  • — Coiffait, 1978: 9, 13, 15 (subgenus of Quedius; characters; key to species of western Palaearctic region).

  • — Shibata, 1984: 119 (subgenus of Quedius; checklist of species of Japan).

  • — Ádám, 1987: 152 (cited as valid genus; list of four species with collecting notes; Hungary).

  • — Smetana, 1988: 185 (subgenus of Quedius; characters; key to species of Himalayan region).

  • — Lohse and Lucht, 1989: 171 (subgenus of Quedius; notes).

  • — Smetana, 1995: 24, 25 (subgenus of Quedius; characters; revision of and key to species of Taiwan).

  • — Smetana, 1996: 19 (subgenus of Quedius; key to species of apicicornis and beesoni species groups from China).

Aemulus Gistel, 1834: 8 (species included: lateralis, fuliginosus, molochinus, nitidus, scitus, boops). Type species: Staphylinus fuliginosus Gravenhorst, fixed by subsequent designation by Blackwelder, 1952: 40.

  • — Blackwelder, 1952: 40 (synonym of Quedius).

  • — Smetana, 1971: 38 (synonym of Quedius).

Thanatomanes Gistel, 1856: 388 (species included: lateralis, fulgidus, xanthopus, impressus, molochinus, fuliginosus, boops, carbonarius, attenuatus). Type species: Staphylinus impressus Panzer, fixed by subsequent designation by Blackwelder, 1952: 384 [Note: Blackwelder cited Gravenhorst as the author of the type species of Thanatomanes; Gravenhorst did not describe impressus, he attributed it to Panzer, furthermore, Gistel cited Panzer as the author].

  • — Blackwelder, 1952: 384 (synonym of Quedius).

  • — Smetana, 1971: 38 (synonym of Quedius).

Ediquus Mulsant and Rey, 1876: 615, 616 [= 1877: 471, 472] (subgenus of Quedius; species included: longicornis, microps). Type species: Quedius microps Gravenhorst, fixed by subsequent designation by Tottenham, 1939d: 229.

  • — Ganglbauer, 1895: 391 (characters; subgenus of Quedius).

  • — Reitter, 1909: 108, 109 (subgenus of Quedius; characters; key to species).

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Gridelli, 1924d: 18 (synonym of Microsaurus).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Scheerpeltz, 1933: 1428 (synonym of Quedius).

  • — Tottenham, 1939d: 229 (type species designation: microps).

  • — Tottenham, 1949: 376, 422 (type species: microps Gravenhorst; subgenus of Quedius; checklist of British species).

  • — Blackwelder, 1952: 144 (type species).

  • — Smetana, 1958: 327 (synonym of Quedius).

  • — Smetana, 1971: 38 (synonym of Quedius).

  • — Coiffait, 1978: 9 (synonym of Quedius subgenus Microsaurus).

  • — Smetana, 1988: 181 (synonym of Quedius subgenus Microsaurus).

  • — Smetana, 1995: 24 (synonym of Quedius).

Sauridus Mulsant and Rey, 1876: 616, 700 [= 1877: 472, 556] (subgenus of Quedius; species included: ochropterus, picipes, nigriceps, robustus, peltatus, dubius, montanus, anceps, cyanescens, umbrinus, suturalis, limbatus, bicolor, oblitteratus, maurorufus, riparius, kraatzi, lucidulus, scintillans). Type species: Staphylinus picipes Mannerheim, fixed by subsequent designation by Tottenham, 1940: 50.

  • — Ganglbauer, 1895: 391 (characters; subgenus of Quedius).

  • — Reitter, 1909: 109 (characters).

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Scheerpeltz, 1933: 1429 (synonym of Quedius).

  • — Tottenham, 1940: 50 (type species: picipes).

  • — Tottenham, 1949: 377, 423 (type species: picipes Mannerheim; subgenus of Quedius; checklist of British species).

  • — Blackwelder, 1952: 344 (synonym of Raphirus; type species).

  • — Hatch, 1957: 215, 226 (subgenus of Quedius; characters; key to species of Pacific Northwest).

  • — Coiffait, 1963a: 373 (subgenus of Quedius; key to species; key to species groups and species using aedeagus).

  • — Coiffait, 1978: 9, 13, 67 (subgenus of Quedius; characters; key to species of western Palaearctic region).

  • — Ádám, 1987: 153 (cited as valid genus; list of two species with collecting notes; Hungary).

  • — Smetana, 1988: 181 (synonym of Quedius subgenus Raphirus).

  • — Lohse and Lucht, 1989: 171 (subgenus of Quedius; notes).

  • — Smetana, 1993b: 50 (synonym of Quedius subgenus Raphirus).

Quedionuchus Sharp, 1884: 336 (species included: laevigatus, impunctus, spinipes, femoralis, angustus, nigerrimus). Type species: Quedius impunctus Solsky, fixed by subsequent designation by Blackwelder, 1952: 334.

  • — Sharp, 1888: 245 (characters; key to British species).

  • — Fauvel, 1895: 274 (synonym of Quedius).

  • — Ganglbauer, 1895: 391 (characters; subgenus of Quedius).

  • — Reitter, 1909: 108, 111 (subgenus of Quedius; characters; key to species).

  • — Casey, 1915: 397, 399 (notes; type species: Quedius laevigatus).

  • — Gridelli, 1924d: 74 (subgenus of Quedius; key to species).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Joy, 1932: 107 (key to British species).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Tottenham, 1939d: 229 (type species designation: laevigatus).

  • — Tottenham, 1949: 377, 422 (type species: plagiatus Mannerheim = laevigatus Gyllenhal; subgenus of Quedius; checklist of British species).

  • — Blackwelder, 1952: 334 (subgenus of Quedius).

  • — Hatch, 1957: 215, 229 (subgenus of Quedius; characters; species of Pacific Northwest).

  • — Smetana, 1958: 328, 357 (subgenus of Quedius).

  • — Korge, 1964: 118 (key to species of western Palaearctic).

  • — Kasule, 1970: 57, 59 (cited as valid genus; larval characters).

  • — Smetana, 1971: 39, 147 (subgenus of Quedius; characters; key to species of Canada and USA).

  • — Bordoni, 1973b: 45 (subgenus of Quedius; characters).

  • — Smetana, 1976a (subgenus of Quedius; review of and key to species of Central America).

  • — Coiffait, 1978: 9, 13, 53 (subgenus of Quedius; characters; key to species of western Palaearctic region).

  • — Shibata, 1984: 129 (subgenus of Quedius; checklist of species of Japan).

  • — Lohse and Lucht, 1989: 171 (subgenus of Quedius; notes).

Ediquus Reitter, 1887: 211 (subgenus of Quedius; species included: przewalskii; preoccupied by Mulsant and Rey, 1876). Type species: Quedius (Ediquus) przewalskii Reitter, fixed by monotypy.

  • — Blackwelder, 1952: 144 (type species: przewalskii).

Prionidus Bernhauer, 1907a: 287 (subgenus of Quedius; species included: iheringi; preoccupied by Uhler, 1886). Type species: Quedius (Prionidus) iheringi Bernhauer, fixed by monotypy.

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Blackwelder, 1952: 321 (type species: iheringi).

Euryquedius Reitter, 1909: 108 (subgenus of Quedius; species included: curtus). Type species: Quedius curtus Erichson, fixed by original designation and monotypy.

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Scheerpeltz, 1933: 1427 (Velleiopsis; catalog).

  • — Blackwelder, 1952: 160 (type species: curtus).

  • — Bordoni, 1973b: 42 (subgenus of Quedius; characters).

  • — Coiffait, 1978: 9, 13, 15 (subgenus of Quedius; characters; key to species of western Palaearctic region).

Tenebrobius Rambousek, 1915a: 27, 29 (subgenus of Quedius; species included: bernhaueri). Type species: Quedius (Tenebrobius) bernhaueri Rambousek, fixed by monotypy.

  • — Rambousek, 1916: 130 (description; subgenus of Quedius; cited as a new genus).

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Gridelli, 1924d: 18 (synonym of Microsaurus).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 375 (type species: bernhaueri).

  • — Coiffait, 1978: 9 (synonym of Quedius subgenus Microsaurus).

Quediellus Casey, 1915: 398, 402 (species included: nanulus, debilis, helenae, humilis, densiventris). Type species: Quedius debilis Horn, fixed by original designation.

  • — Casey, 1915: 398 (type species: Quedius debilis).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 334 (type species: debilis).

  • — Smetana, 1971: 39 (synonym of Quedius).

Quediochrus Casey, 1915: 398, 420 (species included: spelaeus, quadriceps). Type species: Quedius spelaeus Horn, fixed by original designation and monotypy.

  • — Casey, 1915: 398 (type species: Quedius spelaeus).

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 334 (type species: spelaeus).

  • — Hatch, 1957: 230 (subgenus of Quedius; characters; species of Pacific Northwest).

  • — Smetana, 1971: 38 (synonym of Quedius).

Paraquedius Casey, 1915: 397, 400 (species included: puncticeps). Type species: Quedius puncticeps Horn, fixed by original designation and monotypy.

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 293 (type species: puncticeps).

  • — Hatch, 1957: 214, 215 (subgenus of Quedius; characters; species of Pacific Northwest).

  • — Smetana, 1971: 40 (subgenus of Quedius).

  • — Korge, 1971a: 36 (subgenus of Quedius; characters).

Distichalius Casey, 1915: 398, 404 (species included: capucinus, nevadensis, marginalis, brunnipennis, agnatus, virginicus, oculeus, sparsus). Type species: Staphylinus capucinus Gravenhorst, fixed by original designation.

  • — Casey, 1915: 398 (type species: Quedius capucinus).

  • — Scheerpeltz, 1933: 1428 (cited as Distichalis; subgenus of Quedius).

  • — Blackwelder, 1952: 130 (type species: capucinus).

  • — Hatch, 1957: 215, 217 (subgenus of Quedius; characters; key to species of Pacific Northwest).

  • — Smetana, 1971: 39, 153 (subgenus of Quedius; characters; key to species of Canada and USA).

  • — Bordoni, 1973b: 48 (subgenus of Quedius; characters).

  • — Coiffait, 1978: 9, 13, 52 (subgenus of Quedius; characters; key to species of western Palaearctic region).

  • — Smetana, 1988: 227 (key to species of Himalayan region).

  • — Lohse and Lucht, 1989: 171 (subgenus of Quedius; notes).

  • — Smetana, 1995: 25, 57 (subgenus of Quedius; characters; key to species of Taiwan).

Megaquedius Casey, 1915: 421 (species included: laxatus, explanatus, manitobensis). Type species: Quedius explanatus LeConte, fixed by original designation.

  • — Casey, 1915: 399, 421 (type species: Quedius explanatus).

  • — Leng, 1920: 110 (catalog of North American species).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 236 (type species: explanatus).

  • — Hatch, 1957: 230 (subgenus of Quedius; characters; species of Pacific Northwest).

  • — Smetana, 1971: 38, 42 (subgenus of Quedius; key to species of USA and Canada).

Anastictodera Casey, 1915: 421 (species included: compransor). Type species: Quedius compransor Fall, fixed by monotypy.

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 50 (type species: compransor).

  • — Smetana, 1971: 38 (synonym of Quedius).

Cyrtoquedius Bernhauer, 1917h: 92 (subgenus of Quedius; species included: labiatus, frenatus, clypealis, basiventris, jocosus, graciliventris, concolor, arrogans, falvicaudus, rufinasus, mexicanus, verecundus, protensus, flavinasus, ochropygus, laeviventris). Type species: Quedius basiventris Sharp, fixed by subsequent designation by Blackwelder, 1952: 115.

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 115 (subgenus of Quedius).

  • — Scheerpeltz, 1955a: 172 (key to neotropical species).

Quediopsis Portevin, 1929: 337 (subgenus of Quedius; species included: riparius, kraatzi; preoccupied by Fauvel, 1878). Type species: Quedius riparius F. Kellner, fixed by subsequent designation by Blackwelder, 1952: 335.

  • — Scheerpeltz, 1933: 1429 (synonym of Quedius).

  • — Smetana, 1958: 328 (synonym of Quedius).

  • — Smetana, 1971: 40 (synonym of Quedius).

  • — Coiffait, 1978: 9 (synonym of Quedius subgenus Quediops).

  • — Lohse and Lucht, 1989: 171 (subgenus of Quedius; notes).

Arphirus Tottenham, 1945: 70 (proposed as replacement for Raphirus Stephens because he considered its type species to be of uncertain identity and the genus to be a “genus dubium”). Type species: Staphylinus semiobscurus Marsham, by original designation.

  • — Tottenham, 1948a: 243 (may be same as Sauridus; characters and key to species).

  • — Tottenham, 1949: 377, 422 (type species: semiobscurus Marsham = rufipes Erichson; subgenus of Quedius; checklist of British species).

  • — Blackwelder, 1952: 63 (type species: semiobscurus).

  • — Hatch, 1957: 228 (synonym of Raphirus).

  • — Smetana, 1958: 328 (synonym of Quedius).

  • — Smetana, 1971: 40 (synonym of Quedius).

Farus Blackwelder, 1952: 165 (replacement name for Ediquus Reitter; subgenus of Quedius). Type species: Quedius przewalskii Reitter, fixed by objective synonymy with Ediquus.

Pridonius Blackwelder, 1952: 321 (subgenus of Quedius; replacement name for Prionidus Bernhauer). Type species: Quedius iheringi Bernhauer, fixed by objective synonymy with Prionidus.

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

Quediops Coiffait, 1963: 373 (subgenus of Quedius; species included: plancus). Type species: Quedius plancus Erichson, fixed by original designation and monotypy.

  • — Bordoni, 1973g: 117 (subgenus of Quedius; characters; key to species).

  • — Coiffait, 1978: 9, 13, 63 (subgenus of Quedius; characters; key to species of western Palaearctic region).

  • — Smetana, 1993b: 50 (synonym of Quedius subgenus Raphirus).

Microquedius Coiffait, 1967e: 419 (subgenus of Quedius; species included: alpestris, spurius, subalpestris, haberfelneri cited as haberfellneri, marcuzzi, xanthippae, auricomus). Type species: Philonthus alpestris Heer, fixed by original designation.

  • — Bordoni, 1974a: 204 (subgenus of Quedius; notes; key to species of Italy).

  • — Coiffait, 1978: 9, 13, 60 (subgenus of Quedius; characters; key to species of western Palaearctic region).

  • — Lohse and Lucht, 1989: 171 (subgenus of Quedius; notes).

  • — Smetana, 1993b: 50 (synonym of Quedius subgenus Raphirus).

Quedionthus Bordoni, 1973b: 53 (subgenus of Quedius; species included: riparius [Note: Replacement name for Quediopsis Portevin but not so stated]). Type species: Quedius riparius F. Kellner, fixed by original designation and objective synonymy with Quediopsis.

  • — Coiffait, 1977c: 136 (synonym of Quediops).

  • — Coiffait, 1978: 9 (synonym of Quedius subgenus Quediops).

abdominalis Eppelsheim, 1878a: 419 (Quedius; Type locality: Caucasus).

  • — Bernhauer and Schubert, 1916: 417 (Quedius; catalog).

  • — Gridelli, 1924d: 25 (Quedius; subgenus Microsaurus; characters; Caucasus).

  • — Scheerpeltz, 1933: 1429 (Quedius; catalog).

  • — Coiffait, 1978: 158 (Quedius; subgenus Microsaurus; characters; Caucasus).

  • Distribution: Caucasus.

aberrans Cameron, 1932, see: Indoquedius.

abietoides Coiffait, 1967 [Note: Nomen nudum], see: abietum Kiesenwetter, 1858.

abietum Kiesenwetter, 1858: 57 (Quedius; Type locality: Parnes bei Athen; Aetolische Alpen).

  • — Fauvel, 1874: 282 [= 1874a: 508] (Quedius; characters; Illyria; Sicily; Greece; Sardinia; Morocco).

  • — Mulsant and Rey, 1876: 664 [= 1877: 520] (Quedius; characters; France).

  • — Fauvel, 1878: 127 [= 1878a: 47] (Quedius; Morocco).

  • — Fairmaire, 1879: 161 (Quedius; characters; Algeria).

  • — Fauvel, 1886: 57 [= 1886a: 49] (Quedius; Morocco; Tunisia; Algeria).

  • — Ganglbauer, 1895: 400 (Quedius; subgenus Microsaurus; characters; Illyria; Mediterranean region).

  • — Xambeu, 1900: 49 (Quedius; larval characters).

  • — Fauvel, 1902b: 114 (Quedius; Morocco; Algeria; Tunisia).

  • — Porta, 1907: 126 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Reitter, 1909: 110 (Quedius; subgenus Microsaurus; characters; not yet in Germany).

  • — Bernhauer and Schubert, 1916: 417 (Quedius; catalog).

  • — Gridelli, 1924d: 57 (Quedius; subgenus Microsaurus; characters; Sardinia; Sicily; Macedonia; Spain; Tunisia).

  • — Gridelli, 1929: 34 (Quedius; characters).

  • — Scheerpeltz, 1933: 1429 (Quedius; catalog).

  • — Roubal, 1941b: 125, 128 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Normand, 1947a: 9 (Quedius; Tunisia).

  • — Scheerpeltz, 1961d: 133 (Quedius; subgenus Microsaurus; Italy).

  • — Scheerpeltz, 1964a: 300 (Quedius; subgenus Microsaurus; Crete).

  • — Scheerpeltz, 1965b: 407 (Quedius; Sicily).

  • — Korge, 1971a: 47 (Quedius; subgenus Microsaurus; Turkey).

  • — Bordoni, 1974: 48 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Coiffait, 1978: 171 (Quedius; subgenus Microsaurus; characters; Mediterranean region; North Africa).

  • — Tóth, 1984: 113 (Quedius; characters).

  • — Sparacio, 1995: 164 (Quedius; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • Distribution: Tunisia, Algeria, Morocco, Spain, France, Italy, “Illyria”, Macedonia, Greece, Turkey.

reboudi Fairmaire, 1877 ([Note: Nomen nudum (Articles 11.6, 12.1); although the original description for reboudi is cited as Petites Nouvelles Entomologiques 2: 111, the species is neither on page 111 nor on any other page of the volume and there is evidently no description of the species. The first publication of the name appears to have been in 1879, when it was cited as a junior synonym; all subsequent uses have been as a junior synonym]).

  • — Fairmaire, 1879: 161 (Quedius; synonym of abietum).

  • — Fauvel, 1897d: 306 (Quedius; species was not described).

  • — Porta, 1907: 126 (Quedius; synonym of abietum).

  • — Bernhauer and Schubert, 1916: 417 (Quedius; synonym of abietum).

  • — Scheerpeltz, 1933: 1429 (Quedius; catalog; the species not described).

  • — Bordoni, 1974: 48 (Quedius; synonym of abietum).

  • — Coiffait, 1978: 171 (Quedius; cited as “nomen nudum ?”; synonym of abietum).

abietoides Coiffait, 1967e: 393 (Quedius; subgenus Microsaurus; variety of abietum; [Note: Nomen nudum (Article 15.2)]; cited from Itxassou, Basses-Pyrénées).

abkasicus Coiffait, 1963a: 390, 410 (Quedius; subgenus Sauridus; [Note: Cited on page 390 in a key to the species of Sauridus but listed in Microsaurus on page 410]; Type locality: U.R.S.S.: Abkasie, Ritsa).

  • — Coiffait, 1978: 220 (Quedius; subgenus Sauridus; characters; Abkhasia).

  • Distribution: Georgia.

abnormalis Sharp, 1889: 30 (Quedius; Type locality: Nikko).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1429 (Quedius; catalog).

  • — Nakane, 1963a: 94 (Quedius; characters; habitus photograph; Japan).

  • Distribution: Japan.

abruzzensis Drugmand, 1988c: 206 (Quedius; subgenus Raphirus; Type locality: Abruzzes: l'Aquila, versant nord du mont Luco, 900 m).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; cited as “specie dubbia”; Italy).

  • Distribution: Italy.

acatl Smetana, 1975: 322 (Quedius; subgenus Raphirus; Type locality: Mexico: Durango: 10 mi. W. El Salto).

  • Distribution: Mexico.

acco Smetana, 1996: 4 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Gongga Shan, 2750 m).

  • — Smetana, 1997d: 452 (Quedius; subgenus Microsaurus; collecting notes; China).

  • — Smetana, 1999b: 216 (Quedius; subgenus Microsaurus; China).

  • Distribution: China.

acuminatus Fairmaire, 1860, see: maurorufus Gravenhorst, 1806.

acuminatus Hochhuth, 1849: 151 (Quedius; Type locality: Kaukasus).

  • — Hochhuth, 1862: 47 (Quedius; Russia).

  • — Fauvel, 1874: 307 [= 1874a: 533] (Quedius; characters; France; Caucasus; Iran).

  • — Mulsant and Rey, 1876: 784 [= 1877: 784] (Quedius; subgenus Raphirus; characters; France).

  • — Porta, 1907: 147 (Quedius; subgenus Raphirus; characters; Italy).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Gridelli, 1925: 153 (Quedius; subgenus Raphirus; characters; Pyrenees; Caucasus; Persia; Germany).

  • — Portevin, 1929: 345 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1429 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 55 (Quedius; aedeagal illustration).

  • — Fagel, 1960b: 120, 123 (Quedius; characters).

  • — Smetana, 1962b: 149 (Quedius; subgenus Raphirus; characters; Europe Caucasus; Iran).

  • — Lohse, 1964: 218 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Korge, 1964: 123 (Quedius; subgenus Raphirus; Turkey).

  • — Horion, 1965: 298 (Quedius; subgenus Raphirus; Caucasus; Iran; Turkey; Austria; Istria; Bosnia).

  • — Coiffait, 1978: 265 (Quedius; subgenus Raphirus; characters; Pyrenees; Greece; Transcaucasus; Lebanon).

  • — Coiffait, 1978d: 165 (Quedius; Turkey).

  • — Tóth, 1984: 125 (Quedius; characters; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Austria).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy ?).

  • Distribution: Europe, Russia, Armenia, Turkey, Lebanon, Iran.

phenicius Coiffait, 1967e: 423 (Quedius; subgenus Raphirus; subspecies of acuminatus; Type locality: Aïn Batara, Liban).

  • — Coiffait, 1978: 265 (Quedius; subgenus Raphirus; subspecies of acuminatus; characters).

khnzoriani Coiffait, 1967e: 423 (Quedius; subgenus Raphirus; subspecies of acuminatus; Type locality: Aragatch, Karagel, 3200 m, Arménie russe).

  • — Coiffait, 1978: 265 (Quedius; subgenus Raphirus; subspecies of acuminatus; characters).

adanensis Coiffait, 1967e: 394 (Quedius; subgenus Microsaurus; replacement name for gridellii Jarrige).

  • — Coiffait, 1978: 172 (Quedius; subgenus Microsaurus; characters; Turkey).

  • Distribution: Turkey.

gridellii Jarrige, 1952a: 134 (Quedius; subgenus Microsaurus; [preoccupied]; Type locality: Adana, Cilicie).

  • — Coiffait, 1967e: 394 (Quedius; synonym of adanensis).

  • — Coiffait, 1978: 172 (Quedius; synonym of adanensis).

adjacens Cameron, 1926b: 368 (Quedius; subgenus Microsaurus; Type locality: Chakrata district, Deoban, 9300 feet; [Note: See lectotype designation by Smetana, 1988]).

  • — Cameron, 1932: 287 (Quedius; subgenus Microsaurus; characters; India).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Smetana, 1988: 205 (Quedius; subgenus Microsaurus; lectotype designation; characters; Kashmir; Uttar Pradesh; Himachal Pradesh).

  • — Smetana, 1996: 1 (Quedius; subgenus Microsaurus; characters; China).

  • — Smetana, 1999b: 214 (Quedius; subgenus Microsaurus; China).

  • Distribution: India, China.

adnexus Smetana, 1965: 44, 45 (Quedius; subgenus Distichalius; Type locality: Newfoundland: Griguet).

  • — Smetana, 1971: 196 (Quedius; subgenus Raphirus; characters; Newfoundland).

  • — Downie and Arnett, 1996: 393 (Quedius; subgenus Raphirus; characters; Canada).

  • Distribution: Canada.

adustus Sharp, 1889: 32 (Quedius; Type locality: Oyama; Nishi; Nara; Chiuzenji; Nishimura; Numata).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Nakane, 1963a: 94 (Quedius; characters; habitus photograph; Japan).

  • — J. Li, 1993: 41 (Quedius; China).

  • — Watanabe, 1996a: 13 (Quedius; cave collection; Japan).

  • — Watanabe, 1998: 314 (Quedius; subgenus Microsaurus; Japan).

  • Distribution: Japan, China.

advena Sharp, 1884: 329 (Quedius; Type locality: Mexico, Las Vigas).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Campbell 1975a: 319 (Quedius; subgenus Raphirus; characters; Mexico).

  • Distribution: Mexico.

aeneipennis Heer, 1834 [Note: Nomen nudum], see: alpestris Heer, 1839.

aeneipennis Fairmaire and Germain, 1861, see: Loncovilius.

aeneiventris Broun, 1910: 14 (Quedius; Type locality: Waimarino).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • Distribution: New Zealand.

aenescens Mäklin, 1852: 315 (Quedius; Type locality: insula Sitkha; [Note: See lectotype designation by Smetana, 1965]).

  • — Mäklin, 1853: 190 (Quedius; characters; Edgecombe Island; Kadjak Island; Sitka Island).

  • — Horn, 1878: 164 (Quedius; synonym of brunnipennis).

  • — Fall and Cockerell, 1907: 166 (Quedius; New Mexico).

  • — Casey, 1915: 406 (Distichalius; cited as brunneipennis; [Note: Misidentified as brunnipennis according to Smetana, 1971: 193]; characters; Alaska; British Columbia).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; cited as aenesceus; catalog).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Smetana, 1965: 45, 58 (Quedius; subgenus Distichalius; lectotype designation; characters).

  • — Smetana, 1971: 193 (Quedius; subgenus Raphirus; characters; Alberta; British Columbia; Alaska; Oregon; Washington).

  • Distribution: Canada, USA.

agnatus Casey, 1915: 406 (Distichalius; Type locality: Manitoba: Aweme).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Smetana, 1964: 38 (Quedius; subgenus Distichalius; Alaska).

  • — Smetana, 1965: 43, 45 (Quedius; subgenus Distichalius; characters; Labrador).

  • — Smetana, 1965a: 13 (Quedius; subgenus Distichalius; Alaska).

  • — Smetana, 1971: 193 (Quedius; synonym of aenescens).

aeneus Fauvel, 1877: 274 (Quedius; Type locality: Australie, Victoria).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • Distribution: Australia.

aereipennis Bernhauer, 1929d: 111 (Quedius; Type locality: Da-laen-saen bei Nong-po; [Note: See lectotype designation by Smetana, 1996a]).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Smetana, 1996a: 49 (Quedius; subgenus Raphirus; lectotype designation; characters; China).

  • Distribution: China.

aetolicus Kraatz, 1858a: 58 (Quedius; Type locality: ätolische Alpen).

  • — Fauvel, 1874: 282 [= 1874a: 508] (Quedius; synonym of cruentus).

  • — Ganglbauer, 1895: 399 (Quedius; synonym of cruentus).

  • — Porta, 1907: 115 (Quedius; synonym of cruentus).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; synonym of cruentus).

  • — Gridelli, 1924d: 54 (Quedius; subgenus Microsaurus; characters; England; Corsica; Algeria; Sardinia; Italy).

  • — Donisthorpe, 1930: 31 (Quedius; characters; Britain).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Roubal, 1941b: 123, 128 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Bordoni, 1974: 45 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 154 (Quedius; subgenus Microsaurus; characters; Europe; Corsica; Sardinia; North Africa; England; France; Portugal).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • — Owen, 1999b: 61 (Quedius; captured underground at roots of old trees; Britain).

  • — Fowles, Alexander, and Key, 1999: 129 (Quedius; saproxylic species; status as endangered or threatened species; Britain).

  • — Owen, 2000: 251 (Quedius; pitfall trap collections from underground at roots of trees; Britain).

  • Distribution: Europe, Algeria.

subapicalis Joy, 1913a: 76 (Quedius; Type locality: Borckenhurst, New Forest).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • — Gridelli, 1924d: 55 (Quedius; synonym of aetolicus).

  • — Scheerpeltz, 1933: 1430 (Quedius; synonym of aetolicus).

  • — Bordoni, 1974: 45 (Quedius; synonym of aetolicus).

  • — Pope, 1977: 31 (Quedius; synonym of aetolicus).

  • — Coiffait, 1978: 154 (Quedius; synonym of aetolicus).

afghanicus Coiffait, 1977c: 139 (Quedius; subgenus Sauridus; Type locality: Afghanistan: Paktui, Khat Gaï, 2600 m).

  • — Coiffait, 1978: 232 (Quedius; subgenus Sauridus; characters; Afghanistan).

  • Distribution: Afghanistan.

africanus Peyerimhoff, 1917, see: cinctus Paykull, 1790.

afrofuliginosus Gusarov, 1991a: 9 (Quedius; Type locality: Algeria).

  • Distribution: Algeria.

agathis Broun, 1893: 1407 (Quedius; Type locality: Hunua Range, near Clevedon and at Maketu).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • Distribution: New Zealand.

agnatus Casey, 1915, see: aenescens Mäklin, 1852.

akalita Smetana, 1977, see: Indoquedius.

alatus Coiffait, 1967, see: asturicus Bernhauer, 1918.

albanicus Bernhauer, 1914d: 3 (Quedius; Type locality: Albanien).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Bernhauer, 1923d: 177 (Quedius; subgenus Raphirus; characters; Albania).

  • — Gridelli, 1925: 166 (Quedius; subgenus Raphirus; characters; Albania).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Coiffait, 1978: 283 (Quedius; subgenus Raphirus; characters; Albania).

  • Distribution: Albania.

albanicus Bernhauer, 1926, see: illyricus Wendeler, 1928.

alentejensis Coiffait, 1963a: 391, 412 (Quedius; subgenus Sauridus; [Note: Cited in the subgenus Microsaurus on page 412 but that assignment is probably erroneous]; Type locality: Portugal: Castro Verde).

  • — Coiffait, 1978: 212 (Quedius; subgenus Sauridus; characters; Portugal).

  • Distribution: Portugal.

aliiceps Cameron, 1948c: 726 (Quedius; subgenus Raphirus; Type locality: New Zealand: Auckland Island: Carnley Harbour, Musgrave Pen).

  • — Gourlay, 1950: 187 (Quedius; notes).

  • Distribution: New Zealand.

almeriensis Coiffait, 1964 [Note: Nomen nudum], see: cobosi Coiffait, 1964.

alpestris Heer, 1839: 280 (Philonthus; Type locality: Mühlebachalp; [Note: See lectotype designation by Assing, 1999b]. Also cited from Flössalp; Val Emet; Beverserthal; Alpen um Genf).

  • — Kiesenwetter, 1848: 63 (Quedius; synonym of satyrus).

  • — Kraatz, 1857d: 517 (Quedius; characters; Alps).

  • — Redtenbacher, 1857: 202 (Quedius; characters; Austria).

  • — Fauvel, 1874: 303 [= 1874a: 529] (Quedius; characters; France; Germany; Switzerland; Austria; Italy).

  • — Redtenbacher, 1874: 198 (Quedius; characters; Austria).

  • — Mulsant and Rey, 1876: 763 [= 1877: 619] (Quedius; subgenus Raphirus; characters; France).

  • — Ganglbauer, 1895: 410 (Quedius; subgenus Raphirus; characters; Alpes; Sudeten; Carpathians).

  • — Porta, 1907: 145 (Quedius; subgenus Raphirus; characters; Italy).

  • — Reitter, 1909: 114 (Quedius; subgenus Raphirus; characters; Germany).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Gridelli, 1925: 142 (Quedius; subgenus Raphirus; characters).

  • — Portevin, 1929: 344 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 46 (Quedius; aedeagal illustration).

  • — Smetana, 1958: 381 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Fagel, 1960b: 120 (Quedius; notes).

  • — Smetana, 1962a: 137 (Quedius; larval characters).

  • — Smetana, 1962b: 140 (Quedius; subgenus Raphirus; characters; Europe).

  • — Coiffait, 1963a: 382 (Quedius; subgenus Sauridus; characters).

  • — Lohse, 1964: 213 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 289 (Quedius; subgenus Raphirus; mountains of central Europe; France; Italy; Slovenia; Switzerland; Germany; Austria; Czechoslovakia; Poland).

  • — Pototskaia, 1967: 83 (Quedius; larval characters).

  • — Coiffait, 1967e: 419 (Quedius; subgenus Microquedius; type species of Microquedius).

  • — Scheerpeltz, 1968a: 92 (Quedius; catalog; Austria).

  • — Tikhomirova, 1973a: 190 (Quedius; checklist; Soviet Union).

  • — Bordoni, 1974a: 205, 206 (Quedius; subgenus Microquedius; characters; Italy).

  • — Coiffait, 1978: 198 (Quedius; subgenus Microquedius; characters; Alpes; Carpathians; Sudeten).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 137 (Quedius; catalog; Poland).

  • — Tóth, 1984: 121 (Quedius; characters).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • — Assing, 1999b: 43 (Quedius; lectotype designation: Mühlebachalp; characters; notes).

  • — Štourač, 1998b: 135 (Quedius; subgenus Raphirus; notes; Czech Republic).

  • Distribution: Europe.

aeneipennis Heer, 1834: 75 (Staphylinus; subgenus Philonthus; [Note: Nomen nudum, published without characters]).

  • — Heer, 1839: 280 (Philonthus; synonym of alpestris).

  • — Kraatz, 1857d: 517 (Quedius; synonym of alpestris).

  • — Ganglbauer, 1895: 410 (Quedius; synonym of alpestris).

  • — Porta, 1907: 145 (Quedius; synonym of alpestris).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; synonym of alpestris).

  • — Gridelli, 1925: 142 (Quedius; synonym of alpestris).

  • — Smetana, 1958: 381 (Quedius; synonym of alpestris).

  • — Coiffait, 1978: 198 (Quedius; synonym of alpestris).

satyrus Kiesenwetter, 1847: 75 (Quedius; Type locality: Riesengebirge).

  • — Kiesenwetter, 1848: 63 (Quedius; characters; Riesengebirge).

  • — Märkel and Kiesenwetter, 1848a: 325 (Quedius; notes; Carpathian Alps).

  • — Redtenbacher, 1849: 825 (Philonthus; characters; Austria).

  • — Kraatz, 1857d: 517 (Quedius; synonym of alpestris).

  • — Redtenbacher, 1857: 203 (Quedius; characters; Austria).

  • — Gemminger and Harold, 1868: 568 (Quedius; synonym of alpestris).

  • — Redtenbacher, 1874: 200 (Quedius; characters; Austria).

  • — Ganglbauer, 1895: 410 (Quedius; synonym of alpestris).

  • — Porta, 1907: 145 (Quedius; synonym of alpestris).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; synonym of alpestris).

  • — Gridelli, 1925: 142 (Quedius; synonym of alpestris).

  • — Scheerpeltz, 1933: 1430 (Quedius; doubtful synonym of alpestris).

  • — Smetana, 1958: 381 (Quedius; synonym of alpestris).

  • — Coiffait, 1978: 198 (Quedius; synonym of alpestris).

biseriatus Stierlin, 1875: 473 (Philonthus; Type locality: St. Bernhard).

  • — Ganglbauer, 1895: 410 (Quedius; synonym of alpestris).

  • — Porta, 1907: 145 (Quedius; synonym of alpestris).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; synonym of alpestris).

  • — Gridelli, 1925: 142 (Quedius; synonym of alpestris).

  • — Scheerpeltz, 1933: 1430 (Quedius; doubtful synonym of alpestris).

  • — Smetana, 1958: 381 (Quedius; synonym of alpestris).

  • — Coiffait, 1978: 198 (Quedius; synonym of alpestris).

spurius Lokay, 1921: 71 (Quedius; variety of alpestris Heer; Type locality: Alpes Styriae, Carinthiae, Carnioliae, Montes Bohemiae, Silesiae).

  • — Gridelli, 1925: 142 (Quedius; subgenus Raphirus; variety of alpestris; characters).

  • — Scheerpeltz, 1933: 1430 (Quedius; variety of alpestris).

  • — Scheerpeltz, 1953a: 133 (Quedius; [Note: Misidentification: Smetana, 1958: 382, cited alpestris Scheerpeltz as a synonym of spurius; Scheerpeltz attributed alpestris to Heer]; characters).

  • — Smetana, 1958: 382 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 140 (Quedius; subgenus Raphirus; characters).

  • — Coiffait, 1963a: 382 (Quedius; subgenus Sauridus; characters).

  • — Lohse, 1964: 213 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Coiffait, 1967e: 419 (Quedius; subgenus Microquedius).

  • — Coiffait, 1978: 198 (Quedius; subgenus Microquedius; characters; Europe).

  • — Tóth, 1984: 122 (Quedius; characters).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria).

  • — Nowosad, 1990: 147 (Quedius; collected from mole nest; notes; Poland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Assing, 1999b: 43 (Quedius; synonym of alpestris).

marcuzzii Scheerpeltz, 1953a: 130 (Quedius; Type locality: Massiccio della Marmolada, 2200 m).

  • — Smetana, 1958: 381 (Quedius; synonym of alpestris).

  • — Smetana, 1962b: 140 (Quedius; synonym of alpestris).

  • — Coiffait, 1963a: 382 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1967e: 419 (Quedius; subgenus Microquedius).

  • — Coiffait, 1978: 198 (Quedius; subgenus Microquedius; characters; Dolomites).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; cited as “specie dubbia”; Italy).

  • — Assing, 1999b: 43 (Quedius; synonym of alpestris).

alpinus Fiori, 1894, see: plagiatus Mannerheim, 1843.

alpinus Helliesen, 1892: 74 (Quedius; Type locality: Mjosvand in Telemarken).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Gridelli, 1925: 168 (Quedius; characters).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • Distribution: Norway.

altaicus Coiffait, 1967, see: centrasiaticus Coiffait, 1969.

altaicus Korge, 1962: 152 (Quedius; subgenus Quedius; Type locality: Zentral-Altai).

  • — Coiffait, 1978: 194 (Quedius; subgenus Quedius; characters; central Asia).

  • Distribution: “Altai region”.

alticola Smetana, 1971: 172 (Quedius; subgenus Distichalius; Type locality: Washington: Mt. Rainier Nat. Pk., Nisqually River, 4000 ft. Also cited from California).

  • — Smetana, 1976c: 175 (Quedius; subgenus Distichalius; Oregon; Washington).

  • — Smetana, 1981b: 640 (Quedius; collecting notes; Oregon).

  • Distribution: USA.

alutaceus Hatch, 1957, see: densiventris Casey, 1915.

amabilis Smetana, 1971: 205 (Quedius; subgenus Raphirus; Type locality: California: nr. Strawberry, El Dorado Co.).

  • — Smetana, 1990d: 100 (Quedius; California).

  • Distribution: USA.

ambiguus Broun, 1894: 423 (Quedius; Type locality: Tarukenga, about 8 miles from Rotorua).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • Distribution: New Zealand.

amethystinus Roubal, 1948 [Note: Nomen nudum], see: scintillans Gravenhorst, 1806.

amicorum Smetana, 1997d: 470 (Quedius; subgenus Microsaurus; Type locality: China: N. Yunnan, Xue Shan, nr. Zhongdian, 4050 m, 27°49 N, 99°34 E).

  • Distribution: China.

amplicollis Scriba 1868, see: crassus Fairmaire, 1860.

amplissimus Bernhauer, 1912k: 262 (Quedius; Type locality: Umgebung von Sebastopol, Krim).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Gridelli, 1924d: 46 (Quedius; subgenus Microsaurus; characters; Crimea).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Jarrige, 1971: 497 (Quedius; Iran).

  • Distribution: Ukraine, Iran.

analis Fabricius, 1787, see: scitus Gravenhorst, 1806.

analis Stephens, 1835, see: cruentus Olivier, 1795.

analis W. J. MacLeay, 1873, see: manducus Herman, 2001.

analoka Smetana, 1975, see: stevensi Cameron, 1932.

anatolicus Korge, 1964, see: suturalis Kiesenwetter, 1845.

anceps Fairmaire and Laboulbène, 1856: 543 (Quedius; Type locality: H. -Pyr., Aragnouet).

  • — Fauvel, 1874: 294 [= 1874a: 520] (Quedius; synonym of umbrinus).

  • — Mulsant and Rey, 1876: 724 [= 1877: 580] (Quedius; subgenus Sauridus; characters; France).

  • — Ganglbauer, 1895: 405 (Quedius; subgenus Sauridus; characters; Pyrenees; France).

  • — Fauvel, 1900d: 233 (Quedius; notes).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Gridelli, 1924d: 111 (Quedius; subgenus Raphirus; characters; Pyrenees).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Jeannel and Jarrige, 1949: 379 (Quedius; subgenus Raphirus; cave species; France).

  • — Coiffait, 1963a: 385 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 230 (Quedius; subgenus Sauridus; characters; Pyrenees).

  • — Tronquet, 1998a: 114 (Quedius; collected from marmot burrow; France).

  • Distribution: Portugal, Spain, France, Italy, Switzerland.

robustus Scriba, 1870: 81 (Quedius; Type locality: Prope Santas Albas, prov. Leon).

  • — Fauvel, 1874: 292 [= 1874a: 518] (Quedius; characters; France; Switzerland; Italy; Spain; Portugal).

  • — Mulsant and Rey, 1876: 710 [= 1877: 566] (Quedius; subgenus Sauridus; characters; France).

  • — Ganglbauer, 1895: 405 (Quedius; subgenus Sauridus; characters; western Alps; Spain; Portugal).

  • — Porta, 1907: 136 (Quedius; subgenus Sauridus; characters; Italy).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; catalog).

  • — Gridelli, 1924d: 111 (Quedius; synonym of anceps).

  • — Portevin, 1929: 342 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1431 (Quedius; synonym of anceps).

parviceps Fauvel, 1871b: 359 (Quedius; [Note: Nomen nudum (Articles 11.6, 12.1); species not described but has been listed as a synonym by other authors]; cited from Monte Rosa, Furca, Faulhorn, Alpen von 4000 bis 8000′).

  • — Fauvel, 1874: 292 [= 1874a: 518] (Quedius; synonym of robustus).

  • — Ganglbauer, 1895: 405 (Quedius; synonym of robustus).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; synonym of robustus).

  • — Porta, 1907: 136 (Quedius; synonym of robustus).

  • — Scheerpeltz, 1933: 1430 (Quedius; synonym of anceps).

  • — Coiffait, 1978: 230 (Quedius; synonym of anceps).

andersoni Blackburn, 1888: 6 (Quedius; Type locality: Port Lincoln district).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

andreinii Gridelli, 1924d: 19, 30 (Quedius; subgenus Microsaurus; Type locality: Santo Stefano d'Aveto, Apennino Ligure).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Coiffait, 1978: 173 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Sparacio, 1995: 165 (Quedius; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • Distribution: Italy.

angaricus Coiffait, 1975: 34 (Quedius; subgenus Sauridus; Type locality: Kistvianka, région sud-ouest du Lac Baïkal).

  • — Coiffait, 1978: 259 (Quedius; subgenus Sauridus; characters; central Asia).

  • Distribution: Russia.

angnimai Smetana, 1988: 223 (Quedius; subgenus Microsaurus; Type locality: Nepal: Prov. Bagmati, below Thare Pati, 3300 m).

  • — Smetana, 1992b: 4 (Quedius; subgenus Microsaurus; Nepal).

  • Distribution: Nepal.

angularis Cameron, 1948: 41 (Quedius; subgenus Sauridus; Type locality: West of Fort Archambault, Sara-Dai).

  • Distribution: Chad.

angulicollis Fauvel, 1891a: 60 (Quedius; Type locality: Drome; Alpes-Maritimes, Venanson; Apennins).

  • — Ganglbauer, 1895: 408 (Quedius; subgenus Sauridus; characters; Maritime Alps; Apennines).

  • — Porta, 1907: 141 (Quedius; subgenus Sauridus; characters; Italy).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Gridelli, 1922b: 128, 131 (Quedius; subgenus Sauridus; characters; notes; Italy).

  • — Scheerpeltz, 1933: 1433 (Quedius; synonym of bicolor).

  • — Coiffait, 1978: 241 (Quedius; synonym of bicolor).

  • — Herman, 2001: 49 (Quedius; valid name for species).

bicolor Mulsant and Rey, 1876: 738 [= 1877: 594] (Quedius; subgenus Sauridus; [preoccupied]; Type locality: Grande Chartreuse).

  • — Ganglbauer, 1895: 407 (Quedius; synonym of humeralis).

  • — Porta, 1907: 140 (Quedius; synonym of humeralis).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; synonym of humeralis).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — Coiffait, 1963a: 393 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1963b: 425 (Quedius; subgenus Sauridus).

  • — Coiffait, 1978: 241 (Quedius; subgenus Sauridus; characters; Alps; Apennines; Massif Central).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • — Herman, 2001: 49 (Quedius; synonym of angulicollis).

  • Distribution: France, Italy.

angustifrons Wollaston, 1864: 563 (Quedius; Type locality: Gran Canaria; [Note: See lectotype designation by Israelson, 1985]. Originally cited from Grand Canary: Teror; Gomera: Hermigua).

  • — Wollaston, 1865: 486 (Quedius; habitat; Canary Islands).

  • — Fauvel, 1902b: 113 (Quedius; Canaries).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Gridelli, 1924d: 38 (Quedius; subgenus Microsaurus; characters; Canaries).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Israelson, 1985: 152 (Quedius; subgenus Microsaurus; lectotype designation: Gran Canaria; characters; Canary Islands).

  • Distribution: Canary Islands.

angustus Sharp, 1884: 338 (Quedionuchus; Type locality: Guatemala, Totonicapam, 8500 to 10,500 feet; [Note: See lectotype designation by Smetana, 1976a]).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Smetana, 1976a: 232 (Quedius; subgenus Quedionuchus; lectotype designation; characters; Guatemala).

  • Distribution: Guatemala.

annectens Sharp, 1889: 32 (Quedius; Type locality: Miyanoshita; [Note: See lectotype designation by Smetana, 1998a]).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Smetana, 1998a: 119 (Quedius; subgenus Distichalius; lectotype designation; characters; Japan).

  • Distribution: Japan.

anomalus Cameron, 1926b: 370 (Quedius; Type locality: Mussorie district: Dhobi Ghat; [Note: See lectotype designation by Smetana, 1988]).

  • — Cameron, 1932: 295 (Quedius; subgenus Raphirus; characters; India).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Smetana, 1988: 274 (Quedius; subgenus Raphirus; lectotype designation; characters; India).

  • Distribution: India.

antarcticus Bernhauer and Schubert, 1916: 418 (Quedius; replacement name for scutellaris Broun).

  • Distribution: New Zealand.

scutellaris Broun, 1894: 422 (Quedius; [preoccupied]; Type locality: Ashburton).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; synonym of antarcticus).

antennalis Cameron, 1932: 285 (Quedius; subgenus Microsaurus; Type locality: Darjeeling: Gopaldhara, alt. 4720 feet).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Smetana, 1988: 201 (Quedius; subgenus Microsaurus; characters; India).

  • Distribution: India.

anthracinus Solsky, 1868a: 125 (Quedius; Type locality: Mexique).

  • — Sharp, 1884: 332 (Quedius; notes; Mexico).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • Distribution: Mexico.

antipodum Sharp, 1886a: 378 (Quedius; Type locality: New Zealand: Dunedin; Greymouth).

  • — Broun, 1893: 1028 (Quedius; characters; New Zealand).

  • — Broun, 1911: 93 (Quedius; Chatham Islands).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Emberson, 1998: 36 (Quedius; collecting notes; Chatham Islands).

  • Distribution: New Zealand, Chatham Islands.

antoni Smetana, 1995f: 233 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Gongga Shan, 3300–3350 m).

  • — Smetana, 1997d: 452 (Quedius; subgenus Microsaurus; characters; collecting notes; China).

  • Distribution: China.

apicalis Cameron, 1941, see: philippinus Cameron, 1941.

apicicornis Eppelsheim, 1895b: 391 (Quedius; subgenus Microsaurus; Type locality: Sikkim im Himalaya).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Cameron, 1932: 286 (Quedius; subgenus Microsaurus; characters; Sikkim: Darjeeling district, Tiger Hill, alt. 8000 feet).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Smetana 1975a: 325 (Quedius; subgenus Microsaurus; characters; Nepal; Sikkim; Bhutan).

  • — Smetana, 1977a: 246 (Quedius; Bhutan).

  • — Coiffait, 1982: 32 (Quedius; characters; notes; Nepal).

  • — Smetana, 1988: 192 (Quedius; subgenus Microsaurus; characters; Bhutan; West Bengal; Sikkim; Nepal).

  • — Smetana, 1992b: 2 (Quedius; subgenus Microsaurus; Nepal; India).

  • Distribution: India, Nepal, Bhutan.

flavoterminatus Cameron, 1932: 290 (Quedius; subgenus Microsaurus; Type locality: Darjeeling Dist.: Rangirum, alt. 6000 feet).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Smetana, 1988: 192 (Quedius; synonym of apicicornis).

cyanopterus Scheerpeltz, 1976a: 150 (Quedius; subgenus Microsaurus; Type locality: Nepal: Umgebung von Kathmandu, 1400 m).

  • — Coiffait, 1982: 32 (Quedius; synonym of apicicornis).

  • — Smetana, 1988: 192 (Quedius; synonym of apicicornis).

sherpai Coiffait, 1982: 77 (Quedius; subgenus Microsaurus; Type locality: Nepal: Trisuli Tal, Gosainkund, 3200 m).

  • — Smetana, 1988: 192 (Quedius; synonym of apicicornis).

apiciflavus Lea, 1925: 248 (Quedius; Type locality: Queensland: Mount Tambourine).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

apicipennis Hellén, 1934 [Note: Nomen nudum], see: tenellus Gravenhorst, 1806.

aplodontiae Smetana, 1971, see: spelaeus Horn, 1871.

arabs Lokay, 1921 [Note: Nomen nudum], see: cinctus Paykull, 1790.

arcticus Munster, 1921, see: sublimbatus Mäklin, 1853.

arctifrons Broun, 1880: 112 (Philonthus; Type locality: Parua).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • Distribution: New Zealand.

arestor Tottenham, 1948, see: boops Gravenhorst, 1802.

aridulus Jansson, 1939, see: persimilis Mulsant and Rey, 1876.

armipes Sharp, 1889: 34 (Quedionuchus; Type locality: Shimidzu; Yuyama; Nikko).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Smetana, 1995a: 80 (Quedius; subgenus Quedionuchus; characters; Russia).

  • Distribution: Japan, Russia.

arrogans Sharp, 1884: 333 (Quedius; Type locality: Panama, Volcan de Chiriqui, at elevations between 2000 and 6000 feet).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 173, 174 (Quedius; subgenus Cyrtoquedius; characters in key; Panama).

  • Distribution: Panama.

arrowi Bernhauer, 1941b: 32 (Quedius; subgenus Sauridus; Type locality: Neu Seeland: Pudding Hill).

  • Distribution: New Zealand.

arvernicus Mulsant and Rey, 1876, see: mesomelinus Marsham, 1802.

arvernicus Coiffait, 1982, see: obscuripennis Bernhauer, 1901.

arvernus Herman, 2001, see: obscuripennis Bernhauer, 1901.

asiaticus Bernhauer, 1918a: 92 (Quedius; subgenus Ediquus; Type locality: Ost-Buchara, Turkestan).

  • — Gridelli, 1924d: 57 (Quedius; subgenus Microsaurus; characters; Turkestan).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Coiffait, 1978: 183 (Quedius; subgenus Microsaurus; characters; Turkestan).

  • Distribution: Uzbekistan.

aspromontanus Bernhauer, 1908b: 34 (Quedius; Type locality: Kalabrien, Aspromonte).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Gridelli, 1922b: 130, 133 (Quedius; subgenus Sauridus; characters; notes; Italy).

  • — Gridelli, 1925: 120 (Quedius; subgenus Raphirus; characters; Italy).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 34 (Quedius; aedeagal illustration).

  • — Coiffait, 1963a: 387 (Quedius; subgenus Sauridus; characters).

  • — Fagel, 1968c: 7 (Quedius; subgenus Sauridus; characters; notes; Italy; Corsica; Greece).

  • — Coiffait, 1978: 215 (Quedius; subgenus Sauridus; characters; Italy; Corsica; Greece).

  • — Zanetti, 1992b: 130, 131 (Quedius; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • Distribution: Italy, Greece.

assamensis Cameron, 1932: 293 (Quedius; subgenus Raphirus; Type locality: Assam: Naga Hills, alt. 4000 feet; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Smetana, 1988: 284 (Quedius; subgenus Raphirus; lectotype designation; characters; Nepal; India).

  • — Smetana, 1996c: 225 (Quedius; subgenus Raphirus; China).

  • Distribution: India, Nepal, China.

assecla Mulsant and Rey, 1876, see: ochripennis Ménétriés, 1832.

assimilis Nordmann, 1837, see: fulgidus Fabricius, 1793.

asturicus Bernhauer, 1918a: 96 (Quedius; subgenus Raphirus; Type locality: Asturien: Caboalles, Cancas).

  • — Gridelli, 1925: 165 (Quedius; subgenus Raphirus; characters).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Fagel, 1960b: 118 (Quedius; characters; Carinthia; Bosnia; Italy).

  • — Smetana, 1962b: 152 (Quedius; subgenus Raphirus; characters; Europe).

  • — Coiffait, 1978: 281 (Quedius; subgenus Raphirus; characters; Pyrenees; Bulgaria).

  • — Drugmand, 1988c: 209 (Quedius; characters).

  • — Tronquet, 1999a: 214 (Quedius; subgenus Raphirus; notes; France).

  • Distribution: Europe.

brevipennis Fairmaire, 1859: 41 (Quedius; [preoccupied]; Type locality: Pyrénées).

  • — Fauvel, 1874: 309 [= 1874a: 535] (Quedius; race of boops; characters; France; Italy; Portugal).

  • — Mulsant and Rey, 1876: 797 [= 1877: 653] (Quedius; subgenus Raphirus; characters; France).

  • — Sharp, 1889: 33 (Quedius; notes; Japan).

  • — Ganglbauer, 1895: 412 (Quedius; variety of boops).

  • — Porta, 1907: 150 (Quedius; variety of boops).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; variety of boops).

  • — Gridelli, 1925: 165 (Quedius; variety of boops).

  • — Scheerpeltz, 1933: 1434 (Quedius; variety of boops).

  • — Fagel, 1960a: 118 (Quedius; synonym of asturicus).

  • — Smetana, 1962b: 153 (Quedius; synonym of asturicus).

  • — Coiffait, 1978: 281 (Quedius; synonym of asturicus).

alatus Coiffait, 1967e: 424 (Quedius; subgenus Raphirus; subspecies of asturicus; Type locality: Vratza, Bulgarie).

  • — Coiffait, 1970e: 63 (Quedius; subspecies of asturicus; Cyprus).

  • — Coiffait, 1978: 281 (Quedius; subgenus Raphirus; subspecies of asturicus; characters).

  • — Tóth, 1984: 129 (Quedius; subspecies of asturicus; characters).

atchala Smetana, 1975a: 341 (Quedius; subgenus Raphirus; Type locality: Central Nepal: Gosaikunde Lakes area 3500–4500 m).

  • — Smetana, 1988: 270 (Quedius; subgenus Raphirus; characters; Nepal).

  • Distribution: Nepal.

ater J. Sahlberg, 1880, see: plagiatus Mannerheim, 1843.

ater Ziegler, 1845, see: capucinus Gravenhorst, 1806.

atl Smetana, 1975: 315 (Quedius; subgenus Quedionuchus; Type locality: Mexico: Oaxaca: 37 mi. S. Valle Nacional, 8500 ft.).

  • — Smetana, 1976a: 244 (Quedius; subgenus Quedionuchus; characters; Mexico).

  • Distribution: Mexico.

atricapillus Reitter, 1900: 273 (Quedius; Type locality: Ober-Syrien: Akbes).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Gridelli, 1924d: 56 (Quedius; subgenus Microsaurus; characters; Syria).

  • — Scheerpeltz, 1933: 1432 (Quedius; catalog).

  • — Jarrige, 1952a: 134 (Quedius; subgenus Microsaurus; characters).

  • — Coiffait, 1978: 170 (Quedius; subgenus Microsaurus; characters; Turkey).

  • Distribution: Syria.

atriceps Stephens, 1832, see: scitus Gravenhorst, 1806.

atricillus Gravenhorst, 1806, see: scitus Gravenhorst, 1806.

atripennis Stephens, 1835, see: mesomelinus Marsham, 1802.

atritarsis Roubal, 1949 [Note: Nomen nudum], see: unicolor Kiesenwetter, 1847.

attenuatus Gravenhorst, 1802, see: limbatus Heer, 1839.

attenuatus Gyllenhal, 1810 [Note: Nomen nudum], see: nitipennis Stephens, 1833.

atticus Coiffait, 1967, see: coxalis Kraatz, 1858.

aucklandicus Cameron, 1947d: 707 (Quedius; subgenus Raphirus; Type locality: New Zealand: N. Auckland: Tangihua Range, 1800–2000 feet).

  • Distribution: New Zealand.

aureipilis Cameron, 1932: 294 (Quedius; subgenus Raphirus; Type locality: Assam: Naga Hills, Laimatak; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1432 (Quedius; catalog).

  • — Smetana, 1988: 288 (Quedius; subgenus Raphirus; lectotype designation; characters; Nagaland).

  • Distribution: India.

aureiventris Bernhauer, 1915b: 56 (Quedius; subgenus Raphirus; Type locality: Zentral-Himalaya: Kumaon, Binsar, 7900 Fuss).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Cameron, 1932: 294 (Quedius; subgenus Raphirus; characters; India).

  • — Scheerpeltz, 1933: 1432 (Quedius; catalog).

  • — Coiffait, 1982a: 234 (Quedius; Pakistan).

  • — Smetana, 1988: 250 (Quedius; subgenus Raphirus; characters; Bhutan; Kashmir; Himachal Pradesh; Uttar Pradesh; West Bengal; Nepal; Pakistan).

  • — Smetana, 1992b: 5 (Quedius; subgenus Raphirus; Nepal).

  • Distribution: Pakistan, India, Nepal, Bhutan.

decipiens Cameron, 1944: 14 (Quedius; subgenus Raphirus; Type locality: Darjeeling: Ghum district: Mangpu).

  • — Smetana 1975a: 333 (Quedius; subgenus Raphirus; characters; Nepal; India; Bhutan).

  • — Smetana, 1977a: 246 (Quedius; Bhutan).

  • — Coiffait, 1982: 33 (Quedius; characters; Nepal).

  • — Coiffait, 1982a: 234 (Quedius; Nepal).

  • — Smetana, 1988: 250 (Quedius; synonym of aureiventris).

auricomus Kiesenwetter, 1850: 220 (Quedius; Type locality: Pyrenaei orientales).

  • — Kiesenwetter, 1851a: 420 (Quedius; characters; Pyrenees).

  • — Fairmaire and Laboulbène, 1856: 540 (Quedius; characters; France).

  • — Kraatz, 1857d: 501 (Quedius; characters).

  • — Fauvel, 1874: 303 [= 1874a: 529] (Quedius; characters; France; Britain).

  • — Redtenbacher, 1874: 202 (Quedius; characters).

  • — Mulsant and Rey, 1876: 765 [= 1877: 621] (Quedius; subgenus Raphirus; characters; France).

  • — Fowler, 1888: 242 (Quedius; subgenus Raphirus; characters; Britain).

  • — Ganglbauer, 1895: 410 (Quedius; subgenus Raphirus; characters; Germany; France; England).

  • — Porta, 1907: 144 (Quedius; subgenus Raphirus; characters; Italy).

  • — Reitter, 1909: 114 (Quedius; subgenus Raphirus; characters; Germany).

  • — Johansen, 1914: 369 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Gridelli, 1925: 144 (Quedius; subgenus Raphirus; characters; Pyrenees; Germany; Ireland; England; France; Alps; Italy; Belgium; Romania).

  • — Portevin, 1929: 344 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1432 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 48 (Quedius; aedeagal illustration).

  • — Tottenham, 1948a: 245 (Quedius; subgenus Arphirus; characters; Britain).

  • — V. Hansen, 1952: 151 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Smetana, 1958: 383 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 139 (Quedius; subgenus Raphirus; characters; Europe).

  • — Coiffait, 1963a: 383 (Quedius; subgenus Sauridus; characters).

  • — Palm, 1963b: 151 (Quedius; subgenus Raphirus; characters).

  • — Lohse, 1964: 214 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 290 (Quedius; subgenus Raphirus; Ireland; Britain; Denmark; Holland; Belgium; France; Switzerland; Germany; Austria).

  • — Coiffait, 1967e: 419 (Quedius; subgenus Microquedius).

  • — Bordoni, 1974a: 205, 207 (Quedius; subgenus Microquedius; characters; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 199 (Quedius; subgenus Microquedius; characters; England; Ireland; Belgium; France; Germany; Italy; Romania; Pyrenees).

  • — Tóth, 1984: 122 (Quedius; characters; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic ?).

  • Distribution: Europe.

tomentosomaculatus Cornelius, 1863: 130 (Quedius; Type locality: Elberfeld).

  • — Gemminger and Harold, 1868: 568 (Quedius; synonym of auricomus).

  • — Fowler, 1888: 242 (Quedius; subgenus Raphirus; synonym of auricomus).

  • — Ganglbauer, 1895: 410 (Quedius; synonym of auricomus).

  • — Porta, 1907: 144 (Quedius; synonym of auricomus).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; synonym of auricomus).

  • — Smetana, 1958: 384 (Quedius; synonym of auricomus).

  • — Coiffait, 1978: 199 (Quedius; synonym of auricomus).

aurofasciatus Bernhauer, 1917h: 91 (Quedius; subgenus Euryquedius; Type locality: Venezuela).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • Distribution: Venezuela.

aurorus Herman, 2001: 49 (Quedius; replacement name for laticollis Sharp).

  • Distribution: Japan.

laticollis Sharp, 1889: 31 (Quedius; [preoccupied]; Type locality: Sapporo).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1446 (Quedius; catalog).

  • — Herman, 2001: 49 (Quedius; synonym of aurorus).

babai Uéno and Watanabe, 1966: 331 (Quedius; subgenus Microsaurus; Type locality: Japan: central Honshu: southwestern Niigata Prefecture: Sekimi-toge, eastern slope of Mt. Myoko).

  • Distribution: Japan.

badius Broun, 1923: 678 (Quedius; Type locality: Gordon's Pyramid, Mount Arthur, 4500 ft.).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • Distribution: New Zealand.

badius Scholz, 1931 [Note: Nomen nudum], see: fuliginosus Gravenhorst, 1802.

bakeri Hatch, 1957: 218 (Quedius; subgenus Distichalius; Type locality: Washington: Mt. Baker, Chain L. Also cited from Washington).

  • — Smetana, 1971: 173 (Quedius; subgenus Distichalius; characters; Washington).

  • — Smetana, 1976c: 175 (Quedius; subgenus Distichalius; Oregon; Washington).

  • Distribution: USA.

balianii Gridelli, 1924, see: nigrocaeruleus Fauvel, 1876.

balcanicus Bernhauer, 1908e: 324 (Quedius; Type locality: Duzi in der Herzegowina).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Gridelli, 1922b: 128, 133 (Quedius; subgenus Sauridus; characters; Italy; Dalmatia; Hercegovina).

  • — Gridelli, 1925: 136 (Quedius; subgenus Raphirus; characters; Albania).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 41 (Quedius; aedeagal illustration).

  • — Coiffait, 1963a: 386 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 233 (Quedius; subgenus Sauridus; characters; Albania; Dalmatia; Istria).

  • — Tóth, 1984: 137 (Quedius; characters).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • Distribution: Italy, Bosnia Hercegovina, Croatia, Albania.

baldiensis Blackburn, 1891: 69 (Quedius; variety of cuprinus; Type locality: Mountains of Victoria, about 5,000 feet).

  • — Blackburn, 1903: 94 (Quedius; distinct species, not variety of cuprinus).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; variety of cuprinus).

  • — Lea, 1925: 242 (Quedius; cited as valid species; characters; notes; Australia).

  • Distribution: Australia.

balthasari Mařan, 1935, see: levicollis Brullé, 1832.

balticus Korge, 1960: 52 (Quedius; Type locality: Südspitze der Insel Hiddensee bei Rügen).

  • — Last, 1952a (Quedius; cited as molochinus; [Note: Misidentified according to Korge, 1960a: 67]; discussion and characters to separate from pallipes).

  • — Korge, 1960a: 67 (Quedius; characters; notes; England).

  • — Coiffait, 1961a: 50 (Quedius; characters).

  • — Smetana, 1962b: 135 (Quedius; subgenus Quedius; characters).

  • — V. Hansen, 1962a: 319 (Quedius; characters; Denmark).

  • — Last, 1963: 43 (Quedius; characters; notes; Britain).

  • — Palm, 1963: 117 (Quedius; characters; Sweden).

  • — Palm, 1963b: 160 (Quedius; characters; Sweden; Denmark).

  • — Lohse, 1964: 212 (Quedius; subgenus Quedius; characters; central Europe).

  • — Horion, 1965: 277 (Quedius; subgenus Quedius; Sweden; Denmark; Britain; France; Germany; Austria).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 192 (Quedius; subgenus Quedius; characters; Germany; Denmark; Britain; border of salt lakes in central Europe).

  • — Tóth, 1984: 119 (Quedius; characters; Hungary).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Austria; Benelux; Denmark; Sweden).

  • — Smetana, 1993b: 50 (Quedius; subgenus Quedius; Czech Republic; Slovakia).

  • — Hodge and Jones, 1995: 41 (Quedius; characters; Britain).

  • Distribution: Europe.

funebris Coiffait, 1971a: 175 (Quedius; subgenus Quedius; variety of balticus; [Note: Nomen nudum (Article 15.2)]; cited from France: La Ferté-Bernard, Sarthe).

  • — Coiffait, 1978: 192 (Quedius; subgenus Quedius; variety of balticus; characters).

  • — Tóth, 1984: 119 (Quedius; variety of balticus; characters).

bardus Melsheimer, 1844, see: capucinus Gravenhorst, 1806.

basiventris Sharp, 1884: 330 (Quedius; Type locality: Guatemala, Volcan de Atitlan about 3000 feet, Mirandilla 1700 feet, Las Mercedes, Pantaleon, San Isidro; Panama, Volcan de Chiriqui between 2500 and 4000 feet).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 174, 175 (Quedius; subgenus Cyrtoquedius; characters in key; Guatemala; Panama).

  • — Frank, 1982: 45 (Quedius; list of parasites).

  • Distribution: Guatemala, Panama.

becvari Smetana, 1996f: 119 (Quedius; subgenus Microsaurus; Type locality: China: Yunnan, Daju, 50 km N of Lijiang, 27.21N 100.19E).

  • — Smetana, 1997b: 130 (Quedius; notes; China).

  • — Smetana, 1997d: 455 (Quedius; subgenus Microsaurus; collecting notes; China).

  • — Smetana, 1999b: 216 (Quedius; subgenus Microsaurus; China).

  • Distribution: China.

beesoni Cameron, 1932: 285 (Quedius; subgenus Microsaurus; Type locality: Deoban, Chakrata, U.P.; [Note: See lectotype designation by Smetana, 1988]. Originally cited from Chakrata district, alt. 7000–9000 feet. Darjeeling Dist.: Ghum, Rongdong Valley).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — Smetana, 1988: 196 (Quedius; subgenus Microsaurus; lectotype designation: Deoban, Chakrata, U.P.; characters; Uttar Pradesh; West Bengal; Nepal).

  • — Smetana, 1995: 31 (Quedius; subgenus of Microsaurus; characters; bionomics; Taiwan).

  • — Smetana, 1996: 2 (Quedius; subgenus Microsaurus; notes; China).

  • — Smetana, 1999b: 214 (Quedius; subgenus Microsaurus; China).

  • Distribution: India, Nepal, China, Taiwan.

mimeticus Cameron, 1932: 286 (Quedius; subgenus Microsaurus; Type locality: Darjeeling Dist.: Ghum, Rongdong Valley; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1450 (Quedius; catalog).

  • — Smetana, 1988: 196 (Quedius; lectotype designation; synonym of beesoni).

  • — Smetana, 1995: 32 (Quedius; synonym of beesoni).

notabilis Cameron, 1932: 286 (Quedius; subgenus Microsaurus; Type locality: Darjeeling Dist.: Ghum, Rongdong Valley; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1452 (Quedius; catalog).

  • — Scheerpeltz, 1965: 111 (Quedius; subgenus Microsaurus; Burma).

  • — Smetana, 1988: 196 (Quedius; lectotype designation; synonym of beesoni).

  • — Smetana, 1995: 32 (Quedius; synonym of beesoni).

peraffinis Cameron, 1932: 286 (Quedius; subgenus Microsaurus; Type locality: Darjeeling Dist.: Ghum, Rongdong Valley; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1456 (Quedius; catalog).

  • — Smetana, 1988: 196 (Quedius; lectotype designation; synonym of beesoni).

  • — Smetana, 1995: 32 (Quedius; synonym of beesoni).

sungkangensis Hayashi, 1992: 11 (Quedius; subgenus Microsaurus; Type locality: Sungkang, Nantou Hsien).

  • — Hayashi, 1992a: 113 (Quedius; synonym of beesoni).

  • — Smetana, 1995: 32 (Quedius; synonym of beesoni).

bohemorum Smetana, 1997d: 461 (Quedius; subgenus Microsaurus; Type locality: China: N. Yunnan, Xue Shan, nr. Zhongdian, 4050 m, 27°49 N, 99°34 E).

  • Distribution: China.

belgravensis Oke, 1928: 1 (Quedius; Type locality: Australia: Victoria: Belgrave).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

bellus Lea, 1925: 243 (Quedius; Type locality: Victoria: Belgrave, Ferntree Gully, Beaconsfield).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

bernhaueri Rambousek, 1915a: 27, 29 (Quedius; subgenus Tenebrobius; Type locality: Peristerii-Gebirge bei Monastir in Mazedonien).

  • — Rambousek, 1916: 130 (Quedius; subgenus Tenebrobius; cited as a new species; Macedonia).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Gridelli, 1924d: 33 (Quedius; subgenus Microsaurus; characters; Macedonia).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — Coiffait, 1978: 181 (Quedius; subgenus Microsaurus; characters; Macedonia).

  • Distribution: Macedonia.

bernhauerianus Korge, 1971a: 56 (Quedius; replacement name for reitteri Bernhauer).

  • Distribution: China.

reitteri Bernhauer, 1933a: 51 (Quedius; subgenus Sauridus; [preoccupied]; Type locality: Tatsienlu-Kiulung).

  • — Korge, 1971a: 56 (Quedius; synonym of bernhauerianus).

berytensis Coiffait, 1954a: 160 (Quedius; subgenus Microsaurus; Type locality: Liban: Beyrouth).

  • — Coiffait, 1955e: 427 (Quedius; cave collection; Lebanon).

  • — Coiffait, 1978: 159 (Quedius; subgenus Microsaurus; characters; Lebanon).

  • Distribution: Lebanon.

beryticus Coiffait, 1977 [Note: Nomen nudum], see: quadriceps Fauvel, 1900.

bhari Smetana, 1988: 256 (Quedius; subgenus Raphirus; Type locality: Nepal: Khandbari District, For. above Ahale, 2400 m).

  • Distribution: Nepal.

bicolor Redtenbacher, 1849, see: fulgidus Fabricius, 1793.

bicolor Mulsant and Rey, 1876, see: angulicollis Fauvel, 1891.

bih Smetana, 1995: 101 (Quedius; subgenus Raphirus; Type locality: Taiwan: Chiai Hsien: Yushan N. P., Mun-Li Cliff, 2700 m).

  • — Smetana, 1998b: 110 (Quedius; subgenus Raphirus; characters).

  • Distribution: Taiwan.

bipunctatus Eppelsheim, 1895, see: Indoquedius.

bipustulatus Fiori, 1894, see: scintillans Gravenhorst, 1806.

birmanus Cameron, 1932: 284 (Quedius; subgenus Microsaurus; Type locality: Burma: Ruby Mines).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — Scheerpeltz, 1965: 111 (Quedius; subgenus Microsaurus; Burma).

  • — Smetana, 1988: 202 (Quedius; subgenus Microsaurus; characters; Burma).

  • Distribution: Myanmar.

biseriatus Stierlin, 1875, see: alpestris Heer, 1839.

bithynicus Fagel, 1968, see: gemellus Eppelsheim, 1889.

bito Smetana, 1996: 7 (Quedius; subgenus Microsaurus; Type locality: China: Gansu, Mts. 25 km E. Xiahe, 2805–2925 m).

  • — Smetana, 1997d: 452 (Quedius; subgenus Microsaurus; characters; collecting notes; China).

  • — Smetana, 1999b: 215 (Quedius; subgenus Microsaurus; collecting notes; China).

  • Distribution: China.

blackburni Bernhauer and Schubert, 1916, see: tepperi Blackburn, 1888.

bohemicus Roubal, 1941, see: puncticollis Thomson, 1867.

bolivianus Scheerpeltz, 1955a: 175, 181 (Quedius; subgenus Cyrtoquedius; Type locality: Bolivien: Yungas de la Paz).

  • Distribution: Bolivia.

boluensis Korge, 1971a: 49 (Quedius; subgenus Sauridus; Type locality: Abant-Gebirge bei Bolu).

  • — Coiffait, 1978: 209 (Quedius; subgenus Sauridus; characters; Anatolia).

  • Distribution: Turkey.

bonapartei Hatch, 1957, see: densiventris Casey, 1915.

bonvouloirii Brisout, 1863: 31 (Quedius; Type locality: Pyrénées-Orientales, Hautes-Pyrénées).

  • — Fauvel, 1869: 486 (Quedius; synonym of attenuatus).

  • — Fauvel, 1869a: 151 (Quedius; synonym of attenuatus).

  • — Fauvel, 1874: 307 [= 1874a: 533] (Quedius; synonym of acuminatus).

  • — Porta, 1907: 147 (Quedius; synonym of acuminatus).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; synonym of acuminatus).

  • — Scheerpeltz, 1933: 1429 (Quedius; synonym of acuminatus).

  • — Fagel, 1960b: 120 (Quedius; characters).

  • — Coiffait, 1978: 270 (Quedius; subgenus Raphirus; characters; Pyrenees).

  • Distribution: France, Spain.

boopoides Munster, 1923: 201, 203 (Quedius; subgenus Raphirus; variety of fulvicollis; [Note: Name cited as subspecies on page 201 and variety on page 203]; Type locality: Lysaker ved Kristiania; Fokstuen paa Dovre; Sebuhaugen saeter i Torpen; Kongsberg; Fyresdal; Grimstad; Mandal; Bø i Vesteraalen; Lødingen; Tromsø; Matjok i Tana; Vardø; Neiden samt fl. st. ved Pasvikelven i Syd Varanger).

  • — Tottenham, 1948a: 251 (Quedius; subgenus Arphirus; valid species; characters; Britain).

  • — V. Hansen, 1952: 156 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Smetana, 1958: 395 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Fagel, 1960b: 116 (Quedius; characters; Norway; Germany; Denmark; England).

  • — Smetana, 1962b: 153 (Quedius; subgenus Raphirus; characters).

  • — Palm, 1963b: 153 (Quedius; subgenus Raphirus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 219 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Smetana, 1964d: 80 (Quedius; subgenus Raphirus; Czechoslovakia).

  • — Horion, 1965: 305 (Quedius; subgenus Raphirus; Britain; Norway; Sweden; Finland; Russia; Denmark; Holland; Belgium; France; Czech Republic; Poland; Germany).

  • — Shilov, 1975a: 377 [= 1975b: 99] (Quedius; Komi region of Russia).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 280 (Quedius; subgenus Raphirus; characters; Europe; Britain; France).

  • — Hinton, 1981: 662 (Quedius; characters of egg).

  • — Tóth, 1984: 130 (Quedius; characters).

  • — Drugmand, 1987a: 312 (Quedius; Belgium).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Benelux; Denmark; Sweden).

  • — Nowosad, 1990: 149 (Quedius; collected from mole nest; notes; Poland).

  • — Whitehead, 1991 (Quedius; collecting notes; Britain).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • Distribution: Europe, Russia.

boops Gravenhorst, 1802: 21 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).

  • — Latreille, 1804: 311 (Staphylinus; characters; Germany; France).

  • — Gravenhorst, 1806: 71 (Staphylinus; characters; France).

  • — Gyllenhal, 1810: 312 (Staphylinus; characters; Sweden).

  • — Zetterstedt, 1828: 72 (Staphylinus; characters; Lapland).

  • — C. Sahlberg, 1830: 318 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 27 (Staphylinus; Finland).

  • — Mannerheim, 1831: 441 (Staphylinus; Finland).

  • — Stephens, 1833: 242 (Raphirus; characters; England; Scotland).

  • — Lacordaire, 1835: 385 (Emus; characters; France).

  • — Nordmann, 1837: 78 (Philonthus; Europe).

  • — Zetterstedt, 1838: 62 (Staphylinus; characters; habitat; Lapland).

  • — Erichson, 1839a: 494 (Quedius; characters; Germany).

  • — Heer, 1839: 280 (Philonthus; characters; Switzerland).

  • — Stephens, 1839: 393 (Quedius; characters; England; Scotland).

  • — Heer, 1839: 279 (Philonthus; variety b of picipennis described by Heer was cited as a synonym of boops by Scheerpeltz, 1933: 1434; characters; Switzerland).

  • — Erichson, 1840: 548 (Quedius; characters; Germany; Sweden).

  • — Kiesenwetter, 1848: 61 (Quedius; characters; Germany; Sweden).

  • — Gravenhorst, 1847: 229 (Quedius; characters; notes).

  • — Redtenbacher, 1849: 709 (Philonthus; characters; Austria).

  • — Kiesenwetter, 1851a: 421 (Quedius; Pyrenees).

  • — Hardy, 1851: 34 (Quedius; notes; England).

  • — Fairmaire and Laboulbène, 1856: 538 (Quedius; characters; France).

  • — Staudinger, 1857: 286 (Quedius; notes; Iceland).

  • — Kraatz, 1857d: 516 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 202 (Quedius; characters; Austria).

  • — Thomson, 1860: 179 (Raphirus; characters; Scandinavia).

  • — Fauvel, 1874: 308 [= 1874a: 534] (Quedius; characters; France; Iceland; Europe; Cyprus).

  • — Redtenbacher, 1874: 199 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 264 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 794 [= 1877: 650] (Quedius; subgenus Raphirus; characters; France).

  • — J. Sahlberg, 1876: 30 (Raphirus; Finland).

  • — Fowler, 1888: 245 (Quedius; subgenus Raphirus; characters; Britain).

  • — Eppelsheim, 1892: 332 (Quedius; Turkestan).

  • — Ganglbauer, 1895: 412 (Quedius; subgenus Raphirus; characters; Europe).

  • — Fauvel, 1902b: 117 (Quedius; Algeria; Tunisia).

  • — Poppius, 1905: 9 (Quedius; Shetland Islands).

  • — Sainte-Claire Deville, 1906: 103 (Quedius; Corsica).

  • — Porta, 1907: 150 (Quedius; subgenus Raphirus; characters; Italy).

  • — Poppius, 1909: 28 (Quedius; subgenus Raphirus; Lena Valley, Siberia).

  • — Reitter, 1909: 115 (Quedius; subgenus Raphirus; characters; Germany).

  • — Poppius, 1913: 10 (Quedius; Iceland).

  • — Johansen, 1914: 371 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Fiori, 1915: 16 (Quedius; Italy).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Henriksen and Lundbeck, 1918: 495 (Quedius; Greenland).

  • — Šulc, 1922 (Quedius; osmeterium; anatomy; histology; physiology).

  • — Gridelli, 1925: 165 (Quedius; subgenus Raphirus; characters).

  • — Portevin, 1929: 345 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — West, 1937: 38 (Quedius; Faroe Islands).

  • — Wüsthoff, 1938a: fig. 64–68 (Quedius; aedeagal illustration).

  • — Henriksen, 1939: 43 (Quedius; Greenland).

  • — V. Hansen, 1952: 156 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Fagel, 1955a: 85 (Quedius; subgenus Raphirus; Greece).

  • — Larsson and Gígja, 1959: 93 (Quedius; biological notes; Iceland).

  • — Fagel, 1960b: 111 (Quedius; characters; Belgium; France; England; Germany; Sweden; Italy; Austria; Yugoslavia; Spain; Portugal; Corsica; Algeria; Iceland; Norway).

  • — Smetana, 1962b: 153 (Quedius; subgenus Raphirus; characters).

  • — Szujecki, 1963a: 35 (Quedius; Poland).

  • — Palm, 1963b: 154 (Quedius; subgenus Raphirus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 218 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Smetana, 1964d: 81 (Quedius; subgenus Raphirus; Czechoslovakia).

  • — Scheerpeltz, 1965b: 408 (Quedius; Sicily).

  • — Horion, 1965: 306 (Quedius; subgenus Raphirus; northern and central Europe; Spain; Portugal; Corsica; Italy).

  • — Szujecki, 1968a: 736 (Quedius; Poland).

  • — Kasule, 1970: 64 (Quedius; larval characters).

  • — Bordoni, 1974b: 13 (Quedius; subgenus Raphirus; notes; Italy).

  • — Shilov, 1975a: 377 [= 1975b: 99] (Quedius; Komi region of Russia).

  • — Smetana, 1976d: 28 (Quedius; subgenus Raphirus; characters; Siberia).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 280 (Quedius; subgenus Raphirus; characters; Palaearctic region; Greenland; Iceland).

  • — Coiffait, 1978d: 165 (Quedius; Turkey).

  • — Tóth, 1984: 131 (Quedius; characters; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Lohse and Lucht, 1989: 175 (Quedius; characters).

  • — Whitehead, 1990e: 11 (Quedius; collecting notes; Britain).

  • — J. Li, 1993: 41 (Quedius; China).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • — Sadler and Dugmore, 1995: 144 (Quedius; habitat notes; Iceland).

  • — Muona and Viramo, 1995: 9 (Quedius; collected from dung of dog; Finland).

  • — Luff and Owen, 2000: 39 (Quedius; subgenus Raphirus; scutellar variation; separation from Q. aridulus; Britain).

  • — Assing, 2001: 75 (Quedius; collected from xerothermous site; Germany).

  • Distribution: Tunisia, Algeria, Greenland, Iceland, Europe, Turkey, Russia.

fuscipennis Stephens, 1832: 222 (Quedius; [preoccupied]; Type locality: Suffolk; Devonshire).

  • — Stephens, 1839: 391 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 23 (Quedius; synonym of boops).

tauricus Nordmann, 1837: 78 (Philonthus; variety of boops Gravenhorst; Type locality: Taurus Chersonesus).

  • — Scheerpeltz, 1933: 1434 (Quedius; variety of boops).

  • — Korge, 1964: 124 (Quedius; subgenus Raphirus; subspecies of boops; characters; Turkey).

coloratus W. Kolbe, 1912: 12 (Quedius; aberration of boops; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; aberration of boops).

  • — Gridelli, 1925: 165 (Quedius; aberration of boops).

  • — Scheerpeltz, 1933: 1434 (Quedius; aberration of boops).

  • — Coiffait, 1978: 281 (Quedius; subgenus Raphirus; variety of boops; characters).

  • — Tóth, 1984: 131 (Quedius; variety of boops; charactes; Hungary).

arestor Tottenham, 1948a: 255 (Quedius; subgenus Arphirus; Type locality: Britain: Chesil Beach).

  • — Smetana, 1958: 397 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Fagel, 1960a: 111 (Quedius; synonym of boops).

  • — Smetana, 1960b: 259 (Quedius; subgenus Raphirus; Czechoslovakia).

  • — Smetana, 1962b: 153 (Quedius; synonym of boops).

  • — Lohse, 1964: 218 (Quedius; synonym of boops).

  • — Pope, 1977: 31 (Quedius; synonym of boops).

  • — Coiffait, 1978: 281 (Quedius; subgenus Raphirus; subspecies of boops; characters).

  • — Smetana, 1993b: 50 (Quedius; synonym of boops).

islandicus Fagel, 1960b: 113 (Quedius; subspecies of boops; Type locality: Island: Reynivellir).

  • — Coiffait, 1978: 281 (Quedius; subgenus Raphirus; subspecies of boops; characters).

borjoniensis Coiffait, 1969 [Note: Nomen nudum], see: grouziacus Coiffait, 1969.

borneensis Cameron, 1933b: 347 (Quedius; Type locality: British North Borneo: Kenokok, 3300 feet).

  • — Korge, 1963a: 87 (Quedius; subgenus Indoquedius).

  • Distribution: Borneo.

bovinus Coquerel, 1860, see: Othius.

brachypterus Coiffait, 1967e: 401 (Quedius; subgenus Sauridus; Type locality: Caucase).

  • — Coiffait, 1978: 220 (Quedius; subgenus Sauridus; characters; Caucasus).

  • Distribution: Caucasus.

brandmayri Zanetti, 1992b: 135 (Quedius; subgenus Sauridus; Type locality: Sicilia: M. Soro).

  • — Sparacio, 1995: 165 (Quedius; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • Distribution: Italy.

breviceps Casey, 1915: 411 (Microsaurus; Type locality: British Columbia: Stikine River).

  • — Scheerpeltz, 1933: 1434 (Quedius; catalog).

  • — Hatch, 1957: 227 (Quedius; subgenus Sauridus; characters; British Columbia; Washington; Oregon).

  • — Smetana, 1964: 38 (Quedius; subgenus Raphirus; Alaska).

  • — Smetana, 1965a: 14 (Quedius; subgenus Raphirus; Alaska).

  • — Smetana, 1971: 202 (Quedius; subgenus Raphirus; characters; British Columbia; Alaska; California; Oregon; Washington).

  • — Smetana, 1971a: 1847 (Quedius; subgenus Raphirus; collecting notes; California).

  • — Smetana, 1971b: 4 (Quedius; subgenus Raphirus; Oregon).

  • — Smetana, 1976c: 181 (Quedius; subgenus Raphirus; California).

  • — Smetana, 1976d: 26 (Quedius; characters).

  • Distribution: Canada, USA.

brevicornis Thomson, 1860: 175 (Microsaurus; Type locality: Not cited but presumably Sweden).

  • — Sharp, 1867: 451 (Quedius; characters; habitat).

  • — Fauvel, 1874: 277 [= 1874a: 503] (Quedius; characters; France; Scandinavia; Britain; Austria; Russia).

  • — Mulsant and Rey, 1876: 636 [= 1877: 492] (Quedius; characters; France).

  • — Fowler, 1888: 233 (Quedius; characters; Britain).

  • — Ganglbauer, 1895: 398 (Quedius; subgenus Microsaurus; characters; north and middle Europe).

  • — Reitter, 1909: 110 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Johansen, 1914: 358 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Gridelli, 1924d: 59 (Quedius; subgenus Microsaurus; characters).

  • — Gridelli, 1929: 35 (Quedius; characters).

  • — Portevin, 1929: 339 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1434 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 12 (Quedius; aedeagal illustration).

  • — Roubal, 1941b: 124, 128 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — V. Hansen, 1952: 134 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Smetana, 1958: 351 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 158 (Quedius; subgenus Microsaurus; characters).

  • — Palm, 1963b: 134 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 208 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Horion, 1965: 265 (Quedius; subgenus Microsaurus; Ireland; Britain; Norway; Sweden; Finland; Denmark; Baltics; Russia; Ukraine; Poland; Czechoslovakia; Hungary; Romania; Bosnia; Austria; Germany; France; Belgium; Holland).

  • — Szujecki, 1968a: 734 (Quedius; Poland).

  • — Legner and Moore, 1977: 176 (Quedius; found under bark).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 160 (Quedius; subgenus Microsaurus; characters; Europe; Britain; France).

  • — Hoebeke, 1978: 508 (Quedius; parasitized by proctotrupid).

  • — Tóth, 1984: 110 (Quedius; characters; Hungary).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Benelux; Denmark; Sweden).

  • — Schülke and Uhlig, 1988: 12 (Quedius; notes; Germany).

  • — Drugmand, 1989a: 326 (Quedius; pupal characters; Belgium).

  • — Nowosad, 1990: 136 (Quedius; collected from mole nest; notes; Poland).

  • — Welch, 1993: 229 (Quedius; ovariole number and ovary structure).

  • — Wittwer, 1993: 251 (Quedius; Switzerland).

  • — Smetana, 1993b: 49 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • — Fowles, Alexander, and Key, 1999: 129 (Quedius; saproxylic species; status as endangered or threatened species; Britain).

  • Distribution: Europe, Russia.

eppelsheimii Seidlitz, 1891: 398 (Quedius; subgenus Microsaurus; [preoccupied]; Type locality: Eur. bis Curl.).

  • — Ganglbauer, 1895: 398 (Quedius; synonym of brevicornis).

  • — Thomson, 1896: 2389 (Microsaurus; synonym of pectinator).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; synonym of brevicornis).

  • — Gridelli, 1924d: 59 (Quedius; synonym of brevicornis).

  • — Smetana, 1958: 351 (Quedius; synonym of brevicornis).

  • — Coiffait, 1978: 160 (Quedius; synonym of brevicornis).

pectinator Thomson, 1896: 2389 (Microsaurus; replacement name for eppelsheimi Seidlitz; Sweden).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; synonym of brevicornis).

  • — Gridelli, 1924d: 59 (Quedius; synonym of brevicornis).

  • — Smetana, 1958: 351 (Quedius; synonym of brevicornis).

  • — Coiffait, 1978: 160 (Quedius; synonym of brevicornis).

elegans Ericson, 1897: 415 (Microsaurus; Type locality: Gernnebo in Vestergötland, Schweden).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Gridelli, 1924d: 59 (Quedius; synonym of brevicornis).

  • — Scheerpeltz, 1933: 1434 (Quedius; synonym of brevicornis).

  • — Smetana, 1958: 351 (Quedius; synonym of brevicornis).

  • — Coiffait, 1978: 160 (Quedius; synonym of brevicornis).

brevipennis Motschulsky, 1858, see: sturanyi Ganglbauer, 1895.

brevipennis Fairmaire, 1859, see: asturicus Bernhauer, 1918.

brevipennis Ihssen, 1939, see: haberfelneri Eppelsheim, 1891.

brevis Erichson, 1840: 535 (Quedius; Type locality: Germania septentrionalis).

  • — Märkel, 1841: 217 (Quedius; in ant nest).

  • — Mannerheim, 1843a: 83 (Quedius; characters).

  • — Grimm, 1845: 135 (Philonthus; with Formica rufa; Germany).

  • — Redtenbacher, 1849: 708 (Philonthus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 535 (Quedius; characters; France).

  • — Kraatz, 1857d: 500 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 200 (Quedius; characters; Austria).

  • — Thomson, 1860: 177 (Microsaurus; characters; Scandinavia).

  • — Hochhuth, 1862: 45 (Quedius; Russia).

  • — Fauvel, 1874: 275 [= 1874a: 501] (Quedius; characters; France).

  • — Redtenbacher, 1874: 197 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 266 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 625 [= 1877: 481] (Quedius; characters; France).

  • — J. Sahlberg, 1876: 28 (Microsaurus; Finland).

  • — Fowler, 1888: 236 (Quedius; characters; Britain).

  • — Ganglbauer, 1895: 397 (Quedius; subgenus Microsaurus; characters; north and middle Europe).

  • — Porta, 1907: 111 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Reitter, 1909: 109 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Johansen, 1914: 353 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • — Munster, 1923: 190 (Quedius; subgenus Microsaurus; Fennoscandia).

  • — Gridelli, 1924d: 23 (Quedius; subgenus Microsaurus; characters; Germany; Bohemia; Silesia; Italy; Istria).

  • — Portevin, 1929: 339 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1434 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 1 (Quedius; aedeagal illustration).

  • — Roubal, 1941b: 122, 126 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — V. Hansen, 1952: 126 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Smetana, 1958: 338 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 152 (Quedius; subgenus Microsaurus; characters).

  • — Palm, 1963b: 128 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 207 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Horion, 1965: 253 (Quedius; subgenus Microsaurus; northern and central Europe; Italy; France; Romania; Hungary; Czechoslovakia; Poland; Ukraine).

  • — Szujecki, 1968a: 734 (Quedius; Poland).

  • — Bordoni, 1974: 54 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 165 (Quedius; subgenus Microsaurus; characters; Europe; Scandinavia; Britain).

  • — Frank, 1982: 45 (Quedius; list of parasites).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Drugmand, 1989: 175 (Quedius; characters).

  • — Janák and Vysoký, 1992: 137 (Quedius; collected with Formica rufa and polyctena; Bohemia).

  • — Franc, 1993: 308 (Quedius; synecthrical myrmecophile; Slovakia).

  • — Smetana, 1993b: 49 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • Distribution: Europe, Russia.

lokayi M. Dvořák, 1967: 86 (Quedius; subgenus Microsaurus; variety of brevis; [Note: Nomen nudum]; cited from Vlašim).

slovenicus M. Dvořák, 1967: 85 (Quedius; subgenus Microsaurus; variety of brevis; [Note: Nomen nudum (Article 15.2)]; cited from Košice - Bankov).

fraudulentus Coiffait, 1967e: 394 (Quedius; subgenus Microsaurus; variety of brevis; [Note: Nomen nudum (Article 15.2)]; cited from Kosternick, Nukwaldy [Note: Correct spelling is Hukvaldy] Moravie, Tchécoslovaquie).

  • — Coiffait, 1978: 165 (Quedius; subgenus Microsaurus; variety of brevis; characters).

breweri† Scudder, 1890: 508 (Quedius; Type locality: Florissant; fossil).

  • — Scudder, 1891: 575 (Quedius; catalog; Oligocene; Colorado).

  • — Handlirsch, 1907: 723 (Quedius; catalog; Miocene; Colorado).

  • Distribution: USA.

brieni Bruge and Drugmand, 1988, see: invreae Gridelli, 1924.

brookesi Cameron, 1947d: 706 (Quedius; subgenus Microsaurus; Type locality: New Zealand: Southland: Hump Ridge).

  • Distribution: New Zealand.

brouni Cameron, 1947d: 706 (Quedius; subgenus Microsaurus; Type locality: New Zealand: Canterbury: Pudding Hill).

  • Distribution: New Zealand.

bruchi Bernhauer, 1934b: 216 (Quedius; subgenus Cyrtoquedius, cited as Cryptoquedius; Type locality: Argentinien: Tucuman).

  • — Scheerpeltz, 1955a: 175 (Quedius; subgenus Cyrtoquedius; characters in key; Argentina).

  • Distribution: Argentina.

brunneorufus Bernhauer, 1941b: 30 (Quedius; subgenus Sauridus; Type locality: Neu Seeland: Greymouth).

  • Distribution: New Zealand.

brunnipennis Mannerheim, 1843b: 232 (Quedius; Type locality: insula Sitkha).

  • — Mannerheim, 1852: 314 (Quedius; characters).

  • — Horn, 1878: 164 (Quedius; characters; Lake Superior region westward to Vancouver and northward to Alaska).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • — Fall, 1926: 150 (Quedius; cited as brunneipennis; Alaska).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Hatch, 1957: 217 (Quedius; subgenus Distichalius; cited as brunneipennis; characters; British Columbia; Washington; Oregon).

  • — Smetana, 1971: 197 (Quedius; subgenus Raphirus; characters; Alberta; British Columbia; Labrador; Newfoundland; Northwest Territories; Ontario; Quebec; Yukon Territory; Alaska; Michigan; New Hampshire; New York).

  • — Smetana, 1973: 1429 (Quedius; notes; Alberta; Northwest Territories; Quebec).

  • — Smetana, 1976c: 181 (Quedius; subgenus Raphirus; Northwest Territories; Ontario).

  • — Smetana, 1976d: 26 (Quedius; characters).

  • — Smetana, 1978a: 830 (Quedius; subgenus Raphirus; collecting notes; Labrador; Newfoundland).

  • — Downie and Arnett, 1996: 393 (Quedius; subgenus Raphirus; characters; USA).

  • Distribution: Canada, USA.

bruttius Zanetti, 1977: 307 (Quedius; Type locality: Gariglione, La Sila).

  • — Coiffait, 1978: 174 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; probably subspecies of latialis; Italy).

  • Distribution: Italy.

bryocharis Broun, 1923: 678 (Quedius; Type locality: Ben Lomond, 4500 ft.).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • Distribution: New Zealand.

bucharensis Bernhauer, 1918a: 93 (Quedius; subgenus Ediquus; Type locality: Karateghingebirge: Baldschuan, 924 m; Buchara; Persien: Kopet-Dagh, Siaret, 1160 m).

  • — Gridelli, 1924d: 56 (Quedius; subgenus Microsaurus; characters; central Asia; Iran).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Coiffait, 1978: 186 (Quedius; subgenus Microsaurus; characters; Transcaspian region).

  • Distribution: “Transcaspia”, Uzbekistan, Iran.

bulgaricus Scheerpeltz, 1937b: 219 (Quedius; Type locality: Bulgarien: Rosalito-polje, Schipka-Balkan, ca. 1900 m).

  • — Coiffait, 1963a: 387 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 262 (Quedius; subgenus Sauridus; characters; Bulgaria).

  • Distribution: Bulgaria.

buphthalmus Erichson, 1840, see: Bolitogyrus.

bureschi Coiffait, 1971b: 283 (Quedius; subgenus Microsaurus; Type locality: Bulgarie: Aleko, 1800 m, Mt. Vitosha, près de Sofia).

  • — Coiffait, 1978: 149 (Quedius; subgenus Microsaurus; characters; Bulgaria).

  • Distribution: Bulgaria.

burrowsi Broun, 1915: 278 (Quedius; Type locality: Pudding Hill, near Methven).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • Distribution: New Zealand.

cachemiricus Coiffait, 1982, see: fluviatilis Cameron, 1926.

caelebs Rottenberg, 1870: 28 (Quedius; Type locality: Sicily: Nicolosi; Palermo; Girgenti).

  • — Ganglbauer, 1895: 404 (Quedius; synonym of picipes).

  • — Porta, 1907: 135 (Quedius; synonym of picipes).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; cited as coelebs; synonym of picipes).

  • — Gridelli, 1924d: 109 (Quedius; subgenus Raphirus; cited as coelebs characters; Sicily).

  • — Scheerpeltz, 1933: 1436 (Quedius; cited as coelebs; catalog).

  • — Coiffait, 1963a: 378 (Quedius; subgenus Sauridus; cited as coelebs; characters).

  • — Coiffait, 1978: 212 (Quedius; subgenus Sauridus; cited as coelebs; characters; Sicily).

  • — Sparacio, 1995: 165 (Quedius; cited as coelebs; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; cited as coelebs; Italy).

  • Distribution: Italy.

caelestis Smetana, 1996a: 54 (Quedius; subgenus Raphirus; Type locality: China: Yunnan, 50 km N Lijiang, Yulongshan Nat. Res.).

  • Distribution: China.

californicus Smetana, 1971, see: desertus Horn, 1878.

calignosus Stephens, 1832 [Note: Stephens, 1839: 392 cited this name as caliginosus], see: Callicerus.

calli Smetana, 1976a: 244 (Quedius; subgenus Quedionuchus; Type locality: Mexico: Nuevo Leon: Cerro Potosi, 12,000 ft.).

  • Distribution: Mexico.

calogaster Lea, 1929: 206 (Quedius; Type locality: Queensland: Ravenshoe and Magnetic Island).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • Distribution: Australia.

campbelli Smetana, 1971: 103 (Quedius; subgenus Microsaurus; Type locality: Quebec: Gatineau Park, Lac Renaud).

  • — Smetana, 1976c: 172 (Quedius; subgenus Microsaurus; Ontario).

  • — Smetana, 1981b: 636 (Quedius; Saskatchewan).

  • — Smetana, 1990d: 97 (Quedius; Maine).

  • — Downie and Arnett, 1996: 390 (Quedius; subgenus Microsaurus; characters; USA).

  • Distribution: Canada, USA.

canadensis Casey, 1915: 409 (Microsaurus; Type locality: Canada: Kazubazua, Province of Quebec).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Smetana, 1971: 53 (Quedius; subgenus Microsaurus; characters; New Brunswick; Ontario; Quebec; Connecticut; Massachusetts; Maine; Michigan; Minnesota; New Hampshire; New Jersey; New York; North Carolina; Ohio; Pennsylvania).

  • — Smetana, 1971b: 2 (Quedius; subgenus Microsaurus; New York).

  • — Smetana, 1973: 1425 (Quedius; Ohio; Ontario; Quebec).

  • — Smetana, 1978a: 823 (Quedius; subgenus Microsaurus; collecting notes; North Carolina; Quebec).

  • — Smetana, 1981b: 633 (Quedius; collecting notes; New York; Ohio; Ontario).

  • — Smetana, 1990d: 96 (Quedius; Massachusetts; New Hampshire; New York; Ohio; Pennsylvania).

  • — Downie and Arnett, 1996: 391 (Quedius; subgenus Microsaurus; characters; USA).

  • Distribution: Canada, USA.

candicus Coiffait, 1976b: 90 (Quedius; subgenus Sauridus; Type locality: Crête: Héraklion, Gonie).

  • — Coiffait, 1978: 228 (Quedius; subgenus Sauridus; characters; Crete).

  • Distribution: Greece.

capitalis Eppelsheim, 1892: 329 (Quedius; Type locality: Taschkent).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • — Gridelli, 1924d: 40 (Quedius; subgenus Microsaurus; characters; Tashkent).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Coiffait, 1978: 148 (Quedius; subgenus Microsaurus; characters; Turkestan).

  • Distribution: Uzbekistan.

capucinus Gravenhorst, 1806: 40 (Staphylinus; Type locality: America septentr.; [Note: See lectotype designation by Smetana, 1971]).

  • — Nordmann, 1837: 79 (Philonthus; America borealis).

  • — Erichson, 1840: 531 (Quedius; characters; America septentrionalis).

  • — Gravenhorst, 1847: 235 (Quedius; characters; notes).

  • — Provancher, 1877: 728 (Quedius; characters; Quebec).

  • — Horn, 1878: 160 (Quedius; characters; California; Atlantic region to Georgia).

  • — Blatchley, 1910: 371 (Quedius; characters; Indiana).

  • — Casey, 1915: 405 (Distichalius; characters; New York to Virginia westward to Iowa).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Voris, 1939: 154 (Quedius; larval and pupal characters; Indiana).

  • — Smetana, 1971: 161 (Quedius; subgenus Distichalius; lectotype designation; characters; Ontario; Quebec; Alabama; Arkansas; Connecticut; Georgia; Illinois; Indiana; Kansas; Kentucky; Louisiana; Maine; Maryland; Massachusetts; Michigan; Mississippi; Missouri; New Hampshire; New Jersey; New York; North Carolina; Ohio; Pennsylvania; South Carolina; Tennessee; Texas; Virginia; West Virginia; Wisconsin).

  • — Smetana, 1971b: 3 (Quedius; subgenus Distichalius; Maine).

  • — Smetana, 1978a: 827 (Quedius; subgenus Distichalius; collecting notes; Louisiana; Texas).

  • — Smetana, 1981b: 640 (Quedius; Ontario).

  • — Hunter, Kincher, Bay, Beerwinkle, 1991: 169 (Quedius; seasonal distribution and diel flight activity; Texas).

  • — Downie and Arnett, 1996: 392 (Quedius; subgenus Distichalius; characters; USA).

  • Distribution: Canada, USA.

inversus Say, 1830: 36 [= 1834: 449] (Staphylinus; Type locality: Indiana).

  • — Erichson, 1840: 531 (Quedius; synonym of capucinus).

  • — Horn, 1878: 160 (Quedius; synonym of capucinus).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; synonym of capucinus).

  • — Smetana, 1971: 161 (Quedius; synonym of capucinus).

bardus Melsheimer, 1844: 39 (Quedius; Type locality: Pennsylvania).

  • — Horn, 1878: 160 (Quedius; synonym of capucinus).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1435 (Quedius; synonym of capucinus).

  • — Smetana, 1971: 161 (Quedius; synonym of capucinus).

ater Ziegler, 1845: 266 (Philonthus; Type locality: Pennsylvania).

  • — Fauvel, 1869: 488 (Quedius; synonym of capucinus).

  • — Horn, 1878: 160 (Quedius; synonym of capucinus).

  • — Bernhauer and Schubert, 1914: 329 (Philonthus; catalog).

  • — Scheerpeltz, 1933: 1435 (Quedius; synonym of capucinus).

  • — Smetana, 1971: 161 (Quedius; synonym of capucinus).

seeversi Korge, 1963: 122 (Quedius; subgenus Distichalius; Type locality: California: Contra Costa).

  • — Smetana, 1971: 161 (Quedius; synonym of capucinus).

carnicus Fiori, 1894, see: cincticollis Kraatz, 1857.

carpaticola Roubal, 1924, see: cincticollis Kraatz, 1857.

caseyi Scheerpeltz, 1933, see: uteanus Casey, 1915.

castaneipennis Hochhuth, 1872, see: puncticollis Thomson, 1867.

castanopterus Stephens, 1832, see: Philonthus.

cavelli Broun, 1893: 1032 (Quedius; Type locality: Boatman's).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • Distribution: New Zealand.

centrasiaticus Coiffait, 1969: 54 (Quedius; subgenus Raphirus; replacement name for altaicus Coiffait).

  • — Coiffait, 1978: 267 (Quedius; subgenus Raphirus; characters; central Asia).

  • Distribution: Russia.

altaicus Coiffait, 1967e: 422 (Quedius; subgenus Raphirus; [preoccupied]; Type locality: Telekschk Ozero, 1000 m, Altaï).

  • — Coiffait, 1978: 267 (Quedius; synonym of centrasiaticus).

cephalotes Uéno and Watanabe, 1966: 326 (Quedius; subgenus Microsaurus; Type locality: Japan: on the central massif of Shikoku: Ehime Prefecture: a limestone cave called “Rakan-ana”, at Koya in Sogawa of Nomura-cho).

  • — Uéno and Watanabe, 1970: 18 (Quedius; subgenus Microsaurus; notes; Japan).

  • — Watanabe, 1996a: 13 (Quedius; cave and collections; Japan).

  • — Bordoni and Oromi, 1998: 1157 (Quedius; subgenus Microsaurus; facultative cave species; Japan).

  • Distribution: Japan.

chalceiventris Fauvel, 1878f: 275 (Quedius; Type locality: Nouvelle-Guinée, Hatam).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • Distribution: New Guinea.

chalybeipennis W. J. MacLeay, 1873, see: ruficollis Gravenhorst, 1806.

chamberlini† Scudder, 1890: 508 (Quedius; Type locality: Florissant; fossil).

  • — Scudder, 1891: 575 (Quedius; catalog; Oligocene; Colorado).

  • — Handlirsch, 1907: 723 (Quedius; catalog; Miocene; Colorado).

  • — Wickham, 1912: 11 (Quedius; Colorado).

  • — Wickham, 1913: 287 (Quedius; Colorado; doubtful identification).

  • Distribution: USA.

championi Sharp, 1884: 329 (Quedius; Type locality: Guatemala, San Geronimo 3000 feet, Purula).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • Distribution: Guatemala.

chang Smetana, 1995: 83 (Quedius; subgenus Raphirus; Type locality: Taiwan: Taichung Hsien: Anmashan, 2230 m).

  • Distribution: Taiwan.

charactus Hatch, 1957: 222 (Quedius; subgenus Microsaurus; Type locality: Oregon: Curry Co.: Pistol River. Also cited from British Columbia).

  • — Smetana, 1971: 79 (Quedius; subgenus Microsaurus; characters; British Columbia; California; Oregon; Washington).

  • — Smetana, 1971b: 2 (Quedius; subgenus Microsaurus; British Columbia).

  • — Legner and Moore, 1977: 176 (Quedius; found under bark).

  • — Smetana, 1981b: 634 (Quedius; collecting notes; Washington).

  • Distribution: Canada, USA.

chatterjeei Cameron, 1926b: 370 (Quedius; Type locality: Woodstock Falls, Mussoorie; [Note: See lectotype designation by Smetana, 1988]. Originally cited from Mussorie and Chakrata districts, 6000 to 9000 feet).

  • — Cameron, 1932: 295 (Quedius; subgenus Raphirus; characters; India).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Smetana, 1988: 229 (Quedius; subgenus Distichalius; lectotype designation: Woodstock Falls, Mussoorie; characters; Himachal Pradesh; Uttar Pradesh).

  • Distribution: India.

chi Smetana, 1995: 76 (Quedius; subgenus Raphirus; Type locality: Taiwan: Taichung Hsien: Hsuehshan, above Shan-Liu-Gieu Hut, 3200 m).

  • Distribution: Taiwan.

chiangi Smetana, 1995: 48 (Quedius; subgenus Microsaurus; Type locality: Taiwan: Nantou Hsien, Yushan N. P., Pai-Yun Hut, 3528 m).

  • Distribution: Taiwan.

chiaw Smetana, 1995: 56 (Quedius; subgenus Microsaurus; Type locality: Taiwan: Meifo Formosa).

  • — Smetana, 1996g: 24 (Quedius; notes; Taiwan).

  • Distribution: Taiwan.

chinensis Bernhauer, 1915c: 74 (Quedius; subgenus Raphirus; Type locality: Kiautschou).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • — Gridelli, 1925: 149 (Quedius; subgenus Raphirus; characters).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 52 (Quedius; aedeagal illustration).

  • — Smetana, 1996c: 226 (Quedius; subgenus Raphirus; characters; China).

  • Distribution: China.

chiricahuensis Smetana, 1971, see: leechi Smetana, 1971.

chlorophanus Erichson, 1840: 532 (Quedius; Type locality: Bengala).

  • — Kraatz, 1859: 66 (Quedius; characters; Bengalia).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • — Cameron, 1932: 283 (Quedius; subgenus Microsaurus; characters; Bengal).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Smetana, 1988: 184 (Quedius; may not be in Quedius; probably not in Himalaya).

  • Distribution: India.

chremes Smetana, 1996: 10 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan Langmusi, 3500–3600 m).

  • — Smetana, 1999b: 215 (Quedius; subgenus Microsaurus; China).

  • Distribution: China.

chrysogonus Smetana, 1997b: 132 (Quedius; subgenus Raphirus; Type locality: China: Mt. Ailao Shan, 2500 m, Jingdong, Yunnan).

  • — Smetana, 1998b: 109 (Quedius; subgenus Raphirus; characters).

  • Distribution: China.

chrysurus Kiesenwetter, 1848, see: microps Gravenhorst, 1847.

cincticollis Kraatz, 1857d: 508 (Quedius; Type locality: Wien).

  • — Redtenbacher, 1857: 985 (Quedius; characters; Austria).

  • — Redtenbacher, 1874: 201 (Quedius; characters; Austria).

  • — Mulsant and Rey, 1876: 718 [= 1877: 574] (Quedius; characters; Austria; Hungary).

  • — Ganglbauer, 1895: 409 (Quedius; subgenus Raphirus; characters; eastern Alps; Carpathians).

  • — Porta, 1907: 144 (Quedius; subgenus Raphirus; characters; Italy).

  • — Reitter, 1909: 114 (Quedius; subgenus Raphirus; characters; Germany).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • — Gridelli, 1925: 141 (Quedius; subgenus Raphirus; characters; Alps; Bosnia).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 45 (Quedius; aedeagal illustration).

  • — Smetana, 1958: 380 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 139 (Quedius; subgenus Raphirus; characters; Alps and Carpathians).

  • — Coiffait, 1963a: 379 (Quedius; subgenus Sauridus; characters).

  • — Lohse, 1964: 213 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 288 (Quedius; subgenus Raphirus; Austria; Slovenia; Italy; Poland; Czechoslovakia; Hungary; Romania; Bosnia Hercegovina; Ukraine).

  • — Shilov, 1975a: 377 [= 1975b: 99] (Quedius; Komi region of Russia).

  • — Coiffait, 1978: 225 (Quedius; subgenus Sauridus; characters; Alps; Carpathians; Bosnia; mountains of central Europe).

  • — Tóth, 1984: 135 (Quedius; characters).

  • — Lucht, 1987: 110 (Quedius; checklist; Poland; Czechoslovakia; Austria).

  • — Lohse and Lucht, 1989: 172 (Quedius; characters).

  • — Nowosad, 1990: 146 (Quedius; collected from mole nest; notes; Poland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • Distribution: Europe, Russia.

carnicus Fiori, 1894: 88 (Quedius; Type locality: Visdende, nelle Alpi carniche).

  • — Porta, 1907: 144 (Quedius; synonym of cincticollis).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; synonym of cincticollis).

  • — Gridelli, 1925: 141 (Quedius; synonym of cincticollis).

  • — Smetana, 1958: 380 (Quedius; synonym of cincticollis).

  • — Coiffait, 1978: 225 (Quedius; synonym of cincticollis).

carpaticola Roubal, 1924: 245 (Quedius; variety of cincticollis; Type locality: Pietros).

  • — Scheerpeltz, 1933: 1435 (Quedius; variety of cincticollis).

  • — Smetana, 1958: 380 (Quedius; cited as carpathicola; variety of cincticollis).

  • — Coiffait, 1978: 225 (Quedius; cited as carpathicola; variety of cincticollis; characters).

  • — Tóth, 1984: 135 (Quedius; cited as carpathicola; subspecies of cincticollis; characters).

cinctipennis Cameron, 1951c: 406 (Quedius; Type locality: Kenya Colony: Kanziko).

  • Distribution: Kenya.

cinctus Paykull, 1790: 137 (Staphylinus; Type locality: Suecia).

  • — Paykull, 1800: 395 (Staphylinus; characters; Sweden).

  • — Gravenhorst, 1802: 35 (Staphylinus; synonym of impressus).

  • — Marsham, 1802: 505 (Staphylinus; synonym of marginellus).

  • — C. Sahlberg, 1830: 317 (Staphylinus; synonym of impressus).

  • — O. Costa, 1839: 47 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1916: 420, cited marginellus Costa as a synonym of cinctus Paykull; Costa did not describe the species as new, he attributed it to Fabricius]; characters; Italy).

  • — Erichson, 1839a: 489 (Quedius; synonym of impressus).

  • — Erichson, 1840: 530 (Quedius; synonym of impressus).

  • — Kiesenwetter, 1848: 50 (Quedius; characters; Europe).

  • — Kraatz, 1857d: 499 (Quedius; synonym of impressus).

  • — Crotch, 1870: 100 (Quedius; valid name).

  • — Fauvel, 1874: 288 [= 1874a: 514] (Quedius; characters; France; Algeria; Turkey).

  • — Seidlitz, 1875: 266 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 676 [= 1877: 532] (Quedius; characters; larval characters; France).

  • — J. Sahlberg, 1876: 24 (Microsaurus; Finland).

  • — Fauvel, 1878: 127 [= 1878a: 47] (Quedius; Algeria).

  • — Fauvel, 1886: 58 [= 1886a: 50] (Quedius; Algeria).

  • — Fowler, 1888: 235 (Quedius; characters; Britain).

  • — Ganglbauer, 1895: 402 (Quedius; subgenus Quedionuchus; characters; Europe; Mediterranean region).

  • — Fauvel, 1902b: 115 (Quedius; synonym of flavescens).

  • — Poppius, 1905: 9 (Quedius; Shetland Islands).

  • — Porta, 1907: 107 (Quedius; subgenus Quedionuchus; characters; Italy).

  • — Sainte-Claire Deville, 1906: 101 (Quedius; Corsica).

  • — Reitter, 1909: 111 (Quedius; subgenus Quedionuchus; characters; Germany).

  • — Johansen, 1914: 362 (Quedius; subgenus Quedionuchus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • — Munster, 1923: 194 (Quedius; subgenus Quedionuchus; Fennoscandia).

  • — Gridelli, 1924d: 75 (Quedius; subgenus Quedionuchus; characters; Albania).

  • — Portevin, 1929: 338 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 18 (Quedius; aedeagal illustration).

  • — Tottenham, 1949: 422 (Quedius; checklist; Britain).

  • — Jeannel and Jarrige, 1949: 38 (Quedius; subgenus Quedionuchus; notes; cave species; Serbia).

  • — V. Hansen, 1952: 139 (Quedius; subgenus Quedionuchus; characters; Denmark).

  • — Coiffait, 1955e: 427 (Quedius; Lebanon).

  • — Scheerpeltz, 1956c: 1102 (Quedius; subgenus Quedionuchus; Cyprus).

  • — Smetana, 1958: 357 (Quedius; subgenus Quedionuchus; characters; ecology; Czechoslovakia).

  • — Smetana, 1959a: 215 (Quedius; Albania).

  • — Smetana, 1960e: 160 (Quedius; subgenus Quedionuchus; characters).

  • — Scheerpeltz, 1960d: 136 (Quedius; subgenus Quedionuchus; Portugal).

  • — Scheerpeltz, 1961d: 133 (Quedius; subgenus Quedionuchus; Italy).

  • — Scheerpeltz, 1961e: 137 (Quedius; subgenus Quedionuchus; Rhodes).

  • — Ferreira, 1962: 19 (Quedius; Portugal).

  • — Scheerpeltz, 1963a: 420 (Quedius; subgenus Quedionuchus; Greece).

  • — Palm, 1963b: 137 (Quedius; subgenus Quedionuchus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 210 (Quedius; subgenus Quedionuchus; characters; central Europe).

  • — Korge, 1964: 118 (Quedius; subgenus Quedionuchus; characters).

  • — Scheerpeltz, 1964a: 300 (Quedius; subgenus Quedionuchus; Crete).

  • — Horion, 1965: 269 (Quedius; subgenus Quedionuchus, cited as Quedionychus; Caucasus; Morocco; Tunisia; Cyprus; Turkey; Scotland; Lapland; Russia; all central and southern Europe).

  • — Szujecki, 1968a: 735 (Quedius; Poland).

  • — Scheerpeltz, 1968a: 90 (Quedius; catalog; Austria).

  • — Kasule, 1970: 59 (Quedionuchus; larval characters).

  • — Smetana, 1971: 165 (Quedius; subgenus Distichalius; characters; Massachusetts).

  • — Korge, 1971a: 47 (Quedius; subgenus Quedionuchus; characters; Turkey).

  • — Bordoni, 1973b: 49 (Quedius; subgenus Distichalius; characters; Italy).

  • — Coiffait, 1976b: 82 (Quedius; Greece; Rhodes; Thasos).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 186 (Quedius; subgenus Distichalius; characters; Europe; Scandinavia; Britain; Asia Minor; Middle East; North Africa).

  • — Smetana, 1978: 123 (Quedius; subgenus Distichalius; collecting notes; Israel).

  • — Muona, 1979: 15 (Quedius; Finland; Russia; Sweden; Norway; Denmark; Baltic States).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 131 (Quedius; catalog; Poland).

  • — Hinton, 1981: 663 (Quedius; characters of egg).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • — Bordoni, 1984: 342 (Quedius; Lebanon).

  • — Tóth, 1984: 103 (Quedius; characters; Hungary).

  • — Muona and Viramo, 1986: 14 (Quedius; Finland).

  • — H. Franz, 1987: 70 (Quedius; Cyprus).

  • — Ádám, 1987: 152 (Distichalius; collecting notes; Hungary).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Lohse and Lucht, 1989: 172 (Quedius; subgenus Distichalius).

  • — Smetana, 1990d: 99 (Quedius; New Jersey; New York; Washington).

  • — Whitehead, 1990e: 11 (Quedius; collecting notes; Britain).

  • — Allen, 1990b: 2 (Quedius; collecting notes; Britain).

  • — Angelini, 1991: 193 (Quedius; Italy).

  • — Silfverberg, 1992: 18 (Quedius; Finland; Russia; Sweden; Norway; Denmark; Estonia; Latvia; Lithuania).

  • — Welch, 1993: 229 (Quedius; ovariole number and ovary structure).

  • — Smetana, 1993b: 50 (Quedius; subgenus Distichalius; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • — Downie and Arnett, 1996: 392 (Quedius; subgenus Distichalius; characters; USA).

  • — M. Hansen, 1996: 107 (Quedius; checklist; Denmark).

  • — Ádám, 1996: 247 (Distichalius; synonym of Staphylinus flavicornis Gmelin; [Note: Until the identity of the Gmelin name has been verified it is best to allow it to remain in Staphylinus as a nomen dubium; furthermore, even if flavicornis and cinctus are synonyms, the latter should be protected via Article 23.9.1]).

  • — Eisinger, 1997: 171 (Quedius; collecting notes; Germany).

  • — T. Wagner, 1997: 235 (Quedius; collecting notes; Germany).

  • — Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 39 (Quedius; Latvia).

  • — Anderson, Nash, and O'Connor, 1997: 19 (Quedius; Ireland).

  • — Dauphin, Laguerre, Tamisier, and Teissier, 1997: 20 (Quedius; France).

  • — Bordoni and Oromi, 1998: 1157 (Quedius; subgenus Quedionuchus; facultative cave species; Serbia).

  • — Owen, 1999b: 61 (Quedius; captured underground at roots of old trees; Britain).

  • — Owen, 2000: 251 (Quedius; pitfall trap collections from underground at roots of trees; Britain).

  • — Assing and Wunderle, 2001: 37 (Quedius; Cyprus).

  • — Herman, 2001: 29 (Quedius; nomen protectum; discussion).

  • Distribution: Tunisia, Algeria, Morocco, Europe, Russia, Georgia, Turkey, Lebanon, Israel, USA.

flavescens Linné, 1758: 422 (Staphylinus; [Note: This name is older than cinctus but has hardly been cited since its original description. In 1775 Fabricius attributed flavescens to Linné, 1767: 684. Most subsequent workers have credited Fabricius with the name]; Type locality: Europa).

  • — Linné, 1760: 232 (Staphylinus; characters; Sweden).

  • — Pontoppidans, 1765: 211 (Staphylinus; Denmark).

  • — Linné, 1767: 684 (Staphylinus; characters; Europe).

  • — Fabricius, 1775: 266 (Staphylinus; characters; Europe).

  • — O. Müller, 1776: 98 (Staphylinus; characters).

  • — Goeze, 1777: 719 (Staphylinus; characters).

  • — Fabricius, 1781: 336 (Staphylinus; characters; Europe).

  • — Fabricius, 1787: 221 (Staphylinus; characters).

  • — Villers, 1789: 415 (Staphylinus; characters).

  • — Gmelin, 1790: 2029 (Staphylinus; characters).

  • — Fabricius, 1793: 526 (Staphylinus; characters).

  • — Panzer, 1795: 355 (Staphylinus; characters; Germany).

  • — Cederhjelm, 1798: 118 (Staphylinus; characters).

  • — Fabricius, 1801: 597 (Staphylinus; characters).

  • — Turton, 1802: 512 (Staphylinus; characters; Europe).

  • — Curtis, 1829: 25 (Quedius; catalog; Britain).

  • — C. Sahlberg, 1830: 324 (Staphylinus; synonym of discoideus Gravenhorst).

  • — Westwood, 1845: 49 (Staphylinus; notes; no specimen in Linnean cabinet).

  • — Kraatz, 1857d: 499 (Quedius; synonym of impressus).

  • — Gemminger and Harold, 1868: 570 (Quedius; synonym of impressus).

  • — Fauvel, 1902b: 115 (Quedius; [Note: Misidentification: Scheerpeltz, 1933: 1436, cited flavescens Fauvel as a synonym of cinctus; Fauvel did not describe the species as new, he attributed it to Linné]; Morocco; Algeria; Tunisia).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; synonym of cinctus).

  • — Gridelli, 1924d: 75 (Quedius; synonym of cinctus).

  • — Scheerpeltz, 1933: 1436 (Quedius; synonym of cinctus).

  • — Smetana, 1958: 358 (Quedius; synonym of cinctus).

  • — Bordoni, 1973b: 49 (Quedius; synonym of cinctus).

  • — Coiffait, 1978: 186 (Quedius; synonym of cinctus).

  • — Herman, 2001: 29 (Quedius; nomen oblitum).

impressus Panzer, 1796d: 21 (Staphylinus; [preoccupied]; Type locality: Neuwaldegg).

  • — Gravenhorst, 1802: 35 (Staphylinus; characters).

  • — Latreille, 1804: 320 (Staphylinus; characters).

  • — Gravenhorst, 1806: 39 (Staphylinus; characters; Germany).

  • — Panzer, 1808a: 21 (Staphylinus; characters; illustrations).

  • — Gyllenhal, 1810: 307 (Staphylinus; characters; Sweden).

  • — Curtis, 1829: 25 (Quedius; catalog; Britain).

  • — C. Sahlberg, 1830: 317 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 26 (Staphylinus; Finland; Sweden).

  • — Mannerheim, 1831: 440 (Staphylinus; Finland; Sweden).

  • — Stephens, 1832: 219 (Quedius; characters; England).

  • — Brullé, 1832: 131 (Staphylinus; Greece).

  • — Lacordaire, 1835: 381 (Emus; characters; France).

  • — Curtis, 1837: pl. 638 (Quedius; characters; Britain).

  • — Nordmann, 1837: 79 (Philonthus; Russia; Germany; Sweden).

  • — Westwood, 1838a: 16 (Quedius; type species).

  • — Erichson, 1839a: 489 (Quedius; characters; Germany).

  • — Heer, 1839: 274 (Philonthus; characters; Switzerland).

  • — Erichson, 1840: 530 (Quedius; characters; Europe).

  • — Stephens, 1839: 391 (Quedius; characters; England).

  • — Laporte, 1840: 177 (Staphylinus; characters; France).

  • — Gravenhorst, 1847: 234 (Quedius; characters; notes).

  • — Hochhuth, 1849: 151 (Quedius; characters; Tiflis).

  • — Redtenbacher, 1849: 709 (Philonthus; characters; Austria).

  • — Kiesenwetter, 1851a: 419 (Quedius; Pyrenees).

  • — Hardy, 1851: 32 (Quedius; notes; England).

  • — Fairmaire and Laboulbène, 1856: 535 (Quedius; characters; France).

  • — Kraatz, 1857d: 499 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 201 (Quedius; characters; Austria).

  • — Kraatz, 1858a: 58 (Quedius; Greece).

  • — Peyron, 1858: 428 (Quedius; Turkey).

  • — Thomson, 1860: 174 (Microsaurus; characters; Scandinavia).

  • — Fauvel, 1865a: 293 [= 1865b: 43] (Quedius; habitat; France).

  • — Baudi, 1870: 383 (Quedius; characters; Sardinia; Italy).

  • — Crotch, 1870: 100 (Quedius; synonym of cinctus).

  • — Fauvel, 1874: 288 [= 1874a: 514] (Quedius; synonym of cinctus).

  • — Redtenbacher, 1874: 198 (Quedius; characters; Austria).

  • — J. Sahlberg, 1876: 24 (Microsaurus; synonym of cinctus).

  • — Fowler, 1888: 235 (Quedius; synonym of cinctus).

  • — Ganglbauer, 1895: 402 (Quedius; synonym of cinctus).

  • — Porta, 1907: 107 (Quedius; synonym of cinctus).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; synonym of cinctus).

  • — Gridelli, 1924d: 75 (Quedius; synonym of cinctus).

  • — Scheerpeltz, 1933: 1436 (Quedius; synonym of cinctus).

  • — Smetana, 1958: 358 (Quedius; synonym of cinctus).

  • — Smetana, 1971: 165 (Quedius; synonym of cinctus).

  • — Bordoni, 1973b: 49 (Quedius; synonym of cinctus).

  • — Coiffait, 1978: 186 (Quedius; synonym of cinctus).

rufocinctus Mannerheim, 1830: 26 (Staphylinus; Type locality: Finlandia australis; Sweden).

  • — Mannerheim, 1831: 440 (Staphylinus; Finland).

  • — Nordmann, 1837: 79 (Philonthus; Sweden; Finland).

  • — Erichson, 1840: 551 (Quedius; characters; Finland).

  • — Kraatz, 1857d: 501 (Quedius; Finland).

  • — Fauvel, 1874: 288 [= 1874a: 514] (Quedius; synonym of cinctus).

  • — J. Sahlberg, 1876: 25 (Microsaurus; synonym of cinctus).

  • — Ganglbauer, 1895: 402 (Quedius; synonym of cinctus).

  • — Porta, 1907: 107 (Quedius; synonym of cinctus).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; synonym of cinctus).

  • — Gridelli, 1924d: 75 (Quedius; synonym of cinctus).

  • — Smetana, 1958: 358 (Quedius; synonym of cinctus).

  • — Smetana, 1971: 165 (Quedius; synonym of cinctus).

  • — Bordoni, 1973b: 49 (Quedius; synonym of cinctus).

  • — Coiffait, 1978: 187 (Quedius; synonym of cinctus).

niger Porta, 1907: 90, 107, 108 (Quedius; subgenus Quedionuchus; aberration of cinctus; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]).

  • — Gridelli, 1924d: 75 (Quedius; aberration of cinctus).

  • — Scheerpeltz, 1933: 1436 (Quedius; aberration of cinctus).

  • — Smetana, 1958: 358 (Quedius; aberration of cinctus).

  • — Smetana, 1960e: 160 (Quedius; aberration of cinctus; characters).

  • — Scheerpeltz, 1965b: 407 (Quedius; aberration of cinctus; Sicily).

  • — Smetana, 1971: 166 (Quedius; synonym of cinctus).

  • — Bordoni, 1973b: 49 (Quedius; synonym of cinctus).

  • — Coiffait, 1978: 187 (Quedius; subgenus Distichalius; variety of cinctus; characters).

  • — Tóth, 1984: 103 (Quedius; aberration of cinctus).

rufipennis Porta, 1907: 90, 107, 108 (Quedius; subgenus Quedionuchus; aberration of cinctus; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; aberration of cinctus).

  • — Gridelli, 1924d: 75 (Quedius; aberration of cinctus).

  • — Scheerpeltz, 1933: 1436 (Quedius; aberration of cinctus).

  • — Mařan, 1935: 9 (Quedius; variety of cinctus; Crete).

  • — Méquignon, 1948: 76 (Quedius; variety of cinctus; characters; France).

  • — Smetana, 1958: 358 (Quedius; aberration of cinctus).

  • — Smetana, 1960e: 160 (Quedius; aberration of cinctus; characters).

  • — Smetana, 1971: 166 (Quedius; synonym of cinctus).

  • — Bordoni, 1973b: 49 (Quedius; synonym of cinctus).

  • — Coiffait, 1978: 187 (Quedius; subgenus Distichalius; variety of cinctus; characters).

  • — Tóth, 1984: 104 (Quedius; variety of cinctus).

africanus Peyerimhoff, 1917: 123 (Quedius; subgenus Microsaurus; subspecies of flavescens; Type locality: Barbarie).

  • — Gridelli, 1924d: 75 (Quedius; synonym of niger).

  • — Scheerpeltz, 1933: 1436 (Quedius; synonym of cinctus).

  • — Smetana, 1958: 358 (Quedius; synonym of cinctus).

  • — Smetana, 1960e: 160 (Quedius; synonym of cinctus niger).

  • — Smetana, 1971: 166 (Quedius; synonym of cinctus).

  • — Bordoni, 1973b: 49 (Quedius; synonym of cinctus).

  • — Coiffait, 1978: 187 (Quedius; synonym of cinctus variety niger).

lividipennis Lokay, 1921: 71 (Quedius; aberration of cinctus; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]; cited from Candia, Kreta).

  • — Gridelli, 1924d: 75 (Quedius; synonym of rufipennis).

  • — Scheerpeltz, 1933: 1436 (Quedius; synonym of cinctus).

  • — Smetana, 1958: 358 (Quedius; synonym of cinctus).

  • — Smetana, 1960e: 160 (Quedius; synonym of cinctus rufipennis).

  • — Smetana, 1971: 166 (Quedius; synonym of cinctus).

  • — Bordoni, 1973b: 49 (Quedius; synonym of cinctus).

arabs Lokay, 1921: 71 (Quedius; aberration of cinctus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Sidi bel Abbes).

  • — Gridelli, 1924d: 75 (Quedius; synonym of niger).

  • — Scheerpeltz, 1933: 1436 (Quedius; synonym of cinctus).

  • — Smetana, 1958: 358 (Quedius; synonym of cinctus).

  • — Smetana, 1960e: 160 (Quedius; synonym of cinctus niger).

  • — Bordoni, 1973b: 49 (Quedius; synonym of cinctus).

  • — Coiffait, 1978: 187 (Quedius; synonym of cinctus variety niger).

cipactli Smetana, 1976a: 231 (Quedius; subgenus Quedionuchus; Type locality: Guatemala: Sacatepéquez: 10 km. W. San Miguel Duenas, 8000 ft.).

  • Distribution: Guatemala.

circumductus Mulsant and Rey, 1876, see: muscorum Brisout, 1863.

citelli Kirshenblat, 1933: 103 (Quedius; subgenus Microsaurus; Type locality: Transbaikalien: Adun-Tshelon).

  • — Coiffait, 1978: 185 (Quedius; subgenus Microsaurus; cited as citellus; characters; Transbaikal).

  • — Boháč, 1988: 554 [= 1989: 38] (Quedius; cited as citelli; characters).

  • Distribution: Russia.

clypealis Sharp, 1876: 100 (Quedius; Type locality: Ega).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1933: 1436 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 177, 178 (Quedius; subgenus Cyrtoquedius; characters in key; Brazil).

  • Distribution: Brazil.

coatl Smetana, 1976a: 246 (Quedius; subgenus Quedionuchus; Type locality: Mexico: Jalisco: E. slope Nevado de Colima, 10,000 ft.).

  • Distribution: Mexico.

cobosi Coiffait, 1964d: 293 (Quedius; subgenus Quedius; Type locality: Espagne: Province de Almeria, Seron).

  • — Coiffait, 1978: 197 (Quedius; subgenus Quedius; characters; Spain).

  • Distribution: Spain.

almeriensis Coiffait, 1964d: 294 (Quedius; subgenus Quedius; aberration of cobosi; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Espagne: Prov. de Almeria, Laujar).

  • — Coiffait, 1978: 197 (Quedius; subgenus Quedius; variety of cobosi; characters).

coeruleipennis Fauvel, 1874, see: curtus Erichson, 1840.

coeruleipennis Bernhauer, 1926, see: cyaneipennis Scheerpeltz, 1933.

coeruleus Coiffait, 1977, see: gardneri Cameron, 1932.

cohaesus Eppelsheim, 1888: 60 (Quedius; subgenus Sauridus; Type locality: Turcmenien).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Gridelli, 1925: 126 (Quedius; subgenus Raphirus; characters; Turkmenia).

  • — Scheerpeltz, 1933: 1436 (Quedius; catalog).

  • — Coiffait, 1963a: 393 (Quedius; subgenus Sauridus; characters).

  • — Korge, 1964: 122 (Quedius; subgenus Sauridus; Turkey).

  • — Smetana, 1967i: 558 (Quedius; subgenus Raphirus; Turkey).

  • — Coiffait, 1978: 248 (Quedius; subgenus Sauridus; characters; Turkestan; Caucasus; Transcaucasus; Turkey; Bulgaria).

  • Distribution: Turkmenistan, Caucasus Turkey, Bulgaria.

coiffaitianus Fagel, 1968b: 11 (Quedius; subgenus Sauridus; Type locality: Liban: Nabeh Safa, 1000 m).

  • — Coiffait, 1978: 245 (Quedius; subgenus Sauridus; characters; Lebanon).

  • Distribution: Lebanon.

collaris Erichson, 1840: 545 (Quedius; Type locality: Volhynia).

  • — Kraatz, 1857d: 515 (Quedius; characters).

  • — Redtenbacher, 1857: 986 (Quedius; characters; Austria).

  • — G. Waterhouse, 1858: 23 (Quedius; synonym of fulvicollis).

  • — Redtenbacher, 1874: 202 (Quedius; characters).

  • — Mulsant and Rey, 1876: 783 [= 1877: 639] (Quedius; characters; Volhynia).

  • — Ganglbauer, 1895: 411 (Quedius; subgenus Raphirus; characters; Alps; Sudeten; Carpathians).

  • — Porta, 1907: 148 (Quedius; subgenus Raphirus; characters; Italy).

  • — Reitter, 1909: 114 (Quedius; subgenus Raphirus; characters; Germany).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Gridelli, 1925: 151 (Quedius; subgenus Raphirus; characters).

  • — Scheerpeltz, 1933: 1436 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 54 (Quedius; aedeagal illustration).

  • — Smetana, 1958: 388 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 149 (Quedius; subgenus Raphirus; characters; Europe).

  • — Lohse, 1964: 218 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 298 (Quedius; subgenus Raphirus; Czechoslovakia; Poland; Romania; Bosnia Hercegovina; Macedonia; Austria).

  • — Coiffait, 1978: 273 (Quedius; subgenus Raphirus; characters; Carpathians; Alps; Apennines).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • — Tóth, 1984: 126 (Quedius; characters).

  • — Lucht, 1987: 110 (Quedius; checklist; Poland; Czechoslovakia; Austria).

  • — Nowosad, 1990: 149 (Quedius; collected from mole nest; notes; Poland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • Distribution: Europe.

maculicollis W. Kolbe, 1909a: 416 (Quedius; aberration of collaris; [Note: Nomen nudum (Articles 1.3.4, 45.5)]).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; synonym of collaris).

  • — Scheerpeltz, 1933: 1436 (Quedius; aberration of collaris).

  • — Smetana, 1958: 388 (Quedius; aberration of collaris).

  • — Smetana, 1962b: 149 (Quedius; aberration of collaris; characters).

  • — Coiffait, 1978: 273 (Quedius; subgenus Raphirus; variety of collaris; characters).

  • — Tóth, 1984: 126 (Quedius; variety of collaris).

niger Gridelli, 1922: 25 (Quedius; subgenus Raphirus; variety of collaris; [preoccupied]; Type locality: Gran Sasso, Appennino centrale).

  • — Scheerpeltz, 1933: 1436 (Quedius; synonym of collaris).

  • — Coiffait, 1978: 274 (Quedius; synonym of collaris italicus).

italicus Gridelli, 1925: 151 (Quedius; replacement name for niger Gridelli; subspecies of collaris).

  • — Scheerpeltz, 1933: 1436 (Quedius; subspecies of collaris).

  • — Korge, 1964: 124 (Quedius; subgenus Raphirus; characters; Macedonia).

  • — Horion, 1965: 298 (Quedius; subgenus Raphirus; subspecies of collaris; Italy; France).

  • — Coiffait, 1978: 274 (Quedius; subgenus Raphirus; subspecies of collaris; characters).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; subspecies of collaris; Italy).

collinus Broun, 1893: 1031 (Quedius; Type locality: Waitakerei Range).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • Distribution: New Zealand.

coloratus Fauvel, 1875a: XXXIII [= 1875b: 235] (Quedius; Type locality: Palestine Naplouse; [Note: See lectotype designation by Smetana, 1978]. Originally cited from Syrie, Naplouse, Djebel-ech-Cheik).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Gridelli, 1924d: 109 (Quedius; subgenus Raphirus; characters; Syria; Albania).

  • — Scheerpeltz, 1933: 1436 (Quedius; catalog).

  • — Scheerpeltz, 1958a: 23 (Quedius; subgenus Raphirus; Turkey).

  • — Coiffait, 1963a: 378 (Quedius; subgenus Sauridus; characters).

  • — Korge, 1964: 119 (Quedius; subgenus Sauridus; Turkey).

  • — Smetana, 1967i: 558 (Quedius; subgenus Raphirus; Turkey).

  • — Coiffait, 1978: 212 (Quedius; subgenus Sauridus; characters; Balkans; Syria; Lebanon).

  • — Smetana, 1978: 123 (Quedius; subgenus Raphirus; lectotype designation: Palestine Naplouse; collecting notes; Israel).

  • Distribution: Israel, Lebanon, Syria, Turkey, Albania.

coloratus W. Kolbe, 1912 [Note: Nomen nudum], see: boops Gravenhorst, 1802.

columbinus Bernhauer, 1917: 91 (Quedius; subgenus Euryquedius; Type locality: West-Kolumbien: Pacho, in den Ost-Kordilleren, 2000 m; Venezuela).

  • — Scheerpeltz, 1933: 1436 (Quedius; catalog).

  • Distribution: Colombia, Venezuela.

compransor Fall, 1912: 40 (Quedius; Type locality: Manhattan, Kansas).

  • — Casey, 1915: 421 (Anastictodera).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1437 (Quedius; catalog).

  • — Smetana, 1971: 61 (Quedius; subgenus Microsaurus; characters; Kansas).

  • — Smetana, 1978a: 823 (Quedius; subgenus Microsaurus; collecting notes; Florida; Kansas).

  • — Smetana, 1981b: 633 (Quedius; characters; collecting notes; Louisiana).

  • Distribution: USA.

concinnus Last, 1987: 52 (Quedius; Type locality: Papua New Guinea NE: Bulldog Road).

  • Distribution: New Guinea.

concolor Sharp, 1884: 332 (Quedius; Type locality: Guatemala, San Geronimo).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1933: 1437 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 173 (Quedius; subgenus Cyrtoquedius; characters in key; Guatemala).

  • Distribution: Guatemala.

concolor Sharp, 1889, see: samuraicus Bernhauer and Schubert, 1916.

conicus Champion, 1922, see: Indoquedius.

conicus Segers, 1987, see: segersi Herman, 2001.

consors Fauvel, 1866, see: Cheilocolpus.

conspicuellus Broun, 1894: 421 (Quedius; Type locality: Te Aroha).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • Distribution: New Zealand.

constantini Jarrige, 1965a: 19 (Quedius; subgenus Raphirus; Type locality: Espagne: Pena Rubia, au N.-O. de Ponferrada, Province de Lugo, 1700 m).

  • — Coiffait, 1978: 230 (Quedius; subgenus Sauridus; characters; Spain).

  • Distribution: Spain.

cordatus Lea, 1925: 245 (Quedius; Type locality: Queensland: National Park).

  • — Scheerpeltz, 1933: 1437 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

corion Tottenham, 1948a: 258 (Quedius; subgenus Arphirus; Type locality: Haxey).

  • — Smetana, 1958: 397 (Quedius; subgenus Raphirus; characters; ecology).

  • — Fagel, 1960b: 114 (Quedius; notes).

  • — Smetana, 1962b: 152 (Quedius; subgenus Raphirus; characters).

  • — Pope, 1977: 31 (Quedius; possibly synonym of aridulus).

  • — Coiffait, 1978: 277 (Quedius; subgenus Raphirus; characters; England).

  • Distribution: Britain, Czechoslovakia.

coronatus Delabie, 1953: 33 (Quedius; Type locality: versant N. du massif du Canigou Pyrénées Orientales, 1900 m).

  • — Coiffait, 1978: 204 (Quedius; subgenus Quediops; characters; Pyrenees).

  • Distribution: France.

delabiei Coiffait, 1977c: 136 (Quedius; subgenus Quediops; variety of coronatus; [Note: Nomen nudum (Article 15.2)]).

  • — Coiffait, 1978: 204 (Quedius; subgenus Quediops; variety of coronatus; characters).

coxalis Kraatz, 1858a: 59 (Quedius; Type locality: Athen).

  • — Fauvel, 1874: 297 [= 1874a: 523] (Quedius; characters; Italy; Greece; Syria; Palestine).

  • — Mulsant and Rey, 1876: 744 [= 1877: 600] (Quedius; characters; Italy; Greece).

  • — Sainte-Claire Deville, 1906: 102 (Quedius; Corsica).

  • — Porta, 1907: 139 (Quedius; subgenus Sauridus; characters; Italy).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Gridelli, 1922b: 130, 134 (Quedius; characters; notes; Hungary; Transylvania; Italy; Balkan Peninsula; Greece; Syria; Palestine).

  • — Gridelli, 1925: 130 (Quedius; synonym of oblitteratus).

  • — Scheerpeltz, 1933: 1453 (Quedius; synonym of oblitteratus).

  • — Smetana, 1958: 378 (Quedius; synonym of oblitteratus).

  • — Fagel, 1968c: 2 (Quedius; subgenus Sauridus; valid species; characters; notes; Greece; Turkey; Israel; Romania).

  • — Coiffait, 1978: 213 (Quedius; subgenus Sauridus; characters; Greece).

  • — Assing and Wunderle, 2001: 37 (Quedius; Cyprus).

  • Distribution: Europe, Turkey, Syria, Lebanon, Israel.

atticus Coiffait, 1967e: 408 (Quedius; subgenus Sauridus; Type locality: Mont Pantélique près d'Athènes).

  • — Coiffait, 1977c: 140 (Quedius; synonym of coxalis).

  • — Coiffait, 1978: 214 (Quedius; synonym of coxalis).

crassus Fairmaire, 1860b: 153 (Quedius; Type locality: Mers-el-Kébir; Médéah).

  • — Fauvel, 1866c: 247 [= 1866d: 105] (Quedius; synonym of curtus).

  • — Fauvel, 1869: 486 (Quedius; valid species).

  • — Fauvel, 1874: 276 [= 1874a: 502] (Quedius; characters; France; Spain; Algeria).

  • — Mulsant and Rey, 1876: 634 [= 1877: 490] (Quedius; characters; France).

  • — Fauvel, 1878: 127 [= 1878a: 47] (Quedius; Algeria).

  • — Fauvel, 1886: 57 [= 1886a: 49] (Quedius; Tunisia; Algeria).

  • — Ganglbauer, 1895: 397 (Quedius; subgenus Microsaurus; characters; France; Algeria).

  • — Fauvel, 1902b: 114 (Quedius; Algeria; Tunisia).

  • — Porta, 1907: 112 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Gridelli, 1924d: 40 (Quedius; subgenus Microsaurus; characters; France; Spain; Portugal; Algeria).

  • — Portevin, 1929: 339 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1437 (Quedius; catalog).

  • — Roubal, 1941b: 123, 127 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Bordoni, 1974: 72 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Coiffait, 1978: 145 (Quedius; subgenus Microsaurus; characters; western Mediterranean region; North Africa; Iberian Peninsula; Italy; France).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Outerelo, Gamarra, and Salgado, 1998: 129 (Quedius; subgenus Microsaurus; cited as crassus rigidicornis Gridelli, 1924 [Note: I have found no use of rigidicornis by Gridelli]; collected from cave; Spain; Portugal).

  • Distribution: Tunisia, Algeria, Portugal, Spain, France, Italy.

amplicollis Scriba, 1868: 155 (Quedius; Type locality: Spanien).

  • — Fauvel, 1874: 276 [= 1874a: 502] (Quedius; synonym of crassus).

  • — Ganglbauer, 1895: 397 (Quedius; synonym of crassus).

  • — Porta, 1907: 112 (Quedius; synonym of crassus).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; synonym of crassus).

  • — Gridelli, 1924d: 40 (Quedius; synonym of crassus).

  • — Scheerpeltz, 1933: 1437 (Quedius; synonym of crassus).

  • — Bordoni, 1974: 72 (Quedius; synonym of crassus).

  • — Coiffait, 1978: 145 (Quedius; subgenus Microsaurus; subspecies of crassus; characters).

nigricans Gridelli, 1924d: 40 (Quedius; subgenus Microsaurus; variety of crassus; Type locality: Monchique, Portogallo).

  • — Scheerpeltz, 1933: 1437 (Quedius; variety of crassus).

  • — Roubal, 1941b: 123 (Quedius; subgenus Microsaurus; variety of crassus; characters).

  • — Bordoni, 1974: 72 (Quedius; variety of crassus).

  • — Coiffait, 1978: 145 (Quedius; subgenus Microsaurus; variety of crassus; characters).

crescenti Hatch, 1957: 220 (Quedius; subgenus Microsaurus; Type locality: Washington: L. Crescent. Also cited from Oregon).

  • — Smetana, 1971: 100 (Quedius; subgenus Microsaurus; characters; Oregon; Washington).

  • — Smetana, 1976c: 170 (Quedius; subgenus Microsaurus; British Columbia).

  • — Smetana, 1981b: 634 (Quedius; collecting notes; Oregon).

  • Distribution: Canada, USA.

creticus Mařan, 1935, see: fulgidus Fabricius, 1793.

criddlei Casey, 1915: 410 (Microsaurus; Type locality: Manitoba: Aweme).

  • — Scheerpeltz, 1933: 1437 (Quedius; catalog).

  • — Smetana, 1971: 107 (Quedius; subgenus Microsaurus; characters; Alberta; British Columbia; Manitoba; Northwest Territories; Yukon Territories; Colorado; Idaho; Montana; Oregon; Utah; Washington; Wyoming).

  • — Smetana, 1978a: 825 (Quedius; subgenus Microsaurus; collecting notes; Washington).

  • — Smetana, 1981b: 636 (Quedius; collecting notes; Alaska; Saskatchewan).

  • — Smetana, 1990d: 97 (Quedius; Alaska; Ontario).

  • Distribution: Canada, USA.

hoodi Hatch, 1957: 221 (Quedius; subgenus Microsaurus; Type locality: Oregon: Mt. Hood, Timberline. Also cited from Washington, Idaho, and British Columbia).

  • — Smetana, 1971: 107 (Quedius; synonym of criddlei).

crius Tottenham, 1948, see: reitteri Gridelli, 1925.

crnagoricus Coiffait, 1980a: 38 (Quedius; subgenus Raphirus; Type locality: Monténégro, Col de Cakor, 1850 m).

  • — Drugmand, 1988c: 204 (Quedius; characters).

  • Distribution: Yugoslavia.

cruentus Olivier, 1795:(42): 27 (Staphylinus; Type locality: Paris).

  • — Gravenhorst, 1802: 25 (Staphylinus; synonym of fulvipes).

  • — Marsham, 1802: 503 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1916: 421 listed fulgidus Marsham as a synonym of cruentus; Marsham did not describe the species as new, he attributed it to Fabricius]; characters; Britain).

  • — Gyllenhal, 1810: 343 (Staphylinus; synonym of fulvipes).

  • — C. Sahlberg, 1830: 329 (Staphylinus; synonym of fulvipes).

  • — Erichson, 1840: 527 (Quedius; characters; Germany; France; Italy; Volhynia).

  • — Hochhuth, 1849: 150 (Quedius; characters; Iberia).

  • — Redtenbacher, 1849: 708 (Philonthus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 540 (Quedius; characters; France).

  • — Kraatz, 1857d: 495 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 200 (Quedius; characters; Austria).

  • — Kraatz, 1858a: 58 (Quedius; Greece).

  • — Kraatz, 1859: 67 (Quedius; characters; India orientalis).

  • — Thomson, 1867: 162 (Microsaurus; characters; Scandinavia).

  • — Fauvel, 1874: 282 [= 1874a: 508] (Quedius; characters; France; Caucasus; East Indies).

  • — Redtenbacher, 1874: 198 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 266 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 661 [= 1877: 517] (Quedius; characters; France).

  • — J. Sahlberg, 1876: 25 (Microsaurus; Finland).

  • — Fauvel, 1878: 127 [= 1878a: 47] (Quedius; Algeria).

  • — Fauvel, 1886: 57 [= 1886a: 49] (Quedius; notes; Algeria).

  • — Fowler, 1888: 234 (Quedius; characters; Britain).

  • — Ganglbauer, 1895: 399 (Quedius; subgenus Microsaurus; characters; Europe; Caucasus; East Indies).

  • — Fauvel, 1902b: 114 (Quedius; Morocco; Algeria; Tunisia).

  • — Porta, 1907: 115 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Sainte-Claire Deville, 1906: 101 (Quedius; Corsica).

  • — Reitter, 1909: 111 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Johansen, 1914: 354 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Munster, 1923: 192 (Quedius; subgenus Microsaurus; Fennoscandia).

  • — Gridelli, 1924d: 53 (Quedius; subgenus Microsaurus; characters; Belgium; Styria; Italy; Albania; Dalmatia).

  • — Portevin, 1929: 341 (Quedius; characters; France).

  • — Cameron, 1932: 289 (Quedius; subgenus Microsaurus; characters; India).

  • — Scheerpeltz, 1933: 1437 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 10 (Quedius; aedeagal illustration).

  • — Roubal, 1941b: 123, 128 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Jeannel and Jarrige, 1949: 378 (Quedius; subgenus Microsaurus; cave species; France).

  • — V. Hansen, 1952: 135 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Smetana, 1958: 350 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 157 (Quedius; subgenus Microsaurus; characters).

  • — Scheerpeltz, 1961d: 132 (Quedius; subgenus Microsaurus; Italy).

  • — Palm, 1963b: 135 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 209 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Horion, 1965: 263 (Quedius; subgenus Microsaurus; Morocco; Algeria; Turkey; all southern and central Europe; Denmark; Britain; Ireland; Norway; Sweden; Finland; Russia).

  • — Szujecki, 1968a: 734 (Quedius; Poland).

  • — Bordoni, 1974: 40 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Legner and Moore, 1977: 176 (Quedius; found under bark).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 152 (Quedius; subgenus Microsaurus; characters; Europe; Britain; Scandinavia; North Africa; Asia Minor; India).

  • — Hinton, 1981: 662 (Quedius; characters of egg).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • — Tóth, 1984: 113 (Quedius; characters; Hungary).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Welch, 1993: 229 (Quedius; ovariole number and ovary structure).

  • — Smetana, 1993b: 49 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Owen, 1999b: 61 (Quedius; captured underground at roots of old trees; Britain).

  • — Owen, 2000: 251 (Quedius; pitfall trap collections from underground at roots of trees; Britain).

  • Distribution: Tunisia, Algeria, Morocco, Europe, Russia, Caucasus, Turkey, India, “East Indies”.

haemorrhous Stephens, 1832: 217 (Quedius; Type locality: London; Norfolk; Suffolk).

  • — Curtis, 1829: 25 (Quedius; nomen nudum; catalog; Britain).

  • — Curtis, 1837: pl. 638 (Quedius; characters; Britain).

  • — Stephens, 1839: 390 (Quedius; variety of nitidus; characters; England).

  • — Hardy, 1851: 32 (Quedius; synonym of fulgidus).

  • — G. Waterhouse, 1858: 22 (Quedius; synonym of cruentus).

  • — Gemminger and Harold, 1868: 568 (Quedius; synonym of cruentus).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; synonym of cruentus).

  • — Cameron, 1932: 289 (Quedius; synonym of cruentus).

  • — Smetana, 1958: 350 (Quedius; synonym of cruentus).

  • — Bordoni, 1974: 41 (Quedius; synonym of cruentus).

  • — Coiffait, 1978: 152 (Quedius; synonym of cruentus).

erythropterus Stephens, 1832: 226 (Quedius; Type locality: London).

  • — Stephens, 1839: 393 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 22 (Quedius; synonym of cruentus).

  • — Gemminger and Harold, 1868: 568 (Quedius; synonym of cruentus).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; synonym of cruentus).

  • — Cameron, 1932: 289 (Quedius; synonym of cruentus).

  • — Smetana, 1958: 350 (Quedius; synonym of cruentus).

  • — Bordoni, 1974: 41 (Quedius; synonym of cruentus).

  • — Coiffait, 1978: 152 (Quedius; synonym of cruentus).

analis Stephens, 1835: 435 (Quedius; [preoccupied]; Type locality: London; Yorkshire).

  • — Curtis, 1837: pl. 638 (Quedius; characters; Britain).

  • — Stephens, 1839: 391 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 22 (Quedius; synonym of cruentus).

  • — Gemminger and Harold, 1868: 568 (Quedius; synonym of cruentus).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; synonym of cruentus).

  • — Cameron, 1932: 289 (Quedius; synonym of cruentus).

  • — Smetana, 1958: 350 (Quedius; synonym of cruentus).

  • — Bordoni, 1974: 41 (Quedius; synonym of cruentus).

  • — Coiffait, 1978: 152 (Quedius; synonym of cruentus).

virens Rottenberg, 1870: 28 (Quedius; variety of fulgidus; Type locality: Sicily).

  • — Fauvel, 1874: [= 1874a:] (Quedius; race of cruentus; characters; Scandinavia; Russia; Tyrol; Italy; Corsica; Sardinia; Sicily).

  • — Duvivier, 1883: 138 (Quedius; catalog; synonym of cruentus).

  • — Ganglbauer, 1895: 399 (Quedius; variety of cruentus).

  • — Porta, 1907: 115 (Quedius; variety of cruentus).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; variety of cruentus).

  • — Gridelli, 1924d: 53 (Quedius; subgenus Microsaurus; variety of cruentus; Styria; Italy; Croatia; Dalmatia).

  • — Scheerpeltz, 1933: 1437 (Quedius; aberration of cruentus).

  • — Roubal, 1941b: 123 (Quedius; subgenus Microsaurus; aberration of cruentus; characters).

  • — Smetana, 1958: 350 (Quedius; aberration of cruentus).

  • — Smetana, 1960e: 157 (Quedius; aberration of cruentus; characters).

  • — Bordoni, 1974: 41 (Quedius; aberration of cruentus).

  • — Pope, 1977: 31 (Quedius; synonym of cruentus).

  • — Coiffait, 1978: 152 (Quedius; subgenus Microsaurus; variety of cruentus; characters).

  • — Tóth, 1984: 114 (Quedius; aberration of cruentus; characters; Hungary).

obscurus Lokay, 1913: 138, 140 (Quedius; variety of cruentus; Type locality: Kreta).

  • — Scheerpeltz, 1933: 1437 (Quedius; variety of cruentus).

  • — Mařan, 1935: 8 (Quedius; variety of cruentus; Crete).

  • — Roubal, 1941b: 123 (Quedius; subgenus Microsaurus; aberration of cruentus; characters).

  • — Smetana, 1958: 350 (Quedius; aberration of cruentus).

  • — Smetana, 1960e: 157 (Quedius; aberration of cruentus; characters).

  • — Scheerpeltz, 1961d: 133 (Quedius; subgenus Microsaurus; aberration of cruentus; Italy).

  • — Bordoni, 1974: 41 (Quedius; aberration of cruentus).

  • — Coiffait, 1978: 152 (Quedius; subgenus Microsaurus; variety of cruentus; characters).

  • — Tóth, 1984: 114 (Quedius; aberration of cruentus).

cuprinus Fauvel, 1877: 274 (Quedius; Type locality: Australie, Nouvelle Galles du Sud, Sydney).

  • — Lea, 1912: 40 (Quedius; with Camponotus nigriceps; Victoria).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Scheerpeltz, 1935a: 655 (Quedius; Australia).

  • Distribution: Australia.

cursor Smetana, 1971: 99 (Quedius; subgenus Microsaurus; Type locality: California: Luther Pass).

  • — Smetana, 1971a: 1839 (Quedius; subgenus Microsaurus; collecting notes).

  • Distribution: USA.

curtidens Smetana, 1967i: 559 (Quedius; subgenus Raphirus; Type locality: Türkei: Suluhan, Toros).

  • — Coiffait, 1978: 279 (Quedius; subgenus Raphirus; characters; Turkey).

  • Distribution: Turkey.

curtipennis Bernhauer, 1908e: 335 (Quedius; form of fuliginosus; Type locality: Buchara; Böhmen, Wran a. Moldau).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; variety of fuliginosus).

  • — Gridelli, 1924d: 79 (Quedius; variety of fuliginosus).

  • — Scheerpeltz, 1933: 1441 (Quedius; variety of fuliginosus).

  • — V. Hansen, 1952: 141 (Quedius; subgenus Quedius; characters; Denmark).

  • — Smetana, 1958: 362 (Quedius; subgenus Quedius; characters; ecology; Czechoslovakia).

  • — Korge, 1962b: 332 (Quedius; subgenus Quedius; synonymic notes).

  • — Smetana, 1962b: 133 (Quedius; subgenus Quedius; characters; Czechoslovakia).

  • — Palm, 1963b: 141 (Quedius; subgenus Quedius; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 211 (Quedius; subgenus Quedius; characters; central Europe).

  • — Korge, 1964: 119 (Quedius; subgenus Quedius; Turkey).

  • — Smetana, 1965: 38 (Quedius; subgenus Quedius; characters).

  • — Smetana, 1965a: 12 (Quedius; British Columbia).

  • — M. Dvořák, 1965: 89 (Quedius; Slovakia).

  • — Horion, 1965: 272 (Quedius; subgenus Quedius; Turkey; Faroes; Britain; Norway; Sweden; Finland; Russia; Denmark; all central Europe; Spain; Azores; Corsica; Italy; Bosnia; Romania).

  • — Szujecki, 1968a: 735 (Quedius; Poland).

  • — Frank, 1969: 267 (Quedius; biological notes; England).

  • — Smetana, 1970: 62 (Quedius; distributional summary; Azores).

  • — Smetana, 1971: 129 (Quedius; subgenus Quedius; characters; British Columbia; Oregon; Washington).

  • — Smetana, 1971b: 3 (Quedius; subgenus Quedius; British Columbia).

  • — Coiffait, 1973a: 275 (Quedius; Morocco).

  • — Bordoni, 1974b: 10 (Quedius; subgenus Quedius; characters; Italy).

  • — Bordoni, 1976b: 92 (Quedius; characters; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 191 (Quedius; subgenus Quedius; characters; Europe; Britain).

  • — Outerelo, 1978a: 281 (Quedius; pupal characters; Spain).

  • — Smetana, 1978a: 826 (Quedius; subgenus Quedius; collecting notes; Oregon).

  • — Tóth, 1984: 118 (Quedius; characters; Hungary).

  • — Drugmand, 1987a: 312 (Quedius; Belgium).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Nowosad, 1990: 145 (Quedius; collected from mole nest; notes; Poland).

  • — Borges, 1990: Table II (Quedius; Azores).

  • — Smetana, 1990d: 98 (Quedius; New Hampshire).

  • — Smetana, 1993b: 50 (Quedius; subgenus Quedius; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Owen, 1997a: 149 (Quedius; collecting sites; Ireland).

  • — Outerelo, Gamarra, and Salgado, 1998: 129 (Quedius; subgenus Quedius; collected from cave; Spain; Portugal).

  • — Assing, 2001: 75 (Quedius; collected from xerothermous site; Germany).

  • Distribution: Morocco, Azores, Europe, Turkey, Uzbekistan, Canada, USA.

parallelus Hatch, 1957: 216 (Quedius; subgenus Quedius; Type locality: Washington: Seattle, U[niversity of] W[ashington] Campus. Also cited from British Columbia).

  • — Korge, 1962b: 332 (Quedius; synonym of curtipennis).

  • — Smetana, 1965a: 12 (Quedius; synonym of curtipennis).

  • — Smetana, 1971: 130 (Quedius; synonym of curtipennis).

curtipennis Casey, 1915, see: uteanus Casey, 1915.

curtus Erichson, 1840: 534 (Quedius; Type locality: Sardinia).

  • — Fairmaire and Laboulbène, 1856: 535 (Quedius; characters; France).

  • — Redtenbacher, 1857: 200 (Quedius; characters; Austria).

  • — Fauvel, 1874: 274 [= 1874a: 500] (Quedius; characters; France; Italy; Sicily; Sardinia).

  • — Redtenbacher, 1874: 197 (Quedius; characters; Austria).

  • — Fauvel, 1876a: 76 [= 1876: 261] (Quedius; Corsica).

  • — Mulsant and Rey, 1876: 623 [= 1877: 479] (Quedius; characters; France).

  • — Ganglbauer, 1895: 397 (Quedius; subgenus Microsaurus; characters; France; Italy; Sardinia; Sicily).

  • — Porta, 1907: 110 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Sainte-Claire Deville, 1906: 101 (Quedius; Corsica).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Gridelli, 1924d: 17 (Quedius; subgenus Euryquedius; characters; Italy).

  • — Portevin, 1929: 337 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1437 (Quedius; catalog).

  • — Bordoni, 1973b: 43 (Quedius; subgenus Euryquedius; characters; Italy).

  • — Coiffait, 1978: 143 (Quedius; subgenus Euryquedius; characters; western Mediterranean; France; Corsica).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: France, Italy.

coeruleipennis Fauvel, 1874: 275 [= 1874a: 501] (Quedius; race of curtus; Type locality: Mont Rosa; Dijon; Hautes-Alpes; rochers de Royan; Dordogne; Nyons; Toulon; St Maximin; Provence, Languedoc; Gênes).

  • — Duvivier, 1883: 138 (Quedius; catalog; variety of curtus).

  • — Ganglbauer, 1895: 388 (Quedius; variety of curtus).

  • — Porta, 1907: 110 (Quedius; variety of curtus).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; variety of curtus).

  • — Scheerpeltz, 1933: 1438 (Quedius; variety of curtus).

  • — Bordoni, 1973b: 43 (Quedius; variety of curtus).

  • — Coiffait, 1978: 143 (Quedius; variety of curtus; characters).

cyaneipennis Scheerpeltz, 1933: 1438 (Quedius; replacement name for coeruleipennis Bernhauer).

  • Distribution: Indonesia.

coeruleipennis Bernhauer, 1926f: 319 (Quedius; [preoccupied]; Type locality: Buru).

  • — Scheerpeltz, 1933: 1438 (Quedius; synonym of cyaneipennis).

cyanellus Fauvel, 1878f: 274 (Quedius; Type locality: Ternate, Acquiconora).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • Distribution: Indonesia.

cyaneorufus Fauvel, 1878f: 274 (Quedius; Type locality: Iles Arou, Wokan).

  • — Fauvel, 1879: 107 (Quedius; Dorei; Amberbaki).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Cameron, 1931: 363 (Quedius; New Guinea).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Cameron, 1937a: 104 (Quedius; New Guinea).

  • — Last, 1975: 439 (Quedius; subgenus Sauridus; New Guinea).

  • — Last, 1987: 54 (Quedius; Papua New Guinea; Irian).

  • Distribution: Indonesia, New Guinea.

gazellensis Last, 1987: 54 (Quedius; variety of cyaneorufus; [Note: Nomen nudum (Article 15.2)]; cited from Papua New Guinea: New Britain, Gazelle Pen., Mt. Sinewit, 900 m; Morobe Dist., Wau, 1200–1300 m; Iongai, 10 km E Mt. Albert Edward, 1450 m; Umboi I., ca. 8 km WNW Lablab, 300 m; New Ireland, ridge above “Camp Bishop”, 15 km up Kait R., 250–750 m).

cyanescens Mulsant and Rey, 1876: 727 [= 1877: 583] (Quedius; subgenus Sauridus; Type locality: environs d'Hyères).

  • — Ganglbauer, 1895: 406 (Quedius; subgenus Sauridus; characters; France).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; variety of umbrinus).

  • — Gridelli, 1925: 113 (Quedius; variety of umbrinus).

  • — Scheerpeltz, 1933: 1464 (Quedius; variety of umbrinus).

  • — Coiffait, 1963a: 385 (Quedius; subgenus Sauridus; valid species; characters).

  • — Coiffait, 1963b: 423 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 237 (Quedius; subgenus Sauridus; characters; Pyrenees; Mediterranean region of western Europe).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: France.

cyanopterus Scheerpeltz, 1976, see: apicicornis Eppelsheim, 1895.

cyprinus H. Franz, 1987, see: scheerpeltzi Gridelli, 1938.

cyprusensis Last, 1955: 251 (Quedius; Type locality: Cyprus: Mount Troodus, 5600–6000 ft.).

  • — Coiffait, 1963a: 385 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1970e: 62 (Quedius; Cyprus).

  • — Coiffait, 1978: 236 (Quedius; subgenus Sauridus; characters; Cyprus; Lebanon).

  • — Bordoni, 1984: 342 (Quedius; Lebanon).

  • — Assing and Wunderle, 2001: 37 (Quedius; Cyprus).

  • Distribution: Cyprus, Lebanon.

freyi Scheerpeltz, 1956c: 1102 (Quedius; subgenus Raphirus; Type locality: Cypern: Pedhoulas, 3600 feet).

  • — Coiffait, 1978: 236 (Quedius; synonym of cyprusensis).

daibosatsunis Watanabe, 1986: 19 (Quedius; subgenus Microsaurus; Type locality: Japan: Honshu: Yamanashi Pref.: Hikawa Valley, 1400 m alt., Enzan-shi).

  • Distribution: Japan.

daksumensis Coiffait, 1982a: 279 (Quedius; subgenus Raphirus; Type locality: Cachemire: Daksum, 2400–2700 m).

  • — Smetana, 1988: 247 (Quedius; subgenus Raphirus; characters; India; Nepal).

  • — Smetana, 1992b: 4 (Quedius; subgenus Raphirus; Nepal).

  • Distribution: India, Nepal.

dalmatinus Korge, 1960, see: meridiocarpathicus Smetana, 1958.

debilis Horn, 1878: 165 (Quedius; Type locality: California; [Note: See lectotype designation by Smetana, 1971]. Originally cited from Clear Lake, California and Vancouver).

  • — Casey, 1915: 403 (Quediellus; characters; California).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Smetana, 1971: 215 (Quedius; subgenus Raphirus; lectotype designtion: Cala [California]; characters; California).

  • — Smetana, 1971a: 1848 (Quedius; subgenus Raphirus; collecting notes; California).

  • Distribution: USA.

deceptor Cameron, 1944: 14 (Quedius; subgenus Sauridus; Type locality: Darjeeling: Ghum district).

  • — Smetana, 1988: 231 (Quedius; subgenus Distichalius; characters; Uttar Pradesh; West Bengal).

  • Distribution: India.

decipiens Cameron, 1944, see: aureiventris Bernhauer, 1915.

decius Smetana, 1996: 12 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan prov., Liziping env. near Shimien, 200 km SW of Ya'an).

  • — Smetana, 1999b: 215 (Quedius; subgenus Microsaurus; notes; China).

  • Distribution: China.

declivus Fauvel, 1878: 128 [= 1878a: 48] (Quedius; Type locality: Bône).

  • — Fauvel, 1886: 58 [= 1886a: 50] (Quedius; notes; Algeria).

  • — Fauvel, 1902b: 116 (Quedius; Algeria; Tunisia).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Gridelli, 1924d: 100 (Quedius; subgenus Raphirus; characters; Tunisia; Algeria).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Jeannel and Jarrige, 1949: 378 (Quedius; subgenus Raphirus; cave species; Algeria).

  • — Coiffait, 1963a: 375 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 206 (Quedius; subgenus Sauridus; characters; Algeria; Tunisia).

  • — Maachi, 1991: 251 (Quedius; Morocco).

  • Distribution: Morocco, Algeria, Tunisia.

decolor Coiffait, 1976 [Note: Nomen nudum], see: fissus Gridelli, 1938.

decoratus Petri 1902, see: haberfelneri Eppelsheim, 1891.

decoui Coiffait, 1963a: 381, 406 (Quedius; subgenus Sauridus; Type locality: Roumanie, Olténie).

  • — Coiffait, 1978: 221 (Quedius; subgenus Sauridus; characters; Romania).

  • — Tóth, 1984: 135 (Quedius; characters).

  • Distribution: Romania.

deficiens Cameron, 1945, see: quadripunctus Bernhauer, 1941.

delabiei Coiffait, 1977 [Note: Nomen nudum], see: coronatus Delabie, 1953.

delahoni H. Wagner, 1916 [Note: Nomen nudum], see: mesomelinus Marsham, 1802.

densiventris Casey, 1915: 404 (Quediellus; Type locality: Montana: Mullan).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Smetana, 1971: 199 (Quedius; subgenus Raphirus; characters; Alberta; British Columbia; Newfoundland; Yukon Territory; Alaska; Colorado; Idaho; New Hampshire; New Mexico; New York; Oregon; Utah; Washington; Wyoming).

  • — Smetana, 1971a: 1846 (Quedius; subgenus Raphirus; collecting notes; New Mexico; California).

  • — Smetana, 1973: 1429 (Quedius; notes; Quebec).

  • — Smetana, 1976c: 181 (Quedius; subgenus Raphirus; Colorado; New Mexico).

  • — Smetana, 1976d: 26 (Quedius; characters).

  • — Smetana, 1978a: 830 (Quedius; subgenus Raphirus; collecting notes; Arizona; Utah).

  • — Downie and Arnett, 1996: 393 (Quedius; subgenus Raphirus; characters; USA).

  • Distribution: USA, Canada.

alutaceus Hatch, 1957: 219 (Quedius; subgenus Distichalius; Type locality: Washington: Mt. Rainier, Sunrise Park. Also cited from British Columbia).

  • — Smetana, 1971: 199 (Quedius; synonym of densiventris).

bonapartei Hatch, 1957: 226 (Quedius; subgenus Sauridus; Type locality: Washington: Mt. Bonaparte. Also cited from Idaho, Oregon, and British Columbia).

  • — Smetana, 1965: 46 (Quedius; subgenus Raphirus; characters; Newfoundland).

  • — Smetana, 1965a: 14 (Quedius; subgenus Raphirus; British Columbia).

  • — Smetana, 1971: 199 (Quedius; synonym of densiventris).

denudatus Stephens, 1832, see: molochinus Gravenhorst, 1806.

depauperatus Wollaston, 1871: 299 (Quedius; variety of fulgidus; Type locality: Teneriffe).

  • — Wollaston, 1864: 563 (Quedius; [Note: Misidentification: Bernhauer and Schubert, 1916: 422, cited fulgidus Wollaston as a synonym of depauperatus; Wollaston did not describe the species as new, he attributed it to Fabricius]; notes; habitat; Canary Islands).

  • — Fauvel, 1876a: 76 [= 1876: 261] (Quedius; synonym of fulgidus).

  • — Ganglbauer, 1895: 399 (Quedius; variety of fulgidus).

  • — Fauvel, 1902b: 115s (Quedius; Canaries).

  • — Porta, 1907: 114 (Quedius; variety of fulgidus).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Gridelli, 1924d: 64 (Quedius; subgenus Microsaurus; characters; Canaries).

  • — Gridelli, 1929: 39 (Quedius; characters; Canaries).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Coiffait, 1978: 155 (Quedius; subgenus Microsaurus; characters; Canary Islands).

  • — Assing and Wunderle, 1999: 435 (Quedius; collecting notes; Canary Islands).

  • — Assing and Wunderle, 1999: 437 (Quedius; Canary Islands).

  • Distribution: Canary Islands.

robustus Wollaston, 1871: 299 (Quedius; variety of fulgidus; [preoccupied]; Type locality: Gomera).

  • — Fauvel, 1876a: 76 [= 1876: 261] (Quedius; synonym of maurus).

  • — Eppelsheim, 1878b: 175 (Quedius; [Note: Scheerpeltz, 1933: 1438, cited robustus Eppelsheim and robustus Wollaston as synonyms of depauperatus; Eppelsheim attributed the name to Wollaston]).

  • — Ganglbauer, 1895: 400 (Quedius; variety of maurus).

  • — Porta, 1907: 125 (Quedius; synonym of maurus).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; synonym of depauperatus).

  • — Gridelli, 1929: 39 (Quedius; synonym of depauperatus).

  • — Scheerpeltz, 1933: 1438 (Quedius; synonym of depauperatus).

  • — Coiffait, 1978: 155 (Quedius; synonym of depauperatus).

deperditus† Scudder, 1900a: 82 (Quedius; pleistocene fossil; Type locality: Scarborough).

  • Distribution: Canada.

deschutesi Hatch, 1957: 220 (Quedius; subgenus Microsaurus; Type locality: Oregon: Deschutes Co.: Camp Abbott).

  • — Smetana, 1971: 102 (Quedius; subgenus Microsaurus; characters; Oregon).

  • — Smetana, 1981b: 634 (Quedius; characters; collecting notes; Arizona).

  • Distribution: USA.

desertus Horn, 1878: 161 (Quedius; Type locality: Pinal Mts., Arizona; [Note: See lectotype designation by Smetana, 1971]. Originally cited from Colorado River in California; Arizona; southwestern Utah).

  • — Fall and Cockerell, 1907: 166 (Quedius; New Mexico).

  • — Casey, 1915: 412 (Microsaurus; characters; Arizona).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Smetana, 1971: 123 (Quedius; subgenus Microsaurus; nominate subspecies; lectotype designation: Pinal Mts., Ariz.; characters; Arizona; Colorado; New Mexico; Utah).

  • — Smetana, 1971a: 1840 (Quedius; subgenus Microsaurus; collecting notes; New Mexico; Arizona).

  • — Smetana, 1973: 1427 (Quedius; Texas).

  • Distribution: USA.

fontinalis Casey, 1915: 414 (Microsaurus; Type locality: New Mexico: Jemez Springs).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Smetana, 1971: 123 (Quedius; synonym of desertus).

pinalicus Casey, 1915: 413 (Microsaurus; Type locality: Arizona: Pinal Mts.; [Note: See lectotype designation by Smetana, 1971]).

  • — Scheerpeltz, 1933: 1458 (Quedius; catalog).

  • — Smetana, 1971: 123 (Quedius; lectotype designation; synonym of desertus).

californicus Smetana, 1971: 125 (Quedius; subgenus Microsaurus; subspecies of desertus; Type locality: California: Volcan Mt., San Diego Co.).

exulans Smetana, 1971: 125 (Quedius; subgenus Microsaurus; subspecies of desertus; Type locality: California: Niles Cñ., Alameda Co.).

  • — Smetana, 1978a: 825 (Quedius; subgenus Microsaurus; subspecies of desertus; collecting notes; California).

deuvei Coiffait, 1978d: 170 (Quedius; subgenus Sauridus; Type locality: Drannaz Dag).

  • — Coiffait, 1978: 226 (Quedius; subgenus Sauridus; characters; Anatolia).

  • — Coiffait, 1984e: 374 (Quedius; Nepal).

  • Distribution: Turkey, Nepal ?.

deuvei Coiffait, 1983, see: kashmirensis Cameron, 1944.

deuveianus Coiffait, 1983, see: kashmirensis Cameron, 1944.

dewar Smetana, 1988: 265 (Quedius; subgenus Raphirus; Type locality: Nepal: Prov. Bagmati: Mere Dara, 3200 m).

  • Distribution: Nepal.

dhaulagirensis Coiffait, 1982, see: kanyasa Smetana, 1975.

dichrous Fauvel, 1878g: 553 (Quedius; Type locality: Australie méridionale: Victoria, Melbourne).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • Distribution: Australia.

diemenensis Blackburn, 1894a: 91 (Quedius; Type locality: Tasmania).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Lea, 1925: 242 (Quedius; characters; Australia).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

dih Smetana, 1996g: 24 (Quedius; subgenus Raphirus; Type locality: Taiwan: Ilan Hsien: Taipingshan, 1820 m).

  • Distribution: Taiwan.

discrepans Broun, 1893: 1030 (Quedius; Type locality: Wellington).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • Distribution: New Zealand.

dispersepunctatus Scheerpeltz, 1965: 293 (Quedius; subgenus Indoquedius; Type locality: N.E. Burma, Kambaiti, 7000 ft.).

  • Distribution: Myanmar.

distans Smetana, 1971: 91 (Quedius; subgenus Microsaurus; Type locality: California: Haypress Mdws., Eldorado Co. Also cited from Nevada).

  • — Smetana, 1971a: 1839 (Quedius; subgenus Microsaurus; collecting notes).

  • — Smetana, 1971b: 2 (Quedius; subgenus Microsaurus; California).

  • — Smetana, 1973: 1426 (Quedius; Nevada).

  • — Smetana, 1976c: 170 (Quedius; subgenus Microsaurus; California).

  • Distribution: USA.

distincticolor Roubal, 1914 [Note: Nomen nudum], see: minor Hochhuth, 1849.

distinctus Runde, 1835, see: fulgidus Fabricius, 1793.

divergens Casey, 1915, see: utenaus Casey, 1915.

diversicollis Cameron, 1945d: 789 (Quedius; subgenus Sauridus; Type locality: New Zealand: Reefton).

  • Distribution: New Zealand.

diversipennis Fauvel, 1878g: 554 (Quedius; Type locality: Australie occidentale, Swan River).

  • — Bernhauer, 1908: 19 (Quedius; Australia).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Bernhauer, 1916a: 7 (Quedius; Australia).

  • Distribution: Australia.

doderoi Gridelli, 1922: 22 (Quedius; subgenus Microsaurus; Type locality: Monte Misurasca [Apennino ligure]).

  • — Gridelli, 1924d: 32 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Bordoni, 1974: 18 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Coiffait, 1978: 182 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Italy.

dohertyi Cameron, 1932, see: muscicola Cameron, 1932.

dolosus Coiffait, 1961 [Note: Nomen nudum], see: unicolor Kiesenwetter, 1847.

dorsalis Stephens, 1835: 436 (Quedius; [Note: Nomen dubium]; Type locality: Coombe Wood).

  • Distribution: Britain.

dorni Korge, 1969, see: ochropterus Erichson, 1840.

douglasi Bernhauer, 1929: 2 (Quedius; Type locality: Nord-China: Shensi, Yung-ting-Shan Mts, 90 m. W of Taiyan-fu, 7000 ft.).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Smetana, 1996f: 113 (Quedius; subgenus Microsaurus; characters; China).

  • Distribution: China.

downiei Hatch, 1957, see: montanicus Casey, 1915.

drannazensis Coiffait, 1978d: 171 (Quedius; subgenus Sauridus; Type locality: Drannaz Dag).

  • — Coiffait, 1978: 252 (Quedius; subgenus Sauridus; characters; Anatolia).

  • Distribution: Turkey.

dryadum Peyerimhoff, 1917: 121 (Quedius; subgenus Microsaurus; Type locality: Boucharen, au sud de Larache, Maroc occidental; Massif des Mouzaia, Alger, 1200 m).

  • — Gridelli, 1924d: 46 (Quedius; subgenus Microsaurus; characters; Algeria).

  • — Gridelli, 1925: 170 (Quedius; characters).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog; Mauritania).

  • — Coiffait, 1978: 172 (Quedius; subgenus Microsaurus; characters; North Africa).

  • Distribution: Mauritania, Morocco, Algeria.

dubius Heer, 1839: 278 (Philonthus; variety b of montanus; [Note: Preoccupied as a secondary homonym in Philonthus; Heer presented characters for montanus variety b; following those characters he equated the variety with Staphylinus dubius Heer. I take the characterization and link established between variety b and dubius to be sufficient to validate the name which Heer first published in 1834 as an unavailable name]; Type locality: Paradies).

  • — Heer, 1834a: 156 (Staphylinus; subgenus Microsaurus; [Note: Unavailable name]; elevational listing).

  • — Gemminger and Harold, 1868: 569 (Quedius; synonym of fimbriatus).

  • — Fauvel, 1874: 293 [= 1874a: 519] (Quedius; characters; France; Switzerland; Italy; Austria).

  • — Mulsant and Rey, 1876: 719 [= 1877: 575] (Quedius; subgenus Sauridus; characters; France).

  • — Ganglbauer, 1895: 405 (Quedius; subgenus Sauridus; characters; Germany; Alps; Carpathians).

  • — Porta, 1907: 135 (Quedius; subgenus Sauridus; characters; Italy).

  • — Reitter, 1909: 112 (Quedius; subgenus Sauridus; characters; Germany).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Gridelli, 1924d: 98 (Quedius; subgenus Raphirus; characters; Italy).

  • — Portevin, 1929: 342 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1438 (Quedius; [Note: Scheerpeltz, 1933: 1439 cited rubromaculatus Gridelli, 1924: 99 as an aberration of dubius. Gridelli, in the reference cited by Scheerpeltz, described an aberration but did not propose a new name. Gridelli began the description with “ab. elytris rubromaculatis” and he did not cite the name in the index to his article]; catalog).

  • — Wüsthoff, 1938a: fig. 25 (Quedius; aedeagal illustration).

  • — Smetana, 1962b: 140 (Quedius; subgenus Raphirus; characters).

  • — Coiffait, 1963a: 375 (Quedius; subgenus Sauridus; characters).

  • — Lohse, 1964: 215 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 277 (Quedius; subgenus Raphirus; Germany; Austria; Switzerland; Italy; France; Slovenia; Croatia; Bosnia Hercegovina; Czechoslovakia; Poland; Romania).

  • — Coiffait, 1978: 207 (Quedius; subgenus Sauridus; characters; Alps; Apennines).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • — Tóth, 1984: 131 (Quedius; characters).

  • — Lucht, 1987: 109 (Quedius; nominate subspecies; checklist; Germany; Austria; Switzerland; France).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Europe.

montanus Heer, 1839: 277 (Philonthus; Type locality: Matt; alle Glarneralpen; Paradies an der Zaportalp; Urserenthal; Furca; am Mt. Rosa).

  • — Küster, 1853a: 63 (Quedius; characters; Switzerland).

  • — Kraatz, 1857d: 506 (Quedius; synonym of fimbriatus).

  • — Fauvel, 1874: 293 [= 1874a: 519] (Quedius; synonym of dubius).

  • — Mulsant and Rey, 1876: 721 [= 1877: 577] (Quedius; subgenus Sauridus; characters; France).

  • — Ganglbauer, 1895: 405 (Quedius; synonym of dubius).

  • — Porta, 1907: 135 (Quedius; synonym of dubius).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; synonym of dubius).

  • — Gridelli, 1924d: 98 (Quedius; subgenus Raphirus; variety of dubius; characters; Italy).

  • — Scheerpeltz, 1933: 1439 (Quedius; variety of dubius).

  • — Wüsthoff, 1938a: fig. 26 (Quedius; form of dubius; aedeagal illustration).

  • — Smetana, 1958: 366 (Quedius; synonym of dubius fimbriatus).

  • — Smetana, 1962b: 141 (Quedius; aberration of dubius).

  • — Lohse, 1964: 214 (Quedius; subgenus Raphirus; aberration of dubius; characters; central Europe).

  • — Coiffait, 1978: 207 (Quedius; subgenus Sauridus; variety of dubius; characters).

  • — Tóth, 1984: 132 (Quedius; variety of dubius; characters).

fimbriatus Erichson, 1840: 539 (Quedius; Type locality: Austria).

  • — Märkel and Kiesenwetter, 1848a: 325 (Quedius; notes; Carpathian Alps).

  • — Redtenbacher, 1849: 710 (Philonthus; characters; Austria).

  • — Kiesenwetter, 1851a: 419 (Quedius; France).

  • — Fairmaire and Laboulbène, 1856: 544 (Quedius; characters; France).

  • — Kraatz, 1857d: 506 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 203 (Quedius; characters; Austria).

  • — Fauvel, 1874: 293 [= 1874a: 519] (Quedius; synonym of dubius).

  • — Redtenbacher, 1874: 200 (Quedius; characters; Austria).

  • — Ganglbauer, 1895: 405 (Quedius; synonym of dubius).

  • — Porta, 1907: 135 (Quedius; synonym of dubius).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; synonym of dubius).

  • — Gridelli, 1924d: 98 (Quedius; subspecies of dubius).

  • — Scheerpeltz, 1933: 1439 (Quedius; subspecies of dubius).

  • — Wüsthoff, 1938a: fig. 27 (Quedius; form of dubius; aedeagal illustration).

  • — Smetana, 1958: 366 (Quedius; subgenus Raphirus; subspecies of dubius; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 141 (Quedius; subgenus Raphirus; subspecies of dubius; characters).

  • — Coiffait, 1963a: 375 (Quedius; subgenus Sauridus; variety of dubius; characters).

  • — Coiffait, 1978: 207 (Quedius; subgenus Sauridus; variety of dubius; characters).

  • — Tóth, 1984: 132 (Quedius; subspecies of dubius; characters).

  • — Lucht, 1987: 110 (Quedius; subspecies of dubius; checklist; Czechoslovakia; Austria).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; subspecies of dubius; Czech Republic).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; subspecies of dubius; Italy).

speculator Kiesenwetter, 1848: 57 (Quedius; Type locality: Kärthner Alpen).

  • — Kraatz, 1857d: 506 (Quedius; synonym of fimbriatus).

  • — Ganglbauer, 1895: 405 (Quedius; synonym of dubius).

  • — Porta, 1907: 135 (Quedius; synonym of dubius).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; synonym of dubius).

  • — Gridelli, 1924d: 98 (Quedius; synonym of dubius fimbriatus).

  • — Scheerpeltz, 1933: 1439 (Quedius; synonym of dubius).

  • — Smetana, 1958: 366 (Quedius; synonym of dubius fimbriatus).

flavolineatus Bernhauer, 1899a: 433 (Quedius; variety of dubius; Type locality: Schweiz).

  • — Porta, 1907: 135 (Quedius; variety of dubius).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; variety of dubius).

  • — Gridelli, 1924d: 98 (Quedius; synonym of dubius variety montanus).

  • — Scheerpeltz, 1933: 1439 (Quedius; synonym of dubius).

  • — Smetana, 1962b: 141 (Quedius; synonym of dubius).

ermischi Korge, 1969: 44 (Quedius; subspecies of dubius; Type locality: Südbulgarien: Musalla, Rhodope-Gebirge, 1600–2400 m).

  • — Coiffait, 1978: 207 (Quedius; subgenus Sauridus; subspecies of dubius; characters).

  • — Tóth, 1984: 132 (Quedius; subspecies of dubius; characters).

dui Smetana, 1988: 204 (Quedius; subgenus Microsaurus; Type locality: Punjab: Kulu Div.).

  • Distribution: India.

duplepunctatus Gridelli, 1922: 21 (Quedius; subgenus Sauridus; Type locality: Francia, Pirenei orient.: Montlouis).

  • — Gridelli, 1925: 138 (Quedius; subgenus Raphirus; probably identical to muscorum; characters; Pyrenees).

  • — Scheerpeltz, 1933: 1450 (Quedius; synonym of muscorum).

  • — Coiffait, 1963a: 381 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 226 (Quedius; subgenus Sauridus; cited as valid species; characters; Pyrenees).

  • Distribution: France.

duplex Fauvel, 1886, see: iridicolor Quedenfeldt, 1882.

duplopunctatus Lea, 1925, see: leai Scheerpeltz, 1933.

durgaa Smetana, 1988: 273 (Quedius; subgenus Raphirus; Type locality: India: W. Bengal, Darjeeling dist., 13 km N Ghoom, 1500).

  • Distribution: India.

dzambulensis Coiffait, 1967e: 403 (Quedius; subgenus Sauridus; Type locality: Aulie-Ata, Dzambul, Kazakstan).

  • — Coiffait, 1978: 229 (Quedius; subgenus Sauridus; characters; Turkestan).

  • — Boháč, 1988: 556 [= 1989: 40] (Quedius; characters; Uzbekistan; Turkestan).

  • Distribution: Kazakhstan, Uzbekistan.

echion Smetana, 1997: 65 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Langmusi, 3500 m).

  • — Smetana, 1999b: 217 (Quedius; subgenus Microsaurus; notes on holotype; characters of male; China).

  • Distribution: China.

edmundi Coiffait, 1969: 54 (Quedius; subgenus Microsaurus; replacement name for reitterianus Coiffait).

  • — Coiffait, 1978: 164 (Quedius; subgenus Microsaurus; characters; Caucasus).

  • Distribution: Caucasus.

microphthalmus Reitter, 1909: 109 (Quedius; subgenus Ediquus; [preoccupied]; Type locality: Kaukasus: Meskisches Gebirge).

  • — Gridelli, 1924d: 71 (Quedius; subgenus Microsaurus; characters; Caucasus).

  • — Coiffait, 1967e: 394 (Quedius; synonym of reitterianus).

  • — Coiffait, 1978: 164 (Quedius; synonym of edmundi).

reitterianus Coiffait, 1967e: 394 (Quedius; subgenus Microsaurus; replacement name for microphthalmus Reitter; [preoccupied]).

  • — Coiffait, 1978: 164 (Quedius; synonym of edmundi).

edwardsi Bernhauer, 1935, see: edwarsianus Korge, 1963.

edwardsi Sharp, 1886a: 378 (Quedius; Type locality: New Zealand: Picton).

  • — Broun, 1893: 1029 (Quedius; characters; New Zealand).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • Distribution: New Zealand.

edwarsianus Korge, 1963a: 84 (Quedius; replacement name for edwardsi, cited as edwarsi for both the Sharp and Bernhauer names).

  • — Coiffait and Saiz, 1966: 412 (Quedius; characters; Chile).

  • Distribution: Chile.

edwardsi Bernhauer, 1935c: 92 (Quedius; [preoccupied]; Type locality: Süd-Chile: Llanquihue Prov., Casa Pangue).

  • — Korge, 1963a: 84 (Quedius; cited as edwarsi; synonym of edwarsianus).

  • — Coiffait and Saiz, 1966: 413 (Quedius; synonym of edwardsianus).

ehecatl Smetana, 1976a: 240 (Quedius; subgenus Quedionuchus; Type locality: Mexico: Oaxaca: 4 mi. W. jct. Mex. 175 Yuvila Turnoff, 9300 ft.).

  • Distribution: Mexico.

eklai Smetana, 1988: 257 (Quedius; subgenus Raphirus; Type locality: Nepal: 172 Parbat Dist., zwischen Chitre und Ghandrung, Chitre-Seite des Passes, 2800–2900 m).

  • Distribution: Nepal.

elegans Ericson, 1897, see: brevicornis Thomson, 1860.

elevatus Hatch, 1957: 228 (Quedius; subgenus Sauridus; Type locality: Washington: Snoqualmie. Also cited from Oregon and British Columbia).

  • — Smetana, 1971: 224 (Quedius; subgenus Raphirus; characters; British Columbia; California; Nevada; Oregon; Washington).

  • — Smetana, 1971a: 1848 (Quedius; subgenus Raphirus; collecting notes; California).

  • Distribution: Canada, USA.

emei Smetana, 1997: 67 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Emei Shan, 3000 m, 29°32′N, 103°21′E).

  • — Smetana, 1997d: 454 (Quedius; subgenus Microsaurus; characters of female; China).

  • Distribution: China.

ennius Smetana, 1996: 16 (Quedius; subgenus Microsaurus; Type locality: China: Yunnan prov., Heishui 35 km N. Lijiang, 27°13′N, 100°19′E).

  • Distribution: China.

ephialtes Smetana, 1997: 60 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Gongga Shan, abv. Camp 3, 3300–3350 m).

  • — Smetana, 1997d: 454 (Quedius; subgenus Microsaurus; collecting notes; China).

  • — Smetana, 1999b: 216 (Quedius; subgenus Microsaurus; notes; China).

  • Distribution: China.

eppelsheimi Quedenfeldt, 1882, see: plancus Erichson, 1840.

eppelsheimii Seidlitz, 1891, see: brevicornis Thomson, 1860.

epytus Smetana, 1995f: 236 (Quedius; subgenus Microsaurus; Type locality: China: Gansu Mts., 25 km E. Xiahe, 3000 m).

  • Distribution: China.

erdciyasicus Korge, 1971, see: limbatus Heer, 1839.

erinci Korge, 1971, see: nemoralis Baudi, 1848.

ermischi Korge, 1969, see: dubius Heer, 1839.

ernestini Fauvel, 1872, see: praecox Gravenhorst, 1802.

erratus Coiffait, 1977 [Note: Nomen nudum], see: fissus Gridelli, 1938.

eruensis Broun, 1912: 398 (Quedius; Type locality: Erua).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • Distribution: New Zealand.

erythras Smetana, 1997: 58 (Quedius; subgenus Microsaurus; Type locality: China: Gansu Mts., 25 km E Xiahe, 3000 m).

  • — Smetana, 1999b: 216 (Quedius; subgenus Microsaurus; China).

  • Distribution: China.

erythroderes Lea, 1925: 247 (Quedius; Type locality: South Australia: Mount Lofty, Wallaroo. Victoria: Forrest).

  • — Scheerpeltz, 1933: 1439 (Quedius; cited as erythroderus; catalog).

  • — Oke, 1933: 123, 124 (Quedius; tarsal characters; Australia).

  • Distribution: Australia.

erythrogaster Mannerheim, 1852: 314 (Quedius; Type locality: insula Sitkha; [Note: See lectotype designation by Smetana, 1971]).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — Horn, 1878: 159 (Quedius; variety of fulgidus; characters).

  • — Porta, 1907: 114 (Quedius; synonym of fulgidus).

  • — Casey, 1915: 410 (Microsaurus; characters; Queen Charolotte Island; California; New York).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Gridelli, 1929: 24 (Quedius; characters).

  • — Scheerpeltz, 1933: 1439 (Quedius; catalog).

  • — Hatch, 1957: 222 (Quedius; subgenus Microsaurus; characters; British Columbia; Washington; Oregon; Idaho).

  • — Smetana, 1971: 80 (Quedius; subgenus Microsaurus; lectotype designation; characters; Alberta; British Columbia; Quebec; Alabama; Arkansas; California; Connecticut; Georgia; Illinois; Kentucky; Maine; Michigan; Nevada; New York; Ohio; Oregon; Pennsylvania; Tennessee; Virginia; Washington; West Virginia).

  • — Smetana, 1971b: 2 (Quedius; subgenus Microsaurus; Alaska).

  • — Smetana, 1973: 1426 (Quedius; Quebec).

  • — Smetana, 1976c: 169 (Quedius; subgenus Microsaurus; Ontario; Quebec).

  • — Smetana, 1978a: 824 (Quedius; subgenus Microsaurus; collecting notes; New York; Oklahoma).

  • — Smetana, 1981b: 634 (Quedius; Saskatchewan).

  • — Dearborn and Donahue, 1994: 15 (Quedius; Maine).

  • — Downie and Arnett, 1996: 390 (Quedius; subgenus Microsaurus; characters; USA).

  • Distribution: Canada, USA.

neotomae Hatch, 1957: 224 (Quedius; subgenus Microsaurus; Type locality: Oregon: Medford. Also cited from Washington).

  • — Graham, 1963: 37 (Quedius; collected from cave; California).

  • — Smetana, 1971: 80 (Quedius; synonym of erythrogaster).

erythropterus Stephens, 1832, see: cruentus Olivier, 1795.

erythropygus Gridelli, 1929, see: fulgidus Fabricius, 1793.

etruscus Mulsant and Rey, 1876, see: scintillans Gravenhorst, 1806.

euander Smetana, 1997: 63 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Gongga Shan, above Camp 3, 3300–3350 m).

  • — Smetana, 1997d: 454 (Quedius; subgenus Microsaurus; characters; collecting notes; China).

  • — Smetana, 1999b: 217 (Quedius; subgenus Microsaurus; China).

  • Distribution: China.

eumerus Smetana, 1971: 182 (Quedius; subgenus Distichalius; Type locality: Oregon: Mt. Hood. Also cited from Washington).

  • — Smetana, 1976c: 178 (Quedius; subgenus Distichalius; collecting notes; characters; Oregon; Washington).

  • Distribution: USA.

euryalus Smetana, 1997: 52 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Gongga Shan, Lake abv. Camp 2, 2750 m).

  • — Smetana, 1997d: 453 (Quedius; subgenus Microsaurus; collecting notes; China).

  • Distribution: China.

excessus Last, 1975 [Note: Nomen nudum], see: splendidus Fauvel, 1878.

expectatus Israelson, 1985: 145 (Quedius; subgenus Microsaurus; [Note: Described for some specimens misidentified as angustifrons by Wollaston, 1864: 563 and 1865: 486 and by Coiffait(see following citations)]; Type locality: Gomera: Bosque del Cedro).

  • — Coiffait, 1954: 167 (Quedius; subgenus Microsaurus; [Note: Misidentified as angustifrons according to Israelson, 1985: 145]; Canary Islands).

  • — Coiffait, 1978: 175 (Quedius; subgenus Microsaurus; [Note: Misidentified as angustifrons according to Israelson, 1985: 145]; characters; Canary Islands).

  • — Assing, 1999a: 346 (Quedius; Canary Islands).

  • Distribution: Canary Islands.

explanatus LeConte, 1858: 61 (Quedius; Type locality: San Diego, California; [Note: See lectotype designation by Smetana, 1971]).

  • — Horn, 1878: 157 (Quedius; characters; California; Montana).

  • — Fall and Cockerell, 1907: 166 (Quedius; New Mexico).

  • — Casey, 1915: 423 (Megaquedius; characters; California).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Gridelli, 1932: 15 (Quedius; subgenus Microsaurus; characters; California; Montana).

  • — Scheerpeltz, 1933: 1439 (Quedius; catalog).

  • — Hatch, 1957: 230 (Quedius; subgenus Megaquedius; characters; British Columbia; Washington; Oregon; Idaho).

  • — Smetana, 1971: 42 (Quedius; subgenus Megaquedius; lectotype designation; characters; British Columbia; Arizona; California; Colorado; Idaho; kansas; Missouri; Montana; New Mexico; Oregon; Utah; Washington; Wyoming).

  • — Smetana, 1971a: 1839 (Quedius; subgenus Megaquedius; collecting notes; New Mexico; Arizona).

  • — Smetana, 1978a: 823 (Quedius; subgenus Megaquedius; collecting notes; Kansas; Nevada).

  • Distribution: USA, Canada.

laxatus Casey, 1915: 422 (Megaquedius; Type locality: California, probably northern).

  • — Scheerpeltz, 1933: 1446 (Quedius; catalog).

  • — Smetana, 1971: 43 (Quedius; synonym of explanatus).

exulans Smetana, 1971, see: desertus Horn, 1878.

faang Smetana, 1999e: 538 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan Pv., Meigu Xian, Wahei, Dafengding, 2320 m).

  • Distribution: China.

fagelianus Coiffait, 1967e: 391 (Quedius; subgenus Microsaurus; Type locality: Safa, Liban).

  • — Coiffait, 1978: 180 (Quedius; subgenus Microsaurus; characters; Lebanon).

  • — Smetana, 1995a: 80 (Quedius; subgenus Distichalius; notes; Lebanon).

  • Distribution: Israel, Lebanon.

loebli Smetana, 1978: 121 (Quedius; subgenus Distichalius; Type locality: Israel: Galilee au-dessus Safad, 500 m).

  • — Smetana, 1995a: 80 (Quedius; synonym of fagelianus).

fageti Thomson, 1867, see: maurus C. Sahlberg, 1830.

fallaciosus Kraatz, 1862, see: nitipennis Stephens, 1833.

fangi Smetana, 1995: 80 (Quedius; subgenus Raphirus; Type locality: Taiwan: Hualien Hsien: Taroko N. P., Nanhushi Hut, 2200 m).

  • Distribution: Taiwan.

farkaci Smetana, 1997d: 464 (Quedius; subgenus Microsaurus; Type locality: China: N. Yunnan, Xue Shan, nr. Zhongdian, 4000-4100 m, 27°49 N, 99°34 E).

  • Distribution: China.

fasciculatus Eppelsheim, 1886: 39 (Quedius; Type locality: Nicolajewsk; Chabarofka).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Gridelli, 1924d: 25 (Quedius; subgenus Microsaurus; characters; Amur region).

  • — Scheerpeltz, 1933: 1439 (Quedius; catalog).

  • — Coiffait, 1978: 159 (Quedius; subgenus Microsaurus; characters; Siberia).

  • — Smetana, 1978b: 84 (Quedius; subgenus Microsaurus; characters; Siberia).

  • Distribution: Russia.

klimai Roubal, 1914a: 164 (Quedius; Type locality: Irkutsk: Narcinck).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Gridelli, 1924d: 73 (Quedius; subgenus Microsaurus; characters; Siberia).

  • — Scheerpeltz, 1933: 1440 (Quedius; synonym of fasciculatus).

  • — Coiffait, 1978: 159 (Quedius; synonym of fasciculatus).

feihuensis Smetana, 1999d: 530 (Quedius; subgenus Microsaurus; Type locality: China: Hunan Pv., Longshan X., Huoyan, Feihu Dong Cave, 260 m).

  • Distribution: China.

fellmani Zetterstedt, 1838: 62 (Staphylinus; Type locality: Lapponia Umensis: Wilhelmina).

  • — Erichson, 1840: 547 (Quedius; synonym of hyperboreus).

  • — Thomson, 1860: 179 (Raphirus; synonym of attenuatus).

  • — Fauvel, 1869: 486 (Quedius; synonym of attenuatus).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — A. Strand, 1937: 110 (Quedius; characters; notes).

  • — Renkonen, 1939: 65 (Quedius; notes; Finland).

  • — Smetana, 1963c: 300 (Quedius; subgenus Raphirus; Mongolia).

  • — Palm, 1963b: 153 (Quedius; subgenus Raphirus; characters; Sweden; Norway; Finland).

  • — Smetana, 1964: 39 (Quedius; subgenus Raphirus; Alaska).

  • — Smetana, 1965: 47 (Quedius; subgenus Raphirus; characters; Labrador; Newfoundland).

  • — Smetana, 1965a: 16 (Quedius; subgenus Raphirus; notes; Alaska).

  • — Smetana, 1967d: 217 (Quedius; subgenus Raphirus; Mongolia).

  • — Smetana, 1995a: 86 (Quedius; subgenus Raphirus; notes; Russia).

  • — Smetana, 1971: 233 (Quedius; subgenus Raphirus; characters; Alberta; British Columbia; Labrador; Manitoba; Newfoundland; Northwest Territories; Quebec; Yukon Territory; Alaska; Colorado; Idaho; New Hampshire).

  • — Smetana, 1973: 1431 (Quedius; notes; Northwest Territories; Alaska).

  • — Smetana 1975c: 176 (Quedius; Mongolia).

  • — Smetana, 1976c: 183 (Quedius; subgenus Raphirus; Colorado).

  • — Smetana, 1976d: 28 (Quedius; subgenus Raphirus; characters; Siberia).

  • — Coiffait, 1978: 268 (Quedius; subgenus Raphirus; characters; Scandinavia).

  • — Smetana, 1978b: 86 (Quedius; subgenus Raphirus; Siberia).

  • — Smetana, 1981b: 643 (Quedius; collecting notes; Manitoba; Saskatchewan).

  • — Downie and Arnett, 1996: 393 (Quedius; subgenus Raphirus; characters; USA).

  • — Ryabukhin, 1999: 74 (Quedius; subgenus Raphirus; catalog; habitat; distribution; northeastern Russia).

  • Distribution: Europe, Russia, Mongolia, Canada, USA.

hyperboreus Erichson, 1840: 547 (Quedius; Type locality: Unalaschka; [Note: See lectotype designation by Smetana, 1971]).

  • — Mannerheim, 1843b: 233 (Quedius; characters; Unalaska).

  • — Fauvel, 1874: 306 [= 1874a: 532] (Quedius; synonym of attenuatus).

  • — Porta, 1907: 151 (Quedius; synonym of fulvicollis).

  • — Fall and Cockerell, 1907: 166 (Quedius; New Mexico).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fulvicollis).

  • — Notman, 1919a: 95 (Quedius; Ontario).

  • — Gridelli, 1925: 161 (Quedius; synonym of attenuatus).

  • — Fall, 1926: 150 (Quedius; tentative identification; Alaska).

  • — Scheerpeltz, 1933: 1432 (Quedius; synonym of attenuatus).

  • — Smetana, 1958: 393 (Quedius; synonym of nitipennis).

  • — Smetana, 1971: 233 (Quedius; lectotype designation; synonym of fellmani).

  • — Coiffait, 1978: 270 (Quedius; [Note: I assume Coiffait overlooked the change of synonymy by Smetana, 1971: 233]; synonym of nitipennis).

femoralis Sharp, 1884: 338 (Quedionuchus; Type locality: Guatemala, Totonicapam, 8500 to 10,500 feet; [Note: See lectotype designation by Smetana, 1976a]. Also cited from Quiche Mountains 7000 to 9000 feet).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Smetana, 1976a: 234 (Quedius; subgenus Quedionuchus; lectotype designation: Totonicapam, 8500–10,500 ft., Guatemala; characters; Guatemala).

  • Distribution: Guatemala.

fen Smetana, 1996c: 229 (Quedius; subgenus Raphirus; Type locality: China: Sichuan: Emei Shan).

  • Distribution: China.

fenderi Hatch, 1957: 228 (Quedius; subgenus Sauridus; Type locality: Oregon: Lake of Woods, Ashland Rd.).

  • — Smetana, 1971: 178 (Quedius; subgenus Distichalius; characters; Oregon).

  • — Smetana, 1978a: 828 (Quedius; subgenus Distichalius; collecting notes; Oregon).

  • Distribution: USA.

fenestratus Scholz, 1928 [Note: Nomen nudum], see: paradisianus Heer, 1839.

ferox Blackburn, 1888, see: tepperi Blackburn, 1888.

ferox LeConte, 1878, see: Hemiquedius.

filicornis Eppelsheim, 1895, see: Indoquedius.

fimbriatus Erichson, 1840, see: dubius Heer, 1839.

fiorii Korge, 1961, see: mesomelinus Marsham, 1802.

fissus Gridelli, 1938: 13 (Quedius; subgenus Microsaurus; Type locality: Asia Minore meridionale: Monti Salbak nel “Karisch. Taurus. Isola di Lesbo. Asia Minore, a Smirne).

  • — Coiffait, 1970e: 62 (Quedius; Lesbos).

  • — Coiffait, 1976b: 90 (Quedius; subgenus Microsarus; Rhodes).

  • — Bordoni, 1976: 236 (Quedius; characters; notes; Turkey).

  • — Coiffait, 1978: 178 (Quedius; subgenus Microsaurus; characters; Turkey; Rhodes; Lesbos).

  • Distribution: Turkey, Greece.

decolor Coiffait, 1976b: 90 (Quedius; subgenus Microsarus; variety of fissus; [Note: Nomen nudum (Article 15.2)]; cited from Rhodes: Mont Profitis Ilias).

  • — Coiffait, 1978: 178 (Quedius; subgenus Microsaurus; variety of fissus; characters).

erratus Coiffait, 1977: 136 (Quedius; subgenus Microsaurus; variety of fissus; [Note: Nomen nudum (Article 15.2)]; cited from Rhodes).

  • — Coiffait, 1978: 178 (Quedius; synonym of fissus).

flavescens Linné, 1758, see: cinctus Paykull, 1790.

flavicaudus Sharp, 1884: 333 (Quedius; Type locality: Panama, Volcan de Chiriqui between 2000 and 4000 feet, Bugaba).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; catalog; Bolivia; Argentina).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1955a: 177 (Quedius; subgenus Cyrtoquedius; characters in key; Panama).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • Distribution: Panama, Bolivia, Argentina.

flavicornis Sharp, 1889: 30 (Quedius; Type locality: Ishiuchi; Hitoyoshi).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Nakane, 1963a: 94 (Quedius; characters; habitus photograph; Japan).

  • — Smetana, 1995: 55 (Quedius; subgenus of Microsaurus; characters; bionomics; Taiwan).

  • Distribution: Japan, Taiwan.

flavinasus Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius; Type locality: Iatahy, Goyaz).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 178 (Quedius; subgenus Cyrtoquedius; characters in key; Argentina; Paraguay; Brazil).

  • Distribution: Brazil, Paraguay, Argentina.

nigrifrons Bernhauer, 1934b: 217 (Quedius; variety of rufinasus; [Note: Cited as rufinasus Bernhauer but that species was described by Sharp. Bernhauer probably intended flavinasus Bernhauer]; Type locality: Argentinien: Misiones: Loreto).

  • — Scheerpeltz, 1955a: 176 (Quedius; subgenus Cyrtoquedius; variety of flavinasus; characters in key; Argentina).

flavipennis Baudi, 1848, see: virgulatus Erichson, 1840.

flavocaudatus Cameron, 1944: 13 (Quedius; subgenus Microsaurus; Type locality: United Provinces: Naini Tal, altitude 6000–8000 feet).

  • — Smetana, 1988: 200 (Quedius; subgenus Microsaurus; characters; Uttar Pradesh; Nepal).

  • Distribution: India, Nepal.

flavocaudatus Hayashi, 1992, see: noboruitoi Hayashi, 1992.

flavolineatus Bernhauer, 1899, see: dubius Heer, 1839.

flavomarginatus Gridelli, 1925, see: fumatus Stephens, 1833.

flavopilus Gerhardt, 1899, see: paradisianus Heer, 1839.

flavopterus Geoffroy, 1785, see: plagiatus Mannerheim, 1843.

flavoterminatus Cameron, 1932, see: apicicornis Eppelsheim, 1895.

floralis Lacordaire, 1835, see: ochripennis Ménétriés, 1832.

fluviatilis Cameron, 1926b: 369 (Quedius; subgenus Raphirus; Type locality: Chakrata Dist., Dodora Khud, 8000′; [Note: See lectotype designation by Smetana, 1988]. Originally cited from Chakrata district, 6000 to 9300 feet).

  • — Cameron, 1932: 297 (Quedius; subgenus Raphirus; characters; India).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Smetana, 1988: 239 (Quedius; subgenus Raphirus; lectotype designation: Chakrata Dist., Dodora Khud, 8000′; characters; Kashmir; Himachal Pradesh; Uttar Pradesh).

  • Distribution: India.

kashmiricus Cameron, 1932: 296 (Quedius; subgenus Raphirus; Type locality: Kashmir: Gulmarg, alt. 8000 feet; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1445 (Quedius; catalog).

  • — Smetana, 1988: 239 (Quedius; lectotype designation; synonym of fluviatilis).

cachemiricus Coiffait, 1982a: 277 (Quedius; subgenus Sauridus; Type locality: Cachemire: Sonamarg, 2600–2750 m).

  • — Smetana, 1988: 239 (Quedius; synonym of fluviatilis).

nichinaiensis Coiffait, 1982: 79 (Quedius; subgenus Sauridus; Type locality: Inde: Cachemire, Sonamarg, Himalaya-Hauptkette, Nichinai Tal, 3100–3200 m).

  • — Smetana, 1988: 239 (Quedius; synonym of fluviatilis).

fodinarum Uéno and Watanabe, 1966: 323 (Quedius; subgenus Microsaurus; Type locality: Japan: central Kyushu: Oita Prefecture: an abandoned mine called “Otani-ko”, at Kiura-Kozan in Onoichi of Umé-mura).

  • — Watanabe, 1996a: 13 (Quedius; mine collection; Japan).

  • — Bordoni and Oromi, 1998: 1157 (Quedius; subgenus Microsaurus; facultative cave species).

  • Distribution: Japan.

fonteius Smetana, 1995f: 247 (Quedius; subgenus Microsaurus; Type locality: China E Quinghai prov. pass 30 km S of Huizo, 3600 m).

  • Distribution: China.

fontinalis Casey, 1915, see: desertus Horn, 1878.

formosae Cameron, 1949, see: Indoquedius.

forsteri Scheerpeltz, 1960c: 129 (Quedius; subgenus Microsaurus; Type locality: Bolivien: Songtal, St. Rosa, 2800 m).

  • Distribution: Bolivia.

franzi Smetana, 1975a: 326 (Quedius; subgenus Microsaurus; Type locality: Central Nepal: Trail from Gosaikunde to Fulung monastery).

  • — Smetana, 1988: 217 (Quedius; subgenus Microsaurus; characters; Nepal).

  • Distribution: Nepal, India.

trisulensis Coiffait, 1982: 79 (Quedius; subgenus Microsaurus; Type locality: Nepal: Trisuli Tal, Gosainkund, 3200 m).

  • — Smetana, 1988: 217 (Quedius; synonym of franzi).

najik Smetana, 1992b: 4 (Quedius; subgenus Microsaurus; subspecies of franzi; Type locality: India: Sikkim, “Dzongri, 3970 m, Kangchendzonga Area”).

frater Smetana, 1971: 182 (Quedius; subgenus Distichalius; Type locality: Washington: Olympic Hot Springs).

  • — Smetana, 1976c: 178 (Quedius; subgenus Distichalius; Washington).

  • — Smetana, 1978a: 829 (Quedius; subgenus Distichalius; collecting notes; Alberta; British Columbia).

  • Distribution: Canada, USA.

fraternus Bernhauer, 1914, see: suturalis Kiesenwetter, 1845.

fraudulentum Coiffait, 1967 [Note: Nomen nudum], see: brevis Erichson, 1840.

frenatus Erichson, 1840: 533 (Quedius; Type locality: La Guayram, Columbia).

  • — Fauvel, 1891: 119 (Quedius; Venezuela).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; catalog).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 179 (Quedius; subgenus Cyrtoquedius; characters in key; Venezuela; Ecuador; Colombia).

  • Distribution: Venezuela, Colombia, Ecuador.

freyanus Smetana, 1999: 79 (Quedius; subgenus Raphirus; replacement name for freyi Smetana).

  • Distribution: China.

freyi Smetana, 1998b: 108 (Quedius; subgenus Raphirus; [preoccupied]. Type locality: China: Lungai, 2000 m, Wassuland, W. Szechuan, Sankiangkou).

  • — Smetana, 1999: 79 (Quedius; synonym of freyanus).

  • Distribution: China.

freyi Scheerpeltz, 1956, see: cyprusensis Last, 1955.

freyi Smetana, 1998, see: freyanus Smetana, 1999.

frigidus Smetana, 1971: 230 (Quedius; subgenus Raphirus; Type locality: New York: Mt. Whiteface. Also cited from Maine, New Hampshire, and Alberta).

  • — Smetana, 1973: 1431 (Quedius; notes; Newfoundland; Northwest Territories).

  • — Smetana, 1976c: 183 (Quedius; subgenus

  • — Smetana, 1976c: 183 (Quedius; subgenus Raphirus; collecting notes; Ontario).

  • — Smetana, 1978a: 830 (Quedius; subgenus Raphirus; collecting notes; British Columbia; Newfoundland; New Hampshire).

  • — Smetana, 1981b: 642 (Quedius; collecting notes; Alaska).

  • — Smetana, 1990d: 101 (Quedius; Alberta; New Hampshire).

  • — Downie and Arnett, 1996: 393 (Quedius; subgenus Raphirus; characters; USA).

  • Distribution: Canada, USA.

frontalis Nordmann, 1837, see: levicollis Brullé, 1832.

fuchsii Scriba, 1866, see: mesomelinus Marsham, 1802.

fulgidus Fabricius, 1793: 525 (Staphylinus; [preoccupied]; [Note: Nearly all writers (see for example Erichson, 1839a: 486, 1840: 525; Bernhauer and Schubert, 1916: 423; Smetana, 1958: 349; Smetana, 1971: 66; Coiffait, 1978: 146) have cited Fabricius, 1787: 220 as the original reference for Quedius fulgidus. Tottenham (1939d: 237) in his discussion of fulgidus Fabricius, 1787: 220, and fulgidus Fabricius, 1793: 525, concluded that the earlier description referred to the xantholine, Gauropterus fulgidus, and the later to the quediine, Quedius fulgidus]; Type locality: Germania).

  • — Panzer, 1795: 355 (Staphylinus; characters; Germany).

  • — Fabricius, 1801: 596 (Staphylinus; characters; Germany).

  • — Turton, 1802: 512 (Staphylinus; characters; Germany).

  • — Gyllenhal, 1810: 304 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1916: 423 cited variabilis variety e Gyllenhal as a synonym of fulgidus; there is no available name]; characters; Sweden).

  • — Erichson, 1839a: 486 (Quedius; characters; Germany).

  • — Heer, 1839: 273 (Philonthus; characters; Switzerland).

  • — Erichson, 1840: 525 (Quedius; characters; Europe; America septentrionalis).

  • — Redtenbacher, 1849: 708 (Philonthus; characters; Austria).

  • — Kiesenwetter, 1851a: 419 (Quedius; Pyrenees).

  • — Küster, 1853a: 60 (Quedius; characters; Europe).

  • — Hardy, 1851: 31 (Quedius; notes; England).

  • — Rosenhauer, 1856: 71 (Quedius; Granada; Sierra Nevada).

  • — Fairmaire and Laboulbène, 1856: 539 (Quedius; characters; France).

  • — Staudinger, 1857: 285 (Quedius; notes; Iceland).

  • — Kraatz, 1857d: 492 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 201 (Quedius; characters; Austria).

  • — Peyron, 1858: 428 (Quedius; Turkey).

  • — Kraatz, 1859: 67 (Quedius; characters; India).

  • — Thomson, 1860: 174 (Microsaurus; characters; Scandinavia).

  • — Hochhuth, 1862: 44 (Quedius; notes; Russia).

  • — Wollaston, 1865: 486 (Quedius; habitat; Canary Islands).

  • — Rottenberg, 1870: 28 (Quedius; notes; Sicily).

  • — Wollaston, 1871: 299 (Quedius; notes).

  • — Hochhuth, 1872: 131 (Quedius; notes; characters; Ukraine).

  • — Fauvel, 1874: 279 [= 1874a: 505] (Quedius; characters; France; Madeira and Canary Islands; Algeria; Caramania; Indonesia; Australia; USA).

  • — Redtenbacher, 1874: 198 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 266 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 657 [= 1877: 513] (Quedius; characters; France).

  • — J. Sahlberg, 1876: 25 (Microsaurus; Finland).

  • — Fauvel, 1877: 268 (Quedius; characters; Australie, Melbourne, Rockhampton; Tasmanie).

  • — Provancher, 1877: 728 (Quedius; characters; Quebec).

  • — Fauvel, 1878: 127 [= 1878a: 47] (Quedius; Algeria).

  • — Fauvel, 1878g: 552 (Quedius; Australia).

  • — Weise, 1884: 423 (Quedius; [Note: Misidentification: Scheerpeltz, 1933: 1440 cited erythrogaster Weise as a variety of fulgidus; Weise did not describe the species as new, he attributed it to Mannerheim]; variety of fulgidus; notes; characters).

  • — Fauvel, 1886: 57 [= 1886a: 49] (Quedius; Algeria).

  • — Fowler, 1888: 232 (Quedius; characters; Britain).

  • — Ganglbauer, 1895: 399 (Quedius; subgenus Microsaurus; characters; Europe; Mediterranean region; Atlantic Islands; India; Australia; New Zealand; North America).

  • — Fauvel, 1902b: 114 (Quedius; Algeria; Tunisia).

  • — Germain, 1903: 437 (Quedius; introduced species; notes; Chile).

  • — Sainte-Claire Deville, 1906: 101 (Quedius; Corsica).

  • — Porta, 1907: 114 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Fall and Cockerell, 1907: 166 (Quedius; New Mexico).

  • — Montandon, 1908: 80 (Quedius; Romania).

  • — Reitter, 1909: 111 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Blatchley, 1910: 371 (Quedius; characters; Indiana).

  • — Johansen, 1914: 355 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; catalog).

  • — Notman, 1919a: 95 (Quedius; Ontario).

  • — Munster, 1923: 192 (Quedius; subgenus Microsaurus; Fennoscandia).

  • — Gridelli, 1924d: 51 (Quedius; subgenus Microsaurus; characters; Germany; Styria; Transylvania; Dalmatia; Italy; Portugal).

  • — Gridelli, 1929: 22 (Quedius; characters; Germany, France, Portugal, Bohemia, Russia).

  • — Portevin, 1929: 341 (Quedius; characters; France).

  • — Cameron, 1932: 288 (Quedius; subgenus Microsaurus; characters; India).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • — Scheerpeltz, 1935a: 655 (Philonthus; Australia).

  • — Wüsthoff, 1938a: fig. 9 (Quedius; aedeagal illustration).

  • — Tottenham, 1939d: 237 (Quedius; synonym of assimilis).

  • — Hellén, 1939: 32 (Quedius; characters; Finland).

  • — Roubal, 1941b: 124, 127 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Paulian, 1941: 264 (Quedius; larval characters).

  • — Brinck, 1948: 34 (Quedius; collecting and ecological notes; Tristan da Cunha).

  • — Jeannel and Jarrige, 1949: 383 (Quedius; subgenus Microsaurus; notes; cave species; France; Spain; Algeria).

  • — Tottenham, 1949: 422 (Quedius; synonym of assimilis).

  • — V. Hansen, 1952: 133 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Hatch, 1957: 221 (Quedius; subgenus Microsaurus; characters; British Columbia; Washington; Oregon; Idaho).

  • — Smetana, 1958: 349 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 157 (Quedius; subgenus Microsaurus; characters).

  • — Palm, 1963b: 134 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 209 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Raianu, 1964: 102 (Quedius; Romania).

  • — Horion, 1965: 262 (Quedius; subgenus Microsaurus; all central and southern Europe; Finland; Sweden; Norway; Britain; Denmark; Ukraine).

  • — Coiffait and Saiz, 1966: 412 (Quedius; subgenus Microsaurus; characters; Chile).

  • — Pototskaia, 1967: 83 (Quedius; larval characters).

  • — Korge, 1968: 51 (Quedius; notes).

  • — Szujecki, 1968a: 734 (Quedius; Poland).

  • — Smetana, 1971: 66 (Quedius; subgenus Microsaurus; characters; British Columbia; Manitoba; Arizona; California; Colorado; Georgia; Idaho; Illinois; Indiana; Iowa; Kansas; Kentucky; Maryland; Massachusetts; Michigan; Minnesota; Missouri; New Jersey; New Mexico; New York; Oregon; Pennsylvania; Texas; Virginia; Washington; West Virginia; Wisconsin).

  • — Smetana, 1971b: 2 (Quedius; subgenus Microsaurus; California).

  • — Saiz, 1971: 385 (Quedius; Chile).

  • — Tikhomirova, 1973a: 188 (Quedius; checklist USSR).

  • — Bordoni, 1973d (Quedius; subgenus Microsaurus; aedeagal anomaly).

  • — Bordoni, 1974: 65 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Bordoni, 1974b: 10 (Quedius; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Bordoni, 1977a: 18 (Quedius; subgenus Microsaurus; Spain).

  • — Coiffait, 1978: 146 (Quedius; subgenus Microsaurus; characters; Palaearctic region).

  • — Smetana, 1978a: 824 (Quedius; subgenus Microsaurus; collecting notes; Tennessee).

  • — Bordoni, 1978a: 63 (Quedius; cave collection; Turkey).

  • — Muona, 1979: 15 (Quedius; synonym of assimilis).

  • — Bordoni, 1981a: 14 (Quedius; pupal characters; Italy).

  • — Uhlig and Vogel, 1981: 123 (Quedius; notes; Germany).

  • — Bordoni, 1982a: 140 (Quedius; cave collection; Sardinia).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • — Tóth, 1984: 115 (Quedius; characters; Hungary).

  • — Segers, 1986: 49 (Quedius; Belgium).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; France; Benelux; Denmark; Sweden).

  • — Smetana, 1988: 209 (Quedius; subgenus Microsaurus; characters).

  • — Lohse and Lucht, 1989: 172 (Quedius; synonym of assimilis).

  • — Schülke and Uhlig, 1988: 12 (Quedius; Germany).

  • — M. Hansen, 1991: 107 (Quedius; synonym of assimilis).

  • — Elias, 1991: 557 (Quedius; Quaternary fossil collection; Montana).

  • — Campbell and Davies, 1991: 121 (Quedius; Canada).

  • — Silfverberg, 1992: 17 (Quedius; synonym of assimilis).

  • — Smetana, 1993b: 50 (Quedius; synonym of rufitarsis).

  • — Dearborn and Donahue, 1994: 16 (Quedius; Maine).

  • — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 29 (Quedius; synonym of assimilis).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; synonym of rufitarsis).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 245 (Quedius; synonym of assimilis).

  • — M. Hansen, 1996: 107 (Quedius; synonym of assimilis).

  • — Downie and Arnett, 1996: 391 (Quedius; subgenus Microsaurus; characters; USA).

  • — Ádám, 1996: 247 (Microsaurus; synonym of rufitarsis).

  • — Emberson, 1998: 36 (Quedius; collecting notes; Chatham Islands).

  • — Bordoni and Oromi, 1998: 1157 (Quedius; subgenus Microsaurus; facultative cave species).

  • — Zerche, 2000: 52 (Quedius; synonym of assimilis).

  • — Herman, 2001: 29, 83 (Quedius; case presented for citing assimilis as valid in application to Commission under Article 23.9.3, but opinion reversed and petition will request that fulgidus be cited as name of species).

  • Distribution: Libya, Tunisia, Algeria, Tristan da Cunha, Canary Islands, Madeira, Iceland, Europe, Russia, Turkey, India, Indonesia, Australia, New Zealand, Chatham Islands, Canada, USA, Chile.

fuscipennis Block, 1799: 116 (Staphylinus; [Note: Type locality: Tharand, Plauischer Grund bei Dresden).

  • — Erichson, 1840: 526 (Quedius; synonym of fulgidus).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — Fauvel, 1869: 486 (Quedius; synonym of fulgidus).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; synonym of mesomelinus).

  • — Smetana, 1958: 353 (Quedius; synonym of mesomelinus).

  • — Bordoni, 1974: 33 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 150 (Quedius; synonym of mesomelinus).

  • — Ádám, 1996: 247 (Microsaurus; valid name; Hungary).

  • — Zerche, 2000: 52, 53 (Quedius; synonym of assimilis; discussion; nomen oblitum).

rufitarsis Marsham, 1802: 512 (Staphylinus; Type locality: Britain).

  • — Gyllenhal, 1810: 304 (Staphylinus; synonym of variabilis Gyllenhal).

  • — Curtis, 1829: 25 (Quedius; catalog; Britain).

  • — C. Sahlberg, 1830: 316 (Staphylinus; variety of variabilis Gyllenhal; characters; Finland).

  • — Stephens, 1832: 220 (Quedius; characters; England).

  • — Curtis, 1837: pl. 638 (Quedius; characters; Britain).

  • — Stephens, 1839: 391 (Quedius; characters; England).

  • — Erichson, 1840: 526 (Quedius; synonym of fulgidus).

  • — Hardy, 1851: 32 (Quedius; synonym of fulgidus).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — Fauvel, 1869: 486 (Quedius; synonym of fulgidus).

  • — Fauvel, 1874: 280 [= 1874a: 506] (Quedius; synonym of fulgidus).

  • — Ganglbauer, 1895: 399 (Quedius; synonym of fulgidus).

  • — Porta, 1907: 114 (Quedius; synonym of fulgidus).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fulgidus).

  • — Cameron, 1932: 288 (Quedius; synonym of fulgidus).

  • — Smetana, 1958: 349 (Quedius; synonym of fulgidus).

  • — Bordoni, 1974: 65 (Quedius; synonym of fulgidus).

  • — Coiffait, 1978: 146 (Quedius; synonym of fulgidus).

  • — Nowosad, 1990: 136 (Quedius; cited as valid species; collected from mole nest; notes; Poland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Microsaurus; cited as valid species; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; cited as valid species; Italy).

  • — Ádám, 1996: 247 (Microsaurus; valid species; Hungary).

  • — Herman, 2001: 29 (Quedius; synonym of assimilis).

iracundus Say, 1830: 35 [= 1834: 449] (Staphylinus; Type locality: Indiana).

  • — Erichson, 1840: 526 (Quedius; synonym of fulgidus).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — LeConte, 1859: 564 (Quedius).

  • — Fauvel, 1869: 486 (Quedius; synonym of fulgidus).

  • — Fauvel, 1874: 280 [= 1874a: 506] (Quedius; synonym of fulgidus).

  • — Horn, 1878: 158 (Quedius; [Note: Misidentification: Scheerpeltz, 1933: 1444 cited fulgidus Horn as a synonym of iracundus; Horn did not describe the species as new, he attributed it to Say]; characters; California; Georgia).

  • — Horn, 1878: 158 (Quedius; synonym of fulgidus).

  • — Ganglbauer, 1895: 399 (Quedius; synonym of fulgidus).

  • — Porta, 1907: 114 (Quedius; synonym of fulgidus).

  • — Casey, 1915: 411 (Microsaurus; characters; Iowa; Indiana).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fulgidus).

  • — Cameron, 1932: 288 (Quedius; synonym of fulgidus).

  • — Scheerpeltz, 1933: 1444 (Quedius; catalog).

  • — Voris, 1939: 188 (Quedius; larval characters; Indiana).

  • — Smetana, 1971: 66 (Quedius; synonym of fulgidus).

  • — Herman, 2001: 29 (Quedius; synonym of assimilis).

haemopterus Stephens, 1832: 217 (Quedius; Type locality: London; Suffolk; Devonshire).

  • — Curtis, 1829: 25 (Quedius; nomen nudum; catalog; Britain).

  • — Stephens, 1839: 390 (Quedius; variety of nitidus; characters; England).

  • — Hardy, 1851: 32 (Quedius; synonym of fulgidus).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — Fauvel, 1869: 486 (Quedius; synonym of fulgidus).

  • — Fauvel, 1874: 280 [= 1874a: 506] (Quedius; synonym of fulgidus).

  • — Ganglbauer, 1895: 399 (Quedius; synonym of fulgidus).

  • — Porta, 1907: 114 (Quedius; synonym of fulgidus).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fulgidus).

  • — Cameron, 1932: 288 (Quedius; synonym of fulgidus).

  • — Tottenham, 1939d: 237 (Quedius; possible synonym of picipes [Note: Species not specifically listed as a synonym of picipes by Tottenham, 1949, Kloet and Hincks, 1945, or Pope, 1977]).

  • — Smetana, 1958: 349 (Quedius; synonym of fulgidus).

  • — Smetana, 1971: 66 (Quedius; synonym of fulgidus).

  • — Bordoni, 1974: 65 (Quedius; synonym of fulgidus).

  • — Coiffait, 1978: 146 (Quedius; synonym of fulgidus).

  • — Herman, 2001: 29 (Quedius; synonym of assimilis).

haemorrhoidalis Stephens, 1832: 218 (Quedius; Type locality: London; Devonshire).

  • — Stephens, 1839: 390 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 22 (Quedius; synonym of fulgidus).

  • — Herman, 2001: 29 (Quedius; synonym of assimilis).

nigricornis Stephens, 1832: 222 (Quedius; Type locality: London).

  • — Stephens, 1839: 392 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 22 (Quedius; synonym of fulgidus).

  • — Rye, 1870a: 82 (Quedius; synonymn of fulgidus; see last sentence under notes on nigricornis Holme which begins on page 81).

  • — Herman, 2001: 29 (Quedius; synonym of assimilis).

distinctus Runde, 1835: 6 (Staphylinus; Type locality: Halae).

  • — Erichson, 1839a: 486 (Quedius; synonym of fulgidus).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — Herman, 2001: 29 (Quedius; synonym of assimilis).

assimilis Nordmann, 1837: 78 (Philonthus; name given to variabilis variety f of Gyllenhal; Type locality: Europa).

  • — Erichson, 1839a: 486 (Quedius; synonym of fulgidus).

  • — Heer, 1839: 273 (Philonthus; synonym of fulgidus).

  • — Erichson, 1840: 526 (Quedius; synonym of fulgidus).

  • — Fauvel, 1869: 486 (Quedius; synonym of fulgidus).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fulgidus).

  • — Cameron, 1932: 288 (Quedius; synonym of fulgidus).

  • — Scheerpeltz, 1933: 1453 (Quedius; synonym of ochripennis).

  • — Tottenham, 1939d: 237 (Quedius; valid name to replace fulgidus).

  • — Kloet and Hincks, 1945: 167 (Quedius; cited as valid name to replace fulgidus; Britain).

  • — Hinton, 1945: 70 (Quedius; characters).

  • — Tottenham, 1949: 422 (Quedius; checklist; Britain).

  • — Scheerpeltz, 1968a: 89 (Quedius; synonym of ochripennis).

  • — Pope, 1977: 31 (Quedius; synonym of fulgidus).

  • — Muona, 1979: 15 (Quedius; cited as valid name; Denmark; Norway; Sweden; Finland; Russia; Baltic States).

  • — Lohse and Lucht, 1989: 172 (Quedius; note).

  • — M. Hansen, Kristensen, Mahler, and Pedersen, 1991: 107 (Quedius; Denmark).

  • — Silfverberg, 1992: 17 (Quedius; valid species; Finland; Russia; Sweden; Norway; Denmark; Estonia; Lithuania).

  • — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 29 (Quedius; cited as valid; Denmark).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 245 (Quedius; cited as valid name; Denmark).

  • — M. Hansen, 1996: 107 (Quedius; cited as valid name; Denmark).

  • — Telnov, Barsevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 39 (Quedius; valid species; Latvia).

  • — Anderson, Nash, and O'Connor, 1997: 19 (Quedius; Ireland).

  • — Assing, 1998e: 140 (Quedius; Canary Islands).

  • — Zerche, 2000: 52, 53 (Quedius; discussion; nomen protectum).

  • — Herman, 2001: 29, 83 (Quedius; case presented for citing name as valid in petition to Commission under Article 23.9.3, but opinion reversed and petition will request that fulgidus be cited as name of species).

bicolor Redtenbacher, 1849: 710 (Philonthus; [Note: Bernhauer and Schubert, 1916: 423 cited Redtenbacher, 1872 (= 1874): 201 as the first reference to this name]; Type locality: Wien).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — Redtenbacher, 1857: 204 (Quedius; characters; Austria).

  • — Fauvel, 1866c: 247 [= 1866d: 105] (Quedius; synonym of fulgidus).

  • — Fauvel, 1869: 486 (Quedius; synonym of fulgidus).

  • — Fauvel, 1874: 280 [= 1874a: 506] (Quedius; synonym of fulgidus).

  • — Redtenbacher, 1874: 201 (Quedius; characters; Austria).

  • — Fowler, 1888: 232 (Quedius; synonym of fulgidus).

  • — Ganglbauer, 1895: 399 (Quedius; synonym of fulgidus).

  • — Porta, 1907: 114 (Quedius; synonym of fulgidus).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fulgidus).

  • — Cameron, 1932: 288 (Quedius; synonym of fulgidus).

  • — Scheerpeltz, 1933: 1440 (Quedius; synonym of fulgidus).

  • — Smetana, 1958: 349 (Quedius; synonym of fulgidus).

  • — Smetana, 1971: 66 (Quedius; synonym of fulgidus).

  • — Bordoni, 1974: 65 (Quedius; synonym of fulgidus).

  • — Coiffait, 1978: 146 (Quedius; synonym of fulgidus).

quadripunctatus Thomson, 1867: 159 (Microsaurus; Type locality: Scandinavia).

  • — Fauvel, 1876a: 76 [= 1876: 261] (Quedius; synonym of fulgidus).

  • — J. Sahlberg, 1876: 25 (Microsaurus; synonym of fulgidus).

  • — Fowler, 1888: 232 (Quedius; synonym of fulgidus).

  • — Ganglbauer, 1895: 399 (Quedius; synonym of fulgidus).

  • — Porta, 1907: 114 (Quedius; synonym of fulgidus).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fulgidus).

  • — Cameron, 1932: 288 (Quedius; synonym of fulgidus).

  • — Smetana, 1958: 349 (Quedius; synonym of fulgidus).

  • — Smetana, 1971: 66 (Quedius; synonym of fulgidus).

  • — Bordoni, 1974: 65 (Quedius; synonym of fulgidus).

  • — Coiffait, 1978: 146 (Quedius; synonym of fulgidus).

peranxius Mulsant and Rey, 1876: 661 [= 1877: 517] (Quedius; Type locality: Not cited).

  • — Duvivier, 1883: 138 (Quedius; catalog; synonym of fulgidus).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fulgidus).

  • — Cameron, 1932: 288 (Quedius; synonym of fulgidus).

erythropygus Gridelli, 1929: 24 (Quedius; variety of fulgidus; Type locality: Cirenaica: Golfo di Bomba; Bengasi).

  • — Gridelli, 1930: 77 (Quedius; variety of fulgidus; [Note: Listed as a new variety but it has the same locality as the variety of the same name described in 1929 so I list this citation as a subsequent reference]; Libya).

  • — Scheerpeltz, 1933: 1440 (Quedius; variety of fulgidus).

  • — Roubal, 1941b: 124 (Quedius; subgenus Microsaurus; variety of fulgidus; characters).

  • — Normand, 1947a: 9 (Quedius; variety of fulgidus; Tunisia).

  • — Coiffait, 1978: 146 (Quedius; variety of fulgidus; characters).

creticus Mařan, 1935: 8 (Quedius; variety of fulgidus; Type locality: Nida, Creta centr.).

  • — Gridelli, 1938: 16 (Quedius; subspecies of fulgidus; characters; Crete).

  • — Roubal, 1941b: 124 (Quedius; subgenus Microsaurus; variety of fulgidus; characters).

  • — Coiffait, 1978: 146 (Quedius; subgenus Microsaurus; subspecies of fulgidus; characters; Crete).

fuliginosus Gravenhorst, 1802: 34 (Staphylinus; Type locality: Not cited; [Note: See lectotype designation by Gusarov, 1993]).

  • — Marsham, 1802: 504 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1916: 423, cited dilatatus Marsham as a synonym of fulginosus; Marsham did not describe the species as new, he attributed it to Fabricius]; characters; Britain).

  • — Latreille, 1804: 319 (Staphylinus; characters; Germany).

  • — Gravenhorst, 1806: 48 (Staphylinus).

  • — Gyllenhal, 1810: 301 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1916: 423, cited tristis Gyllenhal as a synonym of fuliginosus Gravenhorst; Gyllenhal did not describe the species as new, he attributed it to Gravenhorst who attributed it to Fabricius]; characters; Sweden).

  • — Mannerheim, 1830: 25 (Staphylinus; Finland).

  • — Mannerheim, 1831: 439 (Staphylinus; Finland).

  • — Nordmann, 1837: 78 (Philonthus; Germany; Finland; Wolhynia; Tauria; Odessa).

  • — Erichson, 1839a: 490 (Quedius; characters; Germany).

  • — Heer, 1839: 276 (Philonthus; characters; Switzerland).

  • — Erichson, 1840: 537 (Quedius; characters; Europe).

  • — Dufour, 1843: 33 (Staphylinus; characters; France).

  • — Kolenati, 1846: 22 (Quedius; Caucasus; Transcaucasus).

  • — Gravenhorst, 1847: 232 (Quedius; characters; notes).

  • — Hochhuth, 1849: 151 (Quedius; characters; Lenkoran).

  • — Redtenbacher, 1849: 710 (Philonthus; characters; Austria).

  • — Kiesenwetter, 1851a: 419 (Quedius; Pyrenees).

  • — Küster, 1853a: 62 (Quedius; characters; Europe).

  • — Hardy, 1851: 33 (Quedius; notes; England).

  • — Fairmaire and Laboulbène, 1856: 539 (Quedius; characters; France).

  • — Kraatz, 1857d: 503 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 203 (Quedius; characters; Austria).

  • — Kraatz, 1858a: 58 (Quedius; Greece; Crete).

  • — Peyron, 1858: 428 (Quedius; Turkey).

  • — Thomson, 1860: 173 (Quedius; characters; Scandinavia).

  • — Hochhuth, 1862: 46 (Quedius; Russia).

  • — Fauvel, 1865a: 293 [= 1865b: 43] (Quedius; habitat; France).

  • — Fauvel, 1874: 289 [= 1874a: 515] (Quedius; characters; France; Algeria; Turkey).

  • — Redtenbacher, 1874: 200 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 267 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 686 [= 1877: 542] (Quedius; subgenus Microsaurus; characters; France).

  • — J. Sahlberg, 1876: 24 (Quedius; Finland).

  • — Fauvel, 1878: 128 [= 1878a: 48] (Quedius; Algeria).

  • — Fauvel, 1886: 58 [= 1886a: 50] (Quedius; Algeria).

  • — Fowler, 1888: 237 (Quedius; subgenus Microsaurus; characters; Britain).

  • — Ganglbauer, 1895: 403 (Quedius; characters; Europe; Mediterranean region).

  • — Fauvel, 1902b: 115 (Quedius; Algeria; Tunisia).

  • — Poppius, 1905: 9 (Quedius; Orkney Islands).

  • — Sainte-Claire Deville, 1906: 102 (Quedius; Corsica).

  • — Porta, 1907: 128 (Quedius; subgenus Quedius; characters; Italy).

  • — Reitter, 1909: 111 (Quedius; characters; Germany).

  • — Poppius, 1913: 10 (Quedius; Faroes).

  • — Johansen, 1914: 363 (Quedius; subgenus Quedius; characters; Denmark).

  • — Eichelbaum, 1914: 339 (Quedius; abdominal segmentation; apical segements).

  • — Eichelbaum, 1915a: 91 (Quedius; abdominal segmentation; apical segements).

  • — Fiori, 1915: 16 (Quedius; Italy).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; catalog).

  • — Munster, 1923: 195 (Quedius; Fennoscandia).

  • — Gridelli, 1924d: 79 (Quedius; subgenus Quedius; characters; Europe; Mediterranean region).

  • — Portevin, 1929: 341 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1441 (Quedius; catalog).

  • — West, 1937: 37 (Quedius; Faroe Islands).

  • — Wüsthoff, 1938a: fig. 21 (Quedius; aedeagal illustration).

  • — Bernhauer, 1940c: 5 (Quedius; Azores).

  • — Paulian, 1941: 263 (Quedius; larval characters).

  • — Méquignon, 1942a: 22 (Quedius; Azores).

  • — Hinton, 1945: 70 (Quedius; characters).

  • — Fagel, 1948a (Quedius; characters; discussion of identity).

  • — Jeannel and Jarrige, 1949: 378 (Quedius; cave species; Germany).

  • — V. Hansen, 1952: 140 (Quedius; subgenus Quedius; characters; Denmark).

  • — Smetana, 1958: 361 (Quedius; subgenus Quedius; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 133 (Quedius; subgenus Quedius; characters; Europe; Mediterranean region; Turkey; Caucasus).

  • — Palm, 1963b: 139 (Quedius; subgenus Quedius; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 211 (Quedius; subgenus Quedius; characters; central Europe).

  • — Smetana, 1964d: 79 (Quedius; subgenus Quedius; Czechoslovakia).

  • — Horion, 1965: 272 (Quedius; subgenus Quedius; all Europe; North Africa; Turkey; Caucasus; Siberia; northern China).

  • — Pototskaia, 1967: 84, 87 (Quedius; larval characters).

  • — Coiffait, 1968c: 505 (Quedius; Corsica).

  • — Kasule, 1968a: 53 (Quedius; life history).

  • — Korge, 1968: 52 (Quedius; characters; notes).

  • — Szujecki, 1968a: 735 (Quedius; Poland).

  • — Frank, 1969: 266 (Quedius; biological notes; England).

  • — Kasule, 1970: 63 (Quedius; larval characters).

  • — Bordoni, 1974b: 12 (Quedius; subgenus Quedius; characters).

  • — Osella and Zanetti, 1975: 120 (Quedius; collected from nest of Talpa europaea; notes; Italy).

  • — Smetana, 1976d: 22 (Quedius; subgenus Quedius; Siberia).

  • — Bordoni, 1976: 238 (Quedius; Turkey).

  • — Bordoni, 1976b: 89 (Quedius; characters; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 190 (Quedius; subgenus Quedius; characters; Europe; Britain; Scandinavia; North Africa; Asia Minor).

  • — Smetana, 1978b: 85 (Quedius; subgenus Quedius; Siberia).

  • — Topp, 1979: 19 (Quedius; development; diapause).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • — Tóth, 1984: 117 (Quedius; characters; Hungary).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Borges, 1990: Table II (Quedius; Azores).

  • — Nowosad, 1990: 143 (Quedius; collected from mole nest; notes; Poland).

  • — Allen, 1990b: 4 (Quedius; collecting notes; Britain).

  • — Whitehead, 1991: 6 (Quedius; collecting notes; Britain).

  • — Gusarov, 1991a: 9 (Quedius; notes).

  • — Gusarov, 1993: 72 (Quedius; notes; lectotype designation).

  • — Welch, 1993: 229 (Quedius; ovariole number and ovary structure).

  • — Smetana, 1993b: 50 (Quedius; subgenus Quedius; Czech Republic; Slovakia).

  • — Hodge and Jones, 1995: 42 (Quedius; characters; Britain).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Staniec, 1996: 117 (Quedius; pupal characters).

  • — Owen, 1997a: 149 (Quedius; collecting sites; Ireland).

  • — Staniec, 1999a: 52 (Quedius; characters of pupa; Poland).

  • Distribution: Tunisia, Algeria, Azores, Europe, Russia, Azerbaijan, Turkey.

gracilis Stephens, 1832: 215 (Quedius; Type locality: London).

  • — Curtis, 1837: pl. 638 (Quedius; characters; Britain).

  • — Stephens, 1839: 390 (Quedius; variety of tristis; characters; England).

  • — Hardy, 1851: 33 (Quedius; synonym of fuliginosus).

  • — G. Waterhouse, 1858: 23 (Quedius; synonym of fuliginosus).

  • — Porta, 1907: 128 (Quedius; synonym of fuliginosus).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fuliginosus).

  • — Smetana, 1958: 361 (Quedius; synonym of fuliginosus).

picicornis Stephens, 1832: 215 (Quedius; Type locality: London).

  • — Curtis, 1829: 25 (Quedius; nomen nudum; catalog; Britain).

  • — Stephens, 1839: 390 (Quedius; variety of tristis; characters; England).

  • — Hardy, 1851: 33 (Quedius; synonym of fuliginosus).

  • — G. Waterhouse, 1858: 23 (Quedius; synonym of fuliginosus).

  • — Porta, 1907: 128 (Quedius; synonym of fuliginosus).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fuliginosus).

  • — Smetana, 1958: 361 (Quedius; synonym of fuliginosus).

  • — Coiffait, 1978: 191 (Quedius; synonym of fuliginosus).

granulipennis Motschulsky, 1858: 656 (Quedius; Type locality: Autriche; [Note: See lectotype designation by Gusarov, 1993]).

  • — Fauvel, 1874: 289 [= 1874a: 515] (Quedius; synonym of fuliginosus).

  • — Ganglbauer, 1895: 403 (Quedius; synonym of fuliginosus).

  • — Porta, 1907: 128 (Quedius; synonym of fuliginosus).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fuliginosus).

  • — Smetana, 1958: 361 (Quedius; synonym of fuliginosus).

  • — Gusarov, 1993: 72 (Quedius; lectotype designation; verification of synonymy with fuliginosus).

  • — Coiffait, 1978: 191 (Quedius; synonym of fuliginosus).

jelineki Krása, 1904: 81 (Quedius; variety of fuliginosus; Type locality: Vrané n. Vlt [Bohemia]).

  • — Scheerpeltz, 1933: 1441 (Quedius; variety of fuliginosus).

  • — Smetana, 1958: 361 (Quedius; synonym of fuliginosus).

  • — Smetana, 1962b: 133 (Quedius; aberration of fuliginosus).

  • — Coiffait, 1978: 191 (Quedius; subgenus Quedius; variety of fulginosus; characters).

  • — Tóth, 1984: 118 (Quedius; variety of fuliginosus; characters).

sublateralis Reitter, 1909: 111 (Quedius; aberration of fuliginosus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Paskau).

  • — Gridelli, 1924d: 79 (Quedius; variety of fuliginosus).

  • — Scheerpeltz, 1933: 1441 (Quedius; variety of fuliginosus).

  • — Smetana, 1958: 361 (Quedius; synonym of fuliginosus).

  • — Smetana, 1962b: 133 (Quedius; aberration of fuliginosus).

  • — Lohse, 1964: 211 (Quedius; subgenus Quedius; aberration of fuliginosus; characters).

  • — Coiffait, 1978: 191 (Quedius; subgenus Quedius; variety of fuliginosus; characters).

  • — Tóth, 1984: 118 (Quedius; variety of fuliginosus; characters; Hungary).

impunctifrons Delahon, 1915: 395 (Quedius; variety of fuliginosus; Type locality: Mark Brandenburg).

  • — Gridelli, 1924d: 79 (Quedius; variety of fuliginosus).

  • — Scheerpeltz, 1933: 1441 (Quedius; aberration of fuliginosus).

  • — Smetana, 1958: 361 (Quedius; synonym of fuliginosus).

  • — Coiffait, 1978: 191 (Quedius; synonym of fuliginosus variety jelineki).

molochinicolor Roubal, 1931: 1 (Quedius; form of fuliginosus; Type locality: RCS.: Harmanec).

  • — Scholz, 1933: 95 (Quedius; synonym of badius).

  • — Smetana, 1958: 361 (Quedius; synonym of fuliginosus).

  • — Smetana, 1962b: 133 (Quedius; aberration of fuliginosus).

  • — Lohse, 1964: 211 (Quedius; subgenus Quedius; aberration of fuliginosus; characters).

  • — Szujecki, 1968a: 735 (Quedius; aberration of fuliginosus; Poland).

  • — Coiffait, 1978: 191 (Quedius; subgenus Quedius; variety of fuliginosus; characters).

  • — Tóth, 1984: 118 (Quedius; variety of fuliginosus; characters; Hungary).

badius Scholz, 1931: 137 (Quedius; aberration of fuliginosus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Liegnitz).

  • — Scheerpeltz, 1933: 1441 (Quedius; synonym of fuliginosus).

  • — Smetana, 1958: 361 (Quedius; synonym of fuliginosus).

  • — Smetana, 1962b: 133 (Quedius; synonym of fuliginosus).

  • — Coiffait, 1978: 191 (Quedius; synonym of fuliginosus variety molochinicolor).

scholzi Scheerpeltz, 1933: 1441 (Quedius; replacement name for badius Scholz; [Note: This name was proposed as a replacement for an aberration that was preoccupied but the replacement name was cited as a variety and so is available]; variety of fuliginosus).

  • — Smetana, 1958: 361 (Quedius; synonym of fuliginosus).

  • — Smetana, 1962b: 133 (Quedius; synonym of fuliginosus).

  • — Coiffait, 1978: 191 (Quedius; synonym of fuliginosus variety molochinicolor).

subfuliginosus Britten, 1944: 289, 290 (Quedius; Type locality: England: Cotterill Clough, Cheshire).

  • — Fagel, 1948a: 199 (Quedius; characters; cited as valid species).

  • — Smetana, 1958: 361 (Quedius; synonym of fuliginosus).

  • — Smetana, 1962b: 133 (Quedius; synonym of fuliginosus).

  • — Lohse, 1964: 211 (Quedius; synonym of fuliginosus).

  • — Horion, 1965: 272 (Quedius; synonym of fuliginosus).

  • — Pope, 1977: 31 (Quedius; synonym of fuliginosus).

  • — Coiffait, 1978: 191 (Quedius; synonym of fuliginosus).

fultoni Cameron, 1945d: 787 (Quedius; subgenus Ediquus; Type locality: New Zealand: Taieri).

  • Distribution: New Zealand.

fulvicollis Stephens, 1833: 244 (Raphirus; Type locality: Edinburgh).

  • — Stephens, 1839: 394 (Quedius; characters; England).

  • — Gemminger and Harold, 1868: 568 (Quedius; synonym of collaris).

  • — Fauvel, 1869: 486 (Quedius; valid species).

  • — Fauvel, 1874: 307 [= 1874a: 533] (Quedius; characters; Lake Baikal).

  • — Mulsant and Rey, 1876: 790 [= 1877: 646] (Quedius; subgenus Raphirus; [Note: Misidentification: Bernhauer and Schubert, 1916: 423, cited picipennis Mulsant and Rey as a synonym of fulvicollis; Mulsant and Rey did not describe the species as new, they attributed it to Heer]; characters; France).

  • — Horn, 1878: 164 (Quedius; [Note: Misidentified as hyperboreus according to Smetana, 1971: 226]; characters; Maine to Vancouver).

  • — Fowler, 1888: 244 (Quedius; subgenus Raphirus; characters; Britain).

  • — Ganglbauer, 1895: 412 (Quedius; subgenus Raphirus; characters; Thuringia; France; Scotland; Sweden; Lapland; Baikal; North America).

  • — Poppius, 1905: 9 (Quedius; Shetland Islands).

  • — Porta, 1907: 151 (Quedius; subgenus Raphirus; characters; Italy).

  • — Poppius, 1909: 29 (Quedius; subgenus Raphirus; Lena Valley, Siberia).

  • — Reitter, 1909: 115 (Quedius; subgenus Raphirus; characters; Germany).

  • — Blatchley, 1910: 372 (Quedius; [Note: Misidentified as hyperboreus according to Smetana, 1971: 226]; characters; Indiana).

  • — Casey, 1915: 419 (Raphirus; characters; Maine to Lake Superior).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; catalog).

  • — Van Dyck, 1921: 162 (Quedius; Pribilof Islands, Alaska).

  • — Gridelli, 1925: 163 (Quedius; subgenus Raphirus; characters).

  • — Bernhauer, 1926g: 2 (Quedius; Kamchatka region).

  • — Scheerpeltz, 1933: 1441 (Quedius; catalog).

  • — West, 1937: 38 (Quedius; Faroe Islands).

  • — Wüsthoff, 1938a: fig. 63(Quedius; aedeagal illustration).

  • — Tottenham, 1948a: 250 (Quedius; subgenus Arphirus; characters; Britain).

  • — V. Hansen, 1952: 154 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Smetana, 1958: 394 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Larsson and Gígja, 1959: 92 (Quedius; biological notes; Iceland).

  • — Smetana, 1962b: 150 (Quedius; subgenus Raphirus; characters; Europe).

  • — Palm, 1963b: 152 (Quedius; subgenus Raphirus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 219 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Smetana, 1964: 39 (Quedius; subgenus Raphirus; [Note: Misidentified as hyperboreus according to Smetana, 1971: 226]; characters; Alaska).

  • — Smetana, 1965a: 16 (Quedius; subgenus Raphirus; [Note: Misidentified as hyperboreus according to Smetana, 1971: 226]; Alaska).

  • — Smetana, 1965: 48 (Quedius; subgenus Raphirus; characters; Labrador; St. Pierre; Newfoundland).

  • — Horion, 1965: 304 (Quedius; subgenus Raphirus; widespread in northern Europe; Poland; Czechoslovakia; Romania; Bosnia; Croatia; Istria; Italy; Switzerland; Austria; France; Belgium; Holland ?; Germany).

  • — Smetana, 1966: 332 (Quedius; subgenus Raphirus; notes; Czechoslovakia).

  • — Smetana, 1971: 226 (Quedius; subgenus Raphirus; characters; Alberta; British Columbia; Labrador; Manitoba; Newfoundland; Ontario; Quebec; Yukon Territory; Alaska; Colorado; Maine; Maryland; Massachusetts; Michigan; Montana; New Hampshire; New York; Pennsylvania; Utah; Vermont; Washington; Wyoming).

  • — Smetana, 1971a: 1848 (Quedius; subgenus Raphirus; collecting notes; New Mexico).

  • — Smetana, 1973: 1430 (Quedius; notes; Alberta; Ontario; Quebec).

  • — Smetana, 1976c: 182 (Quedius; subgenus Raphirus; Ontario; Quebec).

  • — Smetana, 1976d: 26, 27 (Quedius; subgenus Raphirus; characters; Siberia).

  • — Shilov, 1975a: 377 [= 1975b: 99] (Quedius; Komi region of Russia).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 276 (Quedius; subgenus Raphirus; characters; Europe; Britain; France).

  • — Tóth, 1984: 129 (Quedius; characters).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; France; Benelux; Denmark; Sweden).

  • — Drugmand, 1988c: 209 (Quedius; characters).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Sadler and Dugmore, 1995: 144 (Quedius; habitat notes; Iceland).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Downie and Arnett, 1996: 393 (Quedius; subgenus Raphirus; characters; USA).

  • — Ryabukhin, 1999: 74 (Quedius; subgenus Raphirus; catalog; habitat; distribution; northeastern Russia).

  • — Eyre, Luff, and Lott, 2000: 34 (Quedius; Britain).

  • Distribution: Iceland, Europe, Russia, Canada, USA.

picipennis Heer, 1839: 279 (Philonthus; [preoccupied]; Type locality: Val Emet; Urscheinalp; Lavirumsalp).

  • — Kraatz, 1857d: 515 (Quedius; synonym of attenuatus).

  • — Fauvel, 1874: 306 [= 1874a: 532] (Quedius; synonym of attenuatus). Raphirus; characters; France).

  • — Fowler, 1888: 244 (Quedius; subgenus Raphirus; variety of attenuatus; characters; Britain).

  • — Porta, 1907: 149 (Quedius; subgenus Raphirus; characters; Italy).

  • — Poppius, 1909: 28 (Quedius; subgenus Raphirus; Lena Valley, Siberia).

  • — Reitter, 1909: 115 (Quedius; subgenus Raphirus; characters; Germany).

  • — Poppius, 1913: 10 (Quedius; Iceland).

  • — Johansen, 1914: 370 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Fiori, 1915: 16 (Quedius; Italy).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; synonym of nitipennis).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; synonym of scribae).

  • — Gridelli, 1925: 160 (Quedius; synonym of scribae).

  • — Scheerpeltz, 1933: 1461 (Quedius; synonym of scribae).

  • — Smetana, 1962b: 150 (Quedius; synonym of scribae).

  • — Coiffait, 1978: 268 (Quedius; synonym of scribae). — Assing, 1999b: 37 (Quedius; lectotype designation; synonym of fulvipennis).

nigricollis W. Kolbe, 1909: 20 (Quedius; aberration of fulvicollis; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fulvicollis).

  • — Gridelli, 1925: 163 (Quedius; aberration of fulvicollis).

  • — Scheerpeltz, 1933: 1442 (Quedius; aberration of fulvicollis).

  • — Smetana, 1958: 395 (Quedius; aberration of fulvicollis).

  • — Smetana, 1962b: 150 (Quedius; aberration of fulvicollis; characters).

  • — Smetana, 1971: 226 (Quedius; synonym of fulvicollis).

  • — Coiffait, 1978: 276 (Quedius; subgenus Raphirus; variety of fulvicollis; characters).

pugetanus Casey, 1915: 419 (Raphirus; Type locality: Washington State: Soda Springs).

  • — Scheerpeltz, 1933: 1458 (Quedius; catalog).

  • — Hatch, 1957: 229 (Quedius; subgenus Raphirus; characters; British Columbia; Washington).

  • — Smetana, 1971: 226 (Quedius; synonym of fulvicollis).

raphiroides Hatch, 1957: 219 (Quedius; subgenus Distichalius; Type locality: British Columbia: Barkerville).

  • — Smetana, 1971: 226 (Quedius; synonym of fulvicollis).

fulvipennis Hochhuth, 1851: 28 (Quedius; Type locality: Dahurien).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; catalog).

  • — Gridelli, 1925: 168 (Quedius; characters).

  • — Scheerpeltz, 1933: 1442 (Quedius; catalog).

  • Distribution: Russia.

fumatus Stephens, 1833: 245 (Raphirus; Type locality: Suffolk; near London; Devonshire).

  • — Stephens, 1839: 394 (Quedius; characters; England).

  • — Erichson, 1839a: 492 (Quedius; [Note: Misidentification: Bernhauer and Schubert, 1916: 424, cited praecox Erichson as a synonym of fumatus Stephens; Erichson did not describe the species as new, he attributed it to Gravenhorst]; characters; Germany).

  • — Fauvel, 1874: 296 [= 1874a: 522] (Quedius; characters; France; Europe; Algeria).

  • — Fowler, 1888: 239 (Quedius; subgenus Sauridus; characters; Britain).

  • — Ganglbauer, 1895: 406 (Quedius; subgenus Sauridus; characters; Europe; Algeria).

  • — Sainte-Claire Deville, 1906: 102 (Quedius; Corsica).

  • — Porta, 1907: 133 (Quedius; subgenus Sauridus; characters; Italy).

  • — Reitter, 1909: 112 (Quedius; subgenus Sauridus; characters; Germany).

  • — Johansen, 1914: 365 (Quedius; subgenus Sauridus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • — Munster, 1923: 197 (Quedius; subgenus Sauridus; Fennoscandia).

  • — Gridelli, 1925: 137 (Quedius; subgenus Raphirus; characters; Italy).

  • — Portevin, 1929: 341 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1442 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 42 (Quedius; aedeagal illustration).

  • — Jeannel and Jarrige, 1949: 381 (Quedius; subgenus Raphirus; cave species; France).

  • — V. Hansen, 1952: 144 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Smetana, 1958: 379 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1961d: 134 (Quedius; subgenus Raphirus; Italy).

  • — Smetana, 1962b: 139 (Quedius; subgenus Raphirus; characters; Europe; North Africa).

  • — Coiffait, 1963a: 379 (Quedius; subgenus Sauridus; characters).

  • — Palm, 1963b: 146 (Quedius; subgenus Raphirus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 214 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Focarile, 1964: 60 (Quedius; Italy).

  • — Horion, 1965: 287 (Quedius; subgenus Raphirus; Algeria; Portugal; Spain; Corsia; Sicily; Italy; Slovenia; Dalmatia; Bosnia Hercegovina; Romania; Hungary; Czechoslovakia; Poland; France; Switzerland; Austria; Germany; Belgium; Holland; Britain; Norway; Sweden; Denmark).

  • — Coiffait, 1968c: 505 (Quedius; Corsica).

  • — Kasule, 1968a: 75 (Quedius; life history).

  • — Kasule, 1970: 63 (Quedius; larval characters).

  • — Shilov, 1975a: 376 [= 1975b: 99] (Quedius; Komi region of Russia).

  • — Legner and Moore, 1977: 176 (Quedius; found under bark).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 243 (Quedius; subgenus Sauridus; characters; Europe; Britain; Scandinavia; Algeria; Tunisia).

  • — Hinton, 1981: 662 (Quedius; characters of egg).

  • — Tóth, 1984: 136 (Quedius; characters; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Outerelo, Gamarra, and Salgado, 1998: 130 (Quedius; subgenus Sauridus; collected from cave; Spain).

  • — Staniec, 1999a: 49 (Quedius; characters of pupa; Poland).

  • — Owen, 2000: 252 (Quedius; pitfall trap collections from underground at roots of trees; Britain).

  • Distribution: Tunisia, Algeria, Europe, Russia.

peltatus Erichson, 1840: 540 (Quedius; proposed for Erichson's, 1839a misidentification of praecox; Type locality: Germania; Hungaria).

  • — Redtenbacher, 1849: 825 (Philonthus; characters; Austria).

  • — Hardy, 1851: 33 (Quedius; notes; England).

  • — G. Waterhouse, 1858: 23 (Quedius; synonym of fumatus).

  • — Fairmaire and Laboulbène, 1856: 543 (Quedius; characters; France).

  • — Kraatz, 1857d: 507 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 204 (Quedius; characters; Austria).

  • — Fairmaire and Coquerel, 1860: 154 (Quedius; Algeria).

  • — Thomson, 1867: 169 (Raphirus; characters; Scandinavia).

  • — Gemminger and Harold, 1868: 569 (Quedius; synonym of fumatus).

  • — Fauvel, 1874: 296 [= 1874a: 522] (Quedius; synonym of fumatus).

  • — Redtenbacher, 1874: 202 (Quedius; characters; Austria).

  • — Mulsant and Rey, 1876: 715 [= 1877: 571] (Quedius; subgenus Sauridus; characters; France).

  • — Fowler, 1888: 239 (Quedius; subgenus Sauridus; synonym of fumatus).

  • — Ganglbauer, 1895: 406 (Quedius; synonym of fumatus).

  • — Porta, 1907: 133 (Quedius; synonym of fumatus).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; synonym of fumatus).

  • — Gridelli, 1925: 137 (Quedius; synonym of fumatus).

  • — Scheerpeltz, 1933: 1442 (Quedius; synonym of fumatus).

  • — Smetana, 1958: 379 (Quedius; synonym of fumatus).

  • — Coiffait, 1978: 243 (Quedius; synonym of fumatus).

irideus Miller, 1851: 10, 110 (Quedius; Type locality: Dornbach bei Wien).

  • — Kraatz, 1857d: 507 (Quedius; synonym of peltatus).

  • — Gemminger and Harold, 1868: 569 (Quedius; synonym of fumatus).

  • — Ganglbauer, 1895: 406 (Quedius; synonym of fumatus).

  • — Porta, 1907: 133 (Quedius; synonym of fumatus).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; synonym of fumatus).

  • — Gridelli, 1925: 137 (Quedius; synonym of fumatus).

  • — Smetana, 1958: 379 (Quedius; synonym of fumatus).

  • — Coiffait, 1978: 243 (Quedius; synonym of fumatus).

flavomarginatus Gridelli, 1925: 137 (Quedius; variety of fumatus; Type locality: Sicilia: Ficuzza).

  • — Scheerpeltz, 1933: 1442 (Quedius; synonym of fumatus).

  • — Fagel, 1957e: 333 (Quedius; subgenus Sauridus; subspecies of fumatus; notes).

  • — Coiffait, 1978: 243 (Quedius; subgenus Sauridus; variety of fumatus; characters).

  • — Tóth, 1984: 136 (Quedius; variety of fumatus; characters; Hungary).

funebris Coiffait, 1971 [Note: Nomen nudum], see: balticus Korge, 1960.

fuscatus Broun, 1893: 1027 (Quedius; Type locality: Mount Arthur).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • Distribution: New Zealand.

fusicornis Luze, 1904: 101 (Quedius; subgenus Microsaurus; Type locality: Seravschan: Putschin-Pass, Boschara).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • — Gridelli, 1924d: 69 (Quedius; characters; Turkestan).

  • — Scheerpeltz, 1933: 1442 (Quedius; catalog).

  • Distribution: Uzbekistan.

fuscipennis Block, 1799, see: fulgidus Fabricius, 1793.

fuscipennis Stephens, 1832, see: boops Gravenhorst, 1802.

fuscipes Stephens, 1833, see: scintillans Gravenhorst, 1806.

gaarho Smetana, 1988: 246 (Quedius; subgenus Raphirus; Type locality: Nepal: Parbat Distr., Ridge E Ghoropani Pass, 3150 m).

  • — Smetana, 1992b: 4 (Quedius; subgenus Raphirus; Nepal).

  • Distribution: Nepal.

galaecianus Outerelo, 1976: 262 (Quedius; subgenus Quedionthus; Type locality: Moreda del Caurel, Sierra del Caurel, Lugo, 720 m).

  • — Coiffait, 1978: 204 (Quedius; subgenus Quediops; characters; Spain).

  • Distribution: Spain.

gang Smetana, 1996c: 232 (Quedius; subgenus Raphirus; Type locality: China: Kuatun, 2300 m, 27,40 n. Br. 117, 40 ö, Fukien).

  • Distribution: China.

gardneri Cameron, 1932: 292 (Quedius; subgenus Sauridus; Type locality: Darjeeling Dist.: Lepchajagat, alt. 7000 feet; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1442 (Quedius; catalog).

  • — Smetana, 1988: 295 (Quedius; subgenus Raphirus; lectotype designation; characters; Bhutan; West Bengal).

  • Distribution: India, Bhutan.

coeruleus Coiffait, 1977b: 241 (Quedius; subgenus Sauridus; Type locality: Bhutan: Nobding, 40 km Ouest de Wangdi-Phodrang, 2800 m).

  • — Smetana, 1988: 295 (Quedius; synonym of gardneri).

gautardi Fauvel, 1898: 100 (Quedius; Type locality: Canaries: Tenerife).

  • — Fauvel, 1902b: 114 (Quedius; Canaries).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • — Gridelli, 1924d: 39 (Quedius; subgenus Microsaurus; characters; Canaries).

  • — Scheerpeltz, 1933: 1431 (Quedius; synonym of angustifrons).

  • — Coiffait, 1967e: 395 (Quedius; subgenus Microsaurus; valid species; characters; Canary Islands).

  • — Coiffait, 1978: 176 (Quedius; subgenus Microsaurus; characters; Canary Islands).

  • Distribution: Canary Islands.

gazellensis Last, 1987 [Note: Nomen nudum], see: cyaneorufus Fauvel, 1878.

gemellus Eppelsheim, 1889d: 18 (Quedius; subgenus Sauridus; Type locality: Circassien).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • — Gridelli, 1922b: 130 (Quedius; characters; Circassia; Caucasus; Turkey).

  • — Gridelli, 1925: 122 (Quedius; subgenus Raphirus; characters; Caucasus).

  • — Scheerpeltz, 1933: 1442 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 36 (Quedius; aedeagal illustration).

  • — Coiffait, 1963a: 387 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1967e: 416 (Quedius; subgenus Sauridus; characters; Asia Minor; Caucasus; Turkestan).

  • — Coiffait, 1978: 257 (Quedius; subgenus Sauridus; characters; Asia Minor; Caucasus; Turkestan).

  • Distribution: Russia, Caucasus, Turkey, “Turkestan”.

bithynicus Fagel, 1968: 124 (Quedius; subgenus Sauridus; subspecies of gemellus; Type locality: Anatolie occidentale: Abant Dagh, 1450 m).

  • — Korge, 1971a: 52 (Quedius; subgenus Sauridus; subspecies of gemellus; Turkey).

  • — Coiffait, 1978: 257 (Quedius; subgenus Sauridus; subspecies of gemellus; characters).

germaini Bernhauer, 1917h: 90 (Quedius; subgenus Euryquedius; Type locality: Bolivien: Yuracaris).

  • — Scheerpeltz, 1933: 1442 (Quedius; catalog).

  • Distribution: Bolivia.

germanorum Smetana, 1997d: 457 (Quedius; subgenus Microsaurus; Type locality: china: Shaanzi, Qin Ling Shan 108.47 E, 33.51 N, Mountain W Pass at Autoroute km 70, 47 km S Xian 2500–2600 m).

  • Distribution: China.

gestroi Gridelli, 1922, see: humeralis Stephens, 1832.

ghilarovi Coiffait, 1967e: 405 (Quedius; subgenus Sauridus; Type locality: Maykop, Russie méridionale).

  • — Pototskaia, 1967: 83, 93 (Quedius; larval characters).

  • — Coiffait, 1978: 240 (Quedius; subgenus Sauridus; characters; Russia).

  • Distribution: Russia.

glaber O. Müller, 1776, see: plagiatus Mannerheim, 1843.

goang Smetana, 1995: 75 (Quedius; subgenus Raphirus; Type locality: Taiwan: Nantou Hsien, Houhuanshan Kuenyang, 3050 m).

  • Distribution: Taiwan.

goropanus Smetana, 1975a: 327 (Quedius; subgenus Microsaurus; Type locality: Central Nepal: near Goropani, W. Pokhara).

  • — Smetana, 1988: 220 (Quedius; subgenus Microsaurus; characters; Nepal).

  • Distribution: Nepal.

gracilicornis Heer, 1839, see: picipes Mannerheim, 1830.

gracilicornis Coiffait, 1977 [Note: Nomen nudum], see: tschinganensis Coiffait, 1969.

gracilis Stephens, 1832, see: fuliginosus Gravenhorst, 1802.

graciliventris Sharp, 1884: 331 (Quedius; Type locality: Panama, Volcan de Chiriqui between 2000 and 6000 feet).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1933: 1442 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 177 (Quedius; subgenus Cyrtoquedius; characters in key; Panama).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • Distribution: Panama.

grandiceps Kraatz, 1879, see: Indoquedius.

granulipennis Motschulsky, 1858, see: fuliginosus Gravenhorst, 1802.

gridellii Scheerpeltz, 1933: 1442 (Quedius; replacement name for picicornis Gridelli).

  • — Coiffait, 1978: 262 (Quedius; synonym of luteomaculatus).

  • — Herman, 2001: 30 (Quedius; valid name for species).

  • Distribution: Iran, Turkey.

picicornis Gridelli, 1922b: 128, 131 (Quedius; subgenus Sauridus; [preoccupied]; Type locality: Persia merid.).

  • — Gridelli, 1925: 127 (Quedius; subgenus Raphirus; characters; Persia).

  • — Scheerpeltz, 1933: 1442 (Quedius; synonym of gridellii).

  • — Coiffait, 1963a: 387 (Quedius; subgenus Sauridus; probably synonym of luteomaculatus).

  • — Coiffait, 1978: 262 (Quedius; synonym of luteomaculatus).

luteomaculatus Scheerpeltz, 1958a: 23 (Quedius; subgenus Raphirus; Type locality: bei Ahlat, Nordwest-Ufer des Van-Sees in Armenien).

  • — Coiffait, 1963a: 387 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 262 (Quedius; subgenus Sauridus; characters; Turkey).

  • — Herman, 2001: 30 (Quedius; synonym of gridellii).

gridellii Jarrige, 1952, see: adanensis Coiffait, 1967.

griffinae Hatch, 1957: 225 (Quedius; subgenus Microsaurus; Type locality: Washington: Olympia).

  • — Smetana, 1971: 70 (Quedius; subgenus Microsaurus; characters; British Columbia; California; Washington).

  • — Smetana, 1978a: 824 (Quedius; subgenus Microsaurus; collecting notes; Oregon).

  • — Smetana, 1990d: 97 (Quedius; California; Oregon; Washington).

  • Distribution: Canada, USA.

groenlandicus Zetterstedt, 1838, see: mesomelinus Marsham, 1802.

grootaerti Drugmand, 1988c: 209 (Quedius; subgenus Raphirus; Type locality: Liban: Nebeh Safa).

  • Distribution: Lebanon.

grouziacus Coiffait, 1969: 45 (Quedius; subgenus Sauridus; Type locality: Lagodeki Zapovedink, Georgie [Note: Correct spelling is Zapovednik]).

  • — Coiffait, 1978: 207 (Quedius; subgenus Sauridus; characters; Georgia; Anatolia).

  • — Coiffait, 1978d: 165 (Quedius; Turkey).

  • Distribution: Georgia, Turkey.

borjoniensis Coiffait, 1969: 46 (Quedius; subgenus Sauridus; variety of grouziacus; [Note: Nomen nudum (Article 15.2)]; cited from Borjoni, Akaldaba, Georgie).

  • — Coiffait, 1978: 207 (Quedius; subgenus Sauridus; variety of grouziacus; characters).

  • — Coiffait, 1978d: 165 (Quedius; variety of grouziacus; Turkey).

rizensis Coiffait, 1977c: 136 (Quedius; subgenus Sauridus; variety of grouziacus; [Note: Nomen nudum (Article 15.2)]; cited from Rize, Nord-Est de l'Anatolie).

  • — Coiffait, 1978: 207 (Quedius; subgenus Sauridus; variety of grouziacus; characters).

gueorguievi Coiffait, 1967e: 399 (Quedius; subgenus Sauridus; Type locality: Grotte Porojnata Dupka, Bulgarie).

  • — Coiffait, 1978: 224 (Quedius; subgenus Sauridus; characters; Bulgaria).

  • Distribution: Bulgaria.

haafi Scheerpeltz, 1956c: 1107 (Quedius; subgenus Raphirus; Type locality: Cypern: Pedhoulas, 3600 feet).

  • — Coiffait, 1978: 283 (Quedius; subgenus Raphirus; characters; Cyprus).

  • Distribution: Cyprus.

haberfelneri Eppelsheim, 1891: 200 (Quedius; subgenus Raphirus; Type locality: Lunz in Nieder-Oesterreich).

  • — Ganglbauer, 1895: 410 (Quedius; subgenus Raphirus; characters; Austria).

  • — Bernhauer, 1899: 22 (Quedius; notes).

  • — Bernhauer, 1900: 46 (Quedius; notes).

  • — Reitter, 1909: 114 (Quedius; subgenus Raphirus; characters; Germany).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • — Gridelli, 1925: 143 (Quedius; subgenus Raphirus; characters).

  • — Scheerpeltz, 1933: 1442 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 47 (Quedius; cited as haberfellneri; aedeagal illustration).

  • — Smetana, 1958: 382 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 138 (Quedius; subgenus Raphirus; characters; Europe).

  • — Lohse, 1964: 214 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Smetana, 1964c: 168 (Quedius; subgenus Raphirus; Czechoslovakia).

  • — Horion, 1965: 290 (Quedius; subgenus Raphirus; Austria; Germany; Italy; Switzerland; France; Czech Republic; Slovakia; Poland; Ukraine; Russia).

  • — Coiffait, 1967e: 419 (Quedius; subgenus Microquedius).

  • — Bordoni, 1974a: 205 (Quedius; subgenus Microquedius; characters; Italy).

  • — Coiffait, 1978: 200 (Quedius; subgenus Microquedius; characters; Alps).

  • — Zanetti, 1979a: 542 (Quedius; characters; Italy).

  • — Tóth, 1984: 122 (Quedius; characters; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic ?; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Europe.

decoratus Petri, 1902: 153 (Quedius; Type locality: Negoi; Paringulgebirge).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; synonym of auricomus).

  • — Scheerpeltz, 1933: 1443 (Quedius; synonym of haberfelneri).

  • — Smetana, 1958: 383 (Quedius; synonym of haberfelneri).

  • — Coiffait, 1978: 200 (Quedius; synonym of haberfelneri).

brevipennis Ihssen, 1939a: 311 (Quedius; variety of haberfelneri; [preoccupied]; Type locality: bayerische Hochgebirge).

  • — Coiffait, 1978: 200 (Quedius; subgenus Microquedius; variety of haberfelneri; characters).

  • — Tóth, 1984: 122 (Quedius; aberration of haberfelneri; characters).

hackeri Lea, 1925: 245 (Quedius; Type locality: Queensland: National Park).

  • — Scheerpeltz, 1933: 1443 (Quedius; catalog).

  • — Oke, 1933: 123, 124 (Quedius; tarsal characters; Australia).

  • Distribution: Australia.

haemon Smetana, 1995f: 239 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Gongga Shan, 2850 m).

  • — Smetana, 1997d: 453 (Quedius; subgenus Microsaurus; characters of female; collecting notes; China).

  • Distribution: China.

haemopterus Stephens, 1832, see: fulgidus Fabricius, 1793.

haemorrhoidalis Stephens, 1832, see: fulgidus Fabricius, 1793.

haemorrhous Stephens, 1832, see: cruentus Olivier, 1795.

hailuogou Smetana, 1999b: 232 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Gongga Shan, above Camp 3, 3050 m).

  • Distribution: China.

hakusanus Watanabe, 1987: 326 (Quedius; subgenus Microsaurus; Type locality: Japan: Honshu: Midori-ga-ike on Mt. Hakusan, 2500 m).

  • Distribution: Japan.

hallianus Broun, 1917: 372 (Quedius; Type locality: Dyer's Pass, near Christchurch).

  • — Scheerpeltz, 1933: 1443 (Quedius; cited as hollianus; catalog).

  • Distribution: New Zealand.

hammianus Sharp, 1911, see: simplicifrons Fairmaire, 1861.

hariyo Smetana, 1988: 291 (Quedius; subgenus Raphirus; Type locality: Nepal: Khandbari District, above Sheduwa, 3000 m).

  • — Smetana, 1998b: 110 (Quedius; subgenus Raphirus; characters).

  • Distribution: Nepal.

hauseri Bernhauer, 1918a: 94 (Quedius; subgenus Raphirus; Type locality: Turkestan. Karateghingebirge: Baldschuan, 924 m; Sary-pul, 1482 m; Ghissargebirge; Ost-Buchara: Tschitschantan, Karatag, und Repetek).

  • — Gridelli, 1925: 154 (Quedius; subgenus Raphirus; characters).

  • — Scheerpeltz, 1933: 1443 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 56 (Quedius; aedeagal illustration).

  • — Coiffait, 1978: 264 (Quedius; subgenus Raphirus; characters; Transcaspian region).

  • — Tronquet, 1981: 71 (Quedius; Afghanistan).

  • Distribution: Afghanistan, Turkmenistan, Kyrgyzstan, Uzbekistan, Tajikistan.

peneckei Bernhauer, 1918a: 95 (Quedius; variety of hauseri; Type locality: Tienschangebirge: Przewalsk, Karakoltal).

  • — Gridelli, 1925: 154 (Quedius; variety of hauseri).

  • — Scheerpeltz, 1933: 1443 (Quedius; variety of hauseri).

  • — Coiffait, 1978: 264 (Quedius; subgenus Raphirus; variety of hauseri; characters).

heeri Blackwelder, 1944, see: Loncovilius.

hei Smetana, 1995: 29 (Quedius; subgenus Microsaurus; Type locality: Taiwan: Sun Kang, alt. 220 m, Nan two Hsien [Note: Correct spelling is Nantou).

  • Distribution: Taiwan.

heidenreichi Heinemann, 1910, see: puncticollis Thomson, 1867.

heidenreichi Bernhauer, 1914, see: puncticollis Thomson, 1867.

heinzi Korge, 1964: 117 (Quedius; subgenus Quedionuchus; Type locality: Ilgaz Dag).

  • — Korge, 1971a: 47 (Quedius; subgenus Quedionuchus; characters; Turkey).

  • — Coiffait, 1978: 187 (Quedius; subgenus Distichalius; characters; Anatolia).

  • Distribution: Turkey.

helenae Casey, 1915, see: nanulus Casey, 1915.

henroti Coiffait, 1970e: 66 (Quedius; subgenus Raphirus; Type locality: Mt. Olimbos, 1200 m, Ile de Lesbos, Grèce).

  • — Coiffait, 1978: 269 (Quedius; subgenus Raphirus; characters; Greece).

  • Distribution: Greece.

hermonensis Coiffait, 1963a: 393, 417 (Quedius; subgenus Sauridus; Type locality: Liban: Hasbaya).

  • — Coiffait, 1978: 244 (Quedius; subgenus Sauridus; characters; Lebanon).

  • — Smetana, 1978: 124 (Quedius; subgenus Raphirus; collecting notes; Israel).

  • Distribution: Lebanon, Israel.

heterodoxus Eppelsheim, 1890a: 208 (Quedius; subgenus Ediquus; Type locality: Bohemia).

  • — Ganglbauer, 1895: 396 (Quedius; subgenus Ediquus; characters; Europe).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • — Gridelli, 1924d: 35 (Quedius; subgenus Microsaurus; characters; Bohemia).

  • — Scheerpeltz, 1933: 1443 (Quedius; catalog).

  • — Roubal, 1941b: 122, 126 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Smetana, 1958: 343 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 154 (Quedius; subgenus Microsaurus; characters).

  • — Lohse, 1964: 207 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Horion, 1965: 257 (Quedius; subgenus Microsaurus; Bosnia; Czechoslovakia; Poland).

  • — Coiffait, 1978: 156 (Quedius; subgenus Microsaurus; characters; Bohemia; Bosnia).

  • — Tóth, 1984: 109 (Quedius; characters).

  • — Lucht, 1987: 109 (Quedius; checklist; Czechoslovakia; Austria).

  • — Lohse and Lucht, 1989: 171 (Quedius; Austria).

  • — Smetana, 1993b: 49 (Quedius; subgenus Microsaurus; Czech Republic).

  • Distribution: Europe.

heterogaster Cameron, 1944, see: muscicola Cameron, 1932.

higonis Uéno and Watanabe, 1970: 14 (Quedius; subgenus Microsaurus; Type locality: Japan: at the southwestern part of Kyushu: Kumamoto Prefecture: Fûshin-dô Cave, at Gokasé of Mizukoshi, in Mifuné-machi).

  • — Watanabe, 1996a: 14 (Quedius; cave collection; Japan).

  • Distribution: Japan.

hilaris Broun, 1909: 226 (Quedius; Type locality: Broken River; Invercargill).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • Distribution: New Zealand.

himalayicus Bernhauer, 1915b: 55 (Quedius; subgenus Sauridus; Type locality: Zentral-Himalaya: Kumaon, Bhowali).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • — Cameron, 1932: 293 (Quedius; subgenus Raphirus; characters; India).

  • — Scheerpeltz, 1933: 1443 (Quedius; catalog).

  • — Smetana, 1988: 285 (Quedius; subgenus Raphirus; characters; Nagaland; Himachal Pradesh; Uttar Pradesh; West Bengal; Nepal).

  • Distribution: Nepal, India.

hirticornis Sharp, 1889: 31 (Quedius; Type locality: Mitzusawa).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1443 (Quedius; catalog).

  • — Maruyama and Toyoda, 2000: 65 (Quedius; from nest of Lasius spathepus and Lasius fuliginosus; Japan).

  • Distribution: Japan.

hirtipennis Stephen, 1832, see: Philonthus.

hirtipennis Broun, 1915: 279 (Quedius; [preoccupied]; Type locality: Pudding Hill, near Methven).

  • — Scheerpeltz, 1933: 1443 (Quedius; catalog).

  • — Herman, 2001: 49 (Quedius; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: New Zealand.

hispanicus Bernhauer, 1898, see: simplicifrons Fairmaire, 1861.

hollandicus Bernhauer, 1908: 19 (Quedius; Type locality: Neuholland).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • Distribution: Australia.

holzschuhi Smetana, 1999b: 220 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Emeishan, 160 km SSW Chengdu, 1530–1700 m).

  • Distribution: China.

hoodi Hatch, 1957, see: criddlei Casey, 1915.

horni Hatch, 1957: 216 (Quedius; subgenus Quedius; Type locality: Washington: Snohomish Co.: L. Ballinger. Also cited from British Columbia, Oregon).

  • — Korge, 1962b: 333 (Quedius; characters; notes).

  • — Smetana, 1965: 39 (Quedius; subgenus Quedius; characters).

  • — Smetana, 1971: 146 (Quedius; subgenus Quedius; characters; British Columbia; Oregon; Washington).

  • Distribution: Canada, USA.

horsti Coiffait, 1978, see: smetanai Korge, 1971.

hospes Smetana, 1967d: 214 (Quedius; subgenus Microsaurus; Type locality: Mongolei: Cojbalsan aimak, 80 km NW von Cojbalsan, 700 m).

  • — Smetana 1975c: 176 (Quedius; Mongolia).

  • — Coiffait, 1978: 181 (Quedius; subgenus Microsaurus; characters; Mongolia).

  • Distribution: Mongolia.

howdeni Smetana, 1976: 114 (Quedius; subgenus Microsaurus; Type locality: Mexico: Durango: Buenos Aires, 9000 ft., 10 mi W. La Ciudad).

  • Distribution: Mexico.

huann Smetana, 1995: 98 (Quedius; subgenus Raphirus; Type locality: Taiwan: Kaohsiung Hsien: Kuanshan trail at Kaunshanchi Riv., 2400 m).

  • — Smetana, 1998b: 110 (Quedius; subgenus Raphirus; characters).

  • Distribution: Taiwan.

humeralis Stephens, 1832: 220 (Quedius; Type locality: Metropolitan district [London] near Barham).

  • — Curtis, 1829: 25 (Quedius; nomen nudum; catalog; Britain).

  • — Stephens, 1839: 391 (Quedius; characters; England).

  • — Thomson, 1867: 171 (Raphirus; [Note: Misidentification: Bernhauer and Schubert, 1916: 424, cited marginalis as a synonym of humeralis; Thomson did not describe the species as new, he attributed it to Kraatz]; characters; Scandinavia).

  • — Thomson, 1867a: 44 (Raphirus; [Note: Misidentification: Bernhauer and Schubert, 1916: 424, cited marginalis as a synonym of humeralis; Thomson did not describe the species as new, he attributed it to Kraatz]; characters; Scandinavia).

  • — Fauvel, 1874: 298 [= 1874a: 524] (Quedius; synonym of suturalis).

  • — Porta, 1907: 140 (Quedius; subgenus Sauridus; characters; Italy).

  • — Montandon, 1908: 80 (Quedius; Romania).

  • — Reitter, 1909: 113 (Quedius; subgenus Sauridus; characters; Germany).

  • — Johansen, 1914: 368 (Quedius; subgenus Sauridus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • — Gridelli, 1922b: 129, 130 (Quedius; subgenus Sauridus; characters; Italy; Sardinia; Turkey).

  • — Gridelli, 1925: 119 (Quedius; subgenus Raphirus; characters; Germany; Austria; Croatia; Italy; Transylvania; Bulgaria; Bosnia; Macedonia).

  • — Donisthorpe, 1930: 94 (Quedius; characters; Britain).

  • — Scheerpeltz, 1933: 1443 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 33 (Quedius; aedeagal illustration).

  • — Jeannel and Jarrige, 1949: 379 (Quedius; subgenus Raphirus; cave species; Herzegowina).

  • — Smetana, 1958: 374 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 146 (Quedius; subgenus Raphirus; characters; Europe).

  • — Coiffait, 1963a: 393 (Quedius; subgenus Sauridus; characters).

  • — Palm, 1963b: 149 (Quedius; subgenus Raphirus; characters; Sweden; Denmark).

  • — Lohse, 1964: 216 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Scheerpeltz, 1964a: 300 (Quedius; subgenus humeralis; Crete).

  • — Horion, 1965: 283 (Quedius; subgenus Raphirus; Ukraine; Poland; Czechoslovakia; Hungary; Romania; Slovenia; Croatia; Dalmatia; Bosnia Hercegovina; Bulgaria; Greece; Macedonia; Italy; Spain; France; Belgium; Holland; Britain; Denmark; Sweden; Germany; Austria).

  • — Pototskaia, 1967: 83, 89 (Quedius; larval characters).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Muona, 1979: 16 (Quedius; Sweden; Denmark).

  • — H. Franz, 1987: 70 (Quedius; Cyprus).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Lohse and Lucht, 1989: 175 (Quedius; characters).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Staniec, 1999a: 50 (Quedius; characters of pupa; Poland).

  • Distribution: Tunisia, Algeria, Morocco, Europe, Turkey, Uzbekistan.

oblitteratus Erichson, 1840: 549 (Quedius; Type locality: Sardinia).

  • — Gravenhorst, 1847: 227 (Quedius; characters; notes).

  • — Küster, 1853a: 65 (Quedius; characters; Sardinia).

  • — Fairmaire and Laboulbène, 1856: 544 (Quedius; characters; France).

  • — Kraatz, 1857d: 501 (Quedius; notes; Italy).

  • — Fauvel, 1874: 297 [= 1874a: 523] (Quedius; characters; France; Scandinavia; Britain; Italy; Corsica; Sardinia; Algeria; Cyprus; Caramania).

  • — Mulsant and Rey, 1876: 741 [= 1877: 597] (Quedius; subgenus Sauridus; characters; France).

  • — Fauvel, 1878: 129 [= 1878a: 49] (Quedius; Algeria).

  • — Fauvel, 1886: 60 [= 1886a: 52] (Quedius; Morocco; Algeria).

  • — Eppelsheim, 1892: 332 (Quedius; Turkestan).

  • — Ganglbauer, 1895: 408 (Quedius; subgenus Sauridus; characters; Europe; Mediterranean region).

  • — Fauvel, 1902b: 117 (Quedius; cited as obliteratus; Morocco; Algeria; Tunisia).

  • — Sainte-Claire Deville, 1906: 102 (Quedius; cited as obliteratus; Corsica).

  • — Porta, 1907: 138 (Quedius; subgenus Sauridus; characters; Italy).

  • — Montandon, 1908: 80 (Quedius; cited as obliteratus; Romania).

  • — Reitter, 1909: 113 (Quedius; subgenus Sauridus; cited as obliteratus; characters; Germany).

  • — Johansen, 1914: 368 (Quedius; subgenus Sauridus; cited as obliteratus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; catalog).

  • — Gridelli, 1922b: 130, 134 (Quedius; subgenus Sauridus; characters; Europe; Mediterranean region; Turkestan).

  • — Gridelli, 1925: 130 (Quedius; subgenus Raphirus; cited as obliteratus; characters).

  • — Portevin, 1929: 343 (Quedius; cited as obliteratus; characters; France).

  • — Scheerpeltz, 1933: 1452 (Quedius; cited as obliteratus; catalog).

  • — Wüsthoff, 1938a: fig. 39(Quedius; cited as obliteratus; aedeagal illustration).

  • — Jarrige, 1952a: 136 (Quedius; subgenus Raphirus; North Africa).

  • — V. Hansen, 1952: 150 (Quedius; subgenus Raphirus; cited as obliteratus; characters; Denmark).

  • — Smetana, 1958: 378 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 147 (Quedius; subgenus Raphirus; characters).

  • — Coiffait, 1963a: 386 (Quedius; subgenus Sauridus; characters).

  • — Palm, 1963b: 150 (Quedius; subgenus Raphirus; cited as obliteratus; characters; Sweden).

  • — Lohse, 1964: 217 (Quedius; subgenus Raphirus; cited as obliteratus; characters; central Europe).

  • — Scheerpeltz, 1965b: 408 (Quedius; cited as obliteratus; Sicily).

  • — Horion, 1965: 287 (Quedius; subgenus Raphirus, cited as obliteratus; France; Spain; Italy; Balkans; Romania; Slovakia; Austria).

  • — Fagel, 1968c: 2 (Quedius; subgenus Sauridus; characters; notes).

  • — Coiffait, 1970e: 63 (Quedius; Cyprus).

  • — Pope, 1977: 31 (Quedius; cited as obliteratus; synonym of humeralis).

  • — Coiffait, 1978: 213 (Quedius; subgenus Sauridus; characters; Europe).

  • — Tóth, 1984: 138 (Quedius; characters; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Poland; Czechoslovakia; Austria).

  • — Lohse and Lucht, 1989: 175 (Quedius; synonym of humeralis).

  • — Smetana, 1993b: 50 (Quedius; synonym of humeralis).

  • — Hodge and Jones, 1995: 42 (Quedius; cited as obliteratus and as valid species; notes; Britain).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; synonym of humeralis).

  • — Assing, Frisch, Kahlen, et al., 1998: 139 (Quedius; note).

gestroi Gridelli, 1922b: 127, 136 (Quedius; subgenus Sauridus; Type locality: Hispania; Sardegna; Sicilia: Pachino; Napoli: Camaldoli; Splato: Dalmazia).

  • — Gridelli, 1925: 130 (Quedius; synonym of oblitteratus).

  • — Scheerpeltz, 1933: 1452 (Quedius; synonym of oblitteratus).

  • — Smetana, 1958: 378 (Quedius; synonym of oblitteratus).

  • — Smetana, 1962b: 147 (Quedius; synonym of oblitteratus).

  • — Coiffait, 1978: 213 (Quedius; synonym of oblitteratus).

humilis Casey, 1915, see: nanulus Casey, 1915.

hummleri Bernhauer, 1905b: 585 (Quedius; Type locality: Parnass, Griechenland).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • — Gridelli, 1924a: 39 (Quedius; characters; Albania).

  • — Gridelli, 1925: 117 (Quedius; subgenus Raphirus; characters; Albania).

  • — Scheerpeltz, 1933: 1444 (Quedius; catalog).

  • — Coiffait, 1963a: 378 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1970e: 63 (Quedius; Cyprus).

  • — Coiffait, 1978: 261 (Quedius; subgenus Sauridus; characters; Greece; Albania).

  • Distribution: Cyprus, Greece, Albania.

hungaricus Coiffait, 1963a: 393, 418 (Quedius; subgenus Sauridus; Type locality: Hongrie: Herculesbad).

  • — Coiffait, 1978: 250 (Quedius; subgenus Sauridus; characters; Hungary).

  • — Tóth, 1984: 140 (Quedius; characters).

  • Distribution: Hungary, Romania.

hybridus Erichson, 1840: 432 (Philonthus; Type locality: Terra Van Diemenii).

  • — Fauvel, 1877: 270 (Philonthus; characters; Australia).

  • — Fauvel, 1878g: 553 (Quedius; characters; Australia).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • — Bernhauer, 1920: 18 (Quedius; Australia).

  • — Lea, 1925: 243 (Quedius; characters; Australia).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

hydrophilus Jarrige, 1972: 156 (Quedius; subgenus Microquedius; Type locality: Sainte Engrâce, Pyr.-Atl., gorges de Kakouetta).

  • — Coiffait, 1978: 199 (Quedius; subgenus Microquedius; characters; Pyrenees).

  • Distribution: France.

hyperboreus Erichson, 1840, see: fellmani Zetterstedt, 1838.

hypocrita Korge, 1971: 15 (Quedius; subgenus Sauridus; Type locality: Iran: Masuleh, 1400–2000, Gilan).

  • — Coiffait, 1978: 260 (Quedius; subgenus Sauridus; characters; Iran).

  • Distribution: Iran.

iablokofi Coiffait, 1967, see: transcaucasicus Iablokov-Khnzorian, 1961.

ibericola H. Franz, 1987, see: latinus Gridelli, 1938.

iheringi Bernhauer, 1907a: 287 (Quedius; subgenus Prionidus; Type locality: Brasilien: Jundiahy, Est. San Paolo).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • Distribution: Brazil.

illatus Sharp, 1884: 330 (Quedius; Type locality: Mexico, Jalapa; Guatemala, Sinanja in Vera Paz; Panama, Volcan de Chiriqui up to 4000 feet).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • Distribution: Mexico, Guatemala, Panama.

illyricus Wendeler, 1928b: 298 (Quedius; replacement name for albanicus Bernhauer).

  • — Scheerpeltz, 1933: 1444 (Quedius; catalog).

  • — Coiffait, 1967e: 400 (Quedius; subgenus Sauridus; characters; Albania).

  • — Korge, 1969: 45 (Quedius; notes; Bulgaria).

  • — Coiffait, 1978: 223 (Quedius; subgenus Sauridus; characters; Albania).

  • Distribution: Albania, Bulgaria.

albanicus Bernhauer, 1926e: 267 (Quedius; subgenus Sauridus; [preoccupied]; Type locality: Pashtrik und Gjalica in Albanien).

  • — Scheerpeltz, 1933: 1444 (Quedius; synonym of illyricus).

  • — Coiffait, 1967e: 400 (Quedius; synonym of illyricus).

  • — Coiffait, 1978: 223 (Quedius; synonym of illyricus).

imbecillis Sharp, 1889: 31 (Quedius; Type locality: Miyanoshita).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1444 (Quedius; catalog).

  • Distribution: Japan.

imitator Luze, 1904: 102 (Quedius; subgenus Sauridus; Type locality: Seravschan: Putschin-Pass, Darch, Obburden, Urmitan, Kumar; Jagnob: Varsaut; Iskander-kul: Iskanderdarja).

  • — Bernhauer, 1905b: 596 (Quedius; form of oblitteratus, cited as obliteratus).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; synonym of oblitteratus).

  • — Scheerpeltz, 1933: 1444 (Quedius; catalog).

  • — Coiffait, 1967e: 406 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 237 (Quedius; subgenus Sauridus; characters; Turkestan).

  • — Boháč, 1988: 556 [= 1989: 40] (Quedius; Uzbekistan; Turkestan).

  • Distribution: Uzbekistan, Tajikistan.

impar Smetana, 1971: 231 (Quedius; subgenus Raphirus; Type locality: Canada: British Columbia: Slide Mts., 10 mi. E. Barkerville, 6000 ft. Also cited from Alberta and Colorado).

  • — Smetana, 1973: 1431 (Quedius; notes; Alberta; British Columbia).

  • Distribution: Canada, USA.

impressifrons Solier, 1849, see: Cheilocolpus.

impressithorax Scheerpeltz, 1965: 281 (Quedius; subgenus Microsaurus; Type locality: N.E. Burma, Kambaiti, 7000 ft).

  • Distribution: Myanmar.

impressus Panzer, 1796, see: cinctus Paykull, 1790.

impunctifrons Delahon, 1915, see: fuliginosus Gravenhorst, 1802.

impunctus Solsky, 1868a: 124 (Quedius; Type locality: Mexique).

  • — Sharp, 1884: 336 (Quedionuchus; characters; notes; Mexico; Guatemala).

  • — Sharp, 1887: 783 (Quedionuchus; characters; Mexico).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Smetana, 1971: 153 (Quedius; subgenus Quedionuchus; characters; California; Guatemala; Mexico).

  • — Smetana, 1976a: 237 (Quedius; subgenus Quedius; characters; Mexico).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • Distribution: Guatemala, Mexico, USA.

inaequalipennis Lea, 1925: 244 (Quedius; Type locality: Tasmania: Waratah, Qilmot, Zeehan, Huon River. Victoria: Mount Donna Buang near Warburton. New South Wales: Blue Mountains).

  • — Scheerpeltz, 1933: 1444 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

incensus Smetana, 1959a: 215 (Quedius; subgenus Raphirus; Type locality: Albanien: Tiranë).

  • — Korge, 1964: 124 (Quedius; subgenus Raphirus; subspecies of attenuatus; Macedonia; Turkey).

  • — Smetana, 1967i: 559 (Quedius; subgenus Raphirus; notes; Turkey).

  • — Coiffait, 1970e: 63 (Quedius; Cyprus).

  • — Coiffait, 1978: 270 (Quedius; subgenus Raphirus; characters; Albania; Corfu).

  • Distribution: Macedonia, Albania, Greece, Cyprus, Turkey.

incisus Hatch, 1957: 224 (Quedius; subgenus Microsaurus; Type locality: Oregon: Blue Mts., Tollgate Road. Also cited from Washington, Idaho, and British Columbia).

  • — Smetana, 1971: 95 (Quedius; subgenus Microsaurus; characters; Alberta; British Columbia; Idaho; Montana; Oregon; Utah; Washington; Wyoming).

  • — Smetana, 1971b: 2 (Quedius; subgenus Microsaurus; British Columbia).

  • — Smetana, 1981b: 634 (Quedius; collecting notes; Alberta).

  • — Smetana, 1990d: 97 (Quedius; Utah).

  • Distribution: Canada, USA.

incompletus Hochhuth, 1849, see: Philonthus.

inconspicuus Blackburn, 1888: 5 (Quedius; Type locality: Wallaroo).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Lea, 1925: 242 (Quedius; characters; Australia).

  • — Oke, 1933: 123, 124 (Quedius; tarsal characters; Australia).

  • Distribution: Australia.

infernus Smetana, 1976c: 170 (Quedius; subgenus Microsaurus; Type locality: United States: Oregon: Mt. Hood, Cloud Cap Road, Tilly Jane Creek, 4100 ft. Also cited from Washington).

  • Distribution: USA.

infimus Cameron, 1952: 255 (Quedius; subgenus Microsaurus; Type locality: New Guinea: Snow Mountains: Scree Valley Camp, 3800 m).

  • Distribution: New Guinea.

inflatus Fauvel, 1875a: XXXIII [= 1875b: 235] (Quedius; Type locality: Naplouse, Wadi-el-Mekek).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Gridelli, 1925: 167 (Quedius; subgenus Raphirus; characters).

  • — Scheerpeltz, 1933: 1444 (Quedius; catalog).

  • — Coiffait, 1967e: 419 (Quedius; subgenus Raphirus; characters).

  • — Coiffait, 1978: 263 (Quedius; subgenus Raphirus; characters; Palestine; Lebanon).

  • — Smetana, 1978: 124 (Quedius; subgenus Raphirus; characters; collecting notes; Israel).

  • Distribution: Lebanon, Israel.

infuscatus Erichson, 1840: 543 (Quedius; Type locality: Lutetiae).

  • — Fairmaire and Laboulbène, 1856: 537 (Quedius; characters; France).

  • — Redtenbacher, 1857: 202 (Quedius; characters; Austria).

  • — Peyron, 1858: 428 (Quedius; Turkey).

  • — Fauvel, 1874: 285 [= 1874a: 511] (Quedius; characters; France; Germany; Austria).

  • — Redtenbacher, 1874: 198 (Quedius; characters; Austria).

  • — Mulsant and Rey, 1876: 670 [= 1877: 526] (Quedius; characters; France).

  • — Ganglbauer, 1895: 401 (Quedius; subgenus Microsaurus; characters; middle Europe).

  • — Porta, 1907: 109 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Reitter, 1909: 109 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Johansen, 1914: 361 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Gridelli, 1924d: 29 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Portevin, 1929: 339 (Quedius; characters; France).

  • — Donisthorpe, 1930: 30 (Quedius; characters; Britain).

  • — Scheerpeltz, 1933: 1444 (Quedius; catalog).

  • — Roubal, 1941b: 122, 126 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — V. Hansen, 1952: 127 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Smetana, 1958: 340 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 152 (Quedius; subgenus Microsaurus; characters).

  • — Palm, 1963b: 130 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Lohse, 1964: 207 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Horion, 1965: 254 (Quedius; subgenus Microsaurus; Britain; Denmark; Holland; Belgium; France; Corsica; Sicily; Romania; Czech Republic; Slovakia; Russia; Germany; Austria).

  • — Bordoni, 1974: 57 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Coiffait, 1978: 167 (Quedius; subgenus Microsaurus; characters; Europe; Britain; Scandinavia; Denmark; France; Corsica; Sicily).

  • — Tóth, 1984: 109 (Quedius; characters).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Benelux; Denmark).

  • — Smetana, 1993b: 49 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy ?).

  • Distribution: Europe, Turkey.

inquietus Champion, 1925: 107 (Velleius; Type locality: Sunderdhunga, W. Almora Division of Kumaon, alt. 9000 ft.).

  • — Cameron, 1932: 279 (Velleius; characters; India).

  • — Scheerpeltz, 1933: 1427 (Velleius; catalog).

  • — Smetana, 1988: 189 (Quedius; subgenus Microsaurus; characters; Uttar Pradesh; West Bengal; Nepal).

  • — Smetana, 1995g: 236 (Quedius; subgenus Microsaurus; characters; China).

  • — Smetana, 1997b: 129 (Quedius; notes; China).

  • — Smetana, 1999b: 214 (Quedius; subgenus Microsaurus; China).

  • Distribution: India, Nepal, China.

leptocephalus Coiffait, 1982a: 276 (Quedius; subgenus Microsaurus; Type locality: Nepal: Manigow, 1200–1900 m).

  • — Smetana, 1988: 189 (Quedius; synonym of inquietus).

  • — Smetana, 1995g: 236 (Quedius; synonym of inquietus).

inquinatus Stephens, 1832, see: Philonthus.

insignis Lea, 1929: 206 (Quedius; Type locality: Victoria: Beaconsfield).

  • — Scheerpeltz, 1933: 1444 (Quedius; catalog).

  • Distribution: Australia.

insolitus Sharp, 1886, see: Quediocafius.

insulanus Cameron, 1949: 176 (Quedius; subgenus Microsaurus; Type locality: Formosa: near Mt. Ari).

  • — Smetana, 1995: 27 (Quedius; subgenus of Microsaurus; characters; bionomics; Taiwan).

  • Distribution: Taiwan.

insularis Bernhauer and Schubert, 1916, see: Philonthus.

insularis Smetana, 1971, see: labradorensis Smetana, 1965.

intricatus Fauvel, 1895: 274 (Quedius; Type locality: Bírmanie, Carin Ghécu, 1300–1400 m; [Note: See lectotype designation by Smetana, 1988]).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Cameron, 1932: 298 (Quedius; subgenus Raphirus; characters; Burma).

  • — Scheerpeltz, 1933: 1444 (Quedius; catalog).

  • — Scheerpeltz, 1965: 112 (Quedius; subgenus Raphirus; Burma).

  • — Smetana, 1988: 295 (Quedius; subgenus Raphirus; lectotype designation; characters; Burma).

  • Distribution: Myanmar.

inversus Say, 1830, see: capucinus Gravenhorst, 1806.

invreae Gridelli, 1924d: 21, 47, 172 (Quedius; subgenus Microsaurus; Type locality: Belvedere presso Genova).

  • — Gridelli, 1929: 31 (Quedius; characters).

  • — Scheerpeltz, 1933: 1444 (Quedius; catalog).

  • — Roubal, 1941b: 124, 127 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — V. Hansen, 1952: 132 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Smetana, 1958: 347 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 156 (Quedius; subgenus Microsaurus; characters).

  • — Korge, 1962: 152 (Quedius; synonym of assecla).

  • — Palm, 1963b: 133 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark; Norway).

  • — Lohse, 1964: 210 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Horion, 1965: 261 (Quedius; subgenus Microsaurus; cited as invreai; Italy; Czechoslovakia; Poland; Germany; Belgium; Norway; Sweden; Denmark).

  • — Coiffait, 1967e: 394 (Quedius; subgenus Microsaurus; cited as invrae; notes).

  • — Bordoni, 1974: 28 (Quedius; subgenus Microsaurus; cited as invreai; characters; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 155 (Quedius; subgenus Microsaurus; characters; Europe; Italy; Corsica; France).

  • — Tóth, 1984: 116 (Quedius; characters).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Benelux; Denmark; Sweden).

  • — Bruge and Drugmand 1988: 169 (Quedius; characters).

  • — Nowosad, 1990: 135 (Quedius; collected from mole nest; notes; Poland).

  • — Wittwer, 1993: 251 (Quedius; Switzerland).

  • — Smetana, 1993b: 49 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Luff, 1996: 30 (Quedius; collecting notes; Isle of Man).

  • — Gerend and Braunert, 1997: 200 (Quedius; cited as invreai; notes; Luxembourg).

  • Distribution: Europe.

vesparum Rüschkamp, 1933: 170, 171 (Quedius; subspecies of othiniensis; Type locality: Bonnae in Rhenania).

  • — Roubal, 1941b: 122, 127 (Quedius; subgenus Microsaurus; subspecies of othiniensis; characters; collecting notes).

  • — Smetana, 1958: 347 (Quedius; synonym of invreae).

  • — Lohse, 1964: 210 (Quedius; synonym of invreae).

  • — Horion, 1965: 261 (Quedius; synonym of invreae).

  • — Bordoni, 1974: 28 (Quedius; synonym of invreae).

  • — Coiffait, 1978: 155 (Quedius; synonym of invreae).

brieni Bruge and Drugmand, 1988: 166 (Quedius; subgenus Microsaurus; Type locality: Belgique: Province de Namur: Vierves-sur-Viroin).

  • — Gerend and Braunert, 1997: 200 (Quedius; synonym of invreae).

iracundus Say, 1830, see: fulgidus Fabricius, 1793.

iranicus Coiffait, 1976a: 62 (Quedius; subgenus Microsaurus; Type locality: Iran: Grotte de Shapur).

  • — Coiffait, 1978: 153 (Quedius; subgenus Microsaurus; characters; Iran).

  • — Bordoni and Oromi, 1998: 1157 (Quedius; facultative cave species).

  • Distribution: Iran.

iridescens Broun, 1921: 480 (Quedius; Type locality: Glenhope).

  • — Scheerpeltz, 1933: 1444 (Quedius; catalog).

  • Distribution: New Zealand.

irideus Miller, 1851, see: fumatus Stephens, 1833.

iridicolor Quedenfeldt, 1882: 181 (Quedius; Type locality: Marocco, Tetuan).

  • — Quedenfeldt, 1884: 371 (Quedius; characters; Morocco).

  • — Fauvel, 1886: 59 [= 1886a: 51] (Quedius; Morocco).

  • — Fauvel, 1902b: 116 (Quedius; Morocco).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1444 (Quedius; catalog).

  • — Coiffait, 1963a: 379 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1973a: 275 (Quedius; Morocco).

  • — Coiffait, 1978: 251 (Quedius; subgenus Sauridus; characters; North Africa; Iberian Peninsula).

  • — Outerelo, 1980a: 57 (Quedius; Spain).

  • Distribution: Algeria, Morocco, Spain.

duplex Fauvel, 1886: 58 [= 1886a: 50] (Quedius; Type locality: Algérie: Teniet-el-Had).

  • — Fauvel, 1902b: 116 (Quedius; Morocco; Algeria).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1439 (Quedius; catalog).

  • — Coiffait, 1978: 251 (Quedius; synonym of iridicolor).

iridiventris Fauvel, 1877: 272 (Quedius; Type locality: Australie, Nouvelle Galles du Sud, Sydney).

  • — Fauvel, 1878g: 555 (Quedius).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Lea, 1925: 241 (Quedius; characters; Australia).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

iriei Uéno and Watanabe, 1970: 10 (Quedius; subgenus Microsaurus; Type locality: Japan: at the southwestern part of Kyushu: Kamamoto Prefecture: Takasawa-dô Cave, at Takasawa of Kônosé, in Kuma-mura).

  • — Watanabe, 1996a: 14 (Quedius; cave collection; Japan).

  • Distribution: Japan.

islamita Saulcy, 1864, see: scintillans Gravenhorst, 1806.

islandicus Fagel, 1960, see: boops Gravenhorst, 1802.

italicus Gridelli, 1925, see: collaris Erichson, 1840.

jacobi Scheerpeltz, 1955a: 176, 179 (Quedius; subgenus Cyrtoquedius; Type locality: Umgebung der Kolonie Hohenau in Suedwest-Paraguay).

  • Distribution: Paraguay.

jaenneri Hubenthal, 1902, see: mesomelinus Marsham, 1802.

japonicus Sharp, 1874: 26 (Quedius; Type locality: Japan; [Note: See lectotype designation by Smetana, 1998a]. Originally cited from Hiogo and Nagasaki).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • — Cameron, 1949b: 469 (Quedius; Japan).

  • — Nakane, 1963a: 94 (Quedius; characters; habitus photograph; Japan).

  • — Watanabe and Shibata, 1965: 321 (Quedius; subgenus Quedionuchus; Japan).

  • — Cho, 1996b: 110 (Quedius; subgenus Quedionuchus; characters; Korea).

  • — Smetana, 1998a: 121 (Quedius; subgenus Distichalius; lectotype designation: Japan; characters; Japan).

  • — Watanabe, 1998: 314 (Quedius; subgenus Distichalius, cited as Distichalis; Japan).

  • Distribution: Japan, Korea.

sachonensis Cho, 1996b: 111 (Quedius; subgenus Quedionuchus; subspecies of japonicus; Type locality: Korea: Kyongsangnam-do, Sachon-gun, Kuam-myon).

jarrigei Coiffait, 1963a: 391, 413 (Quedius; subgenus Sauridus; [Note: Cited in the subgenus Microsaurus on page 413 but this assignment is probably erroneous]; Type locality: Espagne: Sierra de Gredos, Almanzor).

  • — Coiffait, 1978: 227 (Quedius; subgenus Sauridus; characters; Spain).

  • Distribution: Spain.

javanus Cameron, 1937: 29 (Quedius; subgenus Indoquedius; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft).

  • Distribution: Indonesia.

jeanneli Coiffait, 1963a: 381, 407 (Quedius; subgenus Sauridus; Type locality: Roumanie: Banat, Val Comarnica).

  • — Coiffait, 1978: 222 (Quedius; subgenus Sauridus; characters; Romania).

  • — Tóth, 1984: 136 (Quedius; characters).

  • Distribution: Romania.

jeau Smetana, 1995, see: Quetarsis.

jelineki Krása, 1904, see: fuliginosus Gravenhorst, 1802.

jenisseensis J. Sahlberg, 1880: 72 (Raphirus; Type locality: Sibir. Jeniseie; [Note: See lectotype designation by Smetana, 1976d]. Originally cited from Nikulina, Tschulkova, Fatjanovsk, Imbatsk, Kruejka, Polovinka, Chantaika).

  • — Poppius, 1909: 28 (Quedius; subgenus Sauridus; Lena Valley, Siberia).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Gridelli, 1925: 167 (Quedius; subgenus Raphirus; characters).

  • — Scheerpeltz, 1933: 1444 (Quedius; catalog).

  • — Coiffait, 1967e: 417 (Quedius; subgenus Sauridus; [Note: Misidentified as as lederi according to Smetana, 1995a: 85]; characters).

  • — Smetana, 1976d: 24 (Quedius; subgenus Raphirus; lectotype designation: Sibir. Jeniseie; characters; Siberia).

  • — Smetana, 1995a: 85 (Quedius; subgenus Raphirus; notes; Russia).

  • Distribution: Russia.

jindrai Smetana, 1998b: 110 (Quedius; subgenus Raphirus; Type locality: China: Sichuan: Dayi Env., 110 km W. Chengdu).

  • Distribution: China.

job Coiffait, 1963a: 391, 415 (Quedius; subgenus Sauridus; Type locality: Liban: Nahr es Safa).

  • — Fagel, 1968c: 13 (Quedius; subgenus Sauridus; characters).

  • — Korge, 1971a: 52 (Quedius; subgenus Sauridus; Turkey).

  • — Coiffait, 1978: 238 (Quedius; subgenus Sauridus; characters; Lebanon).

  • Distribution: Lebanon, Turkey.

jocosus Sharp, 1884: 331 (Quedius; Type locality: Guatemala, near the city, at an elevation of 5000 feet).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1933: 1445 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 176 (Quedius; subgenus Cyrtoquedius; characters in key; Guatemala).

  • Distribution: Guatemala.

josefi Fleischer, 1925 [Note: Nomen nudum], see: mesomelinus Marsham, 1802.

josue Saulcy, 1864: 636 (Quedius; Type locality: Arag-el-Emir).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; synonym of nivicola).

  • — Coiffait, 1963a: 390 (Quedius; subgenus Sauridus; cited as valid species; characters).

  • — Fagel, 1968b: 5 (Quedius; subgenus Sauridus; characters; Lebanon).

  • — Korge, 1971a: 51 (Quedius; subgenus Sauridus; characters; Turkey).

  • — Coiffait, 1978: 218 (Quedius; subgenus Sauridus; characters; Palestine; Lebanon; Turkey).

  • Distribution: Israel, Lebanon, Turkey.

joyi Fagel, 1946, see: persimilis Mulsant and Rey, 1876.

juno Sharp, 1874, see: Indoquedius.

jureceki Rambousek, 1913, see: Philonthus.

kaalo Smetana, 1988: 243 (Quedius; subgenus Raphirus; Type locality: Nepal: Prov. Bagmati: Malemchi, 2800 m. Also cited from India).

  • Distribution: Nepal, India.

kabateki Smetana, 1997d: 468 (Quedius; subgenus Microsaurus; Type locality: China: N. Yunnan, Xue Shan, nr. Zhongdian, 4050 m, 27°49 N, 99°34 E).

  • Distribution: China.

kailo Smetana, 1988: 223 (Quedius; subgenus Microsaurus; Type locality: Nepal: Janakpur Thodung, 3100 m).

  • Distribution: Nepal.

kalabi Smetana, 1995a: 77 (Quedius; subgenus Microsaurus; Type locality: Kyrgyzstan: SU-Tien Shan, NE part of Terskey Ala Too ridge 3000–3600 m).

  • Distribution: Kyrgyzstan.

kalganensis Bernhauer, 1933: 41 (Quedius; subgenus Sauridus; Type locality: Mongolei: Kalgan).

  • — Smetana, 1996a: 51 (Quedius; subgenus Raphirus; characters; China).

  • Distribution: Mongolia, China.

kambaitiensis Scheerpeltz, 1965: 287 (Quedius; subgenus Raphirus; Type locality: N.E. Burma, Kambaiti, 2000 m).

  • Distribution: Myanmar.

kamchaticus Smetana, 1976d: 22 (Quedius; subgenus Distichalius; Type locality: Kamtschatka: Ozernaja).

  • — Smetana, 1978b: 86 (Quedius; subgenus Distichalius; Siberia).

  • — Ryabukhin, 1999: 74 (Quedius; subgenus Distichalius; catalog; habitat; distribution; northeastern Russia).

  • Distribution: Russia.

kanyasa Smetana, 1975a: 338 (Quedius; subgenus Raphirus; Type locality: West Nepal: Dsunda Khola valley near Talphi, 3000–3500 m).

  • — Smetana, 1988: 262 (Quedius; subgenus Raphirus; characters; Nepal).

  • Distribution: Nepal.

dhaulagirensis Coiffait, 1982: 80 (Quedius; subgenus Raphirus; Type locality: Nepal: südl. Dhaulagiri, Dhorpatan, 3000–3200 m).

  • — Smetana, 1988: 262 (Quedius; synonym of kanyasa).

lama Coiffait, 1982a: 234, 278 (Quedius; subgenus Raphirus; Type locality: Nepal, Padmara-Khari Lagna-Bumra, 3400–2750 m).

  • — Smetana, 1988: 262 (Quedius; synonym of kanyasa).

kashmirensis Cameron, 1944: 13 (Quedius; Type locality: Kashmir: Gulmarg).

  • — Smetana, 1988: 227 (Quedius; subgenus Distichalius; characters; Kashmir; Uttar Pradesh; Nepal).

  • — Herman, 2001: 30 (Quedius; synonymic note).

  • Distribution: India, Nepal.

mandra Smetana, 1975a: 332 (Quedius; subgenus Distichalius; Type locality: Central Nepal: near Goropani, W. Pokhara).

  • — Coiffait, 1982: 33 (Quedius; subgenus Quediops; Nepal).

  • — Coiffait, 1983: 274 (Quedius; Kashmir).

  • — Smetana, 1988: 228 (Quedius; synonym of kashmirensis).

deuvei Coiffait, 1983d: 168 (Quedius; subgenus Quediops; [preoccupied]; Type locality: Nepal: Ganesh Himal, Rupchet, 3600 m).

  • — Smetana, 1988: 228 (Quedius; synonym of kashmirensis).

deuveianus Coiffait, 1983b: 346 (Quedius; replacement name for deuvei Coiffait).

  • — Herman, 2001: 30 (Quedius; synonym of kashmirensis).

kashmiricus Cameron, 1932, see: fluviatilis Cameron, 1926.

katerinae Smetana, 1997: 54 (Quedius; subgenus Microsaurus; Type locality: China: C-Sichuan [Kangding] Gongga Shan massive, +2000 m low, Hailuogou vall., [Camp 1], 29°36′N, 102°02′E).

  • Distribution: China.

kawasawai Uéno and Watanabe, 1970: 15 (Quedius; subgenus Microsaurus; Type locality: Japan: Pacific coast of Shikoku: Kôchi Prefecture: Limestone cave called Shôbu-do at Shôbu of Tosayama-mura).

  • — Watanabe, 1996a: 14 (Quedius; cave collection; Japan).

  • Distribution: Japan.

kermanensis Coiffait, 1973: 113 (Quedius; subgenus Sauridus; Type locality: Iran: Province de Kerman).

  • — Coiffait, 1978: 234 (Quedius; subgenus Sauridus; characters; Iran).

  • Distribution: Iran.

khnzoriani Coiffait, 1967, see: acuminatus Hochhuth, 1849.

kiangsiensis Bernhauer, 1916e: 32 (Quedius; Type locality: Kiangsi).

  • — Gridelli, 1924d: 28 (Quedius; subgenus Microsaurus; characters; China).

  • — Scheerpeltz, 1933: 1462 (Quedius; variety of simulans).

  • — Smetana, 1995g: 236 (Quedius; subgenus Microsaurus; cited as valid species; characters; China).

  • Distribution: China.

kiesenwetteri Bernhauer and Schubert, 1916, see: ochropterus Erichson, 1840.

kirklarensis Korge, 1971a: 52 (Quedius; subgenus Sauridus; Type locality: Kirklar-Daglari oberhalb Çamlik, etwa 2200 m).

  • — Coiffait, 1978: 257 (Quedius; subgenus Sauridus; characters; Anatolia).

  • Distribution: Turkey.

kishimotoi Smetana, 1999d: 528 (Quedius; subgenus Microsaurus; Type locality: China: Guizhou, Shuicheng Xian, Shega, Anjia Yan Cave, 2050 m).

  • Distribution: China.

kiuchii Watanabe and Yoshida, 1970: 2 (Quedius; subgenus Microsaurus; Type locality: Japan: Shikoku: Tokushima Pref.: Ryû-no-iwaya Cave at Kurogô of Kamodani-chô in Anan-shi).

  • — Watanabe, 1996a: 14 (Quedius; cave collection; Japan).

  • Distribution: Japan.

klapperichi Smetana, 1996f: 123 (Quedius; subgenus Microsaurus; Type locality: China: Kuatun, Fukien).

  • — Smetana, 1999b: 220 (Quedius; subgenus Microsaurus; China).

  • Distribution: China.

klimai Roubal, 1914, see: fasciculatus Eppelsheim, 1886.

klinckowstroemi Poppius, 1913, see: mesomelinus Marsham, 1802.

kodiakensis Smetana, 1999, see: labradorensis Smetana, 1965.

koebelei Blackburn, 1895: 203 (Quedius; Type locality: N. Queensland).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Lea, 1925: 243 (Quedius; characters; Australia).

  • Distribution: Australia.

koei Smetana, 1999e: 544 (Quedius; subgenus Microsaurus; Type locality: China: Shaanxi, Qin Ling Shan, 108.47 E, 33.51 N, Mt. W pass autoroute km 70, 47 km S Xian, 2500–2600 m).

  • Distribution: China.

koltzei Eppelsheim, 1887: 420 (Quedius; Type locality: Chabarofka).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Gridelli, 1924d: 24 (Quedius; subgenus Microsaurus; characters; Amur region).

  • — Scheerpeltz, 1933: 1445 (Quedius; catalog).

  • — Coiffait, 1978: 164 (Quedius; subgenus Microsaurus; characters; central Asia).

  • — Smetana, 1998a: 115 (Quedius; subgenus Microsaurus; characters; Russia).

  • Distribution: Russia.

korgeanus Fagel, 1968, see: paradisianus Heer, 1839.

kozlovi Boháč, 1988: 555 [= 1989: 38] (Quedius; subgenus Quedius; Type locality: Vost. Tibet, bassein Goluboi reki, r. Go-Chiu, 13,500 feet).

  • Distribution: China.

kraatzii Brisout, 1859, see: plancus Erichson, 1840.

krali Smetana, 1999d: 524 (Quedius; subgenus Microsaurus; Type locality: China: SE Sichuan, Jinfo Shan, 29°01′N, 107°14′E, 2240 m).

  • Distribution: China.

kraussi Penecke, 1904, see: mesomelinus Marsham, 1802.

kuboni Štourač, 1998: 15 (Quedius; subgenus Raphirus; Type locality: Bulgaria: Rodopi Mts., Černatica-Goljam Persenk, 1400–1900 m).

  • Distribution: Bulgaria.

kucerai Smetana, 1996f: 126 (Quedius; subgenus Microsaurus; Type locality: China: Yunnan, Zongdian).

  • Distribution: China.

kuiro Smetana, 1988: 288 (Quedius; subgenus Raphirus; Type locality: Nepal: Khandbari Distr., Induwa Khola Valley, 2000 m).

  • Distribution: Nepal.

kuniakii Watanabe, 1990c: 26 (Quedius; subgenus Microsaurus; Type locality: Japan: Honshu: Nagano Pref.: Mt. Iwô-dake on the Yatsugatake Mts., alt. 2740 m).

  • Distribution: Japan.

kurosawai Shibata, 1986: 170 (Quedius; subgenus Raphirus; Type locality: Taiwan: Nantou Hsien: near Tsuifeng, about 2200 m alt.).

  • — Smetana, 1995: 105 (Quedius; subgenus of Raphirus; characters; bionomics; Taiwan).

  • Distribution: Taiwan.

kuay Smetana, 1995: 77 (Quedius; subgenus Raphirus; Type locality: Taiwan: Taichung Hsien: Lishan, 1500 m).

  • Distribution: Taiwan.

kubani Smetana, 1996f: 121 (Quedius; subgenus Microsaurus; Type locality: China: Yunnan, Dong-Chuan).

  • — Smetana, 1999b: 218 (Quedius; subgenus Microsaurus; characters of male; China).

  • Distribution: China.

labiatus Erichson, 1840: 533 (Quedius; Type locality: Columbia).

  • — Fauvel, 1891: 119 (Quedius; Venezuela).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1955a: 176, 177, 178 (Quedius; subgenus Cyrtoquedius; characters in key; type species of Cyrtoquedius; Mexico; Central America; Venezuela; Ecuador; Colombia).

  • — Marcuzzi, 1977: 14 (Quedius; Venezuela).

  • Distribution: Ecuador, Colombia, Venezuela, “Central America”, Mexico.

rivorum Sharp, 1884: 335 (Quedius; Type locality: Guatemala, near the city; Mexico, Juquila).

  • — Scheerpeltz, 1955a: 178 (Quedius; synonym of labiatus).

labradorensis Smetana, 1965: 37, 40 (Quedius; subgenus Quedius; Type locality: Südost-Labrador: Forteau).

  • — Smetana, 1964: 37 (Quedius; subgenus Quedius; [Note: Misidentified as lanei according to Smetana, 1971: 138]; Alaska).

  • — Smetana, 1965: 37, 40 (Quedius; subgenus Quedius; [Note: Misidentified as lanei according to Smetana, 1971: 138]; characters; notes; Newfoundland).

  • — Smetana, 1965a: 12 (Quedius; [Note: Misidentified as lanei according to Smetana, 1971: 138]; British Columbia; Alaska).

  • — Smetana, 1971: 138 (Quedius; subgenus Quedius; nominate subspecies; characters; Alberta; British Columbia; Manitoba; Newfoundland; Ontario; Quebec; Alaska; Maine; Massachusetts; Michigan; New Hampshire; New York).

  • — Smetana, 1973: 1428 (Quedius; notes; Alberta; Northwest Territories; Quebec).

  • — Smetana, 1976c: 172 (Quedius; subgenus Quedius; Quebec; Ontario).

  • — Smetana, 1978a: 826 (Quedius; subgenus Quedius; collecting notes; Alberta).

  • — Smetana, 1981b: 640 (Quedius; collecting notes).

  • — Downie and Arnett, 1996: 391 (Quedius; subgenus Quedius; characters; USA).

  • Distribution: Canada, USA.

insularis Smetana, 1971: 140 (Quedius; subgenus Quedius; subspecies of labradorensis; [preoccupied]; Type locality: Alaska: Anton Larsen Bay, Kodiak).

  • — Smetana, 1999: 79 (Quedius; synonym of labradoensis kodiakensis).

kodiakensis Smetana, 1999: 79 (Quedius; subspecies of labradorensis; replacement name for insularis Smetana).

laeticulus Sharp, 1884: 329 (Quedius; Type locality: Panama, Volcan de Chiriqui 2000 to 3000 feet).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • Distribution: Panama.

laetus Faldermann, 1835, see: ochripennis Ménétriés, 1832.

laevicollis Runde, 1835, see: molochinus Gravenhorst, 1806.

laevigatus Gyllenhal, 1810, see: plagiatus Mannerheim, 1843.

laevipennis Dufour, 1843, see: plagiatus Mannerheim, 1843.

laeviventris Bernhauer, 1917h: 94 (Quedius; subgenus laeviventris; Type locality: West-Kolumbien: Umgebung Cali am Rio Cauca, St. Antonio, 2000 m).

  • — Scheerpeltz, 1933: 1445 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 174 (Quedius; subgenus Cyrtoquedius; characters in key; Colombia).

  • Distribution: Colombia.

lama Coiffait, 1982, see: kanyasa Smetana, 1975.

lamus Smetana, 1995, see: przewalskii Reitter, 1887

lanei Hatch, 1957: 216 (Quedius; subgenus Quedius; Type locality: Washington: Blue Mts., 3000 ft. Also cited from British Columbia, Idaho, Oregon).

  • — Korge, 1962b: 333 (Quedius; characters; notes).

  • — Smetana, 1971: 144 (Quedius; subgenus Quedius; characters; British Columbia; Colorado; Oregon; Utah; Washington; Wyoming).

  • — Smetana, 1971a: 1844 (Quedius; subgenus Quedius; collecting notes; New Mexico).

  • — Smetana, 1976c: 174 (Quedius; subgenus Quedius; Colorado; Oregon).

  • Distribution: Canada, USA.

langenhani Hubenthal, 1912, see: vexans Eppelsheim, 1881.

lapponicus Zetterstedt, 1838, see: molochinus Gravenhorst, 1806.

lateralis Gravenhorst, 1802: 35 (Staphylinus; Type locality: Castelnuovo d'Istria; [Note: See neotype designation by Gridelli, 1938]).

  • — Latreille, 1804: 319 (Staphylinus; characters; Germany; France).

  • — Gravenhorst, 1806: 42 (Staphylinus; characters; France).

  • — Gyllenhal, 1827: 477 (Staphylinus; characters; Sweden).

  • — Stephens, 1832: 216 (Quedius; characters; England).

  • — Brullé, 1832: 131 (Staphylinus; notes; Greece).

  • — Curtis, 1837: pl. 638 (Quedius; characters; Britain).

  • — Stephens, 1835: 435 (Microsaurus; characters; England; Scotland).

  • — Lacordaire, 1835: 375 (Emus; characters; France).

  • — Nordmann, 1837: 76 (Philonthus; Germany; Sweden).

  • — Westwood, 1838a: 16 (Microsaurus; type species).

  • — Erichson, 1839a: 485 (Quedius; characters; Germany).

  • — Heer, 1839: 273 (Philonthus; characters; Switzerland).

  • — Stephens, 1839: 389 (Microsaurus; characters; England).

  • — Erichson, 1840: 525 (Quedius; characters; Europe).

  • — Dufour, 1843: 34 (Staphylinus; characters; France).

  • — Kiesenwetter, 1848: 49 (Quedius; characters; Germany).

  • — Redtenbacher, 1849: 708 (Philonthus; characters; Austria).

  • — Kiesenwetter, 1851a: 419 (Quedius; Pyrenees).

  • — Hardy, 1851: 31 (Quedius; notes; England).

  • — Fairmaire and Laboulbène, 1856: 539 (Quedius; characters; France).

  • — Kraatz, 1857d: 491 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 201 (Quedius; characters; Austria).

  • — Kraatz, 1858a: 57 (Quedius; Greece).

  • — Thomson, 1859: 25 (type species of Microsaurus).

  • — Thomson, 1860: 174 (Microsaurus; characters; Scandinavia).

  • — Fauvel, 1865a: 292 [= 1865b: 42] (Quedius; France).

  • — Fauvel, 1874: 276 [= 1874a: 502] (Quedius; characters; France; Europe; Japan).

  • — Sharp, 1874: 25 (Quedius; Japan).

  • — Redtenbacher, 1874: 198 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 267 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 631 [= 1877: 487] (Quedius; characters; France).

  • — Fowler, 1888: 231 (Quedius; characters; Britain).

  • — Ganglbauer, 1895: 397 (Quedius; subgenus Microsaurus; characters; Europe).

  • — Bordas, 1899: 230 (Quedius; defensive glands).

  • — Porta, 1907: 111 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Reitter, 1909: 110 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Johansen, 1914: 354 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; catalog).

  • — Gridelli, 1924d: 33 (Quedius; subgenus Microsaurus; characters; Italy; Sicily).

  • — Portevin, 1929: 339 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1445 (Quedius; catalog).

  • — Gridelli, 1938: 10 (Quedius; characters; neotype designation: Castelnuovo d'Istria; France; Denmark; Scandinavia; Germany; Czechoslovakia; Switzerland; Alps; Italy; Balkan Peninsula; Ionian Islands; Asia Minor).

  • — Wüsthoff, 1938a: fig. 4 (Quedius; aedeagal illustration).

  • — Roubal, 1941b: 123, 126 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Paulian, 1941: 268 (Quedius; larval characters).

  • — V. Hansen, 1952: 128 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Smetana, 1958: 342 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 153 (Quedius; subgenus Microsaurus; characters).

  • — Scheerpeltz, 1961e: 137 (Quedius; subgenus Microsaurus; Rhodes).

  • — Scheerpeltz, 1963a: 420 (Quedius; subgenus Microsaurus; Greece).

  • — Palm, 1963b: 130 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark).

  • — Lohse, 1964: 207 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Horion, 1965: 257 (Quedius; subgenus Microsaurus; widespread in southern and central Europe; Denmark; Sweden; Britain).

  • — Pototskaia, 1967: 83 (Quedius; larval characters).

  • — Kasule, 1968a: 68 (Quedius; life history).

  • — Kasule, 1970: 61 (Quedius; larval characters).

  • — Coiffait, 1970e: 62 (Quedius; Taygete).

  • — Bordoni, 1974: 21 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 177 (Quedius; subgenus Microsaurus; characters; Europe; England; Scandinavia; Balkan Peninsula; Asia Minor; Italy).

  • — Tóth, 1984: 109 (Quedius; characters; Hungary).

  • — H. Franz, 1987: 82 (Quedius; characters; notes).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — J. Li, 1993: 41 (Quedius; cited as laterlais; China).

  • — Smetana, 1993b: 49 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Outerelo, Gamarra, and Salgado, 1998: 130 (Quedius; subgenus Microsaurus; collected from cave; Spain; Portugal).

  • — Assing, 2001: 75 (Quedius; collected from xerothermous site; Germany).

  • Distribution: Europe, Turkey.

lateriflavus Scheerpeltz, 1963: 15, 28 (Quedius; subgenus Raphirus; Type locality: Afghanistan: Grotte von Karokh).

  • — Coiffait, 1978: 263 (Quedius; subgenus Sauridus; characters; Afghanistan).

  • Distribution: Afghanistan.

lateroflavus Lea, 1925: 246 (Quedius; Type locality: Queensland: Mount Tambourine; New South Wales: Blue Mountains, Sydney, Qurimbah).

  • — Scheerpeltz, 1933: 1445 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

lathburii Stephens, 1832, see: molochinus Gravenhorst, 1806.

latialis Gridelli, 1924: 22, 66, 172 (Quedius; subgenus Microsaurus; Type locality: Lazio: Filettino; M.te Viglio; Abruzzo: M.te Terminillo).

  • — Scheerpeltz, 1933: 1446 (Quedius; catalog).

  • — Coiffait, 1978: 173 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Italy.

laticollis Gravenhorst, 1802: 173 (Staphylinus; Type locality: Baltimore; [Note: See lectotype designation by Smetana, 1973]).

  • — Gravenhorst, 1806: 47 (Staphylinus).

  • — Erichson, 1840: 536 (Quedius; synonym of molochinus).

  • — Mannerheim, 1843b: 233 (Quedius; synonym of molochinus).

  • — Gravenhorst, 1847: 227 (Quedius; synonym of molochinus).

  • — Kraatz, 1857d: 501 (Quedius; synonym of molochinus).

  • — Ganglbauer, 1895: 403 (Quedius; synonym of molochinus).

  • — Fauvel, 1902b: 115 (Quedius; synonym of picipennis).

  • — Porta, 1907: 129 (Quedius; synonym of molochinus).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; synonym of picipennis).

  • — Gridelli, 1924d: 81 (Quedius; synonym of variety molochinus).

  • — Scheerpeltz, 1933: 1457 (Quedius; synonym of picipennis).

  • — Méquignon, 1937: 3 (Quedius; synonym of molochinus).

  • — Smetana, 1958: 364 (Quedius; synonym of molochinus).

  • — Smetana, 1973: 1427 (Quedius; lectotype designation; valid species; notes).

  • — Smetana, 1978a: 826 (Quedius; subgenus Quedius; collecting notes; Florida; Illinois; Kentucky; Louisiana; Texas).

  • — Smetana, 1981b: 640 (Quedius; Ontario).

  • — Smetana, 1990d: 98 (Quedius; Arkansas; Mississippi; Oklahoma).

  • — Downie and Arnett, 1996: 391 (Quedius; subgenus Quedius; characters; USA).

  • Distribution: Canada, USA.

neomolochinus Korge, 1962b: 333 (Quedius; Type locality: Washington, D.C.).

  • — Smetana, 1965: 39 (Quedius; subgenus Quedius; characters).

  • — Smetana, 1971: 136 (Quedius; subgenus Quedius; characters; Alabama; Arkansas; Florida; Georgia; Illinois; Indiana; Kentucky; Louisiana; Maryland; Massachusetts; Michigan; Missouri; New Hampshire; New Jersey; New York; North Carolina; Ohio; Oklahoma; Pennsylvania; Rhodes Island; South Carolina; Texas; Virginia; West Virginia).

  • — Smetana, 1973: 1427 (Quedius; synonym of laticollis).

marginatus Korge, 1962b: 335 (Quedius; aberration of neomolochinus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]).

  • — Smetana, 1965: 39 (Quedius; subgenus Quedius; aberration of neomolochinus; characters).

  • — Smetana, 1971: 136 (Quedius; synonym of neomolochinus).

laticollis Sharp, 1889, see: aurorus Herman, 2001.

latifrons Sharp, 1886a: 379 (Quedius; Type locality: New Zealand: Bealey).

  • — Broun, 1893: 1030 (Quedius; characters; New Zealand).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; catalog).

  • Distribution: New Zealand.

latimanus Broun, 1893: 1031 (Quedius; Type locality: Otago).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; catalog).

  • Distribution: New Zealand.

latinus Gridelli, 1938: 15 (Quedius; Type locality: Toscana: Lippiano, Arezzo. Also cited from Spain, Portugal, France).

  • — Wüsthoff, 1938a: fig. 5 (Quedius; aedeagal illustration).

  • — Scheerpeltz, 1961d: 132 (Quedius; subgenus Microsaurus; Italy).

  • — Scheerpeltz, 1965b: 406 (Quedius; Sicily).

  • — Bordoni, 1974: 24 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Coiffait, 1978: 179 (Quedius; subgenus Microsaurus; characters; Italy; France; Iberian Peninsula).

  • — H. Franz, 1984: 59 (Quedius; Macedonia).

  • — Sparacio, 1995: 165 (Quedius; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Assing and Wunderle, 2001: 40 (Quedius; synonymic notes).

  • Distribution: Portugal, Spain, France, Italy.

ibericola H. Franz, 1987: 83 (Quedius; subgenus Microsaurus; subspecies of lateralis; Type locality: Serra de Estrela; Sierra de Guadarrama in Mittelspanien).

  • — Assing and Wunderle, 2001: 40 (Quedius; synonym of latinus).

latus Hochhuth, 1851: 30 (Quedius; Type locality: Kaukasien, Alagez-Gebirge).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1446 (Quedius; catalog).

  • Distribution: Caucasus.

laxatus Casey, 1915, see: explanatus LeConte, 1858.

leai Scheerpeltz, 1933: 1446 (Quedius; replacement name for duplopunctatus Lea).

  • Distribution: Australia.

duplopunctatus Lea, 1925: 249 (Quedius; [Note: Preoccupied; duplopunctatus is cited as a homonym of a duplopunctatus Gridelli (1922) who used duplepunctatus not duplopunctatus but they are homonyms according to Article 58.12]; Type locality: Tasmania: Frankford).

  • — Scheerpeltz, 1933: 1446 (Quedius; synonym of leai).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

lederi Bernhauer, 1902, see: sublimbatus Mäklin, 1853.

ledouxi Coiffait, 1977c: 138 (Quedius; subgenus Sauridus; Type locality: Iran septentrional, Abalurni Guilam, 1000 m).

  • — Coiffait, 1978: 259 (Quedius; subgenus Sauridus; characters; Iran).

  • Distribution: Iran.

leechi Smetana, 1971: 127 (Quedius; subgenus Microsaurus; Type locality: Arizona: Mt. Graham, 10,000–10,500 ft, Graham Co.).

  • — Smetana, 1971a: 1841 (Quedius; subgenus Microsaurus; collecting notes; discussion).

  • — Smetana, 1978a: 825 (Quedius; subgenus Microsaurus; collecting notes; Arizona).

  • Distribution: USA.

chiricahuensis Smetana, 1971a: 1843 (Quedius; subgenus Microsaurus; subspecies of leechi; Type locality: Arizona: Cochise Co., Chiricahua Mts., Rustler Park, 8400 ft.).

  • — Smetana, 1978a: 825 (Quedius; subgenus Microsaurus; subspecies of leechi; collecting notes; Arizona).

neomexicanus Smetana, 1971a: 1842 (Quedius; subgenus Microsaurus; subspecies of leechi; Type locality: New Mexico: Lincoln N. F., Sacramento Mts., 2 mi SE Cloudcroft, 8500 ft.).

leeng Smetana, 1995: 76 (Quedius; subgenus Raphirus; Type locality: Taiwan: Nantou Hsien: Yushan N. P., W. slope blw. Yushan Mn. Pk., 3720 m).

  • Distribution: Taiwan.

lenkorianensis Drugmand, 1989: 176 (Quedius; subgenus Microsaurus; Type locality: Lenkoran: Astara, Caucase oriental).

  • Distribution: Azerbaijan.

lentus Motschulsky, 1860: 571 (Quedius; Type locality: Noukha dans la Géorgie Méridionale).

  • — Hochhuth, 1862: 44 (Quedius; characters; Georgia).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1446 (Quedius; catalog).

  • Distribution: Georgia.

leonhardi Bernhauer, 1914, see: molochinus Gravenhorst, 1806.

lepidulus Stephens, 1832, see: Philonthus.

leptocephalus Coiffait, 1982, see: inquietus Champion, 1925.

lesagei Smetana, 1988: 211 (Quedius; subgenus Microsaurus; Type locality: Nepal: Khandbari Dis. Arun Valley at Num main bridge 1050 m).

  • Distribution: Nepal.

levasseuri Coiffait, 1964d: 293 (Quedius; subgenus Quedius; Type locality: Corse, Porto-Vecchio).

  • — Bordoni, 1976b: 102 (Quedius; characters; Italy).

  • — Coiffait, 1978: 197 (Quedius; subgenus Quedius; characters; Corsica).

  • Distribution: Italy.

levicollis Brullé, 1832: 131 (Staphylinus; [Note: The name tristis Gravenhorst has long been applied to this species. Gravenhorst did not describe tristis as a new, he attributed it to Fabricius. The Fabrician name is now in Pseudocypus. The species misidentified as tristis Gravenhorst needed a new name and levicollis was one of the next two available names. All of the citations for tristis Gravenhorst are annotated and included under levicollis]; Type locality: Morée: plaine de Modon).

  • — Gravenhorst, 1802: 34 (Staphylinus; cited as tristis; [Note: Misidentification: This reference is often cited as the original description for tristis Gravenhorst but he attributed the species to Fabricius]; characters; Germany).

  • — Stephens, 1832: 215 (Quedius; cited as tristis; characters; England).

  • — Lacordaire, 1835: 376 (Emus; cited as tristis; characters; France).

  • — Nordmann, 1837: 75 (Philonthus; cited as tristis; Europe).

  • — Holme, 1837: 63 (Quedius; cited as tristis; notes; Scilly Islands).

  • — Westwood, 1838: 167 (Quedius; cited as tristis; larva, pupa).

  • — Erichson, 1839a: 490 (Quedius; cited as tristis; synonym of fuliginosus).

  • — Stephens, 1839: 389 (Quedius; cited as tristis; characters; England).

  • — Erichson, 1840: 537 (Quedius; cited as tristis; characters; Europe).

  • — Kolenati, 1846: 22 (Quedius; cited as tristis; synonym of fuliginosus).

  • — Gravenhorst, 1847: 231 (Quedius; cited as tristis; characters; notes).

  • — Redtenbacher, 1849: 709 (Philonthus; cited as tristis; characters; Austria).

  • — Kraatz, 1857d: 502 (Quedius; cited as tristis; characters; Germany).

  • — Hochhuth, 1862: 46 (Quedius; cited as tristis; character of variety; Russia).

  • — Fauvel, 1865a: 293 [= 1865b: 43] (Quedius; cited as tristis; habitat; France).

  • — Fauvel, 1874: 289 [= 1874a: 515] (Quedius; cited as tristis; characters; France; Germany; Algeria; Cyprus; Turkey; Caucasus; Transcaucasia).

  • — Redtenbacher, 1874: 198 (Quedius; cited as tristis; characters; Austria).

  • — Mulsant and Rey, 1876: 688 [= 1877: 544] (Quedius; subgenus Microsaurus; cited as tristis; characters; France).

  • — Fauvel, 1878: 128 [= 1878a: 48] (Quedius; cited as tristis; Algeria).

  • — Fauvel, 1886: 58 [= 1886a: 50] (Quedius; cited as tristis; Algeria).

  • — Fowler, 1888: 237 (Quedius; subgenus Microsaurus; cited as tristis; characters; Britain).

  • — Ganglbauer, 1895: 403 (Quedius; cited as tristis; characters; middle Europe; Mediterranean region).

  • — Ganglbauer, 1895: 403 (Quedius; cited as laevicollis; synonym of tristis).

  • — Fauvel, 1902b: 115 (Quedius; cited as tristis; Algeria; Tunisia).

  • — Sainte-Claire Deville, 1906: 102 (Quedius; cited as tristis; Corsica).

  • — Porta, 1907: 129 (Quedius; subgenus Quedius; cited as tristis; characters; Italy).

  • — Porta, 1907: 129 (Quedius; cited as laevicollis; synonym of tristis).

  • — Reitter, 1909: 111 (Quedius; cited as tristis; characters; Germany).

  • — Johansen, 1914: 363 (Quedius; subgenus Quedius; cited as tristis; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; cited as tristis; catalog).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; cited as laevicollis; synonym of tristis).

  • — Munster, 1923: 196 (Quedius; cited as tristis; Fennoscandia).

  • — Gridelli, 1924d: 80 (Quedius; subgenus Quedius; cited as tristis; characters; Carniola; Italy; Sicily; Sardinia; Greece; Anatolia; Tunisia; Algeria).

  • — Portevin, 1929: 341 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1463 (Quedius; cited as tristis; catalog).

  • — Wüsthoff, 1938a: fig. 22 (Quedius; cited as tristis; aedeagal illustration).

  • — Tottenham, 1949: 376 (Quedius; cited as type species of Quedius as “Staphylinus laevicollis Brullé, 1832 [= Staphylinus tristis Gravenhorst, 1802, nec Fabricius, 1793]).

  • — V. Hansen, 1952: 142 (Quedius; subgenus Quedius; cited as tristis; characters; Denmark).

  • — Scheerpeltz, 1956c: 1102 (Quedius; cited as tristis; Greece; Cyprus).

  • — Smetana, 1958: 362 (Quedius; subgenus Quedius; cited as tristis; characters; ecology; Czechoslovakia).

  • — Smetana, 1958: 362 (Quedius; cited as laevicollis; synonym of tristis).

  • — Scheerpeltz, 1961d: 133 (Quedius; subgenus Quedius; cited as tristis; Italy).

  • — Smetana, 1962b: 134 (Quedius; subgenus Quedius; cited as tristis; characters; Mediterranean region; Caucasus; Transcaucasus; Germany; Czechoslovakia; Austria).

  • — Smetana, 1963b: 39 (Quedius; subgenus Quedius; cited as tristis; Madeira).

  • — Palm, 1963b: 141 (Quedius; subgenus Quedius; cited as tristis; characters; Sweden; Denmark; Norway).

  • — Lohse, 1964: 211 (Quedius; subgenus Quedius; cited as tristis; characters; central Europe).

  • — Focarile, 1964: 61 (Quedius; cited as tristis; Italy).

  • — Scheerpeltz, 1965b: 407 (Quedius; cited as tristis; Sicily).

  • — Scheerpeltz, 1965c: 511 (Quedius; cited as tristis; Sicily).

  • — Horion, 1965: 273 (Quedius; subgenus Quedius; cited as tristis; Algeria; Tunisia; Caucasus; Iran; Ireland; Britain; Holland; Belgium; Switzerland; France; Portugal; Spain; Balearics; Corsica; Sardinia; Sicily; Italy; Dalmatia; Albania; Crete; Rhodes; Lesbos; Cyprus; Bosnia Hercegovina; Romania; Hungary; Austria; Germany; Czechoslovakia; Poland; Ukraine; Denmark; Sweden; Norway).

  • — Smetana, 1967i: 558 (Quedius; cited as tristis; Turkey).

  • — Coiffait, 1970e: 62 (Quedius; cited as tristis; Cyprus).

  • — Smetana, 1970: 62 (Quedius; cited as tristis; distributional summary; Madeira).

  • — Bordoni, 1976b: 93 (Quedius; cited as tristis; characters; Italy).

  • — Pope, 1977: 31 (Quedius; cited as tristis; cited as a homonym; Britain).

  • — Pope, 1977: 31 (Quedius; synonym of tristis).

  • — Coiffait, 1978: 192 (Quedius; cited as laevicollis; synonym of tristis).

  • — Coiffait, 1978: 192 (Quedius; subgenus Quedius; cited as tristis; characters; Europe; Britain; Scandinavia; Denmark; Asia Minor; North Africa; Crete).

  • — Smetana, 1978: 121 (Quedius; cited as tristis; collecting notes; Israel).

  • — Hinton, 1981: 662 (Quedius; cited as laevicollis; characters of egg).

  • — Bordoni, 1982a: 140 (Quedius; cited as tristis; cave collection; Sardinia).

  • — Bordoni, 1984: 342 (Quedius; cited as tristis; Lebanon).

  • — Tóth, 1984: 118 (Quedius; cited as tristis; characters; Hungary).

  • — H. Franz, 1987: 70 (Quedius; cited as tristis; Cyprus).

  • — Lucht, 1987: 109 (Quedius; cited as tristis; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Gusarov, 1993: 73 (Quedius; cited as tristis; synonyms).

  • — Welch, 1993: 229 (Quedius; cited as tristis; ovariole number and ovary structure).

  • — Smetana, 1993b: 50 (Quedius; subgenus Quedius; cited as laevicollis; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; cited as laevicollis; Italy).

  • — Newton, 1995: 51 (requested that the International Commission of Zoological Nomenclature place levicollis, on the Official List of Specific Names in Zoology).

  • — ICZN, 1996: 215 (placed on Official List of Specific Names in Zoology).

  • — K. Bland, 1997: 204 (Quedius; cited as tristis; eaten by toad; Scotland).

  • — Marchal and Chardonnet, 2000: 72 (Quedius; cited as tristis; Corsica).

  • — Owen, 2000: 252 (Quedius; cited as tristis; pitfall trap collections from underground at roots of trees; Britain).

  • — Assing and Wunderle, 2001: 37 (Quedius; cited as tristis; Cyprus).

  • Distribution: Tunisia, Algeria, Madeira, Europe, Russia, Caucasus, Turkey, Lebanon, Israel, Iran.

nigritarsis Ménétriés, 1832: 144 (Staphylinus; [Note: This name is cited by Bernhauer and Schubert as a valid species of Quedius (1916: 429) and as a junior synonym of Philonthus splendens (1914: 356), although the page number they cited for the original description of the latter is incorrect; it is herein cited in Quedius only]; Type locality: Caucase; [Note: See lectotype designation by Gusarov, 1993]).

  • — Faldermann, 1835: 123 (Staphylinus; characters; Persia-Armenia).

  • — Hochhuth, 1849: 154 (Quedius; characters).

  • — Bernhauer and Schubert, 1914: 356 (Philonthus; synonym of splendens).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1451 (Quedius; catalog).

  • — Smetana, 1958: 155 (Philonthus; synonym of splendens).

  • — Coiffait, 1974: 266 (Philonthus; synonym of splendens).

  • — Gusarov, 1993: 73 (Quedius; lectotype designation; synonym of tristis).

frontalis Nordmann, 1837: 76 (Philonthus; Type locality: Gallia ?; Symperopoli).

  • — Erichson, 1840: 536 (Quedius; characters; France; Italy; Dalmatia; Greece).

  • — Kiesenwetter, 1848: 54 (Quedius; characters).

  • — Gravenhorst, 1847: 231 (Quedius; synonym of tristis; Erichson cited as author, Erichson attributed the species to Nordmann).

  • — Redtenbacher, 1849: 709 (Philonthus; synonym of tristis).

  • — Hardy, 1851: 32 (Quedius; notes; England).

  • — Rosenhauer, 1856: 71 (Quedius; Algeciras).

  • — Fairmaire and Laboulbène, 1856: 536 (Quedius; characters; France).

  • — Kraatz, 1857d: 502 (Quedius; synonym of tristis).

  • — Redtenbacher, 1857: 201 (Quedius; characters; Austria).

  • — Fairmaire and Coquerel, 1860: 154 (Quedius; Algeria).

  • — Fauvel, 1874: 289 [= 1874a: 515] (Quedius; synonym of tristis).

  • — Ganglbauer, 1895: 403 (Quedius; synonym of tristis).

  • — Porta, 1907: 129 (Quedius; synonym of tristis).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; synonym of tristis).

  • — Gridelli, 1924d: 80 (Quedius; synonym of tristis).

  • — Scheerpeltz, 1933: 1464 (Quedius; synonym of tristis).

  • — Smetana, 1958: 362 (Quedius; synonym of tristis).

  • — Coiffait, 1978: 192 (Quedius; synonym of tristis).

balthasari Mařan, 1935: 9 (Quedius; variety of tristis; Type locality: Knossos, Creta centr. septentr.).

  • — Coiffait, 1978: 192 (Quedius; subgenus Quedius; variety of tristis; characters; Crete).

lewisius Sharp, 1874: 27 (Quedius; Type locality: Hiogo; [Note: See lectotype designation by Smetana, 1998a]).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; catalog).

  • — Nakane, 1963a: 94 (Quedius; characters; habitus photograph; Japan).

  • — Smetana, 1998a: 125 (Quedius; subgenus Distichalius; lectotype designation; characters; Japan).

  • Distribution: Japan.

lgockii Roubal, 1911: 9 (Quedius; Type locality: Kr. Poljana).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; catalog).

  • — Gridelli, 1924d: 106 (Quedius; subgenus Raphirus; characters; Caucasus).

  • — Scheerpeltz, 1933: 1446 (Quedius; catalog).

  • — Coiffait, 1963a: 377 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 260 (Quedius; subgenus Sauridus; characters; Caucasus).

  • — Boháč, 1980: 86 (Quedius; Caucasus).

  • Distribution: Caucasus.

liang Smetana, 1995: 40 (Quedius; subgenus Microsaurus; Type locality: Taiwan: near Chihtuan Illan. Also cited from South Korea).

  • — Smetana, 1997: 51 (Quedius; subgenus Microsaurus; China).

  • Distribution: China, Korea, Taiwan.

liau Smetana, 1999b: 229 (Quedius; subgenus Microsaurus; Type locality: China: Shaaxi, Qin Ling Shan, 108.47 E, 33.51 N, mountain pass at autoroute km 70, 47 km S. Xian, 2500–2600 m).

  • Distribution: China.

libanicus Coiffait, 1954, see: vicinus Ménétriés, 1832.

limbatoides Coiffait, 1963, see: limbatus Heer, 1839.

limbatus Heer, 1839: 281 (Heterothops; Type locality: Matt).

  • — Gyllenhal, 1810: 310 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1916: 426, cited praecox Gyllenhal as a synonym of limbatus Heer; Gyllenhal did not describe the species as new, he attributed it to Gravenhorst]; characters; Sweden).

  • — Heer, 1834: 74 (Staphylinus; subgenus Microsaurus; elevational listing; [Note: Unavailable name; this citation has been used by most workers as the original reference for limbatus, but the name was published therein in a list, without characters. Heer validated the name in 1839 when he presented characters]).

  • — Erichson, 1839a: 492 (Quedius; [Note: Misidentification: Bernhauer and Schubert, 1916: 426, cited maurorufus Erichson as a synonym of limbatus Heer; Erichson did not describe the species as new, he attributed it to Gravenhorst]; characters; Germany).

  • — Fauvel, 1874: 299 [= 1874a: 525] (Quedius; characters; France; Lapland; Scandinavia; Britain; Germany; Switzerland; Austria; Russia).

  • — Mulsant and Rey, 1876: 735 [= 1877: 591] (Quedius; subgenus Sauridus; characters; France).

  • — J. Sahlberg, 1876: 29 (Raphirus; Finland).

  • — Eppelsheim, 1892: 332 (Quedius; Turkestan).

  • — Ganglbauer, 1895: 407 (Quedius; subgenus Sauridus; characters; north and middle Europe).

  • — Porta, 1907: 139 (Quedius; subgenus Sauridus; characters; Italy).

  • — Reitter, 1909: 113 (Quedius; subgenus Sauridus; characters; Germany).

  • — Johansen, 1914: 366 (Quedius; subgenus Sauridus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; catalog).

  • — Gridelli, 1922b: 130, 132 (Quedius; subgenus Sauridus; characters; notes; Italy; Romania).

  • — Munster, 1923: 198 (Quedius; subgenus Sauridus; Fennoscandia).

  • — Gridelli, 1925: 121 (Quedius; subgenus Raphirus; characters; Finland; Germany; Italy; Romania; Bosnia; Herzegowinia; Caucasus).

  • — Portevin, 1929: 343 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1446 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 35(Quedius; aedeagal illustration).

  • — Jeannel and Jarrige, 1949: 380 (Quedius; subgenus Raphirus; cave species; France).

  • — V. Hansen, 1952: 148 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Smetana, 1958: 375 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 146 (Quedius; subgenus Raphirus; characters; Europe; Caucasus; Balkans; Turkestan).

  • — Ferreira, 1962: 19 (Quedius; Portugal).

  • — Palm, 1963b: 148 (Quedius; subgenus Raphirus; characters; Sweden; Norway; Finland).

  • — Coiffait, 1963a: 394 (Quedius; subgenus Sauridus; characters).

  • — Lohse, 1964: 216 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 284 (Quedius; subgenus Raphirus; Caucasus; Turkestan; Turkey; Russia; Finland; Switzerland; Norway; Ukraine; Latvia; Poland; Czechoslovakia; Hungary; Romania; Serbia; Bosnia Hercegovina; Bulgaria; Greece; Dalmatia; Slovenia; Italy; France; Switzerland; Austria; Germany).

  • — Szujecki, 1968a: 736 (Quedius; Poland).

  • — Scheerpeltz, 1968a: 94 (Quedius; catalog; Austria).

  • — Bolov, 1969: 514 (Quedius; collecting notes; Russia).

  • — Bolov, 1969a: 329 (Quedius; collecting notes; Russia).

  • — Tikhomirova, 1973a: 190 (Quedius; checklist; Soviet Union).

  • — Coiffait, 1978: 258 (Quedius; subgenus Sauridus; characters; Europe; Caucasus; Transcaucasus; Asia Minor).

  • — Muona, 1979: 16 (Quedius; Finland; Russia; Sweden; Norway; Baltic States).

  • — Uhlig and Vogler, 1981: 125 (Quedius; collecting notes; Germany).

  • — Tóth, 1983: 186 (Quedius; Hungary).

  • — Tóth, 1984: 140 (Quedius; characters; Hungary).

  • — Tóth, 1985a: 102 (Quedius; subgenus Sauridus; Hungary).

  • — Drugmand, 1987a: 312 (Quedius; Belgium).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Sweden).

  • — Gusarov, 1989: 12 (Quedius; habitat notes; Ukraine).

  • — Lohse and Lucht, 1989: 175 (Quedius; characters).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Kocian, 1993b: 96 (Quedius; Czech Republic).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Šustek, 1995: 398 (Quedius; Czech Republic).

  • — Solodovnikov, 1998: 342 (Quedius; bionomic notes; distribution in northwest Caucasus; Russia).

  • — Paśnik, 1998: 73 (Quedius; habitat; Poland).

  • — Assing, 1999b: 38 (Quedius; synonymic notes).

  • — Herman, 2001: 30 (Quedius; nomen protectum).

  • Distribution: Europe, Russia, Caucasus, Turkey, “Turkestan”, Uzbekistan.

attenuatus Gravenhorst, 1802: 27 (Staphylinus; [Note: This name is older than limbatus. Bernhauer and Schubert, 1916: 426, cited Gravenhorst, 1806, as the reference of this name but Gravenhorst, 1806, attributed it to his 1802 work]; Type locality: Brunsvigae).

  • — Latreille, 1804: 316 (Staphylinus; characters; Germany).

  • — Gravenhorst, 1806: 61 (Staphylinus; characters; Germany).

  • — C. Sahlberg, 1830: 318 (Staphylinus; characters; Finland).

  • — Stephens, 1833: 241 (Raphirus; characters; England).

  • — Runde, 1835: 6 (Staphylinus; characters; Germany).

  • — Lacordaire, 1835: 383 (Emus; characters; France).

  • — Westwood, 1838a: 16 (Raphirus; characters; type species).

  • — Zetterstedt, 1838: 62 (Staphylinus; characters; habitat; Lapland).

  • — Erichson, 1839a: 493 (Quedius; synonym of maurorufus).

  • — Stephens, 1839: 393 (Quedius; characters; England).

  • — Erichson, 1840: 542 (Quedius; synonym of maurorufus).

  • — Laporte, 1840: 177 (Staphylinus; characters; France).

  • — Gravenhorst, 1847: 230 (Quedius; synonym of rufipes).

  • — P. Lucas, 1846: 114 (Quedius; synonym of maurorufus).

  • — Kraatz, 1857d: 512 (Quedius; synonym of maurorufus).

  • — Ganglbauer, 1895: 407 (Quedius; synonym of limbatus).

  • — Porta, 1907: 139 (Quedius; synonym of limbatus).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; synonym of limbatus).

  • — Gridelli, 1925: 121 (Quedius; synonym of limbatus).

  • — Scheerpeltz, 1933: 1446 (Quedius; synonym of limbatus).

  • — Smetana, 1958: 375 (Quedius; synonym of limbatus).

  • — Herman, 2001: 30 (Quedius; nomen oblitum).

limbatoides Coiffait, 1963a: 393, 419 (Quedius; subgenus Sauridus; Type locality: Haut-Rhin, Aubure).

  • — Coiffait, 1978: 255 (Quedius; subgenus Sauridus; characters; Czechoslovakia; Romania; Alsace).

  • — Tóth, 1984: 140 (Quedius; characters).

  • — Lohse and Lucht, 1989: 175 (Quedius; characters; Romania; Czechoslovakia; Alps).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic ?; Slovakia ?).

  • — Assing, 1999b: 38 (Quedius; synonym of limbatus).

ponticus Korge, 1964: 121 (Quedius; subgenus Sauridus; subspecies of limbatus; Type locality: Egribel-Pass).

  • — Korge, 1971a: 54 (Quedius; subgenus Sauridus; subspecies of limbatus; characters; Turkey).

  • — Coiffait, 1978: 258 (Quedius; subgenus Sauridus; subspecies of limbatus; characters).

erdciyasicus Korge, 1971a: 55 (Quedius; subgenus Sauridus; subspecies of limbatus; Type locality: Erçiyas-Dag, Anatolia-centr., in der Nähe von Bachläufen in den östlich Ausläufern, 1900–2100 m).

  • — Coiffait, 1978: 258 (Quedius; subgenus Sauridus; subspecies of limbatus; characters).

limbifer Horn, 1878: 162 (Quedius; Type locality: California; [Note: See lectotype designation by Smetana, 1971]. Originally cited from California, Gilroy and Crystal Springs; Mariposa).

  • — Fall and Cockerell, 1907: 166 (Quedius; New Mexico).

  • — Casey, 1915: 412 (Microsaurus; characters; Vancouver Island; California; Guadeloupe Island).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1446 (Quedius; catalog).

  • — Hatch, 1957: 225 (Quedius; subgenus Microsaurus; characters; British Columbia; Washington; Oregon; Idaho).

  • — Smetana, 1971: 111 (Quedius; subgenus Microsaurus; lectotype designation: Cala [California]; characters; British Columbia; California; Idaho; Nevada; Oregon; Washington).

  • — Graham, 1963: 37 (Quedius; collected from cave; California).

  • — Smetana, 1971b: 2 (Quedius; subgenus Microsaurus; California).

  • Distribution: Canada, USA, Mexico.

lin Smetana, 1995: 62 (Quedius; subgenus Distichalius; Type locality: Taiwan: Taichung Hsien, Hsuehshan, above Shan-Liu-Gieu Hut, 3200 m).

  • Distribution: Taiwan.

lindbergi Scheerpeltz, 1963: 15, 25 (Quedius; subgenus Microsaurus; Type locality: Afghanistan: Grotte Kham Zindan).

  • — Coiffait, 1978: 145 (Quedius; subgenus Microsaurus; characters; Afghanistan).

  • Distribution: Afghanistan.

lineipennis Cameron, 1932: 290 (Quedius; subgenus Quedionuchus; Type locality: Burma: Ruby Mines; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1446 (Quedius; catalog).

  • — Smetana, 1988: 297 (Quedius; lectotype designation; characters; Burma).

  • Distribution: Myanmar.

lividipennis Fairmaire and Germain, 1861, see: Loncovilius.

lividipennis Lokay, 1921 [Note: Nomen nudum], see: cinctus Paykull, 1790.

lloydi Donisthorpe, 1945, see: mesomelinus Marsham, 1802.

loebli Smetana, 1978, see: fagelianus Coiffait, 1967.

lokayi M. Dvořák, 1967 [Note: Nomen nudum], see: brevis Erichson, 1840.

longiceps Broun, 1910: 15 (Quedius; Type locality: Raurimu).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; catalog).

  • Distribution: New Zealand.

longicornis Kraatz, 1857d: 494 (Quedius; Type locality: Pommern; Thüringer Wald).

  • — Redtenbacher, 1857: 986 (Quedius; characters; Austria).

  • — Thomson, 1867: 165 (Microsaurus; characters; Scandinavia).

  • — Thomson, 1867a: 43 (Microsaurus; characters; Scandinavia).

  • — Fauvel, 1874: 273 [= 1874a: 499] (Quedius; characters; France; Scandinavia; Britain; Germany; Italy; Spain).

  • — Redtenbacher, 1874: 202 (Quedius; characters).

  • — Mulsant and Rey, 1876: 616 [= 1877: 472] (Quedius; subgenus Ediquus; characters; France).

  • — Fowler, 1888: 228 (Quedius; subgenus Ediquus; characters; Britain).

  • — Ganglbauer, 1895: 396 (Quedius; subgenus Ediquus; characters; Europe).

  • — Porta, 1907: 105 (Quedius; subgenus Ediquus; characters; Italy).

  • — Reitter, 1909: 109 (Quedius; subgenus Ediquus; characters; Germany).

  • — Petri, 1912: 77 (Quedius; cited as new species with Kraatz as author; characters; Romania).

  • — Johansen, 1914: 353 (Quedius; subgenus Ediquus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; catalog).

  • — Gridelli, 1924d: 36 (Quedius; subgenus Microsaurus; characters; Spain; France; Scandinavia; England; Germany; Italy).

  • — Portevin, 1929: 338 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1446 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 6 (Quedius; aedeagal illustration).

  • — Roubal, 1941b: 122, 127 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Paulian, 1941: 261 (Quedius; larval characters).

  • — Jeannel and Jarrige, 1949: 382 (Quedius; subgenus Microsaurus; notes; cave species; Dalmatia).

  • — V. Hansen, 1952: 128 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Smetana, 1958: 344 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 153 (Quedius; subgenus Microsaurus; characters).

  • — Palm, 1963b: 130 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark; Finland).

  • — Lohse, 1964: 207 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Smetana, 1964d: 78 (Quedius; subgenus Microsaurus; Czechoslovakia).

  • — Horion, 1965: 258 (Quedius; subgenus Microsaurus; Ireland; Britain; Sweden; Finland; Holland; Belgium; France; Spain; Portugal; Switzerland; Italy; Romania; Hungary; Czechoslovakia; Poland; Ukraine; Germany; Austria).

  • — Pototskaia, 1967: 83 (Quedius; larval characters).

  • — Szujecki, 1968a: 734 (Quedius; Poland).

  • — Frank, 1969: 266 (Quedius; biological notes; England).

  • — Bordoni, 1973g: 129 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Bordoni, 1974: 76 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Osella and Zanetti, 1975: 114 (Quedius; collected from nest of Talpa europaea; notes; Italy).

  • — Legner and Moore, 1977: 176 (Quedius; found under bark).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 143 (Quedius; subgenus Microsaurus; characters; Europe; England; Iceland; Caucasus).

  • — Frank, 1982: 47 (Quedius; list of parasites).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; France; Benelux; Denmark; Sweden).

  • — Nowosad, 1990: 125 (Quedius; collected from mole nest; notes; Poland).

  • — Welch, 1993: 229 (Quedius; ovariole number and ovary structure).

  • — Wittwer, 1993: 251 (Quedius; Switzerland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Silfverberg, 1999: 50 (Quedius; Estonia).

  • — Eyre, Luff, and Lott, 2000: 34 (Quedius; Britain).

  • Distribution: Iceland ?, Europe, Caucasus.

nigromarginalis Polentz, 1932: 45 (Quedius; form of longicornis; Type locality: Breslau-Dt. Lissa).

  • — Roubal, 1941b: 122 (Quedius; cited as nigromarginata; subgenus Microsaurus; aberration of longicornis; characters).

  • — Smetana, 1958: 344 (Quedius; cited as nigromarginatus; aberration of longicornis).

  • — Smetana, 1960e: 153 (Quedius; aberration of longicornis; cited as nigromarginatus; characters).

longipennis Mannerheim, 1846, see: plagiatus Mannerheim, 1843.

lortetii† Oustalet, 1874: 175 (Quedius; Type locality: Aix en Provence).

  • — Scudder, 1891: 575 (Quedius; catalog; Ligurian; France).

  • — Handlirsch, 1907: 723 (Quedius; catalog; Oligocene; France).

  • Distribution: France.

lucidulus Erichson, 1839a: 495 (Quedius; Type locality: Mark Brandenburg).

  • — Erichson, 1840: 550 (Quedius; characters; Germany).

  • — Redtenbacher, 1849: 826 (Philonthus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 541 (Quedius; characters; France).

  • — Kraatz, 1857d: 520 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 203 (Quedius; characters; Austria).

  • — Fauvel, 1874: 301 [= 1874a: 527] (Quedius; characters; France; Germany; Switzerland; Italy; Austria).

  • — Redtenbacher, 1874: 200 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 264 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 753 [= 1877: 609] (Quedius; subgenus Sauridus; characters; France).

  • — Ganglbauer, 1895: 409 (Quedius; subgenus Sauridus; characters; middle Europe).

  • — Porta, 1907: 143 (Quedius; subgenus Sauridus; characters; Italy).

  • — Reitter, 1909: 114 (Quedius; subgenus Sauridus; characters; Germany).

  • — Johansen, 1914: 369 (Quedius; subgenus Sauridus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; catalog).

  • — Munster, 1923: 199 (Quedius; subgenus Sauridus; Fennoscandia).

  • — Gridelli, 1925: 146 (Quedius; subgenus Raphirus; characters).

  • — Portevin, 1929: 344 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1447 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 50 (Quedius; aedeagal illustration).

  • — V. Hansen, 1952: 151 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Smetana, 1958: 385 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1961d: 134 (Quedius; subgenus Raphirus; Italy).

  • — Smetana, 1962b: 147 (Quedius; subgenus Raphirus; characters; Europe; Caucasus).

  • — Palm, 1963b: 150 (Quedius; subgenus Raphirus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 213 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 294 (Quedius; subgenus Raphirus; Caucasus; Turkey; Russia; Finland; Sweden; Norway; Denmark; all central Europe; Bosnia Hercegovina; Dalmatia; Italy; Spain).

  • — Bordoni, 1973g: 118 (Quedius; subgenus Quediops; characters; Sardinia; Sicily; Italy).

  • — Coiffait, 1978: 203 (Quedius; subgenus Quediops; characters; Europe; Scandinavia; Asia Minor; Caucasus).

  • — Tóth, 1984: 123 (Quedius; characters; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Europe, Caucasus, Turkey.

lugubris Lokay, 1913: 137, 139 (Quedius; subgenus Quedionuchus; Type locality: Czarnohora in den Ost-Karpathen).

  • — Gridelli, 1924d: 77 (Quedius; subgenus Quedionuchus; characters; Carpathians).

  • — Scheerpeltz, 1933: 1447 (Quedius; catalog).

  • — Coiffait, 1978: 190 (Quedius; subgenus Quedionuchus; characters; Carpathians).

  • — Tóth, 1984: 105 (Quedius; characters).

  • Distribution: Ukraine.

lundbergi Palm, 1973: 183 (Quedius; subgenus Microsaurus; Type locality: Kalix, Norrbotten).

  • — Lundberg, 1972: 173 (Quedius; [Note: The name is a nomen nudum here]; Sweden).

  • Distribution: Sweden.

luridipennis W. J. MacLeay, 1873: 142 (Staphylinus; Type locality: Gayndah).

  • — Fauvel, 1877: 274 (Quedius; characters; Australia).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; catalog).

  • — Lea, 1925: 241 (Quedius; characters; Australia).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

semiviolaceus Fauvel, 1878g: 556 (Quedius; Type locality: Australie occidentale et méridionale: Nouvelle-Galles du Sud, Rivière Paroo).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; catalog).

  • — Lea, 1925: 240 (Quedius; synonym of luridipennis).

semiviolaceus Bernhauer and Schubert, 1916: 427 (Quedius; [Note: Nomen nudum (Articles 11.6, 12.1), published without characters and as a junior synonym. Bernhauer and Schubert attributed this name to Blackburn but failed to list a citation for it and I found no description by Blackburn]; synonym of luridipennis).

luridus Fauvel, 1877: 272 (Quedius; Type locality: Australie occidentale, King's George Sound).

  • — Fauvel, 1878g: 554 (Quedius).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; catalog).

  • Distribution: Australia.

luteomaculatus Scheerpeltz, 1958a: 23 (Quedius; subgenus Raphirus; Type locality: bei Ahlat, Nordwest-Ufer des Van-Sees in Armenien).

  • — Coiffait, 1963a: 387 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 262 (Quedius; subgenus Sauridus; characters; Turkey).

  • Distribution: Iran, Turkey.

picicornis Gridelli, 1922b: 128, 131 (Quedius; subgenus Sauridus; [preoccupied]; Type locality: Persia merid.).

  • — Gridelli, 1925: 127 (Quedius; subgenus Raphirus; characters; Persia).

  • — Scheerpeltz, 1933: 1442 (Quedius; synonym of gridellii).

  • — Coiffait, 1963a: 387 (Quedius; subgenus Sauridus; probably synonym of luteomaculatus).

  • — Coiffait, 1978: 262 (Quedius; synonym of luteomaculatus).

gridellii Scheerpeltz, 1933: 1442 (Quedius; replacement name for picicornis Gridelli; [Note: This name is older than luteomaculatus]).

  • — Coiffait, 1978: 262 (Quedius; synonym of luteomaculatus).

lutzi Reitter, 1909, see: semiobscurus Marsham, 1802.

lydus Fagel, 1968a: 193 (Quedius; subgenus Sauridus; Type locality: Anatolie égéenne: Güne Dagh).

  • — Coiffait, 1978: 243 (Quedius; subgenus Sauridus; characters; Anatolia).

  • Distribution: Turkey.

macchabaeus Saulcy, 1864: 637 (Quedius; Type locality: Jérusalem).

  • — Fauvel, 1874: 297 [= 1874a: 523] (Quedius; synonym of coxalis).

  • — Porta, 1907: 139 (Quedius; synonym of coxalis).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; synonym of coxalis).

  • — Gridelli, 1925: 131 (Quedius; synonym of memoralis).

  • — Scheerpeltz, 1933: 1450 (Quedius; synonym of nemoralis).

  • — Smetana, 1958: 377 (Quedius; synonym of nemoralis).

  • — Coiffait, 1963a: 393 (Quedius; subgenus Sauridus; valid species; characters).

  • — Smetana, 1967i: 558 (Quedius; subgenus Raphirus; Turkey).

  • — Fagel, 1968c: 11 (Quedius; subgenus Sauridus; cited as macchabeus; characters; notes; Lebanon; Iran; Syria).

  • — Coiffait, 1978: 246 (Quedius; subgenus Sauridus; characters; Lebanon; Palestine).

  • Distribution: Iran, Turkey, Syria, Lebanon, Israel.

macrops Lea, 1929: 205 (Quedius; Type locality: Queensland: Herberton; New South Wales: Sydney).

  • — Scheerpeltz, 1933: 1447 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

maculatus Korge, 1971 [Note: Nomen nudum], see: vicinus Ménétriés, 1832.

maculicollis W. Kolbe, 1909 [Note: Nomen nudum], see: collaris Erichson, 1840.

maculicornis Mulsant and Rey, 1876, see: molochinus Gravenhorst, 1806.

maculiventris Bernhauer, 1934: 12 (Quedius; subgenus Raphirus; Type locality: Mittel-Szechuan: Hwa-Yin-Shan, 1600 m; [Note: See lectotype designation by Smetana, 1990a]).

  • — Smetana, 1990a: 200 (Quedius; subgenus Raphirus; lectotype designation; characters; China).

  • — Smetana, 1995: 68 (Quedius; subgenus of Raphirus; characters; bionomics; Taiwan).

  • Distribution: China, Taiwan.

maderensis Smetana, 1963, see: nigriceps Kraatz, 1857.

magarasiensis Bordoni, 1978a: 65 (Quedius; subgenus Microsaurus; Type locality: Turchia: Isparta, Anamas, Grotta Zindan Magarasi).

  • — Bordoni and Oromi, 1998: 1157 (Quedius; facultative cave species).

  • Distribution: Turkey.

magniceps Bernhauer, 1914d: 3 (Quedius; Type locality: Sizilien bei Ficuzza).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; catalog).

  • — Gridelli, 1922b: 127 (Quedius; subgenus Sauridus; characters; Sicily).

  • — Bernhauer, 1923d: 177 (Quedius; subgenus Sauridus; characters; Sizilien).

  • — Gridelli, 1925: 148 (Quedius; subgenus Raphirus; characters; Sicily).

  • — Scheerpeltz, 1933: 1447 (Quedius; catalog).

  • — Coiffait, 1978: 282 (Quedius; subgenus Raphirus; characters; Sicily).

  • — Sparacio, 1995: 165 (Quedius; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Italy.

malaisei Scheerpeltz, 1965: 290 (Quedius; subgenus Indoquedius; Type locality: N.E. Burma, Kambaiti, 2000 m).

  • Distribution: Myanmar.

malinalli Smetana, 1975: 320 (Quedius; subgenus Raphirus; Type locality: Mexico: Oaxaca: 52 mi. N. Oaxaca, 9500 ft.).

  • Distribution: Mexico.

mallius Tottenham, 1948a: 256 (Quedius; subgenus Arphirus; Type locality: Britain: Welwyn).

  • — Smetana, 1958: 396 (Quedius; subgenus Raphirus; characters; ecology; notes).

  • — Fagel, 1960b: 116 (Quedius; notes).

  • — Smetana, 1962b: 152 (Quedius; cited as possible synonym of aridulus).

  • — Pope, 1977: 31 (Quedius; synonym of aridulus).

  • — Coiffait, 1978: 279 (Quedius; subgenus Raphirus; cited as valid species; characters; England).

  • Distribution: Britain.

manaiaensis Broun, 1910, see: Philonthus.

mandli Bernhauer, 1910b: 257 (Quedius; subgenus Quedionuchus; Type locality: Ostpyrenäen, Plat de Guilhem).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; catalog).

  • — Gridelli, 1924d: 77 (Quedius; subgenus Quedionuchus; characters; Pyrenees).

  • — Scheerpeltz, 1933: 1447 (Quedius; catalog).

  • — Korge, 1964: 118 (Quedius; subgenus Quedionuchus; characters).

  • — Coiffait, 1978: 189 (Quedius; subgenus Quedionuchus; characters; Pyrenees).

  • Distribution: Spain, France.

mandra Smetana, 1975, see: kashmirensis Cameron, 1944.

manducus Herman, 2001: 49 (Quedius; replacement name for analis W. J. MacLeay).

  • Distribution: Australia.

analis W. J. MacLeay, 1873: 142 (Staphylinus; [preoccupied]; Type locality: Gayndah).

  • — Fauvel, 1877: 274 (Quedius; characters; Australia).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Herman, 2001: 49 (Quedius; synonym of manducus).

manitobensis Casey, 1915: 423 (Megaquedius; Type locality: Manitoba: Aweme).

  • — Scheerpeltz, 1933: 1447 (Quedius; catalog).

  • — Smetana, 1971: 47 (Quedius; subgenus Megaquedius; characters; Manitoba).

  • — Smetana, 1981b: 633 (Quedius; Alberta).

  • Distribution: Canada.

mannaiaensis Cameron, 1945d: 788 (Quedius; subgenus Microsaurus; Type locality: “. . . type . . . is without locality label.” Other specimens from Korokoro and Wellington).

  • Distribution: New Zealand.

maorinus Broun, 1923: 679 (Quedius; Type locality: Pakarau and Kaiangaroa).

  • — Scheerpeltz, 1933: 1447 (Quedius; catalog).

  • Distribution: New Zealand.

marcuzzii Scheerpeltz, 1953, see: alpestris Heer, 1839.

marginalis Mäklin, 1852, see: transparens Motschulsky, 1845.

marginalis Kraatz, 1857, see: nemoralis Baudi, 1848.

marginalis Oke, 1933, see: okei Korge, 1963.

marginatus Korge, 1962 [Note: Nomen nudum], see: laticollis Gravenhorst, 1802.

marginipennis Bernhauer, 1939d: 103 (Quedius; subgenus Sauridus; Type locality: Japan: Kamikochi).

  • Distribution: Japan.

maritimus J. Sahlberg, 1876, see: umbrinus Erichson, 1839.

maronitus Coiffait, 1963a: 385, 410 (Quedius; subgenus Sauridus; Type locality: Liban: Aïn Batara).

  • — Coiffait, 1978: 234 (Quedius; subgenus Sauridus; characters; Lebanon).

  • Distribution: Lebanon.

martensi Smetana, 1975a: 329 (Quedius; subgenus Microsaurus; Type locality: Nepal: Khumbu, confluence Imja, Phunki-Drangka, 3150–3300 m).

  • — Smetana, 1988: 203 (Quedius; subgenus Microsaurus; characters; Nepal).

  • Distribution: Nepal.

martini Smetana, 1975b: 35 (Quedius; subgenus Megaquedius; Type locality: Mexico: Durango, 10 mi. W El Salto, 9000 ft.).

  • Distribution: Mexico.

masoni Zanetti, 1992b: 126 (Quedius; subgenus Sauridus; Type locality: Monti dell'Uccellina, Toscana, prov. di Grosseto, dint. di Alberese).

  • — Sparacio, 1995: 165 (Quedius; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Italy.

masuzoi Watanabe, 1989b: 170 (Quedius; subgenus Microsaurus; Type locality: Taiwan: Nan-t'ou Hsien, Mt. Neng-kao-pei-feng, 2870 m).

  • — Smetana, 1995: 45 (Quedius; subgenus of Microsaurus; characters; bionomics; Taiwan).

  • Distribution: Taiwan.

mateui Coiffait, 1967 [Note: Nomen nudum], see: mesomelinus Marsham, 1802.

matinai Roubal, 1950 [Note: Nomen nudum], see: mesomelinus Marsham, 1802.

maurorufus Gravenhorst, 1806: 56 (Staphylinus; Type locality: Brunsvigae).

  • — C. Sahlberg, 1830: 317 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 26 (Staphylinus; Finland).

  • — Mannerheim, 1831: 440 (Staphylinus; Finland).

  • — Runde, 1835: 6 (Staphylinus; characters; Germany).

  • — Lacordaire, 1835: 382 (Emus; characters; France).

  • — Lacordaire, 1835: 382 (Emus; [Note: Misidentification: Bernhauer and Schubert, 1916: 427, cited praecox Boisduval and Lacordaire as a synonym of maurorufus; Boisduval and Lacordaire did not describe the species as new, they attributed it to Gravenhorst]; characters; France).

  • — Nordmann, 1837: 77 (Philonthus; Europe).

  • — Zetterstedt, 1838: 61 (Staphylinus; characters; habitat; Lapland).

  • — Heer, 1839: 278 (Philonthus; characters; Switzerland).

  • — Stephens, 1839: 393 (Quedius; characters; England).

  • — Erichson, 1840: 542 (Quedius; characters; Germany; Sweden).

  • — Kolenati, 1846: 21 (Quedius; characters; Somchetiae).

  • — P. Lucas, 1846: 114 (Quedius; notes; Algeria).

  • — Gravenhorst, 1847: 228 (Quedius; characters; notes).

  • — Kiesenwetter, 1848: 60 (Quedius; characters; Germany).

  • — Hochhuth, 1849: 152 (Quedius; characters; Iberia).

  • — Redtenbacher, 1849: 711 (Philonthus; characters; Austria).

  • — Hardy, 1851: 34 (Quedius; notes; England).

  • — Kraatz, 1857d: 512 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 204 (Quedius; characters; Austria).

  • — Thomson, 1860: 178 (Raphirus; characters; Scandinavia).

  • — Fauvel, 1874: 300 [= 1874a: 526] (Quedius; characters; France; Scandinavia; Britain; Germany; Switzerland; Austria; Algeria; Morocco; Caucasus).

  • — Redtenbacher, 1874: 201 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 265 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 744 [= 1877: 600] (Quedius; subgenus Sauridus; characters; France).

  • — Fauvel, 1878: 129 [= 1878a: 49] (Quedius; Morocco; Algeria).

  • — Fauvel, 1886: 60 [= 1886a: 52] (Quedius; Morocco; Algeria).

  • — Fowler, 1888: 240 (Quedius; subgenus Sauridus; characters; Britain).

  • — Ganglbauer, 1895: 408 (Quedius; subgenus Sauridus; characters; north and middle Europe).

  • — Fauvel, 1902b: 117 (Quedius; Morocco; Algeria).

  • — Porta, 1907: 141 (Quedius; subgenus Sauridus; characters; Italy).

  • — Reitter, 1909: 113 (Quedius; subgenus Sauridus; characters; Germany).

  • — Johansen, 1914: 367 (Quedius; subgenus Sauridus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; synonym of limbatus).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; catalog).

  • — Gridelli, 1922b: 127, 138 (Quedius; subgenus Sauridus; characters).

  • — Gridelli, 1925: 128 (Quedius; subgenus Raphirus; characters; Alps; Pyrenees; Germany; Scandinavia; England; Austria; France).

  • — Portevin, 1929: 343 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1447 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 38 (Quedius; aedeagal illustration).

  • — Jeannel and Jarrige, 1949: 379 (Quedius; subgenus Raphirus; cave species; Romania).

  • — V. Hansen, 1952: 146 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Smetana, 1958: 376 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 144 (Quedius; subgenus Raphirus; characters; Europe).

  • — Coiffait, 1963a: 386 (Quedius; subgenus Sauridus; characters).

  • — Palm, 1963b: 148 (Quedius; subgenus Raphirus; characters; Sweden; Denmark; Finland).

  • — Lohse, 1964: 216 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Smetana, 1964d: 80 (Quedius; subgenus Raphirus; Czechoslovakia).

  • — Horion, 1965: 285 (Quedius; subgenus Raphirus; Ireland; Britain; Denmark; Sweden; Finland; Holland; Belgium; France; Spain; Portugal; Switzerland; Czechoslovakia; Hungary; Romania; Poland; Italy ?; Germany; Austria).

  • — Szujecki, 1968a: 736 (Quedius; Poland).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 253 (Quedius; subgenus Sauridus; characters; Europe; Britain; Scandinavia; Alps; Pyrenees).

  • — Topp, 1979: 20 (Quedius; development; diapause).

  • — Hinton, 1981: 663 (Quedius; characters of egg).

  • — Frank, 1982: 47 (Quedius; list of parasites).

  • — Tóth, 1984: 139 (Quedius; characters; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Lohse and Lucht, 1989: 173 (Quedius; characters).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Europe, Caucasus.

modestus Kraatz, 1857d: 509 (Quedius; Type locality: Deutschland).

  • — Redtenbacher, 1857: 986 (Quedius; characters; Austria).

  • — Fauvel, 1865a: 293 [= 1865b: 43] (Quedius; habitat; France).

  • — Fauvel, 1874: 300 [= 1874a: 526] (Quedius; synonym of maurorufus).

  • — Redtenbacher, 1874: 202 (Quedius; characters).

  • — Fowler, 1888: 240 (Quedius; subgenus Sauridus; synonym of maurorufus).

  • — Ganglbauer, 1895: 408 (Quedius; synonym of maurorufus).

  • — Porta, 1907: 141 (Quedius; synonym of maurorufus).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; synonym of maurorufus).

  • — Gridelli, 1925: 128 (Quedius; synonym of maurorufus).

  • — Scheerpeltz, 1933: 1447 (Quedius; synonym of maurorufus).

  • — Smetana, 1958: 376 (Quedius; synonym of maurorufus).

  • — Coiffait, 1978: 253 (Quedius; synonym of maurorufus).

acuminatus Fairmaire, 1860a: 155 (Heterothops; [preoccupied]; Type locality: Tanger).

  • — Fauvel, 1874: 300 [= 1874a: 526] (Quedius; synonym of maurorufus).

  • — Fowler, 1888: 240 (Quedius; subgenus Sauridus; synonym of maurorufus).

  • — Ganglbauer, 1895: 408 (Quedius; synonym of maurorufus).

  • — Porta, 1907: 141 (Quedius; synonym of maurorufus).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; synonym of maurorufus).

  • — Scheerpeltz, 1933: 1447 (Quedius; doubtful synonym of maurorufus).

  • — Smetana, 1958: 376 (Quedius; synonym of maurorufus).

  • — Coiffait, 1978: 253 (Quedius; synonym of maurorufus).

maurus C. Sahlberg, 1830: 317 (Staphylinus; Type locality: Willnäs).

  • — Mannerheim, 1830: 26 (Staphylinus; Finland).

  • — Mannerheim, 1831: 440 (Staphylinus; Finland).

  • — Nordmann, 1837: 79 (Philonthus; cited as marus; characters; Finland).

  • — Erichson, 1840: 551 (Quedius; characters; Finland).

  • — Kraatz, 1857d: 501 (Quedius; Finland).

  • — Hochhuth, 1872: 133 (Quedius; notes; characters; Ukraine).

  • — Fauvel, 1874: 278 [= 1874a: 504] (Quedius; synonym of fageti).

  • — Seidlitz, 1875: 266 (Quedius; characters; Baltic region).

  • — J. Sahlberg, 1876: 26 (Microsaurus; Finland).

  • — Ganglbauer, 1895: 400 (Quedius; subgenus Microsaurus; characters; north and middle Europe; Caucasus; Canary Islands).

  • — Bernhauer, 1900: 46 (Quedius; notes).

  • — Newbery, 1905: 197 (Quedius; [Note: Misidentification: Scheerpeltz, 1934: 1448, cited variabilis Newbery as a synonym of maurus; Newbery did not describe the species as new, he attributed it to Heer]; characters; Britain).

  • — Porta, 1907: 125 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Reitter, 1909: 110 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Johansen, 1914: 359 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; catalog).

  • — Munster, 1923: 193 (Quedius; subgenus Microsaurus; Fennoscandia).

  • — Gridelli, 1924d: 63 (Quedius; subgenus Microsaurus; characters; Silesia; Italy; Bosnia; Romania).

  • — Gridelli, 1929: 36 (Quedius; characters).

  • — Scheerpeltz, 1933: 1447 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 14 (Quedius; aedeagal illustration).

  • — Roubal, 1941b: 125, 129 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Jeannel and Jarrige, 1949: 378 (Quedius; subgenus Microsaurus; cave species; Belgium).

  • — V. Hansen, 1952: 136 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Smetana, 1958: 355 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 159 (Quedius; subgenus Microsaurus; characters).

  • — Korge, 1961: 44 (Quedius; characters; Europe).

  • — Palm, 1963b: 135 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 209 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Smetana, 1964d: 79 (Quedius; subgenus Microsaurus; Czechoslovakia).

  • — Horion, 1965: 267 (Quedius; subgenus Microsaurus; Britain; Norway; Sweden; Finland; Russia; Holland; Belgium; Switzerland; France; Spain; Italy; Dalmatia; Bosnia Hercegovina; Romania; Hungary; Czechoslovakia; Poland; Germany; Austria).

  • — Szujecki, 1968a: 735 (Quedius; Poland).

  • — Bordoni, 1974: 42 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Osella and Zanetti, 1975: 120 (Quedius; collected from nest of Talpa europaea; notes; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 151 (Quedius; subgenus Microsaurus; characters; Europe; Scandinavia; Britain; Asia Minor; Caucasus).

  • — Tóth, 1984: 113 (Quedius; characters; Hungary).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Nowosad, 1990: 141 (Quedius; collected from mole nest; notes; Poland).

  • — Whitehead, 1990e: 11 (Quedius; collecting notes; Britain).

  • — Allen, 1990b: 3 (Quedius; collecting notes; Britain).

  • — Smetana, 1993b: 50 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Owen, 1999b: 61 (Quedius; captured underground at roots of old trees; Britain).

  • — Fowles, Alexander, and Key, 1999: 129 (Quedius; saproxylic species; status as endangered or threatened species; Britain).

  • — Owen, 2000: 252 (Quedius; pitfall trap collections from underground at roots of trees; Britain).

  • Distribution: Europe, Russia, Caucasus, Turkey.

fageti Thomson, 1867: 161 (Microsaurus; Type locality: Skane).

  • — Thomson, 1867a: 43 (Microsaurus; characters; Scandinavia).

  • — Fauvel, 1874: 277 [= 1874a: 503] (Quedius; characters; France; Scandinavia; Russia; Belgium).

  • — Fauvel, 1876a: 76 [= 1876: 261] (Quedius; synonym of maurus).

  • — Mulsant and Rey, 1876: 638 [= 1877: 494] (Quedius; characters; France).

  • — Fowler, 1888: 232 (Quedius; variety of mesomelinus; characters; Britain).

  • — Ganglbauer, 1895: 400 (Quedius; synonym of maurus).

  • — Porta, 1907: 125 (Quedius; synonym of maurus).

  • — Gridelli, 1924d: 63 (Quedius; synonym of maurus).

  • — Portevin, 1929: 340 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1448 (Quedius; synonym of maurus).

  • — Smetana, 1958: 355 (Quedius; synonym of maurus).

  • — Bordoni, 1974: 44 (Quedius; synonym of maurus).

  • — Pope, 1977: 31 (Quedius; synonym of maurus).

  • — Coiffait, 1978: 152 (Quedius; synonym of maurus).

pyropterus Eppelsheim, 1880c: 583 (Quedius; form of maurus; Type locality: Caucasus; Schlesien).

  • — Duvivier, 1883: 139 (Quedius; catalog; synonym of maurus).

  • — Ganglbauer, 1895: 400 (Quedius; synonym of maurus).

  • — Porta, 1907: 125 (Quedius; variety of maurus).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; variety of maurus).

  • — Gridelli, 1924d: 63 (Quedius; variety of maurus).

  • — Scheerpeltz, 1933: 1448 (Quedius; variety of maurus).

  • — Roubal, 1941b: 125, 129 (Quedius; subgenus Microsaurus; variety of maurus; characters; collecting notes).

  • — Smetana, 1958: 355 (Quedius; variety of maurus).

  • — Smetana, 1960e: 159 (Quedius; aberration of maurus; characters).

  • — Korge, 1964: 117 (Quedius; subgenus Microsaurus; aberration of maurus; Turkey).

  • — Bordoni, 1974: 44 (Quedius; variety of maurus).

  • — Coiffait, 1978: 152 (Quedius; subgenus Microsaurus; variety of maurus; characters).

mazatl Smetana, 1975: 311 (Quedius; subgenus Microsaurus; Type locality: Mexico: Nuevo Leon: Cerro Potosi).

  • — Smetana, 1976: 117 (Quedius; subgenus Microsaurus; Mexico).

  • Distribution: Mexico.

medicatus Last, 1975: 441 (Quedius; subgenus Sauridus; Type locality: Amusei, 5800 ft., North Fore census division, Okapa subdistrict).

  • Distribution: New Guinea.

mediofuscus Lea, 1910: 121 (Quedius; Type locality: Tasmania: Mount Wellington).

  • — Scheerpeltz, 1933: 1448 (Quedius; catalog).

  • Distribution: Australia.

meei Smetana, 1995: 85 (Quedius; subgenus Raphirus; Type locality: Taiwan: Nantou Hsien: Houhuanshan Kuenyang, 3050 m).

  • Distribution: Taiwan.

megalops Wollaston, 1864: 564 (Quedius; Type locality: Grand Canary: El Monte; Palma; Hierro: El Golfo; Teneriffe).

  • — Wollaston, 1865: 486 (Quedius; habitat; Canary Islands).

  • — Fauvel, 1902b: 117 (Quedius; Canaries).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; catalog).

  • — Gridelli, 1925: 148 (Quedius; subgenus Raphirus; characters; Canaries).

  • — Scheerpeltz, 1933: 1448 (Quedius; catalog).

  • — Coiffait, 1954: 167 (Quedius; subgenus Raphirus; Canary Islands).

  • — Coiffait, 1978: 264 (Quedius; subgenus Raphirus; characters; Canary Islands).

  • — Assing, 1999a: 346 (Quedius; Canary Islands).

  • — Assing and Wunderle, 1999: 437 (Quedius; Canary Islands).

  • Distribution: Canary Islands.

megophthalmus Broun, 1917: 373 (Quedius; Type locality: Routeburn).

  • — Scheerpeltz, 1933: 1448 (Quedius; catalog).

  • Distribution: New Zealand.

melanocephalus Mannerheim, 1852: 315 (Quedius; Type locality: America borealis ad Tschishlkhath).

  • — Horn, 1878: 159 (Quedius; variety of fulgidus; characters).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1448 (Quedius; catalog).

  • Distribution: USA.

melas Lea, 1925: 251 (Quedius; Type locality: New South Wales: Blue Mountains. Victoria: Melbourne, Ringwood, Beaconsfield).

  • — Scheerpeltz, 1933: 1448 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

meledanus Reitter, 1900, see: scitus Gravenhorst, 1806.

melium Walles, 1938, see: puncticollis Thomson, 1867.

meng Smetana, 1995: 59 (Quedius; subgenus Distichalius; Type locality: Taiwan: Nantou Hsien, Houhuanshan Kuenyang, 3050 m).

  • Distribution: Taiwan.

meridiocarpathicus Smetana, 1958: 365 (Quedius; subgenus Quedius; Type locality: Slovenska, Kamenný Most, Slanec).

  • — Coiffait, 1961a: 53 (Quedius; characters; Greece; Slovakia; Turkey).

  • — Korge, 1962: 153 (Quedius; subgenus Quedius; synonymic notes).

  • — Smetana, 1962b: 138 (Quedius; subgenus Quedius; characters).

  • — Lohse, 1964: 212 (Quedius; subgenus Quedius; characters; central Europe).

  • — Horion, 1965: 277 (Quedius; subgenus Quedius; Turkey; Ukraine; Greece; Dalmatia; Italy; Bulgaria; Slovakia; Austria).

  • — Smetana, 1967i: 558 (Quedius; Bulgaria).

  • — Bordoni, 1976b: 99 (Quedius; characters; Italy).

  • — Coiffait, 1978: 195 (Quedius; subgenus Quedius; characters; Europe; Slovenia; Bulgaria; Greece; Corfu; Turkey).

  • — Tóth, 1984: 129 (Quedius; characters; Hungary).

  • — Lucht, 1987: 109 (Quedius; checklist; Czechoslovakia; Austria).

  • — Smetana, 1993b: 50 (Quedius; subgenus Quedius; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Europe, Turkey.

dalmatinus Korge, 1960a: 68 (Quedius; subspecies of molochinus; Type locality: Dalmatia: Castelnuovo).

  • — Coiffait, 1961a: 53 (Quedius; subspecies of molochinus; characters; Dalmatia).

  • — Korge, 1962: 153 (Quedius; synonym of meridiocarpathicus).

  • — Smetana, 1967i: 558 (Quedius; synonym of meridiocarpathicus).

  • — Coiffait, 1978: 194 (Quedius; subgenus Quedius; subspecies of molochinus; [Note: I assume Coiffait overlooked the synonymy by Korge, 1962]; characters).

  • — Tóth, 1984: 120 (Quedius; subspecies of molochinus; characters; Hungary).

merlini Drugmand and Bruge, 1991: 192 (Quedius; subgenus Sauridus; Type locality: Belgique: Tenneville [code U.T.M.: MOMU FR75], Fange Massa).

  • Distribution: Belgium.

mesomelinus Marsham, 1802: 510 (Staphylinus; Type locality: Britain).

  • — Gravenhorst, 1802: 31 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1916: 428, 430, cited nitidus Gravenhorst as a synonym of mesomelinus and ochripennis but Gravenhorst did not describe the species as new, he attributed it to Fabricius; Gravenhorst cited many varieties of the species]; characters).

  • — Gyllenhal, 1810: 341 (Staphylinus; synonym of opacus).

  • — C. Sahlberg, 1830: 328 (Staphylinus; synonym of opacus).

  • — Erichson, 1840: 526 (Quedius; [Note: Misidentification: Bernhauer and Schubert, 1916: 428, cited an unspecified variety of fulgidus Erichson as a synonym of mesomelinus; there is no available name]; characters).

  • — Erichson, 1840: 526 (Quedius; synonym of fulgidus).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — Fauvel, 1874: 280 [= 1874a: 506] (Quedius; characters; France; Europe; USA; Peru).

  • — Mulsant and Rey, 1876: 641 [= 1877: 497] (Quedius; characters; France).

  • — J. Sahlberg, 1876: 25 (Microsaurus; Finland).

  • — Fauvel, 1878g: 552 (Quedius; characters; Australia).

  • — Fowler, 1888: 232 (Quedius; characters; Britain).

  • — Ganglbauer, 1895: 400 (Quedius; subgenus Microsaurus; characters; Europe; North America; Peru; Australia; New Zealand).

  • — Porta, 1907: 115 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Sainte-Claire Deville, 1906: 101 (Quedius; Corsica).

  • — Montandon, 1908: 80 (Quedius; Romania).

  • — Reitter, 1909: 110 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Johansen, 1914: 358 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Casey, 1915: 411 (Microsaurus; characters; British Columbia; California; New York).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; catalog).

  • — Henriksen and Lundbeck, 1918: 494 (Quedius; Greenland).

  • — Šulc, 1922 (Quedius; osmeterium; anatomy; histology; physiology).

  • — Munster, 1923: 192 (Quedius; subgenus Microsaurus; Fennoscandia).

  • — Gridelli, 1924d: 60 (Quedius; subgenus Microsaurus; characters).

  • — Lea, 1925: 240 (Quedius; Australian introduction).

  • — Portevin, 1929: 340 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1448 (Quedius; catalog).

  • — West, 1937: 35 (Quedius; Faroe Islands).

  • — Wüsthoff, 1938a: fig. 13 (Quedius; aedeagal illustration).

  • — Henriksen, 1939: 43 (Quedius; Greenland).

  • — Roubal, 1941b: 125, 129 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Paulian, 1941: 266 (Quedius; larval characters).

  • — Hinton, 1945: 71 (Quedius; characters).

  • — Scheerpeltz and Höfler, 1948: 157 (Quedius; subgenus Microsaurus; fungus inhabitant).

  • — Brinck, 1948: 31 (Quedius; collecting and ecological notes; larval characters; Tristan da Cunha).

  • — Tottenham, 1949: 422 (Quedius; checklist; Britain).

  • — Jeannel and Jarrige, 1949: 384 (Quedius; subgenus Microsaurus; notes; cave species; discussion of distribution; France; Spain; Slovenia; Serbia; Romania).

  • — V. Hansen, 1952: 135 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Hatch, 1957: 223 (Quedius; subgenus Microsaurus; characters; British Columbia; Washington; Oregon; Idaho).

  • — Cerruti, 1957 (Quedius; Italy).

  • — Smetana, 1958: 353 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Larsson and Gígja, 1959: 90 (Quedius; biological notes; Iceland).

  • — Smetana, 1960e: 159 (Quedius; subgenus Microsaurus; characters).

  • — Korge, 1961: 45 (Quedius; characters; discussion of infraspecific names).

  • — Ferreira, 1962: 19 (Quedius; Portugal).

  • — Palm, 1963b: 135 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 209 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Smetana, 1964d: 79 (Quedius; subgenus Microsaurus; Czechoslovakia).

  • — Raianu, 1964: 103 (Quedius; Romania).

  • — Smetana, 1965: 42 (Quedius; subgenus Microsaurus; characters; Labrador; Newfoundland).

  • — Smetana, 1965a: 13 (Quedius; subgenus Microsaurus; Alaska).

  • — Horion, 1965: 266 (Quedius; subgenus Microsaurus; all Europe).

  • — Pototskaia, 1967: 83 (Quedius; larval characters).

  • — Szujecki, 1968a: 734 (Quedius; Poland).

  • — Scheerpeltz, 1968a: 89 (Quedius; catalog; Austria).

  • — Korge, 1969: 43 (Quedius; notes; Bulgaria).

  • — Kasule, 1970: 63 (Quedius; larval characters).

  • — Smetana, 1971: 74 (Quedius; subgenus Microsaurus; characters; bionomics; notes; Alberta; British Columbia; Manitoba; Miquelon; Labrador; Newfoundland; Nova Scotia; Ontario; Quebec; Alaska; California; Colorado; Connecticut; Florida; Georgia; Illinois; Indiana; Iowa; Maine; Massachusetts; Michigan; Missouri; Montana; Nevada; New Hampshire; New Jersey; New York; North Carolina; Ohio; Oregon; Pennsylvania; Utah; Virginia; Washington; West Virginia; Wisconsin).

  • — Smetana, 1971b: 2 (Quedius; subgenus Microsaurus; Washington).

  • — Bordoni, 1974: 33 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Bordoni, 1974b: 9 (Quedius; subgenus Microsaurus; notes; Italy).

  • — Osella and Zanetti, 1975: 119 (Quedius; collected from nest of Talpa europaea; notes; Italy).

  • — Shilov, 1975a: 376 [= 1975b: 99] (Quedius; Komi region of Russia).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Bordoni, 1977a: 18 (Quedius; subgenus Microsaurus; notes; Spain).

  • — Coiffait, 1978: 150 (Quedius; subgenus Microsaurus; characters; Cosmopolitan).

  • — Muona, 1979: 15 (Quedius; Finland; Russia; Sweden; Norway; Denmark; Baltic States).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 124 (Quedius; catalog; Poland).

  • — Hinton, 1981: 662 (Quedius; characters of egg).

  • — Bordoni, 1982a: 140 (Quedius; cave collection; Sardinia).

  • — Frank, 1982: 47 (Quedius; list of parasites).

  • — Tóth, 1984: 112 (Quedius; characters; Hungary).

  • — Muona and Viramo, 1986: 13 (Quedius; Finland).

  • — Lucht, 1987: 109 (Quedius; nominate subspecies; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Nowosad, 1990: 139 (Quedius; collected from mole nest; notes; Poland).

  • — Whitehead, 1990e: 11 (Quedius; collecting notes; Britain).

  • — Allen, 1990b: 3 (Quedius; collecting notes; Britain).

  • — Silfverberg, 1992: 17 (Quedius; Finland; Russia; Sweden; Norway; Denmark; Estonia; Latvia; Lithuania).

  • — Welch, 1993: 229 (Quedius; ovariole number and ovary structure).

  • — Smetana, 1993b: 50 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Dearborn and Donahue, 1994: 15 (Quedius; Maine).

  • — Sadler and Dugmore, 1995: 144 (Quedius; habitat notes; Iceland).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Downie and Arnett, 1996: 391 (Quedius; subgenus Microsaurus; characters; USA).

  • — M. Hansen, 1996: 107 (Quedius; checklist; Denmark).

  • — Franzen, 1996: 207 (Quedius; collecting notes; Germany).

  • — Ádám, 1996: 247 (Microsaurus; synonym of fuscipennis).

  • — Eisinger, 1997: 171 (Quedius; collecting notes; Germany).

  • — T. Wagner, 1997: 235 (Quedius; collecting notes; Germany).

  • — Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 39 (Quedius; Latvia).

  • — Anderson, Nash, and O'Connor, 1997: 19 (Quedius; Ireland).

  • — Hansen, Jørum, Palm, and Pedersen, 1997: 128 (Quedius; checklist; Denmark).

  • — Allen, 1998a: 126 (Quedius; collecting notes; Britain).

  • — Bain, 1998: 39, 43 (Quedius; 17th century remains; introduced; Massachusetts).

  • — Bordoni and Oromi, 1998: 1157 (Quedius; facultative cave species).

  • — Outerelo, Gamarra, and Salgado, 1998: 130 (Quedius; subgenus Microsaurus; collected from cave; notes; Spain; Portugal).

  • — Owen, 1999a: 15 (Quedius; subterranean capture; Britain).

  • — Owen, 1999b: 61 (Quedius; captured underground at roots of old trees; Britain).

  • — Staniec, 1999a: 50 (Quedius; characters of pupa; Poland).

  • — Ryabukhin, 1999: 72 (Quedius; subgenus Microsaurus; catalog; habitat; distribution; northeastern Russia).

  • — Owen, 2000: 252 (Quedius; pitfall trap collections from underground at roots of trees; Britain).

  • Distribution: Tristan da Cunha, Greenland, Iceland, Europe, Russia, Australia, New Zealand, Canada, USA, Peru.

variabilis Gyllenhal, 1810: 303 (Staphylinus; [Note: Bernhauer and Schubert, 1916: 428, 432, cited variabilis Gyllenhal as a synonym of both mesomelinus and puncticollis]; Type locality: Suecia).

  • — Gyllenhal, 1827: 478 (Staphylinus; characters for variety).

  • — Curtis, 1829: 25 (Quedius; synonym of rufitarsis Marsham).

  • — C. Sahlberg, 1830: 315 (Staphylinus; characters; Finland).

  • — Stephens, 1832: 218 (Quedius; characters; England).

  • — Nordmann, 1837: 78 (Philonthus; Europe).

  • — J. Sahlberg, 1876: 25 (Microsaurus; synonym of mesomelinus).

  • — Ganglbauer, 1895: 400 (Quedius; synonym of mesomelinus).

  • — Porta, 1907: 113 (Quedius; synonym of ochripennis).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; synonym of mesomelinus).

  • — Smetana, 1958: 353 (Quedius; synonym of mesomelinus).

  • — Smetana, 1971: 74 (Quedius; synonym of mesomelinus).

  • — Bordoni, 1974: 33 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 150 (Quedius; synonym of mesomelinus).

skrimshiranus Stephens, 1832: 225 (Quedius; Type locality: Norfolk: Fakenham).

  • — Curtis, 1837: pl. 638 (Quedius; notes; Britain).

  • — Stephens, 1839: 392 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 22 (Quedius; synonym of fulgidus).

  • — Gemminger and Harold, 1868: 569 (Quedius; synonym of fulgidus).

  • — Fauvel, 1874: 281 [= 1874a: 507] (Quedius; synonym of mesomelinus).

  • — Ganglbauer, 1895: 400 (Quedius; synonym of mesomelinus).

  • — Porta, 1907: 116 (Quedius; synonym of mesomelinus).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; synonym of mesomelinus).

  • — Smetana, 1958: 353 (Quedius; synonym of mesomelinus).

  • — Smetana, 1971: 74 (Quedius; synonym of mesomelinus).

  • — Bordoni, 1974: 33 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 150 (Quedius; synonym of mesomelinus).

occultus Lacordaire, 1835: 379 (Emus; Type locality: Paris).

  • — Erichson, 1840: 526 (Quedius; synonym of fulgidus).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — Fauvel, 1865a: 292 [= 1865b: 43] (Quedius; habitat; France).

  • — Fauvel, 1869: 486 (Quedius; synonym of mesomelinus).

  • — Fauvel, 1874: 281 [= 1874a: 507] (Quedius; synonym of mesomelinus).

  • — Ganglbauer, 1895: 400 (Quedius; synonym of mesomelinus).

  • — Porta, 1907: 116 (Quedius; synonym of mesomelinus).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; synonym of mesomelinus).

  • — Smetana, 1958: 353 (Quedius; synonym of mesomelinus).

  • — Smetana, 1971: 74 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 150 (Quedius; synonym of mesomelinus).

atripennis Stephens, 1835: 436 (Quedius; Type locality: Scotland).

  • — Curtis, 1837: pl. 638 (Quedius; characters; Britain).

  • — Stephens, 1839: 391 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 22 (Quedius; synonym of fulgidus).

  • — Fauvel, 1869: 486 (Quedius; synonym of mesomelinus).

  • — Ganglbauer, 1895: 400 (Quedius; synonym of mesomelinus).

  • — Porta, 1907: 116 (Quedius; synonym of mesomelinus).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; synonym of mesomelinus).

  • — Smetana, 1958: 353 (Quedius; synonym of mesomelinus).

  • — Smetana, 1971: 74 (Quedius; synonym of mesomelinus).

  • — Bordoni, 1974: 33 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 150 (Quedius; synonym of mesomelinus).

groenlandicus Zetterstedt, 1838: 61 (Staphylinus; Type locality: Groenlandia).

  • — Erichson, 1840: 526 (Quedius; synonym of fulgidus).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — Thomson, 1860: 175 (Microsaurus; synonym of fulgidus).

  • — Fauvel, 1869: 486 (Quedius; synonym of mesomelinus).

  • — Fauvel, 1874: 280 [= 1874a: 506] (Quedius; synonym of mesomelinus).

  • — Horn, 1878: 159 (Quedius; variety of fulgidus; characters).

  • — Ganglbauer, 1895: 400 (Quedius; synonym of mesomelinus).

  • — Porta, 1907: 116 (Quedius; synonym of mesomelinus).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; synonym of mesomelinus).

  • — Gridelli, 1924d: 60 (Quedius; synonym of mesomelinus).

  • — Scheerpeltz, 1933: 1448 (Quedius; synonym of mesomelinus).

  • — Smetana, 1958: 353 (Quedius; synonym of mesomelinus).

  • — Smetana, 1971: 74 (Quedius; synonym of mesomelinus).

  • — Bordoni, 1974: 33 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 150 (Quedius; synonym of mesomelinus).

nigricornis Holme, 1842: 127 (Raphirus; [preoccupied]; Type locality: Not cited).

  • — Hardy, 1851: 35 (Quedius; synonym of fuscipes).

  • — Fauvel, 1869: 486 (Quedius; synonym of mesomelinus).

  • — Rye, 1870a: 81 (Quedius; notes; Britain).

  • — Fauvel, 1874: 281 [= 1874a: 507] (Quedius; synonym of mesomelinus).

  • — Ganglbauer, 1895: 400 (Quedius; synonym of mesomelinus).

  • — Porta, 1907: 116 (Quedius; synonym of mesomelinus).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; synonym of mesomelinus).

  • — Gridelli, 1924d: 60 (Quedius; synonym of mesomelinus).

  • — Smetana, 1958: 353 (Quedius; synonym of mesomelinus).

  • — Smetana, 1971: 74 (Quedius; synonym of mesomelinus).

  • — Bordoni, 1974: 33 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 150 (Quedius; synonym of mesomelinus).

fuchsii Scriba, 1866: 378 (Quedius; Type locality: Roma).

  • — Fauvel, 1874: 281 [= 1874a: 507] (Quedius; synonym of mesomelinus).

  • — Ganglbauer, 1895: 400 (Quedius; synonym of mesomelinus).

  • — Porta, 1907: 116 (Quedius; synonym of mesomelinus).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; synonym of mesomelinus).

  • — Scheerpeltz, 1933: 1448 (Quedius; synonym of mesomelinus).

  • — Smetana, 1958: 353 (Quedius; synonym of mesomelinus).

  • — Smetana, 1971: 74 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 150 (Quedius; synonym of mesomelinus).

temporalis Thomson, 1867: 161 (Microsaurus; Type locality: Scandinavia).

  • — Fauvel, 1874: 281 [= 1874a: 507] (Quedius; synonym of mesomelinus).

  • — Mulsant and Rey, 1876: 641 [= 1877: 497] (Quedius; synonym of mesomelinus).

  • — J. Sahlberg, 1876: 25 (Microsaurus; synonym of mesomelinus).

  • — Ganglbauer, 1895: 400 (Quedius; synonym of mesomelinus).

  • — Porta, 1907: 116 (Quedius; synonym of mesomelinus).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; synonym of mesomelinus).

  • — Gridelli, 1924d: 60 (Quedius; synonym of mesomelinus).

  • — Smetana, 1958: 353 (Quedius; synonym of mesomelinus).

  • — Smetana, 1971: 74 (Quedius; synonym of mesomelinus).

  • — Bordoni, 1974: 33 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 150 (Quedius; synonym of mesomelinus).

arvernicus Mulsant and Rey, 1876: 643 [= 1877: 499] (Quedius; Type locality: la grotte de Crémieux [Isère], près Lyon).

  • — Gridelli, 1924d: 60 (Quedius; synonym of mesomelinus).

  • — Scheerpeltz, 1933: 1448 (Quedius; synonym of mesomelinus).

  • — Smetana, 1958: 353 (Quedius; synonym of mesomelinus).

  • — Smetana, 1971: 74 (Quedius; synonym of mesomelinus).

  • — Bordoni, 1974: 33 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 150 (Quedius; synonym of mesomelinus).

rufithorax Mulsant and Rey, 1876: 643 [= 1877: 499] (Quedius; Type locality: Not cited).

  • — Gridelli, 1924d: 60 (Quedius; synonym of mesomelinus).

  • — Scheerpeltz, 1933: 1449 (Quedius; synonym of mesomelinus).

  • — Smetana, 1958: 353 (Quedius; synonym of mesomelinus).

  • — Smetana, 1971: 74 (Quedius; synonym of mesomelinus).

  • — Bordoni, 1974: 33 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 150 (Quedius; synonym of mesomelinus).

nigripennis J. Sahlberg, 1876: 26 (Microsaurus; Type locality: Pitkäranta i Ladoga Karelen; Helsingfors).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; catalog).

  • — Gridelli, 1924d: 60 (Quedius; synonym of mesomelinus).

  • — Scheerpeltz, 1933: 1448 (Quedius; synonym of mesomelinus).

  • — Roubal, 1941b: 125 (Quedius; subgenus Microsaurus; aberration of mesomelinus; characters).

  • — Smetana, 1958: 353 (Quedius; synonym of mesomelinus).

  • — Smetana, 1971: 74 (Quedius; synonym of mesomelinus).

  • — Bordoni, 1974: 33 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 150 (Quedius; synonym of mesomelinus).

microcephalus Helliesen, 1892: 73 (Quedius; variety of mesomelinus; Type locality: Jaederen).

  • — Gridelli, 1924d: 60 (Quedius; variety of mesomelinus).

  • — Scheerpeltz, 1933: 1449 (Quedius; variety of mesomelinus).

silensis Fiori, 1894: 86 (Quedius; Type locality: Sila, in Calabria; Catanzaro).

  • — Porta, 1907: 116 (Quedius; synonym of mesomelinus).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; variety of mesomelinus).

  • — Gridelli, 1924d: 73 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • — Korge, 1961: 48 (Quedius; subspecies of mesomelinus; characters; Italy).

  • — Bordoni, 1974: 34, 37 (Quedius; subspecies of mesomelinus; characters; Italy).

  • — Coiffait, 1978: 151 (Quedius; subgenus Microsaurus; subspecies of mesomelinus; characters).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; subspecies of mesomelinus; subspecies of doubtful value; Italy).

jaenneri Hubenthal, 1902: 267 (Quedius; variety of mesomelinus; Type locality: Berlin; Gotha; Sächsische Schweiz).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; synonym of mesomelinus).

  • — Gridelli, 1924d: 60 (Quedius; aberration of mesomelinus).

  • — Scheerpeltz, 1933: 1449 (Quedius; aberration of mesomelinus).

  • — Roubal, 1941b: 125 (Quedius; subgenus Microsaurus; aberration of mesomelinus; characters).

  • — Smetana, 1958: 353 (Quedius; aberration of mesomelinus).

  • — Smetana, 1960e: 159 (Quedius; aberration of mesomelinus; characters).

  • — Korge, 1961: 46 (Quedius; aberration of mesomelinus; characters).

  • — Smetana, 1971: 74 (Quedius; synonym of mesomelinus).

  • — Bordoni, 1974: 34 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 150 (Quedius; subgenus Microsaurus; variety of mesomelinus; characters).

  • — Tóth, 1984: 113 (Quedius; aberration of mesomelinus).

kraussi Penecke, 1904: 135 (Quedius; subgenus Microsaurus; Type locality: Nevesinje; Herzegowina: Velez Gebirge).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Gridelli, 1924d: 60 (Quedius; subspecies of mesomelinus).

  • — Scheerpeltz, 1933: 1449 (Quedius; subspecies of mesomelinus).

  • — Roubal, 1941b: 125 (Quedius; subgenus Microsaurus; subspecies of mesomelinus; characters).

  • — Korge, 1961: 47 (Quedius; subspecies of mesomelinus; characters; Herzegowina; Dalmatia).

  • — Bordoni, 1974: 33 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 151 (Quedius; subgenus Microsaurus; subspecies of mesomelinus; characters).

  • — Tóth, 1984: 112 (Quedius; subspecies of mesomelinus; characters).

klinckowstroemi Poppius, 1913: 10 (Quedius; subgenus Microsaurus; Type locality: Faeröer: Trangisvaag).

  • — Gridelli, 1924d: 59 (Quedius; subgenus Microsaurus; characters; Faroe Islands).

  • — Scheerpeltz, 1933: 1445 (Quedius; catalog).

  • — West, 1937: 35 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 161 (Quedius; subgenus Microsaurus; maybe subspecies of brevicornis; characters).

delahoni H. Wagner, 1916: 226 (Quedius; aberration of mesomelinus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]).

  • — Gridelli, 1924d: 60 (Quedius; aberration of mesomelinus).

  • — Scheerpeltz, 1933: 1449 (Quedius; aberration of mesomelinus).

  • — Roubal, 1941b: 125 (Quedius; subgenus Microsaurus; aberration of mesomelinus; characters).

  • — Smetana, 1958: 353 (Quedius; aberration of mesomelinus).

  • — Smetana, 1960e: 159 (Quedius; aberration of mesomelinus; characters).

  • — Korge, 1961: 46 (Quedius; aberration of mesomelinus; characters).

  • — Smetana, 1971: 74 (Quedius; synonym of mesomelinus).

  • — Bordoni, 1974: 33 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 150 (Quedius; subgenus Microsaurus; variety of mesomelinus; characters).

  • — Tóth, 1984: 113 (Quedius; variety of mesomelinus).

josefi Fleischer, 1925: 20 (Quedius; aberration of mesomelinus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from [Slovakia]: Demänovských jeskyň).

  • — Scheerpeltz, 1933: 1449 (Quedius; aberration of mesomelinus).

  • — Roubal, 1941b: 125 (Quedius; subgenus Microsaurus; aberration of mesomelinus; characters).

  • — Smetana, 1958: 353 (Quedius; aberration of mesomelinus).

  • — Smetana, 1960e: 159 (Quedius; aberration of mesomelinus; characters).

  • — Korge, 1961: 46 (Quedius; aberration of mesomelinus; characters).

  • — Smetana, 1971: 74 (Quedius; synonym of mesomelinus).

  • — Bordoni, 1974: 34 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 150 (Quedius; subgenus Microsaurus; variety of mesomelinus; characters).

  • — Tóth, 1984: 113 (Quedius; aberration of mesomelinus; characters; Hungary).

lloydi Donisthorpe, 1945: 165 (Quedius; Type locality: Britain: Herefordshire: Furnace Wood, Treago).

  • — Pope, 1977: 31 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 185 (Quedius; subgenus Microsaurus; [Note: I asssume Coiffait overlooked the earlier synonymy of this name]; characters; England).

matinai Roubal, 1950: 21 (Quedius; aberration of mesomelinus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Boh.: Malá Skála).

  • — Smetana, 1958: 353 (Quedius; aberration of mesomelinus).

  • — Smetana, 1960e: 159 (Quedius; aberration of mesomelinus; characters).

  • — Smetana, 1971: 74 (Quedius; synonym of mesomelinus).

  • — Bordoni, 1974: 34 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 151 (Quedius; subgenus Microsaurus; variety of mesomelinus; characters).

  • — Tóth, 1984: 113 (Quedius; variety of mesomelinus; characters; Hungary).

fiorii Korge, 1961: 50 (Quedius; subspecies of mesomelinus; Type locality: Emilia).

  • — Bordoni, 1974: 34, 38 (Quedius; subspecies of mesomelinus; characters; Italy).

  • — Coiffait, 1978: 151 (Quedius; subgenus Microsaurus; subspecies of mesomelinus; characters).

skoraszewskyi Korge, 1961: 52 (Quedius; subspecies of mesomelinus; Type locality: Eisenkappel, Obir, Kärnten).

  • — Lohse, 1964: 209 (Quedius; subgenus Microsaurus; subspecies of mesomelinus; characters; central Europe).

  • — Bordoni, 1974: 34, 37 (Quedius; subspecies of mesomelinus; characters).

  • — Coiffait, 1978: 151 (Quedius; subgenus Microsaurus; subspecies of mesomelinus; characters).

  • — Tóth, 1984: 112 (Quedius; subspecies of mesomelinus; characters; Hungary).

  • — Lucht, 1987: 109 (Quedius; subspecies of mesomelinus; checklist; Germany; Poland; Czechoslovakia; Austria).

  • — Lohse and Lucht, 1989: 172 (Quedius; note).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; subspecies of mesomelinus; Italy).

mateui Coiffait, 1967e: 393 (Quedius; subgenus Microsaurus; variety of mesomelinus; [Note: Nomen nudum (Article 15.2)]; cited from Aven de Las Palomas, Sierra de Las Nieves, province de Ronda, Andalousie).

  • — Coiffait, 1978: 151 (Quedius; subgenus Microsaurus; variety of mesomelinus; characters).

metallicus Fauvel, 1878g: 556 (Quedius; Type locality: Australie orientale: Queensland, Wide Bay).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; catalog).

  • — Scheerpeltz, 1935a: 656 (Quedius; Australia).

  • Distribution: Australia.

meurguesae Coiffait, 1977c: 138 (Quedius; subgenus Sauridus; Type locality: Iran septentrional, Alam Kouch, Elbourz, 2500–3500 m).

  • — Coiffait, 1978: 232 (Quedius; subgenus Sauridus; characters; Iran).

  • — Coiffait, 1981b: 237 (Quedius; Saudi Arabia).

  • Distribution: Iran, Saudi Arabia.

mexicanus Sharp, 1884: 334 (Quedius; Type locality: Mexico, Cordova, Mirador).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1449 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 174 (Quedius; subgenus Cyrtoquedius; characters in key; Mexico).

  • Distribution: Mexico.

micantiventris Scheerpeltz, 1965: 292 (Quedius; subgenus Indoquedius; Type locality: N.E. Burma, Kambaiti, 7000 ft.).

  • Distribution: Myanmar.

microcapillatus Korge, 1971: 16 (Quedius; subgenus Sauridus; Type locality: Danavar-dagh südöstlich Ushnuiyeh, ca. 1600 m, Azarbaijan. Also cited from Anatolia).

  • — Coiffait, 1978: 242 (Quedius; subgenus Sauridus; characters; Azerbaijan; Anatolia).

  • Distribution: Azerbaijan, Turkey.

microcephalus Helliesen, 1892, see: mesomelinus Marsham, 1802.

microphthalmus Bernhauer, 1900b: 53 (Quedius; Type locality: Bulghar Maaden in Kleinasien).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; catalog).

  • — Gridelli, 1924d: 70 (Quedius; subgenus Microsaurus; characters; Turkey).

  • — Scheerpeltz, 1933: 1449 (Quedius; catalog).

  • — Coiffait, 1978: 183 (Quedius; subgenus Microsaurus; characters; Anatolia).

  • Distribution: Turkey.

microphthalmus Reitter, 1909, see: edmundi Coiffait, 1969.

microps Gravenhorst, 1847: 213 (Quedius; Type locality: England).

  • — Märkel, 1845: 238 (Quedius; [Note: Misidentification: Scheerpeltz, 1933: 1449, cited infuscatus Märkel as a synonym of microps; Märkel did not describe the species as new, he attributed it to Erichson]; with Formica fuliginosa).

  • — Kraatz, 1857d: 501 (Quedius; England).

  • — Thomson, 1867: 166 (Microsaurus; characters; Scandinavia).

  • — Fauvel, 1869: 486 (Quedius; synonym of nitipennis).

  • — Rye, 1870a: 81 (Quedius; notes; Britain).

  • — Fauvel, 1874: 274 [= 1874a: 500] (Quedius; characters; France; Scandinavia; Britain; Germany; Moravia).

  • — Seidlitz, 1875: 266 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 618 [= 1877: 474] (Quedius; subgenus Ediquus; characters; France).

  • — Fowler, 1888: 229 (Quedius; subgenus Ediquus; characters; Britain).

  • — Ganglbauer, 1895: 396 (Quedius; subgenus Ediquus; characters; north and middle Europe).

  • — Porta, 1907: 105 (Quedius; subgenus Ediquus; characters; Italy).

  • — Reitter, 1909: 109 (Quedius; subgenus Ediquus; characters; Germany).

  • — Johansen, 1914: 352 (Quedius; subgenus Ediquus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; catalog).

  • — Munster, 1923: 189 (Quedius; subgenus Ediquus; Fennoscandia).

  • — Gridelli, 1924d: 31 (Quedius; subgenus Microsaurus; characters; Germany; Bohemia).

  • — Portevin, 1929: 338 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1449 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 3 (Quedius; aedeagal illustration).

  • — Roubal, 1941b: 121, 126 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — V. Hansen, 1952: 127 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Smetana, 1958: 341 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 152 (Quedius; subgenus Microsaurus; characters).

  • — Palm, 1963b: 128 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 207 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Horion, 1965: 255 (Quedius; subgenus Microsaurus; Britain; Norway; Sweden; Finland; Poland; Czecholsovakia; Bosnia; Italy; France; Denmark; Germany; Austria).

  • — Szujecki, 1968a: 734 (Quedius; Poland).

  • — Bordoni, 1974: 53 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 166 (Quedius; subgenus Microsaurus; characters; Europe; Britain).

  • — Tóth, 1984: 107 (Quedius; characters; Hungary).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Denmark; Sweden).

  • — Nowosad, 1990: 125 (Quedius; collected from mole nest; notes; Poland).

  • — Whitehead, 1990e: 12 (Quedius; collecting notes; Britain).

  • — Allen, 1990b: 3 (Quedius; collecting notes; Britain).

  • — Smetana, 1993b: 50 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Fowles, Alexander, and Key, 1999: 129 (Quedius; saproxylic species; status as endangered or threatened species; Britain).

  • Distribution: Europe.

quisquilius Gravenhorst, 1847: 214 (Quedius; [Note: Nomen nudum (Article 11.6), published as a synonym of microps]; cited from France).

  • — Gridelli, 1924d: 31 (Quedius; synonym of microps).

  • — Scheerpeltz, 1933: 1450 (Quedius; synonym of microps).

  • — Smetana, 1958: 341 (Quedius; synonym of microps).

  • — Bordoni, 1974: 53 (Quedius; synonym of microps).

  • — Coiffait, 1978: 166 (Quedius; synonym of microps).

chrysurus Kiesenwetter, 1848: 55 (Quedius; Type locality: Gegend von Leipzig; Harze).

  • — Redtenbacher, 1849: 825 (Philonthus; characters; Austria).

  • — Kraatz, 1857d: 520 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 205 (Quedius; characters; Germany).

  • — Thomson, 1867: 167 (Microsaurus; synonym of microps).

  • — Fauvel, 1874: 274 [= 1874a: 500] (Quedius; synonym of microps).

  • — Redtenbacher, 1874: 202 (Quedius; characters).

  • — Ganglbauer, 1895: 396 (Quedius; synonym of microps).

  • — Porta, 1907: 105 (Quedius; synonym of microps).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; synonym of microps).

  • — Gridelli, 1924d: 31 (Quedius; synonym of microps).

  • — Scheerpeltz, 1933: 1449 (Quedius; synonym of microps).

  • — Smetana, 1958: 341 (Quedius; synonym of microps).

  • — Bordoni, 1974: 53 (Quedius; synonym of microps).

  • — Coiffait, 1978: 166 (Quedius; synonym of microps).

milansaar Smetana, 1988: 217 (Quedius; subgenus Microsaurus; Type locality: India: H. P., Macleodganj).

  • Distribution: India.

mimeticus Cameron, 1932, see: beesoni Cameron, 1932.

minor Hochhuth, 1849: 151 (Quedius; variety of impressus; Type locality: Tiflis).

  • — Scheerpeltz, 1933: 1458 (Quedius; synonym of punctatellus).

  • — Korge, 1964: 117, 118 (Quedius; subgenus Quedionuchus; valid species; characters; Turkey).

  • — Korge, 1971a: 47 (Quedius; subgenus Quedionuchus; Turkey).

  • — Bordoni, 1973b: 51 (Quedius; synonym of punctatellus).

  • — Coiffait, 1978: 188 (Quedius; subgenus Distichalius; characters; Caucasus; Anatolia).

  • Distribution: Georgia, Turkey.

rufipennis Roubal, 1911: 9 (Quedius; aberration of punctatellus; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]; cited from Aibhy).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; aberration of punctatellus).

  • — Gridelli, 1924d: 76 (Quedius; synonym of distincticolor).

  • — Scheerpeltz, 1933: 1438 (Quedius; synonym of distincticolor).

  • — Coiffait, 1978: 188 (Quedius; synonym of minor variety distincticolor).

distincticolor Roubal, 1914: 91 (Quedius; replacement name for rufipennis Roubal; [Note: Nomen nudum since the name it replaced is unavailable]).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; aberration of punctatellus).

  • — Gridelli, 1924d: 76 (Quedius; subgenus Quedionuchus; characters; Caucasus).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Coiffait, 1978: 188 (Quedius; subgenus Distichalius; variety of minor; characters).

roubali Bernhauer, 1916: 432 (Quedius; replacement name for rufipennis Roubal; [Note: Nomen nudum since the name it replaced is unavailable]).

  • — Scheerpeltz, 1933: 1438 (Quedius; synonym of distincticolor).

  • — Coiffait, 1978: 188 (Quedius; synonym of minor variety distincticolor).

miquelonensis Smetana, 1965, see: molochinoides Smetana, 1965.

miwai Bernhauer, 1943a: 181 (Quedius; subgenus Ediquus; Type locality: Formosa: Nokosan; [Note: See lectotype designation by Smetana, 1995]).

  • — Smetana, 1995: 50 (Quedius; subgenus of Microsaurus; lectotype designation; characters; bionomics; Taiwan).

  • Distribution: Taiwan.

mixtus Eppelsheim, 1890: 222 (Quedius; subgenus Microsaurus; Type locality: Kaspi-Gebiet bei Lenkoran und Lirik).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; catalog).

  • — Gridelli, 1924d: 66 (Quedius; subgenus Microsaurus; characters; Lenkoran).

  • — Scheerpeltz, 1933: 1450 (Quedius; catalog).

  • — Korge, 1971: 11 (Quedius; subgenus Microsaurus; characters; notes; Iran).

  • — Coiffait, 1978: 179 (Quedius; subgenus Microsaurus; characters; Azerbaijan).

  • Distribution: Azerbaijan, Iran.

mnemon Smetana, 1995f: 246 (Quedius; subgenus Microsaurus; Type locality: China: Beijing Xiaolongmen).

  • Distribution: China.

modestus Kraatz, 1857, see: maurorufus Gravenhorst, 1806.

moeris Smetana, 1995g: 241 (Quedius; subgenus Microsaurus; Type locality: China: Quinhai, Lajishan-Pass s. Xining, 3700–3900 m).

  • — Smetana, 1997d: 455 (Quedius; subgenus Microsaurus; characters of female; China).

  • — Smetana, 1999d: 524 (Quedius; subgenus Microsaurus; characters; collecting notes; Qinghai; Sichuan).

  • Distribution: China.

molochinicolor Roubal, 1931, see: fuliginosus Gravenhorst, 1802.

molochinoides Smetana, 1965: 35, 39 (Quedius; subgenus Quedius; Type locality: Newfoundland: Ha-Ha Bay).

  • — Smetana, 1971: 141 (Quedius; subgenus Quedius; characters; Alberta; British Columbia; Yukon Territory; Alaska; California; Idaho; Montana; Nevada; New Hampshire; New York).

  • — Smetana, 1976c: 174 (Quedius; subgenus Quedius; Labrador; Ontario; Quebec).

  • — Smetana, 1978a: 826 (Quedius; subgenus Quedius; collecting notes; Quebec).

  • — Smetana, 1981b: 640 (Quedius; Ontario).

  • — Downie and Arnett, 1996: 391 (Quedius; subgenus Quedius; characters; USA).

  • Distribution: Canada, USA.

miquelonensis Smetana, 1965: 36, 39 (Quedius; subgenus Quedius; subspecies of molochinoides; Type locality: Miquelon).

  • — Smetana, 1971: 141, 142 (Quedius; subgenus Quedius; subspecies of molochinoides; characters; Miquelon).

molochinus Gravenhorst, 1806: 46 (Staphylinus; Type locality: Brunsvigae).

  • — Paykull, 1800: 373 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1916: 430, and others cited picipennis “Paykull” as a valid species and Smetana, 1971: 133 cited picipennis “Paykull” as a synonym of molochinus but Paykull attributed the species to Fabricius]; characters; Sweden).

  • — Gyllenhal, 1810: 302 (Staphylinus; characters; Sweden).

  • — Gyllenhal, 1827: 477 (Staphylinus; characters for variety).

  • — Curtis, 1829: 25 (Quedius; catalog; Britain).

  • — C. Sahlberg, 1830: 314 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 25 (Staphylinus; Finland; Sweden).

  • — Mannerheim, 1831: 439 (Staphylinus; Finland; Sweden).

  • — Brullé, 1832: 131 (Creophilus; characters; Greece).

  • — Runde, 1835: 5 (Staphylinus; characters; Germany).

  • — Lacordaire, 1835: 377 (Emus; characters; France).

  • — Curtis, 1837: pl. 638 (Quedius; characters; Britain).

  • — Nordmann, 1837: 76 (Philonthus; Sweden; France; England; Tauria; Germany).

  • — Erichson, 1839a: 489 (Quedius; characters; Germany).

  • — Heer, 1839: 276 (Philonthus; characters; Switzerland).

  • — Erichson, 1840: 535 (Quedius; characters; Europe; America septentrionalis).

  • — Laporte, 1840: 177 (Staphylinus; characters; France).

  • — Mannerheim, 1843b: 232 (Quedius; characters; Sitka Island).

  • — Kiesenwetter, 1848: 52 (Quedius; characters; Europe).

  • — Gravenhorst, 1847: 227 (Quedius; characters; notes).

  • — Redtenbacher, 1849: 709 (Philonthus; characters; Austria).

  • — Kiesenwetter, 1851a: 419 (Quedius; France).

  • — Hardy, 1851: 32 (Quedius; notes; England).

  • — Rosenhauer, 1856: 71 (Quedius; P. S. Maria).

  • — Fairmaire and Laboulbène, 1856: 537 (Quedius; characters; France).

  • — Kraatz, 1857d: 500 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 201 (Quedius; characters; Austria).

  • — Kraatz, 1858a: 58 (Quedius; Greece).

  • — Thomson, 1859: 25 (type species of Quedius).

  • — Thomson, 1860: 173 (Quedius; characters; Scandinavia).

  • — Fairmaire and Coquerel, 1860: 154 (Quedius; Algeria).

  • — Hochhuth, 1862: 46 (Quedius; Caspian Sea).

  • — Fauvel, 1874: 290 [= 1874a: 516] (Quedius; characters; France; Europe; Barbary; Cyprus; Turkey; Caucasus; Caspian Sea; North America).

  • — Redtenbacher, 1874: 198 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 267 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 692 [= 1877: 548] (Quedius; subgenus Microsaurus; characters; France).

  • — J. Sahlberg, 1876: 24 (Quedius; Finland).

  • — Provancher, 1877: 244 (Quedius; cited as melochinus; characters; Quebec).

  • — Fauvel, 1878: 128 [= 1878a: 48] (Quedius; Barbary).

  • — Horn, 1878: 163 (Quedius; characters; New Hampshire to Vancouver; northern Georgia).

  • — Fauvel, 1886: 58 [= 1886a: 50] (Quedius; Barbary).

  • — Fowler, 1888: 237 (Quedius; subgenus Microsaurus; characters; Britain).

  • — Ganglbauer, 1895: 403 (Quedius; characters; Palaearctic region; North America).

  • — Fauvel, 1902b: 115 (Quedius; cited as picipennis “Paykull” with molochinus a synonym; Morocco; Algeria; Tunisia).

  • — Poppius, 1905: 9 (Quedius; Shetland Islands).

  • — Porta, 1907: 129 (Quedius; subgenus Quedius; characters; Italy).

  • — Fall and Cockerell, 1907: 166 (Quedius; New Mexico).

  • — Sainte-Claire Deville, 1906: 102 (Quedius; Corsica).

  • — Reitter, 1909: 112 (Quedius; characters; Germany).

  • — Blatchley, 1910: 372 (Quedius; characters; Indiana).

  • — Johansen, 1914: 364 (Quedius; subgenus Quedius; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; variety of picipennis “Paykull”).

  • — Notman, 1919a: 95 (Quedius; Ontario).

  • — Gridelli, 1924d: 81 (Quedius; subgenus Quedius; [Note: Cited as picipennis “Paykull”; molochinus cited as a variety]; characters).

  • — Fall, 1926: 150 (Quedius; Alaska).

  • — Portevin, 1929: 341 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1456 (Quedius; cited as picipennis “Paykull” with molochinus as a variety; catalog).

  • — Méquignon, 1937: 3 (Quedius; synonyms).

  • — Voris, 1939: 152 (Quedius; egg, larval, and pupal characters; Indiana).

  • — Wüsthoff, 1938a: fig. 23 (Quedius; cited as picipennis “Paykull”; aedeagal illustration).

  • — V. Hansen, 1952: 142 (Quedius; subgenus Quedius; cited as picipennis “Paykull”; characters; Denmark).

  • — Smetana, 1958: 364 (Quedius; subgenus Quedius; characters; ecology; Czechoslovakia).

  • — Korge, 1960: 53 (Quedius; cited as picipennis “Paykull” with molochinus as a variety; characters).

  • — Coiffait, 1961a: 52 (Quedius; characters; France; Alps).

  • — Smetana, 1962a: 134 (Quedius; larval characters).

  • — Smetana, 1962b: 136 (Quedius; subgenus Quedius; characters).

  • — V. Hansen, 1962a: 320 (Quedius; characters; Denmark).

  • — Scheerpeltz, 1963a: 420 (Quedius; subgenus Quedius; variety of picipennis; Greece).

  • — Palm, 1963b: 141 (Quedius; subgenus Quedius; cited as picipennis “Paykull”; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 212 (Quedius; subgenus Quedius; characters; central Europe).

  • — Smetana, 1965: 38 (Quedius; subgenus Quedius; characters).

  • — M. Dvořák, 1965: 89 (Quedius; Slovakia).

  • — Horion, 1965: 276 (Quedius; subgenus Quedius; all Europe).

  • — Pototskaia, 1967: 82 (Quedius; larval characters).

  • — Kasule, 1968a: 66 (Quedius; life history).

  • — Kasule, 1970: 61 (Quedius; larval characters).

  • — Smetana, 1971: 133 (Quedius; subgenus Quedius; characters; most North American records are misidentifications; Newfoundland).

  • — Smetana, 1973: 1427 (Quedius; Quebec; introduction).

  • — Osella and Zanetti, 1975: 120 (Quedius; collected from nest of Talpa europaea; notes; Italy).

  • — Smetana, 1976c: 172 (Quedius; subgenus Quedius; Quebec).

  • — Bordoni, 1976b: 95 (Quedius; characters; Italy).

  • — Shilov, 1975a: 376 [= 1975b: 99] (Quedius; Komi region of Russia).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 194 (Quedius; subgenus Quedius; characters; Europe).

  • — Smetana, 1978b: 85 (Quedius; subgenus Quedius; notes; Siberia).

  • — Smetana, 1981b: 638 (Quedius; notes; Newfoundland; Quebec).

  • — Tóth, 1984: 120 (Quedius; characters; Hungary).

  • — Segers, 1987a: 182 (Quedius; elytral aberration; Belgium).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Smetana, 1990d: 98 (Quedius; Newfoundland).

  • — Welch, 1993: 229 (Quedius; ovariole number and ovary structure).

  • — Smetana, 1993b: 50 (Quedius; subgenus Quedius; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Staniec, 1996: 117 (Quedius; pupal characters).

  • — Downie and Arnett, 1996: 391 (Quedius; subgenus Quedius; characters; Canada).

  • — Owen, 2000: 252 (Quedius; pitfall trap collections from underground at roots of trees; Britain).

  • — Assing, 2001: 75 (Quedius; collected from xerothermous site; Germany).

  • Distribution: Tunisia, Algeria, Morocco, Europe, Russia, Caucasus, Canada.

denudatus Stephens, 1832: 216 (Quedius; Type locality: London; Suffolk).

  • — Curtis, 1829: 25 (Quedius; nomen nudum; catalog; Britain).

  • — Curtis, 1837: pl. 638 (Quedius; characters; Britain).

  • — Stephens, 1839: 390 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 23 (Quedius; synonym of molochinus).

  • — Gemminger and Harold, 1868: 570 (Quedius; synonym of molochinus).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; synonym of picipennis “Paykull”).

  • — Gridelli, 1924d: 81 (Quedius; synonym of picipennis “Paykull”).

  • — Méquignon, 1937: 3 (Quedius; variety of molochinus).

  • — Smetana, 1958: 364 (Quedius; synonym of molochinus).

  • — Korge, 1960a: 69 (Quedius; variety of molochinus).

  • — Coiffait, 1961a: 52 (Quedius; “mut” [probably mutation] of molochinus; characters).

  • — Smetana, 1962b: 136 (Quedius; aberration of molochinus).

  • — Lohse, 1964: 212 (Quedius; subgenus Quedius; aberration of molochinus; characters).

  • — Smetana, 1964d: 80 (Quedius; subgenus Quedius; aberration of molochinus; Czechoslovakia).

  • — Smetana, 1965: 35 (Quedius; subgenus Quedius; aberration of molochinus; Newfoundland).

  • — Szujecki, 1968a: 736 (Quedius; aberration of molochinus; Poland).

  • — Smetana, 1971: 133 (Quedius; synonym of molochinus).

  • — Pope, 1977: 31 (Quedius; synonym of molochinus).

  • — Coiffait, 1978: 194 (Quedius; subgenus Quedius; variety of molochinus; characters).

  • — Tóth, 1984: 120 (Quedius; variety of molochinus; characters; Hungary).

lathburii Stephens, 1832: 218 (Quedius; Type locality: London; Suffolk: Orwell).

  • — Curtis, 1829: 25 (Quedius; nomen nudum; catalog; Britain).

  • — Curtis, 1837: pl. 638 (Quedius; characters; Britain).

  • — Stephens, 1839: 390 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 23 (Quedius; synonym of molochinus).

  • — Gemminger and Harold, 1868: 571 (Quedius; synonym of molochinus).

  • — Porta, 1907: 130 (Quedius; synonym of molochinus).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; synonym of picipennis “Paykull”).

  • — Gridelli, 1924d: 81 (Quedius; synonym of variety molochinus).

  • — Scheerpeltz, 1933: 1457 (Quedius; synonym of picipennis “Paykull”).

  • — Méquignon, 1937: 3 (Quedius; synonym of molochinus).

  • — Korge, 1960a: 69 (Quedius; synonym of molochinus).

  • — Coiffait, 1961a: 52 (Quedius; synonym of molochinus).

  • — Smetana, 1971: 133 (Quedius; synonym of molochinus).

  • — Coiffait, 1978: 194 (Quedius; synonym of molochinus).

picipennis Stephens, 1833: 243 (Raphirus; Type locality: London).

  • — Stephens, 1839: 393 (Quedius; characters; England).

  • — Hardy, 1851: 34 (Quedius; notes; England).

  • — G. Waterhouse, 1858: 23 (Quedius; synonym of molochinus).

laevicollis Runde, 1835: 6 (Staphylinus; [preoccupied by levicollis Brullé according to Article 58.1]; Type locality: Halae).

  • — Erichson, 1839a: 490 (Quedius; synonym of molochinus).

  • — Erichson, 1840: 536 (Quedius; synonym of molochinus).

  • — Mannerheim, 1843b: 233 (Quedius; synonym of molochinus).

  • — Kraatz, 1857d: 501 (Quedius; synonym of molochinus).

  • — Ganglbauer, 1895: 403 (Quedius; synonym of molochinus).

  • — Porta, 1907: 129 (Quedius; synonym of molochinus).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; synonym of picipennis “Paykull”).

  • — Smetana, 1958: 364 (Quedius; synonym of molochinus).

  • — Korge, 1960a: 69 (Quedius; synonym of molochinus).

  • — Smetana, 1971: 133 (Quedius; synonym of molochinus).

lapponicus Zetterstedt, 1838: 61 (Staphylinus; Type locality: Lapponia Umensis: Lycksele; Wilhelmina; Gaskelougt).

  • — Erichson, 1840: 536 (Quedius; synonym of molochinus).

  • — Mannerheim, 1843b: 233 (Quedius; synonym of molochinus).

  • — Kraatz, 1857d: 501 (Quedius; synonym of molochinus).

  • — Thomson, 1860: 173 (Quedius; synonym of molochinus).

  • — Ganglbauer, 1895: 403 (Quedius; synonym of molochinus).

  • — Porta, 1907: 129 (Quedius; synonym of molochinus).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; synonym of picipennis “Paykull”).

  • — Gridelli, 1924d: 81 (Quedius; synonym of variety molochinus).

  • — Scheerpeltz, 1933: 1457 (Quedius; synonym of picipennis “Paykull”).

  • — Méquignon, 1937: 3 (Quedius; synonym of molochinus).

  • — Smetana, 1958: 364 (Quedius; synonym of molochinus).

  • — Korge, 1960a: 69 (Quedius; synonym of molochinus).

  • — Coiffait, 1961a: 52 (Quedius; synonym of molochinus).

  • — Smetana, 1971: 133 (Quedius; synonym of molochinus).

  • — Coiffait, 1978: 194 (Quedius; synonym of molochinus).

maculicornis Mulsant and Rey, 1876: 694 [= 1877: 550] (Quedius; subgenus Microsaurus; Type locality: les montagnes du Lyonnais, à la Grande Chartreuse, dans les Pyrénées).

  • — Ganglbauer, 1895: 403 (Quedius; synonym of molochinus).

  • — Porta, 1907: 129 (Quedius; synonym of molochinus).

  • — Gridelli, 1924d: 81 (Quedius; synonym of picipennis “Paykull”).

  • — Méquignon, 1937: 3 (Quedius; synonym of molochinus variety denudatus).

  • — Smetana, 1958: 364 (Quedius; synonym of molochinus).

  • — Korge, 1960a: 69 (Quedius; synonym of molochinus).

  • — Coiffait, 1961a: 52 (Quedius; synonym of molochinus mut. denudatus).

  • — Smetana, 1962b: 136 (Quedius; synonym of molochinus).

  • — Coiffait, 1963b: 423 (Quedius; subgenus Quedius; characters).

  • — Coiffait, 1978: 194 (Quedius; synonym of molochinus variety denudatus).

niger J. Sahlberg, 1876: 24, 229 (Quedius; variety of molochinus; [Note: Cited as “Var c” on page 24 and given a name on page 229]; Type locality: Muonioniska).

  • — Poppius, 1913: 10 (Quedius; variety of molochinus; Faroes).

  • — Scheerpeltz, 1933: 1456 (Quedius; synonym of picipennis “Paykull”).

  • — Smetana, 1958: 364 (Quedius; synonym of molochinus).

  • — Korge, 1960a: 69 (Quedius; synonym of molochinus).

  • — Smetana, 1971: 133 (Quedius; synonym of molochinus).

  • — Coiffait, 1978: 194 (Quedius; synonym of molochinus variety denudatus).

nigrinus Porta, 1907: 94, 130 (Quedius; aberration of molochinus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; synonym of picipennis “Paykull”).

  • — Gridelli, 1924d: 81 (Quedius; synonym of picipennis “Paykull”).

  • — Smetana, 1958: 364 (Quedius; synonym of molochinus).

  • — Korge, 1960a: 69 (Quedius; synonym of molochinus).

  • — Coiffait, 1961a: 52 (Quedius; synonym of molochinus mut. denudatus).

  • — Smetana, 1962b: 136 (Quedius; synonym of molochinus).

  • — Smetana, 1971: 133 (Quedius; synonym of molochinus).

  • — Coiffait, 1978: 194 (Quedius; synonym of molochinus variety denudatus).

nigripennis Hänel, 1913: 98 (Quedius; aberration of molochinus; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]; cited from Glatzer Gebirge; Altvater; Reitzenhain; Dresden; Radeberg; Jeschken).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; synonym of picipennis “Paykull”).

  • — Gridelli, 1924d: 81 (Quedius; synonym of picipennis “Paykull”).

  • — Méquignon, 1937: 3 (Quedius; synonym of molochinus variety denudatus).

  • — Smetana, 1958: 364 (Quedius; synonym of molochinus).

  • — Korge, 1960a: 69 (Quedius; synonym of molochinus).

  • — Coiffait, 1961a: 52 (Quedius; synonym of molochinus mut. denudatus).

  • — Smetana, 1962b: 136 (Quedius; synonym of molochinus).

  • — Smetana, 1971: 133 (Quedius; synonym of molochinus).

  • — Coiffait, 1978: 194 (Quedius; synonym of molochinus variety denudatus).

nigripennis Delahon, 1914: 621 (Quedius; aberration of molochinus; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]; cited from Mark Brandenburg).

  • — Scheerpeltz, 1933: 1456 (Quedius; synonym of picipennis “Paykull”).

  • — Korge, 1960a: 69 (Quedius; synonym of molochinus).

leonhardi Bernhauer, 1914b: 67 (Quedius; Type locality: Gargano).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; catalog).

  • — Gridelli, 1924d: 81 (Quedius; variety of molochinus; characters).

  • — Scheerpeltz, 1933: 1456 (Quedius; variety of picipennis “Paykull”).

  • — Korge, 1960a: 69 (Quedius; variety of molochinus).

  • — Coiffait, 1961a: 53 (Quedius; aberration of molochinus; characters).

  • — Focarile, 1964: 60 (Quedius; subspecies of pseudopicipennis; Italy).

  • — Coiffait, 1978: 194 (Quedius; subgenus Quedius; variety of molochinus; characters).

  • — Tóth, 1984: 120 (Quedius; variety of molochinus; characters; Hungary).

smetanai Roubal, 1949: 45 (Quedius; aberration of unicolor; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Krkoneše: Bílé Labe).

  • — Smetana, 1958: 364 (Quedius; synonym of molochinus).

  • — Coiffait, 1978: 194 (Quedius; synonym of molochinus).

nigribasis Korge, 1960: 53 (Quedius; aberration of picipennis “Paykull”; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Berlin).

  • — Korge, 1960a: 69 (Quedius; aberration of molochinus).

  • — Coiffait, 1961a: 52 (Quedius; aberration of molochinus; characters).

  • — Smetana, 1962b: 136 (Quedius; aberration of molochinus).

  • — Smetana, 1971: 133 (Quedius; synonym of molochinus).

  • — Coiffait, 1978: 194 (Quedius; subgenus Quedius; variety of molochinus; characters).

  • — Tóth, 1984: 120 (Quedius; variety of molochinus; characters; Hungary).

monspeliensis Fairmaire, 1857, see: scintillans Gravenhorst, 1806.

montanicus Casey, 1915: 414 (Microsaurus; Type locality: Montana: Mullan).

  • — Scheerpeltz, 1933: 1450 (Quedius; catalog).

  • — Smetana, 1971: 120 (Quedius; subgenus Microsaurus; characters; Colorado; Idaho; Oregon; Washington; Wyoming).

  • — Smetana, 1971b: 3 (Quedius; subgenus Microsaurus; Washington).

  • — Smetana, 1978a: 825 (Quedius; subgenus Microsaurus; collecting notes; Oregon).

  • Distribution: USA.

downiei Hatch, 1957: 226 (Quedius; subgenus Microsaurus; Type locality: Washington: Pullman. Also cited from Idaho).

  • — Smetana, 1971: 120 (Quedius; synonym of montanicus).

montanus Heer, 1839, see: dubius Heer, 1839.

monticola Erichson, 1840, see: paradisianus Heer, 1839.

montivagus Smetana, 1971: 160 (Quedius; subgenus Distichalius; Type locality: Washington: Mt. Rainier).

  • — Smetana, 1976c: 174 (Quedius; subgenus Distichalius; Washington).

  • — Smetana, 1978a: 827 (Quedius; subgenus Distichalius; collecting notes; British Columbia).

  • Distribution: USA, Canada.

mordax Smetana, 1971: 109 (Quedius; subgenus Microsaurus; Type locality: Alberta: Banff Nat. Pk., Cirrus Mt., 5500 ft. Also cited from British Columbia, Yukon Territory).

  • — Smetana, 1973: 1426 (Quedius; Alberta; British Columbia).

  • Distribution: Canada.

moritai Watanabe, 1990c: 30 (Quedius; subgenus Microsaurus; Type locality: Japan: Honshu: Toyama Pref.: Yakushizawa - Kumonotaira, northern Japanese Alps, alt. 2300 m).

  • Distribution: Japan.

mortuus† Wickham, 1912: 11 (Quedius; Miocene fossil; Type locality: Florissant, Colorado).

  • Distribution: USA.

muelleri Gridelli, 1924d: 89, 96, 104 (Quedius; proposed for misidentification of robustus auct.; Type locality: Gressoney).

  • — Scheerpeltz, 1933: 1450 (Quedius; catalog).

  • — Smetana, 1962b: 141 (Quedius; subgenus Raphirus; characters; Alps; Austria).

  • — Coiffait, 1963a: 375 (Quedius; subgenus Sauridus; characters).

  • — Lohse, 1964: 215 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 280 (Quedius; subgenus Raphirus; Alps; Austria; Switzerland; Italy; France).

  • — Coiffait, 1978: 208 (Quedius; subgenus Sauridus; characters; Alps).

  • — Lucht, 1987: 110 (Quedius; checklist; Austria).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Assing, Frisch, Kahlen, et al., 1998: 139 (Quedius; note).

  • Distribution: France, Switzerland, Austria.

mukuensis Bernhauer, 1933: 40 (Quedius; subgenus Microsaurus; Type locality: Szetschwan: Mukue Tatsienlu).

  • — Smetana, 1995f: 231 (Quedius; subgenus Microsaurus; characters; China).

  • Distribution: China.

multipunctatus Sharp, 1889: 33 (Quedius; Type locality: Hakone).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1450 (Quedius; catalog).

  • — Smetana, 1998b: 110 (Quedius; subgenus Raphirus; characters).

  • Distribution: Japan.

muscicola Cameron, 1932: 295 (Quedius; subgenus Raphirus; Type locality: Ghum dist.; [Note: See lectotype designation by Smetana, 1988]. Originally cited from Darjeeling Dist.: Ghum, Tiger Hill, alt. 8000–9000 feet).

  • — Scheerpeltz, 1933: 1450 (Quedius; catalog).

  • — Scheerpeltz, 1965: 112 (Quedius; subgenus Raphirus; Burma).

  • — Smetana, 1988: 253 (Quedius; subgenus Raphirus; lectotype designation: Ghum dist.; characters; Burma; Himachal Pradesh; Uttar Pradesh; West Bengal; Nepal).

  • — Smetana, 1992b: 5 (Quedius; subgenus Raphirus; Nepal).

  • — Smetana, 1997b: 131 (Quedius; notes; China).

  • Distribution: India, Nepal, Myanmar, China.

dohertyi Cameron, 1932: 297 (Quedius; subgenus Raphirus; Type locality: Burma: Ruby Mines).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Smetana, 1988: 253 (Quedius; synonym of muscicola).

  • — Smetana, 1997b: 131 (Quedius; synonym of muscicola).

heterogaster Cameron, 1944: 14 (Quedius; subgenus Raphirus; Type locality: United Provinces: Chakrata district: Kanasar; [Note: See lectotype designation by Smetana, 1988]).

  • — Smetana, 1988: 253 (Quedius; lectotype designation; synonym of muscicola).

  • — Smetana, 1997b: 131 (Quedius; synonym of muscicola).

muscorum Brisout, 1863: 32 (Quedius; Type locality: Pyrénées-Orientales, Mont-Louis).

  • — Fauvel, 1874: 298 [= 1874a: 524] (Quedius; synonym of suturalis).

  • — Ganglbauer, 1895: 407 (Quedius; synonym of humeralis).

  • — Porta, 1907: 140 (Quedius; synonym of humeralis).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; synonym of humeralis).

  • — Scheerpeltz, 1933: 1450 (Quedius; catalog).

  • — Coiffait, 1963a: 394 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 258 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 258 (Quedius; subgenus Sauridus; characters; Pyrenees).

  • Distribution: Spain, France.

circumductus Mulsant and Rey, 1876: 718 [= 1877: 574] (Quedius; Type locality: Espagne).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; variety of cincticollis).

  • — Scheerpeltz, 1933: 1450 (Quedius; synonym of muscorum).

  • — Jarrige, 1952a: 134 (Quedius; possible synonym of pineti).

  • — Coiffait, 1978: 258 (Quedius; synonym of muscorum).

mussooriensis Cameron, 1932, see: rugosus Cameron, 1921.

mutator Smetana, 1971: 206 (Quedius; subgenus Raphirus; Type locality: California: 8 mi. N. Post Pile Camp, Tehema Co.).

  • — Smetana, 1990d: 101 (Quedius; California).

  • Distribution: USA.

mutatus Kraatz, 1868, see: nemoralis Baudi, 1848.

mutilatus Eppelsheim, 1888: 58 (Quedius; Type locality: Fluss Tangi [sic] Turkestan; [Note: See lectotype designation by Smetana, 1998a]. Also cited from Hochgebirgssee Issyk-Kul in Ost-Turkestan).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; catalog).

  • — Gridelli, 1924d: 23 (Quedius; subgenus Microsaurus; characters; Turkestan).

  • — Scheerpeltz, 1933: 1450 (Quedius; catalog).

  • — Coiffait, 1978: 161 (Quedius; subgenus Microsaurus; characters; Turkestan).

  • — Smetana, 1998a: 118 (Quedius; subgenus Microsaurus; characters; lectotype designation: Fluss Tangi. Turkestan; Kyrgyzstan).

  • Distribution: Kyrgyzstan.

myagrus Rottenberg, 1870, see: semiaeneus Stephens, 1833.

myau Smetana, 1999e: 535 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Gongga Shan, Hailuogou, in front of Glacier 1, 2850 m, 29°35′N, 102°00′E).

  • Distribution: China.

naati Smetana, 1988: 264 (Quedius; subgenus Raphirus; Type locality: Nepal: Prov. Bagmati: Jangtang Ridge, NE Barahbise, 3250 m).

  • Distribution: Nepal.

najik Smetana, 1992, see: franzi Smetana, 1975.

nanulus Casey, 1915: 402 (Quediellus; Type locality: Siskiyou Co., California; [Note: See lectotype designation by Smetana, 1971]. Also cited from California: Humboldt Co.; Oregon: Lane Co.).

  • — Scheerpeltz, 1933: 1450 (Quedius; catalog).

  • — Hatch, 1957: 227 (Quedius; subgenus Sauridus; characters; British Columbia; Oregon).

  • — Smetana, 1971: 216 (Quedius; subgenus Raphirus; lectotype designation: Siskiyou Co., Cal.; characters; British Columbia; California; Idaho; Montana; Oregon; Washington).

  • — Smetana, 1971b: 4 (Quedius; subgenus Raphirus; British Columbia; Idaho).

  • — Smetana, 1973: 1430 (Quedius; notes; Alberta).

  • — Smetana, 1978a: 830 (Quedius; subgenus Raphirus; collecting notes; Alberta; Oregon).

  • Distribution: USA, Canada.

humilis Casey, 1915: 403 (Quediellus; Type locality: California: Soda Spring, Anderson Valley, between Sonoma and Mendocino Cos.).

  • — Scheerpeltz, 1933: 1444 (Quedius; catalog).

  • — Smetana, 1971: 216 (Quedius; synonym of nanulus).

helenae Casey, 1915: 403 (Quediellus; Type locality: Montana: Helena).

  • — Scheerpeltz, 1933: 1443 (Quedius; catalog).

  • — Smetana, 1971: 216 (Quedius; synonym of nanulus).

narada Hatch, 1957: 228 (Quedius; subgenus Sauridus; Type locality: Washington: Mt. Rainier, Paradise Park).

  • — Smetana, 1971: 179 (Quedius; subgenus Distichalius; characters; Oregon; Washington).

  • — Smetana, 1976c: 176 (Quedius; subgenus Distichalius; collecting notes; characters; Oregon; Washington).

  • Distribution: USA.

navaricus Coiffait, 1967e: 397 (Quedius; subgenus Sauridus; Type locality: Cueva Orobe, Alsasua, Navarre espagnole).

  • — Coiffait, 1978: 222 (Quedius; subgenus Sauridus; characters; Spain).

  • Distribution: Spain.

nelsonensis Blackburn, 1903: 93 (Quedius; Type locality: Victoria, Nelson R. District).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; catalog).

  • — Lea, 1925: 242 (Quedius; characters; Australia).

  • — Oke, 1933: 123, 124 (Quedius; tarsal characters; notes and characters; Australia).

  • Distribution: Australia.

nemoralis Baudi, 1848: 131 (Quedius; Type locality: provincia Montisferrati, [Pedemontium]).

  • — Thomson, 1867: 170 (Raphirus; [Note: Misidentification: Scheerpeltz, 1933: 1451, cited suturalis Thomson as a synonym of nemoralis; Thomson did not describe the species as new, he attributed it to Kraatz who attributed it to Kiesenwetter]; characters; Scandinavia).

  • — Thomson, 1867a: 44 (Raphirus; [Note: Misidentification: Scheerpeltz, 1933: 1451, cited suturalis Thomson as a synonym of nemoralis; Thomson did not describe the species as new, he attributed it to Kraatz who attributed it to Kiesenwetter]; characters; Scandinavia).

  • — Fauvel, 1874: 297 [= 1874a: 523] (Quedius; synonym of oblitteratus).

  • — Ganglbauer, 1895: 408 (Quedius; synonym of oblitteratus).

  • — Porta, 1907: 138 (Quedius; synonym of oblitteratus).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; synonym of oblitteratus).

  • — Gridelli, 1925: 131 (Quedius; subgenus Raphirus; characters; Europe; Mediterranean region; Turkestan).

  • — Donisthorpe, 1930: 94 (Quedius; characters; Britain).

  • — Scheerpeltz, 1933: 1450 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 40 (Quedius; aedeagal illustration).

  • — V. Hansen, 1952: 149 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Scheerpeltz, 1956c: 1107 (Quedius; subgenus Raphirus; Cyprus).

  • — Brakman, 1956: 39 (Quedius; notes; Netherlands).

  • — Smetana, 1958: 377 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1961d: 134 (Quedius; subgenus Raphirus; Italy).

  • — Smetana, 1962b: 146 (Quedius; subgenus Raphirus; characters; Europe).

  • — Coiffait, 1963a: 390 (Quedius; subgenus Sauridus; characters).

  • — Scheerpeltz, 1963a: 420 (Quedius; subgenus Raphirus; Greece).

  • — Palm, 1963b: 149 (Quedius; subgenus Raphirus; characters; Sweden; Denmark; Finland).

  • — Lohse, 1964: 216 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Scheerpeltz, 1965b: 408 (Quedius; Sicily).

  • — Horion, 1965: 286 (Quedius; subgenus Raphirus; Caucasus; Iran; Turkestan; Morocco; all southern Europe from Balkans to Iberian Peninsula; Russia; Poland; Czechoslovakia; Hungary; Romania; Switzerland; Austria; Germany; France; Belgium; Holland; Finland; Sweden; Denmark; Britain).

  • — Coiffait, 1968c: 505 (Quedius; Corsica).

  • — Fagel, 1968c: 5 (Quedius; subgenus Sauridus; characters; notes; Dalmatia; Herzegowina; France; England; Belgium; Italy; Sicily; Spain; Sweden; Finland; Greece; Romania).

  • — Coiffait, 1973a: 275 (Quedius; Morocco).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 214 (Quedius; subgenus Sauridus; characters; Europe; England; Scandinavia; Iberian peninsula; Italy; Yugoslavia).

  • — Outerelo, 1980a: 57 (Quedius; Spain).

  • — H. Franz, 1984: 59 (Quedius; Macedonia).

  • — Tóth, 1984: 138 (Quedius; characters; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; Benelux; Denmark; Sweden).

  • — Lohse and Lucht, 1989: 175 (Quedius; characters).

  • — Whitehead, 1990e: 12 (Quedius; collecting notes; Britain).

  • — Allen, 1990b: 3 (Quedius; collecting notes; Britain).

  • — Zanetti, 1992b: 130, 131 (Quedius; characters; Italy).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Europe, Turkey.

marginalis Kraatz, 1857d: 512 (Quedius; [preoccupied]; Type locality: nördliches Deutschland; bei Freienwalde).

  • — Redtenbacher, 1857: 986 (Quedius; characters; Austria).

  • — Gemminger and Harold, 1868: 571 (Quedius; synonym of mutatus).

  • — Fauvel, 1874: 299 [= 1874a: 525] (Quedius; synonym of limbatus).

  • — Redtenbacher, 1874: 202 (Quedius; characters).

  • — Ganglbauer, 1895: 408 (Quedius; synonym of oblitteratus).

  • — Porta, 1907: 138 (Quedius; synonym of oblitteratus).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; synonym of oblitteratus).

  • — Scheerpeltz, 1933: 1450 (Quedius; synonym of nemoralis).

  • — Smetana, 1958: 377 (Quedius; synonym of nemoralis).

mutatus Kraatz, 1868a: 104 (Quedius; replacement name for marginalis Kraatz).

  • — Fauvel, 1874: 299 [= 1874a: 525] (Quedius; synonym of limbatus).

  • — Ganglbauer, 1895: 408 (Quedius; synonym of oblitteratus).

  • — Porta, 1907: 138 (Quedius; synonym of oblitteratus).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; synonym of oblitteratus).

  • — Scheerpeltz, 1933: 1450 (Quedius; synonym of nemoralis).

  • — Smetana, 1958: 377 (Quedius; synonym of nemoralis).

ovaliceps Mulsant and Rey, 1876: 744 [= 1877: 600] (Quedius; Type locality: Not cited).

  • — Duvivier, 1883: 139 (Quedius; catalog; synonym of oblitteratus).

  • — Ganglbauer, 1895: 408 (Quedius; synonym of oblitteratus).

  • — Porta, 1907: 138 (Quedius; synonym of oblitteratus).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; synonym of oblitteratus).

  • — Scheerpeltz, 1933: 1451 (Quedius; synonym of nemoralis).

  • — Smetana, 1958: 377 (Quedius; synonym of nemoralis).

erinci Korge, 1971a: 55 (Quedius; subspecies of nemoralis; Type locality: Bosburun-Gebirge im Pisidischen Taurus in ca. 1700 m).

  • — Coiffait, 1978: 214 (Quedius; subgenus Sauridus; subspecies of nemoralis; characters).

neomexicanus Smetana, 1971, see: leechi Smetana, 1971.

neomolochinus Korge, 1962, see: laticollis Gravenhorst, 1802.

neotomae Hatch, 1957, see: erythrogaster Mannerheim, 1852.

neu Smetana, 1995, see: Quetarsis.

nevadensis Casey, 1915: 405 (Distichalius; Type locality: Nevada: Ormsby Co.).

  • — Scheerpeltz, 1933: 1451 (Quedius; catalog).

  • — Hatch, 1957: 218 (Quedius; subgenus Distichalius; characters; British Columbia; Washington; Oregon; Idaho).

  • — Smetana, 1964: 37 (Quedius; subgenus Distichalius; Alaska).

  • — Smetana, 1965a: 13 (Quedius; subgenus Distichalius; British Columbia; Alaska).

  • — Smetana, 1971: 167 (Quedius; subgenus Distichalius; characters; Alberta; British Columbia; Yukon Territory; California; Idaho; Montana; Nevada; Oregon).

  • — Smetana, 1971a: 1845 (Quedius; subgenus Distichalius; collecting notes; California).

  • — Smetana, 1976c: 174 (Quedius; subgenus Distichalius; Washington).

  • — Smetana, 1978a: 827 (Quedius; subgenus Distichalius; collecting notes; Washington).

  • Distribution: Canada, USA.

nevesi Scheerpeltz, 1951, see: pineti Brisout, 1866.

newtoni Smetana, 1976: 117 (Quedius; subgenus Microsaurus; Type locality: Mexico: Mexico: W. slope Nevado de Toluca, 11,000 ft.).

  • Distribution: Mexico.

nichinaiensis Coiffait, 1982, see: fluviatilis Cameron, 1926.

nidicola Smetana, 1971: 87 (Quedius; subgenus Microsaurus; Type locality: Oregon: Benton Co.: Shotpouch Cr. nr. Mary's Pk, 16 mi. W. Corv. Also cited from Washington).

  • Distribution: USA.

niger J. Sahlberg, 1876, see: molochinus Gravenhorst, 1806.

niger Porta, 1907 [Note: Nomen nudum], see: cinctus Paykull, 1790.

niger Gridelli, 1922, see: collaris Erichson, 1840.

nigerrimus Sharp, 1884: 338 (Quedionuchus; Type locality: Guatemala, Cerro Zunil 4000 to 5000 feet;; [Note: See lectotype designation by Smetana, 1976a]. Also cited from Guatemala: San Geronimo, 3000 feet).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; catalog).

  • — Smetana, 1976a: 228 (Quedius; subgenus Quedionuchus; lectotype designation: Cerro Zunil, 4000 to 5000 ft., Guatemala; characters; Guatemala).

  • Distribution: Guatemala.

nigribasis Korge, 1960 [Note: Nomen nudum], see: molochinus Gravenhorst, 1806.

nigricans Gridelli, 1924, see: crassus Fairmaire, 1860.

nigriceps Kraatz, 1857d: 510 (Quedius; Type locality: südliches Deutschland).

  • — Fairmaire and Laboulbène, 1856: 541 (Quedius; [Note: Misidentification: Bernhauer and Schubert, 1916: 429, cited maurorufus Fairmaire and Laboulbène as a synonym of nigriceps; Fairmaire and Laboulbène did not describe the species as new, they attributed it to Gravenhorst]; characters; France).

  • — Redtenbacher, 1857: 986 (Quedius; characters; Austria).

  • — G. Waterhouse, 1858: 23 (Quedius; synonym of ruficollis).

  • — Thomson, 1867: 169 (Raphirus; characters; Scandinavia).

  • — Thomson, 1867a: 44 (Raphirus; characters; Scandinavia).

  • — Fauvel, 1874: 295 [= 1874a: 521] (Quedius; [Note: Misidentification: Bernhauer and Schubert, 1916: 429, cited praecox Fauvel as a synonym of nigriceps; Fauvel did not describe the species as new, he attributed it to Gravenhorst]; characters; France; Europe).

  • — Fauvel, 1874: 295 [= 1874a: 521] (Quedius; synonym of praecox).

  • — Redtenbacher, 1874: 202 (Quedius; characters).

  • — Seidlitz, 1875: 265 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 707 [= 1877: 563] (Quedius; subgenus Sauridus; characters; France).

  • — Fauvel, 1878: 128 [= 1878a: 48] (Quedius; Algeria).

  • — Fowler, 1888: 239 (Quedius; subgenus Sauridus; characters; Britain).

  • — Ganglbauer, 1895: 407 (Quedius; subgenus Sauridus; characters; Europe).

  • — Porta, 1907: 137 (Quedius; subgenus Sauridus; characters; Italy).

  • — Reitter, 1909: 113 (Quedius; subgenus Sauridus; characters; Germany).

  • — Johansen, 1914: 365 (Quedius; subgenus Sauridus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; catalog).

  • — Gridelli, 1922b: 127 (Quedius; subgenus Sauridus; characters).

  • — Munster, 1923: 197 (Quedius; subgenus Sauridus; Fennoscandia).

  • — Gridelli, 1925: 116 (Quedius; subgenus Raphirus; characters).

  • — Scheerpeltz, 1933: 1451 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 32 (Quedius; aedeagal illustration).

  • — V. Hansen, 1952: 144 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Smetana, 1958: 373 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 144 (Quedius; subgenus Raphirus; characters).

  • — Coiffait, 1963a: 378 (Quedius; subgenus Sauridus; characters).

  • — Palm, 1963b: 146 (Quedius; subgenus Raphirus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 216 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 282 (Quedius; subgenus Raphirus; Britain; Norway; Sweden; Finland; Denmark; Holland; Belgium; France; Portugal; Spain; Switzerland; Czechoslovakia; Poland; Italy; Romania; Germany; Austria).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 251 (Quedius; subgenus Sauridus; characters; Europe; Britain; Scandinavia).

  • — Outerelo, 1980a: 57 (Quedius; Spain).

  • — Tóth, 1984: 133 (Quedius; characters; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; France; Benelux; Denmark; Sweden).

  • — Lohse and Lucht, 1989: 173 (Quedius; characters).

  • — Nowosad, 1990: 146 (Quedius; collected from mole nest; notes; Poland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Gerend and Braunert, 1997: 201 (Quedius; notes; Luxembourg).

  • — Owen, 1999b: 61 (Quedius; captured underground at roots of old trees; Britain).

  • — Owen, 2000: 252 (Quedius; pitfall trap collections from underground at roots of trees; Britain).

  • Distribution: Europe, Madeira.

ruficollis Stephens, 1833: 244 (Raphirus; [preoccupied]; Type locality: Edinburgh; Carlisle; coast of Devonshire; Swansea; Ipswich).

  • — Stephens, 1839: 394 (Quedius; characters; England; Scotland).

  • — Hardy, 1851: 33 (Quedius; notes; England).

  • — Gemminger and Harold, 1868: 571 (Quedius; synonym of nigriceps).

  • — Fauvel, 1874: 295 [= 1874a: 521] (Quedius; synonym of praecox).

  • — Fauvel, 1876a: 77 [= 1876: 262] (Quedius; synonym of nigriceps).

  • — Fowler, 1888: 239 (Quedius; subgenus Sauridus; synonym of nigriceps).

  • — Ganglbauer, 1895: 407 (Quedius; synonym of nigriceps).

  • — Porta, 1907: 137 (Quedius; synonym of nigriceps).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; synonym of nigriceps).

  • — Smetana, 1958: 373 (Quedius; synonym of nigriceps).

maderensis Smetana, 1963b: 40 (Quedius; subgenus Raphirus; subspecies of nigriceps; Type locality: Madeira: Queimadas).

  • — Coiffait, 1978: 251 (Quedius; subgenus Sauridus; subspecies of nigriceps; characters).

nigricollis Fauvel, 1877: 271 (Quedius; Type locality: Australie, Rockhampton).

  • — Fauvel, 1878g: 553 (Quedius; Australia).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; catalog).

  • Distribution: Australia.

nigricollis W. Kolbe, 1909 [Note: Nomen nudum], see: fulvicollis Stephens, 1833.

nigricornis Stephens, 1832, see: fulgidus Fabricius, 1793.

nigricornis Holme, 1842, see: mesomelinus Marsham, 1802.

nigrifrons Bernhauer, 1934, see: flavinasus Bernhauer, 1917.

nigrinus Porta, 1907 [Note: Nomen nudum], see: molochinus Gravenhorst, 1806.

nigripennis J. Sahlberg, 1876, see: mesomelinus Marsham, 1802.

nigripennis Hänel, 1913 [Note: Nomen nudum], see: molochinus Gravenhorst, 1806.

nigripennis Delahon, 1914 [Note: Nomen nudum], see: molochinus Gravenhorst, 1806.

nigritarsis Ménétriés, 1832, see: levicollis Brullé, 1832.

nigritarsis Roubal, 1949 [Note: Nomen nudum], see: unicolor Kiesenwetter, 1847.

nigritulus R. Sahlberg, 1847, see: Heterothops.

nigrocaeruleus Fauvel, 1876a: 76 [= 1876: 261] (Quedius; Type locality: Paris; Orival; Sos; Tarbes; Portugal).

  • — Mulsant and Rey, 1876: 644 [= 1877: 500] (Quedius; characters; larval characters; France).

  • — Ganglbauer, 1895: 398 (Quedius; variety of ochripennis).

  • — Ganglbauer, 1895: 399 (Quedius; subgenus Microsaurus; variety of fulgidus; [Note: Misidentification: Scheerpeltz, 1933: 1452, cited depauperatus Ganglbauer as a synonym of nigrocaeruleus; Ganglbauer cited depauperatus Wollaston as a variety of fulgidus and two lines down the page used the name again as “(variety depauperatus)”; I assume Ganglbauer was referring to Wollaston's variety and not proposing a new variety]; characters).

  • — Porta, 1907: 113 (Quedius; variety of ochripennis).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; cited as nigrocoeruleus; variety of ochripennis).

  • — Gridelli, 1924d: 45 (Quedius; subgenus Microsaurus; variety of ochripennis; France; Portugal; Germany; England).

  • — Gridelli, 1929: 25 (Quedius; cited as nigrocoeruleus; characters; Britain, Germany, Carniola, France).

  • — Scheerpeltz, 1933: 1452 (Quedius; catalog).

  • — Roubal, 1941b: 123, 127 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Paulian, 1941: 265 (Quedius; cited as nigrocoeruleus; larval characters).

  • — V. Hansen, 1952: 133 (Quedius; subgenus Microsaurus; cited as nigrocoeruleus; characters; Denmark).

  • — Smetana, 1958: 346 (Quedius; subgenus Microsaurus; cited as nigrocoeruleus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 156 (Quedius; subgenus Microsaurus; cited as nigrocoeruleus; characters).

  • — Palm, 1963b: 133 (Quedius; subgenus Microsaurus; cited as nigrocoeruleus; characters; Sweden; Denmark).

  • — Lohse, 1964: 209 (Quedius; subgenus Microsaurus; cited as nigrocoeruleus; characters; central Europe).

  • — Smetana, 1964d: 79 (Quedius; subgenus Microsaurus; cited as nigrocoeruleus; Czechoslovakia).

  • — Horion, 1965: 259 (Quedius; subgenus Microsaurus; cited as nigrocoeruleus; Britain; Denmark; Sweden; all central Europe; Portugal; Spain; Italy; Slovenia; Dalmatia; Greece).

  • — Pototskaia, 1967: 83 (Quedius; cited as nigrocoeruleus; larval characters).

  • — Bordoni, 1974: 63 (Quedius; subgenus Microsaurus; cited as nigrocoeruleus; characters; Italy).

  • — Osella and Zanetti, 1975: 117 (Quedius; collected from nest of Talpa europaea; notes; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 147 (Quedius; subgenus Microsaurus; characters; Europe; Britain; France).

  • — Tóth, 1984: 115 (Quedius; cited as nigrocoeruleus; characters).

  • — Lucht, 1987: 109 (Quedius; cited as nigrocoeruleus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Nowosad, 1990: 129 (Quedius; collected from mole nest; notes; Poland).

  • — Welch, 1993: 229 (Quedius; ovariole number and ovary structure).

  • — Smetana, 1993b: 50 (Quedius; subgenus Microsaurus; cited as nigrocoeruleus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Allen, 1996: 294 (Quedius; collecting notes; England).

  • — Assing and Schülke, 1999: 15 (Quedius; correction of spelling of name).

  • — Assing, 2001: 75 (Quedius; cited as nigrocoeruleus; collected from xerothermous site; Germany).

  • Distribution: Europe.

variabilis Mulsant and Rey, 1876: 649 [= 1877: 505] (Quedius; [preoccupied]; Type locality: environs de Lyon et le Beaujolais).

  • — Ganglbauer, 1895: 398 (Quedius; synonym of variabilis).

  • — Porta, 1907: 113 (Quedius; synonym of ochripennis).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; synonym of ochripennis).

  • — Scheerpeltz, 1933: 1452 (Quedius; synonym of nigrocaeruleus).

  • — Smetana, 1958: 346 (Quedius; synonym of nigrocaeruleus).

  • — Bordoni, 1974: 63 (Quedius; synonym of nigrocaeruleus).

balianii Gridelli, 1924d: 45 (Quedius; subgenus Microsaurus; variety of fulgidus; Type locality: Busalla, Stazzano Scrivia [Apennino ligure]; Lippiano, M.ti Pistoiesi, M.te Falterona [Toscana]; Bassano Sutri, M.te Serrasecca, M.te Autore, M.te Cavo, Carpineto [Lazio]; Vallo Lucano [Campania]; Vulture [Basilicata]; M.te Alto [Sicilia]; Cagliari [Sardegna]).

  • — Gridelli, 1929: 29 (Quedius; variety of fulgidus; characters).

  • — Scheerpeltz, 1933: 1452 (Quedius; variety of nigrocaeruleus).

  • — Roubal, 1941b: 124 (Quedius; subgenus Microsaurus; variety of fulgidus; characters).

  • — Smetana, 1958: 349 (Quedius; synonym of nigrocaeruleus).

  • — Smetana, 1960e: 158 (Quedius; aberration of fulgidus; characters).

  • — Scheerpeltz, 1965b: 406 (Quedius; variety of nigrocaeruleus; Sicily).

  • — Bordoni, 1974: 63, 66 (Quedius; synonym of nigrocaeruleus and fulgidus).

  • — Coiffait, 1978: 147 (Quedius; subgenus Microsaurus; variety of nigrocaeruleus; characters).

  • — Tóth, 1984: 115 (Quedius; variety of nigrocaeruleus; characters).

nigromarginalis Polentz, 1932, see: longicornis Kraarz, 1857.

nigropolitus Cameron, 1937: 29 (Quedius; subgenus Sauridus; Type locality: Java: Idjen Plateau, alt. 900–1500 m).

  • Distribution: Indonesia.

nilgiriensis Cameron, 1920, see: Philonthus.

nilo Smetana, 1988: 287 (Quedius; subgenus Raphirus; Type locality: Nepal: Khandbari Dis., Arun Valley at Num main bridge, 1050 m).

  • Distribution: Nepal.

nireus Smetana, 1995f: 240 (Quedius; subgenus Microsaurus; Type locality: China: Gansu, Mts. 25 km E. Xiahe, 2805–2925 m).

  • Distribution: China.

nishikawai Watanabe, 1991b: 225 (Quedius; subgenus Microsaurus; Type locality: southern Taiwan: P'ing-tung Hsien, Tai-wu Hsiang, Mt. Pei-ta-wu Shan, 2520 m alt.).

  • — Smetana, 1995: 48 (Quedius; subgenus of Microsaurus; characters; bionomics; Taiwan).

  • Distribution: Taiwan.

nitescens Stephens, 1832, see: Philonthus.

nitidissimus Lea, 1925: 246 (Quedius; Type locality: Tasmania: Waratah).

  • — Scheerpeltz, 1933: 1452 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

nitidulus Scheerpeltz, 1933: 1452 (Quedius; [Note: Nomen nudum (Article 13.1.1); this name was attributed to Casey, 1915: 413 but neither I nor Smetana, 1971 found the species as cited by Scheerpeltz]).

nitipennis Stephens, 1833: 242 (Raphirus; [Note: Many references to this species were cited as attenuatus Gyllenhal but Gyllenhal's use was a misidentification; the citations of the species as attenuatus Gyllenhal are listed below under that name but that use is an unavailable name]; Type locality: Battersea-fields; Devonshire).

  • — Stephens, 1839: 393 (Quedius; characters; England).

  • — Hardy, 1851: 34 (Quedius; synonym of attenuatus “Gyllenhal”).

  • — Fauvel, 1874: 305 [= 1874a: 531] (Quedius; synonym of semiaeneus).

  • — Porta, 1907: 149 (Quedius; cited as nitidipennis; synonym of picipennis).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; catalog).

  • — Munster, 1923: 200 (Quedius; subgenus Raphirus; Fennoscandia).

  • — Gridelli, 1925: 156 (Quedius; synonym of semiaeneus).

  • — Scheerpeltz, 1933: 1462 (Quedius; synonym of semiaeneus).

  • — Tottenham, 1948a: 249 (Quedius; subgenus Arphirus; valid species; characters; Britain).

  • — Tottenham, 1949: 422 (Quedius; checklist; Britain).

  • — V. Hansen, 1952: 155 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Smetana, 1958: 393 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1959a: 215 (Quedius; compared to incensus).

  • — Smetana, 1962b: 150 (Quedius; subgenus Raphirus; characters; Europe).

  • — Palm, 1963b: 152 (Quedius; subgenus Raphirus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 218 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Smetana, 1964d: 80 (Quedius; subgenus Raphirus; Czechoslovakia).

  • — Horion, 1965: 303 (Quedius; synonym of attenuatus “Gyllenhal”).

  • — Szujecki, 1968a: 736 (Quedius; Poland).

  • — Shilov, 1975a: 377 [= 1975b: 99] (Quedius; Komi region of Russia).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 269 (Quedius; subgenus Raphirus; characters; Europe; Britain; Scandinavia).

  • — Smetana, 1978: 125 (Quedius; subgenus Raphirus; collecting notes; Israel).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 146 (Quedius; catalog; Poland).

  • — Chemini and Zanetti, 1982: 216 (Quedius; habitat notes; Italy).

  • — H. Franz, 1984: 59 (Quedius; cited as nitidipennis; Macedonia).

  • — Tóth, 1984: 127 (Quedius; characters; Hungary).

  • — Muona and Viramo, 1986: 14 (Quedius; Finland).

  • — Ádám, 1987: 153 (Raphirus; collecting notes; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Zanetti, 1989: 112 (Quedius; Italy).

  • — Nowosad, 1990: 149 (Quedius; collected from mole nest; notes; Poland).

  • — Whitehead, 1990e: 13 (Quedius; collecting notes; Britain).

  • — Silfverberg, 1992: 18 (Quedius; Finland; Russia; Sweden; Norway; Denmark; Estonia; Lithuania).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Franzen, 1996: 207 (Quedius; collecting notes; Germany).

  • — M. Hansen, 1996: 108 (Quedius; checklist; Denmark).

  • — Eisinger, 1997: 171 (Quedius; collecting notes; Germany).

  • — Wenzel, 1997: 347 (Quedius; collecting notes; Germany).

  • — Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 39 (Quedius; Latvia).

  • — Anderson, Nash, and O'Connor, 1997: 19 (Quedius; Ireland).

  • — Herman, 2001: 30 (Quedius; nomen protectum).

  • Distribution: Tunisia, Algeria, Morocco, Iceland, Europe, Russia, Turkey, Israel.

attenuatus: Gyllenhal, 1810: 311 (Staphylinus; [Note: Nomen nudum, misidentification: Bernhauer and Schubert, 1916: 429 cited attenuatus as a synonym of nitipennis Stephens; Gyllenhal did not describe the species as new, he attributed it to Gravenhorst]; Type locality: Not cited).

  • — Erichson, 1839a: 483 (Quedius; characters; Germany).

  • — Erichson, 1840: 546 (Quedius; characters; Germany; England; Sweden).

  • — P. Lucas, 1846: 114 (Quedius; notes; Algeria).

  • — Kiesenwetter, 1848: 62 (Quedius; characters; Europe).

  • — Redtenbacher, 1849: 710 (Philonthus; characters; Austria).

  • — Hardy, 1851: 34 (Quedius; notes; England).

  • — Fairmaire and Laboulbène, 1856: 538 (Quedius; characters; France).

  • — Staudinger, 1857: 286 (Quedius; notes; Iceland).

  • — Kraatz, 1857d: 515 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 202 (Quedius; characters; Austria).

  • — Thomson, 1859: 26 (type species of Raphirus).

  • — Thomson, 1860: 179 (Raphirus; characters; Scandinavia).

  • — Fauvel, 1874: 306 [= 1874a: 532] (Quedius; characters; France; Scandinavia; Britain; Germany; Switzerland; Italy; Austria; Russia; Siberia).

  • — Redtenbacher, 1874: 199 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 264 (Quedius; characters; Baltic region).

  • — Fauvel, 1876a: 78 [= 1876: 263] (Quedius; Spain; Portugal; Siberia).

  • — Mulsant and Rey, 1876: 787 [= 1877: 642] (Quedius; subgenus Raphirus; characters; France).

  • — J. Sahlberg, 1876: 29 (Raphirus; Finland).

  • — Fowler, 1888: 243 (Quedius; subgenus Raphirus; characters; Britain).

  • — Ganglbauer, 1895: 411 (Quedius; synonym of picipennis).

  • — Ganglbauer, 1895: 411 (Quedius; subgenus Raphirus; [Note: Misidentification: Scheerpeltz, 1933: 1432, cited picipennis Ganglbauer as a synonym of attenuatus; Ganglbauer did not describe the species as new, he attributed it to Heer]; characters; north and middle Europe).

  • — Porta, 1907: 149 (Quedius; synonym of picipennis).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; synonym of nitipennis).

  • — Gridelli, 1925: 161 (Quedius; subgenus Raphirus; characters; Germany; Silesia; Carniola; Alps; Italy).

  • — Portevin, 1929: 345 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — West, 1937: 38 (Quedius; Faroe Islands).

  • — Tottenham, 1939d: 234 (Quedius; synonym of hyperboreus).

  • — Wüsthoff, 1938a: fig. 61, 62 (Quedius; aedeagal illustration).

  • — C. Koch, 1941: 68 (Quedius; Morocco).

  • — Normand, 1947a: 9 (Quedius; Tunisia).

  • — Tottenham, 1948a: 249 (Quedius; name cited in discussion of nitipennis as available but preoccupied; synonym of nitipennis).

  • — Smetana, 1958: 393 (Quedius; synonym of nitipennis).

  • — Scheerpeltz, 1960d: 136 (Quedius; subgenus Raphirus; Portugal).

  • — Scheerpeltz, 1961d: 135 (Quedius; subgenus Raphirus; Italy).

  • — Smetana, 1962b: 152 (Quedius; synonym of nitipennis).

  • — Brakman, 1962: 46 (Quedius; characters).

  • — Lohse, 1964: 218 (Quedius; synonym of nitipennis).

  • — Korge, 1964: 124 (Quedius; subgenus Raphirus; cited as valid species).

  • — Focarile, 1964: 60 (Quedius; Italy).

  • — Horion, 1965: 303 (Quedius; subgenus Raphirus; Turkey; Caucasus; Morocco; northern through all central Europe; Spain; Italy; Croatia; Bosnia Hercegovina).

  • — Smetana, 1967i: 559 (Quedius; subgenus Raphirus; Turkey).

  • — Scheerpeltz, 1974c: 25 (Quedius; Tunisia).

  • — Coiffait, 1978: 269 (Quedius; synonym of nitipennis).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; synonym of nitipennis; Italy).

quadripunctatus Zetterstedt, 1828: 76 (Staphylinus; [Note: This name is older than nitipennis]; Type locality: Lapponia Tornensis).

  • — Zetterstedt, 1838: 64 (Staphylinus; characters; habitat; Lapland).

  • — Thomson, 1860: 179 (Raphirus; synonym of attenuatus “Gyllenhal”).

  • — Herman, 2001: 30 (Quedius; nomen oblitum).

rufipennis Stephens, 1833: 243 (Raphirus; Type locality: Suffolk).

  • — Holme, 1837: 63 (Raphirus; notes; Scilly Islands).

  • — Stephens, 1839: 393 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 23 (Quedius; synonym of attenuatus “Gyllenhal”).

  • — Gemminger and Harold, 1868: 568 (Quedius; synonym of attenuatus “Gyllenhal”).

  • — Fauvel, 1874: 305 [= 1874a: 531] (Quedius; synonym of semiaeneus).

  • — Porta, 1907: 149 (Quedius; synonym of picipennis).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; synonym of nitipennis).

  • — Scheerpeltz, 1933: 1432 (Quedius; variety of attenuatus “Gyllenhal”).

  • — Tottenham, 1939d: 234 (Quedius; synonym of fulvicollis).

  • — Smetana, 1958: 394 (Quedius; aberration of nitipennis).

  • — Smetana, 1962b: 152 (Quedius; aberration of nitipennis; characters).

  • — Coiffait, 1978: 270 (Quedius; subgenus Raphirus; variety of nitipennis; characters).

  • — Tóth, 1984: 128 (Quedius; variety of nitipennis; characters).

fallaciosus Kraatz, 1862a: 268 (Quedius; Type locality: Nähe Berlins).

  • — Fauvel, 1869: 486 (Quedius; synonym of attenuatus “Gyllenhal”).

  • — Fauvel, 1874: 306 [= 1874a: 532] (Quedius; synonym of attenuatus “Gyllenhal”).

  • — Redtenbacher, 1874: 202 (Quedius; characters).

  • — Ganglbauer, 1895: 412 (Quedius; synonym of boops).

  • — Porta, 1907: 147 (Quedius; synonym of acuminatus).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; synonym of acuminatus).

  • — Gridelli, 1925: 161 (Quedius; synonym of attenuatus “Gyllenhal”).

  • — Scheerpeltz, 1933: 1432 (Quedius; synonym of attenuatus “Gyllenhal”).

  • — Smetana, 1958: 393 (Quedius; synonym of nitipennis).

  • — Smetana, 1962b: 152 (Quedius; synonym of nitipennis).

  • — Scheerpeltz, 1974c: 25 (Quedius; synonym of attenuatus “Gyllenhal”).

  • — Coiffait, 1978: 270 (Quedius; synonym of nitipennis).

platyphthalmus Roubal, 1955: 129 (Quedius; subgenus Raphirus; Type locality: Carpathes, ad urbem Mukacevo).

  • — Smetana, 1966: 332 (Quedius; subgenus Raphirus; synonym of attenuatus “Gyllenhal”).

  • — Coiffait, 1978: 270 (Quedius; synonym of nitipennis).

nivicola Kiesenwetter, 1858: 58 (Quedius; Type locality: Parnes bei Athen).

  • — Fauvel, 1869: 487 (Quedius; synonym of cruentus).

  • — Fauvel, 1874: 282 [= 1874a: 508] (Quedius; synonym of cruentus).

  • — Bernhauer, 1905b: 596 (Quedius; Greece; Syria; Turkey).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1452 (Quedius; catalog).

  • — Coiffait, 1955e: 427 (Quedius; Lebanon).

  • — Coiffait, 1963a: 390 (Quedius; subgenus Sauridus; characters).

  • — Fagel, 1968b: 2 (Quedius; subgenus Sauridus; characters; Greece; Iran).

  • — Coiffait, 1978: 217 (Quedius; subgenus Sauridus; characters; Greece; Israel; Iran).

  • Distribution: Greece, Lebanon, Israel, Iran.

noboruitoi Hayashi, 1992a: 113 (Quedius; subgenus Microsaurus; replacement name for flavocaudatus Hayashi).

  • — Smetana, 1995: 34 (Quedius; subgenus of Microsaurus; characters; bionomics; Taiwan).

  • — Smetana, 1996g: 23 (Quedius; notes; Taiwan).

  • Distribution: Taiwan.

flavocaudatus Hayashi, 1992: 14 (Quedius; subgenus Microsaurus; [preoccupied]; Type locality: Taiwan: Tungpu, Nantou Hsien).

  • — Hayashi, 1992a: 113 (Quedius; synonym of noboruitoi).

  • — Smetana, 1995: 34 (Quedius; synonym of noboruitoi).

noricus Bernhauer, 1927b: 96 (Quedius; subgenus Sauridus; Type locality: Umgebung von Hofgastein).

  • — Scheerpeltz, 1933: 1452 (Quedius; catalog).

  • — Coiffait, 1963a: 387 (Quedius; subgenus Sauridus; characters).

  • — Horion, 1965: 285 (Quedius; subgenus Raphirus; Austria).

  • — Coiffait, 1978: 249 (Quedius; subgenus Sauridus; characters; Alps; mountains of central Europe).

  • — Lucht, 1987: 110 (Quedius; checklist; Austria).

  • — Lohse and Lucht, 1989: 173 (Quedius; characters; mountains of central Europe).

  • Distribution: Austria.

notabilis Cameron, 1932, see: beesoni Cameron, 1932.

nothus Lea, 1925: 250 (Quedius; Type locality: Queensland: National Park. New South Wales: Illawarra).

  • — Scheerpeltz, 1933: 1452 (Quedius; catalog).

  • Distribution: Australia.

nouristanicus Coiffait, 1979, see: nouristanicus Coiffait, 1978.

nouristanicus Coiffait, 1978: 158 (Quedius; subgenus Microsaurus; Type locality: vallée du Parum, 2400–2800 m. Nouristan, Afghanistan).

  • — Herman, 2001: 30 (Quedius; synonymic note).

  • Distribution: Afghanistan.

nouristanicus Coiffait, 1979d: 559 (Quedius; subgenus Microsaurus; [preoccupied]; Type locality: Nouristan, vallée du Parum, 2400–2800 m).

  • — Herman, 2001: 30 (Quedius; synonym of nouristanicus Coiffait, 1978).

novus Eppelsheim, 1892: 331 (Quedius; Type locality: Taschkent; Margelan).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; catalog).

  • — Gridelli, 1925: 125 (Quedius; subgenus Raphirus; characters; Tashkent).

  • — Scheerpeltz, 1933: 1452 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 37 (Quedius; aedeagal illustration).

  • — Coiffait, 1963a: 389 (Quedius; subgenus Sauridus; characters).

  • — Smetana, 1995a: 84 (Quedius; subgenus Raphirus; Uzbekistan).

  • — Coiffait, 1970d: 143 (Quedius; Uzbekistan).

  • — Coiffait, 1978: 228 (Quedius; subgenus Sauridus; characters; mountains of central Asia).

  • — Boháč, 1988: 556 [= 1989: 40] (Quedius; notes).

  • Distribution: Uzbekistan.

obliqueseriatus Eppelsheim, 1889d: 17 (Quedius; subgenus Sauridus; Type locality: Circassien).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; catalog).

  • — Gridelli, 1924d: 106 (Quedius; subgenus Raphirus; characters).

  • — Scheerpeltz, 1933: 1452 (Quedius; catalog).

  • — Coiffait, 1963a: 377 (Quedius; subgenus Sauridus; characters).

  • — Korge, 1964: 119 (Quedius; subgenus Sauridus; Turkey).

  • — Korge, 1971: 13 (Quedius; subgenus Sauridus; characters; notes; Iran).

  • — Coiffait, 1978: 216 (Quedius; subgenus Sauridus; characters; Caucasus).

  • Distribution: Russia, Caucasus, Turkey, Iran.

oblitteratus Erichson, 1840, see: humeralis Stephens, 1832.

obscuriceps Coiffait, 1967e: 404 (Quedius; subgenus Sauridus; Type locality: Caucase).

  • — Coiffait, 1978: 241 (Quedius; subgenus Sauridus; characters; Caucasus).

  • Distribution: Caucasus.

obscuripennis Bernhauer, 1901: 538 (Quedius; variety of ochropterus; Type locality: Negoi in den transsilvanischen Alpen).

  • — Reitter, 1909: 112 (Quedius; subgenus Sauridus; [Note: Misidentification: Scheerpeltz, 1933: 1453, cited brevipennis Reitter as a synonym of obscuripennis; Reitter did not describe the name as a new taxon, he cited it as an aberration of ochropterus and attributed it to Motschulsky]; characters; Germany).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; variety of ochropterus).

  • — Gridelli, 1924d: 102 (Quedius; subgenus Raphirus; valid species; characters; Italy).

  • — Scheerpeltz, 1933: 1453 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 29 (Quedius; aedeagal illustration).

  • — Smetana, 1958: 368 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 142 (Quedius; subgenus Raphirus; characters; Europe).

  • — Coiffait, 1963a: 377 (Quedius; subgenus Sauridus; characters).

  • — Lohse, 1964: 215 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 279 (Quedius; subgenus Raphirus; Alps; Germany; Austria; Switzerland; Italy; France; Slovenia; Croatia; Romania; Czechoslovakia; Poland; Bosnia Hercegovina; Macedonia).

  • — Smetana, 1966: 332 (Quedius; subgenus Raphirus; notes; Czechoslovakia).

  • — Szujecki, 1969a: 258 (Quedius; notes; Poland).

  • — Coiffait, 1978: 210 (Quedius; subgenus Sauridus; characters; Carpathians; Alps; Apennines).

  • — Tóth, 1984: 132 (Quedius; characters).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Europe.

pyrenaeicola Coiffait, 1963: 377, 405 (Quedius; subgenus Sauridus; subspecies of obscuripennis; Type locality: Hautes-Pyrénées, Fabian).

  • — Coiffait, 1978: 210 (Quedius; subgenus Sauridus; subspecies of obscuripennis; characters).

arvernicus Coiffait, 1982c: 233 (Quedius; subgenus Sauridus; subspecies of obscuripennis; [preoccupied]; Type locality: Puy-de-Dôme, Egliseneuve, d'Entraygues, tourbière de Chambedaze, 1200 m).

  • — Herman, 2001: 49 (Quedius; synonym of obscuripennis arvernus).

arvernus Herman, 2001: 49 (Quedius; subspecies of obscuripennis; replacement name for obscuripennis arvernicus).

obscurus Lokay, 1913, see: cruentus Olivier, 1795.

occultus Lacordaire, 1835, see: mesomelinus Marsham, 1802.

ocelotl Smetana, 1976: 116 (Quedius; subgenus Microsaurus; Type locality: Mexico: Michoacan: Cerro de Garnica, Puerto Garnica, 9400 ft.).

  • Distribution: Mexico.

ochripennis Ménétriés, 1832: 145 (Staphylinus; Type locality: Périmbal sur les montagnes de Talyche).

  • — Gravenhorst, 1802: 31 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1916: 428, 430, cited nitidus Gravenhorst as a synonym of mesomelinus and ochripennis but Gravenhorst did not describe the species as new, he attributed it to Fabricius; Gravenhorst cited many varieties of the species]; characters).

  • — Faldermann, 1835: 130 (Staphylinus; characters; Persia-Armenia).

  • — Hochhuth, 1849: 148 (Quedius; [Note: Misidentification: Bernhauer and Schubert, 1916: 430, cited fulgidus Hochhuth as a synonym of ochripennis; Hochhuth did not describe the species as new, he attributed it to Fabricius]; characters; Iberia; Lenkoran; Helenendorf).

  • — Fauvel, 1874: 278 [= 1874a: 504] (Quedius; characters; France; Lapland; Scandinavia; Russia; Britain; Italy; Corsica; Algeria; Caucasus).

  • — Fauvel, 1876a: 76 [= 1876: 261] (Quedius; Portugal; Syria).

  • — Mulsant and Rey, 1876: 651 [= 1877: 507] (Quedius; characters; France).

  • — Fauvel, 1878: 127 [= 1878a: 47] (Quedius; Algeria).

  • — Fowler, 1888: 233 (Quedius; synonym of puncticollis).

  • — Ganglbauer, 1895: 398 (Quedius; subgenus Microsaurus; characters; Europe; Mediterranean region).

  • — Fauvel, 1902b: 114 (Quedius; Algeria).

  • — Porta, 1907: 113 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Sainte-Claire Deville, 1906: 101 (Quedius; Corsica).

  • — Montandon, 1908: 80 (Quedius; Romania).

  • — Reitter, 1909: 111 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Johansen, 1914: 356 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; catalog).

  • — Munster, 1923: 191 (Quedius; subgenus Microsaurus; Fennoscandia).

  • — Gridelli, 1924d: 42 (Quedius; subgenus Microsaurus; characters; Europe; western Mediterranean region).

  • — Gridelli, 1929: 21 (Quedius; characters; Germany, France, Spain, Italy, Romania, Bulgaria, Macedonia, Asia Minor, Turkestan).

  • — Portevin, 1929: 340 (Quedius; characters; France).

  • — Cameron, 1932: 289 (Quedius; subgenus Microsaurus; characters; India).

  • — Scheerpeltz, 1933: 1453 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 7 (Quedius; aedeagal illustration).

  • — Roubal, 1941b: 123, 127 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Paulian, 1941: 266 (Quedius; larval characters).

  • — Jeannel and Jarrige, 1949: 382 (Quedius; subgenus Microsaurus; notes; cave species; France; Spain).

  • — V. Hansen, 1952: 131 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Scheerpeltz, 1958a: 23 (Quedius; subgenus Microsaurus; Turkey).

  • — Smetana, 1958: 345 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 157 (Quedius; subgenus Microsaurus; characters).

  • — Scheerpeltz, 1960b: 68 (Quedius; subgenus Microsaurus; Afghanistan).

  • — Scheerpeltz, 1960d: 136 (Quedius; subgenus Microsaurus; Portugal).

  • — Scheerpeltz, 1963: 15 (Quedius; subgenus Microsaurus; Afghanistan).

  • — Palm, 1963b: 132 (Quedius; subgenus Microsaurus; characters).

  • — Lohse, 1964: 210 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Scheerpeltz, 1965b: 406 (Quedius; Sicily).

  • — Horion, 1965: 258 (Quedius; subgenus Microsaurus; Ireland; Britain; Denmark; all southern and central Europe).

  • — Smetana, 1967i: 558 (Quedius; Turkey).

  • — Pototskaia, 1967: 82 (Quedius; larval characters).

  • — Coiffait, 1968c: 503 (Quedius; cave collection; Corsica).

  • — Korge, 1968: 51 (Quedius; characters; notes).

  • — Bordoni, 1974: 60 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Osella and Zanetti, 1975: 116 (Quedius; collected from nest of Talpa europaea; notes; Italy).

  • — Shilov, 1975a: 376 [= 1975b: 99] (Quedius; Komi region of Russia).

  • — Coiffait, 1978: 157 (Quedius; subgenus Microsaurus; characters; Palaearctic region).

  • — Bordoni and Carfi, 1978 (Quedius; note on peg setae of aedeagus).

  • — Coiffait, 1982a: 234 (Quedius; Pakistan).

  • — Tóth, 1984: 115 (Quedius; characters; Hungary).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark).

  • — Bruge and Drugmand 1988: 169 (Quedius; characters).

  • — Smetana, 1988: 207 (Quedius; subgenus Microsaurus; characters; Himachal Pradesh; Pakistan).

  • — Schülke and Uhlig, 1988: 12 (Quedius; notes; Germany).

  • — Nowosad, 1990: 125 (Quedius; collected from mole nest; notes; Poland).

  • — Wittwer, 1993: 251 (Quedius; Switzerland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Bordoni and Oromi, 1998: 1157 (Quedius; subgenus Microsaurus; facultative cave species).

  • — Assing, 2001: 75 (Quedius; collected from xerothermous site; Germany).

  • Distribution: Algeria, Europe, Russia, Georgia, Azerbaijan, Turkey, Syria, Iran, Tajikistan-Uzbekistan, Afghanistan, Pakistan, India.

floralis Lacordaire, 1835: 380 (Emus; Type locality: Paris).

  • — Erichson, 1839a: 486 (Quedius; synonym of fulgidus).

  • — Heer, 1839: 273 (Philonthus; synonym of fulgidus).

  • — Erichson, 1840: 526 (Quedius; synonym of fulgidus).

  • — Hardy, 1851: 32 (Quedius; synonym of fulgidus).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — Fauvel, 1869: 486 (Quedius; synonym of fulgidus).

  • — Fauvel, 1874: 278 [= 1874a: 504] (Quedius; synonym of ochripennis).

  • — Ganglbauer, 1895: 398 (Quedius; synonym of ochripennis).

  • — Porta, 1907: 113 (Quedius; synonym of ochripennis).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; synonym of ochripennis).

  • — Cameron, 1932: 289 (Quedius; synonym of ochripennis).

  • — Smetana, 1958: 345 (Quedius; synonym of ochripennis).

  • — Bordoni, 1974: 60 (Quedius; synonym of ochripennis).

  • — Coiffait, 1978: 157 (Quedius; synonym of ochripennis).

laetus Faldermann, 1835: 127 (Staphylinus; Type locality: [area] Persico-Armeniaca).

  • — Hochhuth, 1849: 148 (Quedius; synonym of fulgidus).

  • — Fauvel, 1874: 278 [= 1874a: 504] (Quedius; synonym of ochripennis).

  • — Fowler, 1888: 233 (Quedius; synonym of puncticollis).

  • — Ganglbauer, 1895: 398 (Quedius; synonym of ochripennis).

  • — Porta, 1907: 113 (Quedius; synonym of ochripennis).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; synonym of ochripennis).

  • — Cameron, 1932: 289 (Quedius; synonym of ochripennis).

  • — Smetana, 1958: 345 (Quedius; synonym of ochripennis).

  • — Bordoni, 1974: 60 (Quedius; synonym of ochripennis).

  • — Coiffait, 1978: 157 (Quedius; synonym of ochripennis).

assecla Mulsant and Rey, 1876: 655 [= 1877: 511] (Quedius; Type locality: environs de Lyon et dans la Provence).

  • — Fowler, 1888: 233 (Quedius; synonym of puncticollis).

  • — Ganglbauer, 1895: 398 (Quedius; synonym of ochripennis).

  • — Porta, 1907: 113 (Quedius; synonym of ochripennis).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; synonym of ochripennis).

  • — Cameron, 1932: 289 (Quedius; cited as asseda; synonym of ochripennis).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Korge, 1962: 152 (Quedius; subgenus Microsaurus; cited as valid species).

  • — Coiffait, 1963b: 422 (Quedius; subgenus Microsaurus; characters).

  • — Johnson, 1967a: 80 (Quedius; synonym of ochripennis).

  • — Coiffait, 1967e: 395 (Quedius; synonym of ochripennis).

  • — Bordoni, 1974: 60 (Quedius; synonym of ochripennis).

  • — Coiffait, 1978: 157 (Quedius; synonym of ochripennis).

ochropterus Erichson, 1840: 538 (Quedius; Type locality: Austria).

  • — Kiesenwetter, 1848: 58 (Quedius; [Note: Misidentification: Bernhauer and Schubert, 1916: 430, cited fimbriatus Kiesenwetter as a synonym of ochropterus; Kiesenwetter did not describe the species as new, he attributed it to Erichson]; characters; mountains of Kärnthen, Krain, Austria and Switzerland).

  • — Fairmaire and Laboulbène, 1856: 544 (Quedius; characters; France).

  • — Kraatz, 1857d: 505 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 204 (Quedius; characters; Austria).

  • — Fauvel, 1865a: 293 [= 1865b: 43] (Quedius; habitat; France).

  • — Fauvel, 1874: 291 [= 1874a: 517] (Quedius; characters; France; Switzerland; Italy; Austria).

  • — Redtenbacher, 1874: 201 (Quedius; characters; Austria).

  • — Fauvel, 1876a: 77 [= 1876: 262] (Quedius; Spain).

  • — Mulsant and Rey, 1876: 701 [= 1877: 557] (Quedius; subgenus Sauridus; characters; France).

  • — Ganglbauer, 1895: 404 (Quedius; subgenus Sauridus; characters; Pyrenees; Alps; Carpathians).

  • — Porta, 1907: 133 (Quedius; subgenus Sauridus; characters; Italy).

  • — Reitter, 1909: 112 (Quedius; subgenus Sauridus; characters; Germany).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; catalog).

  • — Gridelli, 1924d: 101 (Quedius; subgenus Raphirus; characters; Italy; Bosnia; Albania).

  • — Portevin, 1929: 342 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1454 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 28 (Quedius; aedeagal illustration).

  • — Scheerpeltz and Höfler, 1948: 157 (Quedius; subgenus Raphirus; fungus inhabitant).

  • — Smetana, 1958: 367 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 142 (Quedius; subgenus Raphirus; characters).

  • — Coiffait, 1963a: 377 (Quedius; subgenus Sauridus; characters).

  • — Lohse, 1964: 215 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 278 (Quedius; subgenus Raphirus; all of Alps; Slovenia; France; Italy; Belgium ?; Istria; Bosnia Hercegovina; Montenegro; Turkey; Albania; Macedonia; Romania; Czechoslovakia).

  • — Coiffait, 1978: 209 (Quedius; subgenus Sauridus; characters; Alps; Apennines; Carpathians; Balkans).

  • — Frank, 1982: 47 (Quedius; list of parasites).

  • — Tóth, 1984: 132 (Quedius; characters; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Europe.

kiesenwetteri Bernhauer and Schubert, 1916: 430 (Quedius; variety of ochropterus; [Note: This name was originally cited as an aberration by Ganglbauer, 1895, so was unavailable. The name was made available by Bernhauer and Schubert who cited it as a variety]; Type locality: not cited).

  • — Ganglbauer, 1895: 404 (Quedius; subgenus Sauridus; aberration of ochropterus; [Note: This name is an unavailable name as proposed by Ganglbauer according to Articles 1.3.4, 45.5).

  • — Porta, 1907: 134 (Quedius; aberration of ochropterus).

  • — Gridelli, 1924d: 101 (Quedius; variety of ochropterus).

  • — Scheerpeltz, 1933: 1454 (Quedius; variety of ochropterus).

  • — Smetana, 1958: 367 (Quedius; aberration of ochropterus).

  • — Smetana, 1962b: 142 (Quedius; aberration of ochropterus; characters).

  • — Lohse, 1964: 215 (Quedius; subgenus Raphirus; aberration of ochropterus; characters).

  • — Coiffait, 1978: 209 (Quedius; subgenus Sauridus; variety of ochropterus; characters).

  • — Tóth, 1984: 132 (Quedius; variety of ochropterus; characters).

dorni Korge, 1969: 45 (Quedius; subspecies of ochropterus; Type locality: Südbulgarien: Pamporovo im Rhodopen-Gebirge).

  • — Coiffait, 1978: 209 (Quedius; subgenus Sauridus; subspecies of ochropterus; characters).

ochropygus Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius, cited as Cyrtoquedius; Type locality: Bolivien: Mapiri; Yuracaris).

  • — Scheerpeltz, 1933: 1454 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 175 (Quedius; subgenus Cyrtoquedius; characters in key; Bolivia).

  • Distribution: Bolivia.

oculatus Fauvel, 1895, see: Indoquedius.

oculeus Casey, 1915: 407 (Distichalius; Type locality: British Columbia: Inverness).

  • — Scheerpeltz, 1933: 1454 (Quedius; catalog).

  • — Hatch, 1957: 219 (Quedius; subgenus Distichalius; characters; British Columbia; Washington; Oregon).

  • — Smetana, 1971: 222 (Quedius; subgenus Raphirus; characters; British Columbia; Alaska; California; Oregon; Washington).

  • Distribution: Canada, USA.

sparsus Casey, 1915: 407 (Distichalius; Type locality: California).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • — Smetana, 1971: 222 (Quedius; synonym of oculeus).

ogloblini Bernhauer, 1934b: 217 (Quedius; subgenus Cyrtoquedius, cited as Cryptoquedius; Type locality: Argentinien: Misiones: Loreto).

  • — Scheerpeltz, 1955a: 173 (Quedius; subgenus Cyrtoquedius; cited as oglobini; characters in key; Argentina).

  • Distribution: Argentina.

ohiaensis Broun, 1923: 679 (Quedius; Type locality: Lake Ohia, North Auckland).

  • — Scheerpeltz, 1933: 1454 (Quedius; catalog).

  • Distribution: New Zealand.

okei Korge, 1963a: 84 (Quedius; replacement name for marginalis Oke).

  • Distribution: Australia.

marginalis Oke, 1933: 124 (Quedius; [preoccupied]; Type locality: Australia: Victoria, Belgrave).

ollin Smetana, 1976a: 226 (Quedius; subgenus Quedionuchus; Type locality: Mexico: Guerrero: Chilapa).

  • Distribution: Mexico.

omissus Coiffait, 1977c: 140 (Quedius; subgenus Raphirus; Type locality: Russie méridionale, environs de Maykop).

  • — Coiffait, 1978: 267 (Quedius; subgenus Raphirus; characters; Russia).

  • Distribution: Russia.

onodai Watanabe, 1986: 22 (Quedius; subgenus Microsaurus; Type locality: Japan: Honshu: Yamanashi Pref.: Hirogawara, Mt. Kita-dake).

  • Distribution: Japan.

ophthalmicus Cameron, 1945d: 790 (Quedius; subgenus Raphirus; Type locality: New Zealand: Gollans Valley).

  • Distribution: New Zealand.

optabilis Bernhauer, 1934: 12 (Quedius; subgenus Raphirus; Type locality: Mittel-Szechuan: Hwa-Yin-Shan, 1600 m; [Note: See lectotype designation by Smetana, 1990a]).

  • — Smetana, 1990a: 201 (Quedius; subgenus Raphirus; lectotype designation; characters; China).

  • Distribution: China.

orbiceps Casey, 1915, see: sublimbatus Mäklin, 1853.

orientalis Korge, 1971: 18 (Quedius; subgenus Sauridus; Type locality: Iran: Songhor, ca. 1800 m, Kermanshahan).

  • — Coiffait, 1978: 252 (Quedius; subgenus Sauridus; characters; Iran).

  • Distribution: Iran.

ormanus Fagel, 1971, see: safaensis Fagel, 1968.

ornatipennis Wendeler, 1927: 9 (Quedius; subgenus Raphirus; Type locality: Ost-Java, Kendeng-Gebirge).

  • — Scheerpeltz, 1933: 1454 (Quedius; catalog).

  • Distribution: Indonesia.

ornatocollis Bierig, 1934d: 173 (Quedius; Type locality: Panama: Puerto Armuelles, paz. Kuste, Prov. Chiriqui).

  • Distribution: Panama.

orophilus Drugmand, 1988c: 202 (Quedius; subgenus Raphirus; Type locality: Anatolie occidentale: l'Ulu Dagh, 1800–1900 m).

  • Distribution: Turkey.

ortrudae Korge, 1971a: 51 (Quedius; subgenus Sauridus; Type locality: Pass nördlich Koyulhisar südlich von Ordu, 1800 m, Anatolia bor.).

  • — Coiffait, 1978: 217 (Quedius; subgenus Sauridus; characters; Turkey).

  • Distribution: Turkey.

osellai Bordoni, 1973e: 37 (Quedius; subgenus Sauridus; Type locality: Turchia: Sümela, Trabzon).

  • — Coiffait, 1978: 245 (Quedius; subgenus Sauridus; characters; Turkey).

  • Distribution: Turkey.

othiniensis Johansen, 1907, see: puncticollis Thomson, 1867.

otho Smetana, 1995f: 243 (Quedius; subgenus Microsaurus; Type locality: China: Yunnan prov. Heishui, 35 km N. Lijiang, 27°13′N, 100°19′E).

  • — Smetana, 1999b: 216 (Quedius; subgenus Microsaurus; collecting notes; China).

  • Distribution: China.

ouzbekiscus Coiffait, 1969: 52 (Quedius; subgenus Raphirus; Type locality: Monts Tschingan, environ 1200 m, Ouzbekistan, U.R.S.S.).

  • — Coiffait, 1970d: 143 (Quedius; Uzbekistan).

  • — Coiffait, 1978: 278 (Quedius; subgenus Raphirus; characters; Uzbekistan).

  • Distribution: Uzbekistan.

ovaliceps Mulsant and Rey, 1876, see: nemoralis Baudi, 1848.

ozomatli Smetana, 1975: 316 (Quedius; subgenus Quedionuchus; Type locality: Mexico: Oaxaca: 3 mi. N. Suchixtepec, 9500 ft.).

  • — Smetana, 1976a: 244 (Quedius; subgenus Quedionuchus; Mexico).

  • Distribution: Mexico.

paganettii Bernhauer, 1936g: 308 (Quedius; subgenus Raphirus; Type locality: Albanien: Ljuboten).

  • — Coiffait, 1963a: 379 (Quedius; subgenus Sauridus; characters).

  • — Korge, 1964: 122 (Quedius; subgenus Sauridus; characters; Macedonia).

  • — Coiffait, 1978: 227 (Quedius; subgenus Sauridus; characters; Albania).

  • Distribution: Albania, Macedonia.

pahelo Smetana, 1992b: 5 (Quedius; subgenus Raphirus; Type locality: India: Sikkim, “Dzongri 3970 m Kangchen-dzonga Area”).

  • Distribution: India.

pallens Smetana, 1996f: 128 (Quedius; subgenus Microsaurus; Type locality: China: Kuatun, 2300 m, 27.40 n. Br., 117.40 ö. L.).

  • Distribution: China.

pallipes P. Lucas, 1846: 113 (Quedius; Type locality: Algeria: environs de Philippeville).

  • — Fauvel, 1869: 487 (Quedius; synonym of molochinus).

  • — Fauvel, 1869a: 151 (Quedius; synonym of molochinus).

  • — Fauvel, 1902b: 115 (Quedius; synonym of picipennis “Paykull”).

  • — Porta, 1907: 130 (Quedius; synonym of molochinus).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1457 (Quedius; subspecies of picipennis “Paykull”).

  • — Méquignon, 1937: 3 (Quedius; subspecies of molochinus).

  • — Paulian, 1941: 268 (Quedius; variety of molochinus; larval characters).

  • — Last, 1952a (Quedius; valid species; discussion and characters to separate from molochinus; Britain).

  • — Korge, 1960: 53 (Quedius; characters).

  • — Coiffait, 1961a: 55 (Quedius; characters).

  • — Korge, 1962b: 335 (Quedius; notes; Italy).

  • — Palm, 1963b: 142 (Quedius; subgenus Quedius; characters).

  • — Scheerpeltz, 1964a: 300 (Quedius; subgenus Quedius; Crete).

  • — Scheerpeltz, 1965b: 407 (Quedius; Sicily).

  • — Scheerpeltz, 1965c: 511 (Quedius; Sicily).

  • — Coiffait, 1973a: 275 (Quedius; Morocco).

  • — Bordoni, 1974b: 12 (Quedius; subgenus Quedius; notes; Italy).

  • — Bordoni, 1976b: 100 (Quedius; characters; Italy).

  • — Coiffait, 1978: 196 (Quedius; subgenus Quedius; characters; North Africa; Italy; Sicily; Sardinia; Corsica; Majorca; Albania).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Outerelo, Palmer and Pons, 1995: 80 (Quedius; Mallorca).

  • Distribution: Tunisia, Algeria, Morocco, France, Italy, Greece, Albania.

pallipoides Coiffait, 1961a: 56 (Quedius; mutation of pallipes; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Tunisie: Pont du Fahs. Also cited from Algeria).

  • — Coiffait, 1973a: 275 (Quedius; variety of pallipes; Morocco).

  • — Bordoni, 1976b: 100 (Quedius; synonym of pallipes).

  • — Coiffait, 1978: 196 (Quedius; subgenus Quedius; variety of pallipes; characters).

pallipoides Coiffait, 1961 [Note: Nomen nudum], see: pallipes P. Lucas, 1846.

papuanus Cameron, 1937, see: Heterothops.

paraboops Coiffait, 1975: 34 (Quedius; subgenus Raphirus; Type locality: Litvianska, région sud-ouest du Lac Baïkal).

  • — Smetana, 1976d: 27 (Quedius; subgenus Raphirus; characters; Siberia).

  • — Coiffait, 1978: 279 (Quedius; subgenus Raphirus; characters; central Asia).

  • — Smetana, 1978b: 86 (Quedius; subgenus Raphirus; Siberia).

  • — Drugmand, 1988c: 209 (Quedius; characters).

  • Distribution: Russia.

paradisi Hatch, 1957: 227 (Quedius; subgenus Sauridus; Type locality: Washington: Mt. Rainier, Paradise Park).

  • — Smetana, 1971: 180 (Quedius; subgenus Distichalius; characters; Oregon; Washington).

  • — Smetana, 1971b: 3 (Quedius; subgenus Distichalius; Washington).

  • — Smetana, 1976c: 176 (Quedius; subgenus Distichalius; collecting notes; characters; Oregon; Washington).

  • — Smetana, 1978a: 828 (Quedius; subgenus Distichalius; collecting notes; Oregon).

  • Distribution: USA.

paradisianus Heer, 1839: 278 (Philonthus; Type locality: Paradies an der Zaportalp).

  • — Heer, 1841: 583 (Philonthus; characters; Switzerland).

  • — Heer, 1841: 583 (Philonthus; [Note: Misidentification: Scheerpeltz, 1933: 1456, cited rufipes Heer as a synonym of paradisianus; Heer did not describe the species as new, he attributed it to Gravenhorst]; characters; Switzerland).

  • — Kraatz, 1857d: 513 (Quedius; synonym of monticola).

  • — Fauvel, 1874: 307 [= 1874a: 533] (Quedius; characters; France; Italy; Switzerland; Austria).

  • — Eppelsheim, 1892: 332 (Quedius; Turkestan).

  • — Ganglbauer, 1895: 411 (Quedius; subgenus Raphirus; characters; Alps; Sudeten; Carpathians).

  • — Porta, 1907: 148 (Quedius; subgenus Raphirus; characters; Italy).

  • — Reitter, 1909: 115 (Quedius; subgenus Raphirus; characters; Germany).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; catalog).

  • — Gridelli, 1925: 149 (Quedius; subgenus Raphirus; characters).

  • — Portevin, 1929: 345 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1455 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 53 (Quedius; aedeagal illustration).

  • — Smetana, 1958: 387 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 149 (Quedius; subgenus Raphirus; characters; Europe; Caucasus).

  • — Lohse, 1964: 218 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Korge, 1964: 123 (Quedius; subgenus Raphirus; Macedonia; Turkey).

  • — Horion, 1965: 297 (Quedius; subgenus Raphirus; Turkey; Turkmenistan; Poland; Czechoslovakia; Hungary; Romania; Bosnia Hercegovina; Macedonia; Croatia; Slovenia; Italy; France; Switzerland; Austria; Germany).

  • — Jarrige, 1968b: 74 (Quedius; France).

  • — Coiffait, 1978: 272 (Quedius; subgenus Raphirus; characters; Alps; Carpathians; mountains of central Europe; Pyrenees; Anatolia).

  • — Tóth, 1984: 127 (Quedius; characters; Hungary).

  • — H. Franz, 1987: 70 (Quedius; Cyprus).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France).

  • — Drugmand, 1988c: 204 (Quedius; characters).

  • — Nowosad, 1990: 147 (Quedius; collected from mole nest; notes; Poland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Europe, Caucasus, Turkey.

monticola Erichson, 1840: 544 (Quedius; Type locality: montes Tyrolenses).

  • — Heer, 1841: 583 (Philonthus; synonym of paradisianus).

  • — Märkel and Kiesenwetter, 1848a: 325 (Quedius; notes; Carpathian Alps).

  • — Fairmaire and Laboulbène, 1856: 544 (Quedius; characters; France).

  • — Kraatz, 1857d: 513 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 202 (Quedius; characters; Austria).

  • — Fauvel, 1867c: 112 [= 1868b: 211] (Quedius; synonym of paradisianus).

  • — Fauvel, 1874: 306 [= 1874a: 533] (Quedius; synonym of paradisianus).

  • — Redtenbacher, 1874: 199 (Quedius; characters; Austria).

  • — Mulsant and Rey, 1876: 780 [= 1877: 636] (Quedius; subgenus Raphirus; characters; France).

  • — Ganglbauer, 1895: 411 (Quedius; synonym of paradisianus).

  • — Porta, 1907: 148 (Quedius; synonym of paradisianus).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; synonym of paradisianus).

  • — Gridelli, 1925: 149 (Quedius; synonym of paradisianus).

  • — Scheerpeltz, 1933: 1456 (Quedius; synonym of paradisianus).

  • — Smetana, 1958: 387 (Quedius; synonym of paradisianus).

  • — Coiffait, 1978: 272 (Quedius; synonym of paradisianus).

flavopilus Gerhardt, 1899: 218 (Quedius; variety of paradisianus; Type locality: [Schlesien] Vor- und Hochgebirge).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; variety of paradisianus).

  • — Gridelli, 1925: 149 (Quedius; variety of paradisianus).

  • — Scheerpeltz, 1933: 1456 (Quedius; cited as flavopilosus; variety of paradisianus).

  • — Smetana, 1958: 387 (Quedius; cited as flavopilosus; synonym of paradisianus).

  • — Coiffait, 1978: 272 (Quedius; subgenus Raphirus; variety of paradisianus; cited as flavopilosus; characters).

  • — Tóth, 1984: 127 (Quedius; cited as flavopilosus; variety of paradisianus; characters).

fenestratus Scholz, 1928: 307 (Quedius; aberration of paradisianus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Langenau-Frankental in der Grafschaft Glatz).

  • — Scheerpeltz, 1933: 1456 (Quedius; synonym of paradisianus).

  • — Smetana, 1958: 387 (Quedius; aberration of paradisianus).

  • — Smetana, 1962b: 149 (Quedius; aberration of paradisianus; characters).

  • — Lohse, 1964: 217 (Quedius; subgenus Raphirus; aberration of paradisianus; characters).

  • — Coiffait, 1978: 272 (Quedius; subgenus Raphirus; variety of paradisianus; characters).

  • — Tóth, 1984: 127 (Quedius; variety of paradisianus; characters).

korgeanus Fagel, 1968: 124 (Quedius; subgenus Raphirus; subspecies of paradisianus; Type locality: Anatolie occidentale: Abant Dagh, env. 1450 m).

  • — Coiffait, 1978: 272 (Quedius; subgenus Raphirus; subspecies of paradisianus; characters).

  • — Coiffait, 1978d: 165 (Quedius; subspecies of paradisianus; Turkey).

paralimbatus Coiffait, 1969, see: suturalis Kiesenwetter, 1845.

parallelicollis Scheerpeltz, 1965: 295 (Quedius; subgenus Indoquedius; Type locality: N.E. Burma, Kambaiti, 2000 m).

  • Distribution: Myanmar.

parallelus Hatch, 1957, see: curtipennis Bernhauer, 1908.

paramerus Coiffait, 1967e: 411 (Quedius; subgenus Sauridus; Type locality: environs de Maykop, Russie méridionale).

  • — Coiffait, 1978: 247 (Quedius; subgenus Sauridus; characters; Russia).

  • Distribution: Russia.

parviceps Fauvel, 1871 [Note: Nomen nudum], see: anceps Fairmaire and Laboulbène, 1856.

parviceps Sharp, 1874: 25 (Quedius; Type locality: Japan).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1456 (Quedius; catalog).

  • — Nakane, 1963a: 94 (Quedius; characters; habitus photograph; Japan).

  • — Yuh, Paik, Kwon, and Lee, 1985: 254 (Quedius; Korea).

  • — Cho, 1996b: 113 (Quedius; characters; Korea).

  • Distribution: Japan, Korea.

parvipennis Bernhauer, 1917, see: utenaus Casey, 1915.

paschim Smetana, 1988: 249 (Quedius; subgenus Raphirus; Type locality: Inde: Garhwal, UP, au-dessus Pauri, 1900 m).

  • Distribution: India.

pauliani C. Koch, 1941: 66 (Quedius; subgenus Raphirus; Type locality: Maroc: Djebel M'Goun, Cañon Tessaout, 3000–3200 m).

  • — Coiffait, 1963a: 390 (Quedius; subgenus Sauridus; subspecies of nemoralis; characters).

  • — Fagel, 1968c: 10 (Quedius; subgenus Sauridus; characters; Tunisia; Algeria; Morocco).

  • — Coiffait, 1978: 215 (Quedius; subgenus Sauridus; characters; Morocco; Algeria; Tunisia).

  • Distribution: Morocco, Algeria, Tunisia.

verecundus Normand, 1947a: 9 (Quedius; subspecies of nemoralis [preoccupied]; Type locality: Tunisie: Aïn-Draham; Sakiet. Algeria: Adékar; Constantine; Ouarsénis).

  • — Coiffait, 1978: 215 (Quedius; synonym of pauliani).

pavelmoraveci Štourač, 1998a: 81 (Quedius; subgenus Raphirus; Type locality: Macedonia: Pelister, Mt. Veternica, 2100–2200 m. Also cited from Serbia).

  • Distribution: Macedonia, Yugoslavia.

pecki Smetana, 1975: 318 (Quedius; subgenus Quedionuchus; Type locality: Mexico: Oaxaca: 20 km N. Oaxaca, 9500 ft.).

  • — Smetana, 1976a: 247 (Quedius; subgenus Quedionuchus; Mexico).

  • Distribution: Mexico.

pecoudi Coiffait, 1977c: 135 (Quedius; subgenus Microsaurus; Type locality: Canaries: Ténérife, Agua Mansa).

  • — Coiffait, 1978: 176 (Quedius; subgenus Microsaurus; characters).

  • Distribution: Canary Islands.

pectinator Thomson, 1896, see: brevicornis Thomson, 1860.

pectinatus Lea, 1908: 151 (Quedius; Type locality: King Island).

  • — Scheerpeltz, 1933: 1456 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

pectoralis Boheman, 1858, see: Phucobius.

pediculus Nordmann, 1837: 79 (Philonthus; Type locality: Unalaschka; Amer. boreal.).

  • — Erichson, 1840: 531 (Quedius; characters; Unalaska).

  • — Mannerheim, 1843b: 231 (Quedius; characters; Unalaska).

  • — Gravenhorst, 1847: 234 (Quedius; characters; notes).

  • — Mulsant and Rey, 1876: 681 [= 1877: 537] (Quedius; characters; France).

  • — Horn, 1878: 161 (Quedius; variety of capucinus; characters; Alaska).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; catalog).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; synonym of punctatellus).

  • — Van Dyke, 1924: 18 (Quedius; variety of capucinus; Alaska).

  • — Portevin, 1929: 338 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1456 (Quedius; catalog).

  • — Smetana, 1971: 170 (Quedius; subgenus Distichalius; characters; Alberta; British Columbia; Alaska; California; Oregon; Washington).

  • — Smetana, 1971a: 1845 (Quedius; subgenus Distichalius; collecting notes; California).

  • — Smetana, 1971b: 3 (Quedius; subgenus Distichalius; Unalaska).

  • — Smetana, 1976c: 175 (Quedius; subgenus Distichalius; Oregon; Washington).

  • — Smetana, 1981b: 640 (Quedius; collecting notes; Arizona).

  • Distribution: Canada, USA.

pellax Smetana, 1971: 115 (Quedius; subgenus Microsaurus; Type locality: California: San Diego Co.: San Vicente. Also cited from British Columbia and Oregon).

  • — Smetana, 1971b: 2 (Quedius; subgenus Microsaurus; British Columbia; Washington).

  • — Smetana, 1976: 114 (Quedius; subgenus Microsaurus; characters; Mexico).

  • — Topp and Smetana, 1998 (Quedius; natural history; California; Oregon; British Columbia).

  • Distribution: Canada, USA, Mexico.

peltatus Erichson, 1840, see: fumatus Stephens, 1833.

peneckei Bernhauer, 1918, see: hauseri Bernhauer, 1918.

peraffinis Cameron, 1932, see: beesoni Cameron, 1932.

peranxius Mulsant and Rey, 1876, see: fulgidus Fabricius, 1793.

peregrinus Gravenhorst, 1806: 53 (Staphylinus; Type locality: Carolina).

  • — Erichson, 1840: 550 (Quedius; characters; Carolonia).

  • — Horn, 1878: 160 (Quedius; characters; Canada to the Carolinas).

  • — Blatchley, 1910: 371 (Quedius; characters; Indiana).

  • — Casey, 1915: 408 (Microsaurus; characters; Rhode Island and District of Columbia to Lake Superior).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; catalog).

  • — Fall, 1926: 150 (Quedius; Alaska).

  • — Scheerpeltz, 1933: 1456 (Quedius; catalog).

  • — Smetana, 1971: 56 (Quedius; subgenus Microsaurus; characters; Ontario; Quebec; Connecticut; Florida; Georgia; Illinois; Indiana; Iowa; Kentucky; Maine; Maryland; Massachusetts; Michigan; Missouri; New Hampshire; New Jersey; New York; North Carolina; Ohio; Pennsylvania; Tennessee; Vermont; Virginia; West Virginia).

  • — Smetana, 1971b: 2 (Quedius; subgenus Microsaurus; Missouri; New York).

  • — Smetana, 1973: 1425 (Quedius; Nova Scotia; Quebec).

  • — Legner and Moore, 1977: 176 (Quedius; cited as perigrinus; found under bark).

  • — Frank, 1982: 47 (Quedius; list of parasites).

  • — Smetana, 1990d: 97 (Quedius; Oklahoma).

  • — Dearborn and Donahue, 1994: 15 (Quedius; Maine).

  • — Downie and Arnett, 1996: 390 (Quedius; subgenus Microsaurus; characters; USA).

  • Distribution: Canada, USA.

silvicola Casey, 1884b: 123 (Quedius; Type locality: Fairmount Park, Philadelphia).

  • — Horn, 1885a: 109 (Quedius; synonym of fulgidus).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1456 (Quedius; synonym of peregrinus).

  • — Smetana, 1971: 56 (Quedius; synonym of peregrinus).

perng Smetana, 1995: 71 (Quedius; subgenus Raphirus; Type locality: Taiwan: Nantou Hsien, Yushan N. P., 1.8 km W. Pai-Yun Hut, 3375 m).

  • Distribution: Taiwan.

persicus Korge, 1971: 9 (Quedius; subgenus Microsaurus; Type locality: Iran: Mazanderan: Sardab-rud-Tal bei Rudbarak, 1400–2000 m).

  • — Coiffait, 1978: 178 (Quedius; subgenus Microsaurus; characters; Iran).

  • Distribution: Iran.

persimilis Mulsant and Rey, 1876: 791 [= 1877: 647] (Quedius; subgenus Raphirus; Type locality: les environs de Lyon et les Alpes; [Note: See lectotype designation by Assing, 1999b]).

  • — Ganglbauer, 1895: 412 (Quedius; synonym of boops).

  • — Porta, 1907: 150 (Quedius; synonym of boops).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; synonym of boops).

  • — Gridelli, 1925: 165 (Quedius; synonym of boops).

  • — Fagel, 1960b: 117 (Quedius; notes).

  • — Coiffait, 1963b: 425 (Quedius; subgenus Raphirus; valid species; characters).

  • — Lohse, 1964: 219 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Coiffait, 1978: 278 (Quedius; subgenus Raphirus; characters; France; Belgium; Germany).

  • — Tóth, 1984: 129 (Quedius; characters).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Benelux).

  • — Drugmand, 1993: 170 (Quedius; notes; Luxembourg).

  • — Assing, 1999b: 41 (Quedius; lectotype designation; characters; notes).

  • — Assing, 2001: 75 (Quedius; collected from xerothermous site; Germany).

  • Distribution: Europe.

aridulus Jansson, 1939: 51 (Quedius; subgenus Raphirus; Type locality: G. Sandön; [Note: See lectotype designation by Assing, 1999b]. Originally cited from Inseln Gotland und Öland).

  • — Tottenham, 1948a: 254 (Quedius; subgenus Arphirus; characters; Britain).

  • — V. Hansen, 1952: 157 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Smetana, 1958: 397 (Quedius; subgenus Raphirus; characters; ecology).

  • — Fagel, 1960b: 114 (Quedius; characters).

  • — Smetana, 1962b: 152 (Quedius; subgenus Raphirus; characters; Europe).

  • — Palm, 1963b: 154 (Quedius; subgenus Raphirus; characters; Sweden; Denmark; Norway).

  • — Lohse, 1964: 218 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Smetana, 1964d: 81 (Quedius; subgenus Raphirus; Czechoslovakia).

  • — Horion, 1965: 307 (Quedius; subgenus Raphirus; Britain; Norway; Sweden; Denmark; Holland; Belgium; France; Portugal; Spain; Germany; Austria; Czechoslovakia).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 275 (Quedius; subgenus Raphirus; characters; Europe; Britain; Scandinavia; Iberian Peninsula).

  • — Outerelo, 1980a: 57 (Quedius; Spain).

  • — Tóth, 1984: 131 (Quedius; characters; Hungary).

  • — Drugmand, 1987a: 312 (Quedius; Belgium).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Austria; Benelux; Denmark; Sweden).

  • — Lohse and Lucht, 1989: 175 (Quedius; characters).

  • — Whitehead, 1990e: 11 (Quedius; collecting notes; Britain).

  • — Whitehead, 1991: 5 (Quedius; collecting notes; Britain).

  • — Drugmand, 1993: 170 (Quedius; notes; Luxembourg).

  • — Hodge and Jones, 1995: 41 (Quedius; characters; Britain).

  • — Assing, 1999b: 41 (Quedius; lectotype designation: G. Sandön; synonym of persimilis).

  • — Luff and Owen, 2000: 39 (Quedius; subgenus Raphirus; scutellar variation; separation from Q. boops; Britain).

  • — Burakowski, Mroczkowski, and Stefańska, 2000: 53 (Quedius; subgenus Raphirus; catalog; Poland).

joyi Fagel, 1946: 130 (Quedius; proposed for the misidentification of picipennis by Joy; Type locality: Uccle, vallon d'Engeland; Wemmel, Jauchelette Anderlecht, Neerpede, Schooten).

  • — Fagel, 1960b: 117 (Quedius; characters; Germany; Belgium).

  • — Smetana, 1962b: 153 (Quedius; subgenus Raphirus; characters).

  • — Lohse, 1964: 219 (Quedius; synonym of persimilis).

  • — Coiffait, 1978: 278 (Quedius; synonym of persimilis).

petilius Smetana, 1995g: 242 (Quedius; subgenus Microsaurus; Type locality: China: N. Sichuan prov. Zhangla env., 4200–4700 m).

  • — Smetana, 1999d: 522 (Quedius; subgenus Microsaurus; characters; Sichuan).

  • Distribution: China.

petraensis Fagel, 1968b: 6 (Quedius; subgenus Sauridus; Type locality: Arabie: Pétra).

  • — Coiffait, 1978: 218 (Quedius; subgenus Sauridus; characters; Arabia).

  • Distribution: Jordan.

pharak Smetana, 1988: 269 (Quedius; subgenus Raphirus; Type locality: Nepal: Khandbari District, above Tashigaon, 3500 m).

  • Distribution: Nepal.

phenicius Coiffait, 1967, see: acuminatus Hochhuth, 1849.

philippinus Cameron, 1941c: 382 (Quedius; subgenus Raphirus; Type locality: Philippines: Luzon: Balbalan).

  • — Korge, 1963a: 87 (Quedius; subgenus Indoquedius).

  • Distribution: Philippines.

apicalis Cameron, 1941c: 383 (Quedius; variety of philippinus; Type locality: Philippines: Not cited, presumably Luzon: Balbalan).

philonthoides Lynch, 1884: 120 (Quedius; subgenus Microsaurus; Type locality: Chacabuco).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1456 (Quedius; catalog; Paraguay).

  • Distribution: Argentina, Paraguay.

picatus Stephens, 1832, see: Philonthus.

piceolineatus Scheerpeltz, 1965: 284 (Quedius; subgenus Microsaurus; Type locality: N.E. Burma, Kambaiti, 2000 m).

  • Distribution: Myanmar.

piceolus Fauvel, 1878g: 554 (Quedius; Type locality: Australie méridionale: Victoria).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

picicollis Stephens, 1832, see: Philonthus.

picicornis Stephens, 1832, see: fuliginosus Gravenhorst, 1802.

picicornis Gridelli, 1922, see: gridellii Scheerpeltz, 1933.

piciformis Bernhauer, 1912e: 177 (Quedius; Type locality: Argentinien: Prov. Buenos Aires).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; catalog).

  • — Scheerpeltz, 1972: 25 (Quedius; subgenus Apoquedius; Argentina).

  • Distribution: Argentina.

picipennis: Paykull, 1800 [Note: Nomen nudum; Paykull attributed picipennis to Fabricius but Paykull's misidentification has been cited as a valid species in Quedius; citations to picipennis Paykull are listed under molochinus; picipennis Fabricius is in Pseudocypus.

picipennis Stephens, 1833, see: molochinus Gravenhorst, 1806.

picipennis Heer, 1839, see: fulvicollis Stephens, 1833.

picipes Mannerheim, 1830: 26 (Staphylinus; Type locality: Corfu).

  • — Mannerheim, 1831: 440 (Staphylinus; Corfu).

  • — Nordmann, 1837: 77 (Philonthus; characters; Corfu).

  • — Erichson, 1839a: 491 (Quedius; characters; Germany).

  • — Stephens, 1839: 390 (Quedius; characters; England).

  • — Erichson, 1840: 537 (Quedius; characters; Germany; Austria; Corcyra).

  • — P. Lucas, 1846: 114 (Quedius; notes; Algeria).

  • — Gravenhorst, 1847: 226 (Quedius; characters; notes).

  • — Redtenbacher, 1849: 710 (Philonthus; characters; Austria).

  • — Hardy, 1851: 33 (Quedius; notes; England).

  • — Fairmaire and Laboulbène, 1856: 543 (Quedius; characters; France).

  • — Kraatz, 1857d: 505 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 204 (Quedius; characters; Austria).

  • — Kraatz, 1858a: 58 (Quedius; Greece).

  • — Hochhuth, 1862: 46 (Quedius; Russia).

  • — Thomson, 1867: 172 (Raphirus; characters; Scandinavia).

  • — Thomson, 1867a: 43 (Raphirus; characters; Scandinavia).

  • — Fauvel, 1874: 292 [= 1874a: 518] (Quedius; characters; France; Algeria).

  • — Redtenbacher, 1874: 201 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 265 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 704 [= 1877: 560] (Quedius; subgenus Sauridus; characters; France).

  • — Fowler, 1888: 239 (Quedius; subgenus Sauridus; characters; Britain).

  • — Ganglbauer, 1895: 404 (Quedius; subgenus Sauridus; characters; middle Europe; Algeria).

  • — Porta, 1907: 134 (Quedius; subgenus Sauridus; characters; Italy).

  • — Reitter, 1909: 112 (Quedius; subgenus Sauridus; characters; Germany).

  • — Johansen, 1914: 364 (Quedius; subgenus Sauridus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; catalog).

  • — Munster, 1923: 197 (Quedius; subgenus Sauridus; Fennoscandia).

  • — Gridelli, 1924d: 108 (Quedius; subgenus Raphirus; characters; Italy; Macedonia; Bosnia).

  • — Portevin, 1929: 342 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1457 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 30 (Quedius; aedeagal illustration).

  • — Horion, 1954: 10 (Quedius; Germany).

  • — Smetana, 1958: 369 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1961d: 134 (Quedius; subgenus Raphirus; Italy).

  • — Smetana, 1962b: 143 (Quedius; subgenus Raphirus; characters).

  • — Coiffait, 1963a: 378 (Quedius; subgenus Sauridus; characters).

  • — Palm, 1963b: 146 (Quedius; subgenus Raphirus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 214 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 280 (Quedius; subgenus Raphirus; Ireland; Britain; Norway; Sweden; Finland; Denmark; Latvia; all central Europe; Spain; Corsica; Sardinia; Italy; Dalmatia; Serbia; Bosnia; Albania; Macedonia; Greece; Ukraine; Germany; Austria).

  • — Pototskaia, 1967: 85, 88 (Quedius; larval characters).

  • — Kasule, 1968a: 67 (Quedius; life history).

  • — Frank, 1969: 263 (Quedius; larval characters; biological notes; England).

  • — Kasule, 1970: 61 (Quedius; larval characters).

  • — Bordoni, 1974b: 13 (Quedius; subgenus Sauridus; Italy).

  • — Osella and Zanetti, 1975: 121 (Quedius; collected from nest of Talpa europaea; notes; Italy).

  • — Pototskaia, 1976: 173 (Quedius; larval characters).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 211 (Quedius; subgenus Sauridus; characters; Europe; Britain; Scandinavia).

  • — H. Franz, 1984: 59 (Quedius; Macedonia).

  • — Tóth, 1984: 133 (Quedius; characters).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Schülke and Uhlig, 1988: 12 (Quedius; Germany).

  • — Nowosad, 1990: 146 (Quedius; collected from mole nest; notes; Poland).

  • — Welch, 1993: 229 (Quedius; ovariole number and ovary structure).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Owen, 2000: 252 (Quedius; pitfall trap collections from underground at roots of trees; Britain).

  • Distribution: Algeria, Europe, Russia.

seminitidus Stephens, 1832: 220 (Quedius; Type locality: Suffolk).

  • — Stephens, 1839: 391 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 23 (Quedius; synonym of picipes).

pyrrhopus Stephens, 1832: 216 (Quedius; Type locality: London; Norfolk; Suffolk; Yorkshire).

  • — Curtis, 1837: pl. 638 (Quedius; characters; Britain).

  • — Hardy, 1851: 33 (Quedius; synonym of picipes).

  • — G. Waterhouse, 1858: 23 (Quedius; synonym of picipes).

  • — Porta, 1907: 134 (Quedius; synonym of picipes).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; synonym of picipes).

  • — Gridelli, 1924d: 108 (Quedius; synonym of picipes).

  • — Smetana, 1958: 371 (Quedius; synonym of picipes).

variicolor Nordmann, 1837: 75 (Philonthus; Type locality: Gallia meridionalis).

  • — Erichson, 1840: 537 (Quedius; synonym of picipes).

  • — P. Lucas, 1846: 114 (Quedius; synonym of picipes).

  • — Kraatz, 1857d: 505 (Quedius; synonym of picipes).

  • — Ganglbauer, 1895: 404 (Quedius; synonym of picipes).

  • — Porta, 1907: 135 (Quedius; synonym of picipes).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; synonym of picipes).

  • — Gridelli, 1924d: 108 (Quedius; synonym of picipes).

  • — Smetana, 1958: 371 (Quedius; synonym of picipes).

gracilicornis Heer, 1839: 274 (Philonthus; Type locality: alpes Helvetiae occidentalis).

  • — Hardy, 1851: 33 (Quedius; synonym of picipes).

  • — Fauvel, 1874: 292 [= 1874a: 518] (Quedius; synonym of picipes).

  • — Ganglbauer, 1895: 404 (Quedius; synonym of picipes).

  • — Porta, 1907: 135 (Quedius; synonym of picipes).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; synonym of picipes).

  • — Gridelli, 1924d: 108 (Quedius; synonym of picipes).

  • — Smetana, 1958: 371 (Quedius; synonym of picipes).

provincialis Mulsant and Rey, 1876: 707 [= 1877: 563] (Quedius; Type locality: Provence).

  • — Porta, 1907: 135 (Quedius; synonym of picipes).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; synonym of picipes).

  • — Gridelli, 1924d: 108 (Quedius; synonym of picipes).

  • — Smetana, 1958: 371 (Quedius; synonym of picipes).

picipes Stephens, 1832, see: Philonthus.

pictipennis Blackburn, 1892: 21 (Quedius; Type locality: Victoria; Dandenong Ranges).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; catalog).

  • — Lea, 1925: 243 (Quedius; characters; Australia).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

pignerator Lea, 1925: 250 (Quedius; Type locality: New South Wales: National Park).

  • — Scheerpeltz, 1933: 1458 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

pinalicus Casey, 1915, see: desertus Horn, 1878.

pineti Brisout, 1866: 359 (Quedius; Type locality: Espagne: Las Navas, près l'Escorial).

  • — Fauvel, 1874: 295 [= 1874a: 521] (Quedius; synonym of praecox).

  • — Ganglbauer, 1895: 407 (Quedius; synonym of nigriceps).

  • — Porta, 1907: 137 (Quedius; synonym of nigriceps).

  • — Bernhauer and Schubert, 1916: 429 (Quedius; synonym of nigriceps).

  • — Scheerpeltz, 1933: 1451 (Quedius; synonym of nigriceps).

  • — Jeannel and Jarrige, 1949: 378 (Quedius; subgenus Microsaurus; cave species; Spain).

  • — Jarrige, 1952a: 134 (Quedius; subgenus Raphirus; characters; notes; Spain; Portugal).

  • — Smetana, 1958: 373 (Quedius; synonym of nigriceps).

  • — Coiffait, 1963a: 381 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1973a: 275 (Quedius; Morocco).

  • — Coiffait, 1978: 242 (Quedius; subgenus Sauridus; characters; Iberian Peninsula).

  • Distribution: Morocco, Spain, Portugal.

nevesi Scheerpeltz, 1951: 130 (Quedius; subgenus Raphirus; Type locality: Portugal: Santos Ferreira Neves bei Agrela, Prov. Douro).

  • — Coiffait, 1963a: 381 (Quedius; subgenus Sauridus; synonym of pineti).

  • — Coiffait, 1978: 242 (Quedius; synonym of pineti).

placidus Cameron, 1932: 282 (Quedius; subgenus Ediquus; Type locality: Darjeeling Dist.: Ghum, Rongdong Valley; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1458 (Quedius; catalog).

  • — Smetana, 1977a: 246 (Quedius; Bhutan).

  • — Smetana, 1988: 210 (Quedius; subgenus Microsaurus; lectotype designation; characters; Bhutan; India).

  • — Smetana, 1992b: 2 (Quedius; subgenus Microsaurus; Nepal).

  • Distribution: India, Nepal, Bhutan.

plagiatus Hochhuth, 1849, see: vulneratus Gemminger and Harold, 1868.

plagiatus Mannerheim, 1843b: 231 (Quedius; Type locality: insula Sitkha; [Note: See lectotype designation by Smetana, 1971]).

  • — Mäklin, 1852: 314 (Quedius; characters; Sitka).

  • — Fauvel, 1876a: 77 [= 1876: 262] (Quedius; synonym of laevigatus).

  • — Horn, 1878: 162 (Quedius; synonym of laevigatus).

  • — Ganglbauer, 1895: 402 (Quedius; synonym of laevigatus).

  • — Porta, 1907: 108 (Quedius; synonym of laevigatus).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; synonym of laevigatus).

  • — Tottenham, 1949: 422 (Quedius; checklist; Britain).

  • — Smetana, 1958: 359 (Quedius; subgenus Quedionuchus; cited as valid species; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 160 (Quedius; subgenus Quedionuchus; characters).

  • — Lohse, 1964: 210 (Quedius; subgenus Quedionuchus; characters; central Europe).

  • — Korge, 1964: 118 (Quedius; subgenus Quedionuchus; characters).

  • — Pototskaia, 1967: 82, 87 (Quedius; larval characters).

  • — Smetana, 1967d: 216 (Quedius; synonym of laevigatus).

  • — Szujecki, 1968a: 735 (Quedius; Poland).

  • — Scheerpeltz, 1968a: 91 (Quedius; synonym of laevigatus).

  • — Szujecki, 1969a: 258 (Quedius; notes; Poland).

  • — Smetana, 1971: 147 (Quedius; lectotype designation; synonym of laevigatus).

  • — Bordoni, 1973b: 45 (Quedius; synonym of laevigatus).

  • — Shilov, 1975a: 377 [= 1975b: 99] (Quedius; Komi region of Russia).

  • — Pototskaia, 1976: 173 (Quedius; larval characters).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 189 (Quedius; subgenus Quedionuchus; characters; Palaearctic region).

  • — Muona, 1979: 15 (Quedius; Finland; Russia; Sweden; Norway; Baltic States).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 132 (Quedius; catalog; Poland).

  • — Tóth, 1984: 105 (Quedius; characters; Hungary).

  • — Muona and Viramo, 1986: 14 (Quedius; Finland).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Sweden).

  • — Smetana, 1990d: 99 (Quedius; valid name).

  • — Silfverberg, 1992: 18 (Quedius; Finland; Russia; Sweden; Norway; Estonia; Latvia).

  • — Smetana, 1993b: 50 (Quedius; subgenus Quedionuchus; Czech Republic; Slovakia).

  • — Siitonen, 1993: 230 (Quedius; collecting notes; Finland).

  • — Øland, 1995: 23 (Quedius; host mushroom; Norway).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Staniec, 1996: 114 (Quedius; pupal characters; Poland).

  • — Ádám, 1996: 247 (Quedionuchus; synonym of glaber).

  • — Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 39 (Quedius; Latvia).

  • — Anderson, Nash, and O'Connor, 1997: 20 (Quedius; Ireland).

  • — Outerelo, Gamarra, and Salgado, 1998: 130 (Quedius; subgenus Quedionuchus; collected from cave; Spain).

  • — Staniec, 1999a: 55 (Quedius; characters of pupa; Poland).

  • — Fowles, Alexander, and Key, 1999: 129 (Quedius; saproxylic species; status as endangered or threatened species; Britain).

  • — Ryabukhin, 1999: 73 (Quedius; subgenus Quedionuchus; catalog; habitat; distribution; northeastern Russia).

  • — Hågvar, 1999: 28 (Quedius; collected on Fomes fomentarius; Norway).

  • — Olberg and Andersen, 2000: 223 (Quedius; attracted to trunk-window traps baited with Fomes fomentarius and Phellinus spp.; Norway).

  • — Herman, 2001: 30 (Quedius; prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.3).

  • Distribution: Europe, Russia, Mongolia, Canada, USA.

glaber O. Müller, 1776: 98 (Staphylinus; [Note: This name is older than plagiatus]; Type locality: Dania).

  • — Goeze, 1777: 729 (Staphylinus; characters).

  • — Gmelin, 1790: 2034 (Staphylinus; characters).

  • — Turton, 1802: 518 (Staphylinus; characters; Europe).

  • — Erichson, 1840: 529 (Quedius; synonym of laevigatus).

  • — Kraatz, 1857d: 497 (Quedius; synonym of laevigatus).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; synonym of laevigatus).

  • — Smetana, 1958: 359 (Quedius; synonym of plagiatus).

  • — Bordoni, 1973b: 45 (Quedius; synonym of laevigatus).

  • — Coiffait, 1978: 189 (Quedius; synonym of plagiatus).

  • — Ádám, 1996: 247 (Quedionuchus; valid name; Hungary).

  • — Herman, 2001: 30 (Quedius; synonym of plagiatus).

flavopterus Geoffroy, 1785: 166 (Staphylinus; [Note: This name is older than plagiatus]; Type locality: Paris).

  • — Villers, 1789: 421 (Staphylinus; characters; France).

  • — Olivier, 1795:(42): 27 (Staphylinus; characters; France).

  • — Nordmann, 1837: 89 (Philonthus; characters).

  • — Erichson, 1840: 529 (Quedius; synonym of laevigatus).

  • — Kraatz, 1857d: 497 (Quedius; synonym of laevigatus).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; synonym of laevigatus).

  • — Smetana, 1958: 359 (Quedius; synonym of plagiatus).

  • — Coiffait, 1978: 189 (Quedius; synonym of plagiatus).

  • — Herman, 2001: 30 (Quedius; synonym of plagiatus).

laevigatus Gyllenhal, 1810: 306 (Staphylinus; [preoccupied]; Type locality: Suecia).

  • — Zetterstedt, 1828: 71 (Staphylinus; characters; Lapland).

  • — C. Sahlberg, 1830: 316 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 26 (Staphylinus; Finland; Russia).

  • — Mannerheim, 1831: 440 (Staphylinus; Finland; Russia).

  • — Lacordaire, 1835: 380 (Emus; characters; France).

  • — Nordmann, 1837: 79 (Philonthus; Sweden; Germany; Podolia).

  • — Zetterstedt, 1838: 61 (Staphylinus; characters; habitat; Lapland).

  • — Erichson, 1839a: 488 (Quedius; characters; Germany).

  • — Heer, 1839: 274 (Philonthus; characters; Switzerland).

  • — Erichson, 1840: 529 (Quedius; characters; Sweden; Germany; Lapland).

  • — Gravenhorst, 1847: 233 (Quedius; characters; notes).

  • — Redtenbacher, 1849: 709 (Philonthus; characters; Austria).

  • — Küster, 1853a: 61 (Quedius; characters; Europe).

  • — Fairmaire and Laboulbène, 1856: 536 (Quedius; characters; France).

  • — Kraatz, 1857d: 497 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 201 (Quedius; characters; Austria).

  • — Thomson, 1860: 176 (Microsaurus; characters; Scandinavia).

  • — Fauvel, 1865a: 292 [= 1865b: 42] (Quedius; habitat; France).

  • — Solsky, 1871b: 240 (Quedius; Russia).

  • — Fauvel, 1874: 286 [= 1874a: 512] (Quedius; characters; France; Lapland; Scandinavia; Britain; Germany; Switzerland; Austria; Italy; Russia; Siberia).

  • — Redtenbacher, 1874: 198 (Quedius; characters; Austria).

  • — Fauvel, 1876a: 77 [= 1876: 262] (Quedius; Alaska).

  • — Seidlitz, 1875: 267 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 673 [= 1877: 529] (Quedius; characters; France).

  • — J. Sahlberg, 1876: 28 (Microsaurus; Finland).

  • — Horn, 1878: 162 (Quedius; characters; Alaska to Oregon, Kansas, Canada, and Pennsylvania).

  • — Sharp, 1884: 336 (Quedionuchus).

  • — Fowler, 1888: 245 (Quedionuchus; characters; Britain).

  • — Ganglbauer, 1895: 402 (Quedius; subgenus Quedionuchus; characters; north and middle Europe; Siberia; North America).

  • — Porta, 1907: 108 (Quedius; subgenus Quedionuchus; characters; Italy).

  • — Fall and Cockerell, 1907: 166 (Quedius; New Mexico).

  • — Reitter, 1909: 111 (Quedius; subgenus Quedionuchus; characters; Germany).

  • — Blatchley, 1910: 372 (Quedius; characters; Indiana).

  • — Johansen, 1914: 362 (Quedius; subgenus Quedionuchus; characters; Denmark).

  • — Eichelbaum, 1915a: 91 (s; abdominal segmentation; apical segements).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Munster, 1923: 194 (Quedius; subgenus Quedionuchus; Fennoscandia).

  • — Gridelli, 1924d: 74 (Quedius; subgenus Quedionuchus; characters).

  • — Van Dyke, 1924: 17 (Quedius; Alaska).

  • — Fall, 1926: 150 (Quedius; Alaska).

  • — Portevin, 1929: 339 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1445 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 20 (Quedius; aedeagal illustration).

  • — Jeannel and Jarrige, 1949: 378 (Quedius; subgenus Quedionuchus; cave species; Czechoslovakia).

  • — V. Hansen, 1952: 140 (Quedius; subgenus Quedionuchus; characters; Denmark).

  • — Hatch, 1957: 229 (Quedius; subgenus Quedionuchus; characters; British Columbia; Washington; Oregon).

  • — Smetana, 1958: 359 (Quedius; synonym of plagiatus).

  • — Smetana, 1960e: 160 (Quedius; synonym of plagiatus).

  • — Palm, 1963b: 138 (Quedius; subgenus Quedionuchus; characters; Sweden; Norway; Finland).

  • — Lohse, 1964: 210 (Quedius; synonym of plagiatus).

  • — Smetana, 1965: 41 (Quedius; subgenus Quedionuchus; Newfoundland).

  • — Horion, 1965: 270 (Quedius; subgenus Quedionuchus, cited as Quedionychus; Britain; Norway; Sweden; Finland; Russia; Denmark; Baltics; Poland; Czechoslovakia; Hungary; Romania; Slovenia; Croatia; Bosnia Hercegovina; Italy; France; Switzerland; Austria; Germany).

  • — Smetana, 1967d: 216 (Quedius; subgenus Microsaurus; Mongolia).

  • — Kasule, 1970: 59 (Quedionuchus; larval characters).

  • — Pope, 1977: 31 (Quedius; synonym of plagiatus).

  • — Smetana, 1971: 147 (Quedius; subgenus Quedionuchus; cited as valid species; characters; Alberta; British Columbia; Labrador; Manitoba; Newfoundland; Northewst Territoires; Ontario; Quebec; Saskatchewan; Yukon Territory; Alaska; Arizona; California; Colorado; Connecticut; Georgia; Idaho; Illinois; Iowa; Maine; Maryland; Massachusetts; Michigan; Montana; Nevada; New Hampshire; New Jersey; New Mexico; New York; North Carolina; Ohio; Oregon; Pennsylvania; South Dakota; Utah; Vermont; Washington; Wisconsin; Wyoming).

  • — Smetana, 1971a: 1845 (Quedius; subgenus Quedionuchus; collecting notes; New Mexico).

  • — Smetana, 1971b: 3 (Quedius; subgenus Quedionuchus; New York).

  • — Bordoni, 1973b: 45 (Quedius; subgenus Quedionuchus; cited as valid species; characters; notes; Italy).

  • — Legner and Moore, 1977: 176 (Quedius; found under bark).

  • — Coiffait, 1978: 189 (Quedius; synonym of plagiatus).

  • — Hoebeke and Kovarik, 1988: 217 (Quedius; parasitized by serphid; collecting notes; notes on parasite; Ohio).

  • — Smetana, 1990d: 99 (Quedius; synonym of plagiatus).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; synonym of plagiatus).

  • — Downie and Arnett, 1996: 392 (Quedius; subgenus Quedionuchus; characters; USA).

laevipennis Dufour, 1843: 33 (Staphylinus; Type locality: vallée d'Ossau).

  • — Gemminger and Harold, 1868: 570 (Quedius; synonym of laevigatus).

  • — Fauvel, 1874: 286 [= 1874a: 512] (Quedius; synonym of laevigatus).

  • — Fowler, 1888: 245 (Quedionuchus; synonym of laevigatus).

  • — Ganglbauer, 1895: 402 (Quedius; synonym of laevigatus).

  • — Porta, 1907: 108 (Quedius; synonym of laevigatus).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; synonym of laevigatus).

  • — Smetana, 1958: 359 (Quedius; synonym of plagiatus).

  • — Bordoni, 1973b: 45 (Quedius; synonym of laevigatus).

  • — Coiffait, 1978: 189 (Quedius; synonym of plagiatus).

longipennis Mannerheim, 1846: 509 (Quedius; Type locality: insula Unalaschka; [Note: See lectotype designation by Smetana, 1971]).

  • — Fauvel, 1874: 286 [= 1874a: 512] (Quedius; synonym of laevigatus).

  • — Fauvel, 1876a: 77 [= 1876: 262] (Quedius; synonym of laevigatus).

  • — Horn, 1878: 162 (Quedius; synonym of laevigatus).

  • — Fowler, 1888: 245 (Quedionuchus; synonym of laevigatus).

  • — Porta, 1907: 108 (Quedius; synonym of laevigatus).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; synonym of laevigatus).

  • — Scheerpeltz, 1933: 1445 (Quedius; synonym of laevigatus; valid species?).

  • — Smetana, 1965: 40 (Quedius; subgenus Quedionuchus; cited as valid species; characters; notes; Newfoundland).

  • — Smetana, 1971: 147 (Quedius; lectotype designation; synonym of laevigatus).

  • — Bordoni, 1973b: 45 (Quedius; synonym of laevigatus).

rufipennis Mäklin, 1853: 189 (Quedius; [preoccupied]; Type locality: peninsula Kenai).

  • — Horn, 1878: 162 (Quedius; synonym of laevigatus).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; synonym of laevigatus).

  • — Smetana, 1958: 359 (Quedius; synonym of plagiatus).

  • — Smetana, 1971: 147 (Quedius; synonym of laevigatus).

resplendens Thomson, 1860: 176 (Microsaurus; Type locality: Lappland).

  • — Fauvel, 1874: 286 [= 1874a: 512] (Quedius; synonym of laevigatus).

  • — Ganglbauer, 1895: 402 (Quedius; variety of laevigatus).

  • — Porta, 1907: 108 (Quedius; aberration of laevigatus).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; aberration of laevigatus).

  • — Gridelli, 1924d: 74 (Quedius; aberration of laevigatus).

  • — Scheerpeltz, 1933: 1445 (Quedius; aberration of laevigatus).

  • — Smetana, 1958: 359 (Quedius; aberration of plagiatus).

  • — Smetana, 1960e: 160 (Quedius; aberration of plagiatus; characters).

  • — Smetana, 1971: 147 (Quedius; synonym of laevigatus).

  • — Bordoni, 1973b: 45 (Quedius; synonym of laevigatus).

  • — Coiffait, 1978: 189 (Quedius; subgenus Quedionuchus; variety of plagiatus; characters).

  • — Tóth, 1984: 105 (Quedius; aberration of plagiatus; characters).

ater J. Sahlberg, 1880: 72 (Microsaurus; [preoccupied]; Type locality: Jenisseisk).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; aberration of laevigatus).

  • — Gridelli, 1924d: 74 (Quedius; aberration of laevigatus).

  • — Smetana, 1958: 359 (Quedius; aberration of plagiatus).

  • — Smetana, 1960e: 160 (Quedius; aberration of plagiatus; characters).

  • — Smetana, 1971: 148 (Quedius; synonym of laevigatus).

  • — Coiffait, 1978: 189 (Quedius; subgenus Quedionuchus; variety of plagiatus; characters).

  • — Tóth, 1984: 105 (Quedius; aberration of plagiatus; characters).

alpinus Fiori, 1894: 87 (Quedius; [preoccupied]; Type locality: Alpi carniche, Visdende).

  • — Porta, 1907: 108 (Quedius; synonym of laevigatus).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; synonym of laevigatus).

  • — Gridelli, 1924d: 74 (Quedius; synonym of laevigatus).

  • — Smetana, 1958: 359 (Quedius; synonym of plagiatus).

  • — Bordoni, 1973b: 45 (Quedius; synonym of laevigatus).

  • — Coiffait, 1978: 189 (Quedius; synonym of plagiatus variety resplendens).

plagifer Eppelsheim, 1889, see: vulneratus Gemminger and Harold, 1868.

planatus Sharp, 1889: 34 (Quedionuchus; Type locality: Nikko).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; catalog).

  • Distribution: Japan.

plancus Erichson, 1840: 538 (Quedius; Type locality: Sardinia).

  • — Kraatz, 1857d: 500 (Quedius; Sardinia).

  • — Redtenbacher, 1857: 203 (Quedius; characters; Austria).

  • — Redtenbacher, 1874: 200 (Quedius; characters; Austria).

  • — Ganglbauer, 1895: 408 (Quedius; subgenus Sauridus; characters; France; Pyrenees; Sardinia; Algeria).

  • — Fauvel, 1902b: 113 (Quedius; Algeria; Tunisia).

  • — Sainte-Claire Deville, 1906: 103 (Quedius; Corsica).

  • — Porta, 1907: 142 (Quedius; subgenus Sauridus; characters; Italy; Sardinia; Corsica).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; catalog).

  • — Gridelli, 1925: 146 (Quedius; subgenus Raphirus; characters; Pyrenees; Alps; Caucasus; Turkey; France).

  • — Scheerpeltz, 1933: 1458 (Quedius; catalog).

  • — Jeannel and Jarrige, 1949: 380 (Quedius; subgenus Raphirus; cave species; Spain).

  • — Smetana, 1962b: 147 (Quedius; subgenus Raphirus; characters; Europe).

  • — Coiffait, 1963a: 373 (Quedius; subgenus Quediops).

  • — Lohse, 1964: 212 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 294 (Quedius; subgenus Raphirus; Algeria; Tunisia; France; Sardinia; Greece; Belgium; Britain; Germany ?).

  • — Bordoni, 1973g: 122 (Quedius; subgenus Quediops; characters; Sardinia; Italy; Corsica).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 202 (Quedius; subgenus Quediops; characters; France; Italy; Corsica; Sardinia; Iberian Peninsula; North Africa).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Benelux).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Gerend and Braunert, 1997: 200 (Quedius; notes; Luxembourg).

  • Distribution: Tunisia, Algeria, Europe, Turkey.

kraatzii Brisout, 1859: CCXXXI (Quedius; Type locality: Collioure).

  • — Fauvel, 1874: 272 [= 1874a: 498] (Quedius; characters; France; Austria).

  • — Fauvel, 1876a: 75 [= 1876: 260] (Quedius; Corsica; Spain).

  • — Mulsant and Rey, 1876: 750 [= 1877: 606] (Quedius; subgenus Sauridus; characters; France).

  • — Fauvel, 1878: 127 [= 1878a: 47] (Quedius; Algeria).

  • — Fauvel, 1886: 57 [= 1886a: 49] (Quedius; Algeria).

  • — Ganglbauer, 1895: 408 (Quedius; synonym of plancus).

  • — Fauvel, 1902b: 113 (Quedius; synonym of plancus).

  • — Eppelsheim, 1894b: 12 (Quedius; synonym of plancus).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; synonym of plancus).

  • — Gridelli, 1925: 146 (Quedius; synonym of plancus).

  • — Portevin, 1929: 338 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1458 (Quedius; synonym of plancus).

  • — Smetana, 1962b: 147 (Quedius; synonym of plancus).

  • — Bordoni, 1973g: 122 (Quedius; synonym of plancus).

  • — Coiffait, 1978: 202 (Quedius; synonym of plancus).

eppelsheimi Quedenfeldt, 1882: 181 (Quedius; Type locality: Hispania mer., Algeciras in Andalusia).

  • — Quedenfeldt, 1884: 372 (Quedius; characters; Spain).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1439 (Quedius; catalog).

  • — Coiffait, 1967e: 418 (Quedius; synonym of plancus).

  • — Bordoni, 1973g: 122 (Quedius; synonym of plancus).

  • — Coiffait, 1978: 202 (Quedius; synonym of plancus).

planus Hatch, 1957: 225 (Quedius; subgenus Microsaurus; Type locality: Oregon: Crater L.).

  • — Smetana, 1971: 72 (Quedius; subgenus Microsaurus; characters; California; Oregon).

  • — Smetana, 1971a: 1839 (Quedius; subgenus Microsaurus; collecting notes; California).

  • — Smetana, 1976c: 169 (Quedius; subgenus Microsaurus; California).

  • — Smetana, 1978a: 824 (Quedius; subgenus Microsaurus; collecting notes; Oregon).

  • Distribution: USA.

platyphthalmus Roubal, 1955, see: nitipennis Stephens, 1833.

pluvialis Smetana, 1998b: 99 (Quedius; subgenus Raphirus; Type locality: China: Sichuan Emei Shan, Leidongping, 2500 m, 29°32′N 103°21′E).

  • Distribution: China.

poggii Coiffait, 1972a: 80 (Quedius; subgenus Sauridus; Type locality: Torre Aquila, Feglino, Prov. de Savone, Italie).

  • — Coiffait, 1978: 231 (Quedius; subgenus Sauridus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Herman, 2001: 12 (Quedius; spelling corrected).

  • Distribution: Italy.

politulus W. J. MacLeay, 1873: 140 (Philonthus; Type locality: Gayndah).

  • — Fauvel, 1877: 272 (Quedius; characters; Australia).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; catalog).

  • Distribution: Australia.

polystigma Wankowicz, 1867, see: tenellus Gravenhorst, 1806.

poneli Coiffait, 1984d: 313 (Quedius; subgenus Sauridus; Type locality: Maroc: Grand Atlas, environs d'Oukaïmden, 2260 m).

  • Distribution: Morocco.

ponteuxinus Coiffait, 1970: 100 (Quedius; subgenus Sauridus; Type locality: Maslen Nos Strandja, littoral de la Mer Noire, au Sud-Est de Burgaz, Bulgarie).

  • — Coiffait, 1978: 220 (Quedius; subgenus Sauridus; characters; Bulgaria).

  • Distribution: Bulgaria.

ponticus Korge, 1964, see: limbatus Heer, 1839.

potockajae Coiffait, 1967e: 414 (Quedius; subgenus Sauridus; Type locality: environs de Maykop, Russie méridionale).

  • — Coiffait, 1978: 256 (Quedius; subgenus Sauridus; characters; Russia).

  • Distribution: Russia.

praecox Gravenhorst, 1802: 172 (Staphylinus; Type locality: Lusitania).

  • — Latreille, 1804: 324 (Staphylinus; characters; Portugal).

  • — Gravenhorst, 1806: 57 (Staphylinus).

  • — Curtis, 1829: 26 (Philonthus; catalog; Britain).

  • — C. Sahlberg, 1830: 318 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 27 (Staphylinus; Finland).

  • — Mannerheim, 1831: 441 (Staphylinus; Finland).

  • — Stephens, 1833: 245 (Raphirus; characters; England).

  • — Nordmann, 1837: 78 (Philonthus; Europe).

  • — Zetterstedt, 1838: 61 (Staphylinus; characters; habitat; Lapland).

  • — Heer, 1839: 278 (Philonthus; characters; Switzerland).

  • — Erichson, 1840: 540 (Quedius; characters; Portugal; Sardinia).

  • — Laporte, 1840: 177 (Staphylinus; characters; France).

  • — P. Lucas, 1846: 114 (Quedius; synonym of maurorufus).

  • — Rosenhauer, 1856: 71 (Quedius; P. S. Maria).

  • — Fairmaire and Laboulbène, 1856: 542 (Quedius; characters; France).

  • — Kraatz, 1857d: 507 (Quedius; characters).

  • — Redtenbacher, 1857: 204 (Quedius; characters; Austria).

  • — Peyron, 1858: 428 (Quedius; Turkey).

  • — Redtenbacher, 1874: 201 (Quedius; characters; Austria).

  • — Fauvel, 1876a: 78 [= 1876: 263] (Quedius; Portugal; Sardinia; Sicily).

  • — Fauvel, 1878: 129 [= 1878a: 49] (Quedius; Algeria).

  • — Fauvel, 1886: 59 [= 1886a: 51] (Quedius; Algeria).

  • — Fauvel, 1902b: 116 (Quedius; Morocco; Algeria; Tunisia).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; catalog).

  • — Gridelli, 1922b: 127, 138 (Quedius; subgenus Sauridus; characters).

  • — Gridelli, 1925: 129 (Quedius; subgenus Raphirus; characters).

  • — Portevin, 1929: 343 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1458 (Quedius; catalog).

  • — C. Koch, 1941: 65 (Quedius; Morocco).

  • — Coiffait, 1978: 251 (Quedius; subgenus Sauridus; characters; North Africa; Iberian Peninsula; Sicily; Sardinia; Pyrenees).

  • — Sparacio, 1995: 166 (Quedius; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Outerelo, Gamarra, and Salgado, 1998: 130 (Quedius; subgenus Sauridus; collected from cave; Spain; Portugal).

  • Distribution: Tunisia, Algeria, Morocco, Europe.

ernestini Fauvel, 1872b: 34 (Quedius; Type locality: Oran; Bône).

  • — Fauvel, 1874: 296 [= 1874a: 522] (Quedius; characters; Algeria).

  • — Fauvel, 1876a: 78 [= 1876: 263] (Quedius; synonym of praecox).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; synonym of praecox).

  • — Gridelli, 1925: 129 (Quedius; synonym of praecox).

  • — Coiffait, 1978: 252 (Quedius; synonym of praecox).

praeditus Sharp, 1889, see: Indoquedius.

pretiosus Sharp, 1874: 26 (Quedius; Type locality: Nagasaki).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1458 (Quedius; catalog).

  • — Smetana, 1998c: 317 (Quedius; subgenus Distichalius; lectotype designation; characters; China).

  • Distribution: Japan.

problematicus Fagel, 1968b: 9 (Quedius; subgenus Sauridus; Type locality: Liban: Kartaba, 1200 m).

  • — Coiffait, 1978: 219 (Quedius; subgenus Sauridus; characters; Lebanon).

  • — Smetana, 1978: 124 (Quedius; subgenus Raphirus; collecting notes; Israel).

  • Distribution: Lebanon, Israel.

probus Casey, 1915: 417 (Raphirus; Type locality: California: Lake Tahoe; [Note: See lectotype designation by Smetana, 1971]).

  • — Scheerpeltz, 1933: 1458 (Quedius; catalog).

  • — Smetana, 1971: 213 (Quedius; subgenus Raphirus; lectotype designation; characters; California; Oregon; Washington).

  • — Smetana, 1971a: 1847 (Quedius; subgenus Raphirus; collecting notes; California).

  • — Smetana, 1976c: 181 (Quedius; subgenus Raphirus; Washington).

  • Distribution: USA.

prominens Bernhauer, 1917, see: Rolla.

prostans Horn, 1878: 165 (Quedius; Type locality: California; [Note: See lectotype designation by Smetana, 1971]. Also cited from Vancouver).

  • — Casey, 1915: 418 (Raphirus; characters; California: Sta. Clara to Siskiyou Co., and at Lake Tahoe; Vancouver Island).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1458 (Quedius; catalog).

  • — Hatch, 1957: 229 (Quedius; subgenus Raphirus; characters; British Columbia; Washington; Oregon; Idaho).

  • — Smetana, 1971: 207 (Quedius; subgenus Raphirus; lectotype designation: Cal. [California]; characters; British Columbia; California; Colorado; Idaho; Montana; Nevada; Oregon; Utah; Washington).

  • — Smetana, 1971a: 1847 (Quedius; subgenus Raphirus; collecting notes; California).

  • — Smetana, 1978a: 830 (Quedius; subgenus Raphirus; collecting notes; Arizona).

  • Distribution: Canada, USA.

rupimontis Casey, 1915: 418 (Raphirus; Type locality: Colorado: Boulder Co.).

  • — Scheerpeltz, 1933: 1460 (Quedius; catalog).

  • — Smetana, 1971: 207 (Quedius; synonym of prostans).

protensus Sharp, 1884: 335 (Quedius; Type locality: Guatemala, Dueñas).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; catalog).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1933: 1458 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 175 (Quedius; subgenus Cyrtoquedius; characters in key; Guatemala).

  • Distribution: Guatemala.

provincialis Mulsant and Rey, 1876, see: picipes Mannerheim, 1830.

proximus Kraatz, 1858, see: semiaeneus Stephens, 1833.

przewalskii Reitter, 1887: 211 (Quedius; subgenus Ediquus; Type locality: Quellengebiet des Gelben Flusses, 13,500–14,000 F.).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; catalog).

  • — Gridelli, 1924d: 71 (Quedius; subgenus Microsaurus; characters).

  • — Scheerpeltz, 1933: 1458 (Quedius; catalog).

  • — Boháč, 1988: 554 [= 1989: 37] (Quedius; subgenus Microsaurus; cited as przewalskyi; characters; China).

  • — Smetana, 1999d: 521, 524 (Quedius; subgenus Microsaurus; characters; notes; Tibet; Qinghai; Yunnan).

  • Distribution: China.

lamus Smetana, 1995g: 239 (Quedius; subgenus Microsaurus; Type locality: China: N. Sichuan prov. 60 km S. of Hongyuan, 4200 m).

  • — Smetana, 1999d: 521 (Quedius; synonym of przewalskii).

pseudassecla Coiffait, 1963b: 423 (Quedius; subgenus Microsaurus; Type locality: Nord de l'Italie: Piémont, Valle Strona).

  • — Gridelli, 1924d: 48 (Quedius; subgenus Microsaurus; [Note: Misidentified as assecla according to Coiffait, 1963b: 423]; characters; Italy).

  • — Gridelli, 1929: 34 (Quedius; [Note: Misidentified as assecla according to Coiffait, 1963b: 423]; characters).

  • — Smetana, 1960e: 157 (Quedius; subgenus Microsaurus; [Note: Misidentified as assecla according to Coiffait, 1963b: 423]; characters).

  • — Johnson, 1967a: 80 (Quedius; synonym of invreae).

  • — Coiffait, 1967e: 394 (Quedius; synonym of invreae).

  • — Bordoni, 1974: 29 (Quedius; synonym of invreae).

  • — Coiffait, 1977c: 133 (Quedius; notes; characters; valid species).

  • — Pope, 1977: 31 (Quedius; Gridelli's 1924 use of assecla cited as synonym of invreae).

  • — Coiffait, 1978: 161 (Quedius; subgenus Microsaurus; characters; Italy).

  • Distribution: Italy.

pseudolimbatus A. Strand, 1938: 79 (Quedius; Type locality: Rostasee in Målselv in Nord-Norwegen).

  • — Coiffait, 1963a: 394 (Quedius; subgenus Sauridus; characters).

  • — Palm, 1963b: 149 (Quedius; subgenus Raphirus; characters; Sweden; Norway).

  • — Coiffait, 1967e: 417 (Quedius; subgenus Sauridus; notes).

  • — Coiffait, 1978: 256 (Quedius; subgenus Sauridus; characters; Scandinavia).

  • Distribution: Norway.

pseudonigriceps Reitter, 1909: 113 (Quedius; subgenus Sauridus; variety of humeralis; Type locality: Herzegowina; Türkei; Kleinasien).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; variety of humeralis).

  • — Gridelli, 1922b: 128, 134 (Quedius; subgenus Sauridus; characters; notes; Istria; Hercegovina; Turkey).

  • — Gridelli, 1925: 137 (Quedius; subgenus Raphirus; characters).

  • — Scheerpeltz, 1933: 1458 (Quedius; catalog).

  • — Coiffait, 1963a: 386 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1967e: 409 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1970e: 63 (Quedius; Taygete).

  • — Coiffait, 1978: 248 (Quedius; subgenus Sauridus; characters; Asia Minor; Hercegovina; Balkans).

  • — Tóth, 1984: 140 (Quedius; characters).

  • Distribution: Greece, Croatia, Bosnia Hercegovina, Turkey.

pseudopyrenaeus Coiffait, 1967e: 396 (Quedius; subgenus Sauridus; cited as peudopyrenaeus on page 396 and as pseudopyrenaeus in the captions to the illustration on page 398; [Note: Name emended by Coiffait, 1978: 224]; Type locality: île de Krk, Veglia, Yougoslavie).

  • — Coiffait, 1978: 224 (Quedius; subgenus Sauridus; emendation of original spelling; characters; Dalmatia; Bosnia; Serbia).

  • — Tóth, 1984: 135 (Quedius; characters).

  • Distribution: Yugoslavia, Bosnia Hercegovina, Croatia.

pseudoumbrinus Lohse, 1958, see: umbrinus Erichson, 1839.

puetzi Smetana, 1998b: 106 (Quedius; subgenus Raphirus; Type locality: China: Shaanxi, Qin Ling Shan, 110.06 E, 34.25 N, N Hua Shan Mt., S.-top, 1950–2000 m).

  • Distribution: China.

pugetanus Casey, 1915, see: fulvicollis Stephens, 1833.

pugetensis Hatch, 1957, see: transparens Motschulsky, 1845.

pullmani Hatch, 1957: 222 (Quedius; subgenus Microsaurus; Type locality: Washington: Pullman).

  • — Smetana, 1971: 86 (Quedius; subgenus Microsaurus; characters; Alberta; British Columbia; Manitoba; Alaska; Colorado; Montana).

  • — Smetana, 1973: 1426 (Quedius; California).

  • — Smetana, 1976c: 170 (Quedius; subgenus Microsaurus; Saskatchewan).

  • Distribution: Canada, USA.

punctatellus Heer, 1839: 275 (Philonthus; Type locality: Mühlebach u. Krauchthalalp Ct. Glarus; Gemmi; Urserenthal beim St. Annagletscher; Gotthard; Susten; Engelbergerjoch; Luckmanier; Nufeneralpen; Walliseralpen).

  • — Kiesenwetter, 1848: 51 (Quedius; characters; Germany; Switzerland).

  • — Märkel and Kiesenwetter, 1848a: 324 (Quedius; notes; Carpathian Alps).

  • — Redtenbacher, 1849: 826 (Philonthus; characters; Austria).

  • — Kiesenwetter, 1851a: 419 (Quedius; Pyrenees).

  • — Fairmaire and Laboulbène, 1856: 536 (Quedius; characters; France).

  • — Kraatz, 1857d: 498 (Quedius; characters; Switzerland; Austria).

  • — Redtenbacher, 1857: 201 (Quedius; characters; Austria).

  • — Fauvel, 1865a: 292 [= 1865b: 42] (Quedius; habitat; France).

  • — Fauvel, 1869: 487 (Quedius; synonym of pediculus).

  • — Fauvel, 1874: 287 [= 1874a: 513] (Quedius; [Note: Misidentification: Bernhauer and Schubert, 1916: 432, cited pediculus Fauvel as a synonym of punctatellus; Fauvel did not describe the species as new, he attributed it to Nordmann]; characters; France; Alps; Germany; Switzerland; Austria; Italy; Caucasus).

  • — Fauvel, 1874: 287 [= 1874a: 513] (Quedius; synonym of pediculus).

  • — Redtenbacher, 1874: 198 (Quedius; characters; Austria).

  • — Eppelsheim, 1892: 331 (Quedius; Turkmenistan).

  • — Ganglbauer, 1895: 402 (Quedius; subgenus Quedionuchus; characters; Pyrenees; Alps; Carpathians; Caucasus).

  • — Porta, 1907: 106 (Quedius; subgenus Quedionuchus; characters; Italy).

  • — Reitter, 1909: 111 (Quedius; subgenus Quedionuchus; characters; Germany).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; catalog).

  • — Gridelli, 1924d: 76 (Quedius; subgenus Quedionuchus; characters; Caucasus; Pyrenees; Alps; Carpathians).

  • — Bernhauer, 1926g: 2 (Quedius; Kamchatka region).

  • — Scheerpeltz, 1933: 1458 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 19 (Quedius; aedeagal illustration).

  • — Smetana, 1958: 359 (Quedius; subgenus Quedionuchus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 160 (Quedius; subgenus Quedionuchus; characters).

  • — Lohse, 1964: 210 (Quedius; subgenus Quedionuchus; characters; central Europe).

  • — Korge, 1964: 119 (Quedius; subgenus Quedionuchus; characters).

  • — Horion, 1965: 269 (Quedius; subgenus Quedionuchus, cited as Quedionychus; Alps; Carpathians; central France; Pyrenees; Germany).

  • — Smetana, 1966: 331 (Quedius; subgenus Quedionuchus; notes; Czechoslovakia).

  • — Bordoni, 1973b: 51 (Quedius; subgenus Distichalius; characters; Italy).

  • — Coiffait, 1978: 188 (Quedius; subgenus Distichalius; characters; Alps; Carpathians; mountains of central Europe).

  • — Tóth, 1984: 104 (Quedius; characters).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; France).

  • — Lohse and Lucht, 1989: 172 (Quedius; subgenus Distichalius).

  • — Nowosad, 1990: 143 (Quedius; collected from mole nest; notes; Poland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Distichalius; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Europe.

puncticeps Horn, 1878: 166 (Quedius; Type locality: Vancouver; [Note: See lectotype designation by Smetana, 1971]).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1458 (Quedius; catalog).

  • — Hatch, 1957: 215 (Quedius; subgenus Paraquedius; characters; British Columbia; Washington; Oregon).

  • — Smetana, 1971: 235 (Quedius; subgenus Paraquedius; lectotype designation; characters; British Columbia; California; Oregon; Washington).

  • — Smetana, 1981b: 643 (Quedius; California).

  • Distribution: Canada, USA.

puncticollis Broun, 1894, see: Philonthus.

puncticollis Thomson, 1867: 164 (Microsaurus; Type locality: Bastad i nordvestra Skåne).

  • — Thomson, 1867a: 43 (Microsaurus; characters; Scandinavia).

  • — Fauvel, 1874: 278 [= 1874a: 504] (Quedius; characters; Sweden).

  • — Fowler, 1888: 233 (Quedius; characters; Britain).

  • — Ganglbauer, 1895: 398 (Quedius; synonym of ochripennis).

  • — Joy, 1906a: 200 (Quedius; [Note: Misidentification: Bernhauer and Schubert, 1916: 432, cited vexans Joy as a synonym of puncticollis; Joy did not describe the species as new, he attributed it to Eppelsheim]; characters; notes mole's nest; Britain).

  • — Porta, 1907: 113 (Quedius; synonym of ochripennis).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; catalog).

  • — Gridelli, 1924d: 49 (Quedius; synonym of othiniensis).

  • — Scheerpeltz, 1933: 1455 (Quedius; synonym of othiniensis).

  • — V. Hansen, 1952: 129 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Smetana, 1958: 348 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 156 (Quedius; subgenus Microsaurus; characters).

  • — Palm, 1963b: 132 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark; Finland).

  • — Lohse, 1964: 210 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Horion, 1965: 260 (Quedius; subgenus Microsaurus; Britain; Denmark; Sweden; Finland; Holland; Belgium; France; Germany; Austria; Switzerland; Italy; Czechoslovakia; Poland; Ukraine).

  • — Scheerpeltz, 1968a: 90 (Quedius; catalog; Austria).

  • — Szujecki, 1969a: 258 (Quedius; notes; Poland).

  • — Bolov, 1969: 514 (Quedius; collecting notes; Russia).

  • — Bolov, 1969a: 329 (Quedius; collecting notes; Russia).

  • — Bordoni, 1974: 26 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Osella and Zanetti, 1975: 118 (Quedius; collected from nest of Talpa europaea; notes; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 162 (Quedius; subgenus Microsaurus; characters; Europe; Scandinavia; Britain).

  • — Muona, 1979: 15 (Quedius; Russia; Sweden; Denmark).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 127 (Quedius; catalog; Poland).

  • — Frank, 1982: 47 (Quedius; list of parasites).

  • — Chemini and Zanetti, 1982: 216 (Quedius; habitat notes; Italy).

  • — Tóth, 1984: 116 (Quedius; characters; Hungary).

  • — Zanetti, 1987a (Quedius; allometric variation of head).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; France; Benelux; Denmark; Sweden).

  • — Nowosad, 1990: 131 (Quedius; collected from mole nest; notes; Poland).

  • — Silfverberg, 1992: 18 (Quedius; Finland; Russia; Sweden; Denmark).

  • — Welch, 1993: 229 (Quedius; ovariole number and ovary structure).

  • — Smetana, 1993b: 50 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — M. Hansen, 1996: 107 (Quedius; checklist; Denmark).

  • — Gerend and Braunert, 1997: 200 (Quedius; notes; Luxembourg).

  • — Anderson, Nash, and O'Connor, 1997: 20 (Quedius; Ireland).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (Quedius; Denmark).

  • — Herman, 2001: 30 (Quedius; nomen protectum).

  • Distribution: Europe, Russia.

variabilis Gyllenhal, 1810: 303 (Staphylinus; [Note: This name is older than puncticollis; Bernhauer and Schubert, 1916: 428, 432, cited variabilis Gyllenhal as a synonym of mesomelinus and puncticollis]; Type locality: Suecia).

  • — Zetterstedt, 1828: 71 (Staphylinus; characters; Lapland).

  • — Mannerheim, 1830: 25 (Staphylinus; Finland; Sweden; Russia).

  • — Mannerheim, 1831: 439 (Staphylinus; Finland; Sweden; Russia).

  • — Zetterstedt, 1838: 61 (Staphylinus; characters; habitat; Lapland).

  • — Heer, 1839: 273 (Philonthus; characters; Switzerland).

  • — Gravenhorst, 1847: 214 (Quedius; characters).

  • — Hardy, 1851: 31 (Quedius; synonym of fulgidus).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — Thomson, 1860: 175 (Microsaurus; synonym of fulgidus).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; synonym of puncticollis).

  • — Gridelli, 1924d: 49 (Quedius; synonym of othiniensis).

  • — Scheerpeltz, 1933: 1455 (Quedius; synonym of othiniensis).

  • — Smetana, 1958: 348 (Quedius; synonym of puncticollis).

  • — Herman, 2001: 30 (Quedius; nomen oblitum).

castaneipennis Hochhuth, 1872: 131 (Quedius; variety of fulgidus; Type locality: Not cited, but presumably from Gouvernements Kiew and/or Volhynien).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; variety of fulgidus).

  • — Cameron, 1932: 288 (Quedius; synonym of fulgidus).

  • — Scheerpeltz, 1933: 1455 (Quedius; variety of othiniensis).

  • — Smetana, 1958: 348 (Quedius; synonym of puncticollis).

othiniensis Johansen, 1907: 171, 173 (Quedius; Type locality: Odense).

  • — Johansen, 1914: 356 (Quedius; subgenus Microsaurus; cited as “nov. spec.”; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; synonym of puncticollis).

  • — Gridelli, 1924d: 49 (Quedius; subgenus Microsaurus; characters; England; France; Germany; Denmark).

  • — Gridelli, 1929: 30 (Quedius; characters).

  • — Scheerpeltz, 1933: 1454 (Quedius; cited as valid species; catalog).

  • — Wüsthoff, 1938a: fig. 8 (Quedius; aedeagal illustration).

  • — Roubal, 1941b: 121, 127 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Smetana, 1958: 348 (Quedius; synonym of puncticollis).

  • — Smetana, 1960e: 156 (Quedius; synonym of puncticollis).

  • — Lohse, 1964: 210 (Quedius; synonym of puncticollis).

  • — Bordoni, 1974: 26 (Quedius; synonym of puncticollis).

  • — Pope, 1977: 31 (Quedius; synonym of puncticollis).

  • — Coiffait, 1978: 162 (Quedius; synonym of puncticollis).

heidenreichi Heinemann, 1910: 163 (Quedius; Type locality: Braunschweig).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; synonym of puncticollis).

  • — Gridelli, 1924d: 49 (Quedius; synonym of othiniensis).

  • — Scheerpeltz, 1933: 1454 (Quedius; synonym of othiniensis).

  • — Smetana, 1958: 348 (Quedius; synonym of puncticollis).

  • — Bordoni, 1974: 26 (Quedius; synonym of puncticollis).

  • — Coiffait, 1978: 162 (Quedius; synonym of puncticollis).

talparum Sainte-Claire Deville, 1910: 158 (Quedius; Type locality: Gallia borealis et orientalis; Anglia).

  • — Johansen, 1914: 356 (Quedius; synonym of othiniensis).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; synonym of puncticollis).

  • — Gridelli, 1924d: 49 (Quedius; synonym of othiniensis).

  • — Scheerpeltz, 1933: 1455 (Quedius; synonym of othiniensis).

  • — Paulian, 1941: 260 (Quedius; larval characters).

  • — Smetana, 1958: 348 (Quedius; synonym of puncticollis).

  • — Smetana, 1960e: 156 (Quedius; synonym of puncticollis).

  • — Bordoni, 1974: 26 (Quedius; synonym of puncticollis).

  • — Pope, 1977: 31 (Quedius; synonym of puncticollis).

  • — Coiffait, 1978: 162 (Quedius; synonym of puncticollis).

heidenreichi Bernhauer, 1914e: 35 (Quedius; [preoccupied]; Type locality: Dessau).

  • — Gridelli, 1924d: 49 (Quedius; synonym of othiniensis).

  • — Scheerpeltz, 1933: 1455 (Quedius; synonym of othiniensis).

  • — Smetana, 1958: 348 (Quedius; synonym of puncticollis).

  • — Smetana, 1960e: 156 (Quedius; synonym of puncticollis).

  • — Bordoni, 1974: 26 (Quedius; synonym of puncticollis).

melium Walles, 1938: 68, 70 (Quedius; Type locality: Polen: Spas [Bezirk Dolina]).

  • — Roubal, 1941b: 122, 127 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Nowosad and Smetana, 1990: 828 (Quedius; notes; synonym of puncticollis).

bohemicus Roubal, 1941b: 121, 131 (Quedius; subgenus Microsaurus; variety of othiniensis; Type locality: Not cited).

  • — Smetana, 1958: 348 (Quedius; synonym of puncticollis).

  • — Smetana, 1960e: 156 (Quedius; aberration of puncticollis; characters).

  • — Bordoni, 1974: 26 (Quedius; synonym of puncticollis).

  • — Coiffait, 1978: 162 (Quedius; subgenus Microsaurus; variety of puncticollis; characters).

  • — Tóth, 1984: 116 (Quedius; aberration of puncticollis; characters).

punctifrons Fauvel, 1886: 59 [= 1886a: 51] (Quedius; Type locality: L'Edough; Bône).

  • — Fauvel, 1902b: 117 (Quedius; Algeria; Tunisia).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1459 (Quedius; catalog).

  • — Jarrige, 1952a: 135 (Quedius; subgenus Raphirus; characters; Algeria).

  • — Coiffait, 1978: 202 (Quedius; subgenus Quediops; characters; North Africa).

  • Distribution: Algeria, Tunisia.

puniceipennis Solsky, 1868, see: Quediomacrus.

puthzi Korge, 1964a: 179 (Quedius; subgenus Raphirus; Type locality: Theresienfeld, Niederösterreich).

  • Distribution: Austria.

pygmaeus Gravenhorst, 1806, see: scitus Gravenhorst, 1806.

pyrenaeicola Coiffait, 1963, see: obscuripennis Bernhauer, 1901.

pyrenaeus Brisout, 1863: 33 (Quedius; Type locality: Cambredaze).

  • — Fauvel, 1874: 302 [= 1874a: 528] (Quedius; characters; France).

  • — Fauvel, 1876a: 78 [= 1876: 263] (Quedius; Spain).

  • — Mulsant and Rey, 1876: 760 [= 1877: 616] (Quedius; subgenus Raphirus; characters; France).

  • — Ganglbauer, 1895: 409 (Quedius; subgenus Raphirus; characters; Pyrenees; eastern Carpathians; Balkans).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; catalog).

  • — Gridelli, 1925: 139 (Quedius; subgenus Raphirus; characters; Pyrenees).

  • — Portevin, 1929: 344 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1459 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 44 (Quedius; aedeagal illustration).

  • — Jeannel and Jarrige, 1949: 380 (Quedius; subgenus Raphirus; cave species; France; Spain).

  • — Coiffait, 1963a: 381 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 223 (Quedius; subgenus Sauridus; characters; Pyrenees).

  • Distribution: Spain, France.

pyropterus Eppelsheim, 1880, see: maurus C. Sahlberg, 1830.

pyrrhopus Stephens, 1832, see: picipes Mannerheim, 1830.

quadriceps Casey, 1915, see: spelaeus Horn, 1871.

quadriceps Fauvel, 1900d: 232 (Quedius; Type locality: Syrie: Akbes).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; catalog).

  • — Gridelli, 1924d: 73 (Quedius; subgenus Microsaurus; characters; Syria).

  • — Scheerpeltz, 1933: 1459 (Quedius; catalog).

  • — Coiffait, 1977c: 134 (Quedius; notes).

  • — Coiffait, 1978: 163 (Quedius; subgenus Microsaurus; characters; Syria).

  • Distribution: Syria, Lebanon.

rambouseki Gridelli, 1929: 37, 42 (Quedius; Type locality: Siria: Akbes).

  • — Scheerpeltz, 1933: 1459 (Quedius; catalog).

  • — Coiffait, 1977c: 134 (Quedius; synonym of quadriceps).

  • — Coiffait, 1978: 162 (Quedius; synonym of quadriceps).

beryticus Coiffait, 1977c: 135 (Quedius; variety of quadriceps; [Note: Nomen nudum (Article 15.2)]; cited from Beyrouth).

  • — Coiffait, 1978: 163 (Quedius; subgenus Microsaurus; variety of quadriceps; characters).

quadripunctatus Zetterstedt, 1828, see: nitipennis Stephens, 1833.

quadripunctatus Thomson, 1867, see: fulgidus Fabricius, 1793.

quadripunctatus Iablokov-Khnzorian, 1961, see: tetrapunctatus Coiffait, 1977.

quadripunctus Cameron, 1945, see: quadripunctus Bernhauer, 1941.

quadripunctus Bernhauer, 1941b: 32 (Quedius; subgenus Sauridus; Type locality: Neu Seeland: Howard).

  • — Herman, 2001: 31 (Quedius; synonymic note).

  • Distribution: New Zealand.

quadripunctus Cameron, 1945d: 788 (Quedius; subgenus Microsaurus; [preoccupied]; Type locality: Not cited, stated to be without locality label; presumably species is from New Zealand).

  • — Herman, 2001: 31 (Quedius; synonym of quadripunctus).

deficiens Cameron, 1945d: 788 (Quedius; subgenus Microsaurus; variety of quadripunctus; Type locality: Korokoro).

queinneci Coiffait, 1983, see: stevensi Cameron, 1932.

quinctius Smetana, 1998c: 323 (Quedius; subgenus Distichalius; Type locality: China: Beijing, Xiaolongmen).

  • Distribution: China.

quiris Smetana, 1998c: 326 (Quedius; subgenus Distichalius; Type locality: China: Gansu, Dallijia Shan, 48 km w. Linxia, 2980 m).

  • Distribution: China.

quisquilius Gravenhorst, 1847 [Note: Nomen nudum], see: microps Gravenhorst, 1847.

rabirius Smetana, 1998c: 321 (Quedius; subgenus Distichalius; Type locality: China: Zehjiang Tienmushan).

  • Distribution: China.

ragusai Eppelsheim, 1889e: 89 (Quedius; cited as ragusae; Type locality: Not cited).

  • — Porta, 1907: 112 (Quedius; subgenus Microsaurus; cited as ragusai; characters; Sicily).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; catalog).

  • — Gridelli, 1924d: 41 (Quedius; subgenus Microsaurus; characters; Sicily).

  • — Scheerpeltz, 1933: 1459 (Quedius; catalog).

  • — Bordoni, 1974: 69 (Quedius; subgenus Microsaurus; cited as ragusai; [Note: Correction of incorrect original spelling of patronym based on Enrico Ragusa]; characters; Italy).

  • — Coiffait, 1978: 147 (Quedius; subgenus Microsaurus; characters; Sicily).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Italy.

rainieri Hatch, 1957: 222 (Quedius; subgenus Microsaurus; Type locality: Washington: Mt. Rainier, Paradise Park. Also cited from Oregon).

  • — Smetana, 1971: 89 (Quedius; subgenus Microsaurus; characters; British Columbia; Oregon; Washington).

  • — Smetana, 1976c: 170 (Quedius; subgenus Microsaurus; Oregon; Washington).

  • — Legner and Moore, 1977: 176 (Quedius; found under bark).

  • Distribution: Canada, USA.

rambouseki Gridelli, 1929, see: quadriceps Fauvel, 1900.

ramiroi Scheerpeltz, 1951: 136 (Quedius; subgenus Raphirus; Type locality: Portugal: Umgebung Peña maior in Paços de Ferreira, Prov. Douro).

  • — Coiffait, 1963a: 386 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 211 (Quedius; subgenus Sauridus; characters; Portugal).

  • Distribution: Portugal.

raphiroides Hatch, 1957, see: fulvicollis Stephens, 1833.

reboudi Fairmaire, 1877 [Note: Nomen nudum], see: abietum Kiesenwetter, 1858.

recticeps Broun, 1917: 373 (Quedius; Type locality: Mistake Basin, Canterbury).

  • — Scheerpeltz, 1933: 1459 (Quedius; catalog).

  • Distribution: New Zealand.

recticollis Scheerpeltz, 1965: 294 (Quedius; subgenus Indoquedius; Type locality: N.E. Burma, Kambaiti, 2000 m).

  • Distribution: Myanmar.

regularis Bernhauer and Schubert, 1916: 432 (Quedius; replacement name for seriatus Eppelsheim).

  • — Gridelli, 1924d: 77 (Quedius; subgenus Quedionuchus; characters; China).

  • — Scheerpeltz, 1933: 1459 (Quedius; catalog).

  • — Smetana, 1998c: 315 (Quedius; subgenus Distichalius; characters; China).

  • Distribution: China.

seriatus Eppelsheim, 1889a: 169 (Quedius; subgenus Microsaurus; [preoccupied]; Type locality: Amdo).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; synonym of regularis).

  • — Smetana, 1998c: 315 (Quedius; synonym of regularis).

reitteri Gridelli, 1925: 165, 166 (Quedius; variety of boops; proposed for aberration fallaciosus as misidentified by Reitter, 1909: 115; Type locality: Silesia; Carniola; Mte. Bjtoraj, Croazia).

  • — Scheerpeltz, 1933: 1434 (Quedius; variety of boops).

  • — Korge, 1964: 125 (Quedius; subgenus Raphirus; valid species; characters; Turkey).

  • — Horion, 1965: 308 (Quedius; subgenus Raphirus; Turkey; Britain; Germany; Austria; Germany; Slovenia; Croatia; Italy; Bosnia).

  • — Korge, 1971a: 56 (Quedius; subgenus Raphirus; notes; Turkey).

  • — Coiffait, 1978: 282 (Quedius; subgenus Raphirus; characters; Europe; Alps; Italy).

  • — Tóth, 1984: 129 (Quedius; characters).

  • — Drugmand, 1988c: 209 (Quedius; characters).

  • — Lohse and Lucht, 1989: 175 (Quedius; characters).

  • — Wittwer, 1993: 251 (Quedius; Switzerland).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Europe, Turkey.

crius Tottenham, 1948a: 258 (Quedius; subgenus Arphirus; Type locality: Britain: Haxey).

  • — Smetana, 1958: 398 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 153 (Quedius; synonym of asturicus).

  • — Lohse, 1964: 219 (Quedius; subgenus Raphirus; subspecies of asturicus; characters; central Europe).

  • — Pope, 1977: 31 (Quedius; possibly synonym of boops).

  • — Coiffait, 1978: 281 (Quedius; subgenus Raphirus; subspecies of asturicus; characters).

  • — Lucht, 1987: 110 (Quedius; subspecies of asturicus; checklist; Germany; Poland; Austria).

  • — Lohse and Lucht, 1989: 175 (Quedius; synonym of reitteri).

reitteri Bernhauer, 1933, see: bernhauerianus Korge, 1971.

reitteri Bernhauer, 1933, see: reitterianus Bernhauer, 1934.

reitterianus Bernhauer, 1934: 12 (Quedius; replacement name for reitteri Bernhauer, 1933: 39).

  • Distribution: China.

reitteri Bernhauer, 1933: 39 (Quedius; subgenus Quedionochus; [preoccupied]; Type locality: Szetschwan: Tatsienlu Tjiji, Urwald-Rodungen).

  • — Bernhauer, 1934: 12 (Quedius; synonym of reitterianus).

reitterianus Coiffait, 1967, see: edmundi Coiffait, 1969.

repens Smetana, 1981b: 636 (Quedius; subgenus Microsaurus; Type locality: Oregon: Lane Co., Harbor Vista Co. Pk., 2.5 mi. N., 1 mi. W. Florence).

  • Distribution: USA.

resplendens Thomson, 1860, see: plagiatus Mannerheim, 1843.

reynesii† Oustalet, 1874: 172 (Quedius; Type locality: Aix en Provence).

  • — Scudder, 1891: 575 (Quedius; catalog; Ligurian; France).

  • — Handlirsch, 1907: 723 (Quedius; catalog; Oligocene; France).

  • Distribution: France.

rhinton Smetana, 1998c: 324 (Quedius; subgenus Distichalius; Type locality: China: Sichuan, Langmusi, 3500–3600 m).

  • Distribution: China.

rhodicus Coiffait, 1976b: 92 (Quedius; subgenus Sauridus; Type locality: Rhodes: epta Pighès).

  • — Coiffait, 1978: 217 (Quedius; subgenus Sauridus; characters; Rhodes).

  • Distribution: Greece.

richteri Korge, 1966a: 60 (Quedius; subgenus Sauridus; Type locality: Waldschutzgebiet, “Gellmersdorfer Forst” bei Stolpe/Oder, zwischen Oderberg und Schwedt).

  • — Coiffait, 1978: 253 (Quedius; subgenus Sauridus; characters; Europe).

  • — Tóth, 1984: 138 (Quedius; characters).

  • — Lohse and Lucht, 1989: 173 (Quedius; characters; Germany).

  • Distribution: Germany.

riparius F. Kellner, 1843: 31 (Quedius; Type locality: Thüringer-Wald).

  • — Kraatz, 1857d: 518 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 203 (Quedius; characters; Austria).

  • — Redtenbacher, 1874: 200 (Quedius; characters; Austria).

  • — Fauvel, 1874: 272 [= 1874a: 498] (Quedius; characters; France; Germany; Austria; Italy).

  • — Mulsant and Rey, 1876: 747 [= 1877: 603] (Quedius; subgenus Sauridus; characters; France).

  • — Ganglbauer, 1895: 408 (Quedius; subgenus Sauridus; characters; middle Europe; Caucasus).

  • — Porta, 1907: 132 (Quedius; subgenus Sauridus; characters; Italy).

  • — Reitter, 1909: 113 (Quedius; subgenus Sauridus; characters; Germany).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; catalog).

  • — Gridelli, 1925: 147 (Quedius; subgenus Raphirus; characters; England; central Europe; Pyrenees; Alps; Caucasus; France; Turkey).

  • — Portevin, 1929: 338 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1459 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 51 (Quedius; aedeagal illustration).

  • — Smetana, 1958: 386 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 138 (Quedius; subgenus Raphirus; characters; Europe).

  • — Lohse, 1964: 214 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Smetana, 1964c: 168 (Quedius; subgenus Raphirus; Czechoslovakia).

  • — Horion, 1965: 295 (Quedius; subgenus Raphirus; Britain; Belgium; France; Switzerland; Italy; Slovenia; Czechoslovakia; Poland; Bosnia Hercegovina; Germany; Austria).

  • — Korge, 1971: 19 (Quedius; subgenus Quediops; Iran).

  • — Korge, 1971a: 7 (Quedius; Turkey).

  • — Bordoni, 1973b: 53 (Quedius; subgenus Quedionthus; characters; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 203 (Quedius; subgenus Quediops; characters; Britain; Europe; Caucasus; Balkans).

  • — Bordoni, 1982b: 29 (Quedius; subgenus Quedionuthus; collected from moss; ecological notes; Italy).

  • — Tóth, 1984: 123 (Quedius; characters).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; Benelux).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; Italy).

  • — Gerend and Braunert, 1997: 201 (Quedius; notes; Luxembourg).

  • Distribution: Europe, Caucasus Turkey, Iran.

ripicola Cameron, 1926b: 368 (Quedius; subgenus Sauridus; Type locality: Keyarkuli Mussoorie; [Note: See lectotype designation by Smetana, 1988]. Also cited from Mussorie, Chakrata and Simla districts, 6000 to 9000 feet).

  • — Cameron, 1932: 291 (Quedius; subgenus Sauridus; characters; India).

  • — Scheerpeltz, 1933: 1459 (Quedius; catalog).

  • — Scheerpeltz, 1976: 43 (Quedius; Nepal).

  • — Smetana, 1988: 215 (Quedius; subgenus Microsaurus; lectotype designation: Keyarkuli Mussoorie; characters; Himachal Pradesh; Uttar Pradesh; Nepal).

  • — Smetana, 1992b: 2 (Quedius; subgenus Microsaurus; Nepal).

  • Distribution: India, Nepal.

sodalis Cameron, 1926b: 367 (Quedius; subgenus Microsaurus; Type locality: Simla Hills, Fagu, 8000 feet).

  • — Cameron, 1932: 287 (Quedius; subgenus Microsaurus; characters; India).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • — Smetana, 1988: 215 (Quedius; synonym of ripicola).

rivorum Sharp, 1884, see: labiatus Erichson, 1840.

rizensis Coiffait, 1977 [Note: Nomen nudum], see: grouziacus Coiffait, 1969.

robustus Scriba, 1870, see: anceps Fairmaire and Laboulbène, 1856.

robustus Wollaston, 1871, see: depauperatus Wollaston, 1871.

rodopianus Coiffait, 1971: 284 (Quedius; subgenus Raphirus; Type locality: Bulgarie: Zabârdo, Rodopes).

  • — Coiffait, 1978: 274 (Quedius; subgenus Raphirus; characters; Bulgaria).

  • Distribution: Bulgaria.

rou Smetana, 1995: 92 (Quedius; subgenus Raphirus; Type locality: Taiwan: Taichung Hsien, Anmashan, 2230 m).

  • Distribution: Taiwan.

roubali Bernhauer, 1916 [Note: Nomen nudum], see: minor Hochhuth, 1849.

rougemonti Smetana, 1990a: 202 (Quedius; subgenus Raphirus; Type locality: Afghanistan: Salang Pass).

  • Distribution: Afghanistan.

rubidulus Casey, 1915: 413 (Microsaurus; Type locality: Arizona).

  • — Scheerpeltz, 1933: 1459 (Quedius; catalog).

  • — Smetana, 1971: 122 (Quedius; subgenus Microsaurus; nominate subspecies; characters; Arizona).

  • — Smetana, 1981b: 638 (Quedius; collecting notes; Arizona).

  • Distribution: USA.

socius Smetana, 1971: 123 (Quedius; subgenus Microsaurus; subspecies of rubidulus; Type locality: Utah: Salt L. City).

rubirius Smetana, 1998c: 321 (Quedius; subgenus Distichalius; Type locality: China: Zhejiang, Tienmushan).

  • Distribution: China.

rubricollis Fauvel, 1878g: 554 (Quedius; Type locality: Australie méridionale: Victoria intérieur).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; catalog).

  • Distribution: Australia.

rubripennis Bernhauer, 1901b: 652 (Quedius; Type locality: Caucasus).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; catalog).

  • — Gridelli, 1924d: 72 (Quedius; subgenus Microsaurus; characters; Caucasus).

  • — Scheerpeltz, 1933: 1459 (Quedius; catalog).

  • — Coiffait, 1978: 30 (Quedius; characters from original description).

  • Distribution: Caucasus.

ruficeps Scheerpeltz, 1965: 286 (Quedius; subgenus Microsaurus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).

  • Distribution: Myanmar.

ruficollis Gravenhorst, 1806: 71 (Staphylinus; Type locality: Not cited).

  • — Erichson, 1840: 431 (Philonthus; characters; Tasmania).

  • — Fauvel, 1877: 270 (Quedius; characters; Australia).

  • — Fauvel, 1878g: 553 (Quedius).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; catalog).

  • — Bernhauer, 1920: 18 (Quedius; Australia).

  • — Lea, 1925: 241 (Quedius; from nest of Camponotus sp.; Australia).

  • — Scheerpeltz, 1933: 1460 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • — Frank, 1982: 47 (Quedius; list of parasites).

  • Distribution: Australia.

chalybeipennis W. J. MacLeay, 1873: 141 (Philonthus; Type locality: Gayndah).

  • — Fauvel, 1877: 271 (Philonthus; characters; Australia).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • — Lea, 1925: 241 (Quedius; synonym of ruficollis).

  • — Scheerpeltz, 1933: 1460 (Quedius; synonym of ruficollis).

ruficollis Stephens, 1833, see: nigriceps Kraatz, 1857.

ruficornis Gravenhorst, 1806, see: semiobscurus Marsham, 1802.

rufilabris Luze, 1904: 100 (Quedius; subgenus Ediquus; Type locality: Seravschan: Putschin-Pass).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; catalog).

  • — Gridelli, 1924d: 72 (Quedius; subgenus Microsaurus; characters; Turkestan).

  • — Scheerpeltz, 1933: 1460 (Quedius; catalog).

  • Distribution: Tajikistan-Uzbekistan.

rufinasus Sharp, 1884: 333 (Quedius; Type locality: Panama, David).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; catalog).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1933: 1460 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 177 (Quedius; subgenus Cyrtoquedius; characters in key; Panama).

  • Distribution: Panama.

rufipennis Stephens, 1833, see: nitipennis Stephens, 1833.

rufipennis Mäklin, 1853, see: plagiatus Mannerheim, 1843.

rufipennis Porta, 1907 [Note: Nomen nudum], see: cinctus Paykull, 1790.

rufipennis Roubal, 1911 [Note: Nomen nudum], see: minor Hochhuth, 1849.

rufipennis Trella, 1930 [Note: Nomen nudum], see: truncicola Fairmaire and Laboulbène, 1856.

rufipes Gravenhorst, 1802, see: semiobscurus Marsham, 1802.

rufipes Stephens, 1832, see: Philonthus.

rufitarsis Marsham, 1802, see: fulgidus Fabricius, 1793.

rufithorax Mulsant and Rey, 1876, see: mesomelinus Marsham, 1802.

rufocinctus Mannerheim, 1830, see: cinctus Paykull, 1790.

rufulus Blümml, 1898, see: simplicifrons Fairmaire, 1861.

rugosipennis Fagel, 1969a: 115 (Quedius; subgenus Sauridus; Type locality: Anatolie méridionale: Antalya, route de Lara).

  • — Coiffait, 1978: 210 (Quedius; subgenus Sauridus; characters; Anatolia).

  • — Assing and Wunderle, 2001: 37 (Quedius; Cyprus).

  • Distribution: Turkey.

rugosus Cameron, 1921: 273 (Quedius; subgenus Raphirus; Type locality: Lebong, Bengal, alt. 5000 feet; [Note: See lectotype designation by Smetana, 1988]).

  • — Cameron, 1932: 298 (Quedius; subgenus Raphirus; characters; India).

  • — Scheerpeltz, 1933: 1460 (Quedius; catalog).

  • — Scheerpeltz, 1965: 112 (Quedius; subgenus Raphirus; Burma).

  • — Smetana, 1988: 292 (Quedius; subgenus Raphirus; lectotype designation; characters; Burma; Nagaland; Uttar Pradesh; West Bengal; Nepal).

  • — Smetana, 1992b: 10 (Quedius; subgenus Raphirus; Nepal).

  • Distribution: India, Nepal, Myanmar.

mussooriensis Cameron, 1932: 299 (Quedius; subgenus Raphirus; Type locality: Arni Gad, Mussoorie; [Note: See lectotype designation by Smetana, 1988]. Originally cited from Mussoorie district).

  • — Scheerpeltz, 1933: 1450 (Quedius; catalog).

  • — Smetana, 1988: 292 (Quedius; lectotype designation: Arni Gad, Mussoorie; synonym of rugosus).

rupimontis Casey, 1915, see: prostans Horn, 1878.

rusticus Smetana, 1971: 190 (Quedius; subgenus Raphirus; Type locality: Canada: British Columbia: Mcleod Meadows, Kootenay Nat. Pk., 3800 ft. Also cited from Alberta, Manitoba, Quebec, New Hampshire, Maine, Massachusetts, New York, Michigan, New Mexico).

  • — Smetana, 1971a: 1846 (Quedius; subgenus Raphirus; collecting notes).

  • — Smetana, 1973: 1428 (Quedius; notes; Alberta; Quebec).

  • — Smetana, 1976c: 180 (Quedius; subgenus Raphirus; Alberta; Northwest Territories; Ontario; Quebec).

  • — Legner and Moore, 1977: 176 (Quedius; found under bark).

  • — Smetana, 1978a: 829 (Quedius; subgenus Raphirus; collecting notes; Arizona; New Hampshire; North Carolina; Quebec).

  • — Smetana, 1990d: 100 (Quedius; Saskatchewan; New Hampshire; South Dakota).

  • — Levesque and Levesque, 1996: 292 (Quedius; seasonal abundance; Quebec).

  • — Downie and Arnett, 1996: 393 (Quedius; subgenus Raphirus; characters; USA).

  • Distribution: Canada, USA.

rutilans Casey, 1915: 409 (Microsaurus; Type locality: Siskiyou California; [Note: See lectotype designation by Smetana, 1971]. Also cited from California: Lake Tahoe).

  • — Scheerpeltz, 1933: 1460 (Quedius; catalog).

  • — Smetana, 1971: 106 (Quedius; subgenus Microsaurus; lectotype designation: Siskiyou Cal.; characters; California; Nevada).

  • — Smetana, 1971a: 1840 (Quedius; subgenus Microsaurus; collecting notes; California).

  • — Smetana, 1976c: 172 (Quedius; subgenus Microsaurus; California).

  • — Smetana, 1978a: 825 (Quedius; subgenus Microsaurus; collecting notes; California).

  • — Smetana, 1990d: 97 (Quedius; Oregon).

  • Distribution: USA.

rutilipennis Scheerpeltz, 1965: 283 (Quedius; subgenus Microsaurus; Type locality: N.E. Burma, Kambaiti, 2000 m).

  • Distribution: Myanmar.

sachonensis Cho, 1996, see: japonicus Sharp, 1874.

safaensis Fagel, 1968c: 8 (Quedius; subgenus Sauridus; Type locality: Liban: Nabeh Safa, 1000 m).

  • — Coiffait, 1978: 215 (Quedius; subgenus Sauridus; characters; Lebanon; Turkey).

  • — Bordoni, 1984: 343 (Quedius; Lebanon).

  • Distribution: Lebanon, Turkey.

ormanus Fagel, 1971: 129 (Quedius; subgenus Sauridus; subspecies of safaensis; Type locality: Anatolie occidentale: vilayet de Bursa, Uludagh, 1800–1900 m).

samuraicus Bernhauer and Schubert, 1916: 433 (Quedius; replacement name for concolor Sharp).

  • — Scheerpeltz, 1933: 1460 (Quedius; catalog).

  • — Nakane, 1963a: 94 (Quedius; characters; habitus photograph; Japan).

  • — Coiffait, 1974a: 199 (Quedius; subgenus Microsaurus; Siberia).

  • Distribution: Japan, Russia.

concolor Sharp, 1889: 34 (Quedionuchus; [preoccupied]; Type locality: Chiuzenji; Nikko).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; synonym of samuraicus).

sanguinipennis Scheerpeltz, 1965: 296 (Quedius; subgenus Indoquedius; Type locality: N.E. Burma, Kambaiti, 7000 ft.).

  • Distribution: Myanmar.

sardous Gridelli, 1924, see: simplicifrons Fairmaire, 1861.

satanas Bernhauer, 1899a: 432 (Quedius; Type locality: Italien).

  • — Porta, 1907: 106 (Quedius; subgenus Ediquus; characters; Italy).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; catalog).

  • — Gridelli, 1922: 24 (Quedius; Italy).

  • — Gridelli, 1924d: 37 (Quedius; subgenus Microsaurus; characters; Italy; Spain).

  • — Scheerpeltz, 1933: 1460 (Quedius; catalog).

  • — Bordoni, 1973g: 126 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Bordoni, 1974: 74 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Osella and Zanetti, 1975: 115 (Quedius; collected from nest of Talpa europaea; notes; Italy).

  • — Coiffait, 1978: 144 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; Italy).

  • Distribution: Italy, Spain.

satoi Smetana, 1988: 260 (Quedius; subgenus Raphirus; Type locality: Nepal: Khandbari District, For. NE Kuwapani, 2450 m).

  • — Smetana, 1992b: 5 (Quedius; subgenus Raphirus; Nepal).

  • Distribution: Nepal.

satyrus Kiesenwetter, 1847, see: alpestris Heer, 1839.

scandens Penecke, 1911: 197 (Quedius; subgenus Microsaurus; Type locality: Czernowitz, Bukowina).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; catalog).

  • — Gridelli, 1924d: 47 (Quedius; subgenus Microsaurus; characters).

  • — Scheerpeltz, 1933: 1460 (Quedius; catalog).

  • — Roubal, 1941b: 124, 128 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • Distribution: Ukraine.

schatzmayri Gridelli, 1922b: 139 (Quedius; subgenus Raphirus; Type locality: Evora, Portogallo).

  • — Fauvel, 1874: 305 [= 1874a: 531] (Quedius; [Note: Misidentification: Scheerpeltz, 1933: 1460, cited semiaeneus Fauvel as a synonym of schatzmayri; Fauvel did not describe the species as new, he attributed it to Stephens]; characters; France; Britain; Algeria; Syria; Caucasus).

  • — Gridelli, 1925: 158 (Quedius; subgenus Raphirus; characters; Portugal; Mediterranean region; Tunisia; Algeria; France).

  • — Donisthorpe, 1930: 94 (Quedius; characters; Britain).

  • — Scheerpeltz, 1933: 1460 (Quedius; catalog).

  • — Tottenham, 1948a: 247 (Quedius; subgenus Arphirus; characters; Britain).

  • — Smetana, 1958: 391 (Quedius; subgenus Raphirus; characters; ecology).

  • — Smetana, 1962b: 148 (Quedius; subgenus Raphirus; characters; Europe).

  • — Brakman, 1962: 46 (Quedius; characters; Holland).

  • — Lohse, 1964: 217 (Quedius; subgenus Raphirus; cited as schatzmayeri; characters; central Europe).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 265 (Quedius; subgenus Raphirus; characters; Europe; Britain; Iberian Peninsula; North Africa).

  • — Lucht, 1987: 110 (Quedius; checklist; Benelux).

  • — Whitehead, 1990e: 13 (Quedius; collecting notes; Britain).

  • — Allen, 1990b: 4 (Quedius; collecting notes; Britain).

  • — Whitehead, 1991: 6 (Quedius; characters; Britain).

  • — Owen, 1997a: 149 (Quedius; collecting sites; Ireland).

  • Distribution: Tunisia, Algeria, Europe.

scheerpeltzi Gridelli, 1938: 14 (Quedius; subgenus Microsaurus; Type locality: Cipro).

  • — Scheerpeltz, 1956c: 1101 (Quedius; subgenus Microsaurus; Cyprus).

  • — Coiffait, 1970e: 62 (Quedius; Cyprus).

  • — Coiffait, 1978: 179 (Quedius; subgenus Microsaurus; characters; Cyprus).

  • — Assing and Wunderle, 2001: 37, 40 (Quedius; synonymic notes; Cyprus).

  • Distribution: Cyprus.

cyprinus H. Franz, 1987: 81 (Quedius; subgenus Microsaurus; Type locality: Troodosgebirge auf Cypern).

  • — Assing and Wunderle, 2001: 40 (Quedius; synonym of scheerpeltzi).

scheerpeltzianus Fagel, 1968a: 195 (Quedius; subgenus Sauridus; Type locality: Morée: Taygète. Also cited from Anatolia).

  • — Coiffait, 1978: 254 (Quedius; subgenus Sauridus; characters; Greece; Turkey).

  • Distribution: Turkey, Greece.

schipkanus Scheerpeltz, 1937b: 223 (Quedius; Type locality: Bulgarien: Usanna, Schiptschenska-Planina, Schipka-Balkan, ca. 1400 m).

  • — Coiffait, 1963a: 389 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 225 (Quedius; subgenus Sauridus; characters; Bulgaria).

  • Distribution: Bulgaria.

scholzi Scheerpeltz, 1933, see: fuliginosus Gravenhorst, 1802.

schuelkei Smetana, 1997d: 455 (Quedius; subgenus Microsaurus; Type locality: China: China Shaanxi, Qin Ling Shan, 109.16 E, 34.20 N, Li Shan Mt., nr. Lintong, 31 km E Xian, 1000-1200 m).

  • Distribution: China.

schultzei Smetana, 1975: 316 (Quedius; subgenus Quedionuchus; Type locality: Mexico: Guerrero: Lucerna bei Malinaltepec).

  • — Smetana, 1976a: 234 (Quedius; subgenus Quedionuchus; characters; Mexico).

  • Distribution: Mexico.

schulzei Korge, 1966: 385 (Quedius; subgenus Microsaurus; Type locality: Albanien: Bizë bei Shëngjergji).

  • — Coiffait, 1978: 175 (Quedius; subgenus Microsaurus; characters; Albania).

  • Distribution: Albania.

schawalleri Smetana, 1992b: 9 (Quedius; subgenus Raphirus; Type locality: Nepal: 395 Sankhua Sabha Distr., descent from Meropapa La to Yumu-tanga River, 4600 m).

  • Distribution: Nepal.

scintillans Gravenhorst, 1806: 70 (Staphylinus; Type locality: Not cited).

  • — Lacordaire, 1835: 384 (Emus; characters; France).

  • — Erichson, 1839a: 494 (Quedius; characters; Germany).

  • — Erichson, 1840: 549 (Quedius; characters; Germany).

  • — Gravenhorst, 1847: 231 (Quedius; characters; notes).

  • — Redtenbacher, 1849: 825 (Philonthus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 541 (Quedius; characters; France).

  • — Kraatz, 1857d: 519 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 203 (Quedius; characters; Austria).

  • — Kraatz, 1858a: 60 (Quedius; Greece).

  • — G. Waterhouse, 1858: 23 (Quedius; synonym of fuscipes).

  • — Thomson, 1860: 178 (Raphirus; characters; Scandinavia).

  • — Fauvel, 1865a: 293 [= 1865b: 43] (Quedius; habitat; France).

  • — Baudi, 1870: 384 (Quedius; characters).

  • — Fauvel, 1874: 301 [= 1874a: 527] (Quedius; characters; France; Algeria; Syria; Cyprus; Caucasus; Iran).

  • — Redtenbacher, 1874: 200 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 264 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 756 [= 1877: 612] (Quedius; subgenus Sauridus; characters; France).

  • — Fauvel, 1878: 130 [= 1878a: 50] (Quedius; Algeria).

  • — Fauvel, 1886: 60 [= 1886a: 52] (Quedius; Morocco; Algeria).

  • — Fowler, 1888: 241 (Quedius; subgenus Sauridus; characters; Britain).

  • — Eppelsheim, 1892: 332 (Quedius; Taschkent).

  • — Ganglbauer, 1895: 409 (Quedius; subgenus Sauridus; characters; Europe; Mediterranean region).

  • — Fauvel, 1902b: 117 (Quedius; Morocco; Algeria; Tunisia).

  • — Sainte-Claire Deville, 1906: 103 (Quedius; Corsica).

  • — Porta, 1907: 142 (Quedius; subgenus Sauridus; characters; Italy).

  • — Reitter, 1909: 114 (Quedius; subgenus Sauridus; characters; Germany).

  • — Johansen, 1914: 369 (Quedius; subgenus Sauridus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; catalog).

  • — Gridelli, 1925: 145 (Quedius; subgenus Raphirus; characters; Europe; Mediterranean region; Iran).

  • — Portevin, 1929: 344 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1460 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 49 (Quedius; aedeagal illustration).

  • — Jeannel and Jarrige, 1949: 380 (Quedius; subgenus Raphirus; cave species; Spain).

  • — V. Hansen, 1952: 150 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Scheerpeltz, 1956c: 1107 (Quedius; subgenus Raphirus; Cyprus).

  • — Smetana, 1958: 384 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1961d: 134 (Quedius; subgenus Raphirus; Italy).

  • — Scheerpeltz, 1961e: 137 (Quedius; subgenus Raphirus; Rhodes).

  • — Smetana, 1962b: 148 (Quedius; subgenus Raphirus; characters; Europe).

  • — Scheerpeltz, 1963a: 421 (Quedius; subgenus Raphirus; Greece).

  • — Szujecki, 1963a: 35 (Quedius; Poland).

  • — Palm, 1963b: 150 (Quedius; subgenus Raphirus; characters; Sweden; Denmark).

  • — Lohse, 1964: 213 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Scheerpeltz, 1964a: 300 (Quedius; subgenus Raphirus; Crete).

  • — Focarile, 1964: 60 (Quedius; Italy).

  • — Scheerpeltz, 1965c: 512 (Quedius; Sicily).

  • — Horion, 1965: 292 (Quedius; subgenus Raphirus; Turkey; Iran; Afghanistan; Cyprus; Lebanon; Morocco; widespread in southern Europe from Balkans to Iberian Peninsula; Russia; Poland; Czechoslovakia; Hungary; Romania; Switzerland; Austria; Germany; France; Belgium; Holland; Finland; Norway; Denmark; Britain; Ireland).

  • — Coiffait, 1970e: 63 (Quedius; Cyprus).

  • — Bordoni, 1973g: 121 (Quedius; subgenus Quediops; characters; Sardinia; Sicily; Italy).

  • — Shilov, 1975a: 377 [= 1975b: 99] (Quedius; Komi region of Russia).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 201 (Quedius; subgenus Quediops; characters; Europe; North Africa; Asia Minor; Persia; Afghanistan).

  • — Smetana, 1978: 123 (Quedius; subgenus Raphirus; collecting notes; Israel).

  • — Tóth, 1984: 123 (Quedius; characters; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; Italy).

  • — Assing and Wunderle, 2001: 37 (Quedius; Cyprus).

  • Distribution: Tunisia, Algeria, Morocco, Europe, Russia, Caucasus, Turkey, Lebanon, Israel, Syria, Iran, Afghanistan, Uzbekistan.

fuscipes Stephens, 1833: 243 (Raphirus; Type locality: banks of the Thames near Gravesend; Norfolk: Holme-near-the-sea).

  • — Stephens, 1839: 393 (Quedius; characters; England).

  • — Hardy, 1851: 35 (Quedius; characters; notes; England).

  • — Gemminger and Harold, 1868: 572 (Quedius; synonym of scintillans).

  • — Porta, 1907: 142 (Quedius; synonym of scintillans).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; synonym of scintillans).

  • — Gridelli, 1925: 145 (Quedius; synonym of scintillans).

  • — Smetana, 1958: 384 (Quedius; synonym of scintillans).

  • — Coiffait, 1978: 201 (Quedius; synonym of scintillans).

vicinus Lacordaire, 1835: 383 (Emus; [preoccupied]; Type locality: Paris).

  • — Erichson, 1840: 552 (Quedius; characters; France).

  • — Kraatz, 1857d: 501 (Quedius; France).

  • — Fauvel, 1874: 301 [= 1874a: 527] (Quedius; synonym of scintillans).

  • — Ganglbauer, 1895: 409 (Quedius; synonym of scintillans).

  • — Porta, 1907: 142 (Quedius; synonym of scintillans).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; synonym of scintillans).

  • — Gridelli, 1925: 145 (Quedius; synonym of scintillans).

  • — Smetana, 1958: 384 (Quedius; cited as bicinus; synonym of scintillans).

  • — Bordoni, 1973g: 121 (Quedius; synonym of scintillans).

  • — Coiffait, 1978: 201 (Quedius; synonym of scintillans).

monspeliensis Fairmaire, 1857: 636 (Quedius; Type locality: Montpellier).

  • — Fauvel, 1869: 487 (Quedius; synonym of scintillans).

  • — Fauvel, 1874: 301 [= 1874a: 527] (Quedius; synonym of scintillans).

  • — Ganglbauer, 1895: 409 (Quedius; synonym of scintillans).

  • — Porta, 1907: 142 (Quedius; synonym of scintillans).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; synonym of scintillans).

  • — Gridelli, 1925: 145 (Quedius; synonym of scintillans).

  • — Smetana, 1958: 384 (Quedius; synonym of scintillans).

  • — Bordoni, 1973g: 121 (Quedius; synonym of scintillans).

  • — Coiffait, 1978: 201 (Quedius; synonym of scintillans).

islamita Saulcy, 1864: 638 (Quedius; Type locality: Beïroût Djenîn, Jérusalem, Hébron, Arag-el-Emir).

  • — Fauvel, 1874: 301 [= 1874a: 527] (Quedius; synonym of scintillans).

  • — Ganglbauer, 1895: 409 (Quedius; synonym of scintillans).

  • — Porta, 1907: 142 (Quedius; synonym of scintillans).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; synonym of scintillans).

  • — Gridelli, 1925: 145 (Quedius; synonym of scintillans).

  • — Smetana, 1958: 384 (Quedius; synonym of scintillans).

  • — Bordoni, 1973g: 121 (Quedius; synonym of scintillans).

  • — Coiffait, 1978: 201 (Quedius; synonym of scintillans).

etruscus Mulsant and Rey, 1876: 759 [= 1877: 615] note (Quedius; Type locality: Toscane).

  • — Duvivier, 1883: 140 (Quedius; catalog; synonym of scintillans).

  • — Ganglbauer, 1895: 409 (Quedius; synonym of scintillans).

  • — Porta, 1907: 142 (Quedius; synonym of scintillans).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; synonym of scintillans).

  • — Gridelli, 1925: 145 (Quedius; synonym of scintillans).

  • — Smetana, 1958: 384 (Quedius; synonym of scintillans).

  • — Bordoni, 1973g: 121 (Quedius; synonym of scintillans).

  • — Coiffait, 1978: 201 (Quedius; synonym of scintillans).

bipustulatus Fiori, 1894: 88 (Quedius; variety of scintillans; Type locality: Not cited).

  • — Porta, 1907: 142 (Quedius; variety of scintillans).

  • — Scheerpeltz, 1933: 1461 (Quedius; synonym of scintillans).

  • — Smetana, 1958: 384 (Quedius; synonym of scintillans).

  • — Bordoni, 1973g: 121 (Quedius; synonym of scintillans).

  • — Coiffait, 1978: 201 (Quedius; subgenus Microquedius; variety of scintillans; characters).

amethystinus Roubal, 1948: 46 (Quedius; aberration of scintillans, cited as scintilans; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Bohemia: Praha-Břevnov).

  • — Smetana, 1958: 384 (Quedius; aberration of scintillans).

  • — Smetana, 1962b: 148 (Quedius; aberration of scintillans; characters).

  • — Bordoni, 1973g: 121 (Quedius; synonym of scintillans).

  • — Coiffait, 1978: 201 (Quedius; synonym of scintillans).

sciticollis Broun, 1894: 420 (Quedius; Type locality: Waiorongamai, near Mount Te Aroha).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; catalog).

  • Distribution: New Zealand.

scitus Gravenhorst, 1806: 50 (Staphylinus; Type locality: Brunsvigae).

  • — Gyllenhal, 1810: 306 (Staphylinus; characters; Sweden).

  • — Gyllenhal, 1827: 478 (Staphylinus; characters for variety).

  • — C. Sahlberg, 1830: 316 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 26 (Staphylinus; Finland; Sweden).

  • — Mannerheim, 1831: 440 (Staphylinus; Finland; Sweden).

  • — Runde, 1835: 6 (Staphylinus; characters; Germany).

  • — Lacordaire, 1835: 378 (Emus; characters; France).

  • — Nordmann, 1837: 79 (Philonthus; Germany; Sweden; Russia).

  • — Erichson, 1839a: 487 (Quedius; characters; Germany).

  • — Heer, 1839: 274 (Philonthus; characters; Switzerland).

  • — Stephens, 1839: 390 (Quedius; characters; England).

  • — Erichson, 1840: 528 (Quedius; characters; Europe).

  • — Gravenhorst, 1847: 215 (Quedius; characters).

  • — Redtenbacher, 1849: 708 (Philonthus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 540 (Quedius; characters; France).

  • — Kraatz, 1857d: 496 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 200 (Quedius; characters; Austria).

  • — Thomson, 1860: 177 (Microsaurus; characters; Scandinavia).

  • — Fauvel, 1874: 284 [= 1874a: 510] (Quedius; characters; France; Europe).

  • — Redtenbacher, 1874: 198 (Quedius; characters; Austria).

  • — Mulsant and Rey, 1876: 667 [= 1877: 523] (Quedius; characters; France).

  • — J. Sahlberg, 1876: 28 (Microsaurus; Finland).

  • — Fowler, 1888: 235 (Quedius; characters; Britain).

  • — Ganglbauer, 1895: 401 (Quedius; subgenus Microsaurus; characters; north and middle Europe).

  • — Porta, 1907: 127 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Reitter, 1909: 110 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Johansen, 1914: 360 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; catalog).

  • — Munster, 1923: 194 (Quedius; subgenus Microsaurus; Fennoscandia).

  • — Gridelli, 1924d: 67 (Quedius; subgenus Microsaurus; characters; Silesia; Russia; Styria; Carniola; Italy).

  • — Portevin, 1929: 340 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1461 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 16 (Quedius; aedeagal illustration).

  • — Roubal, 1941b: 123, 129 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Paulian, 1941: 270 (Quedius; larval characters).

  • — Tottenham, 1949: 422 (Quedius; checklist; Britain).

  • — V. Hansen, 1952: 138 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Smetana, 1958: 356 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 153 (Quedius; subgenus Microsaurus; characters).

  • — Palm, 1963b: 136 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 208 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Smetana, 1964c: 168 (Quedius; subgenus Microsaurus; Czechoslovakia).

  • — M. Dvořák, 1965: 89 (Quedius; Slovakia).

  • — Horion, 1965: 268 (Quedius; subgenus Microsaurus; Finland; Sweden; Norway; Britain; Denmark; Holland; Belgium; Switzerland; France; Italy; Istria; Slovenia; Dalmatia; Bosnia; Romania; Hungary; Czechoslovakia; Poland; Germany; Austria).

  • — Pototskaia, 1967: 82 (Quedius; larval characters).

  • — Szujecki, 1968a: 735 (Quedius; Poland).

  • — Scheerpeltz, 1968a: 90 (Quedius; catalog; Austria).

  • — Bordoni, 1974: 49 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Pototskaia, 1976: 173 (Quedius; larval characters).

  • — Legner and Moore, 1977: 176 (Quedius; found under bark).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 169 (Quedius; subgenus Microsaurus; characters; Europe; Scandinavia; Britain).

  • — Muona, 1979: 15 (Quedius; Finland; Russia; Sweden; Norway; Denmark; Baltic States).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 129 (Quedius; catalog; Poland).

  • — Frank, 1982: 47 (Quedius; list of parasites).

  • — Tóth, 1984: 111 (Quedius; characters; Hungary).

  • — Ádám, 1987: 152 (Microsaurus; collecting notes; Hungary).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Allen, 1990b: 4 (Quedius; collecting notes; Britain).

  • — Silfverberg, 1992: 18 (Quedius; Finland; Russia; Sweden; Norway; Denmark; Estonia; Lithuania).

  • — Smetana, 1993b: 50 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; Italy).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 245 (Quedius; Denmark).

  • — M. Hansen, 1996: 107 (Quedius; checklist; Denmark).

  • — Ádám, 1996: 247 (Microsaurus; synonym of analis).

  • — Gerend and Braunert, 1997: 201 (Quedius; notes; Luxembourg).

  • — Eisinger, 1997: 171 (Quedius; collecting notes; Germany).

  • — M. Hansen, Jørum, Palm, and Pedersen, 1997: 128 (Quedius; checklist; Denmark).

  • — Owen, 1999b: 61 (Quedius; captured underground at roots of old trees; Britain).

  • — Fowles, Alexander, and Key, 1999: 129 (Quedius; saproxylic species; status as endangered or threatened species; Britain).

  • — Schülke, 1999d: 978, 982 (Quedius; synonym of analis).

  • — Owen, 2000: 252 (Quedius; pitfall trap collections from underground at roots of trees; Britain).

  • — Herman, 2001: 31 (Quedius; prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.3).

  • Distribution: Europe, Russia.

analis Fabricius, 1787: 221 (Staphylinus; [Note: This name is older than scitus; this name was usually misidentified as a species of Bolitobius, Megacronus, or Bryocharis (Schülke, 1999d: 978, 982)]; Type locality: Kiliae [Note: See lectotype designation by Schülke, 1999d]).

  • — Gmelin, 1790: 2029 (Staphylinus; characters).

  • — Fabricius, 1793: 526 (Staphylinus; characters).

  • — Fabricius, 1801: 598 (Staphylinus; characters; Germany).

  • — Turton, 1802: 513 (Staphylinus; characters; Kiel).

  • — Erichson, 1840: 528 (Quedius; synonym of scitus).

  • — Kraatz, 1857d: 496 (Quedius; synonym of scitus).

  • — Fauvel, 1869: 487 (Quedius; synonym of scitus).

  • — Ganglbauer, 1895: 401 (Quedius; synonym of scitus).

  • — Porta, 1907: 127 (Quedius; synonym of scitus).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; synonym of scitus).

  • — Gridelli, 1924d: 67 (Quedius; synonym of scitus).

  • — Smetana, 1958: 356 (Quedius; synonym of scitus).

  • — Ádám, 1996: 247 (Quedius; valid name; Hungary).

  • — Schülke, 1999d: 978, 982 (Quedius; valid name; lectotype designation).

  • — Herman, 2001: 31 (Quedius; synonym of scitus).

atricillus Gravenhorst, 1806: 55 (Staphylinus; Type locality: Not cited).

  • — Erichson, 1839a: 488 (Quedius; synonym of scitus).

  • — Erichson, 1840: 528 (Quedius; synonym of scitus).

  • — Gravenhorst, 1847: 216 (Quedius; characters).

  • — Redtenbacher, 1849: 708 (Philonthus; synonym of scitus).

  • — Kraatz, 1857d: 496 (Quedius; synonym of scitus).

  • — Fauvel, 1869: 487 (Quedius; synonym of scitus).

  • — Ganglbauer, 1895: 401 (Quedius; synonym of scitus).

  • — Porta, 1907: 127 (Quedius; synonym of scitus).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; synonym of scitus).

  • — Gridelli, 1924d: 67 (Quedius; synonym of scitus).

  • — Scheerpeltz, 1933: 1461 (Quedius; synonym of scitus).

  • — Smetana, 1958: 356 (Quedius; synonym of scitus).

pygmaeus Gravenhorst, 1806: 57 (Staphylinus; [preoccupied]; Type locality: Not cited).

  • — Erichson, 1839a: 488 (Quedius; synonym of scitus).

  • — Erichson, 1840: 528 (Quedius; synonym of scitus).

  • — Gravenhorst, 1847: 215 (Quedius; characters).

  • — Redtenbacher, 1849: 708 (Philonthus; synonym of scitus).

  • — Kraatz, 1857d: 496 (Quedius; synonym of scitus).

  • — Fauvel, 1869: 487 (Quedius; synonym of scitus).

  • — Ganglbauer, 1895: 401 (Quedius; synonym of scitus).

  • — Porta, 1907: 127 (Quedius; synonym of scitus).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; synonym of scitus).

  • — Gridelli, 1924d: 67 (Quedius; synonym of scitus).

  • — Smetana, 1958: 356 (Quedius; synonym of scitus).

atriceps Stephens, 1832: 218 (Quedius; Type locality: Suffolk).

  • — Curtis, 1829: 25 (Quedius; nomen nudum; catalog; Britain).

  • — Kraatz, 1857d: 496 (Quedius; synonym of scitus).

  • — G. Waterhouse, 1858: 22 (Quedius; synonym of fulgidus).

  • — Fauvel, 1869: 487 (Quedius; synonym of scitus).

  • — Ganglbauer, 1895: 401 (Quedius; synonym of scitus).

  • — Porta, 1907: 127 (Quedius; synonym of scitus).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; synonym of scitus).

  • — Gridelli, 1924d: 67 (Quedius; synonym of scitus).

  • — Smetana, 1958: 356 (Quedius; synonym of scitus).

meledanus Reitter, 1900a: 288 (Quedius; variety of scitus; Type locality: Insel Meleda).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; variety of scitus).

  • — Gridelli, 1924d: 67, 68 (Quedius; variety of scitus; characters).

  • — Scheerpeltz, 1933: 1461 (Quedius; variety of scitus).

  • — Roubal, 1941b: 123, 129 (Quedius; subgenus Microsaurus; variety of scitus; characters; collecting notes).

  • — Smetana, 1958: 356 (Quedius; synonym of scitus).

  • — Smetana, 1960e: 153 (Quedius; aberration of scitus; characters).

  • — Bordoni, 1974: 49 (Quedius; aberration of scitus).

  • — Coiffait, 1978: 169 (Quedius; subgenus Microsaurus; variety of scitus; characters).

  • — Tóth, 1984: 112 (Quedius; variety of scitus; characters).

scribae Ganglbauer, 1895: 412 (Quedius; subgenus Raphirus; proposed for Scriba's misidentification of picipennis; Type locality: Bayer Kirchb. a. W.; [Note: See lectotype designation by Assing, 1999b]. Originally cited from Alpen von Oesterreich und Steiermark; Karpathen).

  • — Scriba, 1864: 128 (Quedius; [Note: Misidentification: Bernhauer and Schubert, 1916: 434, cited picipennis Scriba as a synonym of scribae; Scriba did not describe the species as new, he attributed it to Heer]; characters; Austria).

  • — Porta, 1907: 149 (Quedius; subgenus Raphirus; characters; Italy).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; catalog).

  • — Gridelli, 1925: 160 (Quedius; subgenus Raphirus; characters; Switzerland; Austria; Italy; Romania).

  • — Scheerpeltz, 1933: 1461 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 60 (Quedius; aedeagal illustration).

  • — Smetana, 1958: 391 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 150 (Quedius; subgenus Raphirus; characters; Europe).

  • — Lohse, 1964: 217 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 303 (Quedius; subgenus Raphirus, cited as scribai; Czechoslovakia; Poland; Romania; Germany; Austria; Switzerland; Italy).

  • — Coiffait, 1978: 268 (Quedius; subgenus Raphirus; characters; Alps; Carpathians; mountains of central Europe).

  • — Tóth, 1984: 127 (Quedius; cited as scribai; characters).

  • — Drugmand, 1987a: 312 (Quedius; Belgium).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Assing, 1999b: 36 (Quedius; lectotype designation: Bayer Kirchb. a. W.; characters).

  • Distribution: Europe.

scutellaris Broun, 1894, see: antarcticus Bernhauer and Schubert, 1916.

scutellaris Eppelsheim, 1890: 224 (Quedius; subgenus Microsaurus; Type locality: Tunis).

  • — Fauvel, 1902b: 114 (Quedius; Algeria; Tunisia).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; catalog).

  • — Gridelli, 1924d: 27 (Quedius; subgenus Microsaurus; characters; Tunisia).

  • — Scheerpeltz, 1933: 1461 (Quedius; catalog).

  • — Coiffait, 1978: 182 (Quedius; subgenus Microsaurus; characters; Tunisia; Algeria).

  • Distribution: Tunisia, Algeria.

secretus Cameron, 1948c: 725 (Quedius; subgenus Ediquus; Type locality: New Zealand: Auckland Island: Carnley Harbour, Musgrave Pen).

  • — Gourlay, 1950: 187 (Quedius; collecting notes).

  • Distribution: New Zealand.

secundus Last, 1952, see: simplicifrons Fairmaire, 1861.

sedlaceki Last, 1987: 51 (Quedius; Type locality: Papua New Guinea NE: Karimui, 1080 m).

  • Distribution: New Guinea.

seeversi Korge, 1963, see: capucinus Gravenhorst, 1806.

segersi Herman, 2001: 49 (Quedius; replacement name for conicus Segers).

  • Distribution: France.

conicus Segers, 1987b: 267 (Quedius; [preoccupied]; Type locality: France: Pyrénées Centrales, St.-Lary, Aragnouet).

  • — Herman, 2001: 49 (Quedius; synonym of segersi).

semiaeneus Stephens, 1833: 243 (Raphirus; Type locality: London; Suffolk; Norfolk).

  • — Heer, 1839: 279 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1916: 434, cited attenuatus Heer as a synonym of semiaeneus Stephens; Heer did not describe the species as new, he attributed it to Gyllenhal who attributed it to Gravenhorst]; characters; Switzerland).

  • — Stephens, 1839: 394 (Quedius; characters; England).

  • — Erichson, 1840: 544 (Quedius; [Note: Misidentification: Bernhauer and Schubert, 1916: 434, cited semiobscurus Erichson as a synonym of semiaeneus Stephens; Erichson did not describe the species as new, he attributed it to Marsham]; characters; England; Sardinia).

  • — G. Waterhouse, 1858: 23 (Quedius; synonym of semiobscurus).

  • — Fauvel, 1869: 487 (Quedius; synonym of nitipennis).

  • — Baudi, 1870: 384 (Quedius; characters; Syria).

  • — Fauvel, 1878: 130 [= 1878a: 50] (Quedius; Algeria).

  • — Fauvel, 1886: 60 [= 1886a: 52] (Quedius; Morocco; Tunisia; Algeria).

  • — Fowler, 1888: 244 (Quedius; subgenus Raphirus; characters; Britain).

  • — Ganglbauer, 1895: 411 (Quedius; subgenus Raphirus; characters; western middle Europe; Mediterranean region).

  • — Fauvel, 1902b: 117 (Quedius; Morocco; Algeria; Tunisia).

  • — Poppius, 1905: 9 (Quedius; Shetland Islands).

  • — Sainte-Claire Deville, 1906: 103 (Quedius; Corsica).

  • — Porta, 1907: 146 (Quedius; subgenus Raphirus; characters; Italy).

  • — Reitter, 1909: 114 (Quedius; subgenus Raphirus; characters; Germany).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; catalog).

  • — Gridelli, 1922b: 139 (Quedius; subgenus Raphirus; characters).

  • — Gridelli, 1925: 156 (Quedius; subgenus Raphirus; characters).

  • — Portevin, 1929: 345 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1461 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 59 (Quedius; aedeagal illustration).

  • — Tottenham, 1948a: 246 (Quedius; subgenus Arphirus; characters; Britain).

  • — V. Hansen, 1952: 154 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Scheerpeltz, 1956c: 1107 (Quedius; subgenus Raphirus; Cyprus).

  • — Smetana, 1958: 390 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 148 (Quedius; subgenus Raphirus; characters; Europe).

  • — Brakman, 1962: 46 (Quedius; characters).

  • — Palm, 1963b: 152 (Quedius; subgenus Raphirus; characters; Sweden; Denmark).

  • — Lohse, 1964: 217 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Scheerpeltz, 1964a: 300 (Quedius; subgenus Raphirus; Crete).

  • — Focarile, 1964: 61 (Quedius; Italy).

  • — Horion, 1965: 301 (Quedius; subgenus Raphirus; Algeria; Tunisia; Cyprus; Britain; Denmark; Sweden; Holland; Belgium; Switzerland; Portugal; Spain; Italy; Dalmatia; Albania; Greece; Bosnia; Czech Republic; Germany; Austria).

  • — Coiffait, 1970e: 63 (Quedius; Cyprus).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 271 (Quedius; subgenus Raphirus; characters; Europe; Britain; Scandinavia; Mediterranean region; North Africa).

  • — Tóth, 1984: 125 (Quedius; characters).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; Benelux; Denmark; Sweden).

  • — Schülke and Uhlig, 1988: 12 (Quedius; notes; Germany).

  • — Whitehead, 1990e: 13 (Quedius; collecting notes; Britain).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic ?).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; Italy).

  • — Owen, 1997a: 149 (Quedius; collecting sites; Ireland).

  • — Assing and Wunderle, 2001: 37 (Quedius; Cyprus).

  • — Assing, 2001: 75 (Quedius; collected from xerothermous site; Germany).

  • Distribution: Tunisia, Algeria, Morocco, Europe.

proximus Kraatz, 1858a: 60 (Quedius; Type locality: Zante).

  • — Fauvel, 1869: 487 (Quedius; synonym of nitipennis).

  • — Fauvel, 1874: 305 [= 1874a: 531] (Quedius; synonym of semiaeneus).

  • — Ganglbauer, 1895: 411 (Quedius; synonym of semiaeneus).

  • — Porta, 1907: 146 (Quedius; synonym of semiaeneus).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; synonym of semiaeneus).

  • — Gridelli, 1925: 156 (Quedius; synonym of semiaeneus).

  • — Smetana, 1958: 390 (Quedius; synonym of semiaeneus).

  • — Coiffait, 1978: 217 (Quedius; synonym of semiaeneus).

myagrus Rottenberg, 1870: 29 (Quedius; Type locality: Sicily: Syracus).

  • — Fauvel, 1874: 305 [= 1874a: 531] (Quedius; synonym of semiaeneus).

  • — Ganglbauer, 1895: 411 (Quedius; synonym of semiaeneus).

  • — Porta, 1907: 146 (Quedius; synonym of semiaeneus).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; synonym of semiaeneus).

  • — Gridelli, 1925: 156 (Quedius; synonym of semiaeneus).

  • — Smetana, 1958: 390 (Quedius; synonym of semiaeneus).

  • — Coiffait, 1978: 217 (Quedius; synonym of semiaeneus).

semiflavus Fairmaire and Germain, 1861, see: Loncovilius.

semilaeviventris Scheerpeltz, 1965: 288 (Quedius; subgenus Raphirus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).

  • Distribution: Myanmar.

seminitidus Stephens, 1832, see: picipes Mannerheim, 1830.

semiobscurus Marsham, 1802: 512 (Staphylinus; Type locality: Britain).

  • — Gyllenhal, 1810: 304 (Staphylinus; synonym of variabilis Gyllenhal).

  • — Curtis, 1829: 26 (Philonthus; catalog; Britain).

  • — Stephens, 1833: 241 (Raphirus; characters; England).

  • — Stephens, 1839: 393 (Quedius; characters; England).

  • — Hardy, 1851: 33 (Quedius; notes; England).

  • — Rosenhauer, 1856: 71 (Quedius; P. S. Maria).

  • — Fairmaire and Laboulbène, 1856: 538 (Quedius; characters; France).

  • — Kraatz, 1857d: 501 (Quedius; England; Sardinia).

  • — Redtenbacher, 1857: 202 (Quedius; characters; Austria).

  • — Kraatz, 1858a: 60 (Quedius; Greece).

  • — Fauvel, 1865a: 293 [= 1865b: 43] (Quedius; habitat; France).

  • — Fauvel, 1874: 304 [= 1874a: 530] (Quedius; synonym of rufipes).

  • — Redtenbacher, 1874: 199 (Quedius; characters; Austria).

  • — Mulsant and Rey, 1876: 774 [= 1877: 630] (Quedius; subgenus Raphirus; characters; France).

  • — Ganglbauer, 1895: 410 (Quedius; synonym of rufipes).

  • — Porta, 1907: 147 (Quedius; synonym of rufipes).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; synonym of rufipes).

  • — Gridelli, 1925: 155 (Quedius; synonym of rufipes).

  • — Tottenham, 1948a: 245 (Quedius; subgenus Arphirus; characters; Britain).

  • — Smetana, 1958: 389 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 149 (Quedius; subgenus Raphirus; characters; Europe).

  • — Lohse, 1964: 217 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Coiffait, 1970e: 63 (Quedius; Cyprus).

  • — Korge, 1971: 19 (Quedius; subgenus Raphrius; Iran).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 275 (Quedius; subgenus Raphirus; characters; Europe; Britain; Scandinavia; Mediterranean region; North Africa; Caucasus; Transcaucasus).

  • — Tóth, 1984: 126 (Quedius; characters; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Austria; France; Benelux; Denmark; Sweden).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic ?).

  • — Assing, 1994a: 13, 24 (Quedius; notes; wing length; xerothermous sites; Germany).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; Italy).

  • — Assing, 1998e: 140 (Quedius; Canary Islands).

  • — Assing and Wunderle, 2001: 37 (Quedius; Cyprus).

  • — Assing, 2001: 75 (Quedius; collected from xerothermous site; Germany).

  • Distribution: Tunisia, Algeria, Morocco, Canary Islands, Europe, Azerbaijan, Iran.

rufipes Gravenhorst, 1802: 171 (Staphylinus; [preoccupied]; [Note: Scheerpeltz, 1933: 1460, changed the author and date of this name to Erichson, 1840. However Erichson (1840: 543) attributed rufipes to Gravenhorst]; Type locality: Bohemia).

  • — Latreille, 1804: 324 (Staphylinus; characters; Germany).

  • — Gravenhorst, 1806: 67 (Staphylinus).

  • — Erichson, 1840: 543 (Quedius; characters; Europe).

  • — Gravenhorst, 1847: 230 (Quedius; characters; notes).

  • — Märkel and Kiesenwetter, 1848a: 325 (Quedius; notes; Carpathian Alps).

  • — Redtenbacher, 1849: 710 (Philonthus; characters; Austria).

  • — Küster, 1853a: 64 (Quedius; characters; Europe).

  • — Fairmaire and Laboulbène, 1856: 537 (Quedius; characters; France).

  • — Redtenbacher, 1857: 202 (Quedius; characters; Austria).

  • — Fauvel, 1865a: 293 [= 1865b: 43] (Quedius; habitat; France).

  • — Fauvel, 1874: 304 [= 1874a: 530] (Quedius; characters; France; Britain; Switzerland; Austria; Italy; Sicily; Corsica; Sardinia; Algeria; Cyprus).

  • — Redtenbacher, 1874: 199 (Quedius; characters; Austria).

  • — Mulsant and Rey, 1876: 769 [= 1877: 625] (Quedius; subgenus Raphirus; characters; larval characters; France).

  • — Fauvel, 1878: 130 [= 1878a: 50] (Quedius; Algeria).

  • — Fauvel, 1886: 60 [= 1886a: 52] (Quedius; Algeria; Tunisia).

  • — Fowler, 1888: 243 (Quedius; subgenus Raphirus; characters; Britain).

  • — Ganglbauer, 1895: 410 (Quedius; subgenus Raphirus; characters; western middle Europe; Mediterranean region).

  • — Fauvel, 1902b: 117 (Quedius; Algeria; Tunisia).

  • — Sainte-Claire Deville, 1906: 103 (Quedius; Corsica).

  • — Porta, 1907: 147 (Quedius; subgenus Raphirus; characters; Italy).

  • — Reitter, 1909: 115 (Quedius; subgenus Raphirus; characters; Germany).

  • — Johansen, 1914: 370 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; catalog).

  • — Gridelli, 1925: 155 (Quedius; subgenus Raphirus; characters).

  • — Portevin, 1929: 344 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1460 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 57, 58 (Quedius; aedeagal illustration).

  • — C. Koch, 1941: 68 (Quedius; Morocco).

  • — V. Hansen, 1952: 152 (Quedius; subgenus Raphirus; Erichson cited as author; characters; Denmark).

  • — Scheerpeltz, 1956c: 1107 (Quedius; subgenus Raphirus; Erichson cited as author of species; Cyprus).

  • — Smetana, 1958: 389 (Quedius; synonym of semiobscurus).

  • — Scheerpeltz, 1961d: 134 (Quedius; subgenus Raphirus; Italy).

  • — Smetana, 1962b: 149 (Quedius; synonym of semiobscurus).

  • — Scheerpeltz, 1963a: 421 (Quedius; subgenus Raphirus; Greece).

  • — Palm, 1963b: 151 (Quedius; subgenus Raphirus; Erichson cited as author; characters; Sweden; Denmark).

  • — Scheerpeltz, 1964a: 300 (Quedius; subgenus Raphirus; Crete).

  • — Focarile, 1964: 60 (Quedius; Italy).

  • — Scheerpeltz, 1965b: 408 (Quedius; Sicily).

  • — Scheerpeltz, 1965c: 512 (Quedius; Sicily).

  • — Horion, 1965: 299 (Quedius; subgenus Raphirus; Morocco; Algeria; Tunisia; Turkey; Syria; Cyprus; Portugal; Spain; Corsica; Sardinia; Sicily; Italy; Istria; Dalmatia; Bosnia Hercegovina; Britain; Denmark; Sweden; Holland; Belgium; France; Switzerland; Germany; Austria).

  • — Pope, 1977: 31 (Quedius; synonym of semiobscurus).

  • — Coiffait, 1978: 275 (Quedius; synonym of semiobscurus).

  • — Boháč, 1988: 557 [= 1989: 40] (Quedius; synonym of semiobscurus).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; synonym of semiobscurus).

ruficornis Gravenhorst, 1806: 50 (Staphylinus; [preoccupied]; Type locality: Parisi).

  • — Gyllenhal, 1810: 302 (Staphylinus; synonym of molochinus Gravenhorst).

  • — C. Sahlberg, 1830: 314 (Staphylinus; synonym of molochinus Gravenhorst).

  • — Nordmann, 1837: 76 (Philonthus; France; England).

  • — Erichson, 1840: 543 (Quedius; synonym of rufipes).

  • — Porta, 1907: 147 (Quedius; synonym of rufipes).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; synonym of rufipes).

  • — Smetana, 1958: 389 (Quedius; synonym of semiobscurus).

  • — Coiffait, 1978: 275 (Quedius; synonym of semiobscurus).

velutinus Motschulsky, 1860: 571 (Quedius; Type locality: Schirvan, Transcaucasie meridionale).

  • — Hochhuth, 1862: 46 (Quedius; characters; Transcaucasian).

  • — Fauvel, 1874: 305 [= 1874a: 531] (Quedius; synonym of semiaeneus).

  • — Ganglbauer, 1895: 411 (Quedius; synonym of semiaeneus).

  • — Porta, 1907: 146 (Quedius; synonym of semiaeneus).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; synonym of semiaeneus).

  • — Scheerpeltz, 1933: 1464 (Quedius; catalog).

  • — Boháč, 1988: 557 [= 1989: 40] (Quedius; synonym of semiobscurus).

lutzi Reitter, 1909: 115 (Quedius; subgenus Raphirus; race of rufipes; Type locality: Transkausasus, Lenkoran).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; variety of rufipes).

  • — Coiffait, 1967e: 423 (Quedius; subgenus Raphirus; subspecies of semiobscurus; notes; characters).

  • — Coiffait, 1978: 275 (Quedius; subgenus Raphirus; subspecies of semiobscurus; characters).

semiruber Fauvel, 1875a: XXXIII [= 1875b: 235] (Quedius; Type locality: Pologne).

  • — Ganglbauer, 1895: 406 (Quedius; subgenus Sauridus; characters; Poland).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; catalog).

  • — Gridelli, 1925: 167 (Quedius; subgenus Raphirus; characters; Macedonia).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • Distribution: Poland, Macedonia.

semirufus Korge, 1971: 14 (Quedius; subgenus Sauridus; Type locality: Iran: Charasu-dagh östlich Heroabad, 2000–2300 m, Azarbaijan).

  • — Coiffait, 1978: 216 (Quedius; subgenus Sauridus; characters; Iran).

  • Distribution: Iran.

semiviolaceus Fauvel 1878, see: luridipennis W. J. MacLeay, 1873.

semiviolaceus Bernhauer and Schubert, 1916 [Note: Nomen nudum], see: luridipennis W. J. MacLeay, 1873.

senjonis Watanabe, 1987: 330 (Quedius; subgenus Microsaurus; Type locality: Japan: Honshu: Yamanashi Pref.: Mt. Senjô-ga-take on the southern Japanese Alps).

  • Distribution: Japan.

seriatus Eppelsheim, 1889, see: regularis Bernhauer and Schubert, 1916.

seriatus Horn, 1878: 166 (Quedius; Type locality: Vancouver).

  • — Casey, 1915: 417 (Raphirus; characters; Vancouver).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • — Hatch, 1957: 229 (Quedius; subgenus Raphirus; characters; British Columbia; Washington; Oregon).

  • — Smetana, 1971: 210 (Quedius; subgenus Raphirus; characters; British Columbia; California; Idaho; Oregon; Washington).

  • — Smetana, 1971b: 4 (Quedius; subgenus Raphirus; British Columbia).

  • Distribution: Canada, USA.

sericans Stephens, 1832, see: Heterothops.

sericeicollis Bernhauer, 1941b: 29 (Quedius; subgenus Sauridus; Type locality: Neu Seeland: Greymouth).

  • Distribution: New Zealand.

sericopterus Stephens, 1832: 219 (Quedius; [Note: Nomen dubium]; Type locality: London).

  • — Stephens, 1839: 291 (Quedius; characters; England).

  • — Herman, 2001: 49 (Quedius; nomen dubium).

  • Distribution: Britain.

sericopterus Stephens, 1833: 244 (Raphirus; [preoccupied]; Type locality: Gravesend).

  • — Stephens, 1839: 394 (Quedius; characters; England).

  • — Herman, 2001: 49 (Quedius; nomen dubium; although a junior primary homonym, it is suggested that the required replacement be ignored).

  • Distribution: Britain.

sexpunctatus Bernhauer, 1917g: 47 (Quedius; subgenus Sauridus; Type locality: Kap der guten Hoffnung).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • — Scheerpeltz, 1974: 171 (Quedius; Raphirus; South Africa).

  • Distribution: South Africa.

shan Smetana, 1995: 88 (Quedius; subgenus Raphirus; Type locality: Taiwan: Pingtung Hsien: Peitawushan, above Kuai-Ku Hut, 2760 m).

  • Distribution: Taiwan.

sherpai Coiffait, 1982, see: apicicornis Eppelsheim, 1895.

shibatai Smetana, 1995: 53 (Quedius; subgenus Microsaurus; Type locality: Taiwan: Lushan-Spa, Nantou-Hsien).

  • Distribution: Taiwan.

shimomurai Watanabe, 1990c: 31 (Quedius; subgenus Microsaurus; Type locality: Japan: Honshu: Saitama Pref.: NE slope on Mt. Kumotori, alt. 1750–1850 m).

  • Distribution: Japan.

shiow Smetana, 1995: 58 (Quedius; subgenus Distichalius; Type locality: Taiwan: Nantou Hsien, Yushan N. P., Mun-Li Cliff, 2700 m).

  • Distribution: Taiwan.

sidneensis Fauvel, 1877: 269 (Quedius; Type locality: Australie, Nouvelle Galles du Sud, Sydney).

  • — Fauvel, 1878g: 552 (Quedius; Australia).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; catalog).

  • — Lea, 1925: 242 (Quedius; characters; Australia).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

signifer Smetana, 1971: 176 (Quedius; subgenus Distichalius; Type locality: California: Tahoe Pines. Also cited from Nevada).

  • — Smetana, 1971a: 1846 (Quedius; subgenus Distichalius; collecting notes).

  • Distribution: USA.

sigwalti Coiffait, 1972a: 82 (Quedius; subgenus Sauridus; Type locality: Meskla, Crete).

  • — Coiffait, 1978: 235 (Quedius; subgenus Sauridus; characters; Crete).

  • Distribution: Greece.

sikkimensis Cameron, 1932, see: Indoquedius.

silensis Fiori, 1894, see: mesomelinus Marsham, 1802.

silvicola Casey, 1884, see: peregrinus Gravenhorst, 1806.

simplex Hatch, 1957: 223 (Quedius; subgenus Microsaurus; Type locality: Oregon: Benton Co.: Marys Peak. Also cited from Washington).

  • — Smetana, 1971: 93 (Quedius; subgenus Microsaurus; characters; Nevada; Oregon; Washington).

  • — Smetana, 1971a: 1839 (Quedius; subgenus Microsaurus; collecting notes; California).

  • — Legner and Moore, 1977: 176 (Quedius; found under bark).

  • — Smetana, 1978a: 825 (Quedius; subgenus Microsaurus; collecting notes; Oregon; Washington).

  • Distribution: USA.

simplicifrons Fairmaire, 1861: 580 (Quedius; Type locality: Corse).

  • — Fauvel, 1865c: 16 [= 1866: 23] (Quedius; synonym of molochinus).

  • — Kraatz, 1867: 414 (Quedius; synonym of molochinus).

  • — Ganglbauer, 1895: 403 (Quedius; synonym of molochinus).

  • — Porta, 1907: 129 (Quedius; synonym of molochinus).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; synonym of pallipes).

  • — Gridelli, 1924d: 81 (Quedius; synonym of subspecies pallipes).

  • — Gridelli, 1924d: 81 (Quedius; subspecies of molochinus; [Note: Misidentified as pallipes according to Smetana, 1962b: 135]; characters).

  • — Scheerpeltz, 1933: 1457 (Quedius; synonym of picipennis).

  • — Méquignon, 1937: 3 (Quedius; synonym of molochinus subspecies pallipes).

  • — Korge, 1960a: 69 (Quedius; synonym of pallipes).

  • — Coiffait, 1961a: 56 (Quedius; characters; France; Portugal; Madeira).

  • — Smetana, 1962b: 134 (Quedius; subgenus Quedius; characters; Europe).

  • — Smetana, 1963b: 39 (Quedius; subgenus Quedius; Madeira).

  • — Bordoni, 1976b: 100 (Quedius; synonym of pallipes).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 196 (Quedius; subgenus Quedius; characters).

  • — Lohse and Lucht, 1989: 172 (Quedius; note).

  • — Israelson, 1990: 3 (Quedius; Azores).

  • — Borges, 1990: Table II (Quedius; Azores).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; Italy).

  • Distribution: Maderia, Azores, Europe.

hispanicus Bernhauer, 1898: 341 (Quedius; Type locality: Cartagena und Cadix in Spanien).

  • — Bernhauer, 1900: 46 (Quedius; Spain; Italy).

  • — Porta, 1907: 131 (Quedius; subgenus Quedius; characters; Italy).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; variety of pallipes).

  • — Gridelli, 1924d: 81 (Quedius; synonym of subspecies pallipes).

  • — Scheerpeltz, 1933: 1457 (Quedius; synonym of picipennis).

  • — Méquignon, 1937: 3 (Quedius; synonym of molochinus subspecies pallipes).

  • — Korge, 1960a: 69 (Quedius; synonym of pallipes).

  • — Coiffait, 1961a: 56 (Quedius; synonym of simplicifrons).

  • — Smetana, 1962b: 134 (Quedius; synonym of simplicifrons).

  • — V. Hansen, 1962a: 319 (Quedius; cited as valid species; characters; Denmark).

  • — Last, 1963: 45 (Quedius; synonym of simplicifrons).

  • — Lohse, 1964: 211 (Quedius; subgenus Quedius; cited as valid species; characters; central Europe).

  • — Horion, 1965: 276 (Quedius; subgenus Quedius; cited as valid species; Iberian Peninsula; Balearics; France; Madeira; Belgium; Holland; Germany; Denmark).

  • — Smetana, 1970: 61 (Quedius; cited as valid species; distributional summary; Azores; Madeira).

  • — Bordoni, 1976b: 100 (Quedius; cited as valid species; characters).

  • — Pope, 1977: 31 (Quedius; synonym of simplicifrons).

  • — Coiffait, 1978: 196 (Quedius; synonym of simplicifrons).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Benelux; Denmark).

  • — Lohse and Lucht, 1989: 172 (Quedius; synonym of simplicifrons).

  • — Welch, 1993: 229 (Quedius; cited as valid species; ovariole number and ovary structure).

rufulus Blümml, 1898: 216 (Quedius; variety of hispanicus; Type locality: Spanien, Granada).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; synonym of pallipes).

  • — Gridelli, 1924d: 81 (Quedius; synonym of subspecies pallipes).

  • — Scheerpeltz, 1933: 1457 (Quedius; synonym of picipennis).

  • — Korge, 1960a: 69 (Quedius; synonym of pallipes).

  • — Coiffait, 1961a: 57 (Quedius; mutation of simplicifrons).

  • — Smetana, 1962b: 135 (Quedius; aberration of simplicifrons).

  • — Smetana, 1963b: 40 (Quedius; subgenus Quedius; aberration of rufulus; Madeira).

  • — Lohse, 1964: 211 (Quedius; subgenus Quedius; aberration of hispanicus; characters; central Europe).

  • — Bordoni, 1976b: 100 (Quedius; aberration of pallipes).

  • — Pope, 1977: 31 (Quedius; synonym of simplicifrons).

  • — Coiffait, 1978: 196 (Quedius; subgenus Quedius; variety of simplicifrons; characters).

hammianus Sharp, 1911b: 57 (Quedius; Type locality: Anglia: Deal, Strood, Lymington, and Hayling Island).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; synonym of pallipes).

  • — Gridelli, 1924d: 81 (Quedius; synonym of subspecies pallipes).

  • — Scheerpeltz, 1933: 1457 (Quedius; synonym of picipennis).

  • — Méquignon, 1937: 3 (Quedius; synonym of molochinus subspecies pallipes).

  • — Korge, 1960a: 69 (Quedius; synonym of pallipes).

  • — Coiffait, 1961a: 57 (Quedius; synonym of simplicifrons mut. rufulus).

  • — Smetana, 1962b: 135 (Quedius; synonym of simplicifrons).

  • — Last, 1963: 45 (Quedius; synonym of simplicifrons aberration rufulus).

  • — Pope, 1977: 31 (Quedius; synonym of simplicifrons).

  • — Coiffait, 1978: 196 (Quedius; cited as hammicus; synonym of simplicifrons variety rufulus).

sardous Gridelli, 1924d: 81, 85 (Quedius; variety of picipennis “Paykull”; Type locality: Sardegna: Cagliari, Assemini, Decimo, Golfo Aranci, Gonnesa).

  • — Scheerpeltz, 1933: 1457 (Quedius; synonym of picipennis).

  • — Korge, 1960a: 69 (Quedius; variety of pallipes).

  • — Coiffait, 1961a: 57 (Quedius; aberration of simplicifrons; characters).

  • — Smetana, 1962b: 135 (Quedius; aberration of simplicifrons).

  • — Lohse, 1964: 211 (Quedius; subgenus Quedius; aberration of hispanicus; characters; central Europe).

  • — Bordoni, 1976b: 100 (Quedius; aberration of pallipes).

  • — Coiffait, 1978: 196 (Quedius; subgenus Quedius; variety of simplicifrons; characters).

secundus Last, 1952a: 150 (Quedius; variety of pallipes; Type locality: Studland, Dorset; Mudeford, Hampshire; Chichester).

  • — Korge, 1960a: 69 (Quedius; variety of pallipes).

  • — Coiffait, 1961a: 56 (Quedius; synonym of simplicifrons).

  • — Smetana, 1962b: 134 (Quedius; synonym of simplicifrons).

  • — Last, 1963: 45 (Quedius; synonym of simplicifrons).

  • — Pope, 1977: 31 (Quedius; synonym of simplicifrons).

simulans Sharp, 1874: 25 (Quedius; Type locality: Japan).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • — Watanabe and Shibata, 1965: 321 (Quedius; subgenus Microsaurus; Japan).

  • — Watanabe and Baba 1973: 260 (Quedius; collected in gold mines; Japan).

  • — Hasegawa and Kanie, 1992: 20 (Quedius; seashore collection; Japan).

  • — J. Li, 1993: 41 (Quedius; China).

  • — Cho, 1996b: 112 (Quedius; subgenus Microsaurus; characters; Korea).

  • — Watanabe, 1996a: 14 (Quedius; cited as simurans; mine and cave collections; Japan).

  • Distribution: Japan, Korea, China.

simulator Smetana, 1971: 194 (Quedius; subgenus Raphirus; Type locality: Alaska: mi 1249 Alaska Hwy, Deadman Lake. Also cited from Michigan, Alberta, Labrador, Manitoba, Northwest Territories, Ontario, Quebec, Yukon Territory).

  • — Smetana, 1973: 1429 (Quedius; notes; Alberta; Northwest Territories; Quebec).

  • — Smetana, 1976c: 181 (Quedius; subgenus Raphirus; Ontario).

  • — Smetana, 1978a: 829 (Quedius; subgenus Raphirus; collecting notes; Quebec).

  • — Smetana, 1981b: 641 (Quedius; collecting notes; Arizona).

  • — Smetana, 1990d: 100 (Quedius; British Columbia).

  • — Downie and Arnett, 1996: 393 (Quedius; subgenus Raphirus; characters; USA).

  • Distribution: USA, Canada.

skoraszewskyi Korge, 1961, see: mesomelinus Marsham, 1802.

skrimshiranus Stephens, 1832, see: mesomelinus Marsham, 1802.

slipsensis Broun, 1923: 680 (Quedius; Type locality: Mounts Slips, Nelson).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • Distribution: New Zealand.

slovenicus M. Dvořák, 1967 [Note: Nomen nudum], see: brevis Erichson, 1840.

smetanai Roubal, 1949 [Note: Nomen nudum], see: molochinus Gravenhorst, 1806.

smetanai Korge, 1971a: 47 (Quedius; subgenus Sauridus; Type locality: Ilica, bzw. Ayder, im Tal südlich Ardesen, 1000–1600 m).

  • — Coiffait, 1978: 222 (Quedius; synonym of horsti).

  • — Herman, 2001: 30 (Quedius; valid name for species).

  • Distribution: Turkey.

horsti Coiffait, 1978: 222 (Quedius; subgenus Sauridus; replacement name for smetanai Korge; [Note: An unnecessary replacement name since smetanai Roubal was described as an aberration and is therefore unavailable]).

  • — Herman, 2001: 30 (Quedius; synonym of smetanai).

socius Smetana, 1971, see: rubidulus Casey, 1915.

sodalis Cameron, 1926, see: ripicola Cameron, 1926.

solarii Gridelli, 1924d: 89, 96, 103, 173 (Quedius; subgenus Raphirus; Type locality: Alpi occidentali: Alpi Marit.: Nava, Mte. Fronte; Val pesio; Massif de l'Authion. Alpi Cozie: Crissolo).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • — Coiffait, 1963a: 375 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 206 (Quedius; subgenus Sauridus; characters; Alps).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; Italy).

  • Distribution: France, Italy.

solitarius Casey, 1915, see: sublimbatus Mäklin, 1853.

solskyi Luze, 1904: 99 (Quedius; subgenus Ediquus; Type locality: Jagnob: Kol, Schach-Sara).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; catalog).

  • — Gridelli, 1924d: 72 (Quedius; subgenus Microsaurus; characters; Turkestan).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • Distribution: Tajikistan.

songpan Smetana, 1999e: 547 (Quedius; subgenus Microsaurus; Type locality: China: province de Sinchuan, région de Songpan, 3250 m, Lacs Erdao).

  • Distribution: China.

sparsiventris Bernhauer, 1917h: 92 (Quedius; subgenus Prionidus; Type locality: Brasilien: Ypiranga).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • Distribution: Brazil.

sparsus Casey, 1915, see: oculeus Casey, 1915.

sparsutus Fauvel, 1875, see: sublimbatus Mäklin, 1853.

speciosus Bernhauer, 1917h: 90 (Quedius; Type locality: West-Kolumbien: am Monte Socorro in den West-Kordilleren, 3600 m).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • Distribution: Colombia.

spectabilis Kraatz, 1859: 66 (Quedius; Type locality: India borealis).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; catalog).

  • — Cameron, 1932: 284 (Quedius; subgenus Microsaurus; characters; India).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • — Smetana, 1988: 183 (Quedius; notes).

  • Distribution: India.

speculator Kiesenwetter 1848, see: dubius Heer, 1839.

spelaeus Horn, 1871a: 332 (Quedius; Type locality: Indiana; [Note: See lectotype designation by Smetana, 1971]. Originally cited from Wyandotte Cave in southern Indiana).

  • — Horn, 1878: 158 (Quedius; characters; Indiana; Colorado).

  • — Blatchley, 1910: 370 (Quedius; characters; Indiana).

  • — Casey, 1915: 420 (Quediochrus; characters; Indiana; Colorado).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • — Voris, 1939: 188 (Quedius; larval characters; Indiana; Kentucky).

  • — Hatch, 1957: 230 (Quedius; subgenus Quediochrus; characters; Washington; Oregon).

  • — Graham, 1963: 34 (Quedius; collected from cave; notes; California).

  • — Smetana, 1971: 62 (Quedius; subgenus Microsaurus; lectotype designation: Ind. [Indiana]; characters; Alberta; British Columbia; Nova Scotia; Saskatchewan; California; Colorado; Idaho; Illinois; Indiana; Iowa; Kentucky; Montana; New Mexico; New York; Oregon; Pennsylvania; Virginia; Washington; West Virginia; Wisconsin).

  • — Downie and Arnett, 1996: 390 (Quedius; subgenus Microsaurus; characters; USA).

  • — Bordoni and Oromi, 1998: 1157 (Quedius; facultative cave species).

  • Distribution: Canada, USA.

quadriceps Casey, 1915: 421 (Quediochrus; subspecies of spelaeus; [preoccupied]; Type locality: Manitoba: Aweme; [Note: See lectotype designation by Smetana, 1971]).

  • — Scheerpeltz, 1933: 1462 (Quedius; variety of spelaeus).

  • — Smetana, 1971: 62 (Quedius; lectotype designation; synonym of spelaeus).

aplodontiae Smetana, 1971: 64 (Quedius; subgenus Microsaurus; subspecies of spelaeus; Type locality: Oregon: Shotpouch Cr. nr. Knights of Pythias Cp.).

spinipes Sharp, 1884: 337 (Quedionuchus; Type locality: Guatemala, Totonicapam 8500 to 10,500 feet; [Note: See lectotype designation by Smetana, 1976a]. Also cited from Guatemala, Quiche Mountains 7000 to 9000; Totonicapam 8500 to 10,500 feet; Mexico, Las Vigas).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • — Smetana, 1976a: 241 (Quedius; subgenus Quedionuchus; lectotype designation: Totonicapam 8500 to 10,500 feet; characters; Guatemala; Mexican record my be erroneous).

  • Distribution: Guatemala, Mexico ?.

splendidus Fauvel, 1878f: 273 (Quedius; Type locality: Nouvelle-Guinée, Hatam).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • — Cameron, 1924: 87 (Quedius; New Guinea).

  • — Cameron, 1931: 363 (Quedius; New Guinea).

  • — Last, 1975: 439 (Quedius; subgenus Sauridus; characters; New Guinea).

  • — Last, 1987: 55 (Quedius; Papua New Guinea).

  • Distribution: New Guinea.

excessus Last, 1975: 439 (Quedius; variety of splendidus; [Note: Nomen nudum (Article 15.2)]; cited from Buana, Sarawaged Range, 3000 ft., 20–30 miles North of Lae).

spurius Lokay, 1921, see: alpestris Heer, 1839.

stenocephalus Lea, 1925: 252 (Quedius; Type locality: Queensland: Mount Tambourine. New South Wales: Kurrajong).

  • — Scheerpeltz, 1933: 1463 (Quedius; catalog).

  • Distribution: Australia.

stevensi Cameron, 1932: 288 (Quedius; subgenus Microsaurus; Type locality: Sikkim: Rungbong Valley).

  • — Scheerpeltz, 1933: 1463 (Quedius; catalog).

  • — Smetana, 1988: 212 (Quedius; subgenus Microsaurus; characters; Sikkim; Nepal).

  • Distribution: India, Nepal.

analoka Smetana, 1975a: 331 (Quedius; subgenus Microsaurus; Type locality: Central Nepal: near Goropani, W. Pokhara).

  • — Coiffait, 1982: 33 (Quedius; Nepal).

  • — Smetana, 1988: 212 (Quedius; synonym of stevensi).

queinneci Coiffait, 1983d: 167 (Quedius; subgenus Sauridus; Type locality: Nepal: Ganesh Himal, entre Salmi et Rupchet, 1500 à 3600 m).

  • — Smetana, 1988: 212 (Quedius; synonym of stevensi).

strenuus Casey, 1915: 401 (Quedius; Type locality: Texas; [Note: See lectotype designation by Smetana, 1971]. Originally cited from Texas: Austin; Arizona: Tucson).

  • — Scheerpeltz, 1933: 1463 (Quedius; catalog).

  • — Smetana, 1971: 132 (Quedius; subgenus Quedius; lectotype designation: Tex [Texas]; characters; Arkansas; California; Kansas; Missouri; Oklahoma; Texas).

  • — Smetana, 1981b: 638 (Quedius; collecting notes; Arizona; Texas).

  • — Smetana, 1990d: 98 (Quedius; Oklahoma).

  • Distribution: USA.

sturanyi Ganglbauer, 1895: 404 (Quedius; subgenus Sauridus; Type locality: östliches Alpengebiet [Tauern, Hochschwab, Wechsel, Bachergebirge]).

  • — Porta, 1907: 136 (Quedius; synonym of robustus).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; synonym of robustus).

  • — Gridelli, 1924d: 110 (Quedius; subgenus Raphirus; characters).

  • — Scheerpeltz, 1933: 1463 (Quedius; catalog).

  • — Smetana, 1962b: 143 (Quedius; subgenus Raphirus; characters; Alps).

  • — Coiffait, 1963a: 377 (Quedius; subgenus Sauridus; characters).

  • — Lohse, 1964: 214 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 280 (Quedius; subgenus Raphirus; Austria; Slovenia; Istria; Croatia).

  • — Scheerpeltz, 1968a: 96 (Quedius; catalog; Austria).

  • — Coiffait, 1978: 211 (Quedius; subgenus Sauridus; characters; Alps).

  • — Tóth, 1984: 133 (Quedius; characters).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Austria).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; Italy).

  • — Herman, 2001: 31 (Quedius; should be maintained as valid pending study of brevipennis).

  • Distribution: Italy, Austria, Slovenia, Croatia.

brevipennis Motschulsky, 1858: 656 (Quedius; [Note: Nomen dubium; this name is older than sturanyi]; Type locality: Alpes de la Carniole).

  • — Ganglbauer, 1895: 404 (Quedius; synonym of ochropterus).

  • — Porta, 1907: 133 (Quedius; synonym of ochropterus).

  • — Scheerpeltz, 1933: 1463 (Quedius; synonym of sturanyi).

  • — Scheerpeltz, 1968a: 96 (Quedius; synonym of sturanyi).

  • — Coiffait, 1978: 211 (Quedius; synonym of sturanyi ?).

  • — Herman, 2001: 31 (Quedius; species may not be synonym of sturanyi).

subalpestris Coiffait, 1963: 382, 399, 409 (Quedius; subgenus Sauridus; Type locality: Alpes-Maritimes: Saint-Martin-Lantosque).

  • — Coiffait, 1967e: 419 (Quedius; subgenus Microquedius).

  • — Coiffait, 1978: 199 (Quedius; subgenus Microquedius; characters; Alps).

  • — Bordoni, 1985b: 249 (Quedius; subgenus Microquedius; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; Italy).

  • Distribution: France, Italy.

subapicalis Joy, 1913, see: aetolicus Kraatz, 1858.

subapterus Cameron, 1950: 23 (Quedius; Type locality: Three Kings Island).

  • Distribution: New Zealand.

subfuliginosus Britten, 1944, see: fuliginosus Gravenhorst, 1802.

sublateralis Reitter, 1909 [Note: Nomen nudum], see: fuliginosus Gravenhorst, 1802.

sublimbatus Coiffait, 1967, see: suturalis Kiesenwetter, 1845.

sublimbatus Mäklin, 1853: 190 (Quedius; [Note: Motschulsky cited as author]; Type locality: insula Kadjak; [Note: See lectotype designation by Smetana, 1971]. Also cited from Sibiria orientalis).

  • — Horn, 1878: 159 (Quedius; characters; Alaska; Lake Superior region).

  • — Blatchley, 1910: 371 (Quedius; characters; Indiana).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • — Fall, 1926: 150 (Quedius; Alaska).

  • — Scheerpeltz, 1933: 1463 (Quedius; catalog).

  • — Hatch, 1957: 229 (Quedius; subgenus Raphirus; characters; British Columbia; Washington; Oregon).

  • — Smetana, 1965: 47 (Quedius; subgenus Raphirus; characters; notes; Labrador; Newfoundland).

  • — Smetana, 1965a: 16 (Quedius; subgenus Raphirus; notes; British Columbia; Yukon Territory; Alaska).

  • — Smetana, 1967d: 216 (Quedius; subgenus Raphirus; Mongolia).

  • — Smetana, 1971: 219 (Quedius; subgenus Raphirus; lectotype designation: Kadjak; characters; Alberta; British Columbia; Labrador; Manitoba; Newfoundland; Northwest Territories; Quebec; Yukon Territory; Alaska; Montana; New Hampshire; New York).

  • — Smetana, 1973: 1430 (Quedius; notes; Alberta; Newfoundland; Northwest Territories; Saskatchewan).

  • — Smetana 1975c: 176 (Quedius; Mongolia).

  • — Smetana, 1976d: 26 (Quedius; subgenus Raphirus; Siberia).

  • — Smetana, 1978a: 830 (Quedius; subgenus Raphirus; collecting notes; Alberta; Labrador; Newfoundland; Quebec; Yukon Territory).

  • — Smetana, 1978b: 86 (Quedius; subgenus Raphirus; Siberia).

  • — Smetana, 1981b: 641 (Quedius; Saskatchewan).

  • — Smetana, 1990d: 101 (Quedius; South Dakota).

  • — Smetana, 1995a: 84 (Quedius; subgenus Raphirus; notes; Russia).

  • — Downie and Arnett, 1996: 393 (Quedius; subgenus Raphirus; cited as sublimatus; characters; USA).

  • — Ryabukhin, 1999: 75 (Quedius; subgenus Raphirus; catalog; habitat; distribution; northeastern Russia).

  • Distribution: USA, Canada, Russia, Mongolia, Europe.

sparsutus Fauvel, 1875a: XXXIV [= 1875b: 236] (Quedius; Type locality: région du Baïkal, Irkutsk).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; catalog).

  • — Gridelli, 1925: 124 (Quedius; subgenus Raphirus; characters; Baikal region).

  • — Bernhauer, 1926g: 2 (Quedius; cited as sparsus; Kamchatka region).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • — Coiffait, 1967e: 412 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 250 (Quedius; subgenus Sauridus; cited as valid species; characters; Siberia).

  • — Smetana, 1978b: 87 (Quedius; may be synonym of sublimbatus).

  • — Smetana, 1995a: 84 (Quedius; synonym of sublimbatus).

lederi Bernhauer, 1902d: 699 (Quedius; Type locality: Baikalgebiet).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; catalog).

  • — Gridelli, 1922b: 129 (Quedius; probably synonym of sparsutus; characters).

  • — Gridelli, 1925: 124 (Quedius; synonym of sparsutus).

  • — Scheerpeltz, 1933: 1462 (Quedius; synonym of sparsutus).

  • — Coiffait, 1978: 205 (Quedius; subgenus Sauridus; characters; central Asia).

  • — Smetana, 1995a: 85 (Quedius; synonym of sublimbatus).

orbiceps Casey, 1915: 418 (Raphirus; Type locality: Montana: Mullan).

  • — Scheerpeltz, 1933: 1454 (Quedius; catalog).

  • — Smetana, 1971: 219 (Quedius; synonym of sublimbatus).

solitarius Casey, 1915: 419 (Raphirus; Type locality: Not cited).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • — Smetana, 1971: 219 (Quedius; synonym of sublimbatus).

arcticus Munster, 1921: 57 (Quedius; subgenus Sauridus; Type locality: Norvegia arctica [many localities]).

  • — Munster, 1923: 198 (Quedius; subgenus Sauridus; Fennoscandia).

  • — Gridelli, 1925: 123 (Quedius; subgenus Raphirus; characters; Norway).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Coiffait, 1963a: 387 (Quedius; subgenus Sauridus; cited as articus; characters).

  • — Smetana, 1963c: 300 (Quedius; subgenus Raphrius; Mongolia).

  • — Palm, 1963b: 148 (Quedius; subgenus Raphirus; characters; Sweden; Norway; Finland).

  • — Smetana, 1965a: 16 (Quedius; synonym of sublimbatus).

  • — Smetana, 1967d: 216 (Quedius; synonym of sublimbatus).

  • — Smetana, 1971: 219 (Quedius; synonym of sublimbatus).

  • — Coiffait, 1978: 247 (Quedius; subgenus Sauridus; cited as valid species; characters; Scandinavia).

  • — Smetana, 1995a: 85 (Quedius; synonym of sublimbatus).

wuorentausi Bernhauer, 1927b: 96 (Quedius; Type locality: Jenisseisk).

  • — Scheerpeltz, 1933: 1465 (Quedius; catalog).

  • — Smetana, 1971: 219 (Quedius; synonym of sublimbatus).

  • — Smetana, 1995a: 85 (Quedius; synonym of sublimbatus).

subniger Normand, 1935 [Note: Nomen nudum], see: ustus Fauvel, 1878.

subnigriceps Coiffait, 1967e: 410 (Quedius; subgenus Sauridus; Type locality: Dalmatie. Also cited from Bosnia-Hercegovina).

  • — Coiffait, 1978: 249 (Quedius; subgenus Sauridus; characters; Dalmatia).

  • — Tóth, 1984: 138 (Quedius; characters).

  • Distribution: Croatia, Bosnia Hercegovina.

subopacus Lea, 1925: 248 (Quedius; Type locality: Tasmania: Launceston).

  • — Scheerpeltz, 1933: 1463 (Quedius; catalog).

  • Distribution: Australia.

subunicolor Korge, 1961a: 81 (Quedius; Type locality: Schweden: Häggenås, Jämtland).

  • — Kraatz, 1857d: 504 (Quedius; cited as unicolor; [Note: Misidentification of subunicolor according to Smetana, 1993b]; characters; Germany).

  • — Palm, 1962: 187 (Quedius; characters; notes).

  • — Palm, 1963b: 142 (Quedius; subgenus Quedius; characters; Sweden; Norway; Finland).

  • — Coiffait, 1978: 193 (Quedius; subgenus Quedius; characters; Scandinavia).

  • — Lohse and Lucht, 1989: 172 (Quedius; note).

  • — Smetana, 1993b: 50 (Quedius; subgenus Quedius; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; “unicolor Auct. nec Kiesenwetter” cited as synonym; Italy).

  • Distribution: Finland, Sweden, Norway.

sugai Uéno and Watanabe, 1966: 333 (Quedius; subgenus Microsaurus; Type locality: Japan: central Honshu: Yamanashi Prefecture: a lava cave called “Nishinoumi-nokomori-ana” or “Saiko-komori-ana”, on the southwestern coast of Lake Saiko or Lake Nishinoumi, in Ashiwada-mura).

  • — Watanabe, 1996a: 14 (Quedius; cave collection; Japan).

  • Distribution: Japan.

sulcicollis Fauvel, 1877: 269 (Quedius; Type locality: Australie septentrionale et orientale: Cap York, Sydney).

  • — Fauvel, 1878g: 552 (Quedius; characters; Australia).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

sundar Smetana, 1988: 258 (Quedius; subgenus Raphirus; Type locality: Nepal: Khandbari District, For. NE Kuwapani, 2500 m).

  • Distribution: Nepal.

sungkangensis Hayashi, 1992, see: beesoni Cameron, 1932.

suramensis Eppelsheim, 1880c: 580 (Quedius; variety of transcaucasicus; Type locality: Suramgebirge; [Note: See lectotype designation by Gusarov, 1993]).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; variety of vicinus).

  • — Gridelli, 1924d: 34 (Quedius; variety of vicinus).

  • — Gridelli, 1924d: 34 (Quedius; subgenus Microsaurus; cited as vicinus; [Note: Reference refers to suramensis according to information published by Gusarov, 1993: 73]; characters; Caucasus).

  • — Scheerpeltz, 1933: 1465 (Quedius; variety of vicinus).

  • — Scheerpeltz, 1933: 1465 (Quedius; cited as vicinus; [Note: Reference refers to suramensis according to information published by Gusarov, 1993: 73]; catalog).

  • — Gridelli, 1938: 12 (Quedius; variety of vicinus; characters; Circassia; Caucasus; Asia Minor).

  • — Gridelli, 1938: 12 (Quedius; cited as vicinus; [Note: Reference refers to suramensis according to information published by Gusarov, 1993: 73]; characters; Circassia; Caucasus; Asia Minor).

  • — Korge, 1964: 117 (Quedius; cited as vicinus; [Note: Reference refers to suramensis according to information published by Gusarov, 1993: 73]; Turkey).

  • — Korge, 1971a: 44 (Quedius; aberration of vicinus; characters).

  • — Korge, 1971a: 43 (Quedius; subgenus Microsaurus; cited as vicinus; [Note: Reference refers to suramensis according to information published by Gusarov, 1993: 73]; characters; Turkey).

  • — Bordoni, 1976: 236 (Quedius; cited as vicinus; [Note: Reference refers to suramensis according to information published by Gusarov, 1993: 73]; notes; Turkey).

  • — Coiffait, 1978: 178 (Quedius; subgenus Microsaurus; variety of vicinus; characters).

  • — Coiffait, 1978: 178 (Quedius; subgenus Microsaurus; cited as vicinus; [Note: Reference refers to suramensis according to information published by Gusarov, 1993: 73]; characters; Turkey; Caucasus).

  • — Gusarov, 1993: 73 (Quedius; lectotype designation; valid species; transcaucasicus Gemminger and Harold is unavailable, Falderman did not describe it).

  • Distribution: Russia, Georgia, Turkey.

transcaucasicus Gemminger and Harold, 1868: 572 (Quedius; [Note: Nomen nudum (Article 12.1); Gemminger and Harold did not cite this name as a new species or replacement name; they attributed it to Faldermann who described no such species]; cited from Caucasus).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; synonym of vicinus).

  • — Gridelli, 1924d: 34 (Quedius; synonym of vicinus).

  • — Gridelli, 1938: 12 (Quedius; nomen nudum).

  • — Gusarov, 1993: 73 (Quedius; nomen nudum).

suturalis Stephens, 1832, see: Philonthus.

suturalis Kiesenwetter, 1845: 225 (Quedius; [preoccupied]; Type locality: Deutschland).

  • — Kiesenwetter, 1848: 59 (Quedius; characters; Germany).

  • — Redtenbacher, 1849: 711 (Philonthus; characters; Austria).

  • — Baudi, 1857: 100 (Quedius; characters; notes).

  • — Kraatz, 1857d: 511 (Quedius; characters; Germany; Austria).

  • — Redtenbacher, 1857: 204 (Quedius; characters; Austria).

  • — G. Waterhouse, 1858: 23 (Quedius; synonym of humeralis).

  • — Gemminger and Harold, 1868: 570 (Quedius; synonym of humeralis).

  • — Fauvel, 1874: 298 [= 1874a: 524] (Quedius; characters; France; Scandinavia; Russia; Britain; Germany; Switzerland; Austria; Italy; Sardinia).

  • — Redtenbacher, 1874: 201 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 265 (Quedius; characters; Baltic region).

  • — Fauvel, 1876a: 78 [= 1876: 263] (Quedius; Spain).

  • — Mulsant and Rey, 1876: 732 [= 1877: 588] (Quedius; subgenus Sauridus; characters; France).

  • — J. Sahlberg, 1876: 29 (Quedius; Finland).

  • — Fowler, 1888: 241 (Quedius; subgenus Sauridus; characters; Britain).

  • — Ganglbauer, 1895: 407 (Quedius; synonym of humeralis).

  • — Ganglbauer, 1895: 407 (Quedius; subgenus Sauridus; cited as humeralis; [Note: Misidentification of suturalis according to Smetana, 1993b]; characters; north and middle Europe; Sardinia).

  • — Porta, 1907: 140 (Quedius; synonym of humeralis).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; synonym of humeralis).

  • — Gridelli, 1925: 119 (Quedius; synonym of humeralis).

  • — Portevin, 1929: 343 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1443 (Quedius; synonym of humeralis).

  • — V. Hansen, 1952: 148 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Smetana, 1958: 374 (Quedius; synonym of humeralis).

  • — Smetana, 1962b: 146 (Quedius; synonym of humeralis).

  • — Lohse, 1964: 216 (Quedius; synonym of humeralis).

  • — Coiffait, 1967e: 408 (Quedius; subgenus Sauridus; valid species).

  • — Scheerpeltz, 1968a: 94 (Quedius; synonym of humeralis).

  • — Tikhomirova, 1973a: 190 (Quedius; in checklist for USSR).

  • — Coiffait, 1978: 239 (Quedius; subgenus Sauridus; characters; Europe; Scandinavia; Turkey).

  • — Muona, 1979: 16 (Quedius; synonym of humeralis).

  • — Bordoni, 1982b: 23 (Quedius; Italy).

  • — Tóth, 1984: 139 (Quedius; characters; Hungary).

  • — Biström and Silfverberg, 1985: 145 (Quedius; correction for Enumeratio Coleopterorum Fennoscandiae et Daniae).

  • — Segers, 1986: 51 (Quedius; Belgium).

  • — Drugmand, 1987b: 61 (Quedius; distribution in Belgium).

  • — Lohse and Lucht, 1989: 174 (Quedius; characters).

  • — K. Koch, 1989: 322 (Quedius; habitat).

  • — Silfverberg, 1992: 18 (Quedius; Sweden; Denmark; Estonia; Lithuania).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Gerardi and Zanetti, 1995: 144 (Quedius; collecting notes; Italy).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; “humeralis Auct. nec Stephens” cited as synonym; Italy).

  • — Franzen, 1996: 207 (Quedius; collecting notes; Germany).

  • — M. Hansen, 1996: 107 (Quedius; checklist; Denmark).

  • — Ádám, 1996: 248 (Raphirus; Hungary).

  • — Ádám, 1996a: 25 (Raphirus; Hungary).

  • — Böhme, 1996: 48 (Quedius; humus collection; Germany; Austria).

  • — Eisinger, 1997: 171 (Quedius; collecting notes; Germany).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (Quedius; Denmark).

  • — Assing, Frisch, Kahlen, et al., 1998: 139 (Quedius; note).

  • — Köhler, 1998: 160 (Quedius; habitat notes; Germany).

  • — Marchal and Chardonnet, 2000: 72 (Quedius; Corsica).

  • — Herman, 2001: 49 (Quedius; nomen protectum).

  • Distribution: Europe, Russia, Turkey.

fraternus Bernhauer, 1914d: 9 (Quedius; subspecies of humeralis; Type locality: Albanien: Orhanie bei Uesküb).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; variety of humeralis).

  • — Gridelli, 1922b: 129 (Quedius; subgenus Sauridus; subspecies of humeralis; characters; Albania).

  • — Bernhauer, 1923d: 178 (Quedius; subspecies of humeralis; characters).

  • — Gridelli, 1925: 119 (Quedius; variety of humeralis).

  • — Scheerpeltz, 1933: 1443 (Quedius; variety of humeralis).

  • — Coiffait, 1978: 240 (Quedius; subgenus Sauridus; variety of suturalis; characters).

anatolicus Korge, 1964: 119 (Quedius; subgenus Sauridus; subspecies of humeralis; Type locality: Abant-Gebirge).

  • — Coiffait, 1978: 240 (Quedius; subgenus Sauridus; subspecies of suturalis; characters).

sublimbatus Coiffait, 1967e: 413 (Quedius; subgenus Sauridus; [preoccupied]; Type locality: environs de Vienne, Autriche).

  • — Pototskaia, 1967: 83, 90 (Quedius; larval characters).

  • — Coiffait, 1978: 253 (Quedius; synonym of paralimbatus).

paralimbatus Coiffait, 1969: 54 (Quedius; subgenus Sauridus; replacement name for sublimbatus Coiffait).

  • — Coiffait, 1978: 253 (Quedius; subgenus Sauridus; characters; Austria).

  • — Tóth, 1984: 139 (Quedius; characters).

  • — Lohse and Lucht, 1989: 172 (Quedius; characters; Austria).

  • — Bruge, Haghebaert, and Drugmand, 1991: 48 (Quedius; characters; notes; Belgium).

  • — Assing, Frisch, Kahlen, et al., 1998: 139 (Quedius; synonym of suturalis).

suzukii Watanabe, 1987: 328 (Quedius; subgenus Microsaurus; Type locality: Japan: Honshu: Gifu Pref:, Mt. Momisawa-dake on the northern Japanese Alps, 2600 m).

  • Distribution: Japan.

svanetianus Coiffait, 1969: 53 (Quedius; subgenus Raphirus; Type locality: Mestia, 2400 m, Svanetia, Caucase).

  • — Coiffait, 1978: 273 (Quedius; subgenus Raphirus; characters; Caucasus).

  • — Drugmand, 1988c: 204 (Quedius; characters).

  • Distribution: Georgia.

syh Smetana, 1995: 38 (Quedius; subgenus Microsaurus; Type locality: Taiwan: Chiai Hsien, Yushan N. P., Mun-Li Cliff, 2700 m).

  • Distribution: Taiwan.

syphax Smetana, 1973: 1423 (Quedius; subgenus Megaquedius; Type locality: Washington: Kooskooskie Walla Walla).

  • Distribution: USA.

szechuanus Bernhauer, 1933: 41 (Quedius; subgenus Microsaurus; Type locality: Szetschwan: Tatsienlu Tjiji, Urwald-Rodungen).

  • — Smetana, 1996f: 115 (Quedius; subgenus Microsaurus; characters; China).

  • Distribution: China.

tadjikiscus Coiffait, 1975: 32 (Quedius; subgenus Microsaurus; Type locality: Tadjikabad, Daran-Nazaran, R.S.S. de Tadjikistan).

  • — Coiffait, 1978: 149 (Quedius; subgenus Microsaurus; characters; Tadjikistan).

  • Distribution: Tajikistan.

tahomae Hatch, 1957: 220 (Quedius; subgenus Microsaurus; Type locality: Washington: Mt. Rainier, Paradise Valley).

  • — Smetana, 1971: 102 (Quedius; subgenus Microsaurus; characters; Washington).

  • — Smetana, 1976c: 170 (Quedius; subgenus Microsaurus; British Columbia).

  • — Smetana, 1981b: 634 (Quedius; Oregon).

  • Distribution: USA, Canada.

taieriensis Broun, 1894, see: Quediocafius.

taiwanensis Shibata, 1986: 173 (Quedius; subgenus Raphirus; Type locality: Taiwan: Nantou Hsien, near Piluchi, about 2300 m alt.).

  • — Smetana, 1995: 106 (Quedius; subgenus of Raphirus; characters; bionomics; Taiwan).

  • Distribution: Taiwan.

talparum Sainte-Claire Deville, 1910, see: puncticollis Thomson, 1867.

tanderi Smetana, 1988: 222 (Quedius; subgenus Microsaurus; Type locality: Nepal: Prov. Bagmati, Jangtang Ridge, NE Barahbise, 3250 m).

  • Distribution: Nepal.

taprobanus Cameron, 1918, see: Philonthus.

tarng Smetana, 1995: 70 (Quedius; subgenus Raphirus; Type locality: Taiwan: Chiai Hsien, Alishan, Sister Ponds, 2180 m).

  • Distribution: Taiwan.

taruni Smetana, 1988: 271 (Quedius; subgenus Raphirus; Type locality: Nepal: Rasuwa Dis., north slope above Syabru, 3800 m).

  • — Smetana, 1992b: 6 (Quedius; subgenus Raphirus; characters; India; Nepal).

  • Distribution: India, Nepal.

tatricus L/ omnicki, 1874:(14) (Quedius; Type locality: Zakopiańska Magóra w Tatra).

  • — Scheerpeltz, 1933: 1463 (Quedius; catalog).

  • Distribution: Poland.

tauricus Nordmann, 1837, see: boops Gravenhorst, 1802.

tecpatl Smetana, 1976a: 229 (Quedius; subgenus Quedionuchus; Type locality: Mexico: Chiapas: Mt. Tzontehuitz, 9500 ft.).

  • Distribution: Mexico.

temporalis Thomson, 1867, see: mesomelinus Marsham, 1802.

tenellus Gravenhorst, 1806: 54 (Staphylinus; Type locality: Not cited).

  • — Erichson, 1840: 551 (Quedius; characters; Germany).

  • — Kraatz, 1857d: 501 (Quedius; Germany).

  • — Ganglbauer, 1895: 401 (Quedius; subgenus Microsaurus; characters; Lithuania; Finland; Siberia).

  • — Reitter, 1909: 111 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Johansen, 1914: 360 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • — Munster, 1923: 190 (Quedius; subgenus Microsaurus; Fennoscandia).

  • — Gridelli, 1924d: 68 (Quedius; subgenus Microsaurus; characters; Russia; Lithuania; Baikal region; Finland; Germany; Norway).

  • — Scheerpeltz, 1933: 1463 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 17 (Quedius; aedeagal illustration).

  • — Roubal, 1941b: 123, 129 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — V. Hansen, 1952: 138 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Smetana, 1960e: 152, 154 (Quedius; subgenus Microsaurus; characters).

  • — Palm, 1963b: 137 (Quedius; subgenus Microsaurus; characters; Sweden; Norway; Finland).

  • — Lohse, 1964: 207 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Horion, 1965: 269 (Quedius; subgenus Microsaurus; Russia; Finland; Norway; Poland; Germany).

  • — Smetana, 1967d: 215 (Quedius; subgenus Microsaurus; Mongolia).

  • — Korge, 1968: 52 (Quedius; characters; notes).

  • — Coiffait, 1974a: 199 (Quedius; Siberia).

  • — Smetana 1975c: 176 (Quedius; Mongolia).

  • — Coiffait, 1978: 168 (Quedius; subgenus Microsaurus; characters; Europe; Russia; Siberia).

  • — Smetana, 1978b: 85 (Quedius; subgenus Microsaurus; Siberia; notes).

  • — Lucht, 1987: 109 (Quedius; checklist; Poland; Sweden).

  • — Lohse and Lucht, 1989: 171 (Quedius; notes).

  • — Smetana, 1995a: 79 (Quedius; subgenus Microsaurus; notes; Russia).

  • — Ryabukhin, 1999: 71 (Quedius; subgenus Microsaurus; catalog; habitat; distribution; northeastern Russia).

  • Distribution: Europe, Russia, Mongolia.

polystigma Wankowicz, 1867: 253 (Quedius; Type locality: Minsk).

  • — Fauvel, 1874: 285 [= 1874a: 511] (Quedius; characters; Lithuania; Lake Baikal).

  • — Mulsant and Rey, 1876: 684 [= 1877: 540] (Quedius; characters; Lithuania).

  • — J. Sahlberg, 1876: 27 (Microsaurus; Finland).

  • — Ganglbauer, 1895: 401 (Quedius; synonym of tenellus).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; synonym of tenellus).

  • — Gridelli, 1924d: 68 (Quedius; synonym of tenellus).

  • — Coiffait, 1978: 169 (Quedius; synonym of tenellus).

apicipennis Hellén, 1934: 58 (Quedius; aberration of tenellus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Tammerfors).

  • — Roubal, 1941b: 123 (Quedius; subgenus Microsaurus; aberration of tenellus; characters).

  • — Smetana, 1960e: 152, 154 (Quedius; aberration of tenellus; characters).

  • — Coiffait, 1978: 169 (Quedius; subgenus Microsaurus; variety of tenellus; characters).

tepperi Blackburn, 1888: 6 (Quedius; Type locality: Mount Lofty).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • — Lea, 1925: 241 (Quedius; characters; Australia).

  • — Scheerpeltz, 1933: 1463 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

ferox Blackburn, 1888c: 66 (Quedius; [preoccupied]; Type locality: near Adelaide).

  • — Lea, 1925: 241 (Quedius; synonym of tepperi).

  • — Scheerpeltz, 1933: 1463 (Quedius; synonym of tepperi).

blackburni Bernhauer and Schubert, 1916: 419 (Quedius; replacement name for ferox Blackburn).

  • — Lea, 1925: 241 (Quedius; synonym of tepperi).

  • — Scheerpeltz, 1933: 1463 (Quedius; synonym of tepperi).

terminatus Melsheimer, 1844: 39 (Quedius; Type locality: Pennsylvania).

  • — Horn, 1878: 160 (Quedius; synonym of peregrinus).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1456 (Quedius; synonym of peregrinus).

  • — Smetana, 1971: 55 (Quedius; subgenus Microsaurus; valid species; characters; Alabama; Illinois; Indiana; Kentucky; Missouri; New York; North Carolina; Pennsylvania).

  • — Smetana, 1978a: 823 (Quedius; subgenus Microsaurus; collecting notes; Virginia).

  • — Smetana, 1981b: 633 (Quedius; Ontario).

  • — Smetana, 1990d: 96 (Quedius; Arkansas).

  • — Downie and Arnett, 1996: 391 (Quedius; subgenus Microsaurus; characters; USA).

  • Distribution: USA.

tetrapunctatus Coiffait, 1977c: 136 (Quedius; subgenus Microsaurus; replacement name for quadripunctatus Iablokov-Khnzorian).

  • — Coiffait, 1978: 154 (Quedius; subgenus Microsaurus; characters; Armenia).

  • Distribution: Armenia.

quadripunctatus Iablokov-Khnzorian, 1961: 144 (Quedius; [preoccupied]; Type locality: Armenie du Nord: Shagali).

  • — Coiffait, 1969: 44 (Quedius; subgenus Microsaurus; characters; Caucasus).

  • — Coiffait, 1978: 154 (Quedius; synonym of tetrapunctatus).

tetrastigma Leinberg, 1900, see: umbrinus Erichson, 1839.

tetricus Smetana, 1971: 169 (Quedius; subgenus Distichalius; Type locality: British Columbia: 12 mi. E. Hope. Also cited from Oregon and Washington).

  • — Smetana, 1976c: 174 (Quedius; subgenus Distichalius; British Columbia; Oregon; Washington).

  • — Smetana, 1978a: 827 (Quedius; subgenus Distichalius; collecting notes; British Columbia; Washington).

  • Distribution: Canada, USA.

thoracicus Fauvel, 1877: 271 (Quedius; Type locality: Australie, Sydney, Rockhampton).

  • — Fauvel, 1878g: 553 (Quedius; characters; Australia).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

tian Smetana, 1996c: 234 (Quedius; subgenus Raphirus; Type locality: China: Sichuan, 103.20 el/29.30 nw Mt. Emei, 500–1200 m).

  • Distribution: China.

tibetanus Boháč, 1988: 555 [= 1989: 39] (Quedius; subgenus Quediops; Type locality: Vost. Tibet, bassein Goluboi reki, r. Go-Chiu, 13,500 feet).

  • Distribution: China.

tikta Smetana, 1975a: 339 (Quedius; subgenus Raphirus; Type locality: Central Nepal: Between Tare-Pati and Gosaikunde).

  • — Smetana, 1988: 266 (Quedius; subgenus Raphirus; characters; Nepal).

  • Distribution: Nepal.

tinctellus Broun, 1910: 14 (Quedius; Type locality: Waimarino).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • Distribution: New Zealand.

tochtli Smetana, 1975: 312 (Quedius; subgenus Microsaurus; Type locality: Mexico: Pueblo: Km. 28, 13,000 ft., V. Popocatepetl).

  • — Smetana, 1976: 117 (Quedius; subgenus Microsaurus; Mexico).

  • Distribution: Mexico.

tokiensis Bernhauer, 1913: 132 (Quedius; Type locality: Japan, Tokio).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • Distribution: Japan.

tomentosomaculatus Cornelius, 1863, see: auricomus Kiesenwetter, 1850.

tonglu Smetana, 1988: 268 (Quedius; subgenus Raphirus; Type locality: Nepal: Khandbari District, above Tashigaon, 3500 m).

  • — Smetana, 1992b: 5 (Quedius; subgenus Raphirus; India; Nepal).

  • Distribution: India, Nepal.

transcaucasicus Gemminger and Harold, 1868 [Note: Nomen nudum], see: suramensis Eppelsheim, 1880.

transcaucasicus Iablokov-Khnzorian, 1961: 146 (Quedius; Type locality: Schischkert, Armenie orientale, 1900 m).

  • — Coiffait, 1978: 184 (Quedius; synonym of iablokofi).

  • — Herman, 2001: 30 (Quedius; valid name for species).

  • Distribution: Armenia.

iablokofi Coiffait, 1967e: 396 (Quedius; subgenus Microsaurus; replacement name for transcaucasicus Iablokov-Khnzorian; [Note: Unnecessary name because the senior names, transcaucasicus Falderman and transcaucasicus Gemminger and Harold, are unavailable]).

  • — Coiffait, 1978: 184 (Quedius; subgenus Microsaurus; characters; Armenia).

  • — Herman, 2001: 30 (Quedius; synonym of transcaucasicus).

transparens Motschulsky, 1845b: 358 (Quedius; Type locality: Californie; [Note: See lectotype designation by Smetana, 1976b]).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1463 (Quedius; catalog).

  • — Smetana, 1976b: 59 (Quedius; subgenus Distichalius; lectotype designation; valid species).

  • — Smetana, 1978a: 826 (Quedius; subgenus Distichalius; valid species).

  • Distribution: USA, Canada.

marginalis Mäklin, 1852: 315 (Quedius; Type locality: insula Sitkha; [Note: See lectotype designation by Smetana, 1971]).

  • — Horn, 1878: 161 (Quedius; variety of capucinus; characters).

  • — Casey, 1915: 405 (Distichalius; characters; Alaska; Queen Charlotte Islands; British Columbia to California).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1447 (Quedius; catalog).

  • — Hatch, 1957: 217 (Quedius; subgenus Distichalius; characters; British Columbia; Washington; Oregon).

  • — Smetana, 1971: 156 (Quedius; subgenus Distichalius; lectotype designation; characters; Alberta; British Columbia; Alaska; California; Nevada; Oregon; Washington).

  • — Smetana, 1971b: 3 (Quedius; subgenus Distichalius; California; British Columbia).

  • — Smetana, 1976b: 60 (Quedius; synonym of transparens).

  • — Smetana, 1978a: 826 (Quedius; synonym of transparens).

pugetensis Hatch, 1957: 218 (Quedius; subgenus Distichalius; Type locality: Washington: King Co.: White River, Slippery Creek. Also cited from Oregon and British Columbia).

  • — Smetana, 1971: 156 (Quedius; synonym of marginalis).

  • — Smetana, 1976b: 60 (Quedius; synonym of transparens).

  • — Smetana, 1978a: 826 (Quedius; synonym of transparens).

transsylvanicus Weise, 1875: 356 (Quedius; Type locality: Hoverla und Mencil).

  • — Ganglbauer, 1895: 405 (Quedius; subgenus Sauridus; cited as transsilvanicus; characters; Europe).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • — Gridelli, 1924d: 105 (Quedius; subgenus Raphirus; cited as transylvanicus; characters).

  • — Scheerpeltz, 1933: 1463 (Quedius; cited as transsilvanicus; catalog).

  • — Smetana, 1958: 369 (Quedius; subgenus Raphirus; cited as transsilvanicus; characters; ecology).

  • — Smetana, 1962b: 139 (Quedius; subgenus Raphirus; characters; Europe).

  • — Coiffait, 1963a: 377 (Quedius; subgenus Sauridus; characters).

  • — Lohse, 1964: 214 (Quedius; subgenus Raphirus; cited as transsilvanicus; characters; central Europe).

  • — Coiffait, 1978: 229 (Quedius; subgenus Sauridus; characters; Carpathians).

  • — Tóth, 1984: 133 (Quedius; characters).

  • — Smetana, 1995a: 84 (Quedius; subgenus Raphirus; notes; Ukraine).

  • Distribution: Ukraine.

triangulum Fauvel, 1891: 119 (Quedius; Type locality: Colonie Tovar).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • Distribution: Venezuela.

tripunctatus Cameron, 1945, see: tripunctatus Bernhauer, 1941.

tripunctatus Bernhauer, 1941b: 31 (Quedius; subgenus Sauridus; Type locality: Greymouth).

  • — Herman, 2001: 31 (Quedius; synonymic note).

  • Distribution: New Zealand.

tripunctatus Cameron, 1945d: 787 (Quedius; subgenus Microsaurus; [preoccupied]; Type locality: Not cited, stated to be without locality label; presumably species is from New Zealand).

  • — Herman, 2001: 31 (Quedius; synonym of tripunctatus Bernhauer).

tristis: Gravenhorst, 1802 [Note: Nomen nudum], see: levicollis Brullé, 1832.

trisulensis Coiffait, 1982, see: franzi Smetana, 1975.

troglodytes Uéno and Watanabe, 1966: 327 (Quedius; subgenus Microsaurus; Type locality: Japan: Oinu-ana Cave, at Nakakoshi-zawa in Ueno-mura).

  • — Uéno and Watanabe, 1970: 18 (Quedius; subgenus Microsaurus; notes; Japan).

  • — Watanabe, 1996a: 14 (Quedius; cave collection; Japan).

  • — Bordoni and Oromi, 1998: 1157 (Quedius; subgenus Microsaurus; facultative cave species).

  • Distribution: Japan.

troglophilus Coiffait, 1969: 46 (Quedius; subgenus Sauridus; Type locality: Grotte Bankovica, Karlukovo, Bulgarie).

  • — Coiffait, 1971b: 383 (Quedius; Bulgaria).

  • — Coiffait, 1978: 239 (Quedius; subgenus Sauridus; characters; Bulgaria).

  • Distribution: Bulgaria.

tronqueti Smetana, 1999b: 238 (Quedius; subgenus Microsaurus; Type locality: China: province du Sinchuan [sic], massif du Jiu Ding-Shan, 3300–3500 m).

  • Distribution: China.

troodites Fagel, 1968b: 7 (Quedius; subgenus Sauridus; Type locality: Chypre: Mt. Troodos).

  • — Coiffait, 1978: 219 (Quedius; subgenus Sauridus; characters; Cyprus).

  • — Assing and Wunderle, 2001: 37 (Quedius; Cyprus).

  • Distribution: Cyprus.

truncicola Fairmaire and Laboulbène, 1856: 538 (Quedius; Type locality: Fontainebleau).

  • — Fauvel, 1874: 281 [= 1874a: 507] (Quedius; synonym of ventralis).

  • — Ganglbauer, 1895: 399 (Quedius; synonym of ventralis).

  • — Porta, 1907: 109 (Quedius; synonym of ventralis).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; synonym of ventralis).

  • — Smetana, 1958: 339 (Quedius; synonym of ventralis).

  • — Bordoni, 1974: 52 (Quedius; synonym of ventralis).

  • — Muona, 1977: 15 (Quedius; valid name for ventralis).

  • — Pope, 1977: 31 (Quedius; synonym of ventralis).

  • — Coiffait, 1978: 168 (Quedius; cited as truncicula; synonym of ventralis).

  • — Lohse and Lucht, 1989: 171 (Quedius; note).

  • — Wittwer, 1993: 251 (Quedius; cited as valid species; Switzerland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; Italy).

  • — Sörensson, 1996 (Quedius; natural history and conservation notes; Sweden).

  • Distribution: Europe.

ventralis Aragona, 1830: 12 (Staphylinus; [preoccupied]; Type locality: Valle Ticini).

  • — Lacordaire, 1835: 378 (Emus; [Note: According to Bernhauer and Schubert, 1916: 436, “Emus nitidus variety a” cited by Boisduval and Lacordaire is a synonym of ventralis Aragona]; characters; France).

  • — Erichson, 1840: 526 (Quedius; [Note: Misidentification: Bernhauer and Schubert, 1916: 436, cited fulgidus variety 3 Erichson as a synonym of ventralis; Erichson did not describe the species as new and the variety is unavailable]; characters).

  • — Fauvel, 1867: LIV (Quedius; notes).

  • — Fauvel, 1874: 281 [= 1874a: 507] (Quedius; characters; France; Britain; Italy).

  • — Mulsant and Rey, 1876: 638 [= 1877: 484] (Quedius; characters; France).

  • — Fowler, 1888: 231 (Quedius; characters; Britain).

  • — Ganglbauer, 1895: 399 (Quedius; subgenus Microsaurus; characters; Germany; Italy; France; England).

  • — Porta, 1907: 109 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Reitter, 1909: 109 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Johansen, 1914: 357 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; catalog).

  • — Gridelli, 1924d: 26 (Quedius; subgenus Microsaurus; characters; France; England; Pyrenees; Germany; Italy).

  • — Portevin, 1929: 339 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1464 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 2 (Quedius; aedeagal illustration).

  • — Roubal, 1941b: 122, 126 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — V. Hansen, 1952: 126 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Smetana, 1958: 339 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 151 (Quedius; subgenus Microsaurus; characters).

  • — Palm, 1963b: 128 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark).

  • — Lohse, 1964: 207 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Horion, 1965: 253 (Quedius; subgenus Microsaurus; Britain; Denmark; Sweden; Norway; Finland; Russia; Holland; Belgium; France; Germany; Austria; Spain; Corsica; Sicily; Italy; Romania; Hungary; Czechoslovakia; Poland).

  • — Bordoni, 1974: 50 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 168 (Quedius; subgenus Microsaurus; characters; Europe; Britain; Scandinavia; Denmark; Sicily; Corsica; Iberian Peninsula).

  • — Hinton, 1981: 663 (Quedius; characters of egg).

  • — Frank, 1982: 47 (Quedius; list of parasites).

  • — Tóth, 1984: 106 (Quedius; characters; Hungary).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; France; Benelux; Denmark; Sweden).

  • — Drugmand, 1989: 177 (Quedius; characters).

  • — Lohse and Lucht, 1989: 171 (Quedius; synonym of truncicola).

  • — Smetana, 1993b: 50 (Quedius; synonym of truncicola).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; synonym of truncicola).

  • — Fowles, Alexander, and Key, 1999: 129 (Quedius; saproxylic species; status as endangered or threatened species; Britain).

rufipennis Trella, 1930: 33 (Quedius; aberration of ventralis; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]; cited from Mazury).

  • — Scheerpeltz, 1933: 1465 (Quedius; aberration of ventralis).

  • — Roubal, 1941b: 122 (Quedius; subgenus Microsaurus; aberration of ventralis; characters).

  • — Smetana, 1958: 339 (Quedius; aberration of ventralis).

  • — Smetana, 1960e: 151 (Quedius; aberration of ventralis).

  • — Bordoni, 1974: 52 (Quedius; aberration of ventralis).

  • — Coiffait, 1978: 168 (Quedius; subgenus Microsaurus; variety of ventralis; characters).

  • — Tóth, 1984: 106 (Quedius; variety of ventralis).

tschinganensis Coiffait, 1969: 50 (Quedius; subgenus Sauridus; Type locality: Monts Tschingan, environ 1200 m, Ouzbekistan, U.R.S.S.).

  • — Coiffait, 1970d: 143 (Quedius; Uzbekistan).

  • — Coiffait, 1978: 237 (Quedius; subgenus Sauridus; characters; Uzbekistan).

  • Distribution: Uzbekistan.

gracilicornis Coiffait, 1977c: 139 (Quedius; subgenus Sauridus; variety of tschinganensis; [Note: Nomen nudum (Article 15.2); preoccupied]; cited from Buchara, Karatak).

  • — Coiffait, 1978: 237 (Quedius; subgenus Sauridus; variety of tschinganensis; cited as debilicornis; characters).

tsurugiensis Watanabe and Yoshida, 1970: 4 (Quedius; subgenus Microsaurus; Type locality: Japan: Shikoku: Tokushima Pref.: near Meoto-ike, about 1300 m alt., in Mt. Tsurugi-san).

  • Distribution: Japan.

turkmenicus Coiffait, 1969: 49 (Quedius; subgenus Sauridus; Type locality: Kouchka Agachle, Turkménie).

  • — Coiffait, 1978: 245 (Quedius; subgenus Sauridus; characters; Turkmenia).

  • Distribution: Turkmenistan.

tylophilus Scheerpeltz, 1933 [Note: Nomen nudum], see: xylophilus Lea, 1908.

turnai Smetana, 1999b: 223 (Quedius; subgenus Microsaurus; Type locality: China: “E Tibet, Tuntala Shan” mts., Road Markam-Zogang, 40 km Zogang, 4500 m, 29°40′N, 98°08′E).

  • Distribution: China.

uchikawai Watanabe, 1981: 19 (Quedius; subgenus Microsaurus; Type locality: Japan: Central Honshu: Nagano Pref.: Kaza-ana Cave, Ryu'unji-yama, Minamimajino, Konami, Suwa-shi).

  • — Watanabe, 1996a: 15 (Quedius; cave collection; Japan).

  • Distribution: Japan.

udagra Smetana, 1975a: 335 (Quedius; subgenus Raphirus; Type locality: Central Nepal: Phulchoki near Kathmandu).

  • — Smetana, 1988: 259 (Quedius; subgenus Raphirus; characters; Nepal).

  • Distribution: Nepal.

uenoi Watanabe, 1979: 12 (Quedius; subgenus Microsaurus; Type locality: Japan: Central Honshu: Hideo-kô Adit of Umegashima Mine, Sujin-zawa, Sekinosawa, Umegashima, Shizuoka-shi).

  • — Watanabe, 1996a: 15 (Quedius; mine collection; Japan).

  • Distribution: Japan.

ulrichi Korge, 1971a: 41 (Quedius; subgenus Microsaurus; Type locality: Pass, 2300 m, zwischen Ispir und Ovacik, in Luftlinie ca. 60 km nordöstlich von Erzurum).

  • — Coiffait, 1978: 185 (Quedius; subgenus Microsaurus; characters; Turkey).

  • Distribution: Turkey.

uludaghensis Drugmand, 1988d: 263 (Quedius; subgenus Microsaurus; Type locality: Anatolie occidentale: Uludagh, 1800–1900 m).

  • — Herman, 2001: 31 (Quedius; synonymic note).

  • Distribution: Turkey.

uludaghensis Drugmand, 1989: 174 (Quedius; subgenus Microsaurus; Type locality: Anatolie occidentale: région de l'Ulu Dagh, 1800–1900 m).

  • — Herman, 2001: 31 (Quedius; synonym of uludagensis).

uludaghensis Drugmand, 1989, see: uludaghensis Drugmand, 1988.

umbratus Uéno and Watanabe, 1966: 335 (Quedius; subgenus Microsaurus; Type locality: Japan: central Honshu: Tokyo Prefecture: a limestone cave called “Kurasawa-do”, at Kurasawa-dani in Okutama-cho).

  • — Watanabe, 1996a: 15 (Quedius; cave collection; Japan).

  • Distribution: Japan.

umbrinus Erichson, 1839a: 491 (Quedius; Type locality: Neustadt-Eberswalde).

  • — Gyllenhal, 1810: 309 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1916: 436, cited maurorufus Gyllenhal as a synonym of umbrinus Erichson; Gyllenhal did not describe the species as new, he attributed it to Gravenhorst]; characters; Sweden).

  • — Heer, 1839: 277 (Philonthus; characters; Switzerland).

  • — Erichson, 1840: 541 (Quedius; characters; Sweden; Germany).

  • — Gravenhorst, 1847: 228 (Quedius; characters; notes).

  • — Redtenbacher, 1849: 711 (Philonthus; characters; Austria).

  • — Hardy, 1851: 33 (Quedius; synonym of semiobscurus).

  • — Fairmaire and Laboulbène, 1856: 542 (Quedius; characters; France).

  • — Kraatz, 1857d: 509 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 204 (Quedius; characters; Austria).

  • — Kraatz, 1858a: 59 (Quedius; notes; Greece).

  • — Peyron, 1858: 428 (Quedius; Turkey).

  • — Thomson, 1860: 178 (Raphirus; characters; Scandinavia).

  • — Fauvel, 1865a: 293 [= 1865b: 43] (Quedius; habitat; France).

  • — Baudi, 1870: 384 (Quedius; characters).

  • — Fauvel, 1874: 294 [= 1874a: 520] (Quedius; characters; France; Scandinavia; Russia; Britain; Germany; Switzerland; Austria; Italy; Spain; Portugal).

  • — Redtenbacher, 1874: 201 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 264 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 730 [= 1877: 586] (Quedius; subgenus Sauridus; characters; France).

  • — J. Sahlberg, 1876: 28 (Raphirus; Finland).

  • — Fowler, 1888: 240 (Quedius; subgenus Sauridus; characters; Britain).

  • — Ganglbauer, 1895: 406 (Quedius; subgenus Sauridus; characters; Europe).

  • — Fauvel, 1900d: 233 (Quedius; notes).

  • — Poppius, 1905: 9 (Quedius; Orkney Islands).

  • — Porta, 1907: 138 (Quedius; subgenus Sauridus; characters; Italy).

  • — Reitter, 1909: 112 (Quedius; subgenus Sauridus; characters; Germany).

  • — Johansen, 1914: 366 (Quedius; subgenus Sauridus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • — Munster, 1923: 197 (Quedius; subgenus Sauridus; Fennoscandia).

  • — Gridelli, 1925: 113 (Quedius; subgenus Raphirus; characters; Austria; Italy; Transcaucasus).

  • — Portevin, 1929: 342 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1464 (Quedius; catalog).

  • — West, 1937: 37 (Quedius; Faroe Islands).

  • — Wüsthoff, 1938a: fig. 31 (Quedius; aedeagal illustration).

  • — Jeannel and Jarrige, 1949: 379 (Quedius; subgenus Raphirus; cave species; France; Herzegowina).

  • — V. Hansen, 1952: 146 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Lohse, 1958a (Quedius; clarification of identity).

  • — Smetana, 1958: 372 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Larsson and Gígja, 1959: 91 (Quedius; biological notes; Iceland).

  • — Smetana, 1962b: 143 (Quedius; subgenus Raphirus; characters; Europe).

  • — Coiffait, 1963a: 385 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1963a: 385 (Quedius; subgenus Sauridus; [Note: Misidentified as pseudoumbrinus according to Coiffait, 1978: 235]; characters).

  • — Palm, 1963b: 147 (Quedius; subgenus Raphirus; characters; Sweden; Norway; Finland).

  • — Lohse, 1964: 215 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Focarile, 1964: 61 (Quedius; Italy).

  • — Horion, 1965: 281 (Quedius; subgenus Raphirus; Russia; Lapland; North Cape; Scotland; all central Europe; Portugal; Spain; Italy; bosnia; Macedonia; Bulgaria; Greece; Cyprus).

  • — Pototskaia, 1967: 85, 88 (Quedius; larval characters).

  • — Szujecki, 1968a: 736 (Quedius; Poland).

  • — Coiffait, 1970e: 63 (Quedius; Cyprus).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 235 (Quedius; subgenus Sauridus; characters; Europe; Britain; Scandinavia; Portugal; Caucasus).

  • — Bordoni, 1982b: 29 (Quedius; subgenus Sauridus; collected from moss; ecological notes; Italy).

  • — Tóth, 1984: 136 (Quedius; characters; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; Italy).

  • — Assing, 1999b: 39 (Quedius; characters; notes).

  • Distribution: Iceland, Europe, Russia, Caucasus, Turkey.

maritimus J. Sahlberg, 1876: 29 (Raphirus; variety of umbrinus; Type locality: Aland; Helsingfors; Ladoga; Paanajärvi i Kuusamo).

  • — Fauvel, 1900d: 233 (Quedius; synonym of umbrinus).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; variety of umbrinus).

  • — Gridelli, 1925: 113 (Quedius; variety of umbrinus).

  • — Renkonen, 1941: 106 (Quedius; characters; notes; Finland).

  • — V. Hansen, 1952: 145 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Horion, 1954: 11 (Quedius; characters; Germany).

  • — Smetana, 1958: 373 (Quedius; subgenus Raphirus; cited as valid species; characters; ecology).

  • — Löbl, 1961: 921 (Quedius; synonym of umbrinus).

  • — Smetana, 1962b: 143 (Quedius; synonym of umbrinus).

  • — Palm, 1963b: 147 (Quedius; subgenus Raphirus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 215 (Quedius; synonym of umbrinus).

  • — Pope, 1977: 31 (Quedius; synonym of umbrinus).

  • — Coiffait, 1978: 235 (Quedius; synonym of umbrinus).

tetrastigma Leinberg, 1900: 79 (Quedius; variety of umbrinus; Type locality: Helsingfors).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; variety of umbrinus).

  • — Scheerpeltz, 1933: 1464 (Quedius; variety of umbrinus).

pseudoumbrinus Lohse, 1958a: 60 (Quedius; proposed for umbrinus of authors; Type locality: Altvater; [Note: See lectotype designation by Assing, 1999b]. Originally cited from nördliches Skandinavien; Grossbritannien; Mitteleuropa).

  • — Renkonen, 1941: 106 (Quedius; cited as umbrinus; [Note: Misidentification of pseudoumbrinus according to Löbl, 1961]; characters; notes; Finland).

  • — Smetana, 1962b: 143 (Quedius; subgenus Raphirus; characters).

  • — Lohse, 1964: 215 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Pope, 1977: 31 (Quedius; synonym of umbrinus).

  • — Coiffait, 1978: 236 (Quedius; subgenus Sauridus; characters; Scandinavia; Britain; central Europe; central Asia).

  • — Tóth, 1984: 136 (Quedius; characters).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia).

  • — Hodge and Jones, 1995: 42 (Quedius; notes; Britain).

  • — Assing, 1999b: 39 (Quedius; lectotype designation: Altvater; synonym of umbrinus).

  • — Burakowski, Mroczkowski, and Stefańska, 2000: 54 (Quedius; subgenus Raphirus; catalog; Poland).

umbripennis Bernhauer and Schubert, 1916: 422 (Quedius; variety of dubius; [Note: Roubal, 1913a: 111, described this species as an aberration of dubius so that usage is unavailable. Bernhauer and Schubert seem to have been the first to make the name available by citing it as a variety of dubius. The type locality is taken from Roubal's publication and the characters for the species are listed therein]; Type locality: Umgebung von Vladikavkaz, Kaukasus).

  • — Roubal, 1913a: 111 (Quedius; aberration of dubius; [Note: This is the original description for the umbripennis but it was cited as an aberration and is unavailable]; characters; Type locality: Umgebung von Vladikavkaz, Kaukasus).

  • — Gridelli, 1925: 113 (Quedius; subgenus Raphirus; characters).

  • — Scheerpeltz, 1933: 1464 (Quedius; catalog).

  • — Coiffait, 1963a: 383 (Quedius; subgenus Sauridus; cited as imbripennis; characters).

  • — Korge, 1964: 123 (Quedius; subgenus Sauridus; characters; Turkey).

  • — Coiffait, 1978: 236 (Quedius; subgenus Sauridus; characters; Caucasus).

  • Distribution: Russia, Turkey.

unicolor Stephens, 1832, see: Philonthus.

unicolor Kiesenwetter, 1847: 75 (Quedius; [preoccupied]; Type locality: Riesengebirge).

  • — Kiesenwetter, 1848: 53 (Quedius; characters; Germany).

  • — Redtenbacher, 1849: 825 (Philonthus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 536 (Quedius; characters; France).

  • — Redtenbacher, 1857: 202 (Quedius; characters; Austria).

  • — Fauvel, 1874: 291 [= 1874a: 517] (Quedius; characters; France; Germany; Austria).

  • — Redtenbacher, 1874: 199 (Quedius; characters; Austria).

  • — Mulsant and Rey, 1876: 697 [= 1877: 553] (Quedius; subgenus Microsaurus; characters; France).

  • — Ganglbauer, 1895: 403 (Quedius; characters; Sudeten; Alps; Pyrenees).

  • — Porta, 1907: 132 (Quedius; subgenus Quedius; characters; Italy).

  • — Reitter, 1909: 112 (Quedius; characters; Germany).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; catalog).

  • — Munster, 1923: 196 (Quedius; Fennoscandia).

  • — Gridelli, 1924d: 80 (Quedius; subgenus Quedius; characters; Alps; Pyrenees; Sudeten).

  • — Portevin, 1929: 341 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1464 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 24 (Quedius; aedeagal illustration).

  • — Smetana, 1955c: 143 (Quedius; notes; Czechoslovakia).

  • — Smetana, 1958: 363 (Quedius; subgenus Quedius; characters; ecology; Czechoslovakia).

  • — Smetana, 1960b: 259 (Quedius; Czechoslovakia).

  • — Coiffait, 1961a: 49 (Quedius; characters; France; Alps; Andalusia).

  • — Smetana, 1962b: 134 (Quedius; subgenus Quedius; characters; Europe).

  • — Palm, 1962: 187 (Quedius; characters; notes).

  • — Lohse, 1964: 211 (Quedius; subgenus Quedius; characters; central Europe).

  • — Smetana, 1964c: 168 (Quedius; subgenus Quedius; Czechoslovakia).

  • — Havelka, 1964: 93 (Quedius; subgenus Quedius; Slovakia).

  • — Horion, 1965: 274 (Quedius; subgenus Quedius; mountains of central Europe; Black Sea; Czechoslovakia; Poland; Hungary; Romania; Austria; Germany; Switzerland; Italy; France; Pyrenees).

  • — Smetana, 1966: 331 (Quedius; subgenus Quedius; notes; Czechoslovakia).

  • — Scheerpeltz, 1968a: 92 (Quedius; catalog; Austria).

  • — Tikhomirova, 1973a: 189 (Quedius; checklist; USSR).

  • — Shilov, 1975a: 376 [= 1975b: 99] (Quedius; Komi region of Russia).

  • — Bordoni, 1976b: 94 (Quedius; characters; Italy).

  • — Zerche, 1977a: 9 (Quedius; Germany).

  • — Coiffait, 1978: 193 (Quedius; subgenus Quedius; characters; Alps; Pyrenees; Massif Central; mountains of central Europe).

  • — Tóth, 1984: 119 (Quedius; characters).

  • — Segers, 1986: 50 (Quedius; Belgium).

  • — Lucht, 1987: 109 (Quedius; Germany; Poland; Czechoslovakia; Austria; Switzerland; Sweden).

  • — Janák, 1992: 91 (Quedius; Czech Republic).

  • — Herman, 2001: 50 (Quedius; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: Europe, Russia.

atritarsis Roubal, 1949: 44 (Quedius; aberration of unicolor; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Carpathians: High and Low Tatras).

  • — Smetana, 1958: 363 (Quedius; aberration of unicolor).

  • — Smetana, 1962b: 134 (Quedius; aberration of unicolor).

  • — Coiffait, 1978: 193 (Quedius; subgenus Quedius; variety of unicolor; characters).

  • — Tóth, 1984: 119 (Quedius; variety of unicolor; characters).

nigritarsis Roubal, 1949: 45 (Quedius; aberration of unicolor; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied; this name may be a lapsus for atritarsis according to Smetana (personal communication)]).

  • — Smetana, 1958: 363 (Quedius; synonym of unicolor).

  • — Smetana, 1962b: 134 (Quedius; synonym of unicolor).

  • — Coiffait, 1978: 193 (Quedius; synonym of unicolor variety atritarsis).

dolosus Coiffait, 1961a: 49 (Quedius; aberration of unicolor; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Andalousie, Sierra de las Nieves).

  • — Smetana, 1962b: 134 (Quedius; aberration of unicolor).

  • — Coiffait, 1978: 193 (Quedius; subgenus Quedius; variety of unicolor; characters).

  • — Tóth, 1984: 119 (Quedius; variety of unicolor; characters).

uniformis Coiffait, 1967e: 420 (Quedius; subgenus Raphirus; Type locality: Castro Verde, Portugal).

  • — Coiffait, 1978: 277 (Quedius; subgenus Raphirus; characters; Portugal).

  • Distribution: Portugal.

urbanus Broun, 1921: 480 (Quedius; Type locality: Glenhope).

  • — Scheerpeltz, 1933: 1464 (Quedius; catalog).

  • Distribution: New Zealand.

ustus Fauvel, 1878: 129 [= 1878a: 49] (Quedius; Type locality: Alger, dans les alluvions de l'Arrach; Bône; Edough).

  • — Fauvel, 1886: 59 [= 1886a: 51] (Quedius; Algeria).

  • — Fauvel, 1902b: 116 (Quedius; Morocco; Algeria; Tunisia).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; catalog).

  • — Gridelli, 1925: 138 (Quedius; subgenus Raphirus; characters; Algeria; Tunisia; Morocco).

  • — Scheerpeltz, 1933: 1464 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 43 (Quedius; aedeagal illustration).

  • — C. Koch, 1941: 66 (Quedius; characters; notes; Morocco).

  • — Jeannel and Jarrige, 1949: 381 (Quedius; subgenus Raphirus; cave species; Algeria).

  • — Fagel, 1957e: 333 (Quedius; subgenus Sauridus; notes).

  • — Coiffait, 1963a: 381 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1973a: 275 (Quedius; Morocco).

  • — Coiffait, 1978: 243 (Quedius; subgenus Sauridus; characters; North Africa).

  • Distribution: Tunisia, Algeria, Morocco.

subniger Normand, 1935: 373 (Quedius; aberration of ustus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]).

  • — Coiffait, 1978: 243 (Quedius; subgenus Sauridus; variety of ustus; characters).

uteanus Casey, 1915: 415 (Microsaurus; Type locality: Utah, southwestern).

  • — Scheerpeltz, 1933: 1464 (Quedius; catalog).

  • — Smetana, 1971: 117 (Quedius; synonym of caseyi).

  • — Herman, 2001: 29 (Quedius; valid name for species).

  • Distribution: Canada, USA.

curtipennis Casey, 1915: 414 (Microsaurus; [preoccupied]; Type locality: Manitoba: Aweme).

  • — Scheerpeltz, 1933: 1435 (Quedius; synonym of caseyi).

  • — Hatch, 1957: 226 (Quedius; subgenus Microsaurus; characters; British Columbia; Idaho; Washington; Oregon).

divergens Casey, 1915: 415 (Microsaurus; Type locality: California: Siskiyou Co.).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Smetana, 1971: 117 (Quedius; subspecies of caseyi; characters; Alberta; British Columbia; California; Colorado; Idaho; Oregon; Utah).

  • — Smetana, 1978a: 825 (Quedius; subgenus Microsaurus; subspecies of caseyi; collecting notes; Arizona).

  • — Smetana, 1981b: 637 (Quedius; subspecies of caseyi; collecting notes; Arizona; Colorado).

  • — Smetana, 1990d: 98 (Quedius; subspecies of caseyi; Utah; Wyoming).

parvipennis Bernhauer, 1917l: 249 (Quedius; subgenus Sauridus; [Note: This name is older than caseyi]; Type locality: Alberta, Banff).

  • — Scheerpeltz, 1933: 1456 (Quedius; catalog).

  • — Smetana, 1971: 117 (Quedius; synonym of caseyi).

caseyi Scheerpeltz, 1933: 1435 (Quedius; replacement name for curtipennis Casey).

  • — Smetana, 1971: 117 (Quedius; subgenus Microsaurus; nominate subspecies; characters; Alberta; Manitoba; Saskatchewan; Michigan; Montana).

  • — Smetana, 1971a: 1840 (Quedius; subgenus Microsaurus; collecting notes; California).

  • — Smetana, 1971b: 3 (Quedius; subgenus Microsaurus; Washington is an erroneous locality).

  • — Smetana, 1973: 1426 (Quedius; Alberta; Northwest Territories).

  • — Smetana, 1976c: 172 (Quedius; subgenus Microsaurus; Ontario).

  • — Morgan and Morgan, 1979: 230 (Quedius; Postglacial fossils; Wisconsin).

  • — Smetana, 1981b: 637 (Quedius; collecting notes; Manitoba; Saskatchewan).

  • — Smetana, 1990d: 98 (Quedius; Newfoundland; Quebec).

  • — Downie and Arnett, 1996: 391 (Quedius; subgenus Microsaurus; characters; USA).

  • — Herman, 2001: 29 (Quedius; synonym of uteanus).

vadhu Smetana, 1975a: 334 (Quedius; subgenus Raphirus; Type locality: Central Nepal: Kali-Gandaki Valley between Lete and Tukche).

  • — Coiffait, 1982: 33 (Quedius; characters; Nepal).

  • — Smetana, 1988: 245 (Quedius; subgenus Raphirus; characters; Nepal).

  • Distribution: Nepal.

vafer Smetana, 1997b: 130 (Quedius; subgenus Raphirus; Type locality: China: Yuzhu Feng, 3350 m, Diancang Shan Mts., Dali Shi, Yunnan).

  • Distribution: China.

vagans Oke, 1933: 125 (Quedius; Type locality: Australia: Victoria: Belgrave; New South Wales: Dorrigo).

  • Distribution: Australia.

validus Smetana, 1971: 48 (Quedius; subgenus Megaquedius; Type locality: California: Stanford U. Cal.).

  • — Smetana, 1973: 1422 (Quedius; characters; California).

  • — Smetana, 1976c: 169 (Quedius; subgenus Megaquedius; California).

  • Distribution: USA.

vandykei Smetana, 1971: 105 (Quedius; subgenus Microsaurus; Type locality: California: Illouette Cñ, Yosemite).

  • — Smetana, 1978a: 825 (Quedius; subgenus Microsaurus; collecting notes; California).

  • Distribution: USA.

variabilis Gyllenhal, 1810, see: mesomelinus Marsham, 1802 and puncticollis Thomson, 1867.

variabilis Mulsant and Rey, 1876, see: nigrocoeruleus Fauvel, 1876.

variegatus Bernhauer, 1941b: 28 (Quedius; subgenus Sauridus; Type locality: Neu Seeland: Greymouth, Picton).

  • Distribution: New Zealand.

variicolor Nordmann, 1837, see: picipes Mannerheim, 1830.

vasconicus Jarrige, 1972: 157 (Quedius; subgenus Raphirus; Type locality: forêt d'Iraty, Pyr.-Atl.).

  • — Coiffait, 1978: 266 (Quedius; subgenus Raphirus; characters; Pyrenees).

  • Distribution: France.

velox Smetana, 1971: 96 (Quedius; subgenus Microsaurus; Type locality: Canada: British Columbia: 21 mi. W. Creston. Also cited from Alberta, Alaska, Idaho, Montana, Oregon, Washington).

  • — Legner and Moore, 1977: 176 (Quedius; found under bark).

  • Distribution: Canada, USA.

velutinus Motschulsky, 1860, see: semiobscurus Marsham, 1802.

ventralis Aragona, 1830, see: truncicola Fairmaire and Laboulbène, 1856.

verecundus Normand, 1947, see: pauliani C. Koch, 1941.

verecundus Sharp, 1884: 334 (Quedius; Type locality: Panama, Volcan de Chiriqui between 2500 and 4000 feet).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; catalog).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1933: 1465 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 174 (Quedius; subgenus Cyrtoquedius; characters in key; Panama).

  • Distribution: Panama.

vernix LeConte, 1878, see: Anaquedius.

verres Smetana, 1971: 174 (Quedius; subgenus Distichalius; Type locality: South Carolina: Florence, 7100 ft. Also cited from Arizona, Arkansas, Louisiana, New Jersey, Mississippi, Pennsylvania, Texas; Mexico).

  • — Smetana, 1971a: 1845 (Quedius; subgenus Distichalius; collecting notes).

  • — Smetana, 1973: 1428 (Quedius; notes; Arizona; North Carolina).

  • — Smetana, 1976c: 176 (Quedius; subgenus Distichalius; Alabama; Florida).

  • — Smetana, 1978a: 827 (Quedius; subgenus Distichalius; collecting notes; Arizona; Florida; Oklahoma; Texas).

  • — Smetana, 1981b: 640 (Quedius; collecting notes; Florida; South Carolina).

  • — Smetana, 1990d: 100 (Quedius; Arkansas; Florida).

  • — Downie and Arnett, 1996: 392 (Quedius; subgenus Distichalius; characters; USA).

  • Distribution: USA, Mexico.

versicolor Fauvel, 1877: 273 (Quedius; Type locality: Australie, Queensland, Gayndah).

  • — Fauvel, 1878g: 556 (Quedius; notes).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; catalog).

  • — Bernhauer, 1920: 18 (Quedius; Australia).

  • Distribution: Australia.

vesparum Rüschkamp, 1933, see: invreae Gridelli, 1924.

vexans Eppelsheim, 1881b: 297 (Quedius; Type locality: Ostpreussen; Königsberg, Danzig und Gilgenburg).

  • — Ganglbauer, 1895: 398 (Quedius; subgenus Microsaurus; characters; east Prussia; Germany).

  • — Reitter, 1909: 110 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Sharp and Muir, 1912: 498 (Quedius; characters of male genitalia).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; catalog).

  • — Gridelli, 1924d: 58 (Quedius; subgenus Microsaurus; characters; Prussia; Silesia; Bohemia; Germany).

  • — Gridelli, 1929: 35 (Quedius; characters).

  • — Scheerpeltz, 1933: 1465 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 11 (Quedius; aedeagal illustration).

  • — Roubal, 1941b: 124, 128 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Smetana, 1957a (Quedius; larval characters; collected from nest of Microtus arvalis).

  • — Smetana, 1958: 352 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 158 (Quedius; subgenus Microsaurus; characters).

  • — Lohse, 1964: 208 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Smetana, 1964c: 168 (Quedius; subgenus Microsaurus; Czechoslovakia).

  • — Horion, 1965: 263 (Quedius; subgenus Microsaurus; Poland; Czechoslovakia; Hungary; Austria; Germany; France).

  • — Pototskaia, 1967: 83 (Quedius; larval characters).

  • — Szujecki, 1968a: 734 (Quedius; Poland).

  • — Coiffait, 1978: 166 Quedius; subgenus Microsaurus; characters; Europe).

  • — Hoebeke, 1978: 508 (Quedius; parasitized by proctotrupid).

  • — Frank, 1982: 47 (Quedius; list of parasites).

  • — Tóth, 1984: 110 (Quedius; characters).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Sweden).

  • — Schülke and Uhlig, 1988: 12 (Quedius; notes; Germany).

  • — Lohse and Lucht, 1989: 172 (Quedius; note).

  • — Nowosad, 1990: 137 (Quedius; collected from mole nest; notes; Poland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; Italy).

  • Distribution: Europe.

langenhani Hubenthal, 1912: 72 (Quedius; variety of vexans; Type locality: Gotha).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; aberration of vexans).

  • — Gridelli, 1924d: 59 (Quedius; aberration of vexans).

  • — Roubal, 1941b: 124 (Quedius; subgenus Microsaurus; variety of vexans; characters).

  • — Smetana, 1958: 352 (Quedius; synonym of vexans).

  • — Smetana, 1960e: 158 (Quedius; aberration of vexans; characters).

  • — Coiffait, 1978: 166 (Quedius; subgenus Microsaurus; variety of vexans; characters).

  • — Tóth, 1984: 110 (Quedius; aberration of vexans).

vicinus Lacordaire, 1835, see: scintillans Gravenhorst, 1806.

vicinus Ménétriés, 1832: 144 (Staphylinus; Type locality: Lenkoran; [Note: See lectotype designation by Gusarov, 1993]. Originally cited from montagnes de Talyche).

  • — Faldermann, 1835: 129 (Staphylinus; characters; Persia-Armenia).

  • — Hochhuth, 1849: 147 (Quedius; characters; Lenkoran; Tiflis [Note: This locality probably refers to Q. suramensis]).

  • — Hochhuth, 1862: 44 (Quedius; notes; Russia).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; catalog).

  • — Gusarov, 1993: 73 (Quedius; lectotype designation: Lenkoran; synonyms).

  • — Assing and Wunderle, 2001: 37 (Quedius; Cyprus).

  • Distribution: Lebanon, Azerbaijan, Iran, Turkmenistan.

libanicus Coiffait, 1954a: 160 (Quedius; Type locality: Liban: grotte de Birket Aanjar).

  • — Coiffait, 1955e: 427 (Quedius; cave collection; Lebanon).

  • — Coiffait, 1961a: 55 (Quedius; characters; Lebanon).

  • — Jarrige, 1971: 497 (Quedius; Iran).

  • — Korge, 1971: 11 (Quedius; subgenus Quedius; characters; notes; Iran).

  • — Coiffait, 1978: 195 (Quedius; subgenus Quedius; characters; Lebanon; central Asia).

  • — Boháč, 1988: 554 [= 1989: 38] (Quedius; characters; Turkmenia; Iran).

  • — Gusarov, 1993: 73 (Quedius; synonym of vicinus).

maculatus Korge, 1971a: 44 (Quedius; aberration of vicinus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Kleinasien: Mahmudiye südlich Ordu, 1500 m and the Caucasus).

viduus Sawada, 1965: 17 (Quedius; subgenus Quedius; Type locality: Japan: Pref. Nagano: Mt. Jônen).

  • Distribution: Japan.

vilis Smetana, 1971: 191 (Quedius; subgenus Raphirus; Type locality: Oregon: 7.5 mi. SW. Fort Klamath. Also cited from California, Nevada, British Columbia).

  • — Smetana, 1971a: 1846 (Quedius; subgenus Raphirus; collecting notes).

  • — Smetana, 1976c: 180 (Quedius; subgenus Raphirus; California; Idaho; Oregon).

  • — Smetana, 1978a: 829 (Quedius; subgenus Raphirus; collecting notes; British Columbia; Oregon).

  • Distribution: Canada, USA.

virens Rottenberg, 1870, see: cruentus Olivier, 1795.

virginicus Casey, 1915: 406 (Distichalius; Type locality: Virginia; [Note: See lectotype designation by Smetana, 1971]. Originally cited from Virginia: Hampton Roads; New Jersey: Atlantic City).

  • — Scheerpeltz, 1933: 1465 (Quedius; catalog).

  • — Smetana, 1971: 188 (Quedius; subgenus Raphirus; lectotype designation: Va [Virginia]; characters; Georgia; Massachusetts; New Jersey; New York; North Carolina; Rhode Island).

  • — Smetana, 1978a: 829 (Quedius; subgenus Raphirus; collecting notes; Massachusetts; New Jersey).

  • — Downie and Arnett, 1996: 393 (Quedius; subgenus Raphirus; characters; USA).

  • Distribution: USA.

virgulatus Erichson, 1840: 547 (Quedius; Type locality: Sardinia).

  • — Kraatz, 1857d: 501 (Quedius; Sardinia).

  • — Fauvel, 1874: 304 [= 1874a: 530] (Quedius; characters; France; Corsica; Sardinia).

  • — Mulsant and Rey, 1876: 778 [= 1877: 634] (Quedius; subgenus Raphirus; characters; France).

  • — Ganglbauer, 1895: 410 (Quedius; subgenus Raphirus; characters; Maritime Alps; Corsica; Sardinia).

  • — Sainte-Claire Deville, 1906: 103 (Quedius; Corsica).

  • — Porta, 1907: 145 (Quedius; subgenus Raphirus; characters; Italy).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; catalog).

  • — Gridelli, 1925: 159 (Quedius; subgenus Raphirus; characters; Sardinia; Corsica).

  • — Portevin, 1929: 344 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1465 (Quedius; catalog).

  • — Coiffait, 1978: 271 (Quedius; subgenus Raphirus; characters; Corsica; Sardinia).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; Italy).

  • Distribution: France, Italy.

flavipennis Baudi, 1848: 132 (Quedius; Type locality: Sardinia).

  • — Baudi, 1857: 101 (Quedius; variety of virgulatus; characters; notes).

  • — Kraatz, 1857d: 501 (Quedius; characters; Sardinia).

  • — Fauvel, 1869: 487 (Quedius; synonym of virgulatus).

  • — Fauvel, 1869a: 151 (Quedius; synonym of virgulatus).

  • — Fauvel, 1874: 304 [= 1874a: 530] (Quedius; synonym of virgulatus).

  • — Ganglbauer, 1895: 410 (Quedius; aberration of virgulatus).

  • — Porta, 1907: 145 (Quedius; synonym of virgulatus).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; variety of virgulatus).

  • — Gridelli, 1925: 159 (Quedius; synonym of virgulatus).

  • — Scheerpeltz, 1933: 1465 (Quedius; synonym of virgulatus).

  • — Coiffait, 1978: 271 (Quedius; synonym of virgulatus).

viridescens Fauvel, 1878g: 555 (Quedius; Type locality: Australie orientale et occidentale: Wide Bay, King's George Sound).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; catalog).

  • Distribution: Australia.

viridipennis Fauvel, 1891: 119 (Quedius; Type locality: Colonie Tovar).

  • — Bernhauer and Schubert, 1916: 437 (Quedius; catalog).

  • Distribution: Venezuela.

viridulus Erichson, 1840: 532 (Quedius; Type locality: Bogatae in Columbia).

  • — Bernhauer and Schubert, 1916: 437 (Quedius; catalog).

  • Distribution: Colombia.

vividus Broun, 1880: 112 (Philonthus; Type locality: Whanagarei Harbour).

  • — Bernhauer and Schubert, 1916: 437 (Quedius; catalog).

  • Distribution: New Zealand.

vulneratus Gemminger and Harold, 1868: 572 (Quedius; replacement name for plagiatus Hochhuth).

  • — Bernhauer and Schubert, 1916: 437 (Quedius; catalog).

  • — Gridelli, 1924d: 107 (Quedius; subgenus Raphirus; characters; Caucasus).

  • — Scheerpeltz, 1933: 1465 (Quedius; catalog).

  • — Coiffait, 1963a: 378 (Quedius; subgenus Sauridus; characters).

  • — Korge, 1964: 119 (Quedius; subgenus Sauridus; Turkey).

  • — Coiffait, 1978: 219 (Quedius; subgenus Sauridus; characters; Caucasus).

  • Distribution: Caucasus, Turkey.

plagiatus Hochhuth, 1849: 149 (Quedius; [preoccupied]; Type locality: Länder Kaukasiens).

  • — Hochhuth, 1862: 45 (Quedius; Russia).

  • — Fauvel, 1875a: XXXIII [= 1875b: 235] (Quedius; synonym of plagiatus).

  • — Bernhauer and Schubert, 1916: 437 (Quedius; synonym of vulneratus).

  • — Gridelli, 1924d: 107 (Quedius; synonym of vulneratus).

  • — Coiffait, 1978: 219 (Quedius; synonym of vulneratus).

plagifer Eppelsheim, 1889d: 15 (Quedius; subgenus Sauridus; Type locality: Circassien).

  • — Bernhauer and Schubert, 1916: 437 (Quedius; synonym of vulneratus).

  • — Gridelli, 1924d: 107 (Quedius; synonym of vulneratus).

  • — Coiffait, 1978: 219 (Quedius; synonym of vulneratus).

vulpinus Smetana, 1971: 214 (Quedius; subgenus Raphirus; Type locality: Washington: Mt. Ranier, Sunbeam Falls, 4000 ft.).

  • — Smetana, 1976c: 182 (Quedius; subgenus Raphirus; Oregon; Washington).

  • Distribution: USA.

wakefieldi Bernhauer, 1941b: 30 (Quedius; subgenus Sauridus; Type locality: Neu Seeland: Pakarau).

  • Distribution: New Zealand.

walteri Korge, 1971a: 44 (Quedius; subgenus Microsaurus; Type locality: Ilica im Tal von Ardesen, Kackar-Daglari, 1600–2000 m).

  • — Smetana, 1995a: 83 (Quedius; subgenus Raphirus; characters; notes; Turkey).

  • — Coiffait, 1978: 182 (Quedius; subgenus Microsaurus; characters; Turkey).

  • Distribution: Turkey.

wanyan Smetana, 1996a: 53 (Quedius; subgenus Raphirus; Type locality: China: [Qinghai], Qinghai-Nanshan ca. 32 km NW Caka, 3700–3800 m, [Wan Yan Tong Bu]).

  • Distribution: China.

wassu Smetana, 1998b: 101 (Quedius; subgenus Raphirus; Type locality: China: Lungai, Wassuland, 2000 m, W. Szechuan).

  • Distribution: China.

weiratheri Gridelli, 1938: 17 (Quedius; subgenus Microsaurus; Type locality: Asia Minore meridionale, aprentesi nella catena del Tauro Pisidico, gruppo montagnoso Dipojras, a nord-ovest di Bey Schehir, sui monti Anamas; una caverna del Tauro Isaurico, gruppo montagnoso Kybelon, a sud-est de Bey Schehir, circa a 60 km in linea d'aria dalla caverna precedente).

  • — Coiffait, 1978: 171 (Quedius; subgenus Microsaurus; characters; Turkey).

  • — Bordoni, 1978a: 59 (Quedius; adult and larval characters; notes; cave collection; Turkey).

  • Distribution: Turkey.

wrasei Smetana, 1999b: 226 (Quedius; subgenus Microsaurus; Type locality: China: W. Sichuan, Daxue Shan, Hailuogou Glacier Park Camp 1, 2100 m, 29.36.00 N, 102.03.35 E).

  • Distribution: China.

wuh Smetana, 1995: 90 (Quedius; subgenus Raphirus; Type locality: Taiwan: Hualien Hsien: Taroko N. P., Nanhushi Hut, 2220 m).

  • Distribution: Taiwan.

wuorentausi Bernhauer, 1927, see: sublimbatus Mäklin, 1853.

xanthippae Lohse, 1963: 177 (Quedius; Type locality: Rosazza, 20 km nördlich von Biella).

  • — Coiffait, 1963a: 383 (Quedius; subgenus Sauridus; [Note: Misidentified as haberfelneri according to Lohse, 1987: 138, 149]; characters).

  • — Lohse, 1964: 214 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Coiffait, 1967e: 419 (Quedius; subgenus Microquedius).

  • — Bordoni, 1974a: 205, 208 (Quedius; subgenus Microquedius; characters; Italy).

  • — Coiffait, 1978: 201 (Quedius; subgenus Microquedius; characters; Alps).

  • — Zanetti, 1979a: 542, 543 (Quedius; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; Italy).

  • Distribution: Italy.

xanthopus Erichson, 1839a: 487 (Quedius; Type locality: Mark Brandenburg).

  • — Heer, 1839: 273 (Philonthus; characters; Switzerland).

  • — Erichson, 1840: 527 (Quedius; characters; Germany).

  • — Kiesenwetter, 1848: 56 (Quedius; characters; Germany; Switzerland).

  • — Redtenbacher, 1849: 709 (Philonthus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 542 (Quedius; characters; France).

  • — Kraatz, 1857d: 495 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 201 (Quedius; characters; Austria).

  • — Thomson, 1860: 175 (Microsaurus; characters; Scandinavia).

  • — Fauvel, 1865a: 292 [= 1865b: 42] (Quedius; habitat; France).

  • — Fauvel, 1874: 284 [= 1874a: 510] (Quedius; characters; France; Lapland; Scandinavia; Britain; Germany; Switzerland; Austria; Italy; Russia; Lake Baikal).

  • — Redtenbacher, 1874: 198 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 267 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 665 [= 1877: 521] (Quedius; characters; France).

  • — J. Sahlberg, 1876: 26 (Microsaurus; Finland).

  • — Fowler, 1888: 234 (Quedius; characters; Britain).

  • — Ganglbauer, 1895: 400 (Quedius; subgenus Microsaurus; characters; Palaearctic region).

  • — Porta, 1907: 127 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Reitter, 1909: 110 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Johansen, 1914: 359 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 437 (Quedius; catalog).

  • — Munster, 1923: 194 (Quedius; subgenus Microsaurus; Fennoscandia).

  • — Gridelli, 1924d: 65 (Quedius; subgenus Microsaurus; characters; Styria; Carniola; Italy; Dalmatia; Bosnia; Transylvania).

  • — Portevin, 1929: 340 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1465 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 15 (Quedius; aedeagal illustration).

  • — Roubal, 1941b: 125, 129 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Hinton, 1945: 71 (Quedius; characters).

  • — V. Hansen, 1952: 137 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Smetana, 1958: 355 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 158 (Quedius; subgenus Microsaurus; characters).

  • — Palm, 1963b: 136 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 209 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Horion, 1965: 267 (Quedius; subgenus Microsaurus; widespread in northern Europe; Russia; Poland; Czechoslovakia; Hungary; Romania; Switzerland; France; Holland; Spain; Italy; Slovenia; Istria; Dalmatia; Bosnia Hercegovina; Germany; Austria).

  • — Pototskaia, 1967: 83, 86 (Quedius; larval characters).

  • — Szujecki, 1968a: 735 (Quedius; Poland).

  • — Bordoni, 1974: 14 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Pototskaia, 1976: 173 (Quedius; larval characters).

  • — Legner and Moore, 1977: 176 (Quedius; found under bark).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 174 (Quedius; subgenus Microsaurus; characters; Palaearctic region).

  • — Frank, 1982: 48 (Quedius; list of parasites).

  • — Tóth, 1984: 112 (Quedius; characters; Hungary).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Smetana, 1993b: 50 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; Italy).

  • — Fowles, Alexander, and Key, 1999: 129 (Quedius; saproxylic species; status as endangered or threatened species; Britain).

  • — Lott, 1999: 72 (Quedius; saproxylic species; habitat; endangered status; Britain).

  • — Hågvar, 1999: 28 (Quedius; collected on Fomitopsis pinicola; Norway).

  • Distribution: Europe, Russia.

xanthurus Iablokov-Khnzorian, 1961: 147 (Quedius; Type locality: Armenie orientale: Nerkin Hand).

  • — Coiffait, 1978: 261 (Quedius; subgenus Sauridus; characters; Armenia).

  • Distribution: Armenia.

xeno Smetana, 1997a: 126 (Quedius; subgenus Raphirus; Type locality: N. Vietnam: Hoang Lien Son Mts., 1920 m, N of Phang Si Pang, Lai Chau Prov.).

  • — Smetana, 1998b: 109 (Quedius; subgenus Raphirus; characters).

  • Distribution: Vietnam.

xenophaenus Broun, 1912: 399 (Quedius; Type locality: Waimarino).

  • — Bernhauer and Schubert, 1916: 437 (Quedius; catalog).

  • Distribution: New Zealand.

xochitl Smetana, 1976a: 239 (Quedius; subgenus Quedionuchus; Type locality: Mexico: Chiapas: 5 mi. W. San Cristobal).

  • Distribution: Mexico.

xylophilus Lea, 1908: 152 (Quedius; Type locality: King Island; Tasmania).

  • — Scheerpeltz, 1933: 1465 (Quedius; catalog).

  • Distribution: Australia.

tylophilus Scheerpeltz, 1933: 1464 ([Note: Nomen nudum (Article 13.1.1), cited without description. Scheerpeltz attributed the species to Lea (1913: 152 [= 1908]) but Lea described no such species, however, he did describe xylophilus in the same reference and on the same page that Scheerpeltz cited for tylophilus so the two are probably the same. Scheerpeltz's tylophilus may be a misspelling of that name but Scheerpeltz cited both spellings as valid species]).

yann Smetana, 1995: 102 (Quedius; subgenus Raphirus; Type locality: Taiwan: Chiai Hsien: Yushan N. P., Ta-Ta Ghia, 2750 m).

  • — Smetana, 1998b: 110 (Quedius; subgenus Raphirus; characters).

  • Distribution: Taiwan.

yaoqi Smetana, 1999e: 542 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan Pv., Baoxing Xian, Yaoqi/Mahuanggou, 2650 m).

  • Distribution: China.

yasuhikoi Sawada, 1964: 37 (Quedius; Type locality: Japan: Honshu: Pref. Nagano: Mt. Kiso-Komagadake, 2100 m).

  • — Watanabe, 1987: 324 (Quedius; characters; Japan).

  • Distribution: Japan.

yean Smetana, 1995: 53 (Quedius; subgenus Microsaurus; Type locality: Taiwan: Pingtung Hsien, Peitawushan Trail at 2000 m).

  • Distribution: Taiwan.

yun Smetana, 1995: 43 (Quedius; subgenus Microsaurus; Type locality: Taiwan: Nantou Hsien Yushan N. P. SW side blw. Yushan Mn. Pk., 3650 m).

  • — Smetana, 1996g: 23 (Quedius; notes; Taiwan).

  • Distribution: Taiwan.

zeuxis Smetana, 1997a: 123 (Quedius; subgenus Microsaurus; Type locality: N. Vietnam: Hoang Lien Son Mts., 1840 m, N of Phang Si Pang, Lao Cai Prov,).

  • Distribution: Vietnam.

zheduo Smetana, 1999b: 235 (Quedius; subgenus Microsaurus; Type locality: China: W. Sichuan, Pass Zheduo Shankou, W. Kangding, W. slope, 4000 m, 29°58 N, 101°47 E).

  • Distribution: China.

Quelaestrygon [1 species; Palaearctic Region]

Quelaestrygon Smetana, 1999c: 241 (species included: puetzi). Type species: Quelaestrygon puetzi Smetana, fixed by original designation and monotypy.

puetzi Smetana, 1999c: 246 (Quelaestrygon; Type locality: China: Sichuan, Daxue Shan Gongga Shan, Mt. Hailuogou Glacier Park, 2620–1940 m).

  • Distribution: China.

Quetarsius [2 species; Palaearctic Region]

Quetarsius Smetana, 1996g: 25 (species included: neu, jeau). Type species: Quedius neu Smetana, fixed by original designation.

jeau Smetana, 1995: 95 (Quedius; subgenus Raphirus; Type locality: Taiwan: Kaohsiung Hsien: Kuanshan trail, above Kaunshanchi Riv., 2550 m).

  • — Smetana, 1996g: 26 (Quetarsius; Taiwan).

  • Distribution: Taiwan.

neu Smetana, 1995: 93 (Quedius; subgenus Raphirus; Type locality: Taiwan: Hualien Hsien, Taroko N.P., Nanhushi Hut, 2220 m).

  • — Smetana, 1996g: 26 (Quetarsius; Taiwan).

  • Distribution: Taiwan.

Rolla [3 species; Neotropical Region]

Rolla Blackwelder, 1952: 342 (replacement name for Leptoparius Bernhauer). Type species: Leptoparius pardoxus Bernhauer, fixed by objective synonymy with Leptoparius.

Leptoparius Bernhauer, 1917h: 87 (species included: paradoxus; preoccupied by Peters, 1864). Type species: Leptoparius paradoxus Bernhauer, fixed by monotypy.

  • — Scheerpeltz, 1933: 1421 (world catalog supplement).

  • — Blackwelder, 1944: 144 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 217 (synonym of Rolla; type species).

paradoxa Bernhauer, 1917h: 88 (Leptoparius; Type locality: West-Kolumbien: Linia, Quindici-Pass, 3600 m).

  • — Scheerpeltz, 1933: 1421 (Leptoparius; catalog).

  • Distribution: Colombia.

prominens Bernhauer, 1917h: 89 (Quedius; subgenus Ediquus; Type locality: Hoch-Kordilleren Kolumbiens: Paramo del Monte Tolima, 4500 m).

  • — Scheerpeltz, 1933: 1421 (Leptoparius; catalog).

  • Distribution: Colombia.

wittei Bernhauer, 1927e: 248 (Heterothops; subgenus Leptoparius; Type locality: Argentinien: Gob. Santa Cruz, Valle Tunel).

  • — Scheerpeltz, 1933: 1421 (Leptoparius; catalog).

  • Distribution: Argentina.

Scariphaeus [3 species; Neotropical Region]

Scariphaeus Erichson, 1839b: 342 (species included: luridipennis). Type species: Velleius luridipennis Runde, fixed by monotypy.

  • — Lacordaire, 1854: 73 (characters; notes; list of species).

  • — Gemminger and Harold, 1868: 574 (catalog).

  • — Lynch, 1884: 117 (characters).

  • — Bernhauer and Schubert, 1916: 410 (world catalog; 1 species).

  • — Bruch, 1928a: 429 (catalog; Argentina).

  • — Scheerpeltz, 1933: 1420 (world catalog supplement).

  • — Scheerpeltz, 1936a: 539 (key to species).

  • — Blackwelder, 1944: 143 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 345 (type species: luridipennis).

bruchi Bernhauer, 1927e: 246 (Scariphaeus; Type locality: Argentinien: Prov. Buenos Aires).

  • — Lynch, 1884: 117 (Scariphaeus; [Note: Misidentified as luridipennis according to Scheerpeltz, 1936a: 539]; characters; Argentina).

  • — Scheerpeltz, 1933: 1420 (Scariphaeus; catalog).

  • — Scheerpeltz, 1936a: 539 (Scariphaeus; characters; Argentina).

  • Distribution: Argentina.

erichsoni Scheerpeltz, 1936a: 532, 540 (Scariphaeus; Type locality: Brasilien).

  • Distribution: Brazil.

luridipennis Runde, 1835: 1, note (Velleius; Type locality: Brasilia).

  • — Erichson, 1839b: 343 (Scariphaeus; characters; Brazil).

  • — Bernhauer and Schubert, 1916: 410 (Scariphaeus; catalog).

  • — Scheerpeltz, 1933: 1420 (Scariphaeus; catalog).

  • — Scheerpeltz, 1936a: 531, 539 (Scariphaeus; characters; natural history; Brazil; Paraguay).

  • Distribution: Brazil, Paraguay.

Smilax [4 species; Neotropical Region]

Smilax Laporte, 1835: 116 (species included: americana). Type species: Smilax americana Laporte, fixed by monotypy.

  • — Laporte, 1840: 175 (characters).

  • — Lacordaire, 1854: 73 (synonym of Cordylaspis and homonym of genus of plants).

  • — Bernhauer and Schubert, 1916: 409 (synonym of Cordylaspis).

  • — Blackwelder, 1944: 143 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 353 (type species; note).

Cordylaspis Nordmann, 1837: 17 (species included: tuberculatus). Type species: Cordylaspis tuberculata Nordmann, fixed by monotypy.

  • — Erichson, 1839b: 341 (characters).

  • — Lacordaire, 1854: 73 (characters; notes; list of species).

  • — Gemminger and Harold, 1868: 573 (catalog).

  • — Sharp, 1876: 101 (notes).

  • — Sharp, 1884: 342 (notes; Mexican and Central American species).

  • — Bernhauer and Schubert, 1916: 409 (world catalog; 1 species).

  • — Bruch, 1928a: 429 (catalog; Argentina).

  • — Scheerpeltz, 1933: 1420 (cited as Cordilaspis; world catalog supplement).

  • — Scheerpeltz, 1936a: 502 (key to species).

  • — Scheerpeltz, 1937c (key to species).

  • — Blackwelder, 1952: 107 (synonym of Smilax; type species).

americana Laporte, 1835, see: pilosa Fabricius, 1787.

cyanea Wasmann, 1902a: 267 (Smilax; variety of pilosa; Type locality: Staate Rio de Janeiro, zu Colonia alpina).

  • — Scheerpeltz, 1936a: 492, 495, 502 (Cordylaspis; valid species; characters; Argentina, Brazil, Bolivia, Paraguay, and Panama).

  • — Scheerpeltz, 1937c: 337 (Cordylaspis; characters in key; Colombia; Venezuela; Bolivia; Guyana; Brazil; Paraguay; Uruguay; Argentina; Central America).

  • — Paulian, 1941: 253 (Cordylaspis; larval characters).

  • Distribution: Uruguay, Argentina, Paraguay, Bolivia, Brazil, Guyana, Venezuela, Colombia, Panama.

kuntzeni Scheerpeltz, 1937c: 330, 336 (Cordylaspis; Type locality: Durango, Mexico).

  • Distribution: Mexico.

lynchi Bruch, 1928: 416 (Cordylaspis; Type locality: Tafí Viejo, provincia de Tucumán).

  • — Scheerpeltz, 1933: 1420 (Cordylaspis, cited as Cordilaspis; catalog).

  • — Scheerpeltz, 1936a: 502 (Cordylaspis; characters in key).

  • — Scheerpeltz, 1937c: 336 (Cordylaspis; characters in key).

  • Distribution: Argentina.

pilosa Fabricius, 1787: 220 (Staphylinus; Type locality: Cayennae).

  • — Gmelin, 1790: 2028 (Staphylinus; characters).

  • — Fabricius, 1793: 523 (Staphylinus; characters).

  • — Olivier, 1795:(42): 24 (Staphylinus; characters; French Guiana).

  • — Fabricius, 1801: 594 (Staphylinus; characters; French Guiana).

  • — Turton, 1802: 511 (Staphylinus; characters; Cayenne).

  • — Erichson, 1839b: 342 (Cordylaspis; characters; Columbia; Cayenne; Surinam; Brazil).

  • — Sharp, 1876: 101 (Cordylaspis; Brazil).

  • — Sharp, 1884: 342 (Cordylaspis; notes; Mexico).

  • — Bernhauer and Schubert, 1916: 409 (Cordylaspis; catalog).

  • — Scheerpeltz, 1936a: 495, 502 (Cordylaspis; characters).

  • — Scheerpeltz, 1937c: 337 (Cordylaspis; characters in key).

  • Distribution: Nicaragua, Costa Rica, Panama, Colombia, Surinam, French Guiana, Brazil.

americana Laporte, 1835: 117 (Smilax; Type locality: Cayenne; Brésil).

  • — Erichson, 1839b: 342 (Cordylaspis; synonym of pilosa).

  • — Laporte, 1840: 175 (Smilax; characters; Cayenne; Brazil).

  • — Gemminger and Harold, 1868: 573 (Cordylaspis; synonym of pilosa).

  • — Sharp, 1884: 342 (Cordylaspis; synonym of pilosa).

  • — Bernhauer and Schubert, 1916: 409 (Cordylaspis; synonym of pilosa).

tuberculata Nordmann, 1837: 17 (Cordylaspis; Type locality: Sumatra; [Note: Type locality erroneous according to Erichson, 1839b: 342]).

  • — Erichson, 1839b: 342 (Cordylaspis; type locality erroneous; synonym of pilosa).

  • — Sharp, 1884: 342 (Cordylaspis; synonym of pilosa).

  • — Bernhauer and Schubert, 1916: 409 (Cordylaspis; synonym of pilosa).

tuberculata Nordmann, 1837, see: pilosa Fabricius, 1787.

Sphingoquedius [1 species; Australian Region]

Sphingoquedius Bernhauer, 1941b: 27 (species included: strandi). Type species: Sphingoquedius strandi Bernhauer, fixed by monotypy.

  • — Blackwelder, 1952: 355 (type species: strandi).

strandi Bernhauer, 1941b: 27 (Sphingoquedius; Type locality: Neu Seeland: Greymouth).

  • — Klimaszewski, Newton, and Thayer, 1996: 151 (Sphingoquedius; endemic to New Zealand).

  • Distribution: New Zealand.

Stevensia [1 species; Palaearctic Region]

Stevensia Cameron, 1932: 162 (species included: longipennis). Type species: Stevensia longipennis Cameron, fixed by monotypy.

  • — Scheerpeltz, 1933: 1374 (world catalog supplement).

  • — Blackwelder, 1952: 360 (type species: longipennis).

longipennis Cameron, 1932: 162 (Stevensia; Type locality: Sikkim: Gopaldhara, Rungbong Valley).

  • — Scheerpeltz, 1933: 1374 (Stevensia; catalog).

  • Distribution: India.

Strouhalium [5 species; Palaearctic Region]

Strouhalium Scheerpeltz, 1962b: 259, 267 (species included: gracilicorne). Type species: Strouhalium gracilicorne Scheerpeltz, fixed by original designation.

  • — Korge, 1971a: 35 (characters).

  • — Smetana, 1977b: 181 (characters).

  • — Smetana, 1993: 85 (characters).

  • — Smetana, 1995j: 139 (key to species).

brezinai Smetana, 1995j: 132 (Strouhalium; Type locality: China: C-Sichuan [Kangding], Gongga Shan massive, + 2900 m, ca. 25 km NNW of Moxi village, 29°49′N, 102°03′E).

  • — Smetana, 1998d: 95 (Strouhalium; characters of female; China).

  • Distribution: China.

calculosum Smetana, 1995j: 138 (Strouhalium; Type locality: India: Burphu, Gori V., 11500 ft.).

  • Distribution: India.

farkaci Smetana, 1995j: 135 (Strouhalium; Type locality: China: W. Sichuan, 29.26N, 102.06E, 1900–2900 m, Gonggashan-Hailuogou).

  • Distribution: China.

gracilicorne Scheerpeltz, 1962b: 268 (Strouhalium; Type locality: Tibet).

  • — Smetana, 1993: 88 (Strouhalium; characters; illustrations; Pakistan; Nepal; Tibet is questioned).

  • — Smetana, 1998d: 97 (Strouhalium; notes; China).

  • — Smetana, 1999d: 519 (Strouhalium; distributional notes; collecting notes; have adaptations to underground life; China).

  • Distribution: Pakistan, Nepal, China, Tibet ?.

sichuanense Smetana, 1995j: 135 (Strouhalium; Type locality: China: Sichuan, Kanding).

  • — Smetana, 1998d: 97 (Strouhalium; characters of female; China).

  • Distribution: China.

Termitoquedius [2 species; Neotropical Region]

Termitoquedius Bernhauer, 1912d: 45 (species included: iheringi). Type species: Termitoquedius iheringi Bernhauer, fixed by monotypy.

  • — Bernhauer and Schubert, 1916: 437 (world catalog; 1 species).

  • — Blackwelder, 1944: 145 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 379 (type species: iheringi).

  • — Seevers, 1965: 329 (characters; myrmecophilus species; list of species).

Ecitolycus Wasmann, 1925: 76, 161 (without characters for it or the one included species, laticollis; [Note: Nomen nudum (Article 12.1)]).

  • — Scheerpeltz, 1933: 1420 (world catalog supplement).

  • — Blackwelder, 1944: 143 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 140 (unavailable name).

  • — Seevers, 1965: 329 (synonym of Termitoquedius; [Note: Seevers contended that Ecitolycus was made available by its synonymy with Termitoquedius; however, the species, laticollis, still lacked a description and I can find no provision in the Code to support making an unavailable name available simply by citing it as a junior synonym]).

gigas Borgmeier, 1959: 4 (Ecitolycus; Type locality: Itatiaia, Estado do Rio de Janeiro).

  • — Seevers, 1965: 329 (Termitoquedius).

  • Distribution: Brazil.

iheringi Bernhauer, 1912d: 46 (Termitoquedius; Type locality: Brasilien: Rio de Janeiro: Serra Macahé).

  • — Bernhauer and Schubert, 1916: 437 (Termitoquedius; catalog).

  • — Seevers, 1957: 318 (Termitoquedius; doubtfully termitophilous).

  • — Seevers, 1965: 329 (Termitoquedius; cited as heringi).

  • Distribution: Brazil.

laticollis Wasmann, 1925: 76, 161 (Ecitolycus; without characters; [Note: Nomen nudum (Article 12.1)]; cited from Minas Geraës und Parana).

  • — Scheerpeltz, 1933: 1420 (Ecitolycus; catalog).

  • — Seevers, 1965: 329 (Termitoquedius; synonym of iheringi).

laticollis Wasmann, 1925 [Note: Nomen nudum], see: iheringi Bernhauer, 1912.

Valdiviodes [1 species; Neotropical Region]

Valdiviodes Smetana, 1981a: 349 (species included: ashworthi). Type species: Valdiviodes ashworthi Smetana, fixed by original designation.

ashworthi Smetana, 1981a: 350 (Valdiviodes; Type locality: Chile: nr. Laguna Espejo P. N. de Puyehue, Osorno Prov., 500 m).

  • Distribution: Chile.

Velleiopsis [2 species; Palaearctic Region]

Velleiopsis Fairmaire, 1882: clxiv (species included: marginiventris). Type species: Velleiopsis marginiventris Fairmaire, fixed by monotypy.

  • — Duvivier, 1883: 137 (catalog).

  • — Heyden, Reitter, and Weise, 1891: 99 (list of species of Europe and Caucasus).

  • — Heyden, Reitter, and Weise, 1906: 176 (list of species of Europe and Caucasus).

  • — Bernhauer and Schubert, 1916: 416 (world catalog; 2 species).

  • — Winkler, 1925: 388 (catalog for Palaearctic region).

  • — Scheerpeltz, 1933: 1427 (world catalog supplement).

  • — Blackwelder, 1952: 402 (type species: marginiventris).

  • — Scheerpeltz, 1962b: 266 (characters).

  • — Tikhomirova, 1973a: 188 (in checklist of species of Russia).

  • — Smetana, 1977b: 181 (characters).

  • — Coiffait, 1978: 288 (characters; key to species of western Palaearctic region).

marginiventris Fairmaire, 1882: clxiv (Velleiopsis; Type locality: Bulgarie, aux environs de Varna).

  • — Bernhauer and Schubert, 1916: 416 (Velleiopsis; catalog).

  • — Scheerpeltz, 1933: 1427 (Velleiopsis; catalog; Asia Minor).

  • — Smetana, 1995a: 86 (Velleiopsis; notes; Turkey).

  • — Coiffait, 1978: 288 (Velleiopsis; characters; Bulgaria; Asia Minor).

  • Distribution: Bulgaria, Turkey.

varendorffi Reitter, 1897: 247 (Velleiopsis; Type locality: Araxesthal bei Ordubad).

  • — Bernhauer and Schubert, 1916: 416 (Velleiopsis; catalog).

  • — Scheerpeltz, 1933: 1427 (Velleiopsis; catalog; Transcaucasus).

  • — Coiffait, 1978: 290 (Velleiopsis; characters; Caucasus).

  • Distribution: Azerbaijan.

Velleius [8 species; Palaearctic Region]

Velleius Leach, 1819: 172 (species included: dilatatus, concolor). Type species: Staphylinus dilatatus Fabricius, fixed by subsequent designation by Westwood, 1838a: 15.

  • — Curtis, 1829: 24 (catalog; Britain).

  • — Stephens, 1829a: 22 (catalog; Britain).

  • — Stephens, 1829: 274 (catalog; Britain).

  • — Mannerheim, 1830: 7, 19 (characters).

  • — Mannerheim, 1831: 421, 433 (characters).

  • — Stephens, 1832: 200, 201 (characters).

  • — Dejean, 1833: 59 (list of species).

  • — Lacordaire, 1835: 356 (characters).

  • — Nordmann, 1837: 16 (characters).

  • — Westwood, 1838a: 15 (characters; type species).

  • — Erichson, 1839a: 483 (synonym of Quedius).

  • — Heer, 1839: 282 (characters).

  • — Stephens, 1839: 384 (characters).

  • — Laporte, 1840: 173 (characters).

  • — Lacordaire, 1854: 84 (synonym of Quedius).

  • — Fairmaire and Laboulbène, 1856: 534 (characters).

  • — Jacquelin du Val, 1856: 36 (characters).

  • — Kraatz, 1857d: 490 (synonym of Quedius).

  • — Thomson, 1858: 30 (characters).

  • — G. Waterhouse, 1858: 22 (subgroup of Quedius; catalog of British species).

  • — Thomson, 1859: 25 (characters; type species: dilatatus).

  • — Schaum, 1859: 25 (synonym of Quedius).

  • — Thomson, 1860: 172 (characters).

  • — Jacquelin du Val, 1868: 69 (catalog; European species).

  • — Fauvel, 1874: 264 [= 1874a: 490] (characters; larval characters).

  • — Fauvel, 1875a: XXXII [= 1875b: 234] (catalog).

  • — Mulsant and Rey, 1876: 608 [= 1877: 464] (characters).

  • — Duvivier, 1883: 137 (catalog).

  • — Sharp, 1888: 226 (characters; key to British species).

  • — Heyden, Reitter, and Weise, 1891: 99 (list of species of Europe and Caucasus).

  • — Ganglbauer, 1895: 388 (adult and larval characters).

  • — Heyden, Reitter, and Weise, 1906: 176 (list of species of Europe and Caucasus).

  • — Porta, 1907: 90 (in key to Italian species).

  • — Reitter, 1908: 106, 115 (characters).

  • — Petri, 1912: 80 (list of species of Siebenbürgen).

  • — Johansen, 1914: 348 (characters).

  • — Bernhauer and Schubert, 1916: 416 (world catalog; 4 species).

  • — Winkler, 1925: 388 (catalog for Palaearctic region).

  • — Porta, 1926: 118 (characters; in list of species of Italy).

  • — Portevin, 1929: 345 (characters).

  • — Roubal, 1930: 386 (catalog; Slovakia).

  • — Cameron, 1932: 279 (characters).

  • — Joy, 1932: 107 (in key to British species).

  • — Scheerpeltz, 1933: 1426 (world catalog supplement).

  • — Kloet and Hincks, 1945: 167 (in list of British species).

  • — Tottenham, 1949: 376, 421 (type species: dilatatus Fabricius; notes; checklist of British species).

  • — V. Hansen, 1952: 113 (characters; in list of Danish species).

  • — Blackwelder, 1952: 402 (type species).

  • — Smetana, 1958: 324 (characters; habitus; type species: dilatatus).

  • — Scheerpeltz, 1962b: 266 (characters).

  • — Székessy, 1963: 61 (characters).

  • — Palm, 1963b: 123 (characters; in list of species of Sweden).

  • — Lohse, 1964: 205 (characters; in key to central European species).

  • — Pototskaia, 1967: 79 (larval characters).

  • — Scheerpeltz, 1968a: 87 (in catalog of Austrian species).

  • — Kasule, 1970: 57 (larval characters).

  • — Tikhomirova, 1973a: 188 (in checklist of species of USSR).

  • — Smetana, 1977b: 181 (characters).

  • — Pope, 1977: 31 (in list of British species).

  • — Coiffait, 1978: 285 (characters).

  • — Topp, 1978: 312 (larval characters).

  • — Muona, 1979: 15 (in list of Scandinavian species).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 118 (catalog; Poland).

  • — Tóth, 1984: 95 (characters; habitus).

  • — Shibata, 1984: 117 (in checklist of species of Japan).

  • — Naomi, 1986b: 239 (characters; natural history notes; habitus; in key to species of Japan).

  • — Shibata, 1986a: 125 (in supplemental list of species from Taiwan).

  • — Drugmand 1987b (in comments on distribution of Belgian species).

  • — Watanabe, 1990b: 59 (characters; in key to species of Japan).

  • — Smetana, 1995: 109 (characters; in list of species of Taiwan).

  • — M. Hansen, Kristensen, Mahler, Pedersen, 1991: 107 (in list of Danish species).

  • — Silfverberg, 1992: 17 (in list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Smetana, 1993b: 49 (in list of species of Czech Republic and Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (in list of species of Italy).

  • — M. Hansen, 1996: 107 (in list of Danish species).

  • — Cho, 1996b: 114 (in key to species of Korea).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (in list of species collected in 1997; Denmark).

  • — Assing, Frisch, Kahlen, et al., 1998: 139 (note).

Laverna Gistel, 1829: 1129 (species included: dilatata). Type species: Staphylinus dilatatus Fabricius, fixed by monotypy.

  • — Blackwelder, 1952: 212 (synonym of Velleius; type species; notes).

amamiensis Watanabe, 1990b: 69 (Velleius; Type locality: Japan: Kagoshima Pref.: Amami-Ohshima Is., Hatsuno).

  • Distribution: Japan.

circumipectus Cho, 1996b: 115 (Velleius; Type locality: Korea: Chugchungnam-do, Kumsan-gun, Poseoksa Temple).

  • Distribution: Korea.

concolor Marsham, 1802, see: dilatatus Fabricius, 1787.

dilatatus Fabricius, 1787: 220 (Staphylinus; Type locality: Halae Saxonum).

  • — Gmelin, 1790: 2027 (Staphylinus; characters).

  • — Fabricius, 1793: 522 (Staphylinus; characters).

  • — Panzer, 1795: 353 (Staphylinus; characters; Germany).

  • — Paykull, 1800: 389 (Staphylinus; characters; Sweden).

  • — Fabricius, 1801: 592 (Staphylinus; characters).

  • — Turton, 1802: 510 (Staphylinus; characters; Saxony).

  • — Latreille, 1804: 326 (Staphylinus; characters; Germany; Sweden; France).

  • — Gravenhorst, 1806: 89 (Staphylinus; characters; France; Germany).

  • — Gyllenhal, 1810: 299 (Staphylinus; characters; Sweden).

  • — Leach, 1819: 172 (Velleius).

  • — Germar, 1823: 13 (Staphylinus; characters; northern Europe).

  • — Curtis, 1829: 24 (Velleius; catalog; Britain).

  • — Gistel, 1829: 1129 (Laverna).

  • — Mannerheim, 1830: 19 (Velleius).

  • — Mannerheim, 1831: 433 (Velleius).

  • — Stephens, 1832: 201 (Velleius; characters; England).

  • — Runde, 1835: 1 (Velleius; characters; Germany).

  • — Lacordaire, 1835: 357 (Velleius; characters; France).

  • — Nordmann, 1837: 16 (Velleius; England; France; Germany; Sweden).

  • — Westwood, 1838: 164 (Velleius; habitat).

  • — Westwood, 1838a: 15 (Velleius; type species).

  • — Erichson, 1839a: 484 (Quedius; characters; Germany).

  • — Heer, 1839: 282 (Velleius; characters; Switzerland).

  • — Stephens, 1839: 386 (Velleius; characters).

  • — Erichson, 1840: 534 (Quedius; characters; Europe).

  • — Laporte, 1840: 173 (Velleius; characters; France).

  • — Holme, 1842: 114 (Velleius; notes).

  • — Redtenbacher, 1849: 707 (Philonthus; characters; Austria).

  • — Küster, 1853a: 59 (Quedius; characters; Europe).

  • — Fairmaire and Laboulbène, 1856: 534 (Velleius; characters; France).

  • — Kraatz, 1857d: 490 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 200 (Quedius; characters; Austria).

  • — Thomson, 1859: 25 (type species of Velleius).

  • — Thomson, 1860: 172 (Velleius; characters; Scandinavia).

  • — Fauvel, 1865a: 292 [= 1865b: 42] (Velleius; habitat; France).

  • — Fauvel, 1874: 265 [= 1874a: 491] (Velleius; characters; France).

  • — Sharp, 1874: 23 (Velleius; doubt concerning presence in Japan).

  • — Redtenbacher, 1874: 197 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 267 (Velleius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 610 [= 1877: 466] (Velleius; characters; France).

  • — Sharp, 1889: 29 (Velleius; Japan).

  • — Fowler, 1888: 226 (Velleius; characters; Britain).

  • — Seidlitz, 1891: 395 (Velleius; characters; Europe).

  • — Seidlitz, 1891a: 421 (Velleius; characters; Europe).

  • — Ganglbauer, 1895: 390 (Velleius; characters; north and middle Europe).

  • — Porta, 1907: 104 (Velleius; characters; Italy).

  • — Reitter, 1909: 115 (Velleius; characters; Germany).

  • — Johansen, 1914: 348 (Velleius; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 416 (Velleius; catalog).

  • — Munster, 1923: 185 (Velleius; Fennoscandia).

  • — Portevin, 1929: 345 (Velleius; characters; France).

  • — Scheerpeltz, 1933: 1426 (Velleius; catalog).

  • — Gruardet, 1937 (Velleius; notes).

  • — Paulian, 1941: 257 (Velleius; larval characters).

  • — V. Hansen, 1952: 114 (Velleius; characters; Denmark).

  • — Steel, 1952a (Velleius; note on habits).

  • — Strassen, 1957 (Velleius; morphology; ecology; behavior; egg; larva; pupa).

  • — Smetana, 1958: 325 (Velleius; characters; ecology; Czechoslovakia).

  • — Nakane, 1963a: 94 (Velleius; characters; habitus photograph; Japan).

  • — Palm, 1963b: 123 (Velleius; characters; Sweden; Denmark; Finland).

  • — Lohse, 1964: 205 (Velleius; characters; central Europe).

  • — Horion, 1965: 252 (Velleius; Denmark; Sweden; Finland; Britain; Holland; Belgium; France; Switzerland; Italy; Slovenia; Croatia; Romania; Hungary; Czechoslovakia; Poland; Russia; Baltics; Ukraine; Germany; Austria).

  • — Pototskaia, 1967: 79 (Velleius; larval characters).

  • — Szujecki, 1968a: 734 (Velleius; Poland).

  • — Korge, 1971a: 7 (Velleius; Turkey).

  • — Pope, 1977: 31 (Velleius; Britain).

  • — Coiffait, 1978: 285 (Velleius; characters; Europe; Britain; Scandinavia).

  • — Hinton, 1981: 663 (Velleius; characters of egg).

  • — Tóth, 1984: 96 (Velleius; characters; Hungary).

  • — Yuh, Paik, Kwon, and Lee, 1985: 249 (Velleius; Korea).

  • — Naomi, 1986b: 242 (Velleius; characters; Japan).

  • — Segers, 1986: 51 (Velleius; checklist of species; Belgium).

  • — Lucht, 1987: 109 (Velleius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Watanabe, 1990b: 61 (Velleius; characters; Japan).

  • — J. Li, 1993: 41 (Velleius; China).

  • — Smetana, 1993b: 49 (Velleius; Czech Republic; Slovakia).

  • — Proess, 1994 (Velleius; Luxembourg).

  • — Sparacio, 1995: 164 (Velleius; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 31 (Velleius; Italy).

  • — Cho, 1996b: 114 (Velleius; characters; Korea).

  • — Fowles, Alexander, and Key, 1999: 129 (Veleius; saproxylic species; status as endangered or threatened species; Britain).

  • Distribution: Europe, Turkey, Korea, Japan, China.

serraticornis Schrank, 1798: 641 (Staphylinus; Type locality: Boia).

  • — Erichson, 1839a: 485 (Quedius; synonym of dilatatus).

  • — Erichson, 1840: 524 (Quedius; synonym of dilatatus).

  • — Kraatz, 1857d: 490 (Quedius; synonym of dilatatus).

  • — Ganglbauer, 1895: 390 (Velleius; synonym of dilatatus).

  • — Porta, 1907: 104 (Velleius; synonym of dilatatus).

  • — Bernhauer and Schubert, 1916: 416 (Velleius; synonym of dilatatus).

  • — Smetana, 1958: 325 (Velleius; synonym of dilatatus).

  • — Coiffait, 1978: 285 (Velleius; synonym of dilatatus).

concolor Marsham, 1802: 498 (Staphylinus; Type locality: Britain).

  • — Leach, 1819: 172 (Velleius).

  • — Germar, 1823: 13 (Staphylinus; synonym of dilatatus).

  • — Erichson, 1839a: 485 (Quedius; synonym of dilatatus).

  • — Erichson, 1840: 524 (Quedius; synonym of dilatatus).

  • — Kraatz, 1857d: 490 (Quedius; synonym of dilatatus).

  • — Ganglbauer, 1895: 390 (Velleius; synonym of dilatatus).

  • — Porta, 1907: 104 (Velleius; synonym of dilatatus).

  • — Bernhauer and Schubert, 1916: 416 (Velleius; synonym of dilatatus).

  • — Smetana, 1958: 325 (Velleius; synonym of dilatatus).

  • — Coiffait, 1978: 285 (Velleius; synonym of dilatatus).

elongatus Naomi, 1986b: 244 (Velleius; Type locality: Mt. Gori, Hiroshima Pref.).

  • — Watanabe, 1990b: 67 (Velleius; characters; Japan).

  • Distribution: Japan.

inquietus Champion, 1925, see: Quedius.

japonicus Watanabe, 1990b: 67 (Velleius; Type locality: Japan: Honshu: Tokyo Pref.: Higashimurayama).

  • Distribution: Japan.

luridipennis Runde, 1835, see: Scariphaeus.

pectinatus Sharp, 1874: 24 (Velleius; Type locality: Hiogo; Nagasaki).

  • — Bernhauer and Schubert, 1916: 416 (Velleius; catalog).

  • — Gridelli, 1932: 18 (Velleius; characters).

  • — Nakane, 1963a: 94 (Velleius; characters; habitus photograph; Japan).

  • — Yuh, Paik, Kwon, and Lee, 1985: 249 (Velleius, cited as Veleius; Korea).

  • — Naomi, 1986b: 244 (Velleius; characters; Japan).

  • — Smetana, 1995: 110 (Velleius; characters; Taiwan).

  • — Watanabe, 1990b: 63 (Velleius; characters; Japan).

  • — J. Li, 1993: 41 (Velleius; China).

  • — Cho, 1996b: 114 (Velleius; characters; Korea).

  • Distribution: Taiwan, Japan, Korea, China.

serraticornis Schrank, 1798, see: dilatatus Fabricius, 1787.

setosus Sharp, 1889: 29 (Velleius; Type locality: Junsai).

  • — Bernhauer and Schubert, 1916: 416 (Velleius; catalog).

  • — Naomi, 1986b: 243 (Velleius; characters; Japan).

  • — Watanabe, 1990b: 65 (Velleius; characters; Japan).

  • — Cho, 1996b: 116 (Velleius; characters; Korea).

  • Distribution: Japan, Korea.

simillimus Fairmaire, 1891a: cxci (Velleius; Type locality: Kiu-Kiang).

  • — Bernhauer and Schubert, 1916: 416 (Velleius; catalog).

  • Distribution: China.

Weiserianum [2 species; Neotropical Region]

Weiserianum Bernhauer, 1927e: 247 (species included: woltersi). Type species: Weiserianum woltersi Bernhauer, fixed by monotypy.

  • — Bruch, 1928a: 429 (catalog; Argentina).

  • — Scheerpeltz, 1933: 1420 (world catalog supplement).

  • — Scheerpeltz, 1937d: 340, 351 (characters; key to species).

  • — Blackwelder, 1944: 143 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 403 (type species: woltersi).

kuntzeni Scheerpeltz, 1937d: 348 (Weiserianum; Type locality: Brasilien, Goyaz, Jatahy, Rolle V.).

  • Distribution: Brazil.

woltersi Bernhauer, 1927e: 248 (Weiserianum; Type locality: Argentinien: Prov. Catamarca, Corral Quemado).

  • — Scheerpeltz, 1933: 1420 (Weiserianum; catalog).

  • — Scheerpeltz, 1937d: 351 (Weiserianum; characters in key).

  • Distribution: Argentina.

Subtribe Staphylinina

Staphylinina Latreille, 1802.

  • — Jakobson, 1908: 447 (characters).

  • — Jakobson, 1909: 502 (key to genera).

  • — Johansen, 1914: 371 (characters; key to Danish genera).

  • — Arnett, 1963: 275 (characters; list of North American genera).

  • — Lohse, 1964: 170, 192 (characters; key to central European species).

  • — Huth and Dettner, 1989 (defensive secretions; gland morphology).

  • — Huth and Dettner, 1990 (defensive chemicals).

  • — Newton and Thayer, 1992: 65 (subtribe of Staphylinini).

  • — Smetana, 1995e: 48 (characters).

  • — Klimaszewski, Newton, and Thayer, 1996: 151 (list of New Zealand genera; number of New Zealand species).

  • — Smetana and Davies, 2000: 12, 16 (characters).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 331, 393 (key to Nearctic genera; characters; notes).

Staphyliniae Latreille, 1802: 124 (genera included: Staphylinus, Lesteva, Anthophagus, Paederus, Stenus, Oxyporus, Astrapraeus). Type genus: Staphylinus (see Newton and Thayer, 1992: 46).

Staphylini Latreille, 1802.

  • — Thomson, 1858: 29 (characters; key to genera).

  • — Cameron, 1932: 54 (characters; key to genera of British India).

  • — Scheerpeltz, 1933: 1326 (world catalog supplement).

  • — Scheerpeltz, 1940: 42 (key to Palaearctic genera).

  • — Blackwelder, 1944: 131 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Coiffait, 1974: 353, 354 (characters; key to genera and species of western Palaearctic region).

Creophilidae Kirby, 1837: 95 (genera included: Creophilus). Type genus: Creophilus (see Newton and Thayer, 1992: 65).

  • — Newton and Thayer, 1992: 65 (synonym of Staphylinina).

Thinopininae Böving and Craighead, 1931: 30, 116, 122 (genera included: Thinopinus). Type genus: Thinopinus (see Newton and Thayer, 1992: 65).

  • — Newton and Thayer, 1992: 65 (synonym of Staphylinina).

Ocypina Hatch, 1957: 173 (genera included: Actobius, Neobisnius, Ocypus, Abemus, Parabemus, Alapsodus, Pseudocypus, Tasgius, Platydracus, Ontholestes, Philonthus, Gabrius, Cafius). Type species: Ocypus (see Newton and Thayer, 1992: 65).

  • — Arnett, 1963: 276 (characters; list of North American genera).

  • — Newton and Thayer, 1992: 65 (synonym of Staphylinina).

Staphylinina Hatch, 1957: 173 (genera included: Hadrotes, Thinopinus, Staphylinus (sensu Creophilus), Xanthopygus). Type genus: Staphylinus (see Newton and Thayer, 1992: 65).

  • — Newton and Thayer, 1992: 65 (synonym of Staphylinina).

Abemus [3 species; Palaearctic Region]

Abemus Mulsant and Rey, 1876: 242 [= 1877: 98] (species included: chloropterus, fossor). Type species: Staphylinus chloropterus Panzer, fixed by subsequent designation by Blackwelder, 1943: 443.

  • — Heyden, Reitter, and Weise, 1891: 101 (subgenus of Staphylinus; list of species of Europe and Caucasus).

  • — Heyden, Reitter, and Weise, 1906: 169 (subgenus of Staphylinus; list of species of Europe and Caucasus).

  • — Reitter, 1909: 119 (subgenus of Staphylinus; characters).

  • — Petri, 1912: 75 (subgenus of Staphylinus; list of species of Siebenbürgen).

  • — Bernhauer and Schubert, 1914: 374 (subgenus of Staphylinus).

  • — Cameron, 1932: 180 (subgenus of Staphylinus).

  • — Scheerpeltz, 1933: 1381 (subgenus of Staphylinus).

  • — Chapman, 1934 (subgenus of Staphylinus; notes on classification and distribution of some species).

  • — Blackwelder, 1943: 443 (subgenus of Staphylinus; type species: chloropterus).

  • — Blackwelder, 1952: 32 (type species).

  • — Coiffait, 1956e: 184, 187, 188 (valid genus; characters; notes).

  • — Smetana, 1958: 260, 265 (subgenus of Staphylinus; characters).

  • — Lohse, 1964: 193 (cited as valid genus; characters; central European species).

  • — Pototskaia, 1966b: 362 (subgenus of Staphylinus; larval characters in key).

  • — Pototskaia, 1967: 66 (subgenus of Staphylinus; larval characters).

  • — Coiffait, 1974: 380 (cited as valid genus; characters; species of western Palaearctic region).

  • — M. Dvořák, 1979: 116 (some species collected in Slovakia).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 92 (catalog; Poland).

  • — Tóth, 1984: 22 (cited as valid genus; characters; habitus; species of Hungary).

  • — Ciceroni and Zanetti, 1995: 30 (list of species of Italy).

  • — Smetana and Davies, 2000: 20, 23, 39 (characters; checklist of species of north temperate region).

chloropterus Panzer, 1796d: 20 (Staphylinus; Type locality: Neuwaldegg).

  • — Fabricius, 1801: 590 (Staphylinus; characters; Austria).

  • — Gravenhorst, 1802: 166 (Staphylinus; characters; Austria).

  • — Latreille, 1804: 306 (Staphylinus; characters; Austria).

  • — Gravenhorst, 1806: 121 (Staphylinus).

  • — Panzer, 1808a: 20 (Staphylinus; characters; illustrations).

  • — Lacordaire, 1835: 364 (Emus; characters; France).

  • — Nordmann, 1837: 33 (Staphylinus; France; Austria; Italy).

  • — Erichson, 1839b: 372 (Staphylinus; characters; Austria).

  • — Redtenbacher, 1849: 694 (Staphylinus; characters; Austria).

  • — Rosenhauer, 1856: 70 (Staphylinus; Andalusia).

  • — Fairmaire and Laboulbène, 1856: 506 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 546 (Staphylinus; characters; Germany; Austria).

  • — Redtenbacher, 1857: 184 (Staphylinus; characters; Austria).

  • — Fauvel, 1874: 177 [= 1874a: 403] (Staphylinus; characters; France; Austria).

  • — Redtenbacher, 1874: 204 (Staphylinus; characters; Austria).

  • — Mulsant and Rey, 1876: 244 [= 1877: 100] (Abemus; characters; France).

  • — Ganglbauer, 1895: 426 (Staphylinus; subgenus Abemus; characters; middle Europe).

  • — Reitter, 1909: 119 (Staphylinus; subgenus Abemus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; catalog).

  • — Portevin, 1929: 350 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1383 (Staphylinus; catalog).

  • — Paulian, 1941: 245 (Staphylinus; subgenus Abemus; larval characters).

  • — Coiffait, 1956e: 189 (Abemus; characters; France).

  • — Smetana, 1958: 266 (Staphylinus; subgenus Abemus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 193 (Abemus; characters; central Europe).

  • — Horion, 1965: 198 (Staphylinus; subgenus Abemus; Greece; Albania; Bosnia Hercegovina; Serbia; Slovenia; Italy ?; Croatia; Romania; Hungary; Czechoslovakia; Poland; Ukraine; Russia; Austria; Germany; France).

  • — Pototskaia, 1967: 66 (Staphylinus; subgenus Abemus; larval characters).

  • — Coiffait, 1974: 381 (Abemus; characters; Europe).

  • — Petrenko, 1980a: 59, 61 (Staphylinus; subgenus Abemus; characters; Ukraine).

  • — Tóth, 1984: 23 (Abemus; characters; Hungary).

  • — Lucht, 1987: 107 (Abemus; checklist; Poland; Czechoslovakia; Austria).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Abemus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 30 (Abemus; Italy).

  • — Ponel and Hébrard, 1997: 137 (Abemus; collecting notes; France).

  • — Pilon, 1998: 68 (Abemus; ecological notes; Italy).

  • — Staniec, 1999a: 56 (Abemus; characters of pupa; Poland).

  • — Smetana and Davies, 2000: 39 (Abemus).

  • Distribution: Spain, France, Italy, Austria, Germany, Poland; Czech Republic, Slovakia, Hungary, Slovenia, Croatia, Bosnia Hercegovina, Yugoslavia, Greece, Romania, Ukraine.

hebraeus Smetana, 1978: 118 (Staphylinus; subgenus Abemus; Type locality: Israel: Galilee Tel Dan).

  • — Smetana and Davies, 2000: 39 (Abemus).

  • Distribution: Israel.

martensi Coiffait, 1982, see: olivaceus Cameron, 1928.

olivaceus Cameron, 1928c: 564 (Staphylinus; Type locality: Sikkim: Lachen, 8500 to 9000 feet; Tangu, 11,500 feet).

  • — Cameron, 1932: 202 (Staphylinus; subgenus Goerius; characters; India).

  • — Scheerpeltz, 1933: 1398 (Staphylinus; catalog).

  • Distribution: India, Nepal.

martensi Coiffait, 1982: 69 (Thoracostrongylus; Type locality: Nepal: nördl. Dhaulagiri, Gompa/Tarakot, 3300–3400).

  • — Smetana and Davies, 2000: 39 (Abemus; synonym of olivaceus).

Agelosus [6 species; Palaearctic Region]

Agelosus Sharp, 1889: 110 (species included: carinatus). Type species: Goerius carinatus Sharp, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 400 (world catalog; 1 species).

  • — G. Müller, 1943: 96 (characters; type species: carinata).

  • — Blackwelder, 1952: 41 (type species: carinatus).

  • — Tikhomirova, 1973a: 186 (subgenus of Ocypus; checklist of species of USSR).

  • — Naomi, 1982: 128 (characters).

  • — Naomi, 1983a: 582 (characters; species of Japan).

  • — Shibata, 1984: 111 (checklist of species of Japan).

  • — Shibata, 1986a: 125 (supplemental list of species from Taiwan).

  • — Hayashi, 1991: 179 (notes).

  • — Hayashi, 1993b: 297 (characters).

  • — Smetana and Davies, 2000: 20, 35, 45 (characters; checklist of species of north temperate region).

Apecholinus Bernhauer, 1933: 36 (species included: kaiseri). Type species: Apecholinus kaiseri Bernhauer, fixed by monotypy.

  • — Blackwelder, 1952: 57 (type species: kaiseri).

  • — Smetana and Davies, 2000: 35, 45 (synonym of Agelosus).

brevipennis Naomi, 1983, see: carinatus Sharp, 1974.

carinatus Sharp, 1874: 32 (Goerius; Type locality: Maiyasama).

  • — Sharp, 1889: 111 (Agelosus; Japan).

  • — Bernhauer and Schubert, 1914: 400 (Agelosus; catalog).

  • — G. Müller, 1943: 96 (Agelosus; characters; notes; Japan).

  • — Nakane, 1963a: 93 (Agelosus; characters; habitus photograph; Japan).

  • — Naomi, 1983a: 583 (Agelosus; characters; Japan).

  • — Yuh, Paik, Kwon, and Lee, 1985: 248 (Agelosus; Korea).

  • — Hayashi, 1991: 179 (Agelosus; characters; Japan).

  • — J. Li, 1993: 37 (Agelosus; China).

  • — Smetana and Davies, 2000: 45 (Agelosus).

  • Distribution: Japan, China.

brevipennis Naomi, 1983a: 585 (Agelosus; subspecies of carinatus; Type locality: Japan: Nagasaki Pref.: Mt. Yasuman, Hirado Is.).

  • — Hayashi, 1991: 179 (Agelosus; subspecies of carinatus).

chinensis J. Li, 1992: 58 (Agelosus; Type locality: China: Liaoning Province: Qianshan).

  • — J. Li, 1993: 37 (Agelosus; China).

  • — Smetana and Davies, 2000: 45 (Agelosus).

  • Distribution: China.

fraternus Fairmaire, 1891a: cxci (Ocypus; Type locality: Tchang-Yang).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; subgenus Ocypus; catalog).

  • — G. Müller, 1943: 95, 97 (Agelosus; characters; notes; China).

  • — Hayashi, 1991: 184 (Agelosus; notes).

  • — Smetana and Davies, 2000: 45 (Agelosus).

  • Distribution: China.

kaiseri Bernhauer, 1933: 37 (Apecholinus; Type locality: Westchina: Szetschwan, Kinfushan).

  • — Smetana and Davies, 2000: 35 (Agelosus).

  • — Smetana and Davies, 2000: 45 (Agelosus).

  • Distribution: China.

masaoi Hayashi, 1991, see: unicolor Naomi, 1983.

ohkurai Hayashi, 1978: 29 (Agelosus; Type locality: Taiwan: Chiai Hsieng).

  • — Hayashi, 1991: 182 (Agelosus; characters; Taiwan).

  • — Smetana and Davies, 2000: 45 (Agelosus).

  • Distribution: Taiwan.

unicolor Naomi, 1983a: 585 (Agelosus; subspecies of ohkurai; Type locality: Japan: Kumamoto Pref.: Shirazô, Itsukimura).

  • — Hayashi, 1991: 183 (Agelosus; valid species; characters; Japan).

  • — Smetana and Davies, 2000: 45 (Agelosus).

  • Distribution: Japan.

masaoi Hayashi, 1991: 183 (Agelosus; subspecies of unicolor; Type locality: Japan: Kochi Pref.: Ashizuri).

  • — Smetana and Davies, 2000: 45 (Agelosus; subspecies of unicolor).

Amichorus [4 species; Neotropical Region]

Amichorus Sharp, 1884: 390 (species included: fauveli, cyaneus, vividus). Type species: Amichorus fauveli Sharp, fixed by subsequent designation by R. Lucas, 1920: 89.

  • — Bernhauer and Schubert, 1914: 366 (world catalog; 3 species).

  • — Scheerpeltz, 1933: 1376 (world catalog supplement).

  • — Blackwelder, 1944: 136 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 49 (type species).

cyaneus Sharp, 1884: 390 (Amichorus; Type locality: Mexico, Mirador).

  • — Bernhauer and Schubert, 1914: 366 (Amichorus; catalog).

  • Distribution: Mexico.

divinus Bernhauer, 1917d: 35 (Amichorus; Type locality: Costa Rica: Orosi, 1200 m; Cartago, 1000 m).

  • — Scheerpeltz, 1933: 1376 (Amichorus; catalog).

  • Distribution: Costa Rica.

fauveli Sharp, 1884: 390 (Amichorus; Type locality: Mexico, Mirador).

  • — Bernhauer and Schubert, 1914: 366 (Amichorus; catalog).

  • Distribution: Mexico.

vividus Sharp, 1884: 391 (Amichorus; Type locality: Panama, Volcan de Chiriqui between 4000 and 6000 feet).

  • — Bernhauer and Schubert, 1914: 366 (Amichorus; catalog).

  • Distribution: Panama.

Apostenolinus [1 species; Palaearctic Region]

Apostenolinus Bernhauer, 1934: 9 (subgenus of Staphylinus; species included: cariniceps). Type species: Staphylinus (Apostenolinus) cariniceps Bernhauer, fixed by monotypy.

  • — Blackwelder, 1943: 443 (subgenus of Staphylinus; type species: cariniceps).

  • — Blackwelder, 1952: 59 (subgenus of Platydracus; type species: cariniceps).

  • — Smetana and Davies, 2000: 20, 29, 42 (valid genus; characters; checklist of species of north temperate region).

cariniceps Bernhauer, 1934: 9 (Staphylinus; subgenus Apostenolinus; Type locality: Szechuan: Giufu-Shan, 1500–2000 m).

  • — Blackwelder, 1952: 59 (type species of Apostenolinus, a subgenus of Platydracus).

  • — Smetana and Davies, 2000: 29, 42 (Apostenolinus; notes).

  • Distribution: China.

Ascialinus [1 species; Palaearctic Region]

Ascialinus Bernhauer, 1933: 34 (subgenus of Staphylinus; species included: beckeri). Type species: Staphylinus (Ascialinus) beckeri Bernhauer, fixed by monotypy.

  • — Blackwelder, 1952: 63 (subgenus of Platydracus; type species: beckeri).

  • — Smetana and Davies, 2000: 20, 29, 42 (valid genus; characters; checklist of species of north temperate region).

beckeri Bernhauer, 1933: 34 (Staphylinus; subgenus Ascialinus; Type locality: Szetschwan, Westchina: Kinfushan).

  • — Blackwelder, 1952: 63 (type species of Ascialinus, a subgenus of Platydracus).

  • — Smetana and Davies, 2000: 29, 42 (Ascialinus; notes).

  • Distribution: China.

Bafutella [1 species; Ethiopian Region]

Bafutella Levasseur, 1968: 1388 (species included: mirei). Type species: Bafutella mirei Levasseur, fixed by original designation and monotypy.

mirei Levasseur, 1968: 1390 (Bafutella; Type locality: Cameroun: forêt de Bafut, Nguemba).

  • Distribution: Cameroon.

Barygnathus [1 species; Oriental Region]

Barygnathus Bernhauer, 1902b: 31 (species included: opacus). Type species: Barygnathus opacus Bernhauer, fixed by monotypy.

  • — Fauvel, 1902a: 41, 42 (characters in key, notes).

  • — Bernhauer and Schubert, 1914: 395 (world catalog; 1 species).

  • — Cameron, 1925: 70 (catalog of Indian species).

  • — Cameron, 1932: 245 (characters).

  • — Scheerpeltz, 1933: 1410 (world catalog supplement).

  • — Blackwelder, 1952: 73 (type species: opacus).

opacus Bernhauer, 1902b: 31 (Barygnathus; Type locality: Nalanda).

  • — Bernhauer and Schubert, 1914: 395 (Barygnathus; catalog).

  • — Cameron, 1932: 246 (Barygnathus; characters; Ceylon).

  • — Scheerpeltz, 1933: 1410 (Barygnathus; catalog).

  • Distribution: Sri Lanka.

Cafioquedius [1 species; Australian Region]

Cafioquedius Sharp, 1886a: 376 (species included: gularis). Type species: Cafioquedius gularis Sharp, fixed by monotypy.

  • — Broun, 1893: 1033 (charactes).

  • — Bernhauer and Schubert, 1916: 437 (world catalog; 1 species).

  • — Blackwelder, 1952: 87 (type species: gularis).

gularis Sharp, 1886a: 377 (Cafioquedius; Type locality: New Zealand, North Island; Bealey).

  • — Broun, 1893: 1033 (Cafioquedius; characters; New Zealand).

  • — Bernhauer and Schubert, 1916: 437 (Cafioquedius; catalog).

  • — Klimaszewski, Newton, and Thayer, 1996: 151 (Cafioquedius; endemic to New Zealand).

  • Distribution: New Zealand.

Chapmaniella [1 species; Neotropical Region]

Chapmaniella Bernhauer, 1934c: 145 (species included: miranda). Type species: Chapmaniella miranda Bernhauer, fixed by monotypy.

  • — Blackwelder, 1944: 145 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 98 (type species: miranda).

miranda Bernhauer, 1934c: 146 (Chapmaniella; Type locality: Brasilien: Santa Catarina, Hansa).

  • Distribution: Brazil.

Creophilus [13 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions]

Creophilus Leach, 1819: 172 (species included: maxillosus). Type species: Staphylinus maxillosus Linné, 1785, fixed by original designation according to action of Commission in Opinion 546 (ICZN, 1959).

  • — Curtis, 1829: 24 (catalog; Britain).

  • — Stephens, 1829a: 22 (catalog; Britain).

  • — Stephens, 1829: 274 (catalog; Britain).

  • — Mannerheim, 1830: 7, 20 (characters).

  • — Mannerheim, 1831: 421, 434 (characters).

  • — Stephens, 1832: 200, 202 (characters).

  • — Lacordaire, 1835: 358 (synonym of Emus).

  • — Nordmann, 1837: 21.

  • — Westwood, 1838a: 15 (characters; type species).

  • — Erichson, 1839b: 347 (synonym of Staphylinus).

  • — Stephens, 1839: 384 (characters).

  • — Laporte, 1840: 173 (characters).

  • — Lacordaire, 1854: 76 (synonym of Staphylinus).

  • — Jacquelin du Val, 1856: 34 (characters).

  • — Kraatz, 1857d: 528 (characters).

  • — Redtenbacher, 1857: 984 (characters).

  • — Thomson, 1858: 29 (characters).

  • — G. Waterhouse, 1858: 23 (catalog of British species).

  • — Thomson, 1859: 23 (characters; type species).

  • — Schaum, 1859: 26 (catalog; European species).

  • — Thomson, 1860: 140 (characters).

  • — LeConte, 1861a: 64 (characters).

  • — Gredler, 1863: 101 (infrageneric group of Staphylinus; in list of species from Tirol).

  • — LeConte, 1863: 23 (in list of North American species).

  • — Wencker and Silbermann, 1866: 28 (in list of species; collecting notes; France).

  • — Gemminger and Harold, 1868: 575 (catalog).

  • — Jacquelin du Val, 1868: 66 (group of Staphylinus; catalog; European species).

  • — Sharp, 1871d: 11 (in list of British species).

  • — Redtenbacher, 1874: 203 (characters).

  • — Mulsant and Rey, 1876: 220 [= 1877: 76] (characters).

  • — Provancher, 1877: 245 (characters).

  • — Horn, 1879: 199 (characters).

  • — Heyden, 1880: 71 (subgenus of Emus; in list of species of Siberian region).

  • — Duvivier, 1883: 141 (catalog).

  • — Sharp, 1884: 371 (notes; Mexican and Central American species).

  • — Lynch, 1884: 128 (characters).

  • — Olliff, 1887: 491 (characters; Australian species).

  • — Sharp, 1888: 247 (characters).

  • — Heyden, Reitter, and Weise, 1891: 101 (in list of species of Europe and Caucasus).

  • — Heyden, 1893: 41 (in list of species of Siberian region).

  • — Ganglbauer, 1895: 414 (adult and larval characters).

  • — Heyden, 1896: 32 (in list of species of Siberian region).

  • — Germain, 1903: 410 (characters).

  • — Heyden, Reitter, and Weise, 1906: 172 (in list of species of Europe and Caucasus).

  • — Reitter, 1909: 116 (characters).

  • — Blatchley, 1910: 374 (characters).

  • — Petri, 1912: 76 (in list of species of Siebenbürgen).

  • — Bernhauer and Schubert, 1914: 397 (world catalog; 13 species).

  • — Johansen, 1914: 372 (characters).

  • — Leng, 1920: 108 (catalog of North American species).

  • — Winkler, 1925: 387 (catalog for Palaearctic region).

  • — Cameron, 1925: 71 (catalog of Indian species).

  • — Porta, 1926: 93 (characters; species of Italy).

  • — Portevin, 1929: 347 (characters).

  • — Roubal, 1930: 385 (catalog; Slovakia).

  • — Cameron, 1932: 258 (characters; key to species of British India).

  • — Joy, 1932: 114 (key to British species).

  • — Scheerpeltz, 1933: 1412 (world catalog supplement).

  • — Normand, 1935: 372 (in list of species of Tunisia).

  • — Blackwelder, 1943: 447 (characters; type species: maxillosus; in keys to species of West Indies).

  • — Blackwelder, 1944: 140 (in checklist of species from Mexico, Central and South America, and the West Indies).

  • — Steel, 1949d: 61 (key to Australian species).

  • — Kloet and Hincks, 1945: 167 (in list of British species).

  • — Tottenham, 1949: 375, 421 (type species: maxillosus Linné; notes; in checklist of British species).

  • — Blackwelder, 1952: 111 (type species: maxillosus).

  • — V. Hansen, 1952: 111 (characters; Danish species).

  • — Coiffait, 1956e: 184, 186, 187 (characters in key; notes; key to species).

  • — Hatch, 1957: 212 (characters; species of Pacific Northwest).

  • — Tottenham, 1957a: 87 (proposed fixation as type species of Creophilus).

  • — Kocher, 1958: 155 (in checklist of species; Morocco).

  • — Smetana, 1958: 309 (characters; type species: maxillosus).

  • — Ferreira, 1962a: 18 (in list of Portuguese species).

  • — Székessy, 1963: 58 (characters).

  • — Palm, 1963b: 121 (characters; species of Sweden).

  • — Lohse, 1964: 192 (characters; central European species).

  • — Moore, 1964c (larval characters).

  • — Scheerpeltz, 1965: 278, 279 (characters; type species: maxillosus).

  • — Pototskaia, 1967: 64 (larval characters).

  • — Coiffait and Saiz, 1968: 363 (notes; key to Chilean species).

  • — Scheerpeltz, 1968a: 86 (catalog of Austrian species).

  • — Kasule, 1970: 65 (larval characters).

  • — Bordoni, 1973j: 697 (in list of species from islands near Sicily).

  • — Tikhomirova, 1973a: 188 (in checklist of species of USSR).

  • — Coiffait, 1974: 558 (characters; key to species of western Palaearctic region).

  • — Moore and Legner, 1975: 41 (catalog; North America north of Mexico).

  • — Pope, 1977: 30 (in list of British species).

  • — Topp, 1978: 311 (larval characters).

  • — Frank, 1979a (some larval characters and their use in classification).

  • — Moore and Legner, 1979: 33 (characters; notes).

  • — Muona, 1979: 15 (in list of Scandinavian species).

  • — Uhlig, Vogel, and Sieber, 1980: 247 (some species collected in Germany).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 88 (catalog; Poland).

  • — Uhlig and Vogler, 1981: 118 (in list of some species collected in Germany; habitat and collecting notes).

  • — Naomi, 1982: 128 (characters; species of Japan).

  • — Tóth, 1984: 33 (characters; habitus; species of Hungary).

  • — Shibata, 1984: 102 (in checklist of species of Japan).

  • — Frank, 1986a: 370 (in checklist of species of Florida).

  • — Muona and Viramo, 1986: 13 (in list of species of northeastern Finland).

  • — Dettner, 1987: 30 (defensive glands).

  • — Drugmand 1987b (in comments on distribution of Belgian species).

  • — Angelini, 1991: 191 (in list of species from high elevations of Calabria, Italy).

  • — Silfverberg, 1992: 17 (in in list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Welch, 1993: 229 (ovariole structure and development).

  • — Smetana, 1993b: 49 (in list of species of Czech Republic and Slovakia).

  • — Kanehisa, Tsumuki, and Kawazu, 1994 (chemical secretions).

  • — Ciceroni and Zanetti, 1995: 29 (in list of species of Italy).

  • — M. Hansen, 1996: 107 (in list of Danish species).

  • — Downie and Arnett, 1996: 387 (characters).

  • — Lundgren, 1998: 51 (in list of species from Florida; records for other States listed).

  • — Smetana and Davies, 2000: 18, 21, 38 (characters; in checklist of species of north temperate region).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 393 (1 Nearctic species; notes; characters in key).

Saprophilus Streubel, 1839: 136 (species included: maxillosus). Type species: Staphylinus maxillosus Linné, fixed by monotypy.

  • — Blackwelder, 1952: 344 (type species; synonym of Staphylinus; [Note: Blackwelder used Staphylinus in the sense of Creophilus]).

acuticollis Bernhauer, 1910b: 377 (Creophilus; Type locality: Mexiko).

  • — Bernhauer and Schubert, 1914: 398 (Creophilus; catalog).

  • Distribution: Mexico.

albertisi Fauvel, 1879: 95 (Emus; Type locality: Nouvelle Guinée, Fly River).

  • — Bernhauer and Schubert, 1914: 398 (Creophilus; catalog).

  • — Scheerpeltz, 1971b: 202 (Creophilus; New Guinea).

  • Distribution: New Guinea.

anonymous Sulzer, 1761, see: maxillosus Linné, 1758.

arcticus Erichson, 1839, see: maxillosus Linné, 1758.

badiipennis Nordmann, 1837, see: Torobus.

balteatus De Geer, 1774, see: maxillosus Linné, 1758.

bicinctus Mannerheim, 1843, see: maxillosus Linné, 1758.

canariensis Bernhauer, 1908, see: maxillosus Linné, 1758.

chloris Nordmann, 1837, see: Torobus.

ciliaris Stephens, 1832, see: maxillosus Linné, 1758.

ciliaroides Hatch, 1938, see: maxillosus Linné, 1758.

cinerarius Erichson, 1839, see: maxillosus Linné, 1758.

coquerelii Fauvel, 1866, see: incanus Klug, 1832.

erithacus Nordmann, 1837, see: Torobus.

erythrocephalus Fabricius, 1775: 265 (Staphylinus; Type locality: Nova Hollandia [Note: See “lectotype” designation by Steel, 1949d]).

  • — Goeze, 1777: 724 (Staphylinus; characters).

  • — Fabricius, 1781: 335 (Staphylinus; characters; New Guinea).

  • — Fabricius, 1787: 220 (Staphylinus; characters).

  • — Gmelin, 1790: 2027 (Staphylinus; characters).

  • — Fabricius, 1793: 523 (Staphylinus; characters; New Guinea).

  • — Olivier, 1795:(42): 12 (Staphylinus; characters; New Guinea).

  • — Fabricius, 1801: 593 (Staphylinus; characters; New Guinea).

  • — Turton, 1802: 511 (Staphylinus; characters; New Holland).

  • — Latreille, 1804: 293 (Staphylinus; characters).

  • — Gravenhorst, 1806: 127 (Staphylinus; characters; New Guinea).

  • — Nordmann, 1837: 23 (Creophilus; characters; New Guinea).

  • — Erichson, 1839b: 351 (Staphylinus; characters; Nova Hollandia).

  • — Fairmaire, 1849: 288 (Staphylinus; notes; Tahiti).

  • — Fauvel, 1877: 249 (Emus; characters; Australia; New Caledonia; Tonga; Tahiti).

  • — Fauvel, 1878g: 541 (Emus; Australia).

  • — Olliff, 1887: 492 (Creophilus; characters; Australia).

  • — Olliff, 1889: 81 (Creophilus; Lord Howe Island; Australia; Norfolk Island; Tonga).

  • — Fauvel, 1889b: 261 (Emus; New Caledonia).

  • — Fauvel, 1903d: 267 (Creophilus; New Caledonia; Ile Nou; Tasmania; Australia; Tongo; Tahiti; Chile).

  • — Bernhauer, 1908: 19 (Creophilus; Australia).

  • — Sharp and Muir, 1912: 498 (Creophilus; characters of male genitalia).

  • — Bernhauer and Schubert, 1914: 398 (Creophilus; catalog).

  • — Bernhauer, 1920: 16 (Creophilus; Australia).

  • — Cameron, 1924: 87 (Creophilus; New Guinea).

  • — Lea, 1925: 229 (Creophilus; Australia).

  • — Cameron, 1933e: 83 (Creophilus; Society Islands).

  • — Cameron, 1937a: 104 (Creophilus; New Guinea).

  • — Steel, 1949d: 57, 58 (Creophilus; characters; lectotype not formally designated in footnote but one of the two specimens “. . . is taken as the type . . .”; Australia).

  • — Coiffait and Saiz, 1968: 364 (Creophilus; cited as erytrocephalus; characters; Chile).

  • — Saiz, 1971b: 349 (Creophilus; cited as erytrocephalus; Chile).

  • — Bellas, Brown, and Moore, 1974 (Creophilus; defensive secretions).

  • — Coiffait, 1976: 218 (Creophilus; Tahiti).

  • — Frank, 1982: 34 (Creophilus; list of parasites).

  • — E. Matthews, 1982: 7 (Creophilus; notes; Australia).

  • — Saiz, Solervicens, and Ojeda, 1989: 50 (Creophilus; collecting notes; habitus; Chile).

  • — Nishida, 1994: 71 (Creophilus; Hawaii).

  • — Desender and Baert, 1996: 43 (Creophilus; Easter Island).

  • — Newton, 1997: 25 (Creophilus; notes; reported but evidently not established in Hawaii).

  • — Smithers, 1998: 16 (Creophilus; references; Norfolk Island).

  • Distribution: Nepal, New Guinea, Australia, Lord Howe Island, New Caledonia, Tonga, Society Islands, Hawaii, Easter Island, Chile.

unipunctatus Hope, 1831: 32 (Staphylinus; Type locality: Nepaul).

  • — Lea, 1925: 229 (Creophilus; synonym of erythrocephalus).

  • — Steel, 1949d: 57 (Creophilus; synonym of erythrocephalus).

fasciatus Laporte, 1835, see: maxillosus Linné, 1758.

fasciatus Füessly, 1775, see: maxillosus Linné, 1758.

fulvago Motschulsky, 1860, see: maxillosus Linné, 1758.

huttoni Broun, 1880, see: oculatus Fabricius, 1775.

imbecillus Sharp, 1874, see: maxillosus Linné, 1758.

imitator Cameron, 1952: 255 (Creophilus; Type locality: New Guinea: Snow Mountains: Top Camp, 2100 m).

  • Distribution: New Guinea.

incanus Klug, 1832: 137 (Staphylinus; Type locality: Madagascar).

  • — Erichson, 1839b: 350 (Staphylinus; characters; Madagascar).

  • — Bernhauer and Schubert, 1914: 398 (Creophilus; catalog).

  • — Lecoq, 1990: 206 (Creophilus; Mascarene Islands).

  • Distribution: Madagascar, Mascarene Islands.

coquerelii Fauvel, 1866e: 317 (Staphylinus; subgenus Creophilus; Type locality: La Réunion: Bourbon).

  • — Fauvel, 1867b: 67 [= 1868a: 71] (Staphylinus; [Note: Cited as though a new species]; subgenus Creophilus; characters; Réunion Island).

  • — Bernhauer and Schubert, 1914: 398 (Creophilus; cited as coquereli; synonym of incanus).

  • — Lecoq, 1990: 206 (Creophilus; cited as coquereli; synonym of incanus).

insularis Fauvel, 1879: 94 (Emus; Type locality: Batcian; Ceram; Sumatra).

  • — Bernhauer and Schubert, 1914: 398 (Creophilus; catalog).

  • — Bernhauer, 1926f: 319 (Creophilus; Buru).

  • — Scheerpeltz, 1933: 1412 (Creophilus; catalog; Moluccas; New Guinea).

  • Distribution: Indonesia, New Guinea.

lanio Erichson, 1839b: 352 (Staphylinus; Type locality: Terra Van Diemenii).

  • — Gravenhorst, 1806: 126 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 398, cited oculatus variety Gravenhorst as a synonym of lanio but Gravenhorst did not describe the species, he attributed it to Fabricius]; characters).

  • — Fauvel, 1877: 250 (Emus; characters; Australia).

  • — Fauvel, 1878g: 541 (Emus; Australia).

  • — Bernhauer and Schubert, 1914: 398 (Creophilus; catalog).

  • — Lea, 1925: 229 (Creophilus; synonym of erythrocephalus).

  • — Steel, 1949d: 58 (Creophilus; characters; Australia).

  • Distribution: Australia.

maxillosus Linné, 1758: 421 (Staphylinus; Type locality: Europa).

  • — Linné, 1760: 230 (Staphylinus; characters; Sweden).

  • — O. Müller, 1764: 23 (Staphylinus; characters).

  • — Pontoppidans, 1765: 211 (Staphylinus; Denmark).

  • — Linné, 1767: 683 (Staphylinus; characters).

  • — Ström, 1765: 403 (Staphylinus; characters; Denmark) [= 1767: 362].

  • — Füessly, 1775: 21 (Staphylinus).

  • — Fabricius, 1775: 265 (Staphylinus; characters).

  • — Goeze, 1777: 715 (Staphylinus; characters).

  • — Fabricius, 1781: 334 (Staphylinus; characters; Europe).

  • — Herbst, 1784: 149 (Staphylinus; Germany).

  • — Fourcroy, 1785: 164 (Staphylinus; characters; France).

  • — Fabricius, 1787: 220 (Staphylinus; characters).

  • — Paykull, 1789: 5 (Staphylinus; redescription; notes; Sweden).

  • — Rossi, 1790: 246 (Staphylinus; characters; Italy).

  • — Villers, 1789: 410 (Staphylinus; characters).

  • — Gmelin, 1790: 2026 (Staphylinus; characters).

  • — Fabricius, 1793: 521 (Staphylinus; characters).

  • — Olivier, 1795:(42): 9 (Staphylinus; characters).

  • — Panzer, 1795: 353 (Staphylinus; characters; Germany).

  • — Schrank, 1798: 639 (Staphylinus; characters; notes).

  • — Cederhjelm, 1798: 117 (Staphylinus; characters).

  • — Paykull, 1800: 369 (Staphylinus; characters; Sweden).

  • — Fabricius, 1801: 592 (Staphylinus; characters).

  • — Turton, 1802: 510 (Staphylinus; characters; Europe).

  • — Gravenhorst, 1802: 3 (Staphylinus; characters).

  • — Marsham, 1802: 498 (Staphylinus; characters; Britain).

  • — Latreille, 1804: 293 (Staphylinus; characters).

  • — Gravenhorst, 1806: 126 (Staphylinus; characters).

  • — Panzer, 1808: 2 (Staphylinus; characters; illustrations).

  • — Gyllenhal, 1810: 279 (Staphylinus; characters; Sweden).

  • — Leach, 1819: 172 (Creophilus; type species of Creophilus).

  • — Zetterstedt, 1828: 69 (Staphylinus; characters; Lapland).

  • — Curtis, 1829: 24 (Creophilus; catalog; Britain).

  • — Latreille, 1829: 434 (Staphylinus; characters).

  • — C. Sahlberg, 1830: 308 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 20 (Creophilus; Finland; North America).

  • — Mannerheim, 1831: 434 (Creophilus; Finland; North America).

  • — Stephens, 1832: 202 (Creophilus; characters; England).

  • — Brullé, 1832: 129 (Creophilus; notes; Greece).

  • — Ménétriés, 1832: 142 (Staphylinus; Caucasus-Persia region).

  • — Griffith and Pidgeon, 1832: 291 (Staphylinus; characters).

  • — Runde, 1835: 1 (Creophilus; characters; Germany).

  • — Lacordaire, 1835: 358 (Emus; characters; France).

  • — Nordmann, 1837: 21 (Creophilus; characters; Europe; Siberia; North America).

  • — Holme, 1837: 63 (Creophilus; notes; Scilly Islands).

  • — Westwood, 1838a: 15 (Creophilus; type species).

  • — Zetterstedt, 1838: 60 (Staphylinus; characters; habitat; Lapland).

  • — Brullé, 1839: 60 (Staphylinus; Canary Islands).

  • — O. Costa, 1839b: 118 (Staphylinus; characters; Italy).

  • — Erichson, 1839a: 432 (Staphylinus; characters; Germany).

  • — Heer, 1839: 249 (Staphylinus; characters; Switzerland).

  • — Erichson, 1839b: 348 (Staphylinus; characters; Europe).

  • — Stephens, 1839: 386 (Creophilus; characters).

  • — Laporte, 1840: 173 (Creophilus; characters; France).

  • — Holme, 1842: 114, 118, 119 (Creophilus; notes).

  • — Dufour, 1843: 32 (Staphylinus; characters; France).

  • — Westwood, 1845: 45 (Creophilus; Linné confused two species under this name; characters; notes).

  • — Kolenati, 1846: 17 (Staphylinus; cis et trans Caucasum).

  • — P. Lucas, 1846: 108 (Staphylinus; notes; Algeria).

  • — Jacobsen, 1849: 681 (Staphylinus; characters; Denmark).

  • — Ménétriés, 1849: 43 (Staphylinus; Turcomanie).

  • — Hochhuth, 1849: 112 (Staphylinus; characters).

  • — Redtenbacher, 1849: 694 (Staphylinus; characters; Austria).

  • — Küster, 1853: 42 (Staphylinus; characters; Europe).

  • — Hardy, 1851: 21 (Staphylinus; subgenus Creophilus; notes; England).

  • — Wollaston, 1854: 579 (Staphylinus; characters; notes; Madeira Islands).

  • — Rosenhauer, 1856: 70 (Staphylinus; notes; Andalusia).

  • — Fairmaire and Laboulbène, 1856: 505 (Staphylinus; characters; France).

  • — Staudinger, 1857: 286 (Staphylinus; notes; Iceland).

  • — Wollaston, 1857: 188 (Staphylinus; habitat; Madeira and Canary Islands).

  • — Kraatz, 1857d: 529 (Creophilus; characters; Germany).

  • — Redtenbacher, 1857: 184 (Staphylinus; characters; Austria).

  • — Kraatz, 1858a: 60 (Creophilus; Greece).

  • — Peyron, 1858: 422 (Staphylinus; Turkey).

  • — Kraatz, 1859: 68 (Creophilus; characters; India orientalis).

  • — Thomson, 1859: 23 (cited as maxillaris; type species of Creophilus).

  • — Thomson, 1860: 141 (Creophilus; characters; Scandinavia).

  • — Hochhuth, 1862: 47 (Creophilus; Russia).

  • — Wollaston, 1864: 564 (Creophilus; notes; Canary Islands).

  • — Wollaston, 1865: 486 (Creophilus; habitat; Madeira Islands; Canary Islands).

  • — Crotch, 1867a: 382 (Creophilus; Azores).

  • — Solsky, 1871: 158 (Creophilus; characters; Russia).

  • — Solsky, 1871c: 346 (Creophilus; notes; Siberia).

  • — Sharp, 1874: 27 (Creophilus; characters; notes; Japan).

  • — Fauvel, 1874: 169 [= 1874a: 395] (Emus; characters; France; Europe; Madeira; Barbary; Egypt; Ethiopia; Syria; Turkey; Caucasus; Iran; Bukara; India; China; Mongolia; Japan; Siberia; Kuriles; Kamchatka; North America; Mexico<†125>.

  • — Redtenbacher, 1874: 203 (Creophilus; characters; Austria).

  • — Solsky, 1874: 171 (Creophilus; characters; Turkestan).

  • — Seidlitz, 1875: 268 (Creophilus; characters; Baltic region).

  • — Fauvel, 1876a: 70 [= 1876: 255] (Emus; Azores; Sandwich Islands).

  • — Mulsant and Rey, 1876: 222 [= 1877: 78] (Creophilus; characters; larval characters; France).

  • — J. Sahlberg, 1876: 9 (Creophilus; Finland).

  • — Wollaston, 1877: 26 (Creophilus; characters; St. Helena).

  • — Fauvel, 1878: 118 [= 1878a: 38] (Emus; North Africa).

  • — J. Sahlberg, 1880: 67 (Creophilus; Siberia).

  • — Blackburn and Sharp, 1885: 222 (Creophilus; habitat; Hawaii).

  • — Fauvel, 1886: 50 [= 1886a: 42] (Emus; Algeria).

  • — Fowler, 1888: 247 (Creophilus; characters; Britain).

  • — Seidlitz, 1891: 395 (Creophilus; characters; Europe).

  • — Seidlitz, 1891a: 420 (Creophilus; characters; Europe).

  • — Ganglbauer, 1895: 415 (Creophilus; characters; Palaearctic, Ethiopian, and Indian regions; North America; Mexico).

  • — Fauvel, 1897d: 293 (Emus; Barbary; Azores; Madeira; Canaries).

  • — Fauvel, 1902b: 102 (Creophilus; Barbary; Egypt; Azores; Madeira; Canary Islands).

  • — Poppius, 1905: 9 (Creophilus; notes; Orkney Islands; Shetland Islands).

  • — Sainte-Claire Deville, 1906: 101 (Creophilus; Corsica).

  • — Sharp, 1908: 548 (Creophilus; Hawaii).

  • — Poppius, 1909: 28 (Ontholestes; Lena Valley, Siberia).

  • — Reitter, 1909: 117 (Creophilus; characters; Germany).

  • — Matsumura, 1911: 112 (Creophilus; Japan).

  • — Poppius, 1913: 10 (Creophilus; Faroes; Iceland).

  • — Bernhauer and Schubert, 1914: 398 (Creophilus; catalog).

  • — Johansen, 1914: 373 (Creophilus; characters; Denmark).

  • — Eichelbaum, 1915a: 92 (Creophilus; abdominal segmentation; apical segements).

  • — Šulc, 1922 (Creophilus; osmeterium; anatomy; histology; physiology).

  • — Fall, 1926: 150 (Creophilus; Alaska).

  • — Portevin, 1929: 347 (Creophilus; characters; France).

  • — Gridelli, 1930: 76 (Creophilus; Libya).

  • — Cameron, 1932: 259 (Creophilus; characters; India).

  • — Scheerpeltz, 1933: 1412 (Creophilus; catalog).

  • — Kano, 1933: 98 (Creophilus; Kurile Islands).

  • — Aleksandrov, 1934: 153 (Creophilus; China).

  • — Cameron, 1935b: 289 (Creophilus; India).

  • — Blackwelder, 1936a (Creophilus; morphology).

  • — C. Koch, 1936a: 75 (Creophilus; Libya).

  • — West, 1937: 39 (Creophilus; Faroe Islands).

  • — Hatch, 1938: 148 (Creophilus; characters; notes; Aleutian Islands).

  • — Bernhauer, 1940c: 5 (Creophilus; Azores; Madeira).

  • — Paulian, 1941: 251 (Creophilus; larval characters).

  • — Méquignon, 1942a: 22 (Creophilus; Azores).

  • — Blackwelder, 1943: 447 (Creophilus; characters; Cuba; Isle of Pines; Jamaica).

  • — Hinton, 1945: 73 (Creophilus; characters).

  • — Cerruti, 1951: 16 (Creophilus; India).

  • — V. Hansen, 1952: 111 (Creophilus; characters; Denmark).

  • — Coiffait, 1954: 166 (Creophilus; Canary Islands).

  • — Coiffait, 1956e: 188 (Creophilus; characters; notes).

  • — Scheerpeltz, 1956c: 1101 (Creophilus; Cyprus).

  • — Smetana, 1958: 311 (Creophilus; characters; ecology; Czechoslovakia).

  • — Larsson and Gígja, 1959: 88 (Creophilus; biological notes; Iceland).

  • — ICZN, 1959 (fixed as type species of Creophilus; name placed on Official List of Specific Names in Zoology).

  • — Nakane, 1963a: 93 (Staphylinus; characters; habitus photograph; Japan).

  • — Smetana, 1963b: 39 (Creophilus; Madeira).

  • — Smetana, 1963c: 299 (Creophilus; Mongolia).

  • — Scheerpeltz, 1963: 14 (Creophilus; Afghanistan).

  • — Palm, 1963b: 121 (Creophilus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 192 (Creophilus; characters; central Europe).

  • — Scheerpeltz, 1964: 28 (Creophilus; Mongolia).

  • — Scheerpeltz, 1965c: 510 (Creophilus; Sicily).

  • — Smetana, 1965e: 175 (Creophilus; Mongolia).

  • — Horion, 1965: 243 (Creophilus; notes; all Europe).

  • — Pototskaia, 1967: 64 (Creophilus; larval characters).

  • — Smetana, 1967d: 210 (Creophilus; Mongolia).

  • — Smetana, 1967i: 557 (Creophilus; Turkey).

  • — Coiffait and Saiz, 1968: 364 (Creophilus; characters; Chile).

  • — Szujecki, 1968a: 733 (Creophilus; Poland).

  • — Dajoz and Caussanel, 1968 (Creophilus; natural history, anatomy, and morphology of larva, pupa, and adult).

  • — Bolov, 1969a: 328 (Creophilus; collecting notes; Russia).

  • — Smetana, 1970: 61 (Creophilus; distributional summary; Azores).

  • — Saiz, 1971b: 350 (Creophilus; notes; Chile).

  • — Fagel, 1972: 104 (Creophilus; notes; Sainte Helene).

  • — Scheerpeltz, 1972: 22 (Creophilus; Argentina).

  • — Coiffait, 1973a: 275 (Creophilus; Morocco).

  • — Coiffait, 1974: 560 (Creophilus; characters; Holarctic region).

  • — Bordoni, 1974b: 6 (Creophilus; Italy).

  • — Smetana 1975c: 173 (Creophilus; Mongolia).

  • — Bordoni, 1975a: 147 (Creophilus; Afghanistan).

  • — Shilov, 1975a: 376 [= 1975b: 99] (Creophilus; Komi region of Russia).

  • — Coiffait, 1977b: 212 (Creophilus; Bhutan).

  • — Pope, 1977: 30 (Creophilus; Britain).

  • — Hoebeke, 1978: 508 (Creophilus; parasitized by proctotrupid).

  • — Coiffait, 1979d: 552 (Creophilus; Afghanistan).

  • — Hinton, 1981: 662 (Creophilus; characters of egg).

  • — Naomi, 1982: 129 (Creophilus; characters; Japan).

  • — Coiffait, 1982: 32 (Creophilus; Nepal).

  • — Coiffait, 1982a: 233 (Creophilus; Nepal).

  • — Bordoni, 1982a: 140 (Creophilus; cave collection; Sardinia).

  • — Frank, 1982: 35 (Creophilus; list of parasites).

  • — Tóth, 1984: 34 (Creophilus; characters; Hungary).

  • — Yuh, Paik, Kwon, and Lee, 1985: 247 (Creophilus; Korea).

  • — Segers, 1986: 49 (Creophilus; checklist of species; Belgium).

  • — Ádám, 1987: 148 (Creophilus; collecting notes; Hungary).

  • — Lucht, 1987: 107 (Creophilus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Huth and Dettner, 1989: 154 (Creophilus; defensive secretions).

  • — Israelson, 1990: 3 (Creophilus; Azores).

  • — Borges, 1990: Table II (Creophilus; Azores).

  • — Huth and Dettner, 1990 (Creophilus; defensive chemicals).

  • — Pilon, 1991: 105 (Creophilus; notes; Italy).

  • — Pilon and Zanetti, 1991: 69 (Creophilus; Italy).

  • — Campbell and Davies, 1991: 120 (Creophilus; checklist of species; Alaska and Canada).

  • — Welch, 1993: 229 (Creophilus; ovariole number and ovary structure).

  • — J. Li, 1993: 39 (Creophilus, cited as Crephilus; cited as maxillatus; China).

  • — Smetana, 1993b: 49 (Creophilus; Czech Republic; Slovakia).

  • — Dearborn and Donahue, 1994: 15 (Creophilus; Maine).

  • — Kanehisa, Tsumuki, and Kawazu, 1994 (Creophilus; chemical secretions).

  • — Nishida, 1994: 71 (Creophilus; Hawaii).

  • — Sadler and Dugmore, 1995: 144 (Creophilus; habitat notes; Iceland).

  • — Ciceroni and Zanetti, 1995: 29 (Creophilus; Italy).

  • — Sparacio, 1995: 163 (Creophilus; characters; notes; Sicily).

  • — Staniec, 1996: 118 (Creophilus; pupal characters).

  • — Newton, 1997: 24 (Creophilus; notes; Hawaii).

  • — K. Bland, 1997: 204 (Creophilus; eaten by toad; Scotland).

  • — Nishida, 1997: 60 (Creophilus; Hawaii).

  • — Mikat, Fremuth, and Prouza, 1997: 111 (Creophilus; Czech Republic).

  • — Bain, 1998: 39, 42, 43 (Creophilus; 17th century remains; introduced; Massachusetts).

  • — Pilon, 1998: 64 (Creophilus; ecological notes; Italy).

  • — Schmidt, 1999: 108 (Creophilus; food for adults and larvae used for rearing species; rearing methods).

  • — Ryabukhin, 1999: 67 (Creophilus; catalog; habitat; distribution; northeastern Russia).

  • — Smetana and Davies, 2000: 38 (Creophilus).

  • — Naomi, Kuranishi, Saito, and Maruyama, 2000: 110 (Creophilus; Kuril Islands and Kamchatka).

  • — Assing and Wunderle, 2001: 37 (Creophilus; Cyprus).

  • Distribution: Wide-spread: northern Africa, Eur-Asia, North America, West Indies, Argentina, Chile and many continental and oceanic islands.

anonymus Sulzer, 1761: 17 (Staphylinus; Type locality: Europa).

  • — Erichson, 1839b: 348 (Staphylinus; synonym of maxillosus).

  • — Kraatz, 1857d: 529 (Creophilus; synonym of maxillosus).

  • — Ganglbauer, 1895: 415 (Creophilus; synonym of maxillosus).

  • — Bernhauer and Schubert, 1914: 398 (Creophilus; synonym of maxillosus).

  • — Cameron, 1932: 259 (Creophilus; synonym of maxillosus).

  • — Blackwelder, 1943: 447 (Creophilus; synonym of maxillosus).

  • — Smetana, 1958: 311 (Creophilus; synonym of maxillosus).

  • — Coiffait, 1974: 560 (Creophilus; synonym of maxillosus).

tertius Schaeffer, 1766: xx, 1 (Staphylinus; [Note: Nomen nudum, Article 11.4]; Type locality: Ratisbon).

  • — Paykull, 1789: 6 (Staphylinus; reference to this name cited under maxillosus).

  • — Erichson, 1839b: 348 (Staphylinus; synonym of maxillosus).

  • — Kraatz, 1857d: 529 (Creophilus; synonym of maxillosus).

  • — Blackwelder, 1943: 447 (Creophilus; synonym of maxillosus).

balteatus De Geer, 1774: 18 (Staphylinus; Type locality: Not cited).

  • — O. Müller, 1776: 98 (Staphylinus; characters).

  • — Fabricius, 1781: 334 (Staphylinus; synonym of maxillosus).

  • — Harrer, 1784: 251 (Staphylinus; characters).

  • — Paykull, 1789: 6 (Staphylinus; synonym of maxillosus).

  • — Gmelin, 1790: 2026 (Staphylinus; synonym of maxillosus).

  • — Olivier, 1795:(42): 10 (Staphylinus; synonym of maxillosus).

  • — Gravenhorst, 1802: 3 (Staphylinus; synonym of maxillosus Linné).

  • — Marsham, 1802: 498 (Staphylinus; synonym of maxillosus Linné).

  • — Gyllenhal, 1810: 279 (Staphylinus; synonym of maxillosus Linné).

  • — C. Sahlberg, 1830: 308 (Staphylinus; synonym of maxillosus Linné).

  • — Erichson, 1839b: 348 (Staphylinus; synonym of maxillosus).

  • — Kraatz, 1857d: 529 (Creophilus; synonym of maxillosus).

  • — Ganglbauer, 1895: 415 (Creophilus; synonym of maxillosus).

  • — Bernhauer and Schubert, 1914: 398 (Creophilus; synonym of maxillosus).

  • — Cameron, 1932: 259 (Creophilus; synonym of maxillosus).

  • — Blackwelder, 1943: 447 (Creophilus; synonym of maxillosus).

  • — Smetana, 1958: 311 (Creophilus; synonym of maxillosus).

  • — Coiffait, 1974: 560 (Creophilus; synonym of maxillosus).

fasciatus Füessly, 1775: 21 (Staphylinus; Type locality: Switzerland).

  • — Goeze, 1777: 724 (Staphylinus; characters).

  • — Gravenhorst, 1802: 3 (Staphylinus; synonym of maxillosus Linné).

  • — Erichson, 1839b: 348 (Staphylinus; synonym of maxillosus).

  • — Kraatz, 1857d: 529 (Creophilus; synonym of maxillosus).

  • — Ganglbauer, 1895: 415 (Creophilus; synonym of maxillosus).

  • — Bernhauer and Schubert, 1914: 398 (Creophilus; synonym of maxillosus).

  • — Cameron, 1932: 259 (Creophilus; synonym of maxillosus).

  • — Blackwelder, 1943: 447 (Creophilus; synonym of maxillosus).

  • — Smetana, 1958: 311 (Creophilus; synonym of maxillosus).

  • — Coiffait, 1974: 560 (Creophilus; synonym of maxillosus).

nebulosus Geoffroy, 1785: 165 (Staphylinus; Type locality: Paris).

  • — Olivier, 1795:(42): 10 (Staphylinus; synonym of maxillosus).

  • — Turton, 1802: 509 (Staphylinus; characters; Germany).

  • — Gravenhorst, 1802: 3 (Staphylinus; synonym of maxillosus Linné).

  • — Erichson, 1839b: 348 (Staphylinus; synonym of maxillosus).

  • — P. Lucas, 1846: 108 (Staphylinus; synonym of nebulosus).

  • — Kraatz, 1857d: 529 (Creophilus; synonym of maxillosus).

  • — Ganglbauer, 1895: 415 (Creophilus; synonym of maxillosus).

  • — Bernhauer and Schubert, 1914: 398 (Creophilus; synonym of maxillosus).

  • — Cameron, 1932: 259 (Creophilus; synonym of maxillosus).

  • — Blackwelder, 1943: 447 (Creophilus; synonym of maxillosus).

  • — Smetana, 1958: 311 (Creophilus; synonym of maxillosus).

  • — Coiffait, 1974: 560 (Creophilus; synonym of maxillosus).

villosus Gravenhorst, 1802: 160 (Staphylinus; Type locality: America septentr.).

  • — Gravenhorst, 1806: 126 (Staphylinus).

  • — Nordmann, 1837: 21 (Creophilus; characters; North America).

  • — Kirby, 1837: 95 (Creophilus; characters; Nova Scotia; Canada).

  • — Erichson, 1839b: 349 (Staphylinus; characters; America septentrionali; Mexico; Cuba).

  • — Mannerheim, 1843b: 228 (Staphylinus; characters; California; Cuba; Mexico).

  • — LeConte, 1850: 220 (Xantholinus; Lake Superior).

  • — Haldeman, 1852: 375 (Staphylinus; Utah).

  • — Guérin-Méneville, 1857: 36 (Staphylinus; characters; Amérique septentrionale; Mexico; Cuba).

  • — Chevrolat and Fauvel, 1863: 432 (Creophilus; characters; Cuba, Mexico).

  • — Provancher, 1877: 245 (Creophilus; characters; Quebec).

  • — Horn, 1879: 200 (Creophilus; characters; Alaska southward over our entire territory, and extends probably into Mexico).

  • — Sharp, 1884: 370 (Creophilus; variety of maxillosus; Mexico; Guatemala; Cuba).

  • — Ganglbauer, 1895: 415 (Creophilus; variety of maxillosus).

  • — Fall and Cockerell, 1907: 166 (Creophilus; New Mexico).

  • — Blatchley, 1910: 374 (Creophilus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 399 (Creophilus; variety of maxillosus).

  • — Notman, 1920a: 180 (Creophilus; variety of maxillosus; New York).

  • — Notman, 1920b: 706 (Creophilus; variety of maxillosus; Florida).

  • — Mutchler, 1925: 235 (Creophilus; Galapagos Islands).

  • — Talbot, 1928 (Creophilus; anatomy of disgestive system; USA).

  • — Cameron, 1932: 259 (Creophilus; variety of maxillosus).

  • — Scheerpeltz, 1933: 1413 (Creophilus; variety of maxillosus).

  • — Abbott, 1934 (Creophilus; behavioral notes).

  • — Hatch, 1938: 148 (Creophilus; subspecies of maxillosus; notes; Alaska; Queen Charolotte Islands; Washington; Idaho; California; Utah; Kansas; Oklahoma; Louisiana; Iowa; Michigan; Ontario; New York).

  • — Voris, 1939a: 292 (Creophilus; variety of maxillosus; egg, larval, and pupal characters).

  • — Blackwelder, 1943: 447 (Creophilus; synonym of maxillosus).

  • — Smetana, 1965: 33 (Creophilus; subspecies of maxillosus; Miquelon; Labrador; Nova Scotia; Newfoundland).

  • — Downie and Arnett, 1996: 387 (Creophilus; subspecies of maxillosus; characters; USA).

  • — Ryabukhin, 1999: 68 (Creophilus; subspecies of maxillosus; catalog; habitat; distribution; northeastern Russia).

  • — Smetana and Davies, 2000: 38 (Creophilus; subspecies of maxillosus).

ciliaris Stephens, 1832: 202 (Creophilus; Type locality: Edinburgh).

  • — Kraatz, 1857d: 529 (Creophilus; synonym of maxillosus).

  • — G. Waterhouse, 1858: 23 (Creophilus; synonym of maxillosus).

  • — Fowler, 1888: 247 (Creophilus; variety of ciliaris; characters; Britain).

  • — Ganglbauer, 1895: 415 (Creophilus; variety of maxillosus).

  • — Cameron, 1932: 259 (Creophilus; variety of maxillosus).

  • — Scheerpeltz, 1933: 1413 (Creophilus; variety of maxillosus).

  • — Aleksandrov, 1934: 153 (Creophilus; subspecies of maxillosus; China).

  • — Blackwelder, 1943: 447 (Creophilus; synonym of maxillosus).

  • — Smetana, 1958: 311 (Creophilus; aberration of maxillosus).

  • — Coiffait, 1974: 561 (Creophilus; variety of maxillosus; characters).

  • — Pope, 1977: 30 (Creophilus; synonym of maxillosus).

  • — Naomi, 1982: 129 (Creophilus; cited as cilialis; variety of maxillosus).

fasciatus Laporte, 1835: 111 (Creophilus; [preoccupied]; Type locality: Amérique du nord).

  • — Chevrolat and Fauvel, 1863: 432 (Xantholinus; synonym of villosus).

  • — Horn, 1879: 200 (Creophilus; synonym of villosus).

  • — Bernhauer and Schubert, 1914: 399 (Creophilus; synonym of maxillosus).

arcticus Erichson, 1839b: 348 (Staphylinus; Type locality: Kamtschatka).

  • — Motschulsky, 1860b: 120 (Creophilus; eastern Siberia).

  • — Hochhuth, 1862: 47 (Creophilus; notes; Kamchatka).

  • — Solsky, 1870a: 260 (Creophilus; notes; characters; Mexico).

  • — Ganglbauer, 1895: 415 (Creophilus; synonym of maxillosus).

  • — Bernhauer and Schubert, 1914: 398 (Creophilus; variety of maxillosus).

  • — Bernhauer, 1926g: 2 (Creophilus; variety of maxillosus; Kamchatka region).

  • — Cameron, 1932: 259 (Creophilus; variety of maxillosus).

  • — Blackwelder, 1943: 447 (Creophilus; synonym of maxillosus).

  • — Ryabukhin, 1999: 68 (Creophilus; subspecies of maxillosus; catalog; habitat; distribution; northeastern Russia).

cinerarius Erichson, 1839b: 350 (Staphylinus; Type locality: Aegyptus).

  • — Bernhauer and Schubert, 1914: 399 (Creophilus; variety of maxillosus).

  • — Cameron, 1932: 259 (Creophilus; variety of maxillosus).

  • — Scheerpeltz, 1933: 1413 (Creophilus; variety of maxillosus).

  • — Blackwelder, 1943: 448 (Creophilus; synonym of maxillosus).

  • — Coiffait, 1974: 561 (Creophilus; variety of maxillosus; characters).

bicinctus Mannerheim, 1843b: 229 (Staphylinus; [preoccupied]; Type locality: insula Sitkha).

  • — Solsky, 1870a: 261 (Creophilus; synonym of arcticus).

  • — Horn, 1879: 200 (Creophilus; synonym of villosus).

  • — Ganglbauer, 1895: 415 (Creophilus; synonym of maxillosus).

  • — Bernhauer and Schubert, 1914: 398 (Creophilus; synonym of maxillosus).

  • — Cameron, 1932: 259 (Creophilus; synonym of maxillosus).

  • — Blackwelder, 1943: 448 (Creophilus; synonym of maxillosus).

orientalis Motschulsky, 1858d: 67 (Staphylinus; Type locality: Illes Kouriles).

  • — Motschulsky, 1859: 491 (Creophilus; Amur River).

  • — Motschulsky, 1860b: 120 (Creophilus; characters; notes; eastern Siberia).

  • — Ganglbauer, 1895: 415 (Creophilus; synonym of maxillosus).

  • — Bernhauer and Schubert, 1914: 398 (Creophilus; synonym of maxillosus).

  • — Cameron, 1932: 259 (Creophilus; synonym of maxillosus).

  • — Blackwelder, 1943: 448 (Creophilus; synonym of maxillosus).

fulvago Motschulsky, 1860b: 120 (Creophilus; Type locality: Mongolie et Chine boréale).

  • — Motschulsky, 1859: 491 (Creophilus; Amur River).

  • — Solsky, 1871b: 240 (Creophilus; notes; Russia).

  • — Ganglbauer, 1895: 415 (Creophilus; synonym of maxillosus).

  • — Blackwelder, 1943: 448 (Creophilus; synonym of maxillosus).

  • — Smetana, 1958: 311 (Creophilus; synonym of maxillosus).

imbecillus Sharp, 1874: 28 (Creophilus; variety of maxillosus; Type locality: Japan).

  • — Ganglbauer, 1895: 415 (Creophilus; synonym of maxillosus).

  • — Bernhauer and Schubert, 1914: 399 (Creophilus; synonym of maxillosus).

  • — Blackwelder, 1943: 448 (Creophilus; synonym of maxillosus).

  • — Smetana, 1958: 311 (Creophilus; synonym of maxillosus).

  • — Coiffait, 1974: 561 (Creophilus; synonym of maxillosus).

medialis Sharp, 1874: 28 (Creophilus; variety of maxillosus; Type locality: Japan).

  • — Ganglbauer, 1895: 415 (Creophilus; synonym of maxillosus).

  • — Bernhauer and Schubert, 1914: 399 (Creophilus; variety of maxillosus).

  • — Cameron, 1932: 259 (Creophilus; variety of maxillosus).

  • — Blackwelder, 1943: 448 (Creophilus; synonym of maxillosus).

  • — Nakane and Sawada, 1960: A124 (Staphylinus; variety of maxillosus; Yakushima Island).

  • — Watanabe and Shibata, 1972: 65 (Creophilus; variety of maxillosus; Yaku-shima Island).

  • — Naomi, 1982: 129 (Creophilus; variety of maxillosus).

subfasciatus Sharp, 1874: 28 (Creophilus; variety of maxillosus; Type locality: Japan).

  • — Ganglbauer, 1895: 415 (Creophilus; synonym of maxillosus).

  • — Bernhauer and Schubert, 1914: 399 (Creophilus; variety of maxillosus).

  • — Cameron, 1932: 259 (Creophilus; variety of maxillosus).

  • — Blackwelder, 1943: 448 (Creophilus; synonym of maxillosus).

  • — Naomi, 1982: 129 (Creophilus; variety of maxillosus).

pulchellus Meier, 1899: 99 (Creophilus; variety of maxillosus; Type locality: Hamburg).

  • — Bernhauer and Schubert, 1914: 399 (Creophilus; variety of maxillosus).

  • — Cameron, 1932: 259 (Creophilus; variety of maxillosus).

  • — Scheerpeltz, 1933: 1413 (Creophilus; variety of maxillosus).

  • — Blackwelder, 1943: 448 (Creophilus; synonym of maxillosus).

  • — Coiffait, 1974: 561 (Creophilus; variety of maxillosus; characters).

canariensis Bernhauer, 1908e: 334 (Creophilus; subspecies of maxillosus; Type locality: kanarische Inseln).

  • — Cameron, 1932: 259 (Creophilus; variety of maxillosus).

  • — Blackwelder, 1943: 448 (Creophilus; synonym of maxillosus).

  • — Coiffait, 1974: 560 (Creophilus; variety of maxillosus; characters).

  • — Hernández, Outerelo, and Gamarra, 1994: 187 (Creophilus; variety of maxillosus; checklist of species; Canary Islands).

ciliaroides Hatch, 1938: 149 (Creophilus; variety of maxillosus villosus; Type locality: Washington: Soap Lake).

  • — Blackwelder, 1943: 448 (Creophilus; synonym of maxillosus).

  • — Hatch, 1957: 213 (Creophilus; aberration of maxillosus; characters; British Columbia; Washington; Oregon; Idaho).

medialis Sharp, 1874, see: maxillosus Linné, 1758.

nebulosus Geoffroy, 1785, see: maxillosus Linné, 1758.

oculatus Fabricius, 1775: 265 (Staphylinus; Type locality: Nova Hollandia and Nova Zelandia).

  • — Goeze, 1777: 724 (Staphylinus; characters).

  • — Fabricius, 1781: 335 (Staphylinus; characters; New Guinea, New Zealand).

  • — Fabricius, 1787: 220 (Staphylinus; characters).

  • — Gmelin, 1790: 2026 (Staphylinus; characters).

  • — Fabricius, 1793: 521 (Staphylinus; characters).

  • — Olivier, 1795:(42): 11 (Staphylinus; characters; New Guinea).

  • — Fabricius, 1801: 592 (Staphylinus; characters; New Guinea).

  • — Turton, 1802: 510 (Staphylinus; characters; New Holland; New Zealand).

  • — Nordmann, 1837: 23 (Creophilus; characters; New Zealand).

  • — Erichson, 1839b: 352 (Staphylinus; characters; Nova Seelandia).

  • — White, 1846: 6 (Staphylinus; characters; New Zealand).

  • — Kiesenwetter and Kirsch, 1877: 154 (Staphylinus; Auckland Islands).

  • — Fauvel, 1877: 250 (Emus; characters; Australia; New Zealand; Auckland Islands).

  • — Broun, 1880: 107 (Staphylinus; characters; New Zealand).

  • — Bernhauer and Schubert, 1914: 399 (Creophilus; catalog).

  • — Lea, 1925: 229 (Creophilus; variety of erythrocephalus).

  • — Steel, 1949d: 59 (Creophilus; characters).

  • — Gourlay, 1950: 185 (Creophilus; list of subsequent references; collecting notes).

  • — Coiffait, 1976: 218 (Creophilus; Tahiti).

  • — Kuschel, 1990: 26 (Creophilus; introduced, Australian species; winged; habitat; New Zealand).

  • — Emberson, 1998: 35 (Creophilus; collecting notes; Chatham Islands).

  • Distribution: Chatham Islands, New Zealand, Australia, New Guinea, Society Islands.

huttoni Broun, 1880: 108 (Staphylinus; Type locality: Dunedin).

  • — Bernhauer and Schubert, 1914: 398 (Creophilus; catalog).

  • — Gourlay, 1950: 186 (Creophilus; synonym of oculatus; illustration).

orientalis Motschulsky, 1858, see: maxillosus Linné, 1758.

pulchellus Meier, 1899, see: maxillosus Linné, 1758.

sikkimensis Wendeler, 1927: 8 (Creophilus; Type locality: Sikkim).

  • — Cameron, 1932: 260 (Creophilus; characters; Sikkim).

  • — Scheerpeltz, 1933: 1413 (Creophilus; catalog).

  • — Smetana and Davies, 2000: 38 (Creophilus).

  • Distribution: India.

subfasciatus Sharp, 1874, see: maxillosus Linné, 1758.

tertius Schaeffer, 1766 [Note: Nomen nudum], see: maxillosus Linné, 1758.

unipunctatus Hope, 1831, see: erythrocephalus Fabricius, 1775.

variegatus Mannerheim, 1830: 20 (Creophilus; Type locality: Brasilia: Rio Janeiro).

  • — Mannerheim, 1831: 434 (Creophilus; Brazil).

  • — Nordmann, 1837: 21 (Creophilus; characters; Brasilia).

  • — Blanchard, 1842: 76 (Staphylinus; Uruguay; Brazil; Argentina).

  • — Erichson, 1839b: 351 (Staphylinus; characters; Brazil).

  • — Laporte, 1840: 173 (Creophilus; characters; Brazil).

  • — Fauvel, 1877a: xxvii (Emus; Brazil).

  • — Lynch, 1884: 129 (Creophilus; characters; Argentina).

  • — Bernhauer and Schubert, 1914: 399 (Creophilus; catalog).

  • — Scheerpeltz, 1933: 1413 (Creophilus; catalog).

  • Distribution: Brazil, Uruguay, Argentina, Paraguay.

villipennis Kraatz, 1859: 67 (Creophilus; Type locality: India borealis; India australis [Decan]).

  • — Bernhauer and Schubert, 1914: 399 (Creophilus; catalog).

  • — Cameron, 1932: 261 (Creophilus; characters; India; Sumatra; Nias).

  • — Scheerpeltz, 1933: 1413 (Creophilus; catalog).

  • — Cameron, 1933d: 392 (Creophilus; Sumatra).

  • — Cerruti, 1951: 17 (Creophilus; India).

  • — Scheerpeltz, 1976a: 92 (Creophilus; Nepal).

  • — Smetana and Davies, 2000: 38 (Creophilus).

  • Distribution: Nepal, India, Indonesia.

villosus Gravenhorst, 1802, see: maxillosus Linné, 1758.

violaceus Fauvel, 1878f: 248 (Emus; Type locality: Moluques, Bouro).

  • — Bernhauer and Schubert, 1914: 399 (Creophilus; catalog).

  • Distribution: Indonesia.

Diatrechus [41 species; Ethiopian and Madagascan Regions]

Diatrechus Bernhauer, 1911: 89 (species included: elatus, anthracinus, raffrayi, humeralis, aethiopicus, bicolor). Type species: Philonthus elatus Erichson, fixed by subsequent designation by Herman, 2001.

  • — Bernhauer and Schubert, 1914: 365 (world catalog; 12 species).

  • — Scheerpeltz, 1933: 1375 (world catalog supplement).

  • — Blackwelder, 1952: 123 (type species cited as Staphylinus compressicollis Klug).

  • — Scheerpeltz, 1970d: 83 (key to species).

  • — Herman, 2001: 9 (type species designation; discussion; earlier designation of Staphylinus compressicollis Klug by Blackwelder, 1952, erroneous).

abdominalis Schubert, 1911, see: bicolor Bernhauer, 1906.

aethiopicus Fauvel, 1907: 47 (Anisolinus; Type locality: Kilimandjaro, 1000–1300 m; Abyssinie; Ethiopie meridionale: Hieka).

  • — Bernhauer, 1911: 89 (Diatrechus).

  • — Bernhauer and Schubert, 1914: 365 (Diatrechus; catalog).

  • — Scheerpeltz, 1970d: 83 (Diatrechus; characters; Ethiopia; Uganda; Kenya).

  • Distribution: Ethiopia, Kenya, Tanzania, Uganda.

angolensis Cameron, 1959: 117 (Diatrechus; Type locality: Angola: Lunda; Dundo).

  • Distribution: Angola.

anthracinus Fauvel, 1905b: 181 (Anisolinus; Type locality: Diego-Suarez; Tananarive).

  • — Bernhauer, 1911: 89 (Diatrechus).

  • — Bernhauer and Schubert, 1914: 365 (Diatrechus; catalog).

  • — Jarrige, 1970: 52 (Diatrechus; Madagascar).

  • — Scheerpeltz, 1970d: 84 (Diatrechus; characters; Madagascar).

  • Distribution: Madagascar.

ater Bernhauer, 1915d: 147 (Diatrechus; Type locality: Abessinien: Harrar, Dire Daoua).

  • — Scheerpeltz, 1933: 1375 (Diatrechus; catalog).

  • — Scheerpeltz, 1970d: 85 (Diatrechus; characters; Ethiopia).

  • Distribution: Ethiopia.

bicolor Bernhauer, 1906a: 189 (Anisolinus; Type locality: Rhodesia: Mashonaland).

  • — Bernhauer, 1911: 89 (Diatrechus).

  • — Bernhauer and Schubert, 1914: 365 (Diatrechus; catalog).

  • — Scheerpeltz, 1970d: 89 (Diatrechus; characters; Rhodesia; Zambia; Tanzania; Kenya; Uganda; Ethiopia).

  • Distribution: Zambia, Zimbabwe, Tanzania, Uganda, Kenya, Ethiopia, Eritrea.

abdominalis Schubert, 1911: 28 (Anisolinus; Type locality: Takkesch, Eritrea).

  • — Bernhauer and Schubert, 1914: 365 (Diatrechus; synonym of bicolor).

  • — Scheerpeltz, 1970d: 89 (Diatrechus; synonym of bicolor).

brachypterus Lecoq, 1996: 151 (Diatrechus; Type locality: l'archipel des Comores: Anjouan, Hombo N'Thindi, 600 m).

  • Distribution: Comoros.

cameroni Tottenham, 1956: 310 (Diatrechus; replacement name for paederomimus Cameron).

  • — Herman, 2001: 24 (Diatrechus; synonymy).

  • Distribution: Rwanda, Congo, Tanzania.

paederomimus Cameron, 1933: 43 (Diatrechus; [preoccupied]; Type locality: Belgian Congo: Mahagi: Atshor).

  • — Tottenham, 1956: 310 (Diatrechus; synonym of cameroni).

nigrescens Tottenham, 1956: 310 (Diatrechus; variety of cameroni; Type locality: Ruanda: terr. Nyanza, Gitarama, 2850 m).

  • — Tottenham, 1962: 143 (Diatrechus; variety of cameroni; Tanganyika).

seminigrescens Tottenham, 1962: 254 (Diatrechus; variety of cameroni; [Note: Nomen nudum (Article 15.2)]; cited from Tanganyika Terr.: Bunduki, Uluguru Mts., moy. Mgeta, 1300 m).

cameroni Scheerpeltz, 1970d: 88 (Diatrechus; [preoccupied]; replacement name for paederomimus Cameron).

  • — Herman, 2001: 24 (Diatrechus; synonym of cameroni Tottenham).

cameroni Scheerpeltz, 1970, see: cameroni Tottenham, 1956.

cinctus Tottenham, 1956: 309 (Diatrechus; Type locality: Ruanda: Rutovu, forêt du Rugege, 2350 m).

  • Distribution: Rwanda.

coelestis Bernhauer, 1938d: 328 (Diatrechus; Type locality: Belgisches Kongo: Rutshuru).

  • — Scheerpeltz, 1970d: 88 (Diatrechus; characters; Congo region).

  • Distribution: Congo.

compressicollis Klug, 1834: 138 (Staphylinus; Type locality: Madagascar).

  • — Erichson, 1840: 508 (Philonthus; characters; Madagascar).

  • — Bernhauer and Schubert, 1914: 365 (Diatrechus; catalog).

  • — Scheerpeltz, 1970d: 87 (Diatrechus; characters; Madagascar).

  • Distribution: Madagascar.

congoensis Bernhauer, 1915k: 303 (Diatrechus; Type locality: Kongstaat).

  • — Scheerpeltz, 1933: 1375 (Diatrechus; cited as kongoensis; catalog).

  • — Tottenham, 1962: 144 (Diatrechus; Tanganyika).

  • — Scheerpeltz, 1970d: 85 (Diatrechus; cited as kongoensis; characters; Congo region).

  • Distribution: Congo, Tanzania.

cordicollis Gerstaecker, 1867: 28 (Philonthus; Type locality: Moschi).

  • — Fauvel, 1907: 48 (Anisolinus; Kibonoto; Abyssinie; Ethiopia meridionale; Usamabara; Zanzibar; Zambese).

  • — Bernhauer and Schubert, 1914: 365 (Diatrechus; catalog).

  • — Scheerpeltz, 1970d: 90 (Diatrechus; characters; Ethiopia; Uganda; Kenya; Tanzania; Rwanda; Burundi; Zambia; Rhodesia; Mozambique).

  • Distribution: Ethiopia, Rwanda, Burundi, Tanzania, Zambia, Zimbabwe, Mozambique.

ebeninus Tottenham, 1962: 254 (Diatrechus; Type locality: Kenya: Muguga, near Kikuyu. Also cited from Tanganyika).

  • Distribution: Kenya, Tanzania.

rubicundus Tottenham, 1962: 254 (Diatrechus; variety of ebeninus; [Note: Nomen nudum (Article 15.2)]; cited from Tanganyika Terr.: Mt. Meru, Olkokola, versant N.O., 2500–2600 m).

elatus Erichson, 1840: 508 (Philonthus; Type locality: Madagascar).

  • — Bernhauer, 1911: 89 (Diatrechus).

  • — Bernhauer and Schubert, 1914: 365 (Diatrechus; catalog).

  • — Scheerpeltz, 1961g: 239 (Diatrechus; Madagascar).

  • — Jarrige, 1970: 52 (Diatrechus; Madagascar).

  • — Scheerpeltz, 1970d: 87 (Diatrechus; characters; Madagascar).

  • — Herman, 2001: 9 (Diatrechus; type species of genus).

  • Distribution: Madagascar.

excavatus Tottenham, 1962: 143 (Diatrechus; without description; [Note: Nomen nudum, Article 13.1.1]; cited from Tanganyika).

fauveli Rambousek, 1922: 1011 (Diatrechus; Type locality: Ethiopie: Tchafianani).

  • — Scheerpeltz, 1933: 1375 (Diatrechus; catalog).

  • — Scheerpeltz, 1970d: 87 (Diatrechus; characters; Ethiopia).

  • Distribution: Ethiopia.

femoratus Schubert, 1911: 27 (Anisolinus; Type locality: Takkesch, Eritrea).

  • — Bernhauer and Schubert, 1914: 365 (Diatrechus; catalog).

  • — Tottenham, 1956: 229 (Diatrechus; Ruanda).

  • — Scheerpeltz, 1970d: 89 (Diatrechus; characters; Erythrea).

  • Distribution: Erithrea, Rwanda.

graueri Bernhauer, 1913, see: Tolmerinus.

griveaudi Jarrige, 1972a: 135 (Diatrechus; Type locality: Madagascar Centre, tampoketsa d'Ambohitantely, 1600 m, 16 km au Nord d'Ankazobe).

  • Distribution: Madagascar.

guttipennis Bernhauer, 1932b: 154 (Diatrechus; Type locality: Mulongo: Niunzu; Stanleyville, Pas-Uele).

  • — Scheerpeltz, 1933: 1375 (Diatrechus; catalog).

  • — Scheerpeltz, 1970d: 85 (Diatrechus; characters; Congo region).

  • Distribution: Congo.

haemorrhoidalis Brancsik, 1893: 220 (Philonthus; [preoccupied]; Type locality: Nossibé).

  • — Bernhauer and Schubert, 1914: 340 (Philonthus; noted to be preoccupied; catalog).

  • — Scheerpeltz, 1933: 1375 (Diatrechus; catalog).

  • — Herman, 2001: 43 (Diatrechus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: Madagascar.

humeralis Fauvel, 1907: 47 (Anisolinus; Type locality: Kilimandjaro, 1000–1300 m; Abyssinie [Bogos, 1500 m]; Senegal).

  • — Bernhauer, 1911: 89 (Diatrechus).

  • — Eichelbaum, 1913: 129 (Diatrechus; Tanzania).

  • — Bernhauer and Schubert, 1914: 365 (Diatrechus; catalog).

  • — Scheerpeltz, 1970d: 87 (Diatrechus; characters; Ethiopia; East and central Africa).

  • Distribution: Tanzania, Ethiopia, Senegal.

koenigsbaueri Scheerpeltz, 1970d: 81 (Diatrechus; Type locality: Tansania: Mokombe).

  • Distribution: Tanzania.

kristenseni Bernhauer, 1915d: 146 (Diatrechus; Type locality: Abessinien: Dire Daoua).

  • — Scheerpeltz, 1933: 1375 (Diatrechus; catalog).

  • — Scheerpeltz, 1970d: 86 (Diatrechus; characters; Ethiopia).

  • Distribution: Ethiopia.

lomaensis Levasseur, 1971: 298 (Diatrechus; Type locality: monts Loma, Sierra Leone, 1600 m).

  • Distribution: Sierra Leone.

maculatus Tottenham, 1962: 143 (Diatrechus; without characters; [Note: Nomen nudum, Article 13.1.1]; cited from Uganda).

methneri Bernhauer, 1937b: 614 (Diatrechus; Type locality: N. W. Uluguru).

  • — Scheerpeltz, 1970d: 89 (Diatrechus; characters; Uganda; Kenya; Tanzania; Zambia).

  • Distribution: Zambia, Tanzania, Uganda, Kenya.

natalensis Scheerpeltz, 1974: 165 (Diatrechus; Type locality: Natal: Durban, Stamford Hill).

  • Distribution: South Africa.

nigrescens Tottenham, 1956, see: cameroni Tottenham, 1956.

nigrolucens Cameron, 1959, see: Philonthus.

obscurus Cameron, 1933: 43 (Diatrechus; Type locality: Belgian Congo: Nizi: Blukwa).

  • — Scheerpeltz, 1970d: 86 (Diatrechus; characters; Congo region).

  • Distribution: Congo.

paederiformis Cameron, 1933: 43 (Diatrechus; Type locality: Belgian Congo: Mahagi: Atshor; Nizi: Kwamba).

  • — Scheerpeltz, 1970d: 86 (Diatrechus; characters; Congo region).

  • Distribution: Congo.

paederinus Bernhauer, 1937b: 615 (Diatrechus; Type locality: Utzungwe: Iringa).

  • — Scheerpeltz, 1970d: 88 (Diatrechus; characters; Uganda; Kenya; Rwanda; Burundi; Tanzania; Malawi; Zambia; Southern Rhodesia; Mozambique; South Africa).

  • Distribution: South Africa, Zimbabwe, Mozambique, Malawi, Zambia, Tanzania, Burundi, Rwanda, Uganda, Kenya.

paederoides Bernhauer, 1913b: 231 (Diatrechus; Type locality: NW. Tanganjika).

  • — Bernhauer and Schubert, 1914: 365 (Diatrechus; catalog).

  • — Bernhauer, 1947: 158 (Diatrechus; Deutsch-Ostafrika).

  • — Scheerpeltz, 1970d: 86 (Diatrechus; characters; Tanzania; Uganda).

  • Distribution: Tanzania, Uganda.

paederomimus Bernhauer, 1913: 231 (Diatrechus; Type locality: Abessinien in der Umgebung von Harrar).

  • — Bernhauer and Schubert, 1914: 365 (Diatrechus; catalog).

  • — Cameron, 1950g: 42 (Diatrechus; Belgian Congo).

  • — Scheerpeltz, 1970d: 88 (Diatrechus; characters; Ethiopia).

  • Distribution: Congo, Ethiopia.

paederomimus Cameron, 1933, see: cameroni Tottenham, 1956.

pauper Tottenham, 1962: 256 (Diatrechus; Type locality: Tanganyika Terr.: Mt. Oldeani, 2300 m. Also cited from South Africa).

  • Distribution: Tanzania, South Africa.

raffrayi Fauvel, 1905b: 180 (Anisolinus; Type locality: Madagascar).

  • — Bernhauer, 1911: 89 (Diatrechus).

  • — Bernhauer and Schubert, 1914: 365 (Diatrechus; catalog).

  • — Scheerpeltz, 1970d: 84 (Diatrechus; characters; Madagascar).

  • Distribution: Madagascar.

rapax Tottenham, 1956: 307 (Diatrechus; Type locality: Ruanda: terr. Nyanza, Gitarama, 1850 m).

  • Distribution: Rwanda.

rothschildi Rambousek, 1922: 1010 (Diatrechus; Type locality: Éthiopie: Kounhi, Tchafianani).

  • — Scheerpeltz, 1933: 1375 (Diatrechus; catalog).

  • — Paulian, 1947: 28 (Diatrechus; cited as rotschildi; Sierra Leone).

  • — Levasseur, 1968a: 52 (Diatrechus; cited as rotschildi; Central African Republic).

  • — Scheerpeltz, 1970d: 90 (Diatrechus; characters; Ethiopia).

  • Distribution: Ethiopia, Central African Republic, Sierra Leone.

rubicundus Tottenham [Note: Nomen nudum], 1962, see: ebeninus Tottenham, 1962.

semiflavus Tottenham, 1956: 307 (Diatrechus; Type locality: Ruanda: Tshuruyaga, forêt du Rugege, 2400 m).

  • Distribution: Rwanda.

seminigrescens Tottenham, 1962 [Note: Nomen nudum], see: cameroni Tottenham, 1956.

separatus Tottenham, 1956: 311 (Diatrechus; Type locality: Ruanda: terr. Nyanza, Shyogwe, 1700 m).

  • Distribution: Rwanda.

splendidus Bernhauer, 1937b: 615 (Diatrechus; Type locality: Utzungwe Berge, 1300–1600 m).

  • — Scheerpeltz, 1970d: 84 (Diatrechus; characters; Uganda; Kenya; Tanzania).

  • Distribution: Kenya, Tanzania, Uganda.

strandi Bernhauer, 1937b: 614 (Diatrechus; Type locality: Mons Gelei im ehemaligen Deutsch-Ostafrika).

  • — Scheerpeltz, 1970d: 84 (Diatrechus; characters; Kenya; Tanzania).

  • Distribution: Kenya, Tanzania.

togatus Tottenham, 1956: 310 (Diatrechus; Type locality: Urundi: Bururi, 2100 m).

  • Distribution: Burundi.

villiersi Cameron, 1949a: 321 (Diatrechus; Type locality: Guinée: Mt. Nimba, 700–900 m. Also cited from Ivory Coast).

  • — Cameron, 1952e: 832 (Diatrechus; Togo).

  • — Levasseur, 1971: 298 (Diatrechus; Cameroon; Sierra Leone).

  • Distribution: Guinea, Sierra Leone, Ivory Coast, Togo, Cameroon.

viridipennis Jarrige, 1978: 290 (Diatrechus; Type locality: Madagascar Est, sud-ouest de Befotaka, mont Papango).

  • Distribution: Madagascar.

Dinothenarus [24 species; Nearctic, Palaearctic, and Oriental Regions]

Dinothenarus Thomson, 1858: 29 (species included: pubescens). Type species: Staphylinus pubescens De Geer, fixed by monotypy.

  • — Heyden, Reitter, and Weise, 1891: 101 (cited as Trichoderma; subgenus of Staphylinus; list of species of Europe and Caucasus).

  • — Heyden, Reitter, and Weise, 1906: 169 (cited as Trichoderma; subgenus of Staphylinus; list of species of Europe and Caucasus).

  • — Reitter, 1909: 118 (synonym of Staphylinus subgenus Trichoderma).

  • — Bernhauer and Schubert, 1914: 374 (synonym of Trichoderma).

  • — Cameron, 1932: 180 (subgenus of Staphylinus).

  • — Blackwelder, 1943: 443 (subgenus of Staphylinus; type species: pubescens).

  • — Tottenham, 1949: 374, 420 (type species: pubescens De Geer; subgenus of Staphylinus; checklist of British species).

  • — Blackwelder, 1952: 128 (type species: pubescens).

  • — Coiffait, 1956e: 194 (synonym of Ontholestes, cited as Dinotheranus).

  • — Smetana, 1958: 260, 263 (subgenus of Staphylinus; characters).

  • — Horion, 1965: 205 (synonym of Staphylinus; subgenus Trichoderma).

  • — Pototskaia, 1967: 67 (cited as Trichoderma; subgenus Staphylinus; [Note: These characters probably refer to Dinothenarus since the species on which they are based is in Dinothenarus]; larval characters).

  • — Tikhomirova, 1973a: 187 (subgenus of Staphylinus; checklist of species of USSR).

  • — Coiffait, 1974: 359 (adult and larval characters; key to species of western Palaearctic region).

  • — Frank, 1979a (some larval characters and their use in classification).

  • — Muona, 1979: 15 (list of Scandinavian species).

  • — M. Dvořák, 1979: 116 (some species collected in Slovakia).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 96 (catalog; Poland).

  • — Uhlig and Vogler, 1981: 119 (cited as Trichoderma; list of some species collected in Germany; habitat and collecting notes).

  • — Tóth, 1984: 17 (characters; habitus; key to species of Hungary).

  • — Drugmand 1987b (comments on distribution of Belgian species).

  • — Smetana, 1992: 187 (notes; key to species).

  • — Silfverberg, 1992: 17 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Ciceroni and Zanetti, 1995: 30 (list of species of Italy).

  • — M. Hansen, 1996: 107 (list of Danish species).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 244 (additions to list of Danish species).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (list of species collected in 1997; Denmark).

  • — Smetana and Davies, 2000: 20, 26, 41 (characters; checklist of species of north temperate region).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 393 (6 Nearctic species; notes; characters in key).

Parabemus Reitter, 1909: 118 (subgenus of Staphylinus; species included: fossor, fodiens). Type species: Staphylinus fossor Scopoli, fixed by subsequent designation by Blackwelder, 1943: 443 but cited erroneously as “monobasic”.

  • — Bernhauer and Schubert, 1914: 374 (synonym of Abemus).

  • — Scheerpeltz, 1933: 1381 (synonym of Staphylinus).

  • — Chapman, 1934 (subgenus of Staphylinus; notes on classification and distribution of some species).

  • — Blackwelder, 1943: 443 (subgenus of Staphylinus; type species: fossor).

  • — Blackwelder, 1952: 288 (type species).

  • — Coiffait, 1956e: 187, 183, 195 (valid genus; characters in key; notes).

  • — Hatch, 1957: 176 (subgenus of Ocypus; characters; key to species of Pacific Northwest).

  • — Smetana, 1958: 260 (synonym of Staphylinus).

  • — Lohse, 1964: 195 (characters; central European species).

  • — Scheerpeltz, 1966a: 116 (subgenus of Staphylinus; characters).

  • — Tikhomirova, 1973a: 187 (subgenus of Staphylinus; checklist of species of USSR).

  • — Coiffait, 1974: 364 (characters; key to species of western Palaearctic region).

  • — Frank, 1979a (some larval characters and their use in classification).

  • — Uhlig, 1979: 254 (some species collected in Germany).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 96 (catalog; Poland).

  • — Uhlig and Vogler, 1981: 119 (list of some species collected in Germany; habitat and collecting notes).

  • — Tóth, 1984: 18 (characters; habitus; species of Hungary).

  • — Drugmand 1987b (comments on distribution of Belgian species).

  • — Lohse and Lucht, 1989: 169 (valid genus).

  • — Ciceroni and Zanetti, 1995: 30 (list of species of Italy).

  • — Smetana and Davies, 2000: 20, 26, 41 (subgenus of Dinothenarus; characters; checklist of species of north temperate region).

Parocypus Bernhauer, 1915b: 52 (subgenus of Staphylinus; species included: dehradunensis). Type species: Staphylinus (Parocypus) dehradunensis Bernhauer, fixed by monotypy.

  • — Cameron, 1932: 180 (subgenus of Staphylinus).

  • — Scheerpeltz, 1933: 1381 (subgenus of Staphylinus).

  • — Blackwelder, 1943: 444, 445 (subgenus of Ocypus; type species: dehradunensis).

  • — Blackwelder, 1952: 295 (subgenus of Ocypus; type species: dehradunensis).

  • — Shibata, 1984: 88 (subgenus of Ocypus; checklist of species of Japan).

  • — Smetana and Davies, 2000: 26, 41 (synonym of Dinothenarus subgenus Parabemus).

Hypabemus Scheerpeltz, 1966a: 112, 116 (subgenus of Staphylinus; species included: chrysocomus, arrosus). Type species: Staphylinus chrysocomus Mannerheim, fixed by original designation.

  • — Smetana and Davies, 2000: 26, 41 (synonym of Dinothenarus subgenus Parabemus).

Protabemus Scheerpeltz, 1966a: 114, 115 (subgenus of Staphylinus; species included: xanthocephalus). Type species: Staphylinus xanthocephalus Kraatz, fixed by original designation and monotypy.

  • — Smetana, 1992: 187 (synonym of Dinothenarus).

  • — Smetana and Davies, 2000: 26, 41 (synonym of Dinothenarus subgenus Dinothenarus).

adonis Coiffait, 1956, see: flavocephalus Goeze, 1777.

arrosus Eppelsheim, 1890: 169 (Staphylinus; Type locality: westlicher Kaukasus).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • — Smetana, 1965c: 26 (Staphylinus; subgenus Parabemus; Anatolia).

  • — Smetana, 1965d: 79 (Staphylinus; subgenus Parabemus; Anatolia).

  • — Scheerpeltz, 1966a: 117 (Staphylinus; subgenus Hypabemus; characters; Caucasus).

  • — Smetana, 1968b: 87 (Staphylinus; subgenus Parabemus; Anatolia).

  • — Coiffait, 1974: 367 (Parabemus; characters; Caucasus; Anatolia).

  • — Solodovnikov, 1998a: 13 (Staphylinus; notes; Georgia; Russia).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; subgenus Parabemus).

  • Distribution: Caucasus, Georgia, Turkey.

ornativentris Reitter, 1911a: 49 (Ocypus; Type locality: Abchasien auf dem Kopschara, 6500′).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; synonym of arrosus).

  • — Smetana, 1965c: 26 (Staphylinus; synonym of arrosus).

  • — Smetana, 1968b: 87 (Staphylinus; synonym of arrosus).

  • — Coiffait, 1974: 367 (Parabemus; synonym of arrosus).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; synonym of arrosus).

aureiventris G. Müller, 1926 [Note: Nomen nudum], see: dehradunensis Bernhauer, 1915.

baderlei Scheerpeltz, 1966, see: fossor Scopoli, 1771.

badipes LeConte, 1863a: 36 (Staphylinus; Type locality: Middle and western States as far north as Pembina, Minnesota).

  • — Provancher, 1877: 247 (Staphylinus; characters; Quebec).

  • — Horn, 1879: 188 (Staphylinus; characters; New England States westward through Canada, New York and Michigan).

  • — Blatchley, 1910: 376 (Staphylinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • — Notman, 1920: 16 (Staphylinus; New York).

  • — Notman, 1920a: 180 (Staphylinus; New York).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • — Smetana, 1965: 32 (Staphylinus; subgenus Abemus; Nova Scotia).

  • — Morgan and Morgan, 1985: 1829 (Staphylinus; middle Holocene fossil; Michigan).

  • — Dearborn and Donahue, 1994: 15 (Staphylinus; Maine).

  • — Downie and Arnett, 1996: 385 (Staphylinus; characters; USA).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; subgenus Parabemus).

  • Distribution: Canada, USA.

carbonatus LeConte, 1863a: 36 (Staphylinus; Type locality: Western States).

  • — Horn, 1879: 188 (Staphylinus; synonym of badipes).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; synonym of badipes).

bodemeyeri Bernhauer, 1913, see: chrysocomus Mannerheim, 1830.

bomfordi Eppelsheim, 1895, see: xanthocephalus Kraatz, 1859.

capitatus J. Bland, 1864: 65 (Staphylinus; Type locality: Canada West).

  • — Provancher, 1877: 247 (Staphylinus; characters; Quebec).

  • — Horn, 1879: 198 (Leistotrophus; characters; Ontario; New York).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; catalog).

  • — Smetana, 1959d: 411 (Dinothenarus; notes).

  • — Smetana, 1981d: 126 (Dinothenarus; generic placement confirmed).

  • — Downie and Arnett, 1996: 387 (Ontholestes; characters; Canada; USA).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; subgenus Dinothenarus).

  • Distribution: Canada, USA.

carbonatus LeConte, 1863, see: badipes LeConte, 1863.

choui Smetana, 1992: 193 (Dinothenarus; Type locality: Taiwan: Chiai Hsien, Alishan, 2200 m).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; subgenus Dinothenarus).

  • Distribution: Taiwan.

chrysocephalus Geoffroy, 1785, see: flavocephalus Goeze, 1777.

chrysocomus Mannerheim, 1830: 21 (Staphylinus; Type locality: Dauria).

  • — Mannerheim, 1831: 435 (Staphylinus; Dauria).

  • — Nordmann, 1837: 41 (Staphylinus; characters; Dauria).

  • — Erichson, 1839b: 394 (Staphylinus; characters; Dauria; Siberia).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; catalog).

  • — Aleksandrov, 1934: 153 (Staphylinus; China).

  • — Scheerpeltz, 1966a: 116 (Staphylinus; subgenus Hypabemus; type species of chrysocomus; characters; east and northeast Asia).

  • — Coiffait, 1974: 367 (Parabemus; characters; Siberia).

  • — Smetana and Davies, 2000: 26, 41 (Dinothenarus; subgenus Parabemus).

  • Distribution: Russia, China.

chrysopterus Gebler, 1832: 41 (Staphylinus; Type locality: Sibérie orientale: District des Mines de Nertchinsk).

  • — Solsky, 1871c: 347 (Staphylinus; synonym of chrysocomus).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; synonym of chrysocomus).

  • — Coiffait, 1974: 367 (Parabemus; synonym of chrysocomus).

fulviceps Motschulsky, 1858b: 17 (Staphylinus; Type locality: Not cited).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; synonym of chrysocomus).

  • — Coiffait, 1974: 367 (Parabemus; synonym of chrysocomus).

erythrocephalus Motschulsky, 1859: 491 (Staphylinus; a catalog listing without a description; [Note: Nomen nudum (Article 12.1)]).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; synonym of chrysocomus).

  • — Coiffait, 1974: 367 (Parabemus; synonym of chrysocomus).

xanthocephalus Motschulsky, 1860b: 120 (Staphylinus; [preoccupied]; Type locality: les bords de l'Amour méridional dans les montagnes Boureïa).

  • — Solsky, 1871c: 348 (Staphylinus; notes; Siberia).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; synonym of chrysocomus).

  • — Coiffait, 1974: 367 (Parabemus; synonym of chrysocomus).

bodemeyeri Bernhauer, 1913a: 223 (Staphylinus; subgenus Abemus; Type locality: Ostsibirien: Chitaizki-Sterena).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; synonym of chrysocomus).

  • — Coiffait, 1974: 367 (Parabemus; synonym of chrysocomus).

chrysopterus Gebler, 1832, see: chrysocomus Mannerheim, 1830.

coiffaiti Bordoni, 1977: 329 (Dinothenarus; Type locality: N-Afganistan: Bala Murghab, Herat, mt. 470).

  • — Smetana, 1992: 188 (Parabemus; transferred from Dinothenarus).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; subgenus Parabemus).

  • Distribution: Afghanistan.

dehradunensis Bernhauer, 1915b: 52 (Staphylinus; subgenus Parocypus; Type locality: Zentral-Himalaya: Dehra Dun).

  • — Cameron, 1932: 203 (Staphylinus; subgenus Parocypus; characters; India).

  • — Scheerpeltz, 1933: 1393 (Staphylinus; catalog).

  • — Blackwelder, 1952: 295 (type species of Parocypus, a subgenus of Ocypus).

  • — Smetana and Davies, 2000: 26, 41 (Dinothenarus; subgenus Parabemus).

  • Distribution: India.

aureiventris G. Müller, 1926b: 42 (Staphylinus; subgenus Parocypus; [Note: Nomen nudum, published without a description (Article 12.1)]).

  • — Scheerpeltz, 1933: 1393 (Staphylinus; synonym of dehradunensis).

erythrocephalus Motschulsky, 1859 [Note: Nomen nudum], see: chrysocomus Mannerheim, 1830.

flavocephalus Goeze, 1777: 730 (Staphylinus; Type locality: Not cited).

  • — Rossi, 1792: 97 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 378 cited pubescens Rossi as a synonym of flavocephalus; Rossi did not describe the species as new, he attributed it to De Geer]; misidentified as pubescens; characters).

  • — Ganglbauer, 1895: 425 (Staphylinus; subgenus Trichoderma; characters; France; Mediterranean region).

  • — Reitter, 1909: 118 (Staphylinus; subgenus Trichoderma; characters; Germany).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; catalog).

  • — Coiffait, 1956e: 194 (Trichoderma; characters; France).

  • — Smetana, 1958: 264 (Staphylinus; subgenus Dinothenarus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 195 (Trichoderma; characters; central Europe).

  • — Scheerpeltz, 1965b: 403 (Staphylinus; Sicily).

  • — Horion, 1965: 205 (Staphylinus; subgenus Trichoderma; Lebanon; Palestine; Rhodes; Greece; Dalmatia; Bosnia Hercegovina; Bulgaria; Italy; France; Serbia; Slovakia; Austria; Germany ?).

  • — Scheerpeltz, 1966a: 114 (Staphylinus; subgenus Trichoderma; type species of Trichoderma; characters; southern Europe; Mediterranean region; Turkey; Mesopotamia; Armenia).

  • — Jarrige, 1971: 496 (Trichoderma; Iran).

  • — Coiffait, 1974: 361 (Dinothenarus; characters; Europe).

  • — Petrenko, 1980a: 62 (Staphylinus; subgenus Dinothenarus; characters).

  • — Tóth, 1984: 18 (Dinothenarus; characters).

  • — Lucht, 1987: 107 (Trichoderma; checklist; Czechoslovakia; Austria; France).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Dinothenarus; Slovakia).

  • — Ciceroni and Zanetti, 1995: 30 (Dinothenarus; Italy).

  • — Pilon, 1998: 72 (Dinothenarus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; subgenus Dinothenarus).

  • Distribution: Czechoslovakia, Germany, France, Italy, Greece, Croatia, Armenia, Turkey, Syria, Israel, Iraq, Iran.

chrysocephalus Geoffroy, 1785: 165 (Staphylinus; Type locality: Paris).

  • — Villers, 1789: 421 (Staphylinus; characters).

  • — Latreille, 1804: 296 (Staphylinus; characters).

  • — Gravenhorst, 1806: 124 (Staphylinus; characters; France).

  • — Mannerheim, 1830: 21 (Emus; Dalmatia).

  • — Mannerheim, 1831: 435 (Emus; Dalmatia).

  • — Brullé, 1832: 129 (Emus; characters; Greece).

  • — Lacordaire, 1835: 362 (Emus; characters; France).

  • — Nordmann, 1837: 31 (Staphylinus; characters; France).

  • — Erichson, 1839b: 371 (Staphylinus; characters; France; Italy; Dalmatia).

  • — Küster, 1853: 45 (Staphylinus; characters; France; Italy; Dalmatia).

  • — Fairmaire and Laboulbène, 1856: 506 (Staphylinus; characters; France).

  • — Kraatz, 1858a: 60 (Staphylinus; Greece).

  • — Peyron, 1858: 422 (Staphylinus; Turkey).

  • — Baudi, 1870: 384 (Staphylinus; Turkey).

  • — Fauvel, 1874: 176 [= 1874a: 402] (Staphylinus; characters; France; Dalmatia; Istria; Italy; Sicily; Sardinia; Greece; Syria; Turkey; Iran).

  • — Mulsant and Rey, 1876: 235 [= 1877: 91] (Trichoderma; characters; France).

  • — Ganglbauer, 1895: 425 (Staphylinus; synonym of flavocephalus).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; synonym of flavocephalus).

  • — Portevin, 1929: 348 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; synonym of flavocephalus).

  • — Smetana, 1958: 264 (Staphylinus; synonym of flavocephalus).

  • — Coiffait, 1974: 362 (Dinothenarus; synonym of flavocephalus).

adonis Coiffait, 1956e: 194 (Trichoderma; subspecies of flavocephalus; Type locality: Liban Jerusalem).

  • — Scheerpeltz, 1966a: 115 (Staphylinus; subgenus Trichoderma; subspecies of flavocephalus; characters; Palestine; Syria).

  • — Coiffait, 1974: 361 (Dinothenarus; subspecies of flavocephalus; characters; Syria; Asia Minor; Persia).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; subgenus Dinothenarus; subspecies of flavocephalus).

fodiens Gravenhorst, 1806, see: fossor Scopoli, 1771.

fokiensis Bernhauer, 1933, see: insignis G. Müller, 1926.

fossor Scopoli, 1771: 109 (Staphylinus; Type locality: Not cited).

  • — Goeze, 1777: 725 (Staphylinus; characters).

  • — Gmelin, 1790: 2032 (Staphylinus; page misnumbered as 1032; characters).

  • — Turton, 1802: 517 (Staphylinus; characters; Germany).

  • — Erichson, 1839b: 377 (Staphylinus; characters; Europe).

  • — Redtenbacher, 1849: 695 (Staphylinus; characters; Austria).

  • — Küster, 1853: 47 (Staphylinus; characters; Europe; Carolina).

  • — Fairmaire and Laboulbène, 1856: 507 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 549 (Staphylinus; characters; Germany).

  • — Redtenbacher, 1857: 184 (Staphylinus; characters; Austria).

  • — Fauvel, 1874: 180 [= 1874a: 406] (Staphylinus; characters; France; Switzerland; Germany; Austria; Italy).

  • — Redtenbacher, 1874: 204 (Staphylinus; characters; Austria).

  • — Seidlitz, 1875: 269 (Leistotrophus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 246 [= 1877: 102] (Abemus; characters; France).

  • — Ganglbauer, 1895: 426 (Staphylinus; subgenus Abemus; characters; middle Europe).

  • — Reitter, 1909: 118 (Staphylinus; subgenus Parabemus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; catalog).

  • — Johansen, 1914: 381 (Staphylinus; subgenus Abemus; characters; Denmark).

  • — G. Müller, 1926: 6 (Staphylinus; subgenus Parabemus; notes).

  • — Portevin, 1929: 349 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; catalog).

  • — Coiffait, 1956e: 195 (Parabemus; characters).

  • — Smetana, 1958: 267 (Staphylinus; subgenus Abemus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 195 (Parabemus; characters; central Europe).

  • — Horion, 1965: 207 (Staphylinus; subgenus Parabemus; Holland; Belgium; Switzerland; France; Spain; Italy; Bosnia Hercegovina; Hungary; Czechoslovakia; Poland; Austria; Germany).

  • — Scheerpeltz, 1966a: 116 (Staphylinus; subgenus Parabemus; characters; Europe; Mediterranean region).

  • — Schwaller, 1973 (Parabemus; larval characters).

  • — Coiffait, 1974: 366 (Parabemus; characters; Europe).

  • — Petrenko, 1980a: 59, 62 (Staphylinus; subgenus Abemus; characters; Ukraine).

  • — Tóth, 1984: 19 (Parabemus; characters; Hungary).

  • — Segers, 1986: 47 (Parabemus; checklist of species; Belgium).

  • — Lucht, 1987: 107 (Parabemus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).

  • — Huth and Dettner, 1989: 154 (Parabemus; defensive secretions).

  • — Huth and Dettner, 1990 (Parabemus; defensive chemicals).

  • — Pilon and Zanetti, 1991: 65 (Parabemus; Italy).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Parabemus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 30 (Parabemus; Italy).

  • — Chabrol, 1998: 38 (Parabemus; notes; France).

  • — Pilon, 1998: 73 (Parabemus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 26, 41 (Dinothenarus; subgenus Parabemus).

  • Distribution: Europe.

fossor Fabricius, 1793: 523 (Staphylinus; [preoccupied]; [Note: Erichson, 1839b: 377, cited the Fabricius, 1793: 523, as referring to Scopoli's fossor along with the other references that are listed herein for fossor Fabricius]; Type locality: Germania).

  • — Panzer, 1795: 353 (Staphylinus; characters; Germany).

  • — Fabricius, 1801: 593 (Staphylinus; characters).

  • — Gravenhorst, 1802: 10 (Staphylinus; characters).

  • — Latreille, 1804: 299 (Staphylinus; characters; Germany).

  • — Gravenhorst, 1806: 117 (Staphylinus; characters).

  • — Panzer, 1808: 6 (Staphylinus; characters; illustrations; Germany).

  • — Lacordaire, 1835: 367 (Emus; characters; France).

  • — Nordmann, 1837: 60 (Staphylinus; characters; Germany; France; Italy).

  • — Heer, 1839: 252 (Staphylinus; characters; Switzerland).

  • — Laporte, 1840: 176 (Staphylinus; characters; France).

  • — Heer, 1841: 582 (Staphylinus; Switzerland).

  • — Smetana and Davies, 2000: 27, 41 (Dinothenarus; synonym of fossor).

germanus Turton, 1802: 511 (Staphylinus; replacement name for fossor Fabricius).

fodiens Gravenhorst, 1806: 116 (Staphylinus; [Note: Because D. fossor has not been reported from North America, the synonymy of D. fodiens with D. fossor may be erroneous]; Type locality: Carolina).

  • — Nordmann, 1837: 62 (Staphylinus; United States).

  • — Erichson, 1839b: 377 (Staphylinus; synonym of fossor Scopoli).

  • — Küster, 1853: 47 (Staphylinus; synonym of fossor).

  • — Kraatz, 1857d: 549 (Staphylinus; synonym of fossor).

  • — Ganglbauer, 1895: 426 (Staphylinus; synonym of fossor).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; synonym of fossor).

  • — Reitter, 1909: 118 (Staphylinus; subgenus Parabemus; synonym of fossor).

  • — Smetana, 1958: 267 (Staphylinus; synonym of fossor).

  • — Coiffait, 1974: 366 (Parabemus; cited as fodieus; synonym of fossor).

baderlei Scheerpeltz, 1966a: 106, 116 (Staphylinus; subgenus Parabemus; Type locality: Col di Tenda in den Ligurischen Alpen, 200 m).

  • — Pilon, 1998: 99 (Parabemus; synonym of fossor).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; synonym of fossor).

fossor Fabricius, 1793, see: fossor Scopoli, 1771.

fulviceps Motschulsky, 1858, see: chrysocomus Mannerheim, 1830.

ganglbauerianus Bernhauer, 1939d: 101 (Staphylinus; subgenus Xanthocypus; Type locality: Japan: Kamikochi).

  • — Hayashi, 1995: 52 (Ocypus; characters; review of type).

  • — Smetana and Davies, 2000: 27, 41 (Dinothenarus; subgenus Parabemus; notes; type may be mislabelled and species may not be from Japan).

  • Distribution: Japan.

glasunovi Luze, 1904, see: sibiricus Gebler, 1829.

insignis G. Müller, 1926b: 41 (Staphylinus; subgenus Parocypus; Type locality: Japan; [Note: See lectotype designation by Smetana and Davies, 2000: 28]. Also cited from Focehow - Atchong nella Cina meridionale).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 27, 28, 41 (Dinothenarus; subgenus Parabemus; notes; lectotype designation: Japan; type series may have been mislabelled and species may not be from Japan).

  • Distribution: China, Japan.

fokiensis Bernhauer, 1933: 32 (Staphylinus; subgenus Parocypus; Type locality: Fokien: Foo-chow, Ostchina).

  • — Smetana and Davies, 2000: 27, 41 (Dinothenarus; subgenus Parabemus; notes; synonym of insignis).

kiautschauensis Bernhauer, 1916e: 31 (Staphylinus; subgenus Parocypus; Type locality: Kiautschau).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; subgenus Parabemus).

  • Distribution: China.

krejcii Coiffait, 1984e: 378 (Parocypus; Type locality: vallée de Kathmandou, Gokarna).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: Nepal.

lacustris Bernhauer, 1917, see: pleuralis LeConte, 1861.

luteipes LeConte, 1861: 342 (Staphylinus; Type locality: San Jose, California).

  • — Horn, 1879: 190 (Staphylinus; characters; California).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; subgenus Parabemus).

  • Distribution: USA.

mediorientalis Bordoni, 1984, see: pubescens De Geer, 1774.

nigrellus Horn, 1879: 188 (Staphylinus; Type locality: Oregon and northern California).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • — Hatch, 1957: 176 (Ocypus; subgenus Abemus; characters; British Columbia; Idaho; Washington; Oregon).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; subgenus Parabemus).

  • Distribution: Canada, USA.

ochrocephalus Gmelin, 1790, see: pubescens De Geer, 1774.

ornativentris Reitter, 1911, see: arrosus Eppelsheim, 1890.

pleuralis LeConte, 1861: 343 (Staphylinus; Type locality: Oregon).

  • — Horn, 1879: 189 (Staphylinus; characters; Utah, Oregon, Vancouver).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • — Hatch, 1957: 177 (Ocypus; subgenus Parabemus; characters; British Columbia; Idaho; Washington; Oregon).

  • — Smetana, 1965a: 10 (Staphylinus; subgenus Abemus; British Columbia).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; subgenus Parabemus).

  • Distribution: Canada, USA.

lacustris Bernhauer, 1917a: 2 (Staphylinus; subgenus Staphylinus; Type locality: Britisch-Columbia: Emld. Lake).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • — Hatch, 1957: 177 (Ocypus; characters; British Columbia; Washington; Oregon).

  • — Smetana, 1965a: 11 (Staphylinus; synonym of pleuralis).

prainae Eppelsheim, 1895c: 58 (Trichocosmetes; Type locality: Sikkim im Himalaya; Sikkim, Darjeeling).

  • — Bernhauer and Schubert, 1914: 394 (Trichocosmetes; catalog).

  • — Cameron, 1928c: 565 (Staphylinus; Sikkim).

  • — Cameron, 1932: 201 (Staphylinus; subgenus Goerius; characters; India).

  • — Scheerpeltz, 1933: 1400 (Staphylinus; catalog).

  • — Scheerpeltz, 1965: 109 (Staphylinus; subgenus Goerius; Burma).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; subgenus Parabemus).

  • Distribution: India, Myanmar.

pubescens De Geer, 1774: 17 (Staphylinus; Type locality: Not cited).

  • — Goeze, 1777: 731 (Staphylinus; characters).

  • — Paykull, 1789: 16 (Staphylinus; redescription; Sweden, France).

  • — Gmelin, 1790: 2034 (Staphylinus; characters).

  • — Fabricius, 1793: 520 (Staphylinus; characters).

  • — Olivier, 1795:(42): 16 (Staphylinus; characters).

  • — Panzer, 1795: 352 (Staphylinus; characters; Austria).

  • — Cederhjelm, 1798: 117 (Staphylinus; characters).

  • — Paykull, 1800: 376 (Staphylinus; characters; Sweden).

  • — Fabricius, 1801: 590 (Staphylinus; characters; Sweden).

  • — Turton, 1802: 509, 518 (Staphylinus; characters; Sweden; Europe).

  • — Gravenhorst, 1802: 12 (Staphylinus; characters).

  • — Marsham, 1802: 500 (Staphylinus; characters; Britain).

  • — Latreille, 1804: 295 (Staphylinus; characters).

  • — Gravenhorst, 1806: 123 (Staphylinus; characters).

  • — Gyllenhal, 1810: 284 (Staphylinus; characters; Sweden).

  • — Zetterstedt, 1828: 67 (Staphylinus; characters; Lapland).

  • — Curtis, 1829: 24 (Staphylinus; catalog; Britain).

  • — C. Sahlberg, 1830: 309 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 21 (Emus; Finland; Siberia).

  • — Mannerheim, 1831: 435 (Emus; Finland; Siberia).

  • — Stephens, 1832: 205 (Staphylinus; characters; England).

  • — Runde, 1835: 2 (Emus; characters; Germany).

  • — Lacordaire, 1835: 361 (Emus; characters; France).

  • — Nordmann, 1837: 31 (Staphylinus; Europe; Siberia).

  • — Zetterstedt, 1838: 59 (Staphylinus; characters; habitat; Lapland).

  • — Curtis, 1839: pl. 758 (Staphylinus; characters; England).

  • — Erichson, 1839a: 434 (Staphylinus; characters; Germany).

  • — Heer, 1839: 250 (Staphylinus; characters; Switzerland).

  • — Erichson, 1839b: 372 (Staphylinus; characters; Europe).

  • — Stephens, 1839: 386 (Trichoderma; characters).

  • — Jacobsen, 1849: 682 (Staphylinus; characters; Denmark).

  • — Redtenbacher, 1849: 695 (Staphylinus; characters; Austria).

  • — Küster, 1853: 46 (Staphylinus; characters; Europe).

  • — Hardy, 1851: 22 (Staphylinus; subgenus Trichoderma; notes; England).

  • — Fairmaire and Laboulbène, 1856: 506 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 546 (Staphylinus; characters; Germany).

  • — Redtenbacher, 1857: 184 (Staphylinus; characters; Austria).

  • — Thomson, 1858: 29 (Dinothenarus; type species).

  • — Thomson, 1859: 23 (type species of Trichoderma).

  • — Thomson, 1860: 142 (Trichoderma; characters; Scandinavia).

  • — Fauvel, 1874: 176 [= 1874a: 402] (Staphylinus; characters; France; Europe).

  • — Redtenbacher, 1874: 204 (Staphylinus; characters; Austria).

  • — Seidlitz, 1875: 269 (Leistotrophus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 238 [= 1877: 94] (Trichoderma; characters; larval characters; France).

  • — J. Sahlberg, 1876: 9 (Trichoderma; Finland).

  • — Fowler, 1888: 250 (Staphylinus; characters; Britain).

  • — Ganglbauer, 1895: 425 (Staphylinus; subgenus Dinothenarus; characters; Europe).

  • — Reitter, 1909: 118 (Staphylinus; subgenus Trichoderma; characters; Germany).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • — Johansen, 1914: 378 (Staphylinus; subgenus Trichoderma; characters; Denmark).

  • — Šulc, 1922 (Staphylinus; osmeterium; anatomy; histology; physiology).

  • — Portevin, 1929: 348 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; catalog).

  • — Paulian, 1941: 238 (Staphylinus; subgenus Trichoderma; larval characters).

  • — V. Hansen, 1952: 97 (Staphylinus; characters; Denmark).

  • — Coiffait, 1956e: 194 (Trichoderma; characters).

  • — Smetana, 1958: 264 (Staphylinus; subgenus Dinothenarus; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 109 (Staphylinus; subgenus Trichoderma; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 195 (Trichoderma; characters; central Europe).

  • — Horion, 1965: 206 (Staphylinus; subgenus Trichoderma; widespread in northern Europe, in all central Europe; France; Italy; Serbia; Dalmatia; Croatia; Bosnia Hercegovina; Ukraine; Urals).

  • — Scheerpeltz, 1966a: 115 (Staphylinus; subgenus Trichoderma; characters; Europe).

  • — Pototskaia, 1967: 67 (Staphylinus; subgenus Trichoderma; larval characters).

  • — Smetana, 1967i: 556 (Staphylinus; subgenus Dinothenarus; Turkey).

  • — Szujecki, 1968a: 732 (Staphylinidae; Poland).

  • — Coiffait, 1974: 363 (Dinothenarus; characters; Europe; Britain; Scandinavia).

  • — Shilov, 1975a: 376 [= 1975b: 99] (Staphylinus; Komi region of Russia).

  • — Pope, 1977: 30 (Platydracus; Britain).

  • — Petrenko, 1980a: 59, 62 (Staphylinus; subgenus Dinothenarus; characters; Ukraine).

  • — Tóth, 1984: 18 (Dinothenarus; characters; Hungary).

  • — Segers, 1986: 46 (Dinothenarus; checklist of species; Belgium).

  • — Lucht, 1987: 107 (Trichoderma; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Pilon and Zanetti, 1991: 65 (Dinothenarus; Italy).

  • — Drugmand, 1993: 170 (Dinothenarus; notes; Luxembourg).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Dinothenarus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 30 (Dinothenarus; Italy).

  • — Pilon, 1998: 73 (Dinothenarus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; subgenus Dinothenarus).

  • Distribution: Europe, Russia, Turkey, Lebanon.

ochrocephalus Gmelin, 1790: 2034 (Staphylinus; Type locality: Gallia).

  • — Rossi, 1792: 97 (Staphylinus; synonym of pubescens De Geer).

  • — Turton, 1802: 518 (Staphylinus; characters; France).

mediorientalis Bordoni, 1984: 339 (Dinothenarus; subspecies of pubescens; Type locality: Libano: Les Cèdres, Biblos, 2200 m).

ruficeps Cameron, 1932: 186 (Staphylinus; subgenus Trichoderma; Type locality: Simla).

  • — Scheerpeltz, 1933: 1388 (Staphylinus; catalog).

  • — Scheerpeltz, 1976a: 89 (Staphylinus; subgenus Trichoderma; Nepal).

  • — Smetana, 1992: 190 (Dinothenarus; characters; illustrations; habitat; India; Nepal).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; subgenus Dinothenarus).

  • Distribution: India, Nepal.

sagaris Smetana, 1992: 192 (Dinothenarus; Type locality: Burma: “N. Burma: Adung Valley, 5000 ft”).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; subgenus Dinothenarus).

  • Distribution: Myanmar.

saphyrinus LeConte, 1861: 342 (Staphylinus; Type locality: San Jose and Fort Tejon, California).

  • — Horn, 1879: 189 (Staphylinus; characters; California).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • — Hatch, 1957: 177 (Ocypus; subgenus Parabemus; characters; British Columbia; Washington; Oregon).

  • — Smetana and Davies, 2000: 42 (Dinothenarus; subgenus Parabemus).

  • Distribution: Canada, USA.

sibiricus Gebler, 1829: 71 (Staphylinus; Type locality: Barnaul).

  • — Erichson, 1839b: 363 (Staphylinus; characters; Sibiria).

  • — Solsky, 1871: 158 (Leistotrophus; characters; of male; Russia).

  • — Solsky, 1874: 172 (Leistotrophus; characters; Turkestan).

  • — J. Sahlberg, 1880: 68 (Trichoderma; Siberia).

  • — Eppelsheim, 1892: 332 (Staphylinus; Uzbekistan).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • — Scheerpeltz, 1963: 13 (Staphylinus; subgenus Trichoderma; Afghanistan).

  • — Scheerpeltz, 1966a: 115 (Staphylinus; subgenus Trichoderma; characters; Siberia; Transcaspian region; Turkestan; Iran; Afghanistan).

  • — Coiffait, 1974: 364 (Dinothenarus; characters; central Asia; Siberia).

  • — Boháč, 1987: 89 (Staphylinus; subgenus Dinothenarus; larval and egg characters; collecting and bionomic notes; Tajikistan).

  • — Smetana, 1992: 188 (Parabemus; transferred from Dinothenarus).

  • — Smetana and Davies, 2000: 42 (Dinothenarus; subgenus Parabemus).

  • Distribution: Russia, Tajikistan, Uzbekistan, Kazakhstan, Afghanistan, Iran.

glasunovi Luze, 1904: 92 (Staphylinus; subgenus Trichoderma; Type locality: Seravschan: Putschin-Pass, Kschtut: Artutsch).

  • — Luze, 1911b: 129 (Staphylinus; synonym of sibiricus).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; synonym of sibiricus).

  • — Coiffait, 1974: 364 (Dinothenarus; synonym of sibiricus).

  • — Tronquet, 1981: 71 (Dinothenarus; Afghanistan).

szechuanensis Bernhauer, 1935: 6 (Staphylinus; subgenus Abemus; Type locality: Mittel-Szechuan: Ginyün Sse, China, 4500 m).

  • — Smetana and Davies, 2000: 42 (Dinothenarus; subgenus Parabemus).

  • Distribution: China.

xanthocephalus Kraatz, 1859: 74 (Staphylinus; Type locality: India borealis).

  • — Fauvel, 1904a: 57 (Staphylinus; Kashmir; Sikkim).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Cameron, 1932: 185 (Staphylinus; subgenus Trichoderma; characters; India).

  • — Scheerpeltz, 1933: 1389 (Staphylinus; catalog).

  • — Paulian, 1941: 239 (Staphylinus; subgenus Trichoderma; larval characters).

  • — Cerruti, 1951: 16 (Staphylinus; subgenus Trichoderma; India).

  • — Scheerpeltz, 1966a: 115 (Staphylinus; subgenus Protabemus; type species of xanthocephalus; characters; Kashmir; Himalayan region).

  • — Scheerpeltz, 1976a: 90 (Staphylinus; subgenus Protabemus; Nepal).

  • — Coiffait, 1982: 31 (Trichoderma; Nepal).

  • — Coiffait, 1984e: 374 (Trichoderma; Nepal).

  • — Smetana, 1992: 194 (Dinothenarus; characters; illustrations; habitat; India; Nepal; Pakistan).

  • — Smetana and Davies, 2000: 41 (Dinothenarus; subgenus Dinothenarus).

  • Distribution: Pakistan, India, Nepal.

bomfordi Eppelsheim, 1895c: 56 (Staphylinus; subgenus Trichoderma; Type locality: Simla im Himalaya).

  • — Fauvel, 1904a: 57 (Staphylinus; synonym of xanthocephalus).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; synonym of xanthocephalus).

  • — Cameron, 1932: 185 (Staphylinus; synonym of xanthocephalus).

  • — Smetana, 1992: 194 (Dinothenarus; synonym of xanthocephalus).

xanthocephalus Motschulsky, 1860, see: chrysocomus Mannerheim, 1830.

Emus [4 species (3 doubtful); Palaearctic Region]

Emus Leach, 1819: 172 (species included: hirtus). Type species: Staphylinus hirtus Linné, fixed by original designation and monotypy.

  • — Curtis, 1829: 24 (catalog).

  • — Stephens, 1829a: 22 (catalog; Britain).

  • — Stephens, 1829: 274 (catalog; Britain).

  • — Curtis, 1835: pl. 534 (characters; type species).

  • — Mannerheim, 1830: 7, 20 (characters).

  • — Mannerheim, 1831: 421, 434 (characters).

  • — Stephens, 1832: 200, 203 (characters).

  • — Dejean, 1833: 59 (in list of species).

  • — Lacordaire, 1835: 358 (characters).

  • — Percheron, 1835: liv. 3, no. 1, pl. 5 (characters).

  • — Westwood, 1838a: 15 (characters; type species).

  • — Erichson, 1839b: 346 (synonym of Staphylinus).

  • — Stephens, 1839: 384 (characters).

  • — Laporte, 1840: 174 (characters).

  • — Lacordaire, 1854: 76 (synonym of Staphylinus).

  • — Jacquelin du Val, 1856: 34 (characters).

  • — Kraatz, 1857d: 530 (characters).

  • — Redtenbacher, 1857: 984 (characters).

  • — Thomson, 1858: 29 (characters).

  • — G. Waterhouse, 1858: 23 (catalog of British species).

  • — Thomson, 1859: 23 (characters; type species: hirtus).

  • — Schaum, 1859: 26 (catalog; European species).

  • — Thomson, 1860: 139 (characters).

  • — Gredler, 1863: 101 (infrageneric group of Staphylinus; in list of species from Tirol).

  • — Wencker and Silbermann, 1866: 28 (in list of species; collecting notes; France).

  • — Gemminger and Harold, 1868: 575 (catalog).

  • — Jacquelin du Val, 1868: 66 (group of Staphylinus; catalog; European species).

  • — Sharp, 1871d: 11 (in list of British species).

  • — Fauvel, 1874: 168 [= 1874a: 394] (characters).

  • — Redtenbacher, 1874: 203 (characters).

  • — Fauvel, 1875a: XXVI [= 1875b: 228] (catalog).

  • — Mulsant and Rey, 1876: 215 [= 1877: 71] (characters; key to French species).

  • — Duvivier, 1883: 142 (catalog).

  • — Sharp, 1888: 248 (characters).

  • — Heyden, Reitter, and Weise, 1891: 101 (in list of species of Europe and Caucasus).

  • — Ganglbauer, 1895: 416 (characters).

  • — Heyden, Reitter, and Weise, 1906: 172 (in list of species of Europe and Caucasus).

  • — Reitter, 1909: 116 (characters).

  • — Petri, 1912: 76 (in list of species of Siebenbürgen).

  • — Bernhauer and Schubert, 1914: 391 (world catalog; 3 species).

  • — Johansen, 1914: 373 (characters).

  • — Winkler, 1925: 386 (catalog for Palaearctic region).

  • — Porta, 1926: 93 (characters; species of Italy).

  • — Portevin, 1929: 347 (characters; key to French genera).

  • — Roubal, 1930: 384 (catalog; Slovakia).

  • — Joy, 1932: 115 (key to British species).

  • — Scheerpeltz, 1933: 1404 (world catalog supplement).

  • — Kloet and Hincks, 1945: 167 (in list of British species).

  • — Tottenham, 1949: 375, 421 (type species: hirtus Linné; notes; in checklist of British species).

  • — Blackwelder, 1952: 147 (type species: hirtus).

  • — V. Hansen, 1952: 110 (characters; Danish species).

  • — Coiffait, 1956e: 184, 186, 190 (characters in key; notes; key to species).

  • — Smetana, 1958: 303 (characters; type species: hirtus; Czechoslovakian species).

  • — Székessy, 1963: 58 (characters).

  • — Palm, 1963b: 121 (characters; species of Sweden).

  • — Lohse, 1964: 193 (characters; central European species).

  • — Pototskaia, 1967: 64 (larval characters).

  • — Scheerpeltz, 1968a: 85 (catalog of Austrian species).

  • — Kasule, 1970: 65 (larval characters).

  • — Tikhomirova, 1973a: 187 (in checklist of species of USSR).

  • — Coiffait, 1974: 383 (characters; species of western Palaearctic region).

  • — Pope, 1977: 30 (in list of British species).

  • — Topp, 1978: 311 (larval characters).

  • — Frank, 1979a (some larval characters and their use in classification).

  • — Muona, 1979: 15 (in list of Scandinavian species).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 91 (catalog; Poland).

  • — Tóth, 1984: 10 (characters; habitus; species of Hungary).

  • — Drugmand 1987b (comments on distribution of Belgian species).

  • — Silfverberg, 1992: 17 (in list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Smetana, 1993b: 49 (in list of species of Czech Republic and Slovakia).

  • — Ciceroni and Zanetti, 1995: 30 (in list of species of Italy).

  • — M. Hansen, 1996: 107 (in list of Danish species).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 244 (additions to in list of Danish species).

  • — Assing, Frisch, Kahlen, et al., 1998: 138 (note).

  • — Smetana and Davies, 2000: 18, 22, 38 (characters; in checklist of species of north temperate region).

Emys Agassiz, 1847: 137 (emendation of Emus). Type species: Staphylinus hirtus Linné, fixed by objective synonymy with Emus.

  • — Blackwelder, 1952: 148 (type species).

Creatophilus Gistel, 1856: 267, 388 (species included: hirtus). Type species: Staphylinus hirtus Linné, fixed by monotypy.

  • — Blackwelder, 1952: 110 (synonym of Emus; type species: hirtus).

aeneicollis Lacordaire, 1835: 374 (Emus; [Note: Nomen dubium]; Type locality: Paris).

  • Distribution: France.

albertisi Fauvel, 1879, see: Creophilus.

angustatus Lacordaire, 1835, see: Tasgius.

bombilius De Geer, 1774, see: hirtus Linné, 1758.

carinthiacus Lacordaire, 1835, see: Platydracus.

figulus Gistel, 1857: 69 (Emus; [Note: Nomen dubium]; Type locality: Chamouny).

  • Distribution: France.

floralis Lacordaire, 1835, see: Quedius.

fuscomaculatus Laporte, 1835, see: Staphylinus.

griseosericans Fairmaire, 1894, see: Rhynchocheilus.

hirtus Linné, 1758: 421 (Staphylinus; Type locality: Not cited).

  • — Linné, 1760: 230 (Staphylinus; characters; Sweden).

  • — Linné, 1767: 683 (Staphylinus; characters).

  • — Füessly, 1775: 21 (Staphylinus).

  • — Fabricius, 1775: 264 (Staphylinus; characters).

  • — O. Müller, 1776: 97 (Staphylinus; characters).

  • — Goeze, 1777: 714 (Staphylinus; characters).

  • — Schrank, 1781: 229 (Staphylinus; characters; Austria).

  • — Fabricius, 1781: 334 (Staphylinus; characters; Europe).

  • — Harrer, 1784: 249 (Staphylinus; characters).

  • — Herbst, 1784: 149 (Staphylinus; Germany).

  • — Fourcroy, 1785: 165 (Staphylinus; characters; France).

  • — Fabricius, 1787: 219 (Staphylinus; characters).

  • — Paykull, 1789: 3 (Staphylinus; redescription; notes; Sweden).

  • — Villers, 1789: 409 (Staphylinus; characters).

  • — Gmelin, 1790: 2025 (Staphylinus; characters; Europe).

  • — Rossi, 1792: 95 (Staphylinus; characters).

  • — Fabricius, 1793: 519 (Staphylinus; characters; Europe).

  • — Panzer, 1795: 351 (Staphylinus; characters; Germany).

  • — Panzer, 1796: 19 (Staphylinus; characters; illustration).

  • — Olivier, 1795:(42): 7 (Staphylinus; characters).

  • — Schrank, 1798: 639 (Staphylinus; characters; notes).

  • — Paykull, 1800: 368 (Staphylinus; characters; Sweden).

  • — Fabricius, 1801: 589 (Staphylinus; characters).

  • — Turton, 1802: 509 (Staphylinus; characters; Europe).

  • — Gravenhorst, 1802: 159 (Staphylinus; characters).

  • — Marsham, 1802: 496 (Staphylinus; characters; Britain).

  • — Latreille, 1804: 292 (Staphylinus; characters).

  • — Gravenhorst, 1806: 125 (Staphylinus; characters).

  • — Latreille, 1806: 286 (Staphylinus; characters).

  • — Gyllenhal, 1810: 280 (Staphylinus; characters; Sweden).

  • — Leach, 1819: 172 (type species of Emus).

  • — Curtis, 1829: 24 (Emus; catalog; Britain).

  • — Latreille, 1829: 434 (Staphylinus; characters; France, Germany).

  • — Mannerheim, 1830: 20 (Emus; Sweden; France).

  • — Mannerheim, 1831: 434 (Emus; Sweden; France).

  • — Stephens, 1832: 203 (Emus; characters; England).

  • — Griffith and Pidgeon, 1832: 291 (Staphylinus; characters; France; Germany).

  • — Ménétriés, 1832: 142 (Staphylinus; Caucasus-Persia region).

  • — Curtis, 1835: pl. 534 (Emus; characters; type species of Emus; notes; habitat; England).

  • — Stephens, 1835: 435 (Emus; notes).

  • — Runde, 1835: 2 (Emus; characters; Germany).

  • — Lacordaire, 1835: 360 (Emus; characters; France).

  • — Nordmann, 1837: 38 (Staphylinus; Europe).

  • — Westwood, 1838a: 15 (Creophilus; type species).

  • — O. Costa, 1839b: 118 (Staphylinus; characters; Italy).

  • — Erichson, 1839a: 431 (Staphylinus; characters; Germany).

  • — Heer, 1839: 248 (Staphylinus; characters; Switzerland).

  • — Erichson, 1839b: 346 (Staphylinus; characters; Europe).

  • — Stephens, 1839: 386 (Emus; characters).

  • — Laporte, 1840: 174 (Emus; characters; France).

  • — Westwood, 1845: 45 (Emus).

  • — Kolenati, 1846: 16 (Staphylinus; Caucasus; Transcaucasus).

  • — Jacobsen, 1849: 881 (Staphylinus; characters; Denmark).

  • — Ménétriés, 1849: 43 (Staphylinus; Bachkirie).

  • — Hochhuth, 1849: 112 (Staphylinus; characters; Caucasus).

  • — Redtenbacher, 1849: 694 (Staphylinus; characters; Austria).

  • — Küster, 1853: 41 (Staphylinus; characters; Europe).

  • — Fairmaire and Laboulbène, 1856: 505 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 531 (Emus; characters; Germany).

  • — Redtenbacher, 1857: 184 (Staphylinus; characters; Austria).

  • — Thomson, 1859: 23 (type species of Emus).

  • — Thomson, 1860: 140 (Emus; characters; Scandinavia).

  • — Fauvel, 1874: 169 [= 1874a: 395] (Emus; characters; France; Caucasus; Turkey).

  • — Redtenbacher, 1874: 203 (Emus; characters; Austria).

  • — Seidlitz, 1875: 268 (Emus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 217 [= 1877: 217] (Emus; characters; France).

  • — Fowler, 1888: 248 (Emus; characters; Britain).

  • — Seidlitz, 1891: 395 (Emus; characters; Europe).

  • — Seidlitz, 1891a: 420 (Emus; characters; Europe).

  • — Ganglbauer, 1895: 416 (Emus; characters; Europe; eastern Mediterranean region).

  • — Reitter, 1909: 117 (Emus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 391 (Emus; catalog).

  • — Johansen, 1914: 374 (Emus; characters; Denmark).

  • — Portevin, 1929: 347 (Emus; characters; France).

  • — Scheerpeltz, 1933: 1405 (Emus; catalog).

  • — V. Hansen, 1952: 110 (Emus; characters; Denmark).

  • — Coiffait, 1956e: 191 (Emus; characters; France).

  • — Smetana, 1958: 304 (Emus; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 121 (Emus; characters; Sweden; Denmark).

  • — Lohse, 1964: 193 (Emus; characters; central Europe).

  • — M. Dvořák, 1965: 89 (Emus; Slovakia).

  • — Horion, 1965: 236 (Emus; notes; discussion; Caucasus; Portugal; Spain; Italy; Dalmatia; Slovenia; Croatia; Bosnia Hercegovina; Serbia; Romania; Hungary; Slovakia; Czech Republic; Poland; France; Germany; Austria; Switzerland; Belgium; Holland; Britain; Denmark; Sweden).

  • — Pototskaia, 1967: 64 (Emus; larval characters).

  • — Szujecki, 1968a: 733 (Emus; Poland).

  • — Bolov, 1969a: 328 (Emus; collecting notes; Russia).

  • — Jarrige, 1971: 497 (Emus; Iran).

  • — Coiffait, 1974: 385 (Emus; characters; Europe; Britain; Scandinavia; Mediterranean region; Asia Minor; Caucasus).

  • — Pope, 1977: 30 (Emus; Britain).

  • — Allen, 1980: 50 (Emus; collecting notes; England).

  • — Frank, 1982: 35 (Emus; list of parasites).

  • — Tóth, 1984: 11 (Emus; characters; Hungary).

  • — Segers, 1986: 47 (Emus; checklist of species; Belgium).

  • — Ádám, 1987: 148 (Emus; collecting notes; Hungary).

  • — Lucht, 1987: 107 (Emus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Pilon and Zanetti, 1991: 66 (Emus; Italy).

  • — Smetana, 1993b: 49 (Emus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 30 (Emus; Italy).

  • — Gerend and Braunert, 1997: 199 (Emus; notes; Luxembourg).

  • — Sterrenburg and Schülke, 1997: 20 (Emus; Netherlands).

  • — Hammond, 1997: 14 (Emus; comments on collecting and natural history; Britain).

  • — Kočárek, 1997 (Emus; collecting notes; Czech Republic).

  • — Allen, 1998: 144 (Emus; notes; Isle of Sheppey).

  • — Pilon, 1998: 68 (Emus; ecological notes; Italy).

  • — Kashcheev, 1999a: 58 (Emus; Kazakhstan).

  • — Smetana and Davies, 2000: 38 (Emus).

  • — Allen, 2000a: 212 (Emus; notes on occurence in Britain).

  • Distribution: Europe, Caucasus, Turkey, Iran, Kazakhstan.

quintus Schaeffer, 1766: xxxvi, 6 (Staphylinus; [Note: Nomen nudum, Article 11.4]; Type locality: Ratisbon).

  • — Erichson, 1839b: 346 (Staphylinus; synonym of hirtus).

  • — Kraatz, 1857d: 531 (Emus; synonym of hirtus).

bombilius De Geer, 1774: 20 (Staphylinus; Type locality: Hollande and Allemagne).

  • — Fabricius, 1781: 334 (Staphylinus; cited as bombylius; synonym of hirtus).

  • — Gmelin, 1790: 2025 (Staphylinus; cited as bombylius; synonym of hirtus).

  • — Panzer, 1796: 19 (Staphylinus; synonym of hirtus Linné).

  • — Olivier, 1795:(42): 8 (Staphylinus; synonym of hirtus).

  • — Marsham, 1802: 496 (Staphylinus; synonym of hirtus Linné).

  • — Gyllenhal, 1810: 281 (Staphylinus; synonym of hirtus Linné).

  • — Erichson, 1839b: 346 (Staphylinus; synonym of hirtus).

  • — Fairmaire and Laboulbène, 1856: 505 (Staphylinus; synonym of hirtus).

  • — Kraatz, 1857d: 531 (Emus; synonym of hirtus).

  • — Ganglbauer, 1895: 416 (Emus; synonym of hirtus).

  • — Bernhauer and Schubert, 1914: 391 (Emus; cited as bombylius; synonym of hirtus).

  • — Smetana, 1958: 304 (Emus; cited as bombylius; synonym of hirtus).

  • — Coiffait, 1974: 385 (Emus; cited as bombylius; synonym of hirtus).

inauratus Mannerheim, 1830, see: Ontholestes.

insularis Fauvel, 1879, see: Creophilus.

marginalis Gené, 1836, see: Ontholestes.

nebulosus Percheron, 1835, see: Staphylinus.

nigripes Lacordaire, 1835, see: Tasgius.

occultus Lacordaire, 1835, see: Quedius.

picipennis Lacordaire, 1835, see: Ocypus.

quintus Schaeffer, 1766 [Note: Nomen nudum], see: hirtus Linné, 1758.

ruficornis Ménétriés, 1832, see: Ontholestes.

rufipalpis Lacordaire, 1835, see: Tasgius.

soropegus Gistel, 1857: 67 (Emus; [Note: Nomen dubium]; Type locality: lacum Thun. Cant. Bern. Septbr.).

  • Distribution: Switzerland.

speciosus Mannerheim, 1830, see: Ontholestes.

vicinus Lacordaire, 1835, see: Quedius.

violaceus Fauvel, 1878, see: Creophilus.

Eucibdelus [33 species; Palaearctic and Oriental Regions]

Eucibdelus Kraatz, 1859: 70 (species included: gracilis). Type species: Eucibdelus gracilis Kraatz, fixed by monotypy.

  • — Gemminger and Harold, 1868: 577 (catalog).

  • — Duvivier, 1883: 142 (catalog).

  • — Bernhauer and Schubert, 1914: 393 (world catalog; 5 species).

  • — Winkler, 1925: 386 (catalog for Palaearctic region).

  • — Cameron, 1925: 68 (catalog of Indian species).

  • — Cameron, 1932: 217 (characters; key to species of British India).

  • — Scheerpeltz, 1933: 1407 (world catalog supplement).

  • — Bernhauer, 1939k: 594 (key to Palaearctic species).

  • — Blackwelder, 1952: 154 (type species: gracilis).

  • — R. Dvořák, 1956: 29 (key to Japanese species).

  • — Shibata, 1973c: 62 (checklist of species of Taiwan).

  • — Tikhomirova, 1973a: 188 (checklist of species of USSR).

  • — Scheerpeltz, 1976a: 142 (key to species of Himalayan region).

  • — Shibata, 1984: 100 (checklist of species of Japan).

  • — Hayashi, 1998b: 103 (Eucibdelus; notes; key to subgenera).

Pareucibdelus Hayashi, 1997: 29 (subgenus of Eucibdelus; species included: japonicus). Type species: Eucibdelus japonicus Sharp, fixed by original designation and monotypy.

  • — Hayashi, 1998b: 104 (subgenus of Eucibdelus; characters).

Nudeucibdelus Hayashi, 1997: 30 (subgenus of Eucibdelus; species included: ishigakiensis). Type species: Eucibdelus (Nudeucibdelus) ishigakiensis Hayashi, fixed by original designation and monotypy.

  • — Hayashi, 1998b: 104 (synonym of Eucibdelus).

Neocibdelus Hayashi, 1998b: 104 (subgenus of Eucibdelus; proposed for misidentification of Eucibdelus Kraatz by Hayashi, 1997; species included: orientalis). Type species: Eucibdelus (Neocibdelus) orientalis Hayashi, fixed by original designation and monotypy.

  • — Hayashi, 1997: 27 (generic characters; [Note: Misidentified as Eucibdelus according to 1998b]; key to subgenera; members of and characters for Eucibdelus group; genera included: Philetaerius, Trichocosmetes, Eucibdelus, Phytolinus, Rhychocheilus).

angusticeps Bernhauer, 1920d: 186 (Eucibdelus; Type locality: Himalaya, Sikkim: Darjeeling).

  • — Cameron, 1932: 219 (Eucibdelus; characters; India).

  • — Scheerpeltz, 1933: 1407 (Eucibdelus; catalog).

  • — Scheerpeltz, 1976a: 144 (Eucibdelus; characters).

  • Distribution: India.

aokii R. Dvořák, 1956, see: japonicus Sharp, 1874.

argentipennis Bernhauer, 1933a: 50 (Eucibdelus; Type locality: Tatsienlu-Kiulung).

  • Distribution: China.

bhutanicus Coiffait, 1977b: 237 (Eucibdelus; Type locality: Bhutan: 87 km de Phuntsholing).

  • Distribution: Bhutan.

bicolor Fauvel, 1895: 248 (Eucibdelus; Type locality: Bírmanie, Carin Cheba, 900–1100 m).

  • — Bernhauer and Schubert, 1914: 393 (Eucibdelus; catalog).

  • — Cameron, 1932: 222 (Eucibdelus; characters; Burma).

  • — Scheerpeltz, 1933: 1407 (Eucibdelus; catalog).

  • Distribution: Myanmar.

birmanus Cameron, 1932: 223 (Eucibdelus; Type locality: Burma: Ruby Mines).

  • — Scheerpeltz, 1933: 1407 (Eucibdelus; catalog).

  • — Scheerpeltz, 1965: 109 (Eucibdelus; Burma).

  • Distribution: Myanmar.

cariniceps Scheerpeltz, 1976a: 137 (Eucibdelus; Type locality: Nepal: Jumbesi, 2750 m).

  • Distribution: Nepal.

chapmani Bernhauer, 1933a: 49 (Eucibdelus; Type locality: Yunnan-Sen).

  • Distribution: China.

chinensis Bernhauer, 1933a: 50 (Eucibdelus; Type locality: Tatsienlu-Kiulung).

  • Distribution: China.

chujoi R. Dvořák, 1956, see: japonicus Sharp, 1874.

elegans Coiffait, 1977b: 238 (Eucibdelus; Type locality: Bhutan: Phuntsholing-Thimphu, 2300 m).

  • Distribution: Bhutan.

feae Fauvel, 1895: 247 (Eucibdelus; Type locality: Bírmanie, Carin Cheba, 900–1100 m).

  • — Bernhauer and Schubert, 1914: 393 (Eucibdelus; catalog).

  • — Cameron, 1932: 220 (Eucibdelus; characters; Burma).

  • — Scheerpeltz, 1933: 1407 (Eucibdelus; catalog).

  • — Scheerpeltz, 1965: 109 (Eucibdelus; Burma).

  • Distribution: Myanmar.

freyi Bernhauer, 1939k: 595 (Eucibdelus; Type locality: N. W. China: Tienmuschan).

  • Distribution: China.

gracilis Kraatz, 1859: 71 (Eucibdelus; Type locality: India borealis).

  • — Bernhauer and Schubert, 1914: 394 (Eucibdelus; catalog).

  • — Cameron, 1932: 219 (Eucibdelus; characters; India).

  • — Scheerpeltz, 1933: 1407 (Eucibdelus; catalog).

  • — Scheerpeltz, 1976a: 144 (Eucibdelus; characters).

  • Distribution: India.

gratus Cameron, 1932: 221 (Eucibdelus; Type locality: Sylhet Dist.: Chandkhira).

  • — Scheerpeltz, 1933: 1407 (Eucibdelus; catalog).

  • — Hayashi, 1998a: 40 (Eucibdelus; subgenus Pareucibdelus; characters).

  • Distribution: India.

hoebarthi Bernhauer, 1933a: 48 (Eucibdelus; Type locality: Himalaja, Kulu).

  • — Scheerpeltz, 1976a: 144 (Eucibdelus; characters).

  • Distribution: India.

ishigakiensis Hayashi, 1997: 30 (Eucibdelus; subgenus Nudeucibdelus; Type locality: Ryûkyû Ils: Is. Ishigakijima, Mt. Omoto).

  • Distribution: Japan.

japonicus Sharp, 1874: 29 (Eucibdelus; Type locality: Hiogo; Nagasaki).

  • — Bernhauer and Schubert, 1914: 394 (Eucibdelus; catalog).

  • — Cameron, 1949b: 469 (Eucibdelus; Japan).

  • — Nakane, 1963a: 93 (Eucibdelus; characters; habitus photograph; Japan).

  • — Yuh, Paik, Kwon, and Lee, 1985: 247 (Eucibdelus; Korea).

  • — Hayashi, 1997: 29 (Eucibdelus; subgenus Pareucibdelus).

  • — Watanabe, 1998: 314 (Eucibdelus, cited as Eucibderus; Japan).

  • Distribution: Japan, Korea.

aokii R. Dvořák, 1956: 30 (Eucibdelus; Type locality: Japan: Nikko, Chuzenji).

  • — Hayashi, 1997: 35 (Eucibdelus; synonym of japonicus).

chujoi R. Dvořák, 1956: 32 (Eucibdelus; Type locality: Japan: Tukushima-Shikoku, Mount Tsuguri-san).

  • — Hayashi, 1997: 36 (Eucibdelus; synonym of japonicus).

verae Dvořák, 1956: 33 (Eucibdelus; Type locality: Japan: Mount Dando).

  • — Hayashi, 1997: 36 (Eucibdelus; synonym of japonicus).

javanicus Eppelsheim, 1895, see: Rhyncocheilus.

kochi Bernhauer, 1939k: 596 (Eucibdelus; Type locality: N. W. China: Tienmuschan).

  • Distribution: China.

kraatzi Eppelsheim, 1895, see: Rhyncocheilus.

limbatus Fauvel, 1895, see: Rhyncocheilus.

maderi Bernhauer, 1939d: 102 (Eucibdelus; Type locality: China: Prov. Yunnan, Vallis flum. Soling-Ho).

  • Distribution: China.

malaisei Scheerpeltz, 1965: 248 (Eucibdelus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).

  • Distribution: Myanmar.

milkensis Coiffait, 1984e: 380 (Eucibdelus; Type locality: Nepal oriental: Milke Himal, 2000–2300 m).

  • Distribution: Nepal.

montanus Coiffait, 1977b: 239 (Eucibdelus; Type locality: Bhutan: Gogona, 3100 m).

  • Distribution: Bhutan.

nepalensis Scheerpeltz, 1976a: 134 (Eucibdelus; Type locality: Nepal: Chizpan Garhi, 1600 m).

  • Distribution: Nepal.

orientalis 1998b: 104 (Eucibdelus; subgenus Neocibdelus; Type locality: N. E. India: West Garo Hills, Nokrek Nat. Park, Meghalaya St., 1100 ± 150 m elevation, 25°29.6′N, 90°19.5′E. Also cited from Laos and Thailand).

  • Distribution: India, Thailand, Laos.

pseudobirmanus Scheerpeltz, 1965: 247 (Eucibdelus; Type locality: N.E. Burma, Kambaiti, 1800 m).

  • Distribution: Myanmar.

quadricallosus Bernhauer, 1916e: 32 (Eucibdelus; Type locality: Kiangsi).

  • — Scheerpeltz, 1933: 1407 (Eucibdelus; catalog).

  • Distribution: China.

rufipennis Coiffait, 1982: 73 (Eucibdelus; Type locality: Nepal: Thakkhola, zw. Lethe und Ghasa, 2150–2450 m).

  • Distribution: Nepal.

sauteri Bernhauer, 1943a: 180 (Eucibdelus; Type locality: Formosa: Takao).

  • Distribution: Taiwan.

shibatai Hayashi, 1998a: 33 (Eucibdelus; subgenus Pareucibdelus; Type locality: Loloshan, Taiwan).

  • Distribution: Taiwan.

stevensi Cameron, 1932: 222 (Eucibdelus; Type locality: Darjeeling: Gopaldhara).

  • — Scheerpeltz, 1933: 1407 (Eucibdelus; catalog).

  • — Scheerpeltz, 1976a: 144 (Eucibdelus; characters).

  • — Coiffait, 1982a: 233 (Eucibdelus; India).

  • Distribution: India.

tibialis Coiffait, 1977b: 240 (Eucibdelus; Type locality: Bhutan: Gogona, 3100 m).

  • Distribution: Bhutan.

transversiceps Scheerpeltz, 1976b: 139 (Eucibdelus; Type locality: Nepal: Jumbesi, 2750 m).

  • Distribution: Nepal.

tsuguriensis R. Dvořák, 1956, see: Phytolinus.

varicornis Coiffait, 1977, see: Trichocosmetes.

varius Fauvel, 1895: 247 (Eucibdelus; Type locality: Bírmanie, Carin Cheba, 900–1100 m).

  • — Bernhauer and Schubert, 1914: 394 (Eucibdelus; catalog).

  • — Cameron, 1932: 220 (Eucibdelus; characters; Burma).

  • — Scheerpeltz, 1933: 1407 (Eucibdelus; catalog).

  • — Scheerpeltz, 1965: 109 (Eucibdelus; Burma).

  • — Scheerpeltz, 1976a: 91, 143 (Eucibdelus; characters; Nepal).

  • Distribution: Myanmar, Nepal.

verae R. Dvořák, 1956, see: japonicus Sharp, 1874.

yunnanensis Hayashi, 1998a: 36 (Eucibdelus; subgenus Pareucibdelus; Type locality: China, Yunnan Prov., Dongchuan, 1500–3200 m, 26.07N, 103.14E).

  • Distribution: China.

Glenothorax [4 species; Neotropical Region]

Glenothorax Bierig, 1937: 199 (species included: viridipennis, laetus, analis, velox). Type species: Glenothorax viridipennis Bierig, by original designation.

  • — Blackwelder, 1944: 136 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 170 (type species: viridipennis).

analis Bierig, 1937: 203 (Glenothorax; Type locality: Panama: Volcan de Chiriqui).

  • Distribution: Panama.

laetus Bierig, 1937: 203 (Glenothorax; Type locality: Panama: Volcan de Chiriqui).

  • Distribution: Panama.

velox Bierig, 1937: 204 (Glenothorax; Type locality: Panama: Volcan de Chiriqui).

  • Distribution: Panama.

viridipennis Bierig, 1937: 202 (Glenothorax; Type locality: Panama: Volcan de Chiriqui).

  • Distribution: Panama.

Hadropinus [1 species; Palaearctic Region]

Hadropinus Sharp, 1889: 115 (species included: fossor). Type species: Hadropinus fossor Sharp, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 399 (world catalog; 1 species).

  • — Winkler, 1925: 387 (catalog for Palaearctic region).

  • — Blackwelder, 1952: 178 (type species: fossor).

  • — Tikhomirova, 1973a: 188 (checklist of species of USSR).

  • — Moore and Legner, 1976: 540 (includes seashore species; notes).

  • — Naomi, 1982: 128 (characters).

  • — Naomi, 1983: 47 (characters; species of Japan).

  • — Shibata, 1984: 106 (checklist of species of Japan).

fossor Sharp, 1889: 116 (Hadropinus; Type locality: Hakodate).

  • — Bernhauer and Schubert, 1914: 399 (Hadropinus; catalog).

  • — Kirshenblat, 1938: 531 (Hadropinus; Russia).

  • — Nakane, 1963: A239 (Hadropinus; Japan).

  • — Nakane, 1963a: 93 (Hadropinus; characters; habitus photograph; Japan).

  • — Naomi, 1983: 48 (Hadropinus; characters; Japan).

  • — J. Li, 1993: 41 (Hadropinus; China).

  • — Herman, 2001: 31 (Staphylinus; synonymic note).

  • Distribution: Japan, China, Russia.

suturalis Matsumura, 1911: 113 (Staphylinus; subgenus Ocypus; [preoccupied]; Type locality: Tonnaitcha, Insel Sachalin).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; catalog).

  • — Nakane, 1963: A239 (Staphylinus; probably a synonym of Hadropinus fossor).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • — Herman, 2001: 31 (Staphylinus; synonym of fossor).

suturalis Matsumura, 1911, see: fossor Sharp, 1889.

Hadrotes [2 species; Neotropical, Nearctic, and Australian]

Hadrotes Mäklin, 1852: 313 (species included: crassus). Type species: Staphylinus crassus Mannerheim, fixed by monotypy.

  • — LeConte, 1861a: 64 (characters).

  • — LeConte, 1863: 23 (list of North American species).

  • — Gemminger and Harold, 1868: 597 (catalog).

  • — Horn, 1879: 199 (characters).

  • — Duvivier, 1883: 153 (catalog).

  • — Bernhauer and Schubert, 1914: 399 (world catalog; 2 species).

  • — Leng, 1920: 109 (catalog of North American species).

  • — Scheerpeltz, 1933: 1414 (world catalog supplement).

  • — Blackwelder, 1952: 178 (type species: crassus).

  • — Hatch, 1957: 212 (characters; species of Pacific Northwest).

  • — Moore, 1964c (larval characters).

  • — Moore and Legner, 1975: 43 (catalog; North America north of Mexico).

  • — Moore and Legner, 1976: 540 (includes seashore species; notes).

  • — Frank, 1979a (some larval characters and their use in classification).

  • — Moore and Legner, 1979: 34 (characters; notes).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 394 (1 Nearctic species; notes; characters in key).

crassus Mannerheim, 1846: 509 (Staphylinus; Type locality: insula Unalaschka).

  • — Mannerheim, 1852: 313 (Staphylinus; characters; notes; Sitka).

  • — Horn, 1879: 199 (Hadrotes; characters; Alaska to middle California).

  • — Bernhauer and Schubert, 1914: 399 (Hadrotes; catalog).

  • — Scheerpeltz, 1933: 1414 (Hadrotes; catalog).

  • — Hatch, 1957: 212 (Hadrotes; characters; British Columbia; Washington; Oregon).

  • — Moore, 1964c (Hadrotes; larval characters).

  • — Smetana, 1965a: 12 (Hadrotes; British Columbia).

  • — Orth, Moore, and Fisher, 1978: 187 (Hadrotes; characters; collecting notes; California).

  • — Moore and Andrews, 1985: 222 (Hadrotes; Baja California Sur).

  • — Campbell and Davies, 1991: 120 (Hadrotes; checklist of species; Alaska and Canada).

  • Distribution: Canada, USA, Mexico.

extensus LeConte, 1861: 342 (Hadrotes; Type locality: Washington Territory).

  • — Horn, 1879: 199 (Hadrotes; synonym of crassus).

  • — Bernhauer and Schubert, 1914: 399 (Hadrotes; catalog).

  • — Scheerpeltz, 1933: 1414 (Hadrotes; aberration of crassus).

  • — Smetana, 1965a: 12 (Hadrotes; synonym of crassus).

extensus LeConte, 1861, see: crassus Mannerheim, 1846.

hilleri Weise, 1877, see: Liusus.

wakefieldi Cameron, 1945d: 786 (Hadrotes; Type locality: New Zealand: Methven).

  • Distribution: New Zealand.

Leistotrophus [2 species; Neotropical and Nearctic Regions]

Leistotrophus Perty, 1830: 32 (species included: gravenhorsti). Type species: Leistotrophus gravenhorsti Perty, fixed by monotypy.

  • — Laporte, 1835: 111 (cited as Leistrophus; characters; notes).

  • — Erichson, 1839b: 358 (synonym of Staphylinus).

  • — Laporte, 1840: 174 (cited as Leistrophus; characters).

  • — Erichson, 1847: 89 (subgenus of Staphylinus).

  • — Lacordaire, 1854: 76 (cited as Leistostrophus; synonym of Staphylinus).

  • — Redtenbacher, 1857: 984 (characters).

  • — G. Waterhouse, 1858: 23 (cited as Leistrophorus; catalog of British species).

  • — Schaum, 1859: 26 (catalog; European species).

  • — LeConte, 1861a: 64 (cited as Listotrophus; characters).

  • — Gredler, 1863: 101 (infrageneric group of Staphylinus; list of species from Tirol).

  • — LeConte, 1863: 23 (list of North American species).

  • — Wencker and Silbermann, 1866: 28 (list of species; collecting notes; France).

  • — Gemminger and Harold, 1868: 576 (catalog).

  • — Sharp, 1871d: 11 (list of British species).

  • — Fauvel, 1874: 170 [= 1874a: 396] (characters).

  • — Redtenbacher, 1874: 203 (characters).

  • — Fauvel, 1875a: XXVI [= 1875b: 228] (catalog).

  • — Mulsant and Rey, 1876: 225 [= 1877: 81] (characters).

  • — Sharp, 1876: 150 (notes).

  • — Horn, 1879: 198 (characters; key to North American species).

  • — Duvivier, 1883: 142 (catalog).

  • — Bernhauer and Schubert, 1914: 393 (world catalog; 1 species).

  • — Blackwelder, 1944: 139 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Kloet and Hincks, 1945: 167 (synonym of Ontholestes).

  • — Blackwelder, 1952: 213 (type species: gravenhorsti).

  • — Smetana, 1959d: 394 (characters).

Schizochilus Gray, 1832: 304 (species included: brasiliensis). Type species: Schizochilus brasiliensis Gray, fixed by monotypy.

  • — Laporte, 1835: 111 (synonym of Leistotrophus but cited as Schyzochilus and Leistrophus).

  • — Erichson, 1839b: 358 (synonym of Staphylinus).

  • — Laporte, 1840: 174 (synonym of Leistotrophus but cited as Schyzochilus).

  • — Erichson, 1847: 89 (synonym of Staphylinus subgenus Leistotrophus).

  • — Lacordaire, 1854: 76 (synonym of Staphylinus).

  • — Kraatz, 1857d: 532 (synonym of Leistotrophus).

  • — Schaum, 1859: 26 (synonym of Leistotrophus).

  • — Fauvel, 1874: 170 [= 1874a: 396] (synonym of Leistotrophus).

  • — Sharp, 1884: 371 (characters; notes; Mexican and Central American species).

  • — Bernhauer and Schubert, 1914: 393 (synonym of Leistotrophus).

  • — Blackwelder, 1952: 346 (synonym of Leistotrophus; type species: brasiliensis).

Discocephalus Nordmann, 1837: 3, tab. 1, fig. 8 (species included: versicolor; [Note: The only species included was on Tab. 1, fig. 8, where the name was cited as Diococephalus in parentheses following the name “Staphylinus versicolor”]; preoccupied by Ehrenberg, 1831). Type species: Staphylinus versicolor Gravenhorst, fixed by monotypy.

  • — Gemminger and Harold, 1868: 576 (synonym of Leistotrophus).

  • — Fauvel, 1874: 170 [= 1874a: 396] (synonym of Leistotrophus).

  • — Bernhauer and Schubert, 1914: 393 (synonym of Leistotrophus).

  • — Blackwelder, 1952: 129 (synonym of Leistotrophus; type species; notes).

africanus Bernhauer, 1904, see: Ontholestes.

aurosparsus Fauvel, 1895, see: Ontholestes.

barthei Chapman, 1922, see: Ontholestes.

birmanus Fauvel, 1895, see: Thoracostrongylus.

brasiliensis Gray, 1832, see: versicolor Gravenhorst, 1806.

giganteus Kraatz, 1899, see: Staphylinus.

gravenhorsti Perty, 1830, see: versicolor Gravenhorst, 1806.

gracilis Sharp, 1874, see: Ontholestes.

haroldi Eppelsheim, 1884, see: Ontholestes.

infuscatus Nordmann, 1837, see: versicolor Gravenhorst, 1806.

moluccarum Fauvel, 1878, see: Ontholestes.

oculatus Sharp, 1874, see: Ontholestes.

patriarchicus† Scudder, 1876: 78 (Leistotrophus; Type locality: White River near junction with the Green [River]; Fossil).

  • — Scudder, 1890: 507 (Leistotrophus; characters; fossil; near the Colorado-Utah boundary).

  • — Scudder, 1891: 546 (Leistotrophus; catalog; Oligocene; Colorado or Utah).

  • — Handlirsch, 1907: 724 (Leistotrophus; catalog; Oligocene; Colorado).

  • Distribution: USA.

tenuicornis Kraatz, 1859, see: Ontholestes.

versicolor Gravenhorst, 1806: 119 (Staphylinus; Type locality: Brasiliae: Para).

  • — Le Peletier and Serville, 1828: 477 (Staphylinus; characters; notes; Brazil).

  • — Nordmann, 1837: 27 (Staphylinus; characters; Brazil).

  • — Blanchard, 1842: 77 (Staphylinus; Mexico, Colombia, Guyana, Brazil).

  • — Erichson, 1839b: 359 (Staphylinus; characters; Brazil; Surinam; Colombia; Mexico).

  • — Erichson, 1847: 89 (Staphylinus; Peru).

  • — Sharp, 1876: 150 (Leistotrophus; Brazil).

  • — Sharp, 1884: 371 (Schizochilus; Mexico; Guatemala; Nicaragua; Costa Rica; Panama; South America to Brazil).

  • — Fauvel, 1891: 106 (Leistotrophus; Venezuela).

  • — Fauvel, 1901: 84 (Leistotrophus; Colombia; Mexico to Peru and Brazil).

  • — Bernhauer and Schubert, 1914: 393 (Leistotrophus; catalog).

  • — Smyth, 1934: 107 (Schizochilus; natural history notes; Mexico).

  • — Scheerpeltz, 1960c: 120 (Leistotrophus; Bolivia).

  • — Alcock and Forsyth, 1988 (Leistotrophus; mating behavior; Costa Rica).

  • — Forsyth and Alcock, 1990 (Leistotrophus; mimicry; defense behavior; Costa Rica).

  • — Forsyth and Alcock, 1990a (Leistotrophus; predatory behavior; Costa Rica).

  • — Dettner and Liepert, 1994 (Leistotrophus; chemical mimicry and camouflage).

  • — Alcock and Forsyth, 1994 (Leistotrophus; natural history).

  • Distribution: Peru, Brazil, Surinam, Guyana, Venezuela, Colombia, Panama, Costa Rica, Nicaragua, Guatemala.

gravenhorsti Perty, 1830: 30 (Leistotrophus; Type locality: Brasil).

  • — Laporte, 1835: 112 (Leistotrophus; characters; Brazil).

  • — Blanchard, 1842: 77 (Staphylinus; synonym of versicolor).

  • — Erichson, 1839b: 359 (Staphylinus; synonym of versicolor).

  • — Laporte, 1840: 174 (Leistotrophus; characters; Brazil).

  • — Erichson, 1847: 89 (Staphylinus; synonym of versicolor).

  • — Bernhauer and Schubert, 1914: 393 (Leistotrophus; synonym of versicolor).

brasiliensis Gray, 1832: 304 (Schizochilus; Type locality: Brazil).

  • — Blanchard, 1842: 77 (Staphylinus; synonym of versicolor).

  • — Erichson, 1839b: 359 (Staphylinus; synonym of versicolor).

  • — Erichson, 1847: 89 (Staphylinus; synonym of versicolor).

  • — Bernhauer and Schubert, 1914: 393 (Leistotrophus; synonym of versicolor).

infuscatus Nordmann, 1837: 29 (Staphylinus; Type locality: Xalappae in Mexico).

  • — Blanchard, 1842: 77 (Staphylinus; synonym of versicolor).

  • — Erichson, 1839b: 359 (Staphylinus; synonym of versicolor).

  • — Erichson, 1847: 89 (Staphylinus; synonym of versicolor).

  • — Bernhauer and Schubert, 1914: 393 (Leistotrophus; synonym of versicolor).

Liusus [2 species; Palaearctic Region]

Liusus Sharp, 1889: 116 (species included: hilleri). Type species: Hadrotes hilleri Weise, fixed by monotypy.

  • — Heyden, 1893: 42 (list of species of Siberian region).

  • — Bernhauer and Schubert, 1914: 399 (world catalog; 1 species).

  • — Winkler, 1925: 387 (catalog for Palaearctic region).

  • — Blackwelder, 1952: 225 (type species: hilleri).

  • — Tikhomirova, 1973a: 188 (checklist of species of USSR).

  • — Naomi, 1982: 131 (characters; species of Japan).

  • — Watanabe, 1983: 214 (characters; species of Japan; key).

  • — Shibata, 1984: 104 (checklist of species of Japan).

hilleri Weise, 1877: 93 (Hadrotes; Type locality: Hagi).

  • — Sharp, 1889: 117 (Liusus; Japan).

  • — Bernhauer and Schubert, 1914: 399 (Liusus; catalog).

  • — Kirshenblat, 1938: 531 (Liusus, cited as Liusius; abberation named; Russia).

  • — Nakane, 1963: A239 (Liusus; Japan).

  • — Nakane, 1963a: 93 (Liusus; characters; habitus photograph; Japan).

  • — Coiffait, 1974a: 198 (Liusus; Siberia).

  • — Naomi, 1982: 132 (Liusus; characters; Japan).

  • — Watanabe, 1983: 215 (Liusus; characters; Japan).

  • — Yuh, Paik, Kwon, and Lee, 1985: 254 (Liusus; Korea).

  • Distribution: Japan, Korea, Russia.

rufipennis Kirshenblat, 1938: 531, 535 (Liusus, cited as Liusius; aberration of hilleri; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Sakhalina at various localities).

humeralis Matsumura, 1911: 113 (Staphylinus; subgenus Ocypus; Type locality: Solwiyofka, Sachalin).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; catalog).

  • — Watanabe, 1983: 217 (Liusus; characters; Japan).

  • — J. Li, 1993: 41 (Liusus; China).

  • — Yuh, Paik, Kwon, and Lee, 1985: 248 (Liusus; Korea).

  • Distribution: Russia, China, Korea, Japan.

rufipennis Kirshenblat, 1938 [Note: Nomen nudum], see: hilleri Weise, 1877.

Menoedius [3 species; Oriental Region]

Menoedius Fauvel, 1903: 155 (species included: andrewesi). Type species: Menoedius andrewesi Fauvel, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 395 (world catalog; 1 species).

  • — Cameron, 1925: 69 (catalog of Indian species).

  • — Cameron, 1932: 239 (characters; key to species of British India).

  • — Scheerpeltz, 1933: 1409 (world catalog supplement).

  • — Blackwelder, 1952: 237 (type species: andrewesi).

abnormalis Cameron, 1932: 240 (Menoedius; Type locality: India).

  • — Scheerpeltz, 1933: 1409 (Menoedius; catalog).

  • Distribution: India.

andrewesi Fauvel, 1903: 157 (Menoedius; Type locality: Coonoor).

  • — Fauvel, 1904a: 57 (Menoedius; Nilghiris).

  • — Bernhauer and Schubert, 1914: 395 (Menoedius; catalog).

  • — Cameron, 1932: 239 (Mendoedius; characters; India).

  • — Scheerpeltz, 1933: 1409 (Menoedius; catalog).

  • Distribution: India.

subsimilis Cameron, 1932: 240 (Menoedius; Type locality: India).

  • — Scheerpeltz, 1933: 1409 (Menoedius; catalog).

  • Distribution: India.

Miobdelus [1 species; Palaearctic Region]

Miobdelus Sharp, 1889: 111 (species included: brevipennis). Type species: Miobdelus brevipennis Sharp, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 393 (world catalog; 1 species).

  • — Winkler, 1925: 386 (catalog for Palaearctic region).

  • — Scheerpeltz, 1933: 1407 (world catalog supplement).

  • — Blackwelder, 1952: 249 (type species: brevipennis).

  • — Tikhomirova, 1973a: 187 (in checklist of species of USSR).

  • — Shibata, 1984: 96 (in checklist of species of Japan).

  • — Smetana and Davies, 2000: 20, 28, 42 (characters; in checklist of species of north temperate region).

brevipennis Sharp, 1889: 112 (Miobdelus; Type locality: Niohozan; Nikko).

  • — Bernhauer and Schubert, 1914: 393 (Miobdelus; catalog).

  • — Scheerpeltz, 1933: 1407 (Miobdelus; catalog; Korea; China).

  • — Yuh, Paik, Kwon, and Lee, 1985: 246 (Miobdelus; Korea).

  • — J. Li, 1993: 38 (Miobdelus, cited as Milbdelus; China).

  • — Smetana and Davies, 2000: 42 (Miobdelus).

  • Distribution: Japan, Korea, China.

Naddia [19 species; Palaearctic and Oriental Regions]

Naddia Fauvel, 1867d: 117 (replacement name for Caranistes Erichson). Type species: Caranistes westermanni Erichson, fixed by objective synonymy with Caranistes.

  • — Gemminger and Harold, 1868: 574 (catalog).

  • — Bernhauer and Schubert, 1914: 395 (world catalog; 2 species).

  • — Cameron, 1925: 69 (catalog of Indian species).

  • — Cameron, 1932: 241 (characters; key to species of British India).

  • — Scheerpeltz, 1933: 1409 (world catalog supplement).

  • — Blackwelder, 1952: 256 (type species).

  • — Hammond, 1984: 206 (checklist of species of Borneo).

  • — Shibata, 1986a: 124 (supplemental list of species from Taiwan).

  • — Schillhammer, 1996a: 421 (characters).

  • — Smetana and Davies, 2000: 19, 22, 38 (characters; checklist of species of north temperate region).

Caranistes Erichson, 1840: 925 (species included: westermanni; preoccupied by Schönherr, 1839). Type species: Caranistes westermanni Erichson, fixed by monotypy.

  • — Lacordaire, 1854: 75 (characters; notes; list of species).

  • — Gemminger and Harold, 1868: 574 (synonym of Naddia).

  • — Bernhauer and Schubert, 1914: 395 (synonym of Naddia).

  • — Cameron, 1932: 241 (synonym of Naddia).

  • — Blackwelder, 1952: 93 (synonym of Naddia; type species: westermanni).

aeneipennis Cameron, 1930b: 166 (Naddia; Type locality: North Borneo: Bettotan).

  • — Scheerpeltz, 1933: 1409 (Naddia; catalog).

  • Distribution: Borneo.

assamensis Cameron, 1932: 243 (Naddia; Type locality: Assam: Shillong, alt. 5000 feet).

  • — Scheerpeltz, 1933: 1409 (Naddia; catalog).

  • — Smetana and Davies, 2000: 38 (Naddia).

  • Distribution: India.

atripes Bernhauer, 1939k: 597 (Naddia; Type locality: N. W. China: Tienmuschan).

  • — Smetana and Davies, 2000: 38 (Naddia).

  • Distribution: China.

borneensis Cameron, 1930b: 165 (Naddia; Type locality: North Borneo: Bettotan).

  • — Scheerpeltz, 1933: 1409 (Naddia; catalog).

  • Distribution: Borneo.

chinensis Bernhauer, 1929d: 110 (Naddia; Type locality: Hweisin: Kansu).

  • — Scheerpeltz, 1933: 1409 (Naddia; catalog).

  • — Smetana and Davies, 2000: 38 (Naddia).

  • Distribution: China.

decipiens Cameron, 1932: 242 (Naddia; Type locality: Assam).

  • — Scheerpeltz, 1933: 1409 (Naddia; catalog).

  • — Smetana and Davies, 2000: 38 (Naddia).

  • Distribution: India.

drescheri Cameron, 1937: 25 (Naddia; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft.).

  • Distribution: Indonesia.

ishiharai Shibata, 1994: 315 (Naddia; Type locality: Taiwan: near Hsileng, Taoyuan Hsien).

  • — Smetana and Davies, 2000: 38 (Naddia).

  • Distribution: Taiwan.

javana Cameron, 1937: 24 (Naddia; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft.).

  • Distribution: Indonesia.

kediriana Cameron, 1938b: 151 (Naddia; Type locality: E. Java: Res. Kediri, Toempoek; Toempa Meije, Kalibamban).

  • Distribution: Indonesia.

malaisei Scheerpeltz, 1965: 255 (Naddia; Type locality: S. Shan-States, Burma, 1500 m, Taunggyi).

  • — Smetana and Davies, 2000: 38 (Naddia).

  • Distribution: Myanmar.

miniata Fauvel, 1895: 250 (Naddia; Type locality: Tenasserim, Thagata).

  • — Bernhauer and Schubert, 1914: 395 (Naddia; catalog).

  • — Bernhauer, 1929d: 111 (Naddia; characters in key).

  • — Cameron, 1932: 243 (Naddia; characters; Burma).

  • — Scheerpeltz, 1933: 1409 (Naddia; catalog).

  • — Smetana and Davies, 2000: 38 (Naddia).

  • Distribution: Myanmar.

monticola Shibata, 1994: 318 (Naddia; Type locality: Taiwan: near Sungkang, about 2,100 m alt., Nantou Hsien).

  • — Smetana and Davies, 2000: 38 (Naddia).

  • Distribution: Taiwan.

rufipennis Bernhauer, 1915b: 54 (Naddia; Type locality: Ober-Birma).

  • — Bernhauer, 1929d: 111 (Naddia; characters in key).

  • — Cameron, 1932: 244 (Naddia; characters; Burma).

  • — Scheerpeltz, 1933: 1409 (Naddia; catalog).

  • — Smetana and Davies, 2000: 38 (Naddia).

  • Distribution: Myanmar.

taiwanensis Shibata, 1979: 19 (Naddia; Type locality: near Lushan, about 1200 m, alt., Nantou Hsien).

  • — Shibata, 1994: 319 (Naddia; characters in key).

  • — Smetana and Davies, 2000: 38 (Naddia).

  • Distribution: Taiwan.

vethi Bernhauer, 1915h: 234 (Naddia; Type locality: Java: Preanger).

  • — Bernhauer, 1929d: 111 (Naddia; characters in key).

  • — Scheerpeltz, 1933: 1409 (Naddia; catalog).

  • — Cameron, 1937: 24 (Naddia; Java).

  • Distribution: Indonesia.

westermanni Erichson, 1840: 925 (Caranistes; Type locality: Bengala).

  • — Kraatz, 1859: 72 (Caranistes; characters; Bengalia).

  • — Gemminger and Harold, 1868: 574 (Naddia).

  • — Bernhauer and Schubert, 1914: 395 (Naddia; catalog).

  • — Bernhauer, 1929d: 111 (Naddia; characters in key).

  • — Cameron, 1932: 242 (Naddia; characters; Bengal; Burma).

  • — Scheerpeltz, 1933: 1409 (Naddia; catalog).

  • Distribution: India, Myanmar.

wielebnowskiae Schillhammer, 1996a: 421 (Naddia; Type locality: S-Vietnam: 12 km N Dalat Lang Bian. Also cited from Laos).

  • Distribution: Laos, Vietnam.

wittmeri Coiffait, 1982a: 264 (Naddia; Type locality: Inde: Meghalaya, Mawphlang, 1850 m).

  • — Smetana and Davies, 2000: 38 (Naddia).

  • Distribution: India.

Ocypus [148 species (2 doubtful); Nearctic, Palaearctic, and Oriental Regions]

Ocypus Leach, 1819: 172 (species included: cyaneus). Type species: Staphylinus cyaneus Paykull, fixed by original designation and monotypy.

  • — Curtis, 1829: 24 (catalog; Britain).

  • — Stephens, 1829a: 22 (catalog; Britain).

  • — Stephens, 1829: 276 (catalog; Britain).

  • — Stephens, 1832: 200, 211 (characters).

  • — Westwood, 1838a: 15 (characters; type species).

  • — Erichson, 1839a: 439 (characters).

  • — Stephens, 1839: 384 (characters).

  • — Erichson, 1840: 403, 416 (adult and larval characters).

  • — Schaum, 1852: 25 (list of species; Europe).

  • — Lacordaire, 1854: 79 (characters; notes; list of species).

  • — Jacquelin du Val, 1856: 35 (characters).

  • — Kraatz, 1857d: 550 (characters).

  • — Redtenbacher, 1857: 185 (characters; key to Austrian species).

  • — Thomson, 1858: 29 (characters).

  • — G. Waterhouse, 1858: 23 (catalog of British species).

  • — Thomson, 1859: 24 (type species; brunnipes).

  • — Schaum, 1859: 26 (catalog; European species).

  • — Thomson, 1860: 147 (characters).

  • — Gredler, 1863: 102 (list of species from Tirol).

  • — LeConte, 1863: 24 (list of North American species).

  • — Fauvel, 1866c: 247 [= 1866d: 105] (synonym of Staphylinus).

  • — Wencker and Silbermann, 1866: 28 (list of species; collecting notes; France).

  • — Gemminger and Harold, 1868: 581 (catalog).

  • — Jacquelin du Val, 1868: 67 (group of Staphylinus; catalog; European species).

  • — Sharp, 1871d: 11 (list of British species).

  • — Fauvel, 1874: 172 [= 1874a: 398] (synonym of Staphylinus).

  • — Redtenbacher, 1874: 205 (characters; key to Austrian species).

  • — Mulsant and Rey, 1876: 274 [= 1877: 130] (characters; key to French species).

  • — Provancher, 1877: 248 (characters).

  • — Duvivier, 1883: 143 (catalog).

  • — Fowler, 1888: 253 (notes; larval characters; key to British species).

  • — Seidlitz, 1891: 391 (key to some European species).

  • — Seidlitz, 1891a: 416 (key to some European species).

  • — Heyden, Reitter, and Weise, 1891: 101 (list of species of Europe and Caucasus).

  • — Heyden, 1893: 41 (subgenus of Staphylinus; list of species of Siberian region).

  • — Fauvel, 1897d: 294 (subgenus of Staphylinus; catalog of species of Barbary; Canaries).

  • — Heyden, Reitter, and Weise, 1906: 171 (subgenus of Staphylinus; list of species of Europe and Caucasus).

  • — Petri, 1912: 76 (subgenus of Staphylinus; list of species of Siebenbürgen).

  • — Bernhauer and Schubert, 1914: 374 (subgenus of Staphylinus; Stephens, 1832 cited as author of Ocypus).

  • — Leng, 1920: 108 (catalog of North American species).

  • — Winkler, 1925: 385 (catalog for Palaearctic region; subgenus of Staphylinus).

  • — Cameron, 1925: 67 (catalog of Indian species).

  • — Cameron, 1932: 180 (subgenus of Staphylinus).

  • — Scheerpeltz, 1933: 1381 (subgenus of Staphylinus).

  • — Blackwelder, 1943: 444, 445 (type species: cyaneus).

  • — G. Müller, 1943: 106 (valid genus; characters; notes).

  • — Steel, 1949: 272 (subgenus of Staphylinus; charaters; key to British species).

  • — Tottenham, 1949: 374, 420 (type species: ophthalmicus Scopoli = cyaneus Paykull; notes; checklist of British species).

  • — Blackwelder, 1952: 267 (type species: cyaneus).

  • — Coiffait, 1956e: 185, 187, 197, 217 (characters in key; notes; subgeneric and species group classifications; catalog of and key to French species).

  • — Hatch, 1957: 175 (characters; key to subgenera and species of Pacific Northwest).

  • — Smetana, 1958: 279 (characters; type species: ophthalmicus; key to Czechoslovakian species).

  • — Smetana, 1959a: 214 (list of species collected in Albania).

  • — Székessy, 1963: 53 (characters).

  • — Lohse, 1964: 196 (characters; key to central European subgenera and species).

  • — Coiffait, 1964: 81, 82 (type species: ophthalmicus, cited as ophtalmicus; characters).

  • — Moore, 1965: 98, 102 (characters; notes).

  • — Pototskaia, 1966b: 362 (subgenus of Staphylinus; larval characters in key).

  • — Pototskaia, 1967: 67 (subgenus of Staphylinus; larval characters).

  • — Scheerpeltz, 1968a: 84 (catalog of Austrian species).

  • — Bolov, 1969a: 328 (collecting notes for some species; Russia).

  • — Shibata, 1973c: 62 (checklist of species of Taiwan).

  • — Bordoni, 1973j: 698 (list of species from islands near Sicily).

  • — Tikhomirova, 1973a: 186 (checklist of species of USSR).

  • — Coiffait, 1974: 420 (adult and larval characters; key to species of western Palaearctic region).

  • — Orth, Moore, Fisher and Legner, 1975: 1115 (key to Nearctic species).

  • — Moore and Legner, 1975: 20 (catalog; North America north of Mexico).

  • — Frank, 1979a (comments on some larval features and classification of genus; separate from Staphylinus).

  • — Moore and Legner, 1979: 25 (characters; notes).

  • — Muona, 1979: 15 (list of Scandinavian species).

  • — Uhlig, 1979: 255 (some species collected in Germany).

  • — M. Dvořák, 1979: 116 (some species collected in Slovakia).

  • — Uhlig, Vogel, and Sieber, 1980: 247 (some species collected in Germany).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 100 (catalog; Poland).

  • — Uhlig and Vogler, 1981: 120 (list of some species collected in Germany; habitat and collecting notes).

  • — Tóth, 1984: 23 (characters; habitus; key to species of Hungary).

  • — Shibata, 1984: 87 (checklist of species of Japan).

  • — Muona and Viramo, 1986: 13 (list of species of northeastern Finland).

  • — Segers, 1986: 47 (checklist of species; Belgium).

  • — Ádám, 1987: 149 (list of five species with collecting notes; Hungary).

  • — Drugmand 1987b (comments on distribution of Belgian species).

  • — Angelini, 1991: 191 (list of species from high elevations of Calabria, Italy).

  • — Naomi, 1992b: 1 (characters and key to species of dorsalis group of Japan).

  • — Silfverberg, 1992: 17 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Kanehisa, Tsumuki, and Kawazu, 1994 (chemical secretions).

  • — Hernández, Outerelo, and Gamarra, 1994: 188 (checklist of species; Canary Islands).

  • — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 29 (additional records to Danish list of species).

  • — Ciceroni and Zanetti, 1995: 30 (list of species of Italy).

  • — M. Hansen, 1996: 106 (list of Danish species).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 244 (additions to list of Danish species).

  • — Downie and Arnett, 1996: 384 (characters; key to species of Northeastern North America).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (list of species collected in 1997; Denmark).

  • — Assing, Frisch, Kahlen, et al., 1998: 139 (note).

  • — Smetana and Davies, 2000: 21, 31, 42 (characters; checklist of species of north temperate region).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 394 (4 Nearctic species; notes; characters in key).

Goerius Westwood, 1827: 58 (without description, mentioned in combination with olens as a misidentification of larval Philonthus politus). Type species: Staphylinus olens O. Müller, fixed by monotypy.

  • — Curtis, 1829: 24 (catalog; Britain).

  • — Stephens, 1829a: 22 (catalog; Britain).

  • — Stephens, 1829: 275 (catalog; Britain).

  • — Stephens, 1832: 200, 208 (characters).

  • — Westwood, 1838a: 15 (characters; type species).

  • — Erichson, 1839a: 439 (synonym of Ocypus).

  • — Erichson, 1840: 404 (synonym of Ocypus).

  • — Stephens, 1839: 384 (characters).

  • — Lacordaire, 1854: 79 (synonym of Ocypus).

  • — Kraatz, 1857d: 550 (synonym of Ocypus).

  • — Thomson, 1859: 24 (characters; type species: olens).

  • — Thomson, 1860: 146 (characters).

  • — Fauvel, 1874: 172 [= 1874a: 398] (synonym of Staphylinus).

  • — Heyden, 1880: 72 (subgenus of Staphylinus; list of species of Siberian region).

  • — Heyden, Reitter, and Weise, 1891: 101 (subgenus of Ocypus; list of species of Europe and Caucasus).

  • — Heyden, Reitter, and Weise, 1906: 169 (subgenus of Staphylinus; list of species of Europe and Caucasus).

  • — Reitter, 1909: 120 (subgenus of Staphylinus; key to species).

  • — Petri, 1912: 75 (subgenus of Staphylinus; list of species of Siebenbürgen).

  • — Bernhauer and Schubert, 1914: 374 (subgenus of Staphylinus).

  • — Cameron, 1932: 180 (subgenus of Staphylinus).

  • — Scheerpeltz, 1933: 1381 (subgenus of Staphylinus).

  • — Blackwelder, 1943: 444, 445 (subgenus of Ocypus; type species: olens).

  • — Blackwelder, 1952: 173 (type species: olens).

  • — Coiffait, 1956e: 185, 198 (subgenus of Ocypus; characters; notes; species group classification; catalog of and key to species).

  • — Smetana, 1958: 279 (synonym of Ocypus).

  • — Lohse, 1964: 196 (subgenus of Ocypus; characters; key to central European species).

  • — Coiffait, 1964: 81 (type species: olens; synonym of Ocypus).

  • — Coiffait, 1974: 420 (synonym of Ocypus).

  • — Smetana, 1993b: 49 (synonym of Ocypus).

  • — Smetana and Davies, 2000: 31, 42 (synonym of Ocypus subgenus Ocypus).

Isopterum Gistel, 1856: 388, 424 (species included: olens, cyaneum, picipes, simile, alpestre, brunneipes, fuscatum, cupreum, atrum, morio, edentulum, aeneocephalus, ater; preoccupied by Agassiz, 1846). Type species: Staphylinus cyaneum Paykull, fixed by subsequent designation by Blackwelder, 1952: 205.

Matidus Motschulsky, 1860: 569 (species included: forficularius). Type species: Matidus forficularius Motschulsky, fixed by monotypy.

  • — Gemminger and Harold, 1868: 584 (catalog).

  • — Fauvel, 1874: 172 [= 1874a: 398] (synonym of Staphylinus).

  • — Heyden, Reitter, and Weise, 1891: 102 (list of species of Europe and Caucasus).

  • — Heyden, Reitter, and Weise, 1906: 169 (list of species of Europe and Caucasus).

  • — Scheerpeltz, 1933: 1381 (synonym of Staphylinus).

  • — Blackwelder, 1943: 444, 445 (subgenus of Goerius; type species: forficularius).

  • — Blackwelder, 1952: 231 (type species: forficularius).

  • — Smetana, 1958: 279 (synonym of Ocypus).

  • — Coiffait, 1974: 420 (synonym of Ocypus).

  • — Smetana and Davies, 2000: 21, 31, 42 (subgenus of Ocypus; characters; checklist of species of north temperate region).

Pseudocypus Mulsant and Rey, 1876: 291 [= 1877: 147] (subgenus of Ocypus; species included: mus, fuscatus, picipennis, aeneocephalus, confusus, obscuroaeneus, fulvipennis). Type species: Staphylinus mus Brullé, fixed by subsequent designation by Tottenham, 1939d: 229.

  • — Duvivier, 1883: 143 (catalog; synonym of Ocypus).

  • — Seidlitz, 1891: 392 (key to some European species).

  • — Seidlitz, 1891a: 417 (key to some European species).

  • — Heyden, Reitter, and Weise, 1891: 101 (subgenus of Ocypus; list of species of Europe and Caucasus).

  • — Heyden, 1893: 42 (subgenus of Staphylinus; list of species of Siberian region).

  • — Heyden, 1896: 32 (subgenus of Staphylinus; list of species of Siberian region).

  • — Heyden, Reitter, and Weise, 1906: 170 (subgenus of Staphylinus; list of species of Europe and Caucasus).

  • — Reitter, 1909: 119 (subgenus of Staphylinus; key to species).

  • — Petri, 1912: 75 (subgenus of Staphylinus; list of species of Siebenbürgen).

  • — Bernhauer and Schubert, 1914: 374 (subgenus of Staphylinus).

  • — Winkler, 1925: 384 (catalog for Palaearctic region; subgenus of Staphylinus).

  • — Cameron, 1932: 180 (subgenus of Staphylinus).

  • — Scheerpeltz, 1933: 1381 (subgenus of Staphylinus).

  • — Tottenham, 1939d: 229 (type species designation: mus).

  • — Blackwelder, 1943: 444, 445 (subgenus of Ocypus; type species: mus).

  • — Tottenham, 1949: 374, 421 (type species: mus Brullé; subgenus of Ocypus; checklist of British species).

  • — V. Hansen, 1952: 103 (subgenus of Staphylinus).

  • — Blackwelder, 1952: 325 (type species).

  • — Coiffait, 1956e: 186, 198, 207 (subgenus of Ocypus; characters; notes; species group classification; catalog of and key to species).

  • — Hatch, 1957: 176, 177 (subgenus of Ocypus; characters; species of Pacific Northwest).

  • — Smetana, 1958: 279, 289 (subgenus of Ocypus; characters).

  • — Lohse, 1964: 199 (subgenus of Ocypus; characters; key to central European species).

  • — Coiffait, 1964: 81, 82, 82 (type species: mus; characters; key to subgenera).

  • — Pototskaia, 1966b: 362 (subgenus of Staphylinus; larval characters in key).

  • — Pototskaia, 1967: 67 (subgenus of Staphylinus; larval characters).

  • — Bordoni, 1973j: 700 (list of species from islands near Sicily).

  • — Tikhomirova, 1973a: 186 (subgenus of Ocypus; checklist of species of USSR).

  • — Coiffait, 1974: 483 (adult and larval characters; key to species of western Palaearctic region).

  • — M. Dvořák, 1979: 116 (some species collected in Slovakia).

  • — Shibata, 1984: 93 (subgenus of Ocypus; checklist of species of Japan).

  • — Segers, 1986: 48 (checklist of species; Belgium).

  • — Drugmand 1987b (comments on distribution of Belgian species).

  • — Ádám, 1987: 148 (list of five species with collecting notes; Hungary).

  • — Angelini, 1991: 193 (list of species from high elevations of Calabria, Italy).

  • — Hernández, Outerelo, and Gamarra, 1994: 188 (checklist of species; Canary Islands).

  • — Smetana and Davies, 2000: 21, 31, 43 (subgenus of Ocypus; characters; checklist of species of north temperate region).

Protocypus G. Müller, 1923: 136 (subgenus of Staphylinus; species included: fulvotomentosus). Type species: Ocypus fulvotomentosus Eppelsheim, fixed by monotypy.

  • — Cameron, 1932: 180 (subgenus of Staphylinus).

  • — Scheerpeltz, 1933: 1381 (subgenus of Staphylinus).

  • — Blackwelder, 1943: 444, 445 (subgenus of Ocypus; type species: fulvotomentosus).

  • — Blackwelder, 1952: 323 (type species: fulvotomentosus).

  • — Shibata, 1984: 89 (subgenus of Ocypus; checklist of species of Japan).

  • — Naomi, 1992a: 54 (subgenus of Ocypus; characters; key to species of scutiger group; Japan).

  • — Smetana and Davies, 2000: 31 (synonym of Ocypus; subgenus Pseudocypus).

Aulacocypus G. Müller, 1925: 40 (subgenus of Staphylinus; species included: gloriosus, parvulus). Type species: Ocypus gloriosus Sharp, fixed by subsequent designation by Blackwelder, 1943: 445.

  • — Cameron, 1932: 180 (subgenus of Staphylinus).

  • — Scheerpeltz, 1933: 1381 (subgenus of Staphylinus).

  • — Blackwelder, 1943: 445 (subgenus of Ocypus; type species: gloriosus).

  • — Blackwelder, 1952: 70 (type species).

  • — Shibata, 1984: 91 (subgenus of Ocypus; checklist of species of Japan).

  • — Smetana and Davies, 2000: 21, 32, 45 (subgenus of Ocypus; characters; notes; checklist of species of north temperate region).

Xanthocypus G. Müller, 1925: 40 (subgenus of Staphylinus; species included: weisei). Type species: Ocypus weisei Harold, fixed by monotypy.

  • — Scheerpeltz, 1933: 1382 (subgenus of Staphylinus).

  • — Blackwelder, 1943: 445 (subgenus of Ocypus; type species: weisei).

  • — Blackwelder, 1952: 404 (type species: weisei).

  • — Tikhomirova, 1973a: 186 (subgenus of Ocypus; checklist of species of USSR).

  • — Shibata, 1984: 94 (subgenus of Ocypus; checklist of species of Japan).

  • — Hayashi, 1993b: 297 (synonym of Agelosus).

  • — Smetana and Davies, 2000: 31, 42 (synonym of Ocypus subgenus Ocypus).

Atlantogoerius Coiffait, 1956e: 185, 198 (species included: sylvaticus, mateui, umbricola, canariensis). Type species: Ocypus sylvaticus Wollaston, fixed by original designation.

  • — Coiffait, 1974: 409 (characters; key to species of western Palaearctic region).

  • — M. Dvořák, 1984: 66 (synonym of Pseudocypus).

  • — Hernández, Outerelo, and Gamarra, 1994: 187 (checklist of species; Canary Islands).

  • — Smetana and Davies, 2000: 31, 43 (synonym of Ocypus; subgenus Pseudocypus).

Fortunocypus Coiffait, 1964: 82 (subgenus of Pseudocypus; species included: fortunatarum, subaenescens). Type species: Ocypus fortunatarum Wollaston, fixed by original designation.

  • — Coiffait, 1974: 483 (synonym of Pseudocypus).

  • — Smetana and Davies, 2000: 31, 43 (synonym of Ocypus; subgenus Pseudocypus).

Nudabemus Coiffait, 1982: 74 (species included: caerulescens, quadrimaculatus). Type species: Nudabemus caerulescens Coiffait, fixed by original designation.

  • — Smetana and Davies, 2000: 31, 43 (synonym of Ocypus; subgenus Pseudocypus; notes).

abbreviatipennis Aubé, 1850, see: nitens Schrank, 1781.

abbreviatus Motschulsky, 1858, see: fuscatus Gravenhorst, 1802.

abbruzzensis G. Müller, 1926, see: picipennis Fabricius, 1793.

addendus Lindberg, 1953: 4 (Staphylinus; subgenus Goerius; Type locality: Gran Canaria: Tamadaba; Las Lagunetas).

  • — Coiffait, 1954: 161 (Staphylinus; undoubtedly a synonym of canariensis).

  • — Coiffait, 1974: 413 (Atlantogoerius; cited as valid species; characters; Canary Islands).

  • — Hernández, Outerelo, and Gamarra, 1994: 192 (Atlantogoerius; Canary Islands).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Pseudocypus).

  • Distribution: Canary Islands.

aeneicollis Gyllenhal, 1827, see: aeneocephalus De Geer, 1774.

aeneocephalus De Geer, 1774: 22 (Staphylinus; Type locality: Not cited).

  • — Goeze, 1777: 731 (Staphylinus; characters).

  • — Paykull, 1789: 11 (Staphylinus; characters).

  • — Villers, 1789: 423 (Staphylinus; characters).

  • — Olivier, 1795:(42): 17 (Staphylinus; synonym of cupreus).

  • — Paykull, 1800: 374 (Staphylinus; characters; Sweden).

  • — Turton, 1802: 510 (Staphylinus; characters; Germany).

  • — Gravenhorst, 1802: 8 (Staphylinus; characters).

  • — Latreille, 1804: 300 (Staphylinus; characters).

  • — Gravenhorst, 1806: 113 (Staphylinus).

  • — Gyllenhal, 1810: 292 (Staphylinus; characters; Sweden).

  • — Gyllenhal, 1827: 475 (Staphylinus; notes).

  • — Zetterstedt, 1828: 68 (Staphylinus; characters; Lapland).

  • — Curtis, 1829: 24 (Staphylinus; catalog; Britain).

  • — C. Sahlberg, 1830: 311 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 23 (Staphylinus; Finland; Sweden; Germany; Dauria).

  • — Mannerheim, 1831: 437 (Staphylinus; Finland; Sweden; Germany; Dauria).

  • — Stephens, 1832: 207 (Staphylinus; characters; England; Scotland).

  • — Ménétriés, 1832: 143 (Staphylinus; Caucasus-Persia region).

  • — Lacordaire, 1835: 373 (Emus; characters; France).

  • — Nordmann, 1837: 70 (Staphylinus; Europe).

  • — Holme, 1837: 63 (Staphylinus; notes; Scilly Islands).

  • — Zetterstedt, 1838: 60 (Staphylinus; characters; habitat; Lapland).

  • — Curtis, 1839: pl. 758 (Staphylinus; notes; England).

  • — Stephens, 1839: 387 (Staphylinus; characters; England; Scotland).

  • — Erichson, 1839a: 443 (Ocypus; synonym of cupreus).

  • — Erichson, 1840: 413 (Ocypus; synonym of cupreus).

  • — Laporte, 1840: 176 (Staphylinus; characters; France).

  • — Holme, 1842: 116, 117 (Staphylinus; notes).

  • — Dufour, 1843: 33 (Staphylinus; characters; France).

  • — Kolenati, 1846: 18 (Ocypus; synonym of cupreus).

  • — P. Lucas, 1846: 111 (Ocypus; synonym of cupreus).

  • — Redtenbacher, 1849: 698 (Ocypus; [Note: Misidentification: Scheerpeltz, 1933: 1389 cited cupreus Redtenbacher as a synonym of aeneocephalus; Redtenbacher did not describe the species as new, he attributed it to Rossi]; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 511 (Staphylinus; synonym of cupreus).

  • — Kraatz, 1857d: 560 (Ocypus; synonym of cupreus).

  • — Fauvel, 1874: 191 [= 1874a: 417] (Staphylinus; characters; France; Barbary; Canary Islands).

  • — Seidlitz, 1875: 271 (Ocypus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 308 [= 1877: 164] (Ocypus; subgenus Pseudocypus; characters; France).

  • — Fauvel, 1878: 119 [= 1878a: 39] (Staphylinus; Morocco; Barbary).

  • — Ganglbauer, 1895: 434 (Staphylinus; subgenus Pseudocypus; characters; Europe; Mediterranean region).

  • — Fauvel, 1902b: 105 (Staphylinus; subgenus Ocypus; Barbary; Canary Islands).

  • — Poppius, 1905: 9 (Staphylinus; Shetland Islands).

  • — Sainte-Claire Deville, 1906: 100 (Staphylinus; Corsica).

  • — Reitter, 1909: 120 (Staphylinus; subgenus Pseudocypus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; subgenus Ocypus; catalog).

  • — Johansen, 1914: 385 (Staphylinus; subgenus Goerius; characters; Denmark).

  • — Fiori, 1915: 16 (Staphylinus; Italy).

  • — G. Müller, 1923: 141 (Staphylinus; subgenus Pseudocypus; characters; Italy).

  • — G. Müller, 1926: 18 (Staphylinus; subgenus Pseudocypus; characters; notes).

  • — Portevin, 1929: 351 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1389 (Staphylinus; catalog).

  • — Méquignon, 1933: 229 (Staphylinus; distinguishing characters).

  • — Tottenham, 1932 (Ocypus; distinguishing characters).

  • — V. Hansen, 1952: 105 (Staphylinus; subgenus Pseudocypus; characters; Denmark).

  • — Coiffait, 1956e: 212, 215 (Ocypus; subgenus Pseudocypus; characters).

  • — Hatch, 1957: 177 (Ocypus; subgenus Pseudocypus; characters; British Columbia).

  • — Smetana, 1958: 292 (Ocypus; subgenus Pseudocypus; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 116 (Staphylinus; subgenus Pseudocypus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 200 (Ocypus; subgenus Pseudocypus; characters; central Europe).

  • — Moore, 1965: 102 (Ocypus; introduced in North America).

  • — Smetana, 1965a: 11 (Ocypus; subgenus Pseudocypus; British Columbia).

  • — Horion, 1965: 223 (Staphylinus; subgenus Pseudocypus; Russia; Finland; Sweden; Norway; Britain; Denmark; all central Europe; Portugal; Spain; Switzerland; Italy; Romania; Hungary; Czechoslovakia; Poland; Dalmatia).

  • — Szujecki, 1968a: 733 (Ocypus; Poland).

  • — Coiffait, 1974: 511 (Pseudocypus; characters; England; France; Spain).

  • — Orth, Moore, Fisher and Legner, 1975: 1115 (Ocypus; characters).

  • — Pope, 1977: 30 (Staphylinus; Britain).

  • — Tóth, 1984: 30 (Ocypus; characters; Hungary).

  • — Newton, 1987: 381 (Staphylinus; introduced species; British Columbia).

  • — Lucht, 1987: 108 (Ocypus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Huth and Dettner, 1989: 154 (Ocypus; cited as aenaeocephalus; defensive secretions).

  • — Welch, 1993: 229 (Staphylinus; ovariole number and ovary structure).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Pseudocypus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; Italy).

  • — Pilon, 1998: 88 (Ocypus; ecological notes; Italy).

  • — Staniec, 1999a: 56 (Ocypus; characters of pupa; Poland).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Pseudocypus).

  • — Assing, 2001: 75 (Ocypus; collected from xerothermous site; Germany).

  • Distribution: Europe.

aeneicollis Gyllenhal, 1827: 475 (Staphylinus; Type locality: Not cited).

  • — C. Sahlberg, 1830: 311 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 23 (Staphylinus; Sweden).

  • — Mannerheim, 1831: 437 (Staphylinus; Sweden).

  • — Stephens, 1835: 435 (Staphylinus; characters; England).

  • — Runde, 1835: 3 (Staphylinus; characters; Germany).

  • — Nordmann, 1837: 70 (Staphylinus; Europe).

  • — Erichson, 1839a: 443 (Ocypus; synonym of cupreus).

  • — Stephens, 1839: 387 (Staphylinus; characters; England).

  • — Erichson, 1840: 413 (Ocypus; synonym of cupreus).

  • — Holme, 1842: 117 (Staphylinus; notes).

  • — P. Lucas, 1846: 111 (Ocypus; synonym of cupreus).

  • — Ménétriés, 1849: 43 (Ocypus; Khirgiz region).

  • — Fairmaire and Laboulbène, 1856: 511 (Staphylinus; synonym of cupreus).

  • — Kraatz, 1857d: 560 (Ocypus; synonym of cupreus).

  • — Thomson, 1860: 149 (Ocypus; synonym of cupreus).

  • — Ganglbauer, 1895: 434 (Staphylinus; synonym of aeneocephalus).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; synonym of aeneocephalus).

  • — Scheerpeltz, 1933: 1389 (Staphylinus; synonym of aeneocephalus).

  • — Smetana, 1958: 293 (Ocypus; synonym of aeneocephalus).

  • — Coiffait, 1974: 512 (Pseudocypus; synonym of aeneocephalus).

puberulus Runde, 1835: 3 (Staphylinus; Type locality: Halae).

  • — Erichson, 1839a: 443 (Ocypus; synonym of cupreus).

  • — Erichson, 1840: 413 (Ocypus; synonym of cupreus).

  • — Kraatz, 1857d: 560 (Ocypus; synonym of cupreus).

  • — Ganglbauer, 1895: 434 (Staphylinus; synonym of aeneocephalus).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; synonym of aeneocephalus).

  • — Smetana, 1958: 293 (Ocypus; synonym of aeneocephalus).

  • — Coiffait, 1974: 512 (Pseudocypus; synonym of aeneocephalus).

strigatus Nordmann, 1837: 70 (Staphylinus; Type locality: Italia prope Pisam).

  • — Erichson, 1839a: 443 (Ocypus; synonym of cupreus).

  • — Erichson, 1840: 413 (Ocypus; synonym of cupreus).

  • — P. Lucas, 1846: 111 (Ocypus; synonym of cupreus).

  • — Kraatz, 1857d: 560 (Ocypus; synonym of cupreus).

  • — Ganglbauer, 1895: 434 (Staphylinus; synonym of aeneocephalus).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; synonym of aeneocephalus).

  • — Smetana, 1958: 293 (Ocypus; synonym of aeneocephalus).

  • — Coiffait, 1974: 512 (Pseudocypus; synonym of aeneocephalus).

cyanescens Gerhardt, 1910: 555 (Staphylinus; aberration of aeneocephalus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Ostpreussen).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; aberration of aeneocephalus).

  • — Smetana, 1958: 293 (Ocypus; aberration of aeneocephalus).

  • — Coiffait, 1974: 512 (Pseudocypus; variety of aeneocephalus).

aenescens Eppelsheim, 1889a: 174 (Ocypus; subgenus Goerius; Type locality: Kan-ssu).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: China.

aereus Cameron, 1928c: 565 (Ocypus; subgenus Pseudocypus; Type locality: Sikkim: Tangu, 11,500 feet).

  • — Cameron, 1932: 204 (Staphylinus; subgenus Pseudocypus; characters; India).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; catalog).

  • — Scheerpeltz, 1976a: 90 (Staphylinus; Nepal).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Pseudocypus).

  • Distribution: India, Nepal.

aethiops Waltl, 1835: 56 (Staphylinus; Type locality: Andalusien).

  • — Rosenhauer, 1856: 70 (Ocypus; Cadiz; Algeciras).

  • — Fairmaire and Laboulbène, 1856: 510 (Staphylinus; characters; France).

  • — Peyron, 1858: 422 (Ocypus; Turkey).

  • — Fairmaire and Coquerel, 1860: 157 (Ocypus; Barbary Coast).

  • — Fauvel, 1874: 186 [= 1874a: 412] (Staphylinus; characters; France; Spain; Portugal; Italy; Sicily; Cyprus; Turkey; Barbary; Azores).

  • — Mulsant and Rey, 1876: 299 [= 1877: 155] (Ocypus; characters; France).

  • — Fauvel, 1878: 119 [= 1878a: 39] (Staphylinus; Morocco; Algeria).

  • — Fauvel, 1886: 50 [= 1886a: 42] (Staphylinus; Barbary).

  • — Ganglbauer, 1895: 432 (Staphylinus; subgenus Goerius; characters; France; Mediterranean region).

  • — Fauvel, 1902b: 104 (Staphylinus; subgenus Ocypus; Morocco; Algeria; Tunisia; Azores).

  • — Reitter, 1909: 120 (Staphylinus; subgenus Goerius; characters; Germany).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; subgenus Ocypus; catalog).

  • — Portevin, 1929: 352 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; catalog).

  • — Bernhauer, 1940c: 4 (Staphylinus; subgenus Ocypus; Azores).

  • — Méquignon, 1942a: 22 (Staphylinus; Azores).

  • — Jarrige, 1947a: 155 (Goerius; characters; France).

  • — Jarrige, 1954a: 163 (Staphylinus; subgenus Goerius; characters; Azores).

  • — Coiffait, 1956e: 208, 214 (Ocypus; subgenus Pseudocypus; characters).

  • — Smetana, 1970: 61 (Ocypus; distributional summary; Azores).

  • — Coiffait, 1974: 499 (Pseudocypus; characters; Europe; Mediterranean region; North Africa).

  • — Israelson, 1981: 3 (Pseudocypus; characters; notes; Madeira).

  • — Borges, 1990: Table II (Pseudocypus; Azores).

  • — Pilon, 1998: 88 (Ocypus; ecological notes; Italy).

  • — Outerelo, Gamarra, and Salgado, 1998: 130 (Pseudocypus; subgenus Pseudocypus; collected from cave; Spain; Portugal).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Pseudocypus).

  • Distribution: Tunisia, Algeria, Morocco, Madeira, Azores, Portugal, Spain, France, Italy.

masculus Nordmann, 1837: 67 (Staphylinus; Type locality: Lusitania).

  • — Erichson, 1840: 409 (Ocypus; characters; Portugal).

  • — Erichson, 1840: 927 (Staphylinus; synonym of aethiops).

  • — P. Lucas, 1846: 110 (Ocypus; notes; Algeria).

  • — Fairmaire and Laboulbène, 1856: 510 (Staphylinus; synonym of aethiops).

  • — Fauvel, 1874: 186 [= 1874a: 412] (Staphylinus; synonym of aethiops).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; synonym of aethiops).

  • — Jarrige, 1947a: 155 (Goerius; synonym of aethiops).

  • — Coiffait, 1974: 499 (Pseudocypus; synonym of aethiops).

hesperus Crotch, 1867a: 391 (Staphylinus; Type locality: Azores: Terceira).

  • — Fauvel, 1874: 186 [= 1874a: 412] (Staphylinus; synonym of aethiops).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; synonym of aethiops).

  • — Jarrige, 1947a: 155 (Goerius; synonym of aethiops).

  • — Coiffait, 1974: 499 (Pseudocypus; synonym of aethiops).

luigionii G. Müller, 1926: 16 (Staphylinus; subgenus Goerius; subspecies of aethiops; Type locality: Sizilien).

  • — G. Müller, 1926b: 43 (Staphylinus; subgenus Goerius; cited as new subspecies of aethiops; characters; Sicily).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; subspecies of aethiops).

  • — Coiffait, 1956e: 208 (Ocypus; subgenus Pseudocypus; subspecies of aethiops; characters).

  • — Coiffait, 1974: 499 (Pseudocypus; cited as luigioni; subspecies of aethiops; Sicily).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; subspecies of aethiops; Italy).

  • — Sparacio, 1995: 162 (Ocypus; subspecies of aethiops; characters; notes; Sicily).

  • — Smetana and Davies, 2000: 43 (Ocypus; synonym of aethiops).

inauratus Normand, 1947a: 8 (Staphylinus; subspecies of aethiops; Type locality: Tunisie: Le Kef).

  • — Coiffait, 1974: 499 (Pseudocypus; variety of aethiops).

affinis Wollaston, 1864: 566 (Ocypus; Type locality: Barranco de Galga; Barranco above Santa Cruz).

  • — Wollaston, 1865: 488 (Ocypus; notes; Canary Islands).

  • — Fauvel, 1902b: 104 (Staphylinus; subgenus Ocypus; Canary Islands).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; catalog).

  • — Coiffait, 1954: 166 (Goerius; Canary Islands).

  • — Jarrige, 1954a: 162 (Staphylinus; subgenus Goerius; characters; Canaries).

  • — Coiffait, 1956e: 203 (Ocypus; subgenus Goerius; characters).

  • — Coiffait, 1974: 471 (Ocypus; characters; Canary Islands).

  • — Assing and Wunderle, 1999: 437 (Ocypus; Canary Islands).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus).

  • Distribution: Canary Islands.

albanicus G. Müller, 1943: 100 (Staphylinus; subgenus Goerius; Type locality: Albania meridionale: Brataj Ramica).

  • — G. Müller, 1950: 10 (Staphylinus; subgenus Goerius; notes).

  • — Coiffait, 1974: 451 (Ocypus; characters; Albania).

  • — M. Dvořák, 1984: 61 (Ocypus; characters).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Matidus).

  • Distribution: Albania.

albanicus Schatzmayr, 1943: 114 (Staphylinus; subspecies of kuntzeni; [preoccupied]; Type locality: Fushes Dukati).

  • — Coiffait, 1974: 451 (Ocypus; synonym of albanicus).

albanicus Schatzmayr, 1943, see: albanicus G. Müller, 1943.

alfierii Bernhauer, 1925a: 37 (Staphylinus; Type locality: Aegypten bei Amrieh, Mariout).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus).

  • Distribution: Egypt.

algiricus G. Müller, 1932, see: obscuroaeneus Fairmaire, 1852.

almensis Coiffait, 1967c: 172 (Ocypus; Type locality: environs de Alma-Ata, R.S.S. du Kazakstan).

  • — Coiffait, 1974: 460 (Ocypus; characters; Kazakhstan).

  • — M. Dvořák, 1984: 62 (Ocypus; Kazakhstan).

  • — Boháč, 1988: 552 [= 1989: 36] (Ocypus; Uzbekistan).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Pseudocypus).

  • Distribution: Kazakhstan, Uzbekistan.

almorensis Cameron, 1932, see: helleni G. Müller, 1926.

alpestris Erichson, 1840: 408 (Ocypus; Type locality: alpes Carinthiaces, Tyrolenses).

  • — Redtenbacher, 1849: 698 (Ocypus; characters; Austria).

  • — Küster, 1853: 56 (Ocypus; characters; Austria).

  • — Kraatz, 1857d: 554 (Ocypus; characters; Germany).

  • — Redtenbacher, 1857: 187 (Ocypus; characters; Austria).

  • — Fauvel, 1865a: 294 [= 1865b: 44] (Ocypus; habitat; France).

  • — Fauvel, 1867: LVI (Ocypus; characters; notes; France; Italy).

  • — Fauvel, 1874: 184 [= 1874a: 410] (Staphylinus; characters; France; Germany; Austria; Switzerland; Italy; Caucasus).

  • — Redtenbacher, 1874: 206 (Ocypus; characters; Austria).

  • — Mulsant and Rey, 1876: 292 [= 1877: 148] (Ocypus; characters; France).

  • — Ganglbauer, 1895: 431 (Staphylinus; variety of macrocephalus).

  • — Reitter, 1909: 121 (Staphylinus; subgenus Goerius; variety of macrocephalus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; variety of macrocephalus).

  • — G. Müller, 1926: 11 (Staphylinus; subgenus Goerius; characters; notes).

  • — G. Müller, 1926c: 74 (Staphylinus; subgenus Goerius; characters; notes).

  • — Portevin, 1929: 353 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; catalog).

  • — G. Müller, 1943: 101 (Staphylinus; subgenus Goerius; characters; notes).

  • — Lohse, 1964: 198 (Ocypus; subgenus Goerius; characters; central Europe).

  • — Horion, 1965: 215 (Staphylinus; subgenus Goerius; Italy; Slovenia; Austria).

  • — Coiffait, 1974: 462 (Ocypus; characters; Alps).

  • — Frank, 1982: 37 (Ocypus; list of parasites).

  • — Lucht, 1987: 108 (Ocypus; checklist; Austria).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; Italy).

  • — Pilon, 1995: 32, 35 (Ocypus; characters; Italy).

  • — Pilon, 1998: 77 (Ocypus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Matidus).

  • Distribution: Austria, Italy, Slovenia.

alpinus Gravenhorst, 1846: 96 (Staphylinus; [preoccupied]; Type locality: Oesterreich).

  • — Eppelsheim, 1887b: 431 (Ocypus; form of macrocephalus).

  • — Ganglbauer, 1895: 431 (Staphylinus; synonym of macrocephalus).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; synonym of macrocephalus).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; synonym of alpestris).

alpicola Erichson, 1840, see: brunnipes Fabricius, 1781.

alpinus Gravenhorst, 1846, see: alpestris Erichson, 1840.

alsaticus Coiffait, 1964, see: chevrolatii Baudi, 1848.

altaiensis G. Müller, 1926, see: picipennis Fabricius, 1793.

altissimus Coiffait, 1977b: 232 (Ocypus; Type locality: Bhutan: Pele-La, 3450 m).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: Bhutan.

ambiguus Coiffait, 1964: 93 (Ocypus; Type locality: Turquie d'Asie: Goek Dagh).

  • — Coiffait, 1974: 477 (Ocypus; characters; Anatolia).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: Turkey.

anatolicus Coiffait, 1964, see: picipennis Fabricius, 1793.

andorranus Coiffait, 1964, see: picipennis Fabricius, 1793.

angustatus Stephens, 1832, see: sericeicollis Ménétriés, 1832.

angustulus Eppelsheim, 1888: 61 (Ocypus; subgenus Goerius; Type locality: Taschkent).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; catalog).

  • — Coiffait, 1974: 460 (Ocypus; characters; Turkestan).

  • — M. Dvořák, 1984: 61 (Ocypus; characters; Kirgizia).

  • — Boháč, 1988: 552 [= 1989: 36] (Ocypus; characters).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Pseudocypus).

  • Distribution: Uzbekistan, Kyrgyzstan.

anophthalmicus Hernández and Aguiar, 1988: 85 (Atlantogoerius; Type locality: Islas Canarias: Tenerife, Monte de Los Silos).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Pseudocypus).

  • Distribution: Canary Islands.

aprutianus G. Müller, 1926, see: picipennis Fabricius, 1793.

apterus Scheerpeltz, 1976: 40 (Staphylinus; subgenus Pseudocypus; Type locality: Nepal: Unterhalb Mingo-Alm, ca. 4500 m).

  • — Scheerpeltz, 1976a: 90 (Staphylinus; Nepal).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Pseudocypus).

  • Distribution: Nepal.

asturicus Coiffait, 1956, see: picipennis Fabricius, 1793.

atavus† Oustalet, 1874: 162 (Staphylinus; subgenus Ocypus; [preoccupied]; Type locality: Aix en Provence).

  • — Scudder, 1891: 584 (Staphylinus; subgenus Ocypus; catalog; Ligurian; France).

  • — Handlirsch, 1907: 725 (Ocypus; catalog; Oligocene; France).

  • — Herman, 2001: 45 (Ocypus; seems appropriate to forgo replacing this name because the two homonyms have been in different genera since 1907).

  • Distribution: France.

atlasicus Coiffait, 1974, see: obscuroaeneus Fairmaire, 1852.

atratus Wollaston, 1864, see: Tasgius.

atrocoerulescens Goeze, 1777, see: ophthalmicus Scopoli, 1763.

atrocyaneus Fairmaire, 1860, see: ophthalmicus Scopoli, 1763.

aureotomentosus Bernhauer, 1900, see: tenebricosus Gravenhorst, 1846.

auricomus Lindberg, 1953, see: mateui Coiffait, 1954.

auroguttatus Cameron, 1932: 206 (Staphylinus; subgenus Tasgius; Type locality: Bengal).

  • — Scheerpeltz, 1933: 1391 (Staphylinus; catalog).

  • — Coiffait, 1982a: 233 (Nudabemus; Nepal; India).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus).

  • Distribution: India, Nepal.

aurulentus Lokay, 1921 [Note: Nomen nudum], see: olens O. Müller, 1764.

australis Redtenbacher, 1867, see: Cafius.

azoricus Méquignon, 1942, see: olens O. Müller, 1764.

azurescens Mannerheim, 1830, see: ophthalmicus Scopoli, 1763.

baicalensis Eppelsheim, 1887, see: ophthalmicus Scopoli, 1763.

balcanicus G. Müller, 1923, see: ophthalmicus Scopoli, 1763.

balearicus G. Müller, 1926, see: ophthalmicus Scopoli, 1763.

barbarus G. Müller, 1926, see: picipennis Fabricius, 1793.

baronii Coiffait, 1982a: 266 (Nudabemus; Type locality: Nepal: Phulchoki, 2600 m).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus).

  • Distribution: Nepal.

baudii Fauvel, 1867, see: rhaeticus Eppelsheim, 1873.

baudii Peyron, 1858, see: syriacus Baudi, 1848.

bellicosus Fairmaire, 1855, see: Tasgius.

benoiti Drugmand, 1998, see: ophthalmicus Scopoli, 1763.

bernhaueri G. Müller, 1925: 44 (Staphylinus; subgenus Goerius; Type locality: Turkestan: Mts. Karateghin, Sary-Pul).

  • — Scheerpeltz, 1933: 1391 (Staphylinus; catalog).

  • — Coiffait, 1974: 477 (Ocypus; characters; Turkestan).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Matidus).

  • Distribution: “Turkestan”.

bhutanicus Coiffait, 1977b: 233 (Ocypus; Type locality: Bhutan: Dorjula, 3100 m).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: Bhutan.

bicolor G. Müller, 1943, see: helleni G. Müller, 1926.

biharicus G. Müller, 1926a: 28 (Staphylinus; subgenus Goerius; subspecies of tenebricosus; Type locality: monti Bihar in Rumenia, Detunata, Vartopul, Scarisoara ecc.).

  • — Scheerpeltz, 1933: 1391 (Staphylinus; catalog).

  • — Smetana, 1958: 285 (Ocypus; subgenus Ocypus; characters; ecology; Czechoslovakia).

  • — Szujecki, 1963a: 34 (Ocypus; Poland).

  • — Lohse, 1964: 199 (Ocypus; subgenus Goerius; characters; central Europe).

  • — M. Dvořák, 1965: 89 (Ocypus; notes; Slovakia).

  • — Horion, 1965: 220 (Staphylinus; subgenus Goerius; Romania; Czechoslovakia; Poland; Austria).

  • — Coiffait, 1971b: 280 (Ocypus; Bulgaria).

  • — Coiffait, 1974: 454 (Ocypus; characters; Romania; Bulgaria; Austria; Czechoslovakia).

  • — Tóth, 1984: 28 (Ocypus; characters; Hungary).

  • — Lucht, 1987: 108 (Ocypus; checklist; Poland; Czechoslovakia; Austria).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Ocypus; Czech Republic; Slovakia).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Matidus).

  • Distribution: Bulgaria, Romania, Hungary, Czechoslovakia, Austria.

bimaculatus Cameron, 1932, see: cameroni Smetana and Davies, 2000.

bisetosus G. Müller, 1943, see: brenskei Reitter, 1884.

brenskei Reitter, 1884a: 44 (Ocypus; Type locality: Kumani).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; variety of similis).

  • — G. Müller, 1926: 14 (Staphylinus; subgenus Goerius; characters; notes).

  • — Scheerpeltz, 1933: 1391 (Staphylinus; catalog).

  • — G. Müller, 1943: 99 (Staphylinus; subgenus Goerius; characters; notes; Albania; Greece; Macedonia).

  • — Coiffait, 1974: 471 (Ocypus; characters; Greece).

  • — M. Dvořák, 1984: 65 (Ocypus; characters; Greece).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Matidus).

  • Distribution: Greece, Madedonia, Albania.

bisetosus G. Müller, 1943: 100 (Staphylinus; subgenus Goerius; form of brenskei; Type locality: Maj'e Tartarit nell'Albania meridionale).

  • — Coiffait, 1974: 472 (Ocypus; variety of brenskei; Albania).

  • — Smetana and Davies, 2000: 42 (Ocypus; synonym of brenskei).

holosetosus G. Müller, 1943: 100 (Staphylinus; subgenus Goerius; subspecies of brenskei; Type locality: Mal i Dajtit; Mal i That, tra i laghi di Ochrida e di Presba; valico di Kulmak nel gruppo del Tomor; Logara, tra Valona e Argirocastro; Mal i Gjer, presso Argirocastro).

  • — Coiffait, 1974: 472 (Ocypus; subspecies of brenskei; characters; Albania).

  • — Smetana and Davies, 2000: 42 (Ocypus; synonym of brenskei).

brevicornis Weise, 1877, see: Platydracus.

brevipennis Heer, 1839: 254 (Staphylinus; Type locality: Pilatus, Beverseralpne).

  • — Redtenbacher, 1849: 698 (Ocypus; synonym of alpestris).

  • — Kraatz, 1857d: 554 (Ocypus; synonym of alpestris).

  • — Fauvel, 1866c: 248 [= 1866d: 106] (Staphylinus; synonym of alpestris).

  • — Fauvel, 1869: 487 (Ocypus; synonym of alpestris).

  • — Ganglbauer, 1895: 431 (Staphylinus; synonym of macrocephalus).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; synonym of macrocephalus).

  • — G. Müller, 1926: 11 (Staphylinus; subgenus Goerius; valid species; characters; notes).

  • — G. Müller, 1926c: 74 (Staphylinus; subgenus Goerius; notes).

  • — Scheerpeltz, 1933: 1391 (Staphylinus; catalog).

  • — Jarrige, 1947a: 156 (Goerius; characters; France; Alps).

  • — Coiffait, 1956e: 205, 207 (Ocypus; subgenus Goerius; characters).

  • — Lohse, 1964: 198 (Ocypus; subgenus Goerius; characters; central Europe).

  • — Horion, 1965: 216 (Staphylinus; subgenus Goerius; France; Switzerland; Italy; Germany).

  • — Coiffait, 1974: 466 (Ocypus; characters; Alps).

  • — Lucht, 1987: 108 (Ocypus; nominate subspecies; checklist; Germany; Austria; Switzerland).

  • — Pilon and Zanetti, 1991: 68 (Ocypus; Italy).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; Italy).

  • — Pilon, 1995: 32, 35 (Ocypus; characters; Italy).

  • — Pilon, 1998: 77 (Ocypus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Matidus).

  • Distribution: Germany, France, Switzerland, Italy, Austria.

tenuipennis Erichson, 1840: 408 (Ocypus; variety of alpestris; Type locality: Austria).

  • — Gravenhorst, 1846: 97 (Staphylinus; characters; Hungary).

  • — Kraatz, 1857d: 554 (Ocypus; synonym of alpestris).

  • — Ganglbauer, 1895: 431 (Staphylinus; synonym of macrocephalus).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; synonym of macrocephalus).

  • — Scheerpeltz, 1933: 1391 (Staphylinus; synonym of brevipennis).

  • — Coiffait, 1974: 467 (Ocypus; synonym of brevipennis).

pseudoalpestris G. Müller, 1926c: 75 (Staphylinus; subspecies of brevipennis; Type locality: Stiria: Teichalpe, Koralpe, Bürgeralpe, Hochschwab; Austria inferior).

  • — G. Müller, 1926: 12 (Staphylinus; subgenus Goerius; subspecies of brevipennis; [Note: Published without characters, unavailable name in this article]; notes).

  • — Scheerpeltz, 1933: 1391 (Staphylinus; subspecies of brevipennis).

  • — Coiffait, 1956e: 205 (Ocypus; subgenus Goerius; subspecies of brevipennis; characters).

  • — Coiffait, 1974: 467 (Ocypus; subspecies of brevipennis).

  • — Lucht, 1987: 108 (Ocypus; subspecies of brevipennis; checklist; Germany; Austria).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Matidus; subspecies of brevipennis).

brigitteae Drugmand, 1998, see: ophthalmicus Scopoli, 1763.

brunneopubens G. Müller, 1926, see: graeseri Eppelsheim, 1887.

brunnipes Fabricius, 1781: 336 (Staphylinus; Type locality: Anglia).

  • — Fabricius, 1787: 220 (Staphylinus; characters).

  • — Villers, 1789: 419 (Staphylinus; characters; England).

  • — Gmelin, 1790: 2028 (Staphylinus; characters).

  • — Fabricius, 1793: 524 (Staphylinus; characters; England).

  • — Olivier, 1795:(42): 13 (Staphylinus; characters; France, England).

  • — Fabricius, 1801: 595 (Staphylinus; characters; England).

  • — Turton, 1802: 511 (Staphylinus; characters; England).

  • — Marsham, 1802: 502 (Staphylinus; characters; Britain).

  • — Latreille, 1804: 326 (Staphylinus; characters; England).

  • — Gravenhorst, 1806: 112 (Staphylinus; characters; France).

  • — Gyllenhal, 1810: 289 (Staphylinus; characters; Sweden).

  • — Curtis, 1829: 24 (Ocypus; catalog; Britain).

  • — C. Sahlberg, 1830: 310 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 24 (Staphylinus; Gotland).

  • — Mannerheim, 1831: 438 (Staphylinus; Gotland).

  • — Stephens, 1832: 207 (Staphylinus; characters; England).

  • — Runde, 1835: 4 (Staphylinus; characters; Germany).

  • — Lacordaire, 1835: 371 (Emus; characters; France).

  • — Nordmann, 1837: 73 (Staphylinus; Germany; England; Sweden; Tauria).

  • — Erichson, 1839a: 441 (Ocypus; characters; Germany).

  • — Heer, 1839: 254 (Staphylinus; characters; Switzerland).

  • — Stephens, 1839: 387 (Staphylinus; characters; England).

  • — Erichson, 1840: 410 (Ocypus; characters; Germany; France; England; Sweden).

  • — Laporte, 1840: 176 (Staphylinus; characters; France).

  • — Holme, 1842: 118 (Staphylinus; notes).

  • — Jacobsen, 1849: 684 (Ocypus; characters; Denmark).

  • — Redtenbacher, 1849: 698 (Ocypus; characters; Austria).

  • — Küster, 1853: 58 (Ocypus; characters; Sweden; Finland; England; France; Germany; Switzerland).

  • — Hardy, 1851: 23 (Ocypus; subgenus Goerius; notes; England).

  • — Fairmaire and Laboulbène, 1856: 510 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 557 (Ocypus; characters; Germany).

  • — Redtenbacher, 1857: 188 (Ocypus; characters; Austria).

  • — Thomson, 1859: 24 (type species of Ocypus).

  • — Thomson, 1860: 148 (Ocypus; characters; Scandinavia).

  • — Hochhuth, 1862: 49 (Ocypus; Russia).

  • — Fauvel, 1874: 184 [= 1874a: 410] (Staphylinus; characters; France; Italy; Caucasus; Turkey).

  • — Redtenbacher, 1874: 207 (Ocypus; characters; Austria).

  • — Seidlitz, 1875: 270 (Ocypus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 296 [= 1877: 152] (Ocypus; characters; France).

  • — J. Sahlberg, 1876: 11 (Ocypus; Finland).

  • — Fowler, 1888: 255 (Ocypus; characters; Britain).

  • — Ganglbauer, 1895: 433 (Staphylinus; subgenus Pseudocypus; characters; north and middle Europe; eastern Mediterranean region).

  • — Reitter, 1909: 120 (Staphylinus; subgenus Pseudocypus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; subgenus Ocypus; catalog).

  • — Johansen, 1914: 384 (Staphylinus; subgenus Goerius; characters; Denmark).

  • — G. Müller, 1926: 17 (Staphylinus; subgenus Pseudocypus; notes).

  • — Portevin, 1929: 352 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1391 (Staphylinus; catalog).

  • — V. Hansen, 1952: 103 (Staphylinus; subgenus Pseudocypus; characters; Denmark).

  • — Coiffait, 1956e: 204, 206 (Ocypus; subgenus Goerius; characters).

  • — Smetana, 1958: 290 (Ocypus; subgenus Pseudocypus; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 115 (Staphylinus; subgenus Pseudocypus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 197 (Ocypus; subgenus Goerius; characters; central Europe).

  • — Horion, 1965: 212 (Staphylinus; subgenus Goerius; Russia; Finland; Sweden; Norway; Britain; Ireland; Denmark; all central Europe to southern France; Italy; Istria; Bosnia; Albania; Romania; Hungary; Czechoslovakia; Poland; Ukraine; Germany; Austria).

  • — Kasule, 1968a: 62 (Staphylinus; life history).

  • — Szujecki, 1968a: 732 (Ocypus; Poland).

  • — Kasule, 1970: 70 (Staphylinus; larval characters).

  • — Coiffait, 1974: 459 (Ocypus; characters; Europe; Britain; Caucasus; Asia Minor).

  • — Pope, 1977: 30 (Staphylinus; Britain).

  • — Hinton, 1981: 662 (Ocypus; characters of egg).

  • — Tóth, 1984: 25 (Ocypus; characters; Hungary).

  • — Newton, 1987: 382 (Staphylinus; introduced species; New Hampshire).

  • — Lucht, 1987: 108 (Ocypus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Pilon, 1991: 104 (Ocypus; notes; Italy).

  • — Pilon and Zanetti, 1991: 68 (Ocypus; Italy).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Ocypus; Czech Republic; Slovakia).

  • — K. Bland, 1997: 204 (Staphylinus; eaten by toad; Scotland).

  • — Pilon, 1995: 32, 37 (Ocypus; characters; Italy).

  • — Pilon, 1998: 78 (Ocypus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Europe, Russia, Caucasus, Turkey, USA.

erythropus Paykull, 1800: 372 (Staphylinus; [preoccupied]; [Note: This name is also cited as a synonym of Platydracus fulvipes by Bernhauer and Schubert, 1914: 379]; Type locality: Kinnakulle).

  • — Germar, 1823: 14 (Staphylinus; characters; Sweden; Germany).

  • — Curtis, 1829: 24 (Ocypus; catalog; Britain).

  • — C. Sahlberg, 1830: 310 (Staphylinus; characters; Finland).

  • — Stephens, 1839: 388 (Goerius; characters; England).

  • — Kraatz, 1857d: 558 (Ocypus; synonym of brunnipes).

  • — Ganglbauer, 1895: 433 (Staphylinus; synonym of brunnipes).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of brunnipes).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; synonym of fulvipes).

  • — Smetana, 1958: 290 (Ocypus; synonym of brunnipes).

  • — Coiffait, 1974: 459 (Ocypus; synonym of brunnipes).

cantianus Stephens, 1832: 207 (Staphylinus; Type locality: London, about Gravesend).

  • — Stephens, 1839: 387 (Staphylinus; characters; England).

  • — Hardy, 1851: 23 (Ocypus; synonym of fuscatus).

  • — G. Waterhouse, 1858: 24 (Ocypus; synonym of brunnipes and fuscatus).

  • — Gemminger and Harold, 1868: 581 (Ocypus; synonym of brunnipes).

  • — Ganglbauer, 1895: 433 (Staphylinus; synonym of brunnipes).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of brunnipes).

  • — Smetana, 1958: 290 (Ocypus; synonym of brunnipes).

  • — Coiffait, 1974: 459 (Ocypus; synonym of brunnipes).

alpicola Erichson, 1840: 411 (Ocypus; Type locality: Carinthiae Carnioliaeque alpes).

  • — Heer, 1841: 582 (Staphylinus; characters; Switzerland).

  • — Küster, 1853: 59 (Ocypus; characters; Austria).

  • — Kraatz, 1857d: 558 (Ocypus; synonym of brunnipes).

  • — Fauvel, 1865a: 295 [= 1865b: 45] (Ocypus; habitat; France).

  • — Gemminger and Harold, 1868: 581 (Ocypus; synonym of brunnipes).

  • — Ganglbauer, 1895: 433 (Staphylinus; synonym of brunnipes).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of brunnipes).

  • — G. Müller, 1926: 17 (Staphylinus; subgenus Pseudocypus; valid species).

  • — Scheerpeltz, 1933: 1392 (Staphylinus; subspecies of brunnipes).

  • — Coiffait, 1956e: 204 (Ocypus; subgenus Goerius; subspecies of brunnipes; characters).

  • — Smetana, 1958: 291 (Ocypus; synonym of brunnipes).

  • — Horion, 1965: 213 (Staphylinus; subgenus Goerius; subspecies of brunnipes; Austria; Slovenia; Italy; Switzerland).

  • — Coiffait, 1974: 459 (Ocypus; subspecies of brunnipes; Italy).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; subspecies of brunnipes; Italy).

intermedius G. Müller, 1911: 58 (Staphylinus; subspecies of brunnipes; [preoccupied]; Type locality: Woltschah, im oberen Isonzotale).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; variety of brunnipes).

  • — G. Müller, 1923: 141 (Staphylinus; subgenus Pseudocypus; synonym of brunnipes alpicola).

  • — G. Müller, 1926: 17 (Staphylinus; subgenus Pseudocypus; subspecies of alpicola).

  • — Scheerpeltz, 1933: 1392 (Staphylinus; synonym of brunnipes).

  • — Smetana, 1958: 291 (Ocypus; synonym of brunnipes).

  • — Coiffait, 1974: 459 (Ocypus; synonym of brunnipes).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus; subspecies of brunnipes).

bulgaricus Coiffait, 1970, see: picipennis Fabricius, 1793.

bureschi Coiffait, 1971b: 280 (Ocypus; Type locality: Kuru Dag, Keschan, Turquie d'Europe).

  • — Coiffait, 1974: 475 (Ocypus; characters; Turkey).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: Turkey.

caeruleus Cameron, 1932: 203 (Staphylinus; subgenus Goerius; Type locality: Darjeeling: Gopaldhara).

  • — Scheerpeltz, 1933: 1392 (Staphylinus; catalog).

  • — Coiffait, 1982a: 233 (Ocypus, cited as coeruleus; India).

  • — Coiffait, 1984e: 374 (Ocypus; cited as coeruleus; Nepal).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: India, Nepal.

caeruleipennis Coiffait, 1954 [Note: Nomen nudum], see: subaenescens Wollaston, 1964.

caerulescens Coiffait, 1982: 76 (Nudabemus; Type locality: Nepal: östl. Dhaulagiri, Thankur, nördl. Dhorpatan, 3350 m).

  • — Smetana and Davies, 2000: 31, 42 (Ocypus; subgenus Pseudocypus; [Note: Cited erroneously in subgenus Ocypus]; notes).

  • Distribution: Nepal.

cameroni Smetana and Davies, 2000: 33, 42 (Ocypus; subgenus Ocypus; replacement name for bimaculatus).

  • Distribution: India.

bimaculatus Cameron, 1932: 207 (Staphylinus; subgenus Tasgius; [preoccupied]; Type locality: northern India).

  • — Scheerpeltz, 1933: 1391 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 33, 42 (Ocypus; synonym of cameroni).

canariensis Gemminger and Harold, 1868: 581 (Ocypus; replacement name for curtipennis Wollaston).

  • — Wollaston, 1871: 300 (Ocypus; notes).

  • — Fauvel, 1902b: 104 (Staphylinus; synonym of curtipennis).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1392 (Staphylinus; catalog).

  • — Jarrige, 1954a: 162 (Staphylinus; subgenus Goerius; characters).

  • — Coiffait, 1974: 412 (Atlantogoerius; characters; Canary Islands).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Pseudocypus).

  • Distribution: Canary Islands.

curtipennis Wollaston, 1864: 567 (Ocypus; [preoccupied]; Type locality: Grand Canary: El Monte; El Dorames on the mountains between Osorio and Guia).

  • — Wollaston, 1865: 488 (Ocypus; notes; Canary Islands).

  • — Fauvel, 1902b: 104 (Staphylinus; subgenus Ocypus; Canary Islands).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of canariensis).

  • — Coiffait, 1954: 166 (Goerius; cited as infraspecific rank of canariensis; Canary Islands).

cantabricus G. Müller, 1926, see: pedemontanus G. Müller, 1924.

cantianus Stephens, 1832, see: brunnipes Fabricius, 1781.

caroli Jarrige, 1943: 146 (Staphylinus; subgenus Goerius; Type locality: Monte, Madère, alt. 600 m).

  • — Jarrige, 1954a: 161 (Staphylinus; subgenus Goerius; characters; Madeira).

  • — Coiffait, 1974: 481 (Ocypus; characters).

  • — Israelson, 1981: 2 (Ocypus; characters; Madeira).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Madeira.

carinatus Sharp, 1874, see: Agelosus.

caucasicus G. Müller, 1926, see: picipennis Fabricius, 1793.

cerceticus Coiffait, 1964: 93 (Ocypus; Type locality: Russie d'Europe environs de Maykop, Nord Caucase).

  • — Smetana, 1965c: 40, 41 (Ocypus; cited as quadraticeps; [Note: Misidentification of cerceticus according to Solodovnikov, 2000]; aedeagal illustration; NW Caucasus).

  • — Pototskaia, 1967: 73 (Staphylinus; subgenus Ocypus; larval characters).

  • — Coiffait, 1974: 474 (Ocypus; characters; Caucasus).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • — Solodovnikov, 2000: 315 (Ocypus; characters; Russia; Georgia; Turkmenistan).

  • Distribution: Russia, Georgia; Turkmenistan.

cerdanicus Coiffait, 1956, see: picipennis Fabricius, 1793.

cerdo Erichson, 1840, see: Tasgius.

chevrolatii Baudi, 1848: 121 (Ocypus; Type locality: Not cited).

  • — Fauvel, 1865a: 294 [= 1865b: 44] (Ocypus; synonym of alpestris).

  • — Ganglbauer, 1895: 431 (Staphylinus; synonym of macrocephalus).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; synonym of macrocephalus).

  • — G. Müller, 1926: 12 (Staphylinus; subgenus Goerius; characters; notes).

  • — G. Müller, 1926c: 75 (Staphylinus; characters; Alps).

  • — G. Müller, 1932: 78 (Staphylinus; subgenus Goerius; notes).

  • — Scheerpeltz, 1933: 1392 (Staphylinus; catalog).

  • — Jarrige, 1947a: 155 (Goerius; characters; French Alps).

  • — Coiffait, 1956e: 204, 206 (Ocypus; subgenus Goerius; characters).

  • — Lohse, 1964: 198 (Ocypus; subgenus Goerius; characters; central Europe).

  • — Coiffait, 1964: 87 (Ocypus; characters; France; Italy).

  • — Korge, 1965: 71 (Ocypus; characters).

  • — Coiffait, 1974: 464 (Ocypus; characters; Alps).

  • — Löser, 1979: 20 (Ocypus; cited as chevrolathi; distribution in central Alps).

  • — Lucht, 1987: 108 (Ocypus; cited as chevrolati; checklist; Germany; Austria; France).

  • — Lohse and Lucht, 1989: 169 (Ocypus; cited as chevrolati; Germany).

  • — Wittwer, 1993: 250 (Ocypus; Switzerland).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; Italy).

  • — Pilon, 1995: 32, 36 (Ocypus; cited as chevrolati; characters; Italy).

  • — Pilon, 1998: 79 (Ocypus; cited as chevrolati; ecological notes; Italy).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Germany, France, Italy, Switzerland.

alsaticus Coiffait, 1964: 87 (Ocypus; subspecies of chevrolatii; Type locality: Bas-Rhin: Forêt de Haguenau).

  • — Korge, 1965: 71 (Ocypus; subspecies of chevrolatii; characters).

  • — Coiffait, 1974: 464 (Ocypus; subspecies of chevrolatii).

  • — Smetana and Davies, 2000: 43 (Ocypus; synonym of chevrolatii).

pseudobrevipennis Coiffait, 1964: 87 (Ocypus; subspecies of chevrolatii; Type locality: Haute-Savoie: Les Contamines, Cugnon, 1400 m).

  • — Korge, 1965: 71 (Ocypus; subspecies of chevrolatii; characters).

  • — Korge, 1968: 53 (Ocypus; subpspecies of chevrolatii; characters; notes).

  • — Coiffait, 1974: 464 (Ocypus; subspecies of chevrolatii).

  • — Smetana and Davies, 2000: 43 (Ocypus; synonym of chevrolatii).

vesubiensis Coiffait, 1964: 89 (Ocypus; subspecies of chevrolatii; Type locality: Alpes-Maritimes: Saint-Martin-Vésubie, vers 2300 m).

  • — Korge, 1965: 71 (Ocypus; subspecies of chevrolatii; characters).

  • — Coiffait, 1974: 464 (Ocypus; subspecies of chevrolatii).

  • — Smetana and Davies, 2000: 43 (Ocypus; synonym of chevrolatii).

zimmermanni Korge, 1965: 70 (Ocypus; subspecies of chevrolatii; Type locality: Engelberg, Obwalden, Helvetia, 1700–1800 m).

  • — Coiffait, 1974: 464 (Ocypus; subspecies of chevrolatii).

  • — Smetana and Davies, 2000: 43 (Ocypus; synonym of chevrolatii).

coerulescens Geoffroy, 1785, see: ophthalmicus Scopoli, 1763.

confinis Stephens, 1832, see: sericeicollis Ménétriés, 1832.

confusus Baudi, 1848, see: fulvipennis Erichson, 1840.

congruus Walker, 1858, see: Phucobius.

coreanus G. Müller, 1925: 42 (Staphylinus; subgenus Goerius; Type locality: Korea: Pu-Ryong).

  • — Scheerpeltz, 1933: 1392 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Korea.

crassicollis Gravenhorst, 1802, see: fuscatus Gravenhorst, 1802.

cubae Jacquelin du Val, 1857, see: Platydracus.

cupreus Rossi, 1790, see: sericeicollis Ménétriés, 1832.

curtipennis Motschulsky, 1849: 87 (Ocypus; Type locality: Mt. Tauria; [Note: See lectotype designation by Gusarov, 1991a]. Originally cited from montagnes de la Tauride et du Caucase).

  • — Fauvel, 1874: 183 [= 1874a: 409] (Staphylinus; synonym of olens).

  • — Fiori, 1894: 90 (Ocypus; variety of olens; characters; Italy).

  • — Ganglbauer, 1895: 430 (Staphylinus; synonym of tenebricosus).

  • — Bernhauer and Schubert, 1914: 391 (Staphylinus; synonym of tenebricosus).

  • — Smetana, 1958: 284 (Ocypus; synonym of tenebricosus).

  • — Gusarov, 1991a: 9 (Ocypus; valid species; notes; lectotype designation: Mt. Tauria).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus).

  • Distribution: Ukraina, Italy, Macedonia, Bulgaria, Turkey, Cyprus, Syria, Lebanon, Iran, Caucasus.

integer Abeille, 1900: 203 (Ocypus; Type locality: Smyrne).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; catalog).

  • — Coiffait, 1970e: 62 (Ocypus; Lesbos).

  • — Coiffait, 1971b: 280 (Ocypus; Bulgaria).

  • — Coiffait, 1974: 450 (Ocypus; characters; Crimea; Bulgaria; Black Sea; Asia Minor; Macedonia).

  • — Gusarov, 1991a: 9 (Ocypus; synonym of curtipennis).

  • — Smetana and Davies, 2000: 42 (Ocypus; synonym of curtipennis).

gridellii G. Müller, 1924: 19, 24 (Staphylinus; subgenus Goerius; Type locality: Crimea; Vodena in Macedonia).

  • — G. Müller, 1926: 7 (Staphylinus; subgenus Goerius; characters; notes).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; catalog).

  • — G. Müller, 1950: 8 (Staphylinus; subgenus Goerius; characters; notes).

  • — Coiffait, 1956e: 201 (Ocypus; subgenus Goerius; characters; Macedonia; Crimea; Asia Minor; Cyprus; Lebanon).

  • — Scheerpeltz, 1956c: 1101 (Staphylinus; subgenus Goerius; Cyprus).

  • — Scheerpeltz, 1961e: 137 (Staphylinus; subgenus Goerius; Rhodes).

  • — Coiffait, 1964: 83 (Ocypus; notes).

  • — Smetana, 1965c: 28 (Ocypus; subgenus Ocypus; Anatolia).

  • — Pototskaia, 1966b: 357 (Staphylinus; subgenus Ocypus; larval characters).

  • — Pototskaia, 1967: 71 (Staphylinus; subgenus Ocypus; larval characters).

  • — Smetana, 1968b: 88 (Ocypus; Anatolia).

  • — Smetana, 1971c: 264 (Ocypus; subgenus Ocypus; notes; Iran).

  • — Coiffait, 1974: 450 (Ocypus; synonym of integer).

  • — Gusarov, 1991a: 9 (Ocypus; synonym of curtipennis).

  • — Smetana and Davies, 2000: 42 (Ocypus; synonym of curtipennis).

libanoticus G. Müller, 1950: 8 (Staphylinus; subgenus Goerius; subspecies of gridellii; Type locality: Libano meridionale, presso Saida, m. 900).

  • — Coiffait, 1964: 83 (Ocypus; subspecies of gridellii; characters).

  • — Coiffait, 1974: 450 (Ocypus; subspecies of integer; Lebanon; Syria).

  • — Bordoni, 1984: 341 (Ocypus; subspecies of integer; Lebanon).

  • — Smetana and Davies, 2000: 42 (Ocypus; synonym of curtipennis).

kyproticus Coiffait, 1964: 85 (Ocypus; subspecies of gridellii; Type locality: Chypre).

  • — Smetana, 1971c: 264 (Ocypus; synonym of gridellii).

  • — Coiffait, 1974: 450 (Ocypus; subspecies of integer; Cyprus).

  • — Smetana and Davies, 2000: 42 (Ocypus; synonym of curtipennis).

pseudolens Coiffait, 1964: 85 (Ocypus; subspecies of gridellii; Type locality: Iran).

  • — Smetana, 1971c: 264 (Ocypus; synonym of gridellii).

  • — Coiffait, 1974: 450 (Ocypus; subspecies of integer; Iran).

  • — Smetana and Davies, 2000: 42 (Ocypus; synonym of curtipennis).

curtipennis Wollaston, 1864, see: canariensis Gemminger and Harold, 1868.

cyaneopubens Reitter, 1913: 65 (Goerius; Type locality: Syrien: Umgebung von Rahobot bei Jaffa).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; subgenus Ocypus; catalog).

  • — Dodero, 1925: 5 (Staphylinus; subgenus Goerius; Libya).

  • — Gridelli, 1930: 75 (Staphylinus; Libya).

  • — Scheerpeltz, 1933: 1393 (Staphylinus; catalog).

  • — Peyerimhoff, 1937: 267 (Staphylinus; subgenus Goerius; Tunisia).

  • — Coiffait, 1974: 470 (Ocypus; characters; Palestine; Cyrenaica).

  • — Coiffait, 1974: 471 (Ocypus; characters; Egypt).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus).

  • Distribution: Israel, Egypt, Libya, Tunisia.

cyanescens Gerhardt, 1910 [Note: Nomen nudum], see: aeneocephalus De Geer, 1774.

cyaneus Paykull, 1789, see: ophthalmicus Scopoli, 1763.

cyanochloris Hochhuth, 1849, see: ophthalmicus Scopoli, 1763.

cyclopus Peyron, 1858: 422 (Ocypus; Type locality: Not cited, presumably Tarsous).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1393 (Staphylinus; catalog).

  • — Coiffait, 1974: 481 (Ocypus; characters).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: Turkey.

decurtatus Mulsant and Rey, 1876, see: nitens Schrank, 1781.

densissimus Bernhauer, 1933: 35 (Staphylinus; subgenus Pseudocypus; Type locality: Kiang-Si: T'en-gan).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Pseudocypus).

  • Distribution: China.

depolii G. Müller, 1924, see: tenebricosus Gravenhorst, 1846.

depressus Hochhuth, 1849, see: Tasgius.

deuvei Coiffait, 1978d: 167 (Ocypus; Type locality: Col de Ciftecoprüler, 1000 m).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Turkey.

dorsalis Sharp, 1889: 109 (Ocypus; Type locality: Niohozan; Chiuzenji).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; subgenus Ocypus; catalog).

  • — G. Müller, 1924: 26 (Staphylinus; notes).

  • — Scheerpeltz, 1933: 1393 (Staphylinus; catalog; China).

  • — Cameron, 1949b: 469 (Ocypus; Japan).

  • — Nakane, 1963a: 93 (Ocypus; characters; habitus photograph; Japan).

  • — Yuh, Paik, Kwon, and Lee, 1985: 245 (Ocypus; Korea).

  • — Naomi, 1992b: 2 (Ocypus; subgenus Protocypus; characters; Japan).

  • — J. Li, 1993: 38 (Ocypus; China).

  • — Watanabe, 1996a: 12 (Ocypus; subgenus Xanthocypus; cave collection; Japan).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan, Korea, China.

duplicatus G. Müller, 1943: 102 (Staphylinus; subgenus Goerius; subspecies of alpestris; Type locality: Altai centr.).

  • — Coiffait, 1974: 463 (Ocypus; valid species; characters; central Asia).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: “Altai region”.

ensifer G. Müller, 1932, see: gracilicornis Hochhuth, 1849.

eppelsheimi Reitter, 1887: 214 (Ocypus; Type locality: Flussthal Bytchu, 13,000 F).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; subgenus Ocypus; catalog).

  • — G. Müller, 1926a: 31 (Staphylinus; subgenus Pseudocypus; notes).

  • — Scheerpeltz, 1933: 1393 (Staphylinus; catalog).

  • — Coiffait, 1974: 513 (Pseudocypus; characters; central Asia; Siberia).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Pseudocypus).

  • Distribution: China, Russia.

erosicollis Reiche and Saulcy, 1856, see: Tasgius.

erythropus Paykull, 1800, see: brunnipes Fabricius, 1781.

etruscus Gautier, 1860, see: Tasgius.

evitendus Tottenham, 1945, see: Tasgius.

excisus G. Müller, 1950: 11 (Staphylinus; subgenus Pseudocypus; subspecies of picipennis; Type locality: Centr. Mesopotamien, r. Djirdjibufer-Steppe).

  • — Coiffait, 1964: 99 (Pseudocypus; valid species; elytral color).

  • — Coiffait, 1974: 510 (Pseudocypus; characters; Lebanon; Syria; Mesopotamia).

  • — Bordoni, 1984: 341 (Pseudocypus; Lebanon).

  • — M. Dvořák, 1984: 66 (Pseudocypus; Syria).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Pseudocypus).

  • Distribution: Lebanon, Syria, Iraq.

libanicus Coiffait, 1956e: 211 (Ocypus; subgenus Pseudocypus; subspecies of picipennis; Type locality: Col de Dahr el Baïdar, 1500 m).

  • — Coiffait, 1964: 99 (Pseudocypus; synonym of excisus).

  • — Coiffait, 1974: 510 (Pseudocypus; synonym of excisus).

  • — Smetana and Davies, 2000: 44 (Ocypus; synonym of excisus).

fallaciosus G. Müller, 1926, see: picipennis Fabricius, 1793.

festae G. Müller, 1925a: 25, 26 (Staphylinus; subgenus Goerius; Type locality: Cirenaica).

  • — G. Müller, 1926: 10 (Staphylinus; subgenus Goerius; characters; notes).

  • — Gridelli, 1930: 75 (Staphylinus; Libya).

  • — Scheerpeltz, 1933: 1394 (Staphylinus; catalog).

  • — Coiffait, 1974: 450 (Ocypus; characters; Cyrenaica).

  • — M. Dvořák, 1984: 60 (Ocypus; characters; Libya).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus).

  • Distribution: Libya.

forficularius Motschulsky, 1860: 569 (Matidus; Type locality: Sagerdan; [Note: See lectotype designation by Gusarov, 1993]. Originally cited from environs de Tiflis à Sagerdan au pieds des Montagnes du Caucase).

  • — Hochhuth, 1862: 50 (Ocypus; characters; Caucasus).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; catalog).

  • — G. Müller, 1926b: 42 (Staphylinus; subgenus Goerius; characters; notes).

  • — Scheerpeltz, 1933: 1394 (Staphylinus; catalog).

  • — Coiffait, 1974: 468 (Ocypus; cited as forficularis; characters; Caucasus).

  • — Boháč, 1988: 553 [= 1989: 36] (Ocypus; characters; synonym of gracilicornis).

  • — Gusarov, 1993: 71 (Ocypus; valid species; synonyms; lectotype designation: Sagerdan; Turkey; Georgia).

  • — Solodovnikov, 1998a: 14 (Ocypus; characters; notes; Russia, Krasnodar territory).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Georgia, Turkey.

korgei Smetana, 1965c: 34 (Ocypus; subgenus Ocypus; Type locality: Anatolia bor., Soganli-Pass bei Bayburt, 1400 m).

  • — Smetana, 1965d: 80 (Ocypus; subgenus Ocypus; Anatolia).

  • — Smetana, 1968a: 156 (Ocypus; subgenus Ocypus; Anatolia).

  • — Coiffait, 1974: 463 (Ocypus; characters; Turkey).

  • — Boháč, 1980: 86 (Ocypus; Caucasus).

  • — Gusarov, 1993: 71 (Ocypus; synonym of forficularius).

  • — Solodovnikov, 1998a: 14 (Ocypus; synonym of forficularius).

  • — Smetana and Davies, 2000: 43 (Ocypus; synonym of forficularius).

fortunatarum Wollaston, 1871: 302 (Ocypus; replacement name for punctatissimus Wollaston).

  • — Fauvel, 1874: 191 [= 1874a: 417] (Staphylinus; synonym of aeneocephalus).

  • — Ganglbauer, 1895: 434 (Staphylinus; synonym of aeneocephalus).

  • — Fauvel, 1902b: 105 (Staphylinus; synonym of aeneocephalus).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of cupreus).

  • — Jarrige, 1954a: 163 (Staphylinus; subgenus Pseudocypus; characters; Canaries; Madeira; Barbary; Iberia; France).

  • — Coiffait, 1956e: 214, 215 (Ocypus; subgenus Pseudocypus; characters).

  • — Coiffait, 1974: 514 (Pseudocypus; subgenus Fortunocypus; characters; Canary Islands; Madeira; North Africa; Iberian Peninsula; French Mediterranean region; Sardinia; Sicily; Italy).

  • — Pope, 1977: 30 (Staphylinus; Britain).

  • — Gusarov, 1993: 72 (Ocypus; subgenus Pseudocypus; synonyms).

  • — Outerelo, Palmer and Pons, 1995: 79 (Pseudocypus; Mallorca).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; Italy).

  • — Pilon, 1998: 89 (Ocypus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Tunisia, Algeria, Morocco, Canary Islands, Madeira, Spain, France, Italy, Britain.

sericeus Motschulsky, 1858: 670 (Ocypus; [preoccupied]; Type locality: Algérie; [Note: See lectotype designation by Gusarov, 1993]).

  • — Fauvel, 1869: 488 (Ocypus; synonym of cupreus).

  • — Fauvel, 1874: 191 [= 1874a: 417] (Staphylinus; synonym of aeneocephalus).

  • — Ganglbauer, 1895: 434 (Staphylinus; synonym of aeneocephalus).

  • — Fauvel, 1902b: 105 (Staphylinus; synonym of aeneocephalus).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of cupreus).

  • — G. Müller, 1923: 141 (Staphylinus; subgenus Pseudocypus; characters; Italy).

  • — G. Müller, 1926: 19 (Staphylinus; subgenus Pseudocypus; notes; Sardinia, Sicily, Italy, Tunisia, Algeria, Morocco, Portugal).

  • — Sainte-Claire Deville, 1926: 120 (Staphylinus; notes; Corsica).

  • — Scheerpeltz, 1933: 1401 (Staphylinus; catalog).

  • — Méquignon, 1933: 229 (Staphylinus; distinguishing characters).

  • — Scheerpeltz, 1961d: 131 (Staphylinus; subgenus Pseudocypus; Italy).

  • — Coiffait, 1974: 514 (Pseudocypus; synonym of fortunatarum).

  • — Gusarov, 1993: 72 (Ocypus; lectotype designation; synonym of fortunatarum).

punctatissimus Wollaston, 1864: 568 (Ocypus; [Note: Preoccupied as a secondary homonym by punctatissimus Dufour which was described in Staphylinus but was listed in Ocypus by Gemminger and Harold, 1868: 581, at the time of replacement]; Type locality: Lanzarote; Fuerteventura).

  • — Wollaston, 1865: 489 (Ocypus; habitat; Canary Islands).

  • — Wollaston, 1867: 278 (Ocypus; notes; may be synonym of cupreus).

  • — Fauvel, 1869: 488 (Ocypus; synonym of cupreus).

  • — Fauvel, 1869a: 151 (Ocypus; synonym of cupreus).

  • — Wollaston, 1871: 301 (Ocypus; characters; notes; valid species).

  • — Ganglbauer, 1895: 434 (Staphylinus; synonym of aeneocephalus).

  • — Fauvel, 1902b: 105 (Staphylinus; synonym of aeneocephalus).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of cupreus).

  • — Coiffait, 1974: 514 (Pseudocypus; synonym of fortunatarum).

  • — Gusarov, 1993: 72 (Ocypus; synonym of fortunatarum).

frater Smetana, 1965c: 31 (Ocypus; subgenus Ocypus; Type locality: Anatolia bor.: Abant-Geb., 1100–1500 m).

  • — Smetana, 1965d: 80 (Ocypus; subgenus Ocypus; Anatolia).

  • — Smetana, 1968b: 88 (Ocypus; Anatolia).

  • — Coiffait, 1974: 456 (Ocypus; subspecies of heinzi; characters).

  • — Coiffait, 1978d: 164 (Ocypus; subspecies of heinzi; Turkey).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus; cited as valid species).

  • Distribution: Turkey.

fraternus Fairmaire, 1891, see: Agelosus.

fujiensis Naomi, 1992b: 6 (Ocypus; subgenus Protocypus; Type locality: Japan: Yamanashi Pref.: Aokigahara, Mt. Fuji).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan.

fulvicupreus Coiffait, 1956, see: sericeicollis Ménétriés, 1832.

fulvipennis Erichson, 1840: 413 (Ocypus; Type locality: Austria).

  • — Nordmann, 1837: 70 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 386 cited chalcocephalus Nordmann as a synonym of fulvipennis Erichson; Nordmann did not describe the species as new, he attributed it to Fabricius]; Sweden; France; Germany).

  • — Heer, 1841: 582 (Staphylinus; synonym of vagans).

  • — Hochhuth, 1849: 124 (Ocypus; characters; Iberia; Lenkoran).

  • — Redtenbacher, 1849: 699 (Ocypus; characters; Austria).

  • — Küster, 1853: 63 (Ocypus; characters; Europe).

  • — Fairmaire and Laboulbène, 1856: 511 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 561 (Ocypus; characters; Germany).

  • — Redtenbacher, 1857: 188 (Ocypus; characters; Austria).

  • — Hochhuth, 1862: 49 (Ocypus; Russia).

  • — Fauvel, 1874: 189 [= 1874a: 415] (Staphylinus; characters; France; Scandinavia; Britain; Germany; Austria; Italy; Corsica; Sardinia; Greece; Algeria).

  • — Redtenbacher, 1874: 207 (Ocypus; characters; Austria).

  • — Mulsant and Rey, 1876: 314 [= 1877: 170] (Ocypus; subgenus Pseudocypus; characters; France).

  • — J. Sahlberg, 1876: 11 (Ocypus; characters; Finland).

  • — Ganglbauer, 1895: 434 (Staphylinus; subgenus Pseudocypus; characters; north and middle Europe; Siberia).

  • — Montandon, 1908: 80 (Staphylinus; Romania).

  • — Reitter, 1909: 120 (Staphylinus; subgenus Pseudocypus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; subgenus Ocypus; catalog).

  • — Portevin, 1929: 351 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1394 (Staphylinus; catalog).

  • — Paulian, 1941: 244 (Staphylinus; subgenus Pseudocypus; larval characters).

  • — Coiffait, 1956e: 209, 215 (Ocypus; subgenus Pseudocypus; characters).

  • — Smetana, 1958: 294 (Ocypus; subgenus Pseudocypus; characters; ecology; Czechoslovakia).

  • — Szujecki, 1963a: 34 (Ocypus; Poland).

  • — Palm, 1963b: 117 (Staphylinus; subgenus Pseudocypus; characters; Sweden; Finland).

  • — Lohse, 1964: 200 (Ocypus; subgenus Pseudocypus; characters; central Europe).

  • — Smetana, 1965c: 45 (Ocypus; subgenus Pseudocypus; Anatolia).

  • — Horion, 1965: 222 (Staphylinus; subgenus Pseudocypus; Russia; Finland; Poland; Czechoslovakia; Hungary; Romania; Bosnia Hercegovina; Bulgaria; Istria; Italy; France; Switzerland; Belgium; Holland; Austria; Germany).

  • — Jarrige, 1968b: 74 (Pseudocypus; France).

  • — Scheerpeltz, 1968a: 83 (Staphylinus; catalog; Austria).

  • — Tikhomirova, 1973a: 186 (Ocypus; subgenus Pseudocypus; checklist; Soviet Union).

  • — Coiffait, 1974: 506 (Pseudocypus; characters; France; Scandinavia; central Europe; Balkans; Greece; Anatolia; Caucasus; central Asia; Kazakhstan).

  • — Raichev, 1979: 83 (Pseudocypus; Bulgaria).

  • — Muona, 1979: 15 (Ocypus; Finland; Russia; Sweden).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 107 (Ocypus; catalog; Poland).

  • — Chemini and Zanetti, 1982: 216 (Ocypus; habitat notes; Italy).

  • — Tóth, 1984: 29 (Ocypus; characters; Hungary).

  • — Tóth, 1985a: 92 (Ocypus; subgenus Pseudocypus; Hungary).

  • — Ádám, 1987: 148 (Pseudocypus; collecting notes; Hungary).

  • — Lucht, 1987: 108 (Ocypus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Sweden).

  • — Gusarov, 1989: 11 (Ocypus; habitat notes; Ukraine).

  • — Silfverberg, 1992: 17 (Ocypus; Finland; Russia; Sweden; Estonia; Latvia; Lithuania).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Pseudocypus; subspecies fulvipennis; Czech Republic; Slovakia).

  • — Kocian, 1993b: 95 (Ocypus; Czech Republic).

  • — Assing, 1994a: 12, 24 (Ocypus; notes; wing length; xerothermous sites; Germany).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; Italy).

  • — Šustek, 1995: 398 (Pseudocypus; Czech Republic).

  • — Eisinger, 1997: 171 (Ocypus; collecting notes; Germany).

  • — Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 38 (Ocypus; Latvia).

  • — Pilon, 1998: 89 (Ocypus; notes; Italy).

  • — Solodovnikov, 1998: 338 (Ocypus; bionomic notes; distribution in northwest Caucasus; Russia).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • — Assing, 2001: 75 (Ocypus; collected from xerothermous site; Germany).

  • — Herman, 2001: 27 (Ocypus; nomen protectum).

  • Distribution: Europe, Russia, Georgia, Azerbaijan, Turkey, Kazakhstan.

picipennis Lacordaire, 1835: 374 (Emus; [preoccupied]; Type locality: Paris).

  • — Ganglbauer, 1895: 434 (Staphylinus; synonym of fulvipennis).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; synonym of fulvipennis).

  • — Smetana, 1958: 294 (Ocypus; synonym of fulvipennis).

  • — Coiffait, 1974: 506 (Pseudocypus; synonym of fulvipennis).

  • — Herman, 2001: 27 (Ocypus; nomen oblitum).

vagans Heer, 1839: 255 (Staphylinus; [Note: This name is older than fulvipennis]; Type locality: Bern; Basel; Neuchatel; Genf).

  • — Hochhuth, 1849: 124 (Ocypus; synonym of fulvipennis).

  • — Kraatz, 1857d: 561 (Ocypus; synonym of fulvipennis).

  • — J. Sahlberg, 1876: 12 (Ocypus; synonym of fulvipennis).

  • — Ganglbauer, 1895: 434 (Staphylinus; synonym of fulvipennis).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; synonym of fulvipennis).

  • — Smetana, 1958: 294 (Ocypus; synonym of fulvipennis).

  • — Coiffait, 1974: 506 (Pseudocypus; synonym of fulvipennis).

  • — Herman, 2001: 27 (Ocypus; nomen oblitum).

ibericus Kolenati, 1846: 17 (Staphylinus; Type locality: Iberia ad urbem Tiflis).

  • — Hochhuth, 1849: 124 (Ocypus; synonym of fulvipennis).

  • — Fauvel, 1874: 189 [= 1874a: 415] (Staphylinus; synonym of fulvipennis).

  • — Ganglbauer, 1895: 434 (Staphylinus; synonym of fulvipennis).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; synonym of fulvipennis).

  • — Smetana, 1958: 294 (Ocypus; synonym of fulvipennis).

  • — Coiffait, 1974: 506 (Pseudocypus; synonym of fulvipennis).

confusus Baudi, 1848: 122 (Ocypus; Type locality: Pedemontio).

  • — Fairmaire and Laboulbène, 1856: 511 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 560, note (Ocypus; characters).

  • — Fauvel, 1874: 189 [= 1874a: 415] (Staphylinus; synonym of fulvipennis).

  • — Mulsant and Rey, 1876: 311 [= 1877: 167] (Ocypus; subgenus Pseudocypus; characters; France).

  • — Ganglbauer, 1895: 434 (Staphylinus; synonym of fulvipennis).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; synonym of fulvipennis).

  • — Scheerpeltz, 1933: 1394 (Staphylinus; subspecies of fulvipennis).

  • — Coiffait, 1956e: 209 (Ocypus; subgenus Pseudocypus; subspecies of confusus; characters).

  • — Smetana, 1958: 295 (Ocypus; synonym of fulvipennis).

  • — Jarrige, 1968b: 74 (Pseudocypus; subspecies of fulvipennis; France).

  • — Coiffait, 1974: 506 (Pseudocypus; variety of fulvipennis; Alps).

  • — Pilon and Zanetti, 1991: 68 (Ocypus; subspecies of fulvipennis; Italy).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Pseudocypus; subspecies of fulvipennis; Slovakia ?).

philonthoides Hochhuth, 1849: 125 (Ocypus; Type locality: Tiflis).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; synonym of fulvipennis).

  • — Smetana, 1958: 294 (Ocypus; synonym of fulvipennis).

  • — Coiffait, 1974: 506 (Pseudocypus; synonym of fulvipennis).

fulvipes Ménétriés, 1849, see: Tasgius.

fulvopilosus Fiori, 1894, see: olens O. Müller, 1764.

fulvotomentosus Eppelsheim, 1889a: 172 (Ocypus; subgenus Goerius; Type locality: Kan-ssu; [Note: See lectotype designation by Smetana and Davies, 2000: 33]).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; subgenus Ocypus; catalog).

  • — G. Müller, 1923: 196 (Staphylinus; subgenus Protocypus).

  • — Scheerpeltz, 1933: 1393 (Staphylinus; synonym of dorsalis).

  • — G. Müller, 1943: 97 (Staphylinus; subgenus Protocypus; [Note: I included this name in Ocypus because it is the type species of Protocypus which is listed as a subgenus of Ocypus by Naomi, 1992a: 54]; valid species; characters).

  • — Smetana and Davies, 2000: 33, 44 (Ocypus; subgenus Pseudocypus; lectotype designation).

  • Distribution: China.

hauseri Bernhauer, 1933: 32 (Staphylinus; Type locality: Kalgan, Inn-shan, 980–2100 m).

  • — Smetana and Davies, 2000: 33, 44 (Ocypus; synonym of fulvotomentosus).

fuscatoides Coiffait, 1964: 97 (Pseudocypus; Type locality: Mongolie: Selenga, au sud du Lac Baikal).

  • — Coiffait, 1974: 503 (Pseudocypus; characters; Mongolia).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Mongolia.

fuscatus Gravenhorst, 1802: 164 (Staphylinus; Type locality: Not cited).

  • — De Geer, 1774: 22 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 386, cited politus De Geer as a synonym of fuscatus; De Geer did not describe the species as new, he attributed it to Linné]; redescription; notes).

  • — Marsham, 1802: 501 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 386, cited punctulatus Marsham as a synonym of fuscatus; Marsham did not describe the species, he attributed it to Paykull]; characters; Britain).

  • — Gravenhorst, 1806: 112 (Staphylinus).

  • — Gyllenhal, 1810: 288 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 386, cited morio Gyllenhal as a synonym of fuscatus Gravenhorst; Gyllenhal did not describe the species as new, he attributed it to Gravenhorst]; characters; Sweden).

  • — Curtis, 1829: 24 (Ocypus; catalog; Britain).

  • — Stephens, 1832: 210 (Goerius; characters; England).

  • — Brullé, 1839: 60 (Staphylinus; Canary Islands [Note: Probably misidentified (Schillhammer, personal communication)]).

  • — Erichson, 1839a: 441 (Ocypus; characters; Germany).

  • — Heer, 1839: 254 (Staphylinus; characters; Switzerland).

  • — Stephens, 1839: 388 (Goerius; characters; England).

  • — Erichson, 1840: 411 (Ocypus; characters; Europe).

  • — Laporte, 1840: 177 (Staphylinus; characters; France).

  • — Jacobsen, 1849: 685 (Ocypus; characters; Denmark).

  • — Redtenbacher, 1849: 698 (Ocypus; characters; Austria).

  • — Küster, 1853: 60 (Ocypus; characters; Europe).

  • — Hardy, 1851: 23 (Ocypus; subgenus Goerius; notes; England).

  • — Fairmaire and Laboulbène, 1856: 510 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 558 (Ocypus; characters; Germany).

  • — Redtenbacher, 1857: 188 (Ocypus; characters; Austria).

  • — Kraatz, 1858a: 61 (Ocypus; Greece).

  • — Thomson, 1860: 148 (Ocypus; characters; Scandinavia).

  • — Hochhuth, 1862: 49 (Ocypus; Russia).

  • — Fauvel, 1874: 188 [= 1874a: 414] (Staphylinus; characters; France; Siberia; Amur region).

  • — Redtenbacher, 1874: 207 (Ocypus; characters; Austria).

  • — Seidlitz, 1875: 279 (Ocypus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 303 [= 1877: 159] (Ocypus; subgenus Pseudocypus; characters; France).

  • — J. Sahlberg, 1876: 11 (Ocypus; Finland).

  • — J. Sahlberg, 1880: 68 (Ocypus; Siberia).

  • — Fowler, 1888: 256 (Ocypus; characters; Britain).

  • — Ganglbauer, 1895: 433 (Staphylinus; subgenus Pseudocypus; characters; Palaearctic region).

  • — Fauvel, 1902b: 105 (Staphylinus; synonym of aeneocephalus).

  • — Reitter, 1909: 119 (Staphylinus; subgenus Pseudocypus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; subgenus Ocypus; catalog).

  • — Johansen, 1914: 384 (Staphylinus; subgenus Goerius; characters; Denmark).

  • — G. Müller, 1926: 17 (Staphylinus; subgenus Pseudocypus; characters; notes).

  • — Portevin, 1929: 351 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1394 (Staphylinus; catalog).

  • — V. Hansen, 1952: 104 (Staphylinus; subgenus Pseudocypus; characters; Denmark).

  • — Coiffait, 1956e: 209, 214 (Ocypus; subgenus Pseudocypus; characters).

  • — Smetana, 1958: 291 (Ocypus; subgenus Pseudocypus; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 116 (Staphylinus; subgenus Pseudocypus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 200 (Ocypus; subgenus Pseudocypus; characters; central Europe).

  • — Horion, 1965: 221 (Staphylinus; subgenus Pseudocypus; Russia; Finland; Sweden; Norway; Denmark; Britain; Holland; Belgium; France; Switzerland; Italy; Slovenia; Dalmatia; Bosnia Hercegovina; Montenegro; Greece; Serbia; Romania; Hungary; Czechoslovakia; Poland; Germany; Austria).

  • — Pototskaia, 1967: 69 (Staphylinus; subgenus Pseudocypus; larval characters).

  • — Szujecki, 1968a: 732 (Ocypus; Poland).

  • — Coiffait, 1974: 502 (Pseudocypus; characters; Europe; Britain; Scandinavia; Russia; Kazakhstan).

  • — Shilov, 1975a: 376 [= 1975b: 99] (Ocypus; Komi region of Russia).

  • — Pope, 1977: 30 (Staphylinus; Britain).

  • — Hozman, 1979: 125 (Pseudocypus; aedeagal aberration and normal form).

  • — Hinton, 1981: 662 (Ocypus; characters of egg).

  • — Tóth, 1984: 29 (Ocypus; characters; Hungary).

  • — Lucht, 1987: 108 (Ocypus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Huth and Dettner, 1989: 154 (Ocypus; defensive secretions).

  • — Nowosad, 1990: 117 (Ocypus; collected from mole nest; notes; Poland).

  • — Gusarov, 1993: 71 (Ocypus; subgenus Pseudocypus).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Pseudocypus; Czech Republic; Slovakia).

  • — Pilon, 1998: 99 (Ocypus; notes; doubtfully in Italy).

  • — Staniec, 1999a: 56 (Ocypus; characters of pupa; Poland).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Europe, Russia, Kazakhstan.

crassicollis Gravenhorst, 1802: 7 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).

  • — Latreille, 1804: 302 (Staphylinus; characters; Germany).

  • — Erichson, 1839a: 441 (Ocypus; synonym of fuscatus).

  • — Heer, 1839: 254 (Staphylinus; synonym of fuscatus).

  • — Erichson, 1840: 411 (Ocypus; synonym of fuscatus).

  • — Küster, 1853: 60 (Ocypus; synonym of fuscatus).

  • — Fairmaire and Laboulbène, 1856: 510 (Staphylinus; synonym of fuscatus).

  • — Kraatz, 1857d: 558 (Ocypus; synonym of fuscatus).

  • — Ganglbauer, 1895: 433 (Staphylinus; synonym of fuscatus).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; synonym of fuscatus).

  • — Smetana, 1958: 292 (Ocypus; synonym of fuscatus).

  • — Coiffait, 1974: 502 (Pseudocypus; synonym of fuscatus).

subpunctatus Gyllenhal, 1827: 474 (Staphylinus; Type locality: Not cited).

  • — C. Sahlberg, 1830: 310 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 24 (Staphylinus; Finland).

  • — Mannerheim, 1831: 438 (Staphylinus; Finland).

  • — Runde, 1835: 3 (Staphylinus; characters; Germany).

  • — Nordmann, 1837: 72 (Staphylinus; Europe).

  • — Erichson, 1839a: 441 (Ocypus; synonym of fuscatus).

  • — Heer, 1839: 254 (Staphylinus; synonym of fuscatus).

  • — Erichson, 1840: 411 (Ocypus; synonym of fuscatus).

  • — Küster, 1853: 60 (Ocypus; synonym of fuscatus).

  • — Kraatz, 1857d: 558 (Ocypus; synonym of fuscatus).

  • — Thomson, 1860: 148 (Ocypus; synonym of fuscatus).

  • — Ganglbauer, 1895: 433 (Staphylinus; synonym of fuscatus).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; synonym of fuscatus).

  • — Smetana, 1958: 292 (Ocypus; synonym of fuscatus).

uralensis Mannerheim, 1830: 24 (Staphylinus; Type locality: Slatoust Sibiriae).

  • — Mannerheim, 1831: 438 (Staphylinus; Finland).

  • — Nordmann, 1837: 72 (Staphylinus; characters; Siberia).

  • — Erichson, 1840: 416 (Ocypus; synonym of ater).

  • — Motschulsky, 1858d: 68 (Staphylinus; synonym of fuscatus).

  • — Kraatz, 1857d: 558 (Ocypus; synonym of fuscatus).

  • — Ganglbauer, 1895: 433 (Staphylinus; synonym of fuscatus).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; synonym of fuscatus).

  • — Smetana, 1958: 292 (Ocypus; synonym of fuscatus).

  • — Coiffait, 1974: 502 (Pseudocypus; synonym of fuscatus).

semipolitus Holme, 1842: 118 (Staphylinus; Type locality: Christ Church Meadow, Oxford; Kemp-Town, near Brighton).

  • — Hardy, 1851: 23 (Staphylinus; synonym of fuscatus).

  • — Ganglbauer, 1895: 433 (Staphylinus; synonym of fuscatus).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; synonym of fuscatus).

  • — Smetana, 1958: 292 (Ocypus; synonym of fuscatus).

  • — Coiffait, 1974: 502 (Pseudocypus; synonym of fuscatus).

abbreviatus Motschulsky, 1858: 669 (Tasgius; Type locality: Carniole; [Note: See lectotype designation by Gusarov, 1993]).

  • — Fauvel, 1874: 184 [= 1874a: 410] (Staphylinus; synonym of brunnipes).

  • — Ganglbauer, 1895: 433 (Staphylinus; synonym of brunnipes).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of brunnipes).

  • — Smetana, 1958: 290 (Ocypus; synonym of brunnipes).

  • — Coiffait, 1974: 459 (Ocypus; synonym of brunnipes).

  • — Gusarov, 1993: 71 (Ocypus; lectotype designation; synonym of fuscatus).

  • — Smetana and Davies, 2000: 44 (Ocypus; synonym of fuscatus).

fuscipes Heer, 1839, see: mus Brullé, 1832.

fuscoaeneus Solsky, 1871: 158 (Ocypus; Type locality: Samarkand).

  • — Solsky, 1874: 173 (Ocypus; characters; Turkestan).

  • — J. Sahlberg, 1880: 68 (Ocypus; Siberia).

  • — Eppelsheim, 1892: 332 (Ocypus; Turkestan).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; subgenus Ocypus; catalog).

  • — Coiffait, 1956e: 209 (Ocypus; subgenus Pseudocypus; characters).

  • — Coiffait, 1974: 504 (Pseudocypus; characters; Transcaspian region).

  • — Boháč, 1987: 95 (Ocypus; subgenus Pseudocypus; larval characters; bionomic notes; Tajikistan).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Uzbekistan, Tajikistan.

gagates Baudi, 1848, see: Tasgius.

gallicus Coiffait, 1956, see: picipennis Fabricius, 1793.

garganicus Fiori, 1894, see: italicus Aragona, 1830.

gloriosus Sharp, 1874: 34 (Ocypus; Type locality: Sakai, near Osaka).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1394 (Staphylinus; catalog).

  • — Nakane, 1963a: 92 (Ocypus; characters; habitus photograph; Japan).

  • — Yuh, Paik, Kwon, and Lee, 1985: 245 (Ocypus; Korea).

  • — J. Li, 1993: 38 (Ocypus; China).

  • — Kanehisa, Tsumuki, and Kawazu, 1994 (Ocypus; chemical secretions).

  • — Smetana and Davies, 2000: 32, 45 (Ocypus; subgenus Aulacocypus).

  • Distribution: Japan, Korea, China.

gracilicornis Hochhuth, 1849, see: Tasgius.

gracilipalpis Eppelsheim, 1878, see: Tasgius.

graecus Scheerpeltz, 1958, see: picipennis Fabricius, 1793.

graeseri Eppelsheim, 1887: 424 (Ocypus; Type locality: Wladiwostok).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; synonym of fuscatus).

  • — G. Müller, 1925: 45 (Staphylinus; subgenus Pseudocypus; characters; Siberia).

  • — Scheerpeltz, 1933: 1401 (Staphylinus; subspecies of rambouseki).

  • — Coiffait, 1974: 503 (Pseudocypus; valid species; characters; Siberia).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Russia, Mongolia.

brunneopubens G. Müller, 1926b: 45 (Staphylinus; subgenus Pseudocypus; subspecies of graeseri; Type locality: Lago di Baical).

  • — Scheerpeltz, 1933: 1401 (Staphylinus; variety of rambouseki).

  • — Coiffait, 1974: 503 (Pseudocypus; subspecies of graeseri; characters; Lake Baikal; Mongolia).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus; subspecies of graeseri).

gridellii G. Müller, 1924, see: curtipennis Motschulsky, 1849.

grigiensis Reitter, 1918, see: nitens Schrank, 1781.

hagai Naomi, 1992: 139 (Ocypus; subgenus Protocypus; Type locality: Japan: Tottori Pref.: Daisen Shrine, Mt. Daisen).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan.

hakusanus Naomi, 1992a: 57 (Ocypus; subgenus Protocypus; Type locality: Japan: Honshu: Fukui Pref.: Chubu district, Mt. Akausagi).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan.

harbinensis J. Li, 1993: 61 (Ocypus; subgenus Neotasgius, cited as Neotasginus; Type locality: city of Harbin).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: China.

hauseri Bernhauer, 1933, see: fulvotomentosus Eppelsheim, 1889.

heinzi Smetana, 1965c: 30 (Ocypus; subgenus Ocypus; Type locality: Anatolia bor.: Ilgaz-dagl., 1800–2200 m).

  • — Smetana, 1968b: 88 (Ocypus; Anatolia).

  • — Coiffait, 1974: 456 (Ocypus; characters; Anatolia).

  • — Coiffait, 1978d: 164 (Ocypus; Turkey).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Turkey.

helleni G. Müller, 1926a: 30 (Staphylinus; subgenus Pseudocypus; Type locality: Aschabad, Transcaspia; Kopet Dagh, Transcaspia).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; catalog).

  • — G. Müller, 1943: 104 (Staphylinus; notes; Turkestan; Transcaspia; Transcaucasia; Asia Minor).

  • — Scheerpeltz, 1958a: 22 (Staphylinus; subgenus Pseudocypus; Armenia).

  • — Smetana, 1963c: 299 (Ocypus; subgenus Pseudocypus; Mongolia).

  • — Scheerpeltz, 1963: 14 (Staphylinus; subgenus Pseudocypus; Afghanistan).

  • — Smetana, 1965d: 81 (Ocypus; subgenus Pseudocypus; characters; notes; Anatolia).

  • — Smetana, 1968a: 158 (Ocypus; subgenus Pseudocypus; Anatolia).

  • — Smetana, 1971c: 268 (Ocypus; subgenus Pseudocypus; Iran).

  • — Coiffait, 1974: 510 (Pseudocypus; characters; Turkmenistan; Transcaucasus; Asia Minor).

  • — Tronquet, 1981: 71 (Pseudocypus; Afghanistan).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Turkey, Armenia, Iran, Pakistan, India, Afghanistan, Turkmenistan, Mongolia.

almorensis Cameron, 1932: 204 (Staphylinus; subgenus Pseudocypus; Type locality: Triun, 9300 ft Dharmsala, Punjab; [Note: See lectotype designation by Smetana and Davies, 2000: 32, 33]. Also cited from West Almora: Kali Valley, alt. 9000 feet).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; catalog).

  • — Cameron, 1935b: 288 (Staphylinus; subgenus Pseudocypus; India).

  • — Scheerpeltz, 1960: 5 (Staphylinus; subgenus Pseudocypus; Pakistan; Afghanistan).

  • — Coiffait, 1979d: 552 (Pseudocypus; Afghanistan).

  • — Smetana and Davies, 2000: 32, 44 (Ocypus; lectotype designation: Triun, 9300 ft Dharmsala, Punjab; synonym of helleni).

himalayicus Cameron, 1935a: 288 (Staphylinus; subgenus Pseudocypus; Type locality: Sind Valley, alt. 2000–2800 m).

  • — Smetana and Davies, 2000: 33, 44 (Ocypus; synonym of helleni).

bicolor G. Müller, 1943: 105 (Staphylinus; subgenus Pseudocypus; variety of helleni; [preoccupied]; Type locality: Caucasus).

  • — Smetana, 1965d: 81 (Ocypus; synonym of helleni).

  • — Coiffait, 1974: 511 (Pseudocypus; variety of helleni; characters; Caucasus).

hercegovinensis G. Müller, 1932, see: picipennis Fabricius, 1793.

hesperus Crotch, 1867, see: aethiops Waltl, 1835.

hidanus Watanabe, 1987a: 43 (Ocypus; subgenus Protocypus; Type locality: Japan: Honshu: Gifu Pref.: Kagami-daira on Mt. Yumiori-dake, N. Jpn. Alps, 2250 m).

  • — Naomi, 1992a: 59 (Ocypus; subgenus Protocypus; characters; Japan).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan.

himalaycus Coiffait, 1982a: 267 (Pseudocypus; cited as himalaycus on page 231 and 267 but as himalayicus on page 233; Type locality: Nepal: Sonamarg, 2600–2750 m).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Nepal.

himalayicus Cameron, 1935, see: helleni G. Müller, 1926.

hissaricus M. Dvořák, 1984: 62 (Ocypus; Type locality: UdSSR: Tadjikistan: Javroz, bei Dusanbe, Hissar).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Tajikistan.

hochhuthi Eppelsheim, 1878: 110 (Ocypus; Type locality: Michailowo).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; catalog).

  • — G. Müller, 1943: 98 (Staphylinus; subgenus Goerius; characters; Caucasus).

  • — Coiffait, 1974: 475 (Ocypus; characters; Caucasus; Transcaucasus).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • — Solodovnikov, 2000: 314 (Ocypus; characters; notes; Georgia; Azerbaijan).

  • Distribution: Georgia, Azerbaijan.

holosetosus G. Müller, 1943, see: brenskei Reitter, 1884.

horsti Coiffait, 1974, see: obscuroaeneus Fairmaire, 1852.

hypsibatus Bernhauer, 1899, see: ophthalmicus Scopoli, 1763.

ibericus Kolenati, 1846, see: fulvipennis Erichson, 1840.

ibericus Coiffait, 1964, see: ophthalmicus Scopoli, 1763.

impennis Fauvel, 1882b: 130 (Staphylinus; subgenus Ocypus; Type locality: Abyssinie).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; subgenus Ocypus; catalog).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Ethiopia.

inauratus Normand, 1947, see: aethiops Waltl, 1835.

inexspectatus Eppelsheim, 1887: 422 (Ocypus; Type locality: Wladiwostok).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; subgenus Ocypus; cited as inexpectatus; catalog).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; cited as inexpectatus; catalog).

  • — G. Müller, 1943: 105 (Staphylinus; cited as inexpectatus; chracters; Siberia).

  • — Coiffait, 1974: 505 (Pseudocypus; cited as inexpectatus; characters; Siberia).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Russia.

integer Abeille, 1900, see: curtipennis Motschulsky, 1849.

intermedius G. Müller, 1911, see: brunnipes Fabricius, 1781.

iranicus Smetana, 1971, see: matilei Jarrige, 1971.

italicus Aragona, 1830: 10 (Staphylinus; Type locality: Apenines: Derthonenses et Bobienses).

  • — Fauvel, 1874: 183 [= 1874a: 409] (Staphylinus; characters; Austria; Italy).

  • — Mulsant and Rey, 1876: 280 [= 1877: 136] (Ocypus; subgenus Goerius; [Note: Mulsant and Rey cited Erichson's (1840: 406) use of italicus as a subsequent reference of italicus Aragona]; characters; France).

  • — Fiori, 1894: 90 (Ocypus; characters; Italy).

  • — Ganglbauer, 1895: 430 (Staphylinus; subgenus Goerius; characters; Alps; Apennines).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; subgenus Ocypus; catalog).

  • — G. Müller, 1924: 25 (Staphylinus; subgenus Goerius; characters is key).

  • — G. Müller, 1926: 9 (Staphylinus; subgenus Goerius; notes).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; catalog).

  • — Coiffait, 1956e: 203, 206 (Ocypus; subgenus Goerius; characters).

  • — Scheerpeltz, 1961d: 130 (Staphylinus; subgenus Goerius; Italy).

  • — Coiffait, 1974: 452 (Ocypus; characters; Italy).

  • — Boháč, 1987: 92 (Ocypus; larval and pupal characters; bionomic notes; Italy).

  • — Pilon, 1994 (Ocypus; hybrid with olens).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; Italy).

  • — Pilon, 1998: 79 (Ocypus; notes; Italy).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Italy.

italicus Erichson, 1840: 406 (Ocypus; [Note: Bernhauer and Schubert, 1914: 387 listed Erichson's use as a subsequent citation for italicus Aragona but Erichson described the species without attribution to another author]; [preoccupied]; Type locality: montes Appenines).

  • — Gravenhorst, 1846: 96 (Staphylinus; characters; Italy).

  • — Redtenbacher, 1857: 186 (Ocypus; characters; Austria).

  • — Redtenbacher, 1874: 206 (Ocypus; characters; Austria).

  • — G. Müller, 1923: 137 (Staphylinus; subgenus Goerius; characters; Italy).

  • — Smetana and Davies, 2000: 43 (Ocypus; synonym of italicus).

garganicus Fiori, 1894: 89 (Ocypus; Type locality: Italia).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; subgenus Ocypus; catalog).

  • — G. Müller, 1926: 9 (Staphylinus; subgenus Goerius; subspecies of italicus).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; subspecies of italicus).

  • — Coiffait, 1956e: 203 (Ocypus; subgenus Goerius; subspecies of italicus; characters).

  • — Coiffait, 1974: 452 (Ocypus; subspecies of italicus; Italy).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; subspecies of italicus; Italy).

  • — Smetana and Davies, 2000: 43 (Ocypus; synonym of italicus).

silensis Fiori, 1894: 90 (Ocypus; variety of italicus; Type locality: Sila).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; variety of italicus).

  • — G. Müller, 1926: 9 (Staphylinus; subgenus Goerius; subspecies of italicus).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; subspecies of italicus).

  • — Coiffait, 1956e: 203 (Ocypus; subgenus Goerius; subspecies of italicus; characters).

  • — Coiffait, 1974: 452 (Ocypus; subspecies of italicus; Italy).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; subspecies of italicus; Italy).

  • — Smetana and Davies, 2000: 43 (Ocypus; synonym of italicus).

mirtensis G. Müller, 1943: 101 (Staphylinus; subgenus Goerius; subspecies of italicus; Type locality: Leoni nei pressi di Poggio Mirteto, Umbria).

  • — Coiffait, 1974: 452 (Ocypus; subspecies of italicus; Italy).

  • — Smetana and Davies, 2000: 43 (Ocypus; synonym of italicus).

italicus Erichson, 1840, see: italicus Aragona, 1830.

iyonus Naomi, 1992: 137 (Ocypus; subgenus Protocypus; Type locality: Japan: Ehime Pref.: Mt. Ishizuchi).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan.

izayoi Naomi, 1992b: 5 (Ocypus; subgenus Protocypus; Type locality: Japan: Nagano Pref.: Mt. Yatsu).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan.

japonicus Sawada, 1965: 16 (Ocypus; subgenus Ocypus; Type locality: Japan: Pref. Nagano: Mt. Jônen).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: Japan.

jeannei Coiffait, 1980a: 37 (Ocypus; Type locality: Roumanie: Scarisoara alba, 1320 m).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Romania.

jureceki Rambousek, 1923, see: parvulus Sharp, 1874.

kansuensis Bernhauer, 1933: 33 (Staphylinus; subgenus Xanthocypus; Type locality: Süd-Kansou: Hoei-Siee).

  • — Smetana and Davies, 2000: 33, 45 (Ocypus; subgenus Aulacocypus; [Note: Species is valid but erroneously indented in list as though a junior synonym of gloriosus).

  • Distribution: China.

kendappa Naomi, 1992: 144 (Ocypus; subgenus Protocypus; Type locality: Japan: Ehime Pref.: Mt. Ishizuchi).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan.

khnzoriani Coiffait, 1967c: 171 (Ocypus; Type locality: Sarkaroi, Tadjikistan).

  • — Coiffait, 1974: 458 (Ocypus; characters; Tadjikistan).

  • — M. Dvořák, 1984: 61 (Ocypus; Tadjikistan).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Tajikistan.

kiimontanus Naomi, 1992b: 11 (Ocypus; subgenus Protocypus; Type locality: Japan: Wakayama Pref.: Mt. Ohto).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan.

kirbyi Stephens, 1832, see: Tasgius.

kisonus Naomi, 1992: 142 (Ocypus; subgenus Protocypus; Type locality: Japan: Nagono Pref.: Mt. Jônen).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan.

kobensis Cameron, 1930, see: lewisius Sharp, 1874.

korgei Smetana, 1965, see: forficularius Motschulsky, 1860.

kuntzeni G. Müller, 1926a: 29 (Staphylinus; subgenus Goerius; Type locality: Kronstadt in Rumenia, Transsilvania).

  • — G. Müller, 1926: 8 (Staphylinus; subgenus Georius; notes).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; catalog).

  • — Coiffait, 1974: 455 (Ocypus; characters; Romania).

  • — Tóth, 1984: 28 (Ocypus; characters; Hungary).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Romania, Hungary.

kyproticus Coiffait, 1964, see: curtipennis Motschulsky, 1849.

latens Solodovnikov, 2000: 319 (Ocypus; Type locality: Western Georgia: valley of river Chkhalta near camping Atsgara).

  • Distribution: Georgia.

leucophthalmus Marsham, 1802, see: picipennis Fabricius, 1793.

lewisius Sharp, 1874: 33 (Ocypus; Type locality: Yokohama, Japan; [Note: See lectotype designation by Smetana and Davies, 2000: 34]. Originally cited from Kobé [Note: This species was described from a single specimen from Kobé according to the original description]).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; catalog).

  • — Nakane, 1963a: 93 (Ocypus; characters; habitus photograph; Japan).

  • — Hayashi, 1993b: 298 (species is neither in Ocypus nor Staphylinus).

  • — J. Li, 1993: 38 (Ocypus; China).

  • — Smetana and Davies, 2000: 34, 44 (Ocypus; lectotype designation: Yokohama, Japan).

  • Distribution: Japan, China.

kobensis Cameron, 1930c: 207 (Ocypus; Type locality: Kobe: Harada).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; catalog).

  • — Cameron, 1949b: 469 (Ocypus; Japan).

  • — Smetana and Davies, 2000: 34, 44 (Ocypus; synonym of lewisius).

liaoningensis J. Li, 1993: 62 (Ocypus; subgenus Neotasgius, cited as Neotasginus; Type locality: Dashiqiao, Liaoning).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: China.

libanicus Coiffait, 1956, see: excisus G. Müller, 1950.

libanoticus G. Müller, 1950, see: curtipennis Motschulsky, 1849.

limbifrons Hochhuth, 1849, see: Tasgius.

lineatus Walker, 1859, see: Staphylinus.

longimanus Smetana, 1965c: 33 (Ocypus; subgenus Ocypus; Type locality: Anatolia bor., Kulakkaya bei Giresun, 1450 m).

  • — Smetana, 1965d: 80 (Ocypus; subgenus Ocypus; Anatolia).

  • — Smetana, 1968a: 156 (Ocypus; subgenus Ocypus; Anatolia).

  • — Coiffait, 1974: 457 (Ocypus; characters; Turkey).

  • — Coiffait, 1978d: 164 (Ocypus; Turkey).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Turkey.

longipennis Walker, 1858, see: Cafius.

ludovicae Mulsant and Godart, 1855, see: syriacus Baudi, 1848.

lugens Nordmann, 1837, see: nitens Schrank, 1781.

luigionii G. Müller, 1926, see: aethiops Waltl, 1835.

lusitanicus G. Müller, 1943, see: Tasgius.

macrocephalus Gravenhorst, 1802: 160 (Staphylinus; Type locality: montes Hercyniae).

  • — Gravenhorst, 1806: 111 (Staphylinus).

  • — Nordmann, 1837: 64 (Staphylinus; characters; Hercyniae).

  • — Erichson, 1840: 407 (Ocypus; characters; Hercyniae Austriaeque montibus, in Carpathibut, et Volhynia).

  • — Gravenhorst, 1846: 96 (Staphylinus; characters; Italy; Germany).

  • — Redtenbacher, 1849: 697 (Ocypus; characters; Austria).

  • — Küster, 1853: 55 (Ocypus; characters; Austria).

  • — Kraatz, 1857d: 554 (Ocypus; characters; Germany).

  • — Redtenbacher, 1857: 187 (Ocypus; characters; Austria).

  • — Redtenbacher, 1874: 206 (Ocypus; characters; Austria).

  • — Mulsant and Rey, 1876: 294 [= 1877: 150] (Ocypus; characters; France).

  • — Ganglbauer, 1895: 431 (Staphylinus; subgenus Goerius; characters; Europe).

  • — Reitter, 1909: 121 (Staphylinus; subgenus Goerius; characters; Germany).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; subgenus Ocypus; catalog).

  • — G. Müller, 1926: 11, 12 (Staphylinus; subgenus Goerius; notes).

  • — G. Müller, 1926c: 73, 75 (Staphylinus; subgenus Goerius; characters; notes).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; catalog).

  • — Coiffait, 1956e: 204 (Ocypus; subgenus Goerius; characters).

  • — Smetana, 1958: 285 (Ocypus; subgenus Ocypus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 198 (Ocypus; subgenus Goerius; characters; central Europe).

  • — Horion, 1965: 214 (Staphylinus; subgenus Goerius; Germany; Czech Republic; Slovakia; Hungary; Romania; France ?; Switzerland; Germany; Austria).

  • — Coiffait, 1974: 461 (Ocypus; characters; Germany; Carpathians).

  • — Tóth, 1984: 26 (Ocypus; characters).

  • — Lucht, 1987: 108 (Ocypus; checklist; Germany; Poland; Czechoslovakia; Austria).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Ocypus; Czech Republic; Slovakia).

  • — Pilon, 1998: 98 (Ocypus; notes; doubtfully in Italy).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Europe.

macrocephalus Stephens, 1832, see: olens O. Müller, 1764.

major De Geer, 1774, see: olens O. Müller, 1764.

manceps Smetana, 1965c: 37 (Ocypus; subgenus Ocypus; Type locality: Anatolia bor., Kulakkaya bei Giresun, 1450 m).

  • — Smetana, 1965d: 80 (Ocypus; subgenus Ocypus; Anatolia).

  • — Smetana, 1968a: 157 (Ocypus; subgenus Ocypus; Anatolia).

  • — Smetana, 1968b: 89 (Ocypus; Anatolia).

  • — Coiffait, 1974: 478 (Ocypus; characters; Anatolia).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Turkey.

mandli Bernhauer, 1906, see: nitens Schrank, 1781.

maritimus Coiffait, 1956, see: picipennis Fabricius, 1793.

marocanus Coiffait, 1964, see: picipennis Fabricius, 1793.

masculus Nordmann, 1837, see: aethiops Waltl, 1835.

mateui Coiffait, 1954: 170 (Staphylinus; subgenus Goerius; Type locality: Bosque del Cedro, île de Gomera, Canaries).

  • — Coiffait, 1974: 413 (Atlantogoerius; synonym of auricomus).

  • — Smetana and Davies, 2000: 33, 44 (Ocypus; subgenus Pseudocypus; valid species).

  • Distribution: Canary Islands.

auricomus Lindberg, 1953: 4 (Staphylinus; subgenus Goerius; [preoccupied]; Type locality: La Gomera: supra Vallehermoso, ca. 900 m).

  • — Coiffait, 1954: 161 (Staphylinus; characters correspond to sylvaticus).

  • — Coiffait, 1974: 413 (Atlantogoerius; characters; Canary Islands).

  • — Smetana and Davies, 2000: 33, 44 (Ocypus; synonym of mateui).

matilei Jarrige, 1971: 487 (Ocypus; Type locality: Iran: Manzandéran, Weischar).

  • — Coiffait, 1974: 465 (Ocypus; characters; Iran).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Iran.

iranicus Smetana, 1971c: 265 (Ocypus; subgenus Ocypus; Type locality: Iran: Mazanderan, Kolehsar s. Alamdeh, 1100 m).

  • — Coiffait, 1974: 465 (Ocypus; synonym of matilei).

  • — Smetana and Davies, 2000: 43 (Ocypus; synonym of matilei).

megalocephalus Nordmann, 1837: 63 (Staphylinus; Type locality: Carinthia).

  • — Erichson, 1840: 407 (Ocypus; cited as megacephalus; characters; Carinthiae montibus).

  • — Kraatz, 1857d: 555 (Ocypus; cited as megacephalus; characters; Alps).

  • — Ganglbauer, 1895: 433 (Staphylinus; subgenus Goerius; cited as megacephalus; characters; Alps).

  • — Reitter, 1909: 121 (Staphylinus; subgenus Goerius; cited as megacephalus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; subgenus Ocypus; cited as megacephalus; catalog).

  • — Scheerpeltz, 1933: 1396 (Staphylinus; cited as megacephalus; catalog).

  • — Lohse, 1964: 198 (Ocypus; subgenus Goerius; cited as megacephalus; characters; central Europe).

  • — Horion, 1965: 213 (Staphylinus; subgenus Goerius; cited as megacephalus; Slovenia; Italy; Austria).

  • — Coiffait, 1974: 467 (Ocypus; characters; eastern Alps from Tyrol to Slovenia).

  • — Lucht, 1987: 108 (Ocypus; cited as megacephalus; checklist; Austria).

  • — Pilon and Zanetti, 1991: 68 (Ocypus; Italy).

  • — Pilon, 1995: 32, 36 (Ocypus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; “megacephalus Auct.” stated to be synonym; Italy).

  • — Pilon, 1998: 81 (Ocypus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Austria, Italy, Slovenia.

meridionalis Fiori, 1894, see: olens O. Müller, 1764.

metallicus O. Costa, 1839, see: sericeicollis Ménétriés, 1832.

micropterus Redtenbacher, 1849, see: tenebricosus Gravenhorst, 1846.

milleri Quedenfeldt, 1882: 182 (Ocypus; Type locality: Syria).

  • — Quedenfeldt, 1884: 372 (Staphylinus; subgenus Ocypus; characters; Syria).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; synonym of quadraticeps).

  • — Scheerpeltz, 1933: 1397 (Staphylinus; catalog).

  • — Coiffait, 1964: 96 (Ocypus; notes).

  • — Coiffait, 1974: 478 (Ocypus; characters; Syria; Asia Minor).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Syria, Turkey, Iran.

robustus Coiffait, 1964: 96 (Ocypus; subspecies of milleri; Type locality: Iran).

  • — Coiffait, 1974: 478 (Ocypus; subspecies of milleri; Iran).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus; subspecies of milleri).

minax Mulsant and Rey, 1861, see: Tasgius.

mirtensis G. Müller, 1943, see: italicus Aragona, 1830.

miwai Bernhauer, 1943a: 178 (Staphylinus; subgenus Xanthocypus; Type locality: Formosa: Nokosan; Numanohira).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus).

  • Distribution: Taiwan.

montanus Cameron, 1942b: 330 (Staphylinus; subgenus Pseudocypus; Type locality: Chyulu Hills, altitude 5600 feet).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Kenya.

mordax Block, 1799, see: ophthalmicus Scopoli, 1763.

murgulensis Coiffait, 1978f: 267 (Ocypus; Type locality: Anatolie orientale: Murgul-Yayla, 1800 m).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: Turkey.

mus Brullé, 1832: 130 (Staphylinus; Type locality: Morée: Messénie).

  • — Kraatz, 1858a: 61 (Ocypus; notes; Greece).

  • — Fauvel, 1874: 187 [= 1874a: 413] (Staphylinus; characters; Austria; Istria; Italy; Sardinia; Ionian Islands; Greece; Russia; Cyprus; Syria; Turkey; Iran).

  • — Fauvel, 1876a: 71 [= 1876: 256] (Staphylinus; Corfu; Turkey).

  • — Mulsant and Rey, 1876: 301 [= 1877: 157] (Ocypus; subgenus Pseudocypus; characters; France).

  • — Ganglbauer, 1895: 433 (Staphylinus; subgenus Pseudocypus; characters; southern middle Europe; Mediterranean region).

  • — Reitter, 1909: 120 (Staphylinus; subgenus Pseudocypus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; subgenus Ocypus; catalog).

  • — Portevin, 1929: 351 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1397 (Staphylinus; catalog).

  • — Coiffait, 1956e: 208 (Ocypus; subgenus Pseudocypus; characters).

  • — Smetana, 1958: 289 (Ocypus; subgenus Pseudocypus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 200 (Ocypus; subgenus Pseudocypus; characters; central Europe).

  • — Horion, 1965: 220 (Staphylinus; subgenus Pseudocypus; Syria; Palestine; Turkey; Cyprus; Rhodes; Ionian Islands; Greece; Crete; Bosnia Hercegovina; Dalmatia; Istria; Italy; Sardinia; Romania; Hungary; Slovakia; Poland ?; Austria).

  • — Smetana, 1966: 331 (Ocypus; subgenus Pseudocypus; notes; Czechoslovakia).

  • — Smetana, 1971c: 268 (Ocypus; subgenus Pseudocypus; characters; notes; Iran).

  • — Coiffait, 1974: 500 (Pseudocypus; characters; Austria; Croatia; Greece; Lebanon; Turkey; Azerbaijan; Rhodes, Cyprus; Syria; Transcaspia; Italy).

  • — Smetana, 1978: 119 (Ocypus; subgenus Pseudocypus; collecting notes; Israel).

  • — Petrenko, 1980: 82 (Ocypus; Caucasus).

  • — Bordoni, 1984: 341 (Pseudocypus; Lebanon).

  • — H. Franz, 1984: 59 (Staphylinus; Macedonia).

  • — Tóth, 1984: 29 (Ocypus; characters; Hungary).

  • — Lucht, 1987: 108 (Ocypus; checklist; Czechoslovakia; Austria).

  • — Šustek, 1992 (Ocypus; habitat; discussion of distribution; Hungary; Slovakia; Moravia).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Pseudocypus; Czech Republic; Slovakia).

  • — Pilon, 1998: 91 (Ocypus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Europe, Russia, Azerbaijan, Turkey, Syria, Lebanon, Iran, “Transcaspia”.

picipes Nordmann, 1837: 68 (Staphylinus; [preoccupied]; Type locality: Austria).

  • — Erichson, 1840: 409 (Ocypus; characters; Austria; Istria; insulae Jonicae).

  • — Kraatz, 1857d: 557 (Ocypus; characters; Germany).

  • — Redtenbacher, 1857: 187 (Ocypus; characters; Austria).

  • — Kraatz, 1858a: 61 (Ocypus; synonym of mus).

  • — Motschulsky, 1859: 491 (Ocypus; Amur River).

  • — Motschulsky, 1860b: 121 (Ocyus; Amur River).

  • — Gemminger and Harold, 1868: 583 (Ocypus; synonym of mus).

  • — Redtenbacher, 1874: 207 (Ocypus; characters; Austria).

  • — Ganglbauer, 1895: 433 (Staphylinus; synonym of mus).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; synonym of mus).

  • — Scheerpeltz, 1933: 1397 (Staphylinus; synonym of mus).

  • — Smetana, 1958: 289 (Ocypus; synonym of mus).

  • — Coiffait, 1974: 500 (Pseudocypus; synonym of mus).

fuscipes Heer, 1839: 254 (Staphylinus; [preoccupied]; Type locality: Bern).

  • — Heer, 1841: 582 (Staphylinus; synonym of picipes).

  • — Kraatz, 1857d: 557 (Ocypus; synonym of picipes).

  • — Gemminger and Harold, 1868: 583 (Ocypus; synonym of mus).

  • — Ganglbauer, 1895: 433 (Staphylinus; synonym of mus).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; synonym of mus).

  • — Smetana, 1958: 289 (Ocypus; synonym of mus).

  • — Coiffait, 1974: 500 (Pseudocypus; synonym of mus).

transadriaticus G. Müller, 1926: 16 (Staphylinus; subgenus Pseudocypus; subspecies of mus; Type locality: Calabrien: Sambiase; Sizilien: Messina).

  • — G. Müller, 1926b: 44 (Staphylinus; subgenus Pseudocypus; cited as new subspecies of mus; characters; Italy; Sicily).

  • — Scheerpeltz, 1933: 1397 (Staphylinus; subspecies of mus).

  • — Coiffait, 1956e: 208 (Ocypus; subgenus Pseudocypus; subspecies of mus; characters).

  • — Coiffait, 1974: 500 (Pseudocypus; subspecies of mus; Sicily; Italy).

  • — Sparacio, 1995: 162 (Ocypus; subspecies of mus; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; subspecies of mus; Italy).

  • — Smetana and Davies, 2000: 44 (Ocypus; synonym of mus).

tauricus G. Müller, 1932: 80 (Staphylinus; subgenus Pseudocypus; subspecies of mus; Type locality: montagne del Tauro della Panfilia, Asia Minore).

  • — Scheerpeltz, 1933: 1397 (Staphylinus; subspecies of mus).

  • — Coiffait, 1956e: 209 (Ocypus; subgenus Pseudocypus; subspecies of mus; characters; Asia Minor; Cyprus; Crete).

  • — Scheerpeltz, 1956c: 1101 (Staphylinus; subspecies of mus; Cyprus).

  • — Scheerpeltz, 1958a: 22 (Staphylinus; subgenus Pseudocypus; subspecies of mus; Turkey).

  • — Smetana, 1965c: 43 (Ocypus; subgenus Pseudocypus; notes; Anatolia).

  • — Smetana, 1968b: 90 (Ocypus; subgenus Pseudocypus; subspecies mus; Anatolia).

  • — Smetana, 1971c: 268 (Ocypus; synonym of mus).

  • — Coiffait, 1974: 500 (Pseudocypus; subspecies of mus; Asia Minor; Rhodes Island; Crete; Cyprus).

  • — H. Franz, 1987: 70 (Pseudocypus; subspecies of mus; Cyprus).

  • — Smetana and Davies, 2000: 44 (Ocypus; synonym of mus).

mutatus Gemminger and Harold, 1868, see: Tasgius.

nemotoi Naomi, 1992a: 60 (Ocypus; subgenus Protocypus; Type locality: Japan: Honshu: Yamanashi Pref.: Chubu district, Hirogawara, Mt. Shiranekita,).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan.

nepalicus Coiffait, 1981d: 327 (Pseudocypus; Type locality: Barabinse [Barahbise], Mt. Kalingchok, 3000 m).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Nepal.

nero Falderman, 1835, see: nitens Schrank, 1781.

nevadensis G. Müller, 1926, see: picipennis Fabricius, 1793.

nigrinus P. Lucas, 1846, see: Tasgius.

nigroaeneus Sharp, 1889: 109 (Ocypus; Type locality: Subashiri, Nanai; Hakodate; Olga Bay, Korea).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; subgenus Ocypus; catalog).

  • — G. Müller, 1925: 45 (Staphylinus; subgenus Pseudocypus; probably not distinct from graeseri).

  • — Scheerpeltz, 1933: 1401 (Staphylinus; variety of rambouseki).

  • — Yuh, Paik, Kwon, and Lee, 1985: 246 (Ocypus; cited as senior name; Korea).

  • — J. Li, 1993: 38 (Ocypus; cited as nigraeneus; China).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Russia, Mongolia, China, Korea, Japan.

rambouseki G. Müller, 1925: 46 (Staphylinus; subgenus Pseudocypus; Type locality: Chitai-ki-Sterana nella Siberia orientale).

  • — Scheerpeltz, 1933: 1401 (Staphylinus; catalog).

  • — Smetana, 1963c: 299 (Ocypus; subgenus Pseudocypus; Mongolia).

  • — Smetana, 1967d: 209 (Ocypus; subgenus Pseudocypus; Mongolia).

  • — Coiffait, 1974: 488 (Pseudocypus; characters).

  • — Smetana 1975c: 173 (Ocypus; Mongolia).

  • — Yuh, Paik, Kwon, and Lee, 1985: 246 (Ocypus; subspecies of nigroaeneus; Korea).

  • — J. Li, 1993: 38 (Ocypus; cited as valid species; China).

  • — Smetana and Davies, 2000: 44 (Ocypus; synonym of nigroaeneus).

nitens Schrank, 1781: 231 (Staphylinus; Type locality: Austria).

  • — Villers, 1789: 423 (Staphylinus; characters).

  • — Olivier, 1795:(42): 14 (Staphylinus; synonym of brunnipes Fabricius).

  • — Schrank, 1798: 640 (Staphylinus; [Note: According to the citation in Schrank, 1798 this may be the same as one of O. Müller's (1776) species]; characters; notes).

  • — Gravenhorst, 1802: 5 (Staphylinus; synonym of similis Fabricius).

  • — Erichson, 1840: 409 (Ocypus; synonym of similis).

  • — Kraatz, 1857d: 556 (Ocypus; synonym of similis).

  • — Fauvel, 1876a: 71 [= 1876: 256] (Staphylinus; France; Corfu).

  • — Ganglbauer, 1895: 431 (Staphylinus; synonym of similis).

  • — Portevin, 1929: 353 (Staphylinus; characters; France).

  • — Ádám, 1996: 246 (Ocypus; valid name; Hungary).

  • — Smetana and Davies, 2000: 34, 43 (Ocypus; valid name for species).

  • Distribution: Europe, Russia, Caucasus, Turkey, Iran, USA.

similis: Fabricius, 1793: 521 (Staphylinus; [Note: Nomen nudum; Fabricius did not propose this name, he attributed it to Paykull, 1789: 10; the Paykull name is now in Alapsodus. The misidentified species said to be authored by Fabricius needed a new name and had been replaced by nero but nitens is older]).

  • — Panzer, 1795: 352 (Staphylinus; characters; Germany).

  • — Fabricius, 1801: 591 (Staphylinus; characters; Germany).

  • — Gravenhorst, 1802: 5 (Staphylinus; characters).

  • — Latreille, 1804: 302 (Staphylinus; characters).

  • — Gravenhorst, 1806: 111 (Staphylinus; characters).

  • — Gyllenhal, 1810: 287 (Staphylinus; characters; Sweden).

  • — Gyllenhal, 1827: 473 (Staphylinus; characters).

  • — Curtis, 1829: 24 (Goerius; catalog; Britain).

  • — Stephens, 1832: 211 (Ocypus; characters; England).

  • — Lacordaire, 1835: 368 (Emus; characters; France).

  • — Holme, 1837: 63 (Ocypus; Scilly Islands).

  • — Westwood, 1838a: 15 (Ocypus; type species).

  • — Erichson, 1839a: 440 (Ocypus; characters; Germany).

  • — Heer, 1839: 253 (Staphylinus; characters; Switzerland).

  • — Stephens, 1839: 388 (Ocypus; characters; England).

  • — Erichson, 1840: 408 (Ocypus; characters; Europe; Siberia).

  • — Laporte, 1840: 177 (Staphylinus; characters; France).

  • — Kolenati, 1846: 18 (Ocypus; cis, trans Caucasum; Persia borealis).

  • — Gravenhorst, 1846: 97 (Staphylinus; cited as similis auct. and similis Erichson; characters; Switzerland; Crimea; Italy; Austria; Germany).

  • — Jacobsen, 1849: 684 (Ocypus; characters; Denmark).

  • — Hochhuth, 1849: 119 (Ocypus; characters; Caucasus).

  • — Redtenbacher, 1849: 697 (Ocypus; characters; Austria).

  • — Kiesenwetter, 1851a: 414 (Ocypus; Pyrenees).

  • — Küster, 1853: 57 (Ocypus; characters; Europe; Siberia).

  • — Fairmaire and Laboulbène, 1856: 510 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 556 (Ocypus; characters; Germany).

  • — Redtenbacher, 1857: 187 (Ocypus; characters; Austria).

  • — Thomson, 1860: 148 (Ocypus; characters; Scandinavia).

  • — Hochhuth, 1862: 49 (Ocypus; Russia).

  • — Fauvel, 1867: LVI (Ocypus; characters; notes).

  • — Fauvel, 1874: 185 [= 1874a: 411] (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 390 cited nitens Fauvel as a synonym of similis; Fauvel did not describe the species as new, he attributed it to Schrank]; characters; France).

  • — Fauvel, 1874: 185 [= 1874a: 411] (Staphylinus; synonym of nitens).

  • — Redtenbacher, 1874: 206 (Ocypus; characters; Austria).

  • — Seidlitz, 1875: 270 (Ocypus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 286 [= 1877: 142] (Ocypus; subgenus Goerius; characters; France).

  • — Fowler, 1888: 254 (Ocypus; characters; Britain).

  • — Ganglbauer, 1895: 431 (Staphylinus; subgenus Goerius; characters; Palaearctic region).

  • — Reitter, 1909: 121 (Staphylinus; subgenus Goerius; characters; Germany).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; subgenus Ocypus; catalog).

  • — Johansen, 1914: 383 (Staphylinus; subgenus Goerius; characters; Denmark).

  • — G. Müller, 1923: 139 (Staphylinus; subgenus Goerius; characters).

  • — G. Müller, 1926: 13 (Staphylinus; subgenus Goerius; characters; notes).

  • — Scheerpeltz, 1933: 1402 (Staphylinus; catalog).

  • — Jarrige, 1947a: 151 (Goerius; characters; Belgium).

  • — V. Hansen, 1952: 103 (Staphylinus; subgenus Goerius; characters; Denmark).

  • — Coiffait, 1956e: 202, 205 (Ocypus; subgenus Goerius; characters).

  • — Scheerpeltz, 1961d: 131 (Staphylinus; subgenus Goerius; Italy).

  • — Palm, 1963b: 115 (Staphylinus; characters; Sweden; Denmark; Finland).

  • — Lohse, 1964: 198 (Ocypus; subgenus Goerius; characters; central Europe).

  • — Horion, 1965: 216 (Staphylinus; subgenus Goerius; Russia; Finland; Sweden; Britain; Ireland; all central Europe; Spain; Portugal; Corsica; Sardinia; Italy; Dalmatia; Croatia; Bosnia; Serbia; Albania; Romania; Bulgaria; Macedonia; Greece; Ukraine; Morocco).

  • — Pototskaia, 1966b: 354 (Staphylinus; larval characters).

  • — Pototskaia, 1967: 73 (Staphylinus; subgenus Ocypus; larval characters).

  • — Coiffait, 1974: 473 (Ocypus; characters; Italy; Slovenia; Austria).

  • — Pope, 1977: 30 (Staphylinus; synonym of nero).

  • — Tóth, 1984: 26 (Ocypus; characters; Hungary).

  • — Newton, 1987: 382 (Staphylinus; introduced species in USA; Massachusetts; New Hampshire).

  • — Huth and Dettner, 1989: 154 (Ocypus; defensive secretions).

  • — Lohse and Lucht, 1989: 169 (Ocypus; synonym of nero).

  • — Huth and Dettner, 1990 (Ocypus; defensive chemicals).

  • — Pilon, 1991: 105 (Ocypus; synonym of nero).

  • — Krooss, 1994 (Ocypus; notes on life cycle in laboratory and in field; Germany).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; synonym of nero).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus; misidentification of similis Paykull).

nero Faldermann, 1835: 118 (Staphylinus; Type locality: [area] Persico-Armeniaca).

  • — Gemminger and Harold, 1868: 584 (Ocypus; synonym of similis).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; synonym of similis).

  • — Pope, 1977: 30 (Staphylinus; Britain).

  • — Lohse and Lucht, 1989: 169 (valid name).

  • — Pilon, 1991: 105 (Ocypus; [Note: Pilon stated that similis Fabricius was preoccupied and was a synonym of nero. Had Fabricius actually proposed the name it would be preoccupied by Herbst, 1784 (see Stenus) and Paykull, 1789 (see Alapsodus) but Fabricius attributed the species to Paykull. The species cited by Fabricius has been regarded by most authors as separate from Paykull's]; notes; Italy).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; “similis Fabricius” cited as synonym; Italy).

  • — Ádám, 1996: 246 (Ocypus; synonym of nitens).

  • — Pilon, 1998: 81 (Ocypus; ecological notes; Italy).

  • — Staniec, 1999a: 56 (Ocypus; characters of pupa; Poland).

  • — Eyre, Luff, and Lott, 2000: 34 (Ocypus; Britain).

  • — Smetana and Davies, 2000: 34, 43 (Ocypus; synonym of nitens).

  • — Assing, 2001: 75 (Ocypus; collected from xerothermous site; Germany).

lugens Nordmann, 1837: 66 (Staphylinus; Type locality: Russia meridionalis: Astrachan).

  • — Erichson, 1840: 409 (Ocypus; synonym of similis).

  • — Küster, 1853: 57 (Ocypus; synonym of similis).

  • — Kraatz, 1857d: 556 (Ocypus; synonym of similis).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; synonym of similis).

  • — Scheerpeltz, 1933: 1402 (Staphylinus; doubtful synonym of similis).

  • — Jarrige, 1947a: 151 (Goerius; synonym of similis).

abbreviatipennis Aubé, 1850: 315 (Ocypus; Type locality: environs de Batoum, Iméritie).

  • — Fauvel, 1874: 186 [= 1874a: 412] (Staphylinus; synonym of nitens).

  • — Ganglbauer, 1895: 431 (Staphylinus; synonym of similis).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; synonym of similis).

  • — Jarrige, 1947a: 151 (Goerius; synonym of similis).

decurtatus Mulsant and Rey, 1876: 289 [= 1877: 145] (Ocypus; subgenus Goerius; Type locality: les Hautes-Alpes, les montagnes de la Lozère et de la Provence).

  • — Ganglbauer, 1895: 431 (Staphylinus; synonym of similis).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; variety of similis).

  • — Scheerpeltz, 1933: 1402 (Staphylinus; variety of similis).

  • — Jarrige, 1947a: 151 (Goerius; variety ? of similis).

  • — Coiffait, 1956e: 205 (Ocypus; subgenus Goerius; variety of similis; characters).

  • — Coiffait, 1974: 473 (Ocypus; variety of similis; France).

semialatus G. Müller, 1904: 172 (Staphylinus; subgenus Ocypus; form of similis; Type locality: Dalmatien, Zara).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; variety of similis).

  • — G. Müller, 1923: 140 (Staphylinus; subgenus Goerius; form of similis; characters; Italy).

  • — Scheerpeltz, 1933: 1402 (Staphylinus; variety of similis).

  • — Schatzmayr, 1943: 115 (Staphylinus; subspecies of similis; Albania).

  • — Jarrige, 1947a: 151 (Goerius; subspecies of similis).

  • — Coiffait, 1956e: 202, 205 (Ocypus; subgenus Goerius; subspecies of similis; characters).

  • — Smetana, 1958: 288 (Ocypus; subgenus Ocypus; subspecies of similis; characters; ecology; Czechoslovakia).

  • — Smetana, 1964d: 78 (Ocypus; subgenus Ocypus; subspecies of similis; Czechoslovakia).

  • — Szujecki, 1966: 172 (Ocypus; subspecies of similis; characters of egg; notes on natural history).

  • — Szujecki, 1966a (Ocypus; subspecies of similis; larval morphology).

  • — Szujecki, 1968a: 732 (Ocypus; subspecies of similis; Poland).

  • — Smetana, 1971c: 267 (Ocypus; subgenus Ocypus; subspecies of similis; Iran).

  • — Jarrige, 1971: 496 (Ocypus; subspecies of similis; Iran).

  • — Coiffait, 1974: 473 (Ocypus; subspecies of similis; Europe; Britain; Scandinavia; Balkans; Crimea; Turkey).

  • — Lucht, 1987: 108 (Ocypus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Drugmand, 1988: 61 (Ocypus; subspecies of similis; Belgium).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Ocypus; subspecies of nero; Czech Republic; Slovakia).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus; subspecies of nitens).

mandli Bernhauer, 1906: 126 (Staphylinus; subgenus Goerius; variety of nitens; Type locality: südliches Alpengebiet, Tirol; Insel Elba).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; variety of similis).

  • — Scheerpeltz, 1933: 1402 (Staphylinus; variety of similis).

  • — Jarrige, 1947a: 151 (Goerius; aberration of similis).

  • — Coiffait, 1956e: 202, 205 (Ocypus; subgenus Goerius; aberration of similis; characters).

  • — Smetana, 1958: 288 (Ocypus; variety of similis semialatus).

  • — Scheerpeltz, 1961d: 131 (Staphylinus; subgenus Goerius; variety of similis; Italy).

  • — Coiffait, 1974: 473 (Ocypus; variety of similis).

grigiensis Reitter, 1918a: 160 (Goerius; subspecies of similis; Type locality: Lessinische Alpen: Grigital, 600 m).

  • — Scheerpeltz, 1933: 1402 (Staphylinus; variety of similis).

  • — Coiffait, 1974: 473 (Ocypus; subspecies of similis; Italy).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus; subspecies of nitens).

subapterus G. Müller, 1923: 140 (Staphylinus; subgenus Goerius; form of similis; [Note: This name was cited twice. Once as a form of typical similis the second as a form of similis variety mandli. If regarded as published in synonymy, then they are nomina nuda (Article 11.6), if as forms of similis and mandli then they are infrasubspecific names and excluded from nomenclatural consideration (Article 1.3.4)]).

ochropus G. Müller, 1950: 10 (Staphylinus; subgenus Goerius; subspecies of similis; [preoccupied]; Type locality: Italia meridionalis: Monte Gargano).

  • — Coiffait, 1974: 473 (Ocypus; subspecies of similis).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; subspecies of nero; Italy).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus; subspecies of nitens).

nubigena Smetana, 1965c: 39 (Ocypus; subgenus Ocypus; Type locality: Anatolia bor., Soganli-Pass bei Bayburt, 2000–2600 m).

  • — Smetana, 1965d: 80 (Ocypus; subgenus Ocypus; Anatolia).

  • — Coiffait, 1974: 476 (Ocypus; characters; Anatolia).

  • — Boháč, 1980: 86 (Ocypus; Caucasus).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Turkey, Caucasus.

nudicollis Coiffait, 1977b: 234 (Ocypus; Type locality: Bhutan: Dorjula, 3100 m).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: Bhutan.

obscuripes Bernhauer, 1900, see: Tasgius.

obscuroaeneus Fairmaire, 1852: 73 (Ocypus; Type locality: Tanger).

  • — Fairmaire and Coquerel, 1860: 157 (Ocypus; characters; Morocco).

  • — Fauvel, 1874: 190 [= 1874a: 416] (Staphylinus; characters; France; Algeria; Morocco).

  • — Mulsant and Rey, 1876: 312 [= 1877: 168] (Ocypus; subgenus Pseudocypus; characters; France).

  • — Fauvel, 1878: 119 [= 1878a: 39] (Staphylinus; Morocco; Algeria).

  • — Fauvel, 1886: 51 [= 1886a: 43] (Staphylinus; Morocco; Algeria).

  • — Ganglbauer, 1895: 434 (Staphylinus; subgenus Pseudocypus; characters; France; Algeria; Morocco).

  • — Fauvel, 1902b: 105 (Staphylinus; subgenus Ocypus; Morocco; Algeria).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; subgenus Ocypus; catalog).

  • — G. Müller, 1926: 17 (Staphylinus; subgenus Pseudocypus; notes).

  • — Portevin, 1929: 351 (Staphylinus; characters; France).

  • — G. Müller, 1932: 79 (Staphylinus; characters; notes).

  • — Scheerpeltz, 1933: 1397 (Staphylinus; catalog).

  • — Coiffait, 1956e: 212 (Ocypus; subgenus Pseudocypus; characters).

  • — Coiffait, 1973a: 274 (Pseudocypus; Morocco).

  • — Coiffait, 1974: 501 (Pseudocypus; characters; Algeria).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Tunisia, Algeria, Morocco, Madeira, Portugal, Spain, France.

schatzmayri G. Müller, 1923: 142 (Staphylinus; subgenus Pseudocypus; subspecies of obscuroaeneus; Type locality: Evora, Portogallo).

  • — G. Müller, 1926: 18 (Staphylinus; subgenus Pseudocypus; subspecies of obscuroaeneus; characters).

  • — Scheerpeltz, 1933: 1397 (Staphylinus; synonym of obscuroaeneus).

  • — G. Müller, 1943: 103 (Staphylinus; subspecies of obscuroaeneus; characters; notes).

  • — Jarrige, 1954a: 163 (Staphylinus; subgenus Pseudocypus; subspecies of obscuroaeneus; characters; Madeira; Iberia; France).

  • — Coiffait, 1956e: 212 (Ocypus; subgenus Pseudocypus; subspecies of obscuroaeneus; characters).

  • — Smetana, 1963b: 39 (Ocypus; subgenus Pseudocypus; subspecies of obscuroaeneus; Madeira).

  • — Coiffait, 1974: 501 (Pseudocypus; subspecies of obscuroaeneus; Iberian Peninsula; Madeira).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus; subspecies of obscuroaeneus).

algiricus G. Müller, 1932: 79 (Staphylinus; subgenus Pseudocypus; subspecies of obscuroaeneus; Type locality: Algeria: Edough; Ain Cheraia).

  • — Scheerpeltz, 1933: 1397 (Staphylinus; subspecies of obscuroaeneus).

  • — G. Müller, 1943: 103 (Staphylinus; synonym of obscuroaeneus).

  • — Coiffait, 1974: 501 (Pseudocypus; synonym of obscuroaeneus).

  • — Smetana and Davies, 2000: 44 (Ocypus; synonym of obscuroaeneus obscuroaeneus).

horsti Coiffait, 1974: 501 (Pseudocypus; subspecies of obscuroaeneus; Type locality: Tunisie: Aïn Draham, Les Chênes).

  • — Smetana and Davies, 2000: 44 (Ocypus; synonym of obscuroaeneus obscuroaeneus).

atlasicus Coiffait, 1974: 501 (Pseudocypus; subspecies of obscuroaeneus; Type locality: Irfane, Moyen Atlas).

  • — Smetana and Davies, 2000: 44 (Ocypus; synonym of obscuroaeneus obscuroaeneus).

ochropus G. Müller, 1950, see: nitens Schrank, 1781.

ohtsukaorum Naomi, 1992a: 63 (Ocypus; subgenus Protocypus; Type locality: Japan: Ohita Pref.: Kyushu: Mt. Sobo).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan.

olens O. Müller, 1764: 23 (Staphylinus; Type locality: Not cited).

  • — O. Müller, 1776: 97 (Staphylinus; characters).

  • — Goeze, 1777: 726 (Staphylinus; characters).

  • — Schrank, 1781: 230 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 388 cited maxillosus Schrank as a synonym of olens but Schrank attributed it to Linné]; characters; Austria).

  • — Fabricius, 1781: 335 (Staphylinus; synonym of Staphylinus maxillosus).

  • — Fabricius, 1787: 219 (Staphylinus; characters).

  • — Paykull, 1789: 9 (Staphylinus; redescription; notes; Sweden).

  • — Villers, 1789: 424 (Staphylinus; characters).

  • — Gmelin, 1790: 2026 (Staphylinus; characters).

  • — Fabricius, 1793: 520 (Staphylinus; characters).

  • — Olivier, 1795:(42): 9 (Staphylinus; characters; France).

  • — Panzer, 1795: 352 (Staphylinus; characters; Germany).

  • — Cederhjelm, 1798: 117 (Staphylinus; characters).

  • — Paykull, 1800: 371 (Staphylinus; characters; Sweden).

  • — Fabricius, 1801: 591 (Staphylinus; characters).

  • — Turton, 1802: 510 (Staphylinus; characters; Europe).

  • — Gravenhorst, 1802: 4 (Staphylinus; characters).

  • — Marsham, 1802: 497 (Staphylinus; characters; Britain).

  • — Latreille, 1804: 302 (Staphylinus; characters).

  • — Panzer, 1808: 1 (Staphylinus; characters; illustrations).

  • — Gyllenhal, 1810: 285 (Staphylinus; characters; Sweden).

  • — Curtis, 1829: 24 (Ocypus; catalog; Britain).

  • — Latreille, 1829: 434 (Staphylinus; characters; France).

  • — Mannerheim, 1830: 23 (Staphylinus; Tauria; Sweden).

  • — Mannerheim, 1831: 437 (Staphylinus; Tauria; Sweden).

  • — Stephens, 1832: 208 (Goerius; characters; England).

  • — Brullé, 1832: 130 (Staphylinus; characters; Greece).

  • — Griffith and Pidgeon, 1832: 291 (Staphylinus; characters; France).

  • — Ménétriés, 1832: 143 (Staphylinus; Caucasus-Persia region).

  • — Runde, 1835: 3 (Staphylinus; characters; Germany).

  • — Lacordaire, 1835: 367 (Emus; characters; France).

  • — Holme, 1837: 63 (Goerius; notes; Scilly Islands).

  • — Westwood, 1838: 164, 165 (Ocypus; habits, eggs, larvae).

  • — Westwood, 1838a: 15 (Goerius; type species).

  • — Brullé, 1839: 59 (Staphylinus; Canary Islands).

  • — Erichson, 1839a: 439 (Ocypus; characters; Germany).

  • — Heer, 1839: 252 (Staphylinus; characters; Switzerland).

  • — Stephens, 1839: 387 (Goerius; characters; England).

  • — Erichson, 1840: 405 (Ocypus; characters; Europe; Siberia; Mauritania; Teneriff).

  • — Laporte, 1840: 176 (Staphylinus; characters; France).

  • — Holme, 1842: 115, 118 (Goerius; notes).

  • — Dufour, 1843: 32 (Staphylinus; characters; France).

  • — Gravenhorst, 1846: 95 (Staphylinus; characters; Germany; Hungary; Italy; Switzerland).

  • — P. Lucas, 1846: 109 (Ocypus; notes; Algeria).

  • — Jacobsen, 1849: 684 (Ocypus; characters; Denmark).

  • — Hochhuth, 1849: 117 (Ocypus; characters; Caucasus).

  • — Redtenbacher, 1849: 698 (Ocypus; characters; Austria).

  • — Kiesenwetter, 1851a: 414 (Ocypus; France; Spain).

  • — Küster, 1853: 53 (Ocypus; characters; Europe; western Asia; northern Africa; Canary Islands).

  • — Hardy, 1851: 23 (Ocypus; subgenus Goerius; notes; England).

  • — Rosenhauer, 1856: 70 (Ocypus; Algeciras; Cadiz).

  • — Fairmaire and Laboulbène, 1856: 508 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 553 (Ocypus; characters; Germany).

  • — Redtenbacher, 1857: 187 (Ocypus; characters; Austria).

  • — Kraatz, 1858a: 60 (Ocypus; Greece, Crete).

  • — Peyron, 1858: 422 (Ocypus; Turkey).

  • — Thomson, 1859: 24 (type species of Goerius).

  • — Thomson, 1860: 146 (Goerius; characters; Scandinavia).

  • — Hochhuth, 1862: 49 (Ocypus; Russia).

  • — Wollaston, 1864: 564 (Ocypus; notes; Canary Islands).

  • — Wollaston, 1865: 487 (Ocypus; notes; Canary Islands).

  • — Crotch, 1867a: 383 (Ocypus; Azores).

  • — Wollaston, 1871: 299 (Ocypus; notes).

  • — Fauvel, 1874: 183 [= 1874a: 409] (Staphylinus; characters; France; Azores; Canary Islands; Barbary; Cyprus; Turkey; Caucasus; Iran).

  • — Redtenbacher, 1874: 207 (Ocypus; characters; Austria).

  • — Seidlitz, 1875: 270 (Ocypus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 278 [= 1877: 134] (Ocypus; subgenus Goerius; characters; France).

  • — Fauvel, 1878: 118 [= 1878a: 38] (Staphylinus; Barbary).

  • — Fauvel, 1886: 50 [= 1886a: 42] (Staphylinus; Barbary).

  • — Fowler, 1888: 254 (Ocypus; characters; Britain).

  • — Fiori, 1894: 90 (Ocypus; characters; Italy).

  • — Ganglbauer, 1895: 430 (Staphylinus; subgenus Goerius; characters; Europe; Mediterranean region).

  • — Bordas, 1899: 230 (Ocypus; defensive glands).

  • — Fauvel, 1902b: 103 (Staphylinus; subgenus Ocypus; Morocco; Algeria; Tunisia; Azores; Canary Islands).

  • — Poppius, 1905: 9 (Staphylinus; Orkney Islands; Shetland Islands).

  • — Sainte-Claire Deville, 1906: 100 (Staphylinus; Corsica).

  • — Reitter, 1909: 120 (Staphylinus; subgenus Goerius; characters; Germany).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; subgenus Ocypus; catalog).

  • — Johansen, 1914: 382 (Staphylinus; subgenus Goerius; characters; Denmark).

  • — Eichelbaum, 1915a: 91 (Staphylinus; abdominal segmentation; apical segements).

  • — Šulc, 1922 (Staphylinus; osmeterium; anatomy; histology; physiology).

  • — G. Müller, 1924: 24 (Staphylinus; subgenus Goerius; characters is key).

  • — Scheerpeltz, 1933: 1397 (Staphylinus; catalog).

  • — Bernhauer, 1940c: 4 (Staphylinus; subgenus Ocypus; Azores).

  • — Paulian, 1941: 240 (Staphylinus; subgenus Goerius; larval characters).

  • — Méquignon, 1942a: 21 (Staphylinus; subgenus Goerius; Azores).

  • — Schatzmayr, 1943: 114 (Staphylinus; Albania).

  • — Jarrige, 1947a: 154 (Goerius; characters; Belgium).

  • — V. Hansen, 1952: 102 (Staphylinus; subgenus Goerius; characters; Denmark).

  • — Coiffait, 1954: 166 (Goerius; Canary Islands).

  • — Jarrige, 1954a: 161 (Staphylinus; subgenus Goerius; characters; Canaries; Azores; Madeira; North Africa).

  • — Coiffait, 1956e: 201, 205 (Ocypus; subgenus Goerius; characters).

  • — Smetana, 1958: 283 (Ocypus; subgenus Ocypus; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1961d: 130 (Staphylinus; subgenus Goerius; Italy).

  • — Palm, 1963b: 114 (Staphylinus; characters; Sweden; Denmark).

  • — Lohse, 1964: 199 (Ocypus; subgenus Goerius; characters; central Europe).

  • — Focarile, 1964: 61 (Staphylinus; Italy).

  • — Raianu, 1964: 103 (Staphylinus; Romania).

  • — Moore, 1965: 102 (Ocypus; introduced in North America).

  • — Scheerpeltz, 1965b: 404 (Staphylinus; Sicily).

  • — Scheerpeltz, 1965c: 508 (Staphylinus; Sicily).

  • — Horion, 1965: 217 (Staphylinus; subgenus Goerius; Morocco; Algeria; Canaries; widespread in western Europe; Britain; Holland; Belgium; France; Switzerland; Portugal; Spain; Balearics; Italy; Istria; Dalmatia; Albania; Greece; Crete; Bosnia Hercegovina; Romania; Hungary; Czechoslovakia; Poland; Denmark; Sweden; Austria; Germany).

  • — Pototskaia, 1967: 71 (Staphylinus; subgenus Ocypus; larval characters).

  • — Smetana, 1967c: 198 (Ocypus; subgenus Ocypus; Spain).

  • — Smetana, 1970: 60 (Ocypus; distributional summary; Azores).

  • — Coiffait, 1974: 448 (Ocypus; characters; Europe; Britain; Scandinavia; North Africa; Canary Islands; Azores).

  • — Bordoni, 1974b: 6 (Ocypus; Italy).

  • — Bellas, Brown, and Moore, 1974 (Staphylinus; defensive secretions).

  • — Fish and Pattenden, 1975 (Staphylinus; defensive secretions).

  • — Orth, Moore, Fisher and Legner, 1975 (Ocypus; biological notes; predator of Brown Garden snail; distribution; characters; California).

  • — Orth, Moore, Fisher and Legner, 1976 (Ocypus; biological notes; predator of Brown Garden snail; larval and pupal characters; California).

  • — Pope, 1977: 30 (Staphylinus; Britain).

  • — Araujo, 1978: 224 (Staphylinus; defensive glands).

  • — Hoebeke, 1978: 508 (Ocypus; parasitized by proctotrupid).

  • — Israelson, 1981: 2, 3 (Ocypus; notes; Madeira).

  • — Hinton, 1981: 662 (Ocypus; characters of egg).

  • — Bordoni, 1982a: 140 (Ocypus; cave collection; Sardinia).

  • — Frank, 1982: 37 (Ocypus; list of parasites).

  • — Tóth, 1984: 27 (Ocypus; characters; Hungary).

  • — Dajoz, 1985: 57 (Ocypus; structure and function of defensive glands).

  • — Segers, 1987a: 184 (Ocypus; pronotal aberration; Belgium).

  • — Newton, 1987: 381 (Staphylinus; introduced species; California; Oregon).

  • — Lucht, 1987: 107 (Ocypus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Huth and Dettner, 1989: 154 (Ocypus; defensive secretions).

  • — Borges, 1990: Table II (Ocypus; Azores).

  • — Huth and Dettner, 1990 (Ocypus; defensive chemicals).

  • — Pilon, 1991: 104 (Ocypus; notes; Italy).

  • — Pilon and Zanetti, 1991: 67 (Ocypus; Italy).

  • — Welch, 1993: 229 (Staphylinus; ovariole number and ovary structure).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Ocypus; Czech Republic; Slovakia).

  • — Pilon, 1994 (Ocypus; hybrid with italicus).

  • — Zanella, 1995 (Ocypus; phenology; Italy).

  • — Outerelo, Palmer and Pons, 1995: 78 (Ocypus; Mallorca).

  • — Sparacio, 1995: 161 (Ocypus; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; Italy).

  • — K. Bland, 1997: 204 (Staphylinus; eaten by toad; Scotland).

  • — Pilon, 1998: 82 (Ocypus; ecological notes; Italy).

  • — Assing, 1999a: 346 (Ocypus; Canary Islands).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus).

  • — Marchal and Chardonnet, 2000: 72 (Ocypus; Corsica).

  • Distribution: Tunisia, Algeria, Morocco, Mauritania, Canary Islands, Madeira, Azores, Europe, Russia, Turkey, USA.

major De Geer, 1774: 17 (Staphylinus; Type locality: Suede, Hollande).

  • — Paykull, 1789: 9 (Staphylinus; synonym of olens).

  • — Villers, 1789: 420 (Staphylinus; characters; Sweden; France).

  • — Gravenhorst, 1802: 3 (Staphylinus; synonym of olens).

  • — Marsham, 1802: 497 (Staphylinus; synonym of olens).

  • — Gyllenhal, 1810: 286 (Staphylinus; synonym of olens).

  • — Erichson, 1840: 405 (Ocypus; synonym of olens).

  • — Kraatz, 1857d: 553 (Ocypus; synonym of olens).

  • — Ganglbauer, 1895: 430 (Staphylinus; synonym of olens).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; synonym of olens).

  • — Smetana, 1958: 283 (Ocypus; synonym of olens).

  • — Coiffait, 1974: 449 (Ocypus; synonym of olens).

unicolor Herbst, 1784: 149 (Staphylinus; Type locality: Berlin).

  • — Gmelin, 1790: 2026 (Staphylinus; synonym of olens).

  • — Erichson, 1840: 405 (Ocypus; synonym of olens).

  • — P. Lucas, 1846: 109 (Ocypus; synonym of olens).

  • — Kraatz, 1857d: 553 (Ocypus; synonym of olens).

  • — Ganglbauer, 1895: 430 (Staphylinus; synonym of olens).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; synonym of olens).

  • — Smetana, 1958: 283 (Ocypus; synonym of olens).

  • — Coiffait, 1974: 449 (Ocypus; synonym of olens).

macrocephalus Stephens, 1832: 208 (Goerius; [preoccupied]; Type locality: Devonshire).

  • — Curtis, 1829: 24 (Ocypus; nomen nudum; catalog; Britain).

  • — Stephens, 1839: 387 (Goerius; characters; England).

  • — G. Waterhouse, 1858: 23 (Ocypus; synonym of olens).

fulvopilosus Fiori, 1894: 90 (Ocypus; variety of olens; Type locality: Not cited).

  • — Bernhauer and Schubert, 1914: 391 (Staphylinus; synonym of tenebricosus).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; synonym of tenebricosus).

  • — Smetana, 1958: 284 (Ocypus; synonym of tenebricosus).

  • — Coiffait, 1974: 449 (Ocypus; variety of olens).

meridionalis Fiori, 1894: 90 (Ocypus; variety of olens; Type locality: Not cited).

  • — Bernhauer and Schubert, 1914: 391 (Staphylinus; synonym of tenebricosus).

  • — Smetana, 1958: 284 (Ocypus; variety of tenebricosus).

  • — Coiffait, 1974: 449 (Ocypus; variety of olens).

aurulentus Lokay, 1921: 71 (Goerius; aberration of olens; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Německa [Germany], Mügl-See).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; synonym of tenebricosus).

  • — Smetana, 1958: 284 (Ocypus; synonym of tenebricosus).

  • — Coiffait, 1974: 449 (Ocypus; synonym of olens).

azoricus Méquignon, 1942a: 22 (Staphylinus; subgenus Goerius; variety of olens; Type locality: Azores: Furnas).

  • — Jarrige, 1954a: 161 (Staphylinus; variety of olens; characters; Azores).

  • — Smetana, 1970: 60 (Ocypus; synonym of olens).

  • — Coiffait, 1974: 449 (Ocypus; subspecies of olens).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus; subspecies of olens).

olivieri Fauvel, 1868, see: Tasgius.

olympicus Baudi, 1857, see: Tasgius.

opacus Roth, 1851: 118 (Ocypus; Type locality: Abyssinien).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: Ethiopia.

ophthalmicus Scopoli, 1763: 99 (Staphylinus; Type locality: Sesona Istrien [Note: See neotype designation by Drugmand, 1998]. Originally cited from Carniolia).

  • — Goeze, 1777: 725 (Staphylinus; characters).

  • — Gmelin, 1790: 2032 (Staphylinus; page misnumbered as 1032; characters).

  • — Turton, 1802: 516 (Staphylinus; characters; Carniola).

  • — Gravenhorst, 1802: 7 (Staphylinus; synonym of cyaneus Paykull).

  • — Erichson, 1840: 405 (Ocypus; synonym of cyaneus).

  • — P. Lucas, 1846: 110 (Ocypus; synonym of cyaneus).

  • — Kraatz, 1857d: 556 (Ocypus; synonym of cyaneus).

  • — Fauvel, 1874: 185 [= 1874a: 411] (Staphylinus; characters; France; Algeria; Caucasus; Transcaucasia).

  • — Seidlitz, 1875: 270 (Ocypus; characters; Baltic region).

  • — Fauvel, 1878: 118 [= 1878a: 38] (Staphylinus; Morocco; Algeria).

  • — Fauvel, 1886: 50 [= 1886a: 42] (Staphylinus; Barbary).

  • — Fowler, 1888: 255 (Ocypus; synonym of cyaneus).

  • — Ganglbauer, 1895: 431 (Staphylinus; subgenus Goerius; characters; Europe; Mediterranean region).

  • — Fauvel, 1902b: 104 (Staphylinus; subgenus Ocypus; Barbary).

  • — Sainte-Claire Deville, 1906: 100 (Staphylinus; Corsica).

  • — Reitter, 1909: 120 (Staphylinus; subgenus Goerius; characters; Germany).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; subgenus Ocypus; catalog).

  • — Johansen, 1914: 383 (Staphylinus; subgenus Goerius; characters; Denmark).

  • — G. Müller, 1926: 15 (Staphylinus; subgenus Goerius; characters; notes).

  • — Portevin, 1929: 352 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1398 (Staphylinus; catalog).

  • — C. Koch, 1941: 64 (Staphylinus; subgenus Goerius; notes; Morocco).

  • — Jarrige, 1947a: 153 (Goerius; cited as ophtalmicus; characters; France).

  • — V. Hansen, 1952: 102 (Staphylinus; subgenus Goerius; characters; Denmark).

  • — Coiffait, 1956e: 201, 207 (Ocypus; subgenus Goerius; cited as ophtalmicus; characters).

  • — Smetana, 1958: 287 (Ocypus; subgenus Ocypus; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1961d: 131 (Staphylinus; subgenus Goerius; Italy).

  • — Palm, 1963b: 115 (Staphylinus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 197 (Ocypus; subgenus Goerius; characters; central Europe).

  • — Coiffait, 1964: 89 (Ocypus; cited as ophtalmicus; notes).

  • — Raianu, 1964: 103 (Staphylinus; Romania).

  • — Scheerpeltz, 1965b: 404 (Staphylinus; Sicily).

  • — Scheerpeltz, 1965c: 509 (Staphylinus; cited as ophtalmicus; Sicily).

  • — Horion, 1965: 211 (Staphylinus; subgenus Goerius; Armenia; Russia; Finland; Sweden; Norway; Denmark; Britain; Holland; Belgium; Switzerland; France; Spain; Portugal; Corsica; Italy; Dalmatia; Bosnia Hercegovina; Montenegro; Bulgaria; Greece; Romania; Hungary; Czechoslovakia; Poland; Ukraine; Austria; Germany).

  • — Coiffait, 1970e: 62 (Ocypus; cited as ophtalmicus; Arcadia).

  • — Coiffait, 1974: 468 (Ocypus; cited as ophtalmicus; characters; Europe; Britain; Scandinavia).

  • — Bordoni, 1974b: 8 (Ocypus; Italy).

  • — Pope, 1977: 30 (Staphylinus; Britain).

  • — Topp, 1979: 23 (Ocypus; development; diapause).

  • — Tóth, 1984: 25 (Ocypus; characters; Hungary).

  • — Lucht, 1987: 108 (Ocypus; distribution of nominate subspecies; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Huth and Dettner, 1989: 154 (Ocypus; defensive secretions).

  • — Pilon and Zanetti, 1991: 68 (Ocypus; cited as ophtalmicus; Italy).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Ocypus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; Italy).

  • — Drugmand, 1998: 47 (Ocypus; neotype designation: Sesona Istrien; characters; key to subspecies; plains of western Europe).

  • — Pilon, 1998: 84 (Ocypus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus).

  • — Assing, 2001: 75 (Ocypus; collected from xerothermous site; Germany).

  • Distribution: Libya, Tunisia, Algeria, Morocco, Europe, Russia, Georgia, Armenia, Azerbaijan, Turkey, Turkmenistan.

atrocoerulescens Goeze, 1777: 729 (Staphylinus; Type locality: Not cited).

  • — Erichson, 1840: 405 (Ocypus; synonym of cyaneus).

  • — Kraatz, 1857d: 556 (Ocypus; synonym of cyaneus).

  • — Ganglbauer, 1895: 431 (Staphylinus; synonym of ophthalmicus).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; synonym of ophthalmicus).

  • — Coiffait, 1956e: 201 (Ocypus; subgenus Goerius; subspecies of ophthalmicus; characters).

  • — Smetana, 1958: 287 (Ocypus; synonym of ophthalmicus).

  • — Coiffait, 1974: 468 (Ocypus; cited as atroceruleus; synonym of ophthalmicus, cited as ophtalmicus).

  • — Drugmand, 1998: 47 (Ocypus; synonym of ophthalmicus ophthalmicus).

coerulescens Geoffroy, 1785: 164 (Staphylinus; Type locality: Paris).

  • — Villers, 1789: 421 (Staphylinus; characters).

  • — Olivier, 1795:(42): 14 (Staphylinus; synonym of cyaneus).

  • — Gravenhorst, 1802: 7 (Staphylinus; synonym of cyaneus).

  • — Erichson, 1840: 405 (Ocypus; synonym of cyaneus).

  • — P. Lucas, 1846: 110 (Ocypus; synonym of cyaneus).

  • — Kraatz, 1857d: 556 (Ocypus; synonym of cyaneus).

  • — Ganglbauer, 1895: 431 (Staphylinus; synonym of ophthalmicus).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; synonym of ophthalmicus).

  • — Jarrige, 1947a: 153 (Goerius; synonym of ophtalmicus).

  • — Smetana, 1958: 287 (Ocypus; synonym of ophthalmicus).

  • — Coiffait, 1974: 468 (Ocypus; synonym of ophthalmicus).

  • — Drugmand, 1998: 47 (Ocypus; synonym of ophthalmicus ophthalmicus).

cyaneus Paykull, 1789: 13 (Staphylinus; Type locality: Vermdon; Finlandia).

  • — Rossi, 1792: 96 (Staphylinus; characters).

  • — Fabricius, 1793: 521 (Staphylinus; characters; Italy).

  • — Olivier, 1795:(42): 14 (Staphylinus; characters; France, Finland).

  • — Paykull, 1800: 370 (Staphylinus; characters; Sweden, Finland).

  • — Fabricius, 1801: 592 (Staphylinus; characters; Italy).

  • — Turton, 1802: 510 (Staphylinus; characters; Italy).

  • — Gravenhorst, 1802: 6 (Staphylinus; characters).

  • — Marsham, 1802: 501 (Staphylinus; characters; Britain).

  • — Latreille, 1804: 304 (Staphylinus; characters).

  • — Gravenhorst, 1806: 110 (Staphylinus).

  • — Panzer, 1808: 3 (Staphylinus; characters; illustrations).

  • — Gyllenhal, 1810: 286 (Staphylinus; characters; Sweden).

  • — Leach, 1819: 172 (type species of Ocypus).

  • — Curtis, 1829: 24 (Ocypus; catalog; Britain).

  • — C. Sahlberg, 1830: 309 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 24 (Staphylinus; Sweden).

  • — Mannerheim, 1831: 438 (Staphylinus; Sweden).

  • — Stephens, 1832: 209 (Goerius; characters; England).

  • — Runde, 1835: 3 (Staphylinus; characters; Germany).

  • — Lacordaire, 1835: 368 (Emus; characters; France).

  • — Nordmann, 1837: 69 (Staphylinus; Europe).

  • — Holme, 1842: 119 (Goerius; notes).

  • — Erichson, 1839a: 440 (Ocypus; characters; Germany).

  • — Heer, 1839: 253 (Staphylinus; characters; Switzerland).

  • — Stephens, 1839: 388 (Goerius; characters; England).

  • — Erichson, 1840: 405 (Ocypus; characters; Europe).

  • — Laporte, 1840: 176 (Staphylinus; characters; Brazil [Note: This is either a misidentification or an erroneous locality citation. All other records are in the Old World]).

  • — Dufour, 1843: 32 (Staphylinus; characters; France).

  • — Kolenati, 1846: 17 (Ocypus; Elisabethopol; Karabagh).

  • — Gravenhorst, 1846: 98 (Staphylinus; characters; Germany; Sweden; Portugal; France; Switzerland; Crimea; Algeria; Austria).

  • — P. Lucas, 1846: 110 (Ocypus; notes; Algeria).

  • — Jacobsen, 1849: 684 (Ocypus; characters; Denmark).

  • — Ménétriés, 1849: 43 (Ocypus; Turcomanie).

  • — Hochhuth, 1849: 117 (Ocypus; characters; Helenendorf).

  • — Redtenbacher, 1849: 698 (Ocypus; characters; Austria).

  • — Kiesenwetter, 1851a: 414 (Ocypus; Pyrenees).

  • — Küster, 1853: 54 (Ocypus; characters; Europe).

  • — Rosenhauer, 1856: 70 (Ocypus; Algeciras; Cadiz).

  • — Fairmaire and Laboulbène, 1856: 509 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 555 (Ocypus; characters; Germany).

  • — Redtenbacher, 1857: 187 (Ocypus; characters; Austria).

  • — Kraatz, 1858a: 61 (Ocypus; Greece).

  • — Thomson, 1860: 147 (Goerius; characters; Scandinavia).

  • — Fauvel, 1865a: 295 [= 1865b: 45] (Ocypus; habitat; France).

  • — Fauvel, 1867: LVI (Ocypus; characters; notes).

  • — Fauvel, 1874: 185 [= 1874a: 411] (Staphylinus; synonym of ophthalmicus).

  • — Redtenbacher, 1874: 207 (Ocypus; characters; Austria).

  • — Mulsant and Rey, 1876: 282 [= 1877: 138] (Ocypus; subgenus Goerius; characters; larval characters; France).

  • — Fowler, 1888: 255 (Ocypus; characters; Britain).

  • — Ganglbauer, 1895: 431 (Staphylinus; synonym of ophthalmicus).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; synonym of ophthalmicus).

  • — Scheerpeltz, 1933: 1398 (Staphylinus; synonym of ophthalmicus).

  • — Jarrige, 1947a: 153 (Goerius; synonym of ophtalmicus).

  • — Smetana, 1958: 287 (Ocypus; synonym of ophthalmicus).

  • — Coiffait, 1974: 468 (Ocypus; synonym of ophthalmicus).

  • — Pope, 1977: 30 (Staphylinus; synonym of ophthalmicus).

  • — Drugmand, 1998: 47 (Ocypus; lectotype designation: Suède; synonym of ophthalmicus ophthalmicus).

mordax Block, 1799: 115 (Staphylinus; Type locality: Tharand, Plauischer Grund bei Dresden).

  • — Gemminger and Harold, 1868: 582 (Ocypus; synonym of cyaneus).

  • — Ganglbauer, 1895: 431 (Staphylinus; synonym of ophthalmicus).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; synonym of ophthalmicus).

  • — Smetana, 1958: 287 (Ocypus; synonym of ophthalmicus).

  • — Coiffait, 1974: 468 (Ocypus; synonym of ophthalmicus).

  • — Drugmand, 1998: 47 (Ocypus; synonym of ophthalmicus ophthalmicus).

  • — Zerche, 2000: 52 (Ocypus; synonym of ophthalmicus).

azurescens Mannerheim, 1830: 23 (Staphylinus; Type locality: Dalmatia).

  • — Mannerheim, 1831: 437 (Staphylinus; Dalmatia).

  • — Nordmann, 1837: 69 (Staphylinus; Dalmatia).

  • — Erichson, 1840: 405 (Ocypus; synonym of cyaneus).

  • — P. Lucas, 1846: 110 (Ocypus; synonym of cyaneus).

  • — Kraatz, 1857d: 556 (Ocypus; synonym of cyaneus).

  • — Ganglbauer, 1895: 431 (Staphylinus; synonym of ophthalmicus).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; synonym of ophthalmicus).

  • — Smetana, 1958: 287 (Ocypus; synonym of ophthalmicus).

  • — Coiffait, 1974: 468 (Ocypus; synonym of ophthalmicus).

  • — Drugmand, 1998: 47 (Ocypus; lectotype designation: Corfu; synonym of ophthalmicus ophthalmicus).

rugifrons Runde, 1835: 4 (Staphylinus; Type locality: Mansfeld).

  • — Kraatz, 1857d: 556 (Ocypus; synonym of cyaneus).

  • — Ganglbauer, 1895: 431 (Staphylinus; synonym of ophthalmicus).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; synonym of ophthalmicus).

  • — Smetana, 1958: 287 (Ocypus; synonym of ophthalmicus).

  • — Coiffait, 1974: 468 (Ocypus; synonym of ophthalmicus).

  • — Drugmand, 1998: 47 (Ocypus; synonym of ophthalmicus ophthalmicus).

subcyaneus Heer, 1839: 253 (Staphylinus; variety of cyaneus; Type locality: Not cited).

cyanochloris Hochhuth, 1849: 118 (Ocypus; Type locality: Armenien; [Note: See lectotype designation by Gusarov, 1992a]).

  • — Fauvel, 1874: 185 [= 1874a: 411] (Staphylinus; synonym of ophthalmicus).

  • — Ganglbauer, 1895: 431 (Staphylinus; synonym of ophthalmicus).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; synonym of ophthalmicus).

  • — Scheerpeltz, 1933: 1398 (Staphylinus; subspecies of ophthalmicus).

  • — Coiffait, 1956e: 202 (Ocypus; subgenus Goerius; subspecies of ophthalmicus).

  • — Smetana, 1965c: 36 (Ocypus; subgenus Ocypus; subspecies of ophthalmicus; Anatolia).

  • — Smetana, 1965d: 80 (Ocypus; subgenus Ocypus; subspecies of ophthalmicus; Anatolia).

  • — Coiffait, 1974: 468 (Ocypus; subspecies of ophthalmicus; Armenia).

  • — Gusarov, 1992a: 786 [= 1993a: 75] (Ocypus; synonym of ophthalmicus; lectotype designation).

  • — Drugmand, 1998: 47 (Ocypus; synonym of ophthalmicus ophthalmicus).

  • — Smetana and Davies, 2000: 42 (Ocypus; synonym of ophthalmicus ophthalmicus).

atrocyaneus Fairmaire, 1860b: 157 (Ocypus; Type locality: Mers-el-Kébir).

  • — Gemminger and Harold, 1868: 582 (Ocypus; synonym of cyaneus).

  • — Ganglbauer, 1895: 431 (Staphylinus; synonym of ophthalmicus).

  • — Fauvel, 1902b: 104 (Staphylinus; synonym of ophthalmicus).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; synonym of ophthalmicus).

  • — G. Müller, 1926: 15 (Staphylinus; subgenus Goerius; subspecies of ophthalmicus; characters; notes; Spain, Portugal, Morocco, Algeria, Tunisia).

  • — Scheerpeltz, 1933: 1398 (Staphylinus; subspecies of ophthalmicus).

  • — Jarrige, 1947a: 153 (Goerius; subspecies of ophthalmicus; North Africa; Spain; France).

  • — Coiffait, 1964: 92 (Ocypus; subspecies of ophthalmicus; characters).

  • — Smetana, 1967c: 199 (Ocypus; subgenus Ocypus; subspecies of ophthalmicus; notes; Spain).

  • — Coiffait, 1973a: 274 (Ocypus; subspecies of ophthalmicus, cited as ophtalmicus; Morocco).

  • — Coiffait, 1974: 469 (Ocypus; subspecies of ophthalmicus; North Africa).

  • — M. Dvořák, 1984: 64 (Ocypus; subspecies of ophthalmicus; Libya).

  • — Drugmand, 1998: 52 (Ocypus; subspecies of ophthalmicus; characters; France, Spain; Portugal; Morocco; Algeria; Tunisia).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus; subspecies of ophthalmicus).

baicalensis Eppelsheim, 1887: 422 (Ocypus; variety of ophthalmicus; Type locality: Baikalgebiet).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; synonym of ophthalmicus).

  • — Scheerpeltz, 1933: 1398 (Staphylinus; subspecies of ophthalmicus).

  • — Coiffait, 1956e: 202 (Ocypus; subgenus Goerius; subspecies of ophthalmicus).

  • — Coiffait, 1974: 469 (Ocypus; subspecies of ophthalmicus; central Asia).

  • — Drugmand, 1998: 47 (Ocypus; lectotype designation: lac Baïkal synonym of ophthalmicus ophthalmicus).

  • — Smetana and Davies, 2000: 42 (Ocypus; synonym of ophthalmicus ophthalmicus).

hypsibatus Bernhauer, 1899a: 433 (Staphylinus; variety of ophthalmicus; Type locality: südliche Alpen).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; synonym of ophthalmicus).

  • — G. Müller, 1926: 15 (Staphylinus; subgenus Goerius; subspecies of ophthalmicus; characters; notes; Alps).

  • — Scheerpeltz, 1933: 1398 (Staphylinus; subspecies of ophthalmicus).

  • — Jarrige, 1947a: 153 (Goerius; subspecies of ophthalmicus; Alps; Massif Central; Pyrenees).

  • — Coiffait, 1956e: 202 (Ocypus; subgenus Goerius; variety of ophthalmicus; characters).

  • — Lohse, 1964: 197 (Ocypus; subgenus Goerius; subspecies of ophthalmicus; characters; central Europe).

  • — Coiffait, 1964: 93 (Ocypus; variety of ophthalmicus; characters).

  • — Horion, 1965: 211 (Staphylinus; subgenus Goerius; subspecies of ophthalmicus; France; Switzerland; Italy; Germany; Austria; Pyrenees).

  • — Coiffait, 1974: 469 (Ocypus; subspecies of ophthalmicus; Alps).

  • — Lucht, 1987: 108 (Ocypus; subspecies of ophthalmicus; checklist; Germany; Austria; Switzerland).

  • — Drugmand, 1998: 50 (Ocypus; subspecies of ophthalmicus; lectotype designation: Mangart; characters; France; Italy).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus; subspecies of ophthalmicus).

balcanicus G. Müller, 1923: 137 (Staphylinus; subgenus Goerius; subspecies of ophthalmicus; [Note: Cited by Scheerpeltz, 1933: 1398 as balearicus]; Type locality: Macedonia, Keretschkoi; Tiflis, Transcaucaso).

  • — G. Müller, 1926: 15 (Staphylinus; subgenus Goerius; subspecies of ophthalmicus; characters; notes; Balkan peninsula, Caucasus).

  • — Scheerpeltz, 1933: 1398 (Staphylinus; subspecies of ophthalmicus).

  • — Schatzmayr, 1943: 114 (Staphylinus; subspecies of ophthalmicus; Albania).

  • — Coiffait, 1956e: 202 (Ocypus; subgenus Goerius; subspecies of ophthalmicus).

  • — Coiffait, 1964: 92 (Ocypus; subspecies of ophthalmicus; cited as balkanicus; characters).

  • — Coiffait, 1974: 469 (Ocypus; subspecies of ophthalmicus; Balkans).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus; subspecies of ophthalmicus).

balearicus G. Müller, 1926a: 28 (Staphylinus; subgenus Goerius; subspecies of ophthalmicus; Type locality: Mallorca).

  • — G. Müller, 1926: 15 (Staphylinus; subgenus Goerius; subspecies of ophthalmicus; characters; notes; Mallorca).

  • — Scheerpeltz, 1933: 1398 (Staphylinus; subspecies of ophthalmicus).

  • — Coiffait, 1956e: 202 (Ocypus; subgenus Goerius; subspecies of ophthalmicus).

  • — Coiffait, 1964: 92 (Ocypus; subspecies of ophthalmicus; characters).

  • — Horion, 1965: 211 (Staphylinus; subgenus Goerius; subspecies of ophthalmicus; Mallorca).

  • — Coiffait, 1974: 469 (Ocypus; subspecies of ophthalmicus; Mallorca).

  • — Pons and Palmer, 1996: 165 (Ocypus; subspecies of ophthalmicus; Balearic endemic; notes; habitat; Mallorca).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus; subspecies of ophthalmicus).

pernigrus Coiffait, 1964: 90 (Ocypus; subspecies of ophthalmicus; Type locality: Espagne: Pico de Europa, Aliva, vers 1700 m).

  • — Coiffait, 1974: 469 (Ocypus; subspecies of ophthalmicus; Spain).

  • — Drugmand, 1998: 52 (Ocypus; subspecies of ophthalmicus; characters; Spain).

  • — Smetana and Davies, 2000: 42 (Ocypus; synonym of ophthalmicus atrocyaneus).

ibericus Coiffait, 1964: 90 (Ocypus; subspecies of ophthalmicus; [preoccupied]; Type locality: Portugal: Cap Saint-Vincent. Also cited from Spain and France).

  • — Coiffait, 1974: 469 (Ocypus; subspecies of ophthalmicus; France).

  • — Drugmand, 1998: 52 (Ocypus; synonym of ophthalmicus atrocyaneus).

  • — Smetana and Davies, 2000: 42 (Ocypus; synonym of ophthalmicus atrocyaneus).

rodopensis Coiffait, 1971b: 280 (Ocypus; subspecies of ophthalmicus, cited as ophtalmicus; Type locality: Bulgarie: Zabârdo, Rodopes).

  • — Coiffait, 1974: 469 (Ocypus; subspecies of ophthalmicus).

  • — Drugmand, 1998: 50 (Ocypus; subspecies of ophthalmicus; characters; Bulgaria).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus; subspecies of ophthalmicus).

benoiti Drugmand, 1998: 52 (Ocypus; subspecies of ophthalmicus; Type locality: Espagne: Galice, Faro Domanyo Ponte Vedre).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus; subspecies of ophthalmicus).

brigitteae Drugmand, 1998: 55 (Ocypus; subspecies of ophthalmicus; Type locality: France: Corse, Ghisonaccia, plage de Gradugine).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus; subspecies of ophthalmicus).

orientalis Bernhauer and Schubert, 1914, see: orientis Smetana and Davies, 2000.

orientis Smetana and Davies, 2000: 34, 44 (Ocypus; subgenus Pseudocypus replacement name for orientalis Bernhauer and Schubert).

  • — Assing and Wunderle, 2001: 37 (Ocypus; Cyprus).

  • Distribution: Cyprus, Turkey, Syria, Lebanon, Israel.

tomentosus Baudi, 1870: 384 (Ocypus; [Note: This name was replaced when it was included in Staphylinus where it was preoccupied by two previous uses]; Type locality: Cypro).

  • — Fauvel, 1878: 119 [= 1878a: 39] (Staphylinus; cited as tomentatus; Algeria).

  • — Fauvel, 1886: 51 [= 1886a: 43] (Staphylinus; Algeria).

  • — Fauvel, 1902b: 105 (Staphylinus; subgenus Ocypus; Morocco; Algeria).

  • — Smetana, 1968b: 90 (Ocypus; synonym of orientalis).

  • — Coiffait, 1974: 511 (Pseudocypus; synonym of orientalis).

orientalis Bernhauer and Schubert, 1914: 389 (Staphylinus; subgenus Ocypus; replacement name for tomentosus Baudi; [preoccupied]).

  • — Scheerpeltz, 1933: 1399 (Staphylinus; catalog).

  • — Coiffait, 1956e: 212 (Ocypus; subgenus Pseudocypus; characters).

  • — Smetana, 1968b: 90 (Ocypus; subgenus Pseudocypus; Anatolia).

  • — Coiffait, 1970e: 62 (Pseudocypus; Cyprus).

  • — Coiffait, 1974: 511 (Pseudocypus; characters; Palestine; Lebanon; Syria; Cyprus; Asia Minor).

  • — Smetana, 1978: 119 (Ocypus; subgenus Pseudocypus; collecting notes; Israel).

  • — H. Franz, 1987: 70 (Pseudocypus; Cyprus).

  • — Smetana and Davies, 2000: 34, 44 (Ocypus; synonym of orientis).

ormayi Reitter, 1887a: 261 (Staphylinus; variety of macrocephalus; Type locality: Transsylvani'sche Alpen, Nähe des Rothenthurm-Passes).

  • — Ganglbauer, 1895: 431 (Staphylinus; variety of macrocephalus).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; synonym of macrocephalus).

  • — G. Müller, 1926: 12 (Staphylinus; subgenus Goerius; characters; notes).

  • — G. Müller, 1926c: 75 (Staphylinus; characters; Romania; Serbia).

  • — Scheerpeltz, 1933: 1399 (Staphylinus; catalog).

  • — Smetana, 1958: 286 (Ocypus; subgenus Ocypus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 198 (Ocypus; subgenus Goerius; characters; central Europe).

  • — Coiffait, 1974: 462 (Ocypus; characters; Carpathians; Romania).

  • — Tóth, 1984: 26 (Ocypus; characters).

  • — Lucht, 1987: 108 (Ocypus; checklist; Poland; Czechoslovakia).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Ocypus; Slovakia).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Slovakia, Romania, Yugoslavia.

ornativentris Reitter, 1911, see: Dinothenarus.

ottomanus Fauvel, 1900d: 230 (Ocypus; Type locality: Constantinople, Gallipoli).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1399 (Staphylinus; catalog).

  • — Coiffait, 1974: 479 (Ocypus; characters; Turkey).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Turkey.

parvulus Sharp, 1874: 33 (Ocypus; Type locality: Maiyasama).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1399 (Staphylinus; catalog).

  • — Nakane, 1963a: 92 (Ocypus; characters; habitus photograph; Japan).

  • — Yuh, Paik, Kwon, and Lee, 1985: 245 (Ocypus; Korea).

  • — J. Li, 1993: 38 (Ocypus; cited as parovulus; China).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Aulacocypus).

  • Distribution: Japan, Korea, China, Russia.

jureceki Rambousek, 1923: 54 (Staphylinus; subgenus Tasgius; Type locality: Vladivostok en Sibéria).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; catalog).

  • — G. Müller, 1943: 98 (Staphylinus; synonym of parvulus).

pecoudi Jarrige, 1954: 75 (Staphylinus; subgenus Goerius; Type locality: Spain: Sierra Nevada).

  • — Coiffait, 1974: 466 (Ocypus; characters; Spain).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: Spain.

pedemontanus G. Müller, 1924: 20, 25 (Staphylinus; subgenus Goerius; Type locality: Graglia, Piemonte; Dintorni Torino; Piemonte).

  • — G. Müller, 1925a: 26 (Staphylinus; characters).

  • — G. Müller, 1926: 9 (Staphylinus; subgenus Goerius; notes).

  • — Scheerpeltz, 1933: 1399 (Staphylinus; catalog).

  • — Jarrige, 1947a: 154 (Goerius; characters; France).

  • — Coiffait, 1956e: 204, 206 (Ocypus; subgenus Goerius; characters).

  • — Coiffait, 1974: 453 (Ocypus; characters; Italy).

  • — Pilon, 1991: 104 (Ocypus; notes; Italy).

  • — Pilon, 1992: 83 (Ocypus; distribution; Italy).

  • — Wittwer, 1993: 250 (Ocypus; Switzerland).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; Italy).

  • — Pilon, 1998: 86 (Ocypus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Italy, Switzerland, France, Spain.

pyrenaeus G. Müller, 1924: 21, 25 (Staphylinus; subgenus Goerius; Type locality: Lac de Estom, Hautes, Pyrenees).

  • — G. Müller, 1926: 9 (Staphylinus; subgenus Goerius; subspecies of pedemontanus; notes).

  • — Scheerpeltz, 1933: 1399 (Staphylinus; subspecies of pedemontanus).

  • — Jarrige, 1947a: 154 (Goerius; subspecies of pedemontanus).

  • — Coiffait, 1956e: 204, 206 (Ocypus; subgenus Goerius; subspecies of pedemontanus; characters).

  • — Coiffait, 1974: 453 (Ocypus; subspecies of pedemontanus; Pyrenees).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus; subspecies of pedemontanus).

cantabricus G. Müller, 1926a: 29 (Staphylinus; subgenus Goerius; subspecies of pedemontanus; Type locality: Hispania, Durango).

  • — G. Müller, 1926: 9 (Staphylinus; subgenus Goerius; subspecies of pedemontanus; notes).

  • — Scheerpeltz, 1933: 1399 (Staphylinus; subspecies of pedemontanus).

  • — Jarrige, 1947a: 154 (Goerius; subspecies of pedemontanus).

  • — Coiffait, 1956e: 204, 206 (Ocypus; subgenus Goerius; subspecies of pedemontanus; characters).

  • — Coiffait, 1974: 453 (Ocypus; subspecies of pedemontanus; France).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus; subspecies of pedemontanus).

penetrans O. Müller, 1776, see: picipennis Fabricius, 1793.

pernigrus Coiffait, 1964, see: ophthalmicus Scopoli, 1763.

petricola Gistel, 1857: 71 (Goerius; [Note: Nomen dubium]; Type locality: Tyrolis).

  • Distribution: Austria.

phaeopus Stephens, 1832, see: Tasgius.

philonthoides Hochhuth, 1849, see: fulvipennis Erichson, 1840.

picicupreus Coiffait, 1974 [Note: Nomen nudum], see: sericeicollis Ménétriés, 1832.

picipennis Fabricius, 1793: 521 (Staphylinus; Type locality: Germania).

  • — Paykull, 1789: 11 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 389, cited aeneocephalus Paykull as a synonym of picipennis; Paykull did not describe the species as new, he attributed it to De Geer]; redescription; notes; Sweden).

  • — Panzer, 1795: 353 (Staphylinus; characters; Germany).

  • — Fabricius, 1801: 591 (Staphylinus; characters; Germany).

  • — Turton, 1802: 510 (Staphylinus; characters; Germany).

  • — Curtis, 1829: 24 (Quedius; catalog; Britain).

  • — Mannerheim, 1830: 23 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 389, cited chalcocephalus Mannerheim as a synonym of picipennis Fabricius; Mannerheim did not describe the species as new, he attributed it to Fabricius]; Sweden).

  • — Mannerheim, 1831: 437 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 389, cited chalcocephalus Mannerheim as a synonym of picipennis Fabricius; Mannerheim did not describe the species as new nor even cite characters for it, he attributed it to Fabricius]; Sweden).

  • — Runde, 1835: 2 (Staphylinus; characters; Germany).

  • — Erichson, 1839a: 442 (Ocypus; characters; Germany).

  • — Heer, 1839: 255 (Staphylinus; characters; Switzerland).

  • — Erichson, 1840: 412 (Ocypus; characters; Europe).

  • — Kolenati, 1846: 18 (Ocypus; Caucasus; Transcaucasia; Iberia).

  • — Jacobsen, 1849: 685 (Ocypus; characters; Denmark).

  • — Ménétriés, 1849: 43 (Ocypus; Samarkand).

  • — Hochhuth, 1849: 123 (Ocypus; characters; Caucasus).

  • — Redtenbacher, 1849: 698 (Ocypus; characters; Austria).

  • — Küster, 1853: 61 (Ocypus; characters; Europe).

  • — Rosenhauer, 1856: 70 (Ocypus; Granada).

  • — Fairmaire and Laboulbène, 1856: 510 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 559 (Ocypus; characters; Germany).

  • — Redtenbacher, 1857: 188 (Ocypus; characters; Austria).

  • — Peyron, 1858: 422 (Ocypus; Turkey).

  • — Thomson, 1860: 148 (Ocypus; characters; Scandinavia).

  • — Hochhuth, 1862: 49 (Ocypus; Russia).

  • — Fauvel, 1865a: 295 [= 1865b: 45] (Ocypus; habitat; France).

  • — Fauvel, 1874: 188 [= 1874a: 414] (Staphylinus; characters; France; Algeria; Turkey; Caucasus; Lake Baikal; Amur region).

  • — Redtenbacher, 1874: 207 (Ocypus; characters; Austria).

  • — Solsky, 1874: 173 (Ocypus; characters; Turkestan).

  • — Seidlitz, 1875: 270 (Ocypus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 306 [= 1877: 162] (Ocypus; subgenus Pseudocypus; characters; France).

  • — J. Sahlberg, 1876: 11 (Ocypus; Finland).

  • — Fauvel, 1878: 119 [= 1878a: 39] (Staphylinus; Algeria).

  • — Fauvel, 1886: 50 [= 1886a: 42] (Staphylinus; Algeria).

  • — Eppelsheim, 1892: 332 (Ocypus; Turkestan).

  • — Ganglbauer, 1895: 433 (Staphylinus; subgenus Pseudocypus; characters; Palaearctic region).

  • — Fauvel, 1902b: 104 (Staphylinus; subgenus Ocypus; Algeria).

  • — Reitter, 1909: 120 (Staphylinus; subgenus Pseudocypus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; subgenus Ocypus; catalog).

  • — Johansen, 1914: 385 (Staphylinus; subgenus Goerius; characters; Denmark).

  • — G. Müller, 1926: 20 (Staphylinus; subgenus Pseudocypus; characters; notes).

  • — G. Müller, 1926b: 45, 46 (Staphylinus; characters; notes; Poland, Norway, Austria, western Alps, Hungary, Serbia, Bulgaria, Macedonia, Urals, Syr-Darja, Tien-Schan).

  • — Portevin, 1929: 351 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1400 (Staphylinus; catalog).

  • — Bernhauer, 1936: 5 (Staphylinus; subgenus Pseudocypus; China).

  • — C. Koch, 1941: 65 (Staphylinus; Morocco).

  • — G. Müller, 1943: 103 (Staphylinus; characters; notes).

  • — V. Hansen, 1952: 104 (Staphylinus; subgenus Pseudocypus; characters; Denmark).

  • — Coiffait, 1956e: 209, 215 (Ocypus; subgenus Pseudocypus; characters).

  • — Smetana, 1958: 293 (Ocypus; subgenus Pseudocypus; characters; ecology; Czechoslovakia).

  • — Kocher, 1958: 153 (Staphylinus; Morocco).

  • — Smetana, 1959a: 214 (Ocypus; subgenus Pseudocypus; Albania).

  • — Ferreira, 1962: 18 (Staphylinus; Portugal).

  • — Palm, 1963b: 116 (Staphylinus; subgenus Pseudocypus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 201 (Ocypus; subgenus Pseudocypus; characters; central Europe).

  • — Coiffait, 1964: 98, 101 (Pseudocypus; note; key to subspecies and varieties).

  • — Smetana, 1965c: 43 (Ocypus; subgenus Pseudocypus; characters; notes; Anatolia).

  • — Smetana, 1965d: 80 (Ocypus; subgenus Pseudocypus; notes; Anatolia).

  • — Horion, 1965: 224 (Staphylinus; subgenus Pseudocypus; Russia; Finland; Sweden; Norway; Denmark; all central Europe; Portugal and Spain to Greece).

  • — Pototskaia, 1966b: 359 (Staphylinus; subgenus Pseudocypus larval characters).

  • — Szujecki, 1966: 173 (Ocypus; characters of egg; notes on natural history).

  • — Smetana, 1967d: 209 (Ocypus; subgenus Pseudocypus; Mongolia).

  • — Smetana, 1967i: 557 (Ocypus; subgenus Pseudocypus; Turkey).

  • — Pototskaia, 1967: 68 (Staphylinus; subgenus Pseudocypus; larval characters).

  • — Smetana, 1968a: 157 (Ocypus; subgenus Pseudocypus; Anatolia).

  • — Smetana, 1968b: 90 (Ocypus; subgenus Pseudocypus; Anatolia).

  • — Szujecki, 1968a: 733 (Ocypus; Poland).

  • — Scheerpeltz, 1968a: 83 (Staphylinus; catalog; Austria).

  • — Bolov, 1969: 514 (Ocypus; collecting notes; Russia).

  • — Bolov, 1969a: 328 (Ocypus; collecting notes; Russia).

  • — Coiffait, 1974: 507 (Pseudocypus; characters; Palaearctic region).

  • — Smetana 1975c: 173 (Ocypus; subgenus Pseudocypus; Mongolia).

  • — Shilov, 1975a: 376 [= 1975b: 99] (Ocypus; Komi region of Russia).

  • — Muona, 1979: 15 (Ocypus; Finland; Russia; Sweden; Norway; Denmark; Baltic States).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 109 (Ocypus; catalog; Poland).

  • — Tóth, 1984: 30 (Ocypus; characters; Hungary).

  • — Ádám, 1987: 149 (Pseudocypus; collecting notes; Hungary).

  • — Lucht, 1987: 108 (Ocypus; nominate subspecies; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Silfverberg, 1992: 17 (Ocypus; Finland; Russia; Sweden; Norway; Denmark; Estonia; Latvia; Lithuania).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Pseudocypus; subspecies picipennis; Czech Republic; Slovakia).

  • — Assing, 1994a: 12, 24 (Ocypus; notes; wing length; xerothermous sites; Germany).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; Italy).

  • — M. Hansen, 1996: 106 (Ocypus; checklist; Denmark).

  • — Ádám, 1996: 246 (Pseudocypus; synonym of penetrans).

  • — Smoleński, 1997: 78 (Ocypus; collecting notes; Poland).

  • — Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 38 (Ocypus; Latvia).

  • — Hansen, Jørum, Palm, and Pedersen, 1997: 128 (Ocypus; checklist; Denmark).

  • — Pilon, 1998: 91 (Ocypus; notes; Italy).

  • — Kashcheev, 1999a: 58 (Ocypus; Kazakhstan).

  • — Smetana and Davies, 2000: 44 (Ocypus; subgenus Pseudocypus).

  • — Herman, 2001: 27 (Ocypus; prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.3).

  • Distribution: Algeria, Morocco, Europe, Russia, Georgia, Armenia, Turkey, Iran, Uzbekistan, Kazakhstan, Mongolia, China.

quartus Schaeffer, 1766: xxxv, 9 (Staphylinus; [Note: Nomen nudum, Article 11.4]; Type locality: Ratisbon).

  • — Erichson, 1840: 412 (Ocypus; synonym of picipennis).

  • — Kraatz, 1857d: 559 (Ocypus; synonym of picipennis).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; synonym of picipennis).

  • — Smetana, 1958: 294 (Ocypus; synonym of picipennis).

penetrans O. Müller, 1776: 97 (Staphylinus; [Note: This name is older than picipennis]; Type locality: Dania).

  • — Goeze, 1777: 727 (Staphylinus; characters).

  • — Harrer, 1784: 251 (Staphylinus; characters).

  • — Gmelin, 1790: 2033 (Staphylinus; characters; Denmark).

  • — Erichson, 1840: 412 (Ocypus; synonym of picipennis).

  • — Kraatz, 1857d: 559 (Ocypus; synonym of picipennis).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; synonym of picipennis).

  • — Smetana, 1958: 294 (Ocypus; synonym of picipennis).

  • — Coiffait, 1974: 507 (Pseudocypus; synonym of picipennis).

  • — Ádám, 1996: 246 (Pseudocypus; valid name; Hungary).

  • — Herman, 2001: 27 (Ocypus; synonym of picipennis).

tristis Fabricius, 1793: 524 (Staphylinus; Type locality: Dania. [Note: Staphylinus tristis was cited by Blackwelder, 1952: 335 as the type species for Quedius. The original citation of type species for Quedius was tristis Gravenhorst but Gravenhorst attributed the species to Fabricius]).

  • — Fabricius, 1801: 594 (Staphylinus; characters; Denmark).

  • — Turton, 1802: 511 (Staphylinus; characters; Denmark).

  • — Gravenhorst, 1802: 34 (Staphylinus; [Note: This reference is listed by Bernhauer and Schubert, 1916: 435 as though Gravenhorst described tristis but Gravenhorst attributed the species to Fabricius; however tristis is the type species of Quedius. I have listed this citation for Quedius tristis also]; characters).

  • — Latreille, 1804: 319 (Staphylinus; characters; Germany; France).

  • — Gravenhorst, 1806: 48 (Staphylinus; characters; Germany; France).

  • — C. Sahlberg, 1830: 314 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 25 (Staphylinus; Finland; Sweden).

  • — Mannerheim, 1831: 439 (Staphylinus; Finland; Sweden).

  • — Runde, 1835: 5 (Staphylinus; characters; Germany).

  • — Erichson, 1839a: 442 (Ocypus; characters; Germany).

  • — Erichson, 1840: 412 (Ocypus; synonym of picipennis).

  • — Kraatz, 1857d: 559 (Ocypus; synonym of picipennis).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; synonym of picipennis).

  • — Coiffait, 1956e: 210 (Ocypus; subgenus Pseudocypus; variety of picipennis).

  • — Smetana, 1958: 294 (Ocypus; synonym of picipennis).

  • — Coiffait, 1964: 98 (Pseudocypus; variety of picipennis; elytral color).

  • — Coiffait, 1974: 507 (Pseudocypus; variety of picipennis).

  • — Tóth, 1984: 31 (Ocypus; variety of picipennis; Hungary).

sericeus Marsham, 1802: 508 (Staphylinus; Type locality: Britain).

  • — Gyllenhal, 1810: 292 (Staphylinus; synonym of aeneocephalus De Geer).

  • — Stephens, 1829: 275 (Staphylinus; synonym of aeneocephalus De Geer).

  • — Erichson, 1840: 412 (Ocypus; synonym of picipennis).

  • — Kraatz, 1857d: 559 (Ocypus; synonym of picipennis).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; synonym of picipennis).

  • — Smetana, 1958: 294 (Ocypus; synonym of picipennis).

  • — Coiffait, 1974: 507 (Pseudocypus; synonym of picipennis).

leucophthalmus Marsham, 1802: 511 (Staphylinus; Type locality: Britain).

  • — Erichson, 1840: 412 (Ocypus; synonym of picipennis).

  • — Kraatz, 1857d: 559 (Ocypus; synonym of picipennis).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; synonym of picipennis).

  • — Smetana, 1958: 294 (Ocypus; synonym of picipennis).

  • — Coiffait, 1974: 507 (Pseudocypus; synonym of picipennis).

nevadensis G. Müller, 1926: 21 (Staphylinus; subgenus Pseudocypus; subspecies of picipennis; Type locality: Spanien: Sierra Nevada).

  • — G. Müller, 1926b: 46 (Staphylinus; cited as new subspecies of picipennis; characters; Spain).

  • — Scheerpeltz, 1933: 1400 (Staphylinus; synonym of picipennis).

  • — Coiffait, 1956e: 212 (Ocypus; subgenus Pseudocypus; subspecies of picipennis).

  • — Coiffait, 1964: 98, 104 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana, 1967c: 199 (Ocypus; subgenus Pseudocypus; notes; Spain).

  • — Coiffait, 1974: 507 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus; subspecies of picipennis).

abbruzzensis G. Müller, 1926: 21 (Staphylinus; subgenus Pseudocypus; subspecies of picipennis; Type locality: Abruzzen: Gran Sasso).

  • — Scheerpeltz, 1933: 1400 (Staphylinus; synonym of picipennis).

  • — G. Müller, 1943: 103 (Staphylinus; subspecies of picipennis; characters; notes).

  • — Coiffait, 1956e: 212 (Ocypus; subgenus Pseudocypus; subspecies of picipennis).

  • — Coiffait, 1964: 98, 104 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana, 1965c: 43 (Ocypus; synonym of picipennis).

  • — Coiffait, 1974: 507 (Pseudocypus; subspecies of picipennis; characters).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; subspecies of picipennis; Italy).

  • — Smetana and Davies, 2000: 44 (Ocypus; synonym of picipennis picipennis).

fallaciosus G. Müller, 1926: 21 (Staphylinus; subgenus Pseudocypus; subspecies of picipennis; Type locality: Italien: vom Latium bis in die Südalpen; Ostalpen nördlich bis Graz).

  • — G. Müller, 1926b: 47 (Staphylinus; cited as new subspecies of picipennis; characters; Italy).

  • — G. Müller, 1932: 81 (Staphylinus; subgenus Pseudocypus; notes).

  • — Scheerpeltz, 1933: 1400 (Staphylinus; synonym of picipennis).

  • — Coiffait, 1956e: 210, 215 (Ocypus; subgenus Pseudocypus; subspecies of picipennis; characters).

  • — Smetana, 1958: 294 (Ocypus; synonym of picipennis).

  • — Lohse, 1964: 201 (Ocypus; subgenus Pseudocypus; subspecies of picipennis; characters; central Europe).

  • — Coiffait, 1964: 98, 104 (Pseudocypus; subspecies of picipennis; characters).

  • — Coiffait, 1974: 507 (Pseudocypus; subspecies of picipennis; characters).

  • — Bordoni, 1974b: 8 (Pseudocypus; subspecies of picipennis; notes; Italy).

  • — Lucht, 1987: 108 (Ocypus; subspecies of picipennis; checklist; Austria).

  • — Pilon and Zanetti, 1991: 69 (Ocypus; subspecies of picipennis; Italy).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; subspecies of picipennis; Italy).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus; subspecies of picipennis).

altaiensis G. Müller, 1926b: 47 (Staphylinus; subspecies of picipennis; Type locality: Altai, Tscholesmantal).

  • — Scheerpeltz, 1933: 1400 (Staphylinus; synonym of picipennis).

  • — Coiffait, 1956e: 212 (Ocypus; subgenus Pseudocypus; subspecies of picipennis).

  • — Coiffait, 1964: 98, 105 (Pseudocypus; subspecies of picipennis; characters).

  • — Coiffait, 1974: 507 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus; subspecies of picipennis).

aprutianus G. Müller, 1926b: 47 (Staphylinus; subspecies of picipennis; Type locality: Gran Sasso, Abruzzi; Abruzzo, Aquilano, Alfedena).

  • — Scheerpeltz, 1933: 1400 (Staphylinus; synonym of picipennis).

  • — Smetana, 1965c: 44 (Ocypus; synonym of picipennis).

  • — Coiffait, 1974: 507 (Pseudocypus; synonym of picipennis).

  • — Smetana and Davies, 2000: 44 (Ocypus; synonym of picipennis picipennis).

barbarus G. Müller, 1926b: 47 (Staphylinus; subspecies of picipennis; Type locality: Algeria: Batna; Marocco: Ras el Ma, 1800 m, Moyen Atlas).

  • — Scheerpeltz, 1933: 1400 (Staphylinus; synonym of picipennis).

  • — Coiffait, 1956e: 212 (Ocypus; subgenus Pseudocypus; subspecies of picipennis).

  • — Coiffait, 1964: 98, 102 (Pseudocypus; subspecies of picipennis; characters; elytral color).

  • — Coiffait, 1974: 507 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus; subspecies of picipennis).

caucasicus G. Müller, 1926b: 47 (Staphylinus; subspecies of picipennis; Type locality: Caucaso: Kasbek; Svanetia).

  • — Scheerpeltz, 1933: 1400 (Staphylinus; synonym of picipennis).

  • — Coiffait, 1956e: 212 (Ocypus; subgenus Pseudocypus; subspecies of picipennis).

  • — Scheerpeltz, 1958a: 22 (Staphylinus; subgenus Pseudocypus; subspecies of picipennis; Turkey).

  • — Coiffait, 1964: 98, 104 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana, 1965d: 81 (Ocypus; subgenus Pseudocypus; subspecies of picipennis; characters; Anatolia).

  • — Smetana, 1968a: 157 (Ocypus; subgenus Pseudocypus; subspecies of picipennis; Anatolia).

  • — Smetana, 1971c: 268 (Ocypus; subgenus Pseudocypus; Iran).

  • — Jarrige, 1971: 497 (Pseudocypus; subspecies of picipennis; Iran).

  • — Coiffait, 1974: 507 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus; subspecies of picipennis).

hercegovinensis G. Müller, 1932: 80 (Staphylinus; subgenus Pseudocypus; subgenus picipennis; Type locality: Erzegovina).

  • — Scheerpeltz, 1933: 1400 (Staphylinus; subspecies of picipennis).

  • — Coiffait, 1956e: 212 (Ocypus; subgenus Pseudocypus; subspecies of picipennis).

  • — Coiffait, 1964: 98, 105 (Pseudocypus; subspecies of picipennis; characters).

  • — Coiffait, 1974: 508 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana and Davies, 2000: 44 (Ocypus; synonym of picipennis picipennis).

maritimus Coiffait, 1956e: 210, 215 (Ocypus; subgenus Pseudocypus; subspecies of picipennis; Type locality: Alpes maritimes: l'Authion).

  • — Coiffait, 1964: 98 (Pseudocypus; synonym of picipennis fallaciosus).

  • — Coiffait, 1974: 507 (Pseudocypus; synonym of picipennis fallaciosus; characters).

  • — Smetana and Davies, 2000: 45 (Ocypus; synonym of picipennis fallaciosus).

gallicus Coiffait, 1956e: 210, 215 (Ocypus; subgenus Pseudocypus; subspecies of picipennis; Type locality: Dourbes, Basse-Alpes).

  • — Coiffait, 1964: 98, 104 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana, 1967c: 199 (Ocypus; subgenus Pseudocypus; synonym of picipennis nevadensis).

  • — Coiffait, 1974: 508 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana and Davies, 2000: 45 (Ocypus; synonym of picipennis nevadensis).

asturicus Coiffait, 1956e: 211 (Ocypus; subgenus Pseudocypus; subspecies of picipennis; Type locality: Aliva, 1400 m).

  • — Coiffait, 1964: 98, 104 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana, 1967c: 199 (Ocypus; subgenus Pseudocypus; synonym of picipennis nevadensis).

  • — Coiffait, 1974: 508 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana and Davies, 2000: 45 (Ocypus; synonym of picipennis nevadensis).

cerdanicus Coiffait, 1956e: 211, 215 (Ocypus; subgenus Pseudocypus; subspecies of picipennis; Type locality: Montlouis Pyrénées-Orientales).

  • — Coiffait, 1964: 98, 104 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana, 1967c: 199 (Ocypus; subgenus Pseudocypus; synonym of picipennis nevadensis).

  • — Coiffait, 1974: 508 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana and Davies, 2000: 45 (Ocypus; synonym of picipennis nevadensis).

rufoasturicus Coiffait, 1956e: 211 (Ocypus; subgenus Pseudocypus; variety of picipennis asturicus; Type locality: Aliva, 1400 m).

  • — Coiffait, 1964: 98, 104 (Pseudocypus; variety of picipennis asturicus; characters).

  • — Smetana, 1967c: 199 (Ocypus; subgenus Pseudocypus; synonym of picipennis nevadensis).

  • — Coiffait, 1974: 508 (Pseudocypus; variety of picipennis).

rufocerdanicus Coiffait, 1956e: 211, 215 (Ocypus; subgenus Pseudocypus; variety of picipennis cerdanicus; Type locality: Montlouis).

  • — Coiffait, 1964: 98, 104 (Pseudocypus; variety of picipennis cerdanicus; p. 104 cited as sufocerdanicus; elytral color).

  • — Smetana, 1967c: 199 (Ocypus; subgenus Pseudocypus; synonym of picipennis nevadensis).

  • — Coiffait, 1974: 508 (Pseudocypus; variety of picipennis).

rufogallicus Coiffait, 1956e: 211, 215 (Ocypus; subgenus Pseudocypus; variety of picipennis gallicus; Type locality: Barre des Cévennes).

  • — Coiffait, 1964: 98, 104 (Pseudocypus; variety of picipennis gallicus; characters).

  • — Smetana, 1967c: 199 (Ocypus; subgenus Pseudocypus; synonym of picipennis nevadensis).

  • — Coiffait, 1974: 508 (Pseudocypus; variety of picipennis).

graecus Scheerpeltz, 1958c: 398 (Staphylinus; subgenus Pseudocypus; subspecies of picipennis; Type locality: Kataphigi-Gebirgsstock in der Tsumerka, 1000 bis 1600 m).

  • — Coiffait, 1964: 98, 102 (Pseudocypus; subspecies of picipennis; characters).

  • — Coiffait, 1974: 508 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana and Davies, 2000: 44 (Ocypus; synonym of picipennis picipennis).

marocanus Coiffait, 1964: 99 (Pseudocypus; subspecies of picipennis; Type locality: Maroc: Ifrane).

  • — Coiffait, 1973a: 274 (Pseudocypus; subspecies of picipennis; Morocco).

  • — Coiffait, 1974: 508 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana and Davies, 2000: 45 (Ocypus; synonym of picipennis barbarus).

pindensis Coiffait, 1964: 99 (Pseudocypus; subspecies of picipennis; Type locality: Grèce, chaîne du Pinde, Col de Metsovon, 2000 m).

  • — Smetana, 1965c: 44 (Ocypus; synonym of picipennis).

  • — Coiffait, 1974: 508 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana and Davies, 2000: 45 (Ocypus; synonym of picipennis picipennis).

rufoandorranus Coiffait, 1964: 99 (Pseudocypus; variety of picipennis andorranus; [Note: Nomen nudum (Article 15.2)]; cited from Andorre, Canillo).

  • — Smetana, 1965c: 44 (Ocypus; synonym of picipennis).

  • — Coiffait, 1974: 508 (Pseudocypus; variety of picipennis).

andorranus Coiffait, 1964: 99 (Pseudocypus; subspecies of picipennis; Type locality: Andorre, Envalira).

  • — Smetana, 1965c: 44 (Ocypus; synonym of picipennis).

  • — Coiffait, 1974: 508 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana and Davies, 2000: 44 (Ocypus; synonym of picipennis picipennis).

rufofallaciosus Coiffait, 1964: 99 (Pseudocypus; variety of picipennis fallaciosus; [Note: Nomen nudum (Article 15.2)]; cited from Sud Tyrol, Bozen).

  • — Coiffait, 1974: 507 (Pseudocypus; variety of picipennis).

rufoanatolicus Coiffait, 1964: 101 (Pseudocypus; variety of picipennis anatolicus; [Note: Nomen nudum (Article 15.2)]; cited from Turquie d'Asie: Villayet d'Isparta, Mont Davras, 1700 m).

  • — Smetana, 1965c: 44 (Ocypus; synonym of picipennis).

  • — Coiffait, 1974: 508 (Pseudocypus; variety of picipennis).

rufocaucasicus Coiffait, 1964: 101 (Pseudocypus; variety of picipennis caucasicus; [Note: Nomen nudum (Article 15.2)]; cited from Arménie Russe: Erevan).

  • — Coiffait, 1974: 507 (Pseudocypus; variety of picipennis).

anatolicus Coiffait, 1964: 101 (Pseudocypus; subspecies of picipennis; Type locality: Turquie d'Asie: Villayet d'Isparta, Arak).

  • — Smetana, 1965c: 44 (Ocypus; synonym of picipennis).

  • — Coiffait, 1974: 508 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana and Davies, 2000: 44 (Ocypus; synonym of picipennis picipennis).

bulgaricus Coiffait, 1970c: 99 (Pseudocypus; subspecies of picipennis; Type locality: Vlacuscata pestera, Vallée Cerni Vit dans la Stara Planina occidentale, Bulgarie).

  • — Smetana and Davies, 2000: 45 (Ocypus; synonym of picipennis picipennis).

rodopensis Coiffait, 1971b: 282 (Pseudocypus, cited as Pseudocipus; subspecies of picipennis; Type locality: Bulgarie: Zabârdo, Rodopes).

  • — Coiffait, 1974: 507 (Pseudocypus; subspecies of picipennis; characters).

  • — Smetana and Davies, 2000: 45 (Ocypus; synonym of picipennis picipennis).

teruelensis Coiffait, 1976c: 56 (Pseudocypus; subspecies of picipennis; Type locality: Prov. de Teruel, Frias).

  • — Smetana and Davies, 2000: 45 (Ocypus; synonym of picipennis nevadensis).

ponticus Coiffait, 1978d: 169 (Pseudocypus; subspecies of picipennis; [preoccupied]; Type locality: Col de Zigana, 2000 m).

  • — Smetana and Davies, 2000: 45 (Ocypus; synonym of picipennis picipennis).

rufoponticus Coiffait, 1978d: 170 (Pseudocypus; variety of picipennis ponticus; [Note: Nomen nudum (Article 15.2)]; cited from Col de Zigana, 2000 m).

picipennis Lacordaire, 1835, see: fulvipennis Erichson, 1840.

picipes Stephens, 1832, see: Tasgius.

picipes Nordmann, 1837, see: mus Brullé, 1832.

pindensis Coiffait, 1964, see: picipennis Fabricius, 1793.

plagiicollis Fairmaire, 1891a: cxci (Ocypus; Type locality: Tchang-Yang).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • — Bernhauer, 1922c: 233 (Staphylinus; Formosa).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus).

  • Distribution: China, Taiwan.

planipennis Aubé, 1842, see: Tasgius.

ponticus Smetana, 1968a: 158 (Ocypus; subgenus Ocypus; Type locality: Anatolia bor.: Tal v. Ardesen, Ayder, 1000–1600 m).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Turkey.

ponticus Coiffait, 1978, see: picipennis Fabricius, 1793.

primigenius G. Müller, 1923, see: syriacus Baudi, 1848.

provincialis† Oustalet, 1874: 159 (Staphylinus; subgenus Ocypus; Type locality: Aix en Provence).

  • — Scudder, 1891: 585 (Staphylinus; subgenus Ocypus; catalog; Ligurian; France).

  • — Handlirsch, 1907: 725 (Ocypus; catalog; Oligocene; France).

  • Distribution: France.

pseudoalpestris G. Müller, 1926, see: brevipennis Heer, 1839.

pseudobrevipennis Coiffait, 1964, see: chevrolatii Baudi, 1848.

pseudolens Coiffait, 1964, see: curtipennis Motschulsky, 1849.

puberulus Runde, 1835, see: aeneocephalus De Geer, 1774.

pullus Hochhuth, 1849: 121 (Ocypus; [Note: This name is older than simulator]; Type locality: Kaukasien).

  • — Eppelsheim, 1887b: 432 (Ocypus; synonym of simulator).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; synonym of simulator).

  • — Smetana and Davies, 2000: 34, 43 (Ocypus; may not be synonym of simulator so should be regarded as valid species pending study of type specimens).

  • Distribution: Caucasus.

punctatissimus Wollaston, 1864, see: fortunatarum Wollaston, 1871.

punctilinea Walker, 1858, see: Phucobius.

pyreneaus G. Müller, 1924, see: pedemontanus G. Müller, 1924.

quadraticeps Ménétriés, 1832: 143 (Staphylinus; Type locality: montagnes de Talyche).

  • — Nordmann, 1837: 62 (Staphylinus; Talych Mountains).

  • — Faldermann, 1835: 119 (Staphylinus; characters; Persia-Armenia).

  • — Hochhuth, 1849: 116 (Staphylinus; characters; notes).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1401 (Staphylinus; catalog).

  • — Coiffait, 1974: 474 (Ocypus; characters; Caucasus).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Iran, Caucasus.

quadrimaculatus Cameron, 1932: 207 (Staphylinus; subgenus Tasgius; Type locality: Assam: Khasi Hills).

  • — Scheerpeltz, 1933: 1401 (Staphylinus; catalog).

  • — Coiffait, 1982: 31 (Nudabemus; Nepal).

  • — Smetana and Davies, 2000: 32 (Ocypus; subgenus Pseudocypus).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus).

  • Distribution: India, Nepal.

quartus Schaeffer, 1766 [Note: Nomen nudum], see: picipennis Fabricius, 1793.

queinneci Coiffait, 1984e: 379 (Ocypus ?; Type locality: Nepal oriental: Milke Himal, 2000–2300 m).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: Nepal.

rambouseki G. Müller, 1925, see: nigroaeneus Sharp, 1889.

reimoseri Bernhauer, 1906: 126 (Staphylinus; subgenus Goerius; Type locality: Bjelasnica; Südgrenze Bosniens).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1401 (Staphylinus; catalog).

  • — Coiffait, 1974: 458 (Ocypus; characters; Bosnia).

  • — M. Dvořák, 1984: 61 (Ocypus; characters; Macedonia).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Macedonia, Bosnia Hercegovina.

rhaeticus Eppelsheim, 1873: 85 (Ocypus; Type locality: Tirol).

  • — Fauvel, 1876a: 71 [= 1876: 256] (Staphylinus; synonym of baudii).

  • — Ganglbauer, 1895: 431 (Staphylinus; synonym of baudii).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1401 (Staphylinus; catalog).

  • — Lohse, 1964: 199 (Ocypus; subgenus Goerius; characters; central Europe).

  • — Horion, 1965: 218 (Staphylinus; subgenus Goerius; Italy; Switzerland; Austria).

  • — Coiffait, 1974: 451 (Ocypus; characters; Alps).

  • — Lucht, 1987: 108 (Ocypus; checklist; Austria).

  • — Pilon and Zanetti, 1991: 68 (Ocypus; Italy).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; Italy).

  • — Pilon, 1998: 86 (Ocypus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus).

  • Distribution: Austria, Italy, Switzerland.

baudii Fauvel, 1867: LV (Ocypus; [preoccupied]; Type locality: Alpes du Valais, mont Rosa).

  • — Fauvel, 1874: 183 [= 1874a: 409] (Staphylinus; characters; France).

  • — Mulsant and Rey, 1876: 277 [= 1877: 133] (Ocypus; subgenus Goerius; characters; France).

  • — Fiori, 1894: 90 (Ocypus; characters; Piemonte).

  • — Ganglbauer, 1895: 431 (Staphylinus; subgenus Goerius; characters; Europe).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; synonym of rhaeticus).

  • — G. Müller, 1923: 138 (Staphylinus; subgenus Goerius; characters; Italy).

  • — G. Müller, 1926: 11 (Staphylinus; subgenus Goerius; notes).

  • — Scheerpeltz, 1933: 1401 (Staphylinus; synonym of rhaeticus).

  • — Lohse, 1964: 199 (Ocypus; synonym of rhaeticus).

  • — Coiffait, 1974: 451 (Ocypus; synonym of rhaeticus).

robustus Coiffait, see: milleri Quedenfeldt, 1882.

rodopensis Coiffait, 1971, see: picipennis Fabricius, 1793.

rossii Jarrige, 1954, see: sericeicollis Ménétriés, 1832.

rubidus Verhoeff, 1890, see: Tasgius.

rubripennis Reiche and Saulcy, 1856, see: Tasgius.

rufipes Eppelsheim, 1884, see: Tasgius.

rufipes Verhoeff, 1890, see: Tasgius.

rufoandorranus Coiffait, 1964 [Note: Nomen nudum], see: picipennis Fabricius, 1793.

rufoantolicus Coiffait, 1964 [Note: Nomen nudum], see: picipennis Fabricius, 1793.

rufoasturicus Coiffait, 1956, see: picipennis Fabricius, 1793.

rufocaucasicus Coiffait, 1964 [Note: Nomen nudum], see: picipennis Fabricius, 1793.

rufocerdanicus Coiffait, 1956, see: picipennis Fabricius, 1793.

rufofallaciosus Coiffiat, 1964 [Note: Nomen nudum], see: picipennis Fabricius, 1793.

rufogallicus Coiffait, 1956, see: picipennis Fabricius, 1793.

rufoponticus Coiffait, 1978 [Note: Nomen nudum], see: picipennis Fabricius, 1793.

rugifrons Runde, 1835, see: ophthalmicus Scopoli, 1763.

sabi Naomi, 1992: 141 (Ocypus; subgenus Protocypus; Type locality: Japan: Yamanashi Pref.: Hirogawara, Nakakoma-gun).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan.

saevus Solodovnikov, 2000: 321 (Ocypus; Type locality: Western Georgia: Egrissky range, Mt. Khvira, 1000-1300 m).

  • Distribution: Georgia.

saulcyi Reiche, 1857, see: Tasgius.

scabrosus Curtis, 1839, see: Staphylinus.

schatzmayri G. Müller, 1923, see: obscuroaeneus Fairmaire, 1852.

scutiger Sharp, 1889: 110 (Ocypus; Type locality: Chiuzenji).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1401 (Staphylinus; catalog).

  • — Nakane, 1963a: 93 (Ocypus; characters; habitus photograph; Japan).

  • — Frank, 1982: 38 (Ocypus; list of parasites).

  • — Yuh, Paik, Kwon, and Lee, 1985: 245 (Ocypus; Korea).

  • — Naomi, 1992a: 55 (Ocypus; subgenus Protocypus; characters; notes; Japan).

  • — Kanehisa, Tsumuki, and Kawazu, 1994 (Ocypus; chemical secretions).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan, Korea.

semenowi Reitter, 1887: 213 (Ocypus; Type locality: zwischen dem Flusse Bytchu und dem Blauen Flusse, im Thale des Bytchu und in den Schluchten Diao-tchu und Bö-tchu, der Wasserscheide zwischen den oberen Theilen des Gelben und Blauen Flusses, 13 bis 14,000 F.).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; synonym of angustulus).

  • — Coiffait, 1974: 461 (Ocypus; cited as semenovi; cited as valid species; characters; central Asia).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus).

  • Distribution: China.

semialatus G. Müller, 1904, see: nitens Schrank, 1781.

semipolitus Holme, 1842, see: fuscatus Gravenhorst, 1802.

septentrionalis Watanabe, 1984: 139 (Ocypus; subgenus Protocypus; Type locality: Japan: Honshu: Yamagata Pref.: Ishikorobi-zawa on Iide).

  • — Naomi, 1992a: 57 (Ocypus; synonym of scutiger).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus; cited as valid species).

  • Distribution: Japan.

sericeicollis Ménétriés, 1832: 143 (Staphylinus; [Note: This name is older than rossii]; Type locality: montagnes de Lenkoran).

  • — Nordmann, 1837: 70 (Staphylinus; Lenkoran).

  • — Faldermann, 1835: 120 (Staphylinus; characters; Persia-Armenia).

  • — Erichson, 1840: 413 (Ocypus; synonym of cupreus).

  • — Hochhuth, 1849: 123 (Ocypus; synonym of picipennis).

  • — Kraatz, 1857d: 560 (Ocypus; synonym of cupreus).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; synonym of picipennis).

  • — Scheerpeltz, 1933: 1393 (Staphylinus; variety of cupreus).

  • — Coiffait, 1956e: 213 (Ocypus; subgenus Pseudocypus; subspecies of cupreus; characters).

  • — Scheerpeltz, 1956c: 1101 (Staphylinus; variety of cupreus; Cyprus).

  • — Focarile, 1964: 61 (Staphylinus; variety of cupreus; Italy).

  • — Coiffait, 1974: 513 (Pseudocypus; subspecies of cupreus; Greece; Asia Minor; Caucasus; Rhodes; Syria).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus; valid name).

  • Distribution: Algeria, Europe, Turkey, Azerbaijan, Iran, Syria.

cupreus Rossi, 1790: 248 (Staphylinus; [preoccupied]; Type locality: Etruria).

  • — Olivier, 1795:(42): 16 (Staphylinus; characters; France, Italy).

  • — Gravenhorst, 1806: 113 (Staphylinus; synonym of aenoecephalus).

  • — Gyllenhal, 1810: 292 (Staphylinus; synonym of aeneocephalus De Geer).

  • — C. Sahlberg, 1830: 311 (Staphylinus; synonym of aeneocephalus).

  • — Erichson, 1839a: 443 (Ocypus; characters; Germany).

  • — Heer, 1839: 255 (Staphylinus; characters; Switzerland).

  • — Erichson, 1840: 412 (Ocypus; characters; Europe).

  • — Kolenati, 1846: 18 (Ocypus; Transcaucasiae).

  • — P. Lucas, 1846: 111 (Ocypus; notes; Algeria).

  • — Jacobsen, 1849: 685 (Ocypus; characters; Denmark).

  • — Hochhuth, 1849: 124 (Ocypus; characters; Caucasus).

  • — Kiesenwetter, 1851a: 414 (Ocypus; Pyrenees).

  • — Küster, 1853: 62 (Ocypus; characters; Europe).

  • — Hardy, 1851: 23 (Ocypus; subgenus Goerius; notes; England).

  • — Rosenhauer, 1856: 70 (Ocypus; P. Real; Algeciras; Yunquera).

  • — Fairmaire and Laboulbène, 1856: 511 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 560 (Ocypus; characters; Germany).

  • — Redtenbacher, 1857: 188 (Ocypus; characters; Austria).

  • — Kraatz, 1858a: 61 (Ocypus; Greece).

  • — Thomson, 1860: 149 (Ocypus; characters; Scandinavia).

  • — Baudi, 1870: 384 (Ocypus; characters; Sardinia; France).

  • — Fauvel, 1874: 191 [= 1874a: 417] (Staphylinus; synonym of aeneocephalus).

  • — Redtenbacher, 1874: 207 (Ocypus; characters; Austria).

  • — J. Sahlberg, 1876: 11 (Ocypus; Finland).

  • — Fowler, 1888: 256 (Ocypus; characters; Britain).

  • — Ganglbauer, 1895: 434 (Staphylinus; synonym of aeneocephalus).

  • — Bernhauer, 1910b: 257 (Staphylinus; valid species; characters; Mediterranean region).

  • — Sharp and Muir, 1912: 498 (Ocypus; characters of male genitalia).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; subgenus Ocypus; catalog).

  • — G. Müller, 1925: 47 (Staphylinus; subgenus Pseudocypus; notes).

  • — G. Müller, 1926: 19 (Staphylinus; subgenus Pseudocypus; characters; notes).

  • — Scheerpeltz, 1933: 1392 (Staphylinus; catalog).

  • — Méquignon, 1933: 229 (Staphylinus; distinguishing characters).

  • — Tottenham, 1932 (Ocypus; distinguishing characters).

  • — Coiffait, 1956e: 213, 214 (Ocypus; subgenus Pseudocypus; [Note: Although cupreus Rossi and cupreus Geoffroy are primary homonyms in Staphylinus, Coiffait regarded the homonymic name change by Jarrige as unnecessary because the two names in question had not been in the same genus for more than 100 years; he suggested that cupreus Rossi be conserved]; characters).

  • — Scheerpeltz, 1961d: 131 (Staphylinus; subgenus Pseudocypus; Italy).

  • — Lohse, 1964: 200 (Ocypus; subgenus Pseudocypus; characters; central Europe).

  • — Scheerpeltz, 1965b: 404 (Staphylinus; Sicily).

  • — Scheerpeltz, 1965c: 509 (Staphylinus; Sicily).

  • — Coiffait, 1968c: 505 (Pseudocypus; Corsica).

  • — Coiffait, 1974: 512 (Pseudocypus; characters; Europe; England).

  • — Pope, 1977: 30 (Staphylinus; synonym of fortunatarum).

  • — H. Franz, 1987: 69 (Pseudocypus; Cyprus).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; Italy).

  • — Pilon, 1998: 88 (Ocypus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 35, 45 (Ocypus; synonym of sericeicollis).

confinis Stephens, 1832: 211 (Goerius; Type locality: Essex; London).

  • — Stephens, 1835: 435 (Tasgius; moved from Goerius).

  • — Hardy, 1851: 23 (Staphylinus; synonym of fuscatus).

  • — Gemminger and Harold, 1868: 582 (Ocypus; synonym of cupreus).

  • — Fauvel, 1869: 488 (Ocypus; synonym of cupreus).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of cupreus).

  • — Smetana and Davies, 2000: 45 (Ocypus; synonym of sericeicollis).

angustatus Stephens, 1832: 212 (Ocypus; Type locality: Suffolk; Yorkshire; London).

  • — Stephens, 1839: 388 (Ocypus; characters; England).

  • — Hardy, 1851: 24 (Staphylinus; synonym of morio).

  • — Gemminger and Harold, 1868: 582 (Ocypus; synonym of cupreus).

  • — Fauvel, 1869: 488 (Ocypus; synonym of cupreus).

  • — Ganglbauer, 1895: 434 (Staphylinus; synonym of aeneocephalus).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of cupreus).

  • — Smetana and Davies, 2000: 45 (Ocypus; synonym of sericeicollis).

metallicus O. Costa, 1839: 48 (Staphylinus; [preoccupied]; Type locality: cratere del Vesuvio).

  • — Ganglbauer, 1895: 434 (Staphylinus; synonym of aeneocephalus).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of cupreus).

rossii Jarrige, 1954a: 164 (Staphylinus; subgenus Pseudocypus; replacement name for cupreus Rossi).

  • — Coiffait, 1974: 513 (Pseudocypus; synonym of cupreus).

  • — Smetana and Davies, 2000: 35, 45 (Ocypus; synonym of sericeicollis).

fulvicupreus Coiffait, 1956e: 213 (Ocypus; subgenus Pseudocypus; subspecies of cupreus; Type locality: Chypre).

  • — Coiffait, 1970e: 62 (Pseudocypus; variety of cupreus; Cyprus).

  • — Coiffait, 1974: 513 (Pseudocypus; subspecies of cupreus; Cyprus).

  • — Smetana and Davies, 2000: 35, 45 (Ocypus; synonym of sericeicollis).

picicupreus Coiffait, 1974: 513 (Pseudocypus; variety of cupreus; [Note: Nomen nudum (Article 15.2)]; cited from Saint Maixent, Deux Sèvres).

sericeomicans Bernhauer, 1931: 2 (Staphylinus; subgenus Ocypus; Type locality: Szetschwan: Kia-ting).

  • — Scheerpeltz, 1933: 1401 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus).

  • Distribution: China.

sericeus Marsham, 1802, see: picipennis Fabricius, 1793.

sericeus Motschulsky, 1858, see: fortunatarum Wollaston, 1871.

serotinus Ádám, 1992: 5 (Pseudocypus; Type locality: Hungary, Bükki N. P., Szilvásvárad, Keskeny-rét, 850 m).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus).

  • Distribution: Hungary.

siculus Aubé, 1842, see: Tasgius.

siculus Stierlin, 1864, see: Tasgius.

sikkimensis Coiffait, 1985b: 63 (Ocypus; Type locality: Inde: Sikkim, Gnatang).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: India.

silensis Fiori, 1894, see: italicus Aragona, 1830.

similis: Fabricius, 1793 [Note: Nomen nudum as published by Fabricius], see: nitens Schrank, 1781.

simulator Eppelsheim, 1878a: 420 (Ocypus; Type locality: Acarnanien; Asia minor).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; synonym of pullus).

  • — G. Müller, 1926: 14 (Staphylinus; subgenus Goerius; characters; notes).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; catalog).

  • — Schatzmayr, 1943: 115 (Staphylinus; Albania).

  • — Coiffait, 1974: 472 (Ocypus; characters; Greece; Macedonia; Dalmatia; Corfu).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Greece, Macedonia, Albania, Croatia, Caucasus, Turkey.

smetanai Naomi, 1992a: 55 (Ocypus; subgenus Protocypus; Type locality: Japan: Honshu: Iwate Pref.: Tohoku district: Mt. Hayachine).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan.

solarii G. Müller, 1923: 138 (Staphylinus; subgenus Goerius; Type locality: Nava, Liguria occ.).

  • — G. Müller, 1925a: 26 (Staphylinus; characters).

  • — G. Müller, 1926: 10 (Staphylinus; subgenus Goerius; characters; notes).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; catalog).

  • — Jarrige, 1947a: 155 (Goerius; characters; France).

  • — Coiffait, 1956e: 203, 206 (Ocypus; subgenus Goerius; characters).

  • — Coiffait, 1974: 452 (Ocypus; characters; France; Italy).

  • — M. Dvořák, 1984: 61 (Ocypus; Italy).

  • — Wittwer, 1993: 250 (Ocypus; Switzerland).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; Italy).

  • — Pilon, 1998: 87 (Ocypus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Italy, Switzerland, France.

stastnyi M. Dvořák, 2000: 333 (Ocypus; Type locality: Kasachstan sept. or., Altaj mer., Amuldaus-kul, 2300 m).

  • Distribution: Kazakhstan.

strigatus Nordmann, 1837, see: aeneocephalus De Geer, 1774.

subaenescens Wollaston, 1864: 567 (Ocypus; Type locality: Grand Canary; Hierro; Teneriffe).

  • — Wollaston, 1865: 489 (Ocypus; habitat; notes; Madeira Islands; Canary Islands).

  • — Fauvel, 1902b: 105 (Staphylinus; variety of aeneocephalus).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of cupreus).

  • — Dodero, 1925: 5 (Staphylinus; subgenus Goerius; Libya).

  • — G. Müller, 1926a: 31 (Staphylinus; subgenus Pseudocypus; characters; notes).

  • — Gridelli, 1930: 76 (Staphylinus; Libya).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; catalog).

  • — Coiffait, 1954: 166 (Pseudocypus; Canary Islands).

  • — Jarrige, 1954a: 163 (Staphylinus; subgenus Pseudocypus; characters; Canaries).

  • — Coiffait, 1956e: 213 (Ocypus; subgenus Pseudocypus; characters).

  • — Coiffait, 1974: 515 (Pseudocypus; characters; Canary Islands).

  • — Assing, 1999a: 346 (Pseudocypus; Canary Islands).

  • — Assing and Wunderle, 1999: 437 (Pseudocypus; Canary Islands).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus).

  • Distribution: Canary Islands.

caeruleipennis Coiffait, 1954: 166 (Pseudocypus; aberration of subaenescens; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Hierro).

  • — Coiffait, 1956e: 213 (Ocypus; subgenus Pseudocypus; cited as cyaneipennis; [Note: The characters and locality cited with cyaneipennis are the same as those given for caeruleipennis and both were listed as aberrations of subaenescens so I assume that cyaneipennis is a lapsus for caeruleipennis. Both names are unavailable]; aberration of subaenescens; characters).

  • — Coiffait, 1974: 515 (Pseudocypus; variety of subaenescens; cited as cyaneipennis; [Note: The characters and locality cited with cyaneipennis are the same as those given for caeruleipennis and both were listed as aberrations of subaenescens so I assume that cyaneipennis is a lapsus for caeruleipennis]).

subaeneus Roth, 1851, see: Platydracus.

subapterus G. Müller, 1923 [Note: Nomen nudum], see: nitens Schrank, 1781.

subcyaneus Heer, 1839, see: ophthalmicus Scopoli.

subpunctatus Gyllenhal, 1827, see: fuscatus Gravenhorst, 1802.

subtilis Tikhomirova, 1973: 171 (Ocypus; Type locality: Suputinskii zapovednik).

  • — Tikhomirova, 1973a: 186 (Ocypus; subgenus Pseudocypus; synonym of orientalis ? [Note: Since this name is cited as a questionable synonym, I have continued to list it as valid]).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Aulacocypus).

  • Distribution: Russia.

svozili M. Dvořák, 1984: 64 (Ocypus; Type locality: UdSSR: Kasachstan: Alma Arasan, Alma-Ata env.).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus).

  • Distribution: Kazakhstan.

sylvaticus Wollaston, 1865: 489, 72 (Ocypus; Type locality: Gomera: Hermigua).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; catalog).

  • — Coiffait, 1954: 166 (Goerius; Canary Islands).

  • — Jarrige, 1954a: 163 (Staphylinus; subgenus Goerius; characters).

  • — Coiffait, 1974: 413 (Atlantogoerius; characters; Canary Islands).

  • — Assing, 1999a: 346 (Atlantogoerius; Canary Islands).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus).

  • Distribution: Canary Islands.

syriacus Baudi, 1848: 124 (Ocypus; Type locality: Syria).

  • — Baudi, 1870: 384 (Ocypus; characters).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; subgenus Ocypus; catalog).

  • — G. Müller, 1923: 138 (Staphylinus; subgenus Goerius; characters; Syria).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; catalog).

  • — Coiffait, 1956e: 202 (Ocypus; subgenus Goerius; characters).

  • — Coiffait, 1974: 480 (Ocypus; characters; Syria; Lebanon).

  • — M. Dvořák, 1984: 65 (Ocypus; Cyprus).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Greece, Cyprus, Lebanon, Syria, Turkey, Armenia, Ukraine.

ludovicae Mulsant and Godart, 1855: 263 [= 1855a: 163] (Staphylinus; Type locality: Crimée).

  • — Fauvel, 1875a: XXVII [= 1875b: 229] (Staphylinus; synonym of syriacus).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; synonym of syriacus).

  • — Coiffait, 1974: 480 (Ocypus; synonym of syriacus).

baudii Peyron, 1858: 424 (Ocypus; Type locality: Not cited, presumably Tarsous).

  • — Fauvel, 1869: 487 (Ocypus; synonym of syriacus).

  • — Fauvel, 1875a: XXVII [= 1875b: 229] (Staphylinus; synonym of syriacus).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; synonym of syriacus).

  • — Coiffait, 1974: 480 (Ocypus; synonym of syriacus).

primigenius G. Müller, 1923: 138 (Staphylinus; subgenus Goerius; subspecies of syriacus; Type locality: Armenia).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; subspecies of syriacus).

  • — Coiffait, 1974: 480 (Ocypus; subspecies of syriacus; Armenia).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus; subspecies of syriacus).

tauricus G. Müller, 1932, see: mus Brullé, 1832.

tenebricosus Gravenhorst, 1846: 95 (Staphylinus; Type locality: Schlesien; Ungarn).

  • — Fairmaire and Laboulbène, 1856: 509 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 391 cited brachypterus Fairmaire and Laboulbène as a synonym of tenebricosus; Fairmaire and Laboulbène did not describe the species as new, they attributed it to Brullé]; characters; France).

  • — Eppelsheim, 1887b: 431 (Ocypus; synonym of olens).

  • — Ganglbauer, 1895: 430 (Staphylinus; subgenus Goerius; characters; middle Europe).

  • — Reitter, 1909: 121 (Staphylinus; subgenus Goerius; characters; Germany).

  • — Kryger, 1913: 380 (Ocypus; [Note: Misidentification: Scheerpeltz, 1933: 1403 cited olens Kryger as a synonym of tenebricosus but although Kryger did not include an author's name with olens he also did not indicate the name to be newly proposed nor did he publish characters with it]; notes).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; subgenus Ocypus; catalog).

  • — G. Müller, 1923: 137 (Staphylinus; subgenus Goerius; Italy).

  • — G. Müller, 1924: 25 (Staphylinus; subgenus Goerius; characters is key).

  • — G. Müller, 1926: 7 (Staphylinus; subgenus Goerius; notes).

  • — Portevin, 1929: 352 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; catalog).

  • — Aleksandrov, 1934: 153 (Staphylinus; China).

  • — Coiffait, 1956e: 203 (Ocypus; subgenus Goerius; characters).

  • — Smetana, 1958: 284 (Ocypus; subgenus Ocypus; characters; ecology; Czechoslovakia).

  • — Szujecki, 1963a: 34 (Ocypus; Poland).

  • — Lohse, 1964: 199 (Ocypus; subgenus Goerius; characters; central Europe).

  • — Horion, 1965: 219 (Staphylinus; subgenus Goerius; Italy; Slovenia; Dalmatia; Croatia; Bosnia Hercegovina; Serbia; Albania; Romania; Hungary; Czechoslovakia; Poland; Austria; Germany).

  • — Coiffait, 1974: 454 (Ocypus; characters; Europe).

  • — Tóth, 1984: 28 (Ocypus; characters; Hungary).

  • — Lucht, 1987: 108 (Ocypus; checklist; Germany; Poland; Czechoslovakia; Austria; France).

  • — Wittwer, 1993: 250 (Ocypus; Switzerland).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Ocypus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; Italy).

  • — Pilon, 1998: 87 (Ocypus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Europe.

micropterus Redtenbacher, 1849: 697 (Ocypus; Type locality: Wien).

  • — Kraatz, 1857d: 553 (Ocypus; synonym of brachypterus).

  • — Fauvel, 1865a: 294 [= 1865b: 44] (Ocypus; habitat; France).

  • — Fauvel, 1867: LV (Ocypus; notes).

  • — Ganglbauer, 1895: 430 (Staphylinus; synonym of tenebricosus).

  • — Bernhauer and Schubert, 1914: 391 (Staphylinus; synonym of tenebricosus).

  • — Smetana, 1958: 284 (Ocypus; synonym of tenebricosus).

  • — Coiffait, 1974: 454 (Ocypus; synonym of tenebricosus).

aureotomentosus Bernhauer, 1900: 46 (Ocypus; variety of brachypterus; Type locality: Croatien, Fuzine; Balkangebirge).

  • — Bernhauer and Schubert, 1914: 391 (Staphylinus; synonym of tenebricosus).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; subspecies of tenebricosus).

  • — Smetana, 1958: 284 (Ocypus; synonym of tenebricosus).

  • — Lohse, 1964: 199 (Ocypus; aberration of tenebricosus).

  • — Coiffait, 1974: 454 (Ocypus; variety of tenebricosus).

  • — Tóth, 1984: 28 (Ocypus; variety of tenebricosus).

depolii G. Müller, 1924: 23, 25 (Staphylinus; subgenus Goerius; Type locality: Liburnia montana).

  • — G. Müller, 1926: 8 (Staphylinus; subgenus Goerius; notes).

  • — Scheerpeltz, 1933: 1393 (Staphylinus; catalog).

  • — Coiffait, 1974: 455 (Ocypus; characters; Istria).

  • — M. Dvořák, 1984: 61 (Ocypus; characters; Serbia).

  • — Tóth, 1984: 27 (Ocypus; characters; Hungary).

  • — Pilon, 1998: 99 (Ocypus; synonym of tenebricosus).

  • — Smetana and Davies, 2000: 43 (Ocypus; synonym of tenebricosus).

tenuipennis Erichson, 1840, see: brevipennis Heer, 1839.

teruelensis Coiffait, 1976, see: picipennis Fabricius, 1793.

testaceipes Fairmaire, 1887, see: Staphylinus.

tomentosus Baudi, 1870, see: orientis Smetana and Davies, 2000.

torvus Smetana, 1965c: 41 (Ocypus; subgenus Ocypus; Type locality: Anatolia bor., Çamlik bei Rize, 1750 m).

  • — Smetana, 1968a: 157 (Ocypus; subgenus Ocypus; notes; Anatolia).

  • — Coiffait, 1974: 482 (Ocypus; characters; Turkey).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Turkey.

transadriaticus G. Müller, 1926, see: mus Brullé, 1832.

trapezensis Coiffait, 1964: 86 (Ocypus; Type locality: Turquie d'Asie, environs de Trabzon).

  • — Smetana, 1968b: 88 (Ocypus; characters; Anatolia).

  • — Coiffait, 1974: 455 (Ocypus; characters; Turkey).

  • — Coiffait, 1978d: 164 (Ocypus; Turkey).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Turkey.

tristis Fabricius, 1793, see: picipennis Fabricius, 1793.

turcicus Bernhauer, 1923a: 125 (Staphylinus; subgenus Ocypus; Type locality: Kolat).

  • — Scheerpeltz, 1933: 1404 (Staphylinus; catalog).

  • — Smetana, 1965c: 28 (Ocypus; subgenus Ocypus; characters; notes; Anatolia).

  • — Smetana, 1965d: 79 (Ocypus; subgenus Ocypus; Anatolia).

  • — Coiffait, 1974: 457 (Ocypus; characters; Turkey).

  • — Smetana and Davies, 2000: 43 (Ocypus; subgenus Matidus).

  • Distribution: Turkey.

umbricola Wollaston, 1864: 566 (Ocypus; Type locality: Teneriffe: Taganana; Agua Garcia; the Agua Mansa; Yeod el Alto; Barao do Castello de Paiva).

  • — Wollaston, 1865: 488 (Ocypus; habitat; Canary Islands).

  • — Fauvel, 1902b: 104 (Staphylinus; subgenus Ocypus; Canary Islands).

  • — Bernhauer and Schubert, 1914: 391 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1404 (Staphylinus; catalog).

  • — Uyttenboogaart, 1937: 81 (Staphylinus; notes; Canary Islands).

  • — Coiffait, 1954: 166 (Goerius, cited as Geerius; Canary Islands).

  • — Jarrige, 1954a: 162 (Staphylinus; subgenus Goerius; characters; Canaries).

  • — Coiffait, 1974: 411 (Atlantogoerius; characters; Canary Islands).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus).

  • Distribution: Canary Islands.

unicolor Herbst, 1784, see: olens O. Müller, 1764.

uralensis Mannerheim, 1830, see: fuscatus Gravenhorst, 1802.

vagans Heer, 1839, see: fulvipennis Erichson, 1840.

vesubiensis Coiffait, 1964, see: chevrolatii Baudi, 1848.

viaticus Gistel, 1857: 72 (Goerius; [Note: Nomen dubium]; Type locality: Tyrolis).

  • Distribution: Austria.

wasmanni Bernhauer, 1920d: 184 (Staphylinus; Type locality: Darjeeling: Sikkim).

  • — Cameron, 1932: 205 (Staphylinus; subgenus Tasgius; characters; India).

  • — Scheerpeltz, 1933: 1404 (Staphylinus; catalog).

  • — Coiffait, 1982a: 233 (Nudabemus, cited as wasmani and wassmani; Nepal).

  • — Smetana and Davies, 2000: 42 (Ocypus; subgenus Ocypus).

  • Distribution: India, Nepal.

weisei Harold, 1877: 344 (Ocypus; Type locality: Tokio).

  • — Sharp, 1889: 109 (Ocypus; Japan).

  • — Bernhauer and Schubert, 1914: 391 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1404 (Staphylinus; catalog).

  • — Nakane, 1963a: 93 (Ocypus; characters; habitus photograph; Japan).

  • — Yuh, Paik, Kwon, and Lee, 1985: 246 (Ocypus; Korea).

  • — J. Li, 1993: 38 (Ocypus; China).

  • — Hayashi, 1993b: 298 (Agelosus).

  • — Watanabe, 1995c: 75 (Ocypus; China).

  • — Smetana and Davies, 2000: 31, 42 (Ocypus; subgenus Ocypus).

  • Distribution: Japan, Korea, China.

yamato Naomi, 1992b: 9 (Ocypus; subgenus Protocypus; Type locality: Japan: Nara Pref.: Mt. Ohdaigahara).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan.

yeti M. Dvořák, 2000: 334 (Ocypus; Type locality: China: Tibet or., Nyingchi environ sept.-or., 3700–4500 m, 29°37′N, 94°40′E).

  • Distribution: China.

yoroi Naomi, 1992a: 59 (Ocypus; subgenus Protocypus; Type locality: Japan: Honshu: Niigata Pref.: Chubu district, Sasagamine, Mt. Myoko).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan.

yuinus Naomi, 1992a: 60 (Ocypus; subgenus Protocypus; Type locality: Japan: Honshu: Shizuoka Pref.: Chubu district, Yui).

  • — Smetana and Davies, 2000: 45 (Ocypus; subgenus Pseudocypus).

  • Distribution: Japan.

zimmermanni Korge, 1965, see: chevrolatii Baudi, 1848.

Ontholestes [31 species; Neotropical, Nearctic, Palaearctic, Ethiopian, and Oriental Regions]

Ontholestes Ganglbauer, 1895: 417 (species included: murinus, nebulosus; [Note: The name was proposed for Leistotrophus as misidentified by Kraatz who attributed it to Perty]). Type species: Staphylinus murinus Linné, fixed by subsequent designation by R. Lucas, 1920: 459.

  • — Kraatz, 1857d: 532 (characters; [Note: Misidentification of Leistotrophus according to Ganglbauer, 1895]).

  • — Thomson, 1858: 29 (cited as Schizochilus; [Note: Thomson's use of Schizochilus is a misidentification of Ontholestes; Tottenham, 1949: 375, cited it as a synonym of Ontholestes and with a type species]; characters).

  • — Thomson, 1859: 23 (cited as Schizochilus; [Note: Thomson's use of Schizochilus is a misidentification of Ontholestes; Tottenham, 1949: 375, cited it as a synonym of Ontholestes and with a type species]; characters; type species).

  • — Heyden, 1880: 71 (cited as Leistotrophus; list of species of Siberian region).

  • — Heyden, Reitter, and Weise, 1891: 101 (cited as Leisototrophus; list of species of Europe and Caucasus).

  • — Heyden, 1893: 41 (cited as Leistotrophus; list of species of Siberian region).

  • — Heyden, 1896: 32 (list of species of Siberian region).

  • — Heyden, Reitter, and Weise, 1906: 171 (list of species of Europe and Caucasus).

  • — Reitter, 1909: 116, 117 (characters; key to species).

  • — Petri, 1912: 76 (list of species of Siebenbürgen).

  • — Bernhauer and Schubert, 1914: 393 (world catalog; 18 species).

  • — Johansen, 1914: 374 (characters; key to species).

  • — Leng, 1920: 109 (catalog of North American species).

  • — Gridelli, 1924f: 206 (key to species).

  • — Winkler, 1925: 386 (catalog for Palaearctic region).

  • — Cameron, 1925: 67 (catalog of Indian species).

  • — Porta, 1926: 94 (characters; key to species of Italy).

  • — Portevin, 1929: 348 (characters; key to French species).

  • — Roubal, 1930: 385 (catalog; Slovakia).

  • — Cameron, 1932: 209 (characters; key to species of British India).

  • — Joy, 1932: 123 (key to British species).

  • — Scheerpeltz, 1933: 1405 (world catalog supplement).

  • — Normand, 1935: 372 (list of species of Tunisia).

  • — Kirshenblat, 1936 (revision of and key to Palaearctic species).

  • — Tottenham, 1940: 49 (type species: murinus).

  • — Paulian, 1941: 248 (larval characters).

  • — Blackwelder, 1944: 139 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Jeannel and Paulian, 1945: 92 (larval characters).

  • — Kloet and Hincks, 1945: 167 (list of British species; type species: murinus).

  • — Tottenham, 1949: 375, 421 (type species: murinus Linné; notes; checklist of British species).

  • — V. Hansen, 1952: 108 (characters; key to Danish species).

  • — Blackwelder, 1952: 275 (type species).

  • — Smetana, 1955a: 283 (notes; key to central European species).

  • — Coiffait, 1956e: 189 (subgenus of Abemus; characters in key; notes; key to French species).

  • — Hatch, 1957: 179 (characters; species of Pacific Northwest).

  • — Kocher, 1958: 155 (checklist of species; Morocco).

  • — Smetana, 1958: 305 (characters; type species: murinus; key to Czechoslovakian species).

  • — Smetana, 1959d (characters; notes on history of name, species groups; key to species).

  • — Ferreira, 1962a: 18 (list of Portuguese species).

  • — Székessy, 1963: 55 (characters).

  • — Lohse, 1964: 192 (characters; key to central European species).

  • — Moore, 1965: 98, 101 (characters; notes).

  • — Pototskaia, 1967: 64 (larval characters).

  • — Scheerpeltz, 1968a: 85 (catalog of Austrian species).

  • — Kasule, 1970: 64, 74 (larval characters).

  • — Tikhomirova, 1973a: 187 (checklist of species of USSR).

  • — Coiffait, 1974: 368 (adult and larval characters; key to species of western Palaearctic region).

  • — Moore and Legner, 1975: 21 (catalog; North America north of Mexico).

  • — Pope, 1977: 30 (list of British species).

  • — Topp, 1978: 311 (larval characters).

  • — Frank, 1979a (some larval characters and their use in classification).

  • — Moore and Legner, 1979: 27 (characters; notes).

  • — Muona, 1979: 15 (list of Scandinavian species).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 89 (catalog; Poland).

  • — Smetana, 1981d: 126 (key to species of North America).

  • — Uhlig and Vogler, 1981: 119 (list of some species collected in Germany; habitat and collecting notes).

  • — Hammond, 1984: 206 (checklist of species of Borneo).

  • — Tóth, 1984: 11 (characters; habitus; key to species of Hungary).

  • — Shibata, 1984: 96 (checklist of species of Japan).

  • — Muona and Viramo, 1986: 13 (list of species of northeastern Finland).

  • — Segers, 1986: 47 (checklist of species; Belgium).

  • — Dettner, 1987: 30 (defensive glands).

  • — Drugmand 1987b (comments on distribution of Belgian species).

  • — Ádám, 1987: 148 (list of two species with collecting notes; Hungary).

  • — Angelini, 1991: 192 (list of species from high elevations of Calabria, Italy).

  • — Campbell and Davies, 1991: 120 (checklist of species; Canada).

  • — Silfverberg, 1992: 17 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Welch, 1993: 229 (ovariole structure and development).

  • — Smetana, 1993b: 49 (list of species of Czech Republic and Slovakia).

  • — Kanehisa, Tsumuki, and Kawazu, 1994 (chemical secretions).

  • — Ciceroni and Zanetti, 1995: 29 (list of species of Italy).

  • — M. Hansen, 1996: 107 (list of Danish species).

  • — Downie and Arnett, 1996: 386 (characters; key to species of Northeastern North America).

  • — Lundgren, 1998: 51 (list of species from Florida; records for other States listed).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (list of species collected in 1997; Denmark).

  • — Ryabukhin, 1999: 67 (represented in northeastern Russia by unidentified specimen).

  • — Smetana and Davies, 2000: 19, 22, 38 (characters; checklist of species of north temperate region).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 394 (2 Nearctic species; notes; characters in key).

Trichoderma Stephens, 1833a: 96 (species included: nebulosa, hybrida, murina, pubescens; preoccupied by Fleming, 1822). Type species: Staphylinus murina Linné, fixed by subsequent designation by Westwood, 1838a: 15.

  • — Stephens, 1835: 435 (characters).

  • — Westwood, 1838a: 15 (characters; type species).

  • — Erichson, 1840: 404 (synonym of Ocypus).

  • — Stephens, 1839: 384 (characters).

  • — Lacordaire, 1854: 79 (synonym of Ocypus).

  • — Kraatz, 1857d: 532 (synonym of Leistotrophus).

  • — Schaum, 1859: 26 (synonym of Leistotrophus).

  • — Thomson, 1859: 23 (type species: pubescens).

  • — Thomson, 1860: 141 (misidentified as Schizochilus according to Coiffait, 1974: 368; characters).

  • — Thomson, 1860: 142 (characters).

  • — Reitter, 1909: 118 (subgenus of Staphylinus; characters).

  • — Petri, 1912: 74 (subgenus of Staphylinus; list of species of Siebenbürgen).

  • — Bernhauer and Schubert, 1914: 374 (subgenus of Staphylinus).

  • — Cameron, 1932: 180 (subgenus of Staphylinus).

  • — Scheerpeltz, 1933: 1381 (subgenus of Staphylinus).

  • — Chapman, 1934 (subgenus of Staphylinus; notes on classification and distribution of some species).

  • — Tottenham, 1949: 375 (synonym of Ontholestes; type species: murinus Linné).

  • — Blackwelder, 1952: 394 (type species).

  • — Coiffait, 1956e: 183, 197, 194 (characters in key; notes; catalog of and key to French species).

  • — Smetana, 1958: 305 (synonym of Ontholestes).

  • — Palm, 1963b: 109 (subgenus of Staphylinus; characters).

  • — Lohse, 1964: 194 (characters; key to central European species).

  • — Scheerpeltz, 1966a: 114 (subgenus of Staphylinus; characters).

  • — Pototskaia, 1966b: 362 (subgenus of Staphylinus; larval characters in key).

  • — Coiffait, 1974: 368 (synonym of Ontholestes).

  • — Lohse and Lucht, 1989: 169 (synonym of Dinothenarus).

  • — Smetana, 1993b: 49 (synonym of Dinothenarus).

adonis Coiffait, 1956, see: Dinothenarus.

africanus Bernhauer, 1904a: 236 (Leistotrophus; Type locality: Kamerun, Barombi).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog; Gabon; Central Africa).

  • — Bernhauer, 1915o: 363 (Ontholestes; Cameroon).

  • — Cameron, 1950g: 43 (Ontholestes; Belgian Congo).

  • Distribution: Congo, Cameroon, Gabon, “Central Africa”.

asiaticus Smetana, 1959d: 404 (Ontholestes; Type locality: Tonkin).

  • Distribution: Vietnam.

assamensis Cameron, 1944, see: aurosparsus Fauvel, 1895.

aurosparsus Fauvel, 1895: 245 (Leistotrophus; Type locality: Bírmanie, Carin Asciuii Ghécu, 1200–1400 m; Bengale, Kurseong; Sikkim, Darjeeling).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog).

  • — Gridelli, 1924f: 207 (Ontholestes; characters in key).

  • — Cameron, 1932: 211 (Ontholestes; characters; Burma; Bengal; India).

  • — Scheerpeltz, 1933: 1405 (Ontholestes; catalog).

  • — Kirshenblat, 1936: 562 (Ontholestes; characters).

  • — Paulian, 1941: 250 (Ontholestes; larval characters; India).

  • — Cerruti, 1951: 16 (Ontholestes; India).

  • — Smetana, 1959d: 403 (Ontholestes; characters; Sikkim; Assam; Burma; Bengal; China).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: India, Myanmar, China.

ophthalmicus Kirshenblat, 1936: 560 (Ontholestes; Type locality: Sikkim, Darjeeling).

  • — Smetana, 1959d: 403 (Ontholestes; synonym of aurosparsus).

assamensis Cameron, 1944: 11 (Ontholestes; Type locality: Assam: Shugan, altitude 3000 feet).

  • — Smetana, 1959d: 403 (Ontholestes; synonym of aurosparsus).

barthei Chapman, 1922, see: murinus Linné, 1758.

borneensis Cameron, 1942a: 137 (Ontholestes; Type locality: Brunei).

  • — Smetana, 1959d: 399 (Ontholestes; characters; Borneo; Siam).

  • Distribution: Borneo, Thailand.

siamensis Cameron, 1946a: 685 (Ontholestes; Type locality: Siam: Renong).

  • — Smetana, 1959d: 399 (Ontholestes; synonym of borneensis).

brasilianus Bernhauer, 1906b: 202 (Ontholestes; Type locality: Brasilia).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog).

  • — Scheerpeltz, 1933: 1405 (Ontholestes; catalog; Argentina).

  • Distribution: Brazil, Argentina.

brevicornis Motschulsky, 1862, see: Platydracus.

callistus Hochhuth, 1849: 114 (Staphylinus; Type locality: not cited, presumably in the Caucasus).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog).

  • — Smetana, 1959d: 410 (Ontholestes; republication of original description; notes).

  • — Petrenko, 1980: 83 (Ontholestes; Azerbaijan).

  • — Gusarov, 1991a: 8 (Ontholestes; notes).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: Azerbaijan, Iran.

talyshensis Kirshenblat, 1936: 554 (Ontholestes; Type locality: Talysh, südöstliches Transkaukasien; [Note: See lectotype designation by Gusarov, 1991a]).

  • — Smetana, 1959d: 408 (Ontholestes; characters; Transcaucasia; Persia).

  • — Coiffait, 1974: 379 (Ontholestes; characters; Transcaucasus; Iran).

  • — Boháč, 1980: 86 (Ontholestes; cited as talyschensis; Iran).

  • — Gusarov, 1991a: 8 (Ontholestes; synonym of callistus; lectotype designation).

chalcopygus Hochhuth, 1849: 113 (Staphylinus; Type locality: Armenien).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog).

  • — Scheerpeltz, 1933: 1405 (Ontholestes; catalog; Transcausus).

  • — Kirshenblat, 1936: 563 (Ontholestes; characters).

  • — Smetana, 1959d: 399 (Ontholestes; characters).

  • — Coiffait, 1974: 376 (Ontholestes; characters; Armenia).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: Armenia.

chrysurus Kirby, 1837, see: cingulatus Gravenhorst, 1802.

cingulatus Gravenhorst, 1802: 166 (Staphylinus; Type locality: America septentr.).

  • — Gravenhorst, 1806: 121 (Staphylinus).

  • — Nordmann, 1837: 31 (Staphylinus).

  • — Erichson, 1839b: 360 (Staphylinus; characters; America septentrionalis).

  • — Provancher, 1877: 245 (Leistotrophus; characters; Quebec).

  • — Horn, 1879: 198 (Leistotrophus; characters; east of the Rocky Mountains and in Vancouver).

  • — Blatchley, 1910: 374 (Leistotrophus; cited as Listotrophus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog).

  • — Notman, 1920a: 180 (Ontholestes; New York).

  • — Scheerpeltz, 1933: 1405 (Ontholestes; catalog).

  • — Voris, 1939a: 290 (Ontholestes; egg, larval, and pupal characters).

  • — Hatch, 1957: 179 (Ontholestes; characters; British Columbia; Idaho; Washington; Oregon).

  • — Smetana, 1959d: 412 (Ontholestes; characters; North America).

  • — Smetana, 1965: 33 (Ontholestes; Newfoundland).

  • — Smetana, 1965a: 11 (Ontholestes; British Columbia).

  • — Smetana, 1981d: 126 (Ontholestes; characters).

  • — Alcock, 1991 (Ontholestes; mating behavior; Virginia).

  • — Dearborn and Donahue, 1994: 15 (Ontholestes; Maine).

  • — Downie and Arnett, 1996: 386 (Ontholestes; characters; USA).

  • — Schmidt, 1999: 108 (Ontholestes; food for adults and larvae used for rearing species; rearing methods).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: Canada, USA.

speciosus Mannerheim, 1830: 21 (Emus; Type locality: America borealis).

  • — Mannerheim, 1831: 435 (Emus; America borealis).

  • — Nordmann, 1837: 31 (Staphylinus; erroneously cited as spinosus; North America).

  • — Erichson, 1839b: 360 (Staphylinus; synonym of cingulatus).

  • — Horn, 1879: 198 (Leistotrophus; synonym of cingulatus).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; synonym of cingulatus).

chrysurus Kirby, 1837: 94 (Staphylinus; Type locality: Nova Scotia).

  • — Erichson, 1840: 926 (Staphylinus; synonym of cingulatus).

  • — Horn, 1879: 198 (Leistotrophus; synonym of cingulatus).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; synonym of cingulatus).

concolor Motschulsky, 1858, see: Staphylinus.

cupreus Geoffroy, 1785, see: murinus Linné 1758.

dieckmanni Smetana, 1958a: 363 (Ontholestes; Type locality: Deutschland: Mark Brandenburg, Beeskow).

  • — Smetana, 1959d: 402 (Ontholestes; characters; Germany).

  • — Coiffait, 1974: 378 (Ontholestes; characters; Germany).

  • — Tóth, 1984: 12 (Ontholestes; characters; Germany).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: Germany.

doriae Gridelli, 1924, see: Amichrotus.

elegans Cameron, 1938b: 151 (Ontholestes; Type locality: E. Java: Res. Kediri, Kalibamban).

  • — Smetana, 1959d: 409 (Ontholestes; republication of original description; notes; Java).

  • Distribution: Indonesia.

gracilentus Cameron, 1951c: 405 (Ontholestes; Type locality: Uganda: Bwamba).

  • Distribution: Uganda.

gracilis Sharp, 1874: 28 (Leistotrophus; Type locality: Japan).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog).

  • — Kirshenblat, 1936: 554 (Ontholestes; characters).

  • — Cameron, 1949b: 469 (Leistotrophus; Japan).

  • — Smetana, 1959d: 409 (Ontholestes; characters; Japan; Siberia).

  • — Nakane, 1963a: 93 (Ontholestes; characters; habitus photograph; Japan).

  • — Yuh, Paik, Kwon, and Lee, 1985: 247 (Ontholestes; Korea).

  • — J. Li, 1993: 40 (Ontholestes; China).

  • — Kanehisa, Tsumuki, and Kawazu, 1994 (Ontholestes; chemical secretions).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: Russia, China, Korea, Japan.

hairaerensis J. Li, 1993: 47 (Ontholestes; Type locality: Hairaerbayinhushe, N. E. China).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: China.

haroldi Eppelsheim, 1884: 170 (Leistotrophus; variety of murinus; Type locality: München).

  • — Ganglbauer, 1895: 418 (Ontholestes; variety of murinus).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; variety of murinus).

  • — Scheerpeltz, 1933: 1406 (Ontholestes; aberration of murinus).

  • — Smetana, 1955a: 287 (Ontholestes; cited as valid species; characters; ecology).

  • — Smetana, 1958: 309 (Ontholestes; characters; ecology; Czechoslovakia).

  • — Smetana, 1958a: 365 (Ontholestes; characters).

  • — Smetana, 1959d: 402 (Ontholestes; characters; Yugoslavia; Hungary; Bosnia; Austria; Czechoslovakia; Slovakia; Germany; Switzerland; Ukraine; Italy; Bulgaria).

  • — Smetana, 1960b: 259 (Ontholestes; Czechoslovakia).

  • — Lohse, 1964: 193 (Ontholestes; characters; central Europe).

  • — Horion, 1965: 245 (Ontholestes; Germany; Austria; Czechoslovakia; Switzerland).

  • — Smetana, 1966: 331 (Ontholestes; notes; Czechoslovakia).

  • — Coiffait, 1974: 379 (Ontholestes; characters; Europe).

  • — Tóth, 1984: 12 (Ontholestes; characters; Hungary).

  • — Lucht, 1987: 107 (Ontholestes; checklist; Germany; Poland; Czechoslovakia; Austria).

  • — Pilon and Zanetti, 1991: 66 (Ontholestes; Italy).

  • — Callot, 1993: 24 (Ontholestes; characters; notes; France).

  • — Černý, 1993: 69 (Ontholestes; collecting notes; Czech Republic).

  • — Smetana, 1993b: 49 (Ontholestes; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 29 (Ontholestes; Italy).

  • — Pilon, 1998: 65 (Ontholestes; ecological notes; Italy).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: Europe.

holdhausi Bernhauer, 1913b: 233 (Ontholestes; Type locality: Ukaika).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog).

  • Distribution: “Central Africa”.

hayashii J. Li, 1992: 59 (Ontholestes; Type locality: China: Liaoning Province: Yingkou Co., Huangtuling).

  • — J. Li, 1993: 39, 51 (Ontholestes; characters; China).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: China.

hybridus Marsham, 1802, see: tessellatus Geoffroy, 1785.

inauratus Mannerheim, 1830: 21 (Emus; Type locality: Dauria).

  • — Mannerheim, 1831: 435 (Emus; Dauria).

  • — Nordmann, 1837: 31 (Staphylinus; Dauria).

  • — Erichson, 1839b: 363 (Staphylinus; characters; Dauria).

  • — Motschulsky, 1858d: 67 (Staphylinus; synonym of murinus).

  • — Kraatz, 1857d: 535 (Leistotrophus; synonym of murinus).

  • — Fauvel, 1869: 487 (Leistotrophus; synonym of murinus).

  • — Ganglbauer, 1895: 418 (Ontholestes; synonym of murinus).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; variety of murinus).

  • — Scheerpeltz, 1933: 1406 (Ontholestes; aberration of murinus).

  • — Aleksandrov, 1934: 153 (Ontholestes; variety of murinus; China).

  • — Kirshenblat, 1936: 560 (Ontholestes; characters).

  • — Smetana, 1959d: 402 (Ontholestes; cited as valid species; characters; Transbaikal; Mongolia).

  • — Smetana, 1967d: 210 (Ontholestes; Mongolia).

  • — Coiffait, 1974: 378 (Ontholestes; characters; Siberia; Mongolia).

  • — J. Li, 1993: 40 (Ontholestes; cited as inauratur; China).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: Russia, Mongolia, China.

javanus Bernhauer, 1915, see: Thoracostrongylus.

louwerensi Cameron, 1936c: 183 (Ontholestes; Type locality: E. Java: Toempoek Bij; Popoh, Zuider Geb).

  • — Smetana, 1959d: 403 (Ontholestes; characters; Java).

  • Distribution: Indonesia.

marginalis Gené, 1836: 171 (Emus; Type locality: Sardinia: Caputerra; Cabras).

  • — Erichson, 1839b: 361 (Staphylinus; characters; Sardinia; Lusitania).

  • — P. Lucas, 1846: 109 (Staphylinus; notes; Algeria).

  • — Mulsant and Rey, 1876: 230 [= 1877: 86] (Leistotrophus; characters; Corsica; Sardinia; Portugal).

  • — Fauvel, 1878: 118 [= 1878a: 38] (Leistotrophus; Algeria).

  • — Fauvel, 1886: 50 [= 1886a: 42] (Leistotrophus; Morocco; Algeria).

  • — Fauvel, 1897d: 293 (Leistotrophus; Barbary).

  • — Fauvel, 1902b: 103 (Leistrotrophus; Morocco; Algeria; Tunisia).

  • — Sainte-Claire Deville, 1906: 100 (Ontholestes; Corsica).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog).

  • — Kirshenblat, 1936: 563 (Ontholestes; characters).

  • — Coiffait, 1956e: 190 (Abemus; subgenus Ontholestes; characters; France).

  • — Smetana, 1958a: 366 (Ontholestes; characters).

  • — Smetana, 1959d: 400 (Ontholestes; characters; Portugal; Spain; Corsica; Sardina; Tunisia; Algeria; Morocco).

  • — Coiffait, 1973a: 274 (Ontholestes; Morocco).

  • — Coiffait, 1974: 376 (Ontholestes; characters; Corsica; Sardinia; North Africa).

  • — Ciceroni and Zanetti, 1995: 30 (Ontholestes; Italy).

  • — Pilon, 1998: 65 (Ontholestes; ecological notes; Italy).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: Algeria, Tunisia, Morocco, Portugal, Spain, France, Italy.

marginatus Cameron, 1946, see: variegatus Cameron, 1932.

marmoratus Erichson, 1840: 926 (Staphylinus; Type locality: Java).

  • — Kraatz, 1859: 68 (Leistotrophus; characters; Ceylon).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog).

  • — Cameron, 1932: 210 (Ontholestes; characters; Ceylon).

  • — Scheerpeltz, 1933: 1405 (Ontholestes; catalog).

  • — Cameron, 1933d: 391 (Ontholestes; Sumatra).

  • — Cameron, 1937: 22 (Ontholestes; Java).

  • — Smetana, 1959d: 404 (Ontholestes; characters; Java; Ceylon; Sumatra).

  • Distribution: Sri Lanka, Indonesia.

modiglianii Gridelli, 1924, see: Thoracostrongylus.

moluccarum Fauvel, 1878f: 249 (Leistotrophus; Type locality: Moluques).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog).

  • — Gridelli, 1924f: 207 (Ontholestes; characters in key).

  • — Scheerpeltz, 1933: 1405 (Ontholestes; catalog).

  • — Smetana, 1959d: 403 (Ontholestes; characters; Molucca Islands; Celebes).

  • Distribution: Indonesia.

purpuripennis Bernhauer, 1915f: 144 (Ontholestes; Type locality: Süd-Celebes: Bonthain).

  • — Gridelli, 1924f: 207 (Ontholestes; characters in key).

  • — Scheerpeltz, 1933: 1406 (Ontholestes; catalog).

  • — Smetana, 1959d: 403 (Ontholestes; synonym of moluccarum).

murinus Linné, 1758: 421 (Staphylinus; Type locality: Not cited).

  • — Linné, 1760: 230 (Staphylinus; characters; Sweden).

  • — Scopoli, 1763: 100 (Staphylinus; characters; Carniola).

  • — O. Müller, 1764: 23 (Staphylinus; characters).

  • — Pontoppidans, 1765: 211 (Staphylinus; Denmark).

  • — Linné, 1767: 683 (Staphylinus; characters).

  • — De Geer, 1774: 13 (Staphylinus; redescription; notes).

  • — Füessly, 1775: 21 (Staphylinus).

  • — Fabricius, 1775: 265 (Staphylinus; characters).

  • — O. Müller, 1776: 98 (Staphylinus; characters).

  • — Goeze, 1777: 714 (Staphylinus; characters).

  • — Fabricius, 1781: 334 (Staphylinus; characters).

  • — Herbst, 1784: 149 (Staphylinus; Germany).

  • — Fabricius, 1787: 219 (Staphylinus; characters).

  • — Paykull, 1789: 14 (Staphylinus; redescription; Sweden).

  • — Rossi, 1790: 247 (Staphylinus; characters; Italy).

  • — Villers, 1789: 409 (Staphylinus; characters).

  • — Gmelin, 1790: 2026 (Staphylinus; characters; larval characters).

  • — Fabricius, 1793: 520 (Staphylinus; characters).

  • — Olivier, 1795:(42): 15 (Staphylinus; characters).

  • — Panzer, 1795: 352 (Staphylinus; characters; Germany).

  • — Schrank, 1798: 641 (Staphylinus; characters; notes).

  • — Cederhjelm, 1798: 116 (Staphylinus; characters).

  • — Panzer, 1799d: 16 (Staphylinus; characters; illustrations).

  • — Paykull, 1800: 376 (Staphylinus; characters; Sweden).

  • — Fabricius, 1801: 590 (Staphylinus; characters).

  • — Turton, 1802: 509 (Staphylinus; characters).

  • — Gravenhorst, 1802: 14 (Staphylinus; characters).

  • — Marsham, 1802: 499 (Staphylinus; characters; Britain).

  • — Latreille, 1804: 294 (Staphylinus; characters).

  • — Gravenhorst, 1806: 122 (Staphylinus).

  • — Gyllenhal, 1810: 283 (Staphylinus; characters; Sweden).

  • — Zetterstedt, 1828: 67 (Staphylinus; characters; Lapland).

  • — Curtis, 1829: 24 (Staphylinus; catalog; Britain).

  • — Latreille, 1829: 434 (Staphylinus; characters).

  • — Mannerheim, 1830: 21 (Emus; Finland; Lapponia).

  • — Mannerheim, 1831: 435 (Emus; Finland; Lapponia).

  • — Stephens, 1832: 205 (Staphylinus; characters; England).

  • — Ménétriés, 1832: 142 (Staphylinus; Caucasus-Persia region).

  • — Griffith and Pidgeon, 1832: 291 (Staphylinus; characters).

  • — Runde, 1835: 2 (Emus; characters; Germany).

  • — Curtis, 1839: pl. 758 (Staphylinus; type species of Staphylinus; notes).

  • — C. Sahlberg, 1830: 308 (Staphylinus; characters; Finland).

  • — Lacordaire, 1835: 363 (Emus; characters; France).

  • — Nordmann, 1837: 31 (Staphylinus; Europe; Asia).

  • — Westwood, 1838a: 15 (Trichoderma; type species).

  • — Zetterstedt, 1838: 59 (Staphylinus; characters; habitat; Lapland).

  • — O. Costa, 1839b: 118 (Staphylinus; characters; Italy).

  • — Erichson, 1839a: 433 (Staphylinus; characters; Germany).

  • — Heer, 1839: 250 (Staphylinus; characters; Switzerland).

  • — Erichson, 1839b: 361 (Staphylinus; characters; Europe).

  • — Stephens, 1839: 386 (Trichoderma; characters).

  • — Laporte, 1840: 174 (Emus; characters; France).

  • — Westwood, 1845: 45 (Staphylinus; subgenus Trichoderma; characters; differs from nebulosus).

  • — Kolenati, 1846: 17 (Staphylinus; Caucasus; Transcaucasus).

  • — Jacobsen, 1849: 682 (Staphylinus; characters; Denmark).

  • — Ménétriés, 1849: 43 (Staphylinus; Khirgiz region).

  • — Hochhuth, 1849: 113 (Staphylinus; characters; Tiflis; Armenia).

  • — Redtenbacher, 1849: 695 (Staphylinus; characters; Austria).

  • — Küster, 1853: 44 (Staphylinus; characters; Europe).

  • — Hardy, 1851: 21 (Staphylinus; subgenus Trichoderma; notes; England).

  • — Rosenhauer, 1856: 70 (Staphylinus; Granada).

  • — Fairmaire and Laboulbène, 1856: 506 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 535 (Leistotrophus; characters; Germany).

  • — Redtenbacher, 1857: 184 (Staphylinus; characters; Austria).

  • — Thomson, 1860: 142 (Schizochilus; characters; Scandinavia).

  • — Hochhuth, 1862: 48 (Leistotrophus; notes; Dauria).

  • — Fauvel, 1865a: 294 [= 1865b: 44] (Staphylinus; habitat; France).

  • — Fauvel, 1874: 171 [= 1874a: 397] (Leistotrophus; characters; France; Turkey; Caucasus; Transcaucasia; Siberia; Dauria; North America).

  • — Redtenbacher, 1874: 203 (Leistotrophus; characters; Austria).

  • — Seidlitz, 1875: 269 (Leistotrophus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 230 [= 1877: 86] (Leistotrophus; characters; France).

  • — J. Sahlberg, 1876: 9 (Leistotrophus; Finland).

  • — J. Sahlberg, 1880: 68 (Leistotrophus; Siberia).

  • — Fowler, 1888: 249 (Leistotrophus; characters; Britain).

  • — Seidlitz, 1891: 394 (Leistotrophus; characters; Europe).

  • — Seidlitz, 1891a: 420 (Leistotrophus; characters; Europe).

  • — Ganglbauer, 1895: 418 (Ontholestes; characters; Palaearctic region).

  • — Poppius, 1909: 28 (Ontholestes; Lena Valley, Siberia).

  • — Reitter, 1909: 117 (Ontholestes; characters; Germany).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog).

  • — Johansen, 1914: 375 (Ontholestes; characters; Denmark).

  • — Portevin, 1929: 348 (Ontholestes; characters; France).

  • — Scheerpeltz, 1933: 1405 (Ontholestes; catalog).

  • — Aleksandrov, 1934: 153 (Ontholestes; China).

  • — Kirshenblat, 1936: 559 (Ontholestes; characters).

  • — Paulian, 1941: 250 (Ontholestes; larval characters).

  • — Hinton, 1945: 72 (Ontholestes; characters).

  • — Jeannel and Jarrige, 1949: 376 (Ontholestes; cave species; Spain).

  • — V. Hansen, 1952: 109 (Ontholestes; characters; Denmark).

  • — Smetana, 1955a: 186 (Ontholestes; characters; ecology).

  • — Coiffait, 1956e: 190 (Abemus; subgenus Ontholestes; characters; France).

  • — Smetana, 1958: 308 (Ontholestes; characters; ecology; Czechoslovakia).

  • — Smetana, 1958a: 365 (Ontholestes; characters).

  • — Smetana, 1959d: 401 (Ontholestes; characters; Siberia; Transbaikal).

  • — Palm, 1963b: 120 (Staphylinus; subgenus Ontholestes; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 193 (Ontholestes; characters; central Europe).

  • — Focarile, 1964: 58 (Ontholestes; Italy).

  • — Scheerpeltz, 1965b: 405 (Ontholestes; Sicily).

  • — Smetana, 1965: 33 (Ontholestes; notes; Newfoundland).

  • — Horion, 1965: 244 (Ontholestes; all Europe).

  • — Pototskaia, 1967: 74 (Ontholestes; larval characters).

  • — Szujecki, 1968a: 733 (Ontholestes; Poland).

  • — Bolov, 1969a: 328 (Ontholestes; collecting notes; Russia).

  • — Kasule, 1970: 74 (Ontholestes; larval characters).

  • — Jarrige, 1971: 497 (Ontholestes; Iran).

  • — Coiffait, 1974: 377 (Ontholestes; characters; Europe; Britain; Scandinavia; Asia Minor; central Asia; Siberia).

  • — Shilov, 1975a: 376 [= 1975b: 99] (Ontholestes; Komi region of Russia).

  • — Pope, 1977: 30 (Ontholestes; Britain).

  • — Smetana, 1981d: 125 (Ontholestes; characters; introduced into North America; collecting notes; Newfoundland).

  • — Hinton, 1981: 662 (Ontholestes; characters of egg).

  • — Tóth, 1984: 12 (Ontholestes; characters; Hungary).

  • — Dettner and Schwinger, 1986: 366 (Ontholestes; defensive secretions).

  • — Lucht, 1987: 107 (Ontholestes; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Huth and Dettner, 1989: 154 (Ontholestes; defensive secretions).

  • — Huth and Dettner, 1990 (Ontholestes; defensive chemicals).

  • — Pilon, 1991: 102 (Ontholestes; notes; Italy).

  • — Pilon and Zanetti, 1991: 66 (Ontholestes; Italy).

  • — Welch, 1993: 229 (Ontholestes; ovariole number and ovary structure).

  • — J. Li, 1993: 40 (Ontholestes; cited as murinous; China).

  • — Smetana, 1993b: 49 (Ontholestes; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 30 (Ontholestes; Italy).

  • — Sparacio, 1995: 161 (Ontholestes; characters; notes; Sicily).

  • — Downie and Arnett, 1996: 387 (Ontholestes; characters; Canada).

  • — Pilon, 1998: 65 (Ontholestes; ecological notes; Italy).

  • — Kashcheev, 1999a: 58 (Ontholestes; Kazakhstan).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: Europe, Russia, Georgia, Armenia, Turkey, Iran, Kazakhstan, Kyrgyzstan, China, Canada.

cupreus Geoffroy, 1785: 173 (Staphylinus; Type locality: Paris).

  • — Villers, 1789: 423 (Staphylinus; characters; France).

  • — Ganglbauer, 1895: 418 (Ontholestes; synonym of murinus).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; synonym of murinus).

  • — Smetana, 1958: 308 (Ontholestes; synonym of murinus).

  • — Coiffait, 1974: 377 (Ontholestes; synonym of murinus).

ruficornis Ménétriés, 1832: 148 (Emus; Type locality: Bakou).

  • — Kirshenblat, 1933: 103 (Ontholestes; transferred from Emus; synonym of murinus).

  • — Coiffait, 1974: 377 (Ontholestes; synonym of murinus).

barthei Chapman, 1922: 17 (Leistotrophus; variety of murinus; Type locality: Bourbon-Lancy [Saone-et-Loire]).

  • — Scheerpeltz, 1933: 1406 (Ontholestes; aberration of murinus).

  • — Coiffait, 1974: 377 (Ontholestes; variety of murinus).

nebulosus Fabricius, 1793, see: tessellatus Geoffroy, 1785.

oculatus Sharp, 1874: 29 (Leistotrophus; Type locality: Nagasaki).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog).

  • — Aleksandrov, 1934: 153 (Ontholestes; notes; China).

  • — Kirshenblat, 1936: 562 (Ontholestes; characters).

  • — Smetana, 1959d: 405 (Ontholestes; characters; Japan).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: Japan, China.

olivaceus Cameron, 1946, see: variegatus Cameron, 1932.

ophthalmicus Kirshenblat, 1936, see: aurosparsus Fauvel, 1895.

orientalis Bernhauer, 1906: 125 (Ontholestes; Type locality: Wladiwostock).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog).

  • — Aleksandrov, 1934: 153 (Ontholestes; China).

  • — Kirshenblat, 1936: 556 (Ontholestes; characters).

  • — Smetana, 1959d: 407 (Ontholestes; characters; Siberia; China).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: Russia, China.

paramurinus J. Li, 1993: 49 (Ontholestes; Type locality: Qian Shan, Liaoning Province).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: China.

proximus Kirshenblat, 1936: 556 (Ontholestes; Type locality: Provinz Sze-tshuan, China occidentalis).

  • — Smetana, 1959d: 407 (Ontholestes; characters; China).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: China.

punctatissimus Motschulsky, 1858, see: Staphylinus.

purpuripennis Bernhauer, 1915, see: moluccarum Fauvel, 1878.

ruficornis Ménétriés 1832, see: murinus Linné, 1758.

secundus Schaeffer, 1766 [Note: Nomen nudum], see: tessellatus Geoffroy, 1785.

siamensis Cameron, 1946, see: borneensis Cameron, 1942.

simulator Kirshenblat, 1936: 564 (Ontholestes; Type locality: Süd-Ussuri Gebiet, Ost-Mongolei; [Note: See lectotype designation by Gusarov, 1991a]).

  • — Smetana, 1959d: 400 (Ontholestes; characters; Siberia; Mongolia; Korea; China).

  • — Smetana, 1967d: 210 (Ontholestes; Mongolia).

  • — Gusarov, 1991a: 8 (Ontholestes; notes; lectotype designation).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: Mongolia, Russia, China, Korea.

speciosus Mannerheim, 1830, see: cingulatus Gravenhorst, 1802.

sumatrensis Gridelli, 1924, see: Thoracostrongylus.

superbus Bernhauer, 1915f: 144 (Ontholestes; Type locality: Borneo: Lundu).

  • — Gridelli, 1924f: 207 (Ontholestes; characters in key).

  • — Scheerpeltz, 1933: 1406 (Ontholestes; catalog).

  • — Smetana, 1959d: 405 (Ontholestes; characters; Borneo).

  • Distribution: Borneo.

talyshensis Kirshenblat, 1936, see: callistus Hochhuth, 1849.

tenuicornis Kraatz, 1859: 68 (Leistotrophus; Type locality: India borealis).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog).

  • — Gridelli, 1924f: 207 (Ontholestes; characters in key).

  • — Cameron, 1932: 211 (Ontholestes; characters; India).

  • — Scheerpeltz, 1933: 1406 (Ontholestes; catalog).

  • — Smetana, 1959d: 408 (Ontholestes; characters; China; India).

  • — Coiffait, 1982: 31 (Ontholestes; Nepal).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: India, Nepal, China.

tessellatus Geoffroy, 1785: 165 (Staphylinus; Type locality: Paris).

  • — Harrer, 1784: 252 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 392, cited murinus Harrer as a synonym of tessellatus; Harrer did not describe the species as new, he attributed it to Linné]; characters).

  • — Olivier, 1795:(42): 15 (Staphylinus; synonym of murinus Linné).

  • — Gravenhorst, 1806: 121 (Staphylinus; synonym of nebulosus Fabricius).

  • — Erichson, 1839b: 361 (Staphylinus; synonym of nebulosus).

  • — Kraatz, 1857d: 534 (Leistotrophus; synonym of nebulosus).

  • — Ganglbauer, 1895: 418 (Ontholestes; characters; north and middle Europe; Siberia).

  • — Poppius, 1909: 28 (Ontholestes; Lena Valley, Siberia).

  • — Reitter, 1909: 117 (Ontholestes; characters; Germany).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog).

  • — Eichelbaum, 1915a: 92 (Ontholestes; abdominal segmentation; apical segements).

  • — Portevin, 1929: 348 (Ontholestes; characters; France).

  • — Scheerpeltz, 1933: 1406 (Ontholestes; catalog).

  • — Aleksandrov, 1934: 153 (Ontholestes; China).

  • — Kirshenblat, 1936: 558 (Ontholestes; cited as tesselatus; characters).

  • — Paulian, 1941: 250 (Ontholestes; larval characters).

  • — Tottenham, 1949: 421 (Ontholestes; checklist; Britain).

  • — V. Hansen, 1952: 108 (Ontholestes; characters; Denmark).

  • — Smetana, 1955a: 285 (Ontholestes; characters; ecology).

  • — Coiffait, 1956e: 190 (Abemus; subgenus Ontholestes; characters; France).

  • — Smetana, 1958: 307 (Ontholestes; cited as tesselatus; characters; ecology; Czechoslovakia).

  • — Smetana, 1958a: 365 (Ontholestes; characters).

  • — Smetana, 1959d: 399 (Ontholestes; cited as tesselatus; characters; Siberia; Mongolia).

  • — Palm, 1963b: 119 (Staphylinus; subgenus Ontholestes; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 192 (Ontholestes; characters; central Europe).

  • — Horion, 1965: 244 (Ontholestes; cited as tesselatus; all northern and central Europe; France; Italy; Istria; Croatia; Bosnia Hercegovina).

  • — Pototskaia, 1967: 74 (Ontholestes; larval characters).

  • — Szujecki, 1968a: 733 (Ontholestes; cited as tesselatus; Poland).

  • — Scheerpeltz, 1968a: 86 (Ontholestes; catalog; Austria).

  • — Kasule, 1970: 74 (Ontholestes; larval characters).

  • — Coiffait, 1974: 374 (Ontholestes; characters; Europe; Britain; Scandinavia; central Asia; Siberia).

  • — Shilov, 1975a: 376 [= 1975b: 99] (Ontholestes; cited as tesselatus; Komi region of Russia).

  • — Pope, 1977: 30 (Ontholestes; Britain).

  • — Muona, 1979: 15 (Ontholestes; cited as tesselatus; Finland; Russia; Sweden; Norway; Denmark; Baltic States).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 90 (Ontholestes; catalog; Poland).

  • — Tóth, 1984: 12 (Ontholestes; cited as tesselatus; characters; Hungary).

  • — Lucht, 1987: 107 (Ontholestes; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Huth and Dettner, 1989: 154 (Ontholestes; cited as tesselatus; defensive secretions).

  • — Huth and Dettner, 1990 (Ontholestes; defensive chemicals).

  • — Pilon and Zanetti, 1991: 65 (Ontholestes; Italy).

  • — Silfverberg, 1992: 17 (Ontholestes; Finland; Russia; Sweden; Norway; Denmark; Estonia; Latvia; Lithuania).

  • — Welch, 1993: 229 (Ontholestes; ovariole number and ovary structure).

  • — Smetana, 1993b: 49 (Ontholestes; cited as tesselatus; Czech Republic; Slovakia).

  • — J. Li, 1993: 39 (Ontholestes; cited as tesstlatus; China).

  • — Ciceroni and Zanetti, 1995: 30 (Ontholestes; Italy).

  • — M. Hansen, 1996: 107 (Ontholestes; checklist; Denmark).

  • — Eisinger, 1997: 170 (Ontholestes; collecting notes; Germany).

  • — Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 38 (Ontholestes; cited as tesselatus; Latvia).

  • — Anderson, Nash, and O'Connor, 1997: 19 (Ontholestes; cited as tesselatus; Ireland).

  • — Pilon, 1998: 67 (Ontholestes; ecological notes; Italy).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (Ontholestes; Denmark).

  • — Assing and Schülke, 1999: 14 (Ontholestes; authorship and spelling of tessellatus).

  • — Smetana and Davies, 2000: 38 (Ontholestes).

  • Distribution: Europe, Russia, Mongolia, China.

secundus Schaeffer, 1766: iv, 11 (Staphylinus; [Note: Nomen nudum, Article 11.4]; Type locality: Ratisbon).

  • — Turton, 1802: 509 (Staphylinus; not cited by name but cited as equivalent to murinus; characters).

  • — Erichson, 1839b: 361 (Staphylinus; synonym of nebulosus).

  • — Kraatz, 1857d: 534 (Leistotrophus; synonym of nebulosus).

nebulosus Fabricius, 1793: 520 (Staphylinus; [preoccupied]; Type locality: Germania).

  • — Panzer, 1795: 352 (Staphylinus; characters; Germany).

  • — Paykull, 1800: 375 (Staphylinus; characters; Sweden).

  • — Fabricius, 1801: 590 (Staphylinus; characters; Germany).

  • — Gravenhorst, 1802: 13 (Staphylinus; characters).

  • — Latreille, 1804: 295 (Staphylinus; characters).

  • — Gravenhorst, 1806: 121 (Staphylinus).

  • — Gyllenhal, 1810: 282 (Staphylinus; characters; Sweden).

  • — Curtis, 1829: 24 (Staphylinus; catalog; Britain).

  • — Mannerheim, 1830: 20 (Emus; Sweden; Siberia).

  • — Mannerheim, 1831: 434 (Emus; Sweden; Siberia).

  • — Stephens, 1832: 204 (Staphylinus; characters; England).

  • — Runde, 1835: 2 (Emus; characters; Germany).

  • — Curtis, 1839: pl. 758 (Staphylinus; notes; England).

  • — Lacordaire, 1835: 361 (Emus; characters; France).

  • — Nordmann, 1837: 30 (Staphylinus; Europe; northern Asia).

  • — Zetterstedt, 1838: 59 (Staphylinus; characters; habitat; Lapland).

  • — Erichson, 1839a: 432 (Staphylinus; characters; Germany).

  • — Heer, 1839: 250 (Staphylinus; characters; Switzerland).

  • — Erichson, 1839b: 360 (Staphylinus; characters; Europe).

  • — Stephens, 1839: 386 (Trichoderma; characters).

  • — Laporte, 1840: 174 (Emus; characters; France).

  • — Jacobsen, 1849: 681 (Staphylinus; characters; Denmark).

  • — Redtenbacher, 1849: 695 (Staphylinus; characters; Austria).

  • — Küster, 1853: 43 (Staphylinus; characters; Europe).

  • — Hardy, 1851: 21 (Staphylinus; subgenus Trichoderma; notes; England).

  • — Fairmaire and Laboulbène, 1856: 505 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 534 (Leistotrophus; characters; Germany).

  • — Redtenbacher, 1857: 184 (Staphylinus; characters; Austria).

  • — Motschulsky, 1859: 491 (Emus; Amur River).

  • — Thomson, 1860: 142 (Schizochilus; characters; Scandinavia).

  • — Motschulsky, 1860b: 120 (Emus; eastern Siberia).

  • — Fauvel, 1865a: 294 [= 1865b: 44] (Staphylinus; habitat; France).

  • — Solsky, 1871c: 346 (Leistotrophus; notes; Siberia).

  • — Fauvel, 1874: 170 [= 1874a: 396] (Leistotrophus; characters; France; Italy; Baikal and Amur regions).

  • — Redtenbacher, 1874: 203 (Leistotrophus; characters; Austria).

  • — Seidlitz, 1875: 268 (Leistotrophus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 227 [= 1877: 83] (Leistotrophus; characters; France).

  • — J. Sahlberg, 1876: 9 (Leistotrophus; Finland).

  • — Fowler, 1888: 248 (Leistotrophus; characters; Britain).

  • — Ganglbauer, 1895: 418 (Ontholestes; synonym of tessellatus).

  • — Bernhauer and Schubert, 1914: 393 (Ontholestes; synonym of tessellatus).

  • — Johansen, 1914: 375 (Ontholestes; characters; Denmark).

  • — Scheerpeltz, 1933: 1406 (Ontholestes; synonym of tessellatus).

  • — Smetana, 1958: 307 (Ontholestes; synonym of tessellatus).

  • — Coiffait, 1974: 374 (Ontholestes; synonym of tessellatus).

  • — Pope, 1977: 30 (Ontholestes; synonym of tessellatus).

hybridus Marsham, 1802: 500 (Staphylinus; Type locality: Britain).

  • — Stephens, 1829: 275 (Staphylinus; synonym of nebulosus Fabricius).

  • — Stephens, 1832: 204 (Staphylinus; synonym of nebulosus Fabricius).

  • — Curtis, 1839: pl. 758 (Staphylinus; synonym of nebulosus).

  • — Erichson, 1839b: 361 (Staphylinus; synonym of nebulosus).

  • — Kraatz, 1857d: 534 (Leistotrophus; synonym of nebulosus).

  • — Ganglbauer, 1895: 418 (Ontholestes; synonym of tessellatus).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; synonym of tessellatus).

  • — Smetana, 1958: 307 (Ontholestes; synonym of tessellatus).

  • — Coiffait, 1974: 375 (Ontholestes; synonym of tessellatus).

variegatus Cameron, 1932: 212 (Ontholestes; Type locality: Burma: Karen Mts.).

  • — Scheerpeltz, 1933: 1406 (Ontholestes; catalog).

  • — Smetana, 1959d: 407 (Ontholestes; characters; Burma; Siam; Indo-China; Laos).

  • Distribution: Myanmar, Thailand, Laos, Vietnam.

olivaceus Cameron, 1946a: 687 (Ontholestes; Type locality: Siam: Renong).

  • — Smetana, 1959d: 407 (Ontholestes; synonym of variegatus).

marginatus Cameron, 1946a: 688 (Ontholestes; Type locality: Siam: Renong).

  • — Smetana, 1959d: 407 (Ontholestes; synonym of variegatus).

Palaestrinus [1 species; Oriental Region]

Palaestrinus Erichson, 1839b: 343 (species included: sykesii, aureus). Type species: Palaestrinus sykesii Erichson, fixed by subsequent designation by Duponchel, 1841: 57.

  • — Lacordaire, 1854: 74 (characters; notes; list of species).

  • — Gemminger and Harold, 1868: 574 (catalog).

  • — Bernhauer and Schubert, 1914: 394 (world catalog; 1 species).

  • — Cameron, 1925: 69 (catalog of Indian species).

  • — Cameron, 1932: 238 (characters).

  • — Scheerpeltz, 1933: 1408 (world catalog supplement).

  • — Blackwelder, 1952: 287 (type species).

kraatzi Fauvel, 1903, see: sykesii Erichson, 1839.

mutillarius Erichson, 1840, see: Parapalaestrinus.

sykesii Erichson, 1839b: 344 (Palaestrinus; Type locality: India orientalis).

  • — Kraatz, 1859: 71 (Palaestrinus; characters; India orientalis; Ceylon).

  • — Bernhauer and Schubert, 1914: 394 (Palaestrinus; cited as sykesi; catalog).

  • — Cameron, 1932: 238 (Palaestrinus; cited as sykesi; characters; India).

  • — Scheerpeltz, 1933: 1408 (Palaestrinus; cited as sykesi; catalog).

  • Distribution: “East Indies”, Sri Lanka, India.

kraatzi Fauvel, 1903: 156 (Palaestrinus; form of sykesii, cited as sykesi; Type locality: Ceylan).

  • — Bernhauer and Schubert, 1914: 394 (Palaestrinus; variety of sykesii, cited as sykesi).

  • — Scheerpeltz, 1933: 1408 (Palaestrinus; variety of sykesii, cited as sykesi).

waageni Eppelsheim, 1895, see: Parapalaestrinus.

Pancarpius [4 species; Ethiopian Region]

Pancarpius Bondroit, 1913: 92 (replacement name for Erichsonius Schubert). Type species: Erichsonius bicolor Schubert, fixed by objective synonymy with Erichsonius Schubert.

  • — Scheerpeltz, 1933: 1404 (synonym of Pancarpius).

  • — Blackwelder, 1952: 288 (cited as valid genus; type species).

Erichsonius Schubert, 1911: 33 (species included: bicolor; preoccupied by Fauvel, 1872). Type species: Erichsonius bicolor Schubert, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 391 (world catalog; 1 species).

  • — Blackwelder, 1952: 151 (synonym of Pancarpius; type species: bicolor).

Erichsonellus Bernhauer and Schubert, 1914: 391 (replacement name for Erichsonius Schubert). Type species: Erichsonius bicolor Schubert, fixed by objective synonymy with Erichsonius Schubert.

  • — Scheerpeltz, 1933: 1404 (world catalog supplement).

  • — Blackwelder, 1952: 151 (synonym of Pancarpius; type species).

  • — Levasseur, 1980: 372 (characters).

bicolor Schubert, 1911: 33 (Erichsonius; [Note: This name was described in Erichsonius Schubert, a homonym of Erichsonius Fauvel but not a synonym, so it is not cross-indexed in Erichsonius Fauvel]; Type locality: Brit. Uganda).

  • — Bernhauer and Schubert, 1914: 391 (Erichsonellus; catalog).

  • — Scheerpeltz, 1933: 1404 (Pancarpius; catalog).

  • Distribution: Uganda, Congo.

katangai Bondroit, 1913: 92 (Pancarpius; variety of bicolor; Type locality: Katanga).

  • — Scheerpeltz, 1933: 1404 (Pancarpius; variety of bicolor).

ertli Bernhauer, 1927: 55 (Pancarpius; Type locality: Angola).

  • — Scheerpeltz, 1933: 1404 (Pancarpius; catalog).

  • Distribution: Angola.

gentyi Levasseur, 1971: 300 (Erichsonellus; Type locality: La Mé près Bingerville, Côte d'Ivoire. Also cited from Guinea, Sierra Leone and Ubangi).

  • Distribution: Guinea, Sierra Leone, Ivory Coast, “Ubangi River”.

katangai Bondroit, 1913, see: bicolor Schubert, 1911.

mirei Levasseur, 1980: 372 (Erichsonellus; Type locality: Cameroun: Ahala).

  • Distribution: Cameroon.

Paragastrisus [7 species; Ethiopian Region]

Paragastrisus Bernhauer, 1923c: 63 (replacement name for Eurycnemus Bernhauer). Type species: Eurycnemus imperialis Bernhauer, fixed by objective synonymy with Eurycnemus.

  • — Scheerpeltz, 1933: 1416 (world catalog supplement).

  • — Blackwelder, 1952: 290 (type species).

  • — Levasseur, 1980: 371 (characters).

Eurycnemus Bernhauer, 1906a: 190 (species included: imperialis; preoccupied by van der Wulp, 1874). Type species: Eurycnemus imperialis Bernhauer, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 405 (world catalog; 1 species).

  • — Scheerpeltz, 1933: 1416 (synonym of Paragastrisus).

  • — Blackwelder, 1952: 159 (synonym of Paragastrisus; type species).

coiffaiti Levasseur, 1980: 378 (Paragastrisus; Type locality: Gabon: Belinga).

  • Distribution: Gabon.

imperialis Bernhauer, 1906a: 191 (Eurycnemus; Type locality: Kamerun).

  • — Bernhauer and Schubert, 1914: 405 (Eurycnemus; catalog).

  • Distribution: Cameroon.

koundenensis Levasseur, 1980: 377 (Paragastrisus; Type locality: Cameroun: Kounden).

  • Distribution: Cameroon.

mireanus Levasseur, 1980: 374 (Paragastrisus; Type locality: Cameroun: Ototomo).

  • Distribution: Cameroon.

pictipennis Bernhauer, 1927: 56 (Eurycnemus; Type locality: Uganda: Kampala).

  • — Scheerpeltz, 1933: 1416 (Paragastrisus; catalog).

  • Distribution: Uganda.

princeps Bernhauer, 1915d: 151 (Eurycnemus; Type locality: Kamerun).

  • — Scheerpeltz, 1933: 1416 (Paragastrisus; catalog).

  • Distribution: Cameroon.

regalis Bernhauer, 1915d: 150 (Eurycnemus; Type locality: Kamerun).

  • — Scheerpeltz, 1933: 1416 (Paragastrisus; catalog).

  • Distribution: Cameroon.

Parapalaestrinus [4 species; Oriental Region]

Parapalaestrinus Bernhauer, 1923c: 63 (replacement name for Seleucus Fauvel). Type species: Palaestrinus mutillarius Erichson, fixed by objective synonymy with Seleucus.

  • — Cameron, 1932: 235 (characters; key to species of British India).

  • — Scheerpeltz, 1933: 1408 (world catalog supplement).

  • — Blackwelder, 1952: 292 (type species).

Seleucus Fauvel, 1903: 157 (species included: mutillarius, waageni; preoccupied by Holmgren, 1858). Type species: Palaestrinus mutillarius Erichson, fixed by subsequent designation by R. Lucas, 1920: 589.

  • — Bernhauer and Schubert, 1914: 394 (world catalog; 2 species).

  • — Cameron, 1925: 69 (catalog of Indian species).

  • — Cameron, 1932: 235 (synonym of Parapalaestrinus).

  • — Scheerpeltz, 1933: 1408 (synonym of Parapalaestrinus).

  • — Blackwelder, 1952: 350 (synonym of Parapalaestrinus; type species).

delaportei Jarrige, 1951: 134 (Parapalaestrinus; Type locality: Lang-Giai, 30 km, N.-E. de Long-Son).

  • Distribution: Vietnam.

mutillarius Erichson, 1840: 924 (Palaestrinus; Type locality: Bengala).

  • — Kraatz, 1859: 72 (Palaestrinus; characters; Bengalia).

  • — Fauvel, 1903: 156, 157 (Seleucus; notes).

  • — Bernhauer and Schubert, 1914: 394 (Seleucus; catalog).

  • — Cameron, 1932: 235 (Parapalaestrinus; characters; Bengal; India).

  • — Scheerpeltz, 1933: 1408 (Parapalaestrinus; catalog).

  • Distribution: India.

paradoxus Cameron, 1932: 237 (Parapalaestrinus; Type locality: India).

  • — Scheerpeltz, 1933: 1408 (Parapalaestrinus; catalog).

  • Distribution: India.

waageni Eppelsheim, 1895b: 396 (Palaestrinus; Type locality: Calcutta).

  • — Fauvel, 1903: 157 (Seleucus).

  • — Bernhauer and Schubert, 1914: 394 (Seleucus; catalog).

  • — Cameron, 1932: 236 (Parapalaestrinus; characters; India).

  • — Scheerpeltz, 1933: 1408 (Parapalaestrinus; catalog).

  • Distribution: India.

Paraphytolinus [1 species; Oriental Region]

Paraphytolinus Hayashi, 1999a: 25 (species included: sumatrensis). Type species: Paraphytolinus sumatrensis Hayashi, fixed by original designation and monotypy.

sumatrensis Hayashi, 1999a: 30 (Paraphytolinus; Type locality: Indonesia: N. Sumatra, Brastagi. Gn. Sinabung, 1500-2000 m).

  • Distribution: Indonesia.

Philetaerius [1 species; Palaearctic Region]

Philetaerius Sharp, 1889: 118 (species included: elegans). Type species: Philetaerius elegans Sharp, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 393 (world catalog; 1 species).

  • — Winkler, 1925: 386 (catalog for Palaearctic region).

  • — Blackwelder, 1952: 300 (type species: elegans).

  • — Tikhomirova, 1973a: 188 (checklist of species of USSR).

  • — Shibata, 1984: 99 (checklist of species of Japan).

elegans Sharp, 1889: 119 (Philetaerius; Type locality: Bukenji).

  • — Bernhauer and Schubert, 1914: 393 (Philetaerius; catalog).

  • — Nakane, 1963a: 93 (Philetaerius; characters; habitus photograph; Japan).

  • — Maruyama, Mizota, and Ôhara, 2000: 67 (Philetaerius; notes; with Lasius spathepus and Lasius fuliginosus; Japan).

  • Distribution: Japan.

Physetops [5 species; Palaearctic Region]

Physetops Mannerheim, 1830: 7, 32 [= 1831: 421, 446] (species included: tataricus, cited as tartaricus). Type species: Staphylinus tataricus Pallas, fixed by monotypy.

  • — Mannerheim, 1831: 421, 446 (characters).

  • — Nordmann, 1837: 14 (characters).

  • — Erichson, 1840: 414 (Ocypus; synonym of Ocypus).

  • — Lacordaire, 1854: 79 (synonym of Ocypus).

  • — Jacquelin du Val, 1856: 35 (characters).

  • — Schaum, 1859: 26 (catalog; European species).

  • — Gemminger and Harold, 1868: 584 (catalog).

  • — Jacquelin du Val, 1868: 67 (group of Staphylinus; catalog; European species).

  • — Fauvel, 1875a: XXVIII [= 1875b: 230] (catalog).

  • — Heyden, 1880: 73 (list of species of Siberian region).

  • — Heyden, Reitter, and Weise, 1891: 102 (list of species of Europe and Caucasus).

  • — Heyden, 1896: 32 (list of species of Siberian region).

  • — Heyden, Reitter, and Weise, 1906: 168 (list of species of Europe and Caucasus).

  • — Bernhauer and Schubert, 1914: 395 (world catalog; 3 species).

  • — Bernhauer, 1923 (key to species).

  • — Winkler, 1925: 387 (catalog for Palaearctic region).

  • — Cameron, 1925: 70 (catalog of Indian species).

  • — Cameron, 1932: 244 (characters).

  • — Scheerpeltz, 1933: 1409 (world catalog supplement).

  • — Blackwelder, 1952: 308 (type species: tataricus).

  • — Tikhomirova, 1973a: 188 (checklist of species of USSR).

  • — Coiffait, 1974: 414 (characters; key to species).

  • — Smetana and Davies, 2000: 19, 28, 42 (characters; checklist of species of north temperate region).

ahrendti Wendeler, 1921: 156 (Physetops; Type locality: Jerusalem).

  • — Bernhauer, 1923: 120 (Physetops; characters in key; Palestine).

  • — Scheerpeltz, 1933: 1409 (Physetops; catalog).

  • — Coiffait, 1974: 418 (Physetops; characters; Palestine; Syria).

  • — Smetana and Davies, 2000: 42 (Physetops).

  • Distribution: Syria, Israel.

giganteus Semenow, 1906: 55 (Physetops; Type locality: australis prov. nostrae Transcaspicae, Turcomaniae; Nuchur pr. Artshman; Aschabad; fl Murgab pr. Merv; Sumbar. Afghanistan septentr.: Herat. Persiam saltem orientalem: Chorassan: Serachs; Zurabad; Meshhed; Langer; Bechars; Kerman mer.: distr. Bampur).

  • — Bernhauer and Schubert, 1914: 395 (Physetops; catalog).

  • — Bernhauer, 1923: 121 (Physetops; characters in key; Transcaspian region; Afghanistan).

  • — Cameron, 1932: 245 (Physetops; characters; Baluchistan; Afghanistan; Transcaspian region).

  • — Scheerpeltz, 1933: 1409 (Physetops; catalog).

  • — Scheerpeltz, 1961b: 34 (Physetops; Afghanistan).

  • — Scheerpeltz, 1963: 14 (Physetops; Afghanistan).

  • — Coiffait, 1974: 417 (Physetops; characters; Transcaspian region; Persia; Afghanistan; Pakistan).

  • — Smetana and Davies, 2000: 42 (Physetops).

  • Distribution: Iran, Afghanistan, Pakistan, Turkmenistan.

herculeanus Semenow, 1906: 55 (Physetops; subspecies giganteus; Type locality: Transcaucasia orientalis: Baku; Aresch prov. Elisabethpolitanae).

  • — Bernhauer and Schubert, 1914: 395 (Physetops; variety of giganteus).

  • — Bernhauer, 1923: 121 (Physetops; characters in key; Transcaucasus).

  • — Scheerpeltz, 1933: 1409 (Physetops; catalog).

  • — Coiffait, 1974: 418 (Physetops; characters; Caucasus; Transcaucasus).

  • — Smetana and Davies, 2000: 42 (Physetops).

  • Distribution: Azerbaijan.

spilleri Bernhauer, 1923: 121 (Physetops; Type locality: Turkestan).

  • — Scheerpeltz, 1933: 1409 (Physetops; catalog).

  • — Coiffait, 1974: 419 (Physetops; characters; Turkestan; Kazakhstan).

  • — Smetana and Davies, 2000: 42 (Physetops).

  • Distribution: Kazakhstan, “Turkestan”.

syriacus Pic, 1907, see: tataricus Pallas, 1773.

tartaricus Mannerheim, 1830, see: tataricus Pallas, 1773.

tataricus Pallas, 1773: 726 (Staphylinus; Type locality: Lacus salsus Inderiensis deserti Tatarici).

  • — Goeze, 1777: 723 (Staphylinus; characters).

  • — Gmelin, 1790: 2034 (Staphylinus; characters).

  • — Turton, 1802: 518 (Staphylinus; cited as tartaricus; characters; Tatary).

  • — Guérin-Méneville, 1829: pl. 9, fig. 3 (Staphylinus; illustration).

  • — Mannerheim, 1830: 33 [= 1831: 447] (Physetops).

  • — Nordmann, 1837: 14 (Physetops; characters; southern Siberia).

  • — Erichson, 1840: 414 (Ocypus; characters; desertis Russiae meridionalis).

  • — Guérin-Méneville, 1844a: 30 (Staphylinus; notes; Tatarie).

  • — Solsky, 1874: 174 (Physetops; characters; Turkestan).

  • — Bernhauer and Schubert, 1914: 395 (Physetops; catalog).

  • — Bernhauer, 1923: 121 (Physetops; characters in key; Russia; Transcaspian region).

  • — Scheerpeltz, 1933: 1409 (Physetops; catalog).

  • — Scheerpeltz, 1963: 14 (Physetops; Afghanistan).

  • — Coiffait, 1974: 419 (Physetops; characters; Transcaspian region).

  • — Smetana and Davies, 2000: 42 (Physetops).

  • Distribution: Syria, Afghanistan, Kazakhstan, “Transcaspian region”.

tartaricus Mannerheim, 1830: 33 (Physetops; [Note: An unjustified emendation]; Type locality: Tartaria).

  • — Mannerheim, 1831: 447 (Physetops; [Note: An unjustified emendation]; Tartaria).

  • — Bernhauer and Schubert, 1914: 395 (Physetops; synonym of tataricus).

  • — Coiffait, 1974: 419 (Physetops; synonym of tataricus).

syriacus Pic, 1907: 161 (Physetops; variety of tataricus; Type locality: Syrie).

  • — Bernhauer and Schubert, 1914: 395 (Physetops; variety of tataricus).

  • — Coiffait, 1974: 419 (Physetops; subspecies of tataricus; Syria).

transcaspius Bernhauer, 1910a: 72 (Physetops; Type locality: Transkaspien, Golodnaja Steppe).

  • — Bernhauer and Schubert, 1914: 395 (Physetops; catalog).

  • — Scheerpeltz, 1933: 1409 (Physetops; synonym of tataricus).

  • — Coiffait, 1974: 419 (Physetops; synonym of tataricus).

transcaspius Bernhauer, 1910, see: tataricus Pallas, 1773.

Phytolinus [4 species; Palaearctic Region]

Phytolinus Sharp, 1889: 119 (species included: lewisii). Type species: Phytolinus lewisii Sharp, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 393 (world catalog; one species).

  • — Winkler, 1925: 386 (catalog for Palaearctic region).

  • — Blackwelder, 1952: 308 (type species: lewisii).

  • — Tikhomirova, 1973a: 188 (checklist of species of USSR).

  • — Naomi, 1984: 382 (characters; notes; key to species).

  • — Shibata, 1984: 98 (checklist of species of Japan).

  • — Shibata, 1986a: 124 (supplemental list of species from Taiwan).

formosanus Naomi, 1984: 388 (Phytolinus; Type locality: Taiwan: Nantou Hsien, Lienhuachih).

  • Distribution: Taiwan.

lewisii Sharp, 1889: 121 (Phytolinus; Type locality: Kashiwagi; Nikko).

  • — Bernhauer and Schubert, 1914: 393 (Phytolinus; cited as lewisi; catalog).

  • — Nakane, 1963a: 93 (Phytolinus; characters; habitus photograph; Japan).

  • — Naomi, 1984: 385 (Phytolinus; characters; Japan).

  • Distribution: Japan.

tsuguriensis R. Dvořák, 1956: 30 (Eucibdelus; Type locality: Japan: Tukushima-Shikoku, Mount Tsuguri-san).

  • — Hayashi, 1997: 36 (Phytolinus; synonym of lewisii).

monticola Naomi, 1984: 386 (Phytolinus; Type locality: Mt. Fuji, Yamanashi Pref.).

  • Distribution: Japan.

tsuguriensis R. Dvořák, 1956, see: lewisii Sharp, 1889.

variegatus Sawada and Nakane, 1954: 8 (Phytolinus; Type locality: Mt. Omogo, Iyo, Shikoku).

  • — Nakane, 1963a: 93 (Phytolinus; characters; habitus photograph; Japan).

  • — Naomi, 1984: 387 (Phytolinus; characters; Japan).

  • Distribution: Japan.

Platydracus [178 species (1 doubtful); Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, and Oriental Regions]

Platydracus Thomson, 1858: 29 (species included: stercorarius). Type species: Staphylinus stercorarius Olivier, fixed by monotypy.

  • — Thomson, 1859: 23 (characters; type species: stercorarius).

  • — Thomson, 1860: 143 (Schizochilus; characters; Scandinavia).

  • — Fauvel, 1874: 172 [= 1874a: 398] (synonym of Staphylinus).

  • — Mulsant and Rey, 1876: 257 [= 1877: 113] (characters; key to French species).

  • — Heyden, Reitter, and Weise, 1891: 191 (subgenus of Staphylinus; list of species of Europe and Caucasus).

  • — Heyden, 1893: 41 (subgenus of Staphylinus; list of species of Siberian region).

  • — Heyden, 1896: 32 (subgenus of Staphylinus; list of species of Siberian region).

  • — Heyden, Reitter, and Weise, 1906: 169 (subgenus of Staphylinus; list of species of Europe and Caucasus).

  • — Reitter, 1909: 118 (subgenus of Staphylinus; key to species).

  • — Petri, 1912: 75 (subgenus of Staphylinus; list of species of Siebenbürgen).

  • — Bernhauer and Schubert, 1914: 374 (subgenus of Staphylinus).

  • — Winkler, 1925: 382 (catalog for Palaearctic region; subgenus of Staphylinus).

  • — Cameron, 1932: 180 (subgenus of Staphylinus).

  • — Scheerpeltz, 1933: 1381 (subgenus of Staphylinus).

  • — Chapman, 1934 (subgenus of Staphylinus; notes on classification and distribution of some species).

  • — Blackwelder, 1943: 443 (subgenus of Staphylinus; type species: stercorarius).

  • — Tottenham, 1949: 374, 420 (type species: stercorarius Olivier; subgenus of Staphylinus; checklist of British species).

  • — Blackwelder, 1952: 312 (type species: stercorarius).

  • — Coiffait, 1956e: 184, 186, 191 (valid genus; characters in key; notes; catalog of and key to French species).

  • — Hatch, 1957: 176, 178 (subgenus of Ocypus; characters; key to species of Pacific Northwest).

  • — Smetana, 1958: 260, 269 (subgenus of Staphylinus; characters).

  • — Palm, 1963b: 109 (subgenus of Staphylinus; characters; key to species of Sweden).

  • — Lohse, 1964: 194 (characters; key to central European species).

  • — Pototskaia, 1966b: 363 (subgenus of Staphylinus; larval characters in key).

  • — Pototskaia, 1967: 64 (subgenus of Staphylinus; larval characters).

  • — Levasseur, 1967b: 1631 (notes; characters).

  • — Kasule, 1970: 64, 72 (larval characters).

  • — Shibata, 1973c: 61 (checklist of species of Taiwan).

  • — Tikhomirova, 1973a: 187 (subgenus of Staphylinus; checklist of species of USSR).

  • — Coiffait, 1974: 385 (adult and larval characters; key to species of western Palaearctic region).

  • — Pope, 1977: 30 (list of British species).

  • — Topp, 1978: 311 (larval characters).

  • — Frank, 1979a (some larval characters and their use in classification).

  • — Muona, 1979: 15 (list of Scandinavian species).

  • — M. Dvořák, 1979: 116 (some species collected in Slovakia).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 92 (catalog; Poland).

  • — Uhlig and Vogler, 1981: 119 (list of some species collected in Germany; habitat and collecting notes).

  • — Hammond, 1984: 207 (checklist of species of Borneo).

  • — Tóth, 1984: 12 (characters; habitus; key to species of Hungary).

  • — Frank, 1986a: 370 (checklist of species of Florida).

  • — Muona and Viramo, 1986: 13 (list of species of northeastern Finland).

  • — Segers, 1986: 47 (checklist of species; Belgium).

  • — Ádám, 1987: 148 (list of two species with collecting notes; Hungary).

  • — Drugmand 1987b (comments on distribution of Belgian species).

  • — Campbell and Davies, 1991: 120 (checklist of species; Alaska and Canada).

  • — Silfverberg, 1992: 17 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Navarrete-Heredia and Márquez-Luna, 1993 (cave collections; Mexico).

  • — Siitonen, 1993: 230 (list of Finnish species).

  • — Smetana, 1993b: 49 (list of species of Czech Republic and Slovakia).

  • — Ciceroni and Zanetti, 1995: 30 (list of species of Italy).

  • — M. Hansen, 1996: 106 (list of Danish species).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 244 (additions to list of Danish species).

  • — Lundgren, 1998: 52 (list of species from Florida; records for other States listed for each species).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (list of species collected in 1997; Denmark).

  • — Smetana and Davies, 2000: 18, 24, 39 (characters; checklist of species of north temperate region).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 394 (27 Nearctic species; notes; characters in key).

Bemasus Mulsant and Rey, 1876: 259 [= 1877: 115] (species included: lutarius, meridionalis). Type species: Staphylinus lutarius Gravenhorst, fixed by subsequent designation by Blackwelder, 1943: 443.

  • — Bernhauer and Schubert, 1914: 374 (synonym of Platydracus).

  • — Blackwelder, 1943: 443 (synonym of Staphylinus; type species: lutarius).

  • — Blackwelder, 1952: 75 (subgenus of Platydracus; type species: lutarius).

  • — Coiffait, 1974: 385 (synonym of Platydracus).

Chitocompsus Bernhauer, 1913b: 232 (subgenus of Staphylinus; species included: polyphemus, giganteus, sjostedti). Type species: Staphylinus (Chitocompsus) polyphemus Bernhauer, fixed by subsequent designation by Blackwelder, 1943: 444.

  • — Bernhauer and Schubert, 1914: 374 (subgenus of Staphylinus).

  • — Cameron, 1932: 180 (subgenus of Staphylinus).

  • — Blackwelder, 1943: 443 (subgenus of Staphylinus; type species: polyphemus).

  • — Blackwelder, 1952: 100 (subgenus of Platydracus; type species: polyphemus).

Nesiolinus Bernhauer, 1915e: 122 (subgenus of Staphylinus; speciesincluded: bakeri, pulcherrimus). Type species: Staphylinus (Nesiolinus) bakeri Bernhauer, fixed by subsequent designation by Blackwelder, 1943: 444.

  • — Cameron, 1932: 180 (subgenus of Staphylinus).

  • — Scheerpeltz, 1933: 1381 (subgenus of Staphylinus).

  • — Blackwelder, 1943: 443 (subgenus of Staphylinus; type species: bakeri).

  • — Blackwelder, 1952: 261 (subgenus of Platydracus; type species: bakeri).

Euryolinus Bernhauer, 1915l: 297 (subgenus of Staphylinus; species included: semicyaneus). Type species: Staphylinus (Euryolinus) semicyaneus Bernhauer, fixed by monotypy.

  • — Scheerpeltz, 1933: 1381 (subgenus of Staphylinus).

  • — Blackwelder, 1943: 444 (subgenus of Staphylinus; type species).

  • — Blackwelder, 1952: 160 (subgenus of Platydracus; type species).

Plesiolinus Bernhauer, 1917: 93 (subgenus of Staphylinus; species included: costaricensis, panamensis, tarsalis, isolitus, cordilleranus, acupunctipennis). Type species: Staphylinus (Plesiolinus) costaricensis Bernhauer, fixed by subsequent designation by Blackwelder, 1943: 444.

  • — Cameron, 1932: 180 (subgenus of Staphylinus).

  • — Scheerpeltz, 1933: 1381 (subgenus of Staphylinus).

  • — Blackwelder, 1943: 443 (subgenus of Staphylinus; type species: costaricensis).

  • — Blackwelder, 1952: 314 (subgenus of Platydracus; type species: costaricensis).

Neotasgius G. Müller, 1925: 41 (subgenus of Staphylinus; species included: brevicornis). Type species: Ocypus brevicornis Weise, fixed by monotypy.

  • — Cameron, 1932: 180 (subgenus of Staphylinus).

  • — Scheerpeltz, 1933: 1381 (subgenus of Staphylinus).

  • — Blackwelder, 1943: 445 (subgenus of Ocypus; type species: brevicornis).

  • — Blackwelder, 1952: 261 (type species: brevicornis).

  • — Shibata, 1984: 87 (subgenus of Ocypus; checklist of species of Japan).

  • — Smetana and Davies, 2000: 24, 39 (synonym of Platydracus).

Chaetodracus G. Müller, 1926a: 27 (subgenus of Staphylinus; species included: patricius). Type species: Staphylinus patricius Bernhauer, fixed by monotypy.

  • — Scheerpeltz, 1933: 1381 (subgenus of Staphylinus).

  • — Blackwelder, 1943: 443 (subgenus of Staphylinus; type species: patricius).

  • — Blackwelder, 1952: 98 (subgenus of Platydracus; type species: patricius).

  • — Smetana and Davies, 2000: 18, 25, 41 (subgenus of Platydracus; characters).

Paraplatydracus Levasseur, 1980: 369 (subgenus of Platydracus; species included: girardi). Type species: Platydracus (Paraplatydracus) girardi Levasseur, fixed by original designation and monotypy.

Tropoplatydracus Levasseur, 1980: 367 (subgenus of Platydracus; species included: lecordieri). Type species: Platydracus (Tropoplatydracus) lecordieri Levasseur, fixed by original designation and monotypy.

acupunctipennis Bernhauer, 1907a: 285 (Staphylinus; Type locality: Brasilien: Ypiranga).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • — Bernhauer, 1917: 93 (Staphylinus; subgenus Plesiolinus).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Brazil.

aeneicollis Bernhauer, 1911, see: asemus Kraatz, 1859.

aeneipennis Cameron, 1930b: 164 (Staphylinus; Type locality: Sarawak, Mt. Matang. Also cited from North Borneo and S.E. Borneo).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • — Hammond, 1984: 195 (Platydracus; Borneo).

  • Distribution: Borneo.

aeneoacreus Cameron, 1930b: 164 (Staphylinus; Type locality: North Borneo: Bettotan).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • — Hammond, 1984: 195 (Platydracus; Borneo).

  • Distribution: Borneo.

aeneoniger Bernhauer, 1933a: 43 (Staphylinus; subgenus Platydracus; Type locality: Tatsienlu-Kiulung).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: China.

aericeps Stephens, 1832, see: latebricola Gravenhorst, 1806.

allardi Levasseur, 1967b: 1632 (Platydracus; Type locality: région de Kolwezi, Katanga, Manika).

  • Distribution: Congo.

amamiensis Ito, 1982: 61 (Platydracus; Type locality: Hatsuno, Amami-Oshima, Is.).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Japan.

armeniacus Sharp, 1873, see: meridionalis Rosenhauer, 1847.

asemus Kraatz, 1859: 77 (Staphylinus; Type locality: India borealis).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • — Cameron, 1932: 194 (Staphylinus; subgenus Platydracus; characters; India).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: India.

aeneicollis Bernhauer, 1911: 87 (Staphylinus; [preoccupied]; Type locality: United Provinces Mussoorie).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; synonym of asemus).

  • — Smetana and Davies, 2000: 39 (Platydracus; synonym of asemus).

associatus Bernhauer, 1937: 21 (Staphylinus; subgenus Platydracus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft.).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Indonesia.

aureofasciatus Motschulsky, 1862: 40 (Staphylinus, cited as Staphilinus; Type locality: Not cited, but Amur according to Motschulsky, 1870: 49).

  • — Fauvel, 1874: 177 [= 1874a: 403] (Staphylinus; synonym of fulvipes).

  • — Ganglbauer, 1895: 426 (Staphylinus; synonym of fulvipes).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; synonym of fulvipes).

  • — G. Müller, 1932: 76 (Staphylinus; subgenus Platydracus; notes).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Russia.

teter Bondroit, 1913: 90 (Staphylinus; Type locality: l'Ussuri).

  • — Gridelli, 1932: 76 (Staphylinus; synonym of aureofasciatus).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; synonym of aureofasciatus).

  • — Smetana and Davies, 2000: 39 (Platydracus; synonym of aureofasciatus).

aureovillosus G. Müller, 1932 [Note: Nomen nudum], see: fulvipes Scopoli, 1763.

aurichalceus Cameron, 1941c: 382 (Staphylinus; subgenus Platydracus; Type locality: Philippines: Samar: Catbalogan).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Philippines.

auripennis Kraatz, 1859, see: goryi Laporte, 1835.

auroaeneus Cameron, 1938b: 150 (Staphylinus; subgenus Platydracus; Type locality: E. Java: Res. Kediri, Pandam Aroem).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Indonesia.

auronotatus Fauvel, 1895: 256 (Staphylinus; Type locality: Bírmanie, Carin Cheba, 900–1100 m).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • — Cameron, 1932: 196 (Staphylinus; subgenus Platydracus; characters; Burma).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Myanmar.

aurosericans Fairmaire, 1891a: cxc (Staphylinus; Type locality: Tchang-Yang).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: China.

azureus Gebler, 1829, see: fulvipes Scopoli, 1763.

badius Mannerheim, 1830, see: zonatus Gravenhorst, 1802.

bakeri Bernhauer, 1915e: 122 (Staphylinus; subgenus Nesiolinus; Type locality: Luzon: Laguna, Mount Maquiling).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • — Blackwelder, 1952: 261 (type species of Nesiolinus, a subgenus of Platydracus).

  • Distribution: Philippines.

banghaasi G. Müller, 1932, see: muellerianus Scheerpeltz, 1933.

basicornis Fauvel, 1895: 255 (Staphylinus; Type locality: Bírmanie, Teinzo; Carin Cheba, 400–1100 m; Carin Ghécu, 1300–1400 m; Inde, Sylhet).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • — Cameron, 1932: 197 (Staphylinus; subgenus Platydracus; characters; Burma; India).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • — Scheerpeltz, 1965: 109 (Staphylinus; subgenus Platydracus; Burma).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Myanmar, India.

basicornis Bernhauer, 1932, see: purpureoaureus Bernhauer, 1915.

becquarti Bernhauer, 1938b: 52 (Staphylinus; subgenus Platydracus; Type locality: Hopeh-Nord: Trappe, Yangkiaping).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: China.

bengalensis Bernhauer, 1914c: 100 (Staphylinus; Type locality: Bengal).

  • — Cameron, 1932: 194 (Staphylinus; subgenus Platydracus; characters; Bengal).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: India.

bicinctus Rossi, 1792, see: fulvipes Scopoli, 1763.

biguttatus Bernhauer, 1937: 304 (Staphylinus; subgenus Platydracus; [preoccupied]; Type locality: Uganda: Gulu).

  • — Herman, 2001: 14, 48 (Platydracus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: Uganda.

birmanus Fauvel, 1895: 255 (Staphylinus; Type locality: Bírmanie, Mandalay; Bhamo, Metanja; Shwego-myo).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • — Cameron, 1932: 198 (Staphylinus; subgenus Platydracus; characters; Burma).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • — Coiffait, 1977b: 210 (Platydracus; Bhutan).

  • — Coiffait, 1982a: 233 (Platydracus; India).

  • Distribution: Myanmar, Bhutan, India.

biseriatus Sharp, 1884: 374 (Staphylinus; Type locality: Guatemala, near the city).

  • — Sharp, 1887: 785 (Staphylinus; Mexico).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • — Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 59 (Platydracus; necrophilous species; distribution in Mexico).

  • Distribution: Guatemala, Mexico.

bocandei Fagel, 1951c: 235 (Staphylinus; subgenus Platydracus; Type locality: Sénégal. Also cited from French Guinea).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Guinea, Senegal.

bodongi Bernhauer, 1906a: 187 (Staphylinus; Type locality: Rhodesia, Mashonaland, Umtali).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • — Gridelli, 1939: 321 (Staphylinus; subgenus Platydracus; characters; Abyssinia).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Zimbabwe, Ethiopia.

brachycerus Smetana and Davies, 2000: 24, 39 (Platydracus; subgenus Platydracus; replacement name for brevicornis Weise).

  • Distribution: Japan, Korea.

brevicornis Weise, 1877a: 367 (Ocypus; [preoccupied]; Type locality: Hakodade).

  • — Sharp, 1889: 110 (Ocypus; Japan).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1391 (Staphylinus; catalog).

  • — Nakane, 1963a: 92 (Ocypus; characters; habitus photograph; Japan).

  • — Yuh, Paik, Kwon, and Lee, 1985: 244 (Ocypus; Korea).

  • — Smetana and Davies, 2000: 24, 39 (Platydracus; synonym of brachycerus).

brachypterus Kraatz, 1859, see: brevipennis Smetana and Davies, 2000.

bredoi Fagel, 1950: 6 (Staphylinus; subgenus Platydracus; Type locality: Tanganyika Territory, lac Rukwa).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Tanzania.

brevicornis Motschulsky, 1862a: 11 (Trichoderma; Type locality: Japan; [Note: See lectotype designation by Gusarov, 1995a]).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog).

  • — Smetana, 1959d: 411 (Ontholestes; republication of original description).

  • — Gusarov, 1995a: 92 [= 1996: 54] (Platydracus; lectotype designation).

  • — Watanabe, 1998: 314 (Platydracus; Japan).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Japan, Korea, China.

paganus Sharp, 1874: 30 (Staphylinus; Type locality: Japan; [Note: See lectotype designation by Gusarov, 1995a]).

  • — Fauvel, 1875a: XXVII [= 1875b: 229] (Staphylinus; synonym of dauricus).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; catalog).

  • — Kamiya, 1936: 1, 4 (Staphylinus; characters; Manchoukuo).

  • — Cameron, 1949b: 468 (Staphylinus; Japan).

  • — Nakane, 1963a: 92 (Platydracus; characters; habitus photograph; Japan).

  • — Yuh, Paik, Kwon, and Lee, 1985: 244 (Staphylinus; Korea).

  • — J. Li, 1993: 38 (Staphylinus; China).

  • — Kanehisa, Tsumuki, and Kawazu, 1994 (Staphylinus; chemical secretions).

  • — Gusarov, 1995a: 92 [= 1996: 55] (Platydracus; lectotype designation; synonym of brevicornis).

  • — Smetana and Davies, 2000: 39 (Platydracus; synonym of brevicornis).

brevicornis Weise, 1877, see: brachycerus Smetana and Davies, 2000.

brevipennis Smetana and Davies, 2000: 25, 39 (Platydracus; subgenus Platydracus; replacement name for brachypterus Kraatz).

  • Distribution: India.

brachypterus Kraatz, 1859: 76 (Staphylinus; [preoccupied]; Type locality: India borealis).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • — Cameron, 1932: 200 (Staphylinus; subgenus Platydracus; characters; India).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 25, 39 (Platydracus; synonym of brevipennis).

bruchi Bernhauer, 1934b: 214 (Staphylinus; subgenus Platydracus; Type locality: Argentinien: Salta).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Argentina.

bryanti Cameron, 1918a: 104 (Staphylinus; Type locality: Ceylon: Kandy).

  • — Cameron, 1932: 190 (Staphylinus; subgenus Platydracus; characters; Ceylon).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Sri Lanka.

caliginosus Erichson, 1839b: 388 (Staphylinus; Type locality: Mexico).

  • — Sharp, 1884: 373 (Staphylinus; characters; notes; Mexico; Guatemala).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; catalog).

  • — Newton, 1997: 23 (Platydracus; notes; Hawaii; characters; also occurs in Nevada, Arizona, New Mexico, Texas; Mexico; Central America; Colombia).

  • — Nishida, 1997: 63 (Platydracus; Hawaii).

  • — Schmidt, 1999: 108 (Platydracus; food for adults and larvae used for rearing species; rearing methods).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: USA, Mexico, Guatemala, Costa Rica, Colombia, Hawaii.

ejulans Tottenham, 1939: 170 (Staphylinus; Type locality: Costa Rica: 1,000–1,200 m).

  • — Newton, 1997: 23 (Platydracus; synonym of caliginosus).

  • — Smetana and Davies, 2000: 39 (Platydracus; synonym of caliginosus).

campestris Coiffait, 1977b: 230 (Platydracus; Type locality: Bhutan: Phuntsholing, 200–400 m).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Bhutan.

cantharophagus Fagel, 1950: 1 (Staphylinus; subgenus Platydracus; Type locality: Congo belge: Parc National de l'Upemba, Lusinga, 1760 m).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Congo.

carinthiacus Lacordaire, 1835, see: chalcocephalus Fabricius, 1801.

caseyi Scheerpeltz, 1933, see: zonatus Gravenhorst, 1802.

castaneus Nordmann, 1837: 54 (Staphylinus; Type locality: Mexico).

  • — Erichson, 1839b: 379 (Staphylinus; characters; Mexico).

  • — Sharp, 1884: 377 (Staphylinus; Mexico).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; catalog).

  • — Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 59 (Platydracus; necrophilous species; distribution in Mexico).

  • Distribution: Mexico.

catalonicus Coiffait, 1967c: 170 (Platydracus; Type locality: Tona, Prov. de Barcelone).

  • — Coiffait, 1974: 398 (Platydracus; characters; Spain).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Spain.

centralis Sharp, 1884: 384 (Staphylinus; Type locality: Mexico, Oaxaca; Guatemala, San Geronimo).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Mexico, Guatemala.

modestus Sharp, 1884: 383 (Staphylinus; Type locality: Mexico, Jalapa).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 39 (Platydracus; synonym of centralis).

fusiformis Casey, 1915: 429 (Staphylinus; Type locality: Mexico: Durango City).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 39 (Platydracus; synonym of centralis).

chalcescens Sharp, 1889: 117 (Staphylinus; Type locality: Nikko).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1383 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Japan.

chalcocephalus Fabricius, 1801: 593 (Staphylinus; [Note: Fabricius attributed this name to De Geer who did not describe the species. The description to which Fabricius referred was that of aeneocephalus De Geer. Fabricius also listed other references but all of them referred to aeneocephalus. Later Gmelin, 1790, cited the species in the same manner]; Type locality: Germania).

  • — Fabricius, 1793: 522 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 376, cited aeneocephalus Fabricius as a synonym of chalcocephalus Fabricius but Fabricius did not describe the species as new, he attributed it to De Geer]; characters; Germany).

  • — Gravenhorst, 1806: 113 (Staphylinus; synonym of aeneocephalus).

  • — Gyllenhal, 1810: 296 (Staphylinus; synonym of stercorarius Olivier).

  • — Curtis, 1829: 24 (Staphylinus; catalog; Britain).

  • — Stephens, 1832: 206 (Staphylinus; characters; England).

  • — Brullé, 1832: 130 (Staphylinus; Greece).

  • — Runde, 1835: 2 (Staphylinus; characters; Germany).

  • — Lacordaire, 1835: 374 (Emus; characters; France).

  • — Curtis, 1839: pl. 758 (Staphylinus; England).

  • — Erichson, 1839a: 436 (Staphylinus; characters; Germany).

  • — Heer, 1839: 252 (Staphylinus; characters; Switzerland).

  • — Erichson, 1839b: 381 (Staphylinus; characters; Germany; Prussia; France).

  • — Stephens, 1839: 387 (Staphylinus; characters; England).

  • — Holme, 1842: 116, 117 (Staphylinus; notes).

  • — Redtenbacher, 1849: 695 (Staphylinus; characters; Austria).

  • — Küster, 1853: 52 (Staphylinus; characters; Prussia; Germany; France; Switzerland).

  • — Fairmaire and Laboulbène, 1856: 508 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 544 (Staphylinus; characters; Germany).

  • — Redtenbacher, 1857: 185 (Staphylinus; characters; Austria).

  • — Fauvel, 1874: 179 [= 1874a: 405] (Staphylinus; characters; France; Austria).

  • — Redtenbacher, 1874: 204 (Staphylinus; characters; Austria).

  • — Seidlitz, 1875: 269 (Leistotrophus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 264 [= 1877: 120] (Platydracus; characters; France).

  • — Ganglbauer, 1895: 428 (Staphylinus; subgenus Platydracus; characters; middle Europe).

  • — Reitter, 1909: 119 (Staphylinus; subgenus Platydracus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; catalog).

  • — Johansen, 1914: 380 (Staphylinus; subgenus Platydracus; characters; Denmark).

  • — Portevin, 1929: 349 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1383 (Staphylinus; catalog).

  • — V. Hansen, 1952: 99 (Staphylinus; characters; Denmark).

  • — Coiffait, 1956e: 193, 194 (Platydracus; characters).

  • — Smetana, 1958: 270 (Staphylinus; subgenus Platydracus; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 110 (Staphylinus; subgenus Platydracus; characters).

  • — Lohse, 1964: 194 (Platydracus; characters; central Europe).

  • — Smetana, 1965c: 27 (Staphylinus; subgenus Platydracus; Anatolia).

  • — Horion, 1965: 202 (Staphylinus; subgenus Platydracus; Denmark; Holland; Belgium; France; Germany; Switzerland; Austria; Italy; Slovenia; Dalmatia; Croatia; Bosnia; Serbia; Bulgaria; Greece; Romania; Hungary; Czechoslovakia; Poland; Russia; Ukraine; Caucasus).

  • — Scheerpeltz, 1968a: 80 (Staphylinus; subgenus Platydracus; Austria).

  • — Smetana, 1968b: 87 (Staphylinus; subgenus Platydracus; Anatolia).

  • — Szujecki, 1968a: 732 (Staphylinus; Poland).

  • — Tikhomirova, 1973a: 187 (Staphylinus; subgenus Platydracus; in checklist of species of USSR).

  • — Coiffait, 1974: 396 (Platydracus; characters; Europe; Britain; Denmark; Caucasus).

  • — Topp, 1979: 26 (Platydracus; development; diapause).

  • — Muona, 1979: 15 (Platydracus; Denmark; Baltic states).

  • — Petrenko, 1980a: 60, 62 (Staphylinus; subgenus Platydracus; characters; Ukraine).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 92 (Platydracus; catalog; Poland).

  • — Uhlig and Vogel, 1981: 119 (Platydracus; Germany).

  • — Tóth, 1984: 15 (Platydracus; characters; Hungary).

  • — Lucht, 1987: 107 (Platydracus; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark).

  • — Schülke and Uhlig, 1988: 12 (Platydracus; notes; Germany).

  • — Silfverberg, 1992: 17 (Platydracus; Denmark; Baltic states).

  • — Smetana, 1993b: 49 (Platydracus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 30 (Platydracus; Italy).

  • — Pilon, 1998: 69 (Platydracus; ecological notes; Italy).

  • — Silfverberg, 1999: 50 (Platydracus; Estonia).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Europe, Caucasus, Turkey.

ochropterus Germar, 1824: 34 (Staphylinus; Type locality: Germania).

  • — Germar, 1823b: 14 (Staphylinus; characters; Germany).

  • — Curtis, 1839: pl. 758 (Staphylinus; synonym of chalcocephalus).

  • — Nordmann, 1837: 59 (Staphylinus; characters; Carinthia; Prussia).

  • — Erichson, 1839a: 437 (Staphylinus; synonym of chalcocephalus).

  • — Heer, 1839: 252 (Staphylinus; synonym of chalcocephalus).

  • — Erichson, 1839b: 381 (Staphylinus; synonym of chalcocephalus).

  • — Küster, 1853: 52 (Staphylinus; synonym of chalcocephalus).

  • — Kraatz, 1857d: 544 (Staphylinus; synonym of chalcocephalus).

  • — Ganglbauer, 1895: 428 (Staphylinus; synonym of chalcocephalus).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; synonym of chalcocephalus).

  • — Smetana, 1958: 270 (Staphylinus; synonym of chalcocephalus).

  • — Coiffait, 1974: 397 (Platydracus; synonym of chalcocephalus).

carinthiacus Lacordaire, 1835: 365 (Emus; Type locality: Paris).

  • — Erichson, 1839a: 437 (Staphylinus; synonym of chalcocephalus).

  • — Heer, 1839: 252 (Staphylinus; synonym of chalcocephalus).

  • — Erichson, 1839b: 381 (Staphylinus; synonym of chalcocephalus).

  • — Küster, 1853: 52 (Staphylinus; synonym of chalcocephalus).

  • — Fairmaire and Laboulbène, 1856: 508 (Staphylinus; synonym of chalcocephalus).

  • — Kraatz, 1857d: 544 (Staphylinus; synonym of chalcocephalus).

  • — Ganglbauer, 1895: 428 (Staphylinus; synonym of chalcocephalus).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; synonym of chalcocephalus).

  • — Smetana, 1958: 270 (Staphylinus; synonym of chalcocephalus).

  • — Coiffait, 1974: 397 (Platydracus; synonym of chalcocephalus).

chapmani Bernhauer, 1933a: 47 (Staphylinus; subgenus Abemus; Type locality: Yunnan-Fou).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: China.

chinensis Bernhauer, 1914c: 101 (Staphylinus; Type locality: China: Chusan Is.).

  • — Scheerpeltz, 1933: 1383 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: China.

chrysopterus Brullé, 1842, see: chrysotrichopterus Scheerpeltz, 1933.

chrysotrichopterus Scheerpeltz, 1933: 1384 (Staphylinus; replacement name for chrysopterus Brullé. Also cited from Argentina).

  • — Scheerpeltz, 1960c: 120 (Staphylinus; subgenus Platydracus; Bolivia).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Bolivia, Argentina.

chrysopterus Brullé, 1842: 80 (Staphylinus; [preoccupied]; Type locality: Bolivia: Chuquisaca).

  • — Erichson, 1839b: 384 (Staphylinus; synonym of nobilis).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; synonym of chrysotrichopterus).

  • — Herman, 2001: 14 (Platydracus; synonym of chrysotrichopterus).

cicatricosus LeConte, 1863, see: praelongus Mannerheim, 1830.

cingulus Comolli, 1837, see: fulvipes Scopoli, 1763.

cinnamopterus Gravenhorst, 1802: 164 (Staphylinus; Type locality: Baltimore).

  • — Gravenhorst, 1806: 113 (Staphylinus; characters).

  • — Mannerheim, 1830: 23 (Staphylinus; North America).

  • — Mannerheim, 1831: 437 (Staphylinus; North America).

  • — Nordmann, 1837: 59 (Staphylinus; America borealis).

  • — Erichson, 1839b: 390 (Staphylinus; characters; America septentrionalis).

  • — Provancher, 1877: 247 (Staphylinus; cited as scinnamopterus; characters; Quebec).

  • — Horn, 1879: 195 (Staphylinus; characters; Maine to California).

  • — Fall and Cockerell, 1907: 166 (Staphylinus; New Mexico).

  • — Blatchley, 1910: 377 (Staphylinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • — Notman, 1919: 135 (Staphylinus; New York).

  • — Lesage, 1977 (Staphylinus; larval and pupal characters; collecting notes; Quebec; Ontario).

  • — Dearborn and Donahue, 1994: 15 (Platydracus; Maine).

  • — Downie and Arnett, 1996: 386 (Staphylinus; characters; USA).

  • — Lundgren, 1998: 52 (Platydracus; Florida).

  • — Schmidt, 1999: 108 (Platydracus; food for adults and larvae used for rearing species; rearing methods).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Canada, USA.

circumcinctus Bernhauer, 1914c: 102 (Staphylinus; Type locality: Ostindien).

  • — Bernhauer, 1922c: 233 (Staphylinus; Formosa).

  • — Cameron, 1932: 187 (Staphylinus; subgenus Platydracus; characters; East Indies).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; catalog).

  • — Nakane, 1963a: 92 (Platydracus; characters; habitus photograph; Japan).

  • — Yuh, Paik, Kwon, and Lee, 1985: 244 (Staphylinus; Korea).

  • Distribution: Japan, Taiwan, “East Indies”.

coeruleipennis Coiffait, 1983d: 167 (Platydracus; Type locality: Nepal: Ganesh Himal, entre Salmi et Rupchet, 1500 à 3600 m).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Nepal.

collaris Coiffait, 1977b: 231 (Platydracus; Type locality: Bhutan: Paro, 2300 m).

  • — Coiffait, 1982a: 233 (Platydracus; characters of male; Bhutan).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Bhutan.

comes LeConte, 1863a: 36 (Staphylinus; Type locality: Southern States).

  • — Horn, 1879: 194 (Staphylinus; characters; Georgia; Florida).

  • — Fall and Cockerell, 1907: 166 (Staphylinus; New Mexico).

  • — Blatchley, 1910: 378 (Staphylinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • — Voris, 1939a: 288, 298 (Staphylinus; larval and pupal characters).

  • — Downie and Arnett, 1996: 386 (Staphylinus; characters; USA).

  • — Lundgren, 1998: 52 (Platydracus; Florida).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: USA, Mexico.

consularis Bernhauer, 1915c: 72 (Staphylinus; subgenus Platydracus; Type locality: Japan: Harima).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Japan.

costaricensis Bernhauer, 1917: 93 (Staphylinus; subgenus Plesiolinus; Type locality: Costa-Rica: Orosi, 1300 m).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; catalog).

  • — Blackwelder, 1952: 314 (type species of Plesiolinus, a subgenus of Platydracus).

  • Distribution: Costa Rica.

contiguus Cameron, 1938b: 151 (Staphylinus; subgenus Platydracus; Type locality: E. Java: Res. Kediri, Pandan Aroem).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Indonesia.

cordilleranus Bernhauer, 1917i: 108 (Staphylinus; subgenus Platydracus; Type locality: Kolumbien: Ostkordilleren, Huaso, 500 m; Muzo, Rio Cantinere, 500 m).

  • — Bernhauer, 1917: 93 (Staphylinus; subgenus Plesiolinus).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Colombia.

crebrepunctatus Motschulsky, 1860, see: stercorarius Olivier, 1795.

cubae Jacquelin du Val, 1857, see: tomentosus Gravenhorst, 1802.

cupripennis Melsheimer, 1844, see: violaceus Olivier, 1795.

curticollis Bernhauer, 1917, see: tarsalis Mannerheim, 1843.

curticornis Fauvel, 1895: 256 (Staphylinus; Type locality: Bírmanie, Carin Cheba, 900–1100 m).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • — Cameron, 1932: 197 (Staphylinus; subgenus Platydracus; characters; Burma).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; catalog).

  • — Coiffait, 1982a: 233 (Platydracus; India).

  • Distribution: Myanmar, India.

darfourensis Levasseur, 1967b: 1636 (Platydracus; Type locality: Sunni, Djebel Marra, province de Darfour, Sudan).

  • Distribution: Sudan.

dauricus Mannerheim, 1830: 22 (Staphylinus; Type locality: Dauria).

  • — Mannerheim, 1831: 436 (Staphylinus; Dauria).

  • — Nordmann, 1837: 62 (Staphylinus; Dauria).

  • — Erichson, 1839b: 386 (Staphylinus; characters; Siberia).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; catalog).

  • — Aleksandrov, 1934: 153 (Staphylinus; China).

  • — Bernhauer, 1936: 5 (Staphylinus; subgenus Platydracus; Siberia; Japan).

  • — Smetana, 1963c: 299 (Staphylinus; subgenus Platydracus; Mongolia).

  • — Scheerpeltz, 1964: 28 (Staphylinus; subgenus Platydracus; Mongolia).

  • — Smetana, 1965e: 174 (Staphylinus; subgenus Platydracus; Mongolia).

  • — Smetana, 1967d: 209 (Staphylinus; subgenus Platydracus; Mongolia).

  • — Coiffait, 1974: 394 (Platydracus; characters; Siberia; Mongolia).

  • — Smetana 1975c: 173 (Staphylinus; subgenus Platydracus; Mongolia).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Russia, Mongolia, China, Japan.

decipiens Kraatz, 1859: 75 (Staphylinus; Type locality: Ceylan).

  • — Fauvel, 1903: 157 (Staphylinus; Ceylon).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • — Cameron, 1932: 189 (Staphylinus; subgenus Platydracus; characters; Ceylon).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Sri Lanka.

demissus G. Müller, 1925: 42 (Staphylinus; subgenus Platydracus; Type locality: Sibiria orientalis, Sotka Gora).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; catalog).

  • — Smetana, 1967d: 209 (Staphylinus; subgenus Platydracus; Mongolia).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Russia, Mongolia.

dohertyi Cameron, 1932: 197 (Staphylinus; subgenus Platydracus; Type locality: Burma: Ruby Mines).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; catalog).

  • — Last, 1961: 306 (Platydracus; Thailand).

  • Distribution: Myanmar, Thailand.

drescheri Bernhauer, 1937: 21 (Staphylinus; subgenus Platydracus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft.).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Indonesia.

dudgeoni Cameron, 1932: 196 (Staphylinus; subgenus Platydracus; Type locality: Punjab: Kangra Valley, alt. 4500 feet).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: India.

ejulans Tottenham, 1939, see: caliginosus Erichson, 1839.

erichsoni Boheman, 1848: 275 (Staphylinus; Type locality: Caffraria interior).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • — Bernhauer, 1947: 158 (Staphylinus; Deutsch-Ostafrika).

  • — Cameron, 1950g: 43 (Staphylinus; subgenus Platydracus; Belgian Congo).

  • — Tottenham, 1956: 229 (Staphylinus; Ruanda).

  • — Tottenham, 1962: 144 (Staphylinus; Tanganyika).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: South Africa, Congo, Rwanda, Tanzania.

erythropennis Mannerheim, 1830, see: maculosus Gravenhorst, 1802.

erythropus Paykull, 1800, see: fulvipes Scopoli, 1763.

exulans Erichson, 1839b: 374 (Staphylinus; Type locality: America septentrionalis).

  • — Horn, 1879: 197 (Staphylinus; characters; Canada to Georgia).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; catalog).

  • — Downie and Arnett, 1996: 386 (Staphylinus; characters; USA).

  • — Lundgren, 1998: 52 (Platydracus; Florida).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: USA.

falcimaculatus Bernhauer, 1937b: 617 (Staphylinus; subgenus Platydracus; Type locality: Deutsch-Ostafrika: Uka).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Tanzania.

fauvelianus Fagel, 1950: 5 (Staphylinus; subgenus Platydracus; Type locality: Guinée).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Guinea.

femoratus Fabricius, 1801: 594 (Staphylinus; Type locality: Carolina).

  • — Gravenhorst, 1806: 122 (Staphylinus; characters; United States).

  • — Nordmann, 1837: 31 (Staphylinus; United States).

  • — Erichson, 1839b: 393 (Staphylinus; characters; United States).

  • — Horn, 1879: 193 (Staphylinus; characters; Georgia).

  • — Blatchley, 1910: 377 (Staphylinus; characters; Ohio).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; catalog).

  • — Downie and Arnett, 1996: 386 (Staphylinus; characters; USA).

  • — Lundgren, 1998: 52 (Platydracus; Florida).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Canada, USA.

varipes Sachse, 1852: 126 (Staphylinus; Type locality: Georgia).

  • — Gemminger and Harold, 1868: 578 (Staphylinus; synonym of varipes).

  • — Provancher, 1877: 729 (Staphylinus; characters; Quebec).

  • — Horn, 1879: 193 (Staphylinus; synonym of femoratus).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; synonym of femoratus).

  • — Smetana and Davies, 2000: 39 (Platydracus; synonym of femoratus).

fenyesi Bernhauer, 1917, see: violaceus Gravenhorst, 1802.

fervidus Sharp, 1884: 377 (Staphylinus; Type locality: Mexico, Cordova, Mirador).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; catalog).

  • — Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 59 (Platydracus; necrophilous species; distribution in Mexico).

  • Distribution: Mexico.

flavopilosus Cameron, 1932: 192 (Staphylinus; subgenus Platydracus; Type locality: Burma: Tenasserim, Tavoy).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Myanmar.

flavopunctatus Latreille, 1804: 297 (Staphylinus; Type locality: France).

  • — Erichson, 1839b: 381 (Staphylinus; synonym of lutarius).

  • — Kraatz, 1857d: 543 (Staphylinus; synonym of lutarius).

  • — Ganglbauer, 1895: 428 (Staphylinus; subgenus Platydracus; characters; Tyrol; middle Europe; Mediterranean region).

  • — Reitter, 1909: 118 (Staphylinus; subgenus Platydracus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; catalog).

  • — Fagel, 1955a: 85 (Platydracus; Greece).

  • — Coiffait, 1956e: 193, 194 (Platydracus; characters).

  • — Smetana, 1958: 269 (Staphylinus; subgenus Platydracus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 194 (Platydracus; characters; central Europe).

  • — Horion, 1965: 205 (Staphylinus; subgenus Platydracus; Turkey; France; Portugal; Switzerland; Sardinia; Italy; Dalmatia; Bosnia Hercegovina; Albania; Greece; Serbia; Slovakia; Austria; Germany ?).

  • — Smetana, 1967i: 556 (Staphylinus; subgenus Platydracus; notes; Turkey).

  • — Smetana, 1968a: 158 (Staphylinus; subgenus Platydracus; Anatolia).

  • — Smetana, 1971c: 263 (Staphylinus; subgenus Platydracus; Iran).

  • — Jarrige, 1971: 496 (Platydracus; Iran).

  • — Coiffait, 1974: 395 (Platydracus; characters; Europe; Mediterranean region; Asia Minor).

  • — Petrenko, 1980: 82 (Staphylinus; Caucasus).

  • — Tóth, 1984: 17 (Platydracus; characters; Hungary).

  • — Lucht, 1987: 107 (Platydracus; checklist; Czechoslovakia; Austria; Switzerland; France).

  • — Smetana, 1993b: 49 (Platydracus; Slovakia).

  • — Ciceroni and Zanetti, 1995: 30 (Platydracus; Italy).

  • — Pilon, 1998: 69 (Platydracus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Europe, Armenia, Turkey, Iran.

lutarius Gravenhorst, 1806: 115 (Staphylinus; Type locality: Not cited).

  • — Mannerheim, 1830: 23 (Staphylinus; France; Yugoslavia).

  • — Mannerheim, 1831: 437 (Staphylinus; France; Yugoslavia).

  • — Nordmann, 1837: 58 (Staphylinus; characters; Europe).

  • — Heer, 1839: 251 (Staphylinus; characters; Switzerland).

  • — Erichson, 1839b: 381 (Staphylinus; characters; Austria).

  • — Laporte, 1840: 176 (Staphylinus; characters; France).

  • — Hochhuth, 1849: 115 (Staphylinus; characters; Armenia).

  • — Redtenbacher, 1849: 695 (Staphylinus; characters; Austria).

  • — Küster, 1853: 51 (Staphylinus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 508 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 543 (Staphylinus; characters; Germany).

  • — Redtenbacher, 1857: 185 (Staphylinus; characters; Austria).

  • — Peyron, 1858: 422 (Staphylinus; Turkey).

  • — Fauvel, 1874: 179 [= 1874a: 405] (Staphylinus; characters; France; Austria; Italy; Sardinia; Greece; Turkey; Armenia).

  • — Redtenbacher, 1874: 204 (Staphylinus; characters; Austria).

  • — Mulsant and Rey, 1876: 260 [= 1877: 116] (Platydracus; subgenus Bemasus; characters; France).

  • — Ganglbauer, 1895: 428 (Staphylinus; synonym of flavopunctatus).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; synonym of flavopunctatus).

  • — Portevin, 1929: 349 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; synonym of flavopunctatus).

  • — Smetana, 1958: 269 (Staphylinus; synonym of flavopunctatus).

  • — Coiffait, 1974: 395 (Platydracus; synonym of flavopunctatus).

florentinus Coiffait, 1974 [Note: Nomen nudum], see: fulvipes Scopoli, 1763.

fluviaticus Casey, 1915, see: mysticus Erichson, 1840.

formosae Bernhauer, 1933a: 44 (Staphylinus; Type locality: Formosa).

  • — Shibata, 1994a: 106 (Platydracus; characters; Taiwan).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Taiwan.

fossator Gravenhorst, 1802: 164 (Staphylinus; Type locality: America septentr.).

  • — Gravenhorst, 1806: 117 (Staphylinus).

  • — Nordmann, 1837: 52 (Staphylinus; characters; America septentrionalis).

  • — Erichson, 1839b: 386 (Staphylinus; characters; America septentrionalis).

  • — Provancher, 1877: 729 (Staphylinus; characters; Quebec).

  • — Horn, 1879: 194 (Staphylinus; characters; Michigan to Georgia).

  • — Fall and Cockerell, 1907: 166 (Staphylinus; New Mexico).

  • — Blatchley, 1910: 377 (Staphylinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; catalog).

  • — Hatch, 1957: 178 (Ocypus; subgenus Platydracus; characters; British Columbia; Washington).

  • — Dearborn and Donahue, 1994: 15 (Platydracus; Maine).

  • — Downie and Arnett, 1996: 386 (Staphylinus; characters; USA).

  • — Lundgren, 1998: 52 (Platydracus; Florida).

  • — Schmidt, 1999: 108 (Platydracus; food for adults and larvae used for rearing species; rearing methods).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Canada, USA.

fulvipes Scopoli, 1763: 99 (Staphylinus; Type locality: Carniolia).

  • — Goeze, 1777: 725 (Staphylinus; characters).

  • — Gmelin, 1790: 2032 (Staphylinus; [Note: Page misnumbered as 1032]; characters).

  • — Turton, 1802: 516 (Staphylinus; cited as fulvus; characters; Carniola).

  • — Erichson, 1839a: 438 (Staphylinus; characters; Germany).

  • — Heer, 1839: 253 (Staphylinus; characters; Switzerland).

  • — Erichson, 1839b: 382 (Staphylinus; characters; Europe; Siberia).

  • — Redtenbacher, 1849: 694 (Staphylinus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 509 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 545 (Staphylinus; characters; Germany).

  • — Redtenbacher, 1857: 184 (Staphylinus; characters; Austria).

  • — Thomson, 1860: 144 (Platydracus; characters; Scandinavia).

  • — Hochhuth, 1872: 135 (Staphylinus; notes; Ukraine).

  • — Fauvel, 1874: 177 [= 1874a: 403] (Staphylinus; characters; France; Europe; Siberia; Altai; Amur region).

  • — Redtenbacher, 1874: 204 (Staphylinus; characters; Austria).

  • — Seidlitz, 1875: 269 (Leistotrophus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 271 [= 1877: 127] (Platydracus; characters; France).

  • — J. Sahlberg, 1876: 9 (Platydracus; Finland).

  • — Fowler, 1888: 250 (Staphylinus; characters; Britain).

  • — Ganglbauer, 1895: 426 (Staphylinus; subgenus Platydracus; characters; Europe; Siberia).

  • — Reitter, 1909: 118 (Staphylinus; subgenus Platydracus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; catalog).

  • — Johansen, 1914: 378 (Staphylinus; subgenus Platydracus; characters; Denmark).

  • — Portevin, 1929: 349 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; catalog).

  • — V. Hansen, 1952: 98 (Staphylinus; characters; Denmark).

  • — Coiffait, 1956e: 192, 193 (Platydracus; characters).

  • — Smetana, 1958: 273 (Staphylinus; subgenus Platydracus; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 109 (Staphylinus; subgenus Platydracus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 194 (Platydracus; characters; central Europe).

  • — Smetana, 1965c: 27 (Staphylinus; subgenus Platydracus; Anatolia).

  • — Horion, 1965: 200 (Staphylinus; subgenus Platydracus; Russia; Finland; Sweden; Norway; Denmark; Britain; all central Europe; Spain; Portugal; Italy; Slovenia; Dalmatia; Bosnia; Albania; Greece; Macedonia; Serbia; Ukraine).

  • — Pototskaia, 1967: 67 (Staphylinus; subgenus Platydracus; larval characters).

  • — Smetana, 1967c: 197 (Staphylinus; subgenus Platydracus; Spain).

  • — Smetana, 1968a: 156 (Staphylinus; subgenus Platydracus; Anatolia).

  • — Szujecki, 1968a: 732 (Staphylinus; Poland).

  • — Coiffait, 1974: 392 (Platydracus; characters; Europe; Britain; Scandinavia; central Asia).

  • — Shilov, 1975a: 376 [= 1975b: 99] (Staphylinus; Komi region of Russia).

  • — Pope, 1977: 30 (Platydracus; Britain).

  • — Petrenko, 1980a: 59, 62 (Staphylinus; subgenus Platydracus; characters; Ukraine).

  • — Tóth, 1984: 14 (Platydracus; characters; Hungary).

  • — Lucht, 1987: 107 (Platydracus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Pilon, 1991: 102 (Platydracus; notes; Italy).

  • — Pilon and Zanetti, 1991: 66 (Platydracus; Italy).

  • — Smetana, 1993b: 49 (Platydracus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 30 (Platydracus; Italy).

  • — Pilon, 1998: 70 (Platydracus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Europe, Russia, Turkey, “Altai region” ?.

bicinctus Rossi, 1792: 96 (Staphylinus; Type locality: Etruria).

  • — Block, 1799: 115 (Staphylinus; characters).

  • — Erichson, 1839a: 438 (Staphylinus; synonym of fulvipes).

  • — Heer, 1839: 253 (Staphylinus; synonym of fulvipes).

  • — Erichson, 1839b: 382 (Staphylinus; synonym of fulvipes).

  • — Fairmaire and Laboulbène, 1856: 509 (Staphylinus; synonym of fulvipes).

  • — Ganglbauer, 1895: 426 (Staphylinus; synonym of fulvipes).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; synonym of fulvipes).

  • — Smetana, 1958: 273 (Staphylinus; synonym of fulvipes).

  • — Coiffait, 1974: 392 (Platydracus; synonym of fulvipes).

erythropus Paykull, 1800: 372 (Staphylinus; [preoccupied]; [Note: Bernhauer and Schubert, 1914, cited this name as a synonym of both Staphylinus brunnipes (p. 385) and Staphylinus fulvipes (p. 379)]; Type locality: Kinnakulle).

  • — Gravenhorst, 1806: 110 (Staphylinus; synonym of tricolor Gravenhorst).

  • — Germar, 1823: 14 (Staphylinus; characters; Sweden; Germany).

  • — Erichson, 1839a: 438 (Staphylinus; synonym of fulvipes).

  • — Heer, 1839: 253 (Staphylinus; synonym of fulvipes).

  • — Erichson, 1839b: 382 (Staphylinus; synonym of fulvipes).

  • — Thomson, 1860: 144 (Platydracus; synonym of fulvipes).

  • — Ganglbauer, 1895: 426 (Staphylinus; synonym of fulvipes).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; synonym of fulvipes).

  • — Smetana, 1958: 273 (Staphylinus; synonym of fulvipes).

  • — Coiffait, 1974: 392 (Platydracus; synonym of fulvipes).

tricolor Gravenhorst, 1802: 7 (Staphylinus; [preoccupied]; Type locality: Not cited, but presumably Brunsvigae).

  • — Latreille, 1804: 305 (Staphylinus; characters).

  • — Gravenhorst, 1806: 110 (Staphylinus; characters).

  • — Erichson, 1839a: 438 (Staphylinus; synonym of fulvipes).

  • — Heer, 1839: 253 (Staphylinus; synonym of fulvipes).

  • — Erichson, 1839b: 382 (Staphylinus; synonym of fulvipes).

  • — Ganglbauer, 1895: 426 (Staphylinus; synonym of fulvipes).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; synonym of fulvipes).

  • — Smetana, 1958: 273 (Staphylinus; synonym of fulvipes).

  • — Coiffait, 1974: 392 (Platydracus; synonym of fulvipes).

azureus Gebler, 1829: 70 (Staphylinus; Type locality: Barnaul).

  • — Erichson, 1839a: 438 (Staphylinus; synonym of fulvipes).

  • — Erichson, 1839b: 382 (Staphylinus; synonym of fulvipes).

  • — Redtenbacher, 1849: 694 (Staphylinus; synonym of fulvipes).

  • — Kraatz, 1857d: 545 (Staphylinus; synonym of fulvipes).

  • — Ganglbauer, 1895: 426 (Staphylinus; synonym of fulvipes).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; synonym of fulvipes).

  • — Smetana, 1958: 273 (Staphylinus; synonym of fulvipes).

  • — Coiffait, 1974: 392 (Platydracus; synonym of fulvipes).

cingulus Comolli, 1837: 14 (Staphylinus; Type locality: Monte Legnone).

  • — Fauvel, 1867: LIV (Staphylinus; synonym of fulvipes).

  • — Ganglbauer, 1895: 426 (Staphylinus; synonym of fulvipes).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; synonym of fulvipes).

  • — Smetana, 1958: 273 (Staphylinus; synonym of fulvipes).

  • — Coiffait, 1974: 392 (Platydracus; synonym of fulvipes).

aureovillosus G. Müller, 1932: 76 (Staphylinus; subgenus Platydracus; aberration of fulvipes; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Zakopane in Polonia).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; aberration of fulvipes).

  • — Smetana, 1958: 273 (Staphylinus; aberration of fulvipes).

  • — Coiffait, 1974: 392 (Platydracus; variety of fulvipes; characters).

florentinus Coiffait, 1974: 392 (Platydracus; variety of fulvipes; [Note: Nomen nudum (Article 15.2)]; cited from Italie: Florence).

fulvomaculatus Nordmann, 1837: 39 (Staphylinus; Type locality: Mexico ?).

  • — Erichson, 1839b: 384 (Staphylinus; characters; Mexico).

  • — Solsky, 1868a: 127 (Staphylinus; Mexico).

  • — Sharp, 1884: 373 (Staphylinus; notes; Mexico; Guatemala).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; catalog).

  • — Smyth, 1934: 107 (Staphylinus; natural history notes; Mexico).

  • — Marquez-Luna and Navarrete Heredia, 1995: 35 (Platydracus; in detritus of Atta mexicana; Mexico).

  • Distribution: Mexico, Guatemala.

fuscofemoratus G. Müller, 1923, see: stercorarius Olivier, 1795.

fuscolineatus Bernhauer, 1934: 8 (Staphylinus; subgenus Platydracus; Type locality: Szechuan: Nitou Tatsienlu).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: China.

fusiformis Casey, 1915, see: centralis Sharp, 1884.

gabiruensis Bernhauer, 1934g: 240 (Staphylinus, cited as Staphilinus; subgenus Platydracus; Type locality: Ruanda: Gabiru).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Rwanda.

gemmatus Fauvel, 1895: 251 (Staphylinus; Type locality: Bírmanie, Carin Cheba, 900–1100 m).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; catalog).

  • — Cameron, 1932: 188 (Staphylinus; subgenus Platydracus; characters; Burma).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Myanmar.

girardi Levasseur, 1980: 369 (Platydracus; subgenus Paraplatydracus; Type locality: Est de la Côte-d-Ivoire: Sipilou).

  • Distribution: Ivory Coast.

goryi Laporte, 1835: 113 (Staphylinus; Type locality: Indes orientales: Décan).

  • — Erichson, 1839b: 395 (Staphylinus; characters; Decan).

  • — Kraatz, 1859: 72 (Staphylinus; characters; India).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; catalog).

  • — Cameron, 1932: 186 (Staphylinus; subgenus Platydracus; characters; India).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: India.

auripennis Kraatz, 1859: 72 (Staphylinus; Type locality: India borealis).

  • — Gemminger and Harold, 1868: 579 (Staphylinus; synonym of goryi).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; synonym of goryi).

  • — Cameron, 1932: 186 (Staphylinus; synonym of goryi).

  • — Smetana and Davies, 2000: 39 (Platydracus; synonym of goryi).

guineensis Cameron, 1950e: 431 (Staphylinus; subgenus Platydracus; Type locality: Cameroon: Sakdje).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Cameroon.

hauserianus Bernhauer, 1933: 31 (Staphylinus; subgenus Platydracus; Type locality: Kiang-Si: Chang-Tsin-cheng).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: China.

hewitti Bernhauer, 1914c: 100 (Staphylinus; Type locality: Sarawak; Kuching).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; catalog).

  • — Hammond, 1984: 195 (Platydracus; Borneo).

  • Distribution: Borneo.

hypocrita G. Müller, 1925: 41 (Staphylinus; subgenus Platydracus; Type locality: Asia minor, Smyrna).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • — G. Müller, 1950: 8 (Staphylinus; subgenus Platydracus; notes).

  • — Coiffait, 1974: 393 (Platydracus; characters; Turkey; Caucasus).

  • — Smetana and Davies, 2000: 39 (Platydracus; subgenus Platydracus).

  • Distribution: Turkey, Caucasus.

immaculatus Mannerheim, 1830: 22 (Staphylinus; [Note: This name is older than vulpinus]; Type locality: America).

  • — Mannerheim, 1831: 436 (Staphylinus; America).

  • — Nordmann, 1837: 62 (Staphylinus; America).

  • — Erichson, 1839b: 379 (Staphylinus; synonym of vulpinus).

  • — Horn, 1879: 191 (Staphylinus; synonym of vulpinus).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; synonym of vulpinus).

  • — Smetana and Davies, 2000: 25, 39 (Staphylinus; cited as a nomen dubium and listed separate from vulpinus; although immaculatus is older than vulpinus and has been listed by most authors as a junior synonym of it, the former should not replace the latter because the status of the species is in doubt).

  • — Herman, 2001: 28 (Platydracus; valid name for species).

  • Distribution: Canada†, USA.

vulpinus Nordmann, 1837: 53 (Staphylinus; Type locality: Philadelphia in Amer. bor.).

  • — Erichson, 1839b: 379 (Staphylinus; characters; United States).

  • — Horn, 1879: 191 (Staphylinus; characters; middle States).

  • — Blatchley, 1910: 376 (Staphylinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1389 (Staphylinus; catalog).

  • — Voris, 1939a: 288, 295 (Staphylinus; larval characters).

  • — Schwert, Anderson, Morgan, Morgan, and Karrow, 1985: 216 (Platydracus; Quaternary fossil; Ontario).

  • — Dearborn and Donahue, 1994: 15 (Platydracus; Maine).

  • — Downie and Arnett, 1996: 385 (Staphylinus; characters; USA).

  • — Smetana and Davies, 2000: 25, 41 (Platydracus; subgenus Platydracus).

  • — Herman, 2001: 28 (Platydracus; synonym of immaculatus).

imperatorius Bernhauer, 1916e: 31 (Staphylinus; subgenus Platydracus; Type locality: Kiautschau, Kiangsi).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: China.

impotens Eppelsheim, 1889a: 170 (Staphylinus; subgenus Platydracus; Type locality: Kan-ssu).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • — Bernhauer, 1936: 5 (Staphylinus; subgenus Platydracus; Mongolia).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: China, Mongolia.

indicus Kraatz, 1859: 75 (Staphylinus; Type locality: Ceylan; India borealis).

  • — Fauvel, 1904a: 57 (Staphylinus; Nilghiris; Monts Anamalai; Ceylan; Himalaya; Malacca).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • — Cameron, 1932: 188 (Staphylinus; subgenus Platydracus; characters; Ceylon; India).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • — Scheerpeltz, 1954: 114 (Staphylinus; subgenus Platydracus; Ceylon).

  • — Scheerpeltz, 1961f: 136 (Staphylinus; subgenus Platydracus; Ceylon).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Sri Lanka, India, Malaysia.

lineatus Walker, 1859: 51 (Ocypus; Type locality: Ceylon).

  • — Fauvel, 1904a: 57 (Staphylinus; synonym of indicus).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; synonym of indicus).

  • — Cameron, 1932: 188 (Staphylinus; synonym of indicus).

  • — Smetana and Davies, 2000: 40 (Platydracus; synonym of indicus).

initicornis Scheerpeltz, 1933, see: purpureoaureus Bernhauer, 1915.

inornatus Sharp, 1874: 31 (Staphylinus; Type locality: Simabara).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • — Nakane, 1963a: 92 (Platydracus; characters; habitus photograph; Japan).

  • — Yuh, Paik, Kwon, and Lee, 1985: 244 (Staphylinus; Korea).

  • — J. Li, 1993: 38 (Staphylinus, cited as Staphylius; China).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Japan, Korea, China.

insolitus Sharp, 1884: 389 (Staphylinus; Type locality: Mexico, Panistlahuaca).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • — Bernhauer, 1917: 93 (Staphylinus; subgenus Plesiolinus).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Mexico.

insularis Cameron, 1941c: 382 (Staphylinus; subgenus Platydracus; Type locality: Philippines: Luzon: Laguna).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Philippines.

javanus Bernhauer, 1934j: 173 (Staphylinus; subgenus Platydracus; Type locality: Ost-Java, Nongkodjadjar, 4000 f.).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Indonesia.

jeanneli Chapman, 1939: 72 (Staphylinus; subgenus Platydracus; Type locality: Kenya colony: Camp III de l'Elgon, alt. 3500 m).

  • — Paulian, 1941: 239 (Staphylinus; subgenus Platydracus; larval characters).

  • — Jeannel and Paulian, 1945: 91 (Staphylinus; subgenus Platydracus; adult and larval characters; Kenya).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Kenya.

kasyi Scheerpeltz, 1962a: 19 (Staphylinus; subgenus Platydracus; Type locality: Syrien: Berge etwa 25 km westlich von Damaskus).

  • — Coiffait, 1974: 394 (Platydracus; [Note: The type locality cited by Coiffait is erroneous]; characters; Lebanon).

  • — M. Dvořák, 1984: 60 (Platydracus; notes; Syria).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Syria.

kiulungensis Bernhauer, 1933a: 46 (Staphylinus; subgenus Platydracus; Type locality: Tatsienlu-Kiulung).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: China.

kiushiuensis Bernhauer, 1939e: 155 (Staphylinus; subgenus Platydracus; Type locality: Japan: Beppu, N. O. Kiushiu).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Japan.

lamottei Levasseur, 1967b: 1637 (Platydracus; Type locality: Côte d'Ivoire, Lamto, Toumodi).

  • Distribution: Ivory Coast.

lamtoensis Levasseur, 1967b: 1640 (Platydracus; Type locality: Côte d'Ivoire, Lamto, Toumodi).

  • Distribution: Ivory Coast.

latebricola Gravenhorst, 1806: 113 (Staphylinus; Type locality: Borussia; Brunsvigae).

  • — Nordmann, 1837: 56 (Staphylinus; characters; Europe).

  • — Erichson, 1839a: 437 (Staphylinus; characters; Germany).

  • — Heer, 1839: 252 (Staphylinus; characters; Switzerland).

  • — Erichson, 1839b: 382 (Staphylinus; characters; Germany).

  • — Märkel, 1845: 236 (Staphylinus; with Formica rufa).

  • — Jacobsen, 1849: 683 (Staphylinus; characters; Denmark).

  • — Redtenbacher, 1849: 695 (Staphylinus; characters; Austria).

  • — Hardy, 1851: 23 (Staphylinus; notes; England).

  • — Fairmaire and Laboulbène, 1856: 508 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 545 (Staphylinus; characters; Germany).

  • — Redtenbacher, 1857: 185 (Staphylinus; characters; Austria).

  • — Thomson, 1860: 143 (Platydracus; characters; Scandinavia).

  • — Fauvel, 1874: 180 [= 1874a: 406] (Staphylinus; characters; France; Europe).

  • — Redtenbacher, 1874: 204 (Staphylinus; characters; Austria).

  • — Seidlitz, 1875: 269 (Leistotrophus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 269 [= 1877: 125] (Platydracus; characters; France).

  • — J. Sahlberg, 1876: 9 (Platydracus; Finland).

  • — J. Sahlberg, 1880: 68 (Platydracus; Siberia).

  • — Fowler, 1888: 251 (Staphylinus; characters; Britain).

  • — Ganglbauer, 1895: 428 (Staphylinus; subgenus Platydracus; characters; Europe).

  • — Reitter, 1909: 118 (Staphylinus; subgenus Platydracus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • — Johansen, 1914: 379 (Staphylinus; subgenus Platydracus; characters; Denmark).

  • — Portevin, 1929: 349 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • — V. Hansen, 1952: 99 (Staphylinus; characters; Denmark).

  • — Coiffait, 1956e: 192, 194 (Platydracus; characters).

  • — Smetana, 1958: 271 (Staphylinus; subgenus Platydracus; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 110 (Staphylinus; subgenus Platydracus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 194 (Platydracus; characters; central Europe).

  • — Horion, 1965: 201 (Staphylinus; subgenus Platydracus; Russia; Finland; Sweden; Norway; Denmark; Britain; Holland; Belgium; France; Switzerland; Italy; Dalmatia; Bosnia; Serbia; Romania; Czechoslovakia; Poland; Austria; Germany).

  • — Coiffait, 1974: 397 (Platydracus; characters; Europe; Britain; Scandinavia; Mediterranean region).

  • — Pope, 1977: 30 (Platydracus; Britain).

  • — Petrenko, 1980a: 60, 62 (Staphylinus; subgenus Platydracus; characters; Ukraine).

  • — Tóth, 1984: 14 (Platydracus; characters; Hungary).

  • — Lucht, 1987: 107 (Platydracus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Pilon and Zanetti, 1991: 67 (Platydracus; Italy).

  • — Smetana, 1993b: 49 (Platydracus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 30 (Platydracus; Italy).

  • — Gerend and Braunert, 1997: 199 (Platydracus; notes; Luxembourg).

  • — Pilon, 1998: 70 (Platydracus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Europe, Russia.

aericeps Stephens, 1832: 206 (Staphylinus; Type locality: London; Suffolk).

  • — Curtis, 1839: pl. 758 (Staphylinus; England).

  • — Stephens, 1839: 387 (Staphylinus; characters; England).

  • — Holme, 1842: 116 (Staphylinus; notes).

  • — Hardy, 1851: 23 (Staphylinus; synonym of latebricola).

  • — G. Waterhouse, 1858: 23 (Staphylinus; synonym of latebricola).

  • — Ganglbauer, 1895: 429 (Staphylinus; synonym of latebricola).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; synonym of latebricola).

  • — Smetana, 1958: 271 (Staphylinus; synonym of latebricola).

  • — Coiffait, 1974: 397 (Platydracus; synonym of latebricola).

rupicola Kiesenwetter, 1850: 219 (Staphylinus; Type locality: Pyrenaei orientales).

  • — Kiesenwetter, 1851a: 413 (Staphylinus; characters; valleé de la Preste).

  • — Fairmaire and Laboulbène, 1856: 508 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 545 (Staphylinus; synonym of latebricola).

  • — Fowler, 1888: 251 (Staphylinus; synonym of latebricola).

  • — Ganglbauer, 1895: 429 (Staphylinus; synonym of latebricola).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; synonym of latebricola).

  • — Smetana, 1958: 271 (Staphylinus; synonym of latebricola).

  • — Coiffait, 1974: 397 (Platydracus; synonym of latebricola).

latecarinatus Bernhauer, 1937: 303 (Staphylinus; subgenus Platydracus; Type locality: Uganda: Entebbe).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Uganda.

lecordieri Levasseur, 1980: 367 (Platydracus; subgenus Tropoplatydracus; Type locality: Côte-d-Ivoire: Lamto).

  • Distribution: Ivory Coast.

lefevrei Bernhauer, 1936a: 23 (Staphylinus; subgenus Platydracus; Type locality: Urundi: Kitega. Rhodesia: Solweri Dist., Lusala).

  • — Cameron, 1950g: 43 (Staphylinus; subgenus Platydracus; Belgian Congo).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Burundi, Congo, Zambia.

leleupi Fagel, 1957c: 265 (Platydracus; subgenus Abemus; Type locality: Congo Belge: Kibali-Ituri: Blukwa, 1820–2100 m).

  • Distribution: Congo.

leroyi Bernhauer, 1938, see: procerus Gahan, 1893.

lewisi Cameron, 1932: 191 (Staphylinus; subgenus Platydracus; Type locality: Ceylon: Dikoya, alt. 3800–4200 feet).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Sri Lanka.

lineatus Walker, 1859, see: indicus Kraatz, 1859.

lomii Cerruti, 1951: 17 (Staphylinus; subgenus Platydracus; Type locality: India: Punjab: Yol, 1300 m).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: India.

lucanus Horn, 1894, see: tarsalis Mannerheim, 1843.

lutarius Gravenhorst, 1806, see: flavopunctatus Latreille, 1804.

macgregori Cooper, 1933, see: phoenicurus Nordmann, 1837.

maculipennis Kraatz, 1859: 77 (Staphylinus; Type locality: India borealis).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • — Cameron, 1932: 196 (Staphylinus; subgenus Platydracus; characters; India).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: India.

maculosus Gravenhorst, 1802: 165 (Staphylinus; Type locality: Baltimore).

  • — Gravenhorst, 1806: 123 (Staphylinus; characters).

  • — Nordmann, 1837: 31 (Staphylinus; Carolina).

  • — Erichson, 1839b: 375 (Staphylinus; characters; America septentrionalis).

  • — Provancher, 1877: 246 (Staphylinus; characters; Quebec).

  • — Horn, 1879: 191 (Staphylinus; characters; entire Atlantic region).

  • — Sharp, 1884: 377 (Staphylinus; characters; Mexico).

  • — Blatchley, 1910: 376 (Staphylinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • — Notman, 1920a: 180 (Staphylinus; New York).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • — Voris, 1939a: 295 (Staphylinus; egg, larval and pupal characters).

  • — Hunter, Kincher, Bay, Beerwinkle, 1991: 169 (Platydracus; seasonal distribution and diel flight activity; Texas).

  • — Dearborn and Donahue, 1994: 15 (Platydracus; Maine).

  • — Downie and Arnett, 1996: 385 (Staphylinus; characters; USA).

  • — Lundgren, 1998: 52 (Platydracus; Florida).

  • — Smetana and Davies, 2000: 41 (Platydracus; synonym of viduatus).

  • — Newton, in press (Platydracus; application to Commission to maintain use as vald name of species).

  • Distribution: Canada, USA, Mexico.

viduatus Fabricius, 1801: 591 (Staphylinus; [Note: Say in 1830 and 1834 noted that viduatus had priority over maculosus but most subsequent authors cited viduatus as the junior synonym]; Type locality: Carolina).

  • — Gravenhorst, 1806: 123 (Staphylinus; synonym of maculosus).

  • — Say, 1830: 38 (Staphylinus; viduatus has priority over maculosus; characters).

  • — Say, 1834: 452 (Staphylinus; viduatus has priority over maculosus).

  • — Erichson, 1839b: 375 (Staphylinus; synonym of maculosus).

  • — Horn, 1879: 191 (Staphylinus; synonym of maculosus).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; synonym of maculosus).

  • — Smetana and Davies, 2000: 41 (Platydracus; subgenus Platydracus).

  • — Newton, in press (Platydracus; application to Commission to list name as junior synonym of maculosus).

erythropennis Mannerheim, 1830: 22 [= 1831: 436] (Staphylinus; Type locality: America borealis).

  • — Mannerheim, 1831: 436 (Staphylinus; characters; America borealis).

  • — Nordmann, 1837: 62 (Staphylinus; America borealis).

  • — Erichson, 1839b: 375 (Staphylinus; synonym of maculosus).

  • — Horn, 1879: 191 (Staphylinus; synonym of maculosus).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; synonym of maculosus).

marmorellus Fauvel, 1895: 253 (Staphylinus; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Carin Asciuii Ghécu, 1400–1500 m; Sumatra, Ajer Mantcior; Java orient. et occid., monts Tengger et Gede, 1300 m).

  • — Fauvel, 1905e: 83 (Staphylinus; Java; Sumatra; Burma; Mt. Khasia; Tonkin).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • — Cameron, 1932: 193 (Staphylinus; subgenus Platydracus; characters; Burma).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • — Cameron, 1933d: 391 (Staphylinus; Sumatra).

  • — Scheerpeltz, 1935a: 634 (Staphylinus; Indonesia).

  • — Cameron, 1937: 21 (Staphylinus; Java).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Myanmar, Vietnam, Indonesia.

mendicus Sharp, 1884: 387 (Staphylinus; Type locality: Guatemala, Zapote; Costa Rica).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • — Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 59 (Platydracus; necrophilous species; distribution in Mexico).

  • Distribution: Guatemala, Costa Rica.

meridionalis Rosenhauer, 1847: 12 (Staphylinus; Type locality: südliches Frankreich).

  • — Fairmaire and Laboulbène, 1856: 508 (Staphylinus; characters; France).

  • — Fauvel, 1874: 178 [= 1874a: 404] (Staphylinus; characters; France; Syria; Armenia).

  • — Mulsant and Rey, 1876: 262 [= 1877: 118] (Platydracus; subgenus Bemasus; characters; France).

  • — Ganglbauer, 1895: 427 (Staphylinus; subgenus Platydracus; characters; French Alps; Pyrenees; Syria; Armenia).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • — Portevin, 1929: 349 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • — Coiffait, 1956e: 192, 193 (Platydracus; characters).

  • — Coiffait, 1974: 393 (Platydracus; characters; western Mediterranean region, eastern Mediterranean sites need confirmation).

  • — Petrenko, 1980: 82 (Staphylinus; Caucasus).

  • — Pilon, 1998: 98 (Platydracus; notes; doubtfully in Italy).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: France, Syria ?, Armenia ?.

mulsanti Godard, 1850: LV (Staphylinus; Type locality: montagnes des Corbières).

  • — Fairmaire and Laboulbène, 1856: 508 (Staphylinus; synonym of meridionalis).

  • — Ganglbauer, 1895: 427 (Staphylinus; synonym of meridionalis).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; synonym of meridionalis).

  • — Coiffait, 1974: 393 (Platydracus; synonym of meridionalis).

armeniacus Sharp, 1873b: 412 (Staphylinus; Type locality: Not cited).

  • — Fauvel, 1874: 178 [= 1874a: 404] (Staphylinus; synonym of meridionalis).

  • — Ganglbauer, 1895: 427 (Staphylinus; synonym of meridionalis).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; synonym of meridionalis).

  • — Coiffait, 1974: 393 (Platydracus; synonym of meridionalis).

mimeticus Bernhauer, 1917i: 108 (Staphylinus; subgenus Platydracus; Type locality: Bolivien: Rio Longo, 750 m; Yuracaris).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Bolivia.

modestus Sharp, 1884, see: centralis Sharp, 1884.

modestus Fall, 1907, see: sepulchralis Erichson, 1839.

mongendensis Bernhauer, 1929f: 122 (Staphylinus; subgenus Platydracus; Type locality: Mongende).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Congo.

montanus Coiffait, 1977b: 228 (Platydracus; Type locality: Bhutan: Nobding, 41 km Ouest de Wangdi Phodrang, 2800 m).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Bhutan.

mortuorum Bernhauer, 1912c: 22 (Staphylinus; subgenus Platydracus; Type locality: Ostsibirien: Suczan).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Russia.

muellerianus Scheerpeltz, 1933: 1387 (Staphylinus; replacement name for banghaasi G. Müller).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Korea.

banghaasi G. Müller, 1932: 75 (Staphylinus; subgenus Platydracus; [preoccupied]; Type locality: Korea: Pu Ryong).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; synonym of muellerianus).

  • — Smetana and Davies, 2000: 40 (Platydracus; synonym of muellerianus).

mulsanti Godart, 1850, see: meridionalis Rosenhauer, 1847.

musonoiensis Levasseur, 1967b: 1634 (Platydracus; Type locality: région de Kolwezi, Katanga, Musonoié).

  • Distribution: Congo.

mysticus Erichson, 1840: 926 (Staphylinus; Type locality: America septentrionalis).

  • — Horn, 1879: 192 (Staphylinus; characters; New York to Missouri and Georgia).

  • — Fall and Cockerell, 1907: 166 (Staphylinus; New Mexico).

  • — Blatchley, 1910: 377 (Staphylinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • — Downie and Arnett, 1996: 386 (Staphylinus; characters; USA).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: USA.

fluviaticus Casey, 1915: 428 (Staphylinus; Type locality: Missouri: St. Louis).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 40 (Platydracus; synonym of mysticus).

neomexicanus Bernhauer and Schubert, 1914, see: sepulchralis Erichson, 1839.

nepalensis Scheerpeltz, 1976: 36 (Staphylinus; subgenus Platydracus; Type locality: Nepal: Changma, 2200 m).

  • — Scheerpeltz, 1976a: 90 (Staphylinus; Nepal).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Nepal.

nigripennis Cameron, 1941c: 381 (Staphylinus; subgenus Platydracus; Type locality: Philippines: Luzon: Imugan).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Philippines.

nigriventris Boheman, 1848: 276 (Staphylinus; Type locality: Portus Natalensis).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • — Scheerpeltz, 1974: 167 (Staphylinus; subgenus Platydracus; South Africa).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: South Africa.

notativentris Fauvel, 1905a: 140 (Staphylinus; Type locality: Abyssinie: Hamacen, Azega, 2500 m; d'Asmara a Keren).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • — Rambousek, 1922: 1012 (Staphylinus; Ethiopia).

  • — Gridelli, 1939: 320 (Staphylinus; subgenus Platydracus; characters; Abyssinia).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Ethiopia, Eritrea.

ochropterus Germar, 1824, see: chalcocephalus Fabricius, 1801.

oculatus Bernhauer, 1929, see: oculosus Smetana and Davies, 2000.

oculosus Smetana and Davies, 2000: 25, 40 (Platydracus; subgenus Platydracus; replacement name for oculatus Bernhauer).

  • Distribution: China.

oculatus Bernhauer, 1929d: 110 (Staphylinus; subgenus Abemus; [preoccupied]; Type locality: Szentschwan).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 25, 40 (Platydracus; synonym of oculosus).

opaciceps Scheerpeltz, 1976a: 130 (Staphylinus; subgenus Platydracus; Type locality: Nepal: Jiri, 2000 m).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Nepal.

paganus Sharp, 1874, see: brevicornis Motschulsky, 1862.

pallidipes Bernhauer, 1917i: 107 (Staphylinus; subgenus Platydracus; Type locality: Bolivien: Rio Longo, 750 m).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Bolivia.

palmarum Jarrige, 1972, see: peyrierasi Jarrige, 1972.

panamensis Bernhauer and Schubert, 1914: 381 (Staphylinus; Type locality: Panama, Volcan de Chiriqui).

  • — Bernhauer, 1917: 93 (Staphylinus; subgenus Plesiolinus).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Panama.

tarsalis Sharp, 1884: 389 (Staphylinus; [preoccupied]; Type locality: Panama, Volcan de Chiriqui).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; synonym of panamensis).

  • — Bernhauer, 1917: 93 (Staphylinus; subgenus Plesiolinus; synonym of panamensis).

  • — Herman, 2001: 14 (Platydracus; synonym of panamensis).

parvulus Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus; replacement name for parvus Cameron).

  • Distribution: India.

parvus Cameron, 1932: 192 (Staphylinus; subgenus Platydracus; [preoccupied]; Type locality: Assam: Naga Hills, alt. 9000 feet).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 40 (Platydracus; synonym of parvulus).

  • Distribution: India.

parvus Cameron, 1932, see: parvulus Smetana and Davies, 2000.

patricius Bernhauer, 1915c: 71 (Staphylinus; subgenus Platydracus; Type locality: China: Shin-Guy-Foo).

  • — G. Müller, 1926a: 27 (Staphylinus; subgenus Chaetodracus).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; catalog).

  • — Blackwelder, 1952: 98 (type species of Chaetodracus, a subgenus of Platydracus).

  • — Smetana and Davies, 2000: 41 (Platydracus; subgenus Chaetodracus).

  • Distribution: China.

perniger Scheerpeltz, 1976: 38 (Staphylinus; subgenus Platydracus; Type locality: Nepal: Manga Deorali-Phaeda Khola, 2300 bis 1500 m [sic]).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Nepal.

perreaui Coiffait, 1984e: 378 (Platydracus ?; Type locality: Nepal oriental: Milke Himal, 2000–2300 m).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Nepal.

peyrierasi Jarrige, 1972a: 136 (Platydracus; Type locality: Madagascar Centre, tampoketsa d'Ambohitantely, 1600 m, 16 km aud Nord d'Ankazobe).

  • Distribution: Madagascar.

palmarum Jarrige, 1972a: 137 (Platydracus; subspecies of peyrierasi; Type locality: Madagascar Centre, route d'Andriamena, Andranobe, 1250 m).

phoenicurus Nordmann, 1837: 57 (Staphylinus; Type locality: Mexico).

  • — Erichson, 1839b: 380 (Staphylinus; characters; Mexico).

  • — Solsky, 1868a: 127 (Staphylinus; cited as foenicurus; Mexico).

  • — Sharp, 1884: 377 (Staphylinus; Mexico).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Mexico, USA.

macgregori Cooper, 1933: 264 (Staphylinus; Type locality: Chiricahuae Mts. Arizona).

  • — Smetana and Davies, 2000: 40 (Platydracus; synonym of phoenicurus).

philippinus Cameron, 1941c: 381 (Staphylinus; subgenus Platydracus; Type locality: Philippines: Mindanao: Pt. Rango).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Philippines.

pinorum Casey, 1915: 427 (Staphylinus; Type locality: North Carolina: Southern Pines).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; catalog).

  • — Lundgren, 1998: 52 (Platydracus; Florida).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: USA.

plebejus Bernhauer, 1915c: 73 (Staphylinus; subgenus Platydracus; Type locality: Insel Quelpaerd im Süden von Korea).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; catalog).

  • — Ito, 1993b: 150 (Platydracus; characters; Korea; Japan).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Korea, Japan.

polyphemus Bernhauer, 1913, see: sjostedti Fauvel, 1903.

praelongus Mannerheim, 1830: 24 (Staphylinus; Type locality: America borealis).

  • — Mannerheim, 1831: 438 (Staphylinus; America borealis).

  • — Nordmann, 1837: 73 (Staphylinus; America septentrionalis).

  • — Erichson, 1839b: 391 (Staphylinus; characters; America septentrionalis).

  • — Horn, 1879: 197 (Staphylinus; characters; Massachusetts to Pennsylvania).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; catalog).

  • — Dearborn and Donahue, 1994: 15 (Platydracus; Maine).

  • — Downie and Arnett, 1996: 386 (Staphylinus; characters; USA).

  • — Lundgren, 1998: 52 (Platydracus; Florida).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: USA.

cicatricosus LeConte, 1863a: 37 (Staphylinus; Type locality: Middle and southern States).

  • — Horn, 1879: 197 (Staphylinus; synonym of praelongus).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; synonym of praelongus).

prasinivariegatus Bernhauer, 1921b: 75 (Staphylinus; subgenus Platydracus; Type locality: Kolumbien).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Colombia.

pratti Scheerpeltz, 1962: 593 (Staphylinus; subgenus Platydracus; Type locality: Kiukiang Fluss, Szechuan, S.W. China).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: China.

preangeranus Bernhauer, 1937: 22 (Staphylinus; subgenus Platydracus; Type locality: Java: Bandoeng Dago; Blawan, Idjen Plateau, alt. 900–1500 m).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Indonesia.

procerus Gahan, 1893: 739 (Staphylinus; Type locality: Zomba; Karague).

  • — Bernhauer, 1908c: 108 (Staphylinus; German East Africa: Nyassaland).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • — Bernhauer, 1947: 158 (Staphylinus; Deutsch-Ostafrika).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Malawi, Congo, Tanzania.

leroyi Bernhauer, 1938c: 315 (Staphylinus; subgenus Platydracus; subspecies of procerus; Type locality: Belgian Congo: Bunia).

  • — Herman, 2001: 14 (Platydracus; subspecies of procerus).

pseudopaganus Bernhauer, 1914b: 65 (Staphylinus; Type locality: Mandschurei: Chi-Kuan-Shan).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: China.

pseudopatricius G. Müller, 1926a: 28 (Staphylinus; subgenus Platydracus; subspecies of pseudopaganus; Type locality: China: Shangai).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; subspecies of pseudopaganus).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus; subspecies of pseudopaganus).

pseudopatricius G. Müller, 1926, see: pseudopaganus Bernhauer, 1914.

purpurascens Cameron, 1920a: 217 (Staphylinus; [preoccupied]; Type locality: South Coorg, Pollibetk).

  • — Cameron, 1932: 195 (Staphylinus; subgenus Platydracus; characters; India).

  • — Scheerpeltz, 1933: 1388 (Staphylinus; catalog).

  • — Herman, 2001: 14, 48 (Platydracus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: India.

purpureoaureus Bernhauer, 1915k: 305 (Staphylinus; subgenus Platydracus; Type locality: Deutsch-Ostafrika: Usagara).

  • — Scheerpeltz, 1933: 1388 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Congo, Tanzania.

basicornis Bernhauer, 1932b: 155 (Staphylinus; variety of purpureoaureus, cited as purpureoauratus; [preoccupied]; Type locality: Kafakumba).

  • — Scheerpeltz, 1933: 1388 (Staphylinus; synonym of purpureoaureus).

  • — Herman, 2001: 14 (Platydracus; synonym of purpureoaureus).

initicornis Scheerpeltz, 1933: 1388 (Staphylinus; variety of purpureoaureus; replacement name for basicornis Bernhauer).

  • — Herman, 2001: 14 (Platydracus; synonym of purpureoaureus).

quadraticeps Casey, 1924, see: zonatus Gravenhorst, 1802.

reitterianus Bernhauer, 1933a: 44 (Staphylinus; subgenus Platydracus; Type locality: Tatsienlu-Kiulung).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: China.

riojanus Hozman, 1977: 437 (Platydracus; Type locality: Spanien: Provinz Logroño, Rioja, Torrecilla en Cameros, ca. 800 m).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Spain.

ruandae Bernhauer, 1934g: 239 (Staphylinus, cited as Staphilinus; subgenus Platydracus; Type locality: Ruanda: Gabiru).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Rwanda.

rupicola Kiesenwetter, 1850, see: latebricola Gravenhorst, 1806.

rutilicauda Horn, 1879: 196 (Staphylinus; Type locality: Oregon).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • — Hatch, 1957: 179 (Ocypus; subgenus Platydracus; characters; British Columbia; Idaho; Washington; Oregon).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Canada, USA.

sachalinensis Matsumura, 1911: 112 (Staphylinus; Type locality: Naiptchi, Insel Sachalin).

  • — Scheerpeltz, 1933: 1388 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Russia.

santacoffiensis Levasseur, 1968: 1391 (Platydracus; Type locality: Cameroun: Santa-Coffie, Estete).

  • Distribution: Cameroon.

semicyaneus Bernhauer, 1915l: 297 (Staphylinus; subgenus Euryolinus; Type locality: Peru: Chanchamayo).

  • — Scheerpeltz, 1933: 1388 (Staphylinus; catalog).

  • — Blackwelder, 1952: 160 (type species of Euryolinus, a subgenus of Platydracus).

  • Distribution: Peru.

semipurpureus Kraatz, 1859: 73 (Staphylinus; Type locality: India borealis).

  • — Gemminger and Harold, 1868: 580 (Staphylinus; cited as semicupreus; [Note: Misspelling: Bernhauer and Schubert, 1914: 382, cited semicupreus Gemminger and Harold as a synonym of semipurpureus and a typographical error).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • — Cameron, 1932: 187 (Staphylinus; subgenus Platydracus; characters; India).

  • — Scheerpeltz, 1933: 1388 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: India.

semiviolaceus Cameron, 1932: 191 (Staphylinus; subgenus Platydracus; Type locality: Ceylon: Colombo).

  • — Scheerpeltz, 1933: 1388 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Sri Lanka.

sepulchralis Erichson, 1839b: 388 (Staphylinus; Type locality: Mexico).

  • — Sharp, 1884: 374 (Staphylinus; characters; Mexico).

  • — Lynch, 1884: 140 (Staphylinus; characters; Argentina).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: USA, Mexico, Argentina.

modestus Fall, 1907: 219 (Staphylinus; [preoccupied]; Type locality: New Mexico, Santa Fe and Las Vegas).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; synonym of neomexicanus).

  • — Smetana and Davies, 2000: 40 (Platydracus; synonym of sepulchralis).

neomexicanus Bernhauer and Schubert, 1914: 381 (Staphylinus; replacement name for modestus Fall).

  • — Smetana and Davies, 2000: 40 (Platydracus; synonym of sepulchralis).

sharpi Fauvel, 1901a: 5 (Staphylinus; replacement name for subaeneus Sharp).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1388 (Staphylinus; catalog).

  • — J. Li, 1993: 38 (Staphylinus; China).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Japan, China.

subaeneus Sharp, 1874: 31 (Staphylinus; [Note: Preoccupied]; Type locality: Nagasaki).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; synonym of sharpi).

sjostedti Fauvel, 1903e: 239 (Staphylinus; cited as sjöstedti; Type locality: Cameroun; Congo: Stanley Pool).

  • — Bernhauer, 1913b: 232 (Staphylinus; subgenus Chitocompsus; cited as sjöstedti).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; cited as sjöstedti; catalog).

  • — Levasseur, 1968a: 54 (Platydracus; cited as sjoestedti; Central African Republic).

  • — Herman, 2001: 12 (Platydracus; spelling corrected to sjostedti).

  • Distribution: Cameroon, Congo, Central African Republic.

polyphemus Bernhauer, 1913b: 232 (Staphylinus; subgenus Chitocompsus; Type locality: Urwald zwischen Ukaika und Mawambi).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • — Fagel, 1950: 8 (Staphylinus; synonym of sjostedti, cited as sjöstedti).

sparsus Cameron, 1932: 199 (Staphylinus; subgenus Platydracus; Type locality: Darjeeling: Rungbong Valley).

  • — Scheerpeltz, 1933: 1388 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: India.

speculifrons Bernhauer, 1939k: 593 (Staphylinus; subgenus Platydracus; Type locality: N. W. China: Tienmuschan).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: China.

stercorarius Olivier, 1795:(42): 18 (Staphylinus; Type locality: Paris).

  • — Gravenhorst, 1802: 11 (Staphylinus; characters).

  • — Marsham, 1802: 499 (Staphylinus; characters; Britain).

  • — Latreille, 1804: 299 (Staphylinus; characters; notes; France; Prussia).

  • — Gravenhorst, 1806: 115 (Staphylinus; characters; France; Germany).

  • — Gyllenhal, 1810: 296 (Staphylinus; characters; Sweden).

  • — Gyllenhal, 1813: 703 (Staphylinus; characters for variety; Sweden).

  • — Curtis, 1829: 24 (Staphylinus; catalog; Britain).

  • — C. Sahlberg, 1830: 312 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 22 (Staphylinus; Sweden).

  • — Mannerheim, 1831: 436 (Staphylinus; Sweden).

  • — Stephens, 1832: 206 (Staphylinus; characters; England).

  • — Runde, 1835: 2 (Staphylinus; characters; Germany).

  • — Lacordaire, 1835: 366 (Emus; characters; France).

  • — Nordmann, 1837: 55 (Staphylinus; characters; Germany; Sweden).

  • — Curtis, 1839: pl. 758 (Staphylinus; notes; habitat; England).

  • — Erichson, 1839a: 436 (Staphylinus; characters; Germany).

  • — Heer, 1839: 251 (Staphylinus; characters; Switzerland).

  • — Erichson, 1839b: 380 (Staphylinus; characters; Europe).

  • — Stephens, 1839: 387 (Staphylinus; characters; England).

  • — Laporte, 1840: 176 (Staphylinus; characters; France).

  • — Holme, 1842: 116 (Staphylinus; notes).

  • — Kolenati, 1846: 17 (Staphylinus; Caucasus; Transcaucasia).

  • — Jacobsen, 1849: 683 (Staphylinus; characters; Denmark).

  • — Ménétriés, 1849: 43 (Staphylinus; Khirgiz steppes).

  • — Hochhuth, 1849: 116 (Staphylinus; characters; Akhaltzik; Lenkoran).

  • — Redtenbacher, 1849: 696 (Staphylinus; characters; Austria).

  • — Küster, 1853: 50 (Staphylinus; characters; Finland; Sweden; Germany; France; Switzerland; Italy).

  • — Hardy, 1851: 22 (Staphylinus; notes; England).

  • — Fairmaire and Laboulbène, 1856: 507 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 543 (Staphylinus; characters; Germany).

  • — Redtenbacher, 1857: 185 (Staphylinus; characters; Austria).

  • — Thomson, 1858: 29 (Platydracus; type species).

  • — Thomson, 1859: 23 (type species of Platydracus).

  • — Thomson, 1860: 143 (Platydracus; characters; Scandinavia).

  • — Fauvel, 1874: 178 [= 1874a: 404] (Staphylinus; characters; France; Europe; Caucasus; Transcaucasia).

  • — Redtenbacher, 1874: 205 (Staphylinus; characters; Austria).

  • — Seidlitz, 1875: 269 (Leistotrophus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 266 [= 1877: 122] (Platydracus; characters; France).

  • — J. Sahlberg, 1876: 9 (Platydracus; Finland).

  • — Fowler, 1888: 251 (Staphylinus; characters; Britain).

  • — Ganglbauer, 1895: 427 (Staphylinus; subgenus Platydracus; characters; Europe; eastern Mediterranean region).

  • — Reitter, 1909: 118 (Staphylinus; subgenus Platydracus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • — Johansen, 1914: 379 (Staphylinus; subgenus Platydracus; characters; Denmark).

  • — G. Müller, 1923: 135 (Staphylinus; subgenus Platydracus; characters; Italy).

  • — G. Müller, 1926: 6 (Staphylinus; subgenus Platydracus; notes).

  • — Portevin, 1929: 349 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1388 (Staphylinus; catalog).

  • — V. Hansen, 1952: 98 (Staphylinus; characters; Denmark).

  • — Coiffait, 1956e: 193 (Platydracus; characters).

  • — Smetana, 1958: 272 (Staphylinus; subgenus Platydracus; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 110 (Staphylinus; subgenus Platydracus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 194 (Platydracus; characters; central Europe).

  • — Smetana, 1965c: 27 (Staphylinus; subgenus Platydracus; Anatolia).

  • — Horion, 1965: 204 (Staphylinus; subgenus Platydracus; Norway; all central Europe; Spain; Portugal; Croatia; Bosnia Hercegovina; Bulgaria; Ukraine; Russia).

  • — Szujecki, 1966c (Staphylinus; larval characters; notes on natural history).

  • — Pototskaia, 1967: 68 (Staphylinus; subgenus Platydracus; larval characters).

  • — Smetana, 1968a: 156 (Staphylinus; subgenus Platydracus; Anatolia).

  • — Kasule, 1970: 72 (Platydracus; larval characters).

  • — Smetana, 1971c: 263 (Staphylinus; subgenus Platydracus; Iran).

  • — Jarrige, 1971: 496 (Platydracus; Iran).

  • — Coiffait, 1974: 395 (Platydracus; characters; Europe; Britain; Scandinavia; Caucasus; Transcaucasus).

  • — Pope, 1977: 30 (Platydracus; Britain).

  • — Petrenko, 1980a: 60, 62 (Staphylinus; subgenus Platydracus; characters; Ukraine).

  • — Tóth, 1984: 16 (Platydracus; characters; Hungary).

  • — Lucht, 1987: 107 (Platydracus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Schülke and Uhlig, 1988: 12 (Platydracus; Germany).

  • — Pilon and Zanetti, 1991: 67 (Platydracus; Italy).

  • — Smetana, 1993b: 49 (Platydracus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 30 (Platydracus; Italy).

  • — Mikat, Fremuth, and Prouza, 1997: 111 (Platydracus; Czech Republic).

  • — Pilon, 1998: 71 (Platydracus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • — Assing, 2001: 75 (Platydracus; collected from xerothermous site; Germany).

  • Distribution: Europe, Georgia, Azerbaijan, Turkey, Iran, Turkmenistan, Kazakhstan.

crebrepunctatus Motschulsky, 1860: 570 (Staphylinus; Type locality: Géorgie; Turcmenie sur la côté orientale de la mer Caspienne).

  • — Hochhuth, 1862: 48 (Staphylinus; characters; Turkmenia; Georgia; Caspian Sea).

  • — Fauvel, 1874: 178 [= 1874a: 404] (Staphylinus; synonym of stercorarius).

  • — Ganglbauer, 1895: 427 (Staphylinus; synonym of stercorarius).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; synonym of stercorarius).

  • — Smetana, 1958: 272 (Staphylinus; synonym of stercorarius).

  • — Coiffait, 1974: 395 (Platydracus; synonym of stercorarius).

fuscofemoratus G. Müller, 1923: 136 (Staphylinus; subgenus Platydracus; variety of stercorarius; Type locality: Val d'Aosta, Piastre - Pistoia, Laigueglia - Liguria occ., N. S. della Vittoria, sull'Appennino ligure).

  • — G. Müller, 1926: 6 (Staphylinus; subgenus Platydracus; infraspecies of stercorarius; notes).

  • — Scheerpeltz, 1933: 1388 (Staphylinus; subspecies of stercorarius).

  • — Smetana, 1958: 272 (Staphylinus; subspecies of stercorarius).

  • — Jarrige, 1968b: 74 (Platydracus; subspecies of stercorarius; France).

  • — Coiffait, 1974: 395 (Platydracus; variety of stercorarius; characters; Italy; France; Spain).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus; subspecies of stercorarius).

subaeneipennis Scheerpeltz, 1976a: 127 (Staphylinus; subgenus Platydracus; Type locality: Nepal: Jiri, 1900 m).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Nepal.

subaeneus Roth, 1851: 118 (Ocypus; Type locality: Abyssinien).

  • — Fauvel, 1907: 36 (Staphylinus; Kilimandjaro; Ethiopia; Congo; Gabon; Natal).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Levasseur, 1966a: 1215 (Platydracus; Congo; from Abyssinia and Gabon to Natal).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus; [Note: Cited erroneously as a Palaearctic species]).

  • Distribution: Ethiopia, Kenya, Congo, Gabon, South Africa.

subaeneus Sharp, 1874, see: sharpi Fauvel, 1901.

subauronotatus Coiffait, 1977b: 229 (Platydracus; Type locality: Bhutan: 87 km de Phuntsholing).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Bhutan.

subirideus Kraatz, 1859: 78 (Staphylinus; Type locality: India orientalis).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Cameron, 1932: 199 (Staphylinus; subgenus Platydracus; characters; East Indies).

  • — Scheerpeltz, 1933: 1389 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: “East Indies”.

submarmorellus Schubert, 1908: 613 (Staphylinus; Type locality: Kulu, Himalaya, ca. 1500 m).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Cameron, 1932: 194 (Staphylinus; subgenus Platydracus; characters; India).

  • — Scheerpeltz, 1933: 1389 (Staphylinus; catalog).

  • — Cerruti, 1951: 16 (Staphylinus; subgenus Platydracus; India).

  • — Coiffait, 1977b: 211 (Platydracus; Bhutan).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: India, Bhutan.

submetallicus LeConte, 1861, see: tarsalis Mannerheim, 1843.

subtilicornis Fagel, 1957c: 263 (Platydracus; subgenus Abemus; Type locality: Congo Belge: Kivu: Mt. Muhi, ruisseau Kanazo, bassin moyenne Ulindi).

  • Distribution: Congo.

subviridis Bernhauer, 1933a: 45 (Staphylinus; subgenus Platydracus; Type locality: Yunnan-Sen: Junnam).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: China.

sumakowi Bernhauer, 1911b: 236 (Staphylinus; Type locality: Samarkand).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1389 (Staphylinus; catalog).

  • — Coiffait, 1974: 396 (Platydracus; characters; Turkestan).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Uzbekistan.

surdus Fagel, 1957c: 267 (Platydracus; Type locality: Congo Belge: Kivu: territoire d'Uvira, Haute Ngovi, 3000–3200 m).

  • Distribution: Congo.

suspectus Fauvel, 1904a: 57 (Staphylinus; Type locality: Nilghiris; Matheran; Mahe).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Cameron, 1932: 193 (Staphylinus; subgenus Platydracus; characters; India).

  • — Scheerpeltz, 1933: 1389 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: India.

suspiciosus Bernhauer, 1937: 22 (Staphylinus; subgenus Platydracus; Type locality: Java: Zuid Bankoemas, Djeroeklegi).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Indonesia.

tarsalis Mannerheim, 1843b: 229 (Staphylinus; Type locality: insula Sitkha).

  • — Horn, 1879: 192 (Staphylinus; characters; Sitka to California and Arizona).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1389 (Staphylinus; catalog).

  • — Hatch, 1957: 178 (Ocypus; subgenus Platydracus; characters; British Columbia; Idaho; Washington; Oregon).

  • — Smetana, 1965a: 10 (Staphylinus; subgenus Platydracus; British Columbia).

  • — Smetana and Davies, 2000: 40 (Platydracus; subgenus Platydracus).

  • Distribution: Canada, USA, Mexico.

submetallicus LeConte, 1861: 342 (Staphylinus; Type locality: Tejon).

  • — Horn, 1879: 192 (Staphylinus; synonym of tarsalis).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1389 (Staphylinus; synonym of tarsalis).

  • — Smetana, 1965a: 10 (Staphylinus; synonym of tarsalis).

  • — Smetana and Davies, 2000: 40 (Platydracus; synonym of tarsalis).

lucanus Horn, 1894: 362 (Staphylinus; Type locality: La Chuparosa, Baja California).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 40 (Platydracus; synonym of tarsalis).

curticollis Bernhauer, 1917a: 3 (Staphylinus; subgenus Platydracus; Type locality: Kalifornien: Pasadena).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 41 (Platydracus; synonym of tarsalis).

  • — Herman, 2001: 14 (Platydracus).

tarsalis Sharp, 1884, see: panamensis Bernhauer and Schubert, 1914.

temporalis Casey, 1915: 429 (Staphylinus; Type locality: Florida: Jacksonville; North Carolina: Southern Pines).

  • — Scheerpeltz, 1933: 1389 (Staphylinus; catalog).

  • — Lundgren, 1998: 52 (Platydracus; Florida).

  • — Smetana and Davies, 2000: 41 (Platydracus; subgenus Platydracus).

  • Distribution: USA.

teter Bondroit, 1913, see: aureofasciatus Motschulsky, 1862.

tomentosus Gravenhorst, 1802: 161 (Staphylinus; [preoccupied]; Type locality: America septentr.).

  • — Gravenhorst, 1806: 111 (Staphylinus).

  • — Nordmann, 1837: 50 (Staphylinus; characters; America septentrionalis).

  • — Erichson, 1839b: 389 (Staphylinus; characters; America septentrionalis).

  • — Provancher, 1877: 729 (Staphylinus; characters; Quebec).

  • — Horn, 1879: 193 (Staphylinus; characters; middle States to Louisiana).

  • — Fall and Cockerell, 1907: 166 (Staphylinus; New Mexico).

  • — Blatchley, 1910: 377 (Staphylinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Voris, 1939a: 288, 297 (Staphylinus; larval and pupal characters).

  • — Hatch, 1957: 178 (Ocypus; subgenus Platydracus; characters; Washington).

  • — Schmidt, 1994a (Platydracus; egg, larval and pupal characters; biological notes).

  • — Downie and Arnett, 1996: 386 (Staphylinus; characters; USA).

  • — Lundgren, 1998: 52 (Platydracus; Florida).

  • — Smetana and Davies, 2000: 41 (Platydracus; subgenus Platydracus).

  • — Herman, 2001: 49 (Platydracus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: Canada, USA.

cubae Jacquelin du Val, 1857a: 36 (Ocypus; Type locality: Not cited, presumably Cuba).

  • — Chevrolat and Fauvel, 1863: 432 (Ocypus; characters; Cuba).

  • — Fauvel, 1869: 488 (Staphylinus).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • — Blackwelder, 1943: 446 (Staphylinus; synonym of antiquus).

  • — Smetana and Davies, 2000: 41 (Platydracus; synonym of tomentosus).

toumodiensis Levasseur, 1967b: 1642 (Platydracus; Type locality: Côte d'Ivoire, Lamto, Toumodi).

  • Distribution: Ivory Coast.

tricolor Gravenhorst, 1802, see: fulvipes Scopoli, 1763.

triplicans Casey, 1924, see: viridanus Horn, 1879.

uheheanus Bernhauer, 1937b: 618 (Staphylinus; subgenus Platydracus; Type locality: Uheheland: Kidugala).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Tanzania.

ussuriensis Solsky, 1871c: 347 (Staphylinus; Type locality: Not cited but presumably in eastern Siberia).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1389 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 41 (Platydracus; subgenus Platydracus).

  • Distribution: Russia.

varipes Sachse, 1852, see: femoratus Fabricius, 1801.

vicarius Sharp, 1889: 117 (Staphylinus; Type locality: Nikko; Awomori).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1389 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 41 (Platydracus; subgenus Platydracus).

  • Distribution: Japan.

viduatus Fabricius, 1801, see: maculosus Gravenhorst, 1802.

violaceovirens Fauvel, 1905b: 176 (Staphylinus; Type locality: forêt de Moramanga).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Jarrige, 1978: 291 (Platydracus; Madagascar).

  • Distribution: Madagascar.

violaceus Gravenhorst, 1802: 162 (Staphylinus; [preoccupied]; Type locality: America septentr.).

  • — Gravenhorst, 1806: 110 (Staphylinus).

  • — Say, 1830: 38 (Staphylinus; characters).

  • — Say, 1834: 451 (Staphylinus; characters).

  • — Nordmann, 1837: 48 (Staphylinus; characters; America septentrionalis).

  • — Erichson, 1839b: 392 (Staphylinus; characters; America septentrionalis).

  • — Melsheimer, 1844: 35 (Staphylinus; characters; USA).

  • — Provancher, 1877: 247 (Staphylinus; characters; Quebec).

  • — Horn, 1879: 196 (Staphylinus; characters; Canada to North Carolina).

  • — Blatchley, 1910: 378 (Staphylinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Hoebeke, 1978: 507 (Platydracus; parasitized by proctotrupid; collecting notes; notes on parasite; Michigan).

  • — Hoebeke and Kovarik, 1988: 218 (Platydracus; parasitized by serphid).

  • — Dearborn and Donahue, 1994: 15 (Platydracus; Maine).

  • — Downie and Arnett, 1996: 386 (Staphylinus; cited as violaceous; characters; USA).

  • — Lundgren, 1998: 52 (Platydracus; Florida).

  • — Smetana and Davies, 2000: 25, 39 (Platydracus; synonym of cupripennis).

  • — Newton, in press (Platydracus; application to Commission to maintain use as vald name of species).

  • Distribution: Canada, USA.

cupripennis Melsheimer, 1844: 35 (Staphylinus; variety of violaceus; [Note: This name was spelled “cupripennis” in the copy available to me as a bound journal from the library of the American Museum of Natural History. It was spelled “cuprepennis” in a copy of the article supplied to me by A. Smetana who queried my spelling of the name. A few other differences included the following. On page 26 of the article sent by Smetana there is an up-side-down male symbol in the last line of the characters for Thermonectus irroratus; in the Museum's copy the female symbol is present. On page 35 of Smetana's copy, in the description of Philonthus harrisii, the male symbol points to about 12: 30, in the Museum's copy it points to about 11: 30. The type face of the italicized words in the two copies differs. I assume that Smetana's copy was separately produced, and probably reset, after the journal was printed. The name “cupripennis” is accepted herein as the correct spelling]; Type locality: Not cited).

  • — Gemminger and Harold, 1868: 580 (Staphylinus; synonym of violaceus).

  • — Horn, 1879: 196 (Staphylinus; synonym of violaceus).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; variety of violaceus).

  • — Smetana and Davies, 2000: 25, 39 (Platydracus; subgenus Platydracus; valid name).

  • — Newton, in press (Platydracus; application to Commission to list name as junior synonym of maculosus).

fenyesi Bernhauer, 1917a: 3 (Staphylinus; subgenus Abemus; Type locality: Nord-Carolina: Murphy).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 39 (Platydracus; synonym of cupripennis).

virgulatus Fauvel, 1895: 254 (Staphylinus; Type locality: Bírmanie, Carin Cheba, 500–1000 m; Carin Asciuii Ghécu, 1400–1500 m).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Cameron, 1932: 190 (Staphylinus; subgenus Platydracus; characters; Burma; India).

  • — Scheerpeltz, 1933: 1389 (Staphylinus; catalog).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Myanmar, India.

viridanus Horn, 1879: 196 (Staphylinus; Type locality: New England States; Canada; New York; Michigan).

  • — Blatchley, 1910: 378 (Staphylinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Voris, 1939a: 288, 297 (Staphylinus; larval and pupal characters).

  • — Downie and Arnett, 1996: 386 (Staphylinus; characters; USA).

  • — Smetana and Davies, 2000: 41 (Platydracus; subgenus Platydracus).

  • Distribution: Canada, USA.

triplicans Casey, 1924: 149 (Staphylinus; Type locality: Maine, Eastbrook).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; catalog).

  • — Downie and Arnett, 1996: 386 (Staphylinus; characters; USA).

  • — Smetana and Davies, 2000: 41 (Platydracus; synonym of viridanus).

vulpinus Nordmann, 1837, see: immaculatus Mannerheim, 1830.

wittei Fagel, 1950: 10 (Staphylinus; subgenus Platydracus; Type locality: Congo belge: Katanga: Mitwaba, Nanono; also listed from Angola).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Angola, Congo.

wittmeri Coiffait, 1977b: 227 (Platydracus; Type locality: Bhutan 87 km de Phuntsholing).

  • — Smetana and Davies, 2000: 41 (Platydracus; subgenus Platydracus).

  • Distribution: Bhutan.

yolensis Cerruti, 1951: 20 (Staphylinus; subgenus Platydracus; Type locality: India: Punjab: Yol, 2000–2500 m).

  • — Smetana and Davies, 2000: 41 (Platydracus; subgenus Platydracus).

  • Distribution: India.

yunnanensis Bernhauer, 1943, see: yunnanicus Smetana and Davies, 2000.

yunnanicus Smetana and Davies, 2000: 25, 41 (Platydracus; replacement name for yunnanensis Bernhauer, 1943).

  • Distribution: China.

yunnanensis Bernhauer, 1943: 76 (Staphylinus; subgenus Platydracus; [preoccupied]; Type locality: China: Prov. Yunnan, Vallis flumin. Soling-ho).

  • — Smetana and Davies, 2000: 25, 41 (Platydracus; synonym of yunnanicus).

zavattarii Gridelli, 1939: 318 (Staphylinus; subgenus Platydracus; Type locality: Neghelli; Arero).

  • — Herman, 2001: 14 (Platydracus).

  • Distribution: Ethiopia.

zonatus Gravenhorst, 1802: 162 (Staphylinus; Type locality: America septentr.).

  • — Gravenhorst, 1806: 113 (Staphylinus).

  • — Nordmann, 1837: 61 (Staphylinus; characters; America septentrionalis).

  • — Erichson, 1839b: 390 (Staphylinus; synonym of cinnamopterus).

  • — Horn, 1879: 195 (Staphylinus; synonym of cinnamopterus).

  • — Smetana and Davies, 2000: 41 (Platydracus; subgenus Platydracus; cited as valid species).

  • Distribution: USA.

badius Mannerheim, 1830: 23 (Staphylinus; Type locality: America borealis).

  • — Mannerheim, 1831: 437 (Staphylinus; America borealis).

  • — Nordmann, 1837: 59 (Staphylinus; America borealis).

  • — Erichson, 1839b: 390 (Staphylinus; synonym of cinnamopterus).

  • — Horn, 1879: 195 (Staphylinus; cited as badipes; synonym of cinnamopterus).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; synonym of cinnamopterus).

  • — Smetana and Davies, 2000: 41 (Platydracus; synonym of zonatus).

quadraticeps Casey, 1924: 149 (Staphylinus; [preoccupied]; Type locality: Florida).

  • — Scheerpeltz, 1933: 1392 (Staphylinus; synonym of caseyi).

  • — Smetana and Davies, 2000: 41 (Platydracus; synonym of zonatus).

caseyi Scheerpeltz, 1933: 1392 (Staphylinus; replacement name for quadraticeps Casey).

  • — Lundgren, 1998: 52 (Platydracus; Florida).

  • — Smetana and Davies, 2000: 41 (Platydracus; synonym of zonatus).

Protogoerius [1 species; Palaearctic Region]

Protogoerius Coiffait, 1956e: 184, 198 (species included: brachypterus). Type species: Staphylinus brachypterus Brullé, fixed by monotypy.

  • — Coiffait, 1974: 407 (characters; species of western Palaearctic region).

  • — Smetana and Davies, 2000: 21, 30, 42 (characters; checklist of species of north temperate region).

brullei Smetana and Davies, 2000: 30, 42 (Protogoerius; replacement name for brachypterus).

  • Distribution: Canary Islands.

brachypterus Brullé, 1839: 59 (Staphylinus; [preoccupied]; Type locality: Canary Islands).

  • — Kraatz, 1857d: 553 (Ocypus; characters; Germany; Austria).

  • — Redtenbacher, 1857: 187 (Ocypus; characters; Austria).

  • — Wollaston, 1864: 565 (Ocypus; characters; Canary Islands).

  • — Wollaston, 1865: 488 (Ocypus; notes; European records are erroneous; Canary Islands).

  • — Fauvel, 1866c: 248 [= 1866d: 106] (Staphylinus; synonym of olens).

  • — Fauvel, 1867: LV (Ocypus).

  • — Redtenbacher, 1874: 206 (Ocypus; characters; Austria).

  • — Mulsant and Rey, 1876: 280 [= 1877: 136] (Ocypus; subgenus Goerius; characters; France).

  • — Fauvel, 1902b: 104 (Staphylinus; subgenus Ocypus; Canary Islands).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1391 (Staphylinus; catalog).

  • — Uyttenboogaart, 1937: 81 (Staphylinus; notes; Canary Islands).

  • — Coiffait, 1954: 166 (Goerius; Canary Islands).

  • — Jarrige, 1954a: 161 (Staphylinus; subgenus Goerius; characters; Canaries).

  • — Coiffait, 1956e: 184, 198 (Protogoerius; characters).

  • — Coiffait, 1974: 407 (Protogoerius; characters; Canary Islands).

  • — Hernández, Outerelo, and Gamarra, 1994: 188 (Protogoerius; checklist of species; Canary Islands).

  • — Smetana and Davies, 2000: 30, 42 (Protogoerius; synonym of brullei).

Rhynchocheilus [1 species; Oriental Region]

Rhynchocheilus Sharp, 1889: 120 (species included: pectoralis). Type species: Rhynchocheilus pectoralis Sharp, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 394 (world catalog; cited as Rhyncocheilus; 9 species).

  • — Cameron, 1925: 68 (catalog of Indian species).

  • — Cameron, 1932: 223 (cited as Rhyncochilus; characters; key to species of British India).

  • — Blackwelder, 1952: 341 (synonym of Rhyncocheilus; type species: pectoralis).

  • — Hayashi, 1999: 553, 554, 563 (valid genus; characters; in Eucibdelus groups; spelling of generic name easily confused with similar name by Fauvel; new name should be substituted under plenary power of Commission; [Note: Although Hayashi stated that R. pectoralis was closely allied to three species included in Rhyncocheilus, he explicitly included only one species, the type species. Perhaps others will be transferred from among the remaining species of Rhyncocheilus]).

pectoralis Sharp, 1889: 120 (Rhynchocheilus; Type locality: Malacca, Malay Pen. [Note: See lectotype designation by Hayashi, 1999]).

  • — Bernhauer and Schubert, 1914: 394 (Rhyncocheilus; catalog).

  • — Chapman, 1933a: 222 (Rhyncocheilus).

  • — Hayashi, 1999: 567 (Rhyncocheilus; lectotype designation; characters; Malaysia).

  • Distribution: Malaysia.

Rhyncocheilus [26 species; Palaearctic and Oriental Regions]

Rhyncocheilus Fauvel, 1882c: 211 (species included: aureus). Type species: Staphylinus aureus Fabricius, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 394 (synonym of Rhynchocheilus Sharp).

  • — Cameron, 1932: 223 (synonym of Rhynchocheilus, cited as Rhyncochilus).

  • — Scheerpeltz, 1933: 1407 (world catalog supplement).

  • — Chapman, 1933a: 220 (characters).

  • — Blackwelder, 1952: 341 (type species: aureus).

  • — Hammond, 1984: 207 (checklist of species of Borneo).

  • — Hayashi, 1999: 554 (characters; distinct from Rhynchocheilus).

andrewesi Cameron, 1918a: 105 (Rhynchocheilus; Type locality: India: Nilgiri Hills).

  • — Cameron, 1932: 226 (Rhyncocheilus, cited as Rhyncochilus; characters; India).

  • — Scheerpeltz, 1933: 1407 (Rhyncocheilus; catalog).

  • — Chapman, 1933a: 222 (Rhyncocheilus).

  • Distribution: India.

antennalis Cameron, 1932, see: Trichocosmetes.

argenteus Fauvel, 1895: 249 (Rhyncocheilus, cited as Rhyncochilus; Type locality: Bírmanie, Teinzo; Carin Cheba, 900–1100 m).

  • — Bernhauer and Schubert, 1914: 394 (Rhyncocheilus; catalog).

  • — Cameron, 1932: 231 (Rhyncocheilus, cited as Rhyncochilus; characters; Burma).

  • — Scheerpeltz, 1933: 1408 (Rhyncocheilus; catalog).

  • — Chapman, 1933a: 222 (Rhyncocheilus).

  • Distribution: Myanmar.

assamensis Cameron, 1932: 229 (Rhyncocheilus, cited as Rhyncochilus; Type locality: Assam: Patkai Mts.).

  • — Scheerpeltz, 1933: 1408 (Rhyncocheilus; catalog).

  • — Chapman, 1933a: 222 (Rhyncocheilus).

  • Distribution: India.

aureus Fabricius, 1787: 219 (Staphylinus; Type locality: Siam).

  • — Gmelin, 1790: 2025 (Staphylinus; characters; Thailand).

  • — Fabricius, 1793: 519 (Staphylinus; characters; Thailand).

  • — Olivier, 1795:(42): 7 (Staphylinus; characters; Thailand).

  • — Fabricius, 1801: 589 (Staphylinus; characters; Thailand).

  • — Turton, 1802: 509 (Staphylinus; characters; Siam).

  • — Erichson, 1839b: 344 (Palaestrinus ?; characters; Siam).

  • — Fauvel, 1882c: 211 (Rhyncocheilus; Siam; Sylhet; Cochinchine).

  • — Fauvel, 1904a: 57 (Rhyncochilus; Kanara; Ceylan; Bírmanie; Tenasserim; Siam; Borneo; Sumatra; Java).

  • — Bernhauer and Schubert, 1914: 394 (Rhyncocheilus; catalog).

  • — Cameron, 1932: 225 (Rhyncocheilus, cited as Rhyncochilus; characters; Ceylon; India; Burma).

  • — Scheerpeltz, 1933: 1408 (Rhyncocheilus; catalog).

  • — Cameron, 1933d: 391 (Rhyncocheilus, cited as Rhyncochilus; Sumatra).

  • — Chapman, 1933a: 222 (Rhyncocheilus).

  • — Hayashi, 1999: 559 (Rhyncocheilus; characters; Thailand).

  • Distribution: Borneo, Indonesia, Vietnam, Thailand, Myanmar, Bangladesh, India, Sri Lanka.

beauchenei Fauvel, 1897: 44 (Rhyncochilus; Type locality: Tonkin).

  • — Bernhauer and Schubert, 1914: 394 (Rhyncocheilus; catalog).

  • — Chapman, 1933a: 222 (Rhyncocheilus).

  • Distribution: Vietnam.

bernhaueri Chapman, 1933a: 220 (Rhyncocheilus; Type locality: Haut-Laos).

  • Distribution: Vietnam.

borneensis Cameron, 1942, see: princeps Bernhauer, 1915.

brachycerus Fauvel, 1897b: 229 (Rhyncochilus; Type locality: Cambodje).

  • — Bernhauer and Schubert, 1914: 394 (Rhyncocheilus; catalog).

  • — Chapman, 1933a: 222 (Rhyncocheilus).

  • Distribution: Cambodia.

cameroni Chapman, 1933a: 221 (Rhyncocheilus; Type locality: Tonkin, Chapa).

  • Distribution: Vietnam.

chrysites Fauvel, 1895: 248 (Rhyncocheilus, cited as Rhyncochilus; Type locality: Bírmanie, Teinzo).

  • — Bernhauer and Schubert, 1914: 394 (Rhyncocheilus; catalog).

  • — Cameron, 1932: 227 (Rhyncocheilus, cited as Rhyncochilus; characters; Burma).

  • — Scheerpeltz, 1933: 1408 (Rhyncocheilus; catalog).

  • — Chapman, 1933a: 222 (Rhyncocheilus).

  • Distribution: Myanmar.

dohertyi Cameron, 1932: 226 (Rhyncocheilus, cited as Rhyncochilus; Type locality: Burma: Ruby Mines).

  • — Scheerpeltz, 1933: 1408 (Rhyncocheilus; catalog).

  • — Chapman, 1933a: 222 (Rhyncocheilus).

  • Distribution: Myanmar.

drescheri Cameron, 1937: 23 (Rhyncocheilus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft.).

  • Distribution: Indonesia.

elegans Scheerpeltz, 1965: 253 (Rhyncocheilus; Type locality: N.E. Burma, Kambaiti, 2000 m).

  • Distribution: Myanmar.

exophthalmus Hayashi, 1998: 87 (Rhyncocheilus, cited as Rhynchocheilus; Type locality: Malacca, Malay Pen.).

  • Distribution: Malaysia.

foersteri Bernhauer, 1915a: 26 (Rhyncocheilus, cited as Rhyncochilus; Type locality: Java).

  • — Scheerpeltz, 1933: 1408 (Rhyncocheilus; catalog).

  • Distribution: Indonesia.

franzi Coiffait, 1977, see: Trichocosmetes.

germanus Cameron, 1932: 232 (Rhyncocheilus, cited as Rhyncochilus; Type locality: Assam: Patkai Mts.).

  • — Scheerpeltz, 1933: 1408 (Rhyncocheilus; catalog).

  • — Chapman, 1933a: 222 (Rhyncocheilus).

  • Distribution: India.

griseosericans Fairmaire, 1894: 217 (Emus; Type locality: Se-pin-lou-shan, Ya Tcheou, Thibet).

  • — Bernhauer and Schubert, 1914: 391 (Emus; catalog).

  • — Scheerpeltz, 1933: 1405 (Emus; catalog).

  • — Coiffait, 1974: 383 (Rhynchocheilus, cited as Rhyncocheilus Sharp).

  • — Smetana and Davies, 2000: 38 (Emus).

  • Distribution: China.

javanicus Eppelsheim, 1895b: 394 (Eucibdelus; Type locality: Ostjava: Montes Tengger, 4000 Fuss).

  • — Bernhauer and Schubert, 1914: 394 (Rhyncocheilus; catalog).

  • — Cameron, 1932: 228 (Rhyncocheilus, cited as Rhyncochilus; characters; Burma).

  • — Scheerpeltz, 1933: 1408 (Rhyncocheilus; catalog).

  • — Chapman, 1933a: 222 (Rhyncocheilus).

  • — Cameron, 1937: 23 (Rhyncocheilus; Java).

  • Distribution: Indonesia, Myanmar.

limbatus Fauvel, 1895: 250 (Rhyncocheilus, cited as Rhyncochilus; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Java, monts Tengger, 1300 m).

  • — Bernhauer and Schubert, 1914: 394 (Rhyncocheilus; catalog).

  • — Cameron, 1932: 228 (Rhyncocheilus, cited as Rhyncochilus; synonym of javanicus).

  • — Scheerpeltz, 1933: 1408 (Rhyncocheilus; synonym of javanicus).

  • — Chapman, 1933a: 222 (Rhyncocheilus; synonym of javanicus).

kraatzi Eppelsheim, 1895b: 392 (Eucibdelus; Type locality: Pegu).

  • — Bernhauer and Schubert, 1914: 394 (Rhyncocheilus; catalog).

  • — Cameron, 1932: 230 (Rhyncocheilus, cited as Rhyncochilus; characters; Burma).

  • — Scheerpeltz, 1933: 1408 (Rhyncocheilus; catalog).

  • — Chapman, 1933a: 222 (Rhyncocheilus).

  • Distribution: Myanmar.

lanosipennis Scheerpeltz, 1965: 252 (Rhyncocheilus; Type locality: N.E. Burma, Kambaiti, 1800 m).

  • Distribution: Myanmar.

limbatus Fauvel, 1895, see: javanicus Eppelsheim, 1895.

magnificus Semenov-Tian-Shanskii and Kirshenblat, 1938: 133 (Rhynchocheilus, cited as Rhynchochilus; Type locality: Prov. Setshuan Chinae interioris: angustiae Cho-dzigou prope oppidum Lun-an-fu).

  • Distribution: China.

malaisei Scheerpeltz, 1965: 250 (Rhyncocheilus; Type locality: Burma, ca. 200 m, Washaung, 20 km east of Myitkina).

  • Distribution: Myanmar.

minor Coiffait, 1982a: 263 (Rhyncocheilus, cited as Rhyncochilus; Type locality: Inde: Meghalaya, Upper Shillong, 1900 m).

  • Distribution: India.

preangeranus Cameron, 1937: 24 (Rhyncocheilus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft.).

  • Distribution: Indonesia.

princeps Bernhauer, 1915a: 26 (Rhyncocheilus, cited as Rhyncochilus; Type locality: Java orient.: Montes Tengger, 4000 Fuss).

  • — Scheerpeltz, 1933: 1408 (Rhyncocheilus; catalog).

  • Distribution: Borneo, Indonesia.

borneensis Cameron, 1942a: 137 (Rhyncocheilus; variety of princeps; Type locality: Borneo: Doesonlanden).

rugulipennis Cameron, 1932: 226 (Rhyncocheilus, cited as Rhyncochilus; Type locality: Burma: Ruby Mines).

  • — Scheerpeltz, 1933: 1408 (Rhyncocheilus; catalog).

  • — Chapman, 1933a: 222 (Rhyncocheilus).

  • — Scheerpeltz, 1965: 110 (Rhyncocheilus; Burma).

  • Distribution: Myanmar.

tibialis Cameron, 1932: 233 (Rhyncocheilus, cited as Rhyncochilus; Type locality: Assam).

  • — Scheerpeltz, 1933: 1408 (Rhyncocheilus; catalog).

  • — Chapman, 1933a: 222 (Rhyncocheilus, cited as tibiatis).

  • — Coiffait, 1982a: 233 (Rhyncocheilus, cited as Rhyncochilus; India).

  • Distribution: India.

Saniderus [2 species; Oriental Region]

Saniderus Fauvel, 1895: 257 (species included: ruficollis, laevicollis). Type species: Saniderus ruficollis Fauvel, fixed by subsequent designation by R. Lucas, 1920: 578.

  • — Bernhauer and Schubert, 1914: 373 (world catalog; 2 species).

  • — Cameron, 1925: 65 (catalog of Indian species).

  • — Cameron, 1932: 177 (characters).

  • — Scheerpeltz, 1933: 1380 (world catalog supplement).

  • — Blackwelder, 1952: 343 (type species).

laevicollis Fauvel, 1895: 258 (Saniderus; Type locality: Java occidental, mont Gede, 1300 m).

  • — Bernhauer and Schubert, 1914: 373 (Saniderus; catalog).

  • Distribution: Indonesia.

ruficollis Fauvel, 1895: 257 (Saniderus; Type locality: Bírmanie, Carin Cheba, 900–1100 m).

  • — Bernhauer and Schubert, 1914: 373 (Saniderus; catalog).

  • — Cameron, 1932: 177 (Saniderus; characters; Burma).

  • — Scheerpeltz, 1933: 1380 (Saniderus; catalog).

  • — Jarrige, 1951: 134 (Saniderus; Tonkin).

  • — Scheerpeltz, 1965: 109 (Saniderus; Burma).

  • Distribution: Myanmar, Vietnam.

Sphaeriolinus [1 species; Oceanic Region]

Sphaeriolinus Steel, 1947: 176 (species included: georgii). Type species: Sphaeriolinus georgii Steel, fixed by original designation and monotypy.

  • — Blackwelder, 1952: 355 (type species: georgii).

georgii Steel, 1947: 176 (Sphaeriolinus; Type locality: Rapa Is., 1,100 feet).

  • Distribution: Austral Islands.

Staphylinus [218 species (78 doubtful); Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, and Australian (?) Regions]

Staphylinus Linné, 1758: 421 (species included: hirtus, murinus, maxillosus, erytropterus (sic), politus, rufus, riparius, lignorum, subterraneus, flavescens, biguttatus, littoreus, sanguineus, caraboides, chrysomelinus, flavipes, fuscipes, rufipes, boleti). Type species: Staphylinus erythropterus Linné, fixed by action of Commission in Opinion 546; emendation of original spelling of type species validated by action of the Commission in same Opinion (ICZN, 1959).

  • — O. Müller, 1764: xv, 23 (characters).

  • — De Geer, 1774: 9 (characters; larvae).

  • — Fabricius, 1777: 84 (characters).

  • — Goeze, 1777: 713 (characters).

  • — Harrer, 1784: 249 (characters; synonyms).

  • — Fourcroy, 1785: 163 (characters).

  • — Fabricius, 1787: 219 (characters).

  • — Paykull, 1789: 1 (characters; discussion).

  • — Villers, 1789: 409 (characters).

  • — Gmelin, 1790: 2025 (characters).

  • — Rossi, 1792: 95 (characters).

  • — Fabricius, 1793: 519 (characters).

  • — Olivier, 1795:(42): 1 (characters).

  • — Latreille, 1797: 78 (characters).

  • — Fabricius, 1801: 589 (characters).

  • — Latreille, 1802: 125 (characters; divisions of genus).

  • — Gravenhorst, 1802: 1 (characters).

  • — Marsham, 1802: 496 (characters).

  • — Latreille, 1804: 272, 284, 289 (description; notes).

  • — Gravenhorst, 1806: 35 (characters; notes).

  • — Latreille, 1806: 285 (characters).

  • — Latreille, 1810: 179, 427 (characters; type species).

  • — Gyllenhal, 1810: 278 (characters).

  • — Leach, 1815: 91 (characters).

  • — Lamarck, 1817: 481 (characters; notes; list of some species).

  • — Samouelle, 1819: 171 (characters; type species: erythropterus).

  • — Billberg, 1820: 14 (characters; list of some species).

  • — Dejean, 1821: 21 (list of species in collection).

  • — Latreille, 1825: 244 (cited as Staphylin; characters).

  • — Berthold, 1827: 331 (characters).

  • — Le Peletier and Serville, 1828: 475 (characters; classification; notes; discussion; list of species).

  • — Zetterstedt, 1828: 66 (characters).

  • — Curtis, 1829: 24 (catalog; Britain).

  • — Stephens, 1829a: 22 (catalog; Britain).

  • — Stephens, 1829: 275 (catalog; Britain).

  • — Latreille, 1829: 431, 434 (characters; notes; larval characters).

  • — C. Sahlberg, 1830: 307 (characters).

  • — Mannerheim, 1830: 7, 21 (characters).

  • — Mannerheim, 1831: 421 (key; description).

  • — Stephens, 1832: 200, 204 (characters).

  • — Griffith and Pidgeon, 1832: 291 (characters; notes).

  • — Dejean, 1833: 61 (list of species).

  • — Gistel, 1834: 8 (list of species).

  • — Lacordaire, 1835: 385 (characters).

  • — Westwood, 1838a: 15 (characters; type species).

  • — Nordmann, 1837: 27.

  • — Erichson, 1839a: 430 (characters).

  • — Erichson, 1839b: 345 (characters).

  • — Stephens, 1839: 384 (characters).

  • — Laporte, 1840: 175 (characters).

  • — Solier, 1849: 312 (characters).

  • — Redtenbacher, 1849: 694 (characters).

  • — Schaum, 1852: 25 (list of species; Europe).

  • — Lacordaire, 1854: 76 (characters; notes; list of species).

  • — Wollaston, 1854: 578 (characters).

  • — Fairmaire and Laboulbène, 1856: 504 (characters).

  • — Jacquelin du Val, 1856: 33, 35 (characters).

  • — Giebel, 1856: 71 (fossils).

  • — Gistel, 1856: 388 (list of species).

  • — Kraatz, 1857d: 536 (characters).

  • — Redtenbacher, 1857: 183, 985 (characters; key to Austrian species).

  • — Thomson, 1858: 29 (characters).

  • — G. Waterhouse, 1858: 23 (catalog of British species).

  • — Thomson, 1859: 23 (characters; type species: erythropterus).

  • — Schaum, 1859: 26 (catalog; European species).

  • — Thomson, 1860: 144 (characters).

  • — LeConte, 1861a: 65 (characters).

  • — Gredler, 1863: 101 (list of species from Tirol).

  • — LeConte, 1863: 23 (list of North American species).

  • — Heyden and Heyden, 1866: 136 (characters of fossil larva; Prussia).

  • — Wencker and Silbermann, 1866: 28 (list of species; collecting notes; France).

  • — Gemminger and Harold, 1868: 577 (catalog).

  • — Jacquelin du Val, 1868: 66 (catalog; European species).

  • — Sharp, 1871d: 11 (list of British species).

  • — Fauvel, 1874: 172 [= 1874a: 398] (characters; larval characters; key to species).

  • — Redtenbacher, 1874: 203 (characters; key to Austrian species).

  • — Oustalet, 1874: 153 (characters; description of fossil species).

  • — Fauvel, 1875a: XXVI [= 1875b: 228] (catalog).

  • — Mulsant and Rey, 1876: 249 [= 1877: 105] (characters; key to French species).

  • — Sharp, 1876: 150 (notes).

  • — Provancher, 1877: 246, 728 (characters; key to species).

  • — Horn, 1879: 186 (characters; notes; key to North American species).

  • — Heyden, 1880: 72 (list of species of Siberian region).

  • — Duvivier, 1883: 142 (catalog).

  • — Sharp, 1884: 371 (notes; Mexican and Central American species).

  • — Lynch, 1884: 135 (characters).

  • — Fowler, 1888: 249 (notes; key to British species).

  • — Seidlitz, 1891: 394 (key to some European species).

  • — Seidlitz, 1891a: 429 (key to some European species).

  • — Heyden, Reitter, and Weise, 1891: 101 (list of species of Europe and Caucasus).

  • — Heyden, 1893: 41 (list of species of Siberian region).

  • — Ganglbauer, 1895: 419 (adult and larval characters; key to middle European species).

  • — Heyden, Reitter, and Weise, 1906: 169 (list of species of Europe and Caucasus).

  • — Reitter, 1909: 116, 117 (characters; key to species).

  • — Petri, 1912: 74 (list of species of Siebenbürgen).

  • — Bernhauer and Schubert, 1914: 375 (world catalog; 260 species).

  • — Johansen, 1914: 376 (characters; key to Danish genera).

  • — Leng, 1920: 108 (catalog of North American species).

  • — Winkler, 1925: 381 (catalog for Palaearctic region).

  • — Cameron, 1925: 68 (catalog of Indian species).

  • — Porta, 1926: 94 (used in broad sense; characters; key to species of Italy).

  • — Portevin, 1929: 348 (characters; key to French species).

  • — Roubal, 1930: 379 (catalog; Slovakia).

  • — Cameron, 1932: 179 (characters; key to subgenera and species of British India).

  • — Joy, 1932: 123 (key to British species).

  • — Scheerpeltz, 1933: 1380 (world catalog supplement).

  • — Chapman, 1934 (notes on classification and distribution of some species).

  • — Normand, 1935: 372 (list of species of Tunisia).

  • — Voris, 1939a: 294 (larval and pupal characters).

  • — Paulian, 1941: 236 (larval characters).

  • — Blackwelder, 1943: 443 (characters; type species: maxillosus; key to species of West Indies).

  • — Blackwelder, 1944: 138 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Kloet and Hincks, 1945: 166 (list of British species).

  • — Tottenham, 1949: 373, 420 (type species: erythropterus Linné; notes and discussion of problems with the type species; [Note: These problems were resolved by the Commission in Opinion 546]; checklist of British species).

  • — Blackwelder, 1952: 357 (type species: maxillosus).

  • — Jarrige, 1952a: 133 (genus must be dismembered).

  • — V. Hansen, 1952: 92 (characters; key to Danish species).

  • — Coiffait, 1954: 167 (key to species of Canary Islands).

  • — Coiffait, 1956e: 178–182, 185, 195 (revision of generic concepts and elevation of some subgenera; history of usage; structure; characters in key).

  • — Tottenham, 1957a: 87 (proposal to place name on Official List of Generic Names in Zoology with Staphylinus erythropterus fixed as type species).

  • — Kocher, 1958: 152 (checklist of species; Morocco).

  • — Smetana, 1958: 260, 274 (characters; type species: erythropterus; key to Czechoslovakian species).

  • — Ferreira, 1962a: 18 (list of Portuguese species).

  • — Székessy, 1963: 53 (characters).

  • — Palm, 1963b: 107 (characters; key to species of Sweden).

  • — Lohse, 1964: 195 (characters; key to central European species).

  • — Moore, 1965: 98, 101 (characters; notes).

  • — Horion, 1965: 197 (note; list of European species).

  • — Pototskaia, 1967: 67 (Staphylinus; larval characters).

  • — Pototskaia, 1967: 64 (larval characters).

  • — Scheerpeltz, 1968a: 79 (catalog of Austrian species).

  • — Bolov, 1969a: 328 (collecting notes for some species; Russia).

  • — Kasule, 1970: 64, 70 (larval characters).

  • — Coiffait, 1971c (notes on name of genus).

  • — Shibata, 1973c: 63 (checklist of species of Taiwan).

  • — Tikhomirova, 1973a: 187 (checklist of species of USSR).

  • — Coiffait, 1974: 399 (adult and larval characters; key to species of western Palaearctic region).

  • — Moore and Legner, 1975: 36 (catalog; North America north of Mexico).

  • — Pope, 1977: 30 (list of British species).

  • — Topp, 1978: 311 (larval characters).

  • — Frank, 1979a (some larval characters and their use in classification).

  • — Moore and Legner, 1979: 30 (characters; notes).

  • — Muona, 1979: 15 (list of Scandinavian species).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 97 (catalog; Poland).

  • — Uhlig and Vogler, 1981: 119 (list of some species collected in Germany; habitat and collecting notes).

  • — Spahr, 1981: 96 (references to specimens from amber and copal).

  • — Tóth, 1984: 19 (characters; habitus; key to species of Hungary).

  • — Shibata, 1984: 81 (checklist of species of Japan).

  • — Yuh, Paik, Kwon, and Lee, 1985: 244 (checklist of species of Korea).

  • — Muona and Viramo, 1986: 13 (list of species of northeastern Finland).

  • — Segers, 1986: 47 (checklist of species; Belgium).

  • — Dettner, 1987: 30 (defensive glands).

  • — Drugmand 1987b (comments on distribution of Belgian species).

  • — Angelini, 1991: 194 (list of species from high elevations of Calabria, Italy).

  • — Campbell and Davies, 1991: 120 (checklist of species; Canada).

  • — Silfverberg, 1992: 17 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Siitonen, 1993: 230 (list of Finnish species).

  • — Welch, 1993: 229 (ovariole structure and development).

  • — Smetana, 1993b: 49 (list of species of Czech Republic and Slovakia).

  • — Ciceroni and Zanetti, 1995: 30 (list of species of Italy).

  • — M. Hansen, 1996: 106 (list of Danish species).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 244 (additions to list of Danish species).

  • — Downie and Arnett, 1996: 384 (characters; key to species of Northeastern North America).

  • — Beutel and Molenda, 1997: 48 (larval morphology and phylogeny).

  • — Smetana and Davies, 2000: 18, 20, 25, 41 (key to genera and subgenera associated with Staphylinus sensu lato; characters; checklist of species of north temperate region).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 394 (1 Nearctic species; notes; characters in key).

Ouchemus Gozis, 1886: 14 (species included: erythropterus, caesareus). Type species: Staphylinus erythropterus Linné, fixed by original designation.

  • — Blackwelder, 1943: 443 (synonym of Staphylinus; type species: erythropterus).

  • — Tottenham, 1949: 374 (synonym of Staphylinus; type species: erythropterus).

  • — Blackwelder, 1952: 280 (type species: erythropterus).

  • — Smetana, 1958: 260 (synonym of Staphylinus).

  • — Ádám, 1987: 148 (cited as valid genus; list of three species with collecting notes; Hungary).

Poikilodracus Scheerpeltz, 1974: 168 (subgenus of Staphylinus; species included: brincki, caffer, afer). Type species: Staphylinus (Poikilodracus) brincki Scheerpeltz, fixed by original designation.

abbruzzensis G. Müller, 1926, see: Ocypus.

acupunctipennis Bernhauer, 1907, see: Platydracus.

addendus Lindberg, 1953, see: Ocypus.

aegyptiacus C. Koch, 1936, see: Tasgius.

aeneicollis Gyllenhal, 1827, see: Ocypus.

aeneicollis Bernhauer, 1911, see: Platydracus.

aeneipennis Cameron, 1930, see: Platydracus.

aeneoacreus Cameron, 1930, see: Platydracus.

aeneocephalus De Geer, 1774, see: Ocypus.

aeneoniger Bernhauer, 1933, see: Platydracus.

aeneus De Geer, 1774, see: Philonthus.

aeneus Rossi, 1790, see: Philonthus.

aeneus Olivier, 1795, see: Taenodema.

aericeps Stephens, 1832, see: Platydracus.

aethiops Waltl, 1835, see: Ocypus.

afer Erichson, 1839b: 389 (Staphylinus; Type locality: Promont. Bonae Spei).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • — Scheerpeltz, 1974: 170 (Staphylinus; subgenus Poikilodracus; South Africa).

  • Distribution: South Africa.

affinis Paykull, 1789, see: Atrecus.

affinis Solsky, 1868a: 126 (Staphylinus; [preoccupied]; Type locality: Mexique).

  • — Sharp, 1884: 379 (Staphylinus; notes; Mexico; Guatemala).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • — Herman, 2001: 50 (Staphylinus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: Mexico, Guatemala.

africanus Cameron, 1942b: 329 (Staphylinus; subgenus Abemus; Type locality: Chyulu Hills, altitude 5600 feet).

  • Distribution: Kenya.

agarici O. Müller, 1776: 97 (Staphylinus; [Note: Nomen dubium]; Type locality: Dania).

  • — Gmelin, 1790: 2033 (Staphylinus; characters; Denmark).

  • — Turton, 1802: 517 (Staphylinus; characters; Denmark).

  • Distribution: Denmark.

agilis Gravenhorst, 1806, see: Philonthus.

alatus Gmelin, 1790: 2037 (Staphylinus; [Note: Nomen dubium]; Type locality: Europa).

  • — Zschach, 1788: 44 (Staphylinus; species number 992; characters stated but not named and referred to as alatus by Gmelin).

  • — Turton, 1802: 520 (Staphylinus; characters; Europe).

  • Distribution: Europe.

albanicus G. Müller, 1943, see: Ocypus.

albanicus Schatzmayr, 1943, see: Ocypus.

albipes Gravenhorst, 1802, see: Philonthus.

alfierii Bernhauer, 1925, see: Ocypus.

algiricus G. Müller, 1932, see: Ocypus.

alluaudi Fauvel, 1905d: 196 (Staphylinus; Type locality: Kilimandjaro, zone inferieure).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • Distribution: Tanzania.

almorensis Cameron, 1932, see: Ocypus.

alpinus Paykull, 1790, see: Anthophagus.

alpinus Fabricius, 1793, see: Anthophagus.

alpinus Gravenhorst, 1846, see: Ocypus.

altaiensis G. Müller, 1926, see: Ocypus.

alternans Gravenhorst, 1802, see: Atrecus.

amazonicus Sharp, 1876: 154 (Staphylinus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • Distribution: Brazil.

amiculus G. Müller, 1925, see: Tasgius.

amoenus Olivier, 1795, see: Philonthus.

amoenus Reitter, 1909, see: Tasgius.

analis Fabricius, 1787 [Note: This name has been listed in two genera Bolitobius and Quedius; Bolitobius analis (Fabricius) has been attributed erroneously to Paykull; this issue has been discussed and resolved by Schülke, 1999d: 978], see: Quedius.

analis W. J. MacLeay, 1873, see: Quedius.

anceps Runde, 1835: 8 (Staphylinus; [Note: Nomen dubium]; Type locality: Halae).

  • Distribution: Germany.

anceps Erichson, 1839, see: Oligotergus.

angularis Paykull, 1800, see: Carphacis.

angustatus Schrank, 1781: 233 (Staphylinus; [Note: Nomen dubium]; Type locality: Austria).

  • Distribution: Austria.

angustatus Geoffroy, 1785, see: Rugilus.

angustatus Paykull, 1789, see: Astenus.

angustatus Solier, 1849, see: Cheilocolpus.

anonymus Sulzer, 1761, see: Creophilus.

angusticeps Sharp, 1884: 376 (Staphylinus; Type locality: Mexico, Puebla and Toluca, Jalapa).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • Distribution: Mexico.

antennalis Cameron, 1932: 208 (Staphylinus; subgenus Tasgius; Type locality: Manipur).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; catalog).

  • Distribution: India.

anthrax Gravenhorst, 1802, see: Philonthus.

antiquus Nordmann, 1837: 35 (Staphylinus; Type locality: Para).

  • — Erichson, 1839b: 373 (Staphylinus; characters; Colombia; Brazil).

  • — R. Sahlberg, 1847: 788 (Staphylinus; Brazil).

  • — P. Lucas, 1857: 50 (Staphylinus; Brazil).

  • — Solsky, 1872: 293 (Staphylinus; Colombia).

  • — Sharp, 1876: 154 (Staphylinus; Brazil; Nicaragua; Colombia).

  • — Sharp, 1884: 385 (Staphylinus; notes; Mexico; Guatemala; Honduras; Nicaragua; Panama; South America).

  • — Fauvel, 1891: 106 (Staphylinus; Venezuela).

  • — Fauvel, 1901: 85 (Staphylinus; Colombia; Mexico to Peru and Brazil).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • — Blackwelder, 1943: 446 (Staphylinus; characters; Cuba; Trinidad).

  • Distribution: Venezuela, Brazil, Peru, Colombia, Panama, Nicaragua, Honduras, Guatemala, Mexico, West Indies.

apicialis Say, 1830, see: Hesperus.

apicicornis Guérin-Méneville, 1830, see: Philonthus.

apicipennis Sharp, 1884: 381 (Staphylinus; Type locality: Panama, Bugaba 800 to 1500 feet).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • Distribution: Panama.

aprutianus G. Müller, 1926, see: Ocypus.

apterus Scheerpeltz, 1976, see: Ocypus.

aquisextanus† Oustalet, 1874: 160 (Staphylinus; Type locality: Aix en Provence).

  • — Scudder, 1891: 584 (Staphylinus; catalog; Ligurian; France).

  • — Handlirsch, 1907: 724 (Staphylinus; catalog; Oligocene; France).

  • Distribution: France.

arcticus Erichson, 1839, see: Creophilus.

arenarius Geoffroy, 1785, see: Platystethus.

arenarius Paykull, 1800, see: Bledius.

armatus Panzer, 1799, see: Bledius.

armeniacus Sharp, 1873, see: Platydracus.

arnicae Scopoli, 1763: 102 (Staphylinus; [Note: Nomen dubium]; Type locality: Carniolia).

  • — Goeze, 1777: 725 (Staphylinus; characters).

  • — Villers, 1789: 423 (Staphylinus; characters).

  • — Gmelin, 1790: 2032 (Staphylinus; page misnumbered as 1032; characters).

  • — Turton, 1802: 516 (Staphylinus; characters; Carniola).

  • Distribution: Austria.

arrosus Eppelsheim, 1890, see: Dinothenarus.

arrowi G. Müller, 1932, see: Tasgius.

asemus Kraatz, 1859, see: Platydracus.

assimilis Paykull, 1800, see: Olophrum.

associatus Bernhauer, 1937, see: Platydracus.

atavus† Heer, 1862: 48 (Staphylinus; Type locality: Oeningen).

  • Distribution: Germany].

  • — Scudder, 1891: 584 (Staphylinus; catalog; Tortonian; Baden).

  • — Handlirsch, 1907: 725 (Staphylinus; catalog; Miocene; Germany).

atavus† Oustalet, 1874, see: Ocypus.

ater Gravenhorst, 1802, see: Tasgius.

aterrimus Gravenhorst, 1802, see: Gabrius.

aterrimus Marsham, 1802, see: Philonthus.

atratus Gravenhorst, 1802, see: Philonthus.

atricapillus Fabricius, 1775, see: Lordithon.

atricillus Gravenhorst, 1806, see: Quedius.

atrocoerulescens Goeze, 1777, see: Ocypus.

atronitidus Reitter, 1909, see: Tasgius.

atrox Nordmann, 1837: 38 (Staphylinus; Type locality: Mexico).

  • — Erichson, 1839b: 383 (Staphylinus; synonym of buqueti).

  • — Sharp, 1884: 372 (Staphylinus; notes; Mexico).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • Distribution: Mexico.

attenuatus Gravenhorst, 1802, see: Quedius.

aureiventris G. Müller, 1926 [Note: Nomen nudum], see: Dinothenarus.

aureofasciatus Motschulsky, 1862, see: Platydracus.

aureovillosus G. Müller, 1932, see: Platydracus.

aureus Fabricius, 1787, see: Rhyncocheilus.

aurichalceus Cameron, 1941, see: Platydracus.

auricomus Brullé, 1842, see: Glenus.

auricomus Cameron, 1929: 65 (Staphylinus; [preoccupied]; Type locality: Not cited, presumably Belgian Congo).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • — Herman, 2001: 50 (Staphylinus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: Congo.

auricomus Lindberg, 1953, see: Ocypus.

aurifluus Erichson, 1839b: 387 (Staphylinus; Type locality: Brasilia).

  • — Blanchard, 1842: 80 (Staphylinus; Brazil).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • Distribution: Brazil.

auripennis Kraatz, 1859, see: Platydracus.

auroaeneus Cameron, 1938, see: Platydracus.

auroguttatus Cameron, 1932, see: Ocypus.

auronotatus Fauvel, 1895, see: Platydracus.

auropubescens Cameron, 1930a: 156 (Staphylinus; Type locality: Malay Peninsula. Selangor; Bukit Kutu. Kedah; Catchment area near Jitra).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • Distribution: Malaysia.

aurosericans Fairmaire, 1891, see: Platydracus.

austriacus Schrank, 1781, see: Paederus.

azoricus Méquignon, 1942, see: Ocypus.

azurescens Mannerheim, 1830, see: Ocypus.

azureus Gebler, 1829, see: Platydracus.

baderlei Scheerpeltz, 1966, see: Dinothenarus.

badipes LeConte, 1863, see: Dinothenarus.

badius Mannerheim, 1830, see: Platydracus.

bakeri Bernhauer, 1915, see: Platydracus.

balcanicus G. Müller, 1923, see: Ocypus.

balearicus G. Müller, 1926, see: Ocypus.

balteatus De Geer, 1774, see: Creophilus.

baltimorensis Gravenhorst, 1802, see: Hesperus.

banghaasi Bernhauer, 1915d: 149 (Staphylinus; Type locality: Senegal).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • Distribution: Senegal.

banghaasi G. Müller, 1932, see: Platydracus.

barbarus G. Müller, 1926, see: Ocypus.

basicornis Bernhauer, 1932, see: Platydracus.

basicornis Fauvel, 1895, see: Platydracus.

batychrus Gyllenhal, 1827, see: Leptacinus.

beckeri Bernhauer, 1933, see: Ascialinus.

becquarti Bernhauer, 1938, see: Platydracus.

belti Sharp, 1884: 387 (Staphylinus; Type locality: Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • Distribution: Nicaragua.

bengalensis Bernhauer, 1914, see: Platydracus.

bernhaueri G. Müller, 1925, see: Ocypus.

bicinctus Rossi, 1792, see: Platydracus.

bicinctus Mannerheim, 1843, see: Creophilus.

biclavatus Gmelin, 1790: 2036 (Staphylinus; [Note: Nomen dubium]; Type locality: Europa).

  • — Zschach, 1788: 44 (Staphylinus; species number 978; characters stated but not named and referred to as biclavatus by Gmelin).

  • — Turton, 1802: 519 (Staphylinus; characters; Europe).

  • Distribution: Europe.

bicolor Paykull, 1789, see: Lesteva.

bicolor Gmelin, 1790: 2027 (Staphylinus; [preoccupied]; Type locality: Europa).

  • — Zschach, 1788: 45 (Staphylinus; species number 997; characters stated but not named and referred to as bicolor by Gmelin).

  • — Herman, 2001: 50 (Staphylinus; nomen dubium; although a junior primary homonym, it is suggested that the required replacement be ignored).

  • Distribution: Europe.

bicolor Laporte, 1835, see: Xenopygus.

bicolor G. Müller, 1943, see: Ocypus.

bicolor Cameron, 1944, see: Tasgius.

bicornis Block, 1799, see: Anthophagus.

biguttatus Linné, 1758, see: Stenus.

biguttatus Bernhauer, 1937, see: Platydracus.

biharicus G. Müller, 1926, see: Ocypus.

bimaculatus Schrank, 1798, see: Lordithon.

bimaculatus Gravenhorst, 1802, see: Philonthus.

bimaculatus Marsham, 1802, see: Philonthus.

bimaculatus Cameron, 1932, see: Ocypus.

binotatus Gravenhorst, 1802, see: Heterothops.

binotatus Gravenhorst, 1806, see: Philonthus.

biplagiatus Perty, 1830, see: Glenus.

bipunctatus Olivier, 1795 [Note: This name was cited by Bernhauer and Schubert (1926: 784, 793) as a synonym of two different species], see: Aleochara.

bipunctatus Turton, 1802, see: Tachinus.

bipustulatus Linné, 1760, see: Aleochara.

birmanus Fauvel, 1895, see: Platydracus.

biseriatus Sharp, 1884, see: Platydracus.

bisetosa G. Müller, 1943, see: Ocypus.

bisulcatus Solier, 1849, see: Cafius.

blandus Gravenhorst, 1806, see: Bisnius.

blattinus Schrank, 1781, see: Tachyporus.

bocandei Fagel, 1951, see: Platydracus.

bodemeyeri Bernhauer, 1913, see: Dinothenarus.

bodongi Bernhauer, 1906, see: Platydracus.

boleti Linné, 1758, see: Gyrophaena.

bolivianus Bernhauer, 1908d: 328 (Staphylinus; Type locality: Bolivien: Coroico, 1800 m).

  • — Bernhauer and Schubert, 1914: 375 (Staphylinus; catalog).

  • Distribution: Bolivia.

bombilius De Geer, 1774, see: Emus.

bomfordi Eppelsheim, 1895, see: Dinothenarus.

bonnairei Reitter, 1909 [Note: Nomen nudum], see: Tasgius.

boops Gravenhorst, 1802, see: Quedius.

borealis Paykull, 1792, see: Olophrum.

bosphoranus G. Müller, 1926, see: Tasgius.

brachypterus Geoffroy, 1785, see: Aleochara.

brachypterus Marsham, 1802, see: Aploderus.

brachypterus Brullé, 1839, see: Protogoerius.

brachypterus Kraatz, 1859, see: Platydracus.

brasiliensis R. Sahlberg, 1847, see: Gastrisus.

brassicae Scopoli, 1763, see: Deleaster.

bredoi Fagel, 1950, see: Platydracus.

brevicollis Paykull, 1800, see: Gymnusa.

brevicornis Gravenhorst, 1802, see: Philonthus.

brevipennis Heer, 1839, see: Ocypus.

brevipes Gmelin, 1790: 2036 (Staphylinus; [Note: Nomen dubium]; Type locality: Europa).

  • — Zschach, 1788: 44 (Staphylinus; species number 981; characters stated but not named and referred to as brevipes by Gmelin).

  • — Turton, 1802: 519 (Staphylinus; characters; Europe).

  • Distribution: Europe.

brincki Scheerpeltz, 1974: 168 (Staphylinus; subgenus Poikilodracus; Type locality: Basutoland: Teyateyaneng, 18 miles NE Maseru).

  • Distribution: Lesotho.

bruchi Bernhauer, 1934, see: Platydracus.

brunneimaculatus Tottenham, 1940 [Note: Nomen nudum], see: dimidiaticornis Gemminger, 1851.

brunneopubens G. Müller, 1926, see: Ocypus.

brunneus Paykull, 1789, see: Phyllodrepa.

brunneus Gravenhorst, 1802, see: Philonthus.

brunnipennis Fabricius, 1801, see: Bledius.

brunnipes Fabricius, 1781, see: Ocypus.

bryanti Cameron, 1918, see: Platydracus.

bucharicus Bernhauer, 1912, see: Tasgius.

buphthalmus Schrank, 1776, see: Stenus.

buquetii Laporte, 1835: 113 (Staphylinus; Type locality: Brésil).

  • — Erichson, 1839b: 383 (Staphylinus; characters; Brazil).

  • — P. Lucas, 1857: 50 (Staphylinus; Brazil).

  • — Solsky, 1868a: 127 (Staphylinus; Mexico).

  • — Solsky, 1875: 15 (Staphylinus; Peru).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog, Argentina).

  • Distribution: Mexico, Peru, Brazil, Argentina.

burgeoni Bernhauer, 1932b: 155 (Staphylinus; Type locality: Haut-Uele: Abimva).

  • — Scheerpeltz, 1933: 1382 (Staphylinus; catalog).

  • Distribution: Congo.

caeruleipennis Mannerheim, 1830, see: Philonthus.

caeruleoides Scheerpeltz, 1976a: 134 (Staphylinus; subgenus Goerius; Type locality: Nepal: Ting-Sang-La, 3800 m).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: Nepal.

caerulescens Lacordaire, 1835, see: Philonthus.

caeruleus Cameron, 1932, see: Ocypus.

caesareus Cederhjelm, 1798: 335 (Staphylinus; Type locality: Praedium imperialis Pawlowska [near St. Petersburg]).

  • — Fabricius, 1775: 265 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 376, cited erythropterus Fabricius as a synonym of caesareus, but Fabricius did not describe the species as new, he attributed it to Linné]; characters).

  • — Panzer, 1795: 353 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1916: 376, cited aeneocephalus Panzer as a synonym of caesareus; Panzer did not describe the species as new, he attributed it to Fabricius who attributed it to De Geer]; characters; Germany).

  • — Panzer, 1796c: 5 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 376, cited aeneocephalus Panzer as a synonym of caesareus; Panzer did not describe the species as new, he attributed it to De Geer]; characters; illustration).

  • — Panzer, 1808: 5 (Staphylinus; [Note: Misidentification: Panzer used chalcocephalus for the same species he cited in 1796c: 5 as aeneocephalus which be attributed to De Geer and which Bernhauer and Schubert, 1914: 376 cited as a synonym of caesareus. Panzer cited Fabricius as the author of chalcocephalus]; characters; illustrations).

  • — Erichson, 1839a: 435 (Staphylinus; characters; Germany).

  • — Erichson, 1839b: 378 (Staphylinus; characters; Europe).

  • — Heer, 1839: 250 (Staphylinus; characters; Switzerland).

  • — P. Lucas, 1846: 109 (Staphylinus; notes; Algeria).

  • — Jacobsen, 1849: 683 (Staphylinus; characters; Denmark).

  • — Hochhuth, 1849: 116 (Staphylinus; characters).

  • — Redtenbacher, 1849: 696 (Staphylinus; characters; Austria).

  • — Kiesenwetter, 1851a: 413 (Staphylinus; Catalonia).

  • — Küster, 1853: 49 (Staphylinus; characters; Europe).

  • — Hardy, 1851: 22 (Staphylinus; notes; England).

  • — Fairmaire and Laboulbène, 1856: 507 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 548 (Staphylinus; characters; Germany).

  • — Redtenbacher, 1857: 185 (Staphylinus; characters; Austria).

  • — Peyron, 1858: 422 (Staphylinus; Turkey).

  • — Thomson, 1860: 146 (Staphylinus; characters; Scandinavia).

  • — Hochhuth, 1862: 49 (Staphylinus; notes; Russia).

  • — Fauvel, 1865a: 294 [= 1865b: 44] (Staphylinus; habitat; France).

  • — Fauvel, 1874: 182 [= 1874a: 408] (Staphylinus; characters; France; Cyprus; Turkey; Caucasus; Transcaucasia; Iran).

  • — Redtenbacher, 1874: 205 (Staphylinus; characters; Austria).

  • — Seidlitz, 1875: 268 (Staphylinus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 251 [= 1877: 107] (Staphylinus; characters; France).

  • — J. Sahlberg, 1876: 10 (Staphylinus; Finland).

  • — Horn, 1879: 191 (Staphylinus; characters; Canada).

  • — Gozis, 1886: 14 (Ouchemus).

  • — Fowler, 1888: 252 (Staphylinus; characters; Britain).

  • — Ganglbauer, 1895: 429 (Staphylinus; characters; Europe; Mediterranean region; North America).

  • — Dierckx, 1899: 311 (Staphylinus; pygidial glands).

  • — Bordas, 1899: 230 (Staphylinus; defensive glands).

  • — Sainte-Claire Deville, 1906: 99 (Staphylinus; Corsica).

  • — Reitter, 1909: 119 (Staphylinus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; catalog).

  • — Johansen, 1914: 380 (Staphylinus; subgenus Staphylinus; characters; Denmark).

  • — Šulc, 1922 (Staphylinus; osmeterium; anatomy; histology; physiology).

  • — G. Müller, 1926: 6 (Staphylinus; subgenus Staphylinus; characters; notes).

  • — Sainte-Claire Deville, 1927: 57, 59 (Staphylinus; characters; Russia, Finland, Sweden, Denmark, Germany, Austria, Hungary, Czechoslovakia, France, Switzerland, Italy).

  • — Portevin, 1929: 350 (Staphylinus; characters; France).

  • — G. Müller, 1932: 77 (Staphylinus; characters; notes).

  • — Scheerpeltz, 1933: 1383 (Staphylinus; catalog).

  • — Tottenham, 1940e: 129 (Staphylinus; collecting notes; Britain).

  • — Tottenham, 1949: 420 (Staphylinus; checklist; Britain).

  • — V. Hansen, 1952: 100 (Staphylinus; characters; Denmark).

  • — Coiffait, 1956e: 196, 197 (Staphylinus; characters).

  • — Smetana, 1958: 275 (Staphylinus; subgenus Staphylinus; characters; ecology; Czechoslovakia).

  • — Ferreira, 1962: 18 (Staphylinus; Portugal).

  • — Fagel, 1963: 430 (Staphylinus; Turkey).

  • — Palm, 1963b: 112 (Staphylinus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 195 (Staphylinus; characters; central Europe).

  • — Smetana, 1965c: 28 (Staphylinus; subgenus Staphylinus; Anatolia).

  • — Smetana, 1965d: 79 (Staphylinus; subgenus Staphylinus; Anatolia).

  • — Horion, 1965: 208 (Staphylinus; subgenus Staphylinus; Lebanon; Armenia; Turkey; Russia; Finland; Sweden; Norway; Britain; Denmark; Holland; Switzerland; France; Spain; Portugal; Italy; Dalmatia; Bosnia; Serbia; Bulgaria; Romania; Hungary; Czech Republic; Slovakia; Poland; Ukraine; Germany; Austria).

  • — Pototskaia, 1967: 67 (Staphylinus; subgenus Staphylinus; larval characters).

  • — Smetana, 1967i: 556 (Staphylinus; Turkey).

  • — Smetana, 1968a: 156 (Staphylinus; subgenus Staphylinus; Anatolia).

  • — Smetana, 1968b: 87 (Staphylinus; subgenus Staphylinus; Anatolia).

  • — Scheerpeltz, 1968a: 80 (Staphylinus; catalog; Austria).

  • — Bolov, 1969: 514 (Staphylinus; collecting notes; Russia).

  • — Bolov, 1969a: 328 (Staphylinus; collecting notes; Russia).

  • — Smetana, 1971c: 264 (Staphylinus; subgenus Staphylinus; Iran).

  • — Jarrige, 1971: 496 (Staphylinus; Iran).

  • — Coiffait, 1974: 404 (Staphylinus; characters; Europe; Britain; Scandinavia; Asia Minor; Lebanon).

  • — Shilov, 1975a: 376 [= 1975b: 99] (Staphylinus; Komi region of Russia).

  • — Pope, 1977: 30 (Staphylinus; Britain).

  • — Coiffait, 1978d: 164 (Staphylinus; Turkey).

  • — Muona, 1979: 15 (Staphylinus; Finland; Russia; Sweden; Norway; Denmark; Baltic States).

  • — Petrenko, 1980a: 60, 63 (Staphylinus; characters; Ukraine).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 97 (Staphylinus; catalog; Poland).

  • — Hinton, 1981: 662 (Staphylinus; characters of egg).

  • — Frank, 1982: 48 (Staphylinus; list of parasites).

  • — Bordoni, 1984: 339 (Staphylinus; Lebanon).

  • — Tóth, 1984: 21 (Staphylinus; characters; Hungary).

  • — Lucht, 1987: 107 (Staphylinus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Huth and Dettner, 1989: 154 (Staphylinus; defensive secretions).

  • — Gusarov, 1989: 12 (Staphylinus; habitat notes; Ukraine).

  • — Huth and Dettner, 1990 (Staphylinus; defensive chemicals).

  • — Pilon, 1991: 103 (Staphylinus; notes; Italy).

  • — Pilon and Zanetti, 1991: 67 (Staphylinus; Italy).

  • — Silfverberg, 1992: 17 (Staphylinus; Finland; Russia; Sweden; Norway; Denmark; Estonia; Latvia; Lithuania).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Staphylinus; Czech Republic; Slovakia).

  • — Allen, 1995: 101 (Staphylinus; notes; England).

  • — Hodge and Jones, 1995: 41 (Staphylinus; characters; Britain).

  • — Ciceroni and Zanetti, 1995: 30 (Staphylinus; Italy).

  • — Downie and Arnett, 1996: 385 (Staphylinus; characters; Canada; USA).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 244 (Staphylinus; Denmark).

  • — M. Hansen, 1996: 106 (Staphylinus; checklist; Denmark).

  • — Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 38 (Staphylinus; Latvia).

  • — Pilon, 1998: 74 (Staphylinus; notes; Italy).

  • — Staniec, 1999a: 56 (Staphylinus; characters of pupa; Poland).

  • — Smetana and Davies, 2000: 41 (Staphylinus).

  • Distribution: Algeria, Europe, Russia, Caucasus, Iran, Turkey, Lebanon.

primus Schaeffer, 1766: ii, 2 (Staphylinus; [Note: Nomen nudum, Article 11.4]; Type locality: Ratisbon).

  • — Turton, 1802: 510 (Staphylinus; characters; cited as equivalent to erythropterus).

  • — Erichson, 1839b: 378 (Staphylinus; synonym of caesareus).

  • — Kraatz, 1857d: 548 (Staphylinus; synonym of caesareus).

corporaali Sainte-Claire Deville, 1927: 58 (Staphylinus; variety of caesareus; Type locality: Dept. des Basses-Pyrenees: Val de Sare; Itxassou. Asturies: Puerto de Pajares).

  • — Scheerpeltz, 1933: 1383 (Staphylinus; variety of caesareus).

  • — Coiffait, 1956e: 196 (Staphylinus; aberration of caesareus; characters).

  • — Smetana, 1958: 275 (Staphylinus; aberration of caesareus).

  • — Smetana, 1967c: 197 (Staphylinus; subgenus Staphylinus; subspecies of caesareus; notes; Spain).

  • — Coiffait, 1974: 404 (Staphylinus; variety of caesareus; characters).

  • — Tóth, 1984: 21 (Staphylinus; variety of caesareus; characters).

  • — Smetana and Davies, 2000: 41 (Staphylinus; subspecies of caesareus).

caffer Boheman, 1848: 277 (Staphylinus; Type locality: Caffraria orientalis).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; catalog).

  • — Bernhauer and Chapman, 1939: 72 (Staphylinus; Kenya).

  • — Bernhauer, 1942c: 364 (Staphylinus; Cameroon).

  • — Scheerpeltz, 1974: 170 (Staphylinus; subgenus Poikilodracus; Bechuanaland; South Africa).

  • Distribution: South Africa, Botswana, Kenya, Cameroon.

calidus Erichson, 1839, see: Xanthopygus.

caliginosus Erichson, 1839, see: Platydracus.

callistus Hochhuth, 1849, see: Ontholestes.

calvus† Oustalet, 1874: 155 (Staphylinus; Type locality: Aix en Provence).

  • — Scudder, 1891: 584 (Staphylinus; catalog; Ligurian; France).

  • — Handlirsch, 1907: 724 (Staphylinus; catalog; Oligocene; France).

  • Distribution: France.

canaliculatus Fabricius, 1787, see: Astilbus.

cantabricus G. Müller, 1926, see: Ocypus.

cantharellus Linné, 1767: 685 (Staphylinus; [Note: Nomen dubium]; Type locality: Suecia).

  • — Goeze, 1777: 720 (Staphylinus; characters).

  • — Villers, 1789: 416 (Staphylinus; characters; Sweden).

  • — Gmelin, 1790: 2029 (Staphylinus; characters).

  • — Turton, 1802: 515 (Staphylinus; characters; Sweden).

  • — Westwood, 1845: 51 (Staphylinus; no specimen in Linnean cabinet; may belong in Malthinus).

  • Distribution: Sweden.

cantharophagus Fagel, 1950, see: Platydracus.

cantianus Stephens, 1832, see: Ocypus.

capensis Sachse, 1852, see: hottentottus Nordmann 1837.

capitatus J. Bland, 1864, see: Dinothenarus.

capucinus Gravenhorst, 1806, see: Quedius.

caraboides Linné, 1758, see: Anthophagus.

carbonarius Gravenhorst, 1802, see: Philonthus.

carbonatus LeConte, 1863, see: Dinothenarus.

carinatus Panzer, 1798, see: Anotylus.

cariniceps Bernhauer, 1934, see: Apostenolinus.

carinthiacus Laporte, 1840: 176 (Staphylinus; [Note: Nomen dubium]; Type locality: Paris).

  • Distribution: France.

caroli Jarrige, 1943, see: Ocypus.

caseyi Scheerpeltz, 1933, see: Platydracus.

caspius Bernhauer, 1906, see: Tasgius.

castaneus Nordmann, 1837, see: Platydracus.

castanopterus Gravenhorst, 1802, see: erythropterus Linné, 1758.

caucasicus G. Müller, 1926, see: Ocypus.

cebuensis Wendeler, 1923: 385 (Staphylinus; Type locality: Philippinen: Cebu).

  • — Scheerpeltz, 1933: 1392 (Staphylinus; catalog).

  • Distribution: Philippines.

celer Gravenhorst, 1806, see: Philonthus.

centralis Sharp, 1884, see: Platydracus.

cephalotes Gmelin, 1790: 2036 (Staphylinus; Type locality: Europa).

  • — Zschach, 1788: 44 (Staphylinus; species number 980; characters stated but not named and referred to as cephalotes by Gmelin).

  • — Turton, 1802: 519 (Staphylinus; characters; Europe).

  • — Herman, 2001: 43 (Staphylinus; Nomen dubium).

  • Distribution: Europe.

cephalotes Gravenhorst, 1802, see: Bisnius.

cerdo Gerstaecker, 1867: 27 (Staphylinus; Type locality: Aruscha).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; catalog).

  • Distribution: Tanzania.

cervinipennis Quedenfeldt, 1888: 159 (Staphylinus; Type locality: Not cited, but presumably Central Africa).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; catalog).

  • Distribution: Africa.

chalcescens Sharp, 1889, see: Platydracus.

chalceus Bernhauer, 1911: 87 (Staphylinus; Type locality: Sumatra: Soekaranda).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; catalog).

  • — Cameron, 1933d: 391 (Staphylinus; Sumatra).

  • — Cameron, 1937: 21 (Staphylinus; Java).

  • Distribution: Indonesia.

chalcocephalus Gmelin, 1790: 2034 (Staphylinus; [Note: Nomen nudum; Gmelin attributed this name to De Geer who did not describe it. Gmelin cited De Geer's species number 8, on page 22 of his 1774 work; the reference cited by Gmelin is Staphylinus aeneocephalus De Geer. It can be argued that the name chalcocephalus was a lapsus, an erroneous spelling of aeneocephalus, or that it was published as a synonym of chalcocephalus; in either case the species is unavailable. However, If Staphylinus chalcocephalus is regarded as available and thereby a senior homonym of the same name proposed by Fabricius, then Gmelin's use can be designated to be a nomen oblitum and the use by Fabricius, a nomen protectum (Articles 23.9.1, 23.9.2)]).

chalcocephalus Fabricius, 1801, see: Platydracus.

chalcopygus Hochhuth, 1849, see: Ontholestes.

chalybeipennis Ménétriés, 1832, see: Tasgius.

championi Sharp, 1884: 382 (Staphylinus; Type locality: Panama, Volcan de Chiriqui 4000 to 6000 feet).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; catalog).

  • Distribution: Panama.

chapmani Bernhauer, 1933, see: Platydracus.

chenui Perroud and Montrousier, 1864, see: Diplostictus.

chilensis Solier, 1849, see: Bisnius.

chinensis Bernhauer, 1914, see: Platydracus.

chiriquensis Sharp, 1884: 388 (Staphylinus; Type locality: Panama, Volcan de Chiriqui 2000 to 3000 feet).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; catalog).

  • Distribution: Panama.

chloropterus Panzer, 1796, see: Abemus.

chloropterus Solier, 1849, see: Bisnius.

chrysaster Erichson, 1839b: 393 (Staphylinus; Type locality: Brazil).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • Distribution: Brazil.

chrysis Gravenhorst, 1806, see: Glenus.

chrysis Bernhauer, 1936a: 24 (Staphylinus; subgenus Abemus; [preoccupied]; Type locality: Lulua: Kapanga).

  • — Herman, 2001: 50 (Staphylinus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: Congo.

chrysocephalus Geoffroy, 1785, see: Dinothenarus.

chrysocomus Mannerheim, 1830, see: Dinothenarus.

chrysomelinus Linné, 1758, see: Tachyporus.

chrysopterus Brullé, 1842, see: Platydracus.

chrysopterus Gebler, 1832, see: Dinothenarus.

chrysopygus Nordmann, 1837, see: Xanthopygus.

chrysostigma Fauvel, 1895: 252 (Staphylinus; Type locality: Bírmanie, Teinzo).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • — Cameron, 1932: 198 (Staphylinus; characters; Burma).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; catalog).

  • Distribution: Myanmar.

chrysotrichopterus Scheerpeltz, 1933, see: Platydracus.

chrysurus Kirby, 1837, see: Ontholestes.

cicatricosus LeConte, 1863, see: Platydracus.

cicindeloides Schaller, 1783, see: Stenus.

cincticollis Cameron, 1951c: 404 (Staphylinus; subgenus Staphylinus; Type locality: N.W. Rhodesia: Solwezi district).

  • Distribution: Zambia.

cinctulus Gravenhorst, 1802, see: Philonthus.

cinctus Paykull, 1790, see: Quedius.

cinctus Marsham, 1802, see: Tachinus.

cinctus Latreille, 1804, see: Neobisnius.

cinctus Kollar and Redtenbacher, 1848, see: Trichocosmetes.

cinctus Solier, 1849, see: Blepharrhymenus.

cinerarius Erichson, 1839, see: Creophilus.

cinerascens Gravenhorst, 1802, see: Erichsonius.

cingulatus Gravenhorst, 1802, see: Ontholestes.

cingulus Comolli, 1837, see: Platydracus.

cinnamopterus Gravenhorst, 1802, see: Platydracus.

circumcinctus Bernhauer, 1914, see: Platydracus.

clavatus Gmelin, 1790: 2027 (Staphylinus; [Note: Nomen dubium]; Type locality: Europa).

  • — Zschach, 1788: 45 (Staphylinus; species number 1001; characters stated but not named and referred to as clavatus by Gmelin).

  • — Turton, 1802: 515 (Staphylinus; characters; Europe).

  • Distribution: Europe.

clavicornis Scopoli, 1763, see: Stenus.

clavicornis Fabricius, 1777, see: Stenus.

coquerelii Fauvel, 1866, see: Creophilus.

coenosus Gravenhorst, 1806, see: Philonthus.

coerulescens Geoffroy, 1785, see: Ocypus.

collaris Paykull, 1789, see: Zyras.

columbinus Erichson, 1839, see: Trigonopselaphus.

comes LeConte, 1863, see: Platydracus.

compressicollis Klug, 1832, see: Diatrechus.

compressus Geoffroy, 1785: 167 (Staphylinus; [Note: Nomen dubium]; Type locality: Paris).

  • — Villers, 1789: 421 (Staphylinus; characters).

  • Distribution: France.

compressus Marsham, 1802, see: Tasgius.

comptus Erichson, 1839, see: Plociopterus.

concinnus Gravenhorst, 1802, see: Philonthus.

concinnus Marsham, 1802, see: Xylodromus.

concolor Marsham, 1802, see: Velleius.

concolor Motschulsky, 1858, see: gravenhorsti Blackwelder, 1944.

conformis Lacordaire, 1835, see: Philonthus.

conicus Villers, 1789, see: Tachyporus.

connexus Say, 1830, see: Belonuchus.

consors Sharp, 1884: 382 (Staphylinus; Type locality: Panama, Volcan de Chiriqui 4000 to 6000 feet).

  • Distribution: Panama.

consularis Bernhauer, 1915, see: Platydracus.

contaminatus Gravenhorst, 1802, see: Philonthus.

contiguus Cameron, 1938, see: Platydracus.

cordilleranus Bernhauer, 1917, see: Platydracus.

coreanus G. Müller, 1925, see: Ocypus.

corporaali Sainte-Claire Deville, 1927, see: caesareus Cederhjelm, 1798.

corruscus Gravenhorst, 1802, see: Philonthus.

costaricensis Bernhauer, 1917, see: Platydracus.

crassicollis Gravenhorst, 1802, see: Ocypus.

crassicornis Fabricius, 1793, see: Atheta.

crassipes Gmelin, 1790: 2036 (Staphylinus; [Note: Nomen dubium]; Type locality: Europa).

  • — Zschach, 1788: 44 (Staphylinus; species number 986; characters stated but not named and referred to as crassipes by Gmelin).

  • — Turton, 1802: 520 (Staphylinus; cited as cassipes; characters; Europe).

  • Distribution: Europe.

crassus Mannerheim, 1846, see: Hadrotes.

crebrepunctatus Motschulsky, 1860, see: Platydracus.

crenatus Fabricius, 1793, see: Acidota.

cribratipennis Blanchard, 1842: 78 (Staphylinus; Type locality: Bolivia: Guarayos).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • Distribution: Bolivia.

cruentatus Gmelin, 1790, see: Philonthus.

cruentatus Marsham, 1802, see: Megalinus.

cruentus Olivier, 1795, see: Quedius.

cubae Jacquelin du Val, 1857, see: Platydracus.

cucullatus Steven, 1829: 28 (Staphylinus; [Note: Nomen dubium]; Type locality: not cited).

  • Distribution: ?.

cupreus Geoffroy, 1785, see: Ontholestes.

cupreus Rossi, 1790, see: Ocypus.

cupreicollis Nordmann, 1837: 34 (Staphylinus; Type locality: Brasilia: Minas-Geraes).

  • — Erichson, 1839b: 390 (Staphylinus; cited as cupricollis; characters; Brazil).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; cited as cupricollis; catalog).

  • Distribution: Brazil.

cupripennis Melsheimer, 1844, see: Platydracus.

cursor O. Müller, 1776, see: Omalium.

cursor Grimmer, 1841: 32 (Staphylinus; [preoccupied]; Type locality: Steiermark).

  • — Gemminger and Harold, 1868: 578 (Staphylinus; checklist; generic assignment uncertain; Styria).

  • — Herman, 2001: 50 (Staphylinus; nomen dubium; although a junior primary homonym, it is suggested that the required replacement be ignored).

  • Distribution: Austria.

curticollis Bernhauer, 1917, see: Platydracus.

curticornis Fauvel, 1895, see: Platydracus.

curtulus Goeze, 1777, see: Aleochara.

cyanelytrius Perty, 1830, see: Xanthopygus.

cyanescens Gerhardt, 1910, see: Ocypus.

cyaneus Paykull, 1789, see: Ocypus.

cyanicollis Laporte, 1835, see: Glenus.

cyanicornis Mannerheim, 1830, see: Philonthus.

cyanipennis Fabricius, 1793, see: Philonthus.

cyanipennis Runde, 1835: 7 (Staphylinus; [preoccupied]; Type locality: Halae).

  • — Herman, 2001: 50 (Staphylinus; nomen dubium; although a junior primary homonym, it is suggested that the required replacement be ignored).

  • Distribution: Germany.

cyanomelas Erichson, 1839b: 392 (Staphylinus; Type locality: Mexico).

  • — Solsky, 1868a: 127 (Staphylinus; Mexico).

  • — Sharp, 1884: 385 (Staphylinus; Mexico).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • Distribution: Mexico.

cylindricus Gmelin, 1790: 2035 (Staphylinus; [Note: Nomen dubium]; Type locality: Europa).

  • — Zschach, 1788: 44 (Staphylinus; species number 967; characters stated but not named and referred to as cylindricus by Gmelin).

  • — Turton, 1802: 519 (Staphylinus; characters; Europe).

  • Distribution: Europe.

daimio Sharp, 1889: 118 (Staphylinus; Type locality: Junsai; Sapporo).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; catalog).

  • — J. Li, 1993: 37 (Staphylinus; cited as dimio; China).

  • — Ito, 1996: 66 (Staphylinus; Japan).

  • — Smetana and Davies, 2000: 41 (Staphylinus).

  • Distribution: Japan, China.

daldorfii Turton, 1802, see: Tachyporus.

dauricus Mannerheim, 1830, see: Platydracus.

debilis Gravenhorst, 1806, see: Philonthus.

deceptivus Cuvier, 1833, see: Othius.

decipiens Kraatz, 1859, see: Platydracus.

decorus Gravenhorst, 1802, see: Philonthus.

dehradunensis Bernhauer, 1915, see: Dinothenarus.

demissus G. Müller, 1925, see: Platydracus.

denigrator Gravenhorst, 1806, see: Bisnius.

densissimus Bernhauer, 1933, see: Ocypus.

dentatus Marsham, 1802, see: Lathrobium.

depolii G. Müller, 1924, see: Ocypus.

depressus Paykull, 1789, see: Megarthrus.

diaphanus Marsham, 1802, see: Leptacinus.

dichrous Gmelin, 1790: 2035 (Staphylinus; [Note: Nomen dubium]; Type locality: Europa).

  • — Zschach, 1788: 44 (Staphylinus; species number 970; characters stated but not named and referred to as dichrous by Gmelin).

  • — Turton, 1802: 519 (Staphylinus; characters; Europe).

  • Distribution: Europe.

dilatatus Fabricius, 1787, see: Velleius.

dimidiaticornis Gemminger, 1851: 59 (Staphylinus; variety of erythropterus; Type locality: not cited, presumably München).

  • — G. Müller, 1943: 95 (Staphylinus; notes).

  • — Smetana, 1958: 276 (Staphylinus; subgenus Staphylinus; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1961d: 130 (Staphylinus; subgenus Staphylinus; Italy).

  • — Palm, 1963b: 112 (Staphylinus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 196 (Staphylinus; characters; central Europe).

  • — Scheerpeltz, 1965b: 404 (Staphylinus; Sicily).

  • — Horion, 1965: 210 (Staphylinus; subgenus Staphylinus; Russia; Finland; Sweden; Norway; Denmark; Britain; Holland; Belgium; Switzerland; France; Italy; Slovenia; Dalmatia; Greece; Czechoslovakia; Poland; Austria; Germany).

  • — Smetana, 1967i: 557 (Staphylinus; Bulgaria).

  • — Szujecki, 1966: 171 (Staphylinus; characters of egg; notes on natural history).

  • — Szujecki, 1966b: 236 (Staphylinus; larval characters).

  • — Szujecki, 1968a: 732 (Staphylinus; Poland).

  • — Coiffait, 1974: 405 (Staphylinus; characters; Europe).

  • — Pope, 1977: 30 (Staphylinus; Britain).

  • — Raichev, 1979: 83 (Staphylinus; Bulgaria).

  • — Petrenko, 1980a: 60, 63 (Staphylinus; subgenus Staphylinus; characters; Ukraine).

  • — Coiffait, 1982a: 233 (Staphylinus; Kashmir).

  • — Frank, 1982: 48 (Staphylinus; list of parasites).

  • — Tóth, 1984: 21 (Staphylinus; characters; Hungary).

  • — Drugmand, 1987a: 312 (Staphylinus; characters; Belgium).

  • — Lucht, 1987: 107 (Staphylinus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Huth and Dettner, 1989: 154 (Staphylinus; defensive secretions).

  • — Nowosad, 1990: 115 (Staphylinus; collected from mole nest; notes; Poland).

  • — Huth and Dettner, 1990 (Staphylinus; defensive chemicals).

  • — Pilon and Zanetti, 1991: 67 (Staphylinus; Italy).

  • — Drugmand, 1993: 170 (Staphylinus; notes; Luxembourg).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Staphylinus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 30 (Staphylinus; Italy).

  • — Owen, 1997a: 149 (Staphylinus; collecting sites; Ireland).

  • — Pilon, 1998: 75 (Staphylinus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 41 (Staphylinus).

  • Distribution: Europe, India.

parumtomentosus Stein, 1903: 128 (Staphylinus; Type locality: Graslitz im Erzgebirge).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; variety of caesareus).

  • — H. Bauer, 1914: 55 (Staphylinus; variety of caesarus; notes).

  • — G. Müller, 1926: 6 (Staphylinus; subgenus Staphylinus; valid species; characters; notes).

  • — G. Müller, 1926b: 42 (Staphylinus; subgenus Staphylinus; valid species; characters; notes).

  • — Sainte-Claire Deville, 1927: 58, 60 (Staphylinus; valid species; characters in key; Sweden, Denmark, Germany, Austria, Belgium, France, Corsica, Britain, Iberian Peninsula, Italy, Czechoslovakia, Switzerland).

  • — Portevin, 1929: 350 (Staphylinus; characters; France).

  • — Donisthorpe, 1930: 95 (Staphylinus; characters; Britain).

  • — G. Müller, 1932: 78 (Staphylinus; characters).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; catalog).

  • — Tottenham, 1940e: 129 (Staphylinus; collecting notes; Britain).

  • — G. Müller, 1943: 95 (Staphylinus; synonym of dimidiaticornis).

  • — V. Hansen, 1952: 100 (Staphylinus; characters; Denmark).

  • — Coiffait, 1956e: 196, 197 (Staphylinus; characters).

  • — Smetana, 1958: 276 (Staphylinus; synonym of dimidiaticornis).

  • — Lohse, 1964: 196 (Staphylinus; synonym of dimidiaticornis).

  • — Horion, 1965: 210 (Staphylinus; synonym of dimidiaticornis).

  • — Smetana, 1967i: 557 (Staphylinus; synonym of dimidiaticornis).

  • — Coiffait, 1974: 405 (Staphylinus; synonym of dimidiaticornis).

  • — Pope, 1977: 30 (Staphylinus; synonym of dimidiaticornis).

  • — Hinton, 1981: 662 (Staphylinus; characters of egg).

ernesti Bernhauer, 1913a: 223 (Staphylinus; variety of caesareus; Type locality: niederösterreichischen Waldviertel).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; variety of caesareus).

  • — H. Bauer, 1914: 55 (Staphylinus; variety of caesareus; notes).

  • — Scheerpeltz, 1933: 1383 (Staphylinus; aberration of caesareus).

  • — Smetana, 1958: 275 (Staphylinus; aberration of caesareus).

  • — Coiffait, 1974: 405 (Staphylinus; variety of dimidiaticornis; Austria).

brunneimaculatus Tottenham, 1940e: 130 (Staphylinus; aberration of parumtomentosus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Hartlebury).

  • — Smetana, 1958: 276 (Staphylinus; aberration of dimidiaticornis).

  • — Coiffait, 1974: 406 (Staphylinus; variety of dimidiaticornis; England; France).

  • — Pope, 1977: 30 (Staphylinus; synonym of dimidiaticornis).

dimidiatus C. Sahlberg, 1830, see: Philonthus.

dimidiatus Say, 1830, see: Philonthus.

dimidiatus Lacordaire, 1835, see: Philonthus.

dimidiatus Laporte, 1835: 115 (Staphylinus; [preoccupied]; Type locality: Cap de Bonne Espérance).

  • — Erichson, 1839b: 385 (Staphylinus; characters; South Africa).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • — Herman, 2001: 51 (Staphylinus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: South Africa.

velutinus Nordmann, 1837: 44 (Staphylinus; Type locality: Promontorium bonae spei).

  • — Erichson, 1839b: 385 (Staphylinus; synonym of dimidiatus).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; synonym of dimidiatus).

discoideus Gravenhorst, 1802, see: Philonthus.

discretus Sharp, 1884: 379 (Staphylinus; Type locality: Mexico, Las Vigas).

  • — Sharp, 1887: 786 (Staphylinus; Mexico).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • Distribution: Mexico.

dispar Paykull, 1789, see: Tachyporus.

dispersus Fauvel, 1907: 36 (Staphylinus; Type locality: Kilimandjaro: Kibonoto, 3000 m).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • — Bernhauer and Chapman, 1939: 73 (Staphylinus; Kilimanjaro).

  • Distribution: Tanzania.

distinctus Runde, 1835, see: Quedius.

divisus Marsham, 1802, see: Bledius.

dohertyi Cameron, 1932, see: Platydracus.

domicella Schrank, 1781: 235 (Staphylinus; [Note: Nomen dubium]; Type locality: Austria superiori).

  • — Herbst, 1784: 151 (Staphylinus; Austria).

  • — Villers, 1789: 423 (Staphylinus; characters).

  • — Gmelin, 1790: 2032 (Staphylinus; page misnumbered as 1032; characters).

  • — Turton, 1802: 516 (Staphylinus; characters; Austria).

  • Distribution: Austria.

doriae G. Müller, 1932, see: Tasgius.

drescheri Bernhauer, 1937, see: Platydracus.

dubius Gravenhorst, 1806, see: Philonthus.

dudgeoni Cameron, 1932, see: Platydracus.

dufouri Blackwelder, 1944: 139 (Staphylinus; replacement name from punctatissimus Motschulsky).

  • Distribution: Brazil.

punctatissimus Motschulsky, 1858: 670 (Trichoderma; [preoccupied]; Type locality: intérieru du Bresil).

  • — Fauvel, 1878f: 249 (Staphylinus).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • — Blackwelder, 1944: 139 (Staphylinus; synonym of dufouri).

duplicatus G. Müller, 1943, see: Ocypus.

dvoraki Roubal, 1940, see: Tasgius.

ebeninus Gravenhorst, 1802, see: Philonthus.

edentulus Block, 1799, see: Tasgius.

ejulans Tottenham, 1939, see: Platydracus.

elegans Olivier, 1795, see: Xantholinus.

elongatus Linné, 1767, see: Lathrobium.

elongatus Geoffroy, 1785, see: Gyrohypnus.

emarginatus Paykull, 1789, see: Atemeles.

emeritus Herman, 2001: 51 (Staphylinus; replacement name for priscus Sharp).

  • Distribution: Brazil.

priscus Sharp, 1876: 155 (Staphylinus; [preoccupied]; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • — Herman, 2001: 51 (Staphylinus; synonym of emeritus).

emmesus Gravenhorst, 1802, see: Neohypnus.

ensifer G. Müller, 1932, see: Tasgius.

ephippiatus Say, 1830, see: Belonuchus.

eppelsheimianus Jakobson, 1909, see: Tasgius.

erichsoni Boheman, 1848, see: Platydracus.

ernesti Bernhauer, 1913, see: dimidiaticornis Gemminger, 1851.

ertli Bernhauer, 1908c: 108 (Staphylinus; Type locality: Deutsch-Ostafrika: Uhehe, Luiba).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • Distribution: Tanzania.

erythrocephalus Fabricius, 1775, see: Creophilus.

erythrocephalus Motschulsky, 1859 [Note: Nomen nudum], see: Dinothenarus.

erythrocnemus Nordmann, 1837: 42 (Staphylinus; Type locality: Promontorium bonae spei).

  • — Erichson, 1839b: 385 (Staphylinus; characters; South Africa).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • Distribution: South Africa.

erythropennis Mannerheim, 1830, see: Platydracus.

erythropterus Linné, 1758: 422 (Staphylinus; [Note: Cited originally as erytropterus but in 1760 as erythropterus; the emendation from erytropterus to erythropterus was validated by action of Commission in Opinion 546 (ICZN, 1959)]; Type locality: Not cited).

  • — Linné, 1760: 231 (Staphylinus; characters; Sweden).

  • — Scopoli, 1763: 101 (Staphylinus; characters; Carniola).

  • — O. Müller, 1764: 23 (Staphylinus; cited as erytropterus; characters).

  • — Pontoppidans, 1765: 211 (Staphylinus; Denmark).

  • — Linné, 1767: 683 (Staphylinus; characters).

  • — Ström, 1767: 362 (Staphylinus; characters; Denmark).

  • — De Geer, 1774: 21 (Staphylinus; cited as erytropterus; redescription; notes).

  • — Füessly, 1775: 21 (Staphylinus; cited as erytropterus).

  • — O. Müller, 1776: 97 (Staphylinus; characters).

  • — Goeze, 1777: 716 (Staphylinus; cited erytropterus references).

  • — Schrank, 1781: 230 (Staphylinus; cited as erytropterus characters; Austria).

  • — Fabricius, 1781: 335 (Staphylinus; characters; Europe).

  • — Harrer, 1784: 250 (Staphylinus; characters).

  • — Herbst, 1784: 149 (Staphylinus; Germany).

  • — Fourcroy, 1785: 166 (Staphylinus; cited as erytropterus; characters; France).

  • — Fabricius, 1787: 220 (Staphylinus; cited as erythropterus; characters).

  • — Paykull, 1789: 7 (Staphylinus; redescription; notes; Sweden).

  • — Rossi, 1790: 247 (Staphylinus; characters; Italy).

  • — Villers, 1789: 411 (Staphylinus; cited as erytropterus; characters).

  • — Gmelin, 1790: 2027 (Staphylinus; characters).

  • — Fabricius, 1793: 522 (Staphylinus; characters).

  • — Olivier, 1795:(42): 12 (Staphylinus; characters).

  • — Panzer, 1795: 12, 353 (Staphylinus; characters; illustration).

  • — Schrank, 1798: 640 (Staphylinus; characters; notes).

  • — Cederhjelm, 1798: 117 (Staphylinus; characters).

  • — Paykull, 1800: 369 (Staphylinus; characters; Sweden).

  • — Fabricius, 1801: 593 (Staphylinus; characters).

  • — Turton, 1802: 510 (Staphylinus; characters; Europe).

  • — Gravenhorst, 1802: 10 (Staphylinus; characters).

  • — Marsham, 1802: 499 (Staphylinus; characters; Britain).

  • — Latreille, 1804: 297 (Staphylinus; characters).

  • — Gravenhorst, 1806: 116 (Staphylinus; characters).

  • — Latreille, 1806: 287 (Staphylinus; characters).

  • — Panzer, 1808: 4 (Staphylinus; characters; illustrations).

  • — Gyllenhal, 1810: 293 (Staphylinus; characters; Sweden).

  • — Leach, 1815: 91 (Staphylinus; characters; Europe).

  • — Samouelle, 1819: 171 (Staphylinus; characters; habitat).

  • — Leach, 1819: 172 (type species of Staphylinus).

  • — Zetterstedt, 1828: 68 (Staphylinus; characters; Lapland).

  • — Curtis, 1829: 24 (Staphylinus; catalog; Britain).

  • — Latreille, 1829: 434 (Staphylinus; characters).

  • — C. Sahlberg, 1830: 311 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 22 (Staphylinus; Sweden; Russia; France).

  • — Mannerheim, 1831: 436 (Staphylinus; Sweden; Russia; France).

  • — Stephens, 1832: 205 (Staphylinus; characters; England; Scotland).

  • — Ménétriés, 1832: 142 (Staphylinus; Caucasus-Persia region).

  • — Griffith and Pidgeon, 1832: 292 (Staphylinus; characters).

  • — Runde, 1835: 2 (Staphylinus; characters; Germany).

  • — Lacordaire, 1835: 365 (Emus; characters; France).

  • — Nordmann, 1837: 55 (Staphylinus; characters; Europe).

  • — Westwood, 1838a: 15 (Staphylinus; type species).

  • — Zetterstedt, 1838: 59 (Staphylinus; characters; habitat; Lapland).

  • — Curtis, 1839: pl. 758 (Staphylinus; characters; habitat; England).

  • — Erichson, 1839a: 434 (Staphylinus; characters; Germany).

  • — Heer, 1839: 251 (Staphylinus; characters; Switzerland).

  • — Erichson, 1839b: 377 (Staphylinus; characters; Europe).

  • — Stephens, 1839: 387 (Staphylinus; characters; England; Scotland).

  • — Laporte, 1840: 176 (Staphylinus; cited as erypthropterus; characters; France).

  • — Holme, 1842: 115, 116 (Staphylinus; notes).

  • — Dufour, 1843: 32 (Staphylinus; cited as erithropterus; characters; France).

  • — Westwood, 1845: 46 (Staphylinus; characters; notes).

  • — Kolenati, 1846: 17 (Staphylinus; Caucasus; Transcaucasus).

  • — Jacobsen, 1849: 682 (Staphylinus; characters; Denmark).

  • — Hochhuth, 1849: 115 (Staphylinus; characters; Lenkoran).

  • — Redtenbacher, 1849: 696 (Staphylinus; characters; Austria).

  • — Küster, 1853: 48 (Staphylinus; characters; Europe).

  • — Hardy, 1851: 22 (Staphylinus; notes; England).

  • — Fairmaire and Laboulbène, 1856: 507 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 547 (Staphylinus; characters; Germany).

  • — Redtenbacher, 1857: 185 (Staphylinus; characters; Austria).

  • — Thomson, 1859: 23 (type species of Staphylinus).

  • — Thomson, 1860: 145 (Staphylinus; characters; Scandinavia).

  • — Hochhuth, 1862: 49 (Staphylinus; Russia).

  • — Fauvel, 1874: 181 [= 1874a: 407] (Staphylinus; characters; France; Caucasus; Transcaucasia; Lake Baikal).

  • — Redtenbacher, 1874: 205 (Staphylinus; characters; Austria).

  • — Seidlitz, 1875: 268 (Staphylinus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 254 [= 1877: 110] (Staphylinus; characters; France).

  • — J. Sahlberg, 1876: 10 (Staphylinus; Finland).

  • — LeConte, 1878b: 598 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 378, cited caesareus LeConte as a synonym of erythropterus; LeConte did not describe the species as new, he attributed it to Cederhjelm, cited as Cederholm]; characters; Michigan).

  • — Horn, 1879: 190 (Staphylinus; LeConte's use of caesareus is a misidentification of erythropterus; characters; Michigan).

  • — J. Sahlberg, 1880: 68 (Staphylinus; Siberia).

  • — Gozis, 1886: 14 (type species of Ouchemus).

  • — Fowler, 1888: 252 (Staphylinus; characters; Britain).

  • — Ganglbauer, 1895: 429 (Staphylinus; characters; Europe; Siberia; Transcaucasian region; North America).

  • — Poppius, 1909: 28 (Staphylinus; Lena Valley, Siberia).

  • — Reitter, 1909: 119 (Staphylinus; characters; Germany).

  • — Blatchley, 1910: 376 (Staphylinus; characters; Michigan).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • — Johansen, 1914: 381 (Staphylinus; subgenus Staphylinus; characters; Denmark).

  • — G. Müller, 1926: 6 (Staphylinus; subgenus Staphylinus; characters).

  • — Portevin, 1929: 350 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; catalog).

  • — V. Hansen, 1952: 101 (Staphylinus; characters; Denmark).

  • — Coiffait, 1956e: 196, 197 (Staphylinus; characters).

  • — Tottenham, 1957a: 87, 88 (proposed emendation of original spelling, erytropterus, to erythropterus and fixation of emended name as type species of Staphylinus; proposal to place name on Official List of Specific Names in Zoology).

  • — Smetana, 1958: 278 (Staphylinus; subgenus Staphylinus; characters; ecology; Czechoslovakia).

  • — ICZN, 1959 (fixed as type species of Staphylinus; name placed on Official List of Specific Names in Zoology).

  • — Szujecki, 1960 (pupal characters).

  • — Palm, 1963b: 112 (Staphylinus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 195 (Staphylinus; characters; central Europe).

  • — Raianu, 1964: 103 (Staphylinus; Romania).

  • — Horion, 1965: 207 (Staphylinus; subgenus Staphylinus; widespread in northern Europe; Holland; Belgium; France; Switzerland; Italy; Bosnia Hercegovina; Romania; Hungary; Czechoslovakia; Poland; Russia; Ukraine; Spain; Austria; Germany).

  • — Smetana, 1968b: 88 (Staphylinus; subgenus Staphylinus; Anatolia).

  • — Szujecki, 1968a: 732 (Staphylinus; Poland).

  • — Szujecki, 1966: 170 (Staphylinus; characters of egg; notes on natural history).

  • — Szujecki, 1966b: 236 (Staphylinus; larval characters).

  • — Kasule, 1970: 70 (Staphylinus; larval characters).

  • — Coiffait, 1974: 402 (Staphylinus; characters; France; Pyrenees; Alps; central Europe; Britain; Scandinavia; Caucasus; Siberia).

  • — Shilov, 1975a: 376 [= 1975b: 99] (Staphylinus; Komi region of Russia).

  • — Pope, 1977: 30 (Staphylinus; Britain).

  • — Hinton, 1981: 662 (Staphylinus; characters of egg).

  • — Petrenko, 1980a: 60, 63 (Staphylinus; subgenus Staphylinus; characters; Ukraine).

  • — Frank, 1982: 48 (Staphylinus; list of parasites).

  • — Tóth, 1984: 20 (Staphylinus; characters; Hungary).

  • — Lucht, 1987: 107 (Staphylinus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Huth and Dettner, 1989: 154 (Staphylinus; defensive secretions).

  • — Huth and Dettner, 1990 (Staphylinus; defensive chemicals).

  • — Nowosad, 1990: 115 (Staphylinus; collected from mole nest; notes; Poland).

  • — Pilon, 1991: 103 (Staphylinus; notes; Italy).

  • — Pilon and Zanetti, 1991: 67 (Staphylinus; Italy).

  • — Welch, 1993: 229 (Staphylinus; ovariole number and ovary structure).

  • — J. Li, 1993: 38 (Staphylinus; cited as erythoropterus; China).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Staphylinus; Czech Republic; Slovakia).

  • — Dearborn and Donahue, 1994: 15 (Staphylinus; Maine).

  • — Ciceroni and Zanetti, 1995: 30 (Staphylinus; Italy).

  • — Staniec, 1996: 118 (Staphylinus; pupal characters).

  • — Downie and Arnett, 1996: 385 (Staphylinus; characters; USA).

  • — Sterrenburg and Schülke, 1997: 20 (Staphylinus; Netherlands).

  • — Pilon, 1998: 76 (Staphylinus; ecological notes; Italy).

  • — Staniec, 1999a: 56 (Staphylinus; characters of pupa; Poland).

  • — Smetana and Davies, 2000: 41 (Staphylinus).

  • Distribution: Europe, Russia, Azerbaijan, Turkey.

castanopterus Gravenhorst, 1802: 10 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).

  • — Latreille, 1804: 298 (Staphylinus; characters; notes; Germany; Italy).

  • — Gravenhorst, 1806: 116 (Staphylinus).

  • — Gyllenhal, 1810: 295 (Staphylinus; characters; Sweden).

  • — Zetterstedt, 1828: 68 (Staphylinus; characters; Lapland).

  • — Curtis, 1829: 24 (Staphylinus; catalog; Britain).

  • — C. Sahlberg, 1830: 312 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 22 (Staphylinus; Finland; Sweden).

  • — Mannerheim, 1831: 436 (Staphylinus; Finland; Sweden).

  • — Stephens, 1832: 206 (Staphylinus; characters; England; Scotland).

  • — Runde, 1835: 2 (Staphylinus; characters; Germany).

  • — Nordmann, 1837: 55 (Staphylinus; characters; Europe).

  • — Zetterstedt, 1838: 60 (Staphylinus; characters; habitat; Lapland).

  • — Curtis, 1839: pl. 758 (Staphylinus; England).

  • — Erichson, 1839a: 434 (Staphylinus; characters; Germany).

  • — Erichson, 1839b: 377 (Staphylinus; synonym of erythropterus).

  • — Stephens, 1839: 387 (Staphylinus; characters; England; Scotland).

  • — Holme, 1842: 116 (Staphylinus; notes).

  • — Küster, 1853: 48 (Staphylinus; synonym of erythropterus).

  • — Hardy, 1851: 22 (Staphylinus; synonym of erythropterus).

  • — Fairmaire and Laboulbène, 1856: 507 (Staphylinus; synonym of erythropterus).

  • — Kraatz, 1857d: 547 (Staphylinus; synonym of erythropterus).

  • — Thomson, 1860: 145 (Staphylinus; synonym of erythropterus).

  • — Ganglbauer, 1895: 429 (Staphylinus; synonym of erythropterus).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; synonym of erythropterus).

  • — Smetana, 1958: 278 (Staphylinus; synonym of erythropterus).

  • — Coiffait, 1974: 402 (Staphylinus; synonym of erythropterus).

springeri G. Müller, 1923: 136 (Staphylinus; subspecies of erythropterus; Type locality: Lubiana).

  • — G. Müller, 1926: 6 (Staphylinus; subgenus Staphylinus; subspecies of erythropterus; characters).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; subspecies of erythropterus).

  • — Coiffait, 1956e: 196 (Staphylinus; subspecies of erythropterus; characters).

  • — Smetana, 1958: 278 (Staphylinus; subspecies of erythropterus).

  • — Horion, 1965: 208 (Staphylinus; subspecies of erythropterus; Slovenia; Austria).

  • — Coiffait, 1974: 402 (Staphylinus; subspecies of erythropterus; characters; Slovenia; Croatia; Austria).

  • — Smetana and Davies, 2000: 41 (Staphylinus; subspecies of erythropterus).

erythropus Gmelin, 1790: 2035 (Staphylinus; [Note: Nomen dubium]; Type locality: Europa).

  • — Zschach, 1788: 44 (Staphylinus; species number 974; characters stated but not named and referred to as erythropus by Gmelin).

  • — Turton, 1802: 519 (Staphylinus; characters; Europe).

  • Distribution: Europe.

erythropus Paykull, 1800 [Note: This name was cited as a synonym of two species by Bernhauer and Schubert, 1914], see: Platydracus and Ocypus.

evansi Bernhauer, 1936a: 23 (Staphylinus; Type locality: Elisabethville).

  • Distribution: Congo.

excisus G. Müller, 1950, see: Ocypus.

extensus Sharp, 1884: 372 (Staphylinus; Type locality: Costa Rica, Cache; Panama, Volcan de Chiriqui 2000 to 3000 feet).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; catalog).

  • Distribution: Costa Rica, Panama.

exulans Erichson, 1839, see: Platydracus.

fabricii Turton, 1802, see: Lathrobium.

facialis Erichson, 1839, see: Nordus.

falcimaculatus Bernhauer, 1937, see: Platydracus.

fallaciosus G. Müller, 1926, see: Ocypus.

fasciatus Füessly, 1775, see: Creophilus.

fasciatus Marsham, 1802, see: Gyrophaena.

fassli Bernhauer, 1908d: 327 (Staphylinus; Type locality: Bolivien: Coroico, 1800 m).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; catalog).

  • Distribution: Bolivia.

faustus Erichson, 1839, see: Xanthopygus.

fauveli Péringuey, 1908: 296 (Staphylinus; Type locality: Southern Rhodesia, Salisbury; Umtali).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; catalog).

  • Distribution: Zimbabwe.

fauvelianus Fagel, 1958, see: Platydracus.

feae Bernhauer, 1915k: 304 (Staphylinus; subgenus Abemus; Type locality: Portugiesisch-Guinea: Rio Cassine).

  • — Scheerpeltz, 1933: 1384 (Staphylinus; catalog).

  • Distribution: Guinea-Bissau.

fellmani Zetterstedt, 1838, see: Quedius.

femoratus Fabricius, 1801, see: Platydracus.

fenyesi Bernhauer, 1917, see: Platydracus.

ferox Nordmann, 1837: 39 (Staphylinus; Type locality: Mexico).

  • — Erichson, 1839b: 384 (Staphylinus; characters; Mexico).

  • — Sharp, 1884: 372 (Staphylinus; Mexico; Panama).

  • — Bernhauer and Schubert, 1914: 378 (Staphylinus; catalog).

  • Distribution: Mexico, Panama.

ferrugineus Rossi, 1790, see: Megalinus.

fervidus Sharp, 1884, see: Platydracus.

festae G. Müller, 1925, see: Ocypus.

fetialis Erichson, 1839, see: Plociopterus.

fimetarius Gravenhorst, 1802, see: Bisnius.

flavescens Linné, 1758, see: Quedius.

flavipes Linné, 1758, see: Phloeostiba and Tachinus.

flavocephalus Goeze, 1777, see: Dinothenarus.

flavicornis Gmelin, 1790: 2036 (Staphylinus; [Note: Nomen dubium]; Type locality: Europa).

  • — Zschach, 1788: 44 (Staphylinus; species number 990; characters stated but not named and referred to as flavicornis by Gmelin).

  • — Turton, 1802: 520 (Staphylinus; characters; Europe).

  • — Ádám, 1996: 247 (Distichalius; valid species with Staphylinus cinctus Paykull as junior synonym; [Note: Until the identity of this name has been verified it is best that it remain in Staphylinus as a nomen dubium]).

  • Distribution: Europe.

flavipennis Hope, 1831: 32 (Staphylinus; [Note: Nomen dubium]; Type locality: Nepaul).

  • Distribution: Nepal.

flavomaculatus Goeze, 1777: 731 (Staphylinus; [Note: Nomen dubium]; Type locality: Not cited).

  • Distribution: ?.

flavomarginatus Goeze, 1777, see: Tachinus.

flavopilosus Cameron, 1932, see: Platydracus.

flavopterus Geoffroy, 1785, see: Quedius.

flavopunctatus Latreille, 1804, see: Platydracus.

flavus Gmelin, 1790: 2027 (Staphylinus; [Note: Nomen dubium]; Type locality: Europa).

  • — Zschach, 1788: 45 (Staphylinus; species number 1004; characters stated but not named and referred to as flavus by Gmelin).

  • — Turton, 1802: 515 (Staphylinus; characters; Europe).

  • Distribution: Europe.

floralis Paykull, 1789, see: Phyllodrepa.

floreus O. Müller, 1776: 99 (Staphylinus; Type locality: Dania).

  • — Gmelin, 1790: 2033 (Staphylinus; characters; Denmark).

  • — Turton, 1802: 517 (Staphylinus; characters; Denmark).

  • Distribution: Denmark.

fluviaticus Casey, 1915, see: mysticus Erichson, 1840.

fodiens Gravenhorst, 1806, see: Dinothenarus.

fodinarum Gmelin, 1790: 2032 (Staphylinus; replacement name for subterraneus Scopoli; [Note: Nomen dubium; Gmelin cited this name with characters and followed it by a reference to Scopoli, 1771: 109, where Sopoli used Staphylinus subterraneus Scopoli. Evidently Gmelin's name is a replacement name for the preoccupied subterraneus Scopoli. Also note that the page for S. fodinarum is misnumbered as 1032]).

  • — Turton, 1802: 517 (Staphylinus; characters; Germany).

  • Distribution: ?.

subterraneus Scopoli, 1771: 109 (Staphylinus; [Note: Nomen dubium; preoccupied]; Type locality: Not cited).

fokiensis Bernhauer, 1933, see: Dinothenarus.

formicarius O. Müller, 1776: 99 (Staphylinus; [Note: Nomen dubium]; Type locality: Dania).

  • — Gmelin, 1790: 2033 (Staphylinus; characters).

  • — Turton, 1802: 517 (Staphylinus; characters; Denmark).

  • Distribution: Denmark.

formosae Bernhauer, 1933, see: Platydracus.

formosus Gravenhorst, 1806, see: Belonuchus.

fossator Gravenhorst, 1802, see: Platydracus.

fossor Scopoli, 1771, see: Dinothenarus.

fossor Fabricius, 1793, see: Dinothenarus.

fracticornis O. Müller, 1776, see: Gyrohypnus.

fracticornis Paykull, 1790, see: Bledius.

fragilis Gravenhorst, 1802, see: Bisnius.

fraternus Bernhauer, 1915d: 150 (Staphylinus; Type locality: Erythraea: Gundet).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; catalog).

  • Distribution: Eritrea.

fugax Faldermann, 1835, see: Philonthus.

fulgens Fabricius, 1793, see: Plochionocerus.

fulgescens Turton, 1802, see: Gauropterus.

fulgidus Fabricius, 1787, see: Gauropterus.

fulgidus Fabricius, 1793, see: Quedius.

fuliginosus Gravenhorst, 1802, see: Quedius.

fulminans Gravenhorst, 1802, see: Othius.

fulviceps Motschulsky, 1858, see: Dinothenarus.

fulvipenne Gravenhorst, 1806, see: Lathrobium.

fulvipes Scopoli, 1763, see: Platydracus.

fulvipes Fabricius, 1793, see: Philonthus.

fulvipilis G. Müller, 1932, see: Tasgius.

fulvomaculatus Nordmann, 1837, see: Platydracus.

fulvopterus Geoffroy, 1785, see: Othius.

fulvopubescens Chapman, 1933, see: Tasgius.

fulvus De Geer, 1774, see: Anthophagus.

fumarius Gravenhorst, 1806, see: Philonthus.

funebris Sharp, 1884: 375 (Staphylinus; Type locality: Guatemala, Purula).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; catalog).

  • Distribution: Guatemala.

fuscatus Gravenhorst, 1802, see: Ocypus.

fuscescens Turton, 1802, see: Tachinus.

fuscicornis O. Müller, 1776: 98 (Staphylinus; [Note: Nomen dubium]; Type locality: Dania).

  • — Gmelin, 1790: 2033 (Staphylinus; characters; Denmark).

  • — Turton, 1802: 517 (Staphylinus; characters; Denmark).

  • Distribution: Denmark.

fuscicornis Germar, 1824, see: scabrosus Curtis, 1839.

fuscipennis Block, 1799, see: Quedius.

fuscipennis Mannerheim, 1830, see: Philonthus.

fuscipes Linné, 1758, see: Gyrohypnus.

fuscipes Heer, 1839, see: Ocypus.

fuscofemoratus G. Müller, 1923, see: Platydracus.

fuscolineatus Bernhauer, 1934, see: Platydracus.

fuscomaculatus Goeze, 1777: 730 (Staphylinus; [Note: Nomen dubium]; Type locality: Not cited).

  • — Herman, 2001: 51 (Staphylinus; nomen dubium).

  • Distribution: ?.

fuscomaculatus Laporte, 1835: 113 (Emus; [preoccupied]; Type locality: Mexique).

  • — Erichson, 1839b: 376 (Staphylinus; characters; Mexico).

  • — Sharp, 1884: 378 (Staphylinus; characters; Mexico; Guatemala).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; catalog).

  • — Herman, 2001: 51 (Staphylinus; although a junior primary homonym, the senior homonym is a nomen dubium, so it is suggested that the required replacement be ignored).

  • Distribution: Mexico, Guatemala.

fuscus Gmelin, 1790: 2027 (Staphylinus; [Note: Nomen dubium]; Type locality: Europa).

  • — Zschach, 1788: 45 (Staphylinus; species number 1000; characters stated but not named and referred to as fuscus by Gmelin).

  • — Turton, 1802: 515 (Staphylinus; characters; Europe).

  • Distribution: Europe.

fuscus Gravenhorst, 1802, see: Bisnius.

fusiformis Casey, 1915, see: Platydracus.

gabiruensis Bernhauer, 1934, see: Platydracus.

ganglbauerianus Bernhauer, 1939, see: Dinothenarus.

gemmatus Fauvel, 1895, see: Platydracus.

gentilis Erichson, 1839b: 387 (Staphylinus; Type locality: Brasilia).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; catalog).

  • Distribution: Brazil.

mrazi Rambousek, 1925: 76 (Staphylinus; variety of gentilis; Type locality: São Paulo).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; variety of gentilis).

geometricus Schrank, 1798, see: Aleochara.

gerardi Bondroit, 1913: 90 (Staphylinus; Type locality: Mufunga, Katanga).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; catalog).

  • Distribution: Congo.

germanus Turton, 1802, see: Dinothenarus.

germarii† Oustalet, 1874: 157 (Staphylinus; Type locality: Aix en Provence).

  • — Scudder, 1891: 585 (Staphylinus; catalog; Ligurian; France).

  • — Handlirsch, 1907: 724 (Staphylinus; catalog; Oligocene; France).

  • Distribution: France.

giganteus Cuvier, 1833: 196 (Staphylinus; [Note: Nomen dubium]; Type locality: Not cited).

  • — Herman, 2001: 51 (Staphylinus; nomen dubium).

  • Distribution: ?.

giganteus Kraatz, 1899: 112 (Leistotrophus; [preoccupied]; Type locality: Africa orientalis, Ukambani).

  • — Bernhauer, 1913b: 232 (Staphylinus; subgenus Chitocompsus).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; catalog).

  • — Herman, 2001: 51 (Staphylinus; although a junior primary homonym, the senior homonym is a nomen dubium, so it is suggested that the required replacement be ignored).

  • Distribution: “East Africa”.

glaber O. Müller, 1776, see: Quedius.

glaber Gmelin, 1790: 2035 (Staphylinus; [preoccupied]; Type locality: Europa).

  • — Zschach, 1788: 44 (Staphylinus; species number 973; characters stated but not named and referred to as glaber by Gmelin).

  • — Turton, 1802: 519 (Staphylinus; characters; Europe).

  • — Herman, 2001: 51 (Staphylinus; nomen dubium; although a junior primary homonym, it is suggested that the required replacement be ignored).

  • Distribution: Europe.

glaber Gravenhorst, 1802, see: Hypnogyra.

glabricollis Lacordaire, 1835, see: Acylophorus.

glaberrimus Herbst, 1784, see: Acylophorus.

glabratus Gravenhorst, 1802, see: Megalinus.

glasunovi Luze, 1904, see: Parabemus.

glaucus Gmelin, 1790: 2034 (Staphylinus; [Note: Nomen dubium]; Type locality: Gallia).

  • — Turton, 1802: 518 (Staphylinus; characters; France).

  • Distribution: France.

globulifer Geoffroy, 1785, see: Tasgius.

gmelini Blackwelder, 1944: 139 (Staphylinus; replacement name for haemorrhoidalis Olivier).

  • Distribution: French Guiana.

haemorrhoidalis Olivier, 1795: (42): 11 (Staphylinus; [preoccupied]; Type locality: Cayenne).

  • — Gravenhorst, 1806: 110 (Staphylinus; synonym of tricolor; Gravenhorst).

  • — Erichson, 1839b: 394 (Staphylinus; characters; Cayenne).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; catalog).

  • — Blackwelder, 1944: 139 (Staphylinus; synonym of gmelini).

goliathus Bernhauer, 1912f: 205 (Staphylinus; Type locality: Senegal).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; catalog).

  • Distribution: Senegal.

goryi Laporte, 1835, see: Platydracus.

gracilipes Sharp, 1884: 388 (Staphylinus; Type locality: Mexico, Cordova, Misantla, Jalapa).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; catalog).

  • Distribution: Mexico.

gracilis Paykull, 1789, see: Astenus.

graecus Scheerpeltz, 1958, see: Ocypus.

gratiosus Sharp, 1876: 152 (Staphylinus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; catalog).

  • Distribution: Brazil.

gratus Sharp, 1876: 153 (Staphylinus; Type locality: Tunantins).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; catalog).

  • Distribution: Brazil.

gravenhorsti Blackwelder, 1944: 139 (Staphylinus; replacement name for concolor Motschulsky).

  • Distribution: Colombia.

concolor Motschulsky, 1858: 670 (Trichoderma; [Note: Preoccupied secondary homonym]; Type locality: Colombie).

  • — Fauvel, 1878f: 249 (Staphylinus).

  • — Bernhauer and Schubert, 1914: 377 (Staphylinus; catalog).

  • — Blackwelder, 1944: 139 (Staphylinus; synonym of gravenhorsti).

gregarius Scopoli, 1763, see: Paederus.

gridellii G. Müller, 1924, see: Ocypus.

griseipennis Fairmaire, 1889: 10 (Staphylinus; Type locality: Moupin).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1385 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: China.

groenlandicus Zetterstedt, 1838, see: Quedius.

guatemalenus Sharp, 1884: 386 (Staphylinus; Type locality: Guatemala, Sinanja).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; catalog).

  • Distribution: Guatemala.

guineensis Cameron, 1950, see: Platydracus.

haemorrhoidalis Gmelin, 1790: 2036 (Staphylinus; Type locality: Europa).

  • — Zschach, 1788: 44 (Staphylinus; species number 983; characters stated but not named and referred to as haemorrhoidalis by Gmelin).

  • — Turton, 1802: 520 (Staphylinus; characters; Europe).

  • — Herman, 2001: 42 (Staphylinus; nomen dubium).

  • Distribution: Europe.

haemorrhoidalis Olivier, 1795, see: gmelini Blackwelder, 1944.

haemorrhoidalis Fabricius, 1801, see: Belonuchus.

haemorrhoidalis Germar, 1824, see: Xanthopygus.

haemorrhoidalis Grimmer, 1841 [Note: Nomen dubium], see: Xanthopygus.

hauseri Bernhauer, 1933, see: Ocypus.

hauserianus Bernhauer, 1933, see: Platydracus.

havaniensis Laporte, 1835, see: Philonthus.

hebraeus Smetana, 1978, see: Abemus.

hellebori Scopoli, 1763: 102 (Staphylinus; [Note: Nomen dubium]; Type locality: Carniolia).

  • — Goeze, 1777: 725 (Staphylinus; characters).

  • — Villers, 1789: 423 (Staphylinus; characters).

  • — Gmelin, 1790: 2032 (Staphylinus; page misnumbered as 1032; characters; France).

  • Distribution: Austria, France.

helleni G. Müller, 1926, see: Ocypus.

hemichrysis Fauvel, 1905d: 196 (Staphylinus; Type locality: Abyssinie: Azega, 2500 m; Afrique orientale allemande: Manow. Kiboscho, Kilimandjaro. Angola).

  • — Bernhauer, 1906a: 188 (Staphylinus; characters; Ethiopia).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; catalog).

  • — Bernhauer and Chapman, 1939: 73 (Staphylinus; Kenya).

  • — Bernhauer, 1940: 136 (Staphylinus; Kenya).

  • — Cameron, 1950g: 43 (Staphylinus; subgenus Abemus; Belgian Congo).

  • — Tottenham, 1956: 229 (Staphylinus; subgenus Abemus; Ruanda; Urundi).

  • — Scheerpeltz, 1957: 154 (Staphylinus; Tanganyika).

  • Distribution: Ethiopia, Kenya, Tanzania, Rwanda, Burundi, Congo, Angola.

hercegovinensis G. Müller, 1932, see: Ocypus.

herculeanus Laporte, 1835, see: Trigonopselaphus.

hesperus Crotch, 1867, see: Ocypus.

hewitti Bernhauer, 1914, see: Platydracus.

hilaris Erichson, 1839, see: Xanthopygus.

himalayicus Cameron, 1935, see: Ocypus.

hirtus Linné, 1758, see: Emus.

holosetosa G. Müller, 1943, see: Ocypus.

homonymus Blackwelder, 1944: 139 (Staphylinus; replacement name for lugubris Le Peletier and Serville).

  • Distribution: French Guiana.

lugubris Le Peletier and Serville, 1828: 477 (Staphylinus; [preoccupied]; Type locality: Cayenne).

  • — Erichson, 1839b: 394 (Staphylinus; characters; Cayenne).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; Olivier cited as author; catalog).

  • — Blackwelder, 1944: 139 (Staphylinus; synonym of homonymus).

hottentottus Nordmann, 1837: 37 (Staphylinus; Type locality: Promontorium bonae spei).

  • — Erichson, 1839b: 390 (Staphylinus; characters; South Africa).

  • — Boheman, 1848: 278 (Staphylinus; characters; South Africa).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; catalog).

  • — Rambousek, 1922: 1012 (Staphylinus; cited as hottentotus; Ethiopia).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • — Scheerpeltz, 1974: 170 (Staphylinus; South Africa).

  • Distribution: South Africa, Ethiopia.

capensis Sachse, 1852: 125 (Staphylinus; Type locality: Vorgeb. d. g. Hoffnung).

  • — Gemminger and Harold, 1868: 579 (Staphylinus; synonym of hottentottus).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; synonym of hottentottus).

humeralis Matsumura, 1911, see: Liusus.

huttoni Broun, 1880, see: Creophilus.

hybridus Marsham, 1802, see: Ontholestes.

hypnorum Fabricius, 1775, see: Tachyporus.

hypocrita G. Müller, 1925, see: Platydracus.

hypsibatus Bernhauer, 1899, see: Ocypus.

ibericus Kolenati, 1846, see: Ocypus.

ignavus Runde, 1835: 9 (Staphylinus; [Note: Nomen dubium]; Type locality: Halae).

  • Distribution: Germany.

ignotus Sharp, 1884: 381 (Staphylinus; Type locality: Mexico, Mirador; Panama, Volcan de Chiriqui 4000 to 6000 feet).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • Distribution: Mexico, Panama.

immaculatus Mannerheim, 1830, see: Platydracus.

immundus Gravenhorst, 1806, see: Philonthus.

immunis Marsham, 1802, see: Stenus.

impennis Fauvel, 1882, see: Ocypus.

imperatorius Bernhauer, 1916, see: Platydracus.

impotens Eppelsheim, 1889, see: Platydracus.

impressifrons Solier, 1849, see: Cheilocolpus.

impressus Olivier, 1795, see: Autalia.

impressus Panzer, 1796, see: Quedius.

inauratus Normand, 1947, see: Ocypus.

incanus Klug, 1832, see: Creophilus.

incognitus Sharp, 1884: 376 (Staphylinus; Type locality: Mexico, Coatepec).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • Distribution: Mexico.

indicus Kraatz, 1859, see: Platydracus.

infuscatus Nordmann, 1837, see: Leistotrophus.

initicornis Scheerpeltz, 1933, see: Platydracus.

inornatus Sharp, 1874, see: Platydracus.

insignis G. Müller, 1926, see: Dinothenarus.

insolitus Sharp, 1884, see: Platydracus.

insularis Cameron, 1941, see: Platydracus.

intermedius Lacordaire, 1835, see: Philonthus.

intermedius G. Müller, 1911, see: Ocypus.

interruptus Brullé, 1842, see: Glenus.

inversus Say, 1830, see: Quedius.

iopterus Erichson, 1839, see: Xanthopygus.

iracundus Say, 1830, see: Quedius.

iridiventris Bernhauer, 1936a: 24 (Staphylinus; subgenus Abemus; Type locality: 18 m. S. W. von Elisabethville).

  • Distribution: Congo.

iringanus Bernhauer, 1937b: 619 (Staphylinus; subgenus Abemus; Type locality: Iringa, 1200–1500 m).

  • Distribution: Tanzania.

irregularis Mannerheim, 1830, see: Philonthus.

italicus Aragona, 1830, see: Ocypus.

janthinipennis Blanchard, 1842, see: Xanthopygus.

javanus Bernhauer, 1934, see: Platydracus.

jeanneli Chapman, 1939, see: Platydracus.

juno Paykull, 1789, see: Stenus.

jureceki Rambousek, 1923, see: Ocypus.

kalisi Bernhauer, 1934j: 174 (Staphylinus; subgenus Abemus; Type locality: Ost-Java, Nongkodjadjar, 4000 f.).

  • Distribution: Indonesia.

kamerunensis Bernhauer, 1912f: 204 (Staphylinus; subgenus Abemus; Type locality: Südost-Kamerun: Lolodorf).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • Distribution: Cameroon.

kasyi Scheerpeltz, 1962, see: Platydracus.

kiautschauensis Bernhauer, 1916, see: Dinothenarus.

kiulungensis Bernhauer, 1933, see: Platydracus.

kiushiuensis Bernhauer, 1939, see: Platydracus.

kraatzi Bernhauer, 1906a: 188 (Staphylinus; Type locality: Kamerun).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • Distribution: Cameroon.

kristenseni Bernhauer, 1915d: 148 (Staphylinus; Type locality: Abessinien: Harrar).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • Distribution: Ethiopia.

kulbaikhani Muona, 1977, see: trimaculatus Fauvel, 1895.

kuntzeni G. Müller, 1926, see: Ocypus.

lacustris Bernhauer, 1917, see: Dinothenarus.

laetulus Say, 1834, see: Laetulonthus.

laetus Faldermann, 1835, see: Quedius.

levicollis Brullé, 1832, see: Quedius.

laevicollis Runde, 1835, see: Quedius.

laevicollis Lacordaire, 1835, see: Philonthus.

laevigatus Marsham, 1802, see: Tachinus.

laevigatus Gyllenhal, 1810, see: Quedius.

laevipennis Dufour, 1843, see: Quedius.

laminatus Creutzer, 1799, see: Philonthus.

langei Bernhauer, 1904a: 235 (Staphylinus; subgenus Ocypus; Type locality: Kongogebiet).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • Distribution: “Congo region”.

lanio Erichson, 1839, see: Creophilus.

lapponicus Zetterstedt, 1838, see: Quedius.

laqueatus Marsham, 1802, see: Oxytelus.

latebricola Gravenhorst, 1806, see: Platydracus.

latecarinatus Bernhaeur, 1937, see: Platydracus.

lateralis Gravenhorst, 1802, see: Quedius.

laticeps Zetterstedt, 1828, see: Philonthus.

laticollis Gravenhorst, 1802, see: Quedius.

latro Erichson, 1839b: 374 (Staphylinus; Type locality: Brasilia).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • Distribution: Brazil.

latus Ström, 1768: 332 [= 1770: 280] (Staphylinus; [Note: Nomen dubium]; Type locality: Norske).

  • — Herman, 2001: 51 (Staphylinus; nomen dubium).

  • Distribution: Norway.

latus O. Müller, 1776: 97 (Staphylinus; [preoccupied]; Type locality: Dania).

  • — Gmelin, 1790: 2033 (Staphylinus; characters; Denmark).

  • — Turton, 1802: 517 (Staphylinus; characters;).

  • — Herman, 2001: 51 (Staphylinus; nomen dubium; although a junior primary homonym, it is suggested that the required replacement be ignored).

  • Distribution: Denmark.

latus Marsham, 1802, see: Tachinus.

lefevrei Bernhauer, 1936, see: Platydracus.

leiocephalus Solier, 1849, see: Loncovilius.

lentus Gravenhorst, 1806, see: Nudobius.

lepidus Gravenhorst, 1802, see: Philonthus.

leroyi Bernhauer, 1938, see: Platydracus.

lesleyi† Scudder, 1900: 51 (Staphylinus; fossil; Type locality: Florissant, Colorado).

  • — Handlirsch, 1907: 724 (Staphylinus; catalog; Miocene; Colorado).

  • — Wickham, 1912: 11 (Staphylinus; Colorado).

  • Distribution: USA.

leucomus Erichson, 1839, see: Trichocosmetes.

leucophthalmus Marsham, 1802, see: Ocypus.

leucopus Marsham, 1802, see: Tachyusa.

lewisi Cameron, 1932, see: Platydracus.

libanoticus G. Müller, 1950, see: Ocypus.

lignorum Linné, 1758, see: Tachinus.

limbatus Paykull, 1789, see: Zyras.

limbatus Fabricius, 1801: 600 (Staphylinus; [Note: Nomen dubium; preoccupied]; Type locality: Germania).

  • — Herman, 2001: 51 (Staphylinus; nomen dubium; although a junior primary homonym, it is suggested that the required replacement be ignored).

  • Distribution: Germany.

limbatus Klug, 1832, see: Eulissus.

linearis Olivier, 1795, see: Xantholinus.

linearis Gravenhorst, 1802, see: Leptacinus.

lineatus Walker, 1859, see: Platydracus.

litoreus Broun, 1880, see: Cafius.

littoreus Linné, 1758, see: Sepedophilus.

lituratus Gmelin, 1790: 2036 (Staphylinus; [Note: Nomen dubium]; Type locality: Europa).

  • — Zschach, 1788: 44 (Staphylinus; species number 991; characters stated but not named and referred to as lituratus by Gmelin).

  • — Turton, 1802: 520 (Staphylinus; characters; Europe).

  • Distribution: Europe.

lomii Cerruti, 1951, see: Platydracus.

longiceps Gravenhorst, 1802, see: Xantholinus.

longicornis Waltl, 1835, see: Philonthus.

longoelytratus Goeze, 1777, see: Lesteva.

longomaculatus Goeze, 1777: 730 (Staphylinus; [Note: Nomen dubium]; Type locality: Not cited).

  • Distribution: ?.

lucanus Horn, 1894, see: Platydracus.

lucens Mannerheim, 1830, see: Philonthus.

lucidus Gravenhorst, 1802, see: Philonthus.

luctuosus Blanchard, 1842, see: Xanthopygus.

ludovicae Mulsant and Godart, 1855, see: Ocypus.

luganensis Heer, 1839, see: Tasgius.

lugens Nordmann, 1837, see: Ocypus.

lugubris Gmelin, 1790: 2036 (Staphylinus; [Note: Nomen dubium]; Type locality: Europa).

  • — Zschach, 1788: 44 (Staphylinus; species number 976; characters stated but not named and referred to as lugubris by Gmelin).

  • — Turton, 1802: 519 (Staphylinus; characters; Europe).

  • Distribution: Europe.

lugubris Le Peletier and Serville, 1828, see: homonymus Blackwelder, 1944.

lugubris Nordmann, 1837, see: scabrosus Curtis, 1839.

luigionii G. Müller, 1926, see: Ocypus.

lunulatus Linné, 1760, see: Lordithon.

lupinus Sharp, 1887: 785 (Staphylinus; Type locality: Mexico, Refugio in Durango).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • Distribution: Mexico.

luridipennis W. J. MacLeay, 1873, see: Quedius.

luridipes Erichson, 1839, see: Oligotergus.

luridus Runde, 1835, see: Philonthus.

lutarius Gravenhorst, 1806, see: Platydracus.

luteipes LeConte, 1861, see: Dinothenarus.

luzonicus Fauvel, 1886b: 149 (Staphylinus; Type locality: Luçon).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • Distribution: Philippines.

macgregori Cooper, 1933, see: Platydracus.

macrocephalus Gravenhorst, 1802, see: Ocypus.

macroclytron Geoffroy, 1785, see: Lesteva.

macropterus Gmelin, 1790: 2034 (Staphylinus; [Note: Nomen dubium]; Type locality: Gallia).

  • — Turton, 1802: 518 (Staphylinus; characters; France).

  • Distribution: France.

maculatus Geoffroy, 1785, see: Sepedophilus.

maculicollis Fauvel, 1895: 253 (Staphylinus; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Carin Asciuii Ghécu, 1400–1500 m).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • — Cameron, 1932: 200 (Staphylinus; characters; Burma; India).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog; India).

  • Distribution: Myanmar, India.

maculipennis Kraatz, 1859, see: Platydracus.

maculiventris Sharp, 1884: 375 (Staphylinus; Type locality: Mexico, Ventanas 2000 feet).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • Distribution: Mexico.

maculosus Gravenhorst, see: Platydracus.

maderae Jarrige, 1943, see: Tasgius.

major De Geer, 1774, see: Ocypus.

mandli Bernhauer, 1906, see: Ocypus.

marani Roubal, 1932, see: Tasgius.

marcidus Sharp, 1884: 379 (Staphylinus; Type locality: Panama, Volcan de Chiriqui 4000 to 6000 feet, Bugaba 800 to 1500 feet; Costa Rica, Irazu 6000 to 7000 feet).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • Distribution: Costa Rica, Panama.

marginalis O. Müller, 1776: 98 (Staphylinus; [Note: Nomen dubium]; Type locality: Dania).

  • — Gmelin, 1790: 2033 (Staphylinus; characters; Denmark).

  • — Schrank, 1798: 646 (Staphylinus; characters; notes).

  • — Turton, 1802: 517 (Staphylinus; characters; Denmark).

  • Distribution: Denmark.

marginatus O. Müller, 1764, see: Philonthus.

marginatus Ström, 1768, see: Philonthus.

marginatus Fabricius, 1775, see: Philonthus.

marginatus Geoffroy, 1785, see: Tachinus.

marginatus Cameron, 1944: 11 (Staphylinus; subgenus Abemus; [preoccupied]; Type locality: Bengal: Kalimpong, Samsingh, altitude 1800 feet).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • — Herman, 2001: 51 (Staphylinus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: India.

marginellus Fabricius, 1781, see: Tachinus.

marginellus Gmelin, 1790: 2036 (Staphylinus; [preoccupied]; Type locality: Europa).

  • — Zschach, 1788: 44 (Staphylinus; species number 982; characters stated but not named and referred to as marginellus by Gmelin).

  • — Turton, 1802: 519 (Staphylinus; characters; Europe).

  • — Herman, 2001: 51 (Staphylinus; nomen dubium; although a junior primary homonym, it is suggested that the required replacement be ignored).

  • Distribution: Europe.

marginosulcatus Goeze, 1777: 730 (Staphylinus; [Note: Nomen dubium]; Type locality: Not cited).

  • Distribution: ?.

marinus Ström, 1883, see: Micralymma.

maritimus Broun, 1880, see: Cafius.

marmoratus Erichson, 1840, see: Ontholestes.

marmorellus Fauvel, 1895, see: Platydracus.

martialis Schrank, 1781, see: Tachyporus.

masculus Nordmann, 1837, see: Ocypus.

mateui Coiffait, 1954, see: Ocypus.

matutinalis Faldermann, 1835, see: Philonthus.

maurorufus Gravenhorst, 1806, see: Quedius.

maurus C. Sahlberg, 1830, see: Quedius.

maxillosus Linné, 1758, see: Creophilus.

medialis Sharp, 1884: 384 (Staphylinus; Type locality: Mexico, Oaxaca).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • Distribution: Mexico.

medioximus Fairmaire, 1852: 73 (Staphylinus; Type locality: Tanger).

  • — Fairmaire and Coquerel, 1860: 156 (Staphylinus; characters; Algeria; Morocco).

  • — Mulsant and Rey, 1876: 254 [= 1877: 110] (Staphylinus; characters).

  • — Fauvel, 1878: 118 [= 1878a: 38] (Staphylinus; Morocco; Algeria).

  • — Fauvel, 1886: 50 [= 1886a: 42] (Staphylinus; Morocco; Algeria).

  • — Fauvel, 1897d: 294 (Staphylinus; Barbary).

  • — Fauvel, 1902b: 103 (Staphylinus; Morocco; Algeria; Tunisia).

  • — Bernhauer and Schubert, 1914: 380 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog; Morocco; Gibraltar; Spain).

  • — Coiffait, 1974: 405 (Staphylinus; characters; North Africa; Iberia; Morocco).

  • — Merceron, 1998 (Staphylinus; France).

  • — Smetana and Davies, 2000: 41 (Staphylinus).

  • Distribution: France, Spain, Morocco, Algeria, Tunisia.

sculpticollis Gaubil, 1852: 7 (Staphylinus; without description; [Note: Nomen nudum (Article 12.1)]; cited from Algeria).

  • — Reiche, 1852: xxxvii (Staphylinus; synonym of medioximus).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; synonym of medioximus).

  • — Coiffait, 1974: 405 (Staphylinus; synonym of medioximus).

megalocephalus Nordmann, 1837, see: Ocypus.

melanarius Heer, 1839, see: Tasgius.

melanocephalus Geoffroy, 1785: 172 (Staphylinus; [Note: Nomen dubium]; Type locality: Paris).

  • — Villers, 1789: 423 (Staphylinus; characters; France).

  • — Herman, 2001: 39 (Staphylinus; nomen dubium).

  • Distribution: France.

melanocephalus Fabricius, 1787, see: Phyllodrepa.

melanocephalus Gravenhorst, 1806, see: Othius.

melanophtalmos Geoffroy, 1785: 171 (Staphylinus; [Note: Nomen dubium]; Type locality: Paris).

  • — Villers, 1789: 422 (Staphylinus; characters; France).

  • Distribution: France.

melanophthalmus Gmelin, 1790: 2027 (Staphylinus; [Note: Nomen dubium]; Type locality: Europa).

  • — Zschach, 1788: 45 (Staphylinus; species number 1005; characters stated but not named and referred to as melanophthalmus by Gmelin).

  • — Turton, 1802: 515 (Staphylinus; characters; Europe).

  • Distribution: Europe.

melanopus Marsham, 1802, see: Stenus.

melanurus Schrank, 1798: 646 (Staphylinus; [Note: Nomen dubium]; Type locality: Boia).

  • Distribution: Germany.

melanurus Marsham, 1802, see: Tachyporus.

memnonius Sharp, 1884: 380 (Staphylinus; Type locality: Panama, Volcan de Chiriqui 2500 to 4000 feet).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • Distribution: Panama.

mendax Runde, 1835, see: Philonthus.

mendicus Sharp, 1884, see: Platydracus.

merdarius Fabricius, 1775, see: Tachyporus.

meridionalis Rosenhauer, 1847, see: Platydracus.

mesomelinus Marsham, 1802, see: Quedius.

metallicus Gravenhorst, 1802, see: Philonthus.

metallicus O. Costa, 1839, see: Ocypus.

methneri Bernhauer, 1915d: 148 (Staphylinus; Type locality: Deutsch-Ostafrika).

  • — Scheerpeltz, 1933: 1386 (Staphylinus; catalog).

  • Distribution: Tanzania.

methnerianus Bernhauer, 1936a: 24 (Staphylinus; Type locality: Mwengwa).

  • Distribution: Congo.

mexicanus Sharp, 1884: 384 (Staphylinus; Type locality: Mexico, Misantla).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • Distribution: Mexico.

micans Gravenhorst, 1802, see: Philonthus.

microcephalus Gravenhorst, 1806, see: Philonthus.

mimeticus Bernhauer, 1917, see: Platydracus.

minor O. Müller, 1776: 97 (Staphylinus; [Note: Nomen dubium]; Type locality: Dania).

  • — Herbst, 1784: 150 (Staphylinus; cited as mikor; Germany).

  • — Gmelin, 1790: 2031 (Staphylinus; characters; northern Europe).

  • — Turton, 1802: 516 (Staphylinus; characters; northern Europe).

  • Distribution: Germany, Denmark.

minimus Schrank, 1781, see: Psocoptera.

minutus Olivier, 1795, see: Acrolocha.

minutus Marsham, 1802: 511 (Staphylinus; [preoccupied]; Type locality: Britain).

  • — Herman, 2001: 52 (Staphylinus; nomen dubium; although a junior primary homonym, it is suggested that the required replacement be ignored).

  • Distribution: Britain.

mirtensis G. Müller, 1943, see: Ocypus.

miwai Bernhauer, 1943, see: Ocypus.

modestus Sharp, 1884, see: Platydracus.

modestus Fall, 1907, see: Platydracus.

moestus Gravenhorst, 1806, see: Philonthus.

molochinus Gravenhorst, 1806, see: Quedius.

mongendensis Bernhauer, 1929, see: Platydracus.

montanus Cameron, 1942, see: Ocypus.

mordax Block, 1799, see: Ocypus.

morio Gravenhorst, 1802, see: Tasgius.

morsitans Rossi, 1790, see: Tasgius.

morsitans Paykull, 1792, see: Platystethus.

mortuorum Bernhauer, 1912, see: Platydracus.

mrazi Rambousek, 1925, see: gentilis Erichson, 1839.

muellerianus Scheerpeltz, 1933, see: Platydracus.

mulsanti Godard, 1850, see: Platydracus.

multipunctatus Block, 1799, see: Lesteva.

multipunctatus Mannerheim, 1830, see: Philonthus.

murinus Linné, 1758, see: Ontholestes.

mus Brullé, 1832, see: Ocypus.

myops G. Müller, 1925, see: Tasgius.

mysticus Erichson, 1840, see: Platydracus.

nanus Paykull, 1800, see: Gyrophaena.

nanus Gravenhorst, 1806, see: Gabrius.

nebulosus Geoffroy, 1785, see: Creophilus.

nebulosus Fabricius, 1793, see: Ontholestes.

nebulosus Percheron, 1835, see: ochropygus Nordmann, 1837.

neomexicanus Bernhauer and Schubert, 1914, see: Platydracus.

nepalensis Scheerpeltz, 1976, see: Platydracus.

nero Faldermann, 1835, see: Ocypus.

nevadensis G. Müller, 1926, see: Ocypus.

niger O. Müller, 1764, see: Philonthus.

nigrellus Horn, 1879, see: Dinothenarus.

nigrescens Blanchard, 1842, see: scabrosus Curtis, 1839.

nigricapillus Turton, 1802, see: Tachyporus.

nigriceps Marsham, 1802, see: Atheta.

nigricollis Paykull, 1800, see: Ilyobates.

nigripennis Cameron, 1941, see: Platydracus.

nigrita Gravenhorst, 1806, see: Philonthus.

nigritarsis Ménétriés, 1832, see: Quedius.

nigritulus Gravenhorst, 1802, see: Gabrius.

nigritus Runde, 1835, see: Philonthus.

nigriventris Boheman, 1848, see: Platydracus.

nigrocephalus Turton, 1802, see: Sunius.

nigrofulvus Geoffroy, 1785, see: Oxyporus.

nigrophthalmus Block, 1799, see: Anthophagus.

nitens Schrank, 1781, see: Ocypus.

nitens Gravenhorst, 1802, see: Philonthus.

nitidicollis Lacordaire, 1835, see: Philonthus.

nitidipennis Solier, 1849, see: Endeius.

nitidulus Fabricius, 1781, see: Tachyporus.

nitidulus Gravenhorst, 1802, see: Bisnius.

nitidus Fabricius, 1787, see: Philonthus.

nobilis Nordmann, 1837: 41 (Staphylinus; Type locality: Brasilia meridionalis).

  • — Erichson, 1839b: 384 (Staphylinus; characters; Brazil).

  • — Lynch, 1884: 137 (Staphylinus; characters; Argentina).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; catalog).

  • Distribution: Brazil, Argentina.

normannicus Cuvier, 1833: 196 (Staphylinus; [Note: Nomen dubium]; Type locality: Not cited).

  • Distribution: ?.

notativentris Fauvel, 1905, see: Platydracus.

notatus Solsky, 1872: 293 (Staphylinus; Type locality: Lima).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • Distribution: Peru.

nudicollis Sharp, 1887: 786 (Staphylinus; Type locality: Mexico, Pastepec).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • Distribution: Mexico.

obscurus Herbst, 1784: 149 (Staphylinus; [Note: Nomen dubium]; Type locality: Berlin).

  • — Gmelin, 1790: 2031 (Staphylinus; characters).

  • — Turton, 1802: 516 (Staphylinus; characters; Germany).

  • — Herman, 2001: 47 (Staphylinus; nomen nudum).

  • Distribution: Germany.

obscurus Paykull, 1800, see: Lesteva.

obscurus Fabricius, 1801, see: Zyras.

obscurus Gravenhorst, 1802, see: Philonthus.

obscurus Marsham, 1802, see: Tasgius.

obscurus Runde, 1835, see: Tasgius.

obsoleticornis Bernhauer, 1911a: 420 (Staphylinus; Type locality: Brasilien, Matto Grosso, Cuyaba).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • Distribution: Brazil.

obtusus Linné, 1767, see: Tachyporus.

ochraceus Gravenhorst, 1802, see: Xantholinus.

ochraceus Gyllenhal, 1810, see: Gyrohypnus.

ochripennis Ménétriés, 1832, see: Quedius.

ochrocephalus Gmelin, 1790, see: Dinothenarus.

ochropterus Germar, 1824, see: Platydracus.

ochropus Gravenhorst, 1802, see: Philonthus.

ochropus G. Müller, 1950, see: Ocypus.

ochropygus Nordmann, 1837: 46 (Staphylinus; Type locality: America ?).

  • — Erichson, 1839b: 373 (Staphylinus; characters; Brazil; Cayenne).

  • — Solsky, 1875: 15 (Staphylinus; Peru).

  • — Sharp, 1876: 152 (Staphylinus; Brazil).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • Distribution: Peru, Brazil, French Guiana.

nebulosus Percheron, 1835: liv. 3, no. 1, pl. 5 (Emus; [preoccupied]; Type locality: Cayenne).

  • — Erichson, 1839b: 373 (Staphylinus; synonym of ochropygus).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; synonym of ochropygus).

octavus Schaeffer, 1769 [Note: Nomen nudum], see: Oxyporus.

oculatus Fabricius, 1775, see: Creophilus.

oculatus O. Müller, 1776: 99 (Staphylinus; [preoccupied]; Type locality: Dania).

  • — Herman, 2001: 52 (Staphylinus; nomen dubium; although a junior primary homonym, it is suggested that the required replacement be ignored).

  • Distribution: Denmark.

oculatus Bernhauer, 1929, see: Platydracus.

olens O. Müller, 1764, see: Ocypus.

olivaceus Cameron, 1928, see: Abemus.

opaciceps Scheerpeltz, 1976, see: Platydracus.

opacus Block, 1799, see: Bledius.

opacus Gravenhorst, 1802, see: Philonthus.

ophthalmicus Scopoli, 1763, see: Ocypus.

ophthalmicus Paykull, 1800, see: Eusphalerum.

optatus Sharp, 1884: 386 (Staphylinus; Type locality: Mexico).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • Distribution: Mexico.

orbiculatus Paykull, 1789, see: Rugilus.

orientalis Motschulsky, 1858, see: Creophilus.

orientalis Bernhauer and Schubert, 1914, see: Ocypus.

orizabae Bernhauer, 1917i: 109 (Staphylinus; subgenus Abemus; Type locality: Mexiko: Orizaba).

  • — Scheerpeltz, 1933: 1387 (Staphylinus; catalog).

  • Distribution: Mexico.

ormayi Reitter, 1887, see: Ocypus.

ornaticauda LeConte, 1863a: 37 (Staphylinus; Type locality: Canada).

  • — Horn, 1879: 191 (Staphylinus; synonym of caesareus).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; synonym of caesareus).

  • — Smetana, 1958: 275 (Staphylinus; synonym of caesareus).

  • — Smetana and Davies, 2000: 41 (Staphylinus; cited as valid species).

  • Distribution: Canada.

ornatus Erichson, 1839, see: Plociopterus.

osculatii Guérin-Méneville, 1855: 583 (Staphylinus; Type locality: Not cited).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog; Brazil).

  • Distribution: Brazil.

ovicollis Broun, 1880, see: Gabrius.

paganus Sharp, 1874, see: Platydracus.

palliatus Gravenhorst, 1806, see: Philonthus.

pallidipennis Panzer, 1795, see: Platystethus.

pallidipes Bernhauer, 1917, see: Platydracus.

pallidus Rossi, 1792, see: Lordithon.

palmula Gravenhorst, 1802, see: Neobisnius.

panamensis Bernhauer and Schubert, 1914, see: Platydracus.

parumpunctatus Gyllenhal, 1827, see: Phacophallus.

parumtomentosus Stein, 1903, see: dimidiaticornis Gemminger, 1851.

parviceps Sharp, 1876: 151 (Staphylinus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • Distribution: Brazil.

parvicornis Gravenhorst, 1802, see: Philonthus.

parvulus Gravenhorst, 1802, see: Gabrius.

parvus Gmelin, 1790, see: Tachyporus.

parvus Solier, 1849: 321 (Staphylinus; [preoccupied]; Type locality: Chile).

  • — Kraatz, 1859a: 5 (Staphylinus; subgenus Cheilocolpus; notes).

  • — Fairmaire and Germain, 1861: 433 (Staphylinus; apparently in Philonthus; characters; Chile).

  • — Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).

  • — Saiz, 1969a: 9 (Staphylinus; notes; species unknown).

  • Distribution: Chile.

parvus Cameron, 1932, see: Platydracus.

patricius Bernhauer, 1915, see: Platydracus.

pecoudi Jarrige, 1954, see: Ocypus.

pedator Gravenhorst, 1802, see: Tasgius.

pedemontanus G. Müller, 1924, see: Ocypus.

pendjabensis Bernhauer, 1915, see: Tasgius.

penetrans O. Müller, 1776, see: Ocypus.

peregrinus Gravenhorst, 1806, see: Quedius.

perniger Scheerpeltz, 1976, see: Platydracus.

peyerimhoffi G. Müller, 1926, see: Tasgius.

philanthus Fabricius, 1798, see: Phloeonomus.

philippinus Cameron, 1941, see: Platydracus.

phoenicurus Nordmann, 1837, see: Platydracus.

picatus Gmelin, 1790: 2035 (Staphylinus; [Note: Nomen dubium]; Type locality: Europa).

  • — Zschach, 1788: 44 (Staphylinus; species number 972; characters stated but not named and referred to as picatus by Gmelin).

  • — Turton, 1802: 519 (Staphylinus; cited as pictus; characters; Europe).

  • Distribution: Europe.

piceus Linné, 1767, see: Oxytelus.

picimanus Ménétriés, 1832, see: Philonthus.

picipennis Fabricius, 1793, see: Ocypus.

picipennis Nordmann, 1837: 71 (Staphylinus; [preoccupied]; Type locality: Europa meridionalis; Australia [?]).

  • — Herman, 2001: 52 (Staphylinus; nomen dubium; although a junior primary homonym, it is suggested that the required replacement be ignored).

  • Distribution: Australia (?), Europe.

picipes Mannerheim, 1830, see: Quedius.

picipes Nordmann, 1837, see: Ocypus.

pictus Boheman, 1860: 14 (Staphylinus; Type locality: laucus N'Gami).

  • — Bernhauer and Schubert, 1914: 381 (Staphylinus; catalog).

  • — Cameron, 1952e: 832 (Staphylinus; Dahomey).

  • Distribution: Botswana, Benin.

pilicornis Paykull, 1790, see: Atrecus.

pilosus Fabricius, 1787, see: Smilax.

pinorum Casey, 1915, see: Platydracus.

plagiatus Fabricius, 1798, see: Geodromicus.

planatus Gravenhorst, 1806, see: Neobisnius.

planus Paykull, 1792, see: Phloeostiba.

planus Lacordaire, 1835, see: Philonthus.

platypus Steven, 1829: 29 (Staphylinus; Type locality: Bras.).

  • Distribution: Brazil.

plebejus Bernhauer, 1915, see: Platydracus.

pleuralis LeConte, 1861, see: Dinothenarus.

pliginskii Bernhauer, 1915, see: Tasgius.

politus Linné, 1758, see: Philonthus.

polyphemus Bernhauer, 1913, see: Platydracus.

porcatus Paykull, 1789, see: Micropeplus.

posticalis Laporte, 1840, see: Belonuchus.

praelongus Mannerheim, 1830, see: Platydracus.

prasinivariegatus Bernhauer, 1921, see: Platydracus.

pratti Scheerpeltz, 1962, see: Platydracus.

preangeranus Bernhauer, 1937, see: Platydracus.

praecox Gravenhorst, 1802, see: Quedius.

praetorius Bernhauer, 1915, see: Tasgius.

primigenius G. Müller, 1923, see: Ocypus.

primus Schaeffer, 1766 [Note: Nomen nudum], see: caesareus Cederhjelm, 1798.

priscus† Oustalet, 1874: 166 (Staphylinus; Type locality: Aix en Provence).

  • — Scudder, 1891: 585 (Staphylinus; catalog; Ligurian; France).

  • — Handlirsch, 1907: 724 (Staphylinus; catalog; Oligocene; France).

  • Distribution: France.

priscus Sharp, 1876, see: emeritus Herman, 2001.

procerulus Gravenhorst, 1806, see: Neobisnius.

procerus Gahan, 1893, see: Platydracus.

prodromus† Oustalet, 1874: 163 (Staphylinus; Type locality: Aix en Provence).

  • — Scudder, 1891: 585 (Staphylinus; catalog; Ligurian; France).

  • — Handlirsch, 1907: 724 (Staphylinus; catalog; Oligocene; France).

  • Distribution: France.

propinquus Runde, 1835: 7 (Staphylinus; [Note: Nomen dubium]; Type locality: Halae).

  • Distribution: Germany.

protensus Ménétriés, 1832, see: Tasgius.

provincialis† Oustalet, 1874, see: Ocypus.

pseudoalpestris G. Müller, 1926, see: Ocypus.

pseudopaganus Bernhauer, 1914, see: Platydracus.

pseudopatricius G. Müller, 1926, see: Platydracus.

puberulus Runde, 1835, see: Ocypus.

pubescens De Geer, 1774, see: Dinothenarus.

pubescens Paykull, 1790, see: Sepedophilus.

pulcherrimus Bernhauer, 1915e: 123 (Staphylinus; subgenus Nesiolinus; Type locality: Luzon: Laguna, Mount Maquiling).

  • — Scheerpeltz, 1933: 1388 (Staphylinus; catalog).

  • Distribution: Philippines.

pullus Runde, 1835, see: Philonthus.

punctatissimus Dufour, 1843, see: Tasgius.

punctatissimus Motschulsky, 1858, see: dufouri Blackwelder, 1944.

punctatus Ström, 1768: 333 [= 1770: 281] (Staphylinus; [Note: Nomen dubium]; Type locality: Norske).

  • — Gmelin, 1790: 2033 (Staphylinus; characters; Denmark).

  • — Turton, 1802: 517 (Staphylinus; characters; Denmark).

  • Distribution: Denmark, Norway.

punctatus Geoffroy, 1785, see: Lathrobium.

punctatostriatus Paykull, 1800, see: Mycetoporus.

puncticeps White, 1846, see: Cafius.

puncticollis Runde, 1835: 5 (Staphylinus; [Note: Nomen dubium]; Type locality: Halae).

  • Distribution: Germany.

punctipennis Lacordaire, 1835, see: Othius.

punctipennis Solier, 1849, see: Endeius.

punctulatus Goeze, 1777, see: Othius.

punctulatus Paykull, 1789, see: Gyrohypnus.

punctulatus Gravenhorst, 1802, see: Xantholinus.

punctus Gravenhorst, 1802, see: Philonthus.

purpurascens Nordmann, 1837, see: Torobus.

purpurascens Cameron, 1920, see: Platydracus.

purpureoaureus Bernhauer, 1915, see: Platydracus.

pusillus Runde, 1835: 10 (Staphylinus; [Note: Nomen dubium]; Type locality: Halae).

  • Distribution: Germany.

pygmaeus Villers, 1789: 420 (Staphylinus; Type locality: Kiliae).

  • — Herman, 2001: 37 (Staphylinus; nomen dubium).

  • Distribution: Germany.

pygmaeus Paykull, 1800, see: Hapalaraea.

pygmaeus Gravenhorst, 1806, see: Quedius.

pygmaeus Snellen van Vollenhoven, 1854, see: Gabrius.

pyraster Erichson, 1839, see: Xanthopygus.

pyrenaeus G. Müller, 1924, see: Ocypus.

pyropterus Latreille, 1804, see: Gauropterus.

pyropterus Gravenhorst, 1806, see: Gauropterus.

pyrostomus Solier, 1849, see: Cheilocolpus.

quadraticeps Ménétriés, 1832, see: Ocypus.

quadriticeps Casey, 1924, see: Platydracus.

quadratus Paykull, 1789, see: Tetartopeus.

quadriimpressus White, 1846, see: Cafius.

quadrimaculatus Cameron, 1932, see: Ocypus.

quadripunctatus Zetterstedt, 1828, see: Quedius.

quartus Schaeffer, 1766 [Note: Nomen nudum], see: Ocypus.

quentzeli Paykull, 1789: 72 (Staphylinus; [Note: Nomen dubium]; Type locality: Scania).

  • Distribution: Sweden.

quintus Schaeffer, 1766 [Note: Nomen nudum], see: Emus.

quisquiliarius Gyllenhal, 1810, see: Philonthus.

raffrayi Fauvel, 1907: 35 (Staphylinus; Type locality: Bura [Wa Taita]; Kilimandjaro; Abyssinie [Azega, 2500 m]).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • — Cameron, 1952e: 832 (Staphylinus; Dahomey).

  • Distribution: Ethiopia, Tanzania, Benin.

rambouseki G. Müller, 1925, see: Ocypus.

regalis Erichson, 1839, see: Glenus.

reimoseri Bernhauer, 1906, see: Ocypus.

reitterianus Bernhauer, 1933, see: Platydracus.

relucens Gravenhorst, 1806, see: Megalinus.

rhodesianus Cameron, 1951c: 404 (Staphylinus; subgenus Abemus, cited as Abernus; Type locality: N.W. Rhodesia: Lusala; Kenya Colony: Ngong).

  • Distribution: Zambia, Kenya.

rhyssostomus Schrank, 1796: 119 (Staphylinus; [Note: Nomen dubium]; Type locality: not cited).

  • — Schrank, 1798: 648 (Staphylinus; characters; Boia).

  • Distribution: Germany.

rigidicornis Gravenhorst, 1802, see: Bisnius.

riparius Linné, 1758, see: Paederus.

rivularis Paykull, 1789, see: Omalium.

rossii Jarrige, 1954, see: Ocypus.

rotundicollis Ménétriés, 1832, see: Philonthus.

ruandae Bernhauer, 1934, see: Platydracus.

rubellus Villers, 1789: 425 (Staphylinus; [Note: Nomen dubium]; Type locality: Gallia).

  • Distribution: France.

rubricollis Gmelin, 1790: 2035 (Staphylinus; [Note: Nomen dubium]; Type locality: Gallia).

  • — Turton, 1802: 519 (Staphylinus; characters; France).

  • Distribution: France.

rubricornis Ádám, 1987: 136 (Ouchemus; replacement name for ruficornis Bernhauer).

  • — Smetana and Davies, 2000: 41 (Staphylinus).

  • Distribution: France, Austria, Czechoslovakia, Hungary, Ukraine, Caucasus, Turkey.

ruficornis Bernhauer, 1913a: 224 (Staphylinus; variety of caesareus; [preoccupied]; Type locality: Niederösterreich: Stockerau, Wienerwald; Ungarn: Ofenpest).

  • — Bernhauer and Schubert, 1914: 376 (Staphylinus; variety of caesareus).

  • — G. Müller, 1932: 78 (Staphylinus; characters).

  • — Scheerpeltz, 1933: 1388 (Staphylinus; catalog).

  • — Smetana, 1958: 277 (Staphylinus; subgenus Staphylinus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 196 (Staphylinus; characters; central Europe).

  • — Horion, 1965: 210 (Staphylinus; subgenus Staphylinus; Austria; Czech Republic; Hungary; Italy; Macedonia; Bulgaria).

  • — Smetana, 1968a: 158 (Staphylinus; Anatolia).

  • — Coiffait, 1974: 406 (Staphylinus; characters; Europe; Caucasus; Transcaucasus).

  • — Petrenko, 1980a: 61, 63 (Staphylinus; characters; Ukraine).

  • — Coiffait, 1984c: 267 (Staphylinus; characters; notes; France).

  • — Tóth, 1984: 22 (Staphylinus; characters; Hungary).

  • — Ádám, 1987: 136 (Ouchemus; synonym of rubricornis).

  • — Lucht, 1987: 107 (Staphylinus; checklist; Czechoslovakia; Austria).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Staphylinus; Czech Republic; Slovakia).

  • — Pilon, 1998: 98 (Staphylinus; notes; doubtfully in Italy).

  • — Smetana and Davies, 2000: 41 (Staphylinus; synonym of rubricornis).

rubripes Bernhauer and Schubert, 1914, see: Tasgius.

rubrothoracicus Goeze, 1777, see: Paederidus.

rufescens Gmelin, 1790: 2036 (Staphylinus; [Note: Nomen dubium]; Type locality: Europa).

  • — Zschach, 1788: 44 (Staphylinus; species number 979; characters stated but not named and referred to as rufescens by Gmelin).

  • — Turton, 1802: 519 (Staphylinus; characters; Europe).

  • Distribution: Europe.

ruficeps Cameron, 1932, see: Dinothenarus.

ruficollis Gravenhorst, 1806, see: Quedius.

ruficornis Latreille, 1804: 326 (Staphylinus; [Note: Nomen dubium]; Type locality: France).

  • Distribution: France.

ruficornis Gravenhorst, 1806, see: Quedius.

ruficornis O. Costa, 1839b: 118 (Staphylinus; [preoccupied]; Type locality: Cannavi).

  • — Herman, 2001: 52 (Staphylinus; nomen dubium; although a junior primary homonym, it is suggested that the required replacement be ignored).

  • Distribution: Italy.

ruficornis Bernhauer, 1913, see: rubricornis Ádám, 1987.

rufipennis Fabricius, 1801, see: Belonuchus.

rufipennis Gravenhorst, 1802, see: Hesperus.

rufipennis Solier, 1849, see: Philonthus.

rufipennis Cameron, 1930a: 156 (Staphylinus; [preoccupied]; Type locality: Malay Peninsula. Kedah; Catchment Area, near Jitra).

  • — Scheerpeltz, 1933: 1388 (Staphylinus; catalog).

  • — Cameron, 1937: 21 (Staphylinus; Java).

  • — Herman, 2001: 52 (Staphylinus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: Malaysia, Indonesia.

rufipes Linné, 1758, see: Tachinus.

rufipes Fabricius, 1793: 529 (Staphylinus; [preoccupied]; Type locality: Germania).

  • — Panzer, 1795: 357 (Staphylinus; characters; Germany).

  • — Fabricius, 1801: 600 (Staphylinus; characters; Germany).

  • — Laporte, 1840: 177 (Staphylinus; characters; France).

  • — Herman, 2001: 52 (Staphylinus; nomen dubium; although a junior primary homonym, it is suggested that the required replacement be ignored).

  • Distribution: France, Germany.

rufipes Gravenhorst, 1802, see: Quedius.

rufitarsis Marsham, 1802, see: Quedius.

rufocinctus Mannerheim, 1830, see: Quedius.

rufus Linné, 1758, see: Oxyporus.

rugifrons Runde, 1835, see: Ocypus.

rufilabris Zetterstedt, 1838, see: Acylophorus.

rugosipennis Schubert, 1911: 34 (Staphylinus; Type locality: Deutsch Ostafrika).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • Distribution: Tanzania.

rugosus Fabricius, 1775, see: Anotylus.

rupicola Kiesenwetter, 1850, see: Platydracus.

rutilus Perty, 1830, see: Eulissus.

rutilicauda Horn, 1879, see: Platydracus.

sachalinensis Matsumura, 1911, see: Platydracus.

sahlbergi G. Müller, 1926, see: Tasgius.

sallaei Sharp, 1884: 383 (Staphylinus; Type locality: Mexico, Parada).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • Distribution: Mexico.

salvinianus Sharp, 1884: 385 (Staphylinus; Type locality: Mexico, Jalapa, Cordova).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • Distribution: Mexico.

sanguineus Linné, 1758, see: Aleochara.

sanguinolentus Gravenhorst, 1802, see: Philonthus.

saphyreus Perty, 1830, see: Eulissus.

saphyrinus LeConte, 1861, see: Dinothenarus.

sapphirinus Erichson, 1839, see: Xanthopygus.

sapphirinus Gistel, 1857: 35 (Staphylinus; [preoccupied]; Type locality: Brasilia).

  • — Herman, 2001: 52 (Staphylinus; nomen dubium; although a junior primary homonym, it is suggested that the required replacement be ignored).

  • Distribution: Brazil.

saxatilis Schrank, 1781, see: Philonthus.

scabrosus Curtis, 1839a: 196 (Ocypus; Type locality: Gorrite).

  • — Erichson, 1840: 927 (Staphylinus; synonym of fuscicornis).

  • — Fauvel, 1877a: xxvii (Staphylinus; synonym of fuscicornis).

  • — Lynch, 1884: 138 (Staphylinus; synonym of fuscicornis).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; synonym of fuscicornis).

  • — Blackwelder, 1944: 139 (Staphylinus; synonym of nigrescens).

  • — Herman, 2001: 52 (Staphylinus; valid name for species).

  • Distribution: Brazil, Uruguay, Argentina.

fuscicornis Germar, 1824: 33 (Staphylinus; [preoccupied]; Type locality: Brasilia).

  • — Blanchard, 1842: 80 (Staphylinus; Argentina, Uruguay; Brazil).

  • — Erichson, 1839b: 389 (Staphylinus; characters; Brasilia meridionalis).

  • — Steinheil, 1869: 253 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 379, cited sepulchralis Steinheil as a synonym of fuscicornis; Steinheil did not describe the species as new, he attributed it to Erichson]; Argentina).

  • — Fauvel, 1877a: xxvii (Staphylinus; Brazil).

  • — Lynch, 1884: 138 (Staphylinus; characters; Argentina).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; catalog).

  • — Blackwelder, 1944: 139 (Staphylinus; synonym of nigrescens).

  • — Herman, 2001: 52 (Staphylinus; synonym of scabrosus).

lugubris Nordmann, 1837: 49 (Staphylinus; [preoccupied]; Type locality: Montevideo; Brasiliae).

  • — Erichson, 1839b: 389 (Staphylinus; synonym of fuscicornis).

  • — Fauvel, 1877a: xxvii (Staphylinus; synonym of fuscicornis).

  • — Lynch, 1884: 138 (Staphylinus; synonym of fuscicornis).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; synonym of fuscicornis).

nigrescens Blanchard, 1842: 79 (Staphylinus; Type locality: Montevideo).

  • — Fauvel, 1877a: xxvii (Staphylinus; synonym of fuscicornis).

  • — Lynch, 1884: 139 (Staphylinus; synonym of fuscicornis).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; synonym of fuscicornis).

  • — Blackwelder, 1944: 139 (Staphylinus; valid species).

  • — Herman, 2001: 52 (Staphylinus; synonym of scabrosus).

tristis Blanchard, 1842: 81 (Staphylinus; [preoccupied]; Type locality: Uruguay: Montevideo; Campos-Geraes).

  • — Fauvel, 1877a: xxvii (Staphylinus; synonym of fuscicornis).

  • — Lynch, 1884: 139 (Staphylinus; synonym of fuscicornis).

  • — Bernhauer and Schubert, 1914: 379 (Staphylinus; synonym of fuscicornis).

  • — Blackwelder, 1944: 139 (Staphylinus; synonym of nigrescens).

scenicus Erichson, 1839, see: Plociopterus.

schatzmayri G. Müller, 1923, see: Ocypus.

schultzei Bernhauer, 1915o: 366 (Staphylinus; Type locality: Belgisch-Kongo: Kimuenza).

  • — Scheerpeltz, 1933: 1388 (Staphylinus; catalog).

  • Distribution: Congo.

scintillans Gravenhorst, 1806, see: Quedius.

scitus Gravenhorst, 1806, see: Quedius.

scriptus Nordmann, 1837: 32 (Staphylinus; Type locality: Promontorium bonae spei).

  • — Erichson, 1839b: 375 (Staphylinus; characters; South Africa).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • Distribution: South Africa.

sculpticollis Gaubil, 1852 [Note: Nomen nudum], see: medioximus Fairmaire, 1852.

secundus Schaeffer, 1766 [Note: Nomen nudum], see: Ontholestes.

sedatus Sharp, 1887: 785 (Staphylinus; Type locality: Mexico, Zapotlan in Colima).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • Distribution: Mexico.

segmentarius Erichson, 1839, see: Oligotergus.

semialatus G. Müller, 1904, see: Ocypus.

semiauratus Schubert, 1911: 34 (Staphylinus; Type locality: Theresopolis, Brasilia).

  • — Bernhauer and Schubert, 1914: 382 (Staphylinus; catalog).

  • Distribution: Brazil.

semicyaneus Bernhauer, 1915, see: Platydracus.

semiobscurus Marsham, 1802, see: Quedius.

semipolitus Holme, 1842, see: Ocypus.

semipurpureus Kraatz, 1859, see: Platydracus.

semiviolaceus Cameron, 1932, see: Platydracus.

semotus Herman, 2001: 53 (Staphylinus; replacement name for subcyaneus Sharp).

  • Distribution: Brazil.

subcyaneus Sharp, 1876: 151 (Staphylinus; [preoccupied]; Type locality: Ega; Tunantins).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Herman, 2001: 53 (Staphylinus; synonym of semotus).

septimus Schaeffer, 1769 [Note: Nomen nudum], see: Paederus.

sepulchralis Erichson, 1839, see: Platydracus.

sericans Gravenhorst, 1802, see: Philonthus.

sericeicollis Ménétriés, 1832, see: Ocypus.

sericeomicans Bernhauer, 1931, see: Ocypus.

sericeus Marsham, 1802, see: Ocypus.

serraticornis Schrank, 1798, see: Velleius.

sextus Schaeffer, 1766 [Note: Nomen nudum], see: Philonthus.

sharpi Fauvel, 1901, see: Platydracus.

sibiricus Gebler, 1829, see: Dinothenarus.

sikkimensis Bernhauer, 1920, see: Tasgius.

silphoides Linné, 1767, see: Cilea.

similis Herbst, 1784, see: Stenus.

similis Paykull, 1789, see: Tasgius.

similis: Fabricius, 1793 [Note: Nomen nudum], see: Ocypus.

simplex Marsham, 1802, see: Philonthus.

singeri Huberthal, 1911, see: Tasgius.

sjostedti Fauvel, 1903, see: Platydracus.

socialis Paykull, 1789 [Note: This name is listed by Bernhauer and Schubert, 1926: 651, under the authorship of Erichson but Erichson, 1840, and Olivier, 1795: 37, cited Paykull as the author], see: Atheta.

solarii G. Müller, 1923, see: Ocypus.

solskyi Fauvel, 1875, see: Tasgius.

solwezianus Cameron, 1951c: 404 (Staphylinus; subgenus Staphylinus; Type locality: N.W. Rhodesia: Solwezi district).

  • Distribution: Zambia.

sordidus Gravenhorst, 1802, see: Bisnius.

sordidus Marsham, 1802, see: Atheta.

sparsus Cameron, 1932, see: Platydracus.

speculifrons Bernhauer, 1939, see: Platydracus.

splendens Fabricius, 1793, see: Philonthus.

splendens Marsham, 1802, see: Mycetoporus.

splendidulus Gravenhorst, 1802, see: Gabrius.

spinipes Pontoppidans, 1765, see: Tachinus.

spinipes Block, 1799, see: Aploderus.

springeri G. Müller, 1923, see: erythropterus Linné, 1758.

stenoides Gravenhorst, 1806, see: Leucitus.

stercorarius Olivier, 1795, see: Platydracus.

stevensi Cameron, 1932, see: Wasmannellus.

striatulus Fabricius, 1793, see: Coprophilus.

striatus Ström, 1768, see: Anotylus.

striatus Olivier, 1795, see: Carphacis.

strigatus Nordmann, 1837, see: Ocypus.

strumosus Fabricius, 1793, see: Lomechusa.

subaeneipennis Scheerpeltz, 1976, see: Platydracus.

subaeneus Sharp, 1874, see: Platydracus.

subapterus G. Müller, 1923 [Note: Nomen nudum], see: Ocypus.

subchalceus Cameron, 1930a: 156 (Staphylinus; Type locality: Malay Peninsula. Pahang; Bentong).

  • — Scheerpeltz, 1933: 1388 (Staphylinus; catalog).

  • Distribution: Malaysia.

subclavatus Goeze, 1777: 731 (Staphylinus; [Note: Nomen dubium]; Type locality: Not cited).

  • Distribution: ?.

subcyaneus Heer, 1839, see: Ocypus.

subcyaneus Sharp, 1876, see: semotus Herman, 2001.

subfuscus Gyllenhal, 1810, see: Philonthus.

subirideus Kraatz, 1859, see: Platydracus.

submarmorellus Schubert, 1908, see: Platydracus.

submetallicus LeConte, 1861, see: Platydracus.

subpunctatus Gyllenhal, 1827, see: Ocypus.

substriatus Paykull, 1790, see: Olisthaerus.

subterraneus Linné, 1758, see: Tachinus.

subterraneus Scopoli, 1771, see: fodinarum Gmelin, 1790.

subuliformis Gravenhorst, 1802, see: Bisnius.

subviridis Bernhauer, 1933, see: Platydracus.

sulcatus O. Müller, 1776, see: Oxytelus.

sulcatus Geoffroy, 1785, see: Anotylus.

sulcatus Paykull, 1789, see: Falagria.

sumakowi Bernhauer, 1911, see: Platydracus.

superbus Erichson, 1839, see: Triacrus.

suspectus Fauvel, 1904, see: Platydracus.

suspiciosus Bernhauer, 1937, see: Platydracus.

suturalis Marsham, 1802, see: Philonthus.

suturalis Turton, 1802, see: Tachinus.

suturalis Matsumura, 1911, see: Hadropinus.

szechuanensis Bernhauer, 1935, see: Dinothenarus.

tachiniformis Say, 1830, see: Philonthus.

tanalensis Fauvel, 1905b: 175 (Staphylinus; Type locality: forêt Tanala: Ivohimanitra).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • Distribution: Madagascar.

tarsalis Mannerheim, 1843, see: Platydracus.

tarsalis Sharp, 1884, see: Platydracus.

tataricus Pallas, 1773, see: Physetops.

tauricus G. Müller, 1932, see: Ocypus.

tectus Paykull, 1789, see: Deliphrum.

tempestivus Sharp, 1884: 377 (Staphylinus; Type locality: Mexico, near the city).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • Distribution: Mexico.

temporalis Casey, 1915, see: Platydracus.

tenebricosus Ganglbauer, 1846, see: Ocypus.

tenellus Gravenhorst, 1806, see: Quedius.

tenuis Fabricius, 1793, see: Rabigus.

tepidus Erichson, 1839, see: Xanthopygus.

terminalis Erichson, 1839, see: Oligotergus.

terminalis Laporte, 1840, see: Belonuchus.

terminatus Gravenhorst, 1802, see: Bisnius.

tertius Schaeffer, 1766 [Note: Nomen nudum], see: Creophilus.

tessellatus Geoffroy, 1785, see: Ontholestes.

testaceipes Fairmaire, 1887: 97 (Ocypus; Type locality: Yunnan).

  • — Bernhauer and Schubert, 1914: 391 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 47 (species unknown to authors).

  • Distribution: China.

testaceus Paykull, 1789, see: Lobrathium.

testaceus Fabricius, 1801, see: Nordus.

teter Bondroit, 1913, see: Platydracus.

tetracarinatus Block, 1799, see: Anotylus.

thoracicus Geoffroy, 1785, see: Paederidus.

thoracicus Villers, 1789: 420 (Staphylinus; [preoccupied]; Type locality: Kiliae).

  • — Herman, 2001: 48, 53 (Staphylinus; nomen dubium; although a junior primary homonym, it is suggested that the required replacement be ignored).

  • Distribution: Germany.

thoracicus Gravenhorst, 1802, see: Philonthus.

tomentosus Rossi, 1792, see: Sepedophilus.

tomentosus Rossi, 1792, see: Platydracus.

transadriaticus G. Müller, 1926, see: Ocypus.

transversiceps Luze, 1904, see: Tasgius.

tricinctus Aragona, 1830, see: Tasgius.

tricornis Herbst, 1784, see: Bledius.

tricolor Fabricius, 1787, see: Xantholinus.

tricolor Gravenhorst, 1802, see: Platydracus.

trilobus Olivier, 1795, see: Platystethus.

trimaculatus Fauvel, 1895: 252 (Staphylinus; Type locality: Bírmanie, Teinzo).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • — Cameron, 1932: 208 (Staphylinus; characters; Burma).

  • — Scheerpeltz, 1933: 1389 (Staphylinus; catalog).

  • — Herman, 2001: 31 (Staphylinus; valid name for species).

  • Distribution: Myanmar.

kublaikhani Muona, 1977: 15 (Staphylinus; replacement name for trimaculatus Fauvel).

  • — Herman, 2001: 31 (Staphylinus; unnecessary replacement name; synonym of trimaculatus).

triplicans Casey, 1924, see: Platydracus.

tristis Fabricius, 1793, see: Ocypus.

tristis Blanchard, 1842, see: scabrosus Curtis, 1839.

tuberculatus Fabricius, 1798, see: Bledius.

turcicus Bernhauer, 1923, see: Ocypus.

turfosus Schrank, 1796: 164 (Staphylinus; [Note: Nomen dubium]; Type locality: not cited).

  • — Schrank, 1798: 645 (Staphylinus; characters; Boia).

  • Distribution: Germany.

uheheanus Bernhauer, 1937, see: Platydracus.

ulmi Rossi, 1790, see: Astrapaeus.

ulmineus Fabricius, 1801, see: Astrapaeus.

umbratilis Gravenhorst, 1802, see: Philonthus.

umbrinus Gravenhorst, 1802, see: Philonthus.

unicolor Herbst, 1784, see: Ocypus.

unipunctatus Hope, 1831, see: Creophilus.

uralensis Mannerheim, 1830, see: Ocypus.

ussuriensis Solsky, 1871, see: Platydracus.

ustulatus Gravenhorst, 1802, see: Othius.

vagans Heer, 1839, see: Ocypus.

vagepunctus Gravenhorst, 1806, see: Xantholinus.

vagopunctatus Latreille, 1804, see: Xantholinus.

variabilis Gyllenhal, 1810, see: Quedius.

varians Paykull, 1789, see: Philonthus.

variegatum Geoffroy, 1785, see: Achenium.

varipes Sachse, 1852, see: Platydracus.

varius Gyllenhal, 1810, see: Philonthus.

velitaris Erichson, 1839, see: Nordus.

velox Runde, 1835: 9 (Staphylinus; [Note: Nomen dubium]; Type locality: Halae).

  • Distribution: Germany.

velutinus Nordmann, 1837, see: dimidiatus Laporte, 1835.

ventralis Gravenhorst, 1802, see: Philonthus.

ventralis Aragona, 1830, see: Quedius.

venustus Erichson, 1839, see: Plociopterus.

vernalis O. Müller, 1776, see: Tachyporus.

vernalis Gravenhorst, 1806, see: Gabrius.

versicolor Gravenhorst, 1806, see: Leistotrophus.

vestitus Sharp, 1884: 378 (Staphylinus; Type locality: Mexico, Cordova).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • Distribution: Mexico.

vethi Bernhauer, 1915k: 303 (Staphylinus; subgenus Abemus; Type locality: Transvaal: Johannesburg).

  • — Scheerpeltz, 1933: 1389 (Staphylinus; catalog).

  • Distribution: South Africa.

vetulus† Scudder, 1900: 52 (Staphylinus; fossil; Type locality: Florissant, Colorado).

  • — Handlirsch, 1907: 724 (Staphylinus; catalog; Miocene; Colorado).

  • Distribution: USA.

vetustus Sharp, 1876: 156 (Staphylinus; Type locality: Tunantins).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • Distribution: Brazil.

vicarius Sharp, 1889, see: Platydracus.

vicinus Ménétriés, 1832, see: Quedius.

viduatus Fabricius, 1801, see: Platydracus.

villosus Gravenhorst, 1802, see: Creophilus.

violaceovirens Fauvel, 1905, see: Platydracus.

violaceus Olivier, 1795, see: Plochionocerus.

violaceus Gravenhorst, 1802, see: Platydracus.

virgo Gravenhorst, 1802, see: Philonthus.

virgulatus Fauvel, 1895, see: Platydracus.

viridanus Horn, 1879, see: Platydracus.

vittatus Sharp, 1884: 374 (Staphylinus; Type locality: Mexico, Juquila, Parada, Uruapan).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • Distribution: Mexico.

volxemi Fauvel, 1877: xxvii (Staphylinus; Type locality: Brésil, Barbacena, S. Joao d'El Rey; Sta. Catharina).

  • — Bernhauer and Schubert, 1914: 383 (Staphylinus; catalog).

  • Distribution: Brazil.

vulcanus† Wickham, 1913: 288 (Staphylinus; fossil; Type locality: Florissant, Colorado).

  • Distribution: USA.

vulpinus Nordmann, 1837, see: Platydracus.

walkeri Fauvel, 1898, see: Tasgius.

wasmanni Bernhauer, 1920, see: Ocypus.

weingaertneri Bernhauer, 1928: 17 (Staphylinus; Type locality: Sumatra: Padangpadjang).

  • — Scheerpeltz, 1933: 1389 (Staphylinus; catalog).

  • — Cameron, 1933d: 391 (Staphylinus; Sumatra).

  • Distribution: Indonesia.

winkleri Bernhauer, 1906, see: Tasgius.

wittei Fagel, 1950, see: Platydracus.

xanthocephalus Kraatz, 1859, see: Dinothenarus.

xanthocephalus Motschulsky, 1860, see: Dinothenarus.

xantholoma Gravenhorst, 1806, see: Cafius.

xanthopygus Nordmann, 1837, see: Xanthopygus.

yolensis Cerruti, 1951, see: Platydracus.

yunnanensis Bernhauer, 1933a: 48 (Staphylinus; subgenus Abemus; Type locality: Yunnan-Sen).

  • Distribution: China.

yunnanensis Bernhauer, 1943, see: Platydracus.

zavattarii Gridelli, 1939, see: Platydracus.

zonatus Gravenhorst, 1802, see: Platydracus.

Tasgius [44 species; Palaearctic and Nearctic Regions]

Tasgius Stephens, 1829a: 22 (species included: rufipes). Type species: Astrapaeus rufipes Latreille, fixed by monotypy.

  • — Stephens, 1829: 276 (catalog; Britain).

  • — Stephens, 1832: 200, 213 (characters).

  • — Curtis, 1833: pl. 438 (characters; type species).

  • — Westwood, 1838a: 16 (characters; type species).

  • — Erichson, 1839a: 439 (synonym of Ocypus).

  • — Erichson, 1840: 415 (synonym of Ocypus).

  • — Stephens, 1839: 384 (characters).

  • — Lacordaire, 1854: 79 (synonym of Ocypus).

  • — Jacquelin du Val, 1856: 35 (characters).

  • — Kraatz, 1857d: 550 (synonym of Ocypus).

  • — Thomson, 1859: 24 (characters; type species: ater).

  • — Thomson, 1860: 149 (characters).

  • — Wencker and Silbermann, 1866: 29 (list of species; collecting notes; France).

  • — Jacquelin du Val, 1868: 67 (group of Staphylinus; catalog; European species).

  • — Fauvel, 1874: 172 [= 1874a: 398] (synonym of Staphylinus).

  • — Mulsant and Rey, 1876: 317 [= 1877: 173] (characters; key to French species).

  • — Heyden, 1880: 72 (subgenus of Staphylinus; list of species of Siberian region).

  • — Heyden, Reitter, and Weise, 1891: 102 (list of species of Europe and Caucasus).

  • — Heyden, 1893: 42 (list of species of Siberian region).

  • — Heyden, 1896: 32 (list of species of Siberian region).

  • — Heyden, Reitter, and Weise, 1906: 171 (subgenus of Staphylinus; list of species of Europe and Caucasus).

  • — Petri, 1912: 76 (subgenus of Staphylinus; list of species of Siebenbürgen).

  • — Bernhauer and Schubert, 1914: 374 (subgenus of Staphylinus).

  • — Scheerpeltz, 1933: 1381 (subgenus of Staphylinus).

  • — Blackwelder, 1943: 444, 445 (subgenus of Ocypus; type species: rufipes).

  • — Tottenham, 1949: 374, 420 (type species: pedator Gravenhorst = rufipes Latreille; subgenus of Ocypus; notes; checklist of British species).

  • — Blackwelder, 1952: 374 (type species: rufipes).

  • — Jarrige, 1952a: 133 (type species: pedator).

  • — Coiffait, 1956e: 185, 198, 216 (subgenus of Ocypus; characters; notes; catalog of and key to species).

  • — Hatch, 1957: 176, 178 (subgenus of Ocypus; characters; species of Pacific Northwest).

  • — Smetana, 1958: 279, 295 (subgenus of Ocypus; characters).

  • — Lohse, 1964: 201 (subgenus of Ocypus; characters; central European species).

  • — Coiffait, 1964: 81, 82 (type species: pedator; characters; key to subgenera).

  • — Pototskaia, 1966b: 362 (subgenus of Staphylinus; larval characters in key).

  • — Pototskaia, 1967: 66 (subgenus of Staphylinus; larval characters).

  • — Scheerpeltz, 1968a: 84 (catalog of Austrian species).

  • — Bordoni, 1973j: 702 (list of species from islands near Sicily).

  • — Tikhomirova, 1973a: 187 (subgenus of Ocypus; checklist of species of USSR).

  • — Coiffait, 1974: 515 (adult and larval characters; key to species of western Palaearctic region).

  • — Segers, 1986: 48 (checklist of species; Belgium).

  • — Drugmand 1987b: 16, 70 (distribution of Belgian species).

  • — Smetana and Davies, 2000: 20, 45 (characters; checklist of species of north temperate region).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 395 (3 Nearctic species; notes; characters in key).

Anodus Nordmann, 1837: 11 (species included: messor, falcifer, morio preoccupied by Spix, 1829). Type species: Staphylinus morio Gravenhorst, fixed by subsequent designation by Thomson, 1859: 24.

  • — Erichson, 1840: 416 (synonym of Ocypus).

  • — Lacordaire, 1854: 79 (synonym of Ocypus).

  • — Kraatz, 1857d: 550 (synonym of Ocypus).

  • — Thomson, 1859: 24 (characters; type species: morio).

  • — Thomson, 1860: 150 (characters).

  • — Wencker and Silbermann, 1866: 29 (list of species; collecting notes; France).

  • — Fauvel, 1874: 172 [= 1874a: 398] (synonym of Staphylinus).

  • — Mulsant and Rey, 1876: 325 [= 1877: 181] (characters; key to French species).

  • — Seidlitz, 1891: 392 (key to some European species).

  • — Seidlitz, 1891a: 418 (key to some European species).

  • — Heyden, Reitter, and Weise, 1891: 102 (subgenus of Ocypus; list of species of Europe and Caucasus).

  • — Blackwelder, 1943: 444, 445 (synonym of Ocypus; type species: falcifer).

  • — G. Müller, 1943: 106 (synonym of Ocypus).

  • — Tottenham, 1949: 374 (synonym of Alapsodus; type species: falcifer Nordmann).

  • — Blackwelder, 1952: 53 (synonym of Ocypus; type species: morio).

  • — Smetana, 1958: 279 (synonym of Ocypus).

  • — Coiffait, 1974: 526 (synonym of Alapsodus).

  • — Smetana and Davies, 2000: 36, 46 (synonym of Tasgius subgenus Rayacheila).

Rayacheila Motschulsky, 1845a: 40 (species included: inderiensis). Type species: Rayacheila inderiensis Motschulsky, fixed by monotypy.

  • — Blackwelder, 1952: 337 (type species: inderiensis).

  • — Smetana, 1958: 279 (synonym of Ocypus).

  • — Smetana and Davies, 2000: 20, 36, 46 (subgenus of Tasgius; characters).

Rhagochila Motschulsky, 1858: 668 (emendation of Rayacheila). Type species: Rayacheila inderiensis Motschulsky, fixed by objective synonymy with Rayacheila.

  • — Gemminger and Harold, 1868: 600 (catalog).

  • — Fauvel, 1874: 172 [= 1874a: 398] (synonym of Staphylinus).

  • — Heyden, 1880: 72 (subgenus of Staphylinus; list of species of Siberian region).

  • — Bernhauer and Schubert, 1914: 374 (synonym of Goerius).

  • — Blackwelder, 1943: 444, 445 (subgenus of Goerius; type species: inderiensis).

  • — Blackwelder, 1952: 336 (type species).

  • — Smetana, 1958: 279 (synonym of Ocypus).

Pseudotasgius Seidlitz, 1891a: 418 (species included: pedator). Type species: Staphylinus pedator Gravenhorst, fixed by monotypy.

  • — Seidlitz, 1891: 392 (characters; key to some European species).

  • — Heyden, Reitter, and Weise, 1891: 102 (subgenus of Ocypus; list of species of Europe and Caucasus).

  • — Reitter, 1909: 122 (synonym of Tasgius).

  • — Blackwelder, 1952: 330 (synonym of Tasgius; type species: pedator).

  • — Smetana and Davies, 2000: 36, 45 (synonym of Tasgius subgenus Tasgius).

Alapsodus Tottenham, 1939d: 225 (replacement name for Anodus Nordmann). Type species: Staphylinus morio Gravenhorst, fixed by objective synonymy with Anodus.

  • — Blackwelder, 1943: 444, 445 (synonym of Anodus; type species: falcifer).

  • — Tottenham, 1949: 374, 420 (type species: falcifer Nordmann; checklist of British species).

  • — Blackwelder, 1952: 42 (type species).

  • — Hatch, 1957: 176, 177 (subgenus of Ocypus; characters).

  • — Smetana, 1958: 279, 298 (subgenus of Ocypus).

  • — Coiffait, 1964: 81, 82, 83 (type species: falcifer; characters; key to subgenera).

  • — Pototskaia, 1966b: 362 (subgenus of Staphylinus; larval characters in key).

  • — Pototskaia, 1967: 66 (subgenus of Staphylinus; larval characters).

  • — Tikhomirova, 1973a: 187 (checklist of species of USSR).

  • — Coiffait, 1974: 526 (characters; key to species of western Palaearctic region).

  • — Segers, 1986: 48 (checklist of species; Belgium).

  • — Drugmand 1987b (comments on distribution of Belgian species).

  • — Silfverberg, 1992: 17 (cited as synonym of Ocypus).

  • — Smetana, 1993b: 49 (subgenus of Staphylinus; list of species of Czech Republic and Slovakia).

  • — Smetana and Davies, 2000: 36, 46 (synonym of Tasgius subgenus Rayacheila).

Paratasgius Jarrige, 1952a: 133 (subgenus of Staphylinus; species included: ater, ater olivieri, peyerimhoffi, atronitidus). Type species: Staphylinus ater Gravenhorst, fixed by original designation.

  • — Coiffait, 1964: 82 (characters; subgenus of Tasgius; type species: ater).

  • — Coiffait, 1974: 515 (subgenus of Tasgius; characters; key to species).

  • — Smetana and Davies, 2000: 36, 45 (synonym of Tasgius subgenus Tasgius).

Allocypus Coiffait, 1964: 83 (subgenus of Alapsodus; species included: winkleri, melanarius, minax, eppelsheimianus). Type species: Staphylinus winkleri Bernhauer, fixed by original designation; [Note: This subgenus is a subjective isogenotypic synonym of Alapsodus].

  • — Coiffait, 1974: 526 (subgenus of Alapsodus; characters; key to species).

  • — Ádám, 1987: 149 (cited as valid genus; list of two species with collecting notes; Hungary).

  • — Smetana and Davies, 2000: 36, 46 (synonym of Tasgius subgenus Rayacheila).

Metocypus Coiffait, 1964: 81, 82, 83 (species included: globulifer, nigrinus, arrowi, lusitanicus, rubripennis, walkeri, caspius, dioriae, pliginskii, bucharicus). Type species: Staphylinus globulifer Geoffroy, fixed by original designation.

  • — Coiffait, 1974: 546 (adult and larval characters; key to species of western Palaearctic region).

  • — M. Dvořák, 1984: 66 (synonym of Alapsodus).

  • — Segers, 1986: 49 (checklist of species; Belgium).

  • — Drugmand 1987b (comments on distribution of Belgian species).

  • — Smetana and Davies, 2000: 36, 46 (synonym of Tasgius subgenus Rayacheila).

Paralapsodus Coiffait, 1974: 526, 527, 541 (subgenus of Alapsodus; species included: solskyi). Type species: Staphylinus solskyi Fauvel, fixed by original designation and monotypy.

  • — Smetana and Davies, 2000: 36, 46 (synonym of Tasgius subgenus Rayacheila).

abbreviatus Motschulsky, 1858, see: Ocypus.

aegyptiacus C. Koch, 1936, see: peyerimhoffi G. Müller, 1926.

algericus Coiffait, 1964: 105 (Alapsodus; Type locality: Algérie: Kabylie, Bou Bérak).

  • — Coiffait, 1974: 539 (Alapsodus; characters; Algeria).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Algeria.

aliquoi Bordoni, 1976, see: falcifer Nordmann, 1837.

amiculus G. Müller, 1925: 47 (Staphylinus; subgenus Tasgius; Type locality: Turkestan: Osch nella provincia Ferganah).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; catalog).

  • — Coiffait, 1964: 83 (Tasgius; subgenus Paratasgius).

  • — Coiffait, 1974: 523 (Tasgius; subgenus Paratasgius; characters; Turkestan).

  • — Smetana and Davies, 2000: 45 (Tasgius; subgenus Tasgius).

  • Distribution: Uzbekistan.

amoenanus Smetana and Davies, 2000: 37, 46 (Tasgius; replacement name for amoenus).

  • Distribution: Russia, Kazakhstan.

amoenus Reitter, 1909: 122 (Staphylinus; subgenus Ocypus; [preoccupied]; Type locality: Südrussland: Uralsk).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; catalog).

  • — Coiffait, 1974: 538, 545 (Alapsodus; characters; Russia; Kazakhstan).

  • — Smetana and Davies, 2000: 37, 46 (Tasgius; synonym of amoenanus).

amoenus Reitter, 1909, see: amoenanus Smetana and Davies, 2000.

angustatus Lacordaire, 1835, see: melanarius Heer, 1839.

arrowi G. Müller, 1932: 84, 87 (Staphylinus; subgenus Ocypus; Type locality: Graecia: Corfu).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; catalog).

  • — G. Müller, 1943: 109 (Ocypus; notes; Albania).

  • — Coiffait, 1964: 83 (Metocypus).

  • — Coiffait, 1974: 553 (Metocypus; characters; Corfu; Greece; Albania; Turkey).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Greece, Albania, Turkey.

ater Gravenhorst, 1802: 161 (Staphylinus; Type locality: America septentr.).

  • — Gravenhorst, 1806: 112 (Staphylinus; characters; Germany).

  • — Gyllenhal, 1827: 474 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 384, cited fuscatus Gyllenhal as a synonym of ater Gravenhorst; Gyllenhal did not describe the species as new, he attributed it to Gravenhorst]; characters).

  • — Say, 1830: 38 (Staphylinus; characters).

  • — Stephens, 1832: 209 (Goerius; [Note: Misidentification: Bernhauer and Schubert, 1914: 384, cited punctulatus Stephens as a synonym of ater Gravenhorst; Stephens did not describe the species as new, he attributed it to Marsham who also misidentified (see Pseudocypus fuscatus) it but had ascribed it to Paykull]; characters; England).

  • — C. Sahlberg, 1830: 309 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 384, cited morio C. Sahlberg as a synonym of ater Gravenhorst; Sahlberg did not describe the species as new, he attributed it to Gravenhorst]; characters; Finland).

  • — Say, 1834: 452 (Staphylinus; characters).

  • — Erichson, 1839a: 444 (Ocypus; characters; Germany).

  • — Heer, 1839: 256 (Staphylinus; characters; Switzerland).

  • — Erichson, 1840: 416 (Ocypus; characters; Europe; montes Uralenses; America septentrionalis).

  • — Jacobsen, 1849: 686 (Ocypus; characters; Denmark).

  • — Motschulsky, 1849: 87 (Tasgius; Spain).

  • — Redtenbacher, 1849: 697 (Ocypus; characters; Austria).

  • — Hardy, 1851: 24 (Ocypus; subgenus Tasgius; notes; England).

  • — Rosenhauer, 1856: 70 (Ocypus; Malaga).

  • — Fairmaire and Laboulbène, 1856: 512 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 562 (Ocypus; characters; Germany).

  • — Redtenbacher, 1857: 186 (Ocypus; characters; Austria).

  • — Thomson, 1859: 24 (type species of Tasgius).

  • — Thomson, 1860: 149 (Tasgius; characters; Scandinavia).

  • — Fairmaire and Coquerel, 1860: 158 (Ocypus; Morocco).

  • — Hochhuth, 1862: 51 (Ocypus; notess; Caucasus).

  • — Fauvel, 1874: 192 [= 1874a: 418] (Staphylinus; characters; France; Caucasus; North America).

  • — Redtenbacher, 1874: 206 (Ocypus; characters; Austria).

  • — Seidlitz, 1875: 270 (Tasgius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 323 [= 1877: 179] (Tasgius; characters; France).

  • — J. Sahlberg, 1876: 12 (Tasgius; Finland).

  • — Provancher, 1877: 248 (Tasgius; characters; Quebec).

  • — Fauvel, 1878: 119 [= 1878a: 39] (Staphylinus; Algeria; Tunisia).

  • — Fauvel, 1886: 51 [= 1886a: 43] (Staphylinus; Morocco; Tunisia; Algeria).

  • — Fowler, 1888: 257 (Ocypus; characters; Britain).

  • — Seidlitz, 1891: 393 (Ocypus; subgenus Tasgius; characters; Europe).

  • — Seidlitz, 1891a: 418 (Ocypus; subgenus Tasgius; characters; Europe).

  • — Ganglbauer, 1895: 435 (Staphylinus; subgenus Tasgius; characters; Palaearctic region; North America).

  • — Fauvel, 1902b: 105 (Staphylinus; subgenus Ocypus; Morocco; Algeria; Tunisia).

  • — Sainte-Claire Deville, 1906: 100 (Staphylinus; Corsica).

  • — Reitter, 1909: 122 (Staphylinus; subgenus Tasgius; characters; Germany).

  • — Blatchley, 1910: 379 (Tasgius; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; subgenus Ocypus; catalog).

  • — Johansen, 1914: 386 (Staphylinus; subgenus Tasgius; characters; Denmark).

  • — G. Müller, 1923: 142 (Staphylinus; subgenus Tasgius; characters; Italy).

  • — Lea, 1925: 228 (Ocypus; notes; Australian introduction).

  • — Portevin, 1929: 354 (Staphylinus; characters; France).

  • — Blair, 1931 (Ocypus; pupation cell).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; catalog).

  • — Aleksandrov, 1934: 153 (Staphylinus; China).

  • — Cameron, 1951: 283 (Staphylinus; subgenus Tasgius; Egypt).

  • — Jarrige, 1952a: 133 (Staphylinus; subgenus Paratasgius; must be an introduction to North America).

  • — V. Hansen, 1952: 105 (Staphylinus; subgenus Tasgius; characters; Denmark).

  • — Coiffait, 1956e: 216, 217 (Ocypus; subgenus Tasgius; characters).

  • — Hatch, 1957: 178 (Ocypus; subgenus Tasgius; characters; British Columbia; Idaho; Washington; Oregon).

  • — Smetana, 1958: 297 (Ocypus; subgenus Tasgius; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 117 (Staphylinus; subgenus Tasgius; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 201 (Ocypus; subgenus Tasgius; characters; central Europe).

  • — Coiffait, 1964: 83 (Tasgius; subgenus Paratasgius).

  • — Moore, 1965: 102 (Ocypus; introduced in North America).

  • — Smetana, 1965: 32 (Ocypus; subgenus Tasgius; characters; Labrador; Nova Scotia; Newfoundland).

  • — Smetana, 1965a: 11 (Ocypus; subgenus Tasgius; British Columbia).

  • — Horion, 1965: 228 (Tasgius; Morocco; Portugal; Spain; Corsica; Sardinia; Sicily; Italy; Istria; Holland; Belgium; France; Switzerland; Asutria; Germany; Czechoslovakia; Hungary; Poland; Russia; Finland; Sweden; Norway; Denmark; Britain).

  • — Szujecki, 1968a: 733 (Ocypus; Poland).

  • — Coiffait, 1974: 522 (Tasgius; subgenus Paratasgius; characters; Holarctic region).

  • — Orth, Moore, Fisher and Legner, 1975: 1115 (Ocypus; characters).

  • — Shilov, 1975a: 376 [= 1975b: 99] (Ocypus; Komi region of Russia).

  • — Pope, 1977: 30 (Staphylinus; Britain).

  • — Hoebeke, 1978: 508 (Ocypus; parasitized by proctotrupid).

  • — Frank, 1982: 37 (Ocypus; list of parasites).

  • — Tóth, 1984: 31 (Ocypus; characters; Hungary).

  • — Newton, 1987: 381 (Staphylinus; an introduction, not an indigenous North American species).

  • — Lucht, 1987: 108 (Ocypus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Welch, 1993: 229 (Staphylinus; ovariole number and ovary structure).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Tasgius; Czech Republic; Slovakia).

  • — Hernández, Outerelo, and Gamarra, 1994: 188 (Tasgius; checklist of species; Canary Islands).

  • — Outerelo, Palmer and Pons, 1995: 79 (Tasgius; Mallorca).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; Italy).

  • — Downie and Arnett, 1996: 384 (Ocypus; characters; USA).

  • — Pilon, 1998: 92 (Ocypus; ecological notes; Italy).

  • — Whitehead, 1999: 22 (Ocypus; habitat notes; Britain).

  • — Owen, 2000: 251 (Staphylinus; pitfall trap collections from underground at roots of trees; Britain).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Tasgius).

  • Distribution: Egypt, Tunisia, Algeria, Morocco, Canary Islands, Europe, Russia, Iran, Syria, Canada, USA.

obscurus Marsham, 1802: 514 (Staphylinus; [preoccupied]; Type locality: Britain).

  • — Gyllenhal, 1827: 474 (Staphylinus; synonym of fuscatus Gravenhorst).

  • — Curtis, 1829: 24 (Ocypus; catalog; Britain).

  • — Erichson, 1840: 416 (Ocypus; synonym of ater).

  • — Kraatz, 1857d: 562 (Ocypus; synonym of ater).

  • — Ganglbauer, 1895: 435 (Staphylinus; synonym of ater).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; synonym of ater).

  • — Smetana, 1958: 297 (Ocypus; synonym of ater).

  • — Coiffait, 1974: 523 (Tasgius; synonym of ater).

protensus Ménétriés, 1832: 144 (Staphylinus; Type locality: Caucase).

  • — Faldermann, 1835: 122 (Staphylinus; characters; Persia-Armenia).

  • — Nordmann, 1837: 73 (Staphylinus; characters; Caucasus).

  • — Hochhuth, 1849: 128 (Ocypus; characters).

  • — Eppelsheim, 1887: 425 (Ocypus; characters; Siberia).

  • — Ganglbauer, 1895: 435 (Staphylinus; synonym of ater).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; synonym of ater).

  • — Scheerpeltz, 1933: 1391 (Staphylinus; doubtful synonym of ater).

  • — Smetana, 1958: 297 (Ocypus; synonym of ater).

confinis Curtis, 1833: pl. 438 (Tasgius; Type locality: England: Essex).

  • — Curtis, 1829: 24 (Goerius; nomen nudum; catalog; Britain).

  • — Stephens, 1839: 389 (Tasgius; characters; England).

  • — Erichson, 1840: 416 (Ocypus; synonym of ater).

  • — Redtenbacher, 1849: 697 (Ocypus; synonym of ater).

  • — Hardy, 1851: 24 (Staphylinus; synonym of ater).

  • — Kraatz, 1857d: 562 (Ocypus; synonym of ater).

  • — Ganglbauer, 1895: 435 (Staphylinus; synonym of ater).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; synonym of ater).

  • — Smetana, 1958: 297 (Ocypus; synonym of ater).

  • — Coiffait, 1974: 523 (Tasgius; synonym of ater).

nigripes Lacordaire, 1835: 370 (Emus; Type locality: Paris).

  • — Erichson, 1839a: 444 (Ocypus; synonym of ater).

  • — Heer, 1839: 256 (Staphylinus; synonym of ater).

  • — Erichson, 1840: 416 (Ocypus; synonym of ater).

  • — Redtenbacher, 1849: 697 (Ocypus; synonym of ater).

  • — Fairmaire and Laboulbène, 1856: 512 (Staphylinus; synonym of ater).

  • — Kraatz, 1857d: 562 (Ocypus; synonym of ater).

  • — Ganglbauer, 1895: 435 (Staphylinus; synonym of ater).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; synonym of ater).

  • — Smetana, 1958: 297 (Ocypus; synonym of ater).

  • — Coiffait, 1974: 523 (Tasgius; synonym of ater).

punctatissimus Dufour, 1843: 33 (Staphylinus; Type locality: vallée d'Ossau).

  • — Gemminger and Harold, 1868: 581 (Ocypus; synonym of ater).

  • — Ganglbauer, 1895: 435 (Staphylinus; synonym of ater).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; synonym of ater).

  • — Smetana, 1958: 297 (Ocypus; synonym of ater).

  • — Coiffait, 1974: 523 (Tasgius; synonym of ater).

gagates Baudi, 1848: 123 (Ocypus; Type locality: Syria prope Damascum).

  • — Baudi, 1870: 385 (Ocypus; Syria).

  • — Fauvel, 1875a: XXVIII [= 1875b: 230] (Staphylinus; synonym of planipennis).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; synonym of ater).

  • — Coiffait, 1974: 523 (Tasgius; synonym of ater).

atratus Wollaston, 1864: 567 (Ocypus; Type locality: Lanzarote; Fuerteventura).

  • — Wollaston, 1865: 489 (Ocypus; habitat; notes; Canary Islands).

  • — Wollaston, 1867: 278 (Ocypus; may be synonym of or near ater; notes).

  • — Fauvel, 1869: 487 (Ocypus; synonym of ater).

  • — Fauvel, 1869a: 151 (Ocypus; synonym of ater).

  • — Wollaston, 1871: 300 (Ocypus; characters; notes).

  • — Fauvel, 1875a: XXVIII [= 1875b: 230] (Staphylinus; synonym of planipennis).

  • — Fauvel, 1902b: 105 (Staphylinus; synonym of ater).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; synonym of ater).

  • — Scheerpeltz, 1933: 1391 (Staphylinus; doubtful synonym of ater).

  • — Smetana, 1958: 297 (Ocypus; synonym of ater).

fulvopubescens Chapman, 1933: 194 (Staphylinus; subgenus Tasgius; variety of ater; Type locality: Seine-Inférieure: Canteleu).

  • — Coiffait, 1956e: 216 (Ocypus; subgenus Tasgius; aberration of ater; characters).

  • — Coiffait, 1974: 523 (Tasgius; variety of ater).

atratus Wollaston, 1864, see: ater Gravenhorst, 1802.

atronitidus Reitter, 1909: 122 (Staphylinus; subgenus Tasgius; Type locality: Cypern).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; subgenus Ocypus; catalog).

  • — Jarrige, 1952a: 133 (Staphylinus; subgenus Paratasgius).

  • — Coiffait, 1964: 83 (Tasgius; subgenus Paratasgius).

  • — Coiffait, 1974: 524 (Tasgius; subgenus Paratasgius; characters; Cyprus).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Tasgius).

  • Distribution: Cyprus.

bellicosus Fairmaire, 1855: 312 (Ocypus; Type locality: Tanger).

  • — Fairmaire and Coquerel, 1860: 158 (Ocypus; characters; Morocco; Algeria).

  • — Fauvel, 1878: 120 [= 1878a: 40] (Staphylinus; Morocco; Algeria).

  • — Fauvel, 1886: 52 [= 1886a: 44] (Staphylinus; Morocco; Algeria).

  • — Fauvel, 1902b: 105 (Staphylinus; subgenus Ocypus; Morocco; Algeria; Tunisia).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1391 (Staphylinus; catalog).

  • — Coiffait, 1973a: 274 (Alapsodus; Morocco).

  • — Coiffait, 1974: 539 (Alapsodus; characters; Iberian Peninsula; Morocco; Algeria; Tunisia).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: “Iberia”, Morocco, Algeria, Tunisia.

bicolor Cameron, 1944, see: dichromus Herman, 2001.

bonnairei Scheerpeltz, 1933, see: pedator Gravenhorst, 1802.

bosphoranus G. Müller, 1926, see: solskyi Fauvel, 1875.

bucharicus Bernhauer, 1912c: 21 (Staphylinus; subgenus Ocypus; Type locality: Buchara).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1392 (Staphylinus; catalog).

  • — Coiffait, 1964: 83 (Metocypus).

  • — Coiffait, 1974: 556 (Metocypus; characters; Buchara).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Uzbekistan.

bulgaricus Coiffait, 1971b: 282 (Alapsodus; subgenus Allocypus; Type locality: Bulgarie: Sofia).

  • — Coiffait, 1974: 545 (Alapsodus; subgenus Allocypus; characters; Bulgaria).

  • — Raichev, 1979: 83 (Alapsodus; Bulgaria).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Bulgaria.

caspius Bernhauer, 1906: 125 (Staphylinus; subgenus Ocypus; Type locality: Kaschgar, Transcaspigebiet).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1392 (Staphylinus; catalog).

  • — G. Müller, 1943: 106 (Ocypus; characters; Transcaspian region).

  • — Coiffait, 1964: 83 (Metocypus).

  • — Coiffait, 1974: 557 (Metocypus; characters; Transcaspian region).

  • — M. Dvořák, 1984: 67 (Alapsodus; Kirghizia).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Kyrgyzstan, “Transcaspia”.

cerdo Erichson, 1840, see: see: morsitans Rossi, 1790.

chalybeipennis Ménétriés 1832, see: pedator Gravenhorst, 1802.

compressus Marsham, 1802, see: morsitans Rossi, 1790.

confinis Curtis, 1833, see: ater Gravenhorst, 1802.

depressus Hochhuth, 1849: 126 (Ocypus; Type locality: Lenkoran).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1393 (Staphylinus; catalog).

  • — Coiffait, 1974: 536 (Alapsodus; characters; Caucasus).

  • — Gusarov, 1993: 72 (Ocypus; subgenus Alapsodus; confirm identification).

  • — Solodovnikov, 1998a: 14 (Ocypus; notes; Russia).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Azerbaijan.

gracilipalpis Eppelsheim, 1878: 109 (Ocypus; Type locality: Hochgebirge östlich vom Kasbek; Michailowo).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of depressus).

  • — Coiffait, 1974: 536 (Alapsodus; synonym of depressus).

deuvei Coiffait, 1984e: 379 (Tasgius; Type locality: Nepal oriental: Jaljale Himal, Deourali Bhanzyand 3500 m).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Tasgius).

  • Distribution: Nepal.

dichromus Herman, 2001: 53 (Staphylinus; replacement name for bicolor Cameron).

  • Distribution: Pakistan.

bicolor Cameron, 1944: 11 (Staphylinus; subgenus Ocypus; [preoccupied]; Type locality: Punjab: Lyallpur).

  • — Smetana and Davies, 2000: 37, 46 (Tasgius; subgenus Tasgius).

  • — Herman, 2001: 53 (Tasgius; synonym of dichromus).

doriae G. Müller, 1932: 82, 83 (Staphylinus; subgenus Ocypus; Type locality: Persia septentrionalis).

  • — Scheerpeltz, 1933: 1393 (Staphylinus; catalog).

  • — Coiffait, 1964: 83 (Metocypus; cited as dioriae).

  • — Coiffait, 1974: 555 (Metocypus; characters; Persia).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Iran.

dvoraki Roubal, 1940, see: melanarius Heer, 1839.

edentulus Block, 1799, see: winkleri Bernhauer, 1906.

eppelsheimianus Jakobson, 1909: 510 (Staphylinus; subgenus Ocypus; replacement name for rufipes Eppelsheim).

  • — Scheerpeltz, 1933: 1393 (Staphylinus; catalog).

  • — Coiffait, 1964: 83 (Alapsodus; subgenus Allocypus).

  • — Smetana, 1965c: 45 (Ocypus; subgenus Alapsodus; Anatolia).

  • — Coiffait, 1974: 542 (Alapsodus; subgenus Allocypus; characters; Caucasus; Transcaucasus; Anatolia).

  • — Solodovnikov, 1998a: 14 (Ocypus; notes; Russia, Krasnodar territory).

  • — Smetana and Davies, 2000: 37, 46 (Tasgius; subgenus Rayacheila).

  • — Herman, 2001: 31 (Tasgius; notes).

  • Distribution: Turkey, Azerbaijan.

rufipes Eppelsheim, 1884b: 15 (Ocypus; subgenus Anodus; [preoccupied]; Type locality: Lenkoran).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; synonym of rubripes).

  • — Scheerpeltz, 1933: 1393 (Staphylinus; synonym of eppelsheimianus).

  • — Smetana, 1965c: 45 (Ocypus; synonym of eppelsheimianus).

  • — Coiffait, 1974: 542 (Alapsodus; synonym of eppelsheimianus).

  • — Smetana and Davies, 2000: 37 (Tasgius; synonym of eppelsheimianus).

obscuripes Bernhauer, 1900b: 55 (Ocypus; variety of rufipes; Type locality: Kleinasien, Sabandja).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; synonym of rubripes).

  • — Scheerpeltz, 1933: 1393 (Staphylinus; synonym of eppelsheimianus).

  • — Coiffait, 1974: 542 (Alapsodus; variety of eppelsheimianus; Anatolia).

  • — Smetana and Davies, 2000: 37 (Tasgius; although this name is older than eppelsheimianus its status is unclear at present so it should remain a variety pending study of the type).

  • — Herman, 2001: 31 (Tasgius; may not be conspecific with eppelsheimianus).

rubripes Bernhauer and Schubert, 1914: 390 (Staphylinus; subgenus Ocypus; replacement name for rufipes Eppelsheim).

  • — Scheerpeltz, 1933: 1393 (Staphylinus; synonym of eppelsheimianus).

  • — Smetana, 1965c: 45 (Ocypus; synonym of eppelsheimianus).

  • — Coiffait, 1974: 542 (Alapsodus; synonym of eppelsheimianus).

erosicollis Reiche and Saulcy, 1856, see: winkleri Bernhauer, 1906.

etruscus Gautier, 1860, see: tricinctus Aragona, 1830.

evitendus Tottenham, 1945, see: globulifer Geoffroy, 1785.

falcifer Nordmann, 1837: 12 (Anodus; Type locality: Italia).

  • — Erichson, 1840: 418 (Ocypus; characters; Italy).

  • — Hochhuth, 1849: 126 (Ocypus; characters; Nähe Akhatziks).

  • — Redtenbacher, 1849: 696 (Ocypus; characters; Austria).

  • — Kraatz, 1857d: 565 (Ocypus; characters; Germany).

  • — Redtenbacher, 1857: 186 (Ocypus; characters; Austria).

  • — Fauvel, 1874: 193 [= 1874a: 419] (Staphylinus; characters; Austria; Italy; Sicily; Sardinia; Caucasus).

  • — Redtenbacher, 1874: 205 (Ocypus; characters; Austria).

  • — Fauvel, 1876a: 72 [= 1876: 257] (Staphylinus; Turkey; Crimea).

  • — Mulsant and Rey, 1876: 335 [= 1877: 191] (Anodus; characters; France).

  • — Ganglbauer, 1895: 435 (Staphylinus; subgenus Ocypus; characters; Austria; Italy; Piemont; Sardinia; Sicily; Crimea; Caucasus).

  • — Reitter, 1909: 122 (Staphylinus; subgenus Ocypus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1393 (Staphylinus; catalog).

  • — Coiffait, 1956e: 218, 219 (Ocypus; subgenus Ocypus; characters).

  • — Lohse, 1964: 201 (Ocypus; subgenus Ocypus; characters; central Europe).

  • — Focarile, 1964: 61 (Staphylinus; Italy).

  • — Scheerpeltz, 1965c: 509 (Ocypus; Sicily).

  • — Smetana, 1965c: 45 (Ocypus; subgenus Alapsodus; Anatolia).

  • — Horion, 1965: 231 (Ocypus; Caucasus; Turkey; Ukraine; Italy; France; Sardinia; Sicily; Austria; Czechoslovakia; Romania).

  • — Coiffait, 1974: 537 (Alapsodus; characters; Europe; Italy; Russia; Caucasus).

  • — Bordoni, 1976: 236 (Alapsodus; Turkey).

  • — Tóth, 1984: 32 (Ocypus; characters; Hungary).

  • — Lucht, 1987: 108 (Ocypus; checklist; Austria).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; Italy).

  • — Pilon, 1998: 95 (Ocypus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Italy, Austria, Ukraine, Russia, Caucasus, Turkey.

aliquoi Bordoni, 1976a: 186 (Alapsodus; subgenus Allocypus; subspecies of falcifer; Type locality: Sicilia: M.te Maganoce, Palermo, 850 m circa).

  • — Sparacio, 1995: 163 (Ocypus; subspecies of falcifer; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; subspecies of falcifer; Italy).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila; subspecies of falcifer).

fulvipes Ménétriés, 1849, see: planipennis Aubé, 1842.

fulvipes Motschulsky, 1858, see: morsitans Rossi, 1790.

fulvipilis G. Müller, 1932, see: morsitans Rossi, 1790.

fulvopubescens Chapman, 1933, see: ater Gravenhorst, 1802.

gagates Baudi, 1848, see: ater Gravenhorst, 1802.

globulifer Geoffroy, 1785: 164 (Staphylinus; Type locality: Paris).

  • — Olivier, 1795:(42): 14 (Staphylinus; synonym of brunnipes).

  • — Gravenhorst, 1802: 5 (Staphylinus; synonym of similis).

  • — Erichson, 1840: 409 (Ocypus; synonym of similis).

  • — Kraatz, 1857d: 556 (Ocypus; synonym of similis).

  • — Mulsant and Rey, 1876: 328 [= 1877: 184] (Anodus; [Note: Misidentified as morio according to Smetana, 1958: 299]; characters; France).

  • — Ganglbauer, 1895: 436 (Staphylinus; subgenus Ocypus; characters; Palaearctic region).

  • — Sainte-Claire Deville, 1906: 100 (Staphylinus; Corsica).

  • — Reitter, 1909: 121 (Staphylinus; subgenus Ocypus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; subgenus Ocypus; catalog).

  • — H. Bauer, 1914: 55 (Staphylinus; characters; notes).

  • — Johansen, 1914: 387 (Staphylinus; subgenus Ocypus; characters; Denmark).

  • — G. Müller, 1923: 143 (Staphylinus; subgenus Anodus; characters; Italy).

  • — G. Müller, 1924: 26, 27 (Staphylinus; subgenus Anodus; notes).

  • — G. Müller, 1926: 22 (Staphylinus; subgenus Ocypus; characters; notes).

  • — Edmonds, 1930: 273 (Staphylinus; characters in key; Britain).

  • — V. Hansen, 1931: 80 (Staphylinus; characters).

  • — G. Müller, 1932: 87 (Staphylinus; characters).

  • — Scheerpeltz, 1933: 1394 (Staphylinus; catalog).

  • — Méquignon, 1933: 228 (Staphylinus; distinguishing characters).

  • — Chagnon, 1936: 116 (Staphylinus; Quebec).

  • — Paulian, 1941: 243 (Staphylinus; subgenus Ocypus; larval characters).

  • — Steel, 1949: 273 (Staphylinus; subgenus Ocypus; characters; Britain).

  • — V. Hansen, 1952: 106 (Staphylinus; subgenus Ocypus; characters; Denmark).

  • — Coiffait, 1956e: 219, 220 (Ocypus; subgenus Ocypus; characters).

  • — Hatch, 1957: 177 (Ocypus; subgenus Alapsodus; characters; British Columbia; Washington).

  • — Smetana, 1958: 299 (Ocypus; subgenus Alapsodus; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 118 (Staphylinus; subgenus Ocypus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 202 (Ocypus; subgenus Ocypus; characters; central Europe).

  • — Coiffait, 1964: 83 (Metocypus).

  • — Moore, 1965: 102 (Ocypus; introduced in North America).

  • — Horion, 1965: 235 (Ocypus; Holland; Belgium; France; Germany; Austria; Switzerland; Corsica; Sicily; Italy; Istria; Dalmatia; Greece; Czechoslovakia; Poland; Ukraine; Denmark; Sweden).

  • — Pototskaia, 1967: 66 (Staphylinus; subgenus Alapsodus; larval characters).

  • — Coiffait, 1974: 551 (Metocypus; characters; Europe; Britain; Scandinavia).

  • — Orth, Moore, Fisher and Legner, 1975: 1115 (Ocypus; characters).

  • — Pope, 1977: 30 (Staphylinus; Britain).

  • — Hinton, 1981: 662 (Ocypus; characters of egg).

  • — Tóth, 1984: 33 (Ocypus; characters).

  • — Segers, 1986: 49 (Metocypus; checklist of species; Belgium).

  • — Newton, 1987: 382 (Staphylinus; reports of species in North America may be erroneous).

  • — Lucht, 1987: 108 (Ocypus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Schülke and Uhlig, 1988: 12 (Ocypus; Germany).

  • — Whitehead, 1989: 9 (Staphylinus; field identification; Britain).

  • — Whitehead, 1990c: 5 (Staphylinus; revised field identification; Britain).

  • — Pilon and Zanetti, 1991: 69 (Ocypus; Italy).

  • — Welch, 1993: 229 (Staphylinus; ovariole number and ovary structure).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Alapsodus; Slovakia).

  • — Dearborn and Donahue, 1994: 15 (Staphylinus; Maine).

  • — Outerelo, Palmer and Pons, 1995: 78 (Metocypus; Mallorca).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; Italy).

  • — M. Hansen, 1996: 106 (Ocypus; Denmark).

  • — Downie and Arnett, 1996: 384 (Ocypus; characters; USA).

  • — K. Bland, 1997: 204 (Staphylinus; eaten by toad; Scotland).

  • — Pilon, 1998: 95 (Ocypus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • — Assing, 2001: 75 (Ocypus; collected from xerothermous site; Germany).

  • Distribution: Europe, Caucasus, Turkey, Syria, Canada ?, USA ?.

siculus Stierlin, 1864: 146 (Ocypus; [preoccupied]; Type locality: Sicilien).

  • — Ganglbauer, 1895: 436 (Staphylinus; subgenus Ocypus; synonym of globulifer).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; synonym of globulifer).

  • — G. Müller, 1926: 23 (Staphylinus; subgenus Ocypus; characters; notes; France, Italy, Sicily, Balkan peninsula, Asia Minor, Austria, Germany).

  • — G. Müller, 1924: 27 (Staphylinus; characters; notes).

  • — Edmonds, 1930: 273 (Staphylinus; characters in key; Britain).

  • — V. Hansen, 1931: 80 (Staphylinus; characters).

  • — Van der Wiel, 1932: 151 (Staphylinus; characters; Holland).

  • — G. Müller, 1932: 87 (Staphylinus; characters; Germany; Holland; England).

  • — Scheerpeltz, 1933: 1401 (Staphylinus; catalog).

  • — Méquignon, 1933: 228 (Staphylinus; identical with globulifer; name preoccupied).

  • — G. Müller, 1943: 107 (Ocypus; form of globulifer; characters; notes).

  • — Steel, 1949: 274 (Staphylinus; subgenus Ocypus; synonym of evitendus).

  • — Tottenham, 1949c: 41 (synonym of globulifer).

  • — V. Hansen, 1952: 107 (Staphylinus; subgenus Ocypus; characters; Denmark).

  • — Smetana, 1958: 299 (Ocypus; synonym of globulifer).

  • — Palm, 1963b: 118 (Staphylinus; subgenus Ocypus; characters; Sweden; Denmark).

  • — Lohse, 1964: 202 (Ocypus; synonym of globulifer).

  • — Focarile, 1964: 61 (Staphylinus; Italy).

  • — Scheerpeltz, 1965b: 405 (Ocypus; Sicily).

  • — Scheerpeltz, 1965c: 510 (Ocypus; Sicily).

  • — Coiffait, 1974: 552 (Metocypus; synonym of globulifer).

evitendus Tottenham, 1945: 71 (Ocypus; replacement name for siculus Stierlin).

  • — Steel, 1949: 274 (Staphylinus; subgenus Ocypus; valid species; characters; Britain).

  • — Tottenham, 1949c: 41 (synonym of globulifer).

  • — Smetana, 1958: 299 (Ocypus; synonym of globulifer).

  • — Coiffait, 1974: 552 (Metocypus; synonym of globulifer).

  • — Pope, 1977: 30 (Staphylinus; synonym of globulifer).

  • — Herman, 2001: 31 (Ocypus; subspecies of globulifer; valid name for subspecies).

sicanus Coiffait, 1964: 106 (Metocypus; subspecies of globulifer; replacement name for siculus Stierlin).

  • — Coiffait, 1974: 552 (Metocypus; subspecies of globulifer; cited as “nom. nov.”; Sicily; Italy).

  • — Sparacio, 1995: 163 (Ocypus; subspecies of globulifer; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; subspecies of globulifer; Italy).

  • — Herman, 2001: 31 (Tasgius; synonym of globulifer evitendus).

gracilicornis Hochhuth, 1849: 120 (Ocypus; Type locality: Lenkoran; [Note: See lectotype designation by Gusarov, 1993]).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; synonym of macrocephalus).

  • — Scheerpeltz, 1933: 1390 (Staphylinus; synonym of alpestris).

  • — Coiffait, 1974: 465 (Ocypus; characters; Caucasus; Transcaucasus).

  • — Gusarov, 1993: 72 (Ocypus; subgenus Metocypus; lectotype designation; synonyms).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Georgia, Azerbaijan, Turkey, Iran.

ensifer G. Müller, 1932: 85, 87 (Staphylinus; subgenus Ocypus; Type locality: Caucasia: Abchasia. Also cited from Persia).

  • — Scheerpeltz, 1933: 1393 (Staphylinus; catalog).

  • — Smetana, 1968a: 160 (Ocypus; subgenus Alapsodus; Anatolia).

  • — Smetana, 1971c: 269 (Ocypus; subgenus Alapsodus; Iran).

  • — Coiffait, 1974: 555 (Metocypus; characters; Caucasus; Transcaucasus; Persia).

  • — Gusarov, 1993: 72 (Ocypus; synonym of gracilicornis).

  • — Smetana and Davies, 2000: 46 (Tasgius; synonym of gracilicornis).

gracilipalpis Eppelsheim, 1878, see: depressus Hochhuth, 1849.

herculeanus Coiffait, 1964: 105 (Metocypus; subspecies of nigrinus; Type locality: Andalousie: Tarifa).

  • — Coiffait, 1970c: 99 (Metocypus; valid species; Spain).

  • — Coiffait, 1974: 554 (Metocypus; characters; Spain).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Spain.

inderiensis Motschulsky, 1845a: 40 (Rayacheila; Type locality: Lac Indersk).

  • — Motschulsky, 1858: 669 (Rhagochila; characters; Turkmenia; Lac Inderskoje).

  • — Hochhuth, 1862: 51 (Ocypus; characters; Siberia).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; catalog).

  • — Blackwelder, 1952: 337, 338 (type species of Rayacheila and Rhagochila).

  • — Coiffait, 1974: 480 (Ocypus; characters).

  • — Boháč, 1988: 553 [= 1989: 36] (Metocypus; characters; Russia).

  • — Smetana and Davies, 2000: 36, 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Russia, Turkmenistan.

kirbyi Stephens, 1832, see: melanarius Heer, 1839.

limbifrons Hochhuth, 1849: 127 (Ocypus; Type locality: Armenien).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; catalog).

  • — Coiffait, 1974: 546 (Alapsodus; characters; Asia Minor; Transcaucasus; Russia).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Turkey, Armenia, Russia.

luganensis Heer, 1839, see: morsitans Rossi, 1790.

lusitanicus G. Müller, 1943: 108 (Ocypus; subspecies of siculus; Type locality: Oporto nel Portogallo).

  • — Coiffait, 1964: 83 (Metocypus; cited as valid species).

  • — Coiffait, 1974: 552 (Metocypus; characters; Portugal).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Portugal.

maderae Jarrige, 1943: 147 (Staphylinus; subgenus Anodus; Type locality: Madère, Encumiada).

  • — Coiffait, 1974: 545 (Alapsodus; characters; Madeira).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Madeira.

marani Roubal, 1932, see: melanarius Heer, 1839.

melanarius Heer, 1839: 256 (Staphylinus; Type locality: Zürich; Matt).

  • — Lacordaire, 1835: 372 (Emus; [Note: Misidentification: Bernhauer and Schubert, 1914: 387, cited fuscatus Boisduval and Lacordaire as a synonym of globulifer; Lacordaire did not describe the species as new, he attributed it to Gravenhorst; Coiffait, 1974: 544 cited this misidentification as referring to melanarius]; characters; France).

  • — Erichson, 1839a: 444 (Ocypus; [Note: Misidentified as morio according to Smetana, 1958: 298]; characters; Germany).

  • — Erichson, 1840: 417 (Ocypus; [Note: Misidentified as morio according to Smetana, 1958: 298]; characters; Europe).

  • — Fairmaire and Laboulbène, 1856: 512 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 563 (Ocypus; [Note: Misidentified as morio according to Smetana, 1958: 298]; characters; Germany).

  • — Hochhuth, 1862: 53 (Ocypus; notes; Russia).

  • — Mulsant and Rey, 1876: 328 [= 1877: 184] (Anodus; [Note: Misidentified as morio according to Smetana, 1958: 298]; characters; France).

  • — Ganglbauer, 1895: 436 (Staphylinus; synonym of globulifer).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; synonym of globulifer).

  • — G. Müller, 1926: 23 (Staphylinus; subgenus Ocypus; characters; notes; Romania; Bosnia; Istria; Piemont).

  • — Van der Wiel, 1932: 149 (Staphylinus; characters; Holland).

  • — Scheerpeltz, 1933: 1396 (Staphylinus; catalog).

  • — Méquignon, 1933: 228 (Staphylinus; distinguishing characters).

  • — Tottenham, 1949: 420 (Staphylinus; checklist; Britain).

  • — Steel, 1949: 274 (Staphylinus; subgenus Ocypus; synonym of evitendus).

  • — Coiffait, 1956e: 219, 220 (Ocypus; subgenus Ocypus; characters).

  • — Smetana, 1958: 298 (Ocypus; subgenus Alapsodus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 202 (Ocypus; subgenus Ocypus; characters; central Europe).

  • — Coiffait, 1964: 83 (Alapsodus; subgenus Allocypus).

  • — Horion, 1965: 233 (Ocypus; Finland; Sweden; Norway; Denmark; Britain; Holland; Belgium; France; Germany; Austria; Switzerland; Czechoslovakia; Poland; Italy; Dalmatia; Bosnia Hercegovina; Albania).

  • — Smetana, 1968b: 90 (Ocypus; subgenus Alapsodus; Anatolia).

  • — Szujecki, 1968a: 733 (Ocypus; Poland).

  • — Scheerpeltz, 1968a: 85 (Ocypus; catalog; Austria).

  • — Coiffait, 1974: 543 (Alapsodus; subgenus Allocypus; characters; Europe; Britain; Scandinavia).

  • — Pope, 1977: 30 (Staphylinus; Britain).

  • — Coiffait, 1978d: 165 (Alapsodus; Turkey).

  • — Muona, 1979: 15 (Ocypus; Finland; Sweden; Norway; Denmark; Baltic States).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 112 (Ocypus; catalog; Poland).

  • — Tóth, 1984: 33 (Ocypus; characters; Hungary).

  • — Newton, 1987: 382 (Staphylinus; introduced species; Quebec; Connecticut; Massachusetts; Washington).

  • — Ádám, 1987: 149 (Allocypus; collecting notes; Hungary).

  • — Lucht, 1987: 108 (Ocypus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Huth and Dettner, 1989: 154 (Ocypus; defensive secretions).

  • — Gusarov, 1989: 11 (Ocypus; habitat notes; Ukraine).

  • — Whitehead, 1989: 9 (Staphylinus; field identification; Britain).

  • — Whitehead, 1990c: 5 (Staphylinus; revised field identification; Britain).

  • — Huth and Dettner, 1990 (Ocypus; defensive chemicals).

  • — Silfverberg, 1992: 17 (Ocypus; Finland; Sweden; Norway; Denmark; Estonia; Latvia).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Alapsodus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; Italy).

  • — M. Hansen, 1996: 106 (Ocypus; checklist; Denmark).

  • — Eisinger, 1997: 171 (Ocypus; collecting notes; Germany).

  • — T. Wagner, 1997: 235 (Ocypus; collecting notes; Germany).

  • — Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 38 (Ocypus; Latvia).

  • — Anderson, Nash, and O'Connor, 1997: 19 (Ocypus; Ireland).

  • — Raitschev, 1997a: 100 (Alapsodus; notes; Bulgaria).

  • — Pilon, 1998: 96 (Ocypus; notes; Italy).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • — Assing, 2001: 75 (Ocypus; collected from xerothermous site; Germany).

  • — Herman, 2001: 31 (Tasgius; nomen protectum).

  • Distribution: Europe, Russia, Caucasus, Turkey, Kazakhstan, Canada, USA.

similis Paykull, 1789: 10 (Staphylinus; [preoccupied]; Type locality: Suecia).

  • — Paykull, 1800: 371 (Staphylinus; [Note: Paykull cited both similis Fabricius, 1793: 521, and similis Paykull, 1789: 10, as referring to the same species]; characters; Sweden).

  • — Turton, 1802: (Staphylinus; characters; Germany).

  • — C. Sahlberg, 1830: 310 (Staphylinus; synonym of subpunctatus).

  • — Mannerheim, 1830: 24 (Staphylinus; Sweden).

  • — Mannerheim, 1831: 438 (Staphylinus; Sweden).

  • — Ménétriés, 1832: 143 (Staphylinus; Caucasus).

  • — Runde, 1835: 3 (Staphylinus; characters; Germany).

  • — Erichson, 1839a: 444 (Ocypus; synonym of morio).

  • — Erichson, 1840: 417 (Ocypus; synonym of morio).

  • — Kraatz, 1857d: 563 (Ocypus; synonym of morio).

  • — Ganglbauer, 1895: 436 (Staphylinus; synonym of globulifer).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; synonym of globulifer).

  • — Scheerpeltz, 1933: 1396 (Staphylinus; synonym of melanarius).

  • — Smetana, 1958: 298 (Ocypus; synonym of melanarius).

  • — Coiffait, 1974: 544 (Alapsodus; synonym of melanarius).

  • — Herman, 2001: 32 (Tasgius; nomen oblitum).

kirbyi Stephens, 1832: 210 (Goerius; [Note: This name is older than melanarius]; Type locality: Devonshire).

  • — Stephens, 1839: 388 (Goerius; characters; England).

  • — Hardy, 1851: 25 (Staphylinus; synonym of ater).

  • — G. Waterhouse, 1858: 24 (Ocypus; synonym of morio).

  • — Gemminger and Harold, 1868: 583 (Ocypus; synonym of morio).

  • — Ganglbauer, 1895: 436 (Staphylinus; synonym of globulifer).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; synonym of globulifer).

  • — Smetana, 1958: 298 (Ocypus; synonym of melanarius).

  • — Coiffait, 1974: 544 (Alapsodus; synonym of melanarius).

  • — Herman, 2001: 32 (Tasgius; nomen oblitum).

angustatus Lacordaire, 1835: 369 (Emus; [Note: This name is older than melanarius]; Type locality: Paris).

  • — Erichson, 1839a: 444 (Ocypus; synonym of morio).

  • — Hardy, 1851: 24 (Staphylinus; synonym of morio).

  • — Kraatz, 1857d: 563 (Ocypus; synonym of morio).

  • — Gemminger and Harold, 1868: 582 (Ocypus; synonym of morio).

  • — Ganglbauer, 1895: 436 (Staphylinus; synonym of globulifer).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; synonym of globulifer).

  • — Smetana, 1958: 298 (Ocypus; synonym of melanarius).

  • — Coiffait, 1974: 544 (Alapsodus; synonym of melanarius).

  • — Herman, 2001: 32 (Tasgius; nomen oblitum).

obscurus Runde, 1835: 4 (Staphylinus; [preoccupied]; Type locality: Halae).

  • — Erichson, 1839a: 444 (Ocypus; synonym of morio).

  • — Erichson, 1840: 417 (Ocypus; synonym of morio).

  • — Kraatz, 1857d: 563 (Ocypus; synonym of morio).

  • — Ganglbauer, 1895: 436 (Staphylinus; synonym of globulifer).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; synonym of globulifer).

  • — Smetana, 1958: 298 (Ocypus; synonym of melanarius).

  • — Coiffait, 1974: 544 (Alapsodus; synonym of melanarius).

  • — Herman, 2001: 32 (Tasgius; nomen oblitum).

saulcyi Reiche, 1857: ix (Ocypus; Type locality: Peterhead in Scotia boreali).

  • — Fauvel, 1874: 193 [= 1874a: 419] (Staphylinus; synonym of edentulus).

  • — Ganglbauer, 1895: 436 (Staphylinus; synonym of globulifer).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; synonym of globulifer).

  • — Smetana, 1958: 298 (Ocypus; synonym of melanarius).

  • — Coiffait, 1974: 544 (Alapsodus; synonym of melanarius).

rubidus Verhoeff, 1890: 17 (Ocypus; variety of edentulus; Type locality: Bonn).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; variety of globulifer).

  • — Scheerpeltz, 1933: 1396 (Staphylinus; variety of melanarius).

  • — Smetana, 1958: 298 (Ocypus; aberration of melanarius).

  • — Coiffait, 1974: 544 (Alapsodus; variety of melanarius).

sahlbergi G. Müller, 1926a: 32 (Staphylinus; subgenus Ocypus; subspecies of globulifer; Type locality: Turkestan: Vernyi).

  • — Scheerpeltz, 1933: 1395 (Staphylinus; synonym of globulifer).

  • — Coiffait, 1974: 544 (Alapsodus; subspecies of melanarius).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila; subspecies of melanarius).

marani Roubal, 1932: 1 (Staphylinus; subspecies of minax; Type locality: Republique Tchecoslovaque: V. Fatra).

  • — Scheerpeltz, 1933: 1397 (Staphylinus; subspecies of minax).

  • — Coiffait, 1956e: 219 (Ocypus; subgenus Ocypus; subspecies of minax; characters).

  • — Lohse, 1964: 202 (Ocypus; synonym of melanarius).

  • — Coiffait, 1974: 544 (Alapsodus; synonym of melanarius).

dvoraki Roubal, 1940: 49, 51 (Staphylinus; variety of minax; Type locality: Bohemia: Brandýs nad Labem).

  • — Smetana, 1958: 298 (Ocypus; synonym of melanarius).

  • — Coiffait, 1974: 544 (Alapsodus; synonym of melanarius).

messor Nordmann, 1837: 11 (Anodus; Type locality: Italia: Sicilia prope Palermo).

  • — Erichson, 1840: 418 (Ocypus; synonym of falcifer).

  • — Kraatz, 1857d: 565 (Ocypus; synonym of falcifer).

  • — Ganglbauer, 1895: 435 (Staphylinus; synonym of falcifer).

  • — Bernhauer and Schubert, 1914: 386 (Staphylinus; synonym of falcifer).

  • — Coiffait, 1974: 537 (Alapsodus; cited as valid species; characters; Caucasus; Transcaucasus).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Italy, Caucasus.

minax Mulsant and Rey, 1861: 139 [= 1862: 123] (Ocypus; Type locality: la vallée de Champsaur, Hautes-Alpes).

  • — Fauvel, 1874: 193 [= 1874a: 419] (Staphylinus; characters; France).

  • — Fauvel, 1876a: 72 [= 1876: 257] (Staphylinus; Spain; Italy).

  • — Mulsant and Rey, 1876: 333 [= 1877: 189] (Anodus; characters; France).

  • — Ganglbauer, 1895: 435 (Staphylinus; subgenus Ocypus; characters; France; Spain; Tuscany; western Germany).

  • — Reitter, 1909: 122 (Staphylinus; subgenus Ocypus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; subgenus Ocypus; catalog).

  • — Portevin, 1929: 353 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1397 (Staphylinus; catalog).

  • — Roubal, 1940a: 48, 50 (Staphylinus; characters in key).

  • — Coiffait, 1956e: 219, 220 (Ocypus; subgenus Ocypus; characters).

  • — Lohse, 1964: 202 (Ocypus; subgenus Ocypus; characters; central Europe).

  • — Coiffait, 1964: 83 (Alapsodus; subgenus Allocypus).

  • — Horion, 1965: 232 (Ocypus; Spain; Italy; Holland ?).

  • — Coiffait, 1974: 541 (Alapsodus; subgenus Allocypus; characters; Europe; Italy; France).

  • — Drugmand, 1987a: 311 (Alapsodus; characters; Belgium).

  • — Lucht, 1987: 108 (Ocypus; checklist; France; Benelux).

  • — Pilon, 1998: 98 (Ocypus; notes; doubtfully in Italy).

  • — Assing, Frisch, Kahlen, et al., 1998: 139 (Tasgius; note).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Holland ?; Germany, Belgium, France, Italy ?, Spain.

morio Gravenhorst, 1802, see: winkleri Bernhauer, 1906.

morsitans Rossi, 1790: 249 (Staphylinus; Type locality: Etruria).

  • — Fauvel, 1874: 194 [= 1874a: 420] (Staphylinus; synonym of compressus).

  • — Ganglbauer, 1895: 436 (Staphylinus; synonym of compressus).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of compressus).

  • — Steel, 1949: 275 (Staphylinus; subgenus Ocypus; valid species; characters; Britain).

  • — Smetana, 1958: 302 (Ocypus; synonym of compressus).

  • — Coiffait, 1974: 535 (Alapsodus; synonym of compressus).

  • — Pope, 1977: 30 (Staphylinus; synonym of compressus).

  • — Allen, 1990a: 9 (Ocypus; valid species).

  • — Ádám, 1996: 247 (Alapsodus; valid name; Hungary).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Europe, Russia, Turkey, Syria.

compressus Marsham, 1802: 503 (Staphylinus; [preoccupied]; Type locality: Britain).

  • — Curtis, 1829: 24 (Staphylinus; catalog; Britain).

  • — Stephens, 1832: 212 (Ocypus; characters; England).

  • — Stephens, 1839: 388 (Ocypus; characters; England).

  • — Erichson, 1840: 418 (Ocypus; characters; England; Holland; France; Hungary).

  • — Kraatz, 1857d: 564 (Ocypus; characters; Germany).

  • — Redtenbacher, 1857: 985 (Staphylinus; characters; Austria).

  • — Thomson, 1860: 150 (Anodus; characters; Scandinavia).

  • — Hochhuth, 1862: 53 (Ocypus; Russia).

  • — Fauvel, 1865a: 295 (Ocypus; habitat; France).

  • — Baudi, 1870: 385 (Ocypus; characters).

  • — Fauvel, 1874: 194 [= 1874a: 420] (Staphylinus; characters; France; Syria).

  • — Seidlitz, 1875: 271 (Ocypus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 330 [= 1877: 186] (Anodus; characters; France).

  • — Fowler, 1888: 258 (Ocypus; characters; Britain).

  • — Ganglbauer, 1895: 436 (Staphylinus; subgenus Ocypus; characters; Europe; Syria).

  • — Sainte-Claire Deville, 1906: 100 (Staphylinus; Corsica).

  • — Reitter, 1909: 121 (Staphylinus; subgenus Ocypus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; subgenus Ocypus; catalog).

  • — Johansen, 1914: 387 (Staphylinus; subgenus Ocypus; characters; Denmark).

  • — G. Müller, 1926: 22 (Staphylinus; subgenus Ocypus; notes).

  • — Portevin, 1929: 353 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1392 (Staphylinus; catalog).

  • — Steel, 1949: 275 (Staphylinus; subgenus Ocypus; synonym of morsitans).

  • — V. Hansen, 1952: 108 (Staphylinus; subgenus Ocypus; characters; Denmark).

  • — Coiffait, 1956e: 218, 219 (Ocypus; subgenus Ocypus; characters).

  • — Smetana, 1958: 302 (Ocypus; subgenus Alapsodus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960b: 259 (Ocypus; Czechoslovakia).

  • — Scheerpeltz, 1961d: 132 (Ocypus; Italy).

  • — Palm, 1963b: 119 (Staphylinus; subgenus Ocypus; characters; Sweden).

  • — Lohse, 1964: 201 (Ocypus; subgenus Ocypus; characters; central Europe).

  • — Horion, 1965: 229 (Ocypus; Nordmann cited as author; Spain; Corsica; Sardinia; Sicily; Italy; Dalmatia; Bosnia; Greece; Serbia; Holland; Belgium; France; Switzerland; Germany; Austria; Romania; Hungary; Czechoslovakia; Poland; Ukraine; Britain; Ireland; Norway; Sweden; Denmark).

  • — Smetana, 1968a: 160 (Ocypus; subgenus Alapsodus; Anatolia).

  • — Coiffait, 1974: 534 (Alapsodus; characters; Europe; Britain; Scandinavia).

  • — Pope, 1977: 30 (Staphylinus; Britain).

  • — Coiffait, 1978d: 165 (Alapsodus; Hochhuth cited as author of species; Turkey).

  • — Tóth, 1984: 32 (Ocypus; characters; Hungary).

  • — Ádám, 1987: 149 (Alapsodus; cited as valid species; collecting notes; Hungary).

  • — Lucht, 1987: 108 (Ocypus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Schülke and Uhlig, 1988: 12 (Ocypus; notes; Germany).

  • — Nowosad, 1990: 115 (Ocypus; collected from mole nest; notes; Poland).

  • — Allen, 1990a: 9 (Ocypus; synonym of morsitans).

  • — Gusarov, 1993: 72 (Ocypus; subgenus Alapsodus; cited as valid species).

  • — Welch, 1993: 229 (Staphylinus; ovariole number and ovary structure).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Alapsodus; Czech Republic; Slovakia).

  • — Ádám, 1996: 247 (Alapsodus; synonym of morsitans).

  • — Pilon, 1998: 94 (Ocypus; ecological notes; Italy).

  • — Owen, 2000: 251 (Staphylinus; pitfall trap collections from underground at roots of trees; Britain).

  • — Smetana and Davies, 2000: 46 (Tasgius; synonym of morsitans).

phaeopus Stephens, 1832: 212 (Ocypus; Type locality: London).

  • — Stephens, 1839: 389 (Ocypus; characters; England).

  • — G. Waterhouse, 1858: 24 (Ocypus; synonym of compressus).

  • — Gemminger and Harold, 1868: 581 (Ocypus; synonym of compressus).

  • — Ganglbauer, 1895: 436 (Staphylinus; synonym of compressus).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of compressus).

  • — Steel, 1949: 275 (Staphylinus; subgenus Ocypus; synonym of morsitans).

  • — Smetana, 1958: 302 (Ocypus; synonym of compressus).

  • — Coiffait, 1974: 535 (Alapsodus; synonym of compressus).

  • — Pope, 1977: 30 (Staphylinus; synonym of compressus).

rufipalpis Lacordaire, 1835: 370 (Emus; Type locality: Paris).

  • — Erichson, 1840: 418 (Ocypus; synonym of compressus).

  • — Fairmaire and Laboulbène, 1856: 512 (Staphylinus; synonym of compressus).

  • — Kraatz, 1857d: 564 (Ocypus; synonym of compressus).

  • — Redtenbacher, 1857: 985 (Ocypus; synonym of compressus).

  • — Ganglbauer, 1895: 436 (Staphylinus; synonym of compressus).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of compressus).

  • — Steel, 1949: 275 (Staphylinus; subgenus Ocypus; synonym of morsitans).

  • — Smetana, 1958: 302 (Ocypus; synonym of compressus).

  • — Coiffait, 1974: 535 (Alapsodus; synonym of compressus).

luganensis Heer, 1839: 256 (Staphylinus; Type locality: M. Salvatore bei Lugano).

  • — Gemminger and Harold, 1868: 581 (Ocypus; synonym of compressus).

  • — Fauvel, 1869: 488 (Quedius; synonym of cerdo).

  • — Fauvel, 1874: 194 [= 1874a: 420] (Staphylinus; synonym of compressus).

  • — Ganglbauer, 1895: 436 (Staphylinus; synonym of compressus).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of compressus).

  • — Steel, 1949: 275 (Staphylinus; subgenus Ocypus; variety of morsitans).

  • — Smetana, 1958: 302 (Ocypus; synonym of compressus).

  • — Coiffait, 1974: 535 (Alapsodus; synonym of compressus).

cerdo Erichson, 1840: 417 (Ocypus; Type locality: Italia: in montibus Appeninis; Sardinia).

  • — Fairmaire and Laboulbène, 1856: 512 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 564 (Ocypus; characters; Germany).

  • — Kraatz, 1858a: 61 (Ocypus; Greece, Syria).

  • — Fauvel, 1867c: 112 [= 1868b: 211] (Ocypus; variety of compressus).

  • — Fauvel, 1874: 194 [= 1874a: 420] (Staphylinus; synonym of compressus).

  • — Ganglbauer, 1895: 436 (Staphylinus; variety of compressus).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; variety of compressus).

  • — Scheerpeltz, 1933: 1392 (Staphylinus; variety of compressus).

  • — Steel, 1949: 275 (Staphylinus; subgenus Ocypus; synonym of morsitans).

  • — Coiffait, 1956e: 218 (Ocypus; subgenus Ocypus; variety of compressus; characters).

  • — Smetana, 1958: 302 (Ocypus; aberration of compressus).

  • — Coiffait, 1974: 535 (Alapsodus; variety of compressus; France; Italy; Sicily; Sardinia; Austria; Greece; Syria).

  • — Tóth, 1984: 32 (Ocypus; variety of compressus; Hungary).

  • — Ciceroni and Zanetti, 1995: 30 (Ocypus; subspecies of compressus; Italy).

fulvipes Motschulsky, 1858: 669 (Anodus; [preoccupied]; Type locality: Lausanne en Suisse; [Note: See lectotype designation by Gusarov, 1993]).

  • — Fauvel, 1874: 194 [= 1874a: 420] (Staphylinus; synonym of compressus).

  • — Ganglbauer, 1895: 436 (Staphylinus; synonym of compressus).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of compressus).

  • — Steel, 1949: 275 (Staphylinus; subgenus Ocypus; synonym of morsitans).

  • — Smetana, 1958: 302 (Ocypus; synonym of compressus).

  • — Coiffait, 1974: 535 (Alapsodus; synonym of compressus).

  • — Gusarov, 1993: 72 (Ocypus; lectotype designation; synonym of compressus).

  • — Smetana and Davies, 2000: 46 (Tasgius; synonym of morsitans).

mutatus Gemminger and Harold, 1868: 583 (Ocypus; replacement name for fulvipes Motschulsky).

  • — Fauvel, 1874: 194 [= 1874a: 420] (Staphylinus; synonym of compressus).

  • — Ganglbauer, 1895: 436 (Staphylinus; synonym of compressus).

  • — Bernhauer and Schubert, 1914: 385 (Staphylinus; synonym of compressus).

  • — Steel, 1949: 275 (Staphylinus; subgenus Ocypus; synonym of morsitans).

  • — Smetana, 1958: 302 (Ocypus; synonym of compressus).

  • — Coiffait, 1974: 535 (Alapsodus; synonym of compressus).

fulvipilis G. Müller, 1932: 88 (Staphylinus; subgenus Ocypus; variety of compressus; Type locality: Germania: Württemberg).

  • — Scheerpeltz, 1933: 1392 (Staphylinus; variety of compressus).

  • — G. Müller, 1950: 11 (Staphylinus; subgenus Ocypus; subspecies of compressus; notes; Germany; Poland).

  • — Coiffait, 1956e: 218 (Ocypus; subgenus Ocypus; variety of compressus; characters).

  • — Smetana, 1958: 302 (Ocypus; aberration of compressus).

  • — Coiffait, 1974: 535 (Alapsodus; variety of compressus).

mutatus Gemminger and Harold, 1868, see: morsitans Rossi, 1790.

myops G. Müller, 1925: 47 (Staphylinus; subgenus Ocypus; subspecies of bellicosus; Type locality: [Algeria]: Constantina).

  • — Scheerpeltz, 1933: 1391 (Staphylinus; subspecies of bellicosus).

  • — Coiffait, 1974: 540 (Alapsodus; cited as valid species; characters; Algeria).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Algeria.

nigrinus P. Lucas, 1846: 109 (Ocypus; Type locality: environs d'Algers).

  • — Fairmaire and Coquerel, 1860: 156 (Ocypus; characters; Algeria; Morocco).

  • — Baudi, 1870: 385 (Ocypus; characters; Spain).

  • — Fauvel, 1878: 120 [= 1878a: 40] (Staphylinus; Morocco; Algeria).

  • — Fauvel, 1886: 52 [= 1886a: 44] (Staphylinus; Algeria).

  • — Fauvel, 1902b: 106 (Staphylinus; subgenus Ocypus; Morocco; Algeria; Tunisia).

  • — Bernhauer and Schubert, 1914: 388 (Staphylinus; subgenus Ocypus; catalog).

  • — G. Müller, 1932: 87 (Staphylinus; characters).

  • — Scheerpeltz, 1933: 1397 (Staphylinus; catalog).

  • — Coiffait, 1964: 83 (Metocypus).

  • — Coiffait, 1973a: 274 (Metocypus; Morocco).

  • — Coiffait, 1974: 553 (Metocypus; characters; Algeria; Tunisia; Morocco).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Spain, Morocco, Algeria, Tunisia.

nigripes Lacordaire, 1835, see: ater Gravenhorst, 1802.

obscuripes Bernhauer, 1900, see: eppelsheimianus Jakobson, 1909.

obscurus Marsham, 1802, see: ater Gravenhorst, 1802.

obscurus Runde, 1835, see: melanarius Heer, 1839.

olivieri Fauvel, 1868, see: planipennis Aubé, 1842.

olympicus Baudi, 1857: 100 (Ocypus; Type locality: Insula Cyprus).

  • — Fauvel, 1875a: XXVIII [= 1875b: 230] (Staphylinus; synonym of rubripennis).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; synonym of rubripennis).

  • — M. Dvořák, 1984: 66 (Alapsodus; valid species; characters; Cyprus).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Cyprus.

pedator Gravenhorst, 1802: 163 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).

  • — Gravenhorst, 1806: 113 (Staphylinus).

  • — Nordmann, 1837: 65 (Staphylinus; characters; Dalmatia).

  • — Heer, 1839: 254 (Staphylinus; characters; Switzerland).

  • — Erichson, 1840: 415 (Ocypus; characters; France; Italy; Dalmatia).

  • — Fairmaire and Laboulbène, 1856: 511 (Staphylinus; characters; France).

  • — Kraatz, 1857d: 561 (Ocypus; characters; Austria, Germany).

  • — Redtenbacher, 1857: 186 (Ocypus; characters; Austria).

  • — Kraatz, 1858a: 61 (Ocypus; Greece).

  • — Thomson, 1860: 149 (Tasgius; characters; Scandinavia).

  • — Fauvel, 1874: 191 [= 1874a: 417] (Staphylinus; characters; France; Scandinavia; Britain; Germany; Austria; Italy; Corsica; Sardinia; Greece; Algeria).

  • — Redtenbacher, 1874: 205 (Ocypus; characters; Austria).

  • — Seidlitz, 1875: 271 (Ocypus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 319 [= 1877: 175] (Tasgius; characters; larval characters; France).

  • — J. Sahlberg, 1876: 12 (Tasgius; Finland).

  • — Fauvel, 1878: 119 [= 1878a: 39] (Staphylinus; Algeria).

  • — Fauvel, 1886: 51 [= 1886a: 43] (Staphylinus; Algeria).

  • — Fowler, 1888: 257 (Ocypus; characters; Britain).

  • — Seidlitz, 1891: 392 (Ocypus; subgenus Pseudotasgius; characters; Europe).

  • — Seidlitz, 1891a: 418 (Ocypus; subgenus Pseudotasgius; characters; Europe).

  • — Ganglbauer, 1895: 435 (Staphylinus; subgenus Tasgius; characters; Europe; Mediterranean region).

  • — Fauvel, 1902b: 105 (Staphylinus; subgenus Ocypus; Algeria).

  • — Sainte-Claire Deville, 1906: 100 (Staphylinus; Corsica).

  • — Reitter, 1909: 122 (Staphylinus; subgenus Tasgius; characters; Germany).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; subgenus Ocypus; catalog).

  • — Johansen, 1914: 386 (Staphylinus; subgenus Tasgius; characters; Denmark).

  • — G. Müller, 1923: 142 (Staphylinus; subgenus Tasgius; characters; Italy).

  • — Portevin, 1929: 354 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1399 (Staphylinus; catalog).

  • — Paulian, 1936 (Staphylinus; subgenus Tasgius; larval description).

  • — Paulian, 1941: 242 (Staphylinus; subgenus Tasgius; larval characters).

  • — Jarrige, 1952a: 133 (Staphylinus; subgenus Tasgius; characters; Sicily).

  • — Coiffait, 1956e: 216, 217 (Ocypus; subgenus Tasgius; characters).

  • — Smetana, 1958: 296 (Ocypus; subgenus Tasgius; characters; ecology; Czechoslovakia).

  • — Smetana, 1960b: 259 (Ocypus; subgenus Tasgius; Czechoslovakia).

  • — Scheerpeltz, 1961d: 131 (Tasgius; Italy).

  • — Palm, 1963b: 117 (Staphylinus; subgenus Tasgius; characters; Sweden).

  • — Scheerpeltz, 1965b: 405 (Tasgius; Sicily).

  • — Scheerpeltz, 1965c: 509 (Tasgius; Sicily).

  • — Horion, 1965: 225 (Tasgius; Turkey; Portugal; Spain; Italy; Dalmatia; Greece; Albania; France; Britain; Holland; Belgium; Germany; Austria; Switzerland; Romania; Hungary; Czech Republic; Slovakia; Poland; Ukraine; Caucasus).

  • — Pototskaia, 1967: 66 (Staphylinus; subgenus Tasgius; larval characters).

  • — Smetana, 1967i: 557 (Ocypus; subgenus Tasgius; Bulgaria; Turkey).

  • — Coiffait, 1974: 521 (Tasgius; characters; Europe; Sweden; Britain; Corsica; Algeria; Caucasus).

  • — Bordoni, 1974b: 8 (Tasgius; notes; Italy).

  • — Pope, 1977: 30 (Staphylinus; Britain).

  • — Tóth, 1984: 31 (Ocypus; characters; Hungary).

  • — Ádám, 1987: 149 (Tasgius; collecting notes; Hungary).

  • — Lucht, 1987: 108 (Ocypus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Sweden).

  • — Pilon and Zanetti, 1991: 69 (Ocypus; Italy).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Tasgius; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; Italy).

  • — Pilon, 1998: 93 (Ocypus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Tasgius).

  • Distribution: Algeria, Europe, Azerbaijan, Iran, Turkey.

rufipes Latreille, 1806: 285 (Astrapaeus; Type locality: Parisii).

  • — Curtis, 1829: 24 (Goerius; catalog; Britain).

  • — Stephens, 1832: 213 (Tasgius; characters; England).

  • — Curtis, 1833: pl. 438 (Tasgius; characters; England).

  • — Stephens, 1835: 435 (Lesteva; England).

  • — Nordmann, 1837: 65 (Staphylinus; synonym of pedator).

  • — Westwood, 1838a: 16 (Tasgius; type species).

  • — Heer, 1839: 254 (Staphylinus; synonym of pedator).

  • — Stephens, 1839: 389 (Tasgius; characters; England).

  • — Erichson, 1840: 415 (Ocypus; synonym of pedator).

  • — Laporte, 1840: 173 (Tasgius).

  • — Fairmaire and Laboulbène, 1856: 511 (Staphylinus; synonym of pedator).

  • — Kraatz, 1857d: 561 (Ocypus; synonym of pedator).

  • — Ganglbauer, 1895: 435 (Staphylinus; synonym of pedator).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; synonym of pedator).

  • — Scheerpeltz, 1933: 1399 (Staphylinus; synonym of pedator).

  • — Smetana, 1958: 296 (Ocypus; synonym of pedator).

  • — Coiffait, 1974: 521 (Tasgius; synonym of pedator).

chalybeipennis Ménétriés, 1832: 143 (Staphylinus; Type locality: montagnes de Talyche).

  • — Nordmann, 1837: 73 (Staphylinus; characters; Talych Mountains).

  • — Faldermann, 1835: 117 (Staphylinus; characters; Persia-Armenia).

  • — Hochhuth, 1849: 119 (Ocypus; characters).

  • — Ganglbauer, 1895: 435 (Staphylinus; synonym of pedator).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; synonym of pedator).

  • — Smetana, 1958: 296 (Ocypus; synonym of pedator).

  • — Coiffait, 1974: 521 (Tasgius; synonym of pedator).

siculus Aubé, 1842: 234 (Ocypus; Type locality: Sicile).

  • — Fauvel, 1866c: 248 [= 1866d: 106] (Staphylinus; synonym of pedator).

  • — Kraatz, 1867: 414 (Ocypus; synonym of pedator).

  • — Ganglbauer, 1895: 435 (Staphylinus; synonym of pedator).

  • — Bernhauer and Schubert, 1914: 389 (Staphylinus; synonym of pedator).

  • — Scheerpeltz, 1933: 1399 (Staphylinus; subspecies of pedator).

  • — Coiffait, 1956e: 216 (Ocypus; subgenus Tasgius; subspecies of pedator; characters).

  • — Coiffait, 1974: 521 (Tasgius; subspecies of pedator).

  • — Sparacio, 1995: 163 (Ocypus; subspecies of pedator; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; subspecies of pedator; Italy).

rufipes Verhoeff, 1890: 17 (Ocypus; variety of ater; [preoccupied]; Type locality: Nähe der Kasselsruhe).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; synonym of ater).

  • — Scheerpeltz, 1933: 1399 (Staphylinus; synonym of pedator).

bonnairei Scheerpeltz, 1933: 1399 (Staphylinus; subspecies of pedator; [Note: This name was proposed and described by Reitter, 1909, as an aberration of S. pedator so his use of the name is unavailable. Scheerpeltz listed the name as a subspecies of S. pedator thereby making the name available under his authorship]; Type locality: Frankreich, Ile de Ré [Note: The locality is taken from Reitter, 1909]).

  • — Reitter, 1909: 122 (Staphylinus; subgenus Tasgius; aberration of pedator; [Note: Name is unavailable name]; France).

  • — Bernhauer and Schubert, 1914: 386, 389 (Staphylinus; aberration of falcifer [386] and pedator [389]).

  • — Coiffait, 1956e: 216, 217 (Ocypus; subgenus Tasgius; variety of pedator; characters).

  • — Coiffait, 1974: 521 (Tasgius; variety of pedator).

pendjabensis Bernhauer, 1915b: 53 (Staphylinus; subgenus Ocypus; Type locality: Pendjab, Shahpur).

  • — Cameron, 1932: 209 (Staphylinus; subgenus Ocypus; characters; India).

  • — Scheerpeltz, 1933: 1399 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: India.

peyerimhoffi G. Müller, 1926b: 47 (Staphylinus; subgenus Tasgius; Type locality: Marocco).

  • — G. Müller, 1932: 81 (Staphylinus; subgenus Tasgius; notes).

  • — Scheerpeltz, 1933: 1399 (Staphylinus; catalog).

  • — Jarrige, 1952a: 133 (Staphylinus; subgenus Paratasgius).

  • — Coiffait, 1964: 83 (Tasgius; subgenus Paratasgius).

  • — Coiffait, 1974: 525 (Tasgius; subgenus Paratasgius; characters; Morocco; Algeria; Tunisia; Egypt).

  • — M. Dvořák, 1984: 66 (Tasgius; Libya).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Tasgius).

  • Distribution: Morocco, Algeria, Tunisia, Libya, Egypt.

aegyptiacus C. Koch, 1936b: 194 (Staphylinus; subgenus Tasgius; subspecies of ater; Type locality: Aegypten: Mex, Aboukir, Dekeihla).

  • — Jarrige, 1952a: 133 (Staphylinus; not aberration of peyerimhoffi; characters).

  • — Coiffait, 1974: 525 (Tasgius; synonym of peyerimhoffi).

phaeopus Stephens, 1832, see: morsitans Rossi, 1790.

picipes Stephens, 1832, see: winkleri Bernhauer, 1906.

planipennis Aubé, 1842: 235 (Ocypus; Type locality: Sicile).

  • — P. Lucas, 1846: 110 (Ocypus; notes; Algeria).

  • — Fauvel, 1878: 119 [= 1878a: 39] (Staphylinus; Morocco; Algeria).

  • — Fauvel, 1886: 51 [= 1886a: 43] (Staphylinus; notes; Morocco; Algeria).

  • — Fauvel, 1902b: 105 (Staphylinus; variety of ater).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; synonym of ater).

  • — G. Müller, 1932: 81 (Staphylinus; subgenus Tasgius; notes).

  • — Scheerpeltz, 1933: 1400 (Staphylinus; catalog).

  • — Coiffait, 1956e: 217 (Ocypus; subgenus Tasgius; characters).

  • — Coiffait, 1964: 83 (Tasgius; subgenus Paratasgius).

  • — Coiffait, 1974: 524 (Tasgius; subgenus Paratasgius; characters; North Africa; Morocco).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; Italy).

  • — Pilon, 1998: 93 (Ocypus; ecological notes; Italy).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Tasgius).

  • Distribution: Italy, Morocco, Algeria.

fulvipes Ménétriés, 1849: 43 (Ocypus; Type locality: Turcomanie).

  • — Fauvel, 1875a: XXVIII [= 1875b: 230] (Staphylinus; synonym of planipennis).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; synonym of ater).

  • — Scheerpeltz, 1933: 1400 (Staphylinus; synonym of planipennis).

  • — Coiffait, 1974: 524 (Tasgius; synonym of planipennis).

olivieri Fauvel, 1868d: 61 (Ocypus; Type locality: mines de Mokta-el-Hadid, près Bône).

  • — Fauvel, 1875a: XXVIII [= 1875b: 230] (Staphylinus; synonym of planipennis).

  • — Fauvel, 1902b: 105 (Staphylinus; subgenus Ocypus; variety of ater; Algeria).

  • — Bernhauer and Schubert, 1914: 384 (Staphylinus; synonym of ater).

  • — Scheerpeltz, 1933: 1400 (Staphylinus; aberration of planipennis).

  • — Jarrige, 1952a: 133 (Staphylinus; subgenus Paratasgius; subspecies of ater).

  • — Coiffait, 1956e: 217 (Ocypus; subgenus Tasgius; variety of planipennis; characters).

  • — Coiffait, 1974: 524 (Tasgius; variety of planipennis).

pliginskii Bernhauer, 1915j: 267 (Staphylinus; subgenus Ocypus; Type locality: Krim, Sebastopol).

  • — Bernhauer, 1923d: 181 (Staphylinus; subgenus Ocypus; characters; Crimea).

  • — Scheerpeltz, 1933: 1400 (Staphylinus; catalog).

  • — Coiffait, 1964: 83 (Metocypus).

  • — Coiffait, 1974: 556 (Metocypus; characters; Crimea; Transcaucasus).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Ukraine, Caucasus.

praetorius Bernhauer, 1915c: 73 (Staphylinus; subgenus Tasgius; Type locality: China: Singley).

  • — Scheerpeltz, 1933: 1400 (Staphylinus; catalog).

  • — Coiffait, 1964: 83 (Tasgius; subgenus Paratasgius).

  • — Smetana 1975c: 173 (Ocypus; subgenus Tasgius; Mongolia).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Tasgius).

  • Distribution: China, Mongolia.

protensus Ménétriés, 1832, see: ater Gravenhorst, 1802.

punctatissimus Dufour, 1843, see: ater Gravenhorst, 1802.

rubidus Verhoeff, 1890, see: melanarius Heer, 1839.

rubripennis Reiche and Saulcy, 1856: 365 (Ocypus; Type locality: Jourdain).

  • — Baudi, 1870: 385 (Ocypus; characters; Cyprus).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1401 (Staphylinus; catalog).

  • — Coiffait, 1964: 83 (Metocypus).

  • — Coiffait, 1974: 555 (Metocypus; characters; Palestine; Persia; Russia).

  • — Smetana, 1978: 119 (Ocypus; subgenus Alapsodus; collecting notes; Israel).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Cyprus, Israel, Jordan, Iran, Russia.

rubripes Bernhauer and Schubert, 1914, see: eppelsheimianus Jakobson, 1909.

rufipalpis Lacordaire, 1835, see: morsitans Rossi, 1790.

rufipes Latreille, 1806, see: pedator Gravenhorst, 1802.

rufipes Eppelsheim, 1883, see: eppelsheimianus Jakobson, 1909.

rufipes Verhoeff, 1890, see: pedator Gravenhorst, 1802.

sahlbergi G. Müller, 1926, see: melanarius Heer, 1839.

saulcyi Reiche, 1857, see: melanarius Heer, 1839.

sicanus Coiffait, 1964, see: globulifer Goeffroy, 1785.

siculus Aubé, 1842, see: pedator Gravenhorst, 1802.

siculus Stierlin, 1864, see: globulifer Geoffroy, 1785.

sikkimensis Bernhauer, 1920d: 185 (Staphylinus; Type locality: Darjeeling: Sikkim).

  • — Cameron, 1932: 206 (Staphylinus; subgenus Tasgius; characters; India).

  • — Scheerpeltz, 1933: 1402 (Staphylinus; catalog).

  • — Coiffait, 1977b: 212 (Tasgius; Bhutan).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Tasgius).

  • Distribution: India, Bhutan.

similis Paykull, 1789, see: melanarius Heer, 1839.

singeri Hubenthal, 1911, see: winkleri Bernhauer, 1906.

solskyi Fauvel, 1875a: XXVIII [= 1875b: 230] (Staphylinus; Type locality: Sarepta).

  • — Bernhauer and Schubert, 1914: 390 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; catalog).

  • — Coiffait, 1974: 540 (Alapsodus; subgenus Paralapsodus; characters; Russia; Crimea).

  • — Smetana and Davies, 2000: 36, 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Ukraine, Russia, Turkey.

bosphoranus G. Müller, 1926: 22 (Staphylinus; subgenus Ocypus; subspecies of solskyi; cited as bosporanus; [Note: G. Müller published two spellings of this name. In the article he evidently intended as the original description (1926b: 48) he used bosphoranus but in a description that appeared six days earlier (1926: 22) he used bosporanus but cited the 1926b article as the original description. Although in English the strait is called Bosporus and Bosphorus is considered an incorrect spelling, in Italian it is Bòsforo. I assume that the intended spelling was bosphoranus, which subsequent authors have used, and that bosporanus was a lapsus]; Type locality: Konstantinopel).

  • — G. Müller, 1926b: 48 (Staphylinus; subgenus Ocypus; subspecies of solskyi; cited as bosphoranus; characters; Turkey).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; variety of solskyi; cited as bosphoranus).

  • — Coiffait, 1974: 540 (Alapsodus; subspecies of solskyi; cited as bosphoranus; Turkey).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila; subspecies of solskyi).

transversiceps Luze, 1904: 93 (Staphylinus; subgenus Tasgius; Type locality: Seravschan: Fl. Sching).

  • — Bernhauer and Schubert, 1914: 391 (Staphylinus; subgenus Ocypus; catalog).

  • — Coiffait, 1964: 83 (Tasgius; subgenus Paratasgius).

  • — Coiffait, 1974: 522 (Tasgius; characters; Turkestan).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Tasgius).

  • Distribution: Uzbekistan.

tricinctus Aragona, 1830: 11 (Staphylinus; Type locality: Apenini Derthonensii et Bobiensii).

  • — Fauvel, 1867: LIV (Staphylinus; notes; Italy).

  • — Fauvel, 1874: 193 [= 1874a: 419] (Staphylinus; characters; Italy).

  • — Bernhauer and Schubert, 1914: 391 (Staphylinus; subgenus Ocypus; catalog).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; catalog).

  • — Coiffait, 1974: 535 (Alapsodus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; Italy).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Italy.

etruscus Gautier, 1860: 368 (Ocypus; Type locality: Florence).

  • — Gemminger and Harold, 1868: 584 (Ocypus; synonym of tricinctus).

  • — Fauvel, 1874: 193 [= 1874a: 419] (Staphylinus; synonym of tricinctus).

  • — Bernhauer and Schubert, 1914: 391 (Staphylinus; synonym of tricinctus).

  • — Coiffait, 1974: 535 (Alapsodus; synonym of tricinctus).

tricolor Coiffait, 1974: 538 (Alapsodus; Type locality: Chiraki Vachlovani, Gruzzia).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Georgia.

walkeri Fauvel, 1898: 99 (Staphylinus; subgenus Ocypus; Type locality: Maroc: Esmir).

  • — Fauvel, 1902b: 106 (Staphylinus; subgenus Ocypus; Morocco).

  • — Scheerpeltz, 1933: 1404 (Staphylinus; catalog).

  • — Coiffait, 1964: 83 (Metocypus).

  • — Coiffait, 1974: 554 (Metocypus; characters; Morocco).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • Distribution: Morocco.

winkleri Bernhauer, 1906: 126 (Staphylinus; subgenus Ocypus; variety of globulifer; Type locality: Metkovisc und Corfu).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; synonym of globulifer).

  • — G. Müller, 1924: 27 (Staphylinus; subgenus Anodus; valid species; notes).

  • — G. Müller, 1926: 23 (Staphylinus; subgenus Ocypus; characters; notes).

  • — Edmonds, 1930: 273 (Staphylinus; characters in key; Britain).

  • — V. Hansen, 1931: 80 (Staphylinus; characters).

  • — Van der Wiel, 1932: 150 (Staphylinus; characters; Holland).

  • — G. Müller, 1932: 86, 87 (Staphylinus; characters; England; Germany).

  • — Scheerpeltz, 1933: 1404 (Staphylinus; catalog).

  • — Méquignon, 1933: 228 (Staphylinus; distinguishing characters).

  • — Tottenham, 1949: 420 (Staphylinus; checklist; Britain).

  • — Steel, 1949: 274 (Staphylinus; subgenus Ocypus; valid species; characters; Britain).

  • — V. Hansen, 1952: 107 (Staphylinus; subgenus Ocypus; characters; Denmark).

  • — Coiffait, 1956e: 219, 220 (Ocypus; subgenus Ocypus; characters).

  • — Smetana, 1958: 301 (Ocypus; subgenus Alapsodus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960b: 259 (Ocypus; Czechoslovakia).

  • — Scheerpeltz, 1961d: 132 (Ocypus; Italy).

  • — Palm, 1963b: 119 (Staphylinus; subgenus Ocypus; characters; Sweden; Denmark).

  • — Smetana, 1963b: 39 (Ocypus; subgenus Alapsodus; Madeira).

  • — Lohse, 1964: 202 (Ocypus; subgenus Ocypus; characters; central Europe).

  • — Coiffait, 1964: 83 (Alapsodus; subgenus Allocypus).

  • — Horion, 1965: 234 (Ocypus; Corsica; Italy; Dalmatia; Greece; Serbia; Holland; Belgium; France; Germany; Austria; Switzerland; Czech Republic; Slovakia; Denmark; Sweden; Britain).

  • — Scheerpeltz, 1968a: 85 (Ocypus; catalog; Austria).

  • — Coiffait, 1974: 542 (Alapsodus; subgenus Allocypus; characters; England; Portugal; Asia Minor; Armenia).

  • — Bordoni, 1974b: 9 (Alapsodus; subgenus Allocypus; notes; Italy).

  • — Bordoni and Carfi, 1978 (Alapsodus; note on peg setae of aedeagus).

  • — Muona, 1979: 15 (Ocypus; Sweden; Denmark).

  • — Tóth, 1984: 33 (Ocypus; characters; Hungary).

  • — Segers, 1986: 48 (Ocypus; checklist of species; Belgium).

  • — Newton, 1987: 382 (Staphylinus; introduced species; New York; Connecticut; Massachusetts; New Jersey; Rhode Island; California; Oregon; British Columbia).

  • — Ádám, 1987: 149 (Allocypus; collecting notes; Hungary).

  • — Lucht, 1987: 108 (Ocypus; checklist; Germany; Czechoslovakia; Austria; France; Benelux; Denmark; Sweden).

  • — Schülke and Uhlig, 1988: 12 (Ocypus; Germany).

  • — Whitehead, 1989: 9 (Staphylinus; field identification; Britain).

  • — Whitehead, 1990c: 5 (Staphylinus; revised field identification; Britain).

  • — Pilon, 1991: 105 (Ocypus; notes; Italy).

  • — Pilon and Zanetti, 1991: 69 (Ocypus; Italy).

  • — Silfverberg, 1992: 17 (Ocypus; Sweden; Denmark).

  • — Smetana, 1993b: 49 (Staphylinus; subgenus Alapsodus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Ocypus; Italy).

  • — Kubisz and Melke, 1995: 18 (Ocypus; Poland).

  • — M. Hansen, 1996: 106 (Ocypus; checklist; Denmark).

  • — Sterrenburg and Schülke, 1997: 20 (Ocypus; Netherlands).

  • — Eisinger, 1997: 171 (Ocypus; collecting notes; Germany).

  • — Pilon, 1998: 97 (Ocypus; notes; Italy).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (Ocypus; Denmark).

  • — Smetana and Davies, 2000: 46 (Tasgius; subgenus Rayacheila).

  • — Zerche, 2000: 52, 53 (Ocypus; discussion; nomen protectum).

  • — Burakowski, Mroczkowski, and Stefańska, 2000: 52 (Ocypus; subgenus Ocypus; catalog; Poland).

  • — Assing, 2001: 75 (Ocypus; collected from xerothermous site; Germany).

  • — Herman, 2001: 32 (Tasgius; nomen protectum).

  • Distribution: Madeira, Europe, Turkey, Armenia, Lebanon, Canada, USA.

edentulus Block, 1799: 115 (Staphylinus; Type locality: Potschapel, Plauischer Grund bei Dresden).

  • — Erichson, 1840: 417 (Ocypus; synonym of morio).

  • — Kraatz, 1857d: 563 (Ocypus; synonym of morio).

  • — Fauvel, 1874: 193 [= 1874a: 419] (Staphylinus; characters; France; Syria; Turkey; Caucasus).

  • — Ganglbauer, 1895: 436 (Staphylinus; synonym of globulifer).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; synonym of globulifer).

  • — Portevin, 1929: 353 (Staphylinus; characters; France).

  • — Scheerpeltz, 1933: 1394 (Staphylinus; synonym of globulifer).

  • — Steel, 1949: 273 (Staphylinus; subgenus Ocypus; synonym of globulifer).

  • — Zerche, 2000: 52, 53 (Ocypus; synonym of winkleri; discussion; nomen oblitum).

morio Gravenhorst, 1802: 6 (Staphylinus; [Note: This name is older than winkleri]; Type locality: Not cited, but presumably Brunsvigae).

  • — Latreille, 1804: 303 (Staphylinus; characters; France).

  • — Gravenhorst, 1806: 112 (Staphylinus).

  • — Gyllenhal, 1827: 473 (Staphylinus; characters; Sweden).

  • — Zetterstedt, 1828: 69 (Staphylinus; characters; Lapland).

  • — Curtis, 1829: 24 (Ocypus; catalog; Britain).

  • — Mannerheim, 1830: 24 (Staphylinus; Sweden; Austria).

  • — Mannerheim, 1831: 438 (Staphylinus; Sweden; Austria).

  • — Stephens, 1832: 209 (Goerius; characters; England).

  • — Runde, 1835: 3 (Staphylinus; characters; Germany).

  • — Lacordaire, 1835: 369 (Emus; characters; France).

  • — Nordmann, 1837: 13 (Anodus; notes; Europe).

  • — Zetterstedt, 1838: 60 (Staphylinus; characters; habitat; Lapland).

  • — Heer, 1839: 256 (Staphylinus; characters; Switzerland).

  • — Stephens, 1839: 388 (Goerius; characters; England).

  • — Laporte, 1840: 176 (Staphylinus; characters).

  • — Dufour, 1843: 33 (Staphylinus; characters; France).

  • — Gravenhorst, 1846: 98 (Staphylinus; characters; discussion; Germany; Switzerland; France; Italy; England; Sweden; Hungary; Africa).

  • — Jacobsen, 1849: 686 (Ocypus; characters; Denmark).

  • — Hochhuth, 1849: 126 (Ocypus; characters; Iberia).

  • — Redtenbacher, 1849: 696 (Ocypus; characters; Austria).

  • — Küster, 1853: 64 (Ocypus; characters; Europe).

  • — Hardy, 1851: 24 (Ocypus; subgenus Anodus; notes; England).

  • — Fairmaire and Laboulbène, 1856: 512 (Staphylinus; characters; France).

  • — Redtenbacher, 1857: 186 (Ocypus; characters; Austria).

  • — Thomson, 1859: 24 (type species of Anodus).

  • — Thomson, 1860: 150 (Anodus; characters; Scandinavia).

  • — Mulsant and Rey, 1876: 328 [= 1877: 184] (Anodus; [Note: According to Smetana, 1958: 298, 299, Mulsant and Rey's use of morio also included misidentifications of globulifer and melanarius]; characters; France).

  • — Fauvel, 1874: 193 [= 1874a: 419] (Staphylinus; synonym of edentulus).

  • — Redtenbacher, 1874: 205 (Ocypus; characters; Austria).

  • — Seidlitz, 1875: 271 (Ocypus; characters; Baltic region).

  • — J. Sahlberg, 1876: 12 (Anodus; Finland).

  • — Fowler, 1888: 258 (Ocypus; characters; Britain).

  • — Ganglbauer, 1895: 436 (Staphylinus; synonym of globulifer).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; synonym of globulifer).

  • — Scheerpeltz, 1933: 1404 (Staphylinus; synonym of winkleri).

  • — Steel, 1949: 273 (Staphylinus; subgenus Ocypus; synonym of globulifer).

  • — Smetana, 1958: 301 (Ocypus; synonym of winkleri).

  • — Coiffait, 1974: 543 (Alapsodus; synonym of winkleri).

  • — Herman, 2001: 32 (Tasgius; nomen oblitum).

picipes Stephens, 1832: 212 (Ocypus; [Note: This name is older than winkleri]; Type locality: London; Devonshire).

  • — Stephens, 1839: 388 (Ocypus; characters; England).

  • — G. Waterhouse, 1858: 24 (Ocypus; synonym of morio).

  • — Herman, 2001: 32 (Tasgius; nomen oblitum).

erosicollis Reiche and Saulcy, 1856: 364 (Ocypus; [Note: This name is older than winkleri]; Type locality: Beyrouth).

  • — Fauvel, 1874: 193 [= 1874a: 419] (Staphylinus; synonym of edentulus).

  • — Ganglbauer, 1895: 436 (Staphylinus; synonym of globulifer).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; synonym of globulifer).

  • — Scheerpeltz, 1933: 1404 (Staphylinus; synonym of winkleri).

  • — Smetana, 1958: 301 (Ocypus; synonym of winkleri).

  • — Coiffait, 1974: 543 (Alapsodus; synonym of winkleri).

  • — Herman, 2001: 32 (Tasgius; nomen oblitum).

singeri Hubenthal, 1911a: 189 (Staphylinus; variety of globulifer; Type locality: Passeierthal, St. Leonhard).

  • — Bernhauer and Schubert, 1914: 387 (Staphylinus; synonym of globulifer).

  • — Scheerpeltz, 1933: 1404 (Staphylinus; variety of winkleri).

  • — Smetana, 1958: 302 (Ocypus; synonym of globulifer).

  • — Coiffait, 1974: 543 (Alapsodus; variety of winkleri).

Thinopinus [1 species; Nearctic Region]

Thinopinus LeConte, 1852: 215 (species included: pictus). Type species: Thinopinus pictus LeConte, fixed by monotypy.

  • — Lacordaire, 1854: 75 (characters; notes; list of species).

  • — LeConte, 1861a: 64 (characters).

  • — LeConte, 1863: 23 (list of North American species).

  • — Gemminger and Harold, 1868: 576 (catalog).

  • — Horn, 1879: 200 (characters).

  • — Bernhauer and Schubert, 1914: 399 (world catalog; 2 species).

  • — Leng, 1920: 109 (catalog of North American species).

  • — Scheerpeltz, 1933: 1413 (world catalog supplement).

  • — Blackwelder, 1952: 387 (type species: pictus).

  • — Hatch, 1957: 212 (characters; species of Pacific Northwest).

  • — Moore, 1964c (larval characters).

  • — Moore and Legner, 1975: 44 (catalog; North America north of Mexico).

  • — Moore and Legner, 1976: 540 (includes seashore species; notes).

  • — Frank, 1979a (some larval characters and their use in classification).

  • — Moore and Legner, 1979: 35 (characters; notes).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 395 (1 Nearctic species; notes; characters in key).

Trichocanthus Motschulsky, 1853a: 78 (species included: variegatus). Type species: Trichocanthus variegatus Motschulsky, fixed by monotypy.

  • — Mannerheim, 1853: 187 (characters).

  • — Gemminger and Harold, 1868: 576 (synonym of Thinopinus).

  • — Bernhauer and Schubert, 1914: 399 (synonym of Thinopinus).

  • — Blackwelder, 1952: 394 (type species: variegatus; synonym of Thinopinus; notes on authorship of name).

pictus LeConte, 1852: 216 (Thinopinus; Type locality: San Diego).

  • — Horn, 1879: 200 (Thinopinus; characters; Pacific sea shore from Alaska to San Diego).

  • — Bernhauer and Schubert, 1914: 399 (Thinopinus; catalog).

  • — Scheerpeltz, 1933: 1413 (Thinopinus; catalog).

  • — Hatch, 1957: 212 (Thinopinus; characters; British Columbia; Washington; Oregon).

  • — Malkin, 1959: 20 (Thinopinus; body color and sand color).

  • — Craig, 1970: 1012 (Thinopinus; behavior; distribution in habitat).

  • — Moore, 1975b: 92 (Thinopinus; habitat; feeding).

  • — Moore, 1977 (Thinopinus; color phases and substrate color).

  • — Orth, Moore, and Fisher, 1978: 187 (Thinopinus; characters; collecting notes; California).

  • — Richards, 1982 (Thinopinus; feeding behavior).

  • — Richards, 1983 (Thinopinus; natural history; feeding behavior; activity patterns; British Columbia).

  • Distribution: Canada, USA.

variegatus Motschulsky, 1853a: 78 (Trichocanthus; Type locality: littoral occidental Amérique septentrionale).

  • — Mäklin, 1853: 188 (Trichocanthus; characters; Americes borealis occidentalis).

  • — LeConte, 1861: 342 (Thinopinus; characters; California).

  • — Bernhauer and Schubert, 1914: 399 (Thinopinus; catalog).

  • — Scheerpeltz, 1933: 1414 (Thinopinus; aberration of pictus).

  • — Moore, 1977: 377 (Thinopinus; synonym of pictus).

  • — Campbell and Davies, 1991: 120 (Thinopinus; cited as valid species; checklist of species; Canada).

variegatus Motschulsky, 1853, see: pictus LeConte, 1852.

Thoracostrongylus [13 species; Palaearctic and Oriental Regions]

Thoracostrongylus Bernhauer, 1915h: 233 (subgenus of Ontholestes; species included: javanus). Type species: Ontholestes (Thoracostrongylus) javanus Bernhauer, fixed by monotypy.

  • — Cameron, 1932: 213 (characters; key to species of British India).

  • — Scheerpeltz, 1933: 1406 (world catalog supplement).

  • — Blackwelder, 1952: 388 (type species: javanus).

  • — Smetana, 1959d: 394 (characters).

  • — Shibata, 1973c: 62 (checklist of species of Taiwan).

  • — Hammond, 1984: 207 (checklist of species of Borneo).

  • — Shibata, 1986a: 124 (supplemental list of species from Taiwan).

  • — Smetana and Davies, 2000: 19, 23, 39 (characters; checklist of species of north temperate region).

Paramichrotus Cameron, 1932: 213 (published as a synonym of Thoracostrongylus; [Note: Nomen nudum (Article 11.6)]).

Parontholestes Coiffait, 1982: 71 (species included: nepalicus). Type species: Parontholestes nepalicus Coiffait, fixed by original designation and monotypy.

  • — Smetana and Davies, 2000: 23, 39 (synonym of Thoracostrongylus).

apicicornis Cameron, 1937: 22 (Thoracostrongylus; Type locality: Java: Batoerraden, G. Slamat).

  • Distribution: Indonesia.

birmanus Fauvel, 1895: 246 (Leistotrophus; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Carin Asciuii Ghécu, 1400–1500 m).

  • — Bernhauer and Schubert, 1914: 392 (Ontholestes; catalog).

  • — Gridelli, 1924f: 207, 208 (Ontholestes; subgenus Thoracostrongylus; characters; Burma).

  • — Cameron, 1932: 214 (Thoracostrongylus; characters; Burma; India).

  • — Scheerpeltz, 1933: 1406 (Thoracostrongylus; catalog).

  • — Smetana and Davies, 2000: 39 (Thoracostrongylus).

  • Distribution: Myanmar, India.

bryanti Cameron, 1937, see: Amichrotus.

costatus Fauvel, 1895: 269 (Amichrotus; Type locality: Bírmanie, Carin Ghécu, 1300–1400 m).

  • — Bernhauer and Schubert, 1914: 367 (Amichrotus; catalog).

  • — Cameron, 1932: 217 (Thoracostrongylus; characters; Burma).

  • — Scheerpeltz, 1933: 1407 (Thoracostrongylus; catalog).

  • — Smetana and Davies, 2000: 39 (Thoracostrongylus).

  • Distribution: Myanmar.

distinctus Cameron, 1932, see: Amichrotus.

formosanus Shibata, 1982: 71 (Thoracostrongylus; Type locality: Taiwan: near Tsuifeng, about 2200 m alt., Nantou Hsien).

  • — Smetana and Davies, 2000: 39 (Thoracostrongylus).

  • Distribution: Taiwan.

javanus Bernhauer, 1915h: 233 (Ontholestes; subgenus Thoracostrongylus; Type locality: Java: Preanger).

  • — Scheerpeltz, 1933: 1407 (Thoracostrongylus; catalog).

  • Distribution: Indonesia.

malaisei Scheerpeltz, 1965: 245 (Thoracostrongylus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).

  • — Smetana and Davies, 2000: 39 (Thoracostrongylus).

  • Distribution: Myanmar.

martensi Coiffait, 1982, see: Abemus.

miyakei Bernhauer, 1943a: 179 (Thoracostrongylus; Type locality: Formosa: Shinchiku).

  • — Shibata, 1982: 74 (Thoracostrongylus; characters; Taiwan).

  • — Smetana and Davies, 2000: 39 (Thoracostrongylus).

  • Distribution: Taiwan.

modiglianii Gridelli, 1924f: 207, 209 (Ontholestes; subgenus Thoracostrongylus; Type locality: Sumatra: Foresta di Si Rambe).

  • — Scheerpeltz, 1933: 1407 (Thoracostrongylus; catalog).

  • — Cameron, 1933d: 391 (Thoracostrongylus; Sumatra).

  • Distribution: Indonesia.

nepalicus Coiffait, 1982: 72 (Parontholestes; Type locality: Nepal: Trisuli Tal, bei Dunche, 2000 m).

  • — Smetana and Davies, 2000: 23, 39 (Thoracostrongylus).

  • Distribution: Nepal.

picticollis Cameron, 1937, see: Amichrotus.

ruficollis Cameron, 1932, see: Amichrotus.

sarawakensis Bernhauer, 1915h: 233 (Amichrotus; Type locality: Sarawak: Mt. Matang).

  • — Cameron, 1928f: 410 (Amichrotus; Borneo).

  • — Scheerpeltz, 1933: 1376 (Amichorus; catalog).

  • — Hammond, 1984: 194, 195 (Thoracostrongylus; Borneo).

  • Distribution: Borneo.

semicyaneus Fauvel, 1895: 269 (Amichrotus; Type locality: Tenasserim, mont Mooleyit, 1800–1900 m).

  • — Bernhauer and Schubert, 1914: 367 (Amichrotus; catalog).

  • — Cameron, 1932: 216 (Thoracostrongylus; characters; Burma).

  • — Scheerpeltz, 1933: 1407 (Thoracostrongylus; catalog).

  • Distribution: Myanmar.

sumatrensis Gridelli, 1924f: 207, 211 (Ontholestes; subgenus Thoracostrongylus; Type locality: Sumatra: Foresta di Si Rambe).

  • — Scheerpeltz, 1933: 1407 (Thoracostrongylus; catalog).

  • — Cameron, 1933d: 391 (Thoracostrongylus; Sumatra).

  • Distribution: Indonesia.

velutinus Scheerpeltz, 1965: 243 (Thoracostrongylus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).

  • — Smetana and Davies, 2000: 39 (Thoracostrongylus).

  • Distribution: Myanmar.

Trichocosmetes [4 species; Palaearctic Region]

Trichocosmetes Kraatz, 1859: 69 (species included: leucomus). Type species: Staphylinus leucomus Erichson, fixed by monotypy.

  • — Gemminger and Harold, 1868: 577 (catalog).

  • — Bernhauer and Schubert, 1914: 394 (world catalog; 2 species).

  • — Cameron, 1925: 69 (catalog of Indian species).

  • — Cameron, 1932: 233 (characters).

  • — Scheerpeltz, 1933: 1408 (world catalog supplement).

  • — Blackwelder, 1952: 394 (type species: leucomus).

antennalis Cameron, 1932: 231 (Rhyncocheilus, cited as Rhyncochilus; Type locality: Shillong).

  • — Scheerpeltz, 1933: 1408 (Rhyncocheilus; catalog).

  • — Chapman, 1933a: 222 (Rhyncocheilus).

  • — Schillhammer, 1999a: 62 (Trichocosmetes).

  • Distribution: India.

cinctus Kollar and Redtenbacher, 1848, see: leucomus Erichson, 1839.

franzi Coiffait, 1977, see: leucomus Erichson, 1839.

leucomus Erichson, 1839b: 362 (Staphylinus; Type locality: Nepal).

  • — Kraatz, 1859: 70 (Trichocosmetes; characters; Nepal).

  • — Bernhauer and Schubert, 1914: 394 (Trichocosmetes; catalog).

  • — Cameron, 1932: 234 (Trichocosmetes; characters; Nepal; India).

  • — Scheerpeltz, 1933: 1408 (Trichocosmetes; catalog).

  • — Coiffait, 1982: 31 (Trichocosmetes; Nepal).

  • Distribution: Nepal, India.

cinctus Kollar and Redtenbacher, 1848: 504 (Staphylinus; [preoccupied]; Type locality: Not cited but presumably Kashmir).

  • — Kraatz, 1859: 70 (Trichocosmetes; synonym of leucomus).

  • — Bernhauer and Schubert, 1914: 394 (Trichocosmetes; synonym of leucomus).

  • — Cameron, 1932: 234 (Trichocosmetes; synonym of leucomus).

franzi Coiffait, 1977: 247 (Rhyncocheilus; Type locality: Environs de Maharigaon, 3000–3500 m, région de Jumla, Népal occidental).

  • — Coiffait, 1981d: 324 (Rhyncocheilus, cited as Rhynchochilus; Nepal).

  • — Schillhammer, 1994: 74 (Trichocosmetes; synonym of leucomus).

prainae Eppelsheim, 1895, see: Dinothenarus.

reitteri Bernhauer, 1934: 10 (Trichocosmetes; Type locality: Szechuan: Giufu-Shan, 1600 m).

  • Distribution: China.

varicornis Coiffait, 1977b: 235 (Eucibdelus; Type locality: Bhutan: 87 km de Phuntsholing).

  • — Schillhammer, 1999a: 62 (Trichocosmetes).

  • Distribution: Bhutan.

Wasmannellus [2 species; Palaearctic Region]

Wasmannellus Bernhauer, 1920d: 186 (species included: tristis). Type species: Wasmanellus tristis Bernhauer, fixed by monotypy.

  • — Cameron, 1925: 70 (catalog of Indian species).

  • — Cameron, 1932: 246 (characters).

  • — Scheerpeltz, 1933: 1410 (world catalog supplement).

  • — Blackwelder, 1952: 403 (type species: tristis).

  • — Smetana and Davies, 2000: 21, 35, 45 (subgenus of Ocypus; characters; checklist of species of north temperate region).

stevensi Cameron, 1932: 208 (Staphylinus; subgenus Tasgius; Type locality: Darjeeling: Gopaldhara).

  • — Scheerpeltz, 1933: 1403 (Staphylinus; catalog).

  • — Smetana and Davies, 2000: 45 (Wasmannellus).

  • Distribution: India.

tristis Bernhauer, 1920d: 187 (Wasmannellus; Type locality: Darjeeling, Sikkim).

  • — Cameron, 1932: 247 (Wasmannellus; characters; Sikkim).

  • — Scheerpeltz, 1933: 1410 (Wasmannellus; catalog).

  • — Smetana and Davies, 2000: 45 (Wasmannellus).

  • Distribution: India.

Subtribe Tanygnathinina

Tanygnathinina Reitter, 1909.

  • — Newton and Thayer, 1992: 65 (subtribe of Staphylinini).

  • — Smetana, 1995e: 48 (characters).

  • — Smetana and Davies, 2000: 12 (characters).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 393 (characters; notes).

Tanygnathinini Reitter, 1909: 105 (genera included: Tanygnathinus). Type genus: Tanygnathinus (see Newton and Thayer, 1992: 65).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 155 (catalog; Poland).

  • — Outerelo and Gamarra, 1985: 67 (genera of Iberian Peninsula).

  • — Smetana, 1988: 368 (synonym of Atanygnathini).

  • — Assing, Frisch, Kahlen, et al., 1998: 139 (note).

Tanygnathini Casey, 1915: 424 (genera included: Tanygnathus and Acylophorus). Type genus: Tanygnathus (see Newton and Thayer, 1992: 66).

  • — Portevin, 1929: 333 (characters).

  • — Newton and Thayer, 1992: 66 (synonym of Tanygnathinina).

  • — Smetana, 1995: 14, 23, 133 (characters; revision of species of Taiwan; key to species of Taiwan; notes).

Atanygnathini Lohse, 1964: 220 (characters; central European species; genera included: Atanygnathus). Type genus: Atanygnathus (see Newton and Thayer, 1992: 66).

  • — Scheerpeltz, 1968a: 97 (catalog of Austrian species).

  • — Muona, 1979: 16 (list of Scandinavian species).

  • — Smetana, 1984c: 279 (characters; valid subtribe).

  • — Smetana, 1988: 368 (characters).

  • — Newton and Thayer, 1992: 66 (synonym of Tanygnathinina).

Atanygnathus [45 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, and Australian Regions]

Atanygnathus Jakobson, 1909: 521 (replacement name for Tanygnathus Erichson). Type species: Tanygnathus terminalis Erichson, fixed by objective synonymy with Tanygnathus.

  • — Bernhauer and Schubert, 1916: 439 (world catalog; 15 species).

  • — Cameron, 1921a: 355, 406 (characters; key to Singapore species; catalog).

  • — Winkler, 1925: 393 (catalog for Palaearctic region).

  • — Cameron, 1925: 74 (catalog of Indian species).

  • — Cameron, 1932: 308 (characters; key to species of British India).

  • — Scheerpeltz, 1933: 1467 (world catalog supplement).

  • — Blackwelder, 1943: 471 (characters; type species: terminalis; key to species of West Indies).

  • — Blackwelder, 1944: 145 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — V. Hansen, 1952: 163 (characters).

  • — Blackwelder, 1952: 67 (type species).

  • — Kocher, 1958: 161 (checklist of species; Morocco).

  • — Smetana, 1958: 401 (characters; habitus of one species; type species: terminalis; Czechoslovakian species).

  • — Scheerpeltz, 1962b: 268 (characters).

  • — Székessy, 1963: 62 (characters).

  • — Palm, 1963b: 159 (characters; key to species of Sweden).

  • — Lohse, 1964: 220 (characters; central European species).

  • — Scheerpeltz, 1968a: 97 (catalog of Austrian species).

  • — Smetana, 1971: 269 (revision of and key to species of Canada and United States; characters).

  • — Last, 1975: 442 (characters).

  • — Moore and Legner, 1975: 52 (catalog; North America north of Mexico).

  • — Tikhomirova, 1973a: 190 (checklist of species of USSR).

  • — Smetana, 1977b: 181 (characters).

  • — Coiffait, 1978: 319 (characters; key to species of western Palaearctic region).

  • — Moore and Legner, 1979: 43 (characters; notes).

  • — Muona, 1979: 16 (list of Scandinavian species).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 155 (catalog; Poland).

  • — Uhlig and Vogler, 1981: 126 (list of some species collected in Germany; habitat and collecting notes).

  • — Hammond, 1984: 205 (checklist of species of Borneo).

  • — Tóth, 1984: 141 (characters; habitus; species of Hungary).

  • — Shibata, 1984: 140 (checklist of species of Japan).

  • — Frank, 1986a: 371 (checklist of species of Florida).

  • — Smetana, 1988: 368 (characters; key to species of Himalayan region).

  • — Silfverberg, 1992: 18 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Smetana, 1993b: 50 (in list of species of Czech Republic and Slovakia).

  • — Smetana, 1995: 134 (notes; key to species of Taiwan).

  • — Ciceroni and Zanetti, 1995: 33 (list of species of Italy).

  • — Downie and Arnett, 1996: 395 (characters).

  • — Lundgren, 1998: 50 (list of species from Florida; records for other States listed for each species).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 393 (3 Nearctic species; notes; characters in key).

Tanygnathus Erichson, 1839a: 417 (species included: terminalis; preoccupied by Wagler, 1832). Type species: Tanygnathus terminalis Erichson, fixed by monotypy.

  • — Erichson, 1839b: 288 (characters).

  • — Redtenbacher, 1849: 689 (characters).

  • — Schaum, 1852: 25 (list of species; Europe).

  • — Lacordaire, 1854: 60 (characters; notes; list of species).

  • — Fairmaire and Laboulbène, 1856: 496 (characters).

  • — Jacquelin du Val, 1856: 29 (characters).

  • — Kraatz, 1857d: 477 (characters).

  • — Redtenbacher, 1857: 178 (characters).

  • — Thomson, 1859: 26 (characters; type species: terminalis).

  • — Schaum, 1859: 25 (catalog; European species).

  • — Thomson, 1860: 182 (characters).

  • — Gemminger and Harold, 1868: 565 (catalog).

  • — Jacquelin du Val, 1868: 65 (catalog; European species).

  • — Fauvel, 1874: 317 [= 1874a: 543] (characters).

  • — Redtenbacher, 1874: 195 (characters).

  • — Fauvel, 1875a: XXXIV [= 1875b: 236] (catalog).

  • — Mulsant and Rey, 1876: 387 [= 1877: 693] (characters).

  • — Sharp, 1876: 96 (notes).

  • — Horn, 1878: 149 (characters).

  • — Duvivier, 1883: 135 (catalog).

  • — Sharp, 1884: 321 (characters; notes; Mexican and Central American species).

  • — Heyden, Reitter, and Weise, 1891: 98 (list of species of Europe and Caucasus).

  • — Ganglbauer, 1895: 379 (characters).

  • — Heyden, Reitter, and Weise, 1906: 177 (list of species of Europe and Caucasus).

  • — Porta, 1907: 88 (key to Italian species).

  • — Petri, 1912: 81 (list of species of Siebenbürgen).

  • — Johansen, 1914: 341 (characters).

  • — Casey, 1915: 424 (characters; key to species).

  • — Bernhauer and Schubert, 1916: 439 (synonym of Atanygnathus).

  • — Leng, 1920: 109 (catalog of North American species).

  • — Porta, 1926: 119 (characters; species of Italy).

  • — Cameron, 1932: 308 (synonym of Atanygnathus).

  • — Scheerpeltz, 1933: 1467 (synonym of Atanygnathus).

  • — Blackwelder, 1943: 471 (synonym of Atanygnathus; type species: terminalis).

  • — Blackwelder, 1952: 373 (type species: terminalis).

  • — Smetana, 1958: 401 (synonym of Atanygnathus).

  • — Coiffait, 1978: 319 (synonym of Atanygnathus).

  • — Smetana, 1988: 368 (synonym of Atanygnathus).

Tanygnathinus Reitter, 1909: 105 (replacement name for Tanygnathus Erichson). Type species: Tanygnathus terminalis Erichson, fixed by objective synonymy with Tanygnathus.

  • — Bernhauer and Schubert, 1916: 439 (synonym of Atanygnathus).

  • — Cameron, 1932: 308 (synonym of Atanygnathus).

  • — Blackwelder, 1943: 471 (synonym of Atanygnathus; type species: terminalis).

  • — Blackwelder, 1952: 373 (type species).

  • — Smetana, 1958: 401 (synonym of Atanygnathus).

  • — Coiffait, 1978: 319 (synonym of Atanygnathus).

  • — Smetana, 1988: 368 (synonym of Atanygnathus).

acuminatus Casey, 1915: 425 (Tanygnathus; Type locality: Texas: Austin; [Note: See lectotype designation by Smetana, 1971]).

  • — Scheerpeltz, 1933: 1468 (Atanygnathus; catalog).

  • — Smetana, 1971: 273 (Atanygnathus; lectotype designation; characters; Arizona; California; Texas).

  • — Smetana, 1978a: 839 (Atanygnathus; collecting notes; Arizona; New Mexico).

  • — Smetana, 1981b: 644 (Atanygnathus; collecting notes; Arizona).

  • Distribution: USA.

africanus Bernhauer, 1929f: 125 (Atanygnathus; Type locality: Manyema: Niembe-Kalembe).

  • — Scheerpeltz, 1933: 1468 (Atanygnathus; catalog).

  • — Cameron, 1950g: 44 (Atanygnathus; Belgian Congo).

  • — Tottenham, 1956: 230, 313 (Atanygnathus; characters; Ruanda).

  • Distribution: Congo, Rwanda.

andamanensis Herman, 2001: 42 (Atanygnathus; replacement name for collaris Coiffait).

  • Distribution: Andaman Islands.

collaris Coiffait, 1981f: 338 (Atanygnathus; [preoccupied]; Type locality: South Andaman, Rutland Island).

  • — Herman, 2001: 42 (Atanygnathus; synonym of andamanensis).

antennalis Sharp, 1884: 321 (Tanygnathus; Type locality: Guatemala, Guatemala city, San Geronimo, San Joaquin in Vera Paz).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; catalog).

  • — Scheerpeltz, 1933: 1468 (Atanygnathus; catalog).

  • Distribution: Guatemala.

aquaticus Last, 1975: 443 (Atanygnathus; Type locality: Henegaru, Yani River, Okapa District).

  • Distribution: New Guinea.

australasiae Fauvel, 1878g: 562 (Tanygnathus; Type locality: Australie méridionale: Victoria).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; catalog).

  • — Bernhauer, 1920: 18 (Atanygnathus; variety of terminalis; Australia).

  • — Lea, 1925: 253 (Atanygnathus; variety of terminalis).

  • — Scheerpeltz, 1933: 1468 (Atanygnathus; catalog; Philippines).

  • Distribution: Australia, Philippines.

bicolor Casey, 1915: 424 (Tanygnathus; Type locality: Florida: Sand Point).

  • — Horn, 1878: 150 (Tanygnathus; [Note: Misidentification: Scheerpeltz, 1933: 1468, cited collaris Horn as a synonym of bicolor; Horn did not describe the species as new, he attributed it to Erichson]; characters; Florida).

  • — Notman, 1920b: 706 (Tanygnathus; Florida).

  • — Scheerpeltz, 1933: 1468 (Atanygnathus; catalog).

  • — Smetana, 1971: 270 (Atanygnathus; characters; Alabama; Arkansas; Florida; Georgia; Louisiana; Massachusetts; North Carolina).

  • — Smetana, 1978a: 838 (Atanygnathus; collecting notes; Alabama; Louisiana; Mississippi; Texas).

  • — Smetana, 1990d: 104 (Atanygnathus; New Hampshire; Oklahoma).

  • — Downie and Arnett, 1996: 395 (Atanygnathus; characters; USA).

  • Distribution: USA.

bicolor Lea, 1929, see: leai Scheerpeltz, 1933.

bindu Smetana, 1988: 376 (Atanygnathus; Type locality: Nepal: Khandbari Dis., Arun Valley at Num main bridge, 1050 m. Also cited from India).

  • Distribution: Nepal.

borealis Mulsant and Rey, 1876, see: terminalis Erichson, 1839.

bredoi Cameron, 1950c: 96 (Atanygnathus; Type locality: Belgian Congo: Lualaba, Maka).

  • Distribution: Congo.

brevicollis Fauvel, 1895: 276 (Tanygnathus; Type locality: Bírmanie, Carin Asciuii Ghécu, 1400–1500 m; [Note: See lectotype designation by Smetana, 1988]. Also cited from Bírmanie, Carin Ghécu, 1300–1400 m; Tenasserim, Thagata).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; catalog).

  • — Cameron, 1932: 311 (Atanygnathus; characters; Burma; India).

  • — Scheerpeltz, 1933: 1468 (Atanygnathus; catalog).

  • — Smetana, 1988: 373 (Atanygnathus; lectotype designation: Carin Asciui Ghécu, 1400–1500 m; characters; localities from India published by Cameron, 1932, apply to other species; Burma).

  • Distribution: Myanmar.

brevipennis Cameron, 1959: 118 (Atanygnathus; Type locality: Angola: Dundo: Dundundo River).

  • Distribution: Angola.

californicus Smetana, 1971: 274 (Atanygnathus; Type locality: Cal.: Palm Spgs.).

  • Distribution: USA.

chiso Smetana, 1988: 377 (Atanygnathus; Type locality: Nepal: Khandbari Dis., Arun Valley at Num main bridge, 1050 m. Also cited from India: Meghalaya).

  • Distribution: India, Nepal.

collaris Erichson, 1839b: 289 (Tanygnathus; Type locality: Guyana Venezuelensis).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; catalog).

  • Distribution: Venezuela.

collaris Coiffait, 1981, see: andamanensis Herman, 2001.

crassicornis Fauvel, 1905b: 182 (Tanygnathus; Type locality: Suberbieville).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; catalog).

  • Distribution: Madagascar.

crassulus Sharp, 1884: 322 (Tanygnathus; Type locality: Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; catalog).

  • Distribution: Nicaragua.

flavicollis Sharp, 1876: 97 (Tanygnathus; Type locality: Tapajos).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; catalog).

  • Distribution: Brazil.

fuscus Kraatz, 1859, see: piceus Motschulsky, 1858.

giganteus Levasseur, 1971: 306 (Atanygnathus; Type locality: Monts Loma, Sierra Leone, 1600 m).

  • Distribution: Sierra Leone.

gomyi Lecoq, 1990: 207 (Atanygnathus; Type locality: La Réunion: Saint André).

  • Distribution: Réunion.

heterocerus Cameron, 1922: 123 (Atanygnathus; variety of antennalis; Type locality: St. Vincent; Grenada).

  • — Scheerpeltz, 1933: 1468 (Atanygnathus; variety of antennalis).

  • — Blackwelder, 1943: 471 (Atanygnathus; valid species; characters; St. Vincent; Grenada).

  • Distribution: West Indies.

juang Smetana, 1995: 136 (Atanygnathus; Type locality: Taiwan: Lushan-Spa, Nantou Hsien).

  • Distribution: Taiwan.

laticollis Erichson, 1839b: 289 (Tanygnathus; Type locality: Amer. mer. ins. St. Vincentii).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; catalog).

  • — Blackwelder, 1943: 472 (Atanygnathus; characters; Cuba; Dominican Republic; Puerto Rico; Antigua; St. Vincent; Trinidad).

  • Distribution: West Indies.

leai Scheerpeltz, 1933: 1468 (Atanygnathus; replacement name for bicolor Lea).

  • Distribution: Australia.

bicolor Lea, 1929: 208 (Atanygnathus; [preoccupied]; Type locality: North Australia: Daly River; Oenpelli; Darwin; Adelaide River; Melville Island).

  • — Scheerpeltz, 1933: 1468 (Atanygnathus; synonym of leai).

lohsei Uhlig, 1995a: 102 (Atanygnathus; Type locality: Namibia: Kavango, Popa Falls, Kavango-Ufer, 18°07′S/21°35′E).

  • Distribution: Namibia.

lomaensis Levasseur, 1971: 304 (Atanygnathus; Type locality: Monts Loma, Sierra Leone, 1600 m).

  • Distribution: Sierra Leone.

longicornis Sharp, 1876: 96 (Tanygnathus; Type locality: Amazons).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; catalog).

  • Distribution: Brazil.

muticus Sharp, 1884: 321 (Tanygnathus; Type locality: Guatemala, Rio Naranja 450 feet; Panama, Bugaba, San Miguel one of the Pearl Islands).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; catalog).

  • Distribution: Guatemala, Panama.

nasutus Sharp, 1876: 96 (Tanygnathus; Type locality: Not cited but presumably the Amazonian region).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; catalog).

  • Distribution: “Amazonian region”.

noxius Last, 1975: 442 (Atanygnathus; Type locality: Okapa).

  • Distribution: New Guinea.

ornatus Cameron, 1949a: 322 (Atanygnathus; Type locality: Portuguese Guinea: Bafata).

  • — Cameron, 1952e: 832 (Atanygnathus; Dahomey).

  • — Levasseur, 1966a: 1216 (Atanygnathus; Portuguese Guinea; Upper Volta; Zaire; Congo).

  • — Levasseur, 1971: 304 (Atanygnathus; Ivory Coast; Sierra Leone).

  • — Last, 1975: 442 (Atanygnathus; New Guinea).

  • — Last, 1987: 55 (Atanygnathus; Papua New Guinea).

  • Distribution: Guinea-Bissau, Sierra Leone, Ivory Coast, Burkina Faso, Benin, Congo Republic, Congo, New Guinea.

paani Smetana, 1988: 375 (Atanygnathus; Type locality: Nepal: Lamjung Distr., Besisahar, 900 m. Also cited from India: Meghalaya; Assam; Uttar Pradesh).

  • Distribution: Nepal, India.

pallidus Last, 1980: 154 (Atanygnathus; Type locality: New Guinea: Kiunga).

  • Distribution: New Guinea.

pauliani Jarrige, 1970: 52 (Atanygnathus; Type locality: Madagascar: Mt. Tsaratanana, 1500 m).

  • Distribution: Madagascar.

piceus Motschulsky, 1858a: 213 (Tanygnathus; Type locality: Indes orientales; [Note: See lectotype designation by Smetana, 1988]).

  • — Fauvel, 1898a: 118 (Tanygnathus; Seychelles).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; catalog).

  • — Bernhauer, 1922b: 177 (Atanygnathus; Seychelles).

  • — Cameron, 1932: 310 (Atanygnathus; characters; India; Singapore; Malay Peninsula).

  • — Scheerpeltz, 1933: 1468 (Atanygnathus; catalog).

  • — Cameron, 1934d: 79 (Tanygnathus; type examined).

  • — Cameron, 1937: 30 (Atanygnathus; Java).

  • — Coiffait, 1981f: 330 (Atanygnathus; Andaman Islands).

  • — Smetana, 1988: 374 (Atanygnathus; lectotype designation; characters).

  • Distribution: Seychelles, India, Andaman Islands, Malaysia, Singapore, Borneo, Indonesia.

fuscus Kraatz, 1859: 65 (Tanygnathus; Type locality: India orientalis; [Note: See lectotype designation by Smetana, 1988]).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; catalog).

  • — Cameron, 1928f: 411 (Atanygnathus; Borneo).

  • — Cameron, 1932: 310 (Atanygnathus; characters; India).

  • — Scheerpeltz, 1933: 1468 (Atanygnathus; catalog).

  • — Cameron, 1934d: 79 (Tanygnathus; synonym of piceus).

  • — Smetana, 1988: 375 (Atanygnathus; lectotype designation; synonym of piceus).

pictus Motschulsky, 1858a: 213 (Tanygnathus; Type locality: Indes orientales; [Note: See lectotype designation by Smetana, 1988]).

  • — Fauvel, 1904a: 60 (Tanygnathus; synonym of terminalis).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; variety of terminalis).

  • — Cameron, 1932: 309 (Atanygnathus; variety of terminalis).

  • — Scheerpeltz, 1933: 1468 (Atanygnathus; variety of terminalis).

  • — Smetana, 1988: 370 (Atanygnathus; lectotype designation; characters; Assam; Sri Lanka).

  • — Smetana, 1995: 134 (Atanygnathus; characters; bionomics; Taiwan).

  • Distribution: India, Sri Lanka, Cambodia, Indonesia, Taiwan.

ruficollis Kraatz, 1859: 64 (Tanygnathus; Type locality: Ceylan; [Note: See lectotype designation by Smetana, 1988]. Also cited from India orientalis).

  • — Fauvel, 1904a: 60 (Tanygnathus; synonym of terminalis).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; variety of terminalis).

  • — Bernhauer, 1922a: 151 (Atanygnathus; Cambodia).

  • — Bernhauer, 1928: 18 (Atanygnathus; Sumatra).

  • — Cameron, 1932: 309 (Atanygnathus; variety of terminalis).

  • — Scheerpeltz, 1933: 1468 (Atanygnathus; variety of terminalis).

  • — Cameron, 1933d: 393 (Atanygnathus; variety of terminalis; Sumatra).

  • — Scheerpeltz, 1957c: 235 (Atanygnathus; variety of terminalis; Sumba).

  • — Frank, 1982: 33 (Atanygnathus; list of parasites).

  • — Smetana, 1988: 370 (Atanygnathus; lectotype designation: Ceylan; synonym of pictus).

  • — Smetana, 1995: 134 (Atanygnathus; synonym of pictus).

purba Smetana, 1988: 379 (Atanygnathus; Type locality: E. Nepal: Kosi Val. Arun ss/Num, 1050 m).

  • Distribution: Nepal.

robustus Cameron, 1950d: 24 (Atanygnathus; Type locality: Selangor: The Gap).

  • Distribution: Malaysia.

ruficollis Kraatz, 1859, see: pictus Motschulsky, 1858.

rufipennis Tottenham, 1961: 212 (Atanygnathus; Type locality: Senegal: Parc National du Niokolo-Koba, Badi).

  • Distribution: Senegal.

sasuraa Smetana, 1988: 372 (Atanygnathus; Type locality: Nepal: Prov. Bagmati, Tarang Marang, 1000 m. Also cited from India: Assam; Meghalaya; Uttar Pradesh; West Bengal).

  • Distribution: Nepal.

terminalis Erichson, 1839a: 418 (Tanygnathus; Type locality: Umgegend von Berlin; [Note: See lectotype designation by Uhlig, 1995a]).

  • — Erichson, 1839b: 288 (Tanygnathus; characters; Germany).

  • — Redtenbacher, 1849: 689 (Tanygnathus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 496 (Tanygnathus; characters; France).

  • — Kraatz, 1857d: 478 (Tanygnathus; characters; Germany).

  • — Redtenbacher, 1857: 178 (Tanygnathus; characters; Austria).

  • — Thomson, 1859: 26 (type species of Tanygnathus).

  • — Thomson, 1860: 183 (Tanygnathus; characters; Scandinavia).

  • — Fauvel, 1874: 318 [= 1874a: 544] (Tanygnathus; characters; France; Scandinavia; Russia; Germany; Austria).

  • — Redtenbacher, 1874: 195 (Tanygnathus; characters; Austria).

  • — Seidlitz, 1875: 263 (Tanygnathus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 839 [= 1877: 695] (Tanygnathus; characters; France).

  • — Seidlitz, 1891: 400 (Tanygnathus; characters; Europe).

  • — Seidlitz, 1891a: 427 (Tanygnathus; characters; Europe).

  • — Ganglbauer, 1895: 380 (Tanygnathus; characters; France; Sweden; Germany; Austria; Russia).

  • — Fauvel, 1897d: 311 (Tanygnathus; Morocco).

  • — Fauvel, 1902b: 119 (Tanygnathus; Morocco).

  • — Fauvel, 1903: 160 (Tanygnathus; India; Ceylon; Celebes; Australia; Madagascar; Natal; Mashunaland; Gold Coast; Gabon; Congo; Cape Verde; Morocco; Europe).

  • — Fauvel, 1904a: 60 (Tanygnathus; Khandesh).

  • — Fauvel, 1905e: 84 (Tanygnathus; Java).

  • — Sainte-Claire Deville, 1906: 104 (Tanygnathus; Corsica).

  • — Porta, 1907: 98 (Tanygnathus; characters; Italy).

  • — Reitter, 1909: 105 (Tanygnathinus; characters; Germany).

  • — Eichelbaum, 1913: 129 (Tanygnathus; Tanzania).

  • — Johansen, 1914: 342 (Tanygnathus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; catalog).

  • — Cameron, 1921a: 379, 406 (Atanygnathus; characters; Singapore).

  • — Lea, 1925: 253 (Atanygnathus; characters; Australia).

  • — Portevin, 1929: 334 (Tanygnathus; characters; France).

  • — Cameron, 1932: 309 (Atanygnathus; characters; India).

  • — Scheerpeltz, 1933: 1468 (Atanygnathus; catalog).

  • — Cameron, 1933d: 393 (Atanygnathus; Sumatra).

  • — Cameron, 1937: 30 (Atanygnathus; Java).

  • — V. Hansen, 1952: 163 (Atanygnathus; characters).

  • — Smetana, 1958: 402 (Atanygnathus; characters; ecology; Czechoslovakia).

  • — Tottenham, 1961: 205 (Atanygnathus; Senegal).

  • — Palm, 1963b: 159 (Atanygnathuss; characters; Sweden).

  • — Lohse, 1964: 220 (Atanygnathus; characters; central Europe).

  • — Scheerpeltz, 1965: 112 (Atanygnathus; Burma).

  • — Horion, 1965: 313 (Atanygnathus; Finland; Sweden; Ukraine; Poland; Czech Republic; Romania; Italy; Austria; Germany; France).

  • — Shibata, 1974: 30 (Atanygnathus; Japan).

  • — Scheerpeltz, 1974c: 25 (Atanygnathus; Sudan).

  • — Watanabe and Shibata, 1976: 333 (Atanygnathus; Malaysia).

  • — Coiffait, 1978: 321 (Atanygnathus; characters; Europe; Scandinavia).

  • — Tóth, 1984: 142 (Atanygnathus; characters; Hungary).

  • — Lucht, 1987: 111 (Atanygnathus; checklist; Germany; Poland; Czechoslovakia; Austria; Sweden).

  • — J. Li, 1993: 41 (Atanygnathus; China).

  • — Smetana, 1993b: 50 (Atanygnathus; Czech Republic).

  • — Uhlig, 1995a: 97 (Atanygnathus; lectotype designation; characters; Germany; Czechoslovakia; Italy).

  • — Ciceroni and Zanetti, 1995: 33 (Atanygnathus; Italy).

  • Distribution: Europe, Russia, Cape Verde, Morocco, Senegal, Ghana, Sudan, Congo, Tanzania, Zimbabwe, South Africa, Madagascar, Sri Lanka, India, Myanmar, Malaysia, Singapore, Indonesia, Australia, Japan.

borealis Mulsant and Rey, 1876: 841 [= 1877: 697 (Tanygnathus; Type locality: Prusse).

  • — Ganglbauer, 1895: 380 (Tanygnathus; synonym of terminalis).

  • — Porta, 1907: 98 (Tanygnathus; variety of terminalis).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; variety of terminalis).

  • — Scheerpeltz, 1933: 1468 (Atanygnathus; variety of terminalis).

  • — Coiffait, 1978: 321 (Atanygnathus; characters; variety of terminalis).

toxopei Cameron, 1952: 256 (Atanygnathus; Type locality: New Guinea: Snow Mountains: Sigi Camp, 1500 m).

  • Distribution: New Guinea.

varicornis Wollaston, 1867: 234 (Tanygnathus; Type locality: Cape Verde Islands: S. Antao; S. Iago).

  • — Fauvel, 1902b: 119 (Tanygnathus; synonym of terminalis).

  • — Fauvel, 1904a: 60 (Tanygnathus; synonym of terminalis).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; variety of terminalis).

  • — Bernhauer, 1942c: 364 (Atanygnathus; variety of terminalis; Cameroon).

  • — Coiffait, 1966b: 338 (Atanygnathus; subspecies of terminalis; Mauritania).

  • — Coiffait, 1978: 321 (Atanygnathus; valid species; characters; Italy; North Africa; Mauritania; Cape Verde Islands).

  • — Ciceroni and Zanetti, 1995: 33 (Atanygnathus; Italy).

  • Distribution: Italy, “North Africa”, Mauritania, Cape Verde Islands.

variegatus Lecoq, 1990: 211 (Atanygnathus; Type locality: La Réunion: bassindes Aigrettes près de Saint Gilles).

  • Distribution: Réunion.

vilis Sharp, 1884: 322 (Tanygnathus; Type locality: Guatemala, Guatemala city, San Geronimo, Rio Naranja 450 feet, Coatepeque 1300 feet; Nicaragua, Chontales; Panama, San Miguel one of the Pearl Islands).

  • — Bernhauer and Schubert, 1916: 440 (Atanygnathus; catalog).

  • Distribution: Guatemala, Nicaragua, Panama.

wauensis Last, 1987: 52 (Atanygnathus; Type locality: Papua New Guinea: New Guinea NE: Wau, Hospital Creek, 1250 m).

  • Distribution: New Guinea.

Subtribe Xanthopygina

Xanthopygina Sharp, 1884: 342 (genera included: Tympanophorus, Selma, Xanthopygus, Lampropygus, Brachydirus, Plociopterus, Philothalpus, Gastrisus, Isanopus, Styngetus, Ocyolinus, Phanolinus, Nausicotus, Glenus). Type genus: Xanthopygus (see Newton and Thayer, 1992: 66).

  • — Naomi, 1982 (characters; key to genera of Japan).

  • — Newton and Thayer, 1995: 66 (subtribe of Staphylinini).

  • — Smetana, 1995e: 48 (characters; includes Triacrina).

  • — Smetana and Davies, 2000: 12, 17 (characters).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 332, 395 (key to Nearctic genera; characters; notes).

Xanthopygi Sharp, 1884.

  • — Cameron, 1932: 54, 251 (characters; key to genera of British India).

  • — Scheerpeltz, 1933: 1410 (world catalog supplement).

  • — Scheerpeltz, 1940: 47 (key to Palaearctic genera).

  • — Blackwelder, 1944: 139 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Coiffait and Saiz, 1968: 363 (characters; Chilean species).

  • — Coiffait, 1974: 353, 557 (characters; genera of western Palaearctic region).

Xanthopyginae Sharp, 1884.

  • — Moore, 1964a: 89 (characters; notes).

  • — Moore and Legner, 1974g: 560 (key to genera of America north of Mexico).

  • — Moore and Legner, 1974h: 472 (catalog of the taxonomy, biology, and ecology of developmental stages of species of America north of Mexico).

  • — Moore and Legner, 1979: 32 (characters; tabular key to genera of North America north of Mexico).

  • — Lawrence and Newton, 1995: 830 (included in Staphylininae).

  • — Downie and Arnett, 1996: 387 (characters; key to genera of Northeastern North America).

Xanthopygini Sharp, 1884.

  • — Eichelbaum, 1909: 181 (list of genera with references; generic synonyms; number of genera; number of species in each genus; general distribution of genera).

  • — Scheerpeltz, 1965: 277 (key to genera of Oriental region).

  • — Scheerpeltz, 1968a: 86 (catalog of Austrian species).

  • — Muona, 1979: 15 (list of Scandinavian species).

  • — Outerelo and Gamarra, 1985: 67 (genera of Iberian Peninsula).

  • — Hayashi, 1997a: 476, 490 (characters; historical review).

Triacrina Bernhauer, 1931.

  • — Newton and Thayer, 1992: 66 (subtribe of Staphylinini).

  • — Smetana, 1995e: 48 (included in Xanthopygina).

  • — Smetana and Davies, 2000: 12 (synonym of Xanthopygina).

Triacri Bernhauer, 1931a: 84 (subtribe; genera included: Hasumius, Polyphematiana, Triacrus). Type genus: Triacus (see Newton and Thayer, 1992: 66).

  • — Scheerpeltz, 1933: 1418 (world catalog supplement).

Algon [23 species; Palaearctic, Ethiopian, and Oriental Regions]

Algon Sharp, 1874: 22 (species included: grandicollis). Type species: Algon grandicollis Sharp, fixed by monotypy.

  • — Duvivier, 1883: 141 (catalog).

  • — Bernhauer and Schubert, 1916: 415 (world catalog; 6 species).

  • — Boháč, 1993a: 447 (key to some Oriental Region species).

  • — Winkler, 1925: 388 (catalog for Palaearctic region).

  • — Cameron, 1925: 71 (catalog of Indian species).

  • — Cameron, 1932: 269 (characters; key to species of British India).

  • — Scheerpeltz, 1933: 1425 (world catalog supplement).

  • — Blackwelder, 1952: 45 (type species: grandicollis).

  • — Scheerpeltz, 1962b: 264 (characters).

  • — Tikhomirova, 1973a: 188 (checklist of species of USSR).

  • — Scheerpeltz, 1974b: 140 (key to species).

  • — Naomi, 1982: 128 (characters).

  • — Smetana, 1977b: 180 (member of Staphylinini, subtribe Xanthopygi).

  • — Coiffait, 1982a: 275 (key to species of Himalayan region).

  • — Naomi, 1983a: 589 (characters; species of Japan).

  • — Hammond, 1984: 205 (checklist of species of Borneo).

  • — Shibata, 1984: 114 (checklist of species of Japan).

  • — Shibata, 1986a: 125 (supplemental list of species from Taiwan).

  • — Dettner, 1987: 30 (defensive glands).

  • — Kanehisa, Tsumuki, and Kawazu, 1994 (chemical secretions).

  • — Schillhammer, 1999b: 95 (genus probably does not occur in Ethiopian region).

Creophilopsis Cameron, 1921: 272 (species included: semiaenea). Type species: Creophilopsis semiaenea Cameron, fixed by monotypy.

  • — Cameron, 1925: 71 (catalog of Indian species).

  • — Cameron, 1932: 269 (synonym of Algon).

  • — Blackwelder, 1952: 111 (synonym of Algon; type species: semiaenea).

Brachycamonthus Bernhauer, 1933: 37 (species included: kaiserianus). Type species: Brachycamonthus kaiserianus Bernhauer, fixed by monotypy.

  • — Blackwelder, 1952: 84 (type species: kaiserianus).

  • — Scheerpeltz, 1962b: 267 (characters).

  • — Korge, 1971a: 35 (characters).

  • — Smetana, 1977b: 180 (member of Staphylinini, subtribe Xanthopygi).

  • — Schillhammer, 1999b: 94 (synonym of Algon).

Allopygus Cameron, 1950d: 21 (species included: malayanus). Type species: Allopygus malayanus Cameron, fixed by original designation and monotypy.

  • — Schillhammer, 1999b: 94 (synonym of Algon).

africanus Bernhauer, 1915k: 308 (Algon; Type locality: Kamerun).

  • — Bernhauer and Schubert, 1916: 415 (Algon; catalog).

  • — Scheerpeltz, 1974b: 144 (Algon; characters; Cameroon).

  • — Schillhammer, 1999b: 95 (Algon; may not belong in Algon).

  • Distribution: Cameroon.

bhutanicus Coiffait, 1982, see: semiaureus Fauvel, 1895.

brevipennis Cameron, 1932: 274 (Algon; Type locality: Ceylon: Bogawantalawa).

  • — Scheerpeltz, 1933: 1425 (Algon; catalog).

  • — Scheerpeltz, 1974b: 142 (Algon; characters; Ceylon).

  • Distribution: Sri Lanka.

brevipennis Cameron, 1933, see: dysanelloides Schillhammer, 1999.

cyanipennis Coiffait, 1982, see: Indoquedius.

deuvei Coiffait, 1984e: 380 (Pammegus, cited as Psammegus; Type locality: Nepal oriental: vallée de Kathmandou, Gokarna).

  • — Schillhammer, 1999b: 95 (Algon; cited as deuvi; probably identical with Algon oculatus).

  • Distribution: Nepal.

dysanelloides Schillhammer, 1999b: 94 (Algon; replacement name for brevipennis Cameron, 1933).

  • Distribution: Borneo.

brevipennis Cameron, 1933b: 347 (Dysanellus; [preoccupied nec brevipennis Cameron, 1932]; Type locality: British North Borneo: Pakka, 10,200 feet).

  • — Hammond, 1984: 194 (Dysanellus; may belong in another genus; Borneo).

  • — Schillhammer, 1999b: 94 (Algon; synonym of dysanelloides; characters).

elegans Bernhauer, 1915f: 145 (Dysanellus, cited as Disanellus; Type locality: Sarawak: Limbang).

  • — Scheerpeltz, 1933: 1412 (Dysanellus; catalog).

  • — Last, 1964: 177 (Palaestrinus; [Note: Last cited Bernhauer as the author of elegans but there is no elegans Bernhauer in Palaestrinus. It is neither clear that Last intended to transfer elegans Bernhauer to Palaestrinus nor which of the five uses of elegans by Bernhauer was meant. Bernhauer used elegans in Dysanellus (it is now in Algon), Scelotrichus, Xanthopygus, Agerodes and Neolosus. Neolosus elegans can be eliminated because it is in the Osoriinae. Agerodes elegans and Xanthopygus elegans are excluded because both are South American species. The species listed in Algon and Scelotrichus remain possibilities; according to Hammond, 1984: 205 the record may apply to Algon elegans]; Sarawak).

  • — Hammond, 1984: 194, 195 (Algon; may belong in another genus; Borneo).

  • Distribution: Borneo.

excellens Cameron, 1944: 13 (Algon; Type locality: Assam: Shillong, altitude 6000 feet).

  • Distribution: India.

grandicollis Sharp, 1874: 23 (Algon; Type locality: Copper Temple, Nagasaki; Maiyasama, Hiogo).

  • — Sharp, 1889: 28 (Algon; Japan).

  • — Bernhauer and Schubert, 1916: 415 (Algon; catalog).

  • — Cameron, 1949b: 469 (Algon; Japan).

  • — Nakane, 1963a: 94 (Algon; characters; habitus photograph; Japan).

  • — Scheerpeltz, 1974b: 141 (Algon; characters; Japan; China).

  • — Watanabe and Shibata, 1972: 65 (Algon; Yaku-shima Island).

  • — Naomi, 1983a: 589 (Algon; characters; Japan).

  • — Yuh, Paik, Kwon, and Lee, 1985: 248 (Algon; Korea).

  • — Dettner, 1987: 30 (defensive glands).

  • — J. Li, 1993: 41 (Algon; cited as gradicolli; China).

  • — Kanehisa, Tsumuki, and Kawazu, 1994 (Algon; chemical secretions).

  • — Watanabe, 1998: 314 (Algon; cited as gradicollis; Japan).

  • Distribution: Japan, Korea, China.

immsi Bernhauer, 1915b: 54 (Algon; Type locality: Zentral-Himalaya: Binsar, Kumaon, 7700 Fuss).

  • — Bernhauer and Schubert, 1916: 415 (Algon; catalog).

  • — Cameron, 1932: 272 (Algon; characters; India).

  • — Scheerpeltz, 1933: 1425 (Algon; catalog).

  • — Scheerpeltz, 1974b: 142 (Algon; characters; India; Nepal).

  • — Scheerpeltz, 1976: 43 (Algon; Nepal).

  • — Scheerpeltz, 1976a: 92 (Algon; Nepal).

  • — Coiffait, 1982a: 276 (Algon; characters).

  • — Boháč, 1993a: 447 (Algon; characters in key).

  • Distribution: India, Nepal.

kaiserianus Bernhauer, 1933: 38 (Brachycamonthus; Type locality: Westchina: Szetschwan: Kinfushan).

  • — Schillhammer, 1999b: 94 (Algon; characters; notes).

  • Distribution: China.

matsukii Shibata, 1979: 23 (Algon; Type locality: Kenting Park, Pingtung Hsien).

  • Distribution: Taiwan.

malayanus Cameron, 1950d: 22 (Allopygus; Type locality: Pahang: Frazer's Hill, 4000 feet).

  • — Schillhammer, 1999b: 94 (Algon; notes).

  • Distribution: Malaysia.

nepalicus Coiffait, 1982a: 274 (Algon; Type locality: Nepal: Manigow, 1200–1900 m).

  • Distribution: Nepal.

oculatus Cameron, 1932: 273 (Algon; Type locality: Darjeeling Dist.: Mangpu).

  • — Scheerpeltz, 1933: 1425 (Algon; catalog).

  • — Scheerpeltz, 1974b: 141 (Algon; characters; India).

  • — Coiffait, 1982a: 234 (Algon; Nepal).

  • — Coiffait, 1982a: 275 (Algon; characters).

  • — Schillhammer, 1999b: 95 (Algon; may not belong in Algon).

  • Distribution: India, Nepal.

pergrandis Scheerpeltz, 1974b: 144 (Algon; Type locality: Nordvietnam im Gebirgszuge des Tamdoo, unmittelbar nördlich von Hanoi).

  • — Boháč, 1993a: 447 (Algon; characters in key).

  • Distribution: Vietnam.

psittacus Fauvel, 1895: 273 (Algon; Type locality: Inde).

  • — Bernhauer and Schubert, 1916: 415 (Algon; catalog).

  • — Cameron, 1932: 271 (Algon; characters; India).

  • — Scheerpeltz, 1933: 1426 (Algon; catalog).

  • — Scheerpeltz, 1974b: 143 (Algon; characters).

  • — Coiffait, 1982a: 276 (Algon; characters).

  • — Boháč, 1993a: 447 (Algon; characters in key).

  • Distribution: India.

robustus Wendeler, 1928a: 35 (Algon; Type locality: Süd-Afrika: Kapland).

  • — Scheerpeltz, 1933: 1426 (Algon; catalog).

  • — Scheerpeltz, 1974b: 144 (Algon; characters; South Africa).

  • Distribution: South Africa.

semiaeneus Cameron, 1921: 273 (Creophilopsis; Type locality: Hakachin Hills, Burma).

  • — Cameron, 1932: 271 (Algon; synonym of semiaureus).

  • — Scheerpeltz, 1933: 1426 (Algon; synonym of semiaureus).

  • — Smetana, 1977a: 243 (Algon; synonym of semiaureus).

  • — Schillhammer, 1999b: 94 (Algon; valid species; notes).

  • Distribution: Myanmar.

semiaereus Cameron, 1928, see: Rientis.

semiaureus Fauvel, 1895: 273 (Algon; Type locality: Bengale, Kurseong, Kombir; Sikkim, Darjeeling).

  • — Bernhauer and Schubert, 1916: 415 (Algon; catalog).

  • — Cameron, 1932: 271 (Algon; characters; Burma; Bengal).

  • — Scheerpeltz, 1933: 1426 (Algon; catalog).

  • — Scheerpeltz, 1974b: 143 (Algon; characters; India).

  • — Smetana, 1977a: 243 (Algon; Bhutan).

  • — Coiffait, 1982a: 234, 276 (Algon; characters; India).

  • Distribution: India, Bhutan, Myanmar.

bhutanicus Coiffait, 1982a: 270 (Algon; Type locality: Bhoutan: Ungar Lhuntsi, 1800–2500 m).

  • — Schillhammer, 1999b: 94 (Algon; synonym of semiaureus).

semicaeruleus Cameron, 1942a: 137 (Algon; Type locality: N. Borneo: Kinabalu).

  • — Hammond, 1984: 194, 195 (Algon; synonym of elegans).

  • — Schillhammer, 1999b: 94 (Algon; valid species; notes).

  • Distribution: Borneo.

spatiosus† Zhang, 1989: 115 (Algon; Miocene fossil; Type locality: China: Shanwang, Shandong).

  • Distribution: China.

tricolor Fauvel, 1895: 272 (Algon; Type locality: Bírmanie, Teinzo).

  • — Bernhauer and Schubert, 1916: 415 (Algon; catalog).

  • — Cameron, 1932: 270 (Algon; characters; Burma).

  • — Scheerpeltz, 1933: 1426 (Algon; catalog).

  • — Scheerpeltz, 1974b: 143 (Algon; characters; Himalaya; Burma).

  • — Coiffait, 1982a: 276 (Algon; characters).

  • — Boháč, 1993a: 447 (Algon; characters in key).

  • Distribution: Myanmar, “Himalayan region”.

uniformis Cameron, 1932: 273 (Algon; Type locality: Darjeeling Dist.: Ghum, Tiger Hill, alt. 8000 feet).

  • — Scheerpeltz, 1933: 1426 (Algon; catalog).

  • — Scheerpeltz, 1974b: 141 (Algon; characters; India).

  • — Coiffait, 1982a: 234 (Algon; India).

  • — Coiffait, 1982a: 275 (Algon; characters).

  • — Coiffait, 1984e: 374 (Algon; Nepal).

  • Distribution: India, Nepal.

viridis Boháč, 1993a: 445 (Algon; Type locality: North Vietnam: 65 km north Ankche Tan Nguen).

  • Distribution: Vietnam.

Allostenopsis [2 species; Neotropical Region]

Allostenopsis Bernhauer, 1921b: 74 (replacement name for Stenopsis Bernhauer). Type species: Stenopsis antennaria Bernhauer, fixed by objective synonymy with Stenopsis.

  • — Scheerpeltz, 1933: 1414 (world catalog supplement).

  • — Blackwelder, 1952: 46 (type species).

Stenopsis Bernhauer, 1907a: 286 (species included: antennaria, kraatzi; preoccupied by Rafinesque, 1815, and Cassini, 1851; [Note: The Zoological Record (1966: 281) erroneously listed Stenopsis pardoi Español (1966, Graellsia, 22: 174) in the Staphylinidae; it is in the Tenebrionidae]). Type species: Stenopsis antennaria Bernhauer, fixed by subsequent designation by R. Lucas, 1920: 610.

  • — Bernhauer and Schubert, 1914: 401 (world catalog; 2 species).

  • — Bernhauer, 1921b: 74 (synonym of Allostenopsis).

  • — Scheerpeltz, 1933: 1414 (synonym of Allostenopsis).

  • — Blackwelder, 1944: 140 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 358 (synonym of Allostenopsis; type species).

antennaria Bernhauer, 1907a: 286 (Stenopsis; Type locality: Brasilia: St. Catharina).

  • — Bernhauer and Schubert, 1914: 401 (Stenopsis; catalog).

  • Distribution: Brazil.

kraatzi Bernhauer, 1907a: 287 (Stenopsis; Type locality: Paraguay).

  • — Bernhauer and Schubert, 1914: 401 (Stenopsis; catalog).

  • Distribution: Paraguay.

Amelinus [19 species; Ethiopian and Australian Regions]

Amelinus Bernhauer, 1915k: 306 (species included: gestroi, praenobilis). Type species: Amelinus gestroi Bernhauer, fixed by subsequent designation by Blackwelder, 1952: 49.

  • — Scheerpeltz, 1933: 1410 (world catalog supplement).

allardi Levasseur, 1968: 1398 (Amelinus; Type locality: Katanga: région de Kolwezi, Ruwe).

  • Distribution: Congo.

alticola Levasseur, 1968: 1393 (Amelinus; Type locality: mont Cameroun).

  • Distribution: Cameroon.

camerounensis Levasseur, 1968: 1395 (Amelinus; Type locality: Cameroun: mont Fébé environs de Yaoundé).

  • Distribution: Cameroon.

collarti Cameron, 1934e: 373 (Amelinus; Type locality: Nizi: Ozeguru).

  • Distribution: Congo.

densior Bernhauer, 1928g: 106 (Amelinus; Type locality: Ostafrika: Unt. Umba).

  • — Scheerpeltz, 1933: 1411 (Amelinus; catalog).

  • — Tottenham, 1962: 144 (Amelinus; Uganda).

  • Distribution: Uganda, Tanzania.

densipennis Bernhauer, 1928g: 107 (Amelinus; Type locality: Belg.-Kongo: Haut Uele, Moto; Boga).

  • — Scheerpeltz, 1933: 1411 (Amelinus; catalog).

  • Distribution: Congo.

dollmani Bernhauer, 1928g: 105 (Amelinus; Type locality: N.W. Rhodesia: Kashitu, N. of Broken Hill).

  • — Scheerpeltz, 1933: 1411 (Amelinus; catalog).

  • Distribution: Zambia.

gestroi Bernhauer, 1915k: 307 (Amelinus; Type locality: Portugiesisch-Guinea: Bolama).

  • — Bernhauer, 1928g: 105 (Amelinus; characters in key).

  • — Scheerpeltz, 1933: 1411 (Amelinus; catalog).

  • — Levasseur, 1968a: 54 (Amelinus; Central African Republic).

  • Distribution: Guinea-Bissau, Central African Republic.

kolweziensis Levasseur, 1968: 1400 (Amelinus; Type locality: Katanga: région de Kolwezi, Ruwe).

  • Distribution: Congo.

koundenensis Levasseur, 1968: 1393 (Amelinus; Type locality: Cameroun: Kounden).

  • Distribution: Cameroon.

kraatzi Bernhauer, 1915k: 305 (Pammegus; Type locality: Kamerun).

  • — Scheerpeltz, 1933: 1411 (Pammegus; catalog).

  • — Schillhammer, 1999b: 95 (Amelinus).

  • Distribution: Cameroon.

lomaensis Levasseur, 1971: 302 (Amelinus; Type locality: Monts Loma, Sierra Leone, 1600 m).

  • Distribution: Sierra Leone.

manikaensis Levasseur, 1968: 1397 (Amelinus; Type locality: Katanga: région de Kolwezi, plateau de Manika).

  • Distribution: Congo.

methneri Bernhauer, 1928g: 105 (Amelinus; Type locality: Ostafrika: Unteres Kihansital).

  • — Scheerpeltz, 1933: 1411 (Amelinus; catalog).

  • Distribution: “East Africa”.

musonoiensis Levasseur, 1968: 1396 (Amelinus; Type locality: Katanga: région de Kolwezi, Musonoie).

  • Distribution: Congo.

praenobilis Bernhauer, 1915k: 308 (Amelinus; Type locality: Portugiesisch-Guinea: Bolama).

  • — Bernhauer, 1928g: 106 (Amelinus; characters in key).

  • — Scheerpeltz, 1933: 1411 (Amelinus; catalog).

  • Distribution: Guinea-Bissau.

punctus Last, 1989: 882 (Amelinus; Type locality: Papua New Guinea: Kainantu, Onerunka).

  • Distribution: New Guinea.

semiruber Tottenham, 1962: 257 (Amelinus; Type locality: Uganda: Bugiri, 1400 m).

  • Distribution: Uganda.

zenckei Bernhauer, 1928g: 106 (Amelinus; Type locality: Tanganyika-Küste: Bez. Bismarck-burg).

  • — Scheerpeltz, 1933: 1411 (Amelinus; catalog).

  • Distribution: Tanzania.

Antimerus [4 species; Australian Region]

Antimerus Fauvel, 1878g: 550 (species included: smaragdinus). Type species: Antimerus smaragdinus Fauvel, fixed by monotypy.

  • — Duvivier, 1883: 141 (catalog).

  • — Bernhauer and Schubert, 1916: 410 (world catalog; 2 species).

  • — Scheerpeltz, 1933: 1420 (world catalog supplement).

  • — Blackwelder, 1952: 57 (type species: smaragdinus).

auricomus Lea, 1925: 229 (Antimerus; Type locality: New South Wales: Dorrigo).

  • — Scheerpeltz, 1933: 1420 (Antimerus; catalog).

  • Distribution: Australia.

posttibialis Lea, 1925: 229 (Antimerus; Type locality: Queensland: Kuranda).

  • — Scheerpeltz, 1933: 1420 (Antimerus; catalog).

  • Distribution: Australia.

punctipennis Lea, 1906: 195 (Antimerus; Type locality: Gosford, N.S.W.).

  • — Bernhauer and Schubert, 1916: 410 (Antimerus; catalog).

  • Distribution: Australia.

smaragdinus Fauvel, 1878g: 550 (Antimerus; Type locality: Australie méridionale: montagnes de Victoria).

  • — Bernhauer and Schubert, 1916: 410 (Antimerus; catalog).

  • Distribution: Australia.

Dysanellus [3 species; Neotropical and Oriental Regions]

Dysanellus Bernhauer, 1911a: 419 (species included: bruchi). Type species: Dysanellus bruchi Bernhauer, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 397 (world catalog; 1 species).

  • — Scheerpeltz, 1933: 1412 (world catalog supplement).

  • — Blackwelder, 1944: 140 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 137 (type species: bruchi).

  • — Scheerpeltz, 1965: 277 (characters; type species: bruchi).

Leptodiastemus Bernhauer, 1934b: 215 (subgenus of Dysanellus; species included: excellens). Type species: Dysanellus (Leptodiastemus) excellens Bernhauer, fixed by monotypy.

  • — Blackwelder, 1952: 216 (subgenus of Dysanellus; type species: excellens).

brevipennis Cameron, 1933, see: Algon.

bruchi Bernhauer, 1911a: 419 (Dysanellus; Type locality: Argentinien: Rep. Buenos Aires).

  • — Bernhauer and Schubert, 1914: 397 (Dysanellus; catalog).

  • Distribution: Argentina.

elegans Bernhauer, 1915, see: Algon.

excellens Bernhauer, 1934b: 215 (Dysanellus; subgenus Leptodiastemus; Type locality: Argentinien: Tucuman).

  • Distribution: Argentina.

transverserugosus Bernhauer, 1921d: 175 (Dysanellus; Type locality: Brasilien: Ypiranga).

  • — Scheerpeltz, 1933: 1412 (Dysanellus; catalog).

  • Distribution: Brazil.

Elecatopselaphus [2 species; Neotropical Region]

Elecatopselaphus Scheerpeltz, 1972a: 39 (species included: violaceus, peruvianus). Type species: Trigonopselaphus violaceus Sharp, fixed by original designation.

peruvianus Scheerpeltz, 1972a: 46 (Elecatopselaphus; Type locality: Peru: Umgebung von Salinas bei Lima).

  • Distribution: Peru.

violaceus Sharp, 1876: 145 (Trigonopselaphus; Type locality: Ega).

  • — Sharp, 1884: 364 (Phanolinus).

  • — Scheerpeltz, 1972a: 39 (Elecatopselaphus).

  • Distribution: Brazil.

Elmas [2 species; Neotropical Region]

Elmas Blackwelder, 1952: 146 (replacement name for Selma Sharp). Type species: Selma modesta Sharp, fixed by objective synonymy with Selma.

Selma Sharp, 1876b: 426 (species included: modesta). Type species: Selma modesta Sharp, fixed by monotypy.

  • — Duvivier, 1883: 155 (catalog).

  • — Sharp, 1884: 343 (note).

  • — Bernhauer and Schubert, 1914: 396 (world catalog; 1 species).

  • — Scheerpeltz, 1933: 1411 (world catalog supplement).

  • — Blackwelder, 1944: 140 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 350 (synonym of Elmas; type species: modesta).

modesta Sharp, 1876b: 427 (Selma; Type locality: Chontales).

  • — Sharp, 1884: 343 (Selma; Nicaragua).

  • — Bernhauer and Schubert, 1914: 396 (Selma; catalog).

  • Distribution: Nicaragua.

strigella Bernhauer, 1915l: 296 (Selma; Type locality: Brasilien).

  • — Scheerpeltz, 1933: 1411 (Selma; catalog).

  • Distribution: Brazil.

Gastrisus [18 species; Nearctic and Neotropical Regions]

Gastrisus Sharp, 1876: 136 (species included: obsoletus, laevigatus, punctatus). Type species: Gastrisus laevigatus Sharp, fixed by original designation.

  • — Duvivier, 1883: 154 (catalog).

  • — Sharp, 1884: 359 (notes; Central American species).

  • — Bernhauer and Schubert, 1914: 405 (world catalog; 8 species).

  • — Scheerpeltz, 1933: 1415 (world catalog supplement).

  • — Blackwelder, 1944: 142 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 167 (type species: laevigatus).

brasiliensis R. Sahlberg, 1847: 787 (Staphylinus; Type locality: Rio Janeiro).

  • — Bernhauer and Schubert, 1914: 405 (Gastrisus; catalog).

  • Distribution: Brazil.

cribrum Fauvel, 1891: 107 (Gastrisus; Type locality: San Esteban).

  • — Bernhauer and Schubert, 1914: 405 (Gastrisus; catalog).

  • Distribution: Venezuela.

herilis Erichson, 1840: 497 (Philonthus; Type locality: Brasilia).

  • — Kraatz, 1857d: 539, 540 (Xanthopygus).

  • — Steinheil, 1869: 253 (Xanthopygus; Argentina).

  • — Lynch, 1884: 131 (Xanthopygus; characters; Argentina).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1933: 1416 (Gastrisus; catalog).

  • Distribution: Brazil, Argentina.

klimai Bernhauer, 1911a: 417 (Trigonopselaphus; Type locality: Brasilien, San Leopoldo).

  • — Bernhauer and Schubert, 1914: 403 (Trigonopselaphus; catalog).

  • — Scheerpeltz, 1933: 1417 (Gastrisus; catalog).

  • Distribution: Brazil.

laevigatus Sharp, 1876: 137 (Gastrisus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 405 (Gastrisus; catalog).

  • Distribution: Brazil.

mimetes Sharp, 1884: 360 (Gastrisus; Type locality: Costa Rica).

  • — Bernhauer and Schubert, 1914: 405 (Gastrisus; catalog).

  • Distribution: Costa Rica.

modestus Bernhauer, 1912d: 39 (Trigonopselaphus; Type locality: Uruguay).

  • — Scheerpeltz, 1933: 1415 (Phanolinus; catalog).

  • — Scheerpeltz, 1972a: 37 (Gastrisus).

  • Distribution: Uruguay.

newtonorum Navarrete and Márquez, 1998: 226 (Gastrisus; Type locality: Mexico: Morelos, Yautepec, Carretera México-Cuautla, km. 19).

  • — Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 62 (Gastrisus; necrophilous species; distribution in Mexico).

  • Distribution: Mexico.

nitidus Bernhauer, 1906b: 198 (Gastrisus; Type locality: Brasilia, São Paulo).

  • — Bernhauer and Schubert, 1914: 405 (Gastrisus; catalog).

  • Distribution: Brazil.

nobilis Wendeler, 1956a: 234 (Trigonopsephalus, cited as Trigonopcephalus; Type locality: Brasilien: Matto Grosso).

  • — Scheerpeltz, 1972a: 37 (Gastrisus).

  • Distribution: Brazil.

obsoletus Sharp, 1876: 136 (Gastrisus; Type locality: Not cited).

  • — Bernhauer and Schubert, 1914: 405 (Gastrisus; catalog).

  • Distribution: Brazil.

opaculus Sharp, 1884: 359 (Gastrisus; Type locality: Panama, Volcan de Chiriqui).

  • — Bernhauer and Schubert, 1914: 405 (Gastrisus; catalog).

  • Distribution: Panama.

opulentus Erichson, 1840: 497 (Philonthus; Type locality: Brasilia).

  • — Bernhauer and Schubert, 1914: 349 (Philonthus; catalog).

  • — Scheerpeltz, 1933: 1417 (Gastrisus; catalog).

  • — Herman, 2001: 25 (Gastrisus; synonymy).

  • Distribution: Brazil.

opulentus Bernhauer, 1911a: 417 (Trigonopselaphus; [preoccupied]; [Note: Scheerpeltz, 1933: 1415 transferred this species to Gastrisus but on p. 1417 he cited Bernhauer's use of opulentus as a subsequent reference to Erichson's opulentus; Bernhauer cited opulentus as a new species]; Type locality: Brasilien).

  • — Bernhauer and Schubert, 1914: 403 (Trigonopselaphus; catalog).

  • — Scheerpeltz, 1833: 1415, 1417 (Gastrisus; Bernhauer's use of opulentus cited as subsequent reference of opulentus Erichson).

  • — Herman, 2001: 25 (Gastrisus; synonym of opulentus Erichson).

opulentus Bernhauer, 1911, see: opulentus Erichson, 1840.

punctatus Sharp, 1876: 138 (Gastrisus; Type locality: St. Paulo).

  • — Bernhauer and Schubert, 1914: 406 (Gastrisus; catalog).

  • Distribution: Brazil.

purpuripennis Bernhauer, 1927d: 168 (Trigonopselaphus; Type locality: Brasilien: Rio Jose Pedros, Minas; Passo Quatro, 915 m; S. Minas Geraes).

  • — Scheerpeltz, 1933: 1415 (Phanolinus; catalog).

  • — Scheerpeltz, 1972a: 37 (Gastrisus).

  • Distribution: Brazil.

putidiusculus Tottenham, 1939: 172 (Trigonopselaphus; Type locality: Peru: Junin, Sani Beni).

  • — Scheerpeltz, 1972a: 37 (Gastrisus).

  • Distribution: Peru.

venezolanus Bernhauer, 1912d: 39 (Gastrisus; Type locality: Venezuela).

  • — Scheerpeltz, 1933: 1417 (Gastrisus; catalog).

  • Distribution: Venezuela.

viridicollis Bernhauer, 1908d: 326 (Trigonophorus; Type locality: Brasilien: Goyaz).

  • — Bernhauer and Schubert, 1914: 403 (Trigonopselaphus; catalog).

  • — Scheerpeltz, 1972a: 37 (Gastrisus).

  • Distribution: Brazil.

Glenus [21 species; Neotropical Region]

Glenus Kraatz, 1857d: 541 (species included: chrysis, regalis, biplagiatus, cyanicollis). Type species: Staphylinus biplagiatus Perty, fixed by subsequent designation by Blackwelder, 1952: 170.

  • — Gemminger and Harold, 1868: 574 (catalog).

  • — Sharp, 1876: 147 (notes; characters).

  • — Duvivier, 1883: 141 (catalog).

  • — Sharp, 1884: 369 (notes; Mexican and Central American species).

  • — Bernhauer and Schubert, 1914: 400 (world catalog; 13 species).

  • — Scheerpeltz, 1933: 1414 (world catalog supplement).

  • — Blackwelder, 1944: 140 (checklist of species from Mexico, Central and South America, and the West Indies).

amazonicus Sharp, 1876: 148 (Glenus; Type locality: Tapajos).

  • — Bernhauer and Schubert, 1914: 400 (Glenus; catalog).

  • Distribution: Brazil.

auricomus Brullé, 1842, see: chrysis Gravenhorst, 1806.

aureicollis Wasmann, 1925: 73 (Glenus; Type locality: Not cited).

  • — Scheerpeltz, 1933: 1414 (Glenus; catalog; Brazil).

  • Distribution: Brazil.

batesi Sharp, 1876: 148 (Glenus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 400 (Glenus; catalog).

  • Distribution: Brazil.

biplagiatus Perty, 1830: 31 (Staphylinus; Type locality: Brasil: Prov. Minarum).

  • — Blanchard, 1842: 79 (Staphylinus; Bolivia).

  • — Erichson, 1839b: 370 (Staphylinus; characters; Brazil).

  • — P. Lucas, 1857: 50 (Staphylinus; Brazil).

  • — Bernhauer and Schubert, 1914: 400 (Glenus; catalog).

  • Distribution: Brazil, Bolivia.

interruptus Brullé, 1842: pl. 5, fig. 7 (Staphylinus; Type locality: Not cited).

  • — Blanchard, 1842: 79 (Staphylinus; synonym of biplagiatus).

  • — Erichson, 1839b: 370 (Staphylinus; synonym of cyanicollis).

  • — Gemminger and Harold, 1868: 574 (Glenus; synonym of biplagiatus).

  • — Bernhauer and Schubert, 1914: 400 (Glenus; synonym of biplagiatus).

brasilianus Bernhauer, 1927d: 166 (Glenus; Type locality: Brasilien: Passo Quatro, 1400 m; Itatiaya, Est. do Rio, 700 m).

  • — Scheerpeltz, 1933: 1414 (Glenus; catalog).

  • Distribution: Brazil.

bucki Wasmann, 1925a: 931 (Glenus; Type locality: Not cited).

  • — Scheerpeltz, 1933: 1414 (Glenus; catalog).

  • Distribution: Brazil.

chrysis Gravenhorst, 1806: 124 (Staphylinus; Type locality: Bahia, Brasiliae provincia).

  • — Nordmann, 1837: 31 (Staphylinus; Brazil).

  • — Blanchard, 1842: 79 (Staphylinus; Bolivia; Brazil).

  • — Erichson, 1839b: 368 (Staphylinus; characters; Bahia).

  • — P. Lucas, 1857: 49 (Staphylinus; Brazil).

  • — Gemminger and Harold, 1868: 574 (Glenus; catalog).

  • — Bernhauer and Schubert, 1914: 400 (Glenus; catalog).

  • — Scheerpeltz, 1933: 1414 (Glenus; catalog; Argentina).

  • Distribution: Brazil, Bolivia, Argentina.

auricomus Brullé, 1842: pl. 5, fig.6 (Staphylinus; Type locality: Not cited).

  • — Erichson, 1839b: 369 (Staphylinus; synonym of chrysis).

  • — Blanchard, 1842: 79 (Staphylinus; synonym of chrysis).

  • — Gemminger and Harold, 1868: 574 (Glenus; synonym of chrysis).

  • — Bernhauer and Schubert, 1914: 400 (Glenus; synonym of chrysis).

condei Wendeler, 1956a: 234 (Glenus; Type locality: Brasilien: Espirito Santo, Sta. Theresa; Sta. Catharina).

  • Distribution: Brazil.

coxalis Sharp, 1876b: 425 (Glenus; Type locality: Panama).

  • — Sharp, 1884: 369 (Glenus; Panama).

  • — Bernhauer and Schubert, 1914: 400 (Glenus; catalog).

  • Distribution: Panama.

cyanicollis Laporte, 1835: 115 (Staphylinus; Type locality: Brésil).

  • — Erichson, 1839b: 370 (Staphylinus; characters; Brazil).

  • — Bernhauer and Schubert, 1914: 400 (Glenus; catalog).

  • — Scheerpeltz, 1933: 1414 (Glenus; catalog; Argentina).

  • Distribution: Brazil, Argentina.

extimus Sharp, 1887: 784 (Glenus; Type locality: Mexico, Durango city).

  • — Bernhauer and Schubert, 1914: 400 (Glenus; catalog).

  • Distribution: Mexico.

flohri Sharp, 1884: 369 (Glenus; Type locality: Mexico, Jalapa; Guatemala, near the city).

  • — Bernhauer and Schubert, 1914: 400 (Glenus; catalog).

  • Distribution: Mexico, Guatemala.

interruptus Brullé, 1842, see: biplagiatus Perty, 1830.

jelskii Solsky, 1872: 291 (Glenus; Type locality: Monte-Rico, Pérou).

  • — Bernhauer and Schubert, 1914: 400 (Glenus; catalog).

  • — Scheerpeltz, 1936a: 491 (Glenus; natural history; Brazil).

  • — Scheerpeltz, 1960c: 120 (Glenus; Bolivia; Paraguay; Argentina; Brazil; Peru).

  • Distribution: Argentina, Paraguay, Bolivia, Peru, Brazil.

kraatzi Sharp, 1876: 147 (Glenus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 400 (Glenus; catalog).

  • Distribution: Brazil.

multipunctatus Wendeler, 1956a: 233 (Glenus; Type locality: Brasilien: São Paulo, Ribeirao Preto, Fazenda Dumont).

  • Distribution: Brazil.

nigricollis Wendeler, 1956a: 233 (Glenus; Type locality: Brasilien: Sta. Leopoldina).

  • Distribution: Brazil.

ohausi Wendeler, 1930: 192 (Glenus; Type locality: Brasilien, São Paulo, Ypiranga, Kamp one Busch).

  • — Scheerpeltz, 1933: 1414 (Glenus; catalog).

  • Distribution: Brazil.

princeps Bernhauer, 1914a: 51 (Glenus; Type locality: Brasilien: Rio Grande do Sul).

  • — Scheerpeltz, 1933: 1414 (Glenus; catalog).

  • Distribution: Brazil.

regalis Erichson, 1839b: 369 (Staphylinus; Type locality: Brasilia).

  • — Bernhauer and Schubert, 1914: 400 (Glenus; catalog).

  • Distribution: Brazil.

setosus Sharp, 1887: 784 (Glenus; Type locality: Mexico, Altisco in Puebla).

  • — Bernhauer and Schubert, 1914: 400 (Glenus; catalog).

  • — Marquez-Luna and Navarrete Heredia, 1995: 35 (Glenus; in detritus of Atta mexicana; Mexico).

  • Distribution: Mexico.

vestitus Sharp, 1876: 149 (Glenus; Type locality: Para; Ega; St. Paulo).

  • — Bernhauer and Schubert, 1914: 400 (Glenus; catalog).

  • Distribution: Brazil.

Hasumius [8 species; Ethiopian Region]

Hasumius Fairmaire, 1891: cclxxxii (species included: validus). Type species: Hasumius validus Fairmaire, fixed by monotypy.

  • — Bernhauer, 1909a: 87 (key to species).

  • — Bernhauer and Schubert, 1914: 397 (world catalog; 6 species).

  • — Bernhauer, 1931a: 84 (characters in key).

  • — Scheerpeltz, 1933: 1418 (world catalog supplement).

  • — Blackwelder, 1952: 181 (type species: validus).

ertli Bernhauer, 1909a: 88 (Hasumius; Type locality: Mangu in der Nähe des Kenia, Landschaft Kikuyu, Britisch-Ostafrika).

  • — Bernhauer, 1912h: 479 (Hasumius; Central Africa).

  • — Bernhauer and Schubert, 1914: 397 (Hasumius; catalog).

  • — Scheerpeltz, 1933: 1418 (Hasumius; catalog).

  • — Gridelli, 1939: 338 (Hasumius; Abyssinia).

  • Distribution: Ethiopia, Kenya, “Central Africa”.

ganglbaueri Bernhauer, 1909a: 88 (Hasumius; Type locality: Britisch-Ostafrika, Kigonsera, Urundi).

  • — Bernhauer and Schubert, 1914: 397 (Hasumius; catalog).

  • — Scheerpeltz, 1933: 1418 (Hasumius; catalog).

  • Distribution: Kenya.

iringanus Bernhauer, 1909a: 89 (Hasumius; Type locality: Deutsch-Ostafrika, Iringa).

  • — Bernhauer and Schubert, 1914: 397 (Hasumius; cited as irringanus; catalog).

  • — Scheerpeltz, 1933: 1418 (Hasumius; cited as irringanus; catalog).

  • Distribution: Tanzania.

lamorali Bernhauer, 1933d: 296 (Hasumius; Type locality: Elisabethville).

  • Distribution: Congo.

puncticollis Bernhauer, 1909a: 89 (Hasumius; Type locality: Deutsch-Ostafrika, Issansu).

  • — Bernhauer and Schubert, 1914: 397 (Hasumius; catalog).

  • — Scheerpeltz, 1933: 1418 (Hasumius; catalog).

  • Distribution: Tanzania.

suturalis Fairmaire, 1901: 279 (Hasumius; Type locality: Ubemba, Afrique centrale [Note: This may be Upemba region of the former Zaire]).

  • — Bernhauer, 1909a: 87 (Hasumius; characters in key).

  • — Bernhauer and Schubert, 1914: 397 (Hasumius; catalog).

  • — Scheerpeltz, 1933: 1419 (Hasumius; catalog).

  • Distribution: Congo ?.

validus Fairmaire, 1891: cclxxxii (Hasumius; Type locality: Not cited).

  • — Eppelsheim, 1895: 200 (Hasumius; characters; Pays des Cormoso).

  • — Bernhauer, 1909a: 87 (Hasumius; characters in key).

  • — Bernhauer and Schubert, 1914: 397 (Hasumius; catalog; Somaliland).

  • — Scheerpeltz, 1933: 1419 (Hasumius; catalog).

  • — Gridelli, 1939: 338 (Hasumius; Abyssinia).

  • Distribution: Ethiopia, “Somaliland”.

zernyi Bernhauer, 1947: 158 (Hasumius; [Note: The original spelling of the name was “zerny” but the species was named for Hans Zerny]; Type locality: Deutsch-Ostafrika: Ugano).

  • Distribution: Tanzania.

Hesperoschema [1 species; Oriental Region]

Hesperoschema Scheerpeltz, 1965: 261 (species included: malaisei). Type species: Hesperoschema malaisei Scheerpeltz, fixed by original designation and monotypy.

malaisei Scheerpeltz, 1965: 263 (Hesperoschema; Type locality: N.E. Burma, 7000 ft.).

  • Distribution: Myanmar.

Isanopus [3 species; Neotropical Region]

Isanopus Sharp, 1876: 141 (species included: tenuicornis). Type species: Isanopus tenuicornis Sharp, fixed by monotypy.

  • — Duvivier, 1883: 154 (catalog).

  • — Bernhauer and Schubert, 1914: 400 (world catalog; 2 species).

  • — Scheerpeltz, 1933: 1414 (world catalog supplement).

  • — Blackwelder, 1944: 140 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 200 (type species: tenuicornis).

sahlbergi Bernhauer, 1917i: 109 (Isanopus; Type locality: Brasilien: Petropolis, Boa Sorta).

  • — Scheerpeltz, 1933: 1414 (Isanopus; catalog).

  • Distribution: Brazil.

sallaei Sharp, 1884: 360 (Isanopus; Type locality: Mexico, Cordova).

  • — Bernhauer and Schubert, 1914: 400 (Isanopus; catalog).

  • Distribution: Mexico.

tenuicornis Sharp, 1876: 141 (Isanopus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 401 (Isanopus; catalog).

  • Distribution: Brazil.

Nausicotus [4 species; Neotropical Region]

Nausicotus Sharp, 1884: 368 (species included: opacipennis, spectabilis). Type species: Trigonopselaphus opacipennis Sharp, fixed by subsequent designation by R. Lucas, 1920: 431.

  • — Bernhauer and Schubert, 1914: 403 (world catalog; 4 species).

  • — Blackwelder, 1944: 141 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 257 (type species).

castaneipennis Lynch, 1884: 144 (Trigonopselaphus; Type locality: Chacabuco).

  • — Bernhauer and Schubert, 1914: 403 (Nausicotus; catalog).

  • Distribution: Argentina.

opacipennis Sharp, 1876: 143 (Trigonopselaphus; Type locality: St. Paulo).

  • — Bernhauer and Schubert, 1914: 403 (Nausicotus; catalog).

  • Distribution: Brazil.

spectabilis Sharp, 1884: 369 (Nausicotus; Type locality: Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1914: 403 (Nausicotus; catalog).

  • Distribution: Nicaragua.

staphylinoides Lynch, 1884: 141 (Trigonopselaphus; Type locality: Insula Antequera; Chacabuco).

  • — Bernhauer and Schubert, 1914: 403 (Nausicotus; catalog).

  • Distribution: Argentina.

Nordus [36 species; Neotropical Region]

Nordus Blackwelder, 1952: 262 (replacement name for Brachydirus Nordmann). Type species: Brachydirus xanthocerus Nordmann, fixed by objective synonymy with Brachydirus.

Brachydirus Nordmann, 1837: 131 (species included: xanthocerus; preoccupied by Smith Woodward, 1811). Type species: Brachydirus xanthocerus Nordmann, fixed by monotypy.

  • — Erichson, 1839b: 353 (synonym of Staphylinus).

  • — Lacordaire, 1854: 76 (synony of Staphylinus).

  • — Kraatz, 1857d: 538, note (characters).

  • — Gemminger and Harold, 1868: 598 (catalog).

  • — Sharp, 1876: 108 (notes).

  • — Duvivier, 1883: 155 (catalog).

  • — Sharp, 1884: 348 (notes; Central American species).

  • — Bernhauer and Schubert, 1914: 404 (world catalog; 24 species).

  • — Scheerpeltz, 1933: 1416 (world catalog supplement).

  • — Blackwelder, 1944: 141 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 85 (synonym of Nordus; type species: xanthocerus).

  • — Scheerpeltz, 1971a (key to species; notes).

aeneiceps Sharp, 1876: 115 (Brachydirus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 101 (Brachydirus; characters; Brazil; Bolivia; Colombia).

  • Distribution: Brazil, Bolivia, Columbia.

aeneipennis Bernhauer, 1917i: 115 (Brachydirus; Type locality: Peru: 1000–1800 m, 11°3′s., 75°17′w).

  • — Scheerpeltz, 1933: 1416 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 95 (Brachydirus; characters; Bolivia).

  • Distribution: Peru, Bolivia.

amazonicus Sharp, 1876: 113 (Brachydirus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 107 (Brachydirus; characters; Brazil; Bolivia).

  • Distribution: Brazil.

antennatus Sharp, 1876: 110 (Brachydirus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1960c: 120 (Brachydirus; Bolivia).

  • — Scheerpeltz, 1971a: 96 (Brachydirus; characters; Bolivia; Ecuador; Colombia).

  • — Frank, 1982: 37 (Nordus; list of parasites).

  • Distribution: Bolivia, Brazil, Ecuador, Colombia.

banghaasi Bernhauer, 1905a: 184 (Brachydirus; Type locality: Bolivia: Mapiri).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 104 (Brachydirus; characters; Bolivia).

  • Distribution: Bolivia.

batesi Sharp, 1876: 114 (Brachydirus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 100 (Brachydirus; characters; Brazil; Bolivia).

  • Distribution: Brazil, Bolivia.

championi Sharp, 1884: 350 (Brachydirus; Type locality: Guatemala, Zapote, San Isidro 1600 feet).

  • — Fauvel, 1891: 107 (Brachydirus; synonym of testaceus).

  • — Bernhauer and Schubert, 1914: 405 (Brachydirus; synonym of testaceus).

  • — Scheerpeltz, 1933: 1416 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 108 (Brachydirus; characters; Guatemala; El Salvador).

  • Distribution: Guatemala, El Salvador.

chapareanus Scheerpeltz, 1971a: 103 (Brachydirus; Type locality: Chaparé in Ost-Bolivien).

  • Distribution: Bolivia.

cribricollis Sharp, 1876: 112 (Brachydirus; Type locality: Ega; St. Paulo).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 107 (Brachydirus; characters; Brazil; Guianas; Venezuela; Colombia).

  • Distribution: Brazil, “Guianas”, Venezuela, Colombia.

densiventris Bernhauer, 1917c: 21 (Brachydirus; Type locality: Columbien: Ost-Cordilleren bei Sosomoco, 900 m).

  • — Scheerpeltz, 1933: 1416 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 104 (Brachydirus; characters; Colombia).

  • Distribution: Colombia.

diversiventris Bernhauer, 1917c: 22 (Brachydirus; Type locality: Peru: Cordilleren in der Umgebung von Cerro de Pasca).

  • — Scheerpeltz, 1933: 1416 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 103 (Brachydirus; characters; Peru; Bolivia).

  • Distribution: Bolivia, Peru.

divisus Sharp, 1884: 349 (Brachydirus; Type locality: Panama, Bugaba).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 99 (Brachydirus; characters; Panama; Costa Rica; Nicaragua).

  • Distribution: Panama, Costa Rica, Nicaragua.

ecuadorensis Scheerpeltz, 1971a: 104 (Brachydirus; Type locality: Südwest-Ecuador).

  • Distribution: Ecuador.

erosus Bernhauer, 1917c: 20 (Brachydirus; Type locality: Brasilien: Santa Catharina: Rio Capivary).

  • — Scheerpeltz, 1933: 1416 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 98 (Brachydirus; characters; Brazil).

  • Distribution: Brazil.

facialis Erichson, 1839b: 354 (Staphylinus; Type locality: Cayenna).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 101 (Brachydirus; characters; French Guiana; Venezuela).

  • Distribution: French Guiana, Venezuela.

fungicola Sharp, 1884: 351 (Brachydirus; Type locality: Panama, Bugaba 800 to 1500 feet, Volcan de Chiriqui 3000 to 4000 feet).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 108 (Brachydirus; characters; Panama; Costa Rica; El Salvador).

  • Distribution: Panama, Costa Rica, El Salvador.

gebieni Bernhauer, 1917c: 20 (Brachydirus; Type locality: Brasilien: Jatahy).

  • — Scheerpeltz, 1933: 1416 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 103 (Brachydirus; characters; Brazil).

  • Distribution: Brazil.

gracilis Sharp, 1884: 349 (Brachydirus; Type locality: Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 105 (Brachydirus; characters; Guatemala; El Salvador).

  • Distribution: Nicaragua, El Salvador, Guatemala.

horni Bernhauer, 1927d: 168 (Brachydirus; Type locality: Kolumbien: Magdalena).

  • — Scheerpeltz, 1933: 1416 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 95 (Brachydirus; characters; Colombia).

  • Distribution: Colombia.

longipes Sharp, 1876: 115 (Brachydirus; Type locality: Para).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 102 (Brachydirus; characters; Bolivia).

  • Distribution: Brazil, Bolivia.

major Bernhauer, 1905a: 186 (Brachydirus; Type locality: Peru: Pachitea; Bolivia: Mapiri).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 105 (Brachydirus; characters; Bolivia; Peru; Ecuador).

  • Distribution: Ecuador, Peru, Bolivia.

maculiceps Sharp, 1876: 109 (Brachydirus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 98 (Brachydirus; characters; Peru; Brazil; Ecuador; Colombia; Bolivia).

  • Distribution: Brazil, Colombia, Ecuador, Peru, Bolivia.

nigroscutellatus Bernhauer, 1905a: 185 (Brachydirus; Type locality: Bolivia: Mapiri).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 101 (Brachydirus; characters).

  • Distribution: Bolivia.

occipitalis Bernhauer, 1905a: 185 (Brachydirus; Type locality: Costa Rica: San Carlos).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 106 (Brachydirus; characters; Costa Rica; El Salvador; Panama; Ecuador; Colombia; Bolivia; Peru).

  • Distribution: Bolivia, Peru, Ecuador, Colombia, Panama, Costa Rica, El Salvador.

picticornis Solsky, 1872: 307 (Brachydirus; Type locality: Not cited).

  • — Solsky, 1875: 18 (Brachydirus; characteres; Peru).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 103 (Brachydirus; characters; Peru).

  • Distribution: Peru.

schadei Scheerpeltz, 1971a: 97 (Brachydirus; Type locality: Paraguay: Umgebung von Villarica).

  • Distribution: Paraguay.

schuberti Wendeler, 1926: 74 (Brachydirus; Type locality: Peru: Oberer Madre de Dios, 500 m).

  • — Scheerpeltz, 1933: 1416 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 97 (Brachydirus; characters; Peru; Bolivia).

  • Distribution: Peru, Bolivia.

simplex Sharp, 1876: 112 (Brachydirus; Type locality: Para; Peru).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 102 (Brachydirus; characters; Brazil; Bolivia; Peru).

  • — Frank, 1982: 37 (Nordus; list of parasites).

  • Distribution: Brazil, Bolivia, Peru.

solitarius Sharp, 1884: 350 (Brachydirus; Type locality: Nicaragua, Fort San Carlos).

  • — Bernhauer and Schubert, 1914: 404 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 106 (Brachydirus; characters; Nicaragua; Costa Rica; El Salvador).

  • Distribution: El Salvador, Nicaragua, Costa Rica.

speciosus Bernhauer, 1905a: 187 (Brachydirus; Type locality: Bolivia).

  • — Bernhauer and Schubert, 1914: 405 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 99 (Brachydirus; characters; Bolivia; Ecuador).

  • Distribution: Bolivia.

styloceros Sharp, 1876: 111 (Brachydirus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 405 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 106 (Brachydirus; characters; Brazil; Bolivia; Peru).

  • Distribution: Brazil, Bolivia, Peru.

terminalis Sharp, 1884: 348 (Brachydirus; Type locality: Guatamala, Panima in Ver Paz).

  • — Bernhauer and Schubert, 1914: 405 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 97 (Brachydirus; characters; Guatemala).

  • Distribution: Guatemala.

testaceus Fabricius, 1801: 595 (Staphylinus; [preoccupied]; Type locality: America meridionalis).

  • — Erichson, 1839b: 354 (Staphylinus; characters; Columbia).

  • — Gemminger and Harold, 1868: 598 (Brachydirus; checklist).

  • — Fauvel, 1891: 107 (Brachydirus; Venezuela).

  • — Bernhauer and Schubert, 1914: 405 (Brachydirus; catalog).

  • — Scheerpeltz, 1933: 1416 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 107 (Brachydirus; characters; Colombia; Ecuador; Venezuela; El Salvador; Guatemala).

  • — Herman, 2001: 45 (Nordus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: Ecuador, Colombia, Venezuela, El Salvador, Guatemala.

velitaris Erichson, 1839b: 354 (Staphylinus; Type locality: Brasilia).

  • — Bernhauer and Schubert, 1914: 405 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 99 (Brachydirus; characters; Brazil).

  • Distribution: Brazil.

xanthocerus Nordmann, 1837: 131 (Brachydirus; Type locality: Bahia).

  • — Erichson, 1839b: 353 (Staphylinus; characters; Brazil).

  • — Solsky, 1872: 307 (Brachydirus; Peru).

  • — Bernhauer and Schubert, 1914: 405 (Brachydirus; catalog).

  • — Scheerpeltz, 1971a: 96 (Brachydirus; characters; Brazil; Peru; Bolivia; Ecuador).

  • Distribution: Brazil, Ecuador, Peru, Bolivia.

zischkai Scheerpeltz, 1971a: 96 (Brachydirus; Type locality: Ost-Bolivien: Yungas de Palmar).

  • Distribution: Bolivia.

Ocyolinus [4 species; Neotropical Region]

Ocyolinus Sharp, 1884: 362 (species included: rugatus, amethystinus). Type species: Ocyolinus rugatus Sharp, fixed by virtual monotypy and direct implication.

  • — Bernhauer and Schubert, 1914: 402 (world catalog; 4 species).

  • — Blackwelder, 1944: 140 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 267 (type species).

amethystinus Sharp, 1884: 363 (Ocyolinus; Type locality: Costa Rica).

  • — Bernhauer and Schubert, 1914: 402 (Ocyolinus; catalog).

  • Distribution: Costa Rica.

ganglbaueri Bernhauer, 1906b: 199 (Ocyolinus; Type locality: Venezuela).

  • — Bernhauer and Schubert, 1914: 402 (Ocyolinus; catalog).

  • Distribution: Venezuela.

rugatus Sharp, 1884: 363 (Ocyolinus; Type locality: Venezuela).

  • — Bernhauer and Schubert, 1914: 402 (Ocyolinus; catalog).

  • Distribution: Venezuela.

vulneratus Bernhauer, 1906b: 199 (Ocyolinus; Type locality: Venezuela).

  • — Bernhauer and Schubert, 1914: 402 (Ocyolinus; catalog).

  • Distribution: Venezuela.

Oligotergus [20 species; Neotropical Region]

Oligotergus Bierig, 1937: 204 (subgenus of Philothalpus; species included: oculatus). Type species: Philothalpus (Oligotergus) oculatus Bierig, fixed by monotypy.

  • — Gemminger and Harold, 1868: 598 (cited as Philothalpus; catalog).

  • — Sharp, 1876: 133 (notes).

  • — Duvivier, 1883: 154 (cited as Philothalpus; catalog).

  • — Sharp, 1884: 354 (cited as Philothalpus; notes; Mexican and Central American species).

  • — Bernhauer and Schubert, 1914: 402 (cited as Philothalpus; world catalog; 16 species).

  • — Scheerpeltz, 1933: 1415 (cited as Philothalpus; world catalog supplement).

  • — Blackwelder, 1943: 451 (cited as Philothalpus; characters; type species: anceps; key to species of West Indies).

  • — Blackwelder, 1943: 451 (synonym of Philothalpus; type species: oculatus).

  • — Blackwelder, 1944: 140 (cited as Philothalpus; checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 303 (cited as Philothalpus; type species).

  • — Blackwelder, 1952: 271 (subgenus of Philothalpus; type species: oculatus).

  • — Herman, 2001: 10 (valid name for species formerly in Philothalpus).

anceps Erichson, 1839, see: fasciatus Nordmann, 1837.

chiriquensis Sharp, 1884: 358 (Philothalpus; Type locality: Panama, Volcan de Chiriqui 2000 to 3000 feet).

  • — Bernhauer and Schubert, 1914: 402 (Philothalpus; catalog).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Panama.

cyanipennis Wendeler, 1956b: 261 (Philothalpus; Type locality: Brasilien: Nova Teutonia).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Brazil.

debilis Sharp, 1884: 355 (Philothalpus; Type locality: Panama, Bugaba).

  • — Bernhauer and Schubert, 1914: 402 (Philothalpus; catalog).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Panama.

deyrollei Solsky, 1867, see: Styngetus.

difficilis Sharp, 1884: 355 (Philothalpus; Type locality: Guatemala, San Isidro 1600 feet, El Tumbador 2500 feet, Mirandilla 1700 feet; Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1914: 402 (Philothalpus; catalog).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Guatemala, Nicaragua.

fasciatus Nordmann, 1837: 112 (Philonthus; Type locality: Mexico).

  • — Erichson, 1839b: 397 (Staphylinus; characters; Mexico).

  • — Kraatz, 1857d: 540 (Philothalpus).

  • — Gemminger and Harold, 1868: 598 (Philothalpus).

  • — Sharp, 1884: 355 (Philothalpus; Mexico).

  • — Fauvel, 1891: 108 (Philothalpus; synonym of anceps).

  • — Bernhauer and Schubert, 1914: 402 (Philothalpus; variety of anceps).

  • — Blackwelder, 1943: 452 (Philothalpus; characters; Trinidad; British Guiana; Venezuela).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: West Indies, Guyana, Venezuela, Brazil, Bolivia, Peru, Colombia, Panama, Guatemala, Mexico.

anceps Erichson, 1839b: 396 (Staphylinus; [preoccupied]; Type locality: Columbia).

  • — Kraatz, 1857d: 540 (Philothalpus).

  • — Gemminger and Harold, 1868: 598 (Philothalpus).

  • — Fauvel, 1891: 108 (Philothalpus; Venezuela).

  • — Gemminger and Harold, 1868: 598 (Philothalpus).

  • — Fauvel, 1901: 85 (Philothalpus; variety of segmentarius; Colombia; Mexico; Guatemala; Panama; Venezuela; Guiana; Brazil; Bolivia; Peru).

  • — Bernhauer and Schubert, 1914: 402 (Philothalpus; catalog).

  • — Blackwelder, 1943: 452 (Philothalpus; synonym of fasciatus).

  • — Marcuzzi, 1977: 13 (Philothalpus; Venezuela).

segmentarius Erichson, 1839b: 397 (Staphylinus; Type locality: Columbia).

  • — Kraatz, 1857d: 540 (Philothalpus).

  • — Gemminger and Harold, 1868: 598 (Philothalpus).

  • — Solsky, 1872: 306 (Philothalpus; Peru).

  • — Sharp, 1884: 355 (Philothalpus; characters; Guatemala; Panama; Colombia).

  • — Fauvel, 1891: 108 (Philothalpus; variety of anceps).

  • — Bernhauer and Schubert, 1914: 402 (Philothalpus; variety of anceps).

  • — Blackwelder, 1943: 452 (Philothalpus; synonym of fasciatus).

luteipes Sharp, 1876: 133 (Philothalpus; Type locality: Para; Ega).

  • — Fauvel, 1891: 108 (Philothalpus; synonym of anceps).

  • — Bernhauer and Schubert, 1914: 402 (Philothalpus; synonym of anceps).

  • — Blackwelder, 1943: 452 (Philothalpus; synonym of fasciatus).

incongruus Sharp, 1876: 135 (Philothalpus; Type locality: Ega).

  • — Duvivier, 1883: 154 (Philothalpus; cited as incongrusus; catalog).

  • — Bernhauer and Schubert, 1914: 402 (Philothalpus; catalog).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Brazil.

latus Sharp, 1876: 134 (Philothalpus; Type locality: St. Paulo).

  • — Bernhauer and Schubert, 1914: 402 (Philothalpus; catalog).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Brazil.

lemoulti Bernhauer, 1915: 14 (Philothalpus; Type locality: Französisch Guyana, Roches de Kouron).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: French Guiana.

luridipes Erichson, 1839b: 395 (Staphylinus; Type locality: Columbia).

  • — Kraatz, 1857d: 540 (Philothalpus).

  • — Gemminger and Harold, 1868: 598 (Philothalpus).

  • — Bernhauer and Schubert, 1914: 402 (Philothalpus; catalog).

  • — Marcuzzi, 1977: 13 (Philothalpus; Venezuela).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Colombia, Venezuela.

luteipes Sharp, 1876, see: fasciatus Nordmann, 1837.

mexicanus Sharp, 1884: 357 (Philothalpus; Type locality: Mexico, Cordova; Guatemala, Cubilguitz in Vera Paz).

  • — Bernhauer and Schubert, 1914: 402 (Philothalpus; catalog).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Mexico, Guatemala. nigriventris Sharp, 1884: 356 (Philothalpus; Type locality: Guatemala, San Geronimo).

  • — Bernhauer and Schubert, 1914: 402 (Philothalpus; catalog).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Guatemala.

oculatus Bierig, 1937: 205 (Philothalpus; subgenus Oligotergus; Type locality: Panama: Puerto Armuelles).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Panama.

ogloblini Bernhauer, 1934b: 216 (Philothalpus; Type locality: Argentinien: Misiones: Loreto).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Argentina.

paederiformis Sharp, 1884: 359 (Philothalpus; Type locality: Mexico, Cuernavaca).

  • — Bernhauer and Schubert, 1914: 402 (Philothalpus; catalog).

  • — Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 62 (Philothalpus; necrophilous species; distribution in Mexico).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Mexico.

pectoralis Sharp, 1884: 356 (Philothalpus; Type locality: Guatemala, San Geronimo).

  • — Bernhauer and Schubert, 1914: 403 (Philothalpus; catalog).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Guatemala.

philonthiformis Sharp, 1884: 358 (Philothalpus; Type locality: Panama, Volcan de Chiriqui 4000 to 6000 feet).

  • — Bernhauer and Schubert, 1914: 403 (Philothalpus; catalog).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Panama.

prolixus Sharp, 1884: 357 (Philothalpus; Type locality: Panama, Bugaba 800 to 1500 feet).

  • — Bernhauer and Schubert, 1914: 403 (Philothalpus; catalog).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Panama.

segmentarius Erichson, 1839, see: fasciatus Nordmann, 1837.

subtilis Sharp, 1884: 358 (Philothalpus; Type locality: Mexico, Cuernavaca, Mirador).

  • — Bernhauer and Schubert, 1914: 403 (Philothalpus; catalog).

  • — Marquez-Luna and Navarrete Heredia, 1995: 35 (Philothalpus; in detritus of Atta mexicana; Mexico).

  • — Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 62 (Philothalpus; necrophilous species; distribution in Mexico).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Mexico.

terminalis Erichson, 1839b: 396 (Staphylinus; Type locality: Brasilia).

  • — Kraatz, 1857d: 540 (Philothalpus).

  • — Gemminger and Harold, 1868: 598 (Philothalpus).

  • — Bernhauer and Schubert, 1914: 403 (Philothalpus; catalog).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Brazil.

tinctiventris Sharp, 1884: 357 (Philothalpus; Type locality: Guatemala, Zapote).

  • — Bernhauer and Schubert, 1914: 403 (Philothalpus; catalog).

  • — Herman, 2001: 10 (Oligotergus).

  • Distribution: Guatemala.

Pammegus [4 species; Palaearctic and Oriental Regions]

Pammegus Fauvel, 1895: 271 (species included: flavipes). Type species: Euryporus flavipes Fauvel, fixed by monotypy.

  • — Fauvel, 1902a: 42 (characters in key; notes).

  • — Bernhauer and Schubert, 1914: 396 (world catalog; 1 species).

  • — Cameron, 1925: 70 (catalog of Indian species).

  • — Cameron, 1932: 253 (characters).

  • — Scheerpeltz, 1933: 1411 (world catalog supplement).

  • — Blackwelder, 1952: 287 (type species: flavipes).

  • — Scheerpeltz, 1965: 278 (characters; type species: flavipes).

  • — Shibata, 1986a: 124 (supplemental list of species from Taiwan).

  • — Schillhammer, 1999b: 95 (genus does not occur in Ethiopian region).

  • — Hayashi, 2000: 24 (notes on empodial setae).

andrewesi Cameron, 1921: 271 (Pammegus; Type locality: Nilgiri Hills).

  • — Cameron, 1932: 254 (Pammegus; variety of flavipes; characters).

  • — Scheerpeltz, 1933: 1411 (Pammegus; variety of flavipes).

  • — Schillhammer, 1999b: 95 (Pammegus; valid species).

  • Distribution: India.

deuvei Coiffait, 1984, see: Algon.

flavipes Fauvel, 1884: 241 (Euryporus; [Note: Bernhauer and Schubert listed this species in Pammegus (1914: 396) and Euryporus (1916: 414) but later workers listed it in Pammegus]; Type locality: Sumatra: Boenga mas, Palembang).

  • — Fauvel, 1895: 272 (Pammegus; characters; Sumatra; Sikkim).

  • — Bernhauer and Schubert, 1914: 396 (Pammegus; catalog).

  • — Bernhauer and Schubert, 1916: 414 (Euryporus; catalog).

  • — Cameron, 1932: 254 (Pammegus; characters; India; Sumatra).

  • — Scheerpeltz, 1933: 1411 (Pammegus; catalog).

  • — Cameron, 1933d: 392 (Pammegus; Sumatra).

  • — Cameron, 1937: 26 (Pammegus; Java).

  • — Shibata, 1979: 22 (Pammegus; characters; Taiwan).

  • Distribution: Taiwan, Indonesia, Myanmar, Nepal, India.

javanus Cameron, 1937: 27 (Pammegus; Type locality: Java: Telok Batong [Note: Sumatra, not Java, see Schillhammer, 1999]).

  • — Schillhammer, 1999b: 95 (Pammegus; type locality is in Sumatra, not Java).

  • Distribution: Indonesia.

ruficollis Fauvel, 1895: 272 (Pammegus; variety of flavipes; Type locality: Bírmanie, Teinzo).

  • — Bernhauer and Schubert, 1914: 396 (Pammegus; synonym of flavipes).

  • — Cameron, 1932: 254 (Pammegus; variety of flavipes; characters; Burma).

  • — Scheerpeltz, 1933: 1411 (Pammegus; variety of flavipes).

  • — Cameron, 1937: 26 (Pammegus; variety of flavipes; Java).

  • — Coiffait, 1982a: 233 (Pammegus, cited as Psammegus; variety of flavipes; Nepal).

  • — Schillhammer, 1999b: 95 (Pammegus; valid species).

  • Distribution: Nepal, Myanmar, Indonesia.

Paraxenopygus [3 species; Neotropical Region]

Paraxenopygus Bernhauer, 1911a: 415 (species included: tremolerasi). Type species: Paraxenopygus tremolerasi Bernhauer, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 400 (world catalog; 1 species).

  • — Bruch, 1928a: 429 (catalog; Argentina).

  • — Scheerpeltz, 1933: 1414 (world catalog supplement).

  • — Blackwelder, 1944: 140 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 294 (type species: tremolerasi).

opacipennis Bernhauer, 1927d: 166 (Paraxenopygus; Type locality: Brasilien: Itatiaya, Est. do Rio, 700 m).

  • — Scheerpeltz, 1933: 1414 (Paraxenopygus; catalog).

  • Distribution: Brazil.

peruvianus Bernhauer, 1915l: 298 (Paraxenopygus; Type locality: Peru: Chanchamayo, 1500 m).

  • — Scheerpeltz, 1933: 1414 (Paraxenopygus; catalog).

  • Distribution: Peru.

tremolerasi Bernhauer, 1911a: 415 (Paraxenopygus; Type locality: Uruguay, Canelones, Progreso).

  • — Bernhauer and Schubert, 1914: 400 (Paraxenopygus; catalog).

  • Distribution: Uruguay.

Phanolinopsis [1 species; Neotropical Region]

Phanolinopsis Scheerpeltz, 1968: 67 (species included: discedens). Type species: Phanolinus discedens Sharp, fixed by original designation and monotypy.

discedens Sharp, 1884: 368 (Phanolinus; Type locality: Panama, Bugaba).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 67 (Phanolinopsis; characters).

  • Distribution: Panama.

Phanolinus [34 species; Neotropical Region]

Phanolinus Sharp, 1884: 363 (species included: obsoletus, flohri, divisus, gravidus, pollens, auratus, godmani, longulus, punctatus, speciosus, discedens, pretiosus, amoenus, cyanescens, cupreus, violaceus, venustus). Type species: Phanolinus flohri Sharp, fixed by subsequent designation by Blackwelder, 1952: 300.

  • — Bernhauer and Schubert, 1914: 401 (world catalog; 25 species).

  • — Scheerpeltz, 1933: 1415 (world catalog supplement).

  • — Blackwelder, 1944: 140 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Scheerpeltz, 1968: 55 (key to species).

amoenus Guérin-Méneville, 1844: 14 (Philonthus; [Note: Preoccupied as secondary homonym]; Type locality: Nouvelle Granade).

  • — Sharp, 1884: 364 (Phanolinus).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 57 (Phanolinus; characters).

  • Distribution: Colombia.

auratus Sharp, 1884: 366 (Phanolinus; Type locality: Guatemala, Purula in Vera Paz).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 65 (Phanolinus; characters).

  • Distribution: Guatemala.

aureiventris Bernhauer, 1906b: 200 (Phanolinus; Type locality: Bolivia).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 65 (Phanolinus; characters).

  • Distribution: Bolivia.

bechyneorum Scheerpeltz, 1968: 51, 64 (Phanolinus; Type locality: Gebiet von Tucano in Brasilien, Est. Amazonas).

  • Distribution: Brazil.

bolivianus Bernhauer, 1935c: 91 (Phanolinus; Type locality: Bolivien).

  • — Scheerpeltz, 1968: 65 (Phanolinus; characters).

  • Distribution: Bolivia.

chiriquensis Bernhauer, 1915: 13 (Phanolinus; Type locality: Vulkan Chiriqui in Panama).

  • — Scheerpeltz, 1933: 1415 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 59 (Phanolinus; characters).

  • Distribution: Panama.

coelestus Solsky, 1875: 17 (Trigonopselaphus; Type locality: Pérou, hacienda Amable Maria).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 58 (Phanolinus; characters).

  • Distribution: Peru.

colombinus Bernhauer, 1917i: 111 (Phanolinus; Type locality: kolumbische Ostkordilleren: Pacho, 1900 m; Caucatal).

  • — Scheerpeltz, 1933: 1415 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 63 (Phanolinus; characters).

  • Distribution: Colombia.

cupreus Guérin-Méneville, 1844: 13 (Philonthus; Type locality: Novuelle Granade).

  • — Sharp, 1884: 364 (Phanolinus).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 59 (Phanolinus; characters).

  • Distribution: Colombia.

cyanescens Guérin-Méneville, 1844: 14 (Philonthus; Type locality: Nouvelle Granade).

  • — Sharp, 1884: 364 (Phanolinus).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 56 (Phanolinus; characters).

  • Distribution: Colombia.

discedens Sharp, 1884, see: Phanolinopsis.

dives Erichson, 1847: 89 (Philonthus; Type locality: Republica Peruana).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 61 (Phanolinus; characters).

  • Distribution: Peru.

divinus Bernhauer, 1911a: 414 (Phanolinus; Type locality: Peru).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 66 (Phanolinus; characters).

  • Distribution: Peru.

divisus Sharp, 1884: 365 (Phanolinus; Type locality: Panama, Volcan de Chiriqui 3000 to 4000 feet).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 60 (Phanolinus; characters).

  • Distribution: Panama.

episcopus Bernhauer, 1906b: 201 (Phanolinus; Type locality: Bolivia).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 58 (Phanolinus; characters).

  • Distribution: Bolivia.

ecuadorensis Bernhauer, 1921: 20 (Phanolinus; Type locality: Ecuador).

  • — Scheerpeltz, 1933: 1415 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 62 (Phanolinus; characters).

  • Distribution: Ecuador.

exornatus Solsky, 1875: 15 (Trigonopselaphus; Type locality: Pérou, hacienda Amable Maria).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 66 (Phanolinus; characters).

  • Distribution: Peru.

flohri Sharp, 1884: 364 (Phanolinus; Type locality: Mexico, Jalapa, Almolonga).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 65 (Phanolinus; characters).

  • Distribution: Mexico.

gloriosus Bernhauer, 1917i: 111 (Phanolinus; Type locality: Westkolumbien: Cali am Rio Cauca, St. Antonio, 2000 m).

  • — Scheerpeltz, 1933: 1415 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 61 (Phanolinus; characters).

  • Distribution: Colombia.

godmani Sharp, 1884: 366 (Phanolinus; Type locality: Panama, Volcan de Chiriqui at various elevations below 6000 feet; Guatemala, Cerro Zunil 4000 to 5000 feet).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 61 (Phanolinus; characters).

  • Distribution: Guatemala, Panama.

gravidus Sharp, 1884: 365 (Phanolinus; Type locality: Guatemala, Sinanja in Vera Paz).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 60 (Phanolinus; characters).

  • Distribution: Guatemala.

ilsae Bernhauer, 1927d: 167 (Phanolinus; Type locality: Kolumbien: Muzo).

  • — Scheerpeltz, 1933: 1415 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 58 (Phanolinus; characters).

  • Distribution: Colombia.

longulus Sharp, 1884: 367 (Phanolinus; Type locality: Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 60 (Phanolinus; characters).

  • Distribution: Nicaragua.

magnificus Bernhauer, 1917i: 112 (Phanolinus; Type locality: Westkolumbien: Umgebung von Cali am Rio Cauca, St. Antonio, 2000 m; Rio Aquatal, 1800 m; Alto de las cruces, 2200 m).

  • — Scheerpeltz, 1933: 1415 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 60 (Phanolinus; characters).

  • Distribution: Colombia.

nolkeni Solsky, 1872: 305 (Trigonopselaphus; Type locality: Nouvelle Grenade).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 62 (Phanolinus; characters).

  • Distribution: Colombia.

obsoletus Sharp, 1884: 364 (Phanolinus; Type locality: Mexico).

  • — Sharp, 1887: 784 (Phanolinus; Mexico).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 66 (Phanolinus; characters).

  • Distribution: Mexico.

peruvianus Bernhauer, 1917i: 112 (Phanolinus; Type locality: Peru: Chanchamayo, 1500 m).

  • — Scheerpeltz, 1933: 1415 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 59 (Phanolinus; characters).

  • Distribution: Peru.

pollens Sharp, 1884: 365 (Phanolinus; Type locality: Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1914: 401 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 64 (Phanolinus; characters).

  • Distribution: Nicaragua.

pretiosus Erichson, 1840: 930 (Philonthus; Type locality: Bogotae in Columbia).

  • — Gemminger and Harold, 1868: 597 (Trigonopselaphus).

  • — Solsky, 1872: 306 (Trigonopselaphus; Colombia).

  • — Sharp, 1884: 364 (Phanolinus).

  • — Fauvel, 1891: 110 (Phanolinus; Venezuela).

  • — Bernhauer and Schubert, 1914: 402 (Phanolinus; catalog; Ecuador).

  • — Bernhauer, 1917i: 110 (Phanolinus; characters).

  • — Scheerpeltz, 1933: 1415 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 63 (Phanolinus; cited as type species of Phanolinus; characters).

  • — Marcuzzi, 1977: 13 (Phanolinus; Venezuela).

  • Distribution: Venezuela, Colombia, Ecuador.

vicarius Bernhauer, 1917i: 110 (Phanolinus; subspecies of pretiosus; Type locality: Venezuela).

  • — Scheerpeltz, 1933: 1415 (Phanolinus; subspecies of pretiosus).

  • — Scheerpeltz, 1968: 63 (Phanolinus; variety of pretiosus; characters).

viridissimus Bernhauer, 1917i: 111 (Phanolinus; subspecies of pretiosus; Type locality: Westkolumbien: Cali am Cauca, Cañon del Mt. Tolima, 1700 m).

  • — Scheerpeltz, 1933: 1415 (Phanolinus; variety of pretiosus).

  • — Scheerpeltz, 1968: 62 (Phanolinus; variety of pretiosus; characters).

princeps Bernhauer, 1906b: 201 (Phanolinus; Type locality: Peruvia, Chanchamajo).

  • — Bernhauer and Schubert, 1914: 402 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 67 (Phanolinus; characters).

  • Distribution: Peru.

punctatus Sharp, 1884: 367 (Phanolinus; Type locality: Panama, Volcan de Chiriqui 2000 to 3000 feet).

  • — Bernhauer and Schubert, 1914: 402 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 57 (Phanolinus; characters).

  • Distribution: Panama.

rambouseki Bernhauer, 1925: 36 (Phanolinus; Type locality: Costa Rica: Turialba, 800 m).

  • — Scheerpeltz, 1933: 1415 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 67 (Phanolinus; characters).

  • Distribution: Costa Rica.

sharpi Scheerpeltz, 1968: 64 (Phanolinus; Type locality: Guatemala).

  • Distribution: Guatemala.

speciosus Sharp, 1884: 367 (Phanolinus; Type locality: Mexico, Cordova).

  • — Bernhauer and Schubert, 1914: 402 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 56 (Phanolinus; characters).

  • Distribution: Mexico.

venustus Sharp, 1876: 146 (Trigonopselaphus; Type locality: Ega; Tapajos).

  • — Sharp, 1884: 364 (Phanolinus).

  • — Bernhauer and Schubert, 1914: 402 (Phanolinus; catalog).

  • — Scheerpeltz, 1968: 56 (Phanolinus; characters).

  • Distribution: Brazil.

vicarius Bernhauer, 1917, see: pretiosus Erichson, 1840.

viridissimus Bernhauer, 1917, see: pretiosus Erichson, 1840.

Philothalpus [3 species; Neotropical Region]

Philothalpus Kraatz, 1857d: 540 (species included: fervidus, egregius, viduus, one unavailable name, sticticus, and the species in Staphylinus Fam. IX of Erichson, 1839b: 395 which were luridipes, anceps, terminalis, fasciatus, segmentarius). Type species: Philonthus fervidus Erichson, fixed by subsequent designation by Herman, 2001.

  • — Herman, 2001: 9 (type species designation; discussion; previous designation by Blackwelder, 1943: 451, is erroneous).

Eugastus Sharp, 1876: 139 (species included: bicolor, mundus). Type species: Eugastus mundus Sharp, fixed by subsequent designation by Blackwelder, 1952: 155 [Note: Blackwelder stated that R. Lucas, 1920: 280, designated the type species but the designation was ambiguous because the first two of the three nominal species listed were cited as “mundus et bicolor”].

  • — Duvivier, 1883: 154 (catalog).

  • — Bernhauer and Schubert, 1914: 401 (world catalog; 3 species).

  • — Blackwelder, 1944: 140 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Herman, 2001: 10 (synonym of Philothalpus).

bicolor Sharp, 1876: 139 (Eugastus; Type locality: St. Paulo).

  • — Bernhauer and Schubert, 1914: 401 (Eugastus; catalog).

  • — Herman, 2001: 10 (Philothalpus).

  • Distribution: Brazil.

chiriquensis Sharp, 1884, see: Oligotergus.

cyanipennis Wendeler, 1956b, see: Oligotergus.

debilis Sharp, 1884, see: Oligotergus.

difficilis Sharp, 1884, see: Oligotergus.

fervidus Erichson, 1840: 505 (Philonthus; Type locality: Columbia).

  • — Kraatz, 1857d: 540 (Philothalpus).

  • — Gemminger and Harold, 1868: 598 (Philothalpus).

  • — Sharp, 1884: 361 (Styngetus).

  • — Fauvel, 1891: 109 (Eugastus; notes; Venezuela).

  • — Bernhauer and Schubert, 1914: 401 (Eugastus; catalog).

  • — Marcuzzi, 1977: 13 (Eugastus; Venezuela).

  • — Herman, 2001: 10 (Philothalpus; type species of genus).

  • Distribution: Colombia, Venezuela.

luteipes Sharp, 1876, see: Oligotergus.

incongruus Sharp, 1876, see: Oligotergus.

latus Sharp, 1876, see: Oligotergus.

lemoulti Bernhauer, 1915, see: Oligotergus.

mexicanus Sharp, 1884, see: Oligotergus.

mundus Sharp, 1876: 140 (Eugastus; Type locality: Lages near Manaos).

  • — Bernhauer and Schubert, 1914: 401 (Eugastus; catalog).

  • — Herman, 2001: 10 (Philothalpus).

  • Distribution: Brazil.

nigriventris Sharp, 1884, see: Oligotergus.

oculatus Bierig, 1937, see: Oligotergus.

ogloblini Bernhauer, 1934, see: Oligotergus.

paederiformis Sharp, 1884, see: Oligotergus.

pectoralis Sharp, 1884, see: Oligotergus.

philonthiformis Sharp, 1884, see: Oligotergus.

prolixus Sharp, 1884, see: Oligotergus.

subtilis Sharp, 1884, see: Oligotergus.

tinctiventris Sharp, 1884, see: Oligotergus.

Plociopterus [31 species; Neotropical Region]

Plociopterus Kraatz, 1857d: 539 (species included are those in Staphylinus Fam. IV of Erichson, 1839b: 355, they are: comptus, ornatus, fetialis, scenicus, venustus; [Note: According to Article 67.2.1, since the preceding five species were not explicitly named by Kraatz in Plociopterus they cannot be cited as the orginally included species. However, Gemminger and Harold (1868: 598) published the first explicit list of species for the genus and cited the same five species]). Type species: Staphylinus fetialis Erichson, fixed by subsequent designation by Blackwelder, 1952: 315.

  • — Gemminger and Harold, 1868: 598 (catalog).

  • — Sharp, 1876: 116 (notes; characters).

  • — Duvivier, 1883: 154 (catalog).

  • — Sharp, 1884: 351 (notes; Mexican and Central American species).

  • — Bernhauer and Schubert, 1914: 406 (world catalog; 25 species).

  • — Scheerpeltz, 1933: 1417 (world catalog supplement).

  • — Blackwelder, 1944: 142 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Scheerpeltz, 1960c: 121 (key to species).

affinis Sharp, 1876: 119 (Plociopterus; Type locality: Para).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 124 (Plociopterus; characters; Brazil).

  • Distribution: Brazil.

atticida Wasmann, 1925: 71 (Plociopterus; Type locality: Not cited).

  • — Scheerpeltz, 1933: 1417 (Plociopterus; cited as atticola; catalog).

  • — Scheerpeltz, 1960c: 121 (Plociopterus; characters; Brazil; Paraguay).

  • Distribution: Brazil, Paraguay.

belti Sharp, 1884: 354 (Plociopterus; Type locality: Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 121 (Plociopterus; characters; Guatemala to Nicaragua and Costa Rica).

  • Distribution: Guatemala, Nicaragua, Costa Rica.

brasilianus Bernhauer, 1906b: 197 (Plociopterus; Type locality: Brasilia).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 124 (Plociopterus; characters; Brazil; Paraguay).

  • Distribution: Brazil, Paraguay.

chlorocephalus Bernhauer, 1917c: 22 (Plociopterus; Type locality: Peru: Chanchamajo; Panama: Chiriqui).

  • — Scheerpeltz, 1933: 1417 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 121 (Plociopterus; characters; Peru; Bolivia; Chile).

  • Distribution: Chile, Bolivia, Peru, Panama.

collaris Sharp, 1884: 352 (Plociopterus; Type locality: Panama, Volcan de Chiriqui).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 121 (Plociopterus; characters; Nicaragua; Costa Rica; Panama).

  • Distribution: Panama, Costa Rica, Nicaragua.

columbinus Bernhauer, 1917c: 22 (Plociopterus; Type locality: Columbien).

  • — Scheerpeltz, 1933: 1417 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 121 (Plociopterus; characters; Colombia; Ecuador).

  • Distribution: Colombia, Ecuador.

comptus Erichson, 1839b: 356 (Staphylinus; Type locality: Brasilia).

  • — Blanchard, 1842: 77 (Staphylinus; Brazil).

  • — Kraatz, 1857d: 539 (Plociopterus).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 124 (Plociopterus; characters; Brazil; Paraguay).

  • Distribution: Brazil, Paraguay.

dimidiatus Sharp, 1876: 120 (Plociopterus; Type locality: Tunantins).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 121 (Plociopterus; characters; Brazil).

  • Distribution: Brazil.

fetialis Erichson, 1839b: 357 (Staphylinus; Type locality: Mexico).

  • — Kraatz, 1857d: 539 (Plociopterus).

  • — Solsky, 1868a: 141 (Plociopterus; Mexico).

  • — Sharp, 1884: 351 (Plociopterus; notes; Mexico; Panama).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 124 (Plociopterus; characters; Mexico and Guatemala to Pamana).

  • Distribution: Mexico, Guatemala, Panama.

forsteri Scheerpeltz, 1960c: 125 (Plociopterus; Type locality: Nordbrasilien: Belem do Para).

  • Distribution: Brazil.

fungi Sharp, 1876: 118 (Plociopterus; Type locality: Para).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 124 (Plociopterus; characters; Brazil).

  • Distribution: Brazil.

goeldii Wasmann, 1925: 72 (Plociopterus; Type locality: Not cited).

  • — Scheerpeltz, 1933: 1417 (Plociopterus; catalog; Brazil).

  • — Scheerpeltz, 1960c: 122 (Plociopterus; characters; Bolivia; Paraguay).

  • Distribution: Brazil, Paraguay, Bolivia.

histrio Sharp, 1884: 353 (Plociopterus; Type locality: Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 121 (Plociopterus; characters; Nicaragua; Costa Rica; Panama).

  • Distribution: Panama, Costa Rica, Nicaragua.

jelskii Solsky, 1872: 306 (Plociopterus; Type locality: Monte Rico).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 121 (Plociopterus; characters; Peru; Bolivia).

  • Distribution: Peru, Bolivia.

laetus Sharp, 1876: 121 (Plociopterus; Type locality: Ega; Garrao).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 123 (Plociopterus; characters; Brazil).

  • — Frank, 1982: 45 (Plociopterus; list of parasites).

  • Distribution: Brazil.

lugubris Sharp, 1884: 352 (Plociopterus; Type locality: Panama, Caldera 1200 feet, Bugaba 800 to 1500 feet).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 125 (Plociopterus; characters; Nicaragua; Costa Rica; Panama).

  • Distribution: Panama, Costa Rica, Nicaragua.

mirandus Sharp, 1876: 124 (Plociopterus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 123 (Plociopterus; characters; Brazil).

  • Distribution: Brazil.

miriventris Bernhauer, 1905: 18 (Plociopterus; Type locality: Peru: Chanchamajo).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 123 (Plociopterus; characters; Peru).

  • Distribution: Peru.

mollis Sharp, 1884: 353 (Plociopterus; Type locality: Panama, Volcan de Chiriqui 2000 to 3000 feet).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 125 (Plociopterus; characters; Guatemala; Honduras; Nicaragua; Costa Rica; Panama).

  • Distribution: Panama, Costa Rica, Nicaragua, Honduras, Guatemala.

nigripes Sharp, 1876: 118 (Plociopterus; Type locality: St. Paulo).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 121 (Plociopterus; characters; Brazil; Paraguay).

  • Distribution: Brazil, Paraguay.

ornatus Erichson, 1839b: 356 (Staphylinus; Type locality: Cayenna).

  • — Kraatz, 1857d: 539 (Plociopterus).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 124 (Plociopterus; characters; Venezuela; Guyana).

  • Distribution: French Guiana, Guyana, Venezuela.

scenicus Erichson, 1839b: 357 (Staphylinus; Type locality: Cayenna).

  • — Kraatz, 1857d: 539 (Plociopterus).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 123 (Plociopterus; characters; Venezuela; Guyana).

  • Distribution: French Guiana, Guyana, Venezuela.

traili Sharp, 1876: 122 (Plociopterus; Type locality: Garrao).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 123 (Plociopterus; characters; Brazil).

  • Distribution: Brazil.

tricolor Sharp, 1876: 117 (Plociopterus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 125 (Plociopterus; characters; Brazil; Bolivia).

  • Distribution: Brazil, Bolivia.

variegatus Sharp, 1884: 353 (Plociopterus; Type locality: Panama, Bugaba 800 to 1500 feet).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 121 (Plociopterus; characters; Nicaragua; Costa Rica; Panama).

  • Distribution: Panama, Costa Rica, Nicaragua.

varipes Sharp, 1884: 351 (Plociopterus; Type locality: Guatemala, Capetillo, Zapote, Las Mercedes 3000 feet, San Isidro 1600 feet).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 125 (Plociopterus; characters; Guatemala; Honduras; Nicaragua).

  • Distribution: Guatemala, Honduras, Nicaragua.

ventralis Sharp, 1876: 122 (Plociopterus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 123 (Plociopterus; characters; Brazil).

  • Distribution: Brazil.

venustus Erichson, 1839b: 357 (Staphylinus; Type locality: Cayenna).

  • — Kraatz, 1857d: 539 (Plociopterus).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 123 (Plociopterus; characters; Venezuela; Guyana).

  • Distribution: French Guiana, Guyana, Venezuela.

virgineus Sharp, 1876: 123 (Plociopterus; Type locality: Fonteboa).

  • — Bernhauer and Schubert, 1914: 406 (Plociopterus; catalog).

  • — Scheerpeltz, 1960c: 123 (Plociopterus; characters; Brazil).

  • Distribution: Brazil.

zischkai Scheerpeltz, 1960c: 125 (Plociopterus; [Note: This species was included in a key to species but not cited as new; Scheerpeltz was listed as author and no earlier reference for the species was given]; Type locality: Ostbolivien).

  • Distribution: Bolivia.

Prianophthalmus [2 species; Ethiopian Region]

Prianophthalmus Bernhauer, 1932b: 155 (species included: hulstaerti). Type species: Prianophthalmus hulstaerti Bernhauer, fixed by monotypy.

  • — Scheerpeltz, 1933: 1419 (world catalog supplement).

  • — Blackwelder, 1952: 321 (type species: hulstaerti).

hulstaerti Bernhauer, 1932b: 156 (Prianophthalmus; Type locality: Equateur: Boende).

  • — Scheerpeltz, 1933: 1419 (Prianophthalmus; catalog).

  • — Fagel, 1952: 2 (Prianophthalmus; characters; Belgian Congo).

  • Distribution: Congo.

ferox Fagel, 1952: 3 (Prianophthalmus; Type locality: Gabon: Loango).

  • Distribution: Gabon.

Prionopedinus [1 species; Neotropical Region]

Prionopedinus Scheerpeltz, 1972a: 36 (replacement name for Prionopus Bernhauer; cited as valid genus). Type species: Trigonopselaphus elvirae Bernhauer, fixed by original designation.

Prionopus Bernhauer, 1921: 20 (subgenus of Trigonopselaphus; species included: elvirae; preoccupied by Billberg, 1820). Type species: Trigonopselaphus elvirae Bernhauer, fixed by monotypy.

  • — Scheerpeltz, 1933: 1415 (subgenus of Trigonopselaphus).

  • — Blackwelder, 1952: 321 (synonym of Trigonopselaphus; type species: elvirae).

elvirae Bernhauer, 1921: 20 (Trigonopselaphus; subgenus Prionopus; Type locality: Kolumbien: Umgebung von Cali am Cauca, Villa Elvira, 1800 m).

  • — Scheerpeltz, 1933: 1415 (Trigonopselaphus; catalog).

  • — Scheerpeltz, 1972a: 36 (Prionopedinus).

  • Distribution: Colombia.

Prionophilonthus [1 species; Ethiopian Region]

Prionophilonthus Scheerpeltz, 1974c: 24 (species included: cervicornis). Type species: Philonthus cervicornis Schubert, fixed by monotypy.

cervicornis Schubert, 1911: 25 (Philonthus; Type locality: Gundet, Eritrea).

  • — Bernhauer and Schubert, 1914: 331 (Philonthus; catalog).

  • — Tottenham, 1949b: 361 (Philonthus; characters).

  • — Scheerpeltz, 1974c: 24 (Prionophilonthus; Ethiopia; Sudan).

  • Distribution: Eritrea, Ethiopia, Sudan.

Rientis [4 species; Palaearctic and Oriental Regions]

Rientis Sharp, 1874: 21 (species included: parviceps). Type species: Rientis parviceps Sharp, fixed by monotypy.

  • — Duvivier, 1883: 141 (catalog).

  • — Bernhauer and Schubert, 1916: 414 (world catalog; 1 species).

  • — Winkler, 1925: 388 (catalog for Palaearctic region).

  • — Blackwelder, 1952: 342 (type species: parviceps).

  • — Scheerpeltz, 1962b: 265 (characters).

  • — Tikhomirova, 1973a: 188 (checklist of species of USSR).

  • — Smetana, 1977b: 180 (member of Staphylinini, subtribe Xanthopygi).

  • — Naomi, 1982: 128 (characters).

  • — Naomi, 1983a: 586 (characters).

  • — Shibata, 1984: 113 (in checklist of species of Japan).

Securipalpus Schubert, 1908: 613 (species included: rudepunctatus). Type species: Securipalpus rudepunctatus Schubert, fixed by monotypy.

  • — Bernhauer and Schubert, 1916: 415 (synonym of Algon).

  • — Cameron, 1932: 274 (characters).

  • — Blackwelder, 1952: 350 (type species: rudepunctatus).

  • — Scheerpeltz, 1962b: 265 (characters).

  • — Scheerpeltz, 1974b: 139 (synonym of Algon).

  • — Smetana, 1977a: 245 (valid genus; characters).

  • — Smetana, 1997b: 180 (synonym of Rientis).

Barypalpus Cameron, 1932: 276 (species included: ruficornis). Type species: Barypalpus ruficornis Cameron, fixed by monotypy.

  • — Scheerpeltz, 1933: 1426 (world catalog supplement).

  • — Blackwelder, 1952: 73 (type species: ruficornis).

  • — Scheerpeltz, 1962b: 265 (characters).

  • — Schillhammer, 1999b: 94 (synonym of Rientis; note).

parviceps Sharp, 1874: 21 (Rientis; Type locality: Copper Temple, Nagasaki).

  • — Bernhauer and Schubert, 1916: 414 (Rientis; catalog).

  • — Nakane, 1963a: 94 (Rientis; characters; habitus photograph; Japan).

  • — Naomi, 1983a: 588 (Rientis; characters; Japan).

  • Distribution: Japan.

rudepunctata Schubert, 1908: 613 (Securipalpus; Type locality: Kulu, Himalaya, ca. 3500 m).

  • — Bernhauer and Schubert, 1916: 415 (Algon; catalog).

  • — Cameron, 1932: 275 (Securipalpus; characters; Himalaya).

  • — Scheerpeltz, 1933: 1426 (Algon; catalog).

  • — Scheerpeltz, 1974b: 142 (Algon; characters; Himalaya).

  • — Smetana, 1977a: 244 (Securipalpus; characters; Bhutan).

  • — Smetana, 1977b: 180 (Rientis).

  • — Coiffait, 1982a: 234 (Securipalpus; Nepal).

  • Distribution: India, Nepal, Bhutan.

ruficornis Cameron, 1932: 276 (Barypalpus; Type locality: Burma: Ruby Mines).

  • — Scheerpeltz, 1933: 1426 (Barypalpus; catalog).

  • — Schillhammer, 1999b: 94 (Rientis).

  • Distribution: Myanmar.

semiaereus Cameron, 1928c: 567 (Algon; Type locality: Sikkim: Darjeeling, 7000 feet).

  • — Cameron, 1932: 272 (Algon; characters; Sikkim).

  • — Scheerpeltz, 1933: 1426 (Algon; catalog).

  • — Scheerpeltz, 1974b: 142 (Algon; characters; India).

  • — Coiffait, 1982a: 275 (Algon; characters).

  • — Boháč, 1993a: 447 (Algon; characters in key).

  • — Schillhammer, 1999b: 94 (Rientis).

  • Distribution: India.

Scaponopselaphus [1 species; Neotropical Region]

Scaponopselaphus Scheerpeltz, 1972a: 38 (species included: mutator). Type species: Scaponopselaphus mutator Sharp, fixed by original designation.

mutator Sharp, 1876: 144 (Trigonopselaphus; Type locality: Pebas).

  • — Bernhauer and Schubert, 1914: 403 (Trigonopselaphus; catalog).

  • — Scheerpeltz, 1972a: 38 (Scaponopselaphus).

  • Distribution: Brazil.

Styngetus [17 species; Neotropical Region]

Styngetus Sharp, 1884: 361 (species included: egregius, fervidus, cupripennis, antennatus, deyrollei, viduus, flavicaudus, championi, anticus). Type species: Philonthus viduus Erichson, fixed by subsequent designation by Blackwelder, 1952: 365.

  • — Bernhauer and Schubert, 1914: 403 (world catalog; 13 species).

  • — Bruch, 1928a: 429 (catalog; Argentina).

  • — Scheerpeltz, 1933: 1416 (world catalog supplement).

  • — Blackwelder, 1944: 141 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Navarrete-Heredia, 1998: 59, 70 (characters; key to species of Mexico).

adrianae Navarrete-Heredia, 1998: 61 (Styngetus; Type locality: Mexico: Morelos: Municipio de Tlayacapa, Camino de San José de los Laureles a Amatlan, 19°00′N, 99°00′W, 1770 m).

  • — Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 62 (Styngetus; necrophilous species; distribution in Mexico).

  • Distribution: Mexico.

analis Bernhauer, 1917, see: viduus Erichson, 1840.

antennatus Guérin-Méneville, 1844: 14 (Philonthus; Type locality: Nouvelle Granade).

  • — Sharp, 1884: 361 (Styngetus).

  • — Bernhauer and Schubert, 1914: 404 (Styngetus; catalog).

  • Distribution: Colombia.

anticus Sharp, 1884: 362 (Styngetus; Type locality: Costa Rica, Volcan de Irazu 6000 to 7000 feet; Panama, Volcan de Chiriqui between 4000 and 6000 feet).

  • — Bernhauer and Schubert, 1914: 404 (Styngetus; catalog).

  • Distribution: Costa Rica, Panama.

argentinus Bernhauer, 1917, see: viduus Erichson, 1840.

championi Sharp, 1884: 362 (Styngetus; Type locality: Guatemala, San Geronimo, Sabo, Cubilguitz, Balheu in Vera Paz).

  • — Bernhauer and Schubert, 1914: 404 (Styngetus; catalog).

  • — Navarrete-Heredia, 1998: 68 (Styngetus; characters; Chiapas).

  • Distribution: Mexico, Guatemala.

cribellatus Solsky, 1868a: 137 (Philonthus; Type locality: Mexique).

  • — Sharp, 1885: 426 (Philonthus; Mexico).

  • — Bernhauer and Schubert, 1914: 404 (Styngetus; catalog).

  • — Navarrete-Heredia, 1998: 69 (notes; not in Styngetus but placement not known).

  • Distribution: Mexico.

cupripennis Guérin-Méneville, 1844: 15 (Philonthus; Type locality: Nouvelle Granade).

  • — Sharp, 1884: 361 (Styngetus).

  • — Fauvel, 1891: 108 (Styngetus; Venezuela).

  • — Bernhauer and Schubert, 1914: 404 (Styngetus; catalog).

  • Distribution: Colombia, Venezuela.

deyrollei Solsky, 1867a: 105 (Philothalpus; Type locality: Méxique).

  • — Gemminger and Harold, 1868: 598 (Philothalpus).

  • — Solsky, 1868a: 141 (Philothalpus; cited as deyrollei mihi; Mexico).

  • — Gemminger and Harold, 1868: 598 (Philothalpus).

  • — Sharp, 1884: 361 (Styngetus; Mexico).

  • — Bernhauer and Schubert, 1914: 404 (Styngetus; catalog).

  • — Navarrete-Heredia, 1998: 66 (Styngetus; characters; notes; Chiapas; Hidalgo; Oaxaca; Puebla; Querétaro; San Luis Potosí; Tamaulipas).

  • Distribution: Mexico.

egregius Erichson, 1840: 505 (Philonthus; Type locality: Bogotae in Columbia).

  • — Kraatz, 1857d: 540 (Philothalpus).

  • — Gemminger and Harold, 1868: 598 (Philothalpus).

  • — Sharp, 1884: 361 (Styngetus).

  • — Bernhauer and Schubert, 1914: 404 (Styngetus; catalog).

  • Distribution: Colombia.

flavicaudus Sharp, 1884: 361 (Styngetus; Type locality: Guatemala, Balheu in Vera Paz).

  • — Bernhauer and Schubert, 1914: 404 (Styngetus; catalog).

  • — Navarrete-Heredia, 1998: 69 (Styngetus; characters; Chiapas).

  • Distribution: Guatemala.

germaini Bernhauer, 1917i: 114 (Styngetus; Type locality: Bolivien: Yuracaris).

  • — Scheerpeltz, 1933: 1416 (Styngetus; catalog).

  • Distribution: Bolivia.

goudoti Fauvel, 1891: 108 (Styngetus; Type locality: Colonie Tovar).

  • — Bernhauer and Schubert, 1914: 404 (Styngetus; catalog).

  • Distribution: Venezuela.

heynei Bernhauer, 1917i: 115 (Styngetus; Type locality: Peru: Chanchamayo, 1500 m).

  • — Scheerpeltz, 1933: 1416 (Styngetus; catalog).

  • Distribution: Peru.

langei Bernhauer, 1905: 18 (Styngetus; Type locality: Peru: Chanchamajo).

  • — Bernhauer and Schubert, 1914: 404 (Styngetus; catalog).

  • Distribution: Peru.

niger Bernhauer, 1906a: 197 (Styngetus; Type locality: Brasilia; Paraguay).

  • — Bernhauer and Schubert, 1914: 404 (Styngetus; catalog).

  • — Scheerpeltz, 1933: 1416 (Styngetus; catalog; Argentina).

  • Distribution: Brazil, Paraguay, Argentina.

nigriventris Bernhauer, 1925: 37 (Styngetus; Type locality: Neu-Granada).

  • — Scheerpeltz, 1933: 1416 (Styngetus; catalog).

  • Distribution: Colombia.

sharpi Fauvel, 1891: 108 (Styngetus; Type locality: Merida).

  • — Bernhauer and Schubert, 1914: 404 (Styngetus; catalog).

  • Distribution: Venezuela.

viduus Erichson, 1840: 506 (Philonthus; Type locality: Columbia).

  • — Kraatz, 1857d: 540 (Philothalpus).

  • — Gemminger and Harold, 1868: 598 (Philothalpus).

  • — Sharp, 1884: 361 (Styngetus; characters; Panama; Colombia).

  • — Fauvel, 1891: 109 (Styngetus; notes; Venezuela).

  • — Bernhauer and Schubert, 1914: 404 (Styngetus; catalog).

  • — Scheerpeltz, 1933: 1416 (Styngetus; catalog).

  • — Marcuzzi, 1977: 14 (Styngetus; Venezuela).

  • Distribution: Guatemala, Costa Rica, Panama, Colombia, Venezuela, Argentina.

analis Bernhauer, 1917i: 114 (Styngetus; variety of viduus; Type locality: Panama: Lino, 800 m; Costa Rica: Carthago, 1000 m und Orosi, 1300 m).

  • — Scheerpeltz, 1933: 1416 (Styngetus; variety of viduus; Guatemala).

  • — Marcuzzi, 1977: 14 (Styngetus; variety of viduus; Venezuela).

argentinus Bernhauer, 1917i: 114 (Styngetus; variety of viduus; Type locality: Provinz Tucuman).

  • — Scheerpeltz, 1933: 1416 (Styngetus; variety of viduus).

Torobus [9 species; Neotropical Region]

Torobus Herman, 2001: 29 (species included: badiipennis, brasilianus, chloris, erithacus, fassli, laetipes, mautnermarkhofi, principalis, purpurascens; characters for genus in Scheerpeltz, 1972: 37–40). Type species: Staphylinus purpurascens Nordmann, fixed by original designation.

badiipennis Nordmann, 1837: 25 (Creophilus; Type locality: Brasilia).

  • — Erichson, 1839b: 400 (Staphylinus; characters; Brazil).

  • — Gemminger and Harold, 1868: 597 (Trigonopselaphus).

  • — Bernhauer and Schubert, 1914: 403 (Trigonopselaphus; catalog).

  • — Scheerpeltz, 1972a: 40 (Trigonopselaphus; characters; Brazil).

  • — Herman, 2001: 29 (Torobus).

  • Distribution: Brazil.

brasilianus Bernhauer, 1906c: 327 (Trigonurus; [Note: Bernhauer described this species in Trigonurus but the citation was probably a lapsus since he compared it with purpurascens which was in Trigonopselaphus at that time]; Type locality: Brasilia).

  • — Bernhauer and Schubert, 1914: 403 (Trigonopselaphus; catalog).

  • — Scheerpeltz, 1972a: 42 (Trigonopselaphus; characters; Paraguay; Brazil).

  • — Herman, 2001: 29 (Torobus).

  • Distribution: Brazil, Paraguay.

chloris Nordmann, 1837: 26 (Creophilus; Type locality: Brasilia ?).

  • — Erichson, 1840: 401 (Staphylinus; characters; Brazil).

  • — P. Lucas, 1857: 50 (Staphylinus; Brazil).

  • — Gemminger and Harold, 1868: 597 (Trigonopselaphus).

  • — Bernhauer and Schubert, 1914: 403 (Trigonopselaphus; catalog).

  • — Scheerpeltz, 1972a: 40 (Trigonopselaphus; characters; Brazil).

  • — Herman, 2001: 29 (Torobus).

  • Distribution: Brazil.

erithacus Nordmann, 1837: 25 (Creophilus; Type locality: Brasilia).

  • — Erichson, 1840: 401 (Staphylinus; characters; Brazil).

  • — Gemminger and Harold, 1868: 597 (Trigonopselaphus).

  • — Bernhauer and Schubert, 1914: 403 (Trigonopselaphus; catalog).

  • — Scheerpeltz, 1972a: 41 (Trigonopselaphus; characters; Brazil).

  • — Herman, 2001: 29 (Torobus).

  • Distribution: Brazil.

fassli Bernhauer, 1917i: 113 (Trigonopselaphus; Type locality: Westkolumbien: Umgebung von Cali am Cauca, Cañon del Monte Tolima, 1700 m).

  • — Scheerpeltz, 1933: 1415 (Phanolinus; catalog).

  • — Scheerpeltz, 1972a: 43 (Trigonopselaphus; characters; Colombia; Ecuador).

  • — Herman, 2001: 29 (Torobus).

  • Distribution: Colombia, Ecuador.

laetipes Bernhauer, 1911a: 418 (Trigonopselaphus; Type locality: ? Paraguay).

  • — Bernhauer and Schubert, 1914: 403 (Trigonopselaphus; catalog).

  • — Scheerpeltz, 1972a: 41 (Trigonopselaphus; characters; Paraguay; Argentina; Brazil).

  • — Herman, 2001: 29 (Torobus).

  • Distribution: Argentina, Paraguay, Brazil.

mautnermarkhofi Scheerpeltz, 1972a: 43 (Trigonopselaphus; Type locality: Ecuador: Rio Napo).

  • — Herman, 2001: 29 (Torobus).

  • Distribution: Ecuador.

principalis Bernhauer, 1911a: 416 (Trigonopselaphus; Type locality: Brasilien).

  • — Bernhauer and Schubert, 1914: 403 (Trigonopselaphus; catalog).

  • — Scheerpeltz, 1972a: 42 (Trigonopselaphus; characters; Brazil).

  • — Herman, 2001: 29 (Torobus).

  • Distribution: Brazil.

purpurascens Nordmann, 1837: 47 (Staphylinus; Type locality: Brasilia).

  • — Erichson, 1840: 402 (Staphylinus; characters; Brazil).

  • — P. Lucas, 1857: 50 (Staphylinus; Brazil).

  • — Gemminger and Harold, 1868: 597 (Trigonopselaphus).

  • — Bernhauer and Schubert, 1914: 403 (Trigonopselaphus; catalog).

  • — Scheerpeltz, 1972a: 41 (Trigonopselaphus; characters; Brazil).

  • — Herman, 2001: 29 (Torobus; type species).

  • Distribution: Brazil.

Triacrus [1 species; Neotropical Region]

Triacrus Nordmann, 1837: 19 (species included: dilatus). Type species: Triacrus dilatus Nordmann, fixed by monotypy.

  • — Erichson, 1839b: 398 (Staphylinus; synonym of Staphylinus).

  • — Lacordaire, 1854: 76 (synonym of Staphylinus).

  • — Gemminger and Harold, 1868: 596 (catalog).

  • — Bernhauer and Schubert, 1914: 397 (world catalog; 1 species).

  • — Bruch, 1928a: 428 (catalog; Argentina).

  • — Bernhauer, 1931a: 84 (characters in key).

  • — Scheerpeltz, 1933: 1419 (world catalog supplement).

  • — Blackwelder, 1944: 140 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 393 (type species: dilatus).

dilatus Nordmann, 1837: 19 (Triacrus; Type locality: Brasilia).

  • — Erichson, 1839b: 398 (Staphylinus; characters; Brazil).

  • — P. Lucas, 1857: 50 (Staphylinus; synonym of superbus).

  • — Bernhauer and Schubert, 1914: 397 (Triacrus; synonym of superbus).

  • — Scheerpeltz, 1933: 1419 (Triacrus; synonym of superbus).

  • — Herman, 2001: 32 (Triacrus; valid name for species).

  • Distribution: Brazil.

superbus Erichson, 1839b: 398 (Staphylinus; Type locality: Brasilia).

  • — P. Lucas, 1857: 50 (Staphylinus; Brazil).

  • — Bernhauer and Schubert, 1914: 397 (Triacrus; catalog).

  • — Scheerpeltz, 1933: 1419 (Triacrus; catalog).

  • — Herman, 2001: 32 (Triacrus; synonym of dilatus).

superbus Erichson, 1839, see: dilatus Nordmann, 1837.

Tricholinus [1 species; Neotropical Region]

Tricholinus Bernhauer, 1922: 15 (species included: pehlkei). Type species: Tricholinus pehlkei Bernhauer, fixed by monotypy.

  • — Scheerpeltz, 1933: 1418 (world catalog supplement).

  • — Blackwelder, 1944: 143 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 395 (type species: pehlkei).

pehlkei Bernhauer, 1922: 16 (Tricholinus; Type locality: Kolumbien: Hac. Pehlke).

  • — Scheerpeltz, 1933: 1418 (Tricholinus; catalog).

  • Distribution: Colombia.

Trigonopalpus [1 species; Ethiopian Region]

Trigonopalpus Cameron, 1951c: 405 (species included: pilosus). Type species: Trigonopalpus pilosus Cameron, fixed by monotypy.

pilosus Cameron, 1951c: 405 (Trigonopalpus; Type locality: Uganda: Kamengo For., 3500 feet).

  • Distribution: Uganda.

Trigonopselaphus [8 species; Neotropical Region]

Trigonopselaphus Gemminger and Harold, 1868: 597 (replacement name for Trigonophorus). Type species: Trigonophorus myrtillinus Nordmann, fixed by objective synonymy with Trigonophorus.

  • — Sharp, 1876: 142 (notes).

  • — Duvivier, 1883: 153 (catalog).

  • — Lynch, 1884: 140 (characters).

  • — Bernhauer and Schubert, 1914: 403 (world catalog; 14 species).

  • — Leng, 1920: 109 (catalog of North American species).

  • — Scheerpeltz, 1933: 1415 (world catalog supplement; [Note: Scheerpeltz cited this genus as valid but included the type species, myrtillinus, in Polyphematiana; if that transfer is accepted then Polyphematiana is a junior synonym of Trigonopselaphus and the remaining species of Trigonopselaphus require a new generic name]).

  • — Blackwelder, 1944: 141 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 397 (type species).

  • — Scheerpeltz, 1972a: 40 (key to species; note; discussion).

  • — Moore and Legner, 1975: 45 (catalog; North America north of Mexico).

  • — Herman, 2001: 28, 29 (other than type species, T. myrtillinus, species formerly in Trigonpselaphus moved to Torobus).

Trigonophorus Nordmann, 1837: 8 (species included: myrtillinus; preoccupied by Stephens, 1829, and Hope, 1831). Type species: Trigonophorus myrtillinus Nordmann, fixed by monotypy.

  • — Erichson, 1839b: 399 (Staphylinus; synonym of Staphylinus).

  • — Lacordaire, 1854: 76 (synonym of Staphylinus).

  • — Kraatz, 1857d: 541, note (characters).

  • — LeConte, 1861a: 64 (characters).

  • — LeConte, 1863: 23 (list of North American species).

  • — Bernhauer and Schubert, 1914: 403 (synonym of Trigonopselaphus).

  • — Blackwelder, 1952: 397 (synonym of Trigonopselaphus; type species: myrtillinus).

Polyphematiana E. Strand, 1915: 122 (replacement name for Polyphemus Bernhauer). Type species: Staphylinus herculeanus Laporte, fixed by objective synonymy with Polyphemus.

  • — Bernhauer, 1931a: 84 (characters in key).

  • — Scheerpeltz, 1933: 1419 (world catalog supplement).

  • — Blackwelder, 1944: 143 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 318 (type species).

  • — Herman, 2001: 28 (synonym of Trigonopselaphus).

Polyphemus Bernhauer, 1914: 397 (species included: herculeanus; [Note: Cited without description but with one available species and so is an available name by indication (Article 12.2.5)]; preoccupied by Mueller, 1776). Type species: Staphylinus herculeanus Laporte, fixed by monotypy [Note: Blackwelder, 1952: 318 erroneously cited the fixation as by subsequent designation].

  • — Bernhauer, 1914a: 51 (characters).

  • — Bernhauer, 1921: 19 (key to species).

  • — Scheerpeltz, 1933: 1419 (synonym of Polyphematiana).

  • — Blackwelder, 1952: 318 (synonym of Polyphematiana; type species).

Lypophemus Bernhauer, 1921b: 74 (replacement name for Polyphemus Bernhauer). Type species: Staphylinus herculeanus Laporte, fixed by objective synonymy with Polyphemus.

  • — Scheerpeltz, 1933: 1419 (synonym of Polyphematiana).

  • — Blackwelder, 1952: 228 (synonym of Polyphematiana; type species).

banghaasi Bernhauer, 1914a: 52 (Polyphemus; Type locality: Brasilien: R. Grande do Sul).

  • — Bernhauer, 1921: 19 (Polyphemus; key characters).

  • — Scheerpeltz, 1933: 1419 (Polyphematiana; catalog).

  • — Herman, 2001: 28 (Trigonopselaphus).

  • Distribution: Brazil.

castaneipennis Lynch, 1884, see: Nausicotus.

coelestis Bernhauer, 1921: 19 (Polyphemus; Type locality: Brasilien).

  • — Scheerpeltz, 1933: 1419 (Polyphematiana; catalog).

  • — Herman, 2001: 28 (Trigonopselaphus).

  • Distribution: Brazil.

coelestus Solsky, 1875, see: Phanolinus.

columbinus Erichson, 1839b: 400 (Staphylinus; Type locality: Brasilia).

  • — Gemminger and Harold, 1868: 597 (Trigonopselaphus).

  • — Bernhauer and Schubert, 1914: 403 (Trigonopselaphus; catalog).

  • — Bernhauer, 1921: 19 (Polyphemus; key characters).

  • — Scheerpeltz, 1933: 1419 (Polyphematiana; catalog).

  • — Scheerpeltz, 1972a: 35 (Polyphematiana).

  • — Herman, 2001: 28 (Trigonopselaphus).

  • Distribution: Brazil.

elvirae Bernhauer, 1921, see: Prionopedinus.

exornatus Solsky, 1875, see: Phanolinus.

fassli Bernhauer, 1917, see: Torobus.

herculeanus Laporte, 1835: 114 (Staphylinus; [Note: Page 113 is numbered twice and there is no page 114; this species is on the second page 113]; Type locality: Brésil).

  • — Erichson, 1839b: 394 (Staphylinus; characters; Brazil).

  • — Bernhauer and Schubert, 1914: 397 (Polyphemus; catalog).

  • — Bernhauer, 1914a: 52 (Polyphemus).

  • — Bernhauer, 1921: 19 (Polyphemus; key characters).

  • — Scheerpeltz, 1933: 1419 (Polyphematiana; catalog).

  • — Herman, 2001: 28 (Trigonopselaphus).

  • Distribution: Brazil.

klimai Bernhauer, 1911, see: Gastrisus.

laetipes Bernhauer, 1911, see: Torobus.

liliputanus Bernhauer, 1923b: 58 (Polyphematiana; Type locality: Brasilien: Minas).

  • — Scheerpeltz, 1933: 1419 (Polyphematiana; catalog).

  • — Herman, 2001: 28 (Trigonopselaphus).

  • Distribution: Brazil.

mautnermarkhofi Scheerpeltz, 1972, see: Torobus.

melzeri Bierig, 1932a: 261 (Polyphemus; Type locality: Brasil-São Paulo, Est. S. Paulo).

  • — Scheerpeltz, 1933: 1419 (Polyphematiana; catalog).

  • — Herman, 2001: 28 (Trigonopselaphus).

  • Distribution: Brazil.

modestus Bernhauer, 1912, see: Gastrisus.

mutator Sharp, 1876, see: Scaponopselaphus.

myrtillinus Nordmann, 1837: 8 (Trigonophorus; Type locality: Brasilia).

  • — Erichson, 1839b: 400 (Staphylinus; characters; Brazil).

  • — Gemminger and Harold, 1868: 597 (Trigonopselaphus; cited as myrtillanus).

  • — Bernhauer and Schubert, 1914: 403 (Trigonopselaphus; cited as myrtillanus; catalog).

  • — Bernhauer, 1921: 19 (Polyphemus; [Note: If this generic assignment is accepted then Polyphematiana is a junior synonym of Trigonopselaphus]; characters).

  • — Scheerpeltz, 1933: 1419 (Polyphematiana; [Note: If this generic assignment is accepted then Polyphematiana is a junior synonym of Trigonopselaphus]; catalog).

  • — Blackwelder, 1944: 141 (Trigonopselaphus).

  • — Scheerpeltz, 1972a: 35 (Polyphematiana; cited as myrtillanus; [Note: If this generic assignment is accepted then Polyphematiana is a junior synonym of Trigonopselaphus]).

  • — Moore and Legner, 1975: 45 (Trigonopselaphus; type species).

  • — Herman, 2001: 28 (Trigonopselaphus).

  • Distribution: Brazil.

nobilis Wendeler, 1956, see: Gastrisus.

nolkeni Solsky, 1872, see: Phanolinus.

opacipennis Sharp, 1876, see: Nausicotus.

opulentus Bernhauer, 1911, see: Gastrisus.

principalis Bernhauer, 1911, see: Torobus.

purpuripennis Bernhauer, 1927, see: Gastrisus.

putidiusculus Tottenham, 1939, see: Gastrius.

staphylinoides Lynch, 1884, see: Nausicotus.

subcoeruleus LeConte, 1863, see: Flohria.

venustus Sharp, 1876, see: Phanolinus.

violaceus Sharp, 1876, see: Elecatopselaphus.

viridicollis Bernhauer, 1908, see: Gastrisus.

zikani Bernhauer, 1927d: 165 (Polyphematiana; Type locality: Brasilien: Mar de Hespanha, E. Minas).

  • — Scheerpeltz, 1933: 1419 (Polyphematiana; catalog).

  • — Herman, 2001: 28 (Trigonopselaphus).

  • Distribution: Brazil.

Tympanophorus [12 species; Neotropical, Nearctic, Palaearctic, Ethiopian, and Oriental Regions]

Tympanophorus Nordmann, 1837: 9 (species included: canaliculatus). Type species: Tympanophorus canaliculatus Nordmann, fixed by monotypy.

  • — Erichson, 1840: 402 (Staphylinus; characters; synonym of Staphylinus).

  • — Lacordaire, 1854: 76 (synonym of Staphylinus).

  • — Kraatz, 1857d: 541, note (notes).

  • — Gemminger and Harold, 1868: 597 (catalog).

  • — Sharp, 1884: 342 (notes; Mexican and Central American species).

  • — Fauvel, 1902a: 42 (characters in key; notes).

  • — Bernhauer and Schubert, 1914: 396 (world catalog; 6 species).

  • — Leng, 1920: 108 (catalog of North American species).

  • — Winkler, 1925: 387 (catalog for Palaearctic region).

  • — Cameron, 1925: 70 (catalog of Indian species).

  • — Cameron, 1932: 251 (characters; key to species of British India).

  • — Scheerpeltz, 1933: 1411 (world catalog supplement).

  • — Blackwelder, 1944: 139 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 400 (type species: canaliculatus).

  • — Scheerpeltz, 1965: 277 (characters; type species: canaliculatus).

  • — Tikhomirova, 1973a: 188 (checklist of species of USSR).

  • — Shibata, 1975: 20 (characters; key to species of Japan and Taiwan).

  • — Moore and Legner, 1975: 47 (catalog; North America north of Mexico).

  • — Moore and Legner, 1979: 36 (characters; notes).

  • — Naomi, 1982: 128 (characters).

  • — Naomi, 1983: 52 (characters; key to species of Japan).

  • — Shibata, 1984: 112 (checklist of species of Japan).

  • — Shibata, 1986a: 125 (supplemental list of species from Taiwan).

  • — Downie and Arnett, 1996: 387 (characters).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 395 (2 Nearctic species; notes; characters in key).

  • — Hayashi, 2000: 24 (notes on empodial setae).

Diaphoetus C. Waterhouse, 1884: 213 (species included: rugosus). Type species: Diaphoetes rugosus C. Waterhouse, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 396 (synonym of Tympanophorus).

  • — Cameron, 1932: 251 (synonym of Tympanophorus).

  • — Blackwelder, 1952: 123 (type species: rugosus).

  • — Scheerpeltz, 1965: 277 (characters; synonym of Tympanophorus).

analis Cameron, 1932: 253 (Tympanophorus; Type locality: Ceylon: Kandy).

  • — Scheerpeltz, 1933: 1411 (Tympanophorus; catalog).

  • Distribution: Sri Lanka.

argentatus Fauvel, 1902 [Note: Nomen nudum], see: rugosus C. Waterhouse, 1884.

canaliculatus Nordmann, 1837: 10 (Tympanophorus; Type locality: Brasilia australior).

  • — Erichson, 1840: 403 (Staphylinus; characters; Brazil).

  • — Bernhauer and Schubert, 1914: 396 (Tympanophorus; catalog).

  • Distribution: Brazil.

concolor Sharp, 1884: 343 (Tympanophorus; Type locality: Costa Rica, Irazu 6000 to 7000 feet).

  • — Sharp, 1887: 784 (Tympanophorus; Mexico).

  • — Bernhauer and Schubert, 1914: 396 (Tympanophorus; catalog).

  • Distribution: Costa Rica, Mexico.

hayashidai Shibata, 1975: 21 (Tympanophorus; Type locality: Mt. Kumotori, Tokyo).

  • — Naomi, 1983: 54 (Tympanophorus; characters; Japan).

  • — J. Li, 1993: 41 (Tympanophorus; China).

  • Distribution: Japan, China.

jacobsoni Bernhauer, 1915h: 235 (Tympanophorus; Type locality: Java: Semarang).

  • — Scheerpeltz, 1933: 1411 (Tympanophorus; catalog).

  • — Cameron, 1933d: 392 (Tympanophorus; [Note: Misidentified as rugosus according to Cameron, 1937: 26]; Sumatra).

  • — Cameron, 1937: 26 (Tympanophorus; Java).

  • — Last, 1987: 55 (Tympanophorus; Papua New Guinea).

  • Distribution: Indonesia, New Guinea.

javanus Cameron, 1937: 26 (Tympanophorus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft.).

  • Distribution: Indonesia.

longiceps Bernhauer, 1915h: 235 (Tympanophorus; Type locality: West Himalaya: Murree).

  • — Cameron, 1932: 253 (Tympanophorus; characters; Himalaya).

  • — Scheerpeltz, 1933: 1411 (Tympanophorus; catalog).

  • Distribution: Pakistan.

puncticollis Erichson, 1840: 554 (Euryporus; Type locality: America septentrionalis).

  • — Solsky, 1868a: 123 (Euryporus; Mexico).

  • — Sharp, 1884: 343 (Tympanophorus; notes; Mexico).

  • — Fall and Cockerell, 1907: 166 (Tympanophorus; New Mexico).

  • — Bernhauer and Schubert, 1914: 396 (Tympanophorus; catalog).

  • — Morgan and Morgan, 1985: 1829 (Tympanophorus; middle Holocene fossil; Michigan).

  • — Campbell and Davies, 1991: 120 (Tympanophorus; checklist of species; Canada).

  • — Downie and Arnett, 1996: 387 (Tympanophorus; characters; Canada; USA).

  • Distribution: Mexico, USA, Canada.

rugosus C. Waterhouse, 1884: 214 (Diaphoetes; Type locality: Larat).

  • — Bernhauer and Schubert, 1914: 396 (Tympanophorus; catalog).

  • — Cameron, 1932: 252 (Tympanophorus; characters; Burma; Sumatra; Timor).

  • — Scheerpeltz, 1933: 1411 (Tympanophorus; catalog).

  • — Scheerpeltz, 1957c: 234 (Tympanophorus; Sumba).

  • Distribution: Myanmar, Indonesia.

argentatus Fauvel, 1902a: 42 (Tympanophorus; published without characters and evidently as a synonym of rugosus; [Note: Nomen nudum (Articles 11.6, 12.1)]; cited from Bírmanie, Sumatra, Java, île Larat).

  • — Bernhauer and Schubert, 1914: 396 (Tympanophorus; synonym of rugosus).

  • — Cameron, 1932: 252 (Tympanophorus; synonym of rugosus).

  • — Scheerpeltz, 1957c: 234 (Tympanophorus; synonym of rugosus).

sauteri Bernhauer, 1907: 385 (Tympanophorus; Type locality: Onsen, 2600 feet).

  • — Bernhauer and Schubert, 1914: 396 (Tympanophorus; catalog).

  • — Shibata, 1975: 24 (Tympanophorus; characters; Japan).

  • — Naomi, 1983: 54 (Tympanophorus; characters; Japan).

  • — Hayashi, 2000: 24 (Tympanophorus; belongs in Anisolinina; notes on empodial setae).

  • Distribution: Japan, Taiwan.

taiwanensis Shibata, 1975: 26 (Tympanophorus; subspecies of sauteri; Type locality: Taiwan: Koantauchi, about 650 m. alt., Nantou Hsien).

schenklingi Bernhauer, 1912f: 204 (Tympanophorus; Type locality: Südost-Kamerun: Lolodorf).

  • — Bernhauer and Schubert, 1914: 396 (Tympanophorus; catalog).

  • Distribution: Cameroon.

taiwanensis Shibata, 1975, see: sauteri Bernhauer, 1907.

ventralis Cameron, 1933d: 392 (Tympanophorus; Type locality: Sumatra: Fort de Kock).

  • Distribution: Indonesia.

Xanthopygoides [1 species; Ethiopian Region]

Xanthopygoides Verdcourt, 1952: 176 (replacement name for Heterogaster Cameron). Type species: Heterogaster niger Cameron, fixed by objective synonymy with Heterogaster.

Heterogaster Cameron, 1951c: 406 (species included: niger; preoccupied by Schilling, 1829, Dejean, 1835, and Macquart, 1844). Type species: Heterogaster niger Cameron, fixed by monotypy.

  • — Verdcourt, 1952: 176 (synonym of Xanthopygoides).

niger Cameron, 1951c: 406 (Heterogaster; Type locality: Kenya Colony: Emali Range, Sultan Hamud, alt. 4900–5900 feet).

  • Distribution: Kenya.

Xanthopygus [43 species (1 doubtful); Nearctic and Neotropical Regions]

Xanthopygus Kraatz, 1857d: 539 (species included: analis, bicolor, calidus, chrysopygus, chrysurus, cyanelytrius, faustus, herilis, hilaris, iopterus, mirabilis, pyraster, sapphirinus, tepidus, xanthopygus along with abdominalis and haemorrhoidalis as listed by Gemminger and Harold, 1868: 597; Note: Kraatz included in the genus the species Erichson listed in “Staphylinus Fam. VII and Philonthus Erichs. pro. parte”. According to Article 67.2.1, the included species must be cited by available names so the first included species were those listed by Gemminger and Harold]). Type species: Staphylinus xanthogygus Nordmann, fixed by absolute tautonymy.

  • — Gemminger and Harold, 1868: 597 (catalog).

  • — Sharp, 1876: 125 (notes).

  • — Duvivier, 1883: 153 (catalog).

  • — Sharp, 1884: 344 (notes; Mexican and Central American species).

  • — Lynch, 1884: 130 (characters).

  • — Bernhauer and Schubert, 1914: 407 (world catalog; 26 species).

  • — Leng, 1920: 109 (catalog of North American species).

  • — Blackwelder, 1943: 449 (characters; type species: xanthopygus; key to species of West Indies).

  • — Blackwelder, 1944: 142 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 405 (type species).

  • — Hatch, 1957: 213 (characters; species of Pacific Northwest).

  • — Scheerpeltz, 1969 (key to Central and South American species).

  • — Moore and Legner, 1979: 37 (characters; notes).

  • — Hayashi, 1997a: 488 (characters).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 395 (1 Nearctic species; notes; characters in key).

Lampropygus Sharp, 1884: 346 (species included; bicolor, xanthopygus, cognatus, analis. Type species: Staphylinus xanthopygus Nordmann, fixed by subsequent designation by R. Lucas, 1920: 361.

  • — Bernhauer and Schubert, 1914: 406 (world catalog; 7 species).

  • — Scheerpeltz, 1933: 1417 (world catalog supplement).

  • — Blackwelder, 1943: 449 (synonym of Xanthopygus; type species: xanthopygus).

  • — Blackwelder, 1952: 208 (type species).

  • — Hayashi, 1997a: 488 (synonym of Xanthopygus).

Heteropygus Bernhauer, 1906b: 195 (subgenus of Lampropygus; species included: giganteus). Type species: Lampropygus (Heteropygus) giganteus Bernhauer, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 406 (subgenus of Lampropygus).

  • — Blackwelder, 1943: 449 (synonym of Xanthopygus; type species: giganteus).

  • — Blackwelder, 1952: 185 (synonym of Xanthopygus; type species: giganteus).

  • — Hayashi, 1997a: 488 (synonym of Xanthopygus).

abdominalis Gemminger and Harold, 1868, see: xanthopygus Nordmann, 1837.

alienus Bernhauer, 1905a: 182 (Xanthopygus; Type locality: Brasilien).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 110 (Xanthopygus; characters; Brazil).

  • Distribution: Brazil.

apicalis Sharp, 1876: 128 (Xanthopygus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 110 (Xanthopygus; characters; Brazil).

  • Distribution: Brazil.

borealis Hatch, 1957: 213 (Xanthopygus; Type locality: Michigan: Cheboygan Co.: Douglas Lake. Also cited from British Columbia).

  • Distribution: Canada, USA.

cacti Horn, 1868: 131 (Xanthopygus; Type locality: Fort Grant, Arizona).

  • — Scheerpeltz, 1933: 1417 (Xanthopygus; catalog).

  • Distribution: USA.

calidus Erichson, 1839b: 364 (Staphylinus; Type locality: Columbia).

  • — Kraatz, 1857d: 539, 540 (Xanthopygus).

  • — Fauvel, 1891: 108 (Xanthopygus; Venezuela).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 113 (Xanthopygus; characters; Mexico; Guatemala; El Salvador; Nicaragua; Costa Rica; Panama; Venezuela; Colombia; Ecuador; Peru; Bolivia; Brazil; Paraguay; Argentina).

  • — Frank, 1982: 48 (Xanthopygus; list of parasites).

  • Distribution: Mexico, Guatemala, El Salvador, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Brazil, Ecuador, Peru, Bolivia, Paraguay, Argentina.

solskyi Sharp, 1876: 126 (Xanthopygus; Type locality: Para).

  • — Sharp, 1884: 344 (Xanthopygus; notes; Guatemala; Nicaragua; Panama; Brazil).

  • — Fauvel, 1891: 108 (Xanthopygus; synonym of calidus).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; synonym of calidus).

chapareanus Scheerpeltz, 1969: 115 (Xanthopygus; Type locality: Bolivia).

  • Distribution: Bolivia.

chrysopygus Nordmann, 1837: 45 (Staphylinus; Type locality: Brasilia).

  • — Erichson, 1839b: 366 (Staphylinus; characters; Brazil).

  • — Kraatz, 1857d: 539, 540 (Xanthopygus).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 117 (Xanthopygus; characters; Brazil; Paraguay).

  • Distribution: Brazil, Paraguay.

chrysurus Nordmann, 1837: 108 (Philonthus; Type locality: Para Americae meridionalis).

  • — Erichson, 1839b: 367 (Staphylinus; characters; Brazil).

  • — Kraatz, 1857d: 539, 540 (Xanthopygus).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1933: 1417 (Xanthopygus; catalog; Brazil; Argentina).

  • — Scheerpeltz, 1969: 114 (Xanthopygus; characters; Brazil; Paraguay; Argentina).

  • Distribution: Brazil, Paraguay, Argentina.

cognatus Sharp, 1876: 131 (Xanthopygus; Type locality: Ega).

  • — Sharp, 1884: 347 (Lampropygus; characters; Mexico; Guatemala; Nicaragua; Panama; Brazil).

  • — Fauvel, 1891: 107 (Lampropygus; Venezuela).

  • — Fauvel, 1901: 85 (Lampropygus; Venezuela; Guyana; Colombia; Ecuador; Bolivia; Peru; Brazil; Central America).

  • — Quezada, Amaya, and Herman, 1970 (Xanthopygus; characters of egg, larva, pupa, and adult; biological notes; use for biological control; El Salvador).

  • — Hayashi, 1997a: 484 (Xanthopygus; characters; Honduras; Nicaragua).

  • Distribution: Mexico, Guatemala, Honduras, El Salvador, Nicaragua, Panama, Colombia, Venezuela, Guyana, Brazil, Ecuador, Peru, Bolivia.

minor Sharp, 1884: 347 (Lampropygus; variety of cognatus; without characters; [Note: Nomen nudum (Article 12.1)]).

collaris Bernhauer, 1925: 38 (Xanthopygus; Type locality: Brasilien: Petropolis).

  • — Scheerpeltz, 1933: 1417 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 111 (Xanthopygus; characters; Brazil).

  • Distribution: Brazil.

corcovadoensis Scheerpeltz, 1969: 111 (Xanthopygus; Type locality: Corcovado bei Rio de Janeiro).

  • Distribution: Brazil.

cyanelytrius Perty, 1830: 31 (Staphylinus; Type locality: Brasil: Prov. Minarum, Serra do Caraça).

  • — Blanchard, 1842: 77 (Staphylinus; Brazil).

  • — Erichson, 1839b: 366 (Staphylinus; characters; Brazil).

  • — Kraatz, 1857d: 539, 540 (Xanthopygus).

  • — Fauvel, 1877a: xxviii (Xanthopygus; Brazil).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1933: 1417 (Xanthopygus; catalog; Brazil; Paraguay).

  • — Scheerpeltz, 1969: 117 (Xanthopygus; characters; Brazil; Paraguay).

  • Distribution: Paraguay, Brazil, Uruguay.

cyanipennis Sharp, 1876: 127 (Xanthopygus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 110 (Xanthopygus; characters; Brazil; Argentina; Paraguay).

  • Distribution: Brazil, Argentina, Paraguay.

depressus Sharp, 1876: 129 (Xanthopygus; Type locality: Para; Ega; St. Paulo; Rio Purus).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1933: 1417 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 115 (Xanthopygus; characters; Brazil).

  • Distribution: Brazil.

dimidiatus Bernhauer, 1917c: 23 (Xanthopygus; Type locality: Bolivien: Yuracares).

  • — Scheerpeltz, 1933: 1417 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 117 (Xanthopygus; characters; Bolivia).

  • Distribution: Bolivia.

elegans Bernhauer, 1905a: 183 (Xanthopygus; Type locality: Brasilien).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 112 (Xanthopygus; characters; Brazil; Paraguay).

  • Distribution: Brazil, Paraguay.

faustus Erichson, 1839b: 368 (Staphylinus; Type locality: Cayenna).

  • — Kraatz, 1857d: 539, 540 (Xanthopygus).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 114 (Xanthopygus; characters; French Guiana, cited as Guyana).

  • Distribution: French Guiana.

flohri Sharp, 1884: 345 (Xanthopygus; Type locality: Mexico, Mirador, Oaxaca; Cordova, Tuxtla; Guatemala, Zapote).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 113 (Xanthopygus; characters; Mexico; Guatemala; Salvador; Panama).

  • Distribution: Mexico, Guatemala, El Salvador, Panama.

giganteus Bernhauer, 1906b: 195 (Lampropygus; subgenus Heteropygus; Type locality: Brasilia).

  • — Bernhauer and Schubert, 1914: 406 (Lampropygus; catalog).

  • Distribution: Brazil.

grimmeri Duvivier, 1883: 153 (Xanthopygus; [Note: Nomen dubium]; replacement name for haemorrhoidalis Grimmer).

  • Distribution: Austria.

haemorrhoidalis Grimmer, 1841: 32 (Staphylinus; [Note: Nomen dubium; preoccupied]; Type locality: Steiermark).

  • — Duvivier, 1883: 153 (Xanthopygus; catalog; synonym of grimmeri).

haemorrhoidalis Germar, 1824: 34 (Staphylinus; [preoccupied]; Type locality: Brasilia).

  • — Blanchard, 1842: 77 (Staphylinus; synonym of pyraster Erichson).

  • — Erichson, 1839b: 367 (Staphylinus; synonym of pyraster).

  • — Fauvel, 1869: 487 (Xanthopygus).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 116 (Xanthopygus; characters; Brazil; Paraguay).

  • — Herman, 2001: 53 (Xanthopygus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: Brazil, Paraguay.

pyraster Erichson, 1839b: 367 (Staphylinus; Type locality: Brasilia).

  • — Blanchard, 1842: 77 (Staphylinus; Brazil).

  • — Kraatz, 1857d: 539, 540 (Xanthopygus).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; synonym of haemorrhoidalis).

  • — Scheerpeltz, 1969: 116 (Xanthopygus; synonym of haemorrhoidalis).

haemorrhoidalis Grimmer, 1841 [Note: Nomen dubium], see: grimmeri Duvivier, 1883 [Note: Nomen dubium].

hilaris Erichson, 1839b: 365 (Staphylinus; Type locality: Cayenna).

  • — Kraatz, 1857d: 539, 540 (Xanthopygus).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 115 (Xanthopygus; characters; French Guiana; Guyana).

  • Distribution: Guyana, French Guiana.

iopterus Erichson, 1839b: 366 (Staphylinus; Type locality: Brasilia).

  • — Kraatz, 1857d: 539, 540 (Xanthopygus).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1933: 1417 (Xanthopygus; catalog; Argentina).

  • — Scheerpeltz, 1969: 111 (Xanthopygus; characters; Venezuela; Brazil; Paraguay; Argentina).

  • Distribution: Venezuela, Brazil, Argentina, Paraguay.

janthinipennis Blanchard, 1842: 78 (Staphylinus; Type locality: Bolivia: Santa-Ana de Chiquitos).

  • — Fauvel, 1869: 487 (Xanthopygus).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 116 (Xanthopygus; characters; Bolivia).

  • Distribution: Bolivia.

luctuosus Blanchard, 1842: 76 (Staphylinus; Type locality: Valle Grande; [Note: See lectotype designation by Newton, 1996a]. Originally cited from entre Chilon et Tasajos, Bolivia).

  • — Fauvel, 1869: 487 (Creophilus).

  • — Bernhauer and Schubert, 1914: 398 (Creophilus; catalog).

  • — Newton, 1996a: 80 (Xanthopygus; lectotype designation: Valle Grande; characters; notes; discussion of collecting locality; listing in Brazil is probably an error; Bolivia).

  • Distribution: Bolivia.

major Bernhauer, 1917i: 116 (Lampropygus; Type locality: Westkolumbien: Villavicentia, 440 m).

  • — Scheerpeltz, 1933: 1417 (Lampropygus; catalog).

  • Distribution: Colombia.

max Blackwelder, 1944: 142 (Xanthopygus; replacement name for peruvianus Bernhauer).

  • Distribution: Peru.

peruvianus Bernhauer, 1916f: 188 (Lampropygus; [Note: Preoccupied as secondary homonym]; Type locality: Peru: Chamchamayo, 1500 m Meereshöhe).

  • — Scheerpeltz, 1933: 1417 (Lampropygus; not preoccupied; catalog).

minor Sharp, 1884 [Note: Nomen nudum], see: cognatus Sharp, 1876.

mirabilis Erichson, 1840: 494 (Philonthus; Type locality: Bogatae in Columbia).

  • — Kraatz, 1857d: 539, 540 (Xanthopygus).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 118 (Xanthopygus; characters; Mexico; Guatemala; Nicaragua; Costa Rica; Panama; Colombia).

  • Distribution: Colombia, Panama, Costa Rica, Nicaragua, Guatemala, Mexico.

morosus Sharp, 1884: 346 (Xanthopygus; Type locality: Mexico, Cordova; Guatemala, Senahu in Vera Paz).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 112 (Xanthopygus; characters; Mexico; Guatemala; El Salvador).

  • Distribution: Mexico, Guatemala, El Salvador.

nigricornis Scheerpeltz, 1969: 111 (Xanthopygus; Type locality: Ost-Brasilien, Sta. Catharina; Hansa Humboldt).

  • Distribution: Brazil.

nigripes Sharp, 1876: 130 (Xanthopygus; Type locality: St. Paulo).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 113 (Xanthopygus; characters; Brazil).

  • Distribution: Brazil.

oliveirae Lynch, 1884: 132 (Xanthopygus; Type locality: Arrecifes).

  • — Bernhauer and Schubert, 1914: 406 (Lampropygus; catalog).

  • Distribution: Argentina.

peruvianus Bernhauer, 1916, see: max Blackwelder, 1944.

peruvianus Solsky, 1872, see: Xenopygus.

pexus Motschulsky, 1858: 667 (Belonuchus; Type locality: Colombie).

  • — Fauvel, 1901: 85 (Lampropygus; Venezuela; Saint Vincent; Dmerara; Guyana; Colombia; Brazil).

  • — Bernhauer and Schubert, 1914: 407 (Lampropygus; catalog).

  • — Blackwelder, 1943: 450 (Xanthopygus; characters; St. Vincent; Trinidad).

  • — Marcuzzi, 1977: 14 (Lampropygus; Venezuela).

  • Distribution: Brazil, Colombia, Venezuela, Guyana, West Indies.

punctatus Bernhauer, 1905a: 183 (Xanthopygus; Type locality: Peru).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 112 (Xanthopygus; characters; Peru; Bolivia).

  • Distribution: Peru, Bolivia.

puncticollis Sharp, 1884: 345 (Xanthopygus; Type locality: Panama, Bugaba, Volcan de Chiriqui 2000 to 3000 feet).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 116 (Xanthopygus; characters; Costa Rica; Panama).

  • Distribution: Panama, Costa Rica.

pyraster Erichson, 1839, see: haemorrhoidalis Germar, 1824.

rufipennis Sharp, 1884: 346 (Xanthopygus; Type locality: Mexico, Oaxaca; Panama, Bugaba, Volcan de Chiriqui 2000 to 4000 feet).

  • — Bernhauer and Schubert, 1914: 407 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 117 (Xanthopygus; characters; Mexico; Guatemala; Nicaragua; Panama).

  • Distribution: Mexico, Guatemala, Nicaragua, Panama.

sapphirinus Erichson, 1839b: 364 (Staphylinus; Type locality: Parà interior; Columbia).

  • — Kraatz, 1857d: 539, 540 (Xanthopygus).

  • — Solsky, 1868a: 141 (Xanthopygus; Mexico).

  • — Solsky, 1872: 306 (Xanthopygus, cited as Xanthopygius; Peru).

  • — Sharp, 1876: 126 (Xanthopygus; notes; Brazil).

  • — Sharp, 1884: 344 (Xanthopygus; characters; Mexico; Guatemala; Panama; Brazil).

  • — Fauvel, 1891: 107 (Xanthopygus; Venezuela).

  • — Fauvel, 1901: 85 (Xanthopygus; Venezuela; Guyana; Colombia; Bolivia; Peru; Brazil; Argentina; Central America).

  • — Bernhauer and Schubert, 1914: 408 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 113 (Xanthopygus; characters; Mexico; Guatemala; El Salvador; Nicaragua; Costa Rica; Panama; Venezuela; Guyana; French Guiana; Colombia; Ecuador; Peru; Bolivia; Brazil; Paraguay; Argentina).

  • Distribution: Mexico, Guatemala, El Salvador, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Guyana, French Guiana, Brazil, Ecuador, Peru, Bolivia, Paraguay, Argentina.

skalitzkyi Bernhauer, 1906b: 195 (Lampropygus; Type locality: Peruvia centr.: Mont. de Nancho).

  • — Bernhauer and Schubert, 1914: 407 (Lampropygus; catalog).

  • Distribution: Peru.

solskyi Sharp, 1876, see: calidus Erichson, 1839.

tepidus Erichson, 1839b: 365 (Staphylinus; Type locality: Cayenna).

  • — Kraatz, 1857d: 539, 540 (Xanthopygus).

  • — Bernhauer and Schubert, 1914: 408 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 115 (Xanthopygus; characters; French Guiana).

  • Distribution: French Guiana.

violaceipennis Bernhauer, 1927d: 169 (Xanthopygus; Type locality: Brasilien: Amazonasgebiet).

  • — Scheerpeltz, 1933: 1417 (Xanthopygus; catalog).

  • Distribution: Brazil.

violaceus Sharp, 1876: 128 (Xanthopygus; Type locality: Conceicao, Rio Mauhes; Tunantins; Ega).

  • — Bernhauer and Schubert, 1914: 408 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 116 (Xanthopygus; characters; Brazil).

  • Distribution: Brazil.

viridipennis Sharp, 1876b: 428 (Xanthopygus; Type locality: Chontales, Nicaragua).

  • — Sharp, 1884: 344 (Xanthopygus; Nicaragua; Panama).

  • — Bernhauer and Schubert, 1914: 408 (Xanthopygus; catalog).

  • — Scheerpeltz, 1969: 114 (Xanthopygus; characters; Guatemala; Nicaragua; Costa Rica; Panama; Colombia; Venezuela; Brazil).

  • Distribution: Guatemala, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Brazil.

xanthopygus Nordmann, 1837: 45 (Staphylinus; Type locality: Mexico: Oaxaca).

  • — Erichson, 1840: 496 (Philonthus; characters; Brazil; Colombia; Mexico).

  • — Kraatz, 1857d: 539, 540 (Xanthopygus).

  • — Redtenbacher, 1867: 29 (Philonthus; Brazil).

  • — Solsky, 1868a: 141 (Xanthopygus; Mexico).

  • — Solsky, 1875: 18 (Xanthopygus, cited as Xanthopygius; Peru).

  • — Sharp, 1876: 131 (Xanthopygus; notes; characters of male; Brazil).

  • — Sharp, 1884: 347 (Lampropygus; characters; Mexico; Guatemala).

  • — Bernhauer and Schubert, 1914: 407 (Lampropygus; catalog).

  • — Hayashi, 1997a: 478 (Xanthopygus; characters; Mexico).

  • Distribution: Mexico, Guatemala, Colombia, Peru, Brazil.

abdominalis Gemminger and Harold, 1868: 597 (Xanthopygus; replacement name for xanthopygus Nordman; [Note: I can find no homonymic use of xanthopygus and rejection as a tautonym is not permitted by the Code]).

  • — Sharp, 1884: 347 (Xanthopygus; synonym of xanthopygus).

  • — Bernhauer and Schubert, 1914: 407 (Lampropygus; synonym of xanthopygus).

  • — Hayashi, 1997a: 478 (Xanthopygus; synonym of xanthopygus).

Xenopygus [5 species; Neotropical and Oceanic Regions]

Xenopygus Bernhauer, 1906b: 196 (species included: analis, bicolor). Type species: Philonthus analis Erichson, fixed by subsequent designation by Blackwelder, 1943: 451.

  • — Bernhauer and Schubert, 1914: 405 (world catalog; 3 species).

  • — Scheerpeltz, 1933: 1416 (world catalog supplement).

  • — Blackwelder, 1943: 451 (synonym of Philothalpus; type species: analis).

  • — Blackwelder, 1944: 142 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 407 (type species).

  • — Irmler, 1979a: 31 (key to species).

  • — Irmler, 1982a: 206 (key to species).

analis Erichson, 1840: 495 (Philonthus; [preoccupied]; Type locality: Cayenna; Columbia).

  • — P. Lucas, 1857: 50 (Philonthus; Brazil).

  • — Kraatz, 1857d: 539, 540 (Xanthopygus).

  • — Solsky, 1868a: 141 (Xanthopygus; Mexico).

  • — Sharp, 1876: 132 (Xanthopygus; characters of male; Brazil).

  • — Sharp, 1884: 348 (Lampropygus; Mexico; Guatemala; Panama; South America).

  • — Fauvel, 1891: 107 (Lampropygus; Venezuela).

  • — Bernhauer, 1906b: 196 (Xenopygus).

  • — Bernhauer and Schubert, 1914: 405 (Xenopygus; catalog).

  • — Scheerpeltz, 1933: 1416 (Xenopygus; catalog; Argentina; Peru).

  • — Blackwelder, 1943: 453 (Philothalpus; characters; Trinidad).

  • — Irmler, 1979a: 31 (Xenopygus; adult and larval characters; notes).

  • — Frank, 1982: 45 (Philothalpus; list of parasites).

  • — Irmler, 1982a: 209 (Xenopygus; characters; Colombia; Guyana).

  • — Nishida, 1994: 73 (Philothalpus; Hawaii).

  • — Nishida, 1997: 62 (Philothalpus; Hawaii).

  • — Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 62 (Xenopygus; necrophilous species; distribution in Mexico).

  • — Herman, 2001: 54 (Xenopygus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: Argentina, Brazil, Peru, French Guiana, Guyana, Venezuela, Colombia, Panama, Guatemala, Mexico, West Indies, Hawaii.

bicolor Laporte, 1835: 115 (Staphylinus; [preoccupied]; Type locality: Cayenne [Note: See neotype designation by Irmler, 1982a]).

  • — Erichson, 1840: 495 (Philonthus; characters; Cayenne).

  • — Laporte, 1840: 177 (Staphylinus; characters; Cayenne).

  • — Kraatz, 1857d: 539, 540 (Xanthopygus).

  • — Solsky, 1872: 306 (Xanthopygus, cited as Xanthopygius; Cayenne).

  • — Sharp, 1876: 132 (Xanthopygus; characters of male; Brazil).

  • — Sharp, 1884: 347 (Lampropygus; [Note: Sharp referred to the species as Philonthus bicolor Erichson; the species to which Erichson referred was that of Laporte).

  • — Bernhauer, 1906b: 196 (Xenopygus).

  • — Bernhauer and Schubert, 1914: 405 (Xenopygus; catalog).

  • — Irmler, 1979a: 31 (Xenopygus; characters).

  • — Irmler, 1982a: 209 (Xenopygus; neotype designation: Cayenne; characters).

  • — Herman, 2001: 54 (Xenopygus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: Guyana, Brazil.

confusus Irmler, 1982a: 208 (Xenopygus; Type locality: Guyane, France Nouveau Chantier).

  • Distribution: Guyana.

cordovensis Bernhauer, 1910b: 377 (Xenopygus; Type locality: Mexiko: Cordoba).

  • — Bernhauer and Schubert, 1914: 405 (Xenopygus; catalog).

  • — Irmler, 1979a: 31 (Xenopygus; characters).

  • — Irmler, 1982a: 209 (Xenopygus; characters; Mexico).

  • Distribution: Mexico.

peruvianus Solsky, 1872: 306 (Xanthopygus, cited as Xanthopygius; Type locality: Monte Rico).

  • — Bernhauer and Schubert, 1914: 406 (Lampropygus; catalog).

  • — Scheerpeltz, 1933: 1416 (Xenopygus; catalog).

  • — Irmler, 1979a: 31 (Xenopygus; characters).

  • Distribution: Peru.

STAPHYLININI: SUBTRIBE INCERTAE SEDIS
Bombylodes [1 species; Oriental Region]

Bombylodes Fauvel, 1904: 43 (replacement name for Bombylius Fauvel). Type species: Bombylius mimeticus Fauvel, fixed by objective synonymy with Bombylius.

  • — Bernhauer and Schubert, 1914: 395 (world catalog; 1 species).

  • — Scheerpeltz, 1933: 1410 (world catalog supplement).

  • — Blackwelder, 1952: 83 (type species).

  • — Hammond, 1984: 205 (checklist of species of Borneo).

Bombylius Fauvel, 1902a: 42 (species included: mimeticus; preoccupied by Linné, 1758). Type species: Bombylius mimeticus Fauvel, fixed by original designation and monotypy.

  • — Bernhauer and Schubert, 1914: 395 (synonym of Bombylodes).

  • — Blackwelder, 1952: 83 (synonym of Bombylodes; type species: mimeticus).

mimeticus Fauvel, 1902a: 42 (Bombylius; Type locality: Sumatra).

  • — Bernhauer and Schubert, 1914: 395 (Bombylodes; catalog).

  • — Scheerpeltz, 1933: 1410 (Bombylodes; catalog; Borneo; Java).

  • Distribution: Borneo, Indonesia.

Descarpentriesiellus [6 species; Madagascan Region]

Descarpentriesiellus Jarrige, 1978: 291 (species included: mirandus, vicinus, cylindricus, frater, strictulus, forticornis). Type species: Descarpentriesiellus mirandus Jarrige, fixed by original designation.

cylindricus Jarrige, 1978: 293 (Descarpentriesiellus; Type locality: Andringitra Est, Marositry Est rivière Antsifotra, 2000 m).

  • Distribution: Madagascar.

forticornis Jarrige, 1978: 295 (Descarpentriesiellus; Type locality: Andringitra Centre, plateau Andohariana, 2000–2100 m).

  • Distribution: Madagascar.

frater Jarrige, 1978: 294 (Descarpentriesiellus; Type locality: Andringitra Est, forêt d'Amindramiova, 1950 m).

  • Distribution: Madagascar.

mirandus Jarrige, 1978: 292 (Descarpentriesiellus; Type locality: Andringitra Est, Anjavidilava, 2000 m).

  • Distribution: Madagascar.

strictulus Jarrige, 1978: 294 (Descarpentriesiellus; Type locality: Andringitra Est, Anjavidilava, 2000 m).

  • Distribution: Madagascar.

vicinus Jarrige, 1978: 293 (Descarpentriesiellus; Type locality: Andringitra Centre, plateau Andohariana, 2000–2100 m).

  • Distribution: Madagascar.

Paradiatrechus [1 species; Madagascan Region]

Paradiatrechus Cameron, 1944d: 315 (species included: niger). Type species: Paradiatrechus niger Cameron, fixed by monotypy.

  • — Blackwelder, 1952: 289 (type species: niger).

niger Cameron, 1944d: 316 (Paradiatrechus; Type locality: S.E. Madagascar: Midongy).

  • Distribution: Madagascar.

Tolmerinus [22 species; Palaearctic, Ethiopian, and Oriental Region]

Tolmerinus Bernhauer, 1923c: 63 (replacement name for Tolmerus Bernhauer). Type species: Anisolinus rufipennis Fauvel, fixed by objective synonymy with Tolmerus.

  • — Cameron, 1932: 163 (characters; key to species of British India).

  • — Scheerpeltz, 1933: 1375 (world catalog supplement).

  • — Blackwelder, 1952: 391 (type species).

  • — Coiffait, 1982a: 262 (key to species of Himalayan region).

Tolmerus Bernhauer, 1911: 89 (species included: rufipennis; preoccupied by Loew, 1849, foerster, 1888, Heine, 1890, and Fairmaire, 1900). Type species: Anisolinus rufipennis Fauvel, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 366 (world catalog; 2 species).

  • — Winkler, 1925: 381 (catalog for Palaearctic region).

  • — Cameron, 1925: 64 (catalog of Indian species).

  • — Cameron, 1932: 163 (synonym of Tolmerinus).

  • — Scheerpeltz, 1933: 1375 (synonym of Tolmerinus).

  • — Blackwelder, 1952: 390 (type species: rufipennis).

  • — Tikhomirova, 1973a: 186 (checklist of species of USSR).

abdominalis Cameron, 1926b: 361 (Tolmerinus; Type locality: Dehra Dun).

  • — Cameron, 1932: 165 (Tolmerinus; characters; India).

  • — Scheerpeltz, 1933: 1375 (Tolmerinus; catalog).

  • — Coiffait, 1982a: 263 (Tolmerinus; characters).

  • Distribution: India.

angusticeps Bernhauer, 1937b: 616 (Tolmerinus; Type locality: Ukerewe Insel im Victoria Nyanza: Muruntuguru).

  • Distribution: Tanzania.

auronotatus Fauvel, 1895: 270 (Anisolinus; Type locality: Bírmanie, Carin Cheba, 900–1100 m).

  • — Bernhauer and Schubert, 1914: 365 (Anisolinus; catalog).

  • — Cameron, 1932: 167 (Tolmerinus; characters; Burma).

  • — Scheerpeltz, 1933: 1375 (Tolmerinus; catalog).

  • — Coiffait, 1982a: 262 (Tolmerinus; characters).

  • Distribution: Myanmar.

baronii Coiffait, 1982a: 257 (Tolmerinus; Type locality: Inde: Meghalaya, Upper Shillong, 1900 m).

  • Distribution: India.

brevipennis Bernhauer, 1915h: 231 (Tolmerus; Type locality: Java: Preanger).

  • — Scheerpeltz, 1933: 1375 (Tolmerinus; catalog).

  • Distribution: Indonesia.

burgeoni Bernhauer, 1932b: 154 (Tolmerinus; Type locality: Haut-Uele: Yebo Moto; Moto).

  • — Scheerpeltz, 1933: 1375 (Tolmerinus; catalog).

  • Distribution: Congo.

cameroni Scheerpeltz, 1933: 1375 (Tolmerinus; replacement name for ruficollis Cameron).

  • Distribution: India.

ruficollis Cameron, 1926b: 364 (Tolmerinus; [preoccupied]; Type locality: Dehra Dun).

  • — Scheerpeltz, 1933: 1375 (Tolmerinus; synonym of cameroni).

conradsi Bernhauer, 1937b: 616 (Tolmerinus; Type locality: Ukerewe Insel im Victoria-Nyanzasee, Muruntuguru).

  • Distribution: Tanzania.

curtipennis Coiffait, 1982a: 258 (Tolmerinus; Type locality: Nepal: Mali bei Jiri).

  • Distribution: Nepal.

fasciiventris Fauvel, 1895: 270 (Anisolinus; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Bhamo; Teinzo; Pegou, Palon).

  • — Fauvel, 1904a: 60 (Anisolinus; Bomanhalli; Bírmanie).

  • — Bernhauer and Schubert, 1914: 365 (Anisolinus; catalog).

  • — Cameron, 1932: 166 (Tolmerinus; characters; Burma).

  • — Scheerpeltz, 1933: 1375 (Tolmerinus; catalog).

  • — Coiffait, 1982a: 263 (Tolmerinus; characters).

  • Distribution: Myanmar.

funestus Bernhauer, 1916e: 31 (Tolmerus; Type locality: Kiautschau).

  • — Scheerpeltz, 1933: 1375 (Tolmerinus; catalog).

  • Distribution: China.

graueri Bernhauer, 1913b: 231 (Diatrechus; Type locality: Westliches Tanganjika-Gebiet).

  • — Bernhauer and Schubert, 1914: 365 (Diatrechus; catalog).

  • — Bernhauer and Chapman, 1939: 72 (Tolmerinus; Kenya).

  • Distribution: Tanzania, Kenya.

inornatus Cameron, 1926b: 362 (Tolmerinus; Type locality: Mussorie district; Arni Gad).

  • — Cameron, 1932: 165 (Tolmerinus; characters; India).

  • — Scheerpeltz, 1933: 1375 (Tolmerinus; catalog).

  • — Coiffait, 1982a: 262 (Tolmerinus; characters).

  • Distribution: India.

obscurus Cameron, 1932: 166 (Tolmerinus; Type locality: Manipur).

  • — Scheerpeltz, 1933: 1375 (Tolmerinus; catalog).

  • — Coiffait, 1982a: 263 (Tolmerinus; characters).

  • Distribution: India.

opacus Bernhauer, 1912f: 206 (Tolmerus; Type locality: Südost Kamerun: Lolodorf).

  • — Bernhauer and Schubert, 1914: 366 (Tolmerus; catalog).

  • Distribution: Cameroon.

ruficollis Cameron, 1926, see: cameroni Scheerpeltz, 1933.

ruficollis Schubert, 1908: 615 (Philonthus; Type locality: Kulu, Himalaya).

  • — Bernhauer and Schubert, 1914: 353 (Philonthus; catalog).

  • — Cameron, 1932: 169 (Tolmerinus; characters; India).

  • — Scheerpeltz, 1933: 1375 (Tolmerinus; catalog).

  • — Coiffait, 1982a: 263 (Tolmerinus; characters).

  • Distribution: India.

rufipennis Fauvel, 1895: 271 (Anisolinus; Type locality: Bírmanie, Rangon; Teinzo; Bhamo; Shwego-myo; Cochinchine, Long-Xuyen).

  • — Bernhauer, 1911: 89 (Tolmerus).

  • — Bernhauer and Schubert, 1914: 366 (Tolmerus; catalog).

  • — Cameron, 1932: 168 (Tolmerinus; characters; Burma).

  • — Scheerpeltz, 1933: 1375 (Tolmerinus; catalog).

  • — Coiffait, 1982a: 263 (Tolmerinus; characters).

  • Distribution: Myanmar, Vietnam.

rufipes Cameron, 1926b: 363 (Tolmerinus; Type locality: Mussorie district; Arni Gad).

  • — Cameron, 1932: 168 (Tolmerinus; characters; India).

  • — Scheerpeltz, 1933: 1375 (Tolmerinus; catalog).

  • — Coiffait, 1982a: 263 (Tolmerinus; characters).

  • Distribution: India.

simulans Cameron, 1932: 169 (Tolmerinus; Type locality: Assam: Patkai Mts.).

  • — Scheerpeltz, 1933: 1375 (Tolmerinus; catalog).

  • — Coiffait, 1982a: 263 (Tolmerinus; characters).

  • Distribution: India.

trifasciatus Cameron, 1926b: 363 (Tolmerinus; Type locality: Not cited, presumably India).

  • — Cameron, 1932: 166 (Tolmerinus; characters).

  • — Scheerpeltz, 1933: 1375 (Tolmerinus; catalog; East Indies).

  • — Coiffait, 1982a: 262 (Tolmerinus; characters).

  • Distribution: India.

villiersi Cameron, 1949a: 321 (Tolmerinus; Type locality: Côte d'Ivoire: Mt. Tonkoui, alt. 900–1200 m. Also cited from Guinea).

  • Distribution: Ivory Coast, Guinea.

wittmeri Coiffait, 1982a: 260 (Tolmerinus; Type locality: Nepal: Mure. Also cited from India).

  • Distribution: Nepal, India.

Turgiditarsus [3 species; Palaearctic and Oriental Regions]

Turgiditarsus Schillhammer, 1997a: 109 (replacement name for Tumiditarsus). Type species: Tumiditarsus ledangensis Schillhammer, fixed by objective synonymy with Tumiditarsus.

Tumiditarsus Schillhammer, 1996: 63 (species included: ledangensis, chinensis). Type species: Tumiditarsus ledangensis Schillhammer, fixed by original designation.

chinensis Schillhammer, 1996: 65 (Tumiditarsus; Type locality: China: Zhejiang, Tienmushan).

  • — Schillhammer, 1997a: 109, 110 (Turgiditarsus; characters).

  • Distribution: China.

kodadai Schillhammer, 1997a: 109 (Turgiditarsus; Type locality: Malaysia: Sabah, Batu Punggul Resort env.).

  • Distribution: Malaysia.

ledangensis Schillhammer, 1996: 64 (Tumiditarsus; Type locality: Malaysia: Prov. Johor, Gg. Ledang, Hutan Lipur, 200 m).

  • — Schillhammer, 1997a: 109, 110 (Turgiditarsus; characters).

  • Distribution: Malaysia.

Tribe Xantholinini

Xantholinini Erichson, 1839b: 28, 291 (genera included: Platyprosopus, Othius, Holisus, Diochus, Sterculia, Scytalinus, Xantholinus, Leptacinus). Type genus: Xantholinus (see Newton and Thayer, 1992: 66).

  • — Thomson, 1858: 30 (characters; key to genera).

  • — Casey, 1906: 356 (characters; key to subtribes and genera).

  • — Reitter, 1908: 102, 108 (characters; key to genera).

  • — Eichelbaum, 1909: 160 (list of genera with references; generic synonyms; number of genera; number of species in each genus; general distribution of genera).

  • — Blatchley, 1910: 394 (characters; key to Indiana genera).

  • — Bernhauer and Schubert, 1914: 289 (world catalog).

  • — Šulc, 1922 (osmeterium; anatomy; histology; physiology).

  • — Porta, 1926: 86 (characters; species of Italy).

  • — Portevin, 1929: 369 (characters; key to French genera).

  • — Cameron, 1932: 2 (characters; characters in key; key to genera of British India).

  • — Joy, 1932: 126 (key to British genera).

  • — Scheerpeltz, 1933: 1297 (world catalog supplement).

  • — Scheerpeltz, 1940: 39 (key to Palaearctic genera).

  • — Blackwelder, 1943: 473 (key to genera of West Indies).

  • — Blackwelder, 1944: 145 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Cameron, 1945d: 782 (key to New Zealand genera).

  • — Jarrige, 1952a: 131 (characters).

  • — V. Hansen, 1952: 6 (characters; key to Danish genera).

  • — Coiffait, 1956a: 37, 43 (review of French taxa; characters; key to genera).

  • — Smetana, 1958: 81 (characters; key to Czechoslovakian genera and species).

  • — Moore, 1963a: 94 (key to Nearctic genera).

  • — Arnett, 1963: 274 (characters; list of North American genera).

  • — Palm, 1963b: 40 (characters; key to species of Sweden).

  • — Lohse, 1964: 156 (characters; key to central European genera and species).

  • — Pototskaia, 1967: 46 (larval characters).

  • — Saiz, 1968a: 94 (list of Chilean species).

  • — Scheerpeltz, 1968a: 58 (catalog of Austrian species).

  • — Kasule, 1970: 56 (larval characters).

  • — Coiffait, 1972: 127, 128 (characters; key to genera and species of western Palaearctic region).

  • — Smetana, 1982: 59 (characters; revision of and key to genera and species of North America north of Mexico).

  • — Bordoni, 1985a (anatomical studies of labrum, aedeagus, and male genital segment).

  • — Outerelo and Gamarra, 1985: 37 (key to genera of Iberian Peninsula).

  • — Costa, Vanin, and Neto, 1986: 101 (bioluminescence reported in larva of Xantholinus; Brazil).

  • — Newton and Thayer, 1992: 13, 66 (tribe of Staphylininae; discussion).

  • — Newton, 1995: 50, 51 (requested that the International Commission of Zoological Nomenclature give Xantholinini Erichson precedence over Agrodini Nordmann and Gyrohypnini Kirby).

  • — Klimaszewski, Newton, and Thayer, 1996: 150 (list of New Zealand genera; number of New Zealand species).

  • — M. Hansen, 1996: 108 (list of Danish species).

  • — ICZN, 1996: 216 (Xantholinini is placed on Official List of Family group names and is given precedence over other names based on Xantholinus, Agrodini and other names based on Agrodes, and Gyrohypnini and other names based on Gyrohypnus).

  • — Downie and Arnett, 1996: 397 (characters; key to genera and species of Northeastern North America).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 328, 390 (key to Nearctic genera; characters; notes).

Xantholininae Erichson, 1839.

  • — Lohse, 1964: 156 (characters; key to central European genera and species).

  • — Moore, 1964a: 88 (characters; notes).

  • — Coiffait and Saiz, 1964: 510 (key to Chilean genera).

  • — Coiffait and Saiz, 1968: 345 (list of Chilean species).

  • — Scheerpeltz, 1968a: 58 (catalog of Austrian species).

  • — Saiz, 1970b: 382 (key to Chilean genera).

  • — Coiffait, 1972: 121 (adult and larval characters; key to tribes and genera of western Palaearctic region).

  • — Moore and Legner, 1974g: 556 (key to genera of America north of Mexico).

  • — Moore and Legner, 1974h: 473 (catalog of the taxonomy, biology, and ecology of developmental stages of species of America north of Mexico).

  • — Moore and Legner, 1975: 72 (catalog; North America north of Mexico).

  • — Topp, 1978: 307, 310 (larval characters; key to genera).

  • — Moore and Legner, 1979: 53 (characters; notes).

  • — Muona, 1979: 16 (list of Scandinavian species).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 5 (catalog; Poland).

  • — Smetana, 1982 (revision of and keys to tribes, genera, and species of North America north of Mexico; characters; historical review).

  • — Bordoni, 1982: 126–160 (key to some larval genera; adult morphology and characters; key to genera of Italy; distribution; ecology).

  • — Pajni and Kohli, 1982: 10 (comparative morphology of aedeagus).

  • — E. Matthews, 1982: 7, 24 (notes; key to genera of Australia).

  • — Tóth, 1983: 182 (list of species of the Hortobágy National Park, Hungary).

  • — Outerelo and Gamarra, 1985: 37 (key to genera of Iberian Peninsula).

  • — Tóth, 1985a: 86 (list of species of Bakony Mountains, Hungary).

  • — Dettner, 1987: 30 (defensive glands).

  • — Tóth, 1989 (characters; key to genera and species of Hungary).

  • — Gusarov, 1989 (list of species from the Crimea; ecological and habitat notes).

  • — Frank, 1991: 346 (larval characters).

  • — Dettner, 1993a (summary and discussion of defensive secretions and exocrine glands and their bearing on phylogeny).

  • — Lawrence and Newton, 1995: 830 (included in Staphylininae).

  • — Sparacio, 1995: 155 (characters; species of Sicily).

  • — M. Hansen, 1996: 37 (may be derived subgroup of Staphylininae).

  • — Downie and Arnett, 1996: 396 (characters; key to genera and species of Northeastern North America).

  • — Solodovnikov, 1998: 343 (list of species in northwest Caucasus; Russia).

Xantholinitae:

  • — Jeannel and Jarrige, 1949: 370 (characters; notes).

  • — Jarrige, 1952a: 131 (characters).

  • — Coiffait, 1956a (review of French taxa; characters).

Xantholini Erichson, 1839.

  • — Casey, 1906: 359, 364 (characters; key to genera).

  • — Muona, 1979: 16 (list of Scandinavian species).

Xantholinina Erichson, 1839.

  • — Jakobson, 1908: 447 (characters).

  • — Jakobson, 1909: 497 (key to genera; list of species of Russia and western Europe).

  • — Johansen, 1914: 426 (characters; key to Danish genera).

Gyrohypnidae Kirby, 1837: 88 (genera included: Gyrohypnus). Type genus: Gyrohypnus (see Newton and Thayer, 1992: 66).

  • — Newton and Thayer, 1992: 66 (synonym of Xantholinini).

  • — Newton, 1995: 51 (requested that the International Commission of Zoological Nomenclature place Gyrohypnini on the Official List of Family-Group Names in Zoology and that it and other family-group names based on Gyrohypnus not be given priority over those based on Xantholinus).

  • — ICZN, 1996: 216 (Gyrohypnini placed on Official List of Family Group Names in Zoology; it and other names based on Gyrohypnus suppressed in favor Xantholinini and other names based on Xantholinus).

Gyrohypninae:

  • — Ádám, 1996: 243 (list of species collected in Bükk National Park, Hungary).

Agraeformes Nordmann, 1837: 7 (genera included: Agrodes, Araeocnemus). Type genus: Agrodes (see Newton and Thayer, 1992:).

  • — Newton and Thayer, 1992: 66 (synonym of Xantholinini).

  • — Newton, 1995: 48, 51 (name emended to Agrodini).

Agrodini Nordmann, 1837.

  • — Newton, 1995: 48, 51 (name emended from Agraeformes; requested that the International Commission of Zoological Nomenclature place Agrodini on the Official List of Family-Group Names in Zoology and that it and other family-group names based on Agrodes not be given priority over Xantholinini Erichson and other family group names based on Xantholinus).

  • — ICZN, 1996: 216 (Agrodini placed on Official List of Family Group Names in Zoology; it and other names based on Agrodes suppressed in favor Xantholinini and other names based on Xantholinus).

Metoponci Casey, 1906: 359, 360 (characters; notes; genera included: Metoponcus). Type genus: Metoponcus (see Newton and Thayer, 1992: 66).

  • — Smetana, 1982: 59 (synonym of Xantholinini).

  • — Newton and Thayer, 1992: 66 (synonym of Xantholinini).

Metoponcina Casey, 1906.

  • — Jakobson, 1909: 497 (characters).

  • — Smetana, 1982: 59 (synonym of Xantholinini).

Araeocnemes Casey, 1906: 359 (notes; genera included: Sterculia, Araeocnemis, Agrodes, Plochionocerus). Type genus: Araeocnemus (see Newton and Thayer, 1992: 66).

  • — Newton and Thayer, 1992: 66 (synonym of Xantholinini).

Gyrohypnini Hatch, 1957: 172, 233 (characters; key to genera of Pacific Northwest; genera included: Megalinus, Xestolinus, Hyponygrus, Gyrohypnus, Gauropterus, Nudobius, Leptacinus, Leptacinoides, Stictolinus, Hesperolinus). Type genus: Gyrohypnus (see Newton and Thayer, 1992: 66).

  • — Newton and Thayer, 1992: 66 (synonym of Xantholinini).

Agerodes [43 species; Neotropical Region]

Agerodes Motschulsky, 1858a: 208 (species included: coeruleus). Type species: Agerodes coeruleus Motschulsky, fixed by monotypy.

  • — Gemminger and Harold, 1868: 599 (catalog).

  • — Fauvel, 1869: 489 (synonym of Xantholinus).

  • — Fauvel, 1873b: 95 [= 1873a: 381] (synonym of Xantholinus).

  • — Bernhauer and Schubert, 1914: 312 (world catalog; 28 species).

  • — Bruch, 1928a: 427 (catalog; Argentina).

  • — Scheerpeltz, 1933: 1317 (world catalog supplement).

  • — Blackwelder, 1944: 148 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 42 (type species: coeruleus).

Hymeneus Sharp, 1885: 487 (species included: impressifrons, temporalis, pygialis, godmani, splendens, divisus, cephalotes, amethystinus, laeticulus). Type species: Hymeneus godmani Sharp, fixed by subsequent designation by Blackwelder, 1952: 194.

  • — Bernhauer and Schubert, 1914: 312 (synonym of Agerodes).

  • — Blackwelder, 1944: 148 (synonym of Agerodes).

  • — Blackwelder, 1952: 194 (synonym of Agerodes; type species).

aluticeps Bernhauer, 1927d: 157 (Agerodes; Type locality: Kolumbien).

  • — Scheerpeltz, 1933: 1317 (Agerodes; catalog).

  • Distribution: Colombia.

alutipennis Bernhauer, 1916f: 181 (Agerodes; Type locality: Costa Rica: Cartago, 1000 m).

  • — Scheerpeltz, 1933: 1317 (Agerodes; catalog).

  • Distribution: Costa Rica.

amethystinus Sharp, 1885: 489 (Hymeneus; Type locality: Mexico, Cordova, Jalapa).

  • — Fauvel, 1891: 105 (Agerodes; Venezuela).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • Distribution: Mexico, Venezuela.

capitalis Bernhauer, 1904, see: insignis Erichson, 1839.

cephalotes Sharp, 1885: 488 (Hymeneus; Type locality: Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • Distribution: Nicaragua.

coelestinus Bernhauer, 1906c: 324 (Agerodes; Type locality: Venezuela).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • — Scheerpeltz, 1933: 1317 (Agerodes; catalog).

  • — Marcuzzi, 1977: 11 (Agerodes; Venezuela).

  • Distribution: Venezuela.

coelestinus Bernhauer, 1911, see: divinus Bernhauer and Schubert, 1914.

coeruleus Motschulsky, 1858a: 209 (Agerodes; Type locality: Colombie).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog; Bolivia; Peru).

  • Distribution: Bolivia, Peru, Colombia.

columbicus Bernhauer, 1916f: 180 (Agerodes; Type locality: bei Pacho in den Ostkordilleren, 2000 m, im westlichen Kolumbien).

  • — Scheerpeltz, 1933: 1317 (Agerodes; catalog).

  • Distribution: Colombia.

costaricensis Wendeler, 1930: 190 (Agerodes; Type locality: Costa Rica, Turrialba, 800 m).

  • — Scheerpeltz, 1933: 1317 (Agerodes; catalog).

  • Distribution: Costa Rica.

denticulatus Bernhauer, 1904a: 228 (Agerodes; Type locality: Brasilia, S. Catharina, Rio Capivary).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • Distribution: Brazil.

minutus Schubert, 1911: 19 (Agerodes; Type locality: Espirito Santo, Brasilia).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; synonym of denticulatus).

divinus Bernhauer and Schubert, 1914: 312 (Agerodes; replacement name for coelestinus Bernhauer).

  • Distribution: Brazil.

coelestinus Bernhauer, 1911a: 413 (Agerodes; [preoccupied]; Type locality: Brasilien, Rio de Janeiro, Serra Macahé).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; synonym of divinus).

divisus Sharp, 1885: 488 (Hymeneus; Type locality: Panama, Volcan de Chiriqui 2000 to 3000 feet).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • Distribution: Panama.

dorsalis Schubert, 1911: 21 (Agerodes; Type locality: Yungas de la Paz, Bolivia).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • Distribution: Bolivia.

elegans Bernhauer, 1916f: 183 (Agerodes; Type locality: westliches Kolumbien, Umgebung von Cali am Cauca: St. Antonio, 2000 m; Rio Vitaco, 2200 m; Alto de las cruces, 2200 m; Villa Elvira, 1800 m).

  • — Scheerpeltz, 1933: 1317 (Agerodes; catalog).

  • Distribution: Colombia.

festivus Erichson, 1839b: 313 (Xantholinus; Type locality: Brasilia).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • Distribution: Brazil.

flavipennis Bernhauer, 1916f: 184 (Agerodes; Type locality: Brasilien, S. Paulo; Paraguay, Hohenau).

  • — Scheerpeltz, 1933: 1317 (Agerodes; catalog).

  • Distribution: Brazil, Paraguay.

frater Bernhauer, 1904a: 229 (Agerodes; Type locality: Brasilia, Espirito Santo).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • Distribution: Brazil.

germaini Bernhauer, 1904a: 225 (Agerodes; Type locality: Bolivien, Chaco).

  • — Schubert, 1911: 17 (Agerodes; [Note: Cited as a “germaini Fvl. i. l. n. sp.” then in a footnote said to be identical with Agerodes germaini Bernhauer so Schubert's name is unavailable (Article 11.6)]; Bolivia).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • Distribution: Bolivia.

gloriosus Bernhauer, 1916f: 182 (Agerodes; Type locality: Ostkordilleren, Pacho im westlichen Kolumbien, 2000 m).

  • — Scheerpeltz, 1933: 1317 (Agerodes; catalog).

  • Distribution: Colombia.

godmani Sharp, 1885: 487 (Hymeneus; Type locality: Costa Rica, Irazu 6000 to 7000 feet; Panama, Volcan de Chiriqui up to 6000 feet, Bugaba).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • Distribution: Costa Rica, Panama.

grandiceps Bernhauer, 1927d: 161 (Agerodes; Type locality: Brazilien: Itatiaya, Est. do Rio, 700 m).

  • — Scheerpeltz, 1933: 1317 (Agerodes; catalog).

  • Distribution: Brazil.

impressifrons Guérin-Méneville, 1844: 11 (Xantholinus; Type locality: Nouvelle Granade).

  • — Sharp, 1885: 487 (Hymeneus).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • Distribution: Colombia.

insignis Erichson, 1839b: 312 (Xantholinus; Type locality: Brasilia).

  • — Blanchard, 1842: 84 (Xantholinus; Brazil).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • — Scheerpeltz, 1933: 1317 (Agerodes; catalog).

  • Distribution: Brazil.

capitalis Bernhauer, 1904a: 227 (Agerodes; Type locality: Brasilia, Espirito Santo).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; synonym of insignis).

  • — Scheerpeltz, 1933: 1317 (Agerodes; synonym of insignis).

janthinus Erichson, 1847, see: Plochionocerus.

kraatzii Kirsch, 1865: 44 (Xantholinus; Type locality: Not cited but presumably Bogota).

  • — Solsky, 1875: 21 (Xantholinus; cited as kraatzi; Peru).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; cited as kraatzi; catalog; Brazil; Bolivia; Peru).

  • Distribution: Bolivia, Peru, Brazil, Colombia.

laeticulus Sharp, 1885: 489 (Hymeneus; Type locality: Panama, Volcan de Chiriqui below 4000 feet).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • Distribution: Panama.

lucidus Erichson, 1839b: 313 (Xantholinus; Type locality: Brasilia).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • — Scheerpeltz, 1933: 1317 (Agerodes; catalog).

  • Distribution: Brazil.

quadriceps Bernhauer, 1904a: 229 (Agerodes; Type locality: Brasilia, Espirto Santo).

  • — Bernhauer, 1908d: 325 (Agerodes; synonym of lucidus).

  • — Schubert, 1911: 15 (Xantholinus; [Note: Cited as “quadriceps Fvl. i. l. n. sp.” then in a footnote said to be identical with Agerodes quadriceps Bernhauer so Schubert's name is unavailable (Article 11.6)]; Brazil).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; synonym of lucidus).

  • — Scheerpeltz, 1933: 1317 (Agerodes; synonym of lucidus).

magniceps Bernhauer, 1927d: 159 (Agerodes; Type locality: Brasilien: S. Catharina, Rio Capivary).

  • — Scheerpeltz, 1933: 1317 (Agerodes; catalog).

  • Distribution: Brazil.

minutus Schubert, 1911, see: denticulatus Bernhauer, 1904.

montanus Bernhauer, 1927d: 160 (Agerodes; Type locality: Brasilien: Virginia, S. Minas Geraes, Faz Campos, 1500 m).

  • — Scheerpeltz, 1933: 1317 (Agerodes; catalog).

  • Distribution: Brazil.

nobilis Bernhauer, 1916f: 182 (Agerodes; Type locality: Peru).

  • — Scheerpeltz, 1933: 1318 (Agerodes; catalog).

  • Distribution: Peru.

opaciceps Bernhauer, 1927d: 158 (Agerodes; Type locality: Costa Rica: Orosi).

  • — Scheerpeltz, 1933: 1318 (Agerodes; catalog).

  • Distribution: Costa Rica.

pulcher Bernhauer, 1904a: 225 (Agerodes; Type locality: Ecuador, Santa Inez).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • Distribution: Ecuador.

puncticollis Bernhauer, 1906c: 326 (Agerodes; Type locality: Peru: Chanchamayo).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • — Scheerpeltz, 1933: 1318 (Agerodes; catalog).

  • Distribution: Peru.

punctipennis Bernhauer, 1906c: 325 (Agerodes; Type locality: Peru: Chanchamayo).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog; Venezuela).

  • — Scheerpeltz, 1933: 1318 (Agerodes; catalog).

  • Distribution: Venezuela, Peru.

pygialis Sharp, 1876: 200 (Xantholinus; Type locality: Ega).

  • — Sharp, 1885: 487 (Hymeneus).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • Distribution: Brazil.

quadriceps Bernhauer, 1904, see: lucidus Erichson, 1839.

semicyaneus Schubert, 1911: 18 (Agerodes; Type locality: Chanchamayo, Peru).

  • — Bernhauer and Schubert, 1914: 312 (Agerodes; catalog).

  • Distribution: Peru.

semicupreus Schubert, 1911, see: semiviolaceus Bernhauer, 1904.

semiviolaceus Bernhauer, 1904a: 226 (Agerodes; Type locality: Bolivia).

  • — Bernhauer and Schubert, 1914: 313 (Agerodes; catalog).

  • Distribution: Bolivia.

semicupreus Schubert, 1911: 19 (Agerodes; Type locality: Yungas de la Paz, Bolivia).

  • — Bernhauer and Schubert, 1914: 313 (Agerodes; synonym of semiviolaceus).

simoni Fauvel, 1891: 105 (Agerodes; Type locality: Caracas).

  • — Bernhauer and Schubert, 1914: 313 (Agerodes; catalog).

  • — Marcuzzi, 1977: 11 (Agerodes; Venezuela).

  • Distribution: Venezuela.

speciosus Erichson, 1839b: 312 (Xantholinus; Type locality: Brasilia).

  • — Bernhauer and Schubert, 1914: 313 (Agerodes; catalog).

  • Distribution: Brazil.

splendens Sharp, 1885: 488 (Hymeneus; Type locality: Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1914: 313 (Agerodes; catalog).

  • Distribution: Nicaragua.

sublaevigatus Schubert, 1911: 20 (Agerodes; Type locality: Yungas de la Paz, Bolivia).

  • — Bernhauer and Schubert, 1914: 313 (Agerodes; catalog).

  • Distribution: Bolivia.

temporalis Sharp, 1876: 201 (Xantholinus; Type locality: Ega).

  • — Sharp, 1885: 487 (Hymeneus).

  • — Bernhauer and Schubert, 1914: 313 (Agerodes; catalog).

  • Distribution: Brazil.

tropicus Wendeler, 1930: 191 (Agerodes; Type locality: Costa Rica, Turrialba, 800 m).

  • — Scheerpeltz, 1933: 1318 (Agerodes; catalog).

  • Distribution: Costa Rica.

turrialbanus Wendeler, 1930: 190 (Agerodes; Type locality: Costa Rica, Turrialba).

  • — Scheerpeltz, 1933: 1318 (Agerodes; catalog).

  • Distribution: Costa Rica.

zikani Bernhauer, 1927d: 159 (Agerodes; Type locality: Brasilien: Itatiaya, Est. da Rio, Maromba; Virginia, S. Minas Geraes, Faz Campos, 1500 m).

  • — Scheerpeltz, 1933: 1318 (Agerodes; catalog).

  • Distribution: Brazil.

Allolinus [6 species; Palaearctic Region]

Allolinus Coiffait, 1966f: 195 (subgenus of Leptacinus; species included: peliopterus, potockajae, laeviusculus). Type species: Xantholinus peliopterus Solsky, fixed by original designation.

  • — Smetana, 1967d: 196 (valid genus; characters; notes).

  • — Coiffait, 1972: 134, 137, 145 (subgenus of Leptacinus; characters; key to species).

  • — Smetana, 1975c: 168 (valid genus).

insolens Smetana, 1967d: 198 (Allolinus; Type locality: Mongolei: Chentej aimak, zw. Somon Zenchermandal u. Somon Zargaltchan, 1400 m).

  • — Coiffait, 1972: 159 (Leptacinus; subgenus Allolinus; characters; Mongolia).

  • Distribution: Mongolia.

laeviusculus Solsky, 1864: 442 (Leptacinus; Type locality: Sarepta, sur le bas Wolga).

  • — Fauvel, 1867c: 114 [= 1868b: 213] (Metoponcus).

  • — Reitter, 1908: 112 (Leptacinus; characters; southern Russia).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; catalog).

  • — Scheerpeltz, 1933: 1301 (Leptacinus; catalog).

  • — Coiffait, 1966f: 196 (Leptacinus; subgenus Allolinus).

  • — Coiffait, 1972: 160 (Leptacinus; subgenus Allolinus; characters; Russia).

  • — Herman, 2001: 83 (Allolinus).

  • Distribution: Russia.

ophtalmicus Coiffait, 1980: 264 (Leptacinus; replacement name for oculatus Coiffait).

  • — Herman, 2001: 83 (Allolinus).

  • Distribution: Bhutan.

oculatus Coiffait, 1977b: 215 (Leptacinus; subgenus Allolinus; [preoccupied]; Type locality: Bhutan: Samchi, 300 m).

  • — Coiffait, 1980: 264 (Leptacinus; synonym of ophtalmicus).

peliopterus Solsky, 1875a: 272 (Xantholinus; Type locality: entre la ville d'Irkoutsk et le village Oussolié).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1313 (Xantholinus; catalog).

  • — Coiffait, 1966f: 195 (Leptacinus; subgenus Allolinus).

  • — Smetana, 1967d: 197 (Allolinus; characters; Mongolia).

  • — Coiffait, 1972: 158 (Leptacinus; subgenus Allolinus; characters; references; region of Lake Baikal).

  • — Smetana 1975c: 168 (Allolinus; Mongolia).

  • Distribution: Russia, Mongolia.

sublaevis Fauvel, 1875a: XXV [= 1875b: 227] (Xantholinus; Type locality: Sibérie, environs d'Irkutsk).

  • — Reitter, 1908: 117 (Xantholinus; subgenus Metacyclinus; characters; Siberia).

  • — Eppelsheim, 1894b: 12 (Xantholinus; synonym of peliopterus).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; synonym of peliopterus).

  • — Scheerpeltz, 1933: 1313 (Xantholinus; synonym of peliopterus).

potockajae Coiffait, 1966f: 196 (Leptacinus; subgenus Allolinus; Type locality: SSSR, Uzbekistan, Bukharskaia obl.).

  • — Coiffait, 1972: 159 (Leptacinus; subgenus Allolinus; characters; Uzbekistan).

  • — Herman, 2001: 83 (Allolinus).

  • Distribution: Uzbekistan.

vuldiformis Coiffait, 1977b: 214 (Leptacinus; subgenus Allolinus; Type locality: Bhutan: Samchi, 300 m).

  • — Herman, 2001: 83 (Allolinus).

  • Distribution: Bhutan.

Allotrichus [1 species; Neotropical Region]

Allotrichus Sharp, 1885: 486 (species included: arenarius). Type species: Allotrichus arenarius Sharp, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 309 (world catalog; 1 species).

  • — Blackwelder, 1944: 148 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 47 (type species: arenarius).

arenarius Sharp, 1885: 486 (Allotrichus; Type locality: Panama, Peña Blanca 3000 to 4000 feet).

  • Distribution: Panama.

Atopolinus [1 species; Palaearctic Region]

Atopolinus Coiffait, 1982a: 246 (species included: manhariensis). Type species: Atopolinus manhariensis Coiffait, fixed by original designation and monotypy.

manhariensis Coiffait, 1982a: 247 (Atopolinus; Type locality: Nepal: 5 km E Manhari, 350 m).

  • Distribution: Nepal.

Caecolinus [1 species; Palaearctic Region]

Caecolinus Jeannel, 1923: 337 (species included: endogaeus). Type species: Caecolinus endogaeus Jeannel, fixed by monotypy.

  • — Winkler, 1925: 373 (catalog for Palaearctic region).

  • — Scheerpeltz, 1933: 1321 (world catalog supplement).

  • — Blackwelder, 1952: 87 (type species: endogaeus).

  • — Coiffait, 1956a: 37 (note).

  • — Székessy, 1963: 51 (characters).

  • — Coiffait, 1972: 365 (characters; type species: endogaeus; key to species of western Palaearctic region; references).

  • — Tikhomirova, 1973a: 180 (checklist of species of USSR).

  • — Tóth, 1989: 41 (cited as Coecolinus; characters).

endogaeus Jeannel, 1923: 341 (Caecolinus; Type locality: vers 1100 m. d'altitude, sur le mont Detunata, comm. de Bucium, jud. Alba-inferioara, Roumanie).

  • — Scheerpeltz, 1933: 1321 (Caecolinus; catalog).

  • — Jeannel and Jarrige, 1949: 371 (Caecolinus; characters; cave species; Romania).

  • — Coiffait, 1972: 366 (Caecolinus; characters; references; Romania).

  • — Janák, 1987: 202 (Caecolinus; notes; Romania).

  • — Tóth, 1989: 41 (Caecolinus, cited as Coecolinus; characters).

  • Distribution: Romania.

Calontholinus [1 species; Palaearctic Region]

Calontholinus Reitter, 1908: 114 (subgenus of Xantholinus; species included: fasciatus). Type species: Xantholinus fasciatus Hochhuth, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 299 (subgenus of Xantholinus).

  • — Cameron, 1932: 22 (subgenus of Xantholinus).

  • — Scheerpeltz, 1933: 1307 (subgenus of Xantholinus).

  • — Blackwelder, 1943: 473 (subgenus of Xantholinus; type species: fasciatus).

  • — Steel, 1949a: 269 (either separated genus or subgenus of Nudobius, not subgenus of Xantholinus).

  • — Blackwelder, 1952: 92 (type species: fasciatus).

  • — Coiffait, 1972: 200 (characters; type species: fasciatus; species of western Palaearctic region).

  • — Bordoni, 1975: 90 (subgenus of Xantholinus; distributional notes).

fasciatus Hochhuth, 1849: 103 (Xantholinus; Type locality: Lenkoran; [Note: See lectotype designation by Gusarov, 1992a]).

  • — Reitter, 1908: 114 (Xantholinus; subgenus Calontholinus; characters; Caucasus).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1309 (Xantholinus; catalog).

  • — Jarrige, 1971: 493 (Nudobius; Iran).

  • — Coiffait, 1972: 200 (Calontholinus; characters; Caucasus).

  • — Bordoni, 1975: 78 (Xantholinus; subgenus Calontholinus; notes; Caspian region; Azerbaijan).

  • — Petrenko, 1980: 82 (Xantholinus; Caucasus).

  • — Gusarov, 1992a: 784 [= 1993a: 73] (Calontholinus; lectotype designation).

  • Distribution: Azerbaijan, Iran.

Crinolinus [1 species; Nearctic Region]

Crinolinus Smetana, 1982: 282 (species included: intonsus). Type species: Crinolinus intonsus Smetana, fixed by original designation and monotypy.

  • — Newton, Thayer, Ashe, and Chandler, 2000: 390 (1 Nearctic species; notes; characters in key).

intonsus Smetana, 1982: 282 (Crinolinus; Type locality: Washington: Tenino. Also cited from Nevada).

  • Distribution: USA.

Dactylaptatus [1 species; Madagascan Region]

Dactylaptatus Lecoq, 1990: 189 (species included: insularis). Type species: Dactylaptatus insularis Lecoq, fixed by original designation and monotypy.

insularis Lecoq, 1990: 189 (Dactylaptatus; Type locality: La Réunion).

  • Distribution: Réunion.

Dibothroglyptus [1 species; Oriental Region]

Dibothroglyptus Scheerpeltz, 1957c: 233, 310 (species included: sutteri). Type species: Dibothroglyptus sutteri Scheerpeltz, fixed by monotypy.

sutteri Scheerpeltz, 1957c: 233, 315 (Dibothroglyptus; Type locality: Flores: Rana Mese, 1300 m).

  • Distribution: Indonesia.

Ehomalolinus [4 species; Neotropical Region]

Ehomalolinus Bierig, 1934: 16 (species included: divisus, sanguineus, punctipennis, mordax). Type species: Ehomalolinus punctipennis Bierig, by original designation.

  • — Blackwelder, 1944: 148 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 145 (type species: punctipennis).

divisus Erichson, 1839b: 308 (Xantholinus; Type locality: Vera Cruce, Mexico).

  • — Sharp, 1885: 474 (Homalolinus; notes; Mexico).

  • — Bernhauer and Schubert, 1914: 309 (Homalolinus; catalog).

  • — Bierig, 1934: 17 (Ehomalolinus).

  • Distribution: Mexico, Guatemala.

guatemalenus Sharp, 1885: 474 (Homalolinus; variety of divisus; Type locality: Guatemala, San Geronimo, Balheu).

  • — Bernhauer and Schubert, 1914: 309 (Homalolinus; synonym of divisus).

  • — Scheerpeltz, 1953: 607 (Ehomalolinus; variety of divisus; Guatemala).

guatemalenus Sharp, 1885, see: divisus Erichson, 1839.

mordax Bierig, 1934: 20 (Ehomalolinus; Type locality: Panama: Volcan de Chiriqui, cerca de 1500 m).

  • Distribution: Panama.

punctipennis Bierig, 1934: 18 (Ehomalolinus; Type locality: Costa Rica: Vara Blanca, 2000 m, entre Barba y Poas).

  • Distribution: Costa Rica.

sanguineus Sharp, 1885: 474 (Homalolinus; Type locality: Guatemala, near the city, Calderas, Dueñas, Capetillo, San Geronimo 3000 feet, Cerro Zunil 4000 to 5000 feet).

  • — Bernhauer and Schubert, 1914: 310 (Homalolinus; catalog).

  • — Bierig, 1934: 17 (Ehomalolinus).

  • Distribution: Guatemala.

Eulissus [46 species; Neotropical, Palaearctic, Ethiopian, Madagascan, Oriental, and Australian Regions]

Eulissus Mannerheim, 1830: 8, 35 (species included: chalybaeus). Type species: Eulissus chalybaeus Mannerheim, fixed by monotypy.

  • — Mannerheim, 1831: 422, 449 (characters).

  • — Dejean, 1833: 63 (list of species).

  • — Nordmann, 1837: 129 (characters).

  • — Erichson, 1839b: 306 (synonym of Xantholinus).

  • — Laporte, 1840: 180 (characters).

  • — Lacordaire, 1854: 68 (synonym of Xantholinus).

  • — Kraatz, 1857d: 629 (synonym of Xantholinus).

  • — Gemminger and Harold, 1868: 600 (catalog).

  • — Fauvel, 1873b: 95 [= 1873a: 381] (synonym of Xantholinus).

  • — Lynch, 1884: 178 (characters; subgenus of Eulissus).

  • — Casey, 1906: 366, 379 (characters).

  • — Bernhauer and Schubert, 1914: 310 (world catalog; 27 species).

  • — Cameron, 1921a: 354, 404 (characters; key to Singapore species; catalog).

  • — Cameron, 1925: 52 (catalog of Indian species).

  • — Scheerpeltz, 1933: 1316 (world catalog supplement).

  • — Steel, 1938b (notes; characters to distinguish from Thyreocephalus).

  • — Blackwelder, 1944: 148 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 156 (type species: chalybaeus).

  • — Tottenham, 1956: 255 (key to some species).

Dinolinus Casey, 1906: 366, 373 (species included: chalybeus, rutilus). Type species: Eulissus chalybaeus Mannerheim, fixed by original designation.

  • — Bernhauer and Schubert, 1914: 310 (synonym of Eulissus).

  • — Blackwelder, 1952: 127 (synonym of Eulissus; type species: chalybaeus).

Xanthohypnus Casey, 1906: 367, 374 (species included: strigiceps). Type species: Xantholinus strigiceps Sharp, fixed by original designation and monotypy.

  • — Bernhauer and Schubert, 1914: 310 (synonym of Eulissus).

  • — Blackwelder, 1952: 404 (synonym of Eulissus; type species: strigiceps).

africanus Bernhauer, 1913b: 230 (Eulissus; Type locality: Zentralafrika: Ukaika – Mawambi, Urwald Moera; Kamerun).

  • — Bernhauer and Schubert, 1914: 310 (Eulissus; catalog).

  • — Tottenham, 1956: 255 (Eulissus; characters in key).

  • Distribution: “Cental Africa”, Cameroon.

alluaudi Fauvel, 1907: 34 (Xantholinus; Type locality: Nairobi; Kilimandjaro; Kiboscho; Kibonoto).

  • — Bernhauer and Schubert, 1914: 300 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1316 (Eulissus; catalog).

  • — Bernhauer and Chapman, 1939: 64 (Xantholinus; East Africa).

  • Distribution: Kenya, Tanzania.

amabilis Boheman, 1848: 274 (Xantholinus; Type locality: montes Makkalienses).

  • — Bernhauer and Schubert, 1914: 300 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1316 (Eulissus; catalog).

  • — Tottenham, 1956: 255 (Eulissus; characters in key).

  • Distribution: South Africa.

amicus Bernhauer, 1916c: 26 (Eulissus; Type locality: Peru: Chanchamayo und Callange).

  • — Scheerpeltz, 1933: 1316 (Eulissus; catalog).

  • Distribution: Peru.

analis Nordmann, 1837: 123 (Gyrohypnus; Type locality: Brasilia meridionalis).

  • — Erichson, 1839b: 314 (Xantholinus; characters; Brazil).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • Distribution: Brazil.

ater Laporte, 1835, see: Thyreocephalus.

atlanticus Bernhauer, 1915k: 298 (Eulissus; Type locality: Insel Fernando Poo; Basile, 400–600 m; Bahia de S. Carlos, 200–300 m).

  • — Scheerpeltz, 1933: 1317 (Eulissus; catalog).

  • Distribution: Equatorial Guinea.

badariottii Wasmann, 1900: 282 (Xantholinus; subgenus Eulissus; Type locality: Lorena, Staat S. Paulo, Brazilien).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • Distribution: Brazil.

brunneiventris Tottenham, 1956: 252 (Eulissus; Type locality: Ruanda: terr. Kibuye, Rubengera, 1900 m).

  • Distribution: Rwanda.

burgeoni Bernhauer, 1929f: 122 (Eulissus; Type locality: Haut-Uele: Kubango).

  • — Scheerpeltz, 1933: 1317 (Eulissus; catalog).

  • Distribution: Congo.

chalcopterus Erichson, 1839b: 312 (Xantholinus; Type locality: Nova Hollandia).

  • — Fauvel, 1877: 244 (Xantholinus; characters; Australia).

  • — Olliff, 1887: 486 (Xantholinus; characters; Australia).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • — Bernhauer, 1920: 15 (Eulissus; Australia).

  • Distribution: Australia.

cyaneipennis W. J. MacLeay, 1873: 139 (Xantholinus; Type locality: Gayndah).

  • — Olliff, 1887: 486 (Xantholinus; synonym of chalcopterus).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; synonym of chalcopterus).

  • — Lea, 1925: 219 (Eulissus; synonym of chalcopterus).

chalybaeus Mannerheim, 1830: 35 (Eulissus; Type locality: Minas Geraës Brasiliae).

  • — Mannerheim, 1831: 449 (Eulissus; Brazil).

  • — Nordmann, 1837: 129 (Eulissus; cited as chalibaeus; Brazil).

  • — Blanchard, 1842: 84 (Xantholinus; Bolivia; Brazil).

  • — Erichson, 1839b: 309 (Xantholinus; characters; Brazil).

  • — Laporte, 1840: 180 (Eulissus; characters; Brazil).

  • — P. Lucas, 1857: 49 (Xantholinus; Brazil).

  • — Redtenbacher, 1867: 28 (Xantholinus; Brazil).

  • — Solsky, 1868a: 141 (Xantholinus; Mexico).

  • — Solsky, 1875: 21 (Xantholinus; Peru).

  • — Fauvel, 1877a: xxvi (Xantholinus; cited as chalybeus; Brazil).

  • — Sharp, 1885: 476 (Xantholinus; Mexico, Guatemala, Nicaragua, Panama, Brazil).

  • — Casey, 1906: 373 (Dinolinus).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • — Scheerpeltz, 1933: 1317 (Eulissus; catalog).

  • — Schuh, Schillhammer, and Zettel, 1992: 220 (Eulissus; notes; introduced in Austria).

  • — O. Young, 1998 (Gauropterus [Note: The author cited this species in Gauropterus because he misinterpreted Smetana's (1982: 71) inclusion of Eppelsheim's misidentification of Eulissus in the synonymic list for Gauropterus. Eulissus has not been synonymized with Gauropterus nor has Eulissus chalybaeus been moved to Gauropterus]; predatory behavior; Panama).

  • — Smetana, 1999a: 55 (Eulissus; correction of error of interpretation by O. Young, 1998).

  • Distribution: Brazil, Bolivia, Peru, Colombia, Panama, Nicaragua, Guatemala, Mexico, Austria.

saphyreus Perty, 1830: 31 (Staphylinus; Type locality: Brasil: Prov. Minarum).

  • — Blanchard, 1842: 84 (Xantholinus; synonym of chalybaeus Mannerheim).

  • — Erichson, 1839b: 309 (Xantholinus; cited as saphyrinus; synonym of chalybaeus).

  • — Fauvel, 1877a: xxvi (Xantholinus; synonym of chalybaeus).

  • — Sharp, 1885: 476 (Xantholinus; synonym of chalybeus).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; cited as sapphyreus; synonym of chalybaeus).

chloropterus Erichson, 1839b: 311 (Xantholinus; Type locality: Terra Van Diemenii).

  • — Fauvel, 1877: 243 (Xantholinus; characters; Australia).

  • — Fauvel, 1878g: 539 (Xantholinus; characters; Australia).

  • — Fauvel, 1878g: 540 (Xantholinus; Australia).

  • — Olliff, 1887: 483 (Xantholinus; characters; Australia).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • — Cameron, 1945d: 786 (Indoscytalinus).

  • — Frank, 1982: 31 (Eulissus; list of parasites).

  • Distribution: Australia.

collarti Cameron, 1932b: 133 (Eulissus; Type locality: Blukwa, Nizi).

  • — Scheerpeltz, 1933: 1317 (Eulissus; catalog).

  • Distribution: Congo.

congoensis Bernhauer, 1932b: 144 (Eulissus; Type locality: Haut-Uele: Yebo Moto, Tora).

  • — Scheerpeltz, 1933: 1317 (Eulissus; catalog).

  • Distribution: Congo.

cyaneipennis W. J. MacLeay, 1873, see: chalcopterus Erichson, 1839.

cyanopterus Erichson, 1839b: 311 (Xantholinus; Type locality: Terra Van Diemenii).

  • — Fauvel, 1877: 243 (Xantholinus; characters; Australia).

  • — Fauvel, 1878g: 539 (Xantholinus; characters).

  • — Olliff, 1887: 485 (Xantholinus; characters; Australia).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • Distribution: Australia.

diabolicus Bernhauer, 1902, see: Gauropterus.

ferox Harold, 1881: 262 (Xantholinus; Type locality: Not cited but presumably from Ostafrika).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • Distribution: “East Africa”.

flavomarginatus Bernhauer, 1929f: 121 (Eulissus; Type locality: Haut-Uele: Watsa).

  • — Scheerpeltz, 1933: 1317 (Eulissus; catalog).

  • — Bernhauer, 1942c: 361 (Eulissus; Cameroon).

  • — Cameron, 1952e: 831 (Eulissus; Dahomey).

  • Distribution: Benin, Cameroon, Congo.

flaviventris Bernhauer, 1939i: 259 (Eulissus; Type locality: Kamerun: Ekona).

  • Distribution: Cameroon.

gerardi Bernhauer, 1929f: 121 (Eulissus; Type locality: Manyema: r. Kasa-Niemba).

  • — Scheerpeltz, 1933: 1317 (Eulissus; catalog).

  • — Tottenham, 1956: 255 (Eulissus; characters in key).

  • Distribution: Congo.

goudoti Fauvel, 1905b: 171 (Xantholinus; Type locality: Suberbieville).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • Distribution: Madagascar.

guineensis Bernhauer, 1912f: 183 (Eulissus; Type locality: Guinea).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • Distribution: Guinea.

haemorrhous Fauvel, 1878g: 538 (Xantholinus; Type locality: Australie orientale: Queensland, Rockhampton).

  • — Olliff, 1887: 482 (Xantholinus; characters; Australia).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • — Bernhauer, 1920: 15 (Eulissus; Australia).

  • — Lea, 1925: 219 (Eulissus; notes).

  • Distribution: Australia.

hummleri Bernhauer, 1908: 18 (Eulissus; Type locality: Queensland).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • — Lea, 1925: 219 (Eulissus; characters; Australia).

  • Distribution: Australia.

indicus Wendeler, 1930: 189 (Eulissus; Type locality: Annam, Phuc).

  • — Scheerpeltz, 1933: 1317 (Eulissus; catalog).

  • Distribution: Vietnam.

lateralis Cameron, 1918: 83 (Eulissus; Type locality: Singapore: Woodlands).

  • — Cameron, 1921a: 375, 404 (Eulissus; characters; Singapore).

  • — Scheerpeltz, 1933: 1317 (Eulissus; catalog).

  • Distribution: Singapore.

limbatus Klug, 1832, see: madagascarensis Steel, 1949.

madagascarensis Steel, 1949a: 270 (Xantholinus; replacement name for limbatus Klug).

  • Distribution: Madagascar.

limbatus Klug, 1832: 138 (Staphylinus; [preoccupied]; Type locality: Madagascar).

  • — Erichson, 1839b: 310 (Xantholinus; characters; Madagascar).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • — Steel, 1949a: 270 (Xantholinus; synonym of madascarensis).

  • — Scheerpeltz, 1961g: 238 (Eulissus; Madagascar).

mannerheimi Laporte, 1835, see: rutilus Perty, 1830.

methneri Bernhauer, 1937b: 610 (Eulissus; Type locality: Deutsch Ostafrika: Mtotshovu).

  • Distribution: Tanzania.

milliaui Bernhauer, 1932b: 144 (Eulissus; Type locality: Mongbwalu, Kilo).

  • — Scheerpeltz, 1933: 1317 (Eulissus; catalog).

  • Distribution: Congo.

mirabilis Jarrige, 1948a: 19 (Eulissus; Type locality: Madagascar: Bekily, région sud de l'île).

  • Distribution: Madagascar.

mokaensis Bernhauer, 1915k: 299 (Eulissus; Type locality: Insel Fernando Poo: Moka, 1300–1500 m).

  • — Scheerpeltz, 1933: 1317 (Eulissus; catalog).

  • Distribution: Equatorial Guinea.

oculatus Sharp, 1885: 477 (Xantholinus; Type locality: Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1317 (Eulissus; catalog).

  • Distribution: Nicaragua.

olliffi Lea, 1895: 589 (Xantholinus; Type locality: New South Wales: Tamworth, Peel River).

  • — Blackburn, 1902: 20 (Xantholinus; near phoenicopterus).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • — Scheerpeltz, 1933: 1317 (Eulissus; catalog).

  • Distribution: Australia.

phoenicopterus Erichson, 1839n: 314 (Xantholinus; Type locality: Nova Hollandia).

  • — Redtenbacher, 1867: 28 (Xantholinus; Australia).

  • — Fauvel, 1877: 242 (Xantholinus; cited as phaenicopterus; characters; Australia).

  • — Olliff, 1887: 483 (Xantholinus; characters; Australia).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • Distribution: Australia.

pilosus Roth, 1851: 118 (Xantholinus; Type locality: Abyssinien).

  • — Eppelsheim, 1895: 208 (Eulissus; Ganale Gudda, Arussi Galla).

  • — Bernhauer, 1912h: 478 (Eulissus; Ruwenzori region).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • — Cameron, 1950g: 33 (Thyreocephalus; Belgian Congo).

  • — Tottenham, 1956: 224, 255 (Eulissus; characters in key; Ruanda).

  • Distribution: Ethiopia, Congo, Rwanda.

proximalis Tottenham, 1956: 255 (Eulissus; Type locality: Ruanda: terr. Nyanza, Gitarama, 1850 m).

  • Distribution: Rwanda.

purpuripennis Bernhauer, 1904a: 231 (Eulissus; Type locality: Shanghai).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • Distribution: China.

riedeli Bernhauer, 1916f: 180 (Eulissus; Type locality: Brasilien: Santo Paulo, São Carlos do Pinhal).

  • — Scheerpeltz, 1933: 1317 (Eulissus; catalog).

  • Distribution: Brazil.

rutilus Perty, 1830: 30 (Staphylinus; Type locality: Brasil: Prov. Minarum).

  • — Erichson, 1839b: 318 (Xantholinus; characters; Brazil).

  • — Sharp, 1876: 199 (Xantholinus; characters; Brazil).

  • — Sharp, 1885: 476 (Xantholinus; Panama, Brazil).

  • — Casey, 1906: 373 (Dinolinus).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • Distribution: Peru, Brazil, French Guiana, Colombia, Panama.

mannerheimii Laporte, 1835: 118 (Eulissus; Type locality: Cayenne).

  • — Erichson, 1839b: 310 (Xantholinus; characters; Cayenne).

  • — Sharp, 1876: 199 (Xantholinus; Brazil).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog; Colombia; Peru).

  • — Steel, 1949a: 269 (Eulissus; synonym of rutilus).

sapphyreus Perty, 1830, see: chalybaeus Mannerheim, 1830.

secretus Bernhauer, 1935d: 101 (Eulissus; Type locality: Lulua: Kapanga).

  • Distribution: Congo.

semiflavus Bernhauer, 1913b: 231 (Eulissus; Type locality: Kongogebiet).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • — Cameron, 1950g: 33 (Thyreocephalus; Belgian Congo).

  • — Tottenham, 1956: 255 (Eulissus; characters in key).

  • Distribution: Congo.

semipurpureus Bernhauer, 1916c: 26 (Eulissus; Type locality: Panama: Lino, 800 m).

  • — Scheerpeltz, 1933: 1317 (Eulissus; catalog).

  • Distribution: Panama.

silvaticus Bernhauer, 1904a: 230 (Eulissus; Type locality: Madagaskar).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • Distribution: Madagascar.

spegazzinii Bernhauer, 1915, see: Thyreocephalus.

strigiceps Sharp, 1885: 476 (Xantholinus; Type locality: Nicaragua, Chontales).

  • — Casey, 1906: 374 (Xanthohypnus).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • Distribution: Nicaragua.

strinatii Scheerpeltz, 1958d: 826 (Eulissus [?]; Type locality: Grotte de Pahou, 2 km Ost-Süd-Ost von Lastoursville in Gabon).

  • — Scheerpeltz, 1971c: 169 (Eulissus; Cameroon; Nigeria; Gabon; Congo).

  • Distribution: Congo, Gabon, Cameroon, Nigeria.

turneri Bernhauer, 1937: 299 (Eulissus; Type locality: Kenya: Nairobi).

  • Distribution: Kenya.

vagus Erichson, 1839b: 315 (Xantholinus; Type locality: Caracas Columbiae).

  • — Fauvel, 1891: 105 (Xantholinus; Venezuela).

  • — Bernhauer and Schubert, 1914: 312 (Eulissus; catalog).

  • Distribution: Venezuela.

Gauropterus [30 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, and Oriental Regions]

Gauropterus Thomson, 1860: 187 (species included: fulgidus). Type species: Staphylinus fulgidus Fabricius, fixed by action of the Commission in Opinion 1250 (ICZN, 1983).

  • — Fauvel, 1873b: 95 [= 1873a: 381] (synonym of Xantholinus).

  • — Mulsant and Rey, 1877a: 252 (characters; key to French species).

  • — Heyden, Reitter, and Weise, 1891: 105 (cited as Eulissus; list of species of Europe and Caucasus).

  • — Heyden, Reitter, and Weise, 1906: 162 (list of species of Europe and Caucasus).

  • — Reitter, 1908: 109, 113 (characters; key to species).

  • — Reitter, 1909: 134 (characters).

  • — Petri, 1912: 70 (list of species of Siebenbürgen).

  • — Bernhauer and Schubert, 1914: 310 (world catalog; 12 species).

  • — Leng, 1920: 105 (catalog of North American species).

  • — Winkler, 1925: 372 (catalog for Palaearctic region).

  • — Cameron, 1925: 51 (catalog of Indian species).

  • — Porta, 1926: 87 (characters; species of Italy).

  • — Roubal, 1930: 364 (catalog; Slovakia).

  • — Cameron, 1932: 30 (characters; key to species of British India).

  • — Joy, 1932: 127 (key to British species).

  • — Scheerpeltz, 1933: 1315 (world catalog supplement).

  • — Normand, 1935: 369 (list of species of Tunisia).

  • — Kloet and Hincks, 1945: 165 (list of British species).

  • — Tottenham, 1949: 370, 417 (type species: fulgidus Fabricius, 1797; notes; checklist of British species).

  • — Blackwelder, 1952: 168 (type species: fulgidus).

  • — V. Hansen, 1952: 6, 20 (characters; Danish species).

  • — Coiffait, 1956a: 44, 57 (characters; notes).

  • — Kocher, 1958: 141 (checklist of species; Morocco).

  • — Smetana, 1958: 115 (characters; habitus of one species; type species: fulgidus; Czechoslovakian species).

  • — Smetana, 1959a: 209 (list of species collected in Albania).

  • — Ferreira, 1962a: 16 (list of Portuguese species).

  • — Székessy, 1963: 50 (characters).

  • — Palm, 1963b: 53 (characters; species of Sweden).

  • — Lohse, 1964: 160 (characters; key to central European species).

  • — Coiffait and Saiz, 1964: 511 (characters).

  • — Horion, 1965: 89 (list of central European species).

  • — Coiffait and Saiz, 1968: 345 (list of Chilean species).

  • — Scheerpeltz, 1968a: 63 (catalog of Austrian species).

  • — Saiz, 1970b: 383, 385 (characters).

  • — Coiffait, 1972: 164 (characters; type species: fulgidus; key to species of western Palaearctic region; references).

  • — Bordoni, 1973j: 692 (list of species from islands near Sicily).

  • — Tikhomirova, 1973a: 181 (checklist of species of USSR).

  • — Pope, 1977: 28 (list of British species).

  • — Muona, 1979: 16 (list of Scandinavian species).

  • — Smetana, 1979: 50 (requested that International Commission on Zoological Nomenclature place Gauropterus on the Official list of Generic Names in Zoology with Staphylinus fulgidus as type species).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 12 (catalog; Poland).

  • — Uhlig and Vogler, 1981: 107 (list of some species collected in Germany; habitat and collecting notes).

  • — Smetana, 1982: 71 (characters; habitus; revision of species of North America north of Mexico).

  • — Bordoni, 1982: 164 (characters; habitus; species of Italy).

  • — ICZN, 1983: 85 (Gauropterus placed on Official List of Generic Names in Zoology and Staphylinus fulgidus is fixed as the type species).

  • — Shibata, 1983: 68 (checklist of species of Japan).

  • — Tóth, 1989: 13 (characters; species of Hungary).

  • — Campbell and Davies, 1991: 123 (checklist of species; Canada).

  • — Silfverberg, 1992: 18 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 28 (additional records to Danish list of species).

  • — Ciceroni and Zanetti, 1995: 25 (list of species of Italy).

  • — M. Hansen, 1996: 108 (list of Danish species).

  • — Downie and Arnett, 1996: 398 (characters).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 390 (1 Nearctic species; notes; characters in key).

abactus Tottenham, 1939c: 221 (Gauropterus; Type locality: S. Rhodesia: Mt. Selinda).

  • Distribution: Zimbabwe.

adjacens Tottenham, 1939c: 222 (Gauropterus; Type locality: S. Rhodesia: Sanyati Valley).

  • Distribution: Zimbabwe.

albocinctus Fauvel, 1905b: 172 (Xantholinus; Type locality: Suberbieville).

  • — Bernhauer and Schubert, 1914: 310 (Gauropterus; catalog).

  • Distribution: Madagascar.

angulatus Küster, 1848, see: fulgidus Fabricius, 1787.

birmanus Cameron, 1932: 32 (Gauropterus; Type locality: Burma: Tavoy).

  • — Scheerpeltz, 1933: 1315 (Gauropterus; catalog).

  • Distribution: Myanmar.

bucharicus Bernhauer, 1905b: 585 (Gauropterus; Type locality: Buchara).

  • — Reitter, 1908: 113 (Gauropterus; characters; Turkestan).

  • — Bernhauer and Schubert, 1914: 310 (Gauropterus; catalog).

  • — Scheerpeltz, 1933: 1315 (Gauropterus; catalog).

  • — Kirshenblat, 1951: 543 (Gauropterus; characters in key).

  • — Scheerpeltz, 1963: 10 (Gauropterus; Afghanistan).

  • — Coiffait, 1972: 170 (Gauropterus; characters; references; Uzbekistan).

  • Distribution: Uzbekistan, Afghanistan.

cinctus Fauvel, 1904, see: fauveli Steel, 1949.

claviger Fauvel, 1905b: 173 (Xantholinus; Type locality: forêt du Moramanga).

  • — Bernhauer and Schubert, 1914: 310 (Gauropterus; catalog).

  • Distribution: Madagascar.

diabolicus Bernhauer, 1902: 167 (Eulissus; Type locality: Port Daufin, Madagaskar).

  • — Bernhauer and Schubert, 1914: 310 (Gauropterus; catalog).

  • Distribution: Madagascar.

evansi Bernhauer, 1937: 298 (Gauropterus; Type locality: Belgisch-Kongo: Elisabethville).

  • Distribution: Congo.

fauveli Steel, 1949a: 270 (Gauropterus; replacement name for cinctus Fauvel).

  • Distribution: India.

cinctus Fauvel, 1904a: 56 (Xantholinus; [preoccupied]; Type locality: Nilghiris).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • — Cameron, 1932: 32 (Gauropterus; characters; India).

  • — Scheerpeltz, 1933: 1315 (Gauropterus; catalog).

  • — Steel, 1949a: 270 (Gauropterus; synonym of fauveli).

fulgescens Turton, 1802, see: fulgidus Fabricius, 1787.

fulgidus Fabricius, 1787: 220 (Staphylinus; [Note: This reference is often cited as the original citation for both Gauropterus fulgidus and Quedius fulgidus but Tottenham, 1939d: 237, demonstrated that fulgidus Fabricius, 1787, and fulgidus Fabricius, 1793, referred to the species in Gauropterus and Quedius respectively]; Type locality: Kiliae).

  • — Gmelin, 1790: 2028 (Staphylinus; characters).

  • — Fabricius, 1793: 537 (Paederus; characters).

  • — Olivier, 1795:(42): 18 (Staphylinus; characters).

  • — Panzer, 1795: 363 (Paederus; characters; Germany).

  • — Panzer, 1797: 20 (Paederus; characters; illustrations).

  • — Fabricius, 1801: 609 (Paederus; characters; Kiliae).

  • — Ménétriés, 1832: 146 (Gyrohypnus; Caucasus-Persia region).

  • — Stephens, 1833: 253 (Othius; characters; England).

  • — Westwood, 1838a: 16 (Othius; type species).

  • — Erichson, 1839a: 423 (Xantholinus; characters; Germany).

  • — Heer, 1839: 244 (Xantholinus; characters; Switzerland).

  • — Erichson, 1839b: 319 (Xantholinus; characters; Europe).

  • — Stephens, 1839: 402 (Xantholinus; characters; England).

  • — Laporte, 1840: 179 (Gyrohypnus; characters; France).

  • — P. Lucas, 1846: 106 (Xantholinus; notes; Algeria).

  • — Küster, 1848: 6 (Xantholinus; characters; Europe; Sardinia).

  • — Hochhuth, 1849: 101 (Xantholinus; characters).

  • — Redtenbacher, 1849: 691 (Xantholinus; characters; Austria).

  • — Hardy, 1851: 37 (Xantholinus; notes; England).

  • — Rosenhauer, 1856: 70 (Xantholinus; P. S. Maria; Algeciras).

  • — Fairmaire and Laboulbène, 1856: 500 (Xantholinus; characters; France).

  • — Kraatz, 1857d: 643 (Xantholinus; characters; Germany).

  • — Redtenbacher, 1857: 180 (Xantholinus; characters; Austria).

  • — Kraatz, 1858a: 65 (Xantholinus; Crete; Greece).

  • — Peyron, 1858: 420 (Xantholinus; Turkey).

  • — Kraatz, 1859: 105 (Xantholinus; characters; Ceylon).

  • — Thomson, 1859: 27 (type species of Xantholinus).

  • — Thomson, 1860: 188 (Gauropterus; characters; Scandinavia).

  • — Hochhuth, 1862: 67 (Xantholinus; Russia).

  • — Fauvel, 1873b: 98 [= 1873a: 384] (Xantholinus; characters; France; Europe; Algeria; Caramania; Syria).

  • — Redtenbacher, 1874: 218 (Philonthus; characters; Austria).

  • — Seidlitz, 1875: 282 (Xantholinus; characters; Baltic region).

  • — J. Sahlberg, 1876: 33 (Gauropterus; Finland).

  • — Mulsant and Rey, 1877a: 254 (Gauropterus; characters; France).

  • — Fauvel, 1878: 116 [= 1878a: 36] (Xantholinus; Barbary).

  • — LeConte, 1880: 171, 172 (Xantholinus; characters; North America).

  • — Fauvel, 1886: 48 [= 1886a: 40] (Xantholinus; North Africa; Arabia).

  • — Fowler, 1888: 288 (Xantholinus; characters; Britain).

  • — Ganglbauer, 1895: 475 (Eulissus; characters; Palaearctic region; Ceylon; North America).

  • — Fauvel, 1902b: 100 (Xantholinus; Barbary).

  • — Casey, 1906: 380 (Eulissus; characters; Rhode Island; New Jersey).

  • — Sainte-Claire Deville, 1906: 93 (Eulissus; Corsica).

  • — Peyerimhoff, 1907: 14 (Gauropterus; Sinai).

  • — Reitter, 1908: 113 (Gauropterus; characters; Europe).

  • — Reitter, 1909: 136 (Gauropterus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 310 (Gauropterus; catalog).

  • — Johansen, 1914: 431 (Xantholinus; subgenus Eulissus; characters; Denmark).

  • — Portevin, 1929: 370 (Gauropterus; characters; France).

  • — Gridelli, 1930: 71 (Gauropterus; Libya).

  • — Peyerimhoff, 1931: 31 (Gauropterus; Algeria).

  • — Cameron, 1932: 31 (Gauropterus; characters; Ceylon).

  • — Scheerpeltz, 1933: 1316 (Gauropterus; catalog; Caucasus; Burma; Tonkin; Sumatra).

  • — C. Koch, 1936b: 161 (Gauropterus; characters; Egypt).

  • — C. Koch, 1941: 61 (Gauropterus; Morocco).

  • — V. Hansen, 1952: 20 (Gauropterus; characters; Denmark).

  • — Coiffait, 1954: 165 (Gauropterus; Canary Islands).

  • — Coiffait, 1956a: 57 (Gauropterus; characters in key).

  • — Hatch, 1957: 237 (Gyrohypnus; characters; Idaho; Washington; Oregon).

  • — Kirshenblat, 1951: 543 (Gauropterus; characters in key).

  • — Smetana, 1958: 116 (Gauropterus; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1961d: 127 (Gauropterus; Italy).

  • — Nakane, 1963a: 90 (Gyrohypnus; characters; habitus photograph; Japan).

  • — Palm, 1963b: 53 (Gauropterus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 160 (Gauropterus; characters; central Europe).

  • — Scheerpeltz, 1964a: 299 (Gauropterus; Crete).

  • — Focarile, 1964: 56 (Gauropterus; Italy).

  • — Raianu, 1964: 104 (Eulissus; Romania).

  • — Scheerpeltz, 1965b: 400 (Gauropterus; Sicily).

  • — Scheerpeltz, 1965c: 504 (Gauropterus; Sicily).

  • — M. Dvořák, 1965: 88 (Gauropterus; Slovakia).

  • — Horion, 1965: 89 (Gauropterus; Canary Island; Turkey; Saudi Arabia; Caucasus; Ceylon; Burma; Indonesia; Vietnam; Iberian Peninsula to Balkans; Czechoslovakia; Poland; Russia; Ukraine; Ireland; England; Scotland; Norway; Sweden; Finland; Denmark; Austria; Germany).

  • — Coiffait, 1966: 21 (Gauropterus; Turkey).

  • — Coiffait, 1972: 166 (Gauropterus; characters; references; Europe; Britain; Scandinavia; Southeast Asia; North Africa; North America; Canaries; Madeira).

  • — Shibata, 1974: 32 (Gyrohypnus; Japan).

  • — Bordoni, 1975e: 139 (Gauropterus; notes; Corfu).

  • — Pope, 1977: 28 (Gauropterus; Britain).

  • — Smetana, 1979: 51 (requested that International Commission on Zoological Nomenclature place fulgidus Fabricius, 1787 on the Official List of Specific Names in Zoology).

  • — Outerelo, 1980a: 56 (Gauropterus; Spain).

  • — Smetana, 1982: 72 (Gauropterus; characters; California; Colorado; Connecticut; Illinois; Maryland; Massachusetts; New Jersey; New York; Oregon; Pennsylvania; Rhode Island; Vermont).

  • — Bordoni, 1982: 167 (Gauropterus; characters; Italy).

  • — ICZN, 1983: 86 (Gauropterus; placed on Official List of Specific Names in Zoology and fixed as the type species of Gauropterus).

  • — Yuh, Paik, Kwon, and Lee, 1985: 239 (Gauropterus; Korea).

  • — Bordoni, 1986b: 385 (Gauropterus; Slavonia; Bohemia; France; Italy; Sicily; Dalmatia; Greece; Crete; Algeria; Tunisia; Iraq).

  • — Segers, 1986: 40 (Gauropopters; checklist of species; Belgium).

  • — Ádám, 1987: 146 (Gyrohypnus; collecting notes; Hungary).

  • — Lucht, 1987: 102 (Gauropterus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Schülke and Uhlig, 1988: 10 (Gauropterus; notes; Germany).

  • — Tóth, 1989: 14 (Gauropterus; characters; Hungary).

  • — Smetana, 1990c: 84 (Gauropterus; Ontario).

  • — Janák, 1993a: 47 (Gauropterus; Czech Republic; Slovakia).

  • — Ciceroni, 1993: 68 (Gauropterus; Italy).

  • — J. Li, 1993: 32 (Gauropterus; China).

  • — Hernández, Outerelo, and Gamarra, 1994: 190 (Gauropterus; checklist of species; Canary Islands).

  • — Sparacio, 1995: 156 (Gauropterus; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 25 (Gauropterus; Italy).

  • — Downie and Arnett, 1996: 398 (Gauropterus; characters; USA).

  • Distribution: Egypt, Libya, Tunisia, Algeria, Morocco, Canary Islands, Madeira, Europe, Turkey, Caucasus, Syria, Saudi Arabia, Iraq, Sri Lanka, Myanmar, Vietnam, Indonesia, Korea, Japan, Canada, USA.

fulgescens Turton, 1802: 523 (Staphylinus; cited as a replacement name for Paederus fulgidus Fabricius which Fabricius (1787: 220) named in Staphylinus and later (1793: 537) moved to Paederus and is currently cited in Gauropterus; Fabricius (1793: 525) proposed another Staphylinus fulgidus which is now in Quedius; G. fulgidus Fabricius, 1787, is the older of the two homonyms and needs no replacement).

pyropterus Latreille, 1804: 334 (Staphylinus; [Note: Latreille attributed the species to Gravenhorst, but Latreille's publication appeared first]; Type locality: France meridionale).

  • — Küster, 1848: 6 (Xantholinus; synonym of fulgidus).

  • — Coiffait, 1972: 167 (Gauropterus; synonym of fulgidus).

  • — Smetana, 1982: 72 (Gauropterus; synonym of fulgidus).

pyropterus Gravenhorst, 1806: 102 (Staphylinus; [preoccupied]; Type locality: Parisiae).

  • — Gyllenhal, 1810: 356 (Staphylinus; characters; Sweden).

  • — Curtis, 1829: 26 (Gyrohypnus; catalog; Britain).

  • — C. Sahlberg, 1830: 334 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 33 (Gyrohypnus; Sweden).

  • — Mannerheim, 1831: 447 (Gyrohypnus; Sweden).

  • — Ménétriés, 1832: 146 (Gyrohypnus; Caucasus-Persia region).

  • — Griffith and Pidgeon, 1832: 292 (Xantholinus).

  • — Stephens, 1833: 258 (Gyrohypnus; characters; England).

  • — Lacordaire, 1835: 412 (Xantholinus; characters; France).

  • — Nordmann, 1837: 119 (Gyrohypnus; Sweden).

  • — Erichson, 1839a: 423 (Xantholinus; synonym of fulgidus).

  • — Erichson, 1839b: 319 (Xantholinus; synonym of fulgidus).

  • — Stephens, 1839: 402 (Xantholinus; synonym of fulgidus).

  • — Laporte, 1840: 179 (Gyrohypnus; characters; France).

  • — Küster, 1848: 6 (Xantholinus; synonym of fulgidus).

  • — Redtenbacher, 1849: 691 (Xantholinus; synonym of fulgidus).

  • — Fairmaire and Laboulbène, 1856: 500 (Xantholinus; synonym of fulgidus).

  • — Kraatz, 1857d: 643 (Xantholinus; synonym of fulgidus).

  • — Thomson, 1860: 188 (Gyrohypnus; synonym of Gauropterus).

  • — Mulsant and Rey, 1877a: 255 (Gauropterus; synonym of fulgidus).

  • — Ganglbauer, 1895: 475 (Eulissus; synonym of fulgidus).

  • — Reitter, 1908: 113 (Gauropterus; synonym of fulgidus).

  • — Bernhauer and Schubert, 1914: 310 (Gauropterus; synonym of fulgidus).

  • — Cameron, 1932: 31 (Gauropterus; synonym of fulgidus).

  • — Smetana, 1958: 116 (Gauropterus; synonym of fulgidus).

angulatus Küster, 1848: 7 (Xantholinus; Type locality: Carthagena).

  • — Kraatz, 1857d: 643 (Xantholinus; synonym of fulgidus).

  • — Ganglbauer, 1895: 475 (Eulissus; synonym of fulgidus).

  • — Reitter, 1908: 113 (Gauropterus; synonym of fulgidus).

  • — Bernhauer and Schubert, 1914: 310 (Gauropterus; synonym of fulgidus).

  • — Cameron, 1932: 31 (Gauropterus; synonym of fulgidus).

  • — Smetana, 1958: 116 (Gauropterus; synonym of fulgidus).

  • — Smetana, 1982: 72 (Gauropterus; synonym of fulgidus).

intermedius Küster, 1848: 8 (Xantholinus; Type locality: Sardinia).

  • — Kraatz, 1857d: 643 (Xantholinus; synonym of fulgidus).

  • — Fauvel, 1867c: 113 [= 1868b: 212] (Xantholinus; synonym of fulgidus).

  • — Fauvel, 1869: 489 (Xantholinus; synonym of fulgidus).

  • — Fauvel, 1873b: 98 [= 1873a: 384] (Xantholinus; synonym of fulgidus).

  • — Ganglbauer, 1895: 475 (Eulissus; synonym of fulgidus).

  • — Reitter, 1908: 113 (Gauropterus; synonym of fulgidus).

  • — Bernhauer and Schubert, 1914: 310 (Gauropterus; synonym of fulgidus).

  • — Cameron, 1932: 31 (Gauropterus; synonym of fulgidus).

  • — Smetana, 1958: 116 (Gauropterus; synonym of fulgidus).

  • — Smetana, 1982: 72 (Gauropterus; synonym of fulgidus).

pseudosanguinipes Coiffait, 1964a: 504 (Gauropterus; subspecies of fulgidus; Type locality: Sahara, Hoggar, Tihentekert).

  • — Coiffait, 1966b: 337 (Gauropterus; subspecies of fulgidus; Sahara).

  • — Coiffait, 1972: 167 (Gauropterus; subspecies of fulgidus; characters; Sahara; Algeria; Morocco).

  • — Coiffait, 1973a: 273 (Gauropterus; subspecies of fulgidus; Morocco).

gagatinus Erichson, 1839b: 316 (Xantholinus; Type locality: Madagascar).

  • — Bernhauer and Schubert, 1914: 310 (Gauropterus; catalog).

  • Distribution: Madagascar.

gedyei Cameron, 1951c: 400 (Gauropterus; Type locality: Uganda: Budongo Forest).

  • Distribution: Uganda.

hauseri Bernhauer, 1937b: 609 (Gauropterus; Type locality: Pugu).

  • Distribution: Tanzania.

hova Fauvel, 1905b: 172 (Xantholinus; Type locality: forêt de Moramanga).

  • — Bernhauer and Schubert, 1914: 310 (Gauropterus; catalog).

  • Distribution: Madagascar.

intermedius Küster, 1848, see: fulgidus Fabricius, 1787.

marginalis Cameron, 1932: 32 (Gauropterus; Type locality: Bengal: E. Duars).

  • — Scheerpeltz, 1933: 1316 (Gauropterus; catalog).

  • Distribution: India.

methneri Bernhauer, 1908c: 107 (Gauropterus; Type locality: Deutsch-Ostafrika: Usaromo).

  • — Bernhauer and Schubert, 1914: 310 (Gauropterus; catalog).

  • Distribution: Tanzania.

nasutus Harold, 1879: 27 (Xantholinus; Type locality: Inneren [Congo]).

  • — Bernhauer and Schubert, 1914: 310 (Gauropterus; catalog).

  • Distribution: Congo.

nepalicus Coiffait, 1982a: 248 (Gauropterus; Type locality: Nepal: Godavari, 1500–1700 m).

  • Distribution: Nepal.

nigroaeneus Cameron, 1920a: 146 (Gauropterus; Type locality: Pusa).

  • — Cameron, 1932: 32 (Gauropterus; characters; India).

  • — Scheerpeltz, 1933: 1316 (Gauropterus; catalog).

  • Distribution: India.

notabilis Kirshenblat, 1951: 541 (Gauropterus; Type locality: Turkmeniia: Kopet-dag, dolina r. Chuli).

  • — Coiffait, 1972: 169 (Gauropterus; characters; Turkmenia).

  • Distribution: Turkmenistan.

pseudosanguinipes Coiffait, 1964, see: fulgidus Fabricius, 1787.

pustulatus Bernhauer, 1937: 298 (Gauropterus; Type locality: Sierra Leone: Pokuma; Riverbank; Falaba).

  • Distribution: Sierra Leone.

pyropterus Latreille, 1804, see: fulgidus Fabricius, 1787.

pyropterus Gravenhorst, 1806, see: fulgidus Fabricius, 1787.

romeralensis Coiffait and Saiz, 1964: 516 (Gauropterus; Type locality: Chile: Romeral Curicó).

  • — Saiz, 1970b: 385 (Gauropterus; Chile).

  • Distribution: Chile.

rufipennis Cameron, 1932: 31 (Gauropterus; Type locality: United Provinces: Dehra Dun, Lachiwala).

  • — Scheerpeltz, 1933: 1316 (Gauropterus; catalog).

  • Distribution: India.

sanguinipennis Kolenati, 1846: 14 (Xantholinus; Type locality: Iberia; Armenia).

  • — Hochhuth, 1849: 101 (Xantholinus; characters; Iberia; Armenia).

  • — Kraatz, 1857d: 642 (Xantholinus; characters; Crete).

  • — Kraatz, 1858a: 65 (Xantholinus; Crete).

  • — Reitter, 1908: 113 (Gauropterus; characters; Caucasus).

  • — Bernhauer and Schubert, 1914: 310 (Gauropterus; catalog).

  • — Scheerpeltz, 1933: 1316 (Gauropterus; catalog).

  • — Kirshenblat, 1951: 543 (Gauropterus; characters in key).

  • — Jarrige, 1971: 494 (Gauropterus; Iran).

  • — Coiffait, 1972: 170 (Gauropterus; characters; references; Armenia; Caucasus; Asia Minor; Persia; Crete).

  • — Bordoni, 1973a: 179 (Gauropterus; notes; Turkey).

  • — Bordoni, 1975e: 140 (Gauropterus; characters; notes; Rodi).

  • — Bordoni, 1976: 229 (Gauropterus; notes; Turkey).

  • — Bordoni, 1978a: 58 (Gauropterus; cave collection; Turkey).

  • Distribution: “Iberian region” [Caucasus], Greece, Turkey, Iran, Armenia.

sanguinipes Reitter, 1889: 275 (Xantholinus; subgenus Eulissus; Type locality: Araxesthal bei Ordubad).

  • — Reitter, 1908: 113 (Gauropterus; characters; Caucasus).

  • — Bernhauer and Schubert, 1914: 310 (Gauropterus; catalog).

  • — Scheerpeltz, 1933: 1316 (Gauropterus; catalog).

  • — Kirshenblat, 1951: 543 (Gauropterus; characters in key).

  • — Coiffait, 1966: 21 (Gauropterus; Turkey).

  • — Jarrige, 1971: 494 (Gauropterus; Iran).

  • — Coiffait, 1972: 168 (Gauropterus; characters; Caucasus; Asia Minor).

  • Distribution: Azerbaijan, Iran, Turkey.

semenovi Kirshenblat, 1951: 542 (Gauropterus; Type locality: Iran: Kerman, strana Bampur).

  • — Coiffait, 1972: 169 (Gauropterus; characters; Iran).

  • Distribution: Iran.

sjostedti Bernhauer, 1928: 14 (Gauropterus; cited as sjöstedti; Type locality: Sumatra: Silinda, Medan).

  • — Scheerpeltz, 1933: 1316 (Gauropterus; cited as sjoestedti; catalog).

  • — Cameron, 1933d: 385 (Gauropterus; cited as sjöstedti; Sumatra).

  • — Herman, 2001: 12 (Gauropterus; spelling corrected to sjostedti).

  • Distribution: Indonesia.

subcylindricus Jarrige, 1948a: 18 (Gauropterus; Type locality: Madagascar).

  • Distribution: Madagascar.

subsimilis Cameron, 1945: 68 (Gauropterus; Type locality: United Provinces: Chakrata district, Mundali).

  • Distribution: India.

umbilicatus Fauvel, 1905b: 173 (Xantholinus; Type locality: Diego-Suarez; baie d'Antongil: Antanambe; forêt de Moramanga; Tananarive).

  • — Bernhauer and Schubert, 1914: 310 (Gauropterus; catalog).

  • Distribution: Madagascar.

Gyrohypnus [15 species; Neotropical, Nearctic, Palaearctic, and Australian Regions]

Gyrohypnus Leach, 1819: 172 (species included: fulgidus). Type species: Staphylinus fracticornis O. Müller, fixed by the Commission in Opinion 1250 (ICZN, 1983).

  • — Curtis, 1829: 26 (catalog; Britain).

  • — Stephens, 1829a: 23 (catalog; Britain).

  • — Stephens, 1829: 285 (catalog; Britain).

  • — Stephens, 1832: 200 (characters).

  • — Stephens, 1833: 258 (characters; nomenclatural notes).

  • — Nordmann, 1837: 113 (characters).

  • — Westwood, 1838a: 16 (characters; type species).

  • — Erichson, 1839b: 306 (synonym of Xantholinus).

  • — Laporte, 1840: 179 (characters; notes).

  • — Gistel, 1856: 387 (list of species).

  • — Kraatz, 1857d: 629 (synonym of Xantholinus).

  • — Thomson, 1858: 30 (characters).

  • — G. Waterhouse, 1858: 26 (synonym of Xantholinus).

  • — Thomson, 1859: 27 (characters; type species: pilicornis).

  • — Thomson, 1860: 186 (characters).

  • — Fauvel, 1873b: 95 [= 1873a: 381] (synonym of Xantholinus).

  • — Heyden, 1880: 75 (subgenus of Xantholinus; list of species of Siberian region).

  • — Lynch, 1884: 188 (characters; subgenus of Xantholinus).

  • — Seidlitz, 1891: 381 (key to some European species).

  • — Seidlitz, 1891a: 405 (key to some European species).

  • — Heyden, Reitter, and Weise, 1891: 105 (subgenus of Xantholinus; list of species of Europe and Caucasus).

  • — Heyden, 1893: 44 (subgenus of Xantholinus; list of species of Siberian region).

  • — Ganglbauer, 1895: 480 (subgenus of Xantholinus).

  • — Heyden, Reitter, and Weise, 1906: 160 (subgenus of Xantholinus; list of species of Europe and Caucasus).

  • — Casey, 1906: 367, 386 (characters; key to species).

  • — Bernhauer and Schubert, 1914: 299 (subgenus of Xantholinus; Mannerheim, 1830 cited as author of the genus).

  • — Peyerimhoff, 1917: 121 (key to North African species).

  • — Leng, 1920: 105 (catalog of North American species).

  • — Winkler, 1925: 371 (catalog for Palaearctic region; subgenus of Xantholinus).

  • — Scheerpeltz, 1926: 90 (key to Palaearctic species).

  • — Cameron, 1932: 22 (subgenus of Xantholinus).

  • — Scheerpeltz, 1933: 1307 (subgenus of Xantholinus).

  • — Kloet and Hincks, 1945: 165 (list of British species; type species: punctulatus).

  • — Tottenham, 1949: 370, 417 (type species: punctulatus Goeze = fulgidus Paykull [Note: fulgidus Paykull is an unavailable name that is regarded as a misidentification of punctulatus; Paykull cited Fabricius, 1787 as the author of the name]; checklist of British species [Note: The species listed by Tottenham in Gyrohypnus are herein listed in Othius and Othiellus]).

  • — Blackwelder, 1952: 175 (type species: fulgidus; discussion).

  • — V. Hansen, 1952: 15 (subgenus of Xantholinus; characters).

  • — Coiffait, 1956a: 44, 53 (characters; key to French species).

  • — Hatch, 1957: 233, 237 (characters; species of Pacific Northwest).

  • — Smetana, 1958: 95 (characters; habitus of type species; type species: fracticornis; key to Czechoslovakian species).

  • — Smetana, 1959a: 208 (list of species collected in Albania).

  • — Coiffait, 1962b: 389, 390 (characters; key to North African species).

  • — Moore, 1963a: 94 (characters).

  • — Palm, 1963b: 45 (characters; key to species of Sweden).

  • — Lohse, 1964: 161 (characters; key to central European species).

  • — Horion, 1965: 92 (list of central European species).

  • — Pototskaia, 1967: 47 (larval characters).

  • — Scheerpeltz, 1968a: 60 (catalog of Austrian species).

  • — Saiz, 1970b: 383, 388 (characters).

  • — Coiffait, 1972: 175 (characters; cited as Gyrohyphus and Gyrohypnus; type species: fracticornis; key to species of western Palaearctic region; references; notes).

  • — Bordoni, 1973j: 691 (list of species from islands near Sicily).

  • — Tikhomirova, 1973a: 181 (checklist of species of USSR).

  • — Moore and Legner, 1975: 73 (catalog; North America north of Mexico).

  • — Pope, 1977: 28 (list of British species).

  • — Topp, 1978: 311 (larval characters).

  • — Moore and Legner, 1979: 56 (characters; notes).

  • — Muona, 1979: 16 (list of Scandinavian species).

  • — Uhlig, 1979: 251 (some species collected in Germany).

  • — Smetana, 1979: 50 (requested that International Commission on Zoological Nomenclature set aside all type species designations, designate Staphylinus fracticornis O. Müller as type species and place Gyrohypnus on the Official list of Generic Names in Zoology).

  • — Uhlig, Vogel, and Sieber, 1980: 245 (some species collected in Germany).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 9 (catalog; Poland).

  • — Uhlig and Vogler, 1981: 108 (list of some species collected in Germany; habitat and collecting notes).

  • — Smetana, 1982: 187 (characters; habitus; revision of and key to species of North America north of Mexico).

  • — Bordoni, 1982: 206 (characters; habitus; key to species of Italy).

  • — ICZN, 1983: 85 (Gyrohypnus placed on Official List of Generic Names in Zoology and Staphylinus fracticornis is fixed as the type species).

  • — Muona and Viramo, 1986: 14 (list of species of northeastern Finland).

  • — Segers, 1986: 40 (checklist of species; Belgium).

  • — Tóth, 1989: 16 (characters; key to species of Hungary).

  • — Angelini, 1991: 191 (list of species from high elevations of Calabria, Italy).

  • — M. Hansen, Kristensen, Mahler, Pedersen, 1991: 107 (list of Danish species).

  • — Silfverberg, 1992: 18 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Welch, 1993: 228 (ovariole structure and development).

  • — Janák, 1993a: 47 (list of species of Czech Republic and Slovakia).

  • — Hernández, Outerelo, and Gamarra, 1994: 190 (checklist of species; Canary Islands).

  • — Ciceroni and Zanetti, 1995: 25 (list of species of Italy).

  • — M. Hansen, 1996: 109 (list of Danish species).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 242 (additions to list of Danish species).

  • — Downie and Arnett, 1996: 400 (characters; key to species of Northeastern North America).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 74 (list of species collected in 1997; Denmark).

  • — Assing, Frisch, Kahlen, et al., 1998: 136 (note).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 390 (3 Nearctic species; notes; characters in key).

Hyponygrus Tottenham, 1940: 49 (replacement name for Gyrohypnus Leach). Type species: Staphylinus fracticornis O. Müller, fixed by original designation.

  • — Tottenham, 1949: 370, 417 (type species: fracticornis O. Müller = punctulatus Paykull; checklist of British species).

  • — Blackwelder, 1952: 196 (type species: fracticornis).

  • — Hatch, 1957: 233, 235 (characters; key to species of Pacific Northwest).

  • — Coiffait, 1962b: 390 (synonym of Gyrohypnus).

  • — Moore, 1963a: 94 (characters).

  • — Coiffait, 1972: 175 (synonym of Gyrohypnus).

  • — Moore and Legner, 1975: 76 (catalog; cited as valid genus; North America north of Mexico).

  • — Moore and Legner, 1976: 539 (includes seashore species; notes).

  • — Moore and Legner, 1979: 59 (cited as valid genus; characters; notes).

  • — Smetana, 1982: 187 (synonym of Gyrohypnus).

  • — Ádám, 1987: 146 (list of three species with collecting notes; Hungary).

Xenoxantholinus Coiffait, 1972: 175, 177, 187 (subgenus of Gyrohypnus; species included: marginalis). Type species: Xantholinus marginalis Wollaston, fixed by original designation and monotypy.

analis Nordmann, 1837, see: Eulissus.

angustatus Stephens, 1833: 263 (Gyrohypnus; Type locality: London; Suffolk; [Note: See lectotype designation by Assing, 1996b]).

  • — Erichson, 1839a: 426 (Xantholinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 300 cited punctulatus Erichson as a synonym of angustatus Stephens; Erichson did not describe the species as new, he attributed it to Paykull]; characters; Germany).

  • — G. Waterhouse, 1858: 26 (Xantholinus; synonym of ochraceus).

  • — Fauvel, 1902b: 100 (Xantholinus; synonym of punctulatus).

  • — Bernhauer and Schubert, 1914: 300 (Xantholinus; catalog).

  • — Johansen, 1914: 434 (Xantholinus; subgenus Gyrohypnus; characters; Denmark).

  • — Scheerpeltz, 1925: 91 (Xantholinus; subgenus Gyrohypnus; characters).

  • — Scheerpeltz, 1926: 92 (Xantholinus; subgenus Gyrohypnus; characters in key).

  • — Scheerpeltz, 1933: 1307 (Xantholinus; catalog).

  • — C. Koch, 1941: 61 (Xantholinus; Morocco).

  • — V. Hansen, 1952: 15 (Xantholinus; subgenus Gyrohypnus; characters; Denmark).

  • — Coiffait, 1956a: 55 (Gyrohypnus; characters in key).

  • — Hatch, 1957: 236 (Hyponygrus; characters; British Columbia; Washington; Idaho).

  • — Smetana, 1958: 97 (Gyrohypnus; characters; ecology; Czechoslovakia).

  • — Larsson and Gígja, 1959: 81 (Xantholinus; biological notes; Iceland).

  • — Palm, 1963b: 47, 159 (Gyrohypnus; characters; Sweden; Denmark; Norway; Finland).

  • — Smetana, 1964d: 72 (Gyrohypnus; Czechoslovakia).

  • — Scheerpeltz, 1965c: 504 (Gyrohypnus; Sicily).

  • — Horion, 1965: 93 (Gyrohypnus; Siberia; all Europe; Austria; Germany).

  • — Pototskaia, 1967: 47 (Gyrohypnus; larval characters).

  • — Szujecki, 1968a: 726 (Gyrohypnus; Poland).

  • — Bolov, 1969a: 328 (Xantholinus; collecting notes; Russia).

  • — Coiffait, 1972: 184 (Gyrohypnus; characters; references; Europe; Britain; Scandinavia; Siberia; Iberia).

  • — Coiffait, 1973a: 273 (Gyrohypnus; Morocco).

  • — Osella and Zanetti, 1975: 105 (Gyrohypnus; collected from nest of Talpa europaea; notes; Italy).

  • — Bordoni, 1976d: 122 (Gyrohypnus; Netherlands).

  • — Pope, 1977: 28 (Gyrohypnus; Britain).

  • — Outerelo, 1980a: 56 (Gyrohypnus; Spain).

  • — Smetana, 1982: 193 (Gyrohypnus; characters; North American introduction; British Columbia; Newfoundland; New Brunswick; Nova Scotia; Ontario; Quebec; California; New York; Oregon; Washington).

  • — Bordoni, 1982: 216 (Gyrohypnus; characters; Italy).

  • — Lucht, 1987: 103 (Gyrohypnus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Tóth, 1989: 19 (Gyrohypnus; characters; Hungary).

  • — Lohse and Lucht, 1989: 158 (Gyrohypnus; characters).

  • — Nowosad, 1990: 91 (Gyrohypnus; collected from mole nest; notes; Poland).

  • — Campbell and Davies, 1991: 124 (Gyrohypnus; checklist of species; Canada).

  • — Janák and Vysoky, 1992: 136 (Gyrohypnus; collected with Formica pratensis; Bohemia).

  • — Janák, 1993a: 47 (Gyrohypnus; Czech Republic; Slovakia).

  • — Ciceroni, 1993: 71 (Gyrohypnus; Italy).

  • — Owen, 1993: 19 (Gyrohypnus; historical account; distinct from scoticus; Britain).

  • — Sparacio, 1995: 157 (Gyrohypnus; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 25 (Gyrohypnus; Italy).

  • — Assing, 1996b (Gyrohypnus; lectotype designation: locality not cited; characters; synonymic notes).

  • — Levesque and Levesque, 1996: 288 (Gyrohypnus; seasonal abundance; Quebec).

  • — Downie and Arnett, 1996: 400 (Gyrohypnus; characters; USA).

  • — Owen, 1997a: 149 (Gyrohypnus; collecting sites; Ireland).

  • — Assing, Frisch, Kahlen, et al., 1998: 136 (Gyrohypnus; notes).

  • — Kashcheev, 1999a: 57 (Xantholinus; Kazakhstan).

  • Distribution: Morocco, Iceland, Europe, Russia, Caucasus, Turkey, Kazakhstan, Canada, USA.

ochraceus Gyllenhal, 1810: 352 (Staphylinus; [preoccupied]; Type locality: Suecia).

  • — Gyllenhal, 1827: 480 (Staphylinus; characters).

  • — C. Sahlberg, 1830: 332 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 33 (Gyrohypnus; Finland; Sweden).

  • — Mannerheim, 1831: 447 (Gyrohypnus; Finland; Sweden).

  • — Runde, 1835: 11 (Gyrohypnus; characters; Germany).

  • — Nordmann, 1837: 113 (Gyrohypnus; notes; Germany).

  • — Zetterstedt, 1838: 66 (Xantholinus; characters; habitat; Lapland).

  • — Erichson, 1839a: 426 (Xantholinus; synonym of punctulatus).

  • — Heer, 1839: 245 (Xantholinus; characters; Switzerland).

  • — Erichson, 1839b: 328 (Xantholinus; synonym of punctulatus).

  • — Stephens, 1839: 403 (Xantholinus; variety of punctulatus; characters; England).

  • — Mannerheim, 1843a: 83 (Xantholinus; characters).

  • — Märkel, 1845: 235 (Xantholinus; with Formica fuliginosa and F. rufa).

  • — Jacobsen, 1849: 677 (Xantholinus; characters notes; Denmark).

  • — Hardy, 1851: 38 (Xantholinus; synonym of punctulatus).

  • — Fairmaire and Laboulbène, 1856: 502 (Xantholinus; synonym of punctulatus).

  • — Kraatz, 1857d: 636 (Xantholinus; characters; Germany).

  • — Hochhuth, 1862: 66 (Xantholinus; Russia).

  • — Fauvel, 1869: 490 (Xantholinus; synonym of punctulatus Paykull).

  • — Fauvel, 1873b: 99 [= 1873a: 385] (Xantholinus; synonym of punctulatus).

  • — Redtenbacher, 1874: 219 (Xantholinus; characters; Austria).

  • — Seidlitz, 1875: 281 (Xantholinus; characters; Baltic region).

  • — J. Sahlberg, 1876: 34 (Xantholinus; Finland).

  • — Mulsant and Rey, 1877a: 294 (Xantholinus; subgenus Gyrohypnus; characters; France).

  • — Fowler, 1888: 289 (Xantholinus; characters; Britain).

  • — Ganglbauer, 1895: 480 (Xantholinus; synonym of angustatus).

  • — Fauvel, 1902b: 100 (Xantholinus; synonym of punctulatus).

  • — Reitter, 1908: 115 (Xantholinus; synonym of angustatus).

  • — Joy, 1913c: 225 (Xantholinus; characters).

  • — Bernhauer and Schubert, 1914: 300 (Xantholinus; synonym of angustatus).

  • — Scheerpeltz, 1933: 1308 (Xantholinus; synonym of angustatus).

  • — Steel, 1938: 173 (Xantholinus; variety of angustatus).

  • — Smetana, 1958: 97 (Gyrohypnus; synonym of angustatus).

  • — Coiffait, 1972: 184 (Gyrohypnus; synonym of angustatus).

  • — Smetana, 1982: 193 (Gyrohypnus; synonym of angustatus).

  • — Bordoni, 1982: 216 (Gyrohypnus; synonym of angustatus).

thomsoni Schwarz, 1872: 154 (Xantholinus; Type locality: Not cited).

  • — Fauvel, 1873b: 99 [= 1873a: 385] (Xantholinus; synonym of punctulatus).

  • — Ganglbauer, 1895: 480 (Xantholinus; variety of punctulatus).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; aberration of punctulatus).

  • — Cameron, 1932: 24 (Xantholinus; variety of punctulatus).

  • — Scheerpeltz, 1933: 1313 (Xantholinus; aberration of punctulatus).

  • — Smetana, 1958: 96 (Gyrohypnus; synonym of fracticornis).

  • — Coiffait, 1972: 182 (Gyrohypnus; variety of fracticornis).

  • — Tóth, 1989: 19 (Gyrohypnus; variety of angustatus; characters).

nitidicollis Reitter, 1908: 115 [= 1908a: 18] (Xantholinus; subgenus Gyrohypnus; variety of angustatus; Type locality: Europe; Caucasus; Siberia).

  • — Scheerpeltz, 1933: 1308 (Xantholinus; synonym of angustatus).

  • — Steel, 1938: 173 (Xantholinus; aberration of angustatus).

  • — Coiffait, 1956a: 55 (Gyrohypnus; subspecies of angustatus; characters; Czechoslovakia; France).

  • — Smetana, 1958: 97 (Gyrohypnus; aberration of angustatus).

  • — Coiffait, 1965e: 119 (Gyrohypnus; Turkey).

  • — Coiffait, 1966: 21 (Gyrohypnus; variety of angustatus; Turkey).

  • — Coiffait, 1972: 184 (Gyrohypnus; synonym of angustatus).

  • — Pope, 1977: 28 (Gyrohypnus; synonym of angustatus).

  • — Smetana, 1982: 193 (Gyrohypnus; synonym of angustatus).

  • — Bordoni, 1982: 216 (Gyrohypnus; synonym of angustatus).

  • — Lohse, 1987: 137 (Gyrohypnus; synonym of scoticus).

  • — Tóth, 1989: 19 (Gyrohypnus; variety of angustatus; characters; Hungary).

  • — Lohse and Lucht, 1989: 158 (Gyrohypnus; synonym of angustatus).

scoticus Joy, 1913: 226 (Xantholinus; Type locality: Struy, Strathglass, Inverness; Altnahara, Sutherland).

  • — Ganglbauer, 1895: 480 (Xantholinus; subgenus Gyrohypnus; [Note: Misidentified as angustatus according to Lohse, 1987: 139]; characters; Europe; Siberia).

  • — Reitter, 1908: 115 (Xantholinus; subgenus Gyrohypnus; [Note: Misidentified as angustatus according to Lohse, 1987: 139]; characters; Europe; Caucasus; Siberia).

  • — Reitter, 1909: 136 (Xantholinus; subgenus Gyrohypnus; [Note: Misidentified as angustatus according to Lohse, 1987: 139]; characters; Germany).

  • — Scheerpeltz, 1933: 1308 (Xantholinus; synonym of angustatus).

  • — Steel, 1938: 173 (Xantholinus; synonym of angustatus variety ochraceus).

  • — Smetana, 1958: 97 (Gyrohypnus; synonym of angustatus).

  • — Lohse, 1964: 162 (Gyrohypnus; [Note: Misidentified as angustatus according to Lohse, 1987: 139]; characters; central Europe).

  • — Coiffait, 1972: 184 (Gyrohypnus; variety of angustatus).

  • — Bordoni, 1975d: 197 (Gyrohypnus; subspecies of angustatus; cited with Bordoni as author; notes; Italy).

  • — Pope, 1977: 28 (Gyrohypnus; synonym of angustatus).

  • — Smetana, 1982: 193 (Gyrohypnus; synonym of angustatus).

  • — Bordoni, 1982: 218 (Gyrohypnus; subspecies of angustatus; characters; Italy).

  • — Lohse and Lucht, 1989: 158 (Gyrohypnus; valid species; characters).

  • — Silfverberg, 1992: 18 (Gyrohypnus; cited as valid species; Finland; Russia; Sweden; Norway; Denmark; Estonia; Lithuania; Latvia).

  • — Owen, 1993: 19 (Gyrohypnus; valid species; discussion).

  • — Assing, 1993: 30 (Gyrohypnus; bionomics; Germany).

  • — Hodge and Jones, 1995: 37 (Gyrohypnus; characters; Britain).

  • — Ciceroni and Zanetti, 1995: 25 (Gyrohypnus; Italy).

  • — M. Hansen, 1996: 109 (Gyrohypnus; cited as valid species; Denmark).

  • — Assing, 1996b: 78, 83 (Gyrohypnus; synonym of angustatus).

  • — Assing, Frisch, Kahlen, et al., 1998: 136 (Gyrohypnus; synonym of angustatus).

  • — Burakowski, Mroczkowski, and Stefańska, 2000: 47 (Gyrohypnus; catalog; Poland).

apicalis Stephens, 1833, see: Leptacinus.

assimilis Kirby, 1837, see: Nudobius.

ater Stephens, 1833, see: fracticornis O. Müller, 1776.

atratus Heer, 1839: 246 (Xantholinus; Type locality: Basel).

  • — Kraatz, 1857d: 636 (Xantholinus; characters; Germany).

  • — Redtenbacher, 1857: 983 (Xantholinus; characters; Austria).

  • — Hochhuth, 1862: 66 (Xantholinus; Russia).

  • — Fauvel, 1865a: 298 [= 1865b: 48] (Xantholinus; habitat; France).

  • — Fauvel, 1869: 489 (Xantholinus; synonym of punctulatus).

  • — Fauvel, 1873b: 99 [= 1873a: 385] (Xantholinus; synonym of punctulatus).

  • — Redtenbacher, 1874: 219 (Xantholinus; characters; Austria).

  • — Seidlitz, 1875: 281 (Xantholinus; characters; Baltic region).

  • — J. Sahlberg, 1876: 34 (Xantholinus; Finland).

  • — Mulsant and Rey, 1877a: 296 (Xantholinus; subgenus Gyrohypnus; characters; France).

  • — Fowler, 1888: 289 (Xantholinus; characters; Britain).

  • — Ganglbauer, 1895: 480 (Xantholinus; subgenus Gyrohypnus; characters; north and middle Europe).

  • — Reitter, 1908: 115 (Xantholinus; subgenus Gyrohypnus; characters; northern and central Europe).

  • — Reitter, 1909: 136 (Xantholinus; subgenus Gyrohypnus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • — Johansen, 1914: 434 (Xantholinus; subgenus Gyrohypnus; characters; Denmark).

  • — J. Sahlberg, 1920: 262 (Xantholinus; [Note: Misidentification: Scheerpeltz, 1933: 1308 cited glaber J. Sahlberg as a synonym of atratus; J. Sahlberg did not describe the species as new it was simply included in a list]).

  • — Scheerpeltz, 1925: 90 (Xantholinus; subgenus Gyrohypnus; characters).

  • — Scheerpeltz, 1926: 91 (Xantholinus; subgenus Gyrohypnus; characters in key).

  • — Scheerpeltz, 1933: 1308 (Xantholinus; catalog).

  • — V. Hansen, 1952: 16 (Xantholinus; subgenus Gyrohypnus; characters; Denmark).

  • — Coiffait, 1956a: 55 (Gyrohypnus; characters in key).

  • — Smetana, 1958: 98 (Gyrohypnus; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 47 (Gyrohypnus; characters; Sweden).

  • — Lohse, 1964: 162 (Gyrohypnus; characters; central Europe).

  • — Horion, 1965: 93 (Gyrohypnus; Russia; Finland; Sweden; Norway; England; Denmark; Poland; Czechoslovakia; Hungary; Romania; Istria; Switzerland; Holland; France; Spain ?; Austria; Germany).

  • — Coiffait, 1972: 187 (Gyrohypnus; characters; references; France).

  • — Pope, 1977: 28 (Gyrohypnus; Britain).

  • — Bordoni, 1982: 210 (Gyrohypnus; characters; Italy).

  • — Lucht, 1987: 103 (Gyrohypnus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Tóth, 1989: 19 (Gyrohypnus; characters; Hungary).

  • — Lohse and Lucht, 1989: 156 (Gyrohypnus; characters).

  • — Janák and Vysoky, 1992: 136 (Gyrohypnus; collected with Formica polyctena and rufa; Bohemia).

  • — Janák, 1993a: 47 (Gyrohypnus; Czech Republic; Slovakia).

  • — Franc, 1993: 308 (Gyrohypnus; synecthrical myrmecophile; Slovakia).

  • — Ciceroni and Zanetti, 1995: 25 (Gyrohypnus; Italy).

  • — Kashcheev, 1999a: 57 (Xantholinus; Kazakhstan).

  • Distribution: Europe, Russia, Kazakhstan.

confusus Mulsant and Rey, 1853: 58 [= 1853a: 71] (Xantholinus; variety of punctulatus; Type locality: Not cited).

  • — Mulsant and Rey, 1853: 58 (Xantholinus; variety of punctulatus; characters; France).

  • — Kraatz, 1855: 23 (Xantholinus; variety of punctulatus; notes).

  • — Ganglbauer, 1895: 481 (Xantholinus; synonym of atratus).

  • — Reitter, 1908: 115 (Xantholinus; synonym of atratus).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; synonym of atratus).

  • — Smetana, 1958: 98 (Gyrohypnus; synonym of atratus).

  • — Coiffait, 1972: 187 (Gyrohypnus; synonym of atratus).

picipes Thomson, 1860: 190 (Xantholinus; Type locality: Not cited, presumably Sweden).

  • — Fauvel, 1869: 490 (Xantholinus; synonym of punctulatus Paykull).

  • — Fauvel, 1873b: 99 [= 1873a: 385] (Xantholinus; synonym of punctulatus).

  • — J. Sahlberg, 1876: 34 (Xantholinus; Finland).

  • — Mulsant and Rey, 1877a: 297 (Xantholinus; subgenus Gyrohypnus; characters; France).

  • — Ganglbauer, 1895: 481 (Xantholinus; synonym of atratus).

  • — Reitter, 1908: 115 (Xantholinus; synonym of atratus).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; synonym of atratus).

  • — Smetana, 1958: 98 (Gyrohypnus; synonym of atratus).

  • — Coiffait, 1972: 187 (Gyrohypnus; synonym of atratus).

bipartitus Casey, 1906, see: Neohypnus.

brouni Steel, 1949, see: Maorothius.

campbelli Smetana, 1982: 196 (Gyrohypnus; Type locality: Quebec: Montreal. Also cited from Manitoba, Ontario, Connecticut, Massachusetts, and Michigan).

  • — Smetana, 1988a: 554 (Gyrohypnus; New Hampshire).

  • — Smetana, 1990c: 85 (Gyrohypnus; New Hampshire; New York; Wisconsin).

  • — Downie and Arnett, 1996: 401 (Gyrohypnus; characters; USA).

  • Distribution: Canada, USA.

confusus Mulsant and Rey, 1853, see: atratus Heer, 1839.

consentaneus Nordmann, 1837, see: Nudobius.

davisi Notman, 1924, see: Oxybleptes.

ebneri Scheerpeltz, 1925: 85, 90 (Xantholinus; subgenus Gyrohypnus; Type locality: Süd-Spanien, Gipfelregion des Muley-Halçen, Sierra Nevada).

  • — Scheerpeltz, 1926: 91 (Xantholinus; subgenus Gyrohypnus; characters in key).

  • — Scheerpeltz, 1933: 1309 (Xantholinus; catalog).

  • — Coiffait, 1972: 183 (Gyrohypnus; characters; references; Spain; Portugal).

  • Distribution: Spain, Portugal.

elongatus Geoffroy, 1785, see: fracticornis O. Müller, 1776.

flavipennis Nordmann, 1837, see: Nudobius.

fracticornis O. Müller, 1776: 99 (Staphylinus; Type locality: Dania).

  • — Goeze, 1777: 728 (Staphylinus; characters).

  • — Gmelin, 1790: 2033 (Staphylinus; characters; Denmark).

  • — Turton, 1802: 517 (Staphylinus; cited as fractricornis; characters; Denmark).

  • — Erichson, 1839b: 328 (Xantholinus; synonym of punctulatus).

  • — Kolenati, 1846: 15 (Xantholinus; synonym of punctulatus).

  • — Kraatz, 1857d: 635 (Xantholinus; synonym of punctulatus).

  • — Thomson, 1860: 189 (Xantholinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 307 cited atratus Thomson as a synonym of punctulatus Paykull and Coiffait, 1972: 182 cited atratus Thomson as a synonymn of fracticornis; Thomson did not describe the species as new, he attributed it to Heer]; characters; Scandinavia).

  • — Ganglbauer, 1895: 480 (Xantholinus; synonym of punctulatus).

  • — Reitter, 1908: 115 (Xantholinus; synonym of punctulatus).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; synonym of punctulatus).

  • — Cameron, 1932: 24 (Xantholinus; synonym of punctulatus).

  • — Lohse, 1958b: 121 (Gyrohypnus; identification).

  • — Smetana, 1958: 96 (Gyrohypnus; characters; ecology; Czechoslovakia).

  • — Palm, 1959: 22 (Gyrohypnus; characters; illustrations).

  • — Coiffait, 1962b: 391 (Gyrohypnus; characters; North Africa).

  • — Smetana, 1963b: 35 (Gyrohypnus; Madeira).

  • — Palm, 1963b: 47 (Gyrohypnus; characters; Sweden).

  • — Lohse, 1964: 162 (Gyrohypnus; characters; central Europe).

  • — Scheerpeltz, 1964a: 299 (Gyrohypnus; Crete).

  • — Coiffait, 1965e: 119 (Gyrohypnus; Turkey).

  • — Scheerpeltz, 1965b: 399 (Gyrohypnus; Sicily).

  • — Horion, 1965: 92 (Gyrohypnus; all northern and central Europe; Austria; Germany).

  • — Szujecki, 1968a: 725 (Gyrohypnus; Poland).

  • — Bolov, 1969a: 328 (Xantholinus; collecting notes; Russia).

  • — Saiz, 1970b: 388 (Gyrohypnus; characters; Chile).

  • — Smetana, 1970: 58 (Gyrohypnus; distributional summary; Azores).

  • — Coiffait, 1972: 182 (Gyrohypnus; characters; references).

  • — Coiffait, 1972: 182 (Gyrohypnus; characters; references; Palaearctic region; India; Argentina).

  • — Bordoni, 1975a: 148 (Gyrohypnus; Afghanistan).

  • — Shilov, 1975a: 375 [= 1975b: 97] (Gyrohypnus, cited as Cyrohypnus; Komi region of Russia).

  • — Bordoni, 1976d: 123 (Gyrohypnus; Netherlands).

  • — Pope, 1977: 28 (Gyrohypnus; Britain).

  • — Coiffait, 1978d: 163 (Gyrohypnus; Turkey).

  • — Smetana, 1978c: 338 (Gyrohypnus; distinct from punctulatus; notes; characters; North America).

  • — Smetana, 1979: 50 (requested that International Commission on Zoological Nomenclature place fracticornis on the Official List of Specific Names in Zoology).

  • — Hinton, 1981: 663 (Xantholinus; characters of egg).

  • — Smetana, 1982: 189 (Gyrohypnus; characters; North American introduction; Alberta; British Columbia; Manitoba; New Brunswick; Newfoundland; Nova Scotia; Ontario; Quebec; Arizona; Arkansas; California; Colorado; Idaho; Illinois; Iowa; Kansas; Maine; Maryland; Massachusetts; Michigan; Montana; Nebraska; Nevada; New Hampshire; New Jersey; New Mexico; New York; North Carolina; North Dakota; Ohio; Oregon; Pennsylvania; Rhode Island; South Dakota; Tennessee; Utah; Vermont; Virginia; Washington; West Virginia; Wisconsin; Wyoming).

  • — Bordoni, 1982: 213 (Gyrohypnus; characters; Italy).

  • — Frank, 1982: 31 (Gyrohypnus; list of parasites).

  • — ICZN, 1983: 85 (Gyrohypnus; placed on Official List of Specific Names in Zoology and fixed as type species of Gyrohypnus).

  • — H. Franz, 1987: 69 (Gyrohypnus; Cyprus).

  • — Lucht, 1987: 103 (Gyrohypnus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Tóth, 1989: 18 (Gyrohypnus; characters; Hungary).

  • — Borges, 1990: Table II (Gyrohypnus; Azores).

  • — Kuschel, 1990: 26 (Gyrohypnus; introduced, Palaearctic species; winged; habitat; New Zealand).

  • — Silfverberg, 1992: 18 (Gyrohypnus; Finland; Russia; Sweden; Norway; Denmark; Latvia; Lithuania).

  • — Janák, 1993a: 47 (Gyrohypnus; Czech Republic; Slovakia).

  • — Ciceroni, 1993: 71 (Gyrohypnus; Italy).

  • — Welch, 1993: 228 (Gyrohypnus; ovariole number and ovary structure).

  • — Hodge and Jones, 1995: 37 (Gyrohypnus; characters; Britain).

  • — Sparacio, 1995: 157 (Gyrohypnus; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 25 (Gyrohypnus; Italy).

  • — Downie and Arnett, 1996: 400 (Gyrohypnus; characters; USA).

  • — Owen, 1997a: 149 (Gyrohypnus; collecting sites; Ireland).

  • — Emberson, 1998: 36 (Gyrohypnus; collecting notes; Chatham Islands).

  • — Kashcheev, 1999a: 57 (Xantholinus; Kazakhstan).

  • Distribution: Azores, Madeira, Europe, Russia, Kazakhstan, Turkey, Afghanistan, India, New Zealand, Chatham Islands, Canada, USA, Argentina, Chile.

elongatus Geoffroy, 1785: 171 (Staphylinus; [preoccupied]; Type locality: Paris).

  • — Gravenhorst, 1802: 45 (Staphylinus; characters).

  • — Latreille, 1804: 330 (Staphylinus; characters; France; Germany).

  • — Gravenhorst, 1806: 98 (Staphylinus; characters).

  • — C. Sahlberg, 1830: 332 (Staphylinus; synonym of ochraceus).

  • — Griffith and Pidgeon, 1832: 292 (Xantholinus).

  • — Erichson, 1839b: 328 (Xantholinus; synonym of punctulatus).

  • — Laporte, 1840: 179 (Gyrohypnus; characters; France).

  • — Kolenati, 1846: 15 (Xantholinus; synonym of punctulatus).

  • — P. Lucas, 1846: 107 (Xantholinus; synonym of punctulatus).

  • — Kraatz, 1857d: 635 (Xantholinus; synonym of punctulatus).

  • — Mulsant and Rey, 1877a: 291 (Xantholinus; synonym of punctulatus).

  • — Ganglbauer, 1895: 480 (Xantholinus; synonym of punctulatus).

  • — Reitter, 1908: 115 (Xantholinus; synonym of punctulatus).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; synonym of punctulatus).

  • — Cameron, 1932: 24 (Xantholinus; synonym of punctulatus).

  • — Smetana, 1958: 96 (Gyrohypnus; synonym of fracticornis).

  • — Coiffait, 1972: 182 (Gyrohypnus; synonym of fracticornis).

ater Stephens, 1833: 255 (Othius; Type locality: Norfolk).

  • — Stephens, 1839: 402 (Othius; characters; England).

  • — Hardy, 1851: 36 (Othius; synonym of melanocephalus).

  • — G. Waterhouse, 1858: 26 (Xantholinus; synonym of punctulatus).

  • — Ganglbauer, 1895: 480 (Xantholinus; synonym of punctulatus).

  • — Reitter, 1908: 115 (Xantholinus; synonym of punctulatus).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; synonym of punctulatus).

  • — Cameron, 1932: 24 (Xantholinus; synonym of punctulatus).

  • — Smetana, 1958: 96 (Gyrohypnus; synonym of fracticornis).

  • — Coiffait, 1972: 182 (Gyrohypnus; synonym of fracticornis).

  • — Muona, 1979: 16 (Gyrohypnus; cited as valid species; [Note: Cited by Muona as the senior synonym of punctulatus Paykull but in a later version of the checklist (Silfverberg, 1992) ater Stephens is not cited at all but “ater auct.” is and as a “synonym” of liebei which is cited as the senior synonym of punctulatus Paykull. I have therefore let stand the earlier synonymy of ater Stephens with fracticornis]).

obscurus Stephens, 1833: 256 (Othius; Type locality: Plyn; Ashburton).

  • — Curtis, 1829: 26 (Xantholinus; nomen nudum; catalog; Britain).

  • — Stephens, 1839: 402 (Othius; characters; England).

  • — G. Waterhouse, 1858: 26 (Xantholinus; synonym of punctulatus).

  • — Ganglbauer, 1895: 480 (Xantholinus; synonym of punctulatus).

  • — Reitter, 1908: 115 (Xantholinus; synonym of punctulatus).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; synonym of punctulatus).

  • — Cameron, 1932: 24 (Xantholinus; synonym of punctulatus).

  • — Smetana, 1958: 96 (Gyrohypnus; synonym of fracticornis).

  • — Coiffait, 1972: 182 (Gyrohypnus; synonym of fracticornis).

obsidianus Melsheimer, 1844: 34 (Xantholinus; Type locality: Pennsylvania; [Note: See lectotype designation by Smetana, 1982]).

  • — LeConte, 1850: 219 (Xantholinus; Lake Superior).

  • — Provancher, 1877: 251 (Xantholinus; characters; Quebec).

  • — Casey, 1906: 387 (Gyrohypnus; characters; New York Iowa; Lake Superior; Nevada; Oregon, Manitoba).

  • — Fall and Cockerell, 1907: 167 (Xantholinus; New Mexico).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Steel, 1949a: 269 (Xantholinus; synonym of fracticornis).

  • — Hatch, 1957: 236 (Hyponygrus; characters; British Columbia; Washington; Oregon; Idaho).

  • — Smetana, 1982: 189 (Gyrohypnus; lectotype designation; synonym of fracticornis).

morio Reitter, 1872: 167 (Xantholinus; [preoccupied]; Type locality: [Oran] Frendah).

  • — Fauvel, 1873b: 99 [= 1873a: 385] (Xantholinus; synonym of punctulatus).

  • — Ganglbauer, 1895: 480 (Xantholinus; synonym of punctulatus).

  • — Fauvel, 1902b: 100 (Xantholinus; synonym of punctulatus).

  • — Reitter, 1908: 115 (Xantholinus; synonym of punctulatus).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; synonym of punctulatus).

  • — Cameron, 1932: 24 (Xantholinus; synonym of punctulatus).

  • — Smetana, 1958: 96 (Gyrohypnus; synonym of fracticornis).

  • — Coiffait, 1962b: 391 (Gyrohypnus; synonym of fracticornis).

  • — Coiffait, 1972: 182 (Gyrohypnus; synonym of fracticornis).

melanarius Fauvel, 1872a: 304 (Xantholinus; replacement name for morio Reitter).

  • — Fauvel, 1873b: 99 [= 1873a: 385] (Xantholinus; synonym of punctulatus).

  • — Ganglbauer, 1895: 480 (Xantholinus; synonym of punctulatus).

  • — Reitter, 1908: 115 (Xantholinus; synonym of punctulatus).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; synonym of punctulatus).

  • — Cameron, 1932: 24 (Xantholinus; synonym of punctulatus).

  • — Smetana, 1958: 96 (Gyrohypnus; synonym of fracticornis).

  • — Coiffait, 1962b: 391 (Gyrohypnus; synonym of fracticornis).

  • — Coiffait, 1972: 182 (Gyrohypnus; synonym of fracticornis).

haroldi Reitter, 1873: 146 (Xantholinus; replacement name for morio Reitter).

  • — Fauvel, 1873b: 99 [= 1873a: 385] (Xantholinus; synonym of punctulatus).

  • — Ganglbauer, 1895: 480 (Xantholinus; synonym of punctulatus).

  • — Reitter, 1908: 115 (Xantholinus; synonym of punctulatus).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; synonym of punctulatus).

  • — Cameron, 1932: 24 (Xantholinus; synonym of punctulatus).

fragilis Casey, 1906, see: Neohypnus.

fuscipes Linné, 1758: 423 (Staphylinus; Type locality: Europa).

  • — Linné, 1760: 233 (Staphylinus; characters; Sweden).

  • — Pontoppidans, 1765: 211 (Staphylinus; Denmark).

  • — Linné, 1767: 685 (Staphylinus; characters; Europe).

  • — Fabricius, 1775: 266 (Staphylinus; characters).

  • — Goeze, 1777: 721 (Staphylinus; characters).

  • — Fabricius, 1781: 336 (Staphylinus; characters; Europe).

  • — Herbst, 1784: 150 (Staphylinus; Germany).

  • — Fabricius, 1787: 221 (Staphylinus; characters).

  • — Paykull, 1789: 53 (Staphylinus; redescription; Sweden).

  • — Villers, 1789: 418 (Staphylinus; characters; France).

  • — Rossi, 1790: 251 (Staphylinus; characters; Italy).

  • — Gmelin, 1790: 2030 (Staphylinus; characters).

  • — Fabricius, 1793: 527 (Staphylinus; characters).

  • — Schrank, 1798: 645 (Staphylinus; characters; notes).

  • — Cederhjelm, 1798: 119 (Staphylinus; characters).

  • — Paykull, 1800: 397 (Staphylinus; characters; Sweden).

  • — Fabricius, 1801: 598 (Staphylinus; characters).

  • — Turton, 1802: 513 (Staphylinus; characters; Europe).

  • — Marsham, 1802: 525 (Staphylinus; characters; Britain).

  • — Westwood, 1845: 53 (Gyrohypnus; based on examination of Linnean type and may be identical to lentus; characters; notes).

  • Distribution: Sweden, Britain, Germany, France, Italy.

gilvipennis Casey, 1906, see: Nudobius.

godeti Laporte, 1840, see: Nudobius.

gredensis Coiffait, 1964, see: wagneri Scheerpeltz, 1926.

haroldi Reitter, 1873, see: fracticornis O. Müller, 1776.

infumatus Casey, 1906, see: Neohypnus.

libanoticus Bordoni, 1984: 336 (Gyrohypnus; Type locality: Libano: Les Cèdres, 2200 m).

  • Distribution: Lebanon.

liebei Scheerpeltz, 1926: 86, 92 (Xantholinus; subgenus Gyrohypnus; Type locality: Umgebung von Funchal, San Antonio da Serra auf Maderia).

  • — Scheerpeltz, 1933: 1311 (Xantholinus; catalog).

  • — Smetana, 1963b: 35 (Gyrohypnus; characters; notes; Madeira).

  • — Coiffait, 1972: 183 (Gyrohypnus; characters; references; Madeira).

  • — Lohse, 1988: 41, 49 (Gyrohypnus; synonymic notes).

  • — Lohse and Lucht, 1989: 156 (Gyrohypnus; characters).

  • — Silfverberg, 1992: 18 (Gyrohypnus; Finland, Russia, Sweden, Norway, Denmark, Estonia, Latvia, Lithuania).

  • — Ciceroni and Zanetti, 1995: 25 (Gyrohypnus; Italy).

  • — M. Hansen, 1996: 109 (Gyrohypnus; Denmark).

  • Distribution: Ethiopia, Algeria, Morocco, Canary Islands, Madeira, Azores, Europe, Russia, Georgia, Iran, Kazakhstan, “central Asia”, “East Indies”.

punctulatus Paykull, 1789: 30 (Staphylinus; [preoccupied]; Type locality: Suecia).

  • — Fabricius, 1793: 528 (Staphylinus; characters).

  • — Panzer, 1795: 357 (Staphylinus; characters; Germany).

  • — Paykull, 1800: 380 (Staphylinus; characters; Sweden).

  • — Turton, 1802: 514 (Staphylinus; characters; Sweden).

  • — Gyllenhal, 1810: 353 (Staphylinus; characters; Sweden).

  • — Curtis, 1829: 27 (Gyrohypnus; catalog; Britain).

  • — C. Sahlberg, 1830: 333 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 33 (Gyrohypnus; Finland; Sweden; Russia).

  • — Mannerheim, 1831: 447 (Gyrohypnus; Finland; Sweden; Russia).

  • — Stephens, 1833: 263 (Gyrohypnus; characters; England).

  • — Runde, 1835: 11 (Gyrohypnus; characters; Germany).

  • — Lacordaire, 1835: 415 (Xantholinus; characters; France).

  • — Nordmann, 1837: 117 (Gyrohypnus; Europe).

  • — Heer, 1839: 245 (Xantholinus; characters; Switzerland).

  • — Erichson, 1839b: 328 (Xantholinus; characters; Europe).

  • — Stephens, 1839: 403 (Xantholinus; characters; England).

  • — Märkel, 1845: 235 (Xantholinus; with Formica fuliginosa and F. rufa).

  • — Kolenati, 1846: 15 (Xantholinus; Caucasus; Transcaucasus).

  • — P. Lucas, 1846: 107 (Xantholinus; notes; Algeria).

  • — Hochhuth, 1849: 107 (Xantholinus; characters; Tiflis).

  • — Redtenbacher, 1849: 692 (Xantholinus; characters; Austria).

  • — Hardy, 1851: 37 (Xantholinus; notes; England).

  • — Wollaston, 1854: 577 (Xantholinus; characters; notes; Madeira Islands).

  • — Fairmaire and Laboulbène, 1856: 502 (Xantholinus; characters; France).

  • — Wollaston, 1857: 188 (Xantholinus; habitat; Madeira Islands).

  • — Kraatz, 1857d: 635 (Xantholinus; characters; Germany).

  • — Redtenbacher, 1857: 182 (Xantholinus; characters; Austria).

  • — Kraatz, 1858a: 64 (Xantholinus; Greece).

  • — Kraatz, 1859: 104 (Xantholinus; characters; India orientalis).

  • — Thomson, 1860: 189 (Xantholinus; characters; Scandinavia).

  • — Wollaston, 1864: 579 (Xantholinus; characters; Canary Islands).

  • — Fauvel, 1865a: 298 [= 1865b: 48] (Xantholinus; habitat; France).

  • — Wollaston, 1865: 497 (Xantholinus; habitat; Madeira Islands; Canary Islands).

  • — Crotch, 1867a: 383 (Xantholinus; Azores).

  • — Fauvel, 1873b: 99 [= 1873a: 385] (Xantholinus; characters; France; Europe; Mediterranean region; Georgia; Caucasus; Iran; Siberia; Madeira; North America).

  • — Redtenbacher, 1874: 219 (Xantholinus; characters; Austria).

  • — Seidlitz, 1875: 281 (Xantholinus; characters; Baltic region).

  • — J. Sahlberg, 1876: 33 (Xantholinus; Finland).

  • — Mulsant and Rey, 1877a: 291 (Xantholinus; subgenus Gyrohypnus; characters; France).

  • — Fauvel, 1878: 117 [= 1878a: 37] (Xantholinus; Algeria).

  • — Fauvel, 1886: 49 [= 1886a: 41] (Xantholinus; North Africa).

  • — Fowler, 1888: 289 (Xantholinus; characters; Britain).

  • — Ganglbauer, 1895: 480 (Xantholinus; subgenus Gyrohypnus; characters; Palaearctic region; North America).

  • — Fauvel, 1902b: 100 (Xantholinus; Barbary; Azores; Madeira; Canary Islands).

  • — Poppius, 1905: 10 (Xantholinus; Orkney Islands; Shetland Islands).

  • — Sainte-Claire Deville, 1906: 92 (Xantholinus; Corsica).

  • — Reitter, 1908: 115 (Xantholinus; subgenus Gyrohypnus; characters; Europe; North Africa; central Asia).

  • — Montandon, 1908: 79 (Xantholinus; Romania).

  • — Reitter, 1909: 136 (Xantholinus; subgenus Gyrohypnus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Johansen, 1914: 433 (Xantholinus; subgenus Gyrohypnus; characters; Denmark).

  • — Eichelbaum, 1915a: 96 (Xantholinus; abdominal segmentation; apical segements).

  • — Šulc, 1922 (Xantholinus; osmeterium; anatomy; histology; physiology).

  • — Scheerpeltz, 1925: 90 (Xantholinus; subgenus Gyrohypnus; characters).

  • — Scheerpeltz, 1926: 92 (Xantholinus; subgenus Gyrohypnus; characters in key).

  • — Portevin, 1929: 371 (Xantholinus; characters; France).

  • — Bernhauer, 1931d: 583 (Xantholinus; Abyssinia).

  • — Cameron, 1932: 24 (Xantholinus; characters; East Indies).

  • — Tottenham, 1939d: 230 (Xantholinus; synonym of fracticornis).

  • — C. Koch, 1941: 61 (Xantholinus; subgenus Gyrohypnus; Morocco).

  • — Méquignon, 1942a: 19 (Gyrohypnus; Azores).

  • — Scheerpeltz and Höfler, 1948: 156 (Xantholinus; subgenus Gyrohypnus; fungus inhabitant).

  • — V. Hansen, 1952: 16 (Xantholinus; subgenus Gyrohypnus; characters; Denmark).

  • — Coiffait, 1956a: 54 (Gyrohypnus; characters in key).

  • — Lohse, 1958b: 120 (Gyrohypnus; identification).

  • — Smetana, 1958: 96 (Gyrohypnus; synonym of fracticornis).

  • — Palm, 1959: 22 (Gyrohypnus; characters; illustrations).

  • — Scheerpeltz, 1960d: 135 (Gyrohypnus; Portugal).

  • — Scheerpeltz, 1961d: 126 (Gyrohypnus; Italy).

  • — Coiffait, 1962b: 391 (Gyrohypnus; synonym of fracticornis).

  • — Scheerpeltz, 1963a: 419 (Gyrohypnus; Greece).

  • — Palm, 1963b: 47 (Gyrohypnus; characters; Sweden).

  • — Lohse, 1964: 161 (Gyrohypnus; characters; central Europe).

  • — Smetana, 1964d: 72 (Gyrohypnus; Czechoslovakia).

  • — Raianu, 1964: 104 (Xantholinus; Romania).

  • — Horion, 1965: 92 (Gyrohypnus; all Europe).

  • — Pototskaia, 1967: 47 (Gyrohypnus; larval characters).

  • — Szujecki, 1968a: 725 (Gyrohypnus; Poland).

  • — Saiz, 1970b: 388 (Gyrohypnus; synonym of fracticornis).

  • — Coiffait, 1972: 182 (Gyrohypnus; synonym of fracticornis; notes).

  • — Pope, 1977: 28 (Gyrohypnus; Britain).

  • — Smetana, 1978c: 338 (Gyrohypnus; distinct from fracticornis; name preoccupied but should be conserved; notes; characters).

  • — Muona, 1979: 16 (Gyrohypnus; synonym of ater).

  • — Bordoni, 1982: 214 (Gyrohypnus; characters; Italy).

  • — Frank, 1982: 31 (Gyrohypnus; list of parasites).

  • — Smetana, 1982: 192 (Gyrohypnus; not in North America).

  • — Lucht, 1987: 103 (Gyrohypnus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Lohse, 1988: 41, 49 (Gyrohypnus; synonym of liebei).

  • — Tóth, 1989: 17 (Gyrohypnus; characters; Hungary).

  • — Lohse and Lucht, 1989: 156 (Gyrohypnus; synonym of liebei).

  • — Nowosad, 1990: 89 (Gyrohypnus; collected from mole nest; notes; Poland).

  • — Silfverberg, 1992: 18 (Gyrohypnus; synonym of liebei).

  • — Janák, 1993a: 47 (Gyrohypnus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 25 (Gyrohypnus; synonym of liebei).

  • — M. Hansen, 1996: 109 (Gyrohypnus; synonym of liebei).

  • — Owen, 1999b: 61 (Gyrohypnus; cited as valid species; captured underground at roots of old trees; Britain).

  • — Kashcheev, 1999a: 57 (Xantholinus; Kazakhstan).

  • — Owen, 2000: 251 (Gyrohypnus; pitfall trap collections from underground at roots of trees; Britain).

longicollis Stephens, 1833, see: Phacophallus.

luteiventris Casey, 1906, see: Neohypnus.

macilentus Casey, 1906, see: Neohypnus.

macrocephalus Nordmann, 1837, see: Atrecus.

marginalis Wollaston, 1862: 187 (Xantholinus; Type locality: Risco at the north of Lanzarote; island of Lobos, off the extreme north of Fuerteventura).

  • — Wollaston, 1864: 578 (Xantholinus; notes; Canary Islands).

  • — Wollaston, 1865: 496 (Xantholinus; habitat; notes; Canary Islands).

  • — Fauvel, 1902b: 101 (Xantholinus; Canary Islands).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • — Coiffait, 1972: 187 (Gyrohypnus; subgenus Xenoxantholinus; characters; references; Canary Islands).

  • — Bordoni, 1976c: 100 (Gyrohypnus; Canary notes; Islands).

  • Distribution: Canary Islands.

melanarius Fauvel, 1872, see: fracticornis O. Müller, 1776.

melanops Casey, 1906, see: Neohypnus.

merdarius Nordmann, 1837, see: Megalinus.

meridionalis Nordmann, 1837, see: Xantholinus.

micans Casey, 1906, see: Hypnogyra.

mollinus Casey, 1906, see: Neohypnus.

morio Reitter, 1872, see: fracticornis O. Müller, 1776.

nigriceps Mannerheim, 1830, see: Atrecus.

nitidicollis Reitter, 1908, see: angustatus Stephens, 1833.

obscurus Stephens, 1833, see: fracticornis O. Müller, 1776.

obsidianus Melsheimier, 1844, see: fracticornis O. Müller, 1776.

ochraceus Gyllenhal, 1810, see: angustatus Stephens, 1833.

ochripennis Eppelsheim, 1892: 334 (Xantholinus; Type locality: Taschkent; Margelan).

  • — Reitter, 1908: 115 (Xantholinus; subgenus Gyrohypnus; characters; Turkestan; Aulie-Ata; Tashkent).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1313 (Xantholinus; catalog).

  • — Pototskaia, 1967: 47 (Gyrohypnus; larval characters).

  • — Coiffait, 1972: 186 (Gyrohypnus; characters; references; Caucasus; southern Russia; Turkestan).

  • — Smetana 1975c: 168 (Gyrohypnus; Mongolia).

  • Distribution: Mongolia, Uzbekistan, Kazakhstan, Russia, Caucasus.

ochropterus Nordmann, 1837, see: Leptophius.

oregonus Casey, 1906, see: Neohypnus.

pallipennis Notman, 1921, see: Xantholinus.

peyerimhoffi Coiffait, 1956a: 54 (Gyrohypnus; subspecies of angustatus; Type locality: Mers el Kabir).

  • — Coiffait, 1962b: 391 (Gyrohypnus; valid species; characters; North Africa).

  • — Coiffait, 1972: 185 (Gyrohypnus; characters; references; North Africa).

  • Distribution: Algeria.

picipes Thomson, 1860, see: atratus Heer, 1839.

procerulus Stephens, 1833, see: Leptacinus.

protractus Casey, 1906, see: Neohypnus.

punctulatus Paykull, 1789, see: liebei Scheerpeltz, 1926.

pusillus Stephens, 1833, see: Leptacinus.

quadratus Stephens, 1833, see: Xantholinus.

quadrisulcus Stephens, 1833, see: Leptacinus.

rotundicollis Stephens, 1833, see: Hypnogyra.

scoticus Joy, 1913, see: angustatus Stephens, 1833.

semistriatus Stephens, 1833, see: Leptacinus.

sichuanensis Zheng, 1995a: 220, 221 (Gyrohypnus; Type locality: Sichuan: Wolong, 1920 m).

  • — Bordoni, 2000: 130 (Gyrohypnus; characters; China).

  • Distribution: China.

silvanus Peyerimhoff, 1917: 120 (Xantholinus; subgenus Gyrohypnus; Type locality: Mont Babor; Edough; Ain-Draham).

  • — Scheerpeltz, 1933: 1314 (Xantholinus; catalog).

  • — Coiffait, 1956a: 55 (Gyrohypnus; characters in key; Algeria; Tunisia).

  • — Coiffait, 1962b: 391, 392 (Gyrohypnus; characters; Algeria; Tunisia).

  • — Coiffait, 1972: 186 (Gyrohypnus; characters; references; Algeria; Tunisia).

  • Distribution: Algeria, Tunisia.

sulcifrons Stephens, 1833, see: Leptacinus.

tenuicornis Nordmann, 1837, see: Linosomus.

thomsoni Schwarz, 1872, see: angustatus Stephens, 1833.

vernicatus Casey, 1906, see: Hypnogyra.

wagneri Scheerpeltz, 1926: 82, 92 (Xantholinus; subgenus Gyrohypnus; Type locality: Ufer des Guadalaviar in der Umgebung von Albarracin in Aragonien, ca. 1200 m).

  • — Scheerpeltz, 1933: 1315 (Xantholinus; catalog).

  • — Coiffait, 1972: 185 (Gyrohypnus; characters; Spain).

  • Distribution: Spain.

gredensis Coiffait, 1964a: 504 (Gyrohypnus; subspecies of wagneri; Type locality: Espagne, Prov. de Avila, Sierra de Gredos).

  • — Coiffait, 1972: 185 (Gyrohypnus; subspecies of wagneri; characters; Spain).

wutaishanensis Bordoni, 2000: 129 (Gyrohypnus; Type locality: N. China: Shaanxi, Wutaishan).

  • Distribution: China.

Habrolinus [8 species; Nearctic Region]

Habrolinus Casey, 1906: 369, 406 (species included: tahoensis). Type species: Habrolinus tahoensis Casey, fixed by original designation and monotypy.

  • — Bernhauer and Schubert, 1914: 295 (world catalog; 1 species).

  • — Leng, 1920: 106 (catalog of North American species).

  • — Blackwelder, 1952: 178 (type species: tahoensis).

  • — Moore, 1963a: 94 (characters).

  • — Moore and Legner, 1975: 74 (catalog; North America north of Mexico).

  • — Moore and Legner, 1979: 57 (characters; notes).

  • — Smetana, 1982: 134 (characters; habitus; revision of and key to species of North America north of Mexico).

  • — Smetana, 1988a: 530 (characters; key to species).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 390 (8 Nearctic species; notes; characters in key).

Timagenes Smetana, 1982: 139 (species included: longipes, brevicornis). Type species: Timagenes longipes Smetana, fixed by original designation.

  • — Smetana, 1988a: 530 (subgenus of Habrolinus).

abdominalis Smetana, 1988a: 537 (Habrolinus; subgenus Habrolinus; Type locality: California: Butte Co.: 4.4 mi SW Rackerby).

  • Distribution: USA.

andrewsi Smetana, 1988a: 540 (Habrolinus; subgenus Timagenes; Type locality: California: Whiskey Town, Shasta Co. Also cited from Oregon).

  • Distribution: USA.

brevicornis Smetana, 1982: 142 (Timagenes; Type locality: California: Plumas Co.: 7 mi. NE Bucks Lk.).

  • — Smetana, 1988a: 539 (Habrolinus; subgenus Timagenes; characters; California).

  • Distribution: USA.

californicus Smetana, 1988a: 533 (Habrolinus; subgenus Habrolinus; Type locality: California: Fresno Co.: 7 mi. SW Auberry).

  • Distribution: USA.

dentifer Smetana, 1982: 137 (Habrolinus; Type locality: California: San Mateo Co.: S. Bruno Hills. Also cited from Oregon).

  • — Smetana, 1988a: 536 (Habrolinus; subgenus Habrolinus; characters).

  • Distribution: USA.

hardyi Smetana, 1988a: 534 (Habrolinus; subgenus Habrolinus; Type locality: California: Butte Co.: 4.4 mi SW Rackerby. Also cited from Oregon).

  • Distribution: USA.

longipes Smetana, 1982: 140 (Timagenes; Type locality: California: Tahoe Pines).

  • — Smetana, 1988a: 538 (Habrolinus; subgenus Timagenes; characters; Calfornia; Oregon).

  • Distribution: USA.

tahoensis Casey, 1906: 406 (Habrolinus; Type locality: California: Lake Tahoe; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 295 (Habrolinus; catalog).

  • — Smetana, 1982: 135 (Habrolinus; lectotype designation; characters; California).

  • — Smetana, 1988a: 532 (Habrolinus; subgenus Habrolinus; characters; California).

  • Distribution: USA.

Hesperolinus [2 species; Neotropical and Nearctic Regions]

Hesperolinus Casey, 1906: 371, 411 (species included: parcus, bicolor, alticola, borealis, pomonae, angustus, piceus). Type species: Leptolinus parcus LeConte, fixed by subsequent designation by R. Lucas, 1920: 324.

  • — Bernhauer and Schubert, 1914: 296 (world catalog; 7 species).

  • — Leng, 1920: 106 (catalog of North American species).

  • — Blackwelder, 1952: 183 (type species).

  • — Hatch, 1957: 234, 239 (characters; key to species of Pacific Northwest).

  • — Moore, 1963a: 95 (characters).

  • — Moore and Legner, 1975: 75 (catalog; North America north of Mexico).

  • — Moore and Legner, 1979: 58 (characters; notes).

  • — Smetana, 1982: 183 (characters; habitus; revision of species of North America north of Mexico).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 390 (1 Nearctic species; notes; characters in key).

Leiolinus Casey, 1906: 371, 416 (species included: uteanus, tarsalis). Type species: Leiolinus tarsalis Casey, fixed by subsequent designation by Blackwelder, 1952: 212 [Note: Blackwelder stated that R. Lucas, 1920: 365, designated the type species but the designation was ambiguous because two nominal species were cited as “tarsalis et uteanus”].

  • — Bernhauer and Schubert, 1914: 297 (world catalog; 2 species).

  • — Leng, 1920: 106 (catalog of North American species).

  • — Moore, 1963a: 95 (characters).

  • — Moore and Legner, 1975: 79 (catalog; North America north of Mexico).

  • — Moore and Legner, 1979: 61 (characters; notes).

  • — Smetana, 1982: 183 (synonym of Hesperolinus).

alticola Casey, 1906, see: Linohesperus.

angustus Casey, 1906, see: Linohesperus.

bicolor Casey, 1906, see: Linohesperus.

borealis Casey, 1906, see: Linohesperus.

elytratus Coiffait, 1981g: 291 (Hesperolinus; Type locality: Ile Galapagos: Ile Santa Cruz entre les Cerros Biento et Puntudo, 450 à 500 m).

  • Distribution: Galapagos Islands.

parcus LeConte, 1863a: 41 (Leptolinus; Type locality: San Jose, California; [Note: See lectotype designation by Smetana, 1982]).

  • — Casey, 1906: 413 (Hesperolinus; characters; California).

  • — Bernhauer and Schubert, 1914: 297 (Hesperolinus; catalog).

  • — Hatch, 1957: 240 (Hesperolinus; characters; British Columbia; Idaho; Washington; Oregon).

  • — Smetana, 1982: 184 (Hesperolinus; lectotype designation; characters; Alberta; California; Nevada; Utah).

  • — Smetana, 1988a: 553 (Hesperolinus; collecting notes; California).

  • — Campbell and Davies, 1991: 124 (Hesperolinus; checklist of species; Canada).

  • Distribution: Canada, USA.

tarsalis Casey, 1906: 417 (Leiolinus; Type locality: Nevada, Elko).

  • — Bernhauer and Schubert, 1914: 297 (Leiolinus; catalog).

  • — Smetana, 1982: 184 (Hesperolinus; synonym of parcus).

uteanus Casey, 1906: 417 (Leiolinus; Type locality: southwestern Utah).

  • — Bernhauer and Schubert, 1914: 297 (Leiolinus; catalog).

  • — Smetana, 1982: 184 (Hesperolinus; synonym of parcus).

piceus Casey, 1906, see: Linohesperus.

pomonae Casey, 1906, see: Linohesperus.

tarsalis Casey, 1906, see: parcus LeConte, 1863.

uteanus Casey, 1906, see: parcus LeConte, 1863.

Heterocinus [6 species; Madagascan Region]

Heterocinus Jarrige, 1970: 44 (species included: pauliani, tenellus). Type species: Leptacinus tenellus Erichson, fixed by original designation.

andringranus Jarrige, 1978: 277 (Heterocinus; Type locality: Andringitra Est, Marositry, Est rivière Antsifotra, 2000 m).

  • Distribution: Madagascar.

betschi Jarrige, 1978: 278 (Heterocinus; Type locality: Andringitra Centre, plateau Andohariana, 2000–2100 m).

  • Distribution: Madagascar.

descarpentriesi Jarrige, 1978: 277 (Heterocinus; Type locality: Andringitra Sud, Andrianony, cirque Manjarivolo, 1800 m).

  • Distribution: Madagascar.

jarrigei Lecoq, 1996: 152 (Heterocinus; Type locality: l'archipel des Comores: Mohéli, Miringoni, 500 m).

  • Distribution: Comoro Islands.

pauliani Jarrige, 1970: 44 (Heterocinus; Type locality: Madagascar: Mt. Tsaratanana, 1500 m).

  • Distribution: Madagascar.

tenellus Erichson, 1839b: 336 (Leptacinus; Type locality: Madagascar; [Note: See lectotype designation by Janák, 1989]).

  • — Bernhauer and Schubert, 1914: 298 (Mitomorphus; catalog).

  • — Steel, 1949a: 269 (not in Mitomorphus; appears to be correctly placed in Leptacinus).

  • — Jarrige, 1970: 44 (Heterocinus; type species of genus).

  • — Janák, 1989: 167 (Leptacinus; subgenus Chaetocinus; lectotype designation; characters; Madagascar; [Note: This species is the type species for Heterocinus so by treating it as a member of Leptacinus Janák, in effect, synonymized Heterocinus with Leptacinus. However, since it is unclear that Janák's intention was any more than to study the type, designate a lectotype, and present characters for tenellus, I continue to treat Heterocinus as a valid genus and list tenellus therein]).

  • Distribution: Madagascar.

Heterolinus [2 species; Neotropical Region]

Heterolinus Coiffait, 1966, see: Xantholinus.

Heterolinus Sharp, 1885: 475 (species included: puncticeps). Type species: Xantholinus puncticeps Guérin-Méneville, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 309 (world catalog; 2 species).

  • — Blackwelder, 1944: 148 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 184 (type species: puncticeps).

distinctus Bernhauer, 1905: 17 (Heterolinus; Type locality: Peru: Chanchamajo).

  • — Bernhauer and Schubert, 1914: 309 (Heterolinus; catalog).

  • Distribution: Peru.

puncticeps Guérin-Méneville, 1844: 11 (Xantholinus; Type locality: Nouvelle Granade).

  • — Sharp, 1885: 475 (Heterolinus; notes; Costa Rica; Colombia).

  • — Bernhauer and Schubert, 1914: 309 (Heterolinus; catalog; Santiago).

  • Distribution: Colombia, Costa Rica.

Holocorynus [3 species; Oceanic Region]

Holocorynus Sharp, 1908: 550 (species included: discedens, subdepressus). Type species: Pachycorynus discedens Sharp, fixed by subsequent designation by R. Lucas, 1920: 331.

  • — Bernhauer and Schubert, 1914: 295 (world catalog; 2 species).

  • — Scheerpeltz, 1933: 1304 (world catalog supplement).

  • — Blackwelder, 1952: 187 (type species).

discedens Sharp, 1880: 50 (Pachycorynus; Type locality: Honolulu).

  • — Blackburn and Sharp, 1885: 222 (Pachycorynus; habitat; Hawaii).

  • — Sharp, 1908: 550 (Holocorynus; characters; Hawaii).

  • — Bernhauer and Schubert, 1914: 295 (Holocorynus; catalog).

  • — Nishida, 1994: 72 (Holocorynus; Hawaii).

  • — Nishida, 1997: 61 (Holocorynus; Hawaii).

  • Distribution: Hawaii.

longiceps Cameron, 1927a: 25 (Holocorynus; Type locality: Samoa: Upolu: Malololelei).

  • — Scheerpeltz, 1933: 1304 (Holocorynus; catalog).

  • Distribution: Samoa.

subdepressus Sharp, 1908: 551 (Holocorynus; Type locality: Kauai, Makaweli, 2500 ft.).

  • — Bernhauer and Schubert, 1914: 295 (Holocorynus; catalog).

  • — Nishida, 1994: 72 (Holocorynus; Hawaii).

  • — Nishida, 1997: 61 (Holocorynus; Hawaii).

  • Distribution: Hawaii.

Homalolinus [9 species; Neotropical Region]

Homalolinus Sharp, 1885: 472 (species included: canaliculatus, flavipennis, affinis, dilutus, apicalis, sanguineus, divisus). Type species: Xantholinus canaliculatus Erichson, fixed by subsequent designation by Casey, 1906: 374.

  • — Casey, 1906: 366, 374 (characters).

  • — Bernhauer and Schubert, 1914: 309 (world catalog; 8 species).

  • — Scheerpeltz, 1933: 1315 (world catalog supplement).

  • — Blackwelder, 1944: 148 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 189 (type species).

affinis Sharp, 1885: 473 (Homalolinus; Type locality: Guatemala, Purula, San Geronimo).

  • — Bernhauer and Schubert, 1914: 309 (Homalolinus; catalog).

  • Distribution: Guatemala.

amazonicus Sharp, 1876: 203 (Xantholinus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 300 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1315 (Homalolinus; catalog).

  • Distribution: Brazil.

apicalis Sharp, 1885: 474 (Homalolinus; Type locality: Mexico).

  • — Bernhauer and Schubert, 1914: 309 (Homalolinus; catalog).

  • Distribution: Mexico.

atronitens Casey, 1906: 374 (Homalolinus; Type locality: Guatemala).

  • — Bernhauer and Schubert, 1914: 309 (Homalolinus; catalog).

  • Distribution: Guatemala.

canaliculatus Erichson, 1839b: 307 (Xantholinus; Type locality: Bogatae in Columbia; Mexico; [Note: See lectotype designation by Smetana, 1977c]).

  • — Blanchard, 1842: 84 (Xantholinus; Mexico; Colombia; Bolivia).

  • — Solsky, 1868a: 141 (Xantholinus; Mexico).

  • — Solsky, 1875: 21 (Xantholinus; Peru).

  • — Sharp, 1885: 472 (Homalolinus; notes; Mexico; Guatemala; Costa Rica; Panama; Brazil).

  • — Bernhauer and Schubert, 1914: 309 (Homalolinus; catalog).

  • — Scheerpeltz, 1933: 1315 (Homalolinus; catalog; Argentina).

  • — Smetana, 1977c: 348 (Homalolinus; lectotype designation: Bogota; characters).

  • Distribution: Mexico, Guatemala, Costa Rica, Colombia, Brazil, Peru, Bolivia, Argentina.

dilutus Sharp, 1885: 473 (Homalolinus; Type locality: Mexico, Parada).

  • — Bernhauer and Schubert, 1914: 309 (Homalolinus; catalog).

  • Distribution: Mexico.

flavipennis Erichson, 1839b: 307 (Xantholinus; Type locality: Mexico).

  • — Sharp, 1885: 473 (Homalolinus; notes; Mexico).

  • — Bernhauer and Schubert, 1914: 310 (Homalolinus; catalog).

  • Distribution: Mexico.

guatemalenus Sharp, 1885, see: Ehomalolinus.

ruficollis Bernhauer, 1929h: 192 (Homalolinus; Type locality: Costa Rica: Palo Verde).

  • — Scheerpeltz, 1933: 1315 (Homalolinus; catalog).

  • Distribution: Costa Rica.

sanguineus Sharp, 1885, see: Ehomalolinus.

tripunctatus Bierig, 1934: 15 (Homalolinus; Type locality: Panama: Volcan de Chiriqui, 3500 p[ies]).

  • Distribution: Panama.

Homorocerus [1 species; Ethiopian Region]

Homorocerus Boheman, 1848: 272 (species included: rufipennis). Type species: Homorocerus rufipennis Boheman, fixed by monotypy.

  • — Lacordaire, 1854: 67 (characters; notes; list of species).

  • — Gemminger and Harold, 1868: 573 (catalog).

  • — Duvivier, 1883: 141 (catalog).

  • — Bernhauer and Schubert, 1916: 410 (world catalog; 1 species).

  • — Scheerpeltz, 1933: 1316 (world catalog supplement).

  • — Blackwelder, 1952: 191 (type species: rufipennis).

puncticollis Kraatz, 1858, see: Moeocerus.

robustus Gestro, 1881, see: Moeocerus.

rufipennis Boheman, 1848: 273 (Homorocerus; Type locality: Caffraria orientalis).

  • — Fauvel, 1899a: 32 (Homorocerus; Cafrerie orientale; Port Natal; Zululand).

  • — Bernhauer and Schubert, 1916: 410 (Homorocerus; catalog).

  • — Scheerpeltz, 1933: 1316 (Homorocerus; catalog).

  • — Bernhauer, 1947: 158 (Homorocerus; Deutsch-Ostafrika; Belgian Congo).

  • Distribution: South Africa, Tanzania, Congo.

spinulosus Solsky, 1864, see: Moeocerus.

Hypnogyra [9 species; Nearctic and Palaearctic Regions]

Hypnogyra Casey, 1906: 394 (subgenus of Gyrohypnus; species included: vernicatus, micans, gularis). Type species: Xantholinus gularis LeConte, fixed by subsequent designation by Blackwelder, 1943: 474.

  • — Bernhauer and Schubert, 1914: 300 (subgenus of Xantholinus).

  • — Cameron, 1932: 22 (subgenus of Xantholinus).

  • — Blackwelder, 1943: 473 (subgenus of Xantholinus; type species: gularis).

  • — Smetana, 1982: 274 (characters; habitus; revision of and key to species of North America north of Mexico).

  • — Angelini, 1991: 191 (in list of species from high elevations of Calabria, Italy).

  • — Silfverberg, 1992: 18 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Janák, 1993a: 47 (in list of species of Czech Republic and Slovakia).

  • — Ciceroni and Zanetti, 1995: 26 (in list of species of Italy).

  • — Downie and Arnett, 1996: 402 (characters; key to species of Northeastern North America).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 391 (2 Nearctic species; notes; characters in key).

Phalacrolinus Coiffait, 1972: 214 (species included: baicalensis, hoffmanni, laevissimus, glaber). Type species: Staphylinus glaber Gravenhorst, fixed by original designation.

  • — Bordoni, 1982: 228 (characters; habitus; species of Italy).

  • — Smetana, 1982: 274 (synonym of Hypnogyra).

  • — Janák, 1993a: 47 (synonym of Hypnogyra).

angularis Ganglbauer, 1895: 482 (Xantholinus; variety of glaber; Type locality: Not cited).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; synonym of glaber).

  • — Scheerpeltz, 1933: 1310 (Xantholinus; synonym of glaber).

  • — Tottenham, 1939d: 230 (Xantholinus; valid name and senior synonym of glaber Gravenhorst; notes).

  • — Smetana, 1958: 105 (Xantholinus; aberration of glaber).

  • — Coiffait, 1972: 215, 216 (Phalacrolinus; variety of glaber; characters; references).

  • — Bordoni, 1972: 208 (Xantholinus; synonym of glaber).

  • — Pope, 1977: 28 (Xantholinus; Britain).

  • — Bordoni, 1982: 230 (Phalacrolinus; synonym of glaber).

  • — Fowles, Alexander, and Key, 1999: 129 (Xantholinus; saproxylic species; status as endangered or threatened species; Britain).

  • Distribution: Tunisia, Algeria, Europe, Russia, Turkey, Japan.

glabra Gravenhorst, 1802: 46 (Staphylinus; [preoccupied; Note: Bernhauer and Schubert, 1914: 303, cited Nordmann's 1837 use of the name as the valid species but Nordmann attributed the species to Gravenhorst. Erichson, 1839a: 425, 426, cited in Xantholinus the two varieties of glaber that Gravenhorst, 1806: 100, described. This name is still treated as valid but it is preoccupied and was replaced by an older name, angularis, by Tottenham, 1939d: 230, and accepted by Pope, 1977: 28. There are two names older than that selected by Tottenham. However one, flavipennis, is preoccupied and the other, rotundicollis, Tottenham (op. cit.) thought was a synonym of angustatus. Subsequent workers have continued to list rotundicollis as a synonym of glaber but it is unclear whether they disagreed with Tottenham or overlooked his work. Since the identity of rotundicollis is unsettled it seems best to accept angularis as the valid name of the species]; Type locality: Not cited, but presumably Brunsvigae).

  • — Latreille, 1804: 330 (Staphylinus; characters; Germany).

  • — Gravenhorst, 1806: 100 (Staphylinus; characters).

  • — Panzer, 1808: 14 (Staphylinus; characters; illustrations).

  • — Nordmann, 1837: 114 (Gyrohypnus; [Note: Nordmann attributed this name to Gravenhorst, but Bernhauer and Schubert, 1914: 303, cited Nordmann as the author; Erichson, 1839a: 425, 426, cited in Xantholinus the two varieties of glaber that Gravenhorst, 1806: 100, described]; characters).

  • — Heer, 1839: 245 (Xantholinus; [Note: Heer cited this species as glaber variety 2 of Gravenhorst, 1806]; characters; Switzerland).

  • — Erichson, 1839a: 425 (Xantholinus; cited glaber Gravenhorst, 1806: 100 variation 2 as the reference; characters; Germany).

  • — Heer, 1839: 245 (Xantholinus; [Note: Heer cited this species as glaber variety 2 of Gravenhorst, 1806; see also Staphylinus glaber Gravenhorst]; characters; Switzerland).

  • — Erichson, 1839b: 325 (Xantholinus; characters; Germany).

  • — Märkel, 1845: 235 (Xantholinus; with Formica fuliginosa).

  • — Redtenbacher, 1849: 692 (Xantholinus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 501 (Xantholinus; characters; France).

  • — Kraatz, 1857d: 640 (Xantholinus; characters; Germany).

  • — Redtenbacher, 1857: 181 (Xantholinus; characters; Austria).

  • — Thomson, 1860: 192 (Xantholinus; characters; Scandinavia).

  • — Fauvel, 1873b: 102 [= 1873a: 388] (Xantholinus; characters; France; Scandinavia; Britain; Germany; Switzerland; Austria; Russia; Algeria).

  • — Redtenbacher, 1874: 219 (Xantholinus; characters; Austria).

  • — Seidlitz, 1875: 281 (Xantholinus; characters; Baltic region).

  • — Mulsant and Rey, 1877a: 267 (Xantholinus; characters; France).

  • — Fauvel, 1878: 117 [= 1878a: 37] (Xantholinus; Algeria).

  • — Fauvel, 1886: 49 [= 1886a: 41] (Xantholinus; Algeria).

  • — Fowler, 1888: 290 (Xantholinus; characters; Britain).

  • — Ganglbauer, 1895: 482 (Xantholinus; characters; north and middle Europe).

  • — Fauvel, 1902b: 101 (Xantholinus; Algeria; Tunisia).

  • — Reitter, 1908: 116 (Xantholinus; subgenus Metacyclinus; characters; northern and central Europe).

  • — Montandon, 1908: 79 (Xantholinus; Nordmann cited as author; Romania).

  • — Reitter, 1909: 137 (Xantholinus; subgenus Metacyclinus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; catalog).

  • — Johansen, 1914: 435 (Xantholinus; subgenus Xantholinus; characters; Denmark).

  • — Portevin, 1929: 372 (Xantholinus; characters; France).

  • — Scheerpeltz, 1933: 1310 (Xantholinus; catalog).

  • — Tottenham, 1939d: 230 (Xantholinus; synonym of angularis; discussion of nomenclature).

  • — V. Hansen, 1952: 17 (Xantholinus; subgenus Xantholinus; cited with Nordman as author; characters).

  • — Coiffait, 1956a: 70 (Xantholinus; subgenus Megalinus; cited with Nordmann as author of species; characters in key).

  • — Smetana, 1958: 105 (Xantholinus; subgenus Megalinus; characters; ecology; Czechoslovakia).

  • — Szujecki, 1963a: 33 (Xantholinus; cited as valid species with Nordmann as author; Poland).

  • — Palm, 1963b: 50 (Xantholinus; cited as glaber Nordmann; characters; Sweden; Finland).

  • — Lohse, 1964: 164 (Xantholinus; subgenus Megalinus; cited as valid species with Nordmann as author; characters; central Europe).

  • — Horion, 1965: 96 (Xantholinus; subgenus Megalinus; Greece; Turkey; Bosnia Hercegovina; Dalmatia; Italy; Spain; Russia; Poland; Czechoslovakia; Hungary; Romania; Switzerland; France; Belgium; Holland; Finland; Sweden; Britain; Austria; Germany).

  • — Szujecki, 1968a: 726 (Xantholinus; cited as valid species with Nordmann as author; Poland).

  • — Bordoni, 1971: 687 (Xantholinus; subgenus Megalinus; cited as valid species with Nordmann as author; Turkey).

  • — Coiffait, 1972: 216 (Phalacrolinus; cited as valid species with Gravenhorst as author; characters; references; Britain; Sweden; Asia Minor; North America).

  • — Bordoni, 1972: 163, 208 (Xantholinus; subgenus Megalinus; cited as valid species with Nordmann as author; characters; Italy).

  • — Bordoni, 1972c: 49 (Xantholinus; subgenus Megalinus; cited as valid species with Nordmann as author; Hungary).

  • — Bordoni, 1973c: 155 (Xantholinus; subgenus Megalinus; cited as valid species with Nordmann as author; Romania).

  • — Shibata, 1974: 33 (Megalinus; cited as valid with Nordmann as author; Japan).

  • — Osella and Zanetti, 1975: 106 (Phalacrolinus; cited as valid species with Nordmann as author; collected from nest of Talpa europaea; notes; Italy).

  • — Bordoni, 1975: 75 (Xantholinus; subgenus Megalinus; cited as valid species with Nordmann as author; notes; Ukraine).

  • — Bordoni, 1976c: 99 (Phalacrolinus; cited as valid species with Nordmann as author; France).

  • — Bordoni, 1976d: 122 (Phalacrolinus; cited as valid species with Nordmann as author; Netherlands).

  • — Legner and Moore, 1977: 176 (Hyponygrus; cited as valid species with Nordmann as author; found under bark).

  • — Pope, 1977: 28 (Xantholinus; synonym of angularis).

  • — Bordoni, 1982: 230 (Phalacrolinus; Nordmann cited as author; characters; Italy).

  • — Smetana, 1982: 274 (Hypnogyra; species not in North America).

  • — Dettner and Schwinger, 1986: 367 (Xantholinus; cited as valid species with Nordmann as author; defensive secretions).

  • — Segers, 1986: 40 (Phalacrolinus; checklist of species; Belgium).

  • — Ádám, 1987: 148 (Hypnogyra; cited as glabra Nordmann; collecting notes; Hungary).

  • — Lucht, 1987: 103 (Xantholinus; cited as glabra Nordmann; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Sweden).

  • — Schülke and Uhlig, 1988: 11 (Xantholinus; cited as valid species with Nordmann as author; notes; Germany).

  • — Tóth, 1989: 21 (Xantholinus; cited as valid species with Nordmann as author; characters; Hungary).

  • — Lohse and Lucht, 1989: 159 (Hypnogyra; cited as valid species with Nordmann as author).

  • — Ciceroni, 1993: 72 (Hypnogyra; cited as valid species with Nordmann as author; Italy).

  • — Wittwer, 1993: 249 (Xantholinus; cited as valid species with Nordmann as author; Switzerland).

  • — Janák, 1993a: 47 (Hypnogyra; cited as valid species with Nordmann as author; Czech Republic; Slovakia).

  • — Ciceroni, 1994: 116 (Hypnogyra; cited as a valid species with Nordmann as author).

  • — Ciceroni and Zanetti, 1995: 26 (Hypnogyra; cited as a valid species with Nordmann as author; Italy).

  • — Assing, Frisch, Kahlen, et al., 1998: 136 (Hypnogyra; cited as valid species with Nordmann as author; note).

  • — Jansson and Lundberg, 2000: 93 (Hypnogyra; reported from Sweden).

rotundicollis Stephens, 1833: 259 (Gyrohypnus; Type locality: Bristol; Suffolk).

  • — Stephens, 1839: 403 (Xantholinus; characters; England).

  • — G. Waterhouse, 1858: 26 (Xantholinus; synonym of glaber).

  • — Ganglbauer, 1895: 482 (Xantholinus; synonym of glaber).

  • — Reitter, 1908: 116 (Xantholinus; synonym of glaber).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; synonym of glaber).

  • — Tottenham, 1939d: 230 (Xantholinus; synonym of angustatus).

  • — Smetana, 1958: 105 (Xantholinus; synonym of glaber).

  • — Coiffait, 1972: 216 (Phalacrolinus; synonym of glaber).

  • — Bordoni, 1972: 208 (Xantholinus; synonym of glaber).

flavipennis Redtenbacher, 1849: 692 (Xantholinus; [preoccupied]; Type locality: Austria).

  • — Kraatz, 1857d: 640 (Xantholinus; synonym of glaber).

  • — Redtenbacher, 1857: 181 (Xantholinus; characters; Austria).

  • — Fauvel, 1867c: 113 [= 1868b: 212] (Xantholinus; synonym of glaber).

  • — Redtenbacher, 1874: 219 (Xantholinus; characters; Austria).

  • — Mulsant and Rey, 1877a: 268 (Xantholinus; synonym of glaber).

  • — Ganglbauer, 1895: 482 (Xantholinus; synonym of glaber).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; synonym of glaber).

  • — Scheerpeltz, 1933: 1310 (Xantholinus; synonym of glaber).

  • — Tottenham, 1939d: 230 (Xantholinus; synonym of angularis).

  • — Smetana, 1958: 105 (Xantholinus; synonym of glaber).

  • — Coiffait, 1972: 216 (Phalacrolinus; synonym of glaber).

  • — Bordoni, 1972: 208 (Xantholinus; synonym of glaber).

  • — Bordoni, 1982: 230 (Phalacrolinus; synonym of glaber).

ornata Smetana, 1948: 35 (Xantholinus; aberration of glaber; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; Type locality: Bohemia, Hrádek).

  • — Smetana, 1958: 105 (Xantholinus; aberration of glaber).

  • — Coiffait, 1972: 215, 217 (Phalacrolinus; variety of glaber; characters; references).

bernaboi Coiffait, 1980: 251 (Lepidophallus; Type locality: Italie: Castelleto d'Orba, province d'Alexandrie).

  • — Ciceroni, 1994: 116 (Hypnogyra; synonym of glabra).

baicalensis Fauvel, 1875a: XXV [= 1875b: 227] (Xantholinus; Type locality: lac Baïkal; [Note: See lectotype designation by Bordoni, 1999]).

  • — Reitter, 1908: 116 (Xantholinus; subgenus Metacyclinus; characters; Lake Baikal).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1308 (Xantholinus; catalog).

  • — Coiffait, 1972: 218 (Phalacrolinus; characters; references; Lake Baikal).

  • — Bordoni, 1975: 84 (Xantholinus; characters; notes; Baikal).

  • — Bordoni, 1999: 5 (Xantholinus; lectotype designation).

  • Distribution: Russia.

bernaboi Coiffait, 1980, see: angularis Ganglbauer, 1895.

glabra of Nordmann, 1837 [Note: Nomen nudum], see: angularis Ganglbauer, 1895 and “Note” for glaber Gravenhorst.

glabra Gravenhorst, 1802, see: angularis Ganglbauer, 1895.

glabroides Coiffait, 1962c: 74 (Xantholinus; subspecies of glaber; Type locality: Tunisie: El Fedja).

  • — Coiffait, 1962b: 398, 404, 409 (Xantholinus; subgenus Megalinus; subspecies of glaber; characters; Tunisia).

  • — Coiffait, 1972: 215, 217 (Phalacrolinus; subspecies of glaber; characters; references).

  • — Bordoni, 1972: 163 (Xantholinus; subgenus Megalinus; subspecies of glaber).

  • — Bordoni, 1972b: 128 (Xantholinus; subgenus Megalinus; valid species; characters; notes; Tunisia).

  • — Bordoni, 1976c: 100 (Phalacrolinus; valid species; notes; characters).

  • Distribution: Tunisia.

gularis LeConte, 1880: 173 (Xantholinus; Type locality: Detroit, Michigan).

  • — Casey, 1906: 395 (Gyrohypnus; characters; Michigan).

  • — Blatchley, 1910: 397 (Xantholinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Legner and Moore, 1977: 176 (Hyponygrus; found under bark).

  • — Smetana, 1982: 275 (Hypnogyra; characters; Illinois; Indiana; Iowa; Maine; Massachusetts; Michigan; New Hampshire; New Jersey; New York; Pennsylvania; Rhode Island).

  • — Smetana, 1988a: 558 (Hypnogyra; Ontario; Pennsylvania).

  • — Smetana, 1990c: 87 (Hypnogyra; Ontario; Illinois; Michigan; New Hampshire).

  • — Campbell and Davies, 1991: 124 (Hypnogyra; checklist of species; Canada).

  • — Downie and Arnett, 1996: 402 (Hypnogyra; characters; USA).

  • Distribution: USA, Canada.

vernicata Casey, 1906: 395 (Gyrohypnus; subgenus Hypnogyra; Type locality: Iowa: Iowa City; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • — Smetana, 1982: 275 (Hypnogyra; lectotype designation; synonym of gularis).

flavipennis Redtenbacher, 1849, see: angularis Ganglbauer, 1895.

hoffmanni Bernhauer, 1928b: 11 (Xantholinus; Type locality: Tientsin).

  • — Scheerpeltz, 1933: 1310 (Xantholinus; catalog).

  • — Coiffait, 1972: 214 (Phalacrolinus; characters; China).

  • — Bordoni, 2000: 124 (Hypnogyra; characters; China).

  • Distribution: China.

micans Casey, 1906: 395 (Gyrohypnus; subgenus Hypnogyra; Type locality: Ohio, Cincinnati; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • — Smetana, 1982: 277 (Hypnogyra; lectotype designation; characters; Illinois; Missouri; Ohio; Pennsylvania; Tennessee; Texas).

  • — Smetana, 1988a: 558 (Hypnogyra; Mississippi).

  • — Smetana, 1990c: 87 (Hypnogyra; Arkansas; Indiana; Michigan; Ohio; Oklahoma).

  • — Downie and Arnett, 1996: 402 (Hypnogyra; characters; USA).

  • Distribution: USA.

monserrati Outerelo, 1976a: 254 (Phalacrolinus; Type locality: San Lorenzo del Escorial en la Sierra de Guadarrama, 1100 m. Also includes larval description).

  • Distribution: Spain.

ornata Smetana, 1948 [Note: Nomen nudum], see: angularis Ganglbauer, 1895.

rotundicollis Stephens, 1833, see: angularis Ganglbauer, 1895.

tenebrosa Bordoni, 1997: 174 (Hypnogyra; Type locality: Japan, Yokohama).

  • Distribution: Japan.

tubula Sharp, 1889: 250 (Xantholinus; Type locality: Miyanoshita; [Note: See lectotype designation by Bordoni, 1997]. Also cited from Nikko).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • — Nakane, 1963a: 90 (Megalinus; characters; habitus photograph; Japan).

  • — Watanabe, 1996a: 12 (Xantholinus; cave collection; Japan).

  • — Bordoni, 1997: 172 (Hypnogyra; lectotype designation: Miyanoshita, Japan; characters; Japan).

  • Distribution: Japan.

vernicata Casey, 1906, see: gularis LeConte, 1880.

Kainolinus [1 species; Neotropical Region]

Kainolinus Coiffait and Saiz, 1964: 511, 519 (species included: approbabilis). Type species: Kainolinus approbabilis Coiffait and Saiz, fixed by original designation and monotypy.

  • — Coiffait and Saiz, 1968: 345 (list of Chilean species).

  • — Saiz, 1970b: 382, 385 (characters).

approbabilis Coiffait and Saiz, 1964: 521 (Kainolinus; Type locality: Chile: Santiago).

  • — Saiz, 1970b: 385 (Kainolinus; Chile).

  • Distribution: Chile.

Lemiganus [1 species; Palaearctic Region]

Lemiganus Bordoni, 1985a: 86 (species included: elianae). Type species: Xantholinus elianae Jarrige, fixed by original designation and monotypy.

elianae Jarrige, 1941: 47 (Xantholinus; Type locality: France: Dorres, Cerdagne, alt. 1500 m).

  • — Coiffait, 1956a: 59 (Xantholinus; subgenus Leptophallus; characters in key; Spain).

  • — Coiffait, 1972: 258 (Xantholinus; subgenus Leptophallus; characters; references; France; Spain).

  • — Bordoni, 1972: 160 (Xantholinus; subgenus Leptophallus).

  • — Bordoni, 1985a: 85 (Lemiganus; anatomical studies of labrum, aedeagus, and male genital segment).

  • Distribution: France, Spain.

Lepidophallus [13 species; Neotropical and Palaearctic Regions]

Lepidophallus Coiffait, 1956a: 58, 59 (subgenus of Xantholinus; species included: hesperius, pseudohesperius). Type species: Xantholinus hesperius Erichson, fixed by original designation.

  • — Coiffait, 1962b: 389, 392 (characters; valid genus; key to North African species).

  • — Coiffait and Saiz, 1964: 518 (key to Chilean species).

  • — Coiffait and Saiz, 1968: 345 (list of Chilean species).

  • — Saiz, 1970b: 383, 385 (characters).

  • — Coiffait, 1972: 171 (characters; type species: hesperius; key to species of western Palaearctic region; references).

  • — Bordoni, 1982: 232 (characters; habitus; key to species of Italy).

  • — Coiffait, 1982: 26 (valid genus; notes).

  • — Ciceroni and Zanetti, 1995: 26 (list of species of Italy).

bernaboi Coiffait, 1980, see: Hypnogyra.

boki Bordoni, 2000: 131 (Lepidophallus; Type locality: N. China: Shaanxi, Nan Wutai).

  • Distribution: China.

cinnamomeus Zheng, 1994a: 351, 352 (Lepidophallus; Type locality: China: Sichuan, Mt. Qincheng, Guanxian, 1000 m).

  • Distribution: China.

coloratus Karsch, 1881: 44 (Xantholinus; Type locality: Djebel Tarrhuna: Bir Milrha).

  • — Fauvel, 1902b: 101 (Xantholinus; synonym of hesperius).

  • — Reitter, 1908: 118 (Xantholinus; synonym of hesperius).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; synonym of hesperius).

  • — C. Koch, 1936a: 75 (Xantholinus; subspecies of hesperius; characters; Libya).

  • — Coiffait, 1972: 174 (Lepidophallus; valid species; characters; references; Libya).

  • — Bordoni, 1976c: 101 (Lepidophallus; notes; Tunisia; Algeria; Tripolitania).

  • — Bordoni, 1976d: 122 (Xantholinus; characters; notes; Algeria).

  • Distribution: Libya, Tunisia, Algeria.

coracinus Zheng, 1994a: 350, 352 (Lepidophallus; Type locality: China: Sichuan, Mt. Jin-yun, Chongqing, 800–919 m).

  • Distribution: China.

dahli Blackwelder, 1944: 147 (Xantholinus; replacement name for elongatus Fauvel).

  • — Saiz, 1970b: 385 (Lepidophallus; Chile).

  • Distribution: Chile, Argentina.

elongatus Fauvel, 1866a: 349 [= 1866b: 100 (Xantholinus; [preoccupied]; Type locality: Santiago).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • — Coiffait and Saiz, 1964: 518 (Lepidophallus; characters; Chile).

  • — Saiz, 1970b: 385 (Lepidophallus; synonym of dahli).

  • — Scheerpeltz, 1972: 16 (Gyrohypnus; Argentina).

elongatus Fauvel, 1866, see: dahli Blackwelder, 1944.

hesperius Erichson, 1839b: 329 (Xantholinus; Type locality: Andalusia; [Note: See lectotype designation by H. Franz, 1993]. Also cited from Ulyssipone, Lusitania).

  • — Rosenhauer, 1856: 70 (Xantholinus; notes).

  • — Wollaston, 1860: 100 (Xantholinus; characters; notes; Madeira).

  • — Wollaston, 1864: 578 (Xantholinus; notes; Canary Islands).

  • — Wollaston, 1865: 497 (Xantholinus; Madeira Islands; Canary Islands).

  • — Crotch, 1867a: 383 (Xantholinus; Azores).

  • — J. Sahlberg, 1876: 35 (Xantholinus; characters; Finland).

  • — Mulsant and Rey, 1877a: 282 (Xantholinus; characters; France).

  • — Fauvel, 1878: 117 [= 1878a: 37] (Xantholinus; Morocco; Algeria; Tunisia).

  • — Fauvel, 1886: 49 [= 1886a: 41] (Xantholinus; Morocco; Tunisia; Algeria).

  • — Fauvel, 1902b: 101 (Xantholinus; Barbary; Azores; Madeira; Canary Islands).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1310 (Xantholinus; catalog).

  • — Bernhauer, 1940c: 3 (Xantholinus; Azores).

  • — Méquignon, 1942a: 19 (Xantholinus; Azores).

  • — Coiffait, 1954: 164 (Xantholinus; Canary Islands).

  • — Coiffait, 1956a: 60 (Xantholinus; subgenus Lepidophallus; characters in key; Spain; Canaries; France).

  • — Coiffait, 1962b: 392 (Lepidophallus; characters; Morocco; Algeria; Tunisia).

  • — Coiffait, 1972: 173 (Lepidophallus; characters; references; France; Spain; Portugal; Morocco; Algeria; Tunisia; Canaries; Madeira).

  • — Bordoni, 1976c: 101 (Lepidophallus; notes; Spain; Malta; Italy; Algeria; Canary Islands; Azores).

  • — Bordoni, 1982: 238 (Lepidophallus; characters; Italy).

  • — Borges, 1990: Table II (Lepidophallus; Azores).

  • — H. Franz, 1993 (Xantholinus; subgenus Lepidophallus; characters; lectotype designation: Andalusia).

  • — Ciceroni, 1993: 72 (Lepidophallus; Italy).

  • — Hernández, Outerelo, and Gamarra, 1994: 190 (Lepidophallus; checklist of species; Canary Islands).

  • — Ciceroni and Zanetti, 1995: 26 (Lepidophallus; Italy).

  • Distribution: Tunisia, Algeria, Morocco, Canary Islands, Madeira, Azores, Malta, Portugal, Spain, France, Italy.

limbatus Waltl, 1835: 57 (Xantholinus; [Note: This name is older than hesperius]; Type locality: Andalusien).

  • — Erichson, 1840: 924 (Xantholinus; synonym of hesperius).

  • — Rosenhauer, 1856: 70 (Xantholinus; synonym of hesperius).

  • — Mulsant and Rey, 1877a: 282 (Xantholinus; synonym of hesperius).

  • — Reitter, 1908: 118 (Xantholinus; synonym of hesperius).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; synonym of hesperius).

  • — Coiffait, 1972: 173 (Lepidophallus; synonym of hesperius).

  • — Bordoni, 1982: 238 (Lepidophallus; synonym of hesperius).

japonicus Sharp, 1874: 52 (Xantholinus; Type locality: Japan [Note: See lectotype designation by Bordoni, 1997]. Originally cited from Simabara; Hiogo;).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Adachi, 1937: 55 (Xantholinus; characters; Izu-Oshima).

  • — Nakane, 1963a: 90 (Megalinus; characters; habitus photograph; Japan).

  • — Shibata, 1985: 289 (Xantholinus; characters; Japan).

  • — J. Li, 1993: 32 (Xantholinus; China).

  • — Bordoni, 1997: 176 (Lepidophallus; lectotype designation: Japan; characters; Japan).

  • Distribution: Japan, China.

limbatus Waltl, 1835, see: hesperius Erichson, 1839.

metallicus Fauvel, 1895: 243 (Xantholinus; Type locality: Bírmanie, Carin Asciuii Ghécu, 1400–1500 m).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • — Bernhauer, 1922c: 231 (Xantholinus; Formosa).

  • — Cameron, 1932: 26 (Xantholinus; characters; Burma; Assam; India).

  • — Scheerpeltz, 1933: 1312 (Xantholinus; catalog).

  • — Scheerpeltz, 1965: 105 (Xantholinus; Burma).

  • — Shibata, 1973a: 126 (Megalinus; Taiwan).

  • — Coiffait, 1982a: 231 (Xantholinus; India).

  • — Coiffait, 1982: 26 (Lepidophallus; characters; Nepal).

  • Distribution: India, Nepal, Myanmar, Taiwan.

mixtus Sharp, 1874: 53 (Xantholinus; Type locality: Nagasaki; [Note: See lectotype designation by Bordoni, 1997]).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • — Watanabe and Shibata, 1972: 63 (Megalinus; Yaku-shima Island).

  • — Bordoni, 1997: 177 (Lepidophallus; lectotype designation: Japan; characters; Japan).

  • Distribution: Japan.

pseudoelongatus Coiffait and Saiz, 1964: 519 (Lepidophallus; Type locality: Chile: Santiago).

  • — Saiz, 1970b: 385 (Lepidophallus; Chile).

  • Distribution: Chile.

pseudohesperius Reitter, 1908: 118 (Xantholinus; variety of hesperius; Type locality: Italy, Sicily, Spain, Portugal, Algeria).

  • — Scheerpeltz, 1933: 1310 (Xantholinus; variety of hesperius).

  • — Coiffait, 1956a: 60 (Xantholinus; subgenus Lepidophallus; valid species; characters in key; Tunisie; Algeria; Sicily).

  • — Coiffait, 1962b: 392 (Lepidophallus; characters; Morocco; Algeria; Tunisia).

  • — Coiffait, 1972: 174 (Lepidophallus; characters; North Africa).

  • — Coiffait, 1973a: 273 (Lepidophallus; Morocco).

  • — Bordoni, 1976c: 101 (Lepidophallus; notes; Morocco; Tunisia).

  • — Bordoni, 1982: 238 (Lepidophallus; characters; Italy).

  • — Sparacio, 1995: 158 (Lepidophallus; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 26 (Lepidophallus; Italy).

  • Distribution: Italy, Spain, Portugal, Morocco, Algeria, Tunisia.

suffusus Sharp, 1874: 52 (Xantholinus; Type locality: Nagasaki; [Note: See lectotype designation by Bordoni, 1997]).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • — Adachi, 1937: 56 (Xantholinus; Izu-Oshima).

  • — Cameron, 1949b: 467 (Xantholinus; Japan).

  • — Watanabe, 1961: 353 (Megalinus; Izu Islands).

  • — Nakane, 1963a: 90 (Megalinus; characters; habitus photograph; Japan).

  • — Watanabe and Shibata, 1972: 63 (Megalinus; Yaku-shima Island).

  • — J. Li, 1993: 32 (Xantholinus; China).

  • — Bordoni, 1997: 174 (Lepidophallus; lectotype designation: Japan, Nagasaki; characters; Japan).

  • — Watanabe, 1998: 313 (Lepidophallus; Japan).

  • Distribution: Japan, China.

zhenyuanensis Zheng, 1994a: 349, 352 (Lepidophallus; Type locality: China: Gansu, Shenyuan County).

  • Distribution: China.

Lepitacnus [1 species; Nearctic Region]

Lepitacnus Smetana, 1982: 103 (species included: pallidulus; habitus). Type species: Leptacinus pallidulus LeConte, fixed by original designation and monotypy.

  • — Lundgren, 1998: 50 (in list of species from Florida; records for other States listed for species).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 391 (1 Nearctic species; notes; characters in key).

pallidulus LeConte, 1880: 169 (Leptacinus; Type locality: Fort Yuma, California).

  • — Casey, 1906: 403 (Leptacinodes; characters; California).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; catalog).

  • — Smetana, 1982: 103 (Lepitacnus; characters; Arizona; California; Texas).

  • — Smetana, 1988a: 527 (Lepitacnus; collecting notes; Florida; Mississippi).

  • — Smetana, 1990c: 84 (Lepitacnus; Alabama; Arkansas; Indiana; Oklahoma).

  • Distribution: USA.

Leptacinus [153 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions]

Leptacinus Erichson, 1839a: 429 (species included: batychrus). Type species: Staphylinus batychrus Gyllenhal, fixed by monotypy.

  • — Heer, 1839: 243 (characters).

  • — Erichson, 1839b: 333 (characters).

  • — Stephens, 1839: 385 (characters).

  • — Redtenbacher, 1849: 693 (characters).

  • — Schaum, 1852: 25 (list of species; Europe).

  • — Lacordaire, 1854: 69 (characters; notes; list of species).

  • — Fairmaire and Laboulbène, 1856: 503 (characters).

  • — Jacquelin du Val, 1856: 32 (characters).

  • — Gistel, 1856: 388 (list of species).

  • — Kraatz, 1857d: 645 (characters).

  • — Redtenbacher, 1857: 182, 983 (characters; key to Austrian species).

  • — Thomson, 1858: 30 (characters).

  • — Thomson, 1859: 27 (characters; type species: parumpunctatus).

  • — Schaum, 1859: 27 (catalog; European species).

  • — Thomson, 1860: 192 (characters).

  • — LeConte, 1861a: 65 (characters).

  • — Gredler, 1863: 108 (list of species from Tirol).

  • — LeConte, 1863: 24 (list of North American species).

  • — Fauvel, 1866a: 351 [= 1866b: 102] (characters).

  • — Wencker and Silbermann, 1866: 30 (list of species; collecting notes; France).

  • — Gemminger and Harold, 1868: 605 (catalog).

  • — Jacquelin du Val, 1868: 66 (catalog; European species).

  • — Sharp, 1871d: 12 (list of British species).

  • — Fauvel, 1873b: 88 [= 1873a: 374] (characters; key to species).

  • — Fauvel, 1875a: XXIV [= 1875b: 226] (catalog).

  • — Redtenbacher, 1874: 221 (characters; key to Austrian species).

  • — Sharp, 1876: 203 (characters; notes).

  • — Mulsant and Rey, 1877a: 315 (characters; key to French species).

  • — Fauvel, 1878f: 242 (characters; notes).

  • — LeConte, 1880: 168 (characters; key to North American species).

  • — Heyden, 1880: 75 (list of species of Siberian region).

  • — Duvivier, 1883: 158 (catalog).

  • — Sharp, 1885: 489 (notes; Mexican and Central American species).

  • — Olliff, 1887: 473 (characters).

  • — Fowler, 1888: 292 (notes; key to British species).

  • — Seidlitz, 1891: 380 (key to some European species).

  • — Seidlitz, 1891a: 405 (key to some European species).

  • — Heyden, Reitter, and Weise, 1891: 105 (list of species of Europe and Caucasus).

  • — Heyden, 1893: 44 (list of species of Siberian region).

  • — Ganglbauer, 1895: 486 (characters; key to middle European species).

  • — Heyden, 1896: 33 (list of species of Siberian region).

  • — Casey, 1906: 368, 399 (characters).

  • — Heyden, Reitter, and Weise, 1906: 160 (list of species of Europe and Caucasus).

  • — Reitter, 1908: 109, 111 (characters; key to species).

  • — Reitter, 1909: 134 (characters).

  • — Blatchley, 1910: 398 (characters; key to Indiana species).

  • — Petri, 1912: 68 (list of species of Siebenbürgen).

  • — Bernhauer and Schubert, 1914: 292 (world catalog; 31 species).

  • — Johansen, 1914: 438 (characters; key to Danish species).

  • — Leng, 1920: 106 (catalog of North American species).

  • — Cameron, 1921a: 354, 404 (characters; key to Singapore species; catalog).

  • — Winkler, 1925: 370 (catalog for Palaearctic region).

  • — Cameron, 1925: 50 (catalog of Indian species).

  • — Porta, 1926: 87 (characters; key to species of Italy).

  • — Bruch, 1928a: 427 (catalog; Argentina).

  • — Portevin, 1929: 370 (key to French species).

  • — Roubal, 1930: 361 (catalog; Slovakia).

  • — Cameron, 1932: 6 (characters; key to species of British India).

  • — Joy, 1932: 128 (key to British species).

  • — Scheerpeltz, 1933: 1300 (world catalog supplement).

  • — Normand, 1935: 369 (list of species of Tunisia).

  • — Paulian, 1941: 206 (larval characters).

  • — Blackwelder, 1943: 493 (characters; type species: brevicornis; key to species of West Indies).

  • — Blackwelder, 1944: 146 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Kloet and Hincks, 1945: 165 (list of British species).

  • — Steel, 1949a: 269 (type species: batychrus).

  • — Tottenham, 1949: 369, 416 (type species: batychrus Gyllenhal; notes; checklist of British species).

  • — Cameron, 1950g: 31 (key to African species).

  • — Blackwelder, 1952: 214 (type species: batychrus).

  • — V. Hansen, 1952: 6, 8 (characters; key to Danish species).

  • — Coiffait, 1956a: 44 (characters; key to French species).

  • — Hatch, 1957: 233, 238 (characters; key to species of Pacific Northwest).

  • — Kocher, 1958: 138 (checklist of species; Morocco).

  • — Smetana, 1958: 86 (characters; habitus of one species; type species: batychrus; key to Czechoslovakian species).

  • — Smetana, 1959a: 208 (list of species collected in Albania).

  • — Ferreira, 1962a: 16 (list of Portuguese species).

  • — Moore, 1963a: 94 (characters).

  • — Székessy, 1963: 49 (characters).

  • — Palm, 1963b: 41 (characters; key to species of Sweden).

  • — Lohse, 1964: 158 (characters; key to central European species).

  • — Coiffait and Saiz, 1964: 510, 512 (characters; key to Chilean species).

  • — Pototskaia, 1967: 46 (larval characters).

  • — Coiffait, 1968: 128, 130 (characters; key to species of Gabon; notes).

  • — Coiffait and Saiz, 1968: 345 (list of Chilean species).

  • — Scheerpeltz, 1968a: 59 (catalog of Austrian species).

  • — Kasule, 1970: 74, 80 (larval characters).

  • — Saiz, 1970b: 382, 383 (characters; key to species of Chile).

  • — Coiffait, 1972: 134 (characters; larvae; type species: batychrus; key to species of western Palaearctic region; references).

  • — Bordoni, 1973j: 692 (list of species from islands near Sicily).

  • — Tikhomirova, 1973a: 181 (checklist of species of USSR).

  • — Moore and Legner, 1975: 80 (catalog; North America north of Mexico).

  • — Pope, 1977: 28 (list of British species).

  • — Topp, 1978: 311 (larval characters).

  • — Moore and Legner, 1979: 62 (characters; notes).

  • — Muona, 1979: 16 (list of Scandinavian species).

  • — Uhlig, 1979: 250 (some species collected in Germany).

  • — M. Dvoř, 1979: 114 (some species collected in Slovakia).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 5 (catalog; Poland).

  • — Uhlig and Vogler, 1981: 107 (list of some species collected in Germany; habitat and collecting notes).

  • — Spahr, 1981: 95 (references to specimens from amber and copal).

  • — Smetana, 1982: 92 (characters; habitus; revision of and key to species of North America north of Mexico).

  • — Bordoni, 1982: 174 (characters; habitus; key to species of Italy).

  • — Coiffait, 1982a: 239 (key to species of Himalayan region).

  • — Shibata, 1983: 72 (checklist of species of Japan).

  • — Hammond, 1984: 206 (checklist of species of Borneo).

  • — Shibata, 1986a: 122 (supplemental list of species from Taiwan).

  • — Muona and Viramo, 1986: 14 (list of species of northeastern Finland).

  • — Segers, 1986: 39 (checklist of species; Belgium).

  • — Ádám, 1987: 145 (list of six species with collecting notes; Hungary).

  • — Tóth, 1989: 10 (characters; key to species of Hungary).

  • — Lohse and Lucht, 1989: 156 (notes on species).

  • — Campbell and Davies, 1991: 123 (checklist of species; Canada).

  • — Silfverberg, 1992: 18 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Ciceroni, 1993: 69 (notes).

  • — Janák, 1993a: 47 (list of species of Czech Republic and Slovakia).

  • — Ciceroni, 1994: 99 (key to Italian species).

  • — Hernández, Outerelo, and Gamarra, 1994: 190 (checklist of species; Canary Islands).

  • — Ciceroni and Zanetti, 1995: 25 (list of species of Italy).

  • — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 28 (additional records to Danish list of species).

  • — M. Hansen, 1996: 108 (list of Danish species).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 242 (additions to list of Danish species).

  • — Downie and Arnett, 1996: 398 (characters; key to species of Northeastern North America).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 74 (list of species collected in 1997; Denmark).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 391 (3 Nearctic species; notes; characters in key).

Xanthophius Motschulsky, 1860a: 75 (species included: serpentarius). Type species: Xanthophius serpentarius Motschulsky, fixed by monotypy.

  • — Gemminger and Harold, 1868: 607 (catalog; cited as Xanthophyus).

  • — Duvivier, 1883: 159 (catalog; cited as Xanthophyus).

  • — Bernhauer and Schubert, 1914: 292 (synonym of Leptacinus).

  • — Cameron, 1932: 6 (synonym of Leptacinus).

  • — Blackwelder, 1943: 493 (synonym of Leptacinus; type species: serpentarius).

  • — Blackwelder, 1952: 405 (type species: serpentarius).

  • — Smetana, 1958: 86 (synonym of Leptacinus).

  • — Coiffait, 1972: 134 (synonym of Leptacinus).

  • — Smetana, 1982: 92 (synonym of Leptacinus).

Leptacinodes Casey, 1906: 369, 401 (species included: batychrus, flavipes, nigritulus, lustrans, pallidulus, perexilis). Type species: Staphylinus batychrus Gyllenhal, fixed by subsequent designation by Blackwelder, 1943: 493.

  • — Bernhauer and Schubert, 1914: 292 (synonym of Leptacinus).

  • — Leng, 1920: 106 (catalog of North American species).

  • — Cameron, 1932: 6 (subgenus of Leptacinus).

  • — Blackwelder, 1943: 493 (synonym of Leptacinus; type species: batychrus).

  • — Steel, 1949a: 269 (synonym of Leptacinus; type species: batychrus).

  • — Blackwelder, 1952: 214 (type species).

  • — Smetana, 1958: 86 (synonym of Leptacinus).

  • — Coiffait, 1972: 134 (synonym of Leptacinus).

  • — Smetana, 1982: 92 (synonym of Leptacinus).

Chaetocinus Clark, et al., 1972: 483 (subgenus of Leptacinus; species included: brunneus, delicatulus, grassei, major, microps, minutus, oculatus, vicinus). Type species: Leptacinus oculatus Coiffait, 1968, fixed by original designation [Note: This name was originally proposed by Coiffait, 1968, but he failed to designate a type species so the name was unavailable. The fixation of the type species by the 10 compilers of the 1968 Insecta section of the Zoological Record made Leptacinus (Chaetocinus) available. Their act is acceptable under Article 69.1 and they are the author of the subgenus (Article 50.1).

  • — Coiffait, 1968: 130, 135 (subgenus of Leptacinus; characters; [Note: Nomen nudum here, no type species fixation (Article 13.3)]).

abactus Tottenham, 1956: 240 (Leptacinus; Type locality: Urundi: Bururi, 2100 m).

  • Distribution: Burundi.

abessinus Bernhauer, 1931d: 581 (Leptacinus; Type locality: Abyssinia: Jem-Jem Forest, 8,000–9,000 ft.).

  • — Scheerpeltz, 1933: 1300 (Leptacinus; catalog).

  • — Cameron, 1950g: 29 (Leptacinus; Belgian Congo).

  • Distribution: Ethiopia, Congo.

adonis Coiffait, 1956, see: batychrus Gyllenhal, 1827.

aethiopicus Bernhauer, 1931d: 582 (Leptacinus; Type locality: Abyssinia: Jem-Jem Forest, 8,000–9,000 ft.).

  • — Scheerpeltz, 1933: 1300 (Leptacinus; catalog).

  • Distribution: Ethiopia.

aethiops Bernhauer, 1920d: 180 (Leptacinus; Type locality: Darjeeling: Sikkim).

  • — Cameron, 1932: 14 (Leptacinus; characters; India).

  • — Scheerpeltz, 1933: 1300 (Leptacinus; catalog).

  • — Coiffait, 1982a: 240 (Leptacinus; characters).

  • Distribution: India.

aethiops Cameron, 1933, see: cameroni Steel, 1948.

africanus Bernhauer, 1929f: 120 (Leptacinus; Type locality: Erythraea: Ghinda).

  • — Scheerpeltz, 1933: 1300 (Leptacinus; catalog).

  • Distribution: Eritrea.

ajax Tottenham, 1949c: 41 (Leptacinus; replacement name for persimilis Tottenham).

  • — Saiz, 1970b: 383, 384 (Leptacinus; characters; Chile).

  • Distribution: Chile.

persimilis Tottenham, 1939c: 223 (Leptacinus; [preoccupied]; Type locality: Chile: Valparaiso).

  • — Saiz, 1970b: 384 (Leptacinus; synonym of ajax).

algiricus Coiffait, 1972, see: faunus Coiffait, 1956.

ampliventris Jacquelin du Val, 1854, see: Phacophallus.

angustatus Grimm, 1845, see: sulcifrons Stephens, 1833.

angusticollis Kolenati, 1846, see: Neobisnius.

angustus Sharp, 1874: 54 (Xanthophius, cited as Xanthophyus; Type locality: Hiogo).

  • — Duvivier, 1883: 159 (Xanthophius; cited as angustatus; catalog).

  • — Bernhauer and Schubert, 1914: 292 (Leptacinus; cited as angustatus; catalog).

  • — Scheerpeltz, 1933: 1300 (Leptacinus; catalog).

  • — Watanabe, 1961: 353 (Leptacinus; Izu Islands).

  • — J. Li, 1993: 32 (Xantholinus; China).

  • Distribution: Japan, China.

anommatophilus Cameron, 1948a: 228 (Leptacinus; Type locality: Ivory Coast: Reserve du Banco).

  • Distribution: Ivory Coast.

antennalis Cameron, 1932: 13 (Leptacinus; Type locality: Burma: Ruby Mines).

  • — Scheerpeltz, 1933: 1300 (Leptacinus; catalog).

  • — Coiffait, 1982a: 240 (Leptacinus; characters).

  • Distribution: Myanmar.

apicalis Stephens, 1833, see: batychrus Gyllenhal, 1827.

apicalis Kolenati, 1846, see: batychrus Gyllenhal, 1827.

apicipennis Fairmaire and Germain, 1861: 434 (Leptacinus; Type locality: Santiago).

  • — Fauvel, 1866a: 352 [= 1866b: 103] (Leptacinus; characters; Chile).

  • — Bernhauer and Schubert, 1914: 292 (Leptacinus; catalog).

  • — Coiffait and Saiz, 1964: 514 (Leptacinus; characters; Chile).

  • — Saiz, 1970b: 384 (Leptacinus; characters; Chile).

  • Distribution: Chile.

apicipennis Bernhauer, 1904, see: eggersi Bernhauer and Schubert, 1914.

aristaeus Coiffait, 1956, see: batychrus Gyllenhal, 1827.

armeniacus Coiffait, 1966f: 196 (Leptacinus; subgenus Leptacinus; Type locality: SSSR, Armeniia, okrestnosti Erevana).

  • — Coiffait, 1972: 149 (Leptacinus; characters; references; Armenia).

  • Distribution: Armenia.

assamensis Coiffait, 1982a: 236 (Leptacinus; Type locality: Assam: Kaziranga, 75 m).

  • Distribution: India.

baronii Coiffait, 1982a: 235 (Leptacinus; Type locality: Assam: Kaziranga, 75 m).

  • Distribution: India.

basalis Aubé, 1850, see: Zeteotomus.

basipennis Bernhauer, 1942c: 361 (Leptacinus; Type locality: Mont Cameroun: 1800–2000 m).

  • Distribution: Cameroon.

batychrus Gyllenhal, 1827: 480 (Staphylinus; Type locality: Suecia).

  • — C. Sahlberg, 1830: 332 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 33 (Gyrohypnus; Sweden).

  • — Mannerheim, 1831: 447 (Gyrohypnus; Sweden).

  • — Ménétriés, 1832: 147 (Gyrohypnus; Caucasus-Persia region).

  • — Stephens, 1833: 263 (Gyrohypnus; characters; England).

  • — Runde, 1835: 11 (Gyrohypnus; characters; Germany).

  • — Nordmann, 1837: 117 (Gyrohypnus; Sweden; Germany; France).

  • — Erichson, 1839a: 429 (Leptacinus; characters; Germany).

  • — Heer, 1839: 243 (Leptacinus; characters; Switzerland).

  • — Erichson, 1839b: 335 (Leptacinus; characters; Europe; Asia minor).

  • — Stephens, 1839: 404 (Leptacinus; characters; England).

  • — Laporte, 1840: 180 (Gyrohypnus; characters; France).

  • — Jacobsen, 1849: 680 (Leptacinus; characters; notes; Denmark).

  • — Hochhuth, 1849: 110 (Leptacinus; characters).

  • — Redtenbacher, 1849: 693 (Leptacinus; characters; Austria).

  • — Hardy, 1851: 39 (Leptacinus; notes; England).

  • — Rosenhauer, 1856: 70 (Leptacinus; Xeriz).

  • — Fairmaire and Laboulbène, 1856: 503 (Leptacinus; characters; France).

  • — Baudi, 1857: 99 (Leptacinus; characters; Italy).

  • — Kraatz, 1857d: 649 (Leptacinus; characters; Germany).

  • — Redtenbacher, 1857: 183 (Leptacinus; characters; Austria).

  • — Thomson, 1860: 193 (Leptacinus; characters; Scandinavia).

  • — Hochhuth, 1862: 69 (Leptacinus; Russia).

  • — Fauvel, 1865a: 298 [= 1865b: 48] (Xantholinus; habitat; France).

  • — Fauvel, 1873b: 89 [= 1873a: 375] (Leptacinus; characters; France; Georgia; Madeira; Iran; Canaries).

  • — Redtenbacher, 1874: 221 (Leptacinus; characters; Austria).

  • — Seidlitz, 1875: 280 (Leptacinus; characters; Baltic region).

  • — J. Sahlberg, 1876: 37 (Leptacinus; Finland).

  • — Mulsant and Rey, 1877a: 321 (Leptacinus; adult and larval characters; France).

  • — Fauvel, 1878: 116 [= 1878a: 36] (Leptacinus; Morocco; Algeria; Tunisia).

  • — Fauvel, 1886: 48 [= 1886a: 40] (Leptacinus; Algeria).

  • — Fowler, 1888: 293 (Leptacinus; characters; Britain).

  • — Eppelsheim, 1892: 334 (Leptacinus; Turkestan).

  • — Ganglbauer, 1895: 487 (Leptacinus; characters; Palaearctic region; North America; China; Australia).

  • — Fauvel, 1897d: 291 (Leptacinus; Barbary; Azores; Madeira; Canaries).

  • — Fauvel, 1902b: 100 (Leptacinus; cited as bathychrus; Barbary; Egypt; Azores; Madeira; Canary Islands).

  • — Sainte-Claire Deville, 1906: 93 (Leptacinus; Corsica).

  • — Reitter, 1908: 112 (Leptacinus; cited as bathychrus; characters; Palaearctic region).

  • — Montandon, 1908: 79 (Leptacinus; cited as bathychrus; Romania).

  • — Poppius, 1909: 23 (Leptacinus; Lena Valley, Siberia).

  • — Reitter, 1909: 135 (Leptacinus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 292 (Leptacinus; catalog).

  • — Johansen, 1914: 439 (Leptacinus; characters; Denmark).

  • — Eichelbaum, 1915a: 96 (Leptacinus; abdominal segmentation; apical segements).

  • — Lea, 1925: 215 (Leptacinus; Australian introduction).

  • — Portevin, 1929: 370 (Leptacinus; characters; France).

  • — Gridelli, 1930: 70 (Leptacinus; Libya).

  • — Bernhauer, 1931d: 582 (Leptacinus; Abyssinia).

  • — Scheerpeltz, 1933: 1300 (Leptacinus; catalog).

  • — Scheerpeltz, 1935a: 653 (Leptacinus; Australia).

  • — Bernhauer, 1940c: 3 (Leptacinus; Azores).

  • — Paulian, 1941: 207 (Leptacinus; larval characters).

  • — Méquignon, 1943: 159 (Leptacinus; characters; notes; synonyms).

  • — Tottenham, 1949: 416 (Leptacinus; checklist; Britain).

  • — V. Hansen, 1952: 10 (Leptacinus; characters; Denmark).

  • — Coiffait, 1956a: 47 (Leptacinus; characters in key; France; Pyrenees; Croatia; Czechoslovakia).

  • — Hatch, 1957: 238 (Leptacinus; characters; British Columbia; Idaho; Washington; Oregon).

  • — Smetana, 1958: 88 (Leptacinus; subgenus Leptacinus; characters; ecology; Czechoslovakia).

  • — Kocher, 1958: 138 (Leptacinus; Morocco).

  • — Smetana, 1959a: 208 (Leptacinus; Albania).

  • — Lohse, 1961: 183, 184, 185 (Leptacinus; characters).

  • — Brakman, 1962: 47 (Leptacinus; characters).

  • — Ferreira, 1962: 16 (Leptacinus; Portugal).

  • — Scheerpeltz, 1963: 10 (Leptacinus; Afghanistan).

  • — Scheerpeltz, 1963b: 428 (Leptacinus; cited as bathychrus; Egypt).

  • — Palm, 1963b: 43 (Leptacinus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 160 (Leptacinus; characters; central Europe).

  • — Raianu, 1964: 104 (Leptacinus; Romania).

  • — Coiffait, 1965e: 119 (Leptacinus; Turkey).

  • — Horion, 1965: 88 (Leptacinus; Palaearctic region; Siberia; all Europe; Germany; Austria).

  • — Coiffait, 1966: 21 (Leptacinus; Turkey).

  • — Szujecki, 1968a: 725 (Leptacinus; Poland).

  • — Scheerpeltz, 1968a: 59 (Leptacinus; catalog; Austria).

  • — Coiffait, 1972: 157 (Leptacinus; characters; comments on synonyms; references; Britain; Scandinavia; southeastern Europe; Asia Minor; France).

  • — Shilov, 1975a: 375 [= 1975b: 97] (Leptacinus; Komi region of Russia).

  • — Pope, 1977: 28 (Leptacinus; Britain).

  • — Scheerpeltz, 1978: 206 (Leptacinus; cited as bathychrus; Timor).

  • — Muona, 1979: 18 (Leptacinus; Finland; Russia; Sweden; Norway; Denmark; Baltic States).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 5 (Leptacinus; catalog; Poland).

  • — Smetana, 1982: 101 (Leptacinus; characters; North American introduction; British Columbia; Manitoba; Ontario; Quebec; Illinois; Iowa; Massachusetts; Michigan; New Jersey; New York; Oregon).

  • — Bordoni, 1982: 185 (Leptacinus; characters; Italy).

  • — Ádám, 1987: 145 (Leptacinus; collecting notes; Hungary).

  • — Lucht, 1987: 102 (Leptacinus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Tóth, 1989: 11 (Leptacinus; characters; Hungary).

  • — Silfverberg, 1992: 18 (Leptacinus; Finland; Russia; Sweden; Norway; Denmark; Estonia; Latvia; Lithuania).

  • — Ciceroni, 1993: 70 (Leptacinus; Italy).

  • — Janák, 1993a: 47 (Leptacinus; Czech Republic; Slovakia).

  • — Ciceroni, 1994: 100 (Leptacinus; aedeagal illustration; Italy; Belgium; Finland; France; Greece; Switzerland).

  • — Gerardi and Zanetti, 1995: 143 (Leptacinus; collecting notes; Italy).

  • — Ciceroni and Zanetti, 1995: 25 (Leptacinus; Italy).

  • — Downie and Arnett, 1996: 399 (Leptacinus; characters; USA).

  • — M. Hansen, 1996: 108 (Leptacinus; checklist; Denmark).

  • — Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 40 (Leptacinus; Latvia).

  • — Anderson, Nash, and O'Connor, 1997: 20 (Leptacinus; Ireland).

  • — Owen, 1997a: 149 (Leptacinus; collecting sites; Ireland).

  • — Hansen, Jørum, Palm, and Pedersen, 1997: 129 (Leptacinus; checklist; Denmark).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 74 (Leptacinus; Denmark).

  • — Ryabukhin, 1999: 57 (Leptacinus; catalog; habitat; distribution; northeastern Russia).

  • — Kashcheev, 1999a: 57 (Leptacinus; Kazakhstan).

  • — Herman, 2001: 26 (Leptacinus; nomen protectum).

  • Distribution: Ethiopia, Egypt, Libya, Tunisia, Algeria, Morocco, Canary Islands, Madeira, Azores, Europe, Russia, Georgia, Turkey, Lebanon, Israel, Iran, Afghanistan, Uzbekistan, Kazakhstan, China, Indonesia, Australia, Canada, USA.

diaphanus Marsham, 1802: 514 (Staphylinus; [Note: This name is older than batychrus]; Type locality: Britain).

  • — Stephens, 1833: 259 (Gyrohypnus; characters; England).

  • — Erichson, 1839b: 325 (Xantholinus; synonym of glaber Nordmann).

  • — Stephens, 1839: 404 (Leptacinus; characters; England).

  • — Hardy, 1851: 39 (Leptacinus; synonym of batychrus).

  • — Kraatz, 1857d: 640 (Xantholinus; synonym of glaber).

  • — G. Waterhouse, 1858: 26 (Leptacinus; synonym of batychrus).

  • — Ganglbauer, 1895: 487 (Leptacinus; synonym of batychrus).

  • — Reitter, 1908: 112 (Leptacinus; synonym of batychrus).

  • — Bernhauer and Schubert, 1914: 292 (Leptacinus; synonym of batychrus).

  • — Méquignon, 1943: 160 (Leptacinus; synonym of batychrus).

  • — Herman, 2001: 26 (Leptacinus; nomen oblitum).

apicalis Stephens, 1833: 260 (Gyrohypnus; Type locality: Norfolk).

  • — Stephens, 1839: 403 (Xantholinus; characters; England).

  • — G. Waterhouse, 1858: 26 (Leptacinus; synonym of batychrus).

  • — Ganglbauer, 1895: 487 (Leptacinus; synonym of batychrus).

  • — Reitter, 1908: 112 (Leptacinus; synonym of batychrus).

  • — Bernhauer and Schubert, 1914: 292 (Leptacinus; synonym of batychrus).

  • — Méquignon, 1943: 160 (Leptacinus; synonym of batychrus).

semistriatus Stephens, 1833: 262 (Gyrohypnus; Type locality: London; Suffolk).

  • — Stephens, 1839: 403 (Xantholinus; characters; England).

  • — Hardy, 1851: 38 (Leptacinus; synonym of parumpunctatus).

  • — G. Waterhouse, 1858: 26 (Leptacinus; synonym of batychrus).

  • — Ganglbauer, 1895: 487 (Leptacinus; synonym of batychrus).

  • — Reitter, 1908: 112 (Leptacinus; synonym of batychrus).

  • — Bernhauer and Schubert, 1914: 292 (Leptacinus; synonym of batychrus).

quadrisulcus Stephens, 1833: 264 (Gyrohypnus; Type locality: Suffolk; Bristol).

  • — Stephens, 1839: 404 (Leptacinus; variety of batychrus; characters; England).

  • — Hardy, 1851: 39 (Leptacinus; synonym of batychrus).

  • — G. Waterhouse, 1858: 26 (Leptacinus; synonym of batychrus).

  • — Ganglbauer, 1895: 487 (Leptacinus; synonym of batychrus).

  • — Reitter, 1908: 112 (Leptacinus; synonym of batychrus).

  • — Bernhauer and Schubert, 1914: 292 (Leptacinus; synonym of batychrus).

  • — Méquignon, 1943: 160 (Leptacinus; synonym of batychrus).

episcopalis Lacordaire, 1835: 416 (Xantholinus; Type locality: Paris).

  • — Erichson, 1839a: 430 (Leptacinus; synonym of batychrus).

  • — Erichson, 1839b: 335 (Leptacinus; synonym of batychrus).

  • — Redtenbacher, 1849: 693 (Leptacinus; synonym of batychrus).

  • — Hardy, 1851: 39 (Leptacinus; synonym of batychrus).

  • — Fairmaire and Laboulbène, 1856: 503 (Leptacinus; synonym of batychrus).

  • — Kraatz, 1857d: 649 (Leptacinus; synonym of batychrus).

  • — Mulsant and Rey, 1877a: 321 (Leptacinus; synonym of batychrus).

  • — Ganglbauer, 1895: 487 (Leptacinus; synonym of batychrus).

  • — Reitter, 1908: 112 (Leptacinus; synonym of batychrus).

  • — Bernhauer and Schubert, 1914: 292 (Leptacinus; synonym of batychrus).

  • — Méquignon, 1943: 160 (Leptacinus; synonym of batychrus).

tener Waltl, 1835: 59 (Xantholinus; Type locality: Andalusien).

  • — Erichson, 1840: 924 (Leptacinus; synonym of batychrus).

  • — Rosenhauer, 1856: 70 (Leptacinus; synonym of batychrus).

  • — Ganglbauer, 1895: 487 (Leptacinus; synonym of batychrus).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; synonym of batychrus).

apicalis Kolenati, 1846: 15 (Leptacinus; [preoccupied]; Type locality: Iberia; Tiflis).

  • — Hochhuth, 1849: 110 (Leptacinus; characters; Iberia, Tiflis).

  • — Fauvel, 1873b: 89 [= 1873a: 375] (Leptacinus; synonym of batychrus).

berytensis Saulcy, 1864: 643 (Leptacinus; Type locality: Beïroût).

  • — Fauvel, 1873b: 89 [= 1873a: 375] (Leptacinus; synonym of batychrus).

  • — Bernhauer and Schubert, 1914: 292 (Leptacinus; synonym of batychrus).

jebusaeus Saulcy, 1864: 643 (Leptacinus; Type locality: Jérusalem).

  • — Fauvel, 1873b: 89 [= 1873a: 375] (Leptacinus; synonym of batychrus).

  • — Bernhauer and Schubert, 1914: 292 (Leptacinus; synonym of batychrus).

triangulum Saulcy, 1864: 643 (Leptacinus; Type locality: Tibériade; Sahel).

  • — Fauvel, 1873b: 89 [= 1873a: 375] (Leptacinus; synonym of batychrus).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; synonym of batychrus).

seriatus LeConte, 1880: 169 (Leptacinus; Type locality: Detroit, Michigan; Canada; [Note: See lectotype designation by Smetana, 1982]).

  • — Casey, 1906: 402 (Leptacinodes; synonym of flavipes).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; synonym of batychrus).

  • — Smetana, 1982: 101 (Leptacinus; lectotype designation: locality not cited; synonym of batychrus).

perexilis Casey, 1906: 403 (Leptacinodes; Type locality: Europe, Caucasus).

  • — Bernhauer and Schubert, 1914: 292 (Leptacinus; synonym of batychrus).

substrigosus Joy, 1913c: 226 (Xantholinus; Type locality: Peckham).

  • — Joy, 1913e: 275 (Leptacinus; synonym of batychrus).

  • — Scheerpeltz, 1933: 1300 (Leptacinus; synonym of batychrus).

  • — Pope, 1977: 28 (Leptacinus; synonym of batychrus).

  • — Lohse and Lucht, 1989: 156 (Leptacinus; synonym of batychrus).

varius H. Bauer, 1914: 56 (Xantholinus; aberration of angustatus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Reichenland im Vogtland).

  • — Scheerpeltz, 1933: 1300 (Leptacinus; synonym of batychrus).

aristaeus Coiffait, 1956a: 47 (Leptacinus; Type locality: Isère: St-Julien-du-Rez).

  • — Coiffait, 1972: 156 (Leptacinus; characters; references; France).

  • — Ciceroni, 1994: 100 (Leptacinus; synonym of batychrus).

adonis Coiffait, 1956a: 47 (Leptacinus; Type locality: Liban: Hasbaya).

  • — Coiffait, 1972: 158 (Leptacinus; characters; Lebanon; Turkey).

  • — Ciceroni, 1994: 100 (Leptacinus; synonym of batychrus).

pan Coiffait, 1956a: 49 (Leptacinus; Type locality: Gironde: Mérignac).

  • — Coiffait, 1972: 152 (Leptacinus; characters; references; Germany; France).

  • — Drugmand and Haghebaert, 1987: 325 (Leptacinus; Belgium).

  • — Ciceroni, 1994: 100 (Leptacinus; synonym of batychrus).

beesoni Cameron, 1932: 12 (Leptacinus; Type locality: Dehra Dun: Nim Nadi).

  • — Scheerpeltz, 1933: 1301 (Leptacinus; catalog).

  • — Coiffait, 1982a: 241 (Leptacinus; characters).

  • Distribution: India.

berytensis Saulcy, 1864, see: batychrus Gyllenhal, 1827.

biroi Last, 1980: 156 (Leptacinus; Type locality: New Guinea: Friedrich-Wilhelmshafen).

  • Distribution: New Guinea.

biskrensis Coiffait, 1972, see: intermedius Donisthorpe, 1936.

bisulciceps Lea, 1925: 216 (Leptacinus; Type locality: Western Australia: Bridgetown, Pinjarrah).

  • — Scheerpeltz, 1933: 1301 (Leptacinus; catalog).

  • Distribution: Australia.

blackburni Lea, 1925: 215 (Leptacinus; replacement name for filum Blackburn).

  • — Scheerpeltz, 1933: 1301 (Leptacinus; catalog).

  • — Herman, 2001: 26 (Leptacinus; synonymy).

  • Distribution: Australia.

filum Olliff, 1887: 477 (Leptacinus; [Note: Preoccupied; although Olliff cited Blackburn as the author of filum he published the name before Blackburn]; Type locality: Port Lincoln, Australia).

  • — Herman, 2001: 26 (Leptacinus; synonym of blackburni).

filum Blackburn, 1888: 7 (Leptacinus; [Note: Preoccupied; Bernhauer and Schubert, 1914: 294 and Scheerpeltz, 1933: 1301 cited Blackburn as the author for this name, and although Olliff cited Blackburn as the author, Olliff's description appeared first]; Type locality: Port Lincoln).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; synonym of socius).

  • — Scheerpeltz, 1933: 1301 (Leptacinus; synonym of blackburni).

borneensis Cameron, 1933, see: cribricollis Fauvel, 1905.

breviceps C. Waterhouse, 1877, see: Phacophallus.

brevicornis Erichson, 1839, see: Zeteotomus.

brevisulcatus Cameron, 1933: 41 (Leptacinus; Type locality: Belgian Congo: Nizi: Blukwa).

  • — Cameron, 1950g: 28 (Leptacinus; Belgian Congo).

  • Distribution: Congo.

brincki Scheerpeltz, 1974: 117 (Leptacinus; Type locality: Cape Prov.: Jonkershoek, 8 miles SE Stellenbosch).

  • Distribution: South Africa.

brunnescens LeConte, 1880, see: Linohesperus.

brunneus Coiffait, 1968: 142 (Leptacinus; subgenus Chaetocinus; Type locality: Gabon: Belinga).

  • Distribution: Gabon.

bulgaricus Coiffait, 1972, see: othioides Baudi, 1870.

caffer Scheerpeltz, 1974: 115 (Leptacinus; Type locality: Cape Prov.: Cape Peninsula, Cape Point Nature Reserve).

  • Distribution: South Africa.

cameroni Steel, 1949a: 270 (Leptacinus; replacement name for aethiops Cameron).

  • — Cameron, 1950g: 29 (Leptacinus; Belgian Congo).

  • Distribution: Congo.

aethiops Cameron, 1933: 41 (Leptacinus; [preoccupied]; Type locality: Belgian Congo: Nizi: Blukwa).

  • — Steel, 1949a: 270 (Leptacinus; synonym of cameroni).

cephalicus LeConte, 1880, see: Phacophallus.

cephalotes Coiffait, 1968: 134 (Leptacinus; subgenus Leptacinus; Type locality: Gabon: Belinga).

  • Distribution: Gabon.

chinensis Cameron, 1940d: 251 (Leptacinus; Type locality: S. China: Sheung Shui).

  • Distribution: China.

circumcaspicus Gusarov, 1993: 70 (Leptacinus; Type locality: Turkmenistan: Ashgabat Prov.: 25 km. W Bakharden, Karayalchi).

  • Distribution: Turkmenistan.

clavicornis Erichson, 1839, see: Somoleptus.

congoensis Bernhauer, 1929f: 120 (Leptacinus; Type locality: Haut-Uele: Moto; Elisabethville: Pweto; Uganda: Bussu Busoga).

  • — Scheerpeltz, 1933: 1301 (Leptacinus; catalog).

  • — Cameron, 1950g: 29 (Leptacinus; Belgian Congo).

  • Distribution: Uganda, Congo.

cooperi Bernhauer, 1931d: 581 (Leptacinus; Type locality: Abyssinia: Hawash Railway Station, ca. 3,500 ft.).

  • — Scheerpeltz, 1933: 1301 (Leptacinus; catalog).

  • Distribution: Ethiopia.

corsicus Coiffait, 1972, see: marocanus Coiffait, 1969.

crepitans Tottenham, 1956: 239 (Leptacinus; Type locality: Ruanda: Tshuruyaga, forêt du Rugege, 2400 m).

  • Distribution: Rwanda.

cribricollis Fauvel, 1905e: 83 (Leptacinus; Type locality: Buitenzorg).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; catalog).

  • — Cameron, 1933d: 383 (Leptacinus; Sumatra).

  • Distribution: Borneo, Indonesia.

borneensis Cameron, 1933b: 344 (Leptacinus; variety of cribricollis; Type locality: British North Borneo: Kenokok, 3300 feet).

curicanus Coiffait and Saiz, 1964: 512 (Leptacinus; Type locality: Chile: Romeral, Curicó).

  • — Saiz, 1970b: 384 (Leptacinus; characters; Chile).

  • Distribution: Chile.

cyaneipennis W. J. MacLeay, 1873, see: Neoxantholinus.

debilis Erichson, 1839, see: Somoleptus.

debilis Cameron, 1950g: 28 (Leptacinus; [preoccupied]; Type locality: Belgian Congo: Kitondo, Gandjo, alt. 2000 m).

  • — Herman, 2001: 45 (Leptacinus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: Congo.

delicatulus Coiffait, 1968: 142 (Leptacinus; subgenus Chaetocinus; Type locality: Gabon: Belinga).

  • Distribution: Gabon.

densus Bernhauer, 1939k: 589 (Leptacinus; Type locality: N.W. China: Chinkiang).

  • Distribution: China.

diaphanus Marsham, 1802, see: batychrus Gyllenhal, 1827.

dieganus Fauvel, 1905b: 169 (Leptacinus; Type locality: Diego-Suarez).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; catalog).

  • Distribution: Madagascar.

difficilis Cameron, 1933d: 384 (Leptacinus; Type locality: Not cited but presumably Sumatra).

  • Distribution: Indonesia.

dubius Bordoni, 1975d: 196 (Leptacinus; Type locality: Russia meridionale).

  • Distribution: Russia.

eggersi Bernhauer and Schubert, 1914: 293 (Leptacinus; replacement name for apicipennis Bernhauer).

  • — Blackwelder, 1943: 493 (Leptacinus; characters; St. Thomas).

  • Distribution: West Indies.

apicipennis Bernhauer, 1904a: 234 (Leptacinus; [preoccupied]; Type locality: Ins. St. Thomas).

  • — Blackwelder, 1943: 493 (Leptacinus; synonym of eggersi).

elgonensis Bernhauer, 1939a: 75 (Leptacinus; Type locality: Kenya: Suam fishing hut, M. Elgon, Osthang, 2400 m).

  • Distribution: Kenya.

encephalus Fauvel, 1907: 33 (Leptacinus; Type locality: Kilimandjaro: Kilema).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; cited as eucephalus; catalog).

  • — Scheerpeltz, 1933: 1301 (Leptacinus; catalog).

  • Distribution: Tanzania.

episcopalis Lacordaire, 1835, see: batychrus Gyllenhal, 1827.

exsucidus† Scudder, 1900: 61 (Leptacinus; fossil; Type locality: Florissant, Colorado).

  • — Handlirsch, 1907: 726 (Leptacinus; catalog; Miocene; Colorado).

  • Distribution: USA.

faunus Coiffait, 1956a: 48 (Leptacinus; Type locality: Espagne: Ségovie).

  • — Coiffait, 1972: 152 (Leptacinus; characters; references; Spain; Portugal).

  • — Ciceroni, 1994: 103 (Leptacinus; aedeagal illustration; Sicily; Algeria; Spain).

  • — Ciceroni and Zanetti, 1995: 25 (Leptacinus; Italy).

  • Distribution: Algeria, Portugal, Spain, Italy.

siculus Coiffait, 1956b: 140 (Leptacinus; Type locality: Sicile: Agrigente).

  • — Coiffait, 1972: 153 (Leptacinus; characters; references; Sicily).

  • — Bordoni, 1982: 190 (Leptacinus; characters; Italy).

  • — Ciceroni, 1994: 103 (Leptacinus; synonym of faunus).

  • — Sparacio, 1995: 156 (Leptacinus; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 25 (Leptacinus; synonym of faunus).

algiricus Coiffait, 1972: 147 (Leptacinus; subgenus Leptacinus; Type locality: Gorges d'El Kantara, Algérie).

  • — Ciceroni, 1994: 103 (Leptacinus; synonym of faunus).

fauveli Cameron, 1922, see: Phacophallus.

ferrugineus Bernhauer, 1927e: 243 (Leptacinus; Type locality: Argentinien: Prov. Cordoba, Alta Gracia).

  • — Scheerpeltz, 1933: 1301 (Leptacinus; catalog).

  • Distribution: Argentina.

filarius Erichson, 1839, see: Neoxantholinus.

filiformis Scheerpeltz, 1957c: 233, 307 (Leptacinus; Type locality: Sumba: Mau Marru).

  • Distribution: Indonesia.

filum Kraatz, 1859: 111 (Leptacinus; Type locality: Ceylan).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; catalog).

  • — Cameron, 1932: 10 (Leptacinus; characters; Ceylon, India).

  • — Scheerpeltz, 1933: 1301 (Leptacinus; catalog; Philippines).

  • — Cameron, 1933d: 383 (Leptacinus; Sumatra).

  • — Coiffait, 1982a: 231 (Leptacinus; Assam).

  • — Coiffait, 1982a: 241 (Leptacinus; characters).

  • Distribution: Sri Lanka, India, Indonesia, Philippines.

filum Olliff, 1887, see: blackburni Lea, 1925.

filum Blackburn, 1888, see: blackburni Lea, 1925.

flavipennis Kraatz, 1859, see: Phacophallus.

flavipes LeConte, 1863, see: Stictolinus.

formicetorum Märkel, 1841: 216 (Leptacinus; Type locality: Not cited but presumably Sächs. Schweiz).

  • — Mannerheim, 1843a: 83 (Leptacinus; characters).

  • — Märkel, 1845: 236 (Leptacinus; notes; with Formica rufa).

  • — Rosenhauer, 1856: 70 (Leptacinus; Sierra Nevada).

  • — Fairmaire and Laboulbène, 1856: 504 (Leptacinus; characters; France).

  • — Kraatz, 1857d: 650 (Leptacinus; characters; Germany).

  • — Thomson, 1860: 193 (Leptacinus; characters; Scandinavia).

  • — Baudi, 1870: 389 (Leptacinus; characters).

  • — Fauvel, 1873b: 91 [= 1873a: 377] (Leptacinus; characters; France; Scandinavia; Britain; Germany; Russia; Cyprus; Spain).

  • — Seidlitz, 1875: 280 (Leptacinus; characters; Baltic region).

  • — J. Sahlberg, 1876: 37 (Leptacinus; Finland).

  • — Mulsant and Rey, 1877a: 329 (Leptacinus; characters; France).

  • — Fowler, 1888: 293 (Leptacinus; characters; Britain).

  • — Ganglbauer, 1895: 488 (Leptacinus; characters; north and middle Europe; Spain; Cyprus).

  • — Reitter, 1908: 112 (Leptacinus; characters; Europe).

  • — Reitter, 1909: 135 (Leptacinus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; catalog).

  • — Johansen, 1914: 440 (Leptacinus; characters; Denmark).

  • — Šulc, 1922 (Leptacinus; osmeterium; anatomy; histology; physiology).

  • — Portevin, 1929: 370 (Leptacinus; characters; France).

  • — Scheerpeltz, 1933: 1301 (Leptacinus; catalog).

  • — V. Hansen, 1952: 11 (Leptacinus; characters; Denmark).

  • — Coiffait, 1956a: 50 (Leptacinus; characters in key).

  • — Smetana, 1958: 92 (Leptacinus; subgenus Leptacinus; characters; ecology; Czechoslovakia).

  • — Lohse, 1961: 183, 184, 186 (Leptacinus; characters).

  • — Palm, 1963b: 44 (Leptacinus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 159 (Leptacinus; characters; central Europe).

  • — Smetana, 1964b: 50 (Leptacinus; larval characters).

  • — Horion, 1965: 87 (Leptacinus; Russia; Lapland; Britain; Spain and Italy to Dalmatia, Bosnia, Greece etc.; Spain; Cyprus; Germany; Austria).

  • — Szujecki, 1968a: 725 (Leptacinus; Poland).

  • — Coiffait, 1972: 149 (Leptacinus; characters; references; Europe to Siberia; France).

  • — Pope, 1977: 28 (Leptacinus; Britain).

  • — Bordoni, 1982: 192 (Leptacinus; characters; Italy).

  • — Lucht, 1987: 102 (Leptacinus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; Benelux; Denmark; Sweden).

  • — Tóth, 1989: 10 (Leptacinus; characters).

  • — Janák and Vysoky, 1992: 136 (Leptacinus; collected with Formica polyctena, rufa, and pratensis; Bohemia).

  • — Franc, 1993: 307 (Leptacinus; synecthrical myrmecophile; Slovakia).

  • — Janák, 1993a: 47 (Leptacinus; Czech Republic; Slovakia).

  • — Ciceroni, 1994: 105 (Leptacinus; aedeagal illustration; Italy; Finland; France; Germany; Switzerland; Hungary [Note: Erroneous locality]).

  • — Ciceroni and Zanetti, 1995: 25 (Leptacinus; Italy).

  • — Silfverberg, 1999: 50 (Leeptacinus; Estonia).

  • — Ryabukhin, 1999: 57 (Leptacinus; catalog; habitat; distribution; northeastern Russia).

  • — Kashcheev, 1999a: 57 (Leptacinus; Kazakhstan).

  • Distribution: Europe, Russia, Kazakhstan.

fossus† Scudder, 1900: 59 (Leptacinus; fossil; Type locality: Florissant, Colorado).

  • — Handlirsch, 1907: 726 (Leptacinus; catalog; Miocene; Colorado).

  • Distribution: USA.

fragilis Cameron, 1932a: 139 (Leptacinus; Type locality: Selangor: The Gap).

  • Distribution: Malaysia.

fraternus Cameron, 1932a: 139 (Leptacinus; Type locality: Selangor: The Gap).

  • Distribution: Malaysia.

fusciceps Eppelsheim, 1895b: 399 (Leptacinus; Type locality: Pegu).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; catalog).

  • — Cameron, 1932: 12 (Leptacinus; characters; Burma).

  • — Scheerpeltz, 1933: 1301 (Leptacinus; catalog).

  • — Coiffait, 1982a: 241 (Leptacinus; cited as fuscipes; characters).

  • Distribution: Myanmar.

gabonicus Coiffait, 1968: 133 (Leptacinus; subgenus Leptacinus; Type locality: Gabon: Belinga).

  • Distribution: Gabon.

galapagoensis Coiffait, 1981g: 291 (Leptacinus; Type locality: Ile Galapagos: Ile San Cristobal).

  • Distribution: Galapagos Islands.

gardneri Cameron, 1945: 68 (Leptacinus; Type locality: United Provinces: Dehra Dun, New Forest).

  • Distribution: India.

gerardi Bernhauer, 1932b: 143 (Leptacinus; Type locality: Katanga: Katompe).

  • — Scheerpeltz, 1933: 1301 (Leptacinus; catalog).

  • Distribution: Congo.

germaini Coiffait and Saiz, 1964: 514 (Leptacinus; Type locality: Chile: Santiago).

  • — Saiz, 1970b: 383, 384 (Leptacinus; characters; Chile).

  • Distribution: Chile.

glabripennis Scriba, 1855, see: Somoleptus.

gomyi Lecoq, 1990: 188 (Leptacinus; Type locality: La Réunion: Saint Paul).

  • Distribution: Réunion.

goropanensis Coiffait, 1982: 49 (Leptacinus; Type locality: Nepal: versant sud-est du Col de Goropani, ouest de Pokhara, 2150 m).

  • — Coiffait, 1982a: 240 (Leptacinus; characters).

  • Distribution: Nepal.

gracilentus Scheerpeltz, 1974: 120 (Leptacinus; Type locality: S. Rhodesia: Victoria Falls).

  • Distribution: Zimbabwe.

gracilis Fauvel, 1895: 240 (Leptacinus; Type locality: Bírmanie, Carin Asciuii Ghécu, 1400–1500 m; Java occidental, Pengalengan, 1300 m).

  • — Fauvel, 1904a: 56 (Leptacinus; Nilghiris; Bírmanie; Java).

  • — Fauvel, 1905e: 82 (Leptacinus; Java; Burma; India).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; catalog).

  • — Cameron, 1932: 9 (Leptacinus; characters; Burma; Assam; India).

  • — Scheerpeltz, 1933: 1301 (Leptacinus; catalog).

  • — Scheerpeltz, 1957c: 233 (Leptacinus; Sumba).

  • — Shibata, 1973a: 122 (Leptacinus; Taiwan).

  • — Scheerpeltz, 1976a: 85 (Leptacinus; Nepal).

  • — Coiffait, 1982a: 240 (Leptacinus; characters).

  • Distribution: India, Nepal, Myanmar, Indonesia, Taiwan.

grandiceps Sharp, 1885, see: lecontei Blackwelder, 1944.

grassei Coiffait, 1968: 140 (Leptacinus; subgenus Chaetocinus; Type locality: Gabon: Belinga).

  • Distribution: Gabon.

guadarramus Outerelo, 1975: 51 (Leptacinus; Type locality: Sierra Guadarrama: Valle de Enmedio en Peguerinos, Avila).

  • Distribution: Spain.

harbinensis Bordoni, 2000: 128 (Leptacinus; Type locality: N. China: Harbin, Heilungjiang).

  • Distribution: China.

hariolus Herman, 2001: 45 (Leptacinus; replacement name for sinuatocollis Scheerpeltz, 1974).

  • Distribution: Zambia.

sinuatocollis Scheerpeltz, 1974: 118 (Leptacinus; [preoccupied]; Type locality: N. Rhodesia: Livingstone).

  • — Herman, 2001: 45 (Leptacinus; synonym of hariolus).

hypsibathus Bernhauer, 1939a: 75 (Leptacinus; Type locality: Kenya: Suam fishing hut, M. Elgon, Osthang, 2400 m).

  • Distribution: Kenya.

intermedius Donisthorpe, 1936: 269 (Leptacinus; Type locality: Windsor Forest, England).

  • — Blatchley, 1910: 399 (Xantholinus; [Note: Misidentified as flavipes according to Smetana, 1982: 97]; characters; Indiana).

  • — Méquignon, 1943: 160 (Leptacinus; synonym of sulcifrons).

  • — V. Hansen, 1952: 10 (Leptacinus; characters; Denmark).

  • — Horion, 1954: 9 (Leptacinus; characters; Germany; Scandinavia; Denmark).

  • — Smetana, 1958: 91 (Leptacinus; synonym of sulcifrons).

  • — Lohse, 1961: 183, 184, 185 (Leptacinus; characters).

  • — Brakman, 1962: 46 (Leptacinus; characters; Holland).

  • — Palm, 1963b: 43 (Leptacinus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 160 (Leptacinus; characters; central Europe).

  • — Horion, 1965: 88 (Leptacinus; Britain; Norway; Sweden; Finland; Denmark; Holland; France; Czechoslovakia; Switzerland; Italy Austria; Germany).

  • — Coiffait, 1972: 11 (Leptacinus; synonym of sulcifrons).

  • — Pope, 1977: 28 (Leptacinus; Britain).

  • — Smetana, 1982: 97 (Leptacinus; characters; North American introduction; Alberta; British Columbia; Manitoba; Ontario; Quebec; Saskatchewan; Arizona; California; Colorado; Connecticut; Illinois; Indiana; Massachusetts; Nebraska; Nevada; New Mexico; New York; Ohio; Oregon; Pennsylvania; Washington).

  • — Smetana, 1988a: 526 (Leptacinus; collecting notes; Ontario; Indiana; New Hampshire).

  • — Allen, 1990: 289 (Leptacinus; discussion on validity of species; characters; Britain).

  • — Ciceroni, 1993: 70 (Leptacinus; Italy).

  • — Janák, 1993a: 47 (Leptacinus; synonym of othioides).

  • — Ciceroni, 1994: 106 (Leptacinus; aedeagal illustration; Italy; Algeria; Cyprus; Finland; Greece; Libya; Czech Republic; Switzerland; Turkey).

  • — Hodge and Jones, 1995: 37 (Leptacinus; characters; Britain).

  • — Ciceroni and Zanetti, 1995: 25 (Leptacinus; “othioides0 Auct. nec Baudi” cited as synonym; Italy).

  • — Downie and Arnett, 1996: 399 (Leptacinus; characters; USA).

  • Distribution: Libya, Algeria, Europe, Turkey, Canada, USA.

silvanus Coiffait, 1956a: 50 (Leptacinus; Type locality: Creuse: Ste. Feyre. Also cited from Portugal and Czechoslovakia).

  • — Smetana, 1958: 92 (Leptacinus; subgenus Leptacinus; characters; ecology; Czechoslovakia).

  • — Coiffait, 1972: 150 (Leptacinus; synonym of othioides).

  • — Janák, 1993a: 47 (Leptacinus; synonym of othioides).

  • — Ciceroni, 1993: 69 (Leptacinus; probably synonym of intermedius).

  • — Ciceroni, 1994: 106 (Leptacinus; synonym of intermedius).

rattii Coiffait, 1969a: 42 (Leptacinus; Type locality: Venise, Lido).

  • — Coiffait, 1972: 148 (Leptacinus; characters; references).

  • — Bordoni, 1982: 189 (Leptacinus; characters; Italy).

  • — Ciceroni, 1994: 106 (Leptacinus; synonym of intermedius).

biskrensis Coiffait, 1972: 154 (Leptacinus; subgenus Leptacinus; Type locality: Biskra, Sud Algérien).

  • — Ciceroni, 1994: 106 (Leptacinus; synonym of intermedius).

irritans Tottenham, 1956: 241 (Leptacinus; Type locality: Ruanda: terr. Kisenyi, Kayove, 2000 m).

  • Distribution: Rwanda.

jacobsoni Cameron, 1933d: 384 (Leptacinus; Type locality: Sumatra: Fort de Kock).

  • Distribution: Indonesia.

japonicus Cameron, 1933a: 169 (Leptacinus; Type locality: Japan: Kobe).

  • — Nakane, 1963a: 89 (Leptacinus; characters; habitus photograph; Japan).

  • — Watanabe and Shibata, 1972: 63 (Leptacinus; Yaku-shima Island).

  • Distribution: Japan.

javanus Bernhauer, 1916h: 423 (Leptacinus; Type locality: Java: Preanger).

  • — Scheerpeltz, 1933: 1301 (Leptacinus; catalog).

  • Distribution: Indonesia.

jebusaeus Saulcy, 1864, see: batychrus Gyllenhal, 1827.

karakorus Coiffait, 1975a: 156 (Leptacinus; Type locality: Karakorum, Nangana-Tal, 3200 m).

  • — Coiffait, 1982a: 242 (Leptacinus; characters).

  • Distribution: “Karakorum region”.

kazirensis Coiffait, 1982a: 237 (Leptacinus; Type locality: Assam: Kaziranga, 75 m).

  • Distribution: India.

klapperichi Coiffait, 1981: 15 (Leptacinus; Type locality: Iran: Golhak, près Téhéran, 1400 m).

  • Distribution: Iran.

ladakhensis Coiffait, 1982: 50 (Leptacinus; Type locality: Inde: Ladakh, Kargil, 2950 m).

  • — Coiffait, 1982a: 241 (Leptacinus; characters).

  • Distribution: India.

laeviusculus Solsky, 1864, see: Allolinus.

lecontei Blackwelder, 1944: 146 (Leptacinus; replacement name for grandiceps Sharp).

  • Distribution: Guatemala, Panama.

grandiceps Sharp, 1885: 490 (Leptacinus; Type locality: Guatemala, Las Mercedes 3000 feet, El Tumbador 2500 feet, Coatepeque 1300 feet; Panama, San Feliz, Bugaba).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; catalog).

leidyi† Scudder, 1900: 61 (Leptacinus; fossil; Type locality: Florissant, Colorado).

  • — Handlirsch, 1907: 726 (Leptacinus; catalog; Miocene; Colorado).

  • — Wickham, 1912: 11 (Leptacinus; Colorado).

  • Distribution: USA.

linearis Gravenhorst, 1802, see: pusillus Stephens, 1833.

lineaticeps Coiffait, 1982: 47 (Leptacinus; Type locality: Nepal: versant sud-est du Col de Goropani, ouest de Pokhara, 2150 m).

  • — Coiffait, 1982a: 241 (Leptacinus; characters).

  • Distribution: Nepal.

lipposus Herman, 2001: 45 (Leptacinus; replacement name for microps Coiffait).

  • Distribution: Gabon.

microps Coiffait, 1968: 139 (Leptacinus; subgenus Chaetocinus; [preoccupied]; Type locality: Gabon: Belinga).

  • — Herman, 2001: 45 (Leptacinus; synonym of lipposus).

longicollis LeConte, 1863, see: Lithocharodes.

longipennis Cameron, 1951c: 398 (Leptacinus; Type locality: Kenya Colony: Kabete, 5800 feet).

  • Distribution: Kenya.

luofuensis Cameron, 1950g: 29 (Leptacinus; Type locality: Belgian Congo: Luofu, alt. 1700 m).

  • Distribution: Congo.

lupercus Coiffait, 1956, see: othioides Baudi, 1870.

luridipennis W. J. MacLeay, 1873: 137 (Leptacinus; Type locality: Gayndah).

  • — Fauvel, 1877: 237 (Metoponcus; characters; Australia).

  • — Olliff, 1887: 474 (Leptacinus; characters; Australia).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; catalog).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; catalog).

  • Distribution: Australia.

novaehollandiae Fauvel, 1877: 236 (Leptacinus; Type locality: Australie orientale, Rockhampton; Victoria).

  • — Fauvel, 1878g: 537 (Leptacinus; characters).

  • — Olliff, 1887: 475 (Leptacinus; characters; Australia).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; catalog).

  • — Lea, 1925: 213 (Leptacinus; synonym of luridipennis).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; synonym of luridipennis).

lustrans Casey, 1906, see: Neohypnus.

machadoi Cameron, 1959: 113 (Leptacinus; Type locality: Angola: near Vila Luso).

  • Distribution: Angola.

maclurei† Scudder, 1900: 60 (Leptacinus; fossil; Type locality: Florissant, Colorado).

  • — Handlirsch, 1907: 726 (Leptacinus; catalog; Miocene; Colorado).

  • Distribution: USA.

macropterus Bernhauer, 1939a: 76 (Leptacinus; Type locality: Kenya: Vallée Koptawelil, Mont Elgon, Osthang, 2300 m).

  • Distribution: Kenya.

magniceps Bernhauer, 1922b: 175 (Leptacinus; Type locality: Seychelles: Silhouette: Mare aux Cochons, about 1000 ft., near sea level at Pointe Étienne).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; catalog).

  • Distribution: Seychelles.

major Coiffait, 1968: 143 (Leptacinus; subgenus Chaetocinus; Type locality: Gabon: Belinga).

  • — Coiffait, 1982a: 240 (Leptacinus; characters).

  • Distribution: Gabon.

major Coiffait, 1977, see: titan Coiffait, 1980.

malayanus Watanabe and Shibata, 1976: 338 (Leptacinus; Type locality: Pasoh Forest, Negeri Sembilan).

  • Distribution: Malaysia.

maledivensis Scheerpeltz, 1965a: 63 (Leptacinus; Type locality: Addu-Atoll, südlichstes Inselchen der Malediven-Gruppe, bei Bushey-Island).

  • Distribution: Maldive Islands.

mandibularis Cameron, 1929: 59 (Leptacinus; Type locality: Kivu: Kibati; Lulenga; Burunga).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; catalog).

  • — Cameron, 1933: 41 (Leptacinus; characters of female).

  • — Bernhauer, 1942c: 360 (Leptacinus; Cameroon).

  • — Cameron, 1950g: 28 (Leptacinus; Belgian Congo).

  • Distribution: Congo, Cameroon.

marocanus Coiffait, 1969a: 41 (Leptacinus; Type locality: Aguelmane Sidi Ali, Maroc).

  • — Coiffait, 1972: 155 (Leptacinus; characters; references; Morocco).

  • — Coiffait, 1973a: 273 (Leptacinus; Morocco).

  • — Ciceroni, 1994: 109 (Leptacinus; aedeagal illustration; Italy; France; Morocco).

  • — Ciceroni and Zanetti, 1995: 25 (Leptacinus; Italy).

  • Distribution: Morocco, France, Italy.

corsicus Coiffait, 1972: 156 (Leptacinus; subgenus Leptacinus; Type locality: Aleria, Corse).

  • — Bordoni, 1982: 188 (Leptacinus; characters; Italy).

  • — Ciceroni, 1994: 109 (Leptacinus; synonym of marocanus).

marshalli Bernhauer, 1938: 39 (Leptacinus; Type locality: China: Foochow).

  • Distribution: China.

megacephalus Lea, 1925: 217 (Leptacinus; Type locality: Lord Howe Island).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; catalog).

  • Distribution: Lord Howe Island.

merkli Ádám, 1987: 146 (Leptacinus; Type locality: Hu. c., Dömsöd, Apajpuszta).

  • Distribution: Hungary.

microparamerus Coiffait, 1977b: 212 (Leptacinus; subgenus Leptacinus; Type locality: Bhutan: 87 km de Phuntsholing).

  • — Coiffait, 1982a: 239 (Leptacinus; characters).

  • Distribution: Bhutan.

microphthalmus Fauvel, 1905b: 169 (Leptacinus; Type locality: Suberbieville).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; catalog).

  • Distribution: Madagascar.

microps Jarrige, 1951, see: Paulianella.

microps Coiffait, 1968, see: lipposus Herman, 2001.

micropterus Bernhauer, 1931d: 580 (Leptacinus; Type locality: Abyssinia: Mt. Chillalo, 12,000–13,000 ft. and ca. 9,000 ft.).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; catalog).

  • Distribution: Ethiopia.

miltoni Tottenham, 1953b: 492 (Leptacinus; Type locality: Kilimanjaro: Shira Plateau, Camp 2, 12,450 ft.).

  • Distribution: Tanzania.

minimus Bernhauer, 1920: 15 (Leptacinus; Type locality: Cedar creek; Malanda).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; catalog).

  • Distribution: Australia.

suturalis Lea, 1925: 217 (Leptacinus; Type locality: New South Wales: Galston).

  • — Scheerpeltz, 1933: 1303 (Leptacinus; catalog).

  • — Steel, 1949a: 269 (Leptacinus; synonym of minimus).

minutissimus Cameron, 1959: 113 (Leptacinus; Type locality: Angola: Dundo; Tala Mungongo).

  • Distribution: Angola.

minutus Lacordaire, 1835, see: pusillus Stephens, 1833.

minutus Coiffait, 1968, see: paulus Herman, 2001.

paulus Herman, 2001: 45 (Leptacinus; replacement name for minutus Coiffait).

  • Distribution: Gabon.

minutus Coiffait, 1968: 138 (Leptacinus; subgenus Chaetocinus; [preoccupied]; Type locality: Gabon: Belinga).

  • — Herman, 2001: 45 (Leptacinus; synonym of paulus).

multipunctatus Scheerpeltz, 1974: 110 (Leptacinus; Type locality: Natal: Royal Natal National Park, Tugela Valley).

  • Distribution: South Africa.

natalensis Scheerpeltz, 1974: 114 (Leptacinus; Type locality: Natal: Royal Natal National Park, Gudu Falls).

  • Distribution: South Africa.

nepalicus Coiffait, 1975a: 156 (Leptacinus; Type locality: entre Sunderijal et Mulkharka, Népal central).

  • — Coiffait, 1982: 26 (Leptacinus; characters; Nepal).

  • — Coiffait, 1982a: 241 (Leptacinus; characters).

  • — Coiffait, 1984e: 373 (Leptacinus; Nepal).

  • Distribution: Nepal.

niger Cameron, 1949a: 319 (Leptacinus; Type locality: Guinée: Mt. Nimba. Also cited from Congo).

  • Distribution: Ivory Coast, Congo.

niger Coiffait, 1966, see: nigerrimus Coiffait, 1971.

nigerrimus Coiffait, 1971: 436 (Leptacinus; replacement for niger Coiffait).

  • — Coiffait, 1972: 154 (Leptacinus; characters; references; Caucasus).

  • — Coiffait, 1973: 109 (Leptacinus; Turkey).

  • Distribution: Caucasus, Turkey.

niger Coiffait, 1966a: 125 (Leptacinus; [preoccupied]; Type locality: Caucase, 2200 m).

  • — Coiffait, 1971: 436 (Leptacinus; synonym of nigerrimus).

  • — Coiffait, 1972: 154 (Leptacinus; synonym of nigerrimus).

nigritulus LeConte, 1982, see: pusillus Stephens, 1833.

nilamburensis Cameron, 1932: 10 (Leptacinus; Type locality: Madras: Nilambur).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; catalog).

  • Distribution: India.

nitidus Sharp, 1876, see: Somoleptus.

notabilis Cameron, 1926b: 342 (Leptacinus; Type locality: Chakrata District; Sijla Gad, 5000 feet).

  • — Cameron, 1932: 11 (Leptacinus; characters; India).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; catalog).

  • — Coiffait, 1982a: 241 (Leptacinus; characters).

  • Distribution: India.

nothus Erichson, 1839, see: Stenistoderus.

novaehollandiae Fauvel, 1877, see: luridipennis W. J. MacLeay, 1873.

oculatus Coiffait, 1968: 136 (Leptacinus; subgenus Chaetocinus; Type locality: Gabon: Belinga).

  • Distribution: Gabon.

oculatus Coiffait, 1977, see: ophtalmicus Coiffait, 1980.

opacipennis Lea, 1925: 218 (Leptacinus; Type locality: Norfolk Island).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; catalog).

  • — Smithers, 1998: 16 (Leptacinus; references; Norfolk Island).

  • Distribution: Norfolk Island.

ophtalmicus Coiffait, 1980, see: Allolinus.

ops Coiffait, 1956, see: sulcifrons Stephens, 1833.

oscillans Sharp, 1885: 490 (Leptacinus; Type locality: Nicaragua, Chontales; Panama, Bugaba).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; catalog).

  • Distribution: Nicaragua, Panama.

othioides Baudi, 1870: 390 (Leptacinus; Type locality: [Italy]: Padus [river]; [Note: See lectotype designation by Bordoni, 1975d]).

  • — Baudi, 1857: 99 (Leptacinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 293 cited formicetorum Baudi as a synonym of othioides Baudi; Baudi did not describe the species as new, he attributed it to Märkel]; characters).

  • — Fauvel, 1873b: 89 [= 1873a: 375] (Leptacinus; characters; Italy).

  • — Ganglbauer, 1895: 488 (Leptacinus; characters; Piemont).

  • — Reitter, 1908: 112 (Leptacinus; characters; Piemont; Hungary; Greece; Caucasus; Turkestan).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; catalog).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; catalog).

  • — Coiffait, 1972: 150 (Leptacinus; characters; references; Iraq; Czechoslovakia; Italy; Greece; Turkey; Spain; Portugal; France).

  • — Bordoni, 1975d: 196 (Leptacinus; lectotype designation).

  • — Bordoni, 1982: 189 (Leptacinus; characters; Italy).

  • — Drugmand and Haghebaert, 1987: 325 (Leptacinus; Belgium).

  • — Tóth, 1989: 11 (Leptacinus; characters; Hungary).

  • — Lohse and Lucht, 1989: 156 (Leptacinus; notes).

  • — Janák, 1993a: 47 (Leptacinus; Czech Republic; Slovakia).

  • — Ciceroni, 1994: 110 (Leptacinus; aedeagal illustration; Italy; Bulgaria; Cyprus; Greece; Libya; Turkey).

  • — Ciceroni and Zanetti, 1995: 25 (Leptacinus; Italy).

  • Distribution: Libya, Europe, Caucasus, Turkey, Lebanon, Israel, Syria, Iraq, “Turkestan”.

lupercus Coiffait, 1956a: 46 (Leptacinus; Type locality: Liban: Hasbaya. Also cited from Syria, Palestine, and Turkey).

  • — Coiffait, 1972: 139 (Leptacinus; characters; Lebanon; Palestine; Syria; Turkey).

  • — Coiffait, 1972: 146 (Leptacinus; characters; references; Palestine; Lebanon; Syria; Turkey).

  • — Ciceroni, 1994: 110 (Leptacinus; synonym of othioides).

bulgaricus Coiffait, 1972: 139, 146 (Leptacinus; subgenus Leptacinus; subspecies of lupercus; Type locality: Bulgarie: Elkovo).

  • — Janák, 1993a: 47 (Leptacinus; subspecies of formicetorum; Czech Republic).

  • — Ciceroni, 1994: 110 (Leptacinus; synonym of othioides).

zanettii Bordoni, 1978: 176 (Leptacinus; Type locality: Veneto, S. Pancrazio, Verona).

  • — Bordoni, 1982: 183 (Leptacinus; characters; Italy).

  • — Ciceroni, 1994: 110 (Leptacinus; synonym of othioides).

  • — Ciceroni and Zanetti, 1995: 25 (Leptacinus; synonym of othioides).

ovaliceps Bernhauer, 1937b: 608 (Leptacinus; Type locality: Nord Uluguru, 1400–1900 m).

  • — Bernhauer, 1942c: 361 (Leptacinus; Cameroon).

  • Distribution: Tanzania.

pakistanus Coiffait, 1982a: 238 (Leptacinus; Type locality: Pakistan: Miandam, 1800–2300 m).

  • Distribution: Pakistan.

pallescens Tottenham, 1956: 242 (Leptacinus; Type locality: Ruanda: Kibuye, 1500 m).

  • Distribution: Rwanda.

pallidipennis Motschulsky, 1858, see: Phacophallus.

pallidulus LeConte, 1880, see: Lepitacnus.

pallidus Cameron, 1927a: 24 (Leptacinus; Type locality: Samoa: Tutuila: Fagasa).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; catalog).

  • Distribution: Samoa.

pan Coiffait, 1956, see: batychrus Gyllenhal, 1827.

papuensis Fauvel, 1878, see: Phacophallus.

parviparamerus Coiffait, 1982: 49 (Leptacinus; Type locality: Inde: Cachemire, Aru).

  • — Coiffait, 1982a: 240 (Leptacinus; characters).

  • Distribution: India.

parvus Cameron, 1950g: 28 (Leptacinus; Type locality: Belgian Congo: Shamuheru, volc. Nyamuragira, alt. 1843 m).

  • Distribution: Congo.

pauliani Jarrige, 1978: 276 (Leptacinus; Type locality: Andringitra Centre, plateau Soanindrana, Andohariana, 2090 m).

  • Distribution: Madagascar.

pennatus Fauvel, 1907: 33 (Leptacinus; Type locality: Kilimandjaro: Kiboscho).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; catalog).

  • Distribution: Tanzania.

perexilis Casey, 1906, see: batychrus Gyllenhal, 1827.

perforatus Tottenham, 1956: 240 (Leptacinus; Type locality: Urundi: Bururi, 2100 m).

  • Distribution: Burundi.

perplexus Cameron, 1950g: 30 (Leptacinus; Type locality: Belgian Congo: Nyasheke, volc. Nyamuragira, alt. 1820 m).

  • Distribution: Congo.

persimilis Cameron, 1932a: 139 (Leptacinus; Type locality: Pahang: Raub).

  • Distribution: Malaysia.

persimilis Tottenham, 1939, see: ajax Tottenham, 1949.

philippinus Bernhauer, 1916h: 423 (Leptacinus; Type locality: Los Baños: P.I.).

  • — Scheerpeltz, 1933: 1303 (Leptacinus; catalog).

  • Distribution: Philippines.

pholeobius Jeannel and Paulian, 1945: 76 (Leptacinus; larval description also published; Type locality: camp II de l'Elgon).

  • Distribution: Kenya.

picticornis Olliff, 1887, see: Notolinus.

picticornis Blackburn, 1888, see: Notolinus.

planulatus Sharp, 1889: 252 (Leptacinus; Type locality: Hitoyoshi; Kuma Kuni).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; catalog).

  • Distribution: Japan.

politus Fauvel, 1905b: 168 (Leptacinus; Type locality: Tananarive).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; catalog).

  • Distribution: Madagascar.

potockajae Coiffait, 1966, see: Allolinus.

priapus Coiffait, 1956a: 46 (Leptacinus; Type locality: Syrie: Hedjaneh).

  • — Coiffait, 1972: 147 (Leptacinus; characters; references; Syria).

  • Distribution: Syria.

procerulus Stephens, 1833, see: pusillus Stephens, 1833.

pubipennis Kolenati, 1846, see: Neobisnius.

pusillimus Kraatz, 1859: 112 (Leptacinus; Type locality: Ceylan).

  • — Bernhauer, 1902b: 32 (Leptacinus; Ceylon).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; catalog).

  • — Cameron, 1932: 12 (Leptacinus; characters; Ceylon).

  • — Scheerpeltz, 1933: 1303 (Leptacinus; catalog).

  • Distribution: Sri Lanka.

pusillus Stephens, 1833: 264 (Gyrohypnus; Type locality: London).

  • — Stephens, 1839: 404 (Leptacinus; characters; England).

  • — Hardy, 1851: 39 (Leptacinus; synonym of linearis).

  • — G. Waterhouse, 1858: 26 (Leptacinus; synonym of linearis).

  • — Crotch, 1867a: 383 (Leptacinus; Azores).

  • — J. Sahlberg, 1876: 37 (Leptacinus; Finland).

  • — Ganglbauer, 1895: 487 (Leptacinus; synonym of batychrus).

  • — Fauvel, 1902b: 100 (Leptacinus; synonym of batychrus).

  • — Reitter, 1908: 112 (Leptacinus; synonym of linearis).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; synonym of linearis).

  • — Méquignon, 1942a: 19 (Leptacinus; valid species; characters; Azores).

  • — Méquignon, 1943: 159, 160 (Leptacinus; valid species; characters; notes; synonyms).

  • — Coiffait, 1956a: 45 (Leptacinus; characters in key; France; Czechoslovakia).

  • — Smetana, 1958: 90 (Leptacinus; subgenus Leptacinus; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1960d: 135 (Leptacinus; Portugal).

  • — Smetana, 1963b: 35 (Leptacinus; Madeira).

  • — Scheerpeltz, 1964a: 299 (Leptacinus; Crete).

  • — Szujecki, 1968a: 725 (Leptacinus; Poland).

  • — Coiffait, 1972: 145 (Leptacinus; characters; references; Europe; Czechoslovakia; Portugal; Britain; Scandinavia; Siberia; France).

  • — Pope, 1977: 28 (Leptacinus; Britain).

  • — Smetana, 1982: 94 (Leptacinus; characters; North American introduction; Ontario; Quebec; Arizona; California; Colorado; Illinois; Iowa; Massachusetts; Michigan; Nevada; New Hampshire; New Jersey; New Mexico; New York; Pennsylvania; South Carolina; Washington; Wisconsin).

  • — Bordoni, 1982: 182 (Leptacinus; characters; Italy).

  • — Tóth, 1989: 11 (Leptacinus; characters; Hungary).

  • — Borges, 1990: Table II (Leptacinus; Azores).

  • — Kuschel, 1990: 26 (Leptacinus; introduced, Palaearctic species; winged; habitat; New Zealand).

  • — Ciceroni, 1993: 70 (Leptacinus; Italy).

  • — Janák, 1993a: 47 (Leptacinus; Czech Republic; Slovakia).

  • — Ciceroni, 1994: 113 (Leptacinus; aedeagal illustration; Italy; France; England; Switzerland).

  • — Ciceroni and Zanetti, 1995: 25 (Leptacinus; Italy).

  • — Downie and Arnett, 1996: 399 (Leptacinus; characters; USA).

  • — Silfverberg, 1999: 50 (Leptacinus; Estonia).

  • Distribution: Sudan, Canary Islands, Madeira, Azores, Europe, Russia, Afghanistan, Australia, New Zealand, Canada, USA.

linearis Gravenhorst, 1802: 43 (Staphylinus; [preoccupied]; Type locality: Not cited, but presumably Brunsvigae).

  • — Latreille, 1804: 329 (Staphylinus; characters; Germany).

  • — Gravenhorst, 1806: 97 (Staphylinus).

  • — Mannerheim, 1830: 34 (Gyrohypnus; Sweden).

  • — Mannerheim, 1831: 448 (Gyrohypnus; Sweden).

  • — Nordmann, 1837: 120 (Gyrohypnus; Germany; Russia).

  • — Erichson, 1839a: 430 (Leptacinus; synonym of batychrus).

  • — Heer, 1839: 243 (Leptacinus; characters; Switzerland).

  • — Erichson, 1839b: 335 (Leptacinus; synonym of batychrus).

  • — Hardy, 1851: 39 (Leptacinus; notes; England).

  • — Kraatz, 1857d: 649 (Leptacinus; characters; Germany).

  • — Thomson, 1860: 193 (Leptacinus; characters; Scandinavia).

  • — Wollaston, 1860: 101 (Leptacinus; characters; notes; Madeira).

  • — Wollaston, 1864: 580 (Leptacinus; characters; notes; Canary Islands).

  • — Fauvel, 1865a: 298 [= 1865b: 48] (Xantholinus; habitat; France).

  • — Wollaston, 1865: 498 (Leptacinus; habitat; Madeira Islands; Canary Islands).

  • — Baudi, 1870: 389 (Leptacinus; characters).

  • — Wollaston, 1871: 305 (Leptacinus; notes; synonym of pusillus).

  • — Fauvel, 1873b: 89 [= 1873a: 375] (Leptacinus; synonym of batychrus).

  • — Redtenbacher, 1874: 221 (Leptacinus; characters; Austria).

  • — Seidlitz, 1875: 280 (Leptacinus; characters; Baltic region).

  • — Mulsant and Rey, 1877a: 326 (Leptacinus; adult and larval characters; France).

  • — Olliff, 1887: 476 (Leptacinus; characters; Australia).

  • — Blackburn, 1888: 7 (Leptacinus; notes; Australia).

  • — Fowler, 1888: 293 (Leptacinus; characters; Britain).

  • — Ganglbauer, 1895: 487 (Leptacinus; variety of batychrus).

  • — Fauvel, 1902b: 100 (Leptacinus; synonym of batychrus).

  • — Reitter, 1908: 112 (Leptacinus; characters; Europe).

  • — Reitter, 1909: 135 (Leptacinus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; variety of batychrus).

  • — Johansen, 1914: 439 (Leptacinus; characters; Denmark).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; catalog).

  • — Tottenham, 1939d: 231 (Leptacinus; synonym of sulcifrons).

  • — Méquignon, 1943: 160 (Leptacinus; synonym of pusillus).

  • — Hinton, 1945: 67 (Leptacinus; characters).

  • — V. Hansen, 1952: 11 (Leptacinus; characters; Denmark).

  • — Smetana, 1958: 90 (Leptacinus; synonym of pusillus).

  • — Scheerpeltz, 1960b: 68 (Leptacinus; Afghanistan).

  • — Scheerpeltz, 1961d: 126 (Leptacinus; Italy).

  • — Lohse, 1961: 183, 184, 186 (Leptacinus; characters).

  • — Brakman, 1962: 47 (Leptacinus; characters).

  • — Smetana, 1963b: 35 (Leptacinus; synonym of pusillus).

  • — Scheerpeltz, 1963: 10 (Leptacinus; Afghanistan).

  • — Palm, 1963b: 44 (Leptacinus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 160 (Leptacinus; characters; central Europe).

  • — Horion, 1965: 88 (Leptacinus; Siberia; all Europe).

  • — Coiffait, 1972: 145 (Leptacinus; synonym of pusillus).

  • — Scheerpeltz, 1974c: 18 (Leptacinus; Sudan).

  • — Pope, 1977: 28 (Leptacinus; synonym of pusillus).

  • — Smetana, 1982: 94 (Leptacinus; synonym of pusillus).

  • — Lucht, 1987: 102 (Leptacinus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Lohse and Lucht, 1989: 156 (Leptacinus; synonym of pusillus).

  • — Ciceroni and Zanetti, 1995: 25 (Leptacinus; synonym of pusillus).

procerulus Stephens, 1833: 264 (Gyrohypnus; Type locality: London).

  • — Stephens, 1839: 403 (Xantholinus; characters; England).

  • — G. Waterhouse, 1858: 26 (Leptacinus; synonym of linearis).

  • — Ganglbauer, 1895: 487 (Leptacinus; synonym of batychrus).

  • — Reitter, 1908: 112 (Leptacinus; synonym of linearis).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; synonym of batychrus).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; synonym of linearis).

minutus Lacordaire, 1835: 417 (Xantholinus; Type locality: Paris).

  • — Erichson, 1839a: 430 (Leptacinus; synonym of batychrus).

  • — Erichson, 1839b: 335 (Leptacinus; synonym of batychrus).

  • — Fairmaire and Laboulbène, 1856: 503 (Leptacinus; synonym of batychrus).

  • — Kraatz, 1857d: 649 (Leptacinus; synonym of linearis).

  • — Ganglbauer, 1895: 487 (Leptacinus; synonym of batychrus).

  • — Reitter, 1908: 112 (Leptacinus; synonym of linearis).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; synonym of batychrus).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; synonym of linearis).

  • — Méquignon, 1943: 160 (Leptacinus; synonym of pusillus).

nigritulus LeConte, 1880: 169 (Leptacinus; Type locality: Michigan; [Note: See lectotype designation by Smetana, 1982]. Also cited from Canada).

  • — Casey, 1906: 402 (Leptacinodes; characters; Iowa; Wisconsin, California).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; catalog).

  • — Hatch, 1957: 239 (Leptacinus; characters; British Columbia; Idaho; Oregon).

  • — Smetana, 1982: 94 (Leptacinus; lectotype designation: Mic. [Michigan]; synonym of pusillus).

quadrisulciceps Lea, 1925: 216 (Leptacinus; Type locality: Tasmania: Launceston; Hobart).

  • — Scheerpeltz, 1933: 1303 (Leptacinus; catalog).

  • Distribution: Australia.

quadrisulcus Stephens, 1833, see: batychrus Gyllenhal, 1827.

raptor Tottenham, 1956: 243 (Leptacinus; Type locality: Urundi: terr. Bururi, Nyamurembe, 900 m).

  • Distribution: Burundi.

rattii Coiffait, 1969, see: intermedius Donisthorpe, 1936.

rigatus† Scudder, 1900: 59 (Leptacinus; fossil; Type locality: Florissant, Colorado).

  • — Handlirsch, 1907: 726 (Leptacinus; catalog; Miocene; Colorado).

  • Distribution: USA.

rubricollis Casey, 1906, see: Phacophallus.

rubricollis Reitter, 1899, see: Phacophallus.

rufonitens Coiffait, 1966a: 124 (Leptacinus; Type locality: Caucase).

  • — Coiffait, 1972: 150 (Leptacinus; characters; references; Caucasus).

  • Distribution: Caucasus.

rufoterminatus Cameron, 1951b: 126 (Leptacinus; Type locality: Sénégal: Linguère).

  • — Tottenham, 1961: 203 (Leptacinus; Senegal).

  • Distribution: Senegal.

sanctaehelenae Steel, 1949, see: Notolinus.

sardous Fiori, 1894, see: Phacophallus.

schoutedeni Cameron, 1929: 59 (Leptacinus; Type locality: Kivu: Lulenga; Burunga).

  • — Scheerpeltz, 1933: 1303 (Leptacinus; catalog).

  • — Cameron, 1950g: 29 (Leptacinus; Belgian Congo).

  • Distribution: Congo.

secretus Cameron, 1933d: 384 (Leptacinus; Type locality: Sumatra: Fort de Kock).

  • Distribution: Indonesia.

semistriatus Stephens, 1833, see: batychrus Gyllenhal, 1827.

seriatus LeConte, 1880, see: batychrus Gyllenhal, 1827.

serpentarius Motschulsky, 1860a: 76 (Xanthophius; Type locality: Ceylan).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; catalog).

  • — Cameron, 1932: 11 (Leptacinus; characters; Ceylon).

  • — Scheerpeltz, 1933: 1303 (Leptacinus; catalog).

  • — Cameron, 1934d: 79 (Leptacinus; type examined; synonym of filum).

  • — Paulian, 1941: 207 (Leptacinus; cited as valid species; larval characters; India).

  • — Scheerpeltz, 1957c: 232 (Leptacinus; cited as valid species; Sumba).

  • Distribution: India, Sri Lanka, Indonesia.

sexsulciceps Lea, 1925: 218 (Leptacinus; Type locality: Victoria: Melbourne; Belgrave. Tasmania: Launceston).

  • — Scheerpeltz, 1933: 1303 (Leptacinus; catalog).

  • Distribution: Australia.

siculus Coiffait, 1956, see: faunus Coiffait, 1956.

silvanus Coiffait, 1956, see: intermedius Donisthorpe, 1936.

simeki Bordoni, 1975a: 149 (Leptacinus; Type locality: N-Afghanistan: Prov. Kataghan: 10 km. sö. Kunduz, 450 m).

  • Distribution: Afghanistan.

sinuatocollis Scheerpeltz, 1957c: 233, 309 (Leptacinus; Type locality: Sumba: Mau Marru).

  • Distribution: Indonesia.

sinuatocollis Scheerpeltz, 1974, see: hariolus Herman, 2001.

sobrinus Tottenham, 1956: 244 (Leptacinus; Type locality: Ruanda: terr. Nyanza, Gitarama, 1850 m).

  • Distribution: Rwanda.

sparsepunctus Coiffait, 1968: 133 (Leptacinus; subgenus Leptacinus; Type locality: Gabon: Belinga).

  • Distribution: Gabon.

spectabilis Bernhauer, 1929f: 118 (Leptacinus; Type locality: Haut-Uele: Watsa).

  • — Scheerpeltz, 1933: 1303 (Leptacinus; catalog).

  • Distribution: Congo.

strangulatus Coiffait, 1968: 132 (Leptacinus; subgenus Leptacinus; Type locality: Gabon: Belinga).

  • Distribution: Gabon.

sublaevis Fauvel, 1875, see: peliopterus Solsky, 1875.

substrigosus Joy, 1913, see: batychrus Gyllenhal, 1827.

subtilepunctatus Scheerpeltz, 1974: 112 (Leptacinus; Type locality: Basutoland: Qachas Nek, 20 miles N Matatiele).

  • Distribution: Lesotho.

subtilis Erichson, 1839, see: Somoleptus.

sulcifrons Stephens, 1833: 260 (Gyrohypnus; Type locality: Suffolk; Devonshire; London; Norfolk).

  • — Stephens, 1839: 404 (Leptacinus; characters; England).

  • — Hardy, 1851: 39 (Leptacinus; synonym of linearis).

  • — G. Waterhouse, 1858: 26 (Leptacinus; synonym of linearis).

  • — Ganglbauer, 1895: 487 (Leptacinus; synonym of batychrus).

  • — Reitter, 1908: 112 (Leptacinus; synonym of linearis).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; synonym of batychrus).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; synonym of linearis).

  • — Méquignon, 1943: 159, 160 (Leptacinus; valid species; characters; notes; synonyms).

  • — Coiffait, 1956a: 49 (Leptacinus; characters in key; France; Czechoslovakia).

  • — A. Strand, 1957 (Leptacinus; identity of species).

  • — Smetana, 1958: 91 (Leptacinus; subgenus Leptacinus; characters; ecology; Czechoslovakia).

  • — Lohse, 1961: 183, 184, 185 (Leptacinus; characters).

  • — Scheerpeltz, 1963: 10 (Leptacinus; Afghanistan).

  • — Palm, 1963b: 44 (Leptacinus; characters; Sweden; Norway).

  • — Lohse, 1964: 159 (Leptacinus; characters; central Europe).

  • — Scheerpeltz, 1964a: 299 (Leptacinus; Crete).

  • — Horion, 1965: 87 (Leptacinus; Britain; Norway; Sweden; France; Germany; Austria; Czechoslovakia; Poland; Russia; Afghanistan).

  • — Smetana, 1967d: 196 (Leptacinus; Mongolia).

  • — Szujecki, 1968a: 725 (Leptacinus; Poland).

  • — Coiffait, 1970d: 143 (Leptacinus; Uzbekistan).

  • — Coiffait, 1972: 151 (Leptacinus; characters; references; England; Czechoslovakia; Croatia; Tyrol; Iberia; France; Norway).

  • — Pope, 1977: 28 (Leptacinus; synonym of pusillus).

  • — Tronquet, 1981: 70 (Leptacinus; Afghanistan).

  • — Bordoni, 1982: 191 (Leptacinus; characters; Italy).

  • — Lucht, 1987: 102 (Leptacinus; checklist; Germany; Poland; Czechoslovakia; Austria; Denmark; Sweden).

  • — Janák, 1989: 167 (Leptacinus; notes; Sudan).

  • — Tóth, 1989: 12 (Leptacinus; characters; Hungary).

  • — Janák, 1993a: 47 (Leptacinus; Czech Republic; Slovakia).

  • — Ciceroni, 1993: 70 (Leptacinus; Italy).

  • — Ciceroni, 1994: 114 (Leptacinus; aedeagal illustration; Italy; Corsica; Spain; Switzerland).

  • — Ciceroni and Zanetti, 1995: 25 (Leptacinus; Italy).

  • — Assing, Frisch, Kahlen, et al., 1998: 136 (Leptacinus; notes).

  • — Kashcheev, 1999a: 57 (Leptacinus; Kazakhstan).

  • Distribution: Sudan, Morocco, Europe, Afghanistan, Uzbekistan, Kazakhstan, Mongolia.

angustatus Grimm, 1845: 134 (Leptacinus; Type locality: Berlin).

  • — Redtenbacher, 1849: 824 (Leptacinus; characters).

  • — Redtenbacher, 1857: 183 (Leptacinus; characters; Germany).

  • — Ganglbauer, 1895: 487 (Leptacinus; synonym of batychrus).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; synonym of batychrus).

  • — Scheerpeltz, 1933: 1302 (Leptacinus; synonym of linearis).

  • — Méquignon, 1943: 160 (Leptacinus; synonym of sulcifrons).

ops Coiffait, 1956a: 48 (Leptacinus; Type locality: Gironde: Merignac).

  • — Coiffait, 1958: 24 (Leptacinus; notes).

  • — Smetana, 1958: 90 (Leptacinus; subgenus Leptacinus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 159 (Leptacinus; characters; central Europe).

  • — Coiffait, 1972: 152 (Leptacinus; characters; references; France; Czechoslovakia; Germany; Ukraine; Morocco).

  • — Coiffait, 1973a: 273 (Leptacinus; Morocco).

  • — Drugmand and Haghebaert, 1987: 325 (Leptacinus; Belgium).

  • — Lucht, 1987: 102 (Leptacinus; checklist; Czechoslovakia; Austria).

  • — Tóth, 1989: 12 (Leptacinus; characters).

  • — Lohse and Lucht, 1989: 156 (Leptacinus; notes).

  • — Janák, 1993a: 47 (Leptacinus; Czech Republic ?; Slovakia ?).

  • — Ciceroni, 1994: 114 (Leptacinus; synonym of sulcifrons).

  • — Assing, Frisch, Kahlen, et al., 1998: 136 (Leptacinus; synonym of sulcifrons).

suturalis Lea, 1925, see: minimus Bernhauer, 1920.

tenellus Erichson, 1839, see: Heterocinus.

tener Waltl, 1835, see: batychrus Gyllenhal, 1827.

tenuis Cameron, 1935a: 373 (Leptacinus; Type locality: Stanleyville).

  • Distribution: Congo.

testaceipennis Fauvel, 1863, see: Neoxantholinus.

testaceipennis Bernhauer, 1929f: 119 (Leptacinus; [preoccupied]; Type locality: Ituri: La Moto).

  • — Scheerpeltz, 1933: 1303 (Leptacinus; catalog).

  • Distribution: Congo.

titan Coiffait, 1980: 264 (Leptacinus; replacement name for major Coiffait).

  • Distribution: Bhutan.

major Coiffait, 1977b: 216 (Leptacinus; [preoccupied]; Type locality: Bhutan: Thimphu).

  • — Coiffait, 1980: 264 (Leptacinus; synonym of titan).

trapeziceps Scheerpeltz, 1974c: 18 (Leptacinus; Type locality: Ethiopia: Mai Chew).

  • Distribution: Ethiopia.

triangulum Saulcy, 1864, see: batychrus Gyllenhal, 1827.

tricolor Kraatz, 1859, see: Phacophallus.

trigonocephalus Kraatz, 1859, see: Phacophallus.

unicolor Cameron, 1937: 1 (Leptacinus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft).

  • Distribution: Indonesia.

varius H. Bauer, 1914 [Note: Nomen nudum], see: batychrus Gyllenhal, 1827.

vicinus Coiffait, 1968: 137 (Leptacinus; subgenus Chaetocinus; Type locality: Gabon: Belinga).

  • Distribution: Gabon.

vuldiformis Coiffait, 1977, see: Allolinus.

wittei Cameron, 1950g: 30 (Leptacinus; Type locality: Belgian Congo: Ngesho, alt. 2000 m).

  • Distribution: Congo.

xanthopygus Scheerpeltz, 1974, see: Phacophallus.

zanettii Bordoni, 1978, see: othioides Baudi, 1870.

Leptomicrus [1 species; Oriental Region]

Leptomicrus Fauvel, 1878f: 240 (species included: teredo). Type species: Leptomicrus teredo Fauvel, fixed by monotypy.

  • — Duvivier, 1883: 159 (catalog).

  • — Bernhauer and Schubert, 1914: 295 (world catalog; 1 species).

  • — Blackwelder, 1952: 217 (type species: teredo).

teredo Fauvel, 1878: 241 (Leptomicrus; Type locality: Nouvelle-Guinée, Hatam).

  • — Bernhauer and Schubert, 1914: 295 (Leptomicrus; catalog).

  • Distribution: New Guinea.

Leptophius [1 species; Palaearctic Region]

Leptophius Coiffait, 1983b: 345 (replacement name for Leptophallus Coiffait). Type species: Xantholinus flavocinctus Hochhuth (= Xantholinus relucens sensu Kraatz), fixed by objective synonymy with Leptophallus.

  • — Bordoni, 1985a: 83 (valid genus; type species flavocinctus).

  • — Janák, 1993a: 47 (in list of species of Czech Republic and Slovakia).

  • — Herman, 2001: 10 (type species).

Leptophallus Coiffait, 1956a: 57, 59 (subgenus of Xantholinus; species included: relucens, elianae; preoccupied by Lueche, 1909). Type species: Xantholinus flavocinctus Hochhuth (= Xantholinus relucens sensu Kraatz), fixed by application of Article 70.3.2 (see Herman, 2001).

  • — Coiffait, 1962b: 394 (subgenus of Xantholinus; characters).

  • — Lohse, 1964: 163 (subgenus of Xantholinus; characters; key to central European species).

  • — Coiffait, 1972: 219, 222 (subgenus of Xantholinus; characters; type species: flavocinctus; key to species of western Palaearctic region).

  • — Bordoni, 1972: 160, 164 (subgenus Xantholinus; characters; notes).

  • — Bordoni, 1975: 87 (subgenus of Xantholinus; distributional notes).

  • — Bordoni, 1982: 240, 248 (subgenus of Xantholinus; flavocinctus cited as type species).

  • — Bordoni, 1985a: 83 (synonym of Leptophius; flavocinctus cited as type species).

  • — Janák, 1993a: 47 (synonym of Leptophius).

  • — Herman, 2001: 10 (type species; discussion; fixed by application of article 70.3.2).

flavocinctus Hochhuth, 1849: 102 (Xantholinus; Type locality: Caucasus; [Note: See lectotype designation by Gusarov, 1992a]. Originally cited from Imeretien).

  • — Kraatz, 1857d: 634 (Xantholinus; [Note: Misidentified as relucens according to Coiffait, 1972: 257, who earlier (1956a: 57, 59) had cited the name as type species of Leptophallus]; characters; Germany).

  • — Fauvel, 1873b: 101 [= 1873a: 387] (Xantholinus; synonym of glabratus).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; synonym of relucens).

  • — Scheerpeltz, 1933: 1314 (Xantholinus; aberration of relucens).

  • — Coiffait, 1966: 21 (Xantholinus; subgenus Leptophallus; Turkey).

  • — Coiffait, 1970e: 61 (Xantholinus; Cyprus).

  • — Coiffait, 1972: 257 (Xantholinus; subgenus Leptophallus; characters; references; Croatia; Austria; Tyrol; Czechoslovakia; Germany).

  • — Bordoni, 1972: 160 (Xantholinus; subgenus Leptophallus; characters; Italy).

  • — Bordoni, 1975: 58 (Xantholinus; subgenus Leptophallus; notes; Crimea).

  • — Bordoni, 1976: 230 (Xantholinus; notes; Turkey).

  • — Bordoni, 1976c: 84 (Xantholinus; Bosnia; Herzegovina; Dalmatia; Macedonia; Crete; Cyprus; Turkey).

  • — Bordoni, 1976d: 120 (Xantholinus; Dalmatia; Bosnia).

  • — Bordoni, 1982: 248 (Xantholinus; subgenus Leptophallus; characters; Italy).

  • — Bordoni, 1985a: 83 (Leptophius; anatomical studies of labrum, aedeagus, and male genital segment).

  • — Bordoni, 1986b: 385 (Leptophius; Dalmatia; Greece; Turkey).

  • — Ádám, 1987: 146 (Leptophallus; collecting notes; Hungary).

  • — Tóth, 1989: 20 (Xantholinus; characters; Hungary).

  • — Lohse and Lucht, 1989: 158 (Xantholinus; notes).

  • — Gusarov, 1992a: 785 [= 1993a: 74] (Xantholinus; subgenus Leptophallus; lectotype designation: Caucasus).

  • — Ciceroni and Zanetti, 1995: 25 (Xantholinus; Italy).

  • — Janák, 1993a: 47 (Leptophius; Czech Republic; Slovakia).

  • — Herman, 2001: 10 (Leptophius; type species of Leptophallus; fixed by application of Article 70.3.2).

  • Distribution: Europe, Turkey, Georgia.

ochropterus Nordmann, 1837: 118 (Gyrohypnus; Type locality: Corcyra; Tauria).

  • — Erichson, 1839b: 320 (Xantholinus; synonym of glabratus).

  • — Küster, 1848: 9 (Xantholinus; synonym of glabratus).

  • — Redtenbacher, 1849: 691 (Xantholinus; characters; Austria).

  • — Kraatz, 1857d: 634 (Xantholinus; synonym of relucens).

  • — Redtenbacher, 1857: 181 (Xantholinus; characters; Austria).

  • — Fauvel, 1869: 490 (Xantholinus; synonym of relucens).

  • — Mulsant and Rey, 1877a: 266 (Xantholinus; synonym of relucens).

  • — Ganglbauer, 1895: 481 (Xantholinus; synonym of relucens).

  • — Reitter, 1908: 116 (Xantholinus; synonym of relucens).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; synonym of relucens).

  • — Scheerpeltz, 1933: 1314 (Xantholinus; synonym of relucens).

  • — Smetana, 1958: 105 (Xantholinus; synonym of relucens).

  • — Bordoni, 1971: 685 (Xantholinus; subgenus Leptophallus; Turkey).

  • — Bordoni, 1973c: 151 (Xantholinus; subgenus Leptophallus; cited as valid species; Romania; Palestine; Lebanon; Turkey; Rhodes; Cyprus; Serbia; Hercegovina; Croatia; Austria; Italy; france; Portugal).

  • — Bordoni, 1975: 58 (Xantholinus; synonym of flavocinctus).

hebraicus Reiche and Saulcy, 1856: 362 (Xantholinus; Type locality: Chypre).

  • — Peyron, 1858: 422 (Xantholinus; Turkey).

  • — Fauvel, 1869: 490 (Xantholinus; synonym of relucens).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; synonym of relucens).

  • — Coiffait, 1972: 257 (Xantholinus; synonym of flavocinctus).

  • — Bordoni, 1972: 164 (Xantholinus; synonym of flavocinctus).

relucens sensu Kraatz, 1857d: 634 (Xantholinus; [Note: Misidentification: Kraatz attributed the species to Gravenhorst]; characters; Dalmatia; Rhodes).

  • — Coiffait, 1956a: 59 (Xantholinus; subgenus Leptophallus; characters; Lebanon; Turkey; Austria; Croatia; Switzerland; Pyrénées).

  • — Coiffait, 1972: 257 (Xantholinus; subgenus Leptophallus; synonym of flavocinctus).

hebraicus Reiche and Saulcy, 1856, see: flavocinctus Hochhuth, 1849.

ochropterus Nordmann, 1837, see: flavocinctus Hochhuth, 1849.

relucens sensu Kraatz, 1857 (Nomen nudum, misidentification), see: flavocinctus Hochhuth, 1849.

Leurocorynus [1 species; Oceanic Region]

Leurocorynus Sharp, 1908: 548 (species included: cephalotes). Type species: Leurocorynus cephalotes Sharp, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 295 (world catalog; 1 species).

  • — Blackwelder, 1952: 220 (type species: cephalotes).

cephalotes Sharp, 1908: 549 (Leurocorynus; Type locality: Kauai: 4000 ft; mts. Waimea, 4000 ft.; Koholuamano, 4000 ft. Maui: Haleakala, 4500 ft. Molokai: 4000 ft. Lanai, 3000 and 2000 ft. Hawaii: Hualalai).

  • — Bernhauer and Schubert, 1914: 295 (Leurocorynus; catalog).

  • — Nishida, 1994: 72 (Leurocorynus; Hawaii).

  • — Nishida, 1997: 61 (Leurocorynus; Hawaii).

  • Distribution: Hawai.

Linohesperus [26 species; Nearctic Region]

Linohesperus Smetana, 1982: 143 (species included: borealis, cuspifer, trihamatus, hamatus, brunnescens, subhamatus, rhomboides, rutilus, nanus, emarginatus, obtusus, piceus, hermani, aculeus, mutilus, similis, indistinctus, obtusus, barbatus, hesperius, pilosus, alticola, effeminatus). Type species: Leptacinus brunnescens LeConte, fixed by original designation.

  • — Smetana, 1988a: 550 (revised key to species).

  • — Campbell and Davies, 1991: 123 (checklist of species; Canada).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 391 (25 Nearctic species; notes; characters in key).

aculeus Smetana, 1982: 152 (Linohesperus; Type locality: California: Inyo Co.: Westguard Pass).

  • — Smetana, 1988a: 542 (Linohesperus; collecting notes; California).

  • Distribution: USA.

alticola Casey, 1906: 414 (Hesperolinus; Type locality: Nevada: Reno).

  • — Bernhauer and Schubert, 1914: 296 (Hesperolinus; catalog).

  • — Hatch, 1957: 240 (Hesperolinus; characters; Washington; Idaho).

  • — Smetana, 1982: 161 (Linohesperus; characters; California; Nevada).

  • Distribution: USA.

angustus Casey, 1906, see: nanus LeConte, 1880.

barbatus Smetana, 1982: 158 (Linohesperus; Type locality: California: Marin Co.: Mill Valley).

  • — Smetana, 1988a: 547 (Linohesperus; California).

  • Distribution: USA.

bicolor Casey, 1906, see: brunnescens LeConte, 1880.

borealis Casey, 1906: 414 (Hesperolinus; Type locality: Victoria, Van. Id.; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 297 (Hesperolinus; catalog).

  • — Hatch, 1957: 240 (Hesperolinus; characters; British Columbia; Washington; Oregon).

  • — Smetana, 1982: 153 (Linohesperus; lectotype designation: Victoria, Van. Id.; characters; British Columbia; California; Oregon; Washington).

  • — Smetana, 1988a: 545 (Linohesperus; collecting notes; Calfornia).

  • Distribution: Canada, USA.

brunnescens LeConte, 1880: 169 (Leptacinus; Type locality: California; [Note: See lectotype designation by Smetana, 1982]. Originally cited from Gilroy, California).

  • — Bernhauer and Schubert, 1914: 297 (Hesperolinus; synonym of parcus).

  • — Smetana, 1982: 163 (Linohesperus; lectotype designation: Cala [California]; characters; California).

  • Distribution: USA.

bicolor Casey, 1906: 413 (Hesperolinus; Type locality: southern California; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 297 (Hesperolinus; catalog).

  • — Smetana, 1982: 163 (Linohesperus; lectotype designation; synonym of brunnescens).

cuspifer Smetana, 1982: 155 (Linohesperus; Type locality: California: Riverside Co.: San Timoteo Can. Also cited from Alberta and British Columbia).

  • — Smetana, 1988a: 545 (Linohesperus; California).

  • Distribution: Canada, USA.

effeminatus Smetana, 1982: 158 (Linohesperus; Type locality: California: Inyo Co.: Cottonwood Cr.).

  • — Smetana, 1988a: 546 (Linohesperus; California).

  • Distribution: USA.

emarginatus Smetana, 1982: 147 (Linohesperus; Type locality: California: Modoc Co.: Saddle Blkt. Flat. Also cited from Oregon).

  • — Smetana, 1988a: 541 (Linohesperus; collecting notes; California).

  • Distribution: USA.

hamatus Smetana, 1982: 170 (Linohesperus; Type locality: California: Inyo Co.: South Fork Oak Cr., 6500 feet. Also cited from Colorado).

  • — Smetana, 1988a: 550 (Linohesperus; California).

  • Distribution: USA.

hermani Smetana, 1982: 150 (Linohesperus; Type locality: Arizona: Graham Co.: Pinaleno Mts., Mi. Post 127, 6500 feet).

  • — Smetana, 1988a: 542 (Linohesperus; collecting notes; Utah).

  • — Smetana, 1990c: 85 (Linohesperus; Arizona).

  • Distribution: USA.

hesperius Smetana, 1982: 159 (Linohesperus; Type locality: California: Monterey).

  • — Smetana, 1988a: 547 (Linohesperus; California).

  • Distribution: USA.

iaculator Smetana, 1988a: 543 (Linohesperus; Type locality: California: San Diego Co.: Cleveland N. F., Mt. Laguna, 5900 feet).

  • Distribution: USA.

indistinctus Smetana, 1982: 155 (Linohesperus; Type locality: California: San Luis Obispo Co.: Baywood Park).

  • Distribution: USA.

montivagus Smetana, 1988a: 546 (Linohesperus; Type locality: California: S. Bernard. Mts., 1 mi. E Fallsvale, 6200 feet).

  • Distribution: USA.

mutilus Smetana, 1982: 151 (Linohesperus; Type locality: Arizona: Walnut).

  • Distribution: USA.

nanus LeConte, 1880: 174 (Xantholinus; Type locality: San Diego, California).

  • — Casey, 1906: 396 (Gyrohypnus; characters; California).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Smetana, 1982: 162 (Linohesperus; characters; California).

  • Distribution: USA.

pomonae Casey, 1906: 414 (Hesperolinus; Type locality: California, Los Angeles Co.; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 297 (Hesperolinus; catalog).

  • — Smetana, 1982: 162 (Linohesperus; lectotype designation; synonym of nanus).

angustus Casey, 1906: 415 (Hesperolinus; Type locality: California: San Diego Co., Coronado; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 297 (Hesperolinus; catalog).

  • — Smetana, 1982: 162 (Linohesperus; lectotype designation; synonym of nanus).

obtusus Smetana, 1982: 156 (Linohesperus; Type locality: California: Riverside Co.: Temescal Canyon).

  • — Smetana, 1988a: 546 (Linohesperus; California).

  • Distribution: USA.

piceus Casey, 1906: 415 (Hesperolinus; Type locality: California, Siskiyou Co.; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 297 (Hesperolinus; catalog).

  • — Hatch, 1957: 240 (Hesperolinus; characters; Washington; Oregon).

  • — Smetana, 1982: 149 (Linohesperus; lectotype designation; characters; California; Oregon).

  • Distribution: USA.

pilosus Smetana, 1982: 160 (Linohesperus; Type locality: Idaho: Latah Co.: Moscow. Also cited from Washington).

  • Distribution: USA.

pomonae Casey, 1906, see: nanus LeConte, 1880.

priapus Smetana, 1988a: 547 (Linohesperus; Type locality: California: Fresno Co.: Redinger Lake Rd., 3.5 mi. NE Auberry).

  • Distribution: USA.

rhomboides Smetana, 1982: 166 (Linohesperus; Type locality: California: Cala. o.).

  • — Smetana, 1988a: 550 (Linohesperus; California; Washington).

  • Distribution: USA.

rutilus Smetana, 1982: 167 (Linohesperus; Type locality: California: Pasadena).

  • Distribution: USA.

similis Smetana, 1982: 152 (Linohesperus; Type locality: Arizona: Pima Co.: 2 mi. E. Tucson).

  • Distribution: USA.

spiculifer Smetana, 1988a: 542 (Linohesperus; Type locality: California: Riverside Co.: Whitewater Canyon).

  • Distribution: USA.

subhamatus Smetana, 1982: 168 (Linohesperus; Type locality: Colorado: Denver. Also cited from Alberta, Nebraska, and New Mexico).

  • — Smetana, 1988a: 550 (Linohesperus; California).

  • Distribution: USA.

trihamatus Smetana, 1982: 170 (Linohesperus; Type locality: California: Contra Costa Co.: Vine Hill).

  • Distribution: USA.

Linosomus [6 species; Ethiopian Region]

Linosomus Kraatz, 1857d: 647 (species included: tenuicornis; [Note: Kraatz included the species cited under “Leptacinus Fam. IV” in Erichson, 1838b: 338, which included only tenuicornis but did not explicitly name the species. According to Article 67.2.1, the included species must be actually cited by available name. Gemminger and Harold, 1868: 605 were the first do this by listing tenuicornis (Article 67.2.2]). Type species: Gyrohypnus tenuicornis Nordmann, fixed by subsequent monotypy by Gemminger and Harold, 1868.

  • — Gemminger and Harold, 1868: 605 (catalog; first included species).

  • — Bernhauer and Schubert, 1914: 296 (world catalog; 1 species).

  • — Blackwelder, 1952: 221 (type species: tenuicornis).

Notolinopsis Casey, 1906: 371, 377 (species included: incultus, latitarsis, languidus, capensis, diligens). Type species: Notolinopsis capensis Casey, fixed by subsequent designation by Blackwelder, 1943: 474.

  • — Bernhauer and Schubert, 1914: 300 (subgenus of Xantholinus).

  • — Cameron, 1932: 22 (subgenus of Xantholinus).

  • — Steel, 1949a: 268 (synonym of Linosomus).

  • — Blackwelder, 1952: 263 (synonym of Linosomus; type species).

Linosoma Eichelbaum, 1909: 168 (emendation of Linosomus). Type species: Linosoma tenuicorne Nordmann, fixed by objective synonymy with Linosomus.

  • — Bernhauer and Schubert, 1914: 296 (synonym of Linosomus).

capensis Casey, 1906: 379 (Notolinopsis; Type locality: Cape Town).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • Distribution: South Africa.

diligens Casey, 1906: 379 (Notolinopsis; Type locality: Cape Town).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • Distribution: South Africa.

incultus Casey, 1906: 378 (Notolinopsis; Type locality: Wellington).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • Distribution: South Africa.

languidus Casey, 1906: 379 (Notolinopsis; Type locality: Wellington).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • Distribution: South Africa.

latitarsis Casey, 1906: 378 (Notolinopsis; Type locality: Wellington).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • Distribution: South Africa.

tenuicornis Nordmann, 1837: 125 (Gyrohypnus; Type locality: Promontorium bon.[ae] sp.[ei]).

  • — Erichson, 1839b: 338 (Leptacinus; characters; Promontorio bonae spei).

  • — Gemminger and Harold, 1868: 605 (Linosomus).

  • Distribution: South Africa.

Liotesba [3 species; Palaearctic and Oriental Region]

Liotesba Scheerpeltz, 1965: 199 (species included: malaisei). Type species: Liotesba malaisei Scheerpeltz, fixed by original designation and monotypy.

malaisei Scheerpeltz, 1965: 201 (Liotesba; Type locality: N.E. Burma, Kambaiti, 7000 ft.).

  • Distribution: Myanmar.

punctiventris Sharp, 1889: 252 (Xantholinus; Type locality: Ichiuchi).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Bordoni, 1997: 171 (Liotesba; characters; Japan).

  • Distribution: Japan.

wittmeri Coiffait, 1977b: 219 (Liotesba; Type locality: Bhutan: Phuntsholing-Thimphu, 1680 m).

  • Distribution: Bhutan.

Lissohypnus [1 species; Neotropical and Nearctic Regions]

Lissohypnus Casey, 1906: 368, 398 (species included: texanus). Type species: Lissohypnus texanus Casey, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 309 (world catalog; 1 species).

  • — Leng, 1920: 105 (catalog of North American species).

  • — Blackwelder, 1952: 223 (type species: texanus).

  • — Moore, 1963a: 94 (characters).

  • — Moore and Legner, 1975: 81 (catalog; North America north of Mexico).

  • — Moore and Legner, 1979: 63 (characters; notes).

  • — Smetana, 1982: 261 (characters; habitus; revision of species of North America north of Mexico).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 391 (1 Nearctic species; notes; characters in key).

texanus Casey, 1906: 399 (Lissohypnus; Type locality: Texas, Austin).

  • — Bernhauer and Schubert, 1914: 309 (Lissohypnus; catalog).

  • — Coiffait, 1981g: 288 (Lissohypnus; Galapagos Islands).

  • — Smetana, 1982: 262 (Lissohypnus; characters; Louisiana; Texas).

  • Distribution: USA, Galapagos Islands.

Lithocharodes [32 species; Neotropical, Nearctic, Oriental, and Oceanic Regions]

Lithocharodes Sharp, 1876: 204 (species included: fuscipennis). Type species: Lithocharodes fuscipennis Sharp, fixed by monotypy.

  • — Duvivier, 1883: 157 (catalog).

  • — Lynch, 1884: 193 (characters).

  • — Sharp, 1885: 491 (notes; Mexican and Central American species).

  • — Casey, 1906: 370, 409 (characters).

  • — Bernhauer and Schubert, 1914: 296 (world catalog; 16 species).

  • — Leng, 1920: 106 (catalog of North American species).

  • — Blackwelder, 1943: 496 (characters; type species: fuscipennis; key to species of West Indies).

  • — Blackwelder, 1944: 146 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 224 (type species: fuscipennis).

  • — Moore, 1963a: 95 (characters).

  • — Moore and Legner, 1975: 82 (catalog; North America north of Mexico).

  • — Moore and Legner, 1979: 64 (characters; notes).

  • — Smetana, 1982: 120 (characters; habitus; revision of and key to species of North America north of Mexico).

  • — Frank, 1986a: 369 (checklist of species of Florida).

  • — Downie and Arnett, 1996: 400 (characters).

  • — Lundgren, 1998: 50 (list of species from Florida; records for other States listed for each species).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 391 (7 Nearctic species; notes; characters in key).

Lithocharitoides Rye, 1878: 36 (emendation of Lithocharodes). Type species: Lithocharodes fuscipennis Sharp, fixed by objective synonymy with Lithocharodes.

  • — Smetana, 1982: 120 (synonym of Lithocharodes).

Nematolinus Casey, 1906: 370, 407 (species included: longicollis). Type species: Leptacinus longicollis LeConte, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 297 (world catalog; 1 species).

  • — Leng, 1920: 106 (catalog of North American species).

  • — Blackwelder, 1952: 257 (type species: longicollis).

  • — Moore, 1963a: 95 (characters).

  • — Moore and Legner, 1975: 85 (catalog; North America north of Mexico).

  • — Moore and Legner, 1979: 68 (characters; notes).

  • — Smetana, 1982: 120 (synonym of Lithocharodes).

Oligolinus Casey, 1906: 372, 420 (species included: floridanus). Type species: Metoponcus floridanus LeConte, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 291 (world catalog; 13 species).

  • — Leng, 1920: 106 (catalog of North American species).

  • — Cameron, 1921a: 354, 404 (characters; key to Singapore species; catalog).

  • — Cameron, 1925: 49 (catalog of Indian species).

  • — Scheerpeltz, 1933: 1299 (world catalog supplement).

  • — Blackwelder, 1943: 504 (characters; type species: floridanus; key to species of West Indies).

  • — Blackwelder, 1944: 145 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Steel, 1949a: 268 (identity of type species uncertain so Oligolinus should be restricted to its type species; without explicitly listing them stated that all South American species of Oligolinus should be transferred to Neoxantholinus).

  • — Blackwelder, 1952: 270 (type species: floridanus).

  • — Scheerpeltz, 1960c: 92 (key to species).

  • — Moore, 1963a: 94 (characters).

  • — Moore and Legner, 1975: 88 (catalog; North America north of Mexico).

  • — Moore and Legner, 1979: 71 (characters; notes).

  • — Smetana, 1982: 120 (synonym of Lithocharodes).

armatus Sharp, 1885: 491 (Lithocharodes; Type locality: Nicaragua, Granada).

  • — Bernhauer and Schubert, 1914: 296 (Lithocharodes; catalog).

  • Distribution: Nicaragua.

beckeri Scheerpeltz, 1953, see: Neoxantholinus.

bidentatus Smetana, 1982: 122 (Lithocharodes; Type locality: Texas: Gonzales Co., 1 mi. NE Palmetto St. Pk. Also cited from Arizona).

  • Distribution: USA.

cameroni Blackwelder, 1943: 497 (Lithocharodes; Type locality: Cuba: Cayamas, 15 miles west-northwest of Cienfuegos, province of Santa Clara).

  • Distribution: Cuba.

cavicola Blackwelder, 1943: 499 (Lithocharodes; Type locality: Trinidad: a cave in Aripo Valley, elevation about 2600 feet).

  • Distribution: West Indies.

claviscapus Cameron, 1922: 114 (Somoleptus; Type locality: Grenada, Mount Gay).

  • — Scheerpeltz, 1933: 1304 (Somoleptus; catalog).

  • — Blackwelder, 1943: 497 (Lithocharodes; characters; Grenada; St. Vincent).

  • Distribution: West Indies.

collaris Sharp, 1885: 494 (Lithocharodes; Type locality: Guatemala, Cubilguitz in Vera Paz 1000 feet).

  • — Bernhauer and Schubert, 1914: 296 (Lithocharodes; catalog).

  • Distribution: Guatemala.

darlingtoni Blackwelder, 1943, see: Neoxantholinus.

densus Smetana, 1982: 129 (Lithocharodes; Type locality: Arizona: S. Rita Mts.).

  • Distribution: USA.

discalis Cameron, 1928f: 410 (Metolinus; Type locality: Mt. Penrissen, 4000 feet).

  • — Cameron, 1930b: 162 (Oligolinus; British North Borneo).

  • — Scheerpeltz, 1933: 1299 (Metolinus; catalog).

  • — Hammond, 1984: 206 (Oligolinus; Borneo).

  • Distribution: Borneo.

excellens Bernhauer, 1915, see: Metolinus.

fijiensis Cameron, 1945e: 49 (Oligolinus; Type locality: Fiji).

  • Distribution: Fiji.

flohri Sharp, 1885: 492 (Lithocharodes; Type locality: Mexico, Uruapan).

  • — Bernhauer and Schubert, 1914: 296 (Lithocharodes; catalog).

  • Distribution: Mexico.

floridanus LeConte, 1880: 170 (Metoponcus, cited as Metaponcus; Type locality: Enterprize, Florida).

  • — Casey, 1906: 420 (Oligolinus; characters; Florida).

  • — Bernhauer and Schubert, 1914: 291 (Oligolinus; catalog).

  • — Steel, 1949a: 268 (Oligolinus should be restricted to include only floridanus; identity uncertain; may be congeneric with varians and therefore would be transferred to Neoxantholinus).

  • — Scheerpeltz, 1953: 605 (Oligolinus; characters in key; Florida; Cuba; Mexico).

  • — Scheerpeltz, 1960c: 93 (Oligolinus; characters; Florida; Mexico; South America).

  • — Smetana, 1982: 130 (Lithocharodes; characters; Florida).

  • Distribution: USA, Mexico, West Indies, “South America”.

foresteri Scheerpeltz, 1960, see: Neoxantholinus.

fuscipennis Sharp, 1876: 205 (Lithocharodes; Type locality: Tapajos).

  • — Lynch, 1884: 194 (Lithocharodes; characters; Argentina).

  • — Sharp, 1885: 492 (Lithocharodes; Guatemala; Brazil).

  • — Bernhauer and Schubert, 1914: 296 (Lithocharodes; catalog).

  • Distribution: Brazil, Argentina, Guatemala.

fusciventris Sharp, 1885: 492 (Lithocharodes; Type locality: Guatemala, San Geronimo).

  • — Bernhauer and Schubert, 1914: 296 (Lithocharodes; catalog).

  • Distribution: Guatemala.

fusculus Sharp, 1885: 494 (Lithocharodes; Type locality: Guatemala, near the city, San Geronimo).

  • — Bernhauer and Schubert, 1914: 296 (Lithocharodes; catalog).

  • — Marcuzzi, 1977: 10 (Lithocharodes; Venezuela).

  • Distribution: Guatemala, Venezuela.

germanus Sharp, 1885: 492 (Lithocharodes; Type locality: Guatemala, San Geronimo).

  • — Bernhauer and Schubert, 1914: 296 (Lithocharodes; catalog).

  • Distribution: Guatemala.

gracilentus Scheerpeltz, 1953, see: Neoxantholinus.

gracilis Sharp, 1885: 493 (Lithocharodes; Type locality: Nicaragua, Chinandega; Panama, Tolé, Peña Blance 3000 to 4000 feet, Bugaba).

  • — Bernhauer and Schubert, 1914: 296 (Lithocharodes; catalog).

  • Distribution: Nicaragua, Panama.

grandis Bernhauer, 1919: 364 (Oligolinus; Type locality: Luzon: Mt. Makiling).

  • — Scheerpeltz, 1933: 1299 (Oligolinus; catalog).

  • Distribution: Philippines.

hubbardi Blackwelder, 1943, see: Neoxantholinus.

longicollis LeConte, 1863a: 41 (Leptacinus; Type locality: Middle States; [Note: See lectotype designation by Smetana, 1982]).

  • — Casey, 1906: 409 (Nematolinus; characters; Rhode Island and Pennsylvania to Iowa and Missouri).

  • — Blatchley, 1910: 399 (Xantholinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 297 (Nematolinus; catalog).

  • — Smetana, 1982: 132 (Lithocharodes; lectotype designation; characters; Ontario; Quebec; Connecticut; Illinois; Indiana; Iowa; Kentucky; Louisiana; Massachusetts; Michigan; Minnesota; Missouri; Nebraska; New Hampshire; New Jersey; New York; North Carolina; Ohio; Pennsylvania; Rhode Island; Virginia; West Virginia).

  • — Smetana, 1988a: 530 (Lithocharodes; collecting notes; Quebec; Maryland; New Hampshire).

  • — Smetana, 1990c: 85 (Lithocharodes; Arkansas).

  • — Campbell and Davies, 1991: 123 (Lithocharodes; checklist of species; Canada).

  • — Downie and Arnett, 1996: 400 (Lithocharodes; characters; USA).

  • Distribution: Canada, USA.

nigripennis LeConte, 1863a: 42 (Leptolinus; Type locality: Louisiana).

  • — Casey, 1906: 411 (Lithocharodes; characters; Louisiana).

  • — Bernhauer and Schubert, 1914: 296 (Lithocharodes; catalog).

  • — Notman, 1920b: 704 (Lithocharodes; Florida).

  • — Smetana, 1982: 126 (Lithocharodes; characters; Florida; Louisiana; South Carolina; Texas).

  • — Smetana, 1988a: 529 (Lithocharodes; collecting notes; Louisiana; Mississippi).

  • — Smetana, 1990c: 85 (Lithocharodes; Arkansas; Oklahoma).

  • Distribution: USA.

pallidus Casey, 1906: 410 (Lithocharodes; Type locality: Texas, Austin; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 296 (Lithocharodes; catalog).

  • — Smetana, 1982: 126 (Lithocharodes; lectotype designation; synonym of nigripennis).

pallidus Casey, 1906, see: nigripennis LeConte, 1863.

parvus Cameron, 1918: 81 (Oligolinus; Type locality: Singapore: Keppel Harbour; Woodlands).

  • — Cameron, 1921a: 375, 404 (Oligolinus; characters; Singapore).

  • — Scheerpeltz, 1933: 1299 (Oligolinus; catalog).

  • Distribution: Singapore.

puncticeps Sharp, 1885: 493 (Lithocharodes; Type locality: Mexico, Cordova, Jalapa).

  • — Bernhauer and Schubert, 1914: 296 (Lithocharodes; catalog).

  • Distribution: Mexico.

rambouseki Blackwelder, 1943: 500 (Lithocharodes; Type locality: Cuba: Rio Almendares, province of Habana).

  • Distribution: West Indies.

ruficollis LeConte, 1863a: 42 (Leptolinus; Type locality: Louisiana; [Note: See lectotype designation by Smetana, 1982]).

  • — Casey, 1906: 411 (Lithocharodes; characters; Texas; Louisiana).

  • — Blatchley, 1910: 399 (Xantholinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 296 (Lithocharodes; catalog).

  • — Smetana, 1982: 124 (Lithocharodes; lectotype designation; characters; Alabama; Arkansas; California; Georgia; Louisiana; Mississippi; Tennessee; Texas).

  • — Smetana, 1990c: 85 (Lithocharodes; Oklahoma).

  • Distribution: USA.

rufulus Sharp, 1885: 494 (Lithocharodes; Type locality: Guatemala, near the city, Pantaleon, Rio Naranjo).

  • — Bernhauer and Schubert, 1914: 296 (Lithocharodes; catalog).

  • Distribution: Guatemala.

scutifer Smetana, 1988a: 527 (Lithocharodes; Type locality: Arizona: Pima Co.: Tucson).

  • Distribution: USA.

simillimus Sharp, 1885: 491 (Lithocharodes; Type locality: Guatemala, Cahabon).

  • — Bernhauer and Schubert, 1914: 296 (Lithocharodes; catalog).

  • Distribution: Guatemala.

sjostedti Bernhauer, 1928: 15 (Oligolinus; cited as sjöstedti; Type locality: Sumatra: Brastagi).

  • — Scheerpeltz, 1933: 1299 (Oligolinus; cited as sjoestedti; catalog).

  • — Cameron, 1933d: 383 (Metolinus; cited as sjöstedti; Sumatra).

  • — Herman, 2001: 12 (Lithocharodes; spelling corrected to sjostedti).

  • Distribution: Indonesia.

sordidus Sharp, 1885: 493 (Lithocharodes; Type locality: Panama, Bugaba 800 to 1500 feet).

  • — Bernhauer and Schubert, 1914: 296 (Lithocharodes; catalog; Nicaragua).

  • Distribution: Nicaragua, Panama.

spiniger Sharp, 1885: 491 (Lithocharodes; Type locality: Guatemala, near the city).

  • — Bernhauer and Schubert, 1914: 296 (Lithocharodes; catalog).

  • Distribution: Guatemala.

strigulatus Blackwelder, 1943: 498 (Lithocharodes; Type locality: Dominican Republic: Loma Rucilla and the mountains to the north, elevation 5000 to 8000 feet).

  • Distribution: West Indies.

trinitatis Blackwelder, 1943: 500 (Lithocharodes; Type locality: Trinidad: St. Augustine).

  • Distribution: West Indies.

unicolor Cameron, 1922: 115 (Somoleptus; Type locality: Jamaica).

  • — Scheerpeltz, 1933: 1305 (Somoleptus; catalog).

  • — Blackwelder, 1943: 501 (Lithocharodes; characters; Jamaica).

  • Distribution: West Indies.

variabilis Blackwelder, 1943, see: Neoxantholinus.

Manilla [4 species; Oriental Region]

Manilla Bordoni, 1990: 172 (species included: orousseti, bipatea, emussitata, deharvengi). Type species: Manilla orousseti Bordoni, fixed by original designation.

bipatea Bordoni, 1990: 176 (Manilla; Type locality: Filippine: Luzon, Baguio, Mte. Santo Thomas).

  • Distribution: Philippines.

deharvengi Bordoni, 1990: 179 (Manilla; Type locality: Filippine: Luzon, Baguio, 1000 m).

  • Distribution: Philippines.

emussitata Bordoni, 1990: 176 (Manilla; Type locality: Filippine: Luzon, Mountain Province, Sagada).

  • Distribution: Philippines.

orousseti Bordoni, 1990: 174 (Manilla; Type locality: Filippine: Luzon, Mount Data, 2200–2300 m).

  • Distribution: Philippines.

Megalinus [9 species; Palaearctic Region]

Megalinus Mulsant and Rey, 1877a: 261 (species included: glabratus). Type species: Staphylinus glabratus Gravenhorst, fixed by monotypy.

  • — Lynch, 1884: 182 (characters; subgenus of Xantholinus).

  • — Bernhauer and Schubert, 1914: 300 (subgenus of Xantholinus).

  • — Cameron, 1932: 22 (subgenus of Xantholinus).

  • — Blackwelder, 1943: 473 (subgenus of Xantholinus; type species: glabratus).

  • — Tottenham, 1949: 369, 417 (type species: glabratus Gravenhorst; subgenus of Xantholinus; checklist of British species).

  • — Blackwelder, 1952: 234 (type species: glabratus).

  • — Coiffait, 1956a: 58, 69 (subgenus of Xantholinus; characters in key; key to species).

  • — Hatch, 1957: 233, 234 (characters; key to species of Pacific Northwest).

  • — Smetana, 1958: 99 (synonym of Xantholinus).

  • — Coiffait, 1962b: 394, 395 (subgenus of Xantholinus; characters).

  • — Moore, 1963a: 94 (characters).

  • — Lohse, 1964: 163 (subgenus of Xantholinus; characters; key to central European species).

  • — Coiffait, 1972: 209 (characters; type species: glabratus; valid genus; key to species of western Palaearctic region; references).

  • — Bordoni, 1972: 163 (subgenus of Xantholinus; characters).

  • — Shibata, 1973c: 55 (checklist of species of Taiwan).

  • — Bordoni, 1973j: 697 (list of species from islands near Sicily).

  • — Bordoni, 1975: 89 (subgenus of Xantholinus; distributional notes).

  • — Moore and Legner, 1975: 83 (catalog; North America north of Mexico).

  • — Moore and Legner, 1979: 65 (characters; notes).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 15 (subgenus of Xantholinus; catalog; Poland).

  • — Bordoni, 1982: 200 (characters; habitus; key to species of Italy).

  • — Silfverberg, 1992: 18 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Ciceroni and Zanetti, 1995: 25 (list of species of Italy).

  • — M. Hansen, 1996: 109 (list of Danish species).

Metacyclinus Reitter, 1908: 115 [= 1908a: 18] (species included: glabratus, relucens, glaber, scutellaris, baicalensis, laevissimus, sublaevis). Type species: Staphylinus glabratus Gravenhorst, fixed by subsequent designation by Blackwelder, 1943: 473.

  • — Reitter, 1909: 137 (subgenus of Xantholinus; key to species).

  • — Bernhauer and Schubert, 1914: 300 (subgenus of Xantholinus).

  • — Cameron, 1932: 22 (subgenus of Xantholinus).

  • — Scheerpeltz, 1933: 1307 (subgenus of Xantholinus).

  • — Blackwelder, 1943: 473 (synonym of Megalinus; type species: glabratus).

  • — Tottenham, 1949c: 41 (type species: relucens).

  • — Tottenham, 1949: 369, 416 (type species: relucens Gravenhorst; subgenus of Xantholinus; checklist of British species).

  • — Blackwelder, 1952: 239 (type species).

  • — Smetana, 1958: 99 (synonym of Xantholinus).

  • — Coiffait, 1972: 209 (synonym of Nudobius).

  • — Bordoni, 1982: 200 (synonym of Megalinus).

cadaverinus Lacordaire, 1835, see: glabratus Gravenhorst, 1802.

cruentatus Marsham, 1802, see: glabratus Gravenhorst, 1802.

davisi Hatch, 1957, see: Oxybleptes.

ferrugineus Rossi, 1790, see: glabratus Gravenhorst, 1802.

formosae Cameron, 1949: 175 (Xantholinus; Type locality: Formosa: Funkiko: near Mt. Ari).

  • — Shibata, 1973a: 126 (Megalinus; Taiwan).

  • Distribution: Taiwan.

glabratus Gravenhorst, 1802: 178 (Staphylinus; Type locality: Lusitania).

  • — Panzer, 1796c: 8 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 303, cited nitidus Panzer as a synonym of glabratus Gravenhorst; Panzer did not describe the species as new, he attributed it to Fabricius]; characters; illustration).

  • — Gravenhorst, 1802: 48 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 303, listed fulgidus Gravenhorst as a synonym of glabratus; Gravenhorst did not describe a new species he attributed it Fabricius]; characters).

  • — Gravenhorst, 1806: 101 (Staphylinus).

  • — Nordmann, 1837: 113 (Gyrohypnus; characters; Portugal).

  • — Nordmann, 1837: 115 (Gyrohypnus; [Note: Misidentification: Bernhauer and Schubert, 1914: 303, cited relucens Nordmann as a synonym of glabratus Gravenhorst; Nordmann did not describe the species as new, he attributed it to Gravenhorst]; characters; France; Germany).

  • — Erichson, 1839a: 424 (Xantholinus; characters; Germany).

  • — Heer, 1839: 244 (Xantholinus; characters; Switzerland).

  • — Erichson, 1839b: 319 (Xantholinus; characters; Europe).

  • — Stephens, 1839: 402 (Xantholinus; characters; England).

  • — P. Lucas, 1846: 106 (Xantholinus; notes; Algeria).

  • — Küster, 1848: 9 (Xantholinus; characters; Europe).

  • — Jacobsen, 1849: 676 (Xantholinus; characters; notes; Denmark).

  • — Motschulsky, 1849: 87 (Xantholinus; Spain).

  • — Redtenbacher, 1849: 691 (Xantholinus; characters; Austria).

  • — Hardy, 1851: 37 (Xantholinus; notes; England).

  • — Rosenhauer, 1856: 70 (Xantholinus; Cadiz; Malaga).

  • — Fairmaire and Laboulbène, 1856: 500 (Xantholinus; characters; France).

  • — Kraatz, 1857d: 633 (Xantholinus; characters; Germany).

  • — Redtenbacher, 1857: 181 (Xantholinus; characters; Austria).

  • — Hochhuth, 1862: 65 (Xantholinus; Russia).

  • — Thomson, 1867: 177 (Xantholinus; characters; Scandinavia).

  • — Crotch, 1867a: 383 (Xantholinus; Azores).

  • — Fauvel, 1873b: 101 [= 1873a: 387] (Xantholinus; characters; France; Europe; Barbary).

  • — Redtenbacher, 1874: 218 (Philonthus; characters; Austria).

  • — Seidlitz, 1875: 281 (Xantholinus; characters; Baltic region).

  • — Mulsant and Rey, 1877a: 262 (Xantholinus; subgenus Megalinus; characters; France).

  • — Fauvel, 1878: 117 [= 1878a: 37] (Xantholinus; Morocco; Algeria).

  • — Fauvel, 1886: 49 [= 1886a: 41] (Xantholinus; North Africa).

  • — Fowler, 1888: 288 (Xantholinus; characters; Britain).

  • — Ganglbauer, 1895: 481 (Xantholinus; characters; Europe; North Africa).

  • — Fauvel, 1902b: 101 (Xantholinus; Morocco; Algeria; Tunisia; Azores).

  • — Poppius, 1905: 10 (Xantholinus; Orkney Islands).

  • — Sainte-Claire Deville, 1906: 92 (Xantholinus; Corsica).

  • — Reitter, 1908: 116 (Xantholinus; subgenus Metacyclinus; characters; Europe; Sudan).

  • — Reitter, 1909: 137 (Xantholinus; subgenus Metacyclinus; characters; Germany).

  • — Sharp and Muir, 1912: 499 (Xantholinus; characters of male genitalia).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; catalog).

  • — Johansen, 1914: 435 (Xantholinus; subgenus Xantholinus; characters; Denmark).

  • — Dodero, 1925: 5 (Xantholinus; Libya).

  • — Portevin, 1929: 372 (Xantholinus; characters; France).

  • — Gridelli, 1930: 70 (Xantholinus; Libya).

  • — Scheerpeltz, 1933: 1310 (Xantholinus; catalog).

  • — Bernhauer, 1940c: 3 (Xantholinus; Azores).

  • — Méquignon, 1942a: 19 (Xantholinus; Azores).

  • — Tottenham, 1949: 417 (Xantholinus; checklist; Britain).

  • — Coiffait, 1956a: 70 (Xantholinus; subgenus Megalinus; characters in key).

  • — Smetana, 1958: 102 (Xantholinus; subgenus Megalinus; characters; ecology; Czechoslovakia).

  • — Kocher, 1958: 139 (Xantholinus; Morocco).

  • — Ferreira, 1962: 16 (Xantholinus; Portugal).

  • — Coiffait, 1962b: 398, 404 (Xantholinus; subgenus Megalinus; characters; Morocco; Algeria; Tunisia).

  • — Palm, 1963b: 50 (Xantholinus; characters; Sweden; Denmark).

  • — Lohse, 1964: 163 (Xantholinus; subgenus Megalinus; characters; central Europe).

  • — Focarile, 1964: 64 (Xantholinus; Italy).

  • — Scheerpeltz, 1965b: 400 (Xantholinus; Sicily).

  • — Horion, 1965: 95 (Xantholinus; subgenus Megalinus; Morocco; Tunisia; Iberian Peninsula; Corsica; Sardinia; Sicily; Italy; France; Belgium; Holland; Switzerland; Czech Republic; Slovakia; Britain; Denmark; Sweden; Germany; Austria).

  • — Scheerpeltz, 1968a: 62 (Xantholinus; catalog; Austria).

  • — Kasule, 1970: 79 (Xantholinus; larval characters).

  • — Coiffait, 1972: 211 (Megalinus; characters; references; Europe; Britain; Scandinavia; Corsica; North Africa).

  • — Bordoni, 1972: 163, 210 (Xantholinus; subgenus Megalinus; characters; Italy).

  • — Bordoni, 1975: 78 (Xantholinus; subgenus Megalinus; Switzerland).

  • — Bordoni, 1976c: 98 (Megalinus; notes; France; Italy; Portugal; Spain; Majorca; Morocco; Algeria; Tunisia).

  • — Bordoni, 1976d: 122 (Megalinus; Algeria; Netherlands; Austria; Italy).

  • — Pope, 1977: 28 (Xantholinus; Britain).

  • — Muona, 1979: 16 (Xantholinus; Sweden; Denmark).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 16 (Xantholinus; catalog; Poland).

  • — Hinton, 1981: 663 (Xantholinus; characters of egg).

  • — Bordoni, 1982: 200 (Megalinus; characters; Italy).

  • — Bordoni, 1985a: 87 (Megalinus; anatomical studies of labrum, aedeagus, and male genital segment).

  • — Bordoni, 1986b: 386 (Megalinus; Portugal; Tunisia).

  • — Segers, 1986: 40 (Megalinus; checklist of species; Belgium).

  • — Lucht, 1987: 103 (Xantholinus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Tóth, 1989: 21 (Xantholinus; characters; Hungary).

  • — Lohse and Lucht, 1989: 159 (Xantholinus; subgenus Megalinus).

  • — Silfverberg, 1992: 18 (Megalinus; Sweden; Denmark; Lithuania).

  • — Welch, 1993: 228 (Xantholinus; ovariole number and ovary structure).

  • — Ciceroni, 1993: 71 (Megalinus; Italy).

  • — Janák, 1993a: 47 (Megalinus; Czech Republic; Slovakia).

  • — Hernández, Outerelo, and Gamarra, 1994: 190 (Megalinus; checklist of species; Canary Islands).

  • — Sparacio, 1995: 156 (Megalinus; characters; summary of distribution; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 25 (Megalinus; Italy).

  • — M. Hansen, 1996: 109 (Megalinus; checklist; Denmark).

  • — Sterrenburg and Schülke, 1997: 18 (Xantholinus; Netherlands).

  • — Anderson, Nash, and O'Connor, 1997: 20 (Xantholinus; Ireland).

  • — Owen, 1999b: 61 (Xantholinus; captured underground at roots of old trees; Britain).

  • — Assing and Schülke, 1999: 8 (Megalinus; notes).

  • — Owen, 2000: 251 (Xantholinus; pitfall trap collections from underground at roots of trees; Britain).

  • — Herman, 2001: 26 (Megalinus; nomen protectum).

  • Distribution: Sudan, Libya, Tunisia, Algeria, Morocco, Azores, Europe, Russia, Turkey, Syria, Lebanon.

ferrugineus Rossi, 1790: 248 (Staphylinus; [Note: This name is older than glabratus]; Type locality: Florentiae).

  • — Gyllenhal, 1810: 315 (Staphylinus; synonym of nitidus Fabricius).

  • — C. Sahlberg, 1830: 320 (Staphylinus; synonym of nitidus Fabricius).

  • — Erichson, 1840: 924 (Xantholinus; synonym of glabratus).

  • — Fauvel, 1869: 490 (Xantholinus; synonym of glabratus).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; variety of glabratus).

  • — Herman, 2001: 26 (Megalinus; nomen oblitum).

cruentatus Marsham, 1802: 516 (Staphylinus; [preoccupied]; Type locality: Britain).

  • — Stephens, 1833: 260 (Gyrohypnus; characters; England).

  • — Erichson, 1839b: 319 (Xantholinus; synonym of glabratus).

  • — Küster, 1848: 9 (Xantholinus; synonym of glabratus).

  • — Hardy, 1851: 37 (Xantholinus; synonym of glabratus).

  • — Kraatz, 1857d: 633 (Xantholinus; synonym of glabratus).

  • — G. Waterhouse, 1858: 26 (Xantholinus; synonym of glabratus).

  • — Mulsant and Rey, 1877a: 263 (Xantholinus; synonym of glabratus).

  • — Ganglbauer, 1895: 481 (Xantholinus; synonym of glabratus).

  • — Reitter, 1908: 116 (Xantholinus; synonym of glabratus).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; synonym of glabratus).

  • — Smetana, 1958: 103 (Xantholinus; synonym of glabratus).

  • — Coiffait, 1972: 211 (Nudobius; synonym of glabratus).

  • — Bordoni, 1972: 210 (Xantholinus; synonym of glabratus).

  • — Bordoni, 1982: 202 (Megalinus; synonym of glabratus).

relucens Gravenhorst, 1806: 101 (Staphylinus; Type locality: Parisiae).

  • — Erichson, 1839b: 320 (Xantholinus; synonym of glabratus).

  • — Küster, 1848: 9 (Xantholinus; synonym of glabratus).

  • — Fairmaire and Laboulbène, 1856: 501 (Xantholinus; synonym of glabratus).

  • — Kraatz, 1858a: 63 (Xantholinus; notes; Greece).

  • — Hochhuth, 1862: 65 (Xantholinus; Russia).

  • — Baudi, 1870: 387 (Xantholinus; Syria).

  • — Fauvel, 1873b: 101 [= 1873a: 387] (Xantholinus; characters; Germany; Austria; Tirol; Greece; Crete; Cyprus; Turkey; Rhodes; Syria).

  • — Redtenbacher, 1874: 218 (Philonthus; characters; Austria).

  • — Fauvel, 1876a: 70 [= 1876: 255] (Xantholinus; Turkey).

  • — J. Sahlberg, 1876: 34 (Xantholinus; characters; Finland).

  • — Mulsant and Rey, 1877a: 266 (Xantholinus; characters; France).

  • — Ganglbauer, 1895: 481 (Xantholinus; characters; Germany; Austria; central Europe; eastern Mediterranean region).

  • — Fauvel, 1902b: 101 (Xantholinus; Egypt).

  • — Reitter, 1908: 116 (Xantholinus; subgenus Metacyclinus; characters; central and southern Europe; Mediterranean region).

  • — Montandon, 1908: 79 (Xantholinus; Romania).

  • — Reitter, 1909: 137 (Xantholinus; subgenus Metacyclinus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; catalog).

  • — Portevin, 1929: 372 (Xantholinus; characters; France).

  • — Scheerpeltz, 1933: 1314 (Xantholinus; catalog).

  • — Coiffait, 1956a: 59 (Xantholinus; subgenus Leptophallus; characters in key; Lebanon; Turkey; Croatia; Pyrenees).

  • — Scheerpeltz, 1956c: 1099 (Xantholinus; subgenus Leptophallus; Cyprus).

  • — Smetana, 1958: 103 (Xantholinus; subgenus Megalinus; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1961e: 136 (Xantholinus; subgenus Leptophallus; Rhodes).

  • — Lohse, 1964: 163 (Xantholinus; subgenus Leptophallus; characters; central Europe).

  • — Scheerpeltz, 1964a: 299 (Xantholinus; subgenus Leptophallus; Crete).

  • — Horion, 1965: 94 (Xantholinus; subgenus Leptophallus; Turkey; Lebanon; Cyprus; Greece; Crete; Bosnia Hercegovina; Serbia; Croatia; Slovenia; Italy; Romania; Hungary; Slovakia; Czech Republic; Poland; Ukraine; Switzerland; France; Austria; Germany).

  • — Coiffait, 1972: 211 (Nudobius; synonym of glabratus).

  • — Bordoni, 1972: 210 (Xantholinus; synonym of glabratus).

  • — Shilov, 1975a: 375 [= 1975b: 97] (Xantholinus; Komi region of Russia).

  • — Bordoni, 1982: 202 (Megalinus; synonym of glabratus).

  • — Lucht, 1987: 103 (Xantholinus; checklist; Germany; Poland; Czechoslovakia; Austria).

  • — Lohse and Lucht, 1989: 159 (Xantholinus; synonym of flavocinctus).

cadaverinus Lacordaire, 1835: 414 (Xantholinus; Type locality: Paris).

  • — Erichson, 1839b: 320 (Xantholinus; synonym of glabratus).

  • — Küster, 1848: 9 (Xantholinus; synonym of glabratus).

  • — Fairmaire and Laboulbène, 1856: 501 (Xantholinus; synonym of glabratus).

  • — Kraatz, 1857d: 634 (Xantholinus; synonym of relucens).

  • — Fauvel, 1873b: 101 [= 1873a: 387] (Xantholinus; synonym of glabratus).

  • — Mulsant and Rey, 1877a: 263 (Xantholinus; variety of glabratus).

  • — Ganglbauer, 1895: 481 (Xantholinus; synonym of relucens).

  • — Reitter, 1908: 116 (Xantholinus; synonym of relucens).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; synonym of relucens).

  • — Smetana, 1958: 105 (Xantholinus; synonym of relucens).

  • — Coiffait, 1972: 211 (Nudobius; synonym of glabratus).

  • — Bordoni, 1972: 164 (Xantholinus; synonym of flavocinctus).

  • — Bordoni, 1982: 202 (Megalinus; synonym of glabratus).

merdarius Nordmann, 1837: 116 (Gyrohypnus; Type locality: Italia prope Pisam).

  • — Küster, 1848: 9 (Xantholinus; synonym of glabratus).

  • — Fairmaire and Laboulbène, 1856: 501 (Xantholinus; synonym of glabratus).

  • — Kraatz, 1857d: 633 (Xantholinus; synonym of glabratus).

  • — Fauvel, 1869: 490 (Xantholinus; synonym of glabratus).

  • — Mulsant and Rey, 1877a: 263 (Xantholinus; variety of glabratus).

  • — Ganglbauer, 1895: 481 (Xantholinus; synonym of glabratus).

  • — Reitter, 1908: 116 (Xantholinus; variety of glabratus).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; synonym of glabratus).

  • — Scheerpeltz, 1933: 1310 (Xantholinus; synonym of glabratus).

  • — Smetana, 1958: 103 (Xantholinus; variety of glabratus).

  • — Coiffait, 1972: 211 (Nudobius; variety of glabratus).

  • — Bordoni, 1982: 202 (Megalinus; synonym of glabratus).

occidentalis Waltl, 1835: 58 (Xantholinus; Type locality: Andalusien).

  • — Erichson, 1840: 924 (Xantholinus; synonym of glabratus).

  • — Rosenhauer, 1856: 70 (Xantholinus; Cadiz; Malaga).

  • — Ganglbauer, 1895: 481 (Xantholinus; synonym of glabratus).

  • — Reitter, 1908: 116 (Xantholinus; synonym of glabratus).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; synonym of glabratus).

  • — Coiffait, 1972: 211 (Nudobius; synonym of glabratus).

  • — Bordoni, 1972: 210 (Xantholinus; synonym of glabratus).

  • — Bordoni, 1982: 202 (Megalinus; synonym of glabratus).

keijiroi Watanabe and Shibata, 1965: 319 (Megalinus; subgenus Megalinus; Type locality: Japan: Oshidomari, Rishiri Is., near Hokkaido).

  • Distribution: Japan.

laevissimus Reitter, 1898: 116 (Xantholinus; Type locality: Talysch-Gebirge; [Note: See lectotype designation by Bordoni, 1999]).

  • — Reitter, 1908: 117 (Xantholinus; subgenus Metacyclinus; characters; Caspian).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1311 (Xantholinus; catalog).

  • — Coiffait, 1972: 217 (Phalacrolinus; characters; references; Azerbaijan).

  • — Bordoni, 1975: 75 (Xantholinus; subgenus Megalinus; characters; notes; Caspian region).

  • — Bordoni, 1999: 7 (Megalinus; lectotype designation).

  • Distribution: Azerbaijan, Iran.

tenuipunctus Fauvel, 1900d: 229 (Xantholinus; Type locality: Monts Talysch).

  • — Reitter, 1908: 117 (Xantholinus; synonym of laevissimus).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1311 (Xantholinus; synonym of laevissimus).

  • — Coiffait, 1972: 217 (Phalacrolinus; synonym of laevissimus).

merdarius Nordmann, 1837, see: glabratus Gravenhorst, 1802.

nudobiiformis Tikhomirova, 1973: 162 (Xantholinus; subgenus Megalinus; Type locality: pos. Khasan).

  • — Herman, 2001: 83 (Megalinus).

  • Distribution: Russia.

oasis Steel, 1948a: 186 (Xantholinus; subgenus Megalinus; Type locality: Cyrenaica: north of Benghazi-Merg).

  • — Coiffait, 1956a: 70 (Xantholinus; subgenus Megalinus; characters in key).

  • — Bordoni, 1972: 163 (Xantholinus; subgenus Megalinus).

  • — Coiffait, 1972: 213 (Megalinus; characters; references; Cyrenaica).

  • Distribution: Libya.

occidentalis Waltl, 1835, see: glabratus Gravenhorst, 1802.

relucens Gravenhorst, 1806, see: glabratus Gravenhorst, 1806.

sabellai Ciceroni and Zanetti, 1992: 337 (Megalinus; Type locality: Sicilia: Caronia, Messina, 1330 m).

  • — Sparacio, 1995: 157 (Megalinus; characters; summary of distribution; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 25 (Megalinus; Italy).

  • Distribution: Italy.

scutellaris Fauvel, 1900d: 228 (Xantholinus; Type locality: Turquie).

  • — Reitter, 1908: 116 (Xantholinus; subgenus Metacyclinus; characters; Turkey).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1314 (Xantholinus; catalog).

  • — Bordoni, 1971: 682 (Xantholinus; subgenus Heterolinus ?; characters; Turkey).

  • — Coiffait, 1972: 213 (Megalinus; characters; references; Turkey).

  • Distribution: Turkey.

tenuipunctus Fauvel, 1900, see: laevissimus Reitter, 1898.

ussuricus Tikhomirova, 1973: 163 (Xantholinus; subgenus Megalinus; Type locality: Suputinskii zapovednik).

  • — Herman, 2001: 83 (Megalinus).

  • Distribution: Russia.

Metocinus [2 species; Ethiopian Region]

Metocinus Cameron, 1950g: 32 (species included: aethiops). Type species: Metocinus aethiops Cameron, fixed by original designation and monotypy.

aethiops Cameron, 1950g: 32 (Metocinus; Type locality: Belgian Congo: Kanyabayongo, Kabasha, alt. 1760 m).

  • Distribution: Congo.

reductus Tottenham, 1956: 238 (Metocinus; Type locality: Urundi: terr. Bururi, Usumbura, 800 m. Also cited from Ruanda).

  • Distribution: Burundi, Rwanda.

Metolinus [14 species; Palaearctic, Ethiopian, Oriental, and Australian Regions]

Metolinus Cameron, 1920a: 147, 220 (species included: leucocnemis, basalis). Type species: Metoponcus leucocnemis Kraatz, fixed by subsequent designation by Blackwelder, 1952: 241.

  • — Cameron, 1925: 49 (catalog of Indian species).

  • — Cameron, 1932: 4 (characters; key to species of British India).

  • — Scheerpeltz, 1933: 1298 (world catalog supplement).

  • — Shibata, 1983: 68 (checklist of species of Japan).

  • — Hammond, 1984: 206 (checklist of species of Borneo).

apicipennis Cameron, 1950d: 15 (Metolinus; Type locality: Selangor: The Gap).

  • Distribution: Malaysia.

basalis Cameron, 1920, see: excellens Bernhauer, 1915.

bhutanicus Coiffait, 1977b: 217 (Metolinus; Type locality: Bhutan: Nobding, 41 km O. Wangdi Phodrang, 2800 m).

  • Distribution: Bhutan.

borneensis Cameron, 1933b: 343 (Metolinus; Type locality: British North Borneo: Pakka, 10,200 feet).

  • Distribution: Borneo.

differens Cameron, 1932: 5 (Metolinus; Type locality: Chittagong).

  • — Scheerpeltz, 1933: 1299 (Metolinus; catalog).

  • Distribution: India.

discalis Cameron, 1928, see: Lithocharodes.

distinctus Cameron, 1936b: 43 (Metolinus; Type locality: Johore: Kluang).

  • Distribution: Malaysia.

excellens Bernhauer, 1915i: 253 (Oligolinus; Type locality: Südindien: Chambaganoor, Madura).

  • — Cameron, 1932: 5 (Metolinus; characters; India).

  • — Scheerpeltz, 1933: 1299 (Metolinus; catalog).

  • Distribution: India.

basalis Cameron, 1920a: 147 (Metolinus; Type locality: Anamalai Hills, alt. 5500 feet).

  • — Cameron, 1932: 5 (Metolinus; synonym of excellens).

  • — Scheerpeltz, 1933: 1299 (Metolinus; catalog).

exiguus Kraatz, 1859, see: leucocnemis Kraatz, 1859.

femineus Coiffait, 1977b: 218 (Metolinus; Type locality: Bhutan: 21 km Ouest de Wangdi Phodrang, 1700 à 2200 m).

  • Distribution: Bhutan.

fluvius Last, 1980: 157 (Metolinus; Type locality: New Guinea: Wau, Kilolo Creek River).

  • Distribution: New Guinea.

javanicus Cameron, 1937: 1 (Metolinus; Type locality: Java: G. Patoeha, Preanger, alt. 7000 ft.).

  • Distribution: Indonesia.

javanus Cameron, 1937: 4 (Othius; Type locality: Java: G. Tangkoeban Prahoe, alt. 1400 m).

  • — Cameron, 1939e: 1 (Metolinus).

  • Distribution: Indonesia.

leucocnemis Kraatz, 1859: 107 (Metoponcus; Type locality: Ceylan).

  • — Fauvel, 1904a: 56 (Metoponcus; Kanara; Ceylan; Tenasserim; Sikkim; Annam).

  • — Bernhauer and Schubert, 1914: 291 (Oligolinus; catalog).

  • — Cameron, 1932: 5 (Metolinus; characters; India; Ceylon; Sikkim; Burma).

  • — Scheerpeltz, 1933: 1299 (Metolinus; catalog).

  • — Cameron, 1933d: 383 (Metolinus; Sumatra).

  • — Paulian, 1941: 208 (Metolinus; larval characters; India).

  • — Bernhauer, 1942c: 360 (Metolinus; cited as bucocnemis; Cameroon).

  • — Nakane and Sawada, 1960: A123 (Metolinus; Yakushima Island).

  • — Watanabe and Shibata, 1972: 63 (Metolinus; Yaku-shima Island).

  • — Coiffait, 1975a: 154 (Metolinus; Nepal).

  • Distribution: Cameroon [?], Sri Lanka, India, Nepal, Myanmar, Vietnam, Indonesia, Japan.

exiguus Kraatz, 1859: 108 (Metoponcus; Type locality: India orientalis).

  • — Fauvel, 1904a: 56 (Metoponcus; synonym of leucocnemis).

  • — Bernhauer and Schubert, 1914: 291 (Oligolinus; synonym of leucocnemis).

  • — Cameron, 1932: 5 (Metolinus; synonym of leucocnemis).

  • — Scheerpeltz, 1933: 1299 (Metolinus; synonym of leucocnemis).

malaisei Scheerpeltz, 1965: 191 (Metolinus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).

  • Distribution: Myanmar.

marginalis Cameron, 1933d: 383 (Metolinus; Type locality: Tandjunggadang, Sumatra's Westkust, alt. 1000 m).

  • Distribution: Indonesia.

sutteri Scheerpeltz, 1957c: 232, 305 (Metolinus; Type locality: Sumba: Lokolengo).

  • Distribution: Indonesia.

Microleptus [3 species; Madagascan and Oriental Regions]

Microleptus Jarrige, 1963a: 122 (species included: vadoni). Type species: Microleptus vadoni Jarrige, fixed by original designation and monotypy.

gomyi Lecoq, 1990: 187 (Microleptus; Type locality: La Réunion: Saint Gilles les Bains).

  • Distribution: Réunion.

jeanneli Jarrige, 1963a: 123 (Microleptus; Type locality: Ceylan: Meratuwa. Also cited from India).

  • Distribution: Sri Lanka, India.

vadoni Jarrige, 1963a: 122 (Microleptus; Type locality: Madagascar: d'Andapa).

  • Distribution: Madagascar.

Microlinus [1 species; Neotropical and Nearctic Regions]

Microlinus Casey, 1906: 372, 419 (species included: pusio). Type species: Leptolinus pusio LeConte, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 292 (world catalog; 1 species).

  • — Leng, 1920: 106 (catalog of North American species).

  • — Blackwelder, 1943: 502 (characters; type species: pusio).

  • — Blackwelder, 1944: 146 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 246 (type species: pusio).

  • — Moore, 1963a: 94 (characters).

  • — Moore and Legner, 1975: 84 (catalog; North America north of Mexico).

  • — Moore and Legner, 1979: 67 (characters; notes).

  • — Smetana, 1982: 112 (characters; habitus; revision of species of North America north of Mexico).

  • — Frank, 1986a: 369 (checklist of species of Florida).

  • — Lundgren, 1998: 50 (in list of species from Florida; records for other States listed for species).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 391 (1 Nearctic species; notes; characters in key).

pusio LeConte, 1880: 171 (Leptolinus; Type locality: Columbia, South Carolina).

  • — Casey, 1906: 419 (Microlinus; characters; South Carolina).

  • — Bernhauer and Schubert, 1914: 292 (Microlinus; catalog).

  • — Blackwelder, 1943: 502 (Microlinus; characters; Jamaica; Trinidad).

  • — Legner and Moore, 1977: 176 (Microlinus; found under bark).

  • — Smetana, 1982: 113 (Microlinus; characters; Florida; South Carolina).

  • — Smetana, 1990c: 85 (Microlinus; Florida).

  • Distribution: USA, West Indies.

Mitomorphus [41 species; Palaearctic, Ethiopian, Oriental, and Australian Regions]

Mitomorphus Kraatz, 1859: 105 (species included: nigroaeneus, indicus). Type species: Mitomorphus indicus Kraatz, fixed by subsequent designation by R. Lucas, 1920: 420.

  • — Gemminger and Harold, 1868: 604 (catalog).

  • — Duvivier, 1883: 157 (catalog).

  • — Bernhauer and Schubert, 1914: 298 (world catalog; 5 species).

  • — Cameron, 1925: 51 (catalog of Indian species).

  • — Cameron, 1932: 16 (characters; key to species of British India).

  • — Scheerpeltz, 1933: 1305 (world catalog supplement).

  • — Blackwelder, 1952: 249 (type species).

  • — Coiffait, 1968: 128, 147 (characters; notes).

  • — Coiffait, 1972: 129 (characters).

  • — Shibata, 1973c: 55 (checklist of species of Taiwan).

  • — Coiffait, 1975a: 162 (key to species of Nepal).

africanus Bernhauer, 1929f: 120 (Mitomorphus; Type locality: Ituri: La Moto, Madyu; Haut-Uele: Moto; Stanleyville a Kilo).

  • — Scheerpeltz, 1933: 1305 (Mitomorphus; catalog).

  • Distribution: Congo.

angolensis Cameron, 1959: 114 (Mitomorphus; Type locality: Angola: Dundo, forêt de La Luachimo).

  • Distribution: Angola.

anthracinus Scheerpeltz, 1976a: 122 (Mitomorphus; Type locality: Nepal: Ting-Sang-La, 3800 m).

  • Distribution: Nepal.

armatus Coiffait, 1981d: 325 (Mitomorphus; Type locality: environs de Shermathang, Helamub, Nepal central).

  • Distribution: Nepal.

asymetricus Coiffait, 1977: 245 (Mitomorphus; Type locality: Vallée Kali-Gandaki, entre Lete et Tuchke, Népal central).

  • Distribution: Nepal.

baloghi Last, 1980: 158 (Mitomorphus; Type locality: New Guinea: Wau, Kilolo Creek River, 900 m).

  • Distribution: New Guinea.

basalis Cameron, 1937: 3 (Mitomorphus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–6000 ft).

  • Distribution: Indonesia.

bispinosus Coiffait, 1975a: 158 (Mitomorphus; Type locality: Kali-Gandaki-Tal, entre Ghasa et Lete, Népal central).

  • Distribution: Nepal.

brachypterus Cameron, 1943: 127 (Mitomorphus; Type locality: Darjeeling: Ghum district).

  • Distribution: India.

brachypterus Coiffait, 1975, see: devillei Coiffait, 1987.

brevipennis Cameron, 1943: 127 (Mitomorphus; Type locality: Darjeeling: Ghum district: Tiger Hill, altitude 8500–10,000 feet).

  • Distribution: India.

caeruleipennis Coiffait, 1982: 52 (Mitomorphus; Type locality: Nepal: Weg von Jiri nach Shivalaya, 1800–2500 m).

  • Distribution: Nepal.

cribriceps Cameron, 1926b: 341 (Mitomorphus; Type locality: Mohan Rau, Siwaliks).

  • — Cameron, 1932: 21 (Mitomorphus; characters; India).

  • — Scheerpeltz, 1933: 1305 (Mitomorphus; catalog).

  • Distribution: India.

cupreolus Fauvel, 1895: 240 (Mitomorphus; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Carin Asciuii Ghécu, 1400–1500 m).

  • — Bernhauer and Schubert, 1914: 298 (Mitomorphus; catalog).

  • — Cameron, 1932: 18 (Mitomorphus; characters; Burma).

  • — Scheerpeltz, 1933: 1305 (Mitomorphus; catalog).

  • Distribution: Myanmar.

devillei Coiffait, 1987: 497 (Mitomorphus; replacement name for brachypterus Coiffait).

  • Distribution: Nepal.

brachypterus Coiffait, 1975a: 160 (Mitomorphus; [preoccupied]; Type locality: Phulchoki près Katmandou).

  • — Coiffait, 1987: 497 (Mitomorphus; synonym of devillei).

diabolicus Cameron, 1926b: 343 (Xantholinus; Type locality: Chakrata District, Kanasar, 7050 feet).

  • — Cameron, 1932: 20 (Mitomorphus; characters; India).

  • — Scheerpeltz, 1933: 1305 (Mitomorphus; catalog).

  • Distribution: India.

drescheri Cameron, 1937: 3 (Mitomorphus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft).

  • Distribution: Indonesia.

dubitans Tottenham, 1956: 245 (Mitomorphus; Type locality: Ruanda: terr. Nyanza, Mahembe, 1400 m).

  • Distribution: Rwanda.

evanescens Tottenham, 1956: 247 (Mitomorphus; Type locality: Ruanda: terr. Nyanza, Gitarama, 1850 m).

  • Distribution: Rwanda.

fauveli Last, 1980: 158 (Mitomorphus; Type locality: New Guinea: Erima, Astrolabe Bay).

  • Distribution: New Guinea.

formosae Bernhauer, 1943: 75 (Mitomorphus; Type locality: Formosa: Akau, Kankau).

  • — Shibata, 1973a: 122 (Mitomorphus; Taiwan).

  • Distribution: Taiwan.

fuscipes Cameron, 1932a: 140 (Mitomorphus; Type locality: Pahang: Cameron Highlands, alt. 4800 feet).

  • Distribution: Malaysia.

gabonicus Coiffait, 1968: 147 (Mitomorphus; Type locality: Gabon: Belinga).

  • Distribution: Gabon.

gladiator Coiffait, 1982a: 242 (Mitomorphus; Type locality: Nepal: Phulchoki, 2600 m).

  • Distribution: Nepal.

indicus Kraatz, 1859: 107 (Mitomorphus; Type locality: India orientalis).

  • — Bernhauer and Schubert, 1914: 298 (Mitomorphus; catalog).

  • — Bernhauer, 1922c: 231 (Mitomorphus; Formosa).

  • — Cameron, 1932: 19 (Mitomorphus; characters; India).

  • — Scheerpeltz, 1933: 1305 (Mitomorphus; catalog).

  • — Shibata, 1973a: 123 (Mitomorphus; Taiwan).

  • Distribution: India orientalis, Taiwan.

javanus Cameron, 1937: 2 (Mitomorphus; Type locality: G. Tangkoeban Prahoe, alt. 4000–5000 ft; G. Pateoha, alt. 7000 ft).

  • Distribution: Indonesia.

loksai Last, 1980: 159 (Mitomorphus; Type locality: New Guinea: Wau).

  • Distribution: New Guinea.

minusculus Coiffait, 1975a: 161 (Mitomorphus; Type locality: Barahbise près Ting-Sang-La, Népal).

  • — Coiffait, 1982: 26 (Mitomorphus; Nepal).

  • Distribution: Nepal.

minutissimus Last, 1980: 159 (Mitomorphus; Type locality: New Guinea: Kiunga).

  • Distribution: New Guinea.

nepalicus Coiffait, 1975a: 160 (Mitomorphus; Type locality: Chemin de Pokhara à Goropani, Népal central).

  • Distribution: Nepal.

nigerrimus Bernhauer, 1915i: 253 (Mitomorphus; Type locality: Südliches Indien: Chambaganoor, Madura).

  • — Cameron, 1932: 18 (Mitomorphus; characters; India).

  • — Scheerpeltz, 1933: 1305 (Mitomorphus; catalog).

  • Distribution: India.

ovaliceps Cameron, 1920a: 147 (Mitomorphus; Type locality: Nilgiri Hills).

  • — Cameron, 1932: 18 (Mitomorphus; synonym of nigerrimus).

  • — Scheerpeltz, 1933: 1305 (Mitomorphus; synonym of nigerrimus).

nigroaeneus Kraatz, 1859: 106 (Mitomorphus; Type locality: Ceylan).

  • — Bernhauer and Schubert, 1914: 298 (Mitomorphus; catalog).

  • — Cameron, 1932: 17 (Mitomorphus; characters; Ceylon).

  • — Scheerpeltz, 1933: 1305 (Mitomorphus; catalog).

  • Distribution: Ceylon.

obsoletus Fauvel, 1904a: 56 (Mitomorphus; Type locality: Nilghiris).

  • — Bernhauer and Schubert, 1914: 298 (Mitomorphus; catalog).

  • — Cameron, 1932: 19 (Mitomorphus; characters; India).

  • — Scheerpeltz, 1933: 1305 (Mitomorphus; catalog).

  • Distribution: India.

osellai Coiffait, 1981f: 332 (Mitomorphus; Type locality: South Andaman, Port Blair, Mont Harriet).

  • Distribution: Andaman Islands.

ovaliceps Cameron, 1920, see: nigerrimus Bernhauer, 1915.

pubiventris Cameron, 1937: 3 (Mitomorphus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft).

  • Distribution: Indonesia.

punctulatus Last, 1980: 159 (Mitomorphus; Type locality: New Guinea: Kilolo Creek).

  • Distribution: New Guinea.

purpurescens Cameron, 1926b: 341 (Mitomorphus; Type locality: Mussorie District; Arni Gad and Kaligad).

  • — Cameron, 1932: 20 (Mitomorphus; characters; India).

  • — Scheerpeltz, 1933: 1305 (Mitomorphus; catalog).

  • Distribution: India.

sattelbergenis Last, 1980: 160 (Mitomorphus; Type locality: New Guinea: Sattelberg, Huon Golf).

  • Distribution: New Guinea.

sedatus Cameron, 1932a: 140 (Mitomorphus; Type locality: Pahang: Fraser's Hill, alt. 4000 feet).

  • Distribution: Malaysia.

similis Last, 1980: 160 (Mitomorphus; Type locality: New Guinea: Sattelberg, Huon Golf).

  • Distribution: New Guinea.

subnigroaeneus Coiffait, 1982: 51 (Mitomorphus; Type locality: Nepal: Nagarkatpedi).

  • — Coiffait, 1975a: 154 (Mitomorphus; [Note: Misidentified as nigroaeneus according to Coiffait, 1982: 51]; Nepal).

  • Distribution: Nepal.

sumatrensis Cameron, 1933d: 385 (Mitomorphus; Type locality: Gunung Singgalang, Sumatra Westkust).

  • Distribution: Indonesia.

Neohypnus [32 species; Neotropical, Nearctic, Ethiopian, and Australian Regions]

Neohypnus Coiffait and Saiz, 1964: 512, 522 (species included: chilensis, andinus). Type species: Neohypnus chilensis Coiffait and Saiz, fixed by original designation.

  • — Coiffait and Saiz, 1968: 345 (list of Chilean species).

  • — Saiz, 1970b: 383, 385 (characters; key to species of Chile).

  • — Smetana, 1982: 198 (characters; habitus; revision of and key to species of North America north of Mexico; [Note: According to Smetana, 1982: 198, 199, most of the Central and South American species of Xantholinus belong in this genus but the species to be moved were not named]).

  • — Frank, 1986a: 369 (checklist of species of Florida).

  • — Campbell and Davies, 1991: 124 (Neohypnus; checklist of species; Canada).

  • — Downie and Arnett, 1996: 401 (characters; list of species of Northeastern North America).

  • — Lundgren, 1998: 50 (list of species from Florida; records for other States listed for each species).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 391 (28 Nearctic species; notes; characters in key).

andinus Fauvel, 1866a: 350 [= 1866b: 101 (Xantholinus; Type locality: Santiago).

  • — Fauvel, 1878f: 243 (Xantholinus; synonym of puncticeps).

  • — Lynch, 1884: 189 (Xantholinus; subgenus Gyrohypnus; characters; Argentina).

  • — Bernhauer and Schubert, 1914: 300 (Xantholinus; catalog; Uruguay).

  • — Coiffait and Saiz, 1964: 524 (Neohypnus, cited as Neotyphlus; characters; Chile).

  • — Saiz, 1970b: 385, 386 (Neohypnus; characters; Chile).

  • — Frank, 1982: 32 (Xantholinus; list of parasites).

  • — Kuschel, 1990: 26 (Neohypnus; introduced, South American; winged; habitat; New Zealand).

  • Distribution: Chile, Argentina, Uruguay, New Zealand.

attenuatus Erichson, 1839b: 330 (Xantholinus; Type locality: Americae insulae St. Thoma; Puerto Rico; St. Vincentii; Brasilia; [Note: See lectotype designation by Smetana, 1982]).

  • — Scriba, 1855: 299 (Xantholinus; Venezuela).

  • — Guérin-Méneville, 1857: 34 (Xantholinus; characters; Cuba; Puerto Rico; St. Thomas; St. Vincent; Brazil).

  • — Chevrolat and Fauvel, 1863: 430 (Xantholinus; characters; Cuba, St. Vincent, St. Thomas, Puerto Rico).

  • — Solsky, 1872: 308 (Xantholinus; Cayenne).

  • — Sharp, 1876: 203 (Xantholinus; Brazil).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog; Mexico; Argentina; Paraguay).

  • — Blackwelder, 1943: 478 (Xantholinus; characters; Cuba; Jamaica; Puerto Rico; St. Thomas; St. Kitts; Antigua; Montserrat; Guadeloupe; Dominica; Barbados; St. Vincent; Grenada; Tobago; Trinidad).

  • — Fagel, 1972: 104 (Gyrohypnus; notes; Sainte Helene).

  • — Smetana, 1982: 202 (Neohypnus; lectotype designation: [Note: Locality not cited]; characters; California; Texas).

  • — Smetana, 1988a: 554 (Neohypnus; Mississippi).

  • Distribution: Saint Helena, Argentina, Paraguay, Brazil, French Guiana, Venezuela, West Indies, Mexico, USA.

armatus Wollaston, 1877: 32 (Xantholinus; Type locality: St. Helena: Plantation).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • — Fagel, 1972: 104 (Gyrohypnus; synonym of attenuatus).

  • — Smetana, 1982: 202 (Neohypnus; synonym of attenuatus).

temporalis LeConte, 1880: 172 (Xantholinus; [preoccupied]; Type locality: Cedar Keys, Florida; [Note: See lectotype designation by Smetana, 1982]).

  • — Casey, 1906: 387 (Gyrohypnus; characters; Florida).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; synonym of floridae).

  • — Notman, 1920b: 704 (Gyrohypnus; Florida).

  • — Smetana, 1982: 202 (Neohypnus; lectotype designation; synonym of attenuatus).

floridae Duvivier, 1883: 156 (Xantholinus; replacement name for temporalis LeConte).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • — Smetana, 1982: 202 (Neohypnus; synonym of attenuatus).

armatus Wollaston, 1877, see: attenuatus Erichson, 1839.

beckeri Smetana, 1982: 222 (Neohypnus; Type locality: Quebec: Rigaud. Also cited from Ontario, Arkansas, Illinois, Maine, Massachusetts, Minnesota, New Hampshire, New Jersey, Pennsylvania, Vermont).

  • — Downie and Arnett, 1996: 401 (Neohypnus; USA).

  • Distribution: Canada, USA.

bicolor Smetana, 1982: 223 (Neohypnus; Type locality: Alabama: University. Also cited from Florida).

  • — Smetana, 1990c: 86 (Neohypnus; Florida).

  • Distribution: USA.

bipartitus Casey, 1906, see: dimidiatus LeConte, 1880.

chilensis Coiffait and Saiz, 1964: 522 (Neohypnus, cited as Nehoypnus; Type locality: Chile: Romeral, Curicó).

  • — Saiz, 1970b: 386 (Neohypnus; characters; Chile).

  • Distribution: Chile.

coloratus Smetana, 1988a: 554 (Neohypnus; Type locality: Florida: Marion County, Ocala).

  • Distribution: USA.

cuspifer Smetana, 1982: 252 (Neohypnus; Type locality: New York: Peekskill).

  • — Downie and Arnett, 1996: 401 (Neohypnus; USA).

  • Distribution: USA.

dimidiatus LeConte, 1880: 173 (Xantholinus; Type locality: California; [Note: See lectotype designation by Smetana, 1982]. Originally cited from San Bernardino, and Mojave Desert, California).

  • — Casey, 1906: 388 (Gyrohypnus; characters; California).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • — Smetana, 1982: 204 (Neohypnus; lectotype designation: Cal. [California]; characters; Arizona; California; Baja California).

  • Distribution: USA, Mexico.

bipartitus Casey, 1906: 388 (Gyrohypnus; Type locality: Arizona; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • — Smetana, 1982: 204 (Neohypnus; lectotype designation; synonym of dimidiatus).

infumatus Casey, 1906: 388 (Gyrohypnus; Type locality: California: Pasadena; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Smetana, 1982: 204 (Neohypnus; lectotype designation; synonym of dimidiatus).

emmesus Gravenhorst, 1802: 176 (Staphylinus; Type locality: America septentr.; [Note: See lectotype designation by Smetana, 1982]).

  • — Gravenhorst, 1806: 98 (Staphylinus).

  • — Say, 1830: 39 (Xantholinus; characters).

  • — Say, 1834: 452 (Xantholinus; characters).

  • — Erichson, 1839b: 324 (Xantholinus; characters; America septentrionalis).

  • — Casey, 1906: 389 (Gyrohypnus; characters; New York; Virginia; Ohio; Louisiana).

  • — Blatchley, 1910: 396 (Xantholinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • — Smetana, 1982: 248 (Neohypnus; lectotype designation; characters; Ontario; Quebec; Alabama; Arkansas; Florida; Georgia; Illinois; Indiana; Iowa; Kansas; Kentucky; Louisiana; Maine; Maryland; Massachusetts; Michigan; Minnesota; Missouri; New Hampshire; New Jersey; New York; North Carolina; Ohio; Oklahoma; Pennsylvania; South Carolina; Tennessee; Texas; Virginia; West Virginia).

  • — Smetana, 1988a: 556 (Neohypnus; Mississippi).

  • — Downie and Arnett, 1996: 401 (Neohypnus; USA).

  • Distribution: Canada, USA.

sanguinolentus Melsheimer, 1844: 34 (Xantholinus; Type locality: Pennsylvania; [Note: See lectotype designation by Smetana, 1982]).

  • — Casey, 1906: 389 (Gyrohypnus; synonym of emmesus).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; synonym of emmesus).

  • — Smetana, 1982: 248 (Neohypnus; lectotype designation; synonym of emmesus).

luteiventris Casey, 1906: 389 (Gyrohypnus; Type locality: Louisiana: Ponchatoula; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • — Notman, 1920b: 704 (Gyrohypnus; Florida).

  • — Smetana, 1982: 248 (Neohypnus; lectotype designation; synonym of emmesus).

floridae Duvivier, 1883, see: attenuatus Erichson, 1839.

fragilis Casey, 1906: 394 (Gyrohypnus; Type locality: New Mexico; [Note: See lectotype designation by Smetana, 1982]. Originally cited from Montana to Arizona and New Mexico).

  • — Fall and Cockerell, 1907: 167 (Xantholinus; New Mexico).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; catalog).

  • — Hatch, 1957: 237 (Hyponygrus; characters; British Columbia; Washington; Oregon).

  • — Smetana, 1982: 236 (Neohypnus; lectotype designation: N.M. [New Mexico]; characters; British Columbia; Arizona; California; Colorado; Illinois; Indiana; Kansas; Nevada; New Jersey; New Mexico; New York; Oklahoma; Oregon; Pennsylvania; Texas; Utah; Virginia; Washington; Baja California).

  • — Smetana, 1988a: 556 (Neohypnus; Ontario; Arizona; Nebraska).

  • — Downie and Arnett, 1996: 401 (Neohypnus; USA).

  • — Schmidt, 1999: 108 (Neohypnus; food for adults and larvae used for rearing species; rearing methods).

  • Distribution: Canada, USA.

fucosus Casey, 1885, see: fusciceps LeConte, 1880.

fusciceps LeConte, 1880: 173 (Xantholinus; Type locality: Pennsylvania; [Note: See lectotype designation by Smetana, 1982]. Originally cited from Middle, western, and southern States).

  • — Casey, 1906: 390 (Gyrohypnus; characters; New York to Iowa).

  • — Blatchley, 1910: 396 (Xantholinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; variety of emmesus).

  • — Notman, 1920a: 180 (Gyrohypnus; New York).

  • — Smetana, 1982: 240 (Neohypnus; lectotype designation: Penn. [Pennsylvania]; characters; Florida; North Carolina; Virginia; Pennsylvania; New Jersey).

  • — Smetana, 1988a: 556 (Neohypnus; North Carolina).

  • — Downie and Arnett, 1996: 401 (Neohypnus; USA).

  • Distribution: USA.

fucosus Casey, 1885: 126 (Xantholinus; Type locality: Cape May, New Jersey).

  • — Horn, 1885a: 109 (Xantholinus; synonym of emmesus variety fusciceps).

  • — Casey, 1906: 390 (Gyrohypnus; characters; New Jersey; Virginia).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; synonym of emmesus).

  • — Smetana, 1982: 240 (Neohypnus; synonym of fusciceps).

giulianii Smetana, 1982: 227 (Neohypnus; Type locality: California: Inyo Co.: Deep Springs Vy.).

  • — Smetana, 1988a: 555 (Neohypnus; California).

  • Distribution: USA.

grandis Smetana, 1982: 212 (Neohypnus; Type locality: Arizona: Cochise Co.: Chiric. Mts., Rustler Park, 8400 feet. Also cited from New Mexico).

  • Distribution: USA.

hamatus Say, 1830: 39 [= 1834: 453] (Xantholinus; Type locality: [Note: See neotype designation by Smetana, 1982]. Originally cited from the United States).

  • — Erichson, 1840: 924 (Xantholinus; synonym of obscurus).

  • — LeConte, 1850: 219 (Xantholinus; Lake Superior).

  • — Provancher, 1877: 251 (Xantholinus; characters; Quebec).

  • — Casey, 1906: 391 (Gyrohypnus; characters; Rhode Island and Ontario to northern New Mexico, Colorado, Nevada, Idaho, and Manitoba).

  • — Fall and Cockerell, 1907: 167 (Xantholinus; New Mexico).

  • — Blatchley, 1910: 397 (Xantholinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Hatch, 1957: 236 (Hyponygrus; characters; British Columbia; Washington; Oregon; Idaho).

  • — Smetana, 1982: 228 (Neohypnus; neotype designation: [Note: Locality not cited; specimen selected from LeConte collection with “grey round disc”]; characters; Alberta; British Columbia; Manitoba; Northwest Territories; Nova Scotia; Ontario; Quebec; Saskatchewan; Colorado; Michigan; Minnesota; Montana; Nevada?; New York; North Dakota; Oregon; Vermont; Wyoming).

  • — Smetana, 1988a: 555 (Neohypnus; New Hampshire).

  • — Levesque and Levesque, 1996: 288 (Neohypnus; seasonal abundance; Quebec).

  • — Downie and Arnett, 1996: 401 (Neohypnus; USA).

  • Distribution: Canada, USA.

macilentus Casey, 1906: 394 (Gyrohypnus; Type locality: Nevada; [Note: See lectotype designation by Smetana, 1982]. Originally cited from Nevada: Reno; Wyoming: Laramie; Manitoba).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • — Smetana, 1982: 228 (Neohypnus; lectotype designation: Nev. [Nevada]; synonym of hamatus).

infumatus Casey, 1906, see: dimidiatus LeConte, 1880.

lecontei Duvivier, 1883: 156 (Xantholinus; replacement name for sanguinipennis LeConte).

  • — Casey, 1906: 393 (Gyrohypnus; synonym of sanguinipennis).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Smetana, 1982: 211 (Neohypnus; characters; Connecticut; Georgia; Massachusetts; New Jersey; New York; Virginia).

  • — Downie and Arnett, 1996: 401 (Neohypnus; USA).

  • Distribution: USA.

sanguinipennis LeConte, 1880: 174 (Xantholinus; [preoccupied]; Type locality: Pennsylvania; [Note: See lectotype designation by Smetana, 1982]).

  • — Casey, 1906: 393 (Gyrohypnus; characters; New Jersey).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; synonym of lecontei).

  • — Smetana, 1982: 211 (Neohypnus; lectotype designation; synonym of lecontei).

linearis Blatchley, 1910, see: lineatus Wendeler, 1924.

lineatus Wendeler, 1924a: 344 (Xantholinus; replacement name for linearis Blatchley).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1311 (Xantholinus; catalog).

  • — Smetana, 1982: 239 (Neohypnus; characters; Ontario; llinois; Indiana; Kentucky; Michigan; Nebraska; New York; Ohio).

  • — Smetana, 1988a: 556 (Neohypnus; Ontario).

  • — Smetana, 1990c: 86 (Neohypnus; Arkansas).

  • — Downie and Arnett, 1996: 401 (Neohypnus; USA).

  • Distribution: Canada, USA.

linearis Blatchley, 1910: 395 (Xantholinus; [preoccupied]; Type locality: Putnam County, Indiana; [Note: See lectotype designation by Smetana, 1982]).

  • — Scheerpeltz, 1933: 1312 (Xantholinus; synonym of lineatus).

  • — Smetana, 1982: 239 (Neohypnus; lectotype designation; synonym of lineatus).

lobatus Smetana, 1982: 216 (Neohypnus; Type locality: Arizona: Madera Co.: Santa Rita Mts., 5500–6000 feet).

  • Distribution: USA.

luteiventris Casey, 1906, see: emmesus Gravenhorst, 1802.

lustrans Casey, 1906: 403 (Leptacinodes; Type locality: Colorado: Denver; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; catalog).

  • — Smetana, 1982: 231 (Neohypnus; lectotype designation; characters; Arizona; California; Colorado; New Mexico; Utah).

  • Distribution: USA.

macilentus Casey, 1906, see: hamatus Say, 1830.

mediocris Smetana, 1982: 220 (Neohypnus; Type locality: Arizona: Cochise Co.: Chiric. Mts., Rucker Cn., 5500 feet).

  • — Smetana, 1990c: 86 (Neohypnus; Arizona).

  • Distribution: USA.

melanops Casey, 1906: 390 (Gyrohypnus; Type locality: Virginia; [Note: See lectotype designation by Smetana, 1982]. Also cited from New York, Long Island, and Virginia to Kansas and Colorado, Greeley).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • — Notman, 1920a: 180 (Gyrohypnus; New York).

  • — Smetana, 1982: 242 (Neohypnus; lectotype designation: Va. [Virginia]; characters; Ontario; Quebec; Colorado; Florida; Illinois; Indiana; Iowa; Kansas; Maryland; Massachusetts; Michigan; New Jersey; New York; Pennsylvania; Texas; Virginia).

  • — Smetana, 1990c: 86 (Neohypnus; Oklahoma).

  • — Downie and Arnett, 1996: 401 (Neohypnus; USA).

  • Distribution: Canada, USA.

miles Smetana, 1982: 214 (Neohypnus; Type locality: Arizona: Chiric. Mts., Ash Spg., 7 mi. SW Portal, 6000 feet. Also cited from New Mexico).

  • Distribution: USA.

mollinus Casey, 1906, see: pusillus Sachse, 1852.

monstrosus Smetana, 1982: 245 (Neohypnus; Type locality: New York: N. Lebanon. Also cited from Ontario, Illinois, Indiana, Iowa, Massachusetts, Michigan, Missouri, New Hampshire, New Jersey, Pennsylvania, Vermont).

  • — Smetana, 1988a: 556 (Neohypnus; Ontario; New York).

  • — Smetana, 1990c: 86 (Neohypnus; Indiana; Iowa).

  • — Downie and Arnett, 1996: 401 (Neohypnus; USA).

  • Distribution: Canada, USA.

obscurus Erichson, 1839b: 330 (Xantholinus; Type locality: Pensylvania; [Note: See lectotype designation by Smetana, 1982]).

  • — LeConte, 1850: 220 (Xantholinus; Lake Superior).

  • — LeConte, 1859: 568 (synonym of hamatus Say).

  • — LeConte, 1880: 172, 173 (Xantholinus; characters; Lake Superior and Canada to California).

  • — Casey, 1906: 391 (Gyrohypnus; synonym of hamatus).

  • — Blatchley, 1910: 397 (Xantholinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; synonym of hamatus).

  • — Notman, 1919: 136 (Xantholinus; New York).

  • — Smetana, 1982: 225 (Neohypnus; lectotype designation; characters; Alberta; British Columbia; Manitoba; New Brunswick; Newfoundland; Nova Scotia; Ontario; Quebec; Saskatchewan; Arizona; Arkansas; Colorado; Connecticut; Idaho; Illinois; Indiana; Iowa; Maine; Massachusetts; Michigan; Minnesota; Nevada; New Hampshire; New Jersey; New Mexico; New York; North Dakota; Ohio; Oregon; Pennsylvania; South Dakota; Texas; Utah; Washington, Wisconsin).

  • — Smetana, 1988a: 555 (Neohypnus; California).

  • — Levesque and Levesque, 1996: 288 (Neohypnus; seasonal abundance; Quebec).

  • — Downie and Arnett, 1996: 401 (Neohypnus; USA).

  • — Bain, 1998: 39 (Neohypnus; 17th century remains; Massachusetts).

  • — Schmidt, 1999: 108 (Neohypnus; food for adults and larvae used for rearing species; rearing methods).

  • Distribution: USA, Canada.

oregonus Casey, 1906: 392 (Gyrohypnus; Type locality: Oregon, Portland; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Smetana, 1982: 225 (Neohypnus; lectotype designation; synonym of obscurus).

protractus Casey, 1906: 392 (Gyrohypnus; Type locality: Wisconsin, Bayfield; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Smetana, 1982: 225 (Neohypnus; lectotype designation; synonym of obscurus).

onustus Smetana, 1982: 247 (Neohypnus; Type locality: New Jersey: Phillipsburg. Also cited from Massachusetts, New York, Pennsylvania).

  • — Downie and Arnett, 1996: 401 (Neohypnus; USA).

  • Distribution: USA.

oregonus Casey, 1906, see: obscurus Erichson, 1839.

picipennis LeConte, 1880: 172 (Xantholinus; Type locality: California; [Note: See lectotype designation by Smetana, 1982]).

  • — Casey, 1906: 385 (Xantholinus; characters; California; British Colombia).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Hatch, 1957: 235 (Hyponygrus; characters; British Columbia; Washington; Oregon).

  • — Smetana, 1982: 207 (Neohypnus; lectotype designation; characters; British Columbia; California; Oregon; Washington; Baja California).

  • Distribution: Canada, USA.

pirum Smetana, 1982: 216 (Neohypnus; Type locality: Arizona: Cochise Co.: Chiricahua Mts., Portal, 5000 feet. Also cited from Texas).

  • — Smetana, 1990c: 86 (Neohypnus; Texas).

  • Distribution: USA.

praegnans Smetana, 1982: 219 (Neohypnus; Type locality: Arizona: Cochise Co.: Huachuca Mts., Ramsey Cn., 5200 feet).

  • Distribution: USA.

protractus Casey, 1906, see: obscurus Erichson, 1839.

pusillus Sachse, 1852: 124 (Xantholinus; Type locality: Savannah, Georgia [Note: See neotype designation by Smetana, 1982]. Originally cited from Georgia).

  • — Casey, 1906: 393 (Gyrohypnus; characters; Virginia to Lake Superior and Texas, Arizona and Mexico).

  • — Blatchley, 1910: 398 (Xantholinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1314 (Xantholinus; catalog).

  • — Hu and Frank, 1995a (Neohypnus; life cycle; feeding; biological notes).

  • — Scheerpeltz, 1971b: 205 (Xantholinus; Guatemala).

  • — Smetana, 1981: 385 (Neohypnus).

  • — Smetana, 1982: 233 (Neohypnus; neotype designation: Savannah, Georgia; characters; Alabama; Arizona; Arkansas; California; Colorado; Florida; Georgia; Illinois; Indiana; Kansas; Louisiana; Maine; Maryland; Michigan; Missouri; New Mexico; North Carolina; Ohio; Oklahoma; Pennsylvania; South Carolina; Tennessee; Texas; Virginia; West Virginia).

  • — Smetana, 1988a: 555 (Neohypnus; Alabama; North Carolina; Mississippi).

  • — Downie and Arnett, 1996: 401 (Neohypnus; USA).

  • Distribution: USA, Mexico, Guatemala.

mollinus Casey, 1906: 392 (Gyrohypnus; Type locality: California: Elsinore, Riverside Co.; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • — Smetana, 1982: 233 (Neohypnus; lectotype designation; synonym of pusillus).

sagittifer Smetana, 1982: 217 (Neohypnus; Type locality: Arizona: Cochise Co.: Chiric. Mts., Rustler Park, 8400 feet).

  • — Smetana, 1990c: 86 (Neohypnus; Arizona).

  • Distribution: USA.

sanguinipennis LeConte, 1880, see: lecontei Duvivier, 1883.

sanguinolentus Melsheimer, 1844, see: emmesus Gravenhorst, 1802.

septentrionalis Saiz, 1970b: 387 (Neohypnus; Type locality: Chile: Azapa, Tarapacá).

  • Distribution: Chile.

solus Smetana, 1982: 220 (Neohypnus; Type locality: Arizona: Dragoon).

  • Distribution: USA.

temporalis LeConte, 1880, see: attenuatus Erichson, 1839.

vilis Sharp, 1885: 483 (Xantholinus; Type locality: San Geronimo, Vera Paz; [Note: See lectotype designation by Smetana, 1981]. Originally cited from Mexico, Guanajuato, Cordova, Jalapa; Guatemala, near the city, San Geronimo, Torola, Capetillo, Panajachel).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; synonym of pusillus).

  • — Smetana, 1981: 385 (Neohypnus; lectotype designation: San Geronimo, Vera Paz; valid species; characters).

  • Distribution: Mexico, Guatemala.

Neoleptacinus [2 species; Neotropical Region]

Neoleptacinus Coiffait and Saiz, 1964: 511, 515 (species included: cribripennis). Type species: Leptolinus cribripennis Fairmaire and Germain, fixed by original designation and monotypy.

  • — Coiffait and Saiz, 1968: 345 (list of Chilean species).

  • — Saiz, 1968a: 89 (characters).

  • — Saiz, 1970b: 382, 384 (characters).

cribripennis Fairmaire and Germain, 1861: 433 (Leptolinus; Type locality: Concepcion).

  • — Fauvel, 1866a: 351 [= 1866b: 102] (Leptacinus; characters; Chile).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; catalog).

  • — Coiffait and Saiz, 1964: 515 (Neoleptacinus; characters; Chile).

  • — Saiz, 1968a: 90 (Neoleptacinus; characters).

  • — Saiz, 1970b: 384 (Neoleptacinus; Chile).

  • Distribution: Chile.

microphtalmus Saiz, 1968a: 90 (Neoleptacinus; Type locality: Chile: Cordillera Pelada, Valdivia).

  • — Saiz, 1970b: 384 (Neoleptacinus; Chile).

  • Distribution: Chile.

Neolinus [1 species; Neotropical Region]

Neolinus Coiffait, 1981g: 292 (species included: galapagoensis). Type species: Neolinus galapagoensis Coiffait, fixed by original designation and monotypy.

galapagoensis Coiffait, 1981g: 292 (Neolinus; Type locality: Ile Galapagos: Ile Isabela, environs de Santa Tomas).

  • Distribution: Galapagos Islands.

Neoxantholinus [21 species; Neotropical, Nearctic, and Australian Regions]

Neoxantholinus Cameron, 1945d: 783 (species included: rufulus, brouni). Type species: Metoponcus rufulus Broun, fixed by original designation.

  • — Steel, 1949a: 268 (without explicitly listing them stated that all South American species of Oligolinus should be transferrd to Neoxantholinus).

  • — Blackwelder, 1952: 261 (type species: rufulus).

  • — Smetana, 1977c: 361 (all species included in Oligolinus by Scheerpeltz, 1953 should be moved to Neoxantholinus).

  • — Smetana, 1982: 62 (characters; habitus; revision of species of North America north of Mexico).

  • — Frank, 1986a: 369 (checklist of species of Florida).

  • — Lundgren, 1998: 50 (species from Florida; records for other States listed).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 391 (1 Nearctic species; notes; characters in key).

basiventris Sharp, 1876: 206 (Metoponcus; Type locality: St. Paulo).

  • — Bernhauer and Schubert, 1914: 291 (Oligolinus; catalog).

  • — Steel, 1949a: 268 (stated that all South American species of Oligolinus should be transferred to Neoxantholinus but did not explicitly list the species).

  • — Scheerpeltz, 1953: 604 (Oligolinus; characters in key; Brazil).

  • — Scheerpeltz, 1960c: 92 (Oligolinus; characters; Brazil).

  • — Smetana, 1977c: 361 (Neoxantholinus; all species included in Oligolinus by Scheerpeltz, 1953 should be moved to Neoxantholinus).

  • Distribution: Brazil.

beckeri Scheerpeltz, 1953: 602 (Oligolinus; Type locality: Guatemala: San Marcos).

  • — Scheerpeltz, 1960c: 94 (Oligolinus; characters; Guatemala).

  • — Smetana, 1977c: 361 (Neoxantholinus; all species included in Oligolinus by Scheerpeltz, 1953 should be moved to Neoxantholinus).

  • Distribution: Guatemala.

brouni Sharp, 1876a: 22 (Metoponcus; Type locality: Not cited, presumably New Zealand).

  • — Fauvel, 1877: 239 (Metoponcus; characters; New Zealand).

  • — Broun, 1880: 106 (Metoponcus; characters).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; catalog).

  • — Cameron, 1945d: 784 (Neoxantholinus).

  • — Steel, 1949a: 268 (Neoxantholinus).

  • — Kuschel, 1990: 26 (Neoxantholinus; endemic; winged; habitat; New Zealand).

  • — Emberson, 1998: 36 (Neoxantholinus; collecting notes; Chatham Islands).

  • Distribution: New Zealand, Chatham Islands.

cairnsensis Blackburn, 1895, see: cyaneipennis W. J. MacLeay, 1873.

cingulatus Sharp, 1885: 503 (Metoponcus; Type locality: Panama, Volcan de Chiriqui 2500 to 4000 feet).

  • — Bernhauer and Schubert, 1914: 291 (Oligolinus; catalog).

  • — Steel, 1949a: 268 (stated that all South American species of Oligolinus should be transferred to Neoxantholinus but did not explicitly list the species).

  • — Scheerpeltz, 1953: 605 (Oligolinus; characters in key; Costa Rica; Panama).

  • — Scheerpeltz, 1960c: 92 (Oligolinus; characters; Central America).

  • — Smetana, 1977c: 361 (Neoxantholinus; all species included in Oligolinus by Scheerpeltz, 1953 should be moved to Neoxantholinus).

  • Distribution: Panama, Costa Rica.

cristatus Smetana, 1982: 64 (Neoxantholinus; Type locality: Florida: Key West).

  • — Smetana, 1990c: 84 (Neoxantholinus; Arkansas).

  • Distribution: USA.

cyaneipennis W. J. MacLeay, 1873: 137 (Leptacinus; Type locality: Gayndah).

  • — Fauvel, 1877: 237 (Metoponcus; characters; Australia).

  • — Olliff, 1887: 477 (Metoponcus; characters; Australia).

  • — Olliff, 1889: 81 (Metoponcus; Lord Howe Island).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; cited as cyanipennis; catalog).

  • — Bernhauer, 1920: 14 (Metoponcus; cited as cyanipennis; Australia).

  • — Lea, 1925: 213 (Metoponcus; notes; nest of Euponera lutea; Australia).

  • — Scheerpeltz, 1933: 1298 (Metoponcus; catalog).

  • — Steel, 1949a: 268 (Neoxantholinus).

  • — Smetana, 1977c: 361 (Neoxantholinus; characters; differs from others in genus).

  • Distribution: Australia, Lord Howe Island.

cairnsensis Blackburn, 1895: 204 (Metoponcus; Type locality: N. Queensland, near Cairns).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; cited as cairusensis; catalog).

  • — Lea, 1925: 213 (Metoponcus; variety of cyaneipennis).

  • — Scheerpeltz, 1933: 1298 (Metoponcus; variety of cyaneipennis).

nigricollis Bernhauer, 1920: 15 (Metoponcus; aberration of cyaneipennis; [Note: Nomen nudum (Articles 1.3.4, 45.5)]).

  • — Lea, 1925: 213 (Metoponcus; synonym of cyaneipennis).

  • — Scheerpeltz, 1933: 1298 (Metoponcus; aberration of cyaneipennis).

howensis Lea, 1925: 214 (Metoponcus; variety of cyaneipennis; Type locality: Lord Howe Island).

  • — Scheerpeltz, 1933: 1298 (Metoponcus; variety of cyaneipennis).

darlingtoni Blackwelder, 1943: 506 (Oligolinus; Type locality: Dominican Republic: Loma Rucilla and the mountains to the north, elevation 5000 to 8000 feet).

  • — Steel, 1949a: 268 (stated that all South American species of Oligolinus should be transferred to Neoxantholinus but did not explicitly list the species).

  • Distribution: West Indies.

filarius Erichson, 1839b: 334 (Leptacinus; Type locality: Columbia; [Note: See lectotype designation by Smetana, 1978c]).

  • — Kraatz, 1857d: 652 (Metoponcus).

  • — Sharp, 1876: 206 (Lithocharodes; Brazil).

  • — Sharp, 1885: 503 (Metoponcus; Guatemala, Nicaragua, Colombia, Brazil).

  • — Fauvel, 1891: 104 (Metoponcus; Venezuela).

  • — Bernhauer and Schubert, 1914: 291 (Oligolinus; catalog).

  • — Blackwelder, 1943: 506 (Oligolinus; characters; St. Vincent).

  • — Steel, 1949a: 268 (stated that all South American species of Oligolinus should be transferred to Neoxantholinus but did not explicitly list the species).

  • — Scheerpeltz, 1953: 604 (Oligolinus; characters in key; Mexico; Guatemala; Honduras; Nicaragua; Costa Rica; Panama; Antilles; Colombia; Venezuela; Ecuador; Brazil).

  • — Scheerpeltz, 1960c: 92 (Oligolinus; characters; Mexico; Central America; Brazil; northern South America).

  • — Smetana, 1977c: 361 (Neoxantholinus; all species included in Oligolinus by Scheerpeltz, 1953 should be moved to Neoxantholinus).

  • — Legner and Moore, 1977: 176 (Oligolinus; found under bark).

  • — Smetana, 1978c: 337 (Neoxantholinus; lectotype designation).

  • Distribution: Venezuela, Brazil, Ecuador, Colombia, Panama, Nicaragua, Honduras, Guatemala, Mexico, West Indies.

forsteri Scheerpeltz, 1960c: 92, 94 (Oligolinus; Type locality: Nordbrasilien: Belem do Para).

  • — Steel, 1949a: 268 (stated that all South American species of Oligolinus should be transferred to Neoxantholinus but did not explicitly list the species).

  • Distribution: Brazil.

gracilentus Scheerpeltz, 1953: 603 (Oligolinus; Type locality: Guatemala: Calel).

  • — Scheerpeltz, 1960c: 94 (Oligolinus; characters; Guatemala).

  • — Smetana, 1977c: 361 (Neoxantholinus; all species included in Oligolinus by Scheerpeltz, 1953 should be moved to Neoxantholinus).

  • Distribution: Guatemala.

guatemalensis Sharp, 1885: 503 (Metoponcus; Type locality: Guatemala, San Geronimo; [Note: See lectotype designation by Smetana, 1977c]).

  • — Bernhauer and Schubert, 1914: 291 (Oligolinus; catalog).

  • — Steel, 1949a: 268 (stated that all South American species of Oligolinus should be transferred to Neoxantholinus but did not explicitly list the species).

  • — Scheerpeltz, 1953: 604 (Oligolinus; characters in key; Mexico; Guatemala; Honduras; Nicaragua; Costa Rica).

  • — Scheerpeltz, 1960c: 92 (Oligolinus; characters; Mexico; Central America).

  • — Smetana, 1977c: 361 (Neoxantholinus; all species included in Oligolinus by Scheerpeltz, 1953 should be moved to Neoxantholinus; lectotype designation; characters).

  • Distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica.

holisoides Sharp, 1876: 206 (Metoponcus; Type locality: Ega).

  • — Sharp, 1885: 505 (Metoponcus; characters; Guatemala, Zapote, Torola, Chacoj; Brazil).

  • — Bernhauer and Schubert, 1914: 291 (Oligolinus; catalog).

  • — Steel, 1949a: 268 (stated that all South American species of Oligolinus should be transferred to Neoxantholinus but did not explicitly list the species).

  • — Scheerpeltz, 1953: 605 (Oligolinus; characters in key; Mexico; Guatemala; Honduras; Nicaragua; Costa Rica; Panama; Colombia; Ecuador; Venezuela; Brazil).

  • — Scheerpeltz, 1960c: 93 (Oligolinus; characters; South America).

  • — Smetana, 1977c: 361 (Neoxantholinus; all species included in Oligolinus by Scheerpeltz, 1953 should be moved to Neoxantholinus).

  • Distribution: Venezuela, Brazil, Ecuador, Colombia, Panama, Costa Rica, Nicaragua, Honduras, Guatemala, Mexico.

howensis Lea, 1925, see: cyaneipennis W. J. MacLeay, 1873.

hubbardi Blackwelder, 1943: 505 (Oligolinus; Type locality: Montserrat).

  • — Steel, 1949a: 268 (stated that all South American species of Oligolinus should be transferred to Neoxantholinus but did not explicitly list the species).

  • Distribution: West Indies.

lineatus Schubert, 1909: 296 (Metoponcus; Type locality: Jalapa, Mexico; [Note: See lectotype designation by Smetana, 1977c]. Originally cited from südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d'Orizaba).

  • — Bernhauer and Schubert, 1914: 291 (Oligolinus; catalog).

  • — Steel, 1949a: 268 (stated that all South American species of Oligolinus should be transferred to Neoxantholinus but did not explicitly list the species).

  • — Scheerpeltz, 1953: 605 (Oligolinus; characters in key; Mexico).

  • — Scheerpeltz, 1960c: 93 (Oligolinus; characters; Mexico).

  • — Smetana, 1977c: 362 (Neoxantholinus; all species included in Oligolinus by Scheerpeltz, 1953 should be moved to Neoxantholinus; lectotype designation; characters).

  • Distribution: Mexico.

longiceps Sharp, 1885: 504 (Metoponcus; Type locality: Panama, Bugaba, Volcan de Chiriqui 2500 to 4000 feet).

  • — Bernhauer and Schubert, 1914: 291 (Oligolinus; catalog).

  • — Steel, 1949a: 268 (stated that all South American species of Oligolinus should be transferred to Neoxantholinus but did not explicitly list the species).

  • — Scheerpeltz, 1953: 606 (Oligolinus; characters in key; Costa Rica; Panama).

  • — Scheerpeltz, 1960c: 93 (Oligolinus; characters; Central America).

  • — Smetana, 1977c: 361 (Neoxantholinus; all species included in Oligolinus by Scheerpeltz, 1953 should be moved to Neoxantholinus).

  • Distribution: Panama, Costa Rica.

minutus Sharp, 1885: 505 (Metoponcus; Type locality: Panama, Bugaba).

  • — Bernhauer and Schubert, 1914: 291 (Oligolinus; catalog).

  • — Steel, 1949a: 268 (stated that all South American species of Oligolinus should be transferred to Neoxantholinus but did not explicitly list the species).

  • — Scheerpeltz, 1953: 605 (Oligolinus; characters in key; Costa Rica; Panama).

  • — Scheerpeltz, 1960c: 93 (Oligolinus; characters; Cental America).

  • — Smetana, 1977c: 361 (Neoxantholinus; all species included in Oligolinus by Scheerpeltz, 1953 should be moved to Neoxantholinus).

  • Distribution: Panama, Costa Rica.

nigricollis Bernhauer, 1920 [Note: Nomen nudum], see: cyaneipennis W. J. MacLeay, 1873.

norfolcensis Lea, 1925, see: rufulus Broun, 1880.

rufulus Broun, 1880: 106 (Metoponcus; Type locality: Parua).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; catalog).

  • — Scheerpeltz, 1933: 1298 (Metoponcus; catalog).

  • — Cameron, 1945d: 784 (Neoxantholinus).

  • — Steel, 1949a: 268 (Neoxantholinus).

  • — Smetana, 1977c: 361 (Neoxantholinus; characters).

  • — Smithers, 1998: 16 (Zeteotomus; references; Norfolk Island).

  • Distribution: New Zealand, Norfolk Island.

norfolcensis Lea, 1925: 214 (Metoponcus; variety of rufulus; Type locality: Norfolk Island).

  • — Scheerpeltz, 1933: 1298 (Metoponcus; variety of rufulus).

  • — Smithers, 1998: 16 (Zeteotomus; cited as norfolkensis; references; Norfolk Island).

sinuatus Sharp, 1885: 505 (Metoponcus; Type locality: Mexico, Patzcuaro).

  • — Bernhauer and Schubert, 1914: 291 (Oligolinus; catalog).

  • — Steel, 1949a: 268 (stated that all South American species of Oligolinus should be transferred to Neoxantholinus but did not explicitly list the species).

  • — Scheerpeltz, 1953: 601 (Oligolinus; Guatemala).

  • — Scheerpeltz, 1953: 606 (Oligolinus; characters in key; Mexico; Guatemala; Honduras; Costa Rica; Nicaragua; Panama).

  • — Scheerpeltz, 1960c: 94 (Oligolinus; characters; Mexico; Central America).

  • — Smetana, 1977c: 361 (Neoxantholinus; all species included in Oligolinus by Scheerpeltz, 1953 should be moved to Neoxantholinus).

  • Distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama.

testaceipennis Fauvel, 1863: 431 (Leptacinus; Type locality: Cuba; Saint-Domingue).

  • — Fauvel, 1878f: 242 (Metoponcus).

  • — Bernhauer and Schubert, 1914: 291 (Oligolinus; catalog).

  • — Bierig, 1931: 428 (Metoponcus, transferred from Leptacinus; characters; Panama).

  • — Scheerpeltz, 1933: 1299 (Metoponcus; catalog).

  • — Blackwelder, 1943: 507 (Oligolinus; characters; Cuba; Haiti; Dominican Republic).

  • — Steel, 1949a: 268 (stated that all South American species of Oligolinus should be transferred to Neoxantholinus but did not explicitly list the species).

  • — Legner and Moore, 1977: 176 (Oligolinus; found under bark).

  • Distribution: West Indies, Panama.

varians Sharp, 1885: 504 (Metoponcus; Type locality: Mexico, Cordova; Guatemala, Volcan de Fuego 6400 feet, Capetillo, San Geronimo, Cerro Zunil, Quiche Mountains 7000 to 9000 feet, Totonicapam 8500 to 10,500 feet).

  • — Bernhauer and Schubert, 1914: 291 (Oligolinus; catalog).

  • — Steel, 1949a: 268 (Neoxantholinus).

  • — Scheerpeltz, 1953: 601, 606 (Oligolinus; characters in key; Guatemala; Mexico; Honduras).

  • — Scheerpeltz, 1960c: 94 (Oligolinus; characters; Mexico; Central America).

  • — Smetana, 1977c: 361 (Neoxantholinus; all species included in Oligolinus by Scheerpeltz, 1953 should be moved to Neoxantholinus).

  • Distribution: Mexico, Guatemala, Honduras.

variabilis Blackwelder, 1943: 508 (Oligolinus; Type locality: Haiti: La Visite and vicinity in the La Selle Range, elevation 5000 to 7000 feet. Also cited from Cuba).

  • — Steel, 1949a: 268 (stated that all South American species of Oligolinus should be transferred to Neoxantholinus but did not explicitly list the species).

  • Distribution: West Indies.

Nepaliellus [1 species; Palaearctic Region]

Nepaliellus Bordoni, 1989: 2 (species included: absurdus). Type species: Nepaliellus absurdus Bordoni, by original designation and monotypy.

absurdus Bordoni, 1989: 2 (Nepaliellus; Type locality: Nepal: Ilam Distr., Mai Pokhari, 2100 m.).

  • Distribution: Nepal.

Nepalinus [2 species; Palaearctic Regions]

Nepalinus Coiffait, 1975a: 157 (subgenus of Xantholinus; species included: aculeatus). Type species: Xantholinus (Nepalinus) aculeatus Coiffait, fixed by original designation.

  • — Bordoni, 2000: 126 (valid genus).

aculeatus Coiffait, 1975a: 157 (Xantholinus; subgenus Nepalinus; Type locality: Phulchoki près Katmandou).

  • Distribution: Nepal.

parcipennis Bernhauer, 1933: 29 (Xantholinus; Type locality: Grenze Thibet Ost: Tatsienlu).

  • — Bordoni, 2000: 125 (Nepalinus; characters; China).

  • Distribution: China.

Notolinus [6 species; Ethiopian and Australian Regions]

Notolinus Casey, 1906: 371, 375 (species included: grossulus, fumipennis, hottentotus, parvus). Type species: Notolinus fumipennis Casey, fixed by subsequent designation by Blackwelder, 1943: 474.

  • — Bernhauer and Schubert, 1914: 300 (subgenus of Xantholinus).

  • — Cameron, 1932: 22 (subgenus of Xantholinus).

  • — Blackwelder, 1943: 473 (subgenus of Xantholinus; type species: fumipennis).

  • — Steel, 1949a: 268 (distinct from Xantholinus).

anthracinus Broun, 1880, see: socius Fauvel, 1877.

crateris Bernhauer, 1939a: 80 (Xantholinus; Type locality: Kenya: Cratère de l'Elgon, pied de Koitobbos, 4000 m; Camp III de l'Elgon, zone de Bruyères, est, 3500 m).

  • — Jeannel and Jarrige, 1949: 373 (Notolinus).

  • Distribution: Kenya.

fumipennis Casey, 1906: 376 (Notolinus; Type locality: Wellington).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; [Note: This species is moved back to Notolinus because the genus was elevated]; catalog).

  • Distribution: South Africa.

grossulus Casey, 1906: 376 (Notolinus; Type locality: Cape Town).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; [Note: This species is moved back to Notolinus because the genus was elevated]; catalog).

  • Distribution: South Africa.

hottentottus Sachse, 1852: 123 (Xantholinus; Type locality: Vorgeb. d. g. Hoffnung).

  • — Casey, 1906: 376 (Notolinus; characters; South Africa).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Fagel, 1972: 105 (Notolinus; notes; Sainte Helene).

  • Distribution: Saint Helena, South Africa.

morio Wollaston, 1871a: 410 (Xantholinus; [preoccupied]; Type locality: St. Helena).

  • — Wollaston, 1877: 31 (Xantholinus; characters; St. Helena).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • — Steel, 1949a: 270 (Leptacinus; synonym of sanctaehelenae).

  • — Fagel, 1972: 105 (Notolinus; synonym of hottentotus).

sanctaehelenae Steel, 1949a: 270 (Leptacinus; replacement name for morio Wollaston).

  • — Fagel, 1972: 105 (Notolinus; synonym of hottentotus).

morio Wollaston, 1871, see: hottentottus Sachse, 1852.

parvus Casey, 1906: 377 (Notolinus; Type locality: Cape Town and Wellington).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; [Note: This species is moved back to Notolinus because the genus was elevated]; catalog).

  • Distribution: South Africa.

picticornis Olliff, 1887, see: socius Fauvel, 1877.

picticornis Blackburn, 1888, see: socius Fauvel, 1877.

sanctaehelenae Steel, 1948, see: hottentottus Sachse, 1852.

socius Fauvel, 1877: 247 (Xantholinus; Type locality: Australie orientale et meridionale, Victoria, Rockhampton).

  • — Olliff, 1887: 490 (Xantholinus; characters; Australia).

  • — Bernhauer, 1908: 18 (Xantholinus; Australia).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; catalog).

  • — Lea, 1925: 215 (Leptacinus; from nest of Iridomyrmex conifer; Australia).

  • — Scheerpeltz, 1933: 1303 (Leptacinus; catalog).

  • — Kuschel, 1990: 26 (Notolinus; introduced, Australian species; winged; habitat; New Zealand).

  • — Klimaszewski, Newton and Thayer, 1996: 151 (Notolinus; adventive species; New Zeland).

  • — Emberson, 1998: 36 (Notolinus; collecting notes; Chatham Islands).

  • — Herman, 2001: 26 (Notolinus; synonymy).

  • Distribution: Australia, New Zealand, Chatham Islands.

anthracinus Broun, 1880: 105 (Xantholinus; Type locality: Whangarei Harbour).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; synonym of socius).

  • — Lea, 1925: 215 (Leptacinus; synonym of socius).

picticornis Olliff, 1887: 476 (Leptacinus; Type locality: Adelaide, Port Lincoln, South Australia).

  • — Herman, 2001: 26 (Notolinus; synonym of socius).

picticornis Blackburn, 1888: 7 (Leptacinus; [preoccupied]; Type locality: Port Lincoln district; near Adelaide).

  • — Blackburn, 1888d: 190 (Leptacinus; synonym of socius).

  • — Lea, 1925: 215 (Leptacinus; synonym of socius).

  • — Scheerpeltz, 1933: 1303 (Leptacinus; synonym of socius).

Nudobius [43 species; Neotropical, Nearctic, Palaearctic, and Ethiopian Regions]

Nudobius Thomson, 1860: 188 (species included: lentus). Type species: Staphylinus lentus Gravenhorst, fixed by monotypy.

  • — Fauvel, 1873b: 95 [= 1873a: 381] (synonym of Xantholinus).

  • — Mulsant and Rey, 1877a: 299 (characters; key to French species).

  • — Heyden, 1880: 75 (subgenus of Xantholinus; list of species of Siberian region).

  • — Sharp, 1885: 502 (notes; Mexican and Central American species).

  • — Fowler, 1888: 291 (notes; larval characters).

  • — Heyden, Reitter, and Weise, 1891: 105 (list of species of Europe and Caucasus).

  • — Heyden, 1893: 44 (list of species of Siberian region).

  • — Ganglbauer, 1895: 475 (adult and larval characters).

  • — Casey, 1906: 367, 380 (characters; key to species).

  • — Heyden, Reitter, and Weise, 1906: 161 (list of species of Europe and Caucasus).

  • — Reitter, 1908: 114 (subgenus of Xantholinus; key to species).

  • — Petri, 1912: 70 (list of species of Siebenbürgen).

  • — Bernhauer and Schubert, 1914: 298 (world catalog; 18 species).

  • — Leng, 1920: 105 (catalog of North American species).

  • — Winkler, 1925: 370 (catalog for Palaearctic region).

  • — Porta, 1926: 91 (characters; key to species of Italy).

  • — Roubal, 1930: 361 (catalog; Slovakia).

  • — Scheerpeltz, 1933: 1305 (world catalog supplement).

  • — Normand, 1935: 369 (list of species of Tunisia).

  • — Paulian, 1941: 210 (larval characters).

  • — Blackwelder, 1944: 146 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Kloet and Hincks, 1945: 165 (list of British species).

  • — Jeannel and Jarrige, 1949: 373 (cave species; type species: lentus).

  • — Tottenham, 1949: 369, 416 (type species: lentus Gravenhorst; checklist of British species).

  • — Blackwelder, 1952: 264 (type species: lentus).

  • — V. Hansen, 1952: 14 (subgenus of Xantholinus; characters).

  • — Coiffait, 1956a: 44, 55 (characters; key to French species).

  • — Scheerpeltz, 1956b: 11 (key to African species).

  • — Hatch, 1957: 237 (characters; species of Pacific Northwest).

  • — Kocher, 1958: 138 (checklist of species; Morocco).

  • — Smetana, 1958: 93 (characters; habitus of one species; type species: lentus; Czechoslovakian species).

  • — Moore, 1963a: 94 (characters).

  • — Székessy, 1963: 49 (characters).

  • — Palm, 1963b: 44 (characters; species of Sweden).

  • — Lohse, 1964: 161 (characters; key to central European species).

  • — Horion, 1965: 90 (list of central European species).

  • — Pototskaia, 1967: 47 (larval characters).

  • — Scheerpeltz, 1968a: 60 (catalog of Austrian species).

  • — Kasule, 1970: 74, 80 (larval characters).

  • — Coiffait, 1972: 202 (characters; larval characters; type species: lentus; key to species of western Palaearctic region; references).

  • — Tikhomirova, 1973a: 181 (checklist of species of USSR).

  • — Moore and Legner, 1975: 86 (catalog; North America north of Mexico).

  • — Pototskaia, 1976: 171 (larval characters).

  • — Nikitskii, 1976: 182 (larval characters).

  • — Pope, 1977: 28 (list of British species).

  • — Moore and Legner, 1979: 69 (characters; notes).

  • — Muona, 1979: 16 (list of Scandinavian species).

  • — Smetana, 1980a: 159 (review of and key to species of Mexico).

  • — Uhlig, Vogel, and Sieber, 1980: 245 (some species collected in Germany).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 13 (catalog; Poland).

  • — Uhlig and Vogler, 1981: 107 (list of some species collected in Germany; habitat and collecting notes).

  • — Smetana, 1982: 75 (characters; habitus; revision of and key to species of North America north of Mexico).

  • — Bordoni, 1982: 218 (characters; habitus; key to species of Italy).

  • — Shibata, 1983: 70 (checklist of species of Japan).

  • — Frank, 1986a: 369 (checklist of species of Florida).

  • — Shibata, 1986a: 123 (supplemental list of species from Taiwan).

  • — Muona and Viramo, 1986: 14 (list of species of northeastern Finland).

  • — Tóth, 1989: 15 (characters).

  • — Angelini, 1991: 191 (list of species from high elevations of Calabria, Italy).

  • — Campbell and Davies, 1991: 123 (checklist of species; Alaska and Canada).

  • — Silfverberg, 1992: 18 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Welch, 1993: 228 (ovariole structure and development).

  • — Hernández, Outerelo, and Gamarra, 1994: 190 (checklist of species; Canary Islands).

  • — Ciceroni and Zanetti, 1995: 25 (list of species of Italy).

  • — M. Hansen, 1996: 108 (list of Danish species).

  • — Downie and Arnett, 1996: 398 (characters; key to species of Northeastern North America).

  • — Lundgren, 1998: 50 (list of species from Florida; records for other States listed for each species).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 74 (list of species collected in 1997; Denmark).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 391 (5 Nearctic species; notes; characters in key).

Pedinolinus Bernhauer, 1912h: 479 (subgenus of Nudobius; species included: africanus). Type species: Nudobius (Pedinolinus) africanus Bernhauer, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 299 (subgenus of Nudobius).

  • — Scheerpeltz, 1933: 1306 (subgenus of Nudobius).

  • — Scheerpeltz, 1956b: 13 (subgenus of Nudobius; characters; key to African species).

abessinus Bernhauer, 1915d: 134 (Nudobius; Type locality: Abessinien: Boroda).

  • — Bernhauer, 1931d: 582 (Nudobius; Abyssinia).

  • — Scheerpeltz, 1933: 1305 (Nudobius; catalog).

  • — Scheerpeltz, 1956b: 13 (Nudobius; characters; Abyssinia).

  • Distribution: Ethiopia.

africanus Bernhauer, 1912h: 479 (Nudobius; subgenus Pedinolinus; Type locality: Nördlich vom Albert-Eduard-See, Ruwenzori-Fuss).

  • — Bernhauer and Schubert, 1914: 299 (Nudobius; subgenus Pedinolinus, cited as Pedolinus; catalog).

  • — Scheerpeltz, 1956b: 14 (Nudobius; subgenus Pedinolinus; characters).

  • Distribution: Congo/Uganda.

apicipennis Sharp, 1889: 253 (Nudobius; Type locality: Miyanoshita; Kiga; Kuma Kuni; Sendai; Sapporo).

  • — Bernhauer and Schubert, 1914: 298 (Nudobius; catalog).

  • — Nakane, 1963a: 89 (Nudobius; characters; habitus photograph; Japan).

  • — Yuh, Paik, Kwon, and Lee, 1985: 253 (Nudobius; Korea).

  • — J. Li, 1993: 32 (Nudobius; China).

  • Distribution: Japan, Korea, China.

arizonicus Casey, 1906: 382 (Nudobius; Type locality: Arizona, Williams; [Note: See lectotype designation by Smetana, 1982]. Also cited from New Mexico: Las Vegas).

  • — Fall and Cockerell, 1907: 167 (Xantholinus; New Mexico).

  • — Bernhauer and Schubert, 1914: 298 (Nudobius; catalog).

  • — Smetana, 1982: 87 (Nudobius; lectotype designation: Williams, Arizona; characters; Arizona; Colorado; New Mexico; South Dakota; Utah).

  • Distribution: USA.

phoenicis Casey, 1906: 382 (Nudobius; Type locality: Arizona, Phoenix; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 299 (Nudobius; catalog).

  • — Smetana, 1982: 87 (Nudobius; lectotype designation; synonym of arizonicus).

assimilis Kirby, 1837, see: cephalus Say, 1830.

bipustulatus Bernhauer, 1937: 297 (Nudobius; Type locality: Kenya: Ngong; Escarpment, Chania).

  • — Scheerpeltz, 1956a: 484 (Nudobius; Kenya; Uganda; French Congo; Belgian Congo).

  • — Scheerpeltz, 1956b: 12 (Nudobius; characters; Kenya; Uganda; French Congo; Belgian Congo).

  • Distribution: Kenya, Uganda, Congo, French Equatorial Africa.

brincki Scheerpeltz, 1974: 123 (Nudobius; Type locality: Cape Prov.: Langebaan).

  • Distribution: South Africa.

burgeoni Bernhauer, 1934g: 234 (Nudobius; Type locality: Kivu: Mts. Nyamukubi, 2600 m).

  • — Scheerpeltz, 1956b: 12 (Nudobius; characters; French Congo; Belgian Congo).

  • Distribution: Congo, French Equatorial Africa.

canariensis Israelson, 1969: 150 (Nudobius; Type locality: Canary Islands: La Palma, Barranco de los Hombres, about 1000 m).

  • — Assing and Wunderle, 1999: 437 (Nudobius; Canary Islands).

  • Distribution: Canary Islands.

palmensis Coiffait, 1973: 109 (Nudobius; subspecies of collaris; Type locality: environs de Roque del Faro, Ile de la Palma, Canaries).

  • — Assing, 1998e: 141 (Nudobius; synonym of canariensis).

palmensis Coiffait, 1976c: 55 (Nudobius; subspecies of collaris; [Note: Preoccupied; Probably the same as the subspecies of the same name and locality described earlier by Coiffait, 1973: 109 but published and described in 1976 without reference to the earlier name]; Type locality: Canaries: Palma, environs de Roque del Faro).

  • — Assing, 1998e: 141 (Nudobius; synonym of canaiensis).

cavicola Cameron, 1938e: 202 (Xantholinus; subgenus Typhlodes; Type locality: Kenya Colony: Grotte Campell, prairies alpines du mont Kenya, versant nord-ouest, alt. 3480 m).

  • — Paulian, 1938: 210 (Xantholinus; subgenus Typhlodes; larval description; Kenya).

  • — Paulian, 1941: 214 (Xantholinus; larval characters; Kenya).

  • — Jeannel and Jarrige, 1949: 373 (Nudobius; cave species; notes; Kenya).

  • — Bordoni and Oromi, 1998: 1157 (Nudobius; facultative cave species; Kenya).

  • Distribution: Kenya.

excelsus Bernhauer, 1939a: 78 (Xantholinus; Type locality: British-Ostafrika: M. Kenya, vers. Ouest, zone alpine, prairies alpines, reg. Senecio arbor., 4000–4100 m).

  • — Jeannel and Jarrige, 1949: 373 (Nudobius; synonym of cavicola).

celatus Sharp, 1885: 502 (Nudobius; Type locality: Mexico; [Note: See lectotype designation by Smetana, 1980a]. Originally cited from Mexico, San Andreas, Oaxaca; Guatemala, Balheu).

  • — Bernhauer and Schubert, 1914: 298 (Nudobius; catalog).

  • — Scheerpeltz, 1953: 607 (Nudobius; Guatemala; Mexico; Honduras; Costa Rica).

  • — Smetana, 1980a: 160 (Nudobius; lectotype designation: Mexico; characters; Mexico).

  • Distribution: Mexico, Guatemala, Honduras, Costa Rica.

cephalus Say, 1830: 39 [= 1834: 452] (Xantholinus; Type locality: Southern States [Note: See neotype designation by Smetana, 1982]. Originally cited from Virginia: Chinquotege Island).

  • — LeConte, 1850: 219 (Xantholinus; Lake Superior).

  • — Provancher, 1877: 251 (Xantholinus; characters; Quebec).

  • — LeConte, 1880: 172 (Xantholinus; California).

  • — Casey, 1906: 381 (Nudobius; characters).

  • — Fall and Cockerell, 1907: 166 (Xantholinus; New Mexico).

  • — Blatchley, 1910: 395 (Xantholinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 298 (Nudobius; catalog).

  • — Notman, 1919a: 95 (Xantholinus; Ontario).

  • — Scheerpeltz, 1933: 1305 (Nudobius; catalog).

  • — Paulian, 1941: 212 (Nudobius; cited as cephalotes; larval characters; Arizona).

  • — Hatch, 1957: 237 (Nudobius; cited as cephalicus; characters; British Columbia; Idaho; Washington; Oregon).

  • — Legner and Moore, 1977: 176 (Nudobius; found under bark).

  • — Smetana, 1982: 77 (Nudobius; neotype designation: [Note: Locality not cited, but the specimen chosen has an orange disc as one of the labels. Specimens in the LeConte collection with an orange disc are from the “southern States” (Gulf States) according to Smetana (personal communication)]; characters; Alberta; British Columbia; Manitoba; New Brunswick; Newfoundland; Northwest Territories; Nova Scotia; Ontario; Quebec; Saskatchewan; Alaska; Arizona; Arkansas; Colorado; Illinois; Idaho; Indiana; Maine; Maryland; Massachusetts; Michigan; Missouri; Montana; Nebraska; New Hampshire; New Jersey; New Mexico; New York; North Carolina; Ohio; Pennsylvania; Rhode Island; South Dakota; Vermont; Virginia; Washington; West Virginia; Wisconsin; Wyoming).

  • — Morgan and Morgan, 1985: 1829 (Nudobius; middle Holocene fossil; Michigan).

  • — Dearborn and Donahue, 1994: 14 (Nudobius; Maine).

  • — Downie and Arnett, 1996: 398 (Nudobius; characters; USA).

  • Distribution: Canada, USA.

assimilis Kirby, 1837: 88 (Gyrohypnus; Type locality: America borealis: lat. 54°).

  • — LeConte, 1859: 568 (synonym of cephalus Say).

  • — Erichson, 1840: 924 (Xantholinus; synonym of cephalus).

  • — Horn, 1876: 151 (Xantholinus; synonym of cephalus).

  • — Casey, 1906: 381 (Nudobius; synonym of cephalus).

  • — Bernhauer and Schubert, 1914: 298 (Nudobius; synonym of cephalus).

  • — Smetana, 1982: 77 (Nudobius; synonym of cephalus).

flavipennis Nordmann, 1837: 121 (Gyrohypnus; Type locality: America borealis).

  • — Erichson, 1839b: 326 (Xantholinus; synonym of consentaneus).

  • — Casey, 1906: 381 (Nudobius; synonym of cephalus).

  • — Bernhauer and Schubert, 1914: 298 (Nudobius; synonym of cephalus).

  • — Smetana, 1982: 77 (Nudobius; synonym of cephalus).

consentaneus Nordmann, 1837: 122 (Gyrohypnus; Type locality: America borealis).

  • — LeConte, 1859: 568 (synonym of cephalus Say).

  • — Erichson, 1839b: 326 (Xantholinus; characters; America septentrionalis).

  • — Erichson, 1840: 924 (Xantholinus; synonym of cephalus).

  • — Casey, 1906: 381 (Nudobius; synonym of cephalus).

  • — Bernhauer and Schubert, 1914: 298 (Nudobius; synonym of cephalus).

  • — Smetana, 1982: 77 (Nudobius; synonym of cephalus).

palliatus Melsheimer, 1844: 34 (Xantholinus; Type locality: Pennsylvania; [Note: See lectotype designation by Smetana, 1982]).

  • — Casey, 1906: 381 (Nudobius; synonym of cephalus).

  • — Bernhauer and Schubert, 1914: 298 (Nudobius; synonym of cephalus).

  • — Smetana, 1982: 77 (Nudobius; lectotype designation; synonym of cephalus).

kiesenwetteri Sachse, 1852: 123 (Xantholinus; Type locality: Georgia).

  • — LeConte, 1859: 568 (synonym of cephalus Say).

  • — Casey, 1906: 381 (Nudobius; synonym of cephalus).

  • — Bernhauer and Schubert, 1914: 298 (Nudobius; synonym of cephalus).

  • — Smetana, 1982: 77 (Nudobius; synonym of cephalus but probably a different species).

nubipennis Casey, 1906: 381 (Nudobius; Type locality: southwestern Utah; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 299 (Nudobius; catalog).

  • — Smetana, 1982: 77 (Nudobius; lectotype designation; synonym of cephalus).

gilvipennis Casey, 1906: 393 (Gyrohypnus; Type locality: New York, Hudson Valley).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; catalog).

  • — Smetana, 1982: 77 (Nudobius; synonym of cephalus).

cinctipennis Scheerpeltz, 1956b: 14, 18 (Nudobius; subgenus Pedinolinus; Type locality listed in Scheerpeltz, 1956a: Belgisch-Kongo: Hembe Bitale, also cited from French Congo).

  • — Scheerpeltz, 1956a: 484 (Nudobius; listed type locality).

  • Distribution: Congo, French Equatorial Africa.

circularis Smetana, 1980a: 162 (Nudobius; Type locality: Mexico: 10 mi. W El Salto, Dgo., 9000 ft.).

  • Distribution: Mexico.

collaris Erichson, 1839a: 424 (Xantholinus; Type locality: Mark Brandenburg).

  • — Erichson, 1839b: 324 (Xantholinus; characters; Germany; Sardinia).

  • — Redtenbacher, 1849: 824 (Xantholinus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 501 (Xantholinus; characters; habitat; France).

  • — Kraatz, 1857d: 644 (Xantholinus; characters; Germany).

  • — Redtenbacher, 1857: 180 (Xantholinus; characters; Austria).

  • — Fauvel, 1873b: 97 [= 1873a: 383] (Xantholinus; characters; France; Germany; Austria; Italy; Corsica; Sardinia; Algeria; Cyprus; Russia).

  • — Redtenbacher, 1874: 219 (Xantholinus; characters; Austria).

  • — Seidlitz, 1875: 282 (Xantholinus; characters; Baltic region).

  • — Mulsant and Rey, 1877a: 302 (Nudobius; characters; France).

  • — Fauvel, 1878: 116 [= 1878a: 36] (Xantholinus; Algeria).

  • — Fauvel, 1886: 48 [= 1886a: 40] (Xantholinus; Tunisia; Algeria).

  • — Ganglbauer, 1895: 477 (Nudobius; characters; middle Europe; Mediterranean region).

  • — Fauvel, 1902b: 100 (Xantholinus; Algeria; Tunisia).

  • — Sainte-Claire Deville, 1906: 93 (Nudobius; Corsica).

  • — Reitter, 1908: 114 (Xantholinus; subgenus Nudobius; characters; Austria; France; Mediterranean region).

  • — Bernhauer and Schubert, 1914: 298 (Nudobius; catalog).

  • — Portevin, 1929: 371 (Xantholinus; characters; France).

  • — Scheerpeltz, 1933: 1305 (Nudobius; catalog).

  • — Coiffait, 1956a: 56 (Nudobius; characters in key; France).

  • — Lohse, 1964: 161 (Nudobius; characters; central Europe).

  • — Horion, 1965: 91 (Nudobius; Portugal; Spain; Corsica; Sardinia; Sicily; Italy; Greece; Cyprus; France; Austria).

  • — Coiffait, 1972: 207 (Nudobius; characters; references; France; Europe).

  • — Legner and Moore, 1977: 176 (Nudobius; found under bark).

  • — Bordoni, 1982: 225 (Nudobius; characters; Italy).

  • — Lucht, 1987: 102 (Nudobius; checklist; Austria).

  • — Tóth, 1989: 15 (Nudobius; characters).

  • — Ciceroni, 1993: 72 (Nudobius; Italy).

  • — Sparacio, 1995: 157 (Nudobius; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 25 (Nudobius; Italy).

  • Distribution: Tunisia, Algeria, Canary Islands, Europe, Russia.

godeti Laporte, 1840: 180 (Gyrohypnus; Type locality: Volhynie).

  • — Fauvel, 1874: 338 [= 1874b: 44] (Xantholinus; synonym of collaris).

  • — Ganglbauer, 1895: 477 (Nudobius; synonym of collaris).

  • — Reitter, 1908: 114 (Xantholinus; synonym of collaris).

  • — Bernhauer and Schubert, 1914: 298 (Nudobius; synonym of collaris).

ruficollis P. Lucas, 1846: 107 (Xantholinus; Type locality: Algeria: lac Tonga aux environs du cercle de Lacalle).

  • — Fauvel, 1869: 490 (Xantholinus; synonym of ruficollis).

  • — Fauvel, 1869a: 151 (Xantholinus; synonym of collaris).

  • — Fauvel, 1873b: 97 [= 1873a: 383] (Xantholinus; synonym of collaris).

  • — Mulsant and Rey, 1877a: 302 (Nudobius; synonym of collaris).

  • — Ganglbauer, 1895: 477 (Nudobius; synonym of collaris).

  • — Reitter, 1908: 114 (Xantholinus; synonym of collaris).

  • — Bernhauer and Schubert, 1914: 298 (Nudobius; synonym of collaris).

coloriventris Bernhauer, 1915d: 133 (Nudobius; Type locality: Abessinien: Boroda).

  • — Bernhauer, 1931d: 582 (Nudobius; Abyssinia).

  • — Scheerpeltz, 1933: 1306 (Nudobius; catalog).

  • — Scheerpeltz, 1956b: 12 (Nudobius; characters; Abyssinia).

  • Distribution: Ethiopia.

concinnus Tottenham, 1956: 251 (Nudobius; Type locality: Ruanda: terr. Kibuye, Yanina, 2300 m).

  • Distribution: Rwanda.

congoensis Scheerpeltz, 1956b: 13, 16 (Nudobius; Type locality listed in Scheerpeltz, 1956a: Belgisch-Kongo: Burgarama).

  • — Scheerpeltz, 1956a: 484 (Nudobius; listed type locality).

  • Distribution: Congo.

consentaneus Nordmann, 1837, see: cephalus Say, 1830.

corticalis Casey, 1906, see: pugetanus Casey, 1906.

cypriacus Coiffait, 1956a: 57 (Nudobius; Type locality: Chypre).

  • — Coiffait, 1970e: 62 (Nudobius; Cyprus).

  • — Coiffait, 1972: 209 (Nudobius; characters; Cyprus; Croatia).

  • — Bordoni, 1975e: 140 (Nudobius; notes; Greece).

  • — Bordoni and Bordoni, 1989: 745 (Nubodius; Greece).

  • Distribution: Cyprus, Greece, Croatia.

debilis Casey, 1906, see: pugetanus Casey, 1906.

elegantulus Casey, 1906, see: luridipennis Casey, 1906.

excellens Tikhomirova, 1973, see: lentus Gravenhorst, 1806.

excelsus Bernhauer, 1939, see: cavicola Cameron, 1938.

flavipennis Nordmann, 1837, see: cephalus Say, 1830.

formosanus Shibata, 1973a: 123 (Nudobius; Type locality: Taiwan: Meifeng near Wushe, Nantou Hsien).

  • Distribution: Taiwan.

gilvipennis Casey, 1906, see: cephalus Say, 1830.

godeti Laporte, 1840, see: collaris Erichson, 1839.

kiesenwetteri Sachse, 1852, see: cephalus Say, 1830.

lentus Gravenhorst, 1806: 101 (Staphylinus; Type locality: Parisiae).

  • — Gyllenhal, 1810: 354 (Staphylinus; characters; Sweden).

  • — Zetterstedt, 1828: 81 (Xantholinus; characters; Lapland).

  • — C. Sahlberg, 1830: 333 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 33 (Gyrohypnus; Finland; Sweden).

  • — Mannerheim, 1831: 447 (Gyrohypnus; Finland; Sweden).

  • — Nordmann, 1837: 117 (Gyrohypnus; Europe).

  • — Zetterstedt, 1838: 66 (Xantholinus; characters; habitat; Lapland).

  • — Erichson, 1839a: 426 (Xantholinus; cited glaber Gravenhorst, 1806: 100, variation 1 as a synonym of lentus characters; Germany).

  • — Heer, 1839: 245 (Xantholinus; characters; Switzerland).

  • — Erichson, 1839b: 325 (Xantholinus; characters; Germany; Sweden).

  • — Stephens, 1839: 403 (Xantholinus; characters; England).

  • — Redtenbacher, 1849: 824 (Xantholinus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 501 (Xantholinus; characters; France).

  • — Kraatz, 1857d: 644 (Xantholinus; characters; Germany).

  • — Redtenbacher, 1857: 181 (Xantholinus; characters; Austria).

  • — Peyron, 1858: 422 (Xantholinus; Turkey).

  • — Thomson, 1860: 188 (Nudobius; characters; Scandinavia).

  • — Crotch, 1867: 451 (Xantholinus; Britain).

  • — Fauvel, 1873b: 98 [= 1873a: 384] (Xantholinus; characters; France; Italy).

  • — Redtenbacher, 1874: 218 (Philonthus; characters; Austria).

  • — Redtenbacher, 1874: 219 (Philonthus; [Note: Misidentification: Bernhauer and Schubert, 1914: 299, cited rufipennis Redtenbacher as a synonym of lentus; Redtenbacher did not describe the species as new, he attributed it to Erichson]; characters; Austria).

  • — Seidlitz, 1875: 282 (Xantholinus; characters; Baltic region).

  • — J. Sahlberg, 1876: 33 (Nudobius; Finland).

  • — Mulsant and Rey, 1877a: 305 (Nudobius; characters; France).

  • — Fowler, 1888: 292 (Nudobius; characters; Britain).

  • — Ganglbauer, 1895: 476 (Nudobius; characters; north and middle Europe; Siberia).

  • — Reitter, 1908: 114 (Xantholinus; subgenus Nudobius; characters; Europe).

  • — Reitter, 1909: 136 (Xantholinus; subgenus Nudobius; characters; Germany).

  • — Bernhauer and Schubert, 1914: 299 (Nudobius; catalog).

  • — Johansen, 1914: 432 (Xantholinus; subgenus Nudobius; characters; Denmark).

  • — Portevin, 1929: 371 (Xantholinus; characters; France).

  • — Scheerpeltz, 1933: 1306 (Nudobius; catalog).

  • — Paulian, 1941: 211 (Nudobius; larval characters; Germany).

  • — V. Hansen, 1952: 14 (Xantholinus; subgenus Nudobius; characters; Denmark).

  • — Coiffait, 1956a: 56 (Nudobius; characters in key; France).

  • — Smetana, 1958: 94 (Nudobius; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 45 (Nudobius; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 161 (Nudobius; characters; central Europe).

  • — Horion, 1965: 90 (Nudobius; Turkey; Siberia; Russia; Finland; Sweden; Norway; Scotland; Denmark; Holland; Belgium; France; Poland; Pyrenees; Italy; Bosnia; Austria; Germany).

  • — Coiffait, 1965e: 119 (Nudobius; Turkey).

  • — Coiffait, 1966: 21 (Nudobius; Turkey).

  • — Pototskaia, 1967: 47 (Nudobius; larval characters).

  • — Szujecki, 1968a: 725 (Nudobius; Poland).

  • — Coiffait, 1972: 206 (Nudobius; characters; references; France; Europe; Scandinavia; Britain; Siberia; Asia Minor).

  • — Allen, 1974: 161 (Nudobius; collecting notes; England).

  • — Shibata, 1974: 33 (Nudobius; Japan).

  • — Pototskaia, 1976: 172 (Nudobius; larval characters).

  • — Legner and Moore, 1977: 176 (Nudobius; found under bark).

  • — Pope, 1977: 28 (Nudobius; Britain).

  • — Raichev, 1979: 81 (Nudobius; Bulgaria).

  • — Bordoni, 1982: 222 (Nudobius; characters; Italy).

  • — Smetana, 1982a: 310 (Nudobius; characters; Kuril Islands).

  • — Segers, 1986: 40 (Nudobius; checklist of species; Belgium).

  • — Lucht, 1987: 102 (Nudobius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Tóth, 1989: 15 (Nudobius; characters).

  • — Janák, 1993a: 47 (Nudobius; Czech Republic; Slovakia).

  • — Welch, 1993: 228 (Nudobius; ovariole number and ovary structure).

  • — J. Li, 1993: 32 (Nudobius; China).

  • — Zheng, 1994c: 472, 473 (Nudobius; China).

  • — Ciceroni and Zanetti, 1995: 25 (Nudobius; Italy).

  • — Sterrenburg and Schülke, 1997: 16 (Nudobius; Netherlands).

  • — Cibul ¸skis, 1998: 66 (Nudobius; Latvia).

  • — Fowles, Alexander, and Key, 1999: 129 (Nudobius; saproxylic species; status as endangered or threatened species; Britain).

  • — Ryabukhin, 1999: 59 (Nudobius; catalog; habitat; distribution; northeastern Russia).

  • Distribution: Europe, Russia, Turkey, China, Japan.

wingelmuelleri Bernhauer, 1899: 24 (Xantholinus; Type locality: Negoi in den siebenbürgischen Alpen; “Stavnja-Th.b. Saraj”; Umgebung von Kronstadt; Vorarlberg).

  • — Bernhauer, 1900: 47 (Xantholinus; notes).

  • — Reitter, 1908: 114 (Xantholinus; synonym of lentus).

  • — Bernhauer and Schubert, 1914: 299 (Nudobius; synonym of lentus).

  • — Coiffait, 1972: 206 (Nudobius; synonym of lentus).

  • — Bordoni, 1982: 222 (Nudobius; synonym of lentus).

picipennis Everts, 1920a: 288 (Nudobius; aberration of picipennis; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Loosduinen).

  • — Scheerpeltz, 1933: 1306 (Nudobius; aberration of lentus).

  • — Coiffait, 1972: 206 (Nudobius; variety of lentus; characters).

  • — Bordoni, 1982: 222 (Nudobius; synonym of lentus).

excellens Tikhomirova, 1973: 162 (Nudobius; Type locality: Kedrovaia Pad).

  • — Smetana, 1982a: 312 (Nudobius; synonym of lentus).

limbalis Casey, 1906, see: pugetanus Casey, 1906.

luridipennis Casey, 1906: 383 (Nudobius; Type locality: Alabama: Citronelle; [Note: See lectotype designation by Smetana, 1982]. Also cited from Texas: Houston; West Virginia).

  • — Bernhauer and Schubert, 1914: 299 (Nudobius; catalog).

  • — Smetana, 1982: 89 (Nudobius; lectotype designation: Citronelle, Alabama; characters; Alabama; Arkansas; Florida; Georgia; Maryland; Mississippi; New Jersey; New York; North Carolina; Pennsylvania; South Carolina; Tennessee; Texas; Virginia; West Virginia).

  • — Smetana, 1990c: 84 (Nudobius; Texas).

  • — Downie and Arnett, 1996: 398 (Nudobius; characters; USA).

  • Distribution: USA.

elegantulus Casey, 1906: 384 (Nudobius; Type locality: West Virginia; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 298 (Nudobius; catalog).

  • — Smetana, 1982: 89 (Nudobius; lectotype designation: Kanawha Sta. W. Va. [West Virginia]; synonym of luridipennis).

magnus Cameron, 1951c: 399 (Nudobius; Type locality: Kenya Colony: N.W. Mau, 8–10,000 feet).

  • Distribution: Kenya.

meruensis Cameron, 1951c: 398 (Nudobius; Type locality: Kenya Colony: Meru).

  • Distribution: Kenya.

morosus Tottenham, 1939c: 223 (Nudobius; Type locality: S. Rhodesia: Vumba Mts.).

  • Distribution: Zimbabwe.

nigriventris Zheng, 1994c: 471, 473 (Nudobius; Type locality: China: Sichuan Province, Nanjiang forest zone, Mt. Daiba, 32°23′N, 107°E, 1300–1600 m).

  • — Bordoni, 2000: 121 (Nudobius; characters; China).

  • Distribution: China.

nigrocyaneus Chapman, 1939: 62 (Nudobius; subgenus Pedinolinus, cited as Pedolinus; Type locality: Kenya Colony: Massif du Marakwet, alt. 2500 m; forêts inférieures du Kenya, près de la maison forestière, alt. 2400 m).

  • — Scheerpeltz, 1956b: 15 (Nudobius; subgenus Pedinolinus; characters; Kenya).

  • Distribution: Kenya.

nubipennis Casey, 1906, see: cephalus Say, 1830.

opacinus Tottenham, 1956: 249 (Nudobius; Type locality: Ruanda: Rutovu, forêt du Rugege, 2350 m).

  • Distribution: Rwanda.

palliatus Melsheimer, 1844, see: cephalus Say, 1830.

palmensis Coiffait, 1973, see: canariensis Israelson, 1969.

palmensis Coiffait, 1976, see: canariensis Israelson, 1969.

phoenicis Casey, 1906, see: arizonicus Casey, 1906.

picipennis Everts, 1920 [Note: Nomen nudum], see: lentus Gravenhorst, 1806.

pictipennis Fauvel, 1904d: 289 (Xantholinus; subgenus Nudobius; Type locality: Usambara: Derema. Natal).

  • — Bernhauer and Schubert, 1914: 299 (Nudobius; catalog).

  • — Scheerpeltz, 1956b: 14 (Nudobius; subgenus Pedinolinus; characters; Kenya; Uganda; Tanganyika; East Africa; Mozambique; Natal).

  • Distribution: Uganda, Kenya, Tanzania, Mozambique, South Africa.

pleuralis Sharp, 1874: 53 (Xantholinus; Type locality: Nagasaki; [Note: See lectotype designation by Bordoni, 1997]).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Cameron, 1949b: 467 (Xantholinus; Japan).

  • — J. Li, 1993: 32 (Xantholinus; cited as pleuvalis; China).

  • — Bordoni, 1997: 169 (Nudobius; lectotype designation; characters; Japan).

  • Distribution: Japan, China.

praecellens Bernhauer, 1915d: 134 (Nudobius; subgenus Pedinolinus; Type locality: Deutsch-Ostafrika: West-Kilimandscharo, 1500–1700 m).

  • — Scheerpeltz, 1933: 1306 (Nudobius; subgenus Pedinolinus; catalog).

  • — Scheerpeltz, 1956b: 15 (Nudobius; subgenus Pedinolinus; characters).

  • Distribution: Tanzania.

proximus Bernhauer, 1915d: 134 (Nudobius; Type locality: Abessinien: Boroda).

  • — Scheerpeltz, 1933: 1306 (Nudobius; catalog).

  • — Scheerpeltz, 1956b: 13 (Nudobius; characters; Abyssinia).

  • Distribution: Ethiopia.

pugetanus Casey, 1906: 382 (Nudobius; Type locality: Washington Territory; [Note: See lectotype designation by Smetana, 1982]. Originally cited from Washington State; Oregon, The Dalles).

  • — Bernhauer and Schubert, 1914: 299 (Nudobius; catalog).

  • — Struble, 1930: 127 (Nudobius; characters of eggs, larvae, pupa, adults; life history and habits; Washington, Oregon, California).

  • — Scheerpeltz, 1933: 1306 (Nudobius; catalog).

  • — Smetana, 1982: 83 (Nudobius; lectotype designation: W. T. [Washington Territory]; characters; British Columbia; California; Idaho; Nevada; Oregon; Washington).

  • Distribution: Canada, USA.

corticalis Casey, 1906: 383 (Nudobius; Type locality: California, Sta. Clara; [Note: See lectotype designation by Smetana, 1982]. Also cited from California: Sta. Cruz).

  • — Bernhauer and Schubert, 1914: 298 (Nudobius; catalog).

  • — Smetana, 1982: 83 (Nudobius; lectotype designation: Sta. Clara, California; synonym of pugetanus).

debilis Casey, 1906: 383 (Nudobius; Type locality: California: Siskiyou Co.; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 298 (Nudobius; catalog).

  • — Smetana, 1982: 83 (Nudobius; lectotype designation; synonym of pugetanus).

limbalis Casey, 1906: 383 (Nudobius; Type locality: California, Truckee, 5800 ft.; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 299 (Nudobius; catalog).

  • — Smetana, 1982: 83 (Nudobius; lectotype designation; synonym of pugetanus).

pulcher Tottenham, 1956: 247 (Nudobius; Type locality: Ruanda: Rutovu, forêt du Rugege, 2350 m).

  • Distribution: Rwanda.

quadriceps Cameron, 1929: 60 (Nudobius; Type locality: Kivu: Kibati).

  • — Scheerpeltz, 1933: 1306 (Nudobius; catalog).

  • — Scheerpeltz, 1956a: 484 (Nudobius; subgenus Pedinolinus; characters; French Congo; Belgian Congo).

  • Distribution: Congo, French Equatorial Africa.

quadrimaculatus Eichelbaum, 1910: 86 (Xantholinus; variety of pictipennis; Type locality: Kilimandjaro-Meru: Steppe an den Natronseen).

  • — Bernhauer and Schubert, 1914: 299 (Nudobius; variety of pictipennis).

  • — Scheerpeltz, 1956b: 14 (Nudobius; subgenus Pedinolinus; cited as valid species; characters).

  • Distribution: Tanzania.

rudebecki Scheerpeltz, 1974: 125 (Nudobius; Type locality: Cape Prov.: Tzitzikama Forest, Stormsrivierpiek).

  • Distribution: South Africa.

ruficollis P. Lucas, 1846, see: collaris Erichson, 1839.

schedli Scheerpeltz, 1956b: 12, 15 (Nudobius; Type locality cited in Scheerpeltz, 1956a: Belgisch-Kongo: Hembe Bitale, also cited from French Congo).

  • Distribution: Congo, French Equatorial Africa.

schlueteri Scheerpeltz, 1956b: 15 (Nudobius; subgenus Pedinolinus; Type locality: Insel Fernando Poo).

  • Distribution: Equatorial Guinea.

singularis Smetana, 1982: 91 (Nudobius; Type locality: Michigan: Port Huron. Also cited from Indiana).

  • — Downie and Arnett, 1996: 398 (Nudobius; characters; USA).

  • Distribution: USA.

subviridipennis Bernhauer, 1912f: 203 (Pedinolinus; Type locality: Usambara).

  • — Bernhauer and Schubert, 1914: 299 (Nudobius; subgenus Pedinolinus, cited as Pedolinus; catalog).

  • — Scheerpeltz, 1956b: 15 (Nudobius; subgenus Pedinolinus; characters).

  • Distribution: Tanzania.

tuniseus Coiffait, 1956a: 56 (Nudobius; Type locality: Tunisie: Sbeitla).

  • — Coiffait, 1972: 207 (Nudobius; characters; Tunisia).

  • Distribution: Tunisia.

turcicus Coiffait, 1978d: 166 (Nudobius; Type locality: Abant, Bolu).

  • Distribution: Turkey.

umbratus Motschulsky, 1860: 564 (Xantholinus; Type locality: Armenie aux environs d'Akhelzik).

  • — Hochhuth, 1862: 67 (Xantholinus; characters; Armenia; Daghestan).

  • — Reitter, 1908: 114 (Xantholinus; subgenus Nudobius; characters; Caucasus).

  • — Bernhauer and Schubert, 1914: 299 (Nudobius; catalog).

  • — Scheerpeltz, 1933: 1306 (Nudobius; catalog).

  • — Coiffait, 1972: 208 (Nudobius; characters; references; Armenia).

  • — Pototskaia, 1976: 173 (Nudobius; larval characters).

  • Distribution: Armenia, Russia.

wingelmuelleri Bernhauer, 1899, see: lentus Gravenhorst, 1806.

Oculolabrus [1 species; Oriental Region]

Oculolabrus Steel, 1946: 107 (species included: sumatrensis). Type species: Oculolabrus sumatrensis Steel, fixed by original designation and monotypy.

  • — Blackwelder, 1952: 267 (type species: sumatrensis).

sumatrensis Steel, 1946: 108 (Oculolabrus; Type locality: Sumatra).

  • Distribution: Indonesia.

Oxybleptes [5 species; Nearctic Region]

Oxybleptes Smetana, 1982: 253 (species included: davisi, kiteleyi, hatchi, pusio). Type species: Gyrohypnus davisi Notman, fixed by original designation.

  • — Downie and Arnett, 1996: 401 (characters; key to species of Northeastern North America).

  • — Lundgren, 1998: 50 (list of species from Florida; records for other States listed for each species).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 391 (5 Nearctic species; notes; characters in key).

davisi Notman, 1924a: 72 (Gyrohypnus; subgenus Hypnogyra; Type locality: Richmond, Staten Island, New York).

  • — Scheerpeltz, 1933: 1309 (Xantholinus; catalog).

  • — Smetana, 1982: 254 (Oxybleptes; characters; District of Columbia; New Jersey; New York).

  • — Downie and Arnett, 1996: 401 (Oxybleptes; characters; USA).

  • Distribution: USA.

davisi Hatch, 1957, see: hatchi Smetana, 1982.

hatchi Smetana, 1982: 259 (Oxybleptes; replacement name for davisi Hatch).

  • Distribution: USA.

davisi Hatch, 1957: 234 (Megalinus; [preoccupied]; Type locality: Oregon: West Salem).

  • — Smetana, 1982: 259 (Oxybleptes; synonym of hatchi).

kiteleyi Smetana, 1982: 256 (Oxybleptes; Type locality: Quebec: Montreal. Also cited from New Brunswick, Ontario, Quebec, Maine, New York, and North Carolina).

  • — Campbell and Davies, 1991: 124 (Oxybleptes; checklist of species; Canada).

  • — Downie and Arnett, 1996: 403 (Oxybleptes; characters; Canada; USA).

  • Distribution: Canada, USA.

meridionalis Smetana, 1988a: 556 (Oxybleptes; Type locality: Florida: Indian River Co.: Vero Beach).

  • Distribution: USA.

pusio Smetana, 1982: 260 (Oxybleptes; Type locality: North Carolina: Bryson City).

  • — Smetana, 1990c: 86 (Oxybleptes; Oklahoma).

  • Distribution: USA.

Pachycorynus [38 species; Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions]

Pachycorynus Motschulsky, 1858a: 204 (species included: dimidiatus). Type species: Pachycorynus dimidiatus Motschulsky, fixed by monotypy.

  • — Gemminger and Harold, 1868: 595 (catalog).

  • — Fauvel, 1878f: 239 (characters; notes).

  • — Fauvel, 1878g: 535 (Pachycorynus; notes).

  • — Duvivier, 1883: 152 (catalog).

  • — Bernhauer and Schubert, 1914: 295 (world catalog; 12 species).

  • — Cameron, 1925: 50 (catalog of Indian species).

  • — Cameron, 1932: 14 (characters; key to species of British India).

  • — Scheerpeltz, 1933: 1304 (world catalog supplement).

  • — Blackwelder, 1952: 284 (type species: dimidiatus).

  • — Coiffait, 1968: 128, 149 (characters; cited as Pachycorynus and Pachycorinus; key to species of Gabon; notes).

  • — Hammond, 1984: 206 (checklist of species of Borneo).

  • — Shibata, 1986a: 122 (supplemental list of species from Taiwan).

  • — Lecoq, 1990: 185 (key to species of Mascarene Islands).

Holisomorphus Kraatz, 1859: 100 (species included: ceylanensis). Type species: Holisomorphus ceylanensis Kraatz, fixed by monotypy.

  • — Gemminger and Harold, 1868: 595 (synonym of Pachycorynus).

  • — Fauvel, 1878f: 239 (synonym of Pachycorynus).

  • — Fauvel, 1878g: 535 (synonym of Pachycorynus).

  • — Bernhauer and Schubert, 1914: 295 (synonym of Pachycorynus).

  • — Cameron, 1932: 14 (synonym of Pachycorynus).

  • — Blackwelder, 1952: 187 (synonym of Pachycorynus; type species: ceylanensis).

analis Fauvel, 1878f: 240 (Pachycorynus; Type locality: Nouvelle-Guinée, Salvatti).

  • — Bernhauer and Schubert, 1914: 295 (Pachycorynus; catalog).

  • Distribution: New Guinea.

bakeri Bernhauer, 1916h: 421 (Pachycorynus, cited as Pachycorinus; Type locality: Mt. Banahao, P.I.).

  • — Scheerpeltz, 1933: 1304 (Pachycorynus; catalog).

  • Distribution: Philippines.

borneensis Cameron, 1928g: 442 (Pachycorynus; Type locality: Borneo: Mt. Murud, 6000 feet).

  • — Scheerpeltz, 1933: 1304 (Pachycorynus; catalog).

  • Distribution: Borneo.

caledonicus Fauvel, 1878g: 536 (Pachycorynus; Type locality: Nouvelle-Calédonie, Ile des Pins).

  • — Fauvel, 1889b: 259 (Pachynorus; New Caledonia).

  • — Fauvel, 1903d: 266 (Pachycorynus; New Hebrides; Iles des Pins).

  • — Bernhauer and Schubert, 1914: 295 (Pachycorynus; catalog).

  • Distribution: New Caledonia, Vanuatu.

ceylanensis Kraatz, 1859, see: dimidiatus Motschulsky, 1858.

cinctus Walker, 1858, see: dimidiatus Motschulsky, 1858.

caecus Broun, 1893, see: fulvipes Broun, 1880.

conradsi Bernhauer, 1937b: 609 (Pachycorynus, cited as Pachycorinus; Type locality: Ukerewe Insel im Victoria Nyanza, Muruntuguru).

  • Distribution: Tanzania.

delicatulus Cameron, 1927c: 263 (Pachycorynus; Type locality: Fiji: Ovalau, Teveuni, Moturiki, Viti Levu).

  • — Scheerpeltz, 1933: 1304 (Pachycorynus; catalog).

  • Distribution: Fiji.

discedens Sharp, 1880, see: Holocorynus.

dilaticeps Cameron, 1928g: 442 (Pachycorynus; Type locality: Borneo: Mt. Murud, 6000 feet).

  • — Scheerpeltz, 1933: 1304 (Pachycorynus; catalog).

  • — Cameron, 1933b: 344 (Pachycorynus British North Borneo).

  • — Shibata, 1973a: 122 (Pachycorynus; Taiwan).

  • Distribution: Borneo, Taiwan.

dimidiatus Motschulsky, 1858a: 205 (Pachycorynus, cited as Pachycorinus; Type locality: Indes orientales).

  • — Fauvel, 1878f: 239 (Pachycorynus; characters; Aru Islands; Java; Sri Lanka; East Indies).

  • — Fauvel, 1879: 92 (Pachycorynus; Dorei; Gilolo; Pulo Pinang).

  • — Bernhauer, 1902b: 32 (Pachycorynus, cited as Pachycorinus; Ceylon).

  • — Fauvel, 1904a: 56 (Pachycorynus, cited as Pachycorinus; Nilghiris; Indo-Malaysian region; Gilolo; Arou; New Guinea).

  • — Bernhauer and Schubert, 1914: 295 (Pachycorynus; catalog).

  • — Bernhauer, 1922a: 150 (Pachycorynus, cited as Pachycorinus; Indochina).

  • — Bernhauer, 1928: 14 (Pachycorynus, cited as Pachycorinus; Sumatra).

  • — Cameron, 1931: 359 (Pachycorynus; New Guinea).

  • — Cameron, 1932: 15 (Pachycorynus; characters; Ceylon, India).

  • — Scheerpeltz, 1933: 1304 (Pachycorynus; catalog).

  • — Cameron, 1933d: 385 (Pachycorynus; Sumatra).

  • — Cameron, 1937: 2 (Pachycorynus; Java).

  • — Shibata, 1973a: 122 (Pachycorynus; Taiwan).

  • — Ito, 1991: 12 (Pachycorynus; Okinawa).

  • Distribution: India, Sri Lanka, Malaysia, “Indochina”, Indonesia, New Guinea, Taiwan, Japan.

cinctus Walker, 1858: 205 (Xantholinus; Type locality: Ceylon).

  • — Fauvel, 1904a: 56 (Pachycorynus; synonym of dimidiatus).

  • — Bernhauer and Schubert, 1914: 295 (Pachycorynus; synonym of dimidiatus).

  • — Cameron, 1932: 15 (Pachycorynus; synonym of dimidiatus).

ceylanensis Kraatz, 1859: 101 (Holisomorphus; Type locality: Ceylan).

  • — Fauvel, 1878f: 239 (Pachycorynus; synonym of dimidiatus).

  • — Fauvel, 1904a: 56 (Pachycorynus; synonym of dimidiatus).

  • — Bernhauer and Schubert, 1914: 295 (Pachycorynus; synonym of dimidiatus).

  • — Cameron, 1932: 15 (Pachycorynus; synonym of dimidiatus).

dimorphus Fauvel, 1882, see: fulvipes Broun, 1880.

fragilis Cameron, 1936b: 44 (Pachycorynus; Type locality: Selangor: Fraser's Hill, 4200 feet).

  • Distribution: Malaysia.

fulvipes Broun, 1880: 106 (Metoponcus; Type locality: Parua).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; catalog).

  • — Cameron, 1945d: 782 (transferred from Metoponcus to Leptacinus).

  • — Steel, 1949a: 269 (Pachycorynus).

  • — Kuschel, 1990: 26 (Pachycorynus; endemic; flightless; habitat; New Zealand).

  • Distribution: New Zealand.

dimorphus Fauvel, 1882a: 92 (Pachycorynus, cited as Pachycorinus; Type locality: Nouvelle-Zélande, Middle Island, Greymouth).

  • — Bernhauer and Schubert, 1914: 295 (Pachycorynus; catalog).

  • — Steel, 1949a: 269 (Pachycorynus; synonym of fulvipes).

caecus Broun, 1893: 1406 (Xantholinus; Type locality: Hunua Range and Waitakerei).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; cited as coecus; catalog).

  • — Steel, 1949a: 269 (Pachycorynus; cited as coecus; synonym of fulvipes).

ganglbaueri Bernhauer, 1904: 234 (Pachycorynus, cited as Pachycorinus; Type locality: Madagaskar).

  • — Bernhauer and Schubert, 1914: 295 (Pachycorynus; catalog).

  • Distribution: Madagascar.

gracilis Cameron, 1928a: 36 (Pachycorynus; Type locality: Java).

  • — Scheerpeltz, 1933: 1304 (Pachycorynus; catalog).

  • — Cameron, 1933d: 385 (Pachycorynus; Sumatra).

  • Distribution: Indonesia.

sumatrensis Cameron, 1928a: 38 (Pachycorynus; variety of gracilis; Type locality: Sumatra: Mana Riang, Palembang).

  • — Scheerpeltz, 1933: 1304 (Pachycorynus; variety of gracilis).

  • — Cameron, 1933d: 385 (Pachycorynus; variety of gracilis; Sumatra).

grandiceps Bernhauer, 1934j: 172 (Pachycorynus, cited as Pachycorinus; Type locality: Ost-Java, Nongkodjadjar, 4000 f.).

  • Distribution: Indonesia.

infimus Lecoq, 1990: 184 (Pachycorynus; Type locality: Maurice: Macabe).

  • Distribution: Mascarene Islands.

inflatipes Fairmaire, 1893: 527 (Pachycorynus; Type locality: Mayotte).

  • — Bernhauer and Schubert, 1914: 295 (Pachycorynus; catalog).

  • Distribution: Comoros.

insularis Cameron, 1933f: 79 (Pachycorynus; Type locality: Marquesas Islands: Eiao: Vaituha Valley, altitude 200 feet).

  • — Coiffait, 1976: 216 (Pachycorynus, cited as Pachycorinus; Marquesas).

  • — Coiffait, 1980b: 472 (Pachycorynus, cited as Pachycorinus; Marquesas Islands).

  • Distribution: Marquesas Islands.

jarrigei Lecoq, 1990: 185 (Pachycorynus; Type locality: La Réunion: Grand Matarum, Cilaos).

  • Distribution: Mascarene Islands.

lateralis Cameron, 1928g: 443 (Pachycorynus, cited as Pachycorinus; Type locality: Borneo: Tutau River).

  • — Scheerpeltz, 1933: 1304 (Pachycorynus; catalog).

  • Distribution: Borneo.

longiceps Cameron, 1941b: 445 (Pachycorynus; Type locality: Philippines: Luzon: Imugan).

  • Distribution: Philippines.

luteipennis Coiffait, 1968: 149 (Pachycorynus, cited as Pachycorinus; Type locality: Gabon: Belinga).

  • Distribution: Gabon.

marginellus Fauvel, 1905d: 195 (Pachycorynus, cited as Pachycorinus; Type locality: Gabon: Libreville; Loango. Cameroun).

  • — Bernhauer and Schubert, 1914: 295 (Pachycorynus; catalog).

  • — Coiffait, 1968: 149 (Pachycorynus, cited as Pachycorinus; Gabon; Cameroon).

  • — Scheerpeltz, 1971c: 169 (Pachycorynus; Cameroon).

  • Distribution: Congo Republic, Gabon, Cameroon.

mauritianus Lecoq, 1990: 182 (Pachycorynus; Type locality: Maurice: Mont Cocotte).

  • Distribution: Mascarene Islands.

modestus Bernhauer, 1928: 14 (Pachycoryus, cited as Pachycorinus; Type locality: Sumatra: Dodok, 2000 m).

  • — Scheerpeltz, 1933: 1304 (Pachycorynus; catalog).

  • — Cameron, 1933d: 385 (Pachycorynus; Sumatra).

  • Distribution: Indonesia.

niger Cameron, 1932: 16 (Pachycorynus; Type locality: Darjeeling: Lopchu, atl. 5000 feet).

  • — Scheerpeltz, 1933: 1304 (Pachycorynus; catalog).

  • Distribution: India.

ophis Cameron, 1928a: 37 (Pachycorynus; Type locality: W. Preanger. Also cited from Sumatra).

  • — Scheerpeltz, 1933: 1304 (Pachycorynus; catalog).

  • — Cameron, 1933d: 385 (Pachycorynus; Sumatra).

  • — Cameron, 1937: 2 (Pachycorynus; Java).

  • Distribution: Indonesia.

pallidipennis Fauvel, 1903b: 166 (Pachycorynus, cited as Pachycorinus; Type locality: Bussira, Congo belge; Gabon: Libreville, Loango).

  • — Bernhauer and Schubert, 1914: 296 (Pachycorynus; catalog).

  • — Coiffait, 1968: 149, 150 (Pachycorynus, cited as Pachycorinus; characters; notes; Gabon).

  • Distribution: Congo, Congo Republic, Gabon.

pallidus Lea, 1927: 276 (Pachycorynus; Type locality: Fiji: Viti Levu).

  • — Scheerpeltz, 1933: 1304 (Pachycorynus; catalog).

  • Distribution: Fiji.

perakensis Cameron, 1950d: 16 (Pachycorynus; Type locality: Perak: Larut Hills, 3700 feet).

  • Distribution: Malaysia.

puncticollis Fauvel, 1905b: 169 (Pachycorynus, cited as Pachycorinus; Type locality: Diego-Suarez).

  • — Bernhauer and Schubert, 1914: 296 (Pachycorynus; catalog).

  • Distribution: Madagascar.

quadriceps Cameron, 1936d: 202 (Pachycorynus; Type locality: Mauritius: Les Mares).

  • — Lecoq, 1990: 182 (Pachycorynus; Mascarene Islands).

  • Distribution: Mascarene Islands.

rufotestaceus Cameron, 1927c: 263 (Pachycorynus; Type locality: Fiji: Viti Levu).

  • — Scheerpeltz, 1933: 1304 (Pachycorynus; catalog).

  • Distribution: Fiji.

scalabrei Fauvel, 1905b: 170 (Pachycorynus, cited as Pachycorinus; Type locality: Suberbieville; Tsarasaotra; Nord du Pays Androy).

  • — Bernhauer and Schubert, 1914: 296 (Pachycorynus; catalog).

  • Distribution: Madagascar.

selangorensis Cameron, 1936b: 43 (Pachycorynus; Type locality: Selangor: Bukit Kutu).

  • Distribution: Malaysia.

sparsipennis Fairmaire, 1893: 527 (Pachycorynus, cited as Pachycorinus; Type locality: Grande Comore).

  • — Bernhauer and Schubert, 1914: 296 (Pachycorynus; catalog).

  • Distribution: Comoros.

sulciceps Bernhauer, 1937b: 609 (Pachycorynus, cited as Pachycorinus; Type locality: W. Usambara).

  • Distribution: Tanzania.

sumatrensis Cameron, 1928, see: gracilis Cameron, 1928.

tabuensis Fauvel, 1878g: 536 (Pachycorynus; Type locality: Tonga Tabou).

  • — Bernhauer and Schubert, 1914: 296 (Pachycorynus; catalog).

  • Distribution: Tonga.

unicolor Cameron, 1937: 2 (Pachycorynus; Type locality: Java: G. Tangkoeban Prahoe).

  • Distribution: Indonesia.

vinsoni Cameron, 1947c: 115 (Xantholinus; Type locality: Mauritius: Mt. Cocotte).

  • — Lecoq, 1990: 182 (Pachycorynus; Mascarene Islands).

  • Distribution: Mascarene Islands.

Paracorynus [1 species; Australian Region]

Paracorynus Cameron, 1945d: 783, 785 (species included: arecae). Type species: Xantholinus arecae Broun, fixed by original designation.

  • — Blackwelder, 1952: 289 (type species: arecae).

arecae Broun, 1880: 103 (Xantholinus; Type locality: Whangarei Harbour).

  • — Bernhauer and Schubert, 1914: 300 (Xantholinus; catalog).

  • — Cameron, 1945d: 786 (Paracorynus).

  • — Kuschel, 1990: 26 (Paracorynus; endemic; winged; habitat; New Zealand).

  • — Klimaszewski, Newton, and Thayer, 1996: 151 (Paracorynus; endemic to New Zealand).

  • Distribution: New Zealand.

Paratesba [5 species; Oriental Region]

Paratesba Cameron, 1932: 40 (species included: indica). Type species: Tesba indica Eppelsheim, fixed by monotypy.

  • — Scheerpeltz, 1933: 1319 (world catalog supplement).

  • — Blackwelder, 1952: 294 (type species: indica).

  • — Bordoni, 1996: 73 (characters; key to species).

borneensis Bordoni, 1996: 78 (Paratesba; Type locality: Borneo: Brunei, 350 m).

  • Distribution: Borneo.

indica Eppelsheim, 1895b: 398 (Tesba; Type locality: Pegu).

  • — Bernhauer and Schubert, 1914: 314 (Tesba; catalog).

  • — Cameron, 1932: 40 (Paratesba; characters; Burma).

  • — Scheerpeltz, 1933: 1319 (Paratesba; catalog).

  • — Bordoni, 1996: 76, 81 (Paratesba; characters; Burma).

  • Distribution: Myanmar.

karenensis Bordoni, 1996: 76 (Paratesba; Type locality: Birmania: Carin Chebà, 900–1100 m. Also cited from Thailand).

  • Distribution: Myanmar, Thailand.

nagaii Bordoni, 1996: 78 (Paratesba; Type locality: East Malaysia, Borneo, Park Headquarters, Mt. Kinabalu).

  • Distribution: Borneo.

sternalis Bordoni, 1996: 81 (Paratesba; Type locality: W. Sumatra: Indrapura).

  • Distribution: Indonesia.

Paraxantholinus [1 species; Oriental Region]

Paraxantholinus Cameron, 1945, see: Pseudoxantholinus.

Paraxantholinus Bernhauer, 1926f: 316 (species included: corporaali). Type species: Paraxantholinus corporaali Bernhauer, fixed by original designation and monotypy.

  • — Scheerpeltz, 1933: 1304 (world catalog supplement).

  • — Blackwelder, 1952: 294 (type species: corporaali).

corporaali Bernhauer, 1926f: 317 (Paraxantholinus; Type locality: Buru).

  • — Scheerpeltz, 1933: 1304 (Paraxantholinus; catalog).

  • Distribution: Indonesia.

Paulianella [9 species; Madagascan Region]

Paulianella Jarrige, 1951b: 335 (subgenus of Leptacinus; species included: microps). Type species: Leptacinus (Paulianella) microps Jarrige, fixed by original designation and monotypy.

  • — Jarrige, 1970: 40 (valid genus; characters).

aethiops Jarrige, 1978: 273 (Paulianella; Type locality: Andringitra Est, Anjavidilava, 2000 m).

  • Distribution: Madagascar.

angustata Jarrige, 1978: 272 (Paulianella; Type locality: Andringitra Centre, zone sommitale, cirque Boby, 2500 m).

  • Distribution: Madagascar.

longiceps Jarrige, 1978: 275 (Paulianella; Type locality: Andringitra Sud, Andrianony, cirque Manjarivolo, 1600 m).

  • Distribution: Madagascar.

microps Jarrige, 1951b: 335 (Leptacinus; subgenus Paulianella; [Note: This is the type species of Paulianella, which was elevated to the generic level. The species was transferred implicitly]; Type locality: Madagascar: Tsiafajavona, Ankaratra).

  • Distribution: Madagascar.

monticola Jarrige, 1970: 42 (Paulianella; Type locality: Madagascar: Mt. Tsaratanana, 1800 m).

  • — Jarrige, 1978: 273 (Paulianella; Madagascar).

  • Distribution: Madagascar.

subaenea Jarrige, 1970: 42 (Paulianella; Type locality: Madagascar: Mt. Tsaratanana, 1500 m).

  • Distribution: Madagascar.

subnitida Jarrige, 1978: 275 (Paulianella; Type locality: Andringitra, forêt d'Andranomavaha, 2200 m).

  • Distribution: Madagascar.

tristis Jarrige, 1970: 42 (Paulianella; Type locality: Madagascar: Mt. Tsaratanana, 2000 m).

  • Distribution: Madagascar.

turficola Jarrige, 1970: 44 (Paulianella; Type locality: Madagascar: Mt. Tsaratanana, 2300 m).

  • Distribution: Madagascar.

Phacophallus [8 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions]

Phacophallus Coiffait, 1956a: 50 (species included: parumpunctatus, rubricollis). Type species: Staphylinus parumpunctatus Gyllenhal, fixed by original designation and monotypy.

  • — Smetana, 1958: 87 (subgenus of Leptacinus; characters; type species: parumpunctatus; Czechoslovakian species).

  • — Lohse, 1964: 160 (characters; key to central European species).

  • — Smetana, 1967d: 196 (subgenus of Leptacinus; characters in key).

  • — Coiffait, 1968: 128, 144 (characters; notes).

  • — Scheerpeltz, 1968a: 59 (catalog of Austrian species).

  • — Coiffait, 1972: 160 (characters; type species: parumpunctatus; key to species of western Palaearctic region; references).

  • — Shibata, 1973c: 54 (cited as Leptacinus; checklist of species of Taiwan).

  • — Bordoni, 1973j: 693 (list of species from islands near Sicily).

  • — Tikhomirova, 1973a: 181 (cited as subgenus of Leptacinus; checklist of species of USSR).

  • — Pope, 1977: 28 (list of British species).

  • — Muona, 1979: 16 (list of Scandinavian species).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 9 (catalog; Poland).

  • — Smetana, 1982: 106 (characters; habitus; revision of and key to species of North America north of Mexico).

  • — Bordoni, 1982: 194 (characters; habitus; key to species of Italy).

  • — Shibata, 1983: 74 (checklist of species of Japan).

  • — Frank, 1986a: 369 (checklist of species of Florida).

  • — Tóth, 1989: 9 (characters; species of Hungary).

  • — M. Hansen, Kristensen, Mahler, Pedersen, 1991: 107 (list of Danish species).

  • — Silfverberg, 1992: 18 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Hernández, Outerelo, and Gamarra, 1994: 190 (checklist of species; Canary Islands).

  • — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 28 (additional records to Danish list of species).

  • — Ciceroni and Zanetti, 1995: 25 (list of species of Italy).

  • — M. Hansen, 1996: 108 (list of Danish species).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 242 (additions to list of Danish species).

  • — Downie and Arnett, 1996: 399 (characters; key to species of Northeastern North America).

  • — Lundgren, 1998: 50 (list of species from Florida; records for other States listed for each species).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 391 (2 Nearctic species; notes; characters in key).

amissus Coquerel, 1860, see: parumpunctatus Gyllenhal, 1827.

ampliventris Jacquelin du Val, 1854, see: parumpunctatus Gyllenhal, 1827.

arabicus Coiffait, 1979a: 164 (Phacophallus; Type locality: Saudi Arabia: Wadi Hesua nr Wadi ad Villa, 600–700 m).

  • Distribution: Saudi Arabia.

breviceps C. Waterhouse, 1877, see: parumpunctatus Gyllenhal, 1827.

cephalicus LeConte, 1880, see: parumpunctatus Gyllenhal, 1827.

fauveli Cameron, 1922, see: parumpunctatus Gyllenhal, 1827.

flavipennis Kraatz, 1859: 111 (Leptacinus; Type locality: Ceylan; [Note: See lectotype designation by Smetana, 1980]).

  • — Sharp, 1874: 54 (Leptacinus; Japan).

  • — Blackburn and Sharp, 1885: 222 (Leptacinus; Hawaii).

  • — Bernhauer, 1902b: 32 (Leptacinus; Ceylon).

  • — Fauvel, 1903: 155 (Leptacinus; synonym of parumpunctatus).

  • — Fauvel, 1903d: 266 (Leptacinus; synonym of parumpunctatus).

  • — Fauvel, 1904a: 55 (Leptacinus; synonym of parumpunctatus).

  • — Sharp, 1908: 551 (Leptacinus; Hawaii).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; synonym of parumpunctatus).

  • — Bernhauer, 1922c: 231 (Leptacinus; variety of parumpunctatus; Formosa).

  • — Cameron, 1931: 359 (Leptacinus; New Guinea).

  • — Cameron, 1932: 9 (Leptacinus; variety of parumpunctatus).

  • — Scheerpeltz, 1933: 1303 (Leptacinus; cited as valid species; catalog).

  • — Blackwelder, 1943: 494 (Leptacinus; synonym of parumpunctatus).

  • — Scheerpeltz, 1954: 112 (Leptacinus; variety of parumpunctatus; Ceylon).

  • — Shibata, 1973a: 121 (Leptacinus; subgenus Phacophallus; subspecies of parumpunctatus; characters; Taiwan).

  • — Watanabe and Shibata, 1976: 331 (Leptacinus; subspecies of parumpunctatus, cited as parunpunctatus; Malaysia).

  • — Smetana, 1980: 54 (Phacophallus; lectotype designation; valid species; characters; notes on distribution).

  • — Lecoq, 1990: 188 (Phacophallus; Mascarene Islands).

  • Distribution: Mascarene Islands, Sri Lanka, Malaysia, New Guinea, Taiwan, Japan, Hawaii.

longicollis Stephens, 1833, see: parumpunctatus Gyllenhal, 1827.

pallidipennis Motschulsky, 1858, see: parumpunctatus Gyllenhal, 1827.

papuensis Fauvel, 1878, see: parumpunctatus Gyllenhal, 1827.

parumpunctatus Gyllenhal, 1827: 481 (Staphylinus; Type locality: Suecia).

  • — C. Sahlberg, 1830: 333 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 33 (Gyrohypnus; Finland; Russia).

  • — Mannerheim, 1831: 447 (Gyrohypnus; Finland; Russia).

  • — Stephens, 1833: 262 (Gyrohypnus; characters; England).

  • — Runde, 1835: 11 (Gyrohypnus; characters; Germany).

  • — Lacordaire, 1835: 414 (Xantholinus; characters; France).

  • — Nordmann, 1837: 117 (Gyrohypnus; Sweden; Russia).

  • — Erichson, 1839b: 335 (Leptacinus; characters; Germany; Sweden).

  • — Stephens, 1839: 403 (Xantholinus; characters; England).

  • — P. Lucas, 1846: 108 (Leptacinus; notes; Algeria).

  • — Jacobsen, 1849: 680 (Leptacinus; characters; Denmark).

  • — Redtenbacher, 1849: 693 (Leptacinus; characters; Austria).

  • — Hardy, 1851: 38 (Leptacinus; notes; England).

  • — Rosenhauer, 1856: 70 (Leptacinus; Cadiz; Granada).

  • — Fairmaire and Laboulbène, 1856: 503 (Leptacinus; characters; France).

  • — Baudi, 1857: 99 (Leptacinus; characters).

  • — Kraatz, 1857d: 648 (Leptacinus; characters; Germany).

  • — Redtenbacher, 1857: 183 (Leptacinus; characters; Austria).

  • — Thomson, 1859: 27 (type species of Leptacinus).

  • — Thomson, 1860: 193 (Leptacinus; characters; Scandinavia).

  • — Wollaston, 1864: 579 (Leptacinus; characters; Canary Islands).

  • — Wollaston, 1865: 498 (Leptacinus; habitat; Madeira Islands; Canary Islands).

  • — Wollaston, 1867: 241 (Leptacinus; characters; notes; Cape Verde Islands).

  • — Fauvel, 1873b: 88 [= 1873a: 374] (Leptacinus; characters; France; Europe; Asia; North Africa; Madeira; Cape Verde Islands; North America).

  • — Redtenbacher, 1874: 221 (Leptacinus; characters; Austria).

  • — Seidlitz, 1875: 280 (Leptacinus; characters; Baltic region).

  • — J. Sahlberg, 1876: 36 (Leptacinus; Finland).

  • — Mulsant and Rey, 1877a: 318 (Leptacinus; characters; France).

  • — Fauvel, 1878: 116 [= 1878a: 36] (Leptacinus; Morocco; Algeria; Tunisia).

  • — Fauvel, 1878g: 537 (Leptacinus; characters; Australie méridionale: Victoria, Melbourne).

  • — Fauvel, 1886: 48 [= 1886a: 40] (Leptacinus; North Africa).

  • — Olliff, 1887: 474 (Leptacinus; characters; Australia).

  • — Fowler, 1888: 292 (Leptacinus; characters; Britain).

  • — Fauvel, 1889b: 259 (Leptacinus; New Caledonia).

  • — Ganglbauer, 1895: 487 (Leptacinus; characters; Palaearctic and Nearctic regions; Cape Verde; Abyssinia; Australia; New Caledonia; Cuba).

  • — Fauvel, 1897d: 291 (Leptacinus; Barbary; Madeira; Canaries).

  • — Fauvel, 1902b: 100 (Leptacinus; Barbary; Egypt; Madeira; Canary Islands).

  • — Fauvel, 1903: 155 (Leptacinus; India).

  • — Fauvel, 1903d: 266 (Leptacinus; New Caledonia).

  • — Fauvel, 1904a: 55 (Leptacinus; Belgaum; Ghozeh; Khandesh; Nilghiris).

  • — Fauvel, 1907: 33 (Leptacinus; Nairobi).

  • — Sainte-Claire Deville, 1906: 93 (Leptacinus; Corsica).

  • — Reitter, 1908: 111 (Leptacinus; characters; Europe).

  • — Reitter, 1909: 135 (Leptacinus; characters; Germany).

  • — Eichelbaum, 1913: 129 (Leptacinus; Tanzania).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; catalog).

  • — Johansen, 1914: 438 (Leptacinus; characters; Denmark).

  • — Bernhauer, 1922a: 150 (Leptacinus; Samoa).

  • — Rambousek, 1922: 1008 (Leptacinus; Ethiopia).

  • — Lea, 1925: 215, 216 (Leptacinus; characters; Australian introduction).

  • — Cameron, 1928f: 410 (Leptacinus; Borneo).

  • — Portevin, 1929: 370 (Leptacinus; characters; France).

  • — Cameron, 1932: 8 (Leptacinus; characters; India).

  • — Scheerpeltz, 1933: 1303 (Leptacinus; catalog).

  • — Blackwelder, 1943: 494 (Leptacinus; characters; Cuba; Jamaica; Puerto Rico; St. Kitts; Antigua; Barbados; St. Vincent; Grenada).

  • — Cameron, 1950g: 28 (Leptacinus; Belgian Congo).

  • — V. Hansen, 1952: 9 (Leptacinus; characters; Denmark).

  • — Cameron, 1952e: 831 (Leptacinus; Dahomey).

  • — Coiffait, 1956a: 50 (Phacophallus; characters in key; France; Mediterranean region; Turkey).

  • — Hatch, 1957: 238 (Leptacinus; characters; Washington; Oregon).

  • — Coiffait, 1958: 24 (Phacophallus; notes).

  • — Smetana, 1958: 88 (Leptacinus; subgenus Phacophallus; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1961d: 126 (Phacophallus; Italy).

  • — Lohse, 1961: 183, 184, 185 (Phacophallus; characters).

  • — Palm, 1963b: 43 (Leptacinus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 160 (Phacophallus; characters; central Europe).

  • — Focarile, 1964: 57 (Leptacinus; Italy).

  • — Horion, 1965: 89 (Phacophallus; central and southern Europe; sparse in northern Europe).

  • — Coiffait, 1968: 144 (Phacophallus; notes).

  • — Bolov, 1969a: 328 (Leptacinus; collecting notes; Russia).

  • — Coiffait, 1972: 160 (Phacophallus; characters; references; central Europe; Turkey; Iberia; Morocco; Algeria; Gabon; France).

  • — Watanabe and Shibata, 1972: 63 (Phacophallus; Yaku-shima Island).

  • — Bacchus and Hammond, 1972: 63 (Leptacinus; donkey dung; Britain).

  • — Moore and Legner, 1974a: 39 (Leptacinus; spread by commerce).

  • — Scheerpeltz, 1974: 110 (Leptacinus; South Africa; Basutoland).

  • — Osella and Zanetti, 1975: 105 (Phacophallus; collected from nest of Talpa europaea; notes; Italy).

  • — Moore, 1975c: 107 (Phacophallus, cited as Phacolinus; Hawaii).

  • — Pope, 1977: 28 (Phacophallus; Britain).

  • — Scheerpeltz, 1978: 206 (Phacophallus; Timor).

  • — Raichev, 1979: 81 (Phacophallus; Bulgaria).

  • — Hinton, 1981: 663 (Leptacinus; characters of egg).

  • — Smetana, 1982: 107 (Phacophallus; characters; North American introduction; Ontario; Quebec; California; District of Columbia; Illinois; Kansas; Massachusetts; New Hampshire; New Jersey; New York; Pennsylvania).

  • — Bordoni, 1982: 196 (Phacophallus; characters; Italy).

  • — Hammond, 1982: 231 (Phacophallus; characters; may be of West African origin; Britain).

  • — Segers, 1986: 39 (Phacophallus; checklist of species; Belgium).

  • — Ádám, 1987: 146 (Phacophallus; collecting notes; Hungary).

  • — Lucht, 1987: 102 (Phacophallus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Tóth, 1989: 10 (Phacophallus; characters; Hungary).

  • — Janák, 1989: 169 (Phacophallus; notes; Tanzania; Kenya).

  • — Lohse and Lucht, 1989: 156 (Leptacinus; notes).

  • — Borges, 1990: Table II (Phacophallus; Azores).

  • — Campbell and Davies, 1991: 123 (Phacophallus; checklist of species; Canada).

  • — Janák, 1993a: 47 (Phacophallus; Czech Republic; Slovakia).

  • — Ciceroni, 1993: 70 (Phacophallus; Italy).

  • — Nishida, 1994: 73 (Phacophallus; Hawaii).

  • — Ciceroni and Zanetti, 1995: 25 (Phacophallus; Italy).

  • — Nishida, 1997: 62 (Phacophallus; Hawaii).

  • — Downie and Arnett, 1996: 399 (Phacophallus; characters; USA).

  • Distribution: Benin, Gabon, Congo, Lesotho, South Africa, “Zambezi”, Tanzania, Kenya, Ethiopia, Egypt, Tunisia, Algeria, Morocco, Cape Verde Islands, Canary Islands, Madeira, Azores, Europe, Russia, Turkey, India, Borneo, Indonesia, New Guinea, Australia, New Caledonia, Samoa, Japan, Hawaii, Canada, USA, West Indies.

longicollis Stephens, 1833: 259 (Gyrohypnus; Type locality: Suffolk).

  • — Stephens, 1839: 404 (Leptacinus; characters; England).

  • — Hardy, 1851: 39 (Leptacinus; synonym of batychrus).

  • — G. Waterhouse, 1858: 26 (Xantholinus and Leptacinus; synonym of linearis and parumpunctatus).

  • — Ganglbauer, 1895: 487 (Leptacinus; synonym of parumpunctatus).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; synonym of parumpunctatus).

  • — Cameron, 1932: 9 (Leptacinus; synonym of parumpunctatus).

  • — Méquignon, 1943: 160 (Leptacinus; synonym of batychrus).

  • — Blackwelder, 1943: 494 (Leptacinus; synonym of parumpunctatus).

ampliventris Jacquelin du Val, 1854: xxxvii (Leptacinus; Type locality: environs de Bercy).

  • — Fairmaire and Laboulbène, 1856: 503 (Leptacinus; characters; France).

  • — Kraatz, 1857d: 648 (Leptacinus; synonym of parumpunctatus).

  • — Mulsant and Rey, 1877a: 319 (Leptacinus; synonym of parumpunctatus).

  • — Ganglbauer, 1895: 487 (Leptacinus; synonym of parumpunctatus).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; synonym of parumpunctatus).

  • — Cameron, 1932: 9 (Leptacinus; synonym of parumpunctatus).

  • — Blackwelder, 1943: 494 (Leptacinus; synonym of parumpunctatus).

pallidipennis Motschulsky, 1858a: 206 (Leptacinus; Type locality: Indes orientales).

  • — Fauvel, 1903: 155 (Leptacinus; synonym of parumpunctatus).

  • — Fauvel, 1903d: 266 (Leptacinus; synonym of parumpunctatus).

  • — Fauvel, 1904a: 55 (Leptacinus; synonym of parumpunctatus).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; synonym of parumpunctatus).

  • — Cameron, 1932: 9 (Leptacinus; synonym of parumpunctatus).

  • — Blackwelder, 1943: 494 (Leptacinus; synonym of parumpunctatus).

  • — Hodge and Jones, 1995: 37 (Phacophallus; cited as valid species; [Note: These authors listed pallidipennis as a valid species and seem to suggest that the name is synonymous with tricolor Kraatz and therefore the senior name. They do not cite an article or data to justify the suggested changes and the articles they do cite do not make such a change. I have chosen to let stand the synonymy of pallidipennis with parumpunctatus]; notes; characters; Britain).

radiosus Peyron, 1858: 421 (Xantholinus; Type locality: Taurus, au Kuleg. Boghaz).

  • — Fauvel, 1869: 490 (Xantholinus; synonym of parumpunctatus).

  • — Fauvel, 1873b: 89 [= 1873a: 375] (Leptacinus; synonym of parumpunctatus).

  • — Ganglbauer, 1895: 487 (Leptacinus; synonym of parumpunctatus).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; synonym of parumpunctatus).

  • — Bernhauer and Schubert, 1914: 294, 307 (Xantholinus; [Note: The name, radiosus, is listed by Bernhauer and Schubert, as a valid species (p. 307) in Xantholinus and as a synonym of Leptacinus parumpunctatus (p. 294); catalog).

  • — Cameron, 1932: 9 (Leptacinus; synonym of parumpunctatus).

  • — Blackwelder, 1943: 494 (Leptacinus; synonym of parumpunctatus).

amissus Coquerel, 1860: 158 (Xantholinus; Type locality: Mers-el-Kébir).

  • — Fauvel, 1867c: 114 [= 1868b: 212] (Xantholinus; synonym of parumpunctatus).

  • — Fauvel, 1869: 490 (Leptacinus; synonym of parumpunctatus).

  • — Fauvel, 1869a: 151 (Leptacinus; synonym of parumpunctatus).

  • — Fauvel, 1873b: 89 [= 1873a: 375] (Leptacinus; synonym of parumpunctatus).

  • — Ganglbauer, 1895: 487 (Leptacinus; synonym of parumpunctatus).

  • — Fauvel, 1902b: 100 (Leptacinus; synonym of parumpunctatus).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; synonym of parumpunctatus).

  • — Cameron, 1932: 9 (Leptacinus; synonym of parumpunctatus).

  • — Blackwelder, 1943: 494 (Leptacinus; synonym of parumpunctatus).

breviceps C. Waterhouse, 1877: 24 (Leptacinus; Type locality: S. E. Africa, Zambesi).

  • — Fauvel, 1903: 155 (Leptacinus; synonym of parumpunctatus).

  • — Fauvel, 1904a: 55 (Leptacinus; synonym of parumpunctatus).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; synonym of parumpunctatus).

  • — Cameron, 1932: 9 (Leptacinus; synonym of parumpunctatus).

  • — Blackwelder, 1943: 494 (Leptacinus; synonym of parumpunctatus).

papuensis Fauvel, 1878f: 242 (Leptacinus; Type locality: Nouvelle-Guinée, Salvatti).

  • — Fauvel, 1879: 92 (Leptacinus; Dorei; Fly River).

  • — Fauvel, 1903: 155 (Leptacinus; synonym of parumpunctatus).

  • — Fauvel, 1903d: 266 (Leptacinus; synonym of parumpunctatus).

  • — Fauvel, 1904a: 55 (Leptacinus; synonym of parumpunctatus).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; synonym of parumpunctatus).

  • — Cameron, 1932: 9 (Leptacinus; synonym of parumpunctatus).

  • — Blackwelder, 1943: 494 (Leptacinus; synonym of parumpunctatus).

cephalicus LeConte, 1880: 170 (Leptacinus; Type locality: Columbia, South Carolina).

  • — Casey, 1906: 400 (Leptacinus; characters; New York).

  • — Bernhauer and Schubert, 1914: 293 (Leptacinus; catalog).

  • — Scheerpeltz, 1933: 1301 (Leptacinus; catalog).

  • — Smetana, 1982: 107 (Phacophallus; synonym of parumpunctatus).

sardous Fiori, 1894: 94 (Leptacinus; Type locality: Cabras, Sardegna).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; synonym of parumpunctatus).

  • — Cameron, 1932: 9 (Leptacinus; synonym of parumpunctatus).

  • — Bordoni, 1975d: 194 (Xantholinus; synonym of parumpunctatus).

  • — Smetana, 1982: 107 (Phacophallus; synonym of parumpunctatus).

  • — Bordoni, 1982: 196 (Phacophallus; synonym of parumpunctatus).

fauveli Cameron, 1922: 114 (Leptacinus; variety of parumpunctatus; Type locality: St. Vincent and Grenada).

  • — Scheerpeltz, 1933: 1303 (Leptacinus; variety of parumpunctatus).

  • — Blackwelder, 1943: 494 (Leptacinus; synonym of parumpunctatus).

povolnyi Bordoni, 1975a: 148 (Phacophallus; Type locality: O-Afghanistan: Prov. Nengrahar: Jalalabad, 580 m).

  • Distribution: Afghanistan.

radiosus Peyron, 1858, see: parumpunctatus Gyllenhal, 1827.

rubricollis Reitter, 1899, see: tricolor Kraatz, 1859.

rubricollis Casey, 1906, see: tricolor Kraatz, 1859.

sahariensis Coiffait, 1968a: 135, 139 (Phacophallus; Type locality: Tchad: Ennedi, mare de Damas. Also cited from Mauritania).

  • — Coiffait, 1966b: 337 (Phacophallus; Mauritania).

  • — Coiffait, 1972: 164 (Phacophallus; characters; Sahara).

  • — Janák, 1989: 170 (Phacophallus; notes; Sudan).

  • Distribution: Chad, Sudan, Mauritania.

sardous Fiori, 1894, see: parumpunctatus Gyllenhal, 1827.

tricolor Kraatz, 1859: 110 (Leptacinus; Type locality: Ceylan; [Note: See lectotype designation by Smetana, 1980]. Also cited from India orientalis).

  • — Fauvel, 1903: 155 (Leptacinus; synonym of parumpunctatus).

  • — Fauvel, 1903d: 266 (Leptacinus; synonym of parumpunctatus).

  • — Fauvel, 1904a: 55 (Leptacinus; synonym of parumpunctatus).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; synonym of parumpunctatus).

  • — Cameron, 1921a: 375, 404 (Leptacinus; characters; Singapore).

  • — Cameron, 1932: 9 (Leptacinus; variety of parumpunctatus).

  • — Scheerpeltz, 1933: 1303 (Leptacinus; not synonym; catalog).

  • — Cameron, 1937: 1 (Leptacinus; Java).

  • — Blackwelder, 1943: 494 (Leptacinus; synonym of parumpunctatus).

  • — Jarrige, 1952a: 132 (Leptacinus; subspecies of parumpunctatus; Canary Islands).

  • — Coiffait, 1972: 163 (Phacophallus; [Note: Misidentified as trigonocephalus according to Smetana, 1980: 52]; characters; references; Sicily; France; Morocco; Europe; North Africa).

  • — Smetana, 1980: 52 (Phacophallus; lectotype designation: Ceylan; characters; notes on distribution).

  • — Smetana, 1982: 109 (Phacophallus; characters; North American introduction; Arizona; California; Florida; Illinois; Maryland; Massachusetts; New Mexico; New York; Oregon; Pennsylvania; Texas; West Virginia).

  • — Hammond, 1982: 231 (Phacophallus; characters; may be of Asian origin; Britain).

  • — Coiffait, 1982a: 232 (Phacophallus; variety of parumpunctatus; India).

  • — Bordoni, 1982: 198 (Phacophallus; [Note: Cited as trigonocephalus but probably a misidentification of tricolor as suggested by the work of Smetana, 1980: 52]; characters; Italy).

  • — Smetana, 1988a: 527 (Phacophallus; Mississippi).

  • — Lecoq, 1990: 187 (Phacophallus; introduction to Mascarene Islands).

  • — Smetana, 1990c: 84 (Phacophallus; Arkansas; North Carolina; Oklahoma).

  • — Ciceroni and Zanetti, 1995: 25 (Phacophallus; Italy).

  • — Hayashi, 1996a: 22 (Phacophallus; Japan).

  • — Downie and Arnett, 1996: 399 (Phacophallus; characters; USA).

  • Distribution: Morocco, Europe, Mascarene Islands, India, Sri Lanka, Singapore, Indonesia, Japan, USA.

rubricollis Reitter, 1899: 157 (Leptacinus; variety of parumpunctatus; Type locality: Sicilien; [Note: See lectotype designation by Smetana, 1980]).

  • — Fauvel, 1903: 155 (Leptacinus; synonym of parumpunctatus).

  • — Fauvel, 1903d: 266 (Leptacinus; synonym of parumpunctatus).

  • — Fauvel, 1904a: 55 (Leptacinus; synonym of parumpunctatus).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; synonym of parumpunctatus).

  • — Cameron, 1932: 9 (Leptacinus; variety of parumpunctatus).

  • — Scheerpeltz, 1933: 1303 (Leptacinus; variety of parumpunctatus).

  • — Blackwelder, 1943: 494 (Leptacinus; synonym of parumpunctatus).

  • — Jarrige, 1952a: 132 (Leptacinus; synonym of parumpunctatus tricolor).

  • — Coiffait, 1956a: 51 (Phacophallus; characters in key; France).

  • — Coiffait, 1972: 163 (Phacophallus; synonym of trigonocephalus).

  • — Smetana, 1980: 52 (Phacophallus; lectotype designation; synonym of tricolor).

  • — Smetana, 1982: 109 (Phacophallus; synonym of tricolor).

rubricollis Casey, 1906: 400 (Leptacinus; [preoccupied]; [Note: Bernhauer and Schubert, 1914: 294, cited this reference as a subsequent reference of rubricollis Reitter]; Type locality: Missouri, St. Louis; [Note: See lectotype designation by Smetana, 1980]. Also cited from Kentucky).

  • — Scheerpeltz, 1933: 1301 (Leptacinus; aberration of cephalicus).

  • — Smetana, 1980: 52 (Phacophallus; lectotype designation: St. Louis, Missouri; synonym of tricolor).

  • — Smetana, 1982: 109 (Phacophallus; synonym of tricolor).

trigonocephalus Kraatz, 1859: 109 (Leptacinus; Type locality: Ceylan; [Note: See lectotype designation by Smetana, 1980]).

  • — Fauvel, 1904a: 56 (Leptacinus; Nilghiris; Ceylan).

  • — Bernhauer and Schubert, 1914: 294 (Leptacinus; catalog).

  • — Cameron, 1932: 8 (Leptacinus; characters; Ceylon, India).

  • — Scheerpeltz, 1933: 1303 (Leptacinus; catalog).

  • — Coiffait, 1977b: 205 (Phacophallus; Bhutan).

  • — Smetana, 1980: 50 (Leptacinus; lectotype designation; characters; may belong in different genus; Sri Lanka).

  • — Assing, 2000b: 115 (Phacophallus; collecting notes; Canary Islands).

  • Distribution: Sri Lanka, India, Bhutan, Canary Islands.

xanthopygus Scheerpeltz, 1974c: 19 (Leptacinus; Type locality: Sudan: Erkowit).

  • — Janák, 1989: 170 (Phacophallus; characters; notes; Sudan).

  • Distribution: Sudan.

Platydromus [2 species; Madagascan Region]

Platydromus Fauvel, 1905b: 174 (species included: erosus). Type species: Platydromus erosus Fauvel, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 309 (world catalog; 1 species).

  • — Blackwelder, 1952: 313 (type species: erosus).

erosus Fauvel, 1905b: 174 (Platydromus; Type locality: Forêt de Moramanga).

  • — Bernhauer and Schubert, 1914: 309 (Platydromus; catalog).

  • Distribution: Madagascar.

nigripennis Jarrige, 1948a: 18 (Platydromus; Type locality: Madagascar: Antongil).

  • Distribution: Madagascar.

Plochionocerus [29 species; Neotropical Region]

Plochionocerus Sharp, 1885, see: Renda.

Plochionocerus Dejean, 1833: 64 (species included: violaceus and one unavailable name). Type species: Staphylinus violaceus Olivier, fixed by monotypy.

  • — Blackwelder, 1952: 315 (type species: violaceus).

Sterculia Laporte, 1835: 118 (species included: violacea, leprieuri, formicaria). Type species: Staphylinus violacea Olivier, fixed by original designation.

  • — Erichson, 1839b: 301 (characters).

  • — Laporte, 1840: 180 (characters).

  • — Erichson, 1847: 88 (synonym of Agrodes).

  • — Lacordaire, 1854: 66 (synonym of Agrodes).

  • — Gemminger and Harold, 1868: 599 (catalog).

  • — Sharp, 1876: 186 (characters; notes).

  • — Duvivier, 1883: 155 (catalog).

  • — Bernhauer and Schubert, 1914: 314 (world catalog; 24 species).

  • — Scheerpeltz, 1933: 1320 (world catalog supplement).

  • — Blackwelder, 1944: 149 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 360 (synonym of Plochionocerus; type species: violacea).

Agrodes Nordmann, 1837: 161 (species included: elegans). Type species: Agrodes elegans Nordmann, fixed by monotypy.

  • — Lacordaire, 1854: 66 (characters; notes; list of species).

  • — Gemminger and Harold, 1868: 599 (synonym of Sterculia).

  • — Sharp, 1876: 192 (notes).

  • — Bernhauer and Schubert, 1914: 314 (subgenus of Sterculia).

  • — Blackwelder, 1952: 42 (subgenus of Plochionocerus; type species: elegans).

  • — Newton, 1995: 51 (requested that the International Commission of Zoological Nomenclature place Agrodes on the Official List of Generic Names in Zoology).

  • — ICZN, 1996: 215 (placed on Official List of Generic Names in Zoology).

Araeocnemus Nordmann, 1837: 163 (species included: fulgens, flagellicornis, pubescens). Type species: Staphylinus fulgens Fabricius, fixed by subsequent designation (see Blackwelder, 1952: 61).

  • — Erichson, 1839b: 301 (cited as Araeocnemis; synonym of Sterculia).

  • — Lacordaire, 1854: 66 (synonym of Agrodes).

  • — Gemminger and Harold, 1868: 599 (synonym of Sterculia).

  • — Bernhauer and Schubert, 1914: 314 (synonym of Sterculia).

  • — Casey, 1906: 359 (cited both as Araeocnemis and Araeocnemes; cited to replace Sterculia because it is a homonym of a genus in plants; characters).

Plociocerus Agassiz, 1847: 299 (emendation of Plochionocerus). Type species: Staphylinus violaceus Olivier, fixed by objective synonymy with Plochionocerus.

  • — Blackwelder, 1952: 315 (synonym of Plochionocerus; type species).

Callictenus Chevrolat, 1848: 24 (Note: Nomen nudum (Article 11.6, 12.1); the genus was published as synonym of Sterculia, without a description, and without assigned species. Blackwelder, 1952: 89 considered it to be an isogenotypic synonym of Sterculia).

amazonicus Sharp, 1876: 186 (Sterculia; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 314 (Sterculia; catalog).

  • Distribution: Brazil.

aureipennis Bernhauer, 1927d: 164 (Sterculia; Type locality: Brasilien).

  • — Scheerpeltz, 1933: 1320 (Sterculia; catalog).

  • Distribution: Brazil.

basalis Sharp, 1885: 469 (Sterculia; Type locality: Mexico, Jalapa).

  • — Bernhauer and Schubert, 1914: 314 (Sterculia; catalog).

  • Distribution: Mexico.

brasilianus Bernhauer, 1927, see: Renda.

carinatus Bernhauer, 1916f: 185 (Sterculia; Type locality: Bolivien: Rio Gongo, 750 m; Peru: Poguzo, Rolle).

  • — Scheerpeltz, 1933: 1320 (Sterculia; catalog).

  • Distribution: Bolivia, Peru.

clavicornis Sharp, 1876, see: Renda.

coelestinus Erichson, 1839, see: violaceus Olivier, 1795.

conicicollis Sharp, 1876: 192 (Agrodes; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 314 (Sterculia; catalog; Colombia).

  • Distribution: Brazil, Columbia.

dalmasi Fauvel, 1901, see: Renda.

discedens Sharp, 1885: 470 (Sterculia; Type locality: Guatemala, Quiche Mountains 7000–9000 feet).

  • — Bernhauer and Schubert, 1914: 314 (Sterculia; catalog).

  • Distribution: Guatemala.

discolor Sharp, 1876, see: splendens Blanchard, 1842.

elegans Nordmann, 1837: 162 (Agrodes; Type locality: Para).

  • — Erichson, 1839b: 303 (Sterculia; characters; Brazil).

  • — Bernhauer and Schubert, 1914: 314 (Sterculia; catalog).

  • — Newton, 1995: 51 (requested that the International Commission of Zoological Nomenclature place elegans on the Official list of Specific Names in Zoology).

  • — ICZN, 1996: 215 (placed on Official List of Specific Names in Zoology).

  • Distribution: Brazil.

fimetarius Sharp, 1876, see: Renda.

flagellicornis Nordmann, 1837, see: Renda.

formicaria Laporte, 1835, see: Renda.

fulgens Fabricius, 1793: 522 (Staphylinus; Type locality: Cayennae).

  • — Fabricius, 1801: 592 (Staphylinus; characters; French Guiana).

  • — Turton, 1802: 510 (Staphylinus; characters; Cayenne).

  • — Gravenhorst, 1806: 91 (Staphylinus; characters; French Guiana).

  • — Erichson, 1839b: 303 (Sterculia; characters; Mexico; Cayenne; Surinam).

  • — Sharp, 1885: 468 (Sterculia; notes; Mexico; Cayenne).

  • — Fauvel, 1901c: 251 (Sterculia; characters; discussion of identity).

  • — Bernhauer and Schubert, 1914: 314 (Sterculia; catalog).

  • Distribution: Mexico, Surinam, French Guiana, Brazil.

pauloensis Sharp, 1876: 187 (Sterculia; Type locality: St. Paulo).

  • — Fauvel, 1901c: 252 (Sterculia; synonym of fulgens).

  • — Bernhauer and Schubert, 1914: 314 (Sterculia; synonym of fulgens).

fulgens Brullé, 1842, see: splendens Blanchard, 1842.

funebris Sharp, 1876, see: Renda.

holtzi Bernhauer, 1907, see: leprieuri Laporte, 1835.

humeralis Sharp, 1885: 469 (Sterculia; Type locality: Panama, Volcan de Chiriqui 2500 to 6000 feet; Colombia; Venezuela).

  • — Fauvel, 1891: 106 (Sterculia; Venezuela).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog).

  • Distribution: Panama, Columbia, Venezuela.

janthinus Erichson, 1847: 88 (Agrodes; Type locality: Republica Peruana).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog).

  • Distribution: Peru.

igneus Fauvel, 1901c: 252 (Sterculia; Type locality: Amazone).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog).

  • Distribution: Brazil.

impressipennis Sharp, 1891: 41 (Sterculia; Type locality: Nanegal, 3–4,000 feet).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog).

  • Distribution: Ecuador.

kayseri Bernhauer, 1927d: 163 (Sterculia; Type locality: Venezuela).

  • — Scheerpeltz, 1933: 1320 (Sterculia; catalog).

  • Distribution: Venezuela.

lautus Casey, 1906: 360 (Araeocnemus, cited as Araeocnemis; Type locality: Panama).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog).

  • Distribution: Panama.

leprieuri Laporte, 1835: 118 (Sterculia; Type locality: Cayenne).

  • — Erichson, 1839b: 304 (Sterculia; characters; Cayenne).

  • — Laporte, 1840: 180 (Sterculia; characters; Cayenne).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog).

  • Distribution: French Guiana, Guyana.

holtzi Bernhauer, 1907a: 284 (Sterculia; Type locality: Britisch-Guayana).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; synonym of leprieuri).

longiceps Sharp, 1876: 193 (Agrodes; Type locality: Tunantins).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog; Bolivia; Peru).

  • Distribution: Brazil, Bolivia, Peru.

longicornis Sharp, 1885: 467 (Agrodes; Type locality: Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog).

  • Distribution: Nicaragua.

magnificus Bernhauer, 1911a: 412 (Sterculia; Type locality: Amazonas, Obidos).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog).

  • Distribution: Brazil.

mandibularis Sharp, 1885: 468 (Sterculia; Type locality: Mexico, Jalapa; Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog; Guatemala).

  • Distribution: Mexico, Guatemala, Nicaragua.

melancholicus Schubert, 1911: 13 (Sterculia; Type locality: Chanchamayo, Peru, 2000 m).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog).

  • Distribution: Peru.

minor Sharp, 1876, see: Renda.

nevermanni Bernhauer, 1942: 23 (Sterculia; Type locality: Kostarika: San José, 1000–1200 m).

  • Distribution: Costa Rica.

nordmanni Fauvel, 1901c: 252 (Sterculia; proposed for Nordmann's misidentification of fulgens; Type locality: Mexico. Also cited Guatemala).

  • — Nordmann, 1837: 165 (Araeocnemus; [Note: Misidentification: Bernhauer and Schubert, 1914: 315, cited fulgens Nordmann as a synonym of nordmanni Fauvel; Nordmann did not describe the species as new, he attributed it to Fabricius]; characters; Mexico).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog).

  • Distribution: Mexico, Guatemala.

pauloensis Sharp, 1876, see: fulgens Fabricius, 1793

peruvianus Bernhauer, 1907a: 284 (Sterculia; Type locality: Peru: Callanga).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog; Bolivia).

  • Distribution: Peru, Bolivia.

pollens Sharp, 1885: 469 (Sterculia; Type locality: Guatemala, Coban in Vera Paz; Nicaragua, Chontales; Costa Rica, Cache).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog).

  • Distribution: Guatemala, Nicaragua, Costa Rica.

profundepunctatus Bernhauer, 1927, see: Renda.

pubescens Nordmann, 1837, see: Renda.

puncticeps Sharp, 1885: 470 (Sterculia; Type locality: Costa Rica).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog).

  • Distribution: Costa Rica.

punctipennis Bernhauer, 1916f: 186 (Sterculia; Type locality: Westkolumbien: Monte Tolima, 1700 m).

  • — Scheerpeltz, 1933: 1320 (Sterculia; catalog).

  • Distribution: Colombia.

simplicicollis C. Waterhouse, 1879: 421 (Sterculia; Type locality: Medellin, Colombia).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog).

  • Distribution: Columbia.

splendens Blanchard, 1842: 83 (Sterculia; Type locality: Bolivia: Guarayos, province des Moxos, les rives du Mamoré).

  • — Solsky, 1872: 308 (Sterculia; Peru).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog).

  • Distribution: Brazil, Bolivia, Peru.

fulgens Brullé, 1842: pl. 5, fig. 10 (Sterculia; [preoccupied]; Type locality: Mexique et la Guyane [as cited by Blanchard, 1842: 83]).

  • — Blanchard, 1842: 83 (Sterculia; synonym of splendens).

  • — Fauvel, 1901c: 252 (Sterculia; synonym of splendens).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; synonym of splendens).

discolor Sharp, 1876: 188 (Sterculia; Type locality: Ega).

  • — Fauvel, 1901c: 252 (Sterculia; synonym of splendens).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; synonym of splendens).

violaceus Olivier, 1795:(42): 8 (Staphylinus; Type locality: Cayenne, Surinam).

  • — Gravenhorst, 1806: 92 (Staphylinus; synonym of fulgens Fabricius).

  • — Laporte, 1835: 118 (type species of Sterculia).

  • — Erichson, 1839b: 303 (Sterculia; synonym of fulgens).

  • — Laporte, 1840: 180 (Sterculia; characters; Cayenne).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; catalog).

  • Distribution: French Guiana, Surinam, Brazil.

coelestinus Erichson, 1839b: 302 (Sterculia; Type locality: Brasilia).

  • — Fauvel, 1901c: 252 (Sterculia; synonym of violacea).

  • — Bernhauer and Schubert, 1914: 315 (Sterculia; synonym of violacea).

Pseudocorynus [2 species; Australian Region]

Pseudocorynus Cameron, 1945d: 783, 785 (species included: cultus, mediocris). Type species: Xantholinus cultus Broun, fixed by original designation.

  • — Blackwelder, 1952: 325 (type species: cultus).

cultus Broun, 1880: 103 (Xantholinus; Type locality: Parua).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • — Cameron, 1945d: 785 (Pseudocorynus).

  • Distribution: New Zealand.

mediocris Broun, 1880: 104 (Xantholinus; Type locality: Parua).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • — Cameron, 1945d: 785 (Pseudocorynus).

  • Distribution: New Zealand.

Pseudoxantholinus [1 species; Australian Region]

Pseudoxantholinus Cameron, 1946b: 500 (replacement name for Paraxantholinus Cameron). Type species: Xantholinus sharpi Broun, fixed by objective synonymy with Paraxantholinus.

  • — Blackwelder, 1952: 331 (type species).

  • — Klimaszewski, Newton, and Thayer, 1996: 152 (endemic to New Zealand).

Paraxantholinus Cameron, 1945d: 783, 784 (species included: sharpi; preoccupied by Bernhauer, 1926). Type species: Xantholinus sharpi Broun, fixed by original designation and monotypy.

  • — Cameron, 1946b: 500 (synonym of Pseudoxantholinus).

  • — Blackwelder, 1952: 294 (synonym of Pseudoxantholinus; type species: sharpi).

sharpi Broun, 1880: 102 (Xantholinus; Type locality: Dumfriesshire).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • — Cameron, 1945d: 784 (Paraxantholinus).

  • Distribution: New Zealand.

Renda [12 species; Neotropical Region]

Renda Blackwelder, 1952: 338 (replacement name for Plochionocerus Sharp). Type species: Sterculia formicaria Laporte, fixed by objective synonymy with Plochionocerus Sharp.

Plochionocerus Sharp, 1885: 471 (species included: funebris, fimetaria, clavicornis, minor, brachypterus, formicaria, debilis; preoccupied by Dejean, 1833). Type species: Sterculia formicaria Laporte, fixed by original designation.

  • — Bernhauer and Schubert, 1914: 315 (world catalog; 8 species).

  • — Bruch, 1928a: 427 (catalog; Argentina).

  • — Scheerpeltz, 1933: 1320 (world catalog supplement).

  • — Blackwelder, 1943: 491 (characters; type species: formicarius).

  • — Blackwelder, 1944: 149 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 315 (synonym of Renda; type species: formicarius).

brachyptera Sharp, 1885: 471 (Plochionocerus; Type locality: Mexico, Cordova, Misantla; Guatemala, Cerro Zunil 4000 to 5000 feet, Las Mercedes 3000 feet, Zapote; Honduras; Panama, Bugaba).

  • — Bernhauer and Schubert, 1914: 315 (Plochionocerus; catalog).

  • — Blackwelder, 1943: 492 (Plochionocerus; characters; Trinidad).

  • Distribution: Mexico, Guatemala, Honduras, Panama, West Indies.

brasiliana Bernhauer, 1927d: 164 (Plochionocerus; Type locality: Brasilien: Rio de Janeiro).

  • — Scheerpeltz, 1933: 1320 (Plochionocerus; catalog).

  • Distribution: Brazil.

clavicornis Sharp, 1876: 190 (Sterculia; Type locality: Obydos).

  • — Sharp, 1885: 471 (Plochionocerus).

  • — Bernhauer and Schubert, 1914: 315 (Plochionocerus; catalog; Peru).

  • Distribution: Brazil, Peru.

dalmasi Fauvel, 1901: 84 (Plochionocerus; Type locality: Colombie).

  • — Bernhauer and Schubert, 1914: 315 (Plochionocerus; catalog).

  • Distribution: Colombia.

debilis Sharp, 1885: 472 (Plochionocerus; Type locality: Guatemala, San Geronimo and San Joaquin in Vera Paz).

  • — Bernhauer and Schubert, 1914: 315 (Plochionocerus; catalog).

  • Distribution: Guatemala.

fimetaria Sharp, 1876: 190 (Sterculia; Type locality: Ega).

  • — Sharp, 1885: 471 (Plochionocerus).

  • — Bernhauer and Schubert, 1914: 315 (Plochionocerus; catalog).

  • Distribution: Brazil.

flagellicornis Nordmann, 1837: 165 (Araeocnemus; Type locality: Cameta; Surinam).

  • — Blanchard, 1842: 83 (Sterculia; synonym of formicaria).

  • — Erichson, 1839b: 304 (Sterculia; synonym of formicaria).

  • — P. Lucas, 1857: 49 (Agrodes; synonym of formicarius).

  • — Fauvel, 1901: 83 (Plochionocerus; synonym of formicarius).

  • — Bernhauer and Schubert, 1914: 316 (Plochionocerus; synonym of formicarius).

  • — Scheerpeltz, 1933: 1320 (Plochionocerus; catalog).

  • Distribution: Surinam.

formicaria Laporte, 1835: 119 (Sterculia; Type locality: Cayenne).

  • — Blanchard, 1842: 83 (Sterculia; Colombia; Guyana; Brazil).

  • — Erichson, 1839b: 304 (Sterculia; characters; Brazil; Surinam; Cayenne; Colombia).

  • — Laporte, 1840: 181 (Sterculia; characters; Cayenne).

  • — P. Lucas, 1857: 49 (Agrodes; Brazil).

  • — Sharp, 1885: 471 (Plochionocerus; notes; Costa Rica; Colombia; Cayenne).

  • — Fauvel, 1901: 83 (Plochionocerus; Venezuela; Costa Rica; Demerara; Cayenne; Brazil; Peru).

  • — Bernhauer and Schubert, 1914: 315 (Plochionocerus; catalog).

  • — Scheerpeltz, 1933: 1320 (Plochionocerus; catalog).

  • Distribution: Peru, Brazil, French Guiana, Guyana, Venezuela, Colombia, Costa Rica.

funebris Sharp, 1876: 188 (Sterculia; Type locality: Ega).

  • — Sharp, 1885: 471 (Plochionocerus).

  • — Fauvel, 1901: 83 (Plochionocerus; synonym of formicarius).

  • — Bernhauer and Schubert, 1914: 316 (Plochionocerus; catalog).

  • Distribution: Brazil.

minor Sharp, 1876: 191 (Sterculia; Type locality: Fonteboa; Ega).

  • — Sharp, 1885: 471 (Plochionocerus).

  • — Bernhauer and Schubert, 1914: 316 (Plochionocerus; catalog).

  • Distribution: Brazil.

profundepunctata Bernhauer, 1927d: 164 (Plochionocerus; Type locality: Brasilien: Itatiaya, Estacao do Rios, 800 m, Montserrat).

  • Distribution: Brazil.

pubescens Nordmann, 1837: 167 (Araeocnemus; Type locality: Brasilia meridionalis).

  • — Blanchard, 1842: 83 (Sterculia; synonym of formicaria).

  • — Erichson, 1839b: 304 (Sterculia; synonym of formicaria).

  • — P. Lucas, 1857: 49 (Agrodes; synonym of formicarius).

  • — Fauvel, 1901: 83 (Plochionocerus; synonym of formicarius).

  • — Bernhauer and Schubert, 1914: 316 (Plochionocerus; synonym of formicarius).

  • — Scheerpeltz, 1933: 1320 (Plochionocerus; catalog).

  • Distribution: Brazil.

Scytalinus [9 species; Neotropical Region]

Scytalinus Erichson, 1839b: 305 (species included: serpentinus). Type species: Scytalinus serpentinus Erichson, fixed by monotypy.

  • — Lacordaire, 1854: 67 (characters; notes; list of species).

  • — Gemminger and Harold, 1868: 604 (catalog).

  • — Bernhauer and Schubert, 1914: 314 (world catalog; 4 species).

  • — Bruch, 1928a: 427 (catalog; Argentina).

  • — Scheerpeltz, 1933: 1320 (world catalog supplement).

  • — Blackwelder, 1944: 149 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 349 (type species: serpentinus).

azureus Lynch, 1884: 179 (Xantholinus; subgenus Eulissus; Type locality: Republica Argentina in Arrecifes; Republica Uruguayensis).

  • — Bernhauer and Schubert, 1914: 314 (Scytalinus; catalog).

  • Distribution: Argentina, Uruguay.

bruchi Bernhauer, 1921c: 104 (Scytalinus; Type locality: Argentinien: Prov. Santiago d. Estero).

  • — Scheerpeltz, 1933: 1320 (Scytalinus; catalog).

  • Distribution: Argentina.

flavipennis Bernhauer, 1927d: 162 (Scytalinus; Type locality: Brasilien: Petropolis).

  • — Scheerpeltz, 1933: 1320 (Scytalinus; catalog).

  • Distribution: Brazil.

parcepunctatus Bernhauer, 1911a: 412 (Scytalinus; Type locality: Brasilien).

  • — Bernhauer and Schubert, 1914: 314 (Scytalinus; catalog).

  • Distribution: Brazil.

parcus Bernhauer, 1927d: 162 (Scytalinus; Type locality: Brasilien: Virginia, S. Minas Geraes, Faz Campos, 1500 m).

  • — Scheerpeltz, 1933: 1320 (Scytalinus; catalog).

  • Distribution: Brazil.

rugiceps Scriba, 1855: 298 (Scytalinus; Type locality: Venezuela).

  • — Solsky, 1872: 308 (Scytalinus; Cayenne).

  • — Bernhauer and Schubert, 1914: 314 (Scytalinus; catalog).

  • Distribution: Venezuela, French Guiana.

serpentinus Erichson, 1839b: 305 (Scytalinus; Type locality: Brasilia).

  • — Bernhauer and Schubert, 1914: 314 (Scytalinus; catalog).

  • Distribution: Brazil.

spectabilis Bernhauer, 1915l: 295 (Scytalinus, cited as Scitalinus; Type locality: Peru: Chanchamayo, 1500 m).

  • — Scheerpeltz, 1933: 1320 (Scytalinus; catalog).

  • Distribution: Peru.

zikani Bernhauer, 1927d: 162 (Scytalinus; Type locality: Brasilien: Virginia, S. Minas Geraes, Faz Campos, 1500 m).

  • — Scheerpeltz, 1933: 1320 (Scytalinus; catalog).

  • Distribution: Brazil.

Somoleptus [22 species; Neotropical and Oriental Regions]

Somoleptus Sharp, 1885: 494 (species included: clavicornis, subtilis, debilis, nitidus, aenescens, bicolor, gracilis, parvulus, obscurus, obsoletus, punctulatus, sparsus, pallipes, elegans, alticola, agraeformis). Type species: Somoleptus aenescens Sharp, fixed by subsequent designation by Blackwelder, 1952: 354.

  • — Bernhauer and Schubert, 1914: 297 (world catalog; 19 species).

  • — Cameron, 1921a: 353, 404 (characters; key to Singapore species; catalog).

  • — Bruch, 1928a: 427 (catalog; Argentina).

  • — Scheerpeltz, 1933: 1304 (world catalog supplement).

  • — Blackwelder, 1944: 146 (checklist of species from Mexico, Central and South America, and the West Indies).

aenescens Sharp, 1885: 495 (Somoleptus; Type locality: Guatemala, near the city, San Geronimo, San Joaquin in Vera Paz).

  • — Bernhauer and Schubert, 1914: 297 (Somoleptus; catalog).

  • Distribution: Guatemala.

agraeformis Sharp, 1885: 498 (Somoleptus; Type locality: Mexico).

  • — Bernhauer and Schubert, 1914: 297 (Somoleptus; catalog).

  • Distribution: Mexico.

alticola Sharp, 1885: 498 (Somoleptus; Type locality: Guatemala, Totonicapam between 8500 to 10,500 feet).

  • — Bernhauer and Schubert, 1914: 297 (Somoleptus; catalog).

  • Distribution: Guatemala.

bicolor Sharp, 1885: 495 (Somoleptus; Type locality: Guatemala, Capetillo, Quezaltenango 7800 feet).

  • — Sharp, 1887: 791 (Somoleptus; Mexico).

  • — Bernhauer and Schubert, 1914: 297 (Somoleptus; catalog).

  • Distribution: Guatemala, Mexico.

clavicornis Erichson, 1839b: 337 (Leptacinus; Type locality: vallis Araguensis Columbiae).

  • — Scriba, 1855: 299 (Leptacinus; Venezuela).

  • — Sharp, 1885: 495 (Somoleptus).

  • — Bernhauer and Schubert, 1914: 297 (Somoleptus; catalog).

  • Distribution: Colombia, Venezuela.

claviscapus Cameron, 1922, see: Lithocharodes.

columbicus Bernhauer, 1915: 13 (Somoleptus; [Note: Cited as columbiens; I accept Scheerpeltz's 1933 emendation as justified]; Type locality: Columbien: Bogota).

  • — Scheerpeltz, 1933: 1304 (Somoleptus; emendation of columbiens; catalog).

  • Distribution: Colombia.

debilis Erichson, 1839b: 336 (Leptacinus; Type locality: vallis Araguensis Columbiae).

  • — Scriba, 1855: 299 (Leptacinus; Venezuela).

  • — Fauvel, 1878f: 242 (Xantholinus).

  • — Sharp, 1885: 495 (Somoleptus “possibly”).

  • — Bernhauer and Schubert, 1914: 297 (Somoleptus; catalog).

  • Distribution: Columbia, Venezuela.

diochiformis Bernhauer, 1910b: 376 (Somoleptus; Type locality: Mexiko: Vera Cruz, Cordoba).

  • — Bernhauer and Schubert, 1914: 297 (Somoleptus; catalog).

  • Distribution: Mexico.

elegans Sharp, 1885: 498 (Somoleptus; Type locality: Guatemala, Balheu).

  • — Bernhauer and Schubert, 1914: 297 (Somoleptus; catalog).

  • Distribution: Guatemala.

glabripennis Scriba, 1855: 299 (Leptacinus; Type locality: Venezuela).

  • — Fauvel, 1869: 490 (Mitomorphus).

  • — Bernhauer and Schubert, 1914: 297 (Somoleptus; catalog).

  • Distribution: Venezuela.

gracilis Sharp, 1885: 496 (Somoleptus; Type locality: Mexico, Cordova).

  • — Bernhauer and Schubert, 1914: 297 (Somoleptus; catalog).

  • Distribution: Mexico.

laevis Bernhauer, 1908d: 325 (Somoleptus; Type locality: Brasilien: S. Paulo).

  • — Bernhauer and Schubert, 1914: 297 (Somoleptus; catalog).

  • — Silvestri, 1946: 58 (Somoleptus; adult and larval characters; illustrations; Brazil).

  • Distribution: Brazil.

linearis Cameron, 1918: 82 (Somoleptus; Type locality: Singapore: Bukit Panjang).

  • — Cameron, 1921a: 375, 404 (Somoleptus; characters; Singapore).

  • — Scheerpeltz, 1933: 1304 (Somoleptus; catalog).

  • Distribution: Singapore.

nitidus Sharp, 1876: 204 (Leptacinus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 297 (Somoleptus; catalog).

  • Distribution: Brazil.

obscurus Sharp, 1885: 496 (Somoleptus; Type locality: Panama, Peña Blanca 3000 to 4000 feet).

  • — Bernhauer and Schubert, 1914: 297 (Somoleptus; catalog).

  • Distribution: Panama.

obsoletus Sharp, 1885: 496 (Somoleptus; Type locality: Guatemala, near the city).

  • — Bernhauer and Schubert, 1914: 297 (Somoleptus; catalog).

  • Distribution: Guatemala.

pallipes Sharp, 1885: 497 (Somoleptus; Type locality: Guatemala, San Geronimo).

  • — Bernhauer and Schubert, 1914: 297 (Somoleptus; catalog).

  • Distribution: Guatemala.

parvulus Sharp, 1885: 496 (Somoleptus; Type locality: Guatemala, near the city, San Geronimo; Panama, Los Remedios, Tolé).

  • — Bernhauer and Schubert, 1914: 297 (Somoleptus; catalog).

  • Distribution: Guatemala, Panama.

pulcher Bernhauer, 1935c: 90 (Somoleptus; Type locality: Brasilien: Rio de Janeiro).

  • Distribution: Brazil.

punctulatus Sharp, 1885: 497 (Somoleptus; Type locality: Guatemala, near the city, Aceituno).

  • — Bernhauer and Schubert, 1914: 297 (Somoleptus; catalog).

  • Distribution: Guatemala.

sparsus Sharp, 1885: 497 (Somoleptus; Type locality: Guatemala, Chiacam in Vera Paz).

  • — Bernhauer and Schubert, 1914: 298 (Somoleptus; catalog).

  • Distribution: Guatemala.

subtilis Erichson, 1839b: 337 (Leptacinus; Type locality: Columbia).

  • — Sharp, 1885: 495 (Somoleptus).

  • — Fauvel, 1891: 105 (Somoleptus; Venezuela).

  • — Bernhauer and Schubert, 1914: 298 (Somoleptus; catalog).

  • Distribution: Colombia.

unicolor Cameron, 1922, see: Lithocharodes.

Spaniolinus [1 species; Oriental Region]

Spaniolinus Bernhauer, 1916h: 421 (species included: piceorufus). Type species: Spaniolinus piceorufus Bernhauer, fixed by monotypy.

  • — Scheerpeltz, 1933: 1305 (world catalog supplement).

  • — Blackwelder, 1952: 354 (type species: piceorufus).

piceorufus Bernhauer, 1916h: 422 (Spaniolinus; Type locality: Negros: Cuernos Mts.).

  • — Scheerpeltz, 1933: 1305 (Spaniolinus; catalog).

  • Distribution: Philippines.

Stenistoderus [19 species; Nearctic, Palaearctic, and Ethiopian Regions]

Stenistoderus Jacquelin du Val, 1856: 33 (species included: nothus). Type species: Leptacinus nothus Erichson, fixed by monotypy.

  • — Schaum, 1859: 27 (synonym of Leptolinus).

  • — Gemminger and Harold, 1868: 604 (synonym of Leptolinus).

  • — Jacquelin du Val, 1868: 66 (subgenus of Leptacinus; catalog; European species).

  • — Fauvel, 1873b: 91 [= 1873a: 375] (synonym of Leptolinus).

  • — Ganglbauer, 1895: 488 (synonym of Leptolinus).

  • — Bernhauer and Schubert, 1914: 291 (synonym of Leptolinus).

  • — Blackwelder, 1952: 358 (type species: nothus).

  • — Moore, 1963a: 94 (characters).

  • — Székessy, 1963: 48 (characters).

  • — Coiffait, 1972: 188 (synonym of Leptolinus).

  • — Moore and Legner, 1975: 91 (catalog; North America north of Mexico).

  • — Moore and Legner, 1979: 73 (characters; notes).

  • — Smetana, 1982: 117 (characters; habitus; revision of species of North America north of Mexico).

  • — Shibata, 1983: 75 (checklist of species of Japan).

  • — Downie and Arnett, 1996: 399 (characters).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 391 (1 Nearctic species; notes; characters in key).

Leptolinus Kraatz, 1857d: 647 (species included: nothus; [Note: Kraatz included the species listed in “Leptacinus Fam. III” of Erichson, 1839b: 338, but did not include any available names by name, which is required by Article 67.2.1. Redtenbacher, 1858: 984, seems to have been this first to have included species by name when he cited nothus]). Type species: Leptacinus nothus Erichson, fixed by subsequent monotypy.

  • — Redtenbacher, 1858: 983 (characters).

  • — Schaum, 1859: 27 (catalog; European species).

  • — LeConte, 1861a: 65 (characters).

  • — LeConte, 1863: 24 (list of North American species).

  • — Gemminger and Harold, 1868: 604 (catalog).

  • — Fauvel, 1873b: 91 [= 1873a: 377] (characters).

  • — Fauvel, 1875a: XXIV [= 1875b: 226] (catalog).

  • — Redtenbacher, 1874: 220 (characters).

  • — Mulsant and Rey, 1877a: 331 (characters).

  • — LeConte, 1880: 171 (characters; key to North American species).

  • — Duvivier, 1883: 157 (catalog).

  • — Heyden, Reitter, and Weise, 1891: 105 (list of species of Europe and Caucasus).

  • — Ganglbauer, 1895: 488 (characters).

  • — Casey, 1906: 372, 418 (characters).

  • — Heyden, Reitter, and Weise, 1906: 160 (list of species of Europe and Caucasus).

  • — Reitter, 1908: 109, 110 (characters; key to species).

  • — Reitter, 1909: 134 (characters).

  • — Blatchley, 1910: 398 (characters; key to Indiana species).

  • — Bernhauer and Schubert, 1914: 291 (world catalog; 4 species).

  • — Leng, 1920: 106 (catalog of North American species).

  • — Winkler, 1925: 370 (catalog for Palaearctic region).

  • — Porta, 1926: 86 (characters; key to species of Italy).

  • — Portevin, 1929: 370 (key to French species).

  • — Scheerpeltz, 1933: 1299 (world catalog supplement).

  • — Blackwelder, 1952: 216 (type species: nothus).

  • — Coiffait, 1956a: 43, 51 (characters; key to French species).

  • — Kocher, 1958: 137 (checklist of species; Morocco).

  • — Smetana, 1959a: 208 (list of species collected in Albania).

  • — Ferreira, 1962a: 14 (list of Portuguese species).

  • — Coiffait, 1968: 128, 145 (characters; notes).

  • — Coiffait, 1972: 188 (characters; type species: nothus; cited as valid genus; key to species of western Palaearctic region; references).

  • — Tikhomirova, 1973a: 181 (checklist of species of USSR).

  • — Smetana, 1982: 116 (synonym of Stenistoderus).

  • — Bordoni, 1982: 168 (characters; habitus; key to species of Italy).

  • — Tóth, 1989: 7 (characters; species of Hungary).

  • — Ciceroni and Zanetti, 1995: 25 (list of species of Italy).

Leptoglenus Reitter, 1900b: 227 (species included: caecus). Type species: Leptolinus caecus Reitter, fixed by monotypy.

  • — Reitter, 1908: 109 (characters).

  • — Bernhauer and Schubert, 1914: 291 (synonym of Leptolinus).

  • — Scheerpeltz, 1933: 1299 (subgenus of Leptolinus).

  • — Blackwelder, 1952: 216 (type species: coecus).

  • — Coiffait, 1972: 188, 189, 193 (subgenus of Leptolinus; characters; type species: caecus; key to species of western Palaearctic region; references).

  • — Bordoni, 1982: 168, 172 (subgenus of Leptolinus; characters).

Paraleptolinus Coiffait, 1956a: 51 (subgenus of Leptolinus; species included: cephalotes). Type species: Leptolinus cephalotes Kraatz, fixed by monotypy.

  • — Coiffait, 1972: 188 (synonym of Leptolinus subgenus Leptoglenus).

  • — Bordoni, 1982: 168, 172 (synonym of Leptolinus).

algericus Coiffait, 1964a: 506 (Leptolinus; Type locality: Algérie, Kerrata).

  • — Coiffait, 1972: 196 (Leptolinus; characters; Algeria).

  • Distribution: Algeria.

andalusiacus Coiffait, 1956, see: subangulatus Reitter, 1908.

arabicus Coiffait, 1981b: 238 (Leptolinus; Type locality: Arabie Saoudite, Hofuf).

  • Distribution: Saudi Arabia.

armeniacus Coiffait, 1966, see: cephalotes Kraatz, 1858.

berberus Coiffait, 1971, see: cephalotes Kraatz, 1858.

brevicornis Latreille, 1804, see: nothus Erichson, 1839.

caecus Reitter, 1900b: 227 (Leptolinus; subgenus Leptoglenus; Type locality: Samarkand).

  • — Reitter, 1908: 111 (Leptoglenus; characters; Samarkand).

  • — Bernhauer and Schubert, 1914: 291 (Leptolinus; cited as coecus; catalog).

  • — Scheerpeltz, 1933: 1299 (Leptolinus; cited as coecus; catalog).

  • — Coiffait, 1972: 199 (Leptolinus; subgenus Leptoglenus; characters; Turkestan).

  • Distribution: Uzbekistan.

caucasicus Fleischer, 1914: 144 (Leptolinus; Type locality: Kaukasien).

  • — Scheerpeltz, 1933: 1299 (Leptolinus; catalog).

  • — Coiffait, 1972: 199 (Leptolinus; subgenus Leptoglenus; characters; Caucasus).

  • Distribution: Caucasus.

cephalotes Kraatz, 1858a: 65 (Leptolinus; Type locality: Zante).

  • — Fauvel, 1867c: 114 [= 1868b: 213] (Leptolinus; variety of nothus).

  • — Fauvel, 1873b: 92 [= 1873a: 378] (Leptolinus; synonym of nothus).

  • — Ganglbauer, 1895: 489 (Leptolinus; variety of nothus).

  • — Reitter, 1908: 110 (Leptolinus; variety of nothus; Greece; Mesopotamia; Algeria).

  • — Bernhauer and Schubert, 1914: 291 (Leptolinus; variety of nothus).

  • — Scheerpeltz, 1933: 1300 (Leptolinus; variety of nothus).

  • — Coiffait, 1956a: 52 (Leptolinus; subgenus Paraleptolinus; characters in key; Lebanon; Turkey; Dalmatia; Algeria; Tunisia; Morocco; Sicily; Sardinia; Italy; Greece; Mesopotamia).

  • — Scheerpeltz, 1961: 6 (Leptolinus; subgenus Leptolinus; Iran).

  • — Coiffait, 1965e: 119 (Leptolinus; Turkey).

  • — Coiffait, 1972: 197 (Leptolinus; subgenus Leptoglenus; characters; references; Mediterranean region; Mesopotamia; Morocco; Lebanon; Turkey; Greece; Dalmatia; France).

  • — Bordoni, 1982: 173 (Leptolinus; subgenus Leptoglenus; characters; Italy).

  • — Sparacio, 1995: 156 (Leptolinus; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 25 (Leptolinus; Italy).

  • Distribution: Tunisia, Algeria, Morocco, France, Italy, Croatia, Greece, Turkey, Lebanon, Iraq, Iran, Armenia.

armeniacus Coiffait, 1966f: 197 (Leptolinus; subgenus Leptoglenus; subspecies of cephalotes; Type locality: SSSR, Armeniia, okrestnosti, Erevana).

  • — Coiffait, 1972: 197 (Leptolinus; subgenus Leptoglenus; subspecies of cephalotes; characters; Armenia).

berberus Coiffait, 1971: 436 (Leptolinus; subgenus Leptoglenus; subspecies of cephalotes; Type locality: St. Charles près de Philippeville, Algérie).

  • — Coiffait, 1972: 197 (Leptolinus; subgenus Leptoglenus; subspecies of cephalotes; characters; Algeria).

congoensis Bernhauer, 1928f: 118 (Leptolinus; Type locality: Luebo; Irebu).

  • — Scheerpeltz, 1933: 1299 (Leptolinus; catalog).

  • Distribution: Congo.

cribripennis Fairmaire and Germain, 1861, see: Neoleptacinus.

fuscipennis Cameron, 1951c: 398 (Leptolinus; Type locality: Kenya Colony: Naivasha).

  • Distribution: Kenya.

grandiceps LeConte, 1863, see: Stictolinus.

nanus Iablokov-Khnzorian, 1961: 143 (Leptolinus; Type locality: Armenie: Aïguerlitsche, env. d'Etschmiadzin, région désertique).

  • — Coiffait, 1972: 198 (Leptolinus; subgenus Leptoglenus; characters; Armenia).

  • Distribution: Armenia.

nigripennis LeConte, 1863, see: Lithocharodes.

nothus Erichson, 1839b: 338 (Leptacinus; Type locality: Croatia meridionalis; Mesopotamia).

  • — P. Lucas, 1846: 108 (Leptacinus; notes; Algeria).

  • — Hochhuth, 1849: 111 (Leptacinus; characters; Asovischen Meeres).

  • — Fairmaire and Laboulbène, 1856: 504 (Leptacinus; characters; France).

  • — Redtenbacher, 1857: 984 (Leptolinus; characters; Austria).

  • — Kraatz, 1858a: 65 (Leptolinus; Croatia; Greece).

  • — Fauvel, 1873b: 92 [= 1873a: 378] (Leptacinus; characters; France; Croatia; Europe; Mediterranean region; Russia; Mesopotamia).

  • — Redtenbacher, 1874: 220 (Leptolinus; characters).

  • — Mulsant and Rey, 1877a: 334 (Leptolinus; characters; France).

  • — Fauvel, 1878: 116 [= 1878a: 36] (Leptolinus; Algeria).

  • — Fauvel, 1886: 48 [= 1886a: 40] (Leptolinus; Morocco; Algeria).

  • — Ganglbauer, 1895: 489 (Leptolinus; characters; France; Croatia; Mediterranean region).

  • — Fauvel, 1897d: 291 (Leptolinus; Barbary; Canaries).

  • — Fauvel, 1902b: 100 (Leptolinus; Barbary; Canary Islands).

  • — Sainte-Claire Deville, 1906: 93 (Leptolinus; Corsica).

  • — Montandon, 1908: 79 (Leptolinus; Romania).

  • — Reitter, 1908: 110 (Leptolinus; characters; Croatia; Dalmatia; Balkan peninsula; France).

  • — Bernhauer and Schubert, 1914: 291 (Leptolinus; catalog).

  • — Portevin, 1929: 370 (Leptolinus; characters; France).

  • — Scheerpeltz, 1933: 1300 (Leptolinus; catalog).

  • — Coiffait, 1956a: 52 (Leptolinus; characters in key; France; Spain).

  • — Focarile, 1964: 57 (Leptolinus; Italy).

  • — Scheerpeltz, 1965b: 399 (Leptolinus; Sicily).

  • — Scheerpeltz, 1965c: 504 (Leptolinus; Sicily).

  • — Coiffait, 1972: 194 (Leptolinus; characters; references; Mediterranean region; Balkans; Canary Islands; France).

  • — Coiffait, 1973a: 273 (Leptolinus; Morocco).

  • — Bordoni, 1982: 172 (Leptolinus; characters; Italy).

  • — Tóth, 1989: 8 (Leptolinus; characters; Hungary).

  • — Ciceroni, 1993: 68 (Leptolinus; Italy).

  • — J. Li, 1993: 32 (Stenistoderus; China).

  • — Hernández, Outerelo, and Gamarra, 1994: 190 (Leptolinus; checklist of species; Canary Islands).

  • — Sparacio, 1995: 156 (Leptolinus; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 25 (Leptolinus; Italy).

  • — Bordoni, 1995: 85 (Stenistoderus; ecological notes; Italy).

  • Distribution: Algeria, Morocco, Canary Islands, Europe, Iraq, Russia.

brevicornis Latreille, 1804: 342 (Lathrobium; [Note: Bernhauer and Schubert listed this name as a junior synonym of Medon castaneus Gravenhorst (1912: 234) and Leptolinus nothus Erichson (1914: 291)]; Type locality: France).

  • — Bernhauer and Schubert, 1914: 291 (Leptolinus; synonym of nothus).

sareptanus Stierlin, 1867: 219 (Lathrobium; Type locality: Sarepta).

  • — Fauvel, 1873b: 92 [= 1873a: 378] (Leptolinus; synonym of nothus).

  • — Ganglbauer, 1895: 489 (Leptolinus; synonym of nothus).

  • — Reitter, 1908: 110 (Leptolinus; synonym of nothus variety versicolor).

  • — Bernhauer and Schubert, 1914: 291 (Leptolinus; synonym of nothus).

parcus LeConte, 1863, see: Hesperolinus.

pusio LeConte, 1880, see: Microlinus.

rubripennis LeConte, 1880: 171 (Leptolinus; Type locality: District of Columbia; [Note: See lectotype designation by Smetana, 1982]. Originally cited from Middle and western States).

  • — Casey, 1906: 418 (Leptolinus; characters; Ohio; Iowa; Texas).

  • — Blatchley, 1910: 398 (Xantholinus; characters; Indiana).

  • — Bernhauer and Schubert, 1914: 291 (Leptolinus; catalog).

  • — Smetana, 1982: 117 (Stenistoderus; lectotype designation: D.C. [= District of Columbia]; characters; Ontario; Alabama; Arkansas; Illinois; Indiana; Iowa; Kansas; Kentucky; Louisiana; Maryland; Michigan; Missouri; Nebraska; New Jersey; New York; Ohio; South Carolina; Tennessee; Texas; Virginia; Wisconsin).

  • — Smetana, 1988a: 527 (Stenistoderus; collecting notes; Ontario; Mississippi).

  • — Smetana, 1990c: 85 (Stenistoderus; Kentucky; Oklahoma).

  • — Campbell and Davies, 1991: 123 (Stenistoderus; checklist of species; Canada).

  • — Downie and Arnett, 1996: 399 (Stenistoderus; characters; USA).

  • Distribution: Canada, USA.

ruficollis LeConte, 1863, see: Lithocharodes.

rufus Coiffait, 1956a: 53 (Leptolinus; Type locality: Liban: Hasbaya au bord du Jourdain).

  • — Coiffait, 1972: 196 (Leptolinus; characters; Lebanon).

  • Distribution: Lebanon.

sareptanus Stierlin, 1867, see: nothus Erichson, 1839.

sinicus Bordoni, 2000: 123 (Stenistoderus; Type locality: N-China: Beijing, Yingtaogou).

  • Distribution: China.

striolatus Coiffait, 1968: 145 (Leptolinus; subgenus Leptolinus; Type locality: Gabon: Belinga).

  • Distribution: Gabon.

subangulatus Reitter, 1908: 111 [= 1908a: 14] (Leptolinus; variety of nothus; Type locality: Andalusien).

  • — Bernhauer and Schubert, 1914: 291 (Leptolinus; variety of nothus).

  • — Scheerpeltz, 1933: 1300 (Leptolinus; variety of nothus).

  • — Coiffait, 1972: 195 (Leptolinus; cited as valid species; characters; references; Iberia; Morocco).

  • — Coiffait, 1973a: 273 (Leptolinus; Morocco).

  • Distribution: Spain, Morocco.

andalusiacus Coiffait, 1956a: 52 (Leptolinus; Type locality: Espagne: Prov. de Cadix, Lagune de la Janda).

  • — Coiffait, 1972: 195 (Leptolinus; synonym of subangulatus).

subnitidiceps Bernhauer, 1937b: 608 (Leptolinus; Type locality: N. O. Upogoro).

  • Distribution: Tanzania.

syriacus Coiffait, 1956a: 53 (Leptolinus; Type locality: Syrie: Damas, bords du Barada).

  • — Coiffait, 1972: 195 (Leptolinus; characters; Syria).

  • Distribution: Syria.

turcicus Coiffait, 1956b: 142 (Leptolinus; Type locality: Turquie d'Asie, Villayet de Balikesir: bords du Lac Manyas).

  • — Coiffait, 1972: 196 (Leptolinus; characters; Turkey).

  • Distribution: Turkey.

usagarae Bernhauer, 1908c: 107 (Leptolinus; Type locality: Deutsch-Ostafrika: Usagara, Kilossa).

  • — Bernhauer and Schubert, 1914: 292 (Leptolinus; catalog).

  • Distribution: Tanzania.

versicolor Solsky, 1871a: 181 (Leptolinus; Type locality: Astrakhan).

  • — Fauvel, 1873b: 92 [= 1873a: 378] (Leptolinus; synonym of nothus).

  • — Reitter, 1908: 110 (Leptolinus; variety of nothus).

  • — Bernhauer and Schubert, 1914: 291 (Leptolinus; synonym of nothus).

  • — Scheerpeltz, 1933: 1300 (Leptolinus; not synonym of nothus).

  • — Coiffait, 1972: 197 (Leptolinus; characters; Caucasus; Turkestan; Tajikistan).

  • — Ushakov, 1989: 129 (Leptolinus; characters; Russia).

  • Distribution: Russia, Caucasus, Tajikistan.

Stenolinus [1 species; Neotropical Region]

Stenolinus Bierig, 1937: 273 (species included: macrothrichus). Type species: Stenolinus macrothrichus Bierig, fixed by original designation and monotypy.

  • — Blackwelder, 1943: 503 (characters; type species: macrothrichus).

  • — Blackwelder, 1944: 145 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 358 (type species: macrothrichus).

macrothrichus Bierig, 1937: 275 (Stenolinus; Type locality: Cuba: Sierra del Rosario, El Rangel).

  • — Blackwelder, 1943: 503 (Stenolinus; characters; Cuba).

  • — Legner and Moore, 1977: 176 (Stenolinus; found under bark).

  • Distribution: West Indies.

Stictolinus [4 species; Nearctic Region]

Stictolinus Casey, 1906: 369, 404 (species included: grandiceps, aequalis, scolopaceus, franciscanus). Type species: Leptolinus grandiceps LeConte, fixed by subsequent designation by R. Lucas, 1920.

  • — Bernhauer and Schubert, 1914: 294 (world catalog; 4 species).

  • — Leng, 1920: 106 (catalog of North American species).

  • — Blackwelder, 1952: 361 (type species).

  • — Hatch, 1957: 234, 239 (characters; key to species of Pacific Northwest).

  • — Moore, 1963a: 94 (characters).

  • — Moore and Legner, 1975: 92 (catalog; North America north of Mexico).

  • — Moore and Legner, 1979: 74 (characters; notes).

  • — Smetana, 1982: 175 (characters; habitus; revision of and key to species of North America north of Mexico).

  • — Downie and Arnett, 1996: 400 (characters).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 392 (4 Nearctic species; notes; characters in key).

aequalis Casey, 1906, see: grandiceps LeConte, 1863.

flavipes LeConte, 1863a: 41 (Leptacinus; Type locality: Middle and western States; [Note: See lectotype designation by Smetana, 1982]).

  • — Casey, 1906: 402 (Leptacinodes; characters; New York; New Jersey; Iowa).

  • — Bernhauer and Schubert, 1914: 292 (Leptacinus; synonym of batychrus).

  • — Smetana, 1982: 177 (Stictolinus; lectotype designation: Locality not cited; characters; Nova Scotia; Ontario; Quebec; Connecticut; Illinois; Indiana; Maine; Massachusetts; Michigan; New Hampshire; New Jersey; New York; Pennsylvania; Rhode Island; Wisconsin).

  • — Smetana, 1988a: 553 (Stictolinus; New Hampshire).

  • — Campbell and Davies, 1991: 124 (Stictolinus; checklist of species; Canada).

  • — Downie and Arnett, 1996: 400 (Stictolinus; characters; USA).

  • Distribution: Canada, USA.

scolopaceus Casey, 1906: 406 (Stictolinus; Type locality: Rhode Island; [Note: See lectotype designation by Smetana, 1982]. Originally cited from Massachusetts, New York and Pennsylvania to Iowa and Wisconsin [Bayfield]).

  • — Bernhauer and Schubert, 1914: 294 (Stictolinus; catalog).

  • — Smetana, 1982: 177 (Stictolinus; lectotype designation: R.I. [Rhode Island]; synonym of flavipes).

franciscanus Casey, 1906: 406 (Stictolinus; Type locality: California: Lake Co.; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 294 (Stictolinus; catalog).

  • — Hatch, 1957: 239 (Stictolinus; characters; Washington; Oregon).

  • — Smetana, 1982: 181 (Stictolinus; lectotype designation; characters; California; Oregon).

  • — Smetana, 1988a: 553 (Stictolinus; California).

  • Distribution: USA.

grandiceps LeConte, 1863a: 42 (Leptolinus; Type locality: San Francisco, California; [Note: See lectotype designation by Smetana, 1982]).

  • — LeConte, 1880: 168 (Leptacinus; characters).

  • — Casey, 1906: 405 (Stictolinus; characters; California).

  • — Bernhauer and Schubert, 1914: 294 (Stictolinus; catalog).

  • — Smetana, 1982: 175 (Stictolinus; lectotype designation; characters; California; Oregon).

  • — Smetana, 1988a: 553 (Stictolinus; California).

  • Distribution: USA.

aequalis Casey, 1906: 405 (Stictolinus; Type locality: California: Siskiyou Co.; [Note: See lectotype designation by Smetana, 1982]. Also cited from California: Truckee).

  • — Bernhauer and Schubert, 1914: 294 (Stictolinus; catalog).

  • — Hatch, 1957: 239 (Stictolinus; characters; Washington; Oregon).

  • — Smetana, 1982: 175 (Stictolinus; lectotype designation: Siskiyou Co., California; synonym of grandiceps).

nudus Smetana, 1982: 182 (Stictolinus; Type locality: California: Mokel. Hill).

  • Distribution: USA.

scolopaceus Casey, 1906, see: flavipes LeConte, 1863.

Tesba [7 species; Neotropical Region]

Tesba Sharp, 1876: 194 (species included: gigas, laticornis). Type species: Tesba gigas Sharp, fixed by subsequent designation by Blackwelder, 1952: 381 [Note: Blackwelder stated that R. Lucas, 1920: 633, designated the type species but the designation was ambiguous because two nominal species were cited as “gigas et laticornis”].

  • — Duvivier, 1883: 158 (catalog).

  • — Bernhauer and Schubert, 1914: 314 (world catalog; 6 species).

  • — Cameron, 1925: 52 (catalog of Indian species).

  • — Scheerpeltz, 1933: 1319 (world catalog supplement).

  • — Blackwelder, 1944: 149 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Scheerpeltz, 1960c: 99 (key to species).

belti Sharp, 1876b: 431 (Tesba; Type locality: Chontales).

  • — Sharp, 1885: 475 (Tesba; Nicaragua).

  • — Bernhauer and Schubert, 1914: 314 (Tesba; catalog).

  • — Scheerpeltz, 1960c: 100 (Tesba; characters; Brazil).

  • Distribution: Nicaragua, Brazil.

collaris Eppelsheim, 1895b: 399 (Tesba; Type locality: Guatemala).

  • — Bernhauer and Schubert, 1914: 314 (Tesba; catalog).

  • — Scheerpeltz, 1960c: 99 (Tesba; characters; Guatemala).

  • Distribution: Guatemala.

gigas Sharp, 1876: 195 (Tesba; Type locality: St. Paulo).

  • — Bernhauer and Schubert, 1914: 314 (Tesba; catalog; Peru).

  • — Scheerpeltz, 1960c: 99 (Tesba; characters; Brazil).

  • Distribution: Brazil, Peru.

indica Eppelsheim, 1895, see: Paratesba.

iheringi Bernhauer, 1908d: 325 (Tesba; Type locality: Brasilien: S. Paulo, Ypiranga).

  • — Bernhauer and Schubert, 1914: 314 (Tesba; catalog).

  • — Scheerpeltz, 1960c: 100 (Tesba; characters; Brazil; Paraguay; Argentina).

  • Distribution: Brazil, Paraguay, Argentina.

laticornis Sharp, 1876: 195 (Tesba; Type locality: Tunantins).

  • — Bernhauer and Schubert, 1914: 314 (Tesba; catalog).

  • — Scheerpeltz, 1960c: 100 (Tesba; characters; Brazil).

  • Distribution: Brazil.

lizeri Bernhauer, 1923b: 58 (Tesba; Type locality: Bolivien: Lagunillas).

  • — Scheerpeltz, 1933: 1319 (Tesba; catalog).

  • — Scheerpeltz, 1960c: 100 (Tesba; characters; Bolivia).

  • Distribution: Bolivia.

ohausi Wendeler, 1930: 191 (Tesba; Type locality: Brasilien, São Paulo, Ypiranga).

  • — Scheerpeltz, 1933: 1319 (Tesba; catalog).

  • — Scheerpeltz, 1960c: 100 (Tesba; characters; Brazil).

  • Distribution: Brazil.

Thyreocephalus [75 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions]

Thyreocephalus Guérin-Méneville, 1844: 10 (species included: jekelii, lynceus). Type species: Thyreocephalus jekelii Guérin-Méneville, fixed by subsequent designation by R. Lucas, 1920: 640.

  • — Lacordaire, 1854: 67 (synonym of Scytalinus).

  • — Gemminger and Harold, 1868: 600 (catalog).

  • — Sharp, 1885: 498 (characters; Mexican and Central American species).

  • — Bernhauer and Schubert, 1914: 313 (world catalog; 22 species).

  • — Cameron, 1921a: 354, 404 (characters; key to Singapore species; catalog).

  • — Cameron, 1925: 52 (catalog of Indian species).

  • — Scheerpeltz, 1933: 1318 (world catalog supplement; cited as Thyroecephalus).

  • — Steel, 1938b: 55 (notes; characters to distinguish from Eulissus).

  • — Blackwelder, 1943: 490 (characters; type species: jekelii).

  • — Blackwelder, 1944: 149 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 390 (type species).

  • — Tottenham, 1956: 258 (key to African species).

  • — Coiffait, 1968: 128, 152 (characters; key to species of Gabon; notes).

  • — Moore and Legner, 1979: 75 (characters; notes).

  • — Smetana, 1982: 66 (characters; habitus; revision of and key to species of North America north of Mexico).

  • — Dettner, 1987: 30 (defensive glands).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 392 (2 Nearctic species; notes; characters in key).

Linidius Sharp, 1876: 196 (species included: recticollis; tenuipes; extremus). Type species: Linidius recticollis Sharp, fixed by original designation.

  • — Duvivier, 1883: 158 (catalog).

  • — Bernhauer and Schubert, 1914: 314 (world catalog; 1 species).

  • — Steel, 1938b: 56 (synonym of Thyreocephalus).

  • — Blackwelder, 1943: 490 (synonym of Thyreocephalus; type species: recticollis).

  • — Blackwelder, 1952: 221 (type species: recticollis).

  • — Smetana, 1982: 66 (synonym of Thyreocephalus).

Saurohypnus Sharp, 1885: 501 (species included: scutellaris). Type species: Saurohypnus scutellaris Sharp, fixed by monotypy.

  • — Casey, 1906: 366, 373 (characters).

  • — Bernhauer and Schubert, 1914: 299 (world catalog; 3 species).

  • — Leng, 1920: 105 (catalog of North American species).

  • — Blackwelder, 1944: 146 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 344 (type species: scutellaris).

  • — Moore, 1963a: 94 (characters).

  • — Moore and Legner, 1975: 90 (catalog; North America north of Mexico).

  • — Smetana, 1977c: 348 (synonym of Thyreocephalus).

  • — Smetana, 1982: 66 (synonym of Thyreocephalus).

Indoscytalinus Heller, 1900: 5 (subgenus of Xantholinus; species included: albicornis). Type species: Xantholinus (Indoscytalinus) albicornis Heller, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 313 (synonym of Thyreocephalus).

  • — Cameron, 1932: 33 (characters; cited as Indoscitalinus; key to species of British India).

  • — Scheerpeltz, 1933: 1318 (subgenus of Thyreocephalus).

  • — Steel, 1938b: 55 (synonym of Thyreocephalus).

  • — Blackwelder, 1943: 490 (synonym of Thyreocephalus; cited as Indoscitalinus; type species: albicornis).

  • — Blackwelder, 1952: 199 (synonym of Thyreocephalus; type species: albicornis).

  • — Shibata, 1973a: 126 (valid genus; characters).

  • — Smetana, 1982: 66 (cited as Indoscitalinus; synonym of Thyreocephalus).

  • — Hammond, 1984: 206 (cited as valid genus; checklist of species of Borneo).

Dinoxantholinus Heller, 1910: 7 (species included: prodigiosus). Type species: Dinoxantholinus prodigiosus Heller, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 309 (world catalog; 1 species).

  • — Scheerpeltz, 1933: 1315 (world catalog supplement).

  • — Steel, 1938b: 55 (synonym of Thyreocephalus).

  • — Blackwelder, 1943: 490 (synonym of Thyreocephalus; type species: prodigiosus).

  • — Blackwelder, 1952: 128 (type species: prodigiosus).

  • — Smetana, 1982: 66 (synonym of Thyreocephalus).

albertisi Fauvel, 1877: 246 (Xantholinus; Type locality: Australie, Cap York, Somerset; Nouvelle Guinée, aux îles Arou, à Ternate et à Célèbes).

  • — Fauvel, 1878f: 245 (Xantholinus; characters; Aru Islands; Moluccas; New Guinea; Sulawesi; Australia).

  • — Fauvel, 1879: 93 (Xantholinus; Dorei).

  • — Olliff, 1887: 489 (Xantholinus; characters; Australia).

  • — Bernhauer and Schubert, 1914: 300 (Xantholinus; catalog).

  • — Lea, 1925: 218 (Xantholinus; Australia).

  • — Cameron, 1931: 360 (Thyreocephalus; New Guinea).

  • — Scheerpeltz, 1933: 1318 (Thyreocephalus; catalog).

  • — Cameron, 1937a: 100 (Indoscytalinus, cited as Indoscitalinus; New Guinea).

  • — Marucci and Clancy, 1952 (Thyreocephalus; life history and biological notes; released in Hawaii).

  • — Moore, 1975c: 107 (Thyreocephalus; Hawaii).

  • — Nishida, 1994: 74 (Thyreocephalus; Hawaii).

  • — Nishida, 1997: 63 (Thyreocephalus; Hawaii).

  • Distribution: Australia, New Guinea, Indonesia, Hawaii.

albicornis Heller, 1900: 5 (Xantholinus; subgenus Indoscytalinus; Type locality: Celebes septentrionalis, mons Masarang).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • Distribution: Indonesia.

anachoreta Erichson, 1839b: 316 (Xantholinus; Type locality: Bengala).

  • — Kraatz, 1859: 102 (Xantholinus; characters; Ceylon; India borealis).

  • — Cocquerel, 1866: 318 (Xantholinus; characters; Réunion; Bengal).

  • — Fauvel, 1867b: 68 [= 1868a: 72] (Xantholinus; characters; Réunion Islands; Ceylon; Bengal).

  • — Bernhauer, 1902b: 32 (Eulissus; Ceylon).

  • — Fauvel, 1903: 155 (Xantholinus; India; Ceylon; Burma; Annam; Tonkin; China; Sumatra; Mauritius; Réunion; Madagascar).

  • — Fauvel, 1904a: 56 (Xantholinus; Belgaum; Nagargali; Nilghiris; Bírmanie; Annam; Tonkin; China; Sumatra; Maurice; Réunion; Madagascar).

  • — Fauvel, 1905e: 83 (Xantholinus; Java).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog; Philippines).

  • — Bernhauer, 1928: 15 (Eulissus; Sumatra).

  • — Cameron, 1932: 37 (Indoscytalinus, cited as Indoscitalinus; characters; Ceylon; India).

  • — Scheerpeltz, 1933: 1316 (Eulissus; catalog).

  • — Scheerpeltz, 1933: 1318 (Thyreocephalus; catalog).

  • — Cameron, 1933d: 385 (Indoscytalinus, cited as Indoscitalinus; Sumatra).

  • — Cameron, 1937: 3 (Indoscytalinus, cited as Indoscitalinus; Java).

  • — Paulian, 1941: 216 (Thyreocephalus; larval characters).

  • — Jarrige, 1951: 133 (Indoscytalinus; Indochina).

  • — Cerruti, 1951: 15 (Indoscytalinus, cited as Indoscitalinus; India).

  • — Scheerpeltz, 1954: 112 (Thyreocephalus; subgenus Indoscytalinus, cited as Indoscitalinus; India; Ceylon).

  • — Scheerpeltz, 1976a: 86 (Indoscytalinus, cited as Indoscitalinus; Nepal).

  • — Coiffait, 1982a: 232 (Indoscytalinus, cited as Indoscitalinus; India).

  • — Biswas, 1986: 127 (Indoscytalinus, cited as Indoscitalinus; India).

  • — Lecoq, 1990: 188 (Thyreocephalus; Mascarene Islands).

  • Distribution: Madagascar, Mascarene Islands, Sri Lanka, India, Nepal, Myanmar, Vietnam, Indonesia, Philippines, China.

morio Motschulsky, 1858a: 207 (Xantholinus; Type locality: Indes orientales).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; synonym of anachoreta).

  • — Cameron, 1932: 37 (Indoscytalinus, cited as Indoscitalinus; synonym of anachoreta).

  • — Scheerpeltz, 1933: 1318 (Thyreocephalus; synonym of anachoreta).

hongkongensis Redtenbacher, 1867: 28 (Xantholinus; Type locality: Hongkong).

  • — Fauvel, 1903: 155 (Xantholinus; synonym of anachoreta).

  • — Fauvel, 1904a: 56 (Xantholinus; synonym of anachoreta).

  • — Fauvel, 1905e: 83 (Xantholinus; synonym of anachoreta).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; synonym of anachoreta).

  • — Cameron, 1932: 37 (Indoscytalinus, cited as Indoscitalinus; synonym of anachoreta).

  • — Scheerpeltz, 1933: 1318 (Thyreocephalus; synonym of anachoreta).

analis Schubert, 1911: 14 (Thyreocephalus; Type locality: Yungas de la Paz, Bolivia).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • Distribution: Bolivia.

anguliceps Bernhauer, 1916c: 27 (Thyreocephalus; Type locality: West-Kolumbien: Umgebung Cali, Alto de las cruces, 2200 m).

  • — Scheerpeltz, 1933: 1318 (Thyreocephalus; catalog).

  • Distribution: Colombia.

annulatus Fauvel, 1895: 241 (Xantholinus; Type locality: Bírmanie, Bhamo; Carin Cheba, 500–1000 m; Tenasserim, mont Mooleyit, 1800–1900 m; Inde; Pulo-Penang; Sumatra, Ajer, Mantcior; Tandjong Morawa Serdang, Sungei-Bulu; Borneo, Sarawak).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • — Cameron, 1921a: 375, 404 (Thyreocephalus; [Note: Misspelling: Cited on page 375 as annularis; Scheerpeltz, 1933: 1318 cited annularis Cameron as a synonym of annulatus]; characters; Singapore).

  • — Cameron, 1932: 36 (Indoscytalinus, cited as Indoscitalinus; characters; Burma).

  • — Scheerpeltz, 1933: 1318 (Thyreocephalus; catalog).

  • — Cameron, 1933d: 385 (Indoscytalinus, cited as Indoscitalinus; Sumatra).

  • Distribution: Myanmar, Borneo, Singapore, Indonesia.

deficiens Cameron, 1937: 4 (Indoscytalinus; variety of annulatus; Type locality: Java: G. Raoeng, Bajoekdoel, alt. 450–700 m; Batoerraden, G. Slamat; Bogor; Lombok).

arizonicus Smetana, 1982: 67 (Thyreocephalus; Type locality: Arizona: Patagonia).

  • Distribution: USA.

ater Laporte, 1835: 117 (Eulissus; Type locality: Sénégal).

  • — Erichson, 1839b: 318 (Xantholinus; characters; Senegal).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog; Sierra Leone; Guinea; Nyassa Lake).

  • — Bernhauer, 1915o: 362 (Eulissus; Belgian Congo).

  • — Scheerpeltz, 1933: 1316 (Eulissus; catalog).

  • — Cameron, 1952e: 831 (Thyreocephalus; Dahomey).

  • Distribution: Senegal, Guinea, Sierra Leone, Benin, Congo, Malawi.

aureus Bernhauer, 1915g: 187 (Thyreocephalus; Type locality: Umgebung Ralums).

  • — Cameron, 1931: 360 (Thyreocephalus; New Guinea).

  • — Scheerpeltz, 1933: 1318 (Thyreocephalus; catalog).

  • Distribution: New Guinea.

basilewskyi Tottenham, 1956: 258 (Thyreocephalus; Type locality: Ruanda: Tshuruyaga, forêt du Rugege, 2400 m).

  • Distribution: Rwanda.

bordonii Lecoq, 1996: 153 (Thyreocephalus ?; cited in genus provisionally; Type locality: l'archipel des Comores: Mohéli, Miringoni).

  • Distribution: Comoro Islands.

borneensis Bernhauer, 1915f: 140 (Thyreocephalus; Type locality: Sarawak: Mt. Matang, 2000 Fuss).

  • — Cameron, 1928f: 410 (Thyreocephalus; Borneo).

  • — Bernhauer, 1928: 15 (Thyreocephalus; Sumatra).

  • — Scheerpeltz, 1933: 1318 (Thyreocephalus; catalog).

  • — Cameron, 1933b: 344 (Indoscytalinus, cited as Indoscitalinus; British North Borneo).

  • — Cameron, 1933d: 385 (Indoscytalinus, cited as Indoscitalinus; Sumatra).

  • — Cameron, 1937: 4 (Indoscytalinus, cited as Indoscitalinus; Java).

  • Distribution: Borneo, Indonesia.

buruensis Bernhauer, 1926f: 316 (Thyreocephalus; Type locality: Buru).

  • — Scheerpeltz, 1933: 1318 (Thyreocephalus; catalog).

  • Distribution: Indonesia.

caeruleipennis Cameron, 1931: 359 (Dinoxantholinus; Type locality: Mount Lamington, New Guinea).

  • — Scheerpeltz, 1933: 1315 (Dinoxantholinus; catalog).

  • — Bernhauer, 1942c: 362 (Thyreocephalus; cited as coeruleipennis; Cameroon).

  • — Scheerpeltz, 1956a: 484 (Thyreocephalus; cited as coeruleipennis; Belgian Congo).

  • — Scheerpeltz, 1974c: 20 (Thyreocephalus; Sudan).

  • Distribution: New Guinea, Sudan, Congo, Cameroon.

caeruleus Oke, 1933: 126 (Thyreocephalus; Type locality: Australia: Victoria: Hattah, Melton, Sunshine).

  • Distribution: Australia.

castaneipennis Cameron, 1925a: 185 (Thyreocephalus; Type locality: Merauke, S. N. Guinea).

  • — Scheerpeltz, 1933: 1318 (Thyreocephalus; catalog).

  • Distribution: New Guinea.

celebensis Cameron, 1942c: 843 (Thyreocephalus; Type locality: S. Celebes: Tamba).

  • Distribution: Indonesia.

coelestis Fauvel, 1878g: 540 (Xantholinus; Type locality: Australie méridionale: Victoria, Melbourne).

  • — Olliff, 1887: 487 (Xantholinus; characters; Australia).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • — Lea, 1925: 219 (Thyreocephalus; characters; Australia).

  • Distribution: Australia.

coeruleipennis Quedenfeldt, 1881: 293 (Xantholinus; Type locality: Africa occ. Angola).

  • — Bernhauer, 1912h: 479 (Thyreocephalus; Central Africa).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • — Bernhauer, 1915o: 362 (Thyreocephalus; Cameroon).

  • — Cameron, 1950g: 33 (Thyreocephalus; Belgian Congo).

  • — Tottenham, 1956: 259 (Thyreocephalus; characters in key).

  • — Coiffait, 1968: 154 (Thyreocephalus; cited as caeruleipennis; notes).

  • Distribution: Angola, Congo, Cameroon, “Central Africa”.

columbicus Bernhauer, 1918: 84 (Thyreocephalus; Type locality: West-Kolumbien: Rio Vitaco, 2200 m, in der Umgebung von Cali am Rio Cauca).

  • — Scheerpeltz, 1933: 1318 (Thyreocephalus; catalog).

  • Distribution: Colombia.

cribripennis Sharp, 1885: 501 (Thyreocephalus; Type locality: Mexico, Cordova).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • Distribution: Mexico.

cyanipennis Sharp, 1885: 500 (Thyreocephalus; Type locality: Panama, Volcan de Chiriqui 4000 to 6000 feet).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • Distribution: Panama.

deficiens Cameron, 1937, see: annulatus Fauvel, 1895.

dispilus Erichson, 1839b: 317 (Xantholinus; Type locality: Bengala).

  • — Kraatz, 1859: 104 (Xantholinus; characters; Bengal).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • — Cameron, 1932: 39 (Indoscytalinus, cited as Indoscitalinus; characters; India).

  • — Scheerpeltz, 1933: 1318 (Thyreocephalus; catalog).

  • — Cameron, 1937: 3 (Indoscytalinus, cited as Indoscitalinus; Java).

  • — Zheng, 1995b: 95, 96 (Indoscytalinus, cited as Indoscitalinus; China).

  • Distribution: India, Indonesia, China.

diversiceps Bernhauer, 1936a: 21 (Thyreocepalus; Type locality: Lulua: Kapanga).

  • Distribution: Congo.

dohertyi Cameron, 1942c: 842 (Thyreocephalus; Type locality: Boeroe Island: Ilat).

  • Distribution: Indonesia.

dugesi Casey, 1906: 374 (Saurohypnus; Type locality: Mexico: Guanajuato).

  • — Bernhauer and Schubert, 1914: 299 (Saurohypnus; catalog).

  • Distribution: Mexico.

eppelsheimi Bernhauer and Schubert, 1914, see: gestroi Fauvel, 1895.

eppelsheimi Cameron, 1925, see: gestroi Fauvel, 1895.

extremus Sharp, 1876: 198 (Linidius; Type locality: Upper Amazons).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • — Sharp, 1885: 499 (transferred from Linidius to Thyreocephalus).

  • Distribution: Brazil.

fassli Bernhauer, 1916f: 184 (Thyreocephalus; Type locality: Westperu, Huancabamba).

  • — Scheerpeltz, 1933: 1318 (Thyreocephalus; catalog).

  • Distribution: Peru.

feae Fauvel, 1895: 242 (Xantholinus; Type locality: Bírmanie, Carin Cheba, 900–1100 m).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • — Cameron, 1932: 38 (Indoscytalinus, cited as Indoscitalinus; characters; Burma).

  • — Scheerpeltz, 1933: 1318 (Thyreocephalus; catalog).

  • Distribution: Myanmar.

feae Cameron, 1941b: 446 (Thyreocephalus; [preoccupied]; Type locality: Philippines: Luzon: Imugan).

  • — Herman, 2001: 53 (Thyreocephalus; name will be replaced by Bordoni).

  • Distribution: Philippines.

flavus Last, 1980: 157 (Thyreocephalus; Type locality: New Guinea: Simbang, Huon Golf).

  • Distribution: New Guinea.

frater Cameron, 1937a: 100 (Indoscytalinus, cited as Indoscitalinus; Type locality: New Guinea: Kokoda, alt. 1200 ft.).

  • Distribution: New Guinea.

gabonensis Coiffait, 1968: 154 (Thyreocephalus; Type locality: Gabon: Belinga).

  • Distribution: Gabon.

gestroi Fauvel, 1895: 243 (Xantholinus; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Carin Ghécu, 1300–1400 m; Carin Asciuii Ghécu, 1400–1500 m).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • — Cameron, 1932: 39 (Indoscytalinus, cited as Indoscitalinus; synonym of rudis).

  • — Scheerpeltz, 1933: 1319 (Thyreocephalus; synonym of rudis).

  • — Herman, 2001: 32 (Thyreocephalus; valid name for species).

  • Distribution: India, Nepal, Bhutan, Myanmar, Vietnam, Taiwan.

rudis Eppelsheim, 1895c: 62 (Xantholinus; subgenus Eulissus; [preoccupied]; Type locality: Sikkim im Himalaya).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; synonym of eppelsheimi).

  • — Cameron, 1932: 39 (Indoscytalinus, cited as Indoscitalinus; characters; Burma; India).

  • — Scheerpeltz, 1933: 1319 (Thyreocephalus; catalog).

  • — Jarrige, 1951: 133 (Indoscytalinus; Tonkin).

  • — Shibata, 1973a: 126 (Indoscytalinus, cited as Indoscitalinus; cited as valid species; Taiwan).

  • — Coiffait, 1977b: 206 (Indoscytalinus, cited as Indoscitalinus; cited as valid species; Bhutan).

  • — Coiffait, 1982a: 232 (Indoscytalinus, cited as Indoscitalinus; cited as valid species; Nepal).

  • — Coiffait, 1983d: 161 (Indoscytalinus, cited as Indoscitalinus; cited as valid species; Nepal).

eppelsheimi Bernhauer and Schubert, 1914: 302 (Xantholinus; replacement name for rudis Eppelsheim).

  • — Cameron, 1932: 39 (Indoscytalinus, cited as Indoscitalinus; synonym of rudis).

  • — Scheerpeltz, 1933: 1319 (Thyreocephalus; synonym of rudis).

  • — Herman, 2001: 32 (Thyreocephalus; synonym of gestroi).

eppelsheimi Cameron, 1925: 51 (Xantholinus; [preoccupied]; replacement name for rudis Eppelsheim).

gigantulus Bernhauer, 1915g: 186 (Thyreocephalus; Type locality: Deutsch-Neuguinea: Lord-berg).

  • — Cameron, 1931: 360 (Thyreocephalus; New Guinea).

  • — Scheerpeltz, 1933: 1319 (Thyreocephalus; catalog).

  • — Cameron, 1937a: 100 (Indoscytalinus, cited as Indoscitalinus; New Guinea).

  • Distribution: New Guinea.

halffteri Smetana, 1977c: 349 (Thyreocephalus; Type locality: Mexico: Tizapan el Alto Edo. de Jalisco).

  • Distribution: Mexico.

holomelas Perroud and Montrousier, 1864: 84 (Xantholinus; Type locality: Nouvelle Calédonie: prise à Kanala).

  • — Fauvel, 1877: 244 (Xantholinus; characters; Australia; Samoa; New Caledonia; New Guinea).

  • — Fauvel, 1878f: 246 (Xantholinus; characters; Aru Islands; New Guinea; New Caledonia; Samoa; Australia).

  • — Fauvel, 1879: 94 (Xantholinus; Dorei).

  • — Olliff, 1887: 488 (Xantholinus; characters; Australia).

  • — Fauvel, 1889b: 261 (Xantholinus; New Caledonia).

  • — Fauvel, 1903d: 267 (Xantholinus; synonym of taitiensis).

  • — Bernhauer and Schubert, 1914: 312 (Eulissus; synonym of taitiensis).

  • — Lea, 1925: 219 (Eulissus; synonym of taitiensis).

  • — Scheerpeltz, 1933: 1319 (Thyreocephalus; synonym of taitiensis).

  • — Coiffait, 1976: 216 (Thyreocephalus; characters; Tahiti).

  • — Coiffait, 1980b: 472 (Thyreocephalus; valid species; Tahiti).

  • Distribution: Australia, New Guinea, New Caledonia, Samoa, Society Islands.

hongkongensis Redtenbacher, 1867, see: anachoreta Erichson, 1839.

indicus Fauvel, 1895: 242 (Xantholinus; Type locality: Bírmanie, Carin Ghécu, 1300–1400; Malacca, Perak).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • — Cameron, 1932: 37 (Indoscytalinus, cited as Indoscitalinus; characters; Burma).

  • — Scheerpeltz, 1933: 1319 (Thyreocephalus; catalog).

  • — Scheerpeltz, 1965: 106 (Indoscytalinus, cited as Indoscitalinus; Burma).

  • Distribution: Myanmar, Malaysia.

insularis Cameron, 1941b: 446 (Thyreocephalus; Type locality: Philippines: Luzon: Mt. Makiling).

  • Distribution: Philippines.

interocularis Eppelsheim, 1895a: 131 (Xantholinus; Type locality: Togoland).

  • — Fauvel, 1903e: 239 (Xantholinus; Cameroon; Togo; Gabon; Congo; Ogowe).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • — Bernhauer, 1942c: 361 (Thyreocephalus; Cameroon).

  • — Cameron, 1950g: 33 (Thyreocephalus; variety of coeruleipennis; Belgian Congo).

  • — Tottenham, 1956: 259 (Thyreocephalus; characters in key).

  • — Koppenhöfer, 1994 (Thyreocephalus; bionomics; Kenya).

  • Distribution: Togo, Cameroon, Gabon, Congo, Kenya.

javanus Bernhauer, 1915h: 227 (Thyreocephalus; Type locality: Java: Preanger).

  • — Scheerpeltz, 1933: 1319 (Thyreocephalus; catalog).

  • — Cameron, 1937: 4 (Indoscytalinus, cited as Indoscitalinus; Java).

  • Distribution: Indonesia.

jekelii Guérin-Méneville, 1844: 10 (Thyreocephalus; Type locality: Nouvelle Grande: Ibagué, Carthago).

  • — Solsky, 1872: 308 (Thyreocephalus; Colombia).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • Distribution: Colombia.

lateralis Cameron, 1931: 359 (Dinoxantholinus; Type locality: Fak Fak, New Guinea).

  • — Scheerpeltz, 1933: 1315 (Dinoxantholinus; catalog).

  • Distribution: New Guinea.

lorquini Fauvel, 1877: 241 (Xantholinus; Type locality: Australie probablement septentrionale).

  • — Fauvel, 1878f: 245 (Xantholinus; characters; Moluccas; Australia; Sulawesi).

  • — Fauvel, 1879: 93 (Xantholinus; Aru Islands).

  • — Olliff, 1887: 481 (Xantholinus; characters; Australia).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • — Lea, 1925: 219 (Thyreocephalus; characters; Australia).

  • — Scheerpeltz, 1933: 1319 (Thyreocephalus; catalog).

  • — Bellas, Brown, and Moore, 1974 (Thyreocephalus; defensive secretions).

  • Distribution: Australia, Indonesia.

rufitarsis Fauvel, 1877: 241 (Xantholinus; Type locality: Nouvelle Galles du Sud; Queensland, Gayndah).

  • — Fauvel, 1878g: 538 (Xantholinus; Australia).

  • — Olliff, 1887: 481 (Xantholinus; characters; Australia).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • — Bernhauer, 1920: 16 (Thyreocephalus; Australia).

  • — Lea, 1925: 219 (Thyreocephalus; synonym of lorquini).

  • — Scheerpeltz, 1933: 1319 (Thyreocephalus; synonym of lorquini).

lynceus Erichson, 1839b: 309 (Xantholinus; Type locality: Bogotae in Columbia).

  • — Solsky, 1875: 19 (Thyreocephalus; Peru).

  • — Sharp, 1885: 499 (Thyreocephalus; Nicaragua, Panama, Colombia).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog; Bolivia; Peru; Brazil).

  • — Scheerpeltz, 1933: 1319 (Thyreocephalus; catalog).

  • Distribution: Nicaragua, Panama, Colombia, Brazil, Peru, Bolivia.

malaisei Scheerpeltz, 1965: 197 (Indoscytalinus, cited as Indoscitalinus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).

  • Distribution: Myanmar.

menglaensis Zheng, 1995b: 95, 96 (Indoscytalinus, cited as Indoscitalinus; Type locality: Yunnan: Mengla County, 631 m).

  • Distribution: China.

mocquerysi Fauvel, 1903e: 239 (Xantholinus; Type locality: Cameroun; Gabon: Libreville, Loanda).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • — Coiffait, 1968: 155 (Thyreocephalus; notes; Gabon; Cameroon).

  • Distribution: Cameroon, Gabon.

morio Motschulsky, 1858, see: anachoreta Erichson, 1839.

nigerrimus Kraatz, 1859: 103 (Xantholinus; Type locality: India orientalis; Hongkong).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Cameron, 1932: 37 (Indoscytalinus, cited as Indoscitalinus; characters; India).

  • — Scheerpeltz, 1933: 1319 (Thyreocephalus; catalog).

  • — Jarrige, 1951: 133 (Indoscytalinus; Tonkin).

  • Distribution: India, Vietnam, China, “East Indies”.

pallidipennis Kraatz, 1859: 103 (Xantholinus; variety of nigerrimus; Type locality: Not cited).

  • — Scheerpeltz, 1933: 1319 (Thyreocephalus; variety of nigerrimus).

nigerrimus Sharp, 1887: 791 (Saurohypnus; [preoccupied]; Type locality: Mexico, Jalapa; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 299 (Saurohypnus; catalog).

  • — Smetana, 1982: 70 (Thyreocephalus; lectotype designation; characters; Texas).

  • — Herman, 2001: 53 (Thyreocephalus; name will be replaced by Bordoni).

  • Distribution: Mexico, USA.

ohlmusi Last, 1980: 156 (Thyreocephalus; Type locality: New Guinea: Karimui, 1000 m).

  • Distribution: New Guinea.

orthodoxus Olliff, 1887: 484 (Xantholinus; Type locality: Sydney, Port Hacking, New South Wales).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Bellas, Brown, and Moore, 1974: 277 (Eulissus; defensive secretions).

  • — Kuschel, 1990: 26 (Thyreocephalus; introduced, Australian species; winged; habitat; New Zealand).

  • Distribution: Australia, New Zealand.

pallidipennis Kraatz, 1859, see: nigerrimus Kraatz, 1859.

papuensis Bernhauer, 1915g: 187 (Thyreocephalus; Type locality: Deutsch-Neuguinea: Pionnierlager und Dampfer-Ankerplatz).

  • — Scheerpeltz, 1933: 1319 (Thyreocephalus; catalog).

  • Distribution: New Guinea.

parcior Tottenham, 1956: 257 (Thyreocephalus; Type locality: Ruanda: Tshuruyaga, forêt du Rugege, 2400 m).

  • Distribution: Rwanda.

philippinus Bernhauer, 1912g: 252 (Thyreocephalus; Type locality: Luzon: Rizal, Montalban Gorge).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • Distribution: Philippines.

prodigiosus Heller, 1910: 8 (Dinoxantholinus; Type locality: Papua New Guinea).

  • — Bernhauer and Schubert, 1914: 309 (Dinoxantholinus; catalog).

  • — Cameron, 1931: 360 (Dinoxantholinus; New Guinea).

  • Distribution: New Guinea.

puncticeps Sharp, 1885: 501 (Thyreocephalus; Type locality: Mexico, Uruapan, Juquila; [Note: See lectotype designation by Smetana, 1977c]).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • — Smetana, 1977c: 349 (Thyreocephalus; lectotype designation; characters).

  • Distribution: Mexico.

puncticeps Cameron, 1942c: 843 (Thyreocephalus; [preoccupied]; Type locality: S. Celebes: Patutuang).

  • — Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 59 (Thyreocephalus; cited as punticeps; necrophilous species; distribution in Mexico).

  • — Herman, 2001: 53 (Thyreocephalus; name will be placed in synonymy by Bordoni).

  • Distribution: Indonesia.

raptor Tottenham, 1956: 256 (Thyreocephalus; Type locality: Ruanda: Tshuruyaga, forêt du Rugege, 2400 m).

  • Distribution: Rwanda.

recticollis Sharp, 1876: 197 (Linidius; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 314 (Linidius; catalog).

  • — Steel, 1938b: 56 (Thyreocephalus).

  • Distribution: Brazil.

rudis Eppelsheim, 1895, see: gestroi Fauvel, 1895.

rufipennis Sharp, 1885: 500 (Thyreocephalus; Type locality: Mexico, Oaxaca; [Note: See lectotype designation by Smetana, 1977c]. Also cited from Guatemala, San Geronimo).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • — Smetana, 1977c: 349 (Thyreocephalus; lectotype designation: Oaxaca, Mexico; characters).

  • Distribution: Mexico, Guatemala.

rufipennis Coiffait, 1982a: 249 (Indoscytalinus, cited as Indoscitalinus; [preoccupied]; Type locality: Inde: Cinchona Hills).

  • — Herman, 2001: 53 (Thyreocephalus; name will be replaced by Bordoni).

  • Distribution: India.

rufofasciatus Pajni and Kohli, 1980: 1 (Indoscytalinus, cited as Indoscitalinus; Type locality: India: Punjab, Chandigarh, Labour Colony Sector 25).

  • Distribution: India.

rufotarsis Fauvel, 1877, see: lorquini Fauvel, 1877.

rufus Cameron, 1941b: 446 (Thyreocephalus; Type locality: Philippines: Mindanao: Iligan. Tangolan).

  • Distribution: Philippines.

salvini Sharp, 1885: 499 (Thyreocephalus; Type locality: Panama, Volcan de Chiriqui 4000 to 6000 feet).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog).

  • Distribution: Panama.

scutellaris Sharp, 1885: 502 (Saurohypnus; Type locality: Mexico, Pinos Altos in Chihuahua; [Note: See lectotype designation by Smetana, 1977c]).

  • — Bernhauer and Schubert, 1914: 299 (Saurohypnus; catalog).

  • — Smetana, 1977c: 348 (Thyreocephalus; lectotype designation).

  • Distribution: Mexico.

semipiceus Bernhauer, 1937: 610 (Thyreocephalus; Type locality: Daressalam).

  • — Levasseur, 1968a: 52 (Thyreocephalus; Central African Republic).

  • Distribution: Tanzania, Central African Republic.

semirufus Coiffait, 1968: 155 (Thyreocephalus; Type locality: Gabon: Belinga).

  • Distribution: Gabon.

sondaicus Bernhauer, 1915f: 140 (Thyreocephalus; Type locality: Java).

  • — Scheerpeltz, 1933: 1319 (Thyreocephalus; catalog).

  • Distribution: Indonesia.

spegazzinii Bernhauer, 1915k: 299 (Eulissus; Type locality: Senegal: Dakkar).

  • — Scheerpeltz, 1933: 1317 (Eulissus; catalog).

  • — Coiffait, 1968: 156 (Thyreocephalus; cited as spegazziini; notes; Gabon; Senegal).

  • Distribution: Senegal, Gabon.

splendens Cameron, 1939b: 145 (Thyreocephalus; Type locality: New Guinea: Cyclops Mts.: Mt. Lina, altitude 3500 feet).

  • Distribution: New Guinea.

subtilis Sharp, 1885: 499 (Thyreocephalus; Type locality: Panama, Volcan de Chiriqui 3000 to 4000 feet; Venezuela).

  • — Bernhauer and Schubert, 1914: 313 (Thyreocephalus; catalog; Brazil; Peru).

  • — Blackwelder, 1943: 491 (Thyreocephalus; characters; St. Vincent).

  • Distribution: West Indies, Panama, Venezuela, Brazil, Peru.

taitiensis Boheman, 1858: 26 (Xantholinus; Type locality: Insula Taiti).

  • — Redtenbacher, 1867: 28 (Xantholinus; Tahiti).

  • — Fauvel, 1877: 244 (Xantholinus; characters; Tahiti).

  • — Fauvel, 1903d: 267 (Xantholinus; New Caledonia; New Hebrides; Australia; Tahiti; Tonga; Samoa; New Guinea; Aru).

  • — Bernhauer and Schubert, 1914: 312 (Eulissus; catalog).

  • — Bernhauer, 1922a: 150 (Thyreocephalus; Samoa).

  • — Lea, 1925: 219 (Eulissus; Australian introduction).

  • — Cameron, 1927a: 25 (Thyreocephalus; Samoa).

  • — Cameron, 1931: 360 (Thyreocephalus; New Guinea).

  • — Scheerpeltz, 1933: 1319 (Thyreocephalus; catalog).

  • — Cameron, 1934a: 22 (Indoscytalinus, cited as Indoscitalinus; New Hebrides).

  • — Cameron, 1937a: 100 (Indoscytalinus, cited as Indoscitalinus; New Guinea).

  • — Coiffait, 1976: 216 (Thyreocephalus; characters; Tahiti).

  • Distribution: Indonesia, New Guinea, New Caledonia, Vanuatu, Tonga, Samoa, Society Islands.

tenuipes Sharp, 1876: 197 (Linidius; Type locality: Ega).

  • — Sharp, 1885: 499 (Thyreocephalus).

  • Distribution: Brazil.

unicolor Sharp, 1885: 500 (Thyreocephalus; Type locality: Mexico, Jalapa).

  • Distribution: Mexico.

Vulda [21 species; Palaearctic Region]

Vulda Jacquelin du Val, 1852: 695 (species included: gracilipes). Type species: Vulda gracilipes Jacquelin du Val, fixed by monotypy.

  • — Lacordaire, 1854: 67 (characters; notes; list of species).

  • — Fairmaire and Laboulbène, 1856: 499 (characters).

  • — Jacquelin du Val, 1856: 31 (characters).

  • — Redtenbacher, 1857: 180 (characters).

  • — Schaum, 1859: 27 (catalog; European species).

  • — Gemminger and Harold, 1868: 599 (catalog).

  • — Jacquelin du Val, 1868: 65 (catalog; European species).

  • — Fauvel, 1873b: 95 [= 1873a: 381] (synonym of Xantholinus).

  • — Redtenbacher, 1874: 217 (characters).

  • — Mulsant and Rey, 1877a: 308 (characters; key to French species).

  • — Heyden, Reitter, and Weise, 1891: 106 (list of species of Europe and Caucasus).

  • — Ganglbauer, 1895: 486 (subgenus of Xantholinus).

  • — Heyden, Reitter, and Weise, 1906: 161 (subgenus of Xantholinus; list of species of Europe and Caucasus).

  • — Reitter, 1908: 110 (characters).

  • — Reitter, 1909: 134 (characters).

  • — Bernhauer and Schubert, 1914: 300 (subgenus of Xantholinus).

  • — Rambousek, 1914 (notes; key to species; new species).

  • — Winkler, 1925: 372 (catalog for Palaearctic region; subgenus of Xantholinus).

  • — Cameron, 1932: 22 (subgenus of Xantholinus).

  • — Gridelli, 1947a: 97, 104 (subgenus of Xantholinus; characters; notes).

  • — Blackwelder, 1952: 403 (type species: gracilipes).

  • — Coiffait, 1956a: 44, 73 (characters; key to French species).

  • — Coiffait, 1972: 310 (characters; type species: gracilipes; key to species of western Palaearctic region; references).

  • — Bordoni, 1973a: 184 (characters; notes; list of species).

  • — Pace, 1977c: 137 (key to species of Apennines).

  • — Bordoni, 1982: 343 (characters; habitus; key to species of Italy).

  • — Tóth, 1989: 32 (characters).

  • — Ciceroni and Zanetti, 1995: 26 (list of species of Italy).

Typhlodes Sharp, 1873a: 1 (species included: italicus). Type species: Typhlodes italicus Sharp, fixed by monotypy.

  • — Duvivier, 1883: 158 (catalog).

  • — Heyden, Reitter, and Weise, 1891: 106 (subgenus of Vulda; list of species of Europe and Caucasus).

  • — Ganglbauer, 1895: 485 (subgenus of Xantholinus).

  • — Heyden, Reitter, and Weise, 1906: 161 (subgenus of Xantholinus; list of species of Europe and Caucasus).

  • — Reitter, 1908: 123 (subgenus of Xantholinus; characters; key to species).

  • — Bernhauer and Schubert, 1914: 300 (subgenus of Xantholinus).

  • — Cameron, 1932: 22 (subgenus of Xantholinus).

  • — Scheerpeltz, 1933: 1307 (subgenus of Xantholinus).

  • — Gridelli, 1947a: 97, 113 (subgenus of Xantholinus; characters; notes).

  • — Blackwelder, 1952: 401 (type species: italicus).

  • — Coiffait, 1956a: 73 (subgenus of Vulda; characters).

  • — Coiffait, 1962b: 390, 404 (characters; cited as valid genus).

  • — Coiffait, 1972: 310 (subgenus of Vulda; characters; type species: italicus; key to species of western Palaearctic region; references).

  • — Bordoni, 1982: 343, 347 (subgenus of Vulda; characters; key to species from Italy).

angusticollis Fauvel, 1874: 338 [= 1872: 44] (Xantholinus; name proposed for gracilipes misidentified by Fauvel; Type locality: Toscane).

  • — Fauvel, 1873b: 100 [= 1873a: 386] (Xantholinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 300, cited gracilipes Fauvel as a synonym of angusticollis; Fauvel did not describe the species as new, he attributed it to Jacquelin du Val]; characters; France; Italy).

  • — Reitter, 1908: 124 (Vulda; characters; France; Italy).

  • — Cameron, 1912: 2 (Xantholinus; subgenus Vulda; characters in key).

  • — Bernhauer and Schubert, 1914: 300 (Xantholinus; catalog).

  • — Rambousek, 1914: 9 (Xantholinus; subgenus Vulda; characters; Italy).

  • — Scheerpeltz, 1933: 1308 (Xantholinus; catalog).

  • — Gridelli, 1947a: 105 (Xantholinus; subgenus Vulda; characters; Italy).

  • — Coiffait, 1972: 318 (Vulda; characters; references; Italy).

  • — Bordoni, 1973a: 184 (Vulda).

  • — Bordoni, 1982: 352 (Vulda; characters; Italy).

  • — Ciceroni, 1993: 75 (Vulda; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Vulda; Italy).

  • Distribution: Italy.

barbara Fauvel, 1873b: 103 [= 1873a: 389] (Xantholinus; Type locality: Alger).

  • — Fauvel, 1878: 117 [= 1878a: 37] (Xantholinus; Algeria).

  • — Fauvel, 1886: 49 [= 1886a: 41] (Xantholinus; Algeria).

  • — Fauvel, 1902b: 101 (Xantholinus; Algeria).

  • — Reitter, 1908: 123 (Xantholinus; subgenus Typhlodes; characters).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • — Rambousek, 1924: 39 (Xantholinus; reduced eyes; Algeria).

  • — Scheerpeltz, 1933: 1308 (Xantholinus; catalog).

  • — Steel, 1949a: 269 (seems to belong in Xantholinus subgenus Idiolinus).

  • — Coiffait, 1962b: 404 (Typhlodes; Algeria).

  • — Coiffait, 1972: 321 (Vulda; subgenus Typhlodes; characters; references; Algeria).

  • Distribution: Algeria.

belisarius Abeille, 1901, see: myops Fauvel, 1873.

brignolii Bordoni, 1973a: 190 (Vulda; subgenus Typhlodes; Type locality: Turchia: vil. Amasya, Boraby gölü).

  • Distribution: Turkey.

dalmatina Rambousek, 1914: 3, 4 (Xantholinus; subgenus Vulda; Type locality: Dalmatien).

  • — Scheerpeltz, 1933: 1309 (Xantholinus; catalog).

  • — Coiffait, 1972: 319 (Vulda; characters; references; Dalmatia).

  • — Bordoni, 1973a: 184 (Vulda).

  • — Bordoni, 1982: 349 (Vulda; subgenus Vulda; characters).

  • — Tóth, 1989: 33 (Vulda; characters).

  • Distribution: Croatia.

doderoi Rambousek, 1914: 4, 12 (Xantholinus; subgenus Vulda; Type locality: Italien).

  • — Scheerpeltz, 1933: 1309 (Xantholinus; catalog).

  • — Gridelli, 1947a: 111 (Xantholinus; subgenus Vulda; characters; Italy).

  • — Coiffait, 1972: 317 (Vulda; characters; references; Italy).

  • — Bordoni, 1973a: 184 (Vulda).

  • — Bordoni, 1982: 349 (Vulda; subgenus Vulda; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Vulda; Italy).

  • Distribution: Italy.

fallaciosa Gridelli, 1947, see: tenuipes Baudi, 1870.

gabrielei Bordoni and Magrini, 1996: 178 (Vulda; subgenus Typhlodes; Type locality: Umbria: M.te La Pelosa, Colle Bertone, Terni).

  • Distribution: Italy.

garganica Bernhauer, 1908e: 323 (Xantholinus; subgenus Typhlodes; Type locality: Mte. S. Angelo, Gargano).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; catalog).

  • — Rambousek, 1924: 39 (Xantholinus; reduced eyes; Italy).

  • — Scheerpeltz, 1933: 1309 (Xantholinus; catalog).

  • — C. Koch, 1936: 19 (Xantholinus; subgenus Typhlodes; characters in key).

  • — Coiffait, 1972: 323 (Vulda; subgenus Typhlodes; characters; references; Italy).

  • — Bordoni, 1973a: 184 (Vulda).

  • — Bordoni, 1982: 360 (Vulda; subgenus Typhlodes; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Vulda; Italy).

  • Distribution: Italy.

gracilipes Jacquelin du Val, 1852: 698 (Vulda; Type locality: environs de Marseille).

  • — Fairmaire and Laboulbène, 1856: 499 (Vulda; characters; France).

  • — Redtenbacher, 1857: 180 (Vulda; characters; France).

  • — Fauvel, 1874: 338 [= 1874b: 45] (Xantholinus; characters; France).

  • — Redtenbacher, 1874: 218 (Vulda; characters; France).

  • — Mulsant and Rey, 1877a: 309, 339 (Vulda; characters; France).

  • — Ganglbauer, 1895: 486 (Xantholinus; subgenus Vulda; characters; France).

  • — Reitter, 1908: 124 (Vulda; characters; Maritime Alps).

  • — Cameron, 1912: 2 (Xantholinus; subgenus Vulda; characters in key).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; catalog).

  • — Rambousek, 1914: 8 (Xantholinus; subgenus Vulda; characters; France).

  • — Scheerpeltz, 1933: 1310 (Xantholinus; catalog).

  • — Gridelli, 1947a: 109 (Xantholinus; subgenus Vulda; characters; Italy).

  • — Coiffait, 1956a: 73 (Vulda; characters; France).

  • — Coiffait, 1972: 315 (Vulda; characters; references; France).

  • — Bordoni, 1973a: 184 (Vulda; cited as gracilis).

  • — Bordoni, 1982: 350 (Vulda; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Vulda; Italy).

  • Distribution: France, Italy.

holdhausi Bernhauer, 1908e: 323 (Xantholinus; subgenus Typhlodes; Type locality: Insel Elba).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Rambousek, 1924: 39 (Xantholinus; reduced eyes).

  • — Scheerpeltz, 1933: 1310 (Xantholinus; catalog).

  • — C. Koch, 1936: 18 (Xantholinus; subgenus Typhlodes; characters in key).

  • — Coiffait, 1972: 324 (Vulda; subgenus Typhlodes; characters; references; Elba Island).

  • — Bordoni, 1973a: 184 (Vulda).

  • — Bordoni, 1982: 362 (Vulda; subgenus Typhlodes; characters; Italy).

  • — Bordoni, 1984b: 39 (Vulda; notes; Tuscany coast, not Elba; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Vulda; Italy).

  • Distribution: Italy.

istriensis Rambousek, 1924, see: mihoki Bernhauer, 1914.

italica Sharp, 1873a: 2 (Typhlodes; Type locality: Florence).

  • — Fauvel, 1873b: 131 [= 1873c: 24] (Xantholinus; synonym of myops).

  • — Fauvel, 1874: 339 [= 1874b: 45] (Xantholinus; synonym of tenuipes).

  • — Ganglbauer, 1895: 485 (Xantholinus; synonym of tenuipes).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; synonym of tenuipes).

  • — Scheerpeltz, 1933: 1314 (Xantholinus; synonym of tenuipes).

  • — C. Koch, 1936: 17 (Xantholinus; subgenus Typhlodes; characters; notes; Italy).

  • — Gridelli, 1947a: 124 (Xantholinus; subgenus Typhlodes; characters; Italy).

  • — Coiffait, 1972: 321 (Vulda; subgenus Typhlodes; characters; references; Italy).

  • — Bordoni, 1973a: 184 (Vulda).

  • — Pace, 1977c: 131 (Vulda; subgenus Typhlodes; characters).

  • — Bordoni, 1982: 371 (Vulda; subgenus Typhlodes; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Vulda; Italy).

  • Distribution: Italy.

lepinensis Pace, 1977c: 127 (Vulda; subgenus Typhlodes; Type locality: Monti Lepini, Lazio, nei pressi del passo Cona di Selvapiana, verso Carpineto Romano, a circa m 700 d'altitudine).

  • — Bordoni, 1982: 374 (Vulda; subgenus Typhlodes; characters; Italy).

  • — Ciceroni, 1993: 75 (Vulda; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Vulda; Italy).

  • Distribution: Italy.

magrinii Bordoni, 1987: 26 (Vulda; subgenus Typhlodes; Type locality: Campania: M.te Cervialto, Salerno).

  • — Ciceroni and Zanetti, 1995: 26 (Vulda; Italy).

  • Distribution: Italy.

mariellae Merceron, 1999 [Note: Nomen nudum], see: myops Fauvel, 1873.

mihoki Bernhauer, 1914b: 65 (Xantholinus; subgenus Typhlodes; Type locality: Kroatien, Plitvica).

  • — Rambousek, 1924: 39 (Xantholinus; reduced eyes; Croatia).

  • — Scheerpeltz, 1933: 1312 (Xantholinus; catalog).

  • — Gridelli, 1947a: 128 (Xantholinus; subgenus Typhlodes; characters; Istria).

  • — Coiffait, 1972: 325 (Vulda; subgenus Typhlodes; characters; references; Croatia).

  • — Bordoni, 1982: 356 (Vulda; subgenus Typhlodes; characters; Italy).

  • — Tóth, 1989: 33 (Vulda; characters).

  • — Ciceroni and Zanetti, 1995: 26 (Vulda; Italy).

  • Distribution: Italy, Croatia.

istriensis Rambousek, 1924: 40 (Xantholinus; Type locality: Istrie à Monte Maggiore).

  • — Scheerpeltz, 1933: 1312 (Xantholinus; synonym of mihoki).

  • — Bordoni, 1982: 356 (Vulda; synonym of mihoki).

minima Fiori, 1915: 13 (Xantholinus; subgenus Vulda; Type locality: Piazza Armerina).

  • — Scheerpeltz, 1933: 1312 (Xantholinus; catalog).

  • — Bordoni, 1982: 347 (Vulda ?; notes).

  • — Ciceroni and Zanetti, 1995: 26 (Vulda; incertae sedis; Italy).

  • Distribution: Italy.

myops Fauvel, 1873b: 103 [= 1873a: 389] (Xantholinus; Type locality: Alpes Maritimes, près la frontière française).

  • — Fauvel, 1873b: 131 [= 1873c: 24] (Xantholinus; notes; Italy).

  • — Mulsant and Rey, 1877a: 270 (Xantholinus; characters; France).

  • — Ganglbauer, 1895: 485 (Xantholinus; subgenus Typhlodes; characters; Maritime Alps; France).

  • — Reitter, 1908: 123 (Xantholinus; subgenus Typhlodes; characters; Maritime Alps).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Rambousek, 1924: 39 (Xantholinus; reduced eyes; Monaco).

  • — Portevin, 1929: 372 (Xantholinus; characters; France).

  • — Scheerpeltz, 1933: 1312 (Xantholinus; catalog).

  • — C. Koch, 1936: 19 (Xantholinus; subgenus Typhlodes; characters in key).

  • — Gridelli, 1947a: 126 (Xantholinus; subgenus Typhlodes; characters; Italy).

  • — Coiffait, 1956a: 73 (Vulda; subgenus Typhlodes; characters in key; France).

  • — Coiffait, 1972: 324 (Vulda; subgenus Typhlodes; characters; references; Italy).

  • — Bordoni, 1973a: 184 (Vulda).

  • — Bordoni, 1982: 361 (Vulda; subgenus Typhlodes; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Vulda; Italy).

  • Distribution: France, Italy.

belisarius Abeille, 1901: 60 (Xantholinus; Type locality: Sospel; Ste-Baume).

  • — Reitter, 1908: 123 (Xantholinus; synonym of myops).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; synonym of myops).

  • — Coiffait, 1972: 324 (Vulda; synonym of myops).

mariellae Merceron, 1999: 35 (Vulda; aberration of myops; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Castelvittorio, Italie-Ligurie).

ophthalmica Gridelli, 1947a: 115 (Xantholinus; subgenus Typhlodes; Type locality: Cerchio).

  • — Coiffait, 1972: 320 (Vulda; subgenus Typhlodes; characters; Italy).

  • — Bordoni, 1973a: 184 (Vulda).

  • — Pace, 1977c: 137 (Vulda; subgenus Typhlodes; characters).

  • — Bordoni, 1982: 358 (Vulda; subgenus Typhlodes; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Vulda; Italy).

  • Distribution: Italy.

ottomana Cameron, 1912: 1 (Xantholinus; subgenus Vulda; Type locality: Beikos).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Rambousek, 1914: 7 (Xantholinus; subgenus Vulda; republication of original description).

  • — Scheerpeltz, 1933: 1313 (Xantholinus; catalog).

  • — Coiffait, 1972: 319 (Vulda; characters; references; Turkey).

  • Distribution: Turkey.

sbordonii Pace, 1979a: 107 (Vulda; subgenus Typhlodes; Type locality: Foresta Umbra, Gargano, Puglia).

  • — Bordoni, 1982: 407 (Vulda; subgenus Typhlodes; genitalic illustrations).

  • — Ciceroni and Zanetti, 1995: 26 (Vulda; Italy).

  • Distribution: Italy.

simbruinica Pace, 1977c: 132 (Vulda; subgenus Typhlodes; Type locality: Abruzzo, nord di Monte Viglio, strada Filettino-Capistrello, a circa 1500 metri d'altitudine).

  • — Bordoni, 1982: 367 (Vulda; subgenus Typhlodes; characters; Italy).

  • — Ciceroni, 1993: 75 (Vulda; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Vulda; Italy).

  • Distribution: Italy.

tenuipes Baudi, 1870: 388 (Xantholinus; Type locality: Apennini, Alpes Lepontii).

  • — Fauvel, 1873b: 100 [= 1873a: 386] (Xantholinus; characters; Italy).

  • — Ganglbauer, 1895: 485 (Xantholinus; subgenus Typhlodes; characters; Apennines).

  • — Reitter, 1908: 123 (Xantholinus; subgenus Typhlodes; characters; Italy).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • — Rambousek, 1924: 39 (Xantholinus; reduced eyes; Italy).

  • — Scheerpeltz, 1933: 1314 (Xantholinus; catalog).

  • — C. Koch, 1936: 19 (Xantholinus; subgenus Typhlodes; characters in key).

  • — Gridelli, 1947a: 119 (Xantholinus; subgenus Typhlodes; characters).

  • — Coiffait, 1972: 322 (Vulda; subgenus Typhlodes; characters; references; Italy).

  • — Bordoni, 1973a: 184 (Vulda).

  • — Pace, 1977c: 137 (Vulda; subgenus Typhlodes; characters).

  • — Bordoni, 1982: 363 (Vulda; subgenus Typhlodes; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Vulda; Italy).

  • Distribution: Italy.

fallaciosa Gridelli, 1947a: 121 (Xantholinus; subgenus Typhlodes; subspecies of tenuipes; Type locality: Gabellina).

  • — Coiffait, 1972: 322 (Vulda; subgenus Typhlodes; subspecies of tenuipes; characters; references; Italy).

  • — Pace, 1977c: 137 (Vulda; subgenus Typhlodes; subspecies of tenuipes; characters).

  • — Bordoni, 1982: 366 (Vulda; subgenus Typhlodes; subspecies of tenuipes; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Vulda; subspecies of tenuipes; Italy).

vignai Bordoni, 1973a: 186 (Vulda; subgenus Typhlodes; Type locality: Turchia: vil. Bolu, Abant, 1400 m).

  • Distribution: Turkey.

Xanthocorynus [1 species; Oceanic Region]

Xanthocorynus Sharp, 1908: 549 (species included: deceptor). Type species: Xanthocorynus deceptor Sharp, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 295 (world catalog; 1 species).

  • — Blackwelder, 1952: 404 (type species: deceptor).

deceptor Sharp, 1908: 550 (Xanthocorynus; Type locality: Kauai, mts. Waimea, 4000 ft.).

  • — Bernhauer and Schubert, 1914: 295 (Xanthopygus; catalog).

  • — Nishida, 1994: 74 (Xanthocorynus; Hawaii).

  • — Nishida, 1997: 63 (Xanthocorynus; Hawaii).

  • Distribution: Hawaii.

Xantholinus [274 species (7 doubtful); Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australia, and Oceanic Regions]

Xantholinus Dejean, 1821: 23 (species included: fulminans, pyropterus, elegans, lentus, ochraceus, alternans, fulgidus; [Note: The type species of the genus has often been cited as fulgidus; this designation was overturned by the Commission]). Type species: Staphylinus linearis Olivier, fixed by Commission in Opinion 1250 (ICZN, 1983).

  • — Latreille, 1825: 244 (cited as Xantholin; characters).

  • — Berthold, 1827: 331 (cited as Xanthalinus; characters).

  • — Zetterstedt, 1828: 80 (characters).

  • — Curtis, 1829: 26 (catalog; Britain).

  • — Mannerheim, 1830: 7, 33 (characters; [Note: According to Tottenham, 1949: 369, Mannerheim's usage of Gyrohypnus was a misidentification of Xantholinus]).

  • — Mannerheim, 1831: 421, 447 (characters; [Note: According to Tottenham, 1949: 369, Mannerheim's usage of Gyrohypnus was a misidentification of Xantholinus]).

  • — Griffith and Pidgeon, 1832: 292 (characters).

  • — Dejean, 1833: 63 (list of species).

  • — Gistel, 1834: 8 (list of species).

  • — Lacordaire, 1835: 411 (characters).

  • — Zetterstedt, 1838: 66 (characters).

  • — Erichson, 1839a: 422 (characters).

  • — Heer, 1839: 244 (characters).

  • — Erichson, 1839b: 306 (characters).

  • — Stephens, 1839: 385 (characters).

  • — Redtenbacher, 1849: 691 (characters).

  • — Schaum, 1852: 25 (list of species; Europe).

  • — Lacordaire, 1854: 68 (characters; notes; list of species).

  • — Wollaston, 1854: 576 (characters; notes).

  • — Fairmaire and Laboulbène, 1856: 500 (characters).

  • — Jacquelin du Val, 1856: 32 (characters).

  • — Kraatz, 1857d: 629 (characters).

  • — Redtenbacher, 1857: 180 (characters; key to Austrian species).

  • — Thomson, 1858: 30 (characters).

  • — G. Waterhouse, 1858: 26 (catalog of British species).

  • — Thomson, 1859: 27 (characters; type species: fulgidus).

  • — Schaum, 1859: 27 (catalog; European species).

  • — Thomson, 1860: 189 (characters).

  • — LeConte, 1861a: 65 (characters).

  • — Gredler, 1863: 107 (list of species from Tirol).

  • — LeConte, 1863: 24 (list of North American species).

  • — Wencker and Silbermann, 1866: 30 (list of species; collecting notes; France).

  • — Fauvel, 1866a: 348 [= 1866b: 99] (characters).

  • — Gemminger and Harold, 1868: 600 (catalog).

  • — Jacquelin du Val, 1868: 65 (catalog; European species).

  • — Sharp, 1871d: 12 (list of British species).

  • — Fauvel, 1873b: 95 [= 1873a: 381] (characters; larval characters; key to species).

  • — Redtenbacher, 1874: 218 (characters; key to Austrian species).

  • — Oustalet, 1874: 150 (characters; description of fossil species).

  • — Fauvel, 1875a: XXV [= 1875b: 227] (catalog).

  • — Sharp, 1876: 198 (notes).

  • — Mulsant and Rey, 1877a: 257 (characters; key to French species).

  • — Provancher, 1877: 250 (characters).

  • — Fauvel, 1878f: 243 (characters; notes).

  • — Broun, 1880: 102 (characters).

  • — LeConte, 1880: 171 (key to North American species).

  • — Heyden, 1880: 75 (list of species of Siberian region).

  • — Duvivier, 1883: 155 (catalog).

  • — Lynch, 1884: 175 (characters).

  • — Sharp, 1885: 475 (notes; Mexican and Central American species).

  • — Olliff, 1887: 479 (characters).

  • — Fowler, 1888: 287 (notes; key to British species).

  • — Seidlitz, 1891: 380 (key to some European species).

  • — Seidlitz, 1891a: 405 (key to some European species).

  • — Heyden, Reitter, and Weise, 1891: 105 (list of species of Europe and Caucasus).

  • — Heyden, 1893: 44 (list of species of Siberian region).

  • — Ganglbauer, 1895: 477 (adult and larval characters; key to middle European species).

  • — Heyden, 1896: 33 (list of species of Siberian region).

  • — Fauvel, 1897d: 291 (catalog of species of Barbary; Canaries).

  • — Casey, 1906: 367, 384 (characters).

  • — Heyden, Reitter, and Weise, 1906: 160 (list of species of Europe and Caucasus).

  • — Reitter, 1908: 110, 113 (characters; key to species).

  • — Reitter, 1909: 134, 137 (characters; key to species).

  • — Blatchley, 1910: 394 (characters; key to Indiana species).

  • — Petri, 1912: 68 (list of species of Siebenbürgen).

  • — Bernhauer and Schubert, 1914: 299 (world catalog; 187 species).

  • — Johansen, 1914: 430 (characters; key to Danish species).

  • — Leng, 1920: 105 (catalog of North American species).

  • — Winkler, 1925: 371 (catalog for Palaearctic region).

  • — Lea, 1925: 218 (notes on Australian species).

  • — Cameron, 1925: 51 (catalog of Indian species).

  • — Porta, 1926: 88 (characters; key to species of Italy).

  • — Bruch, 1928a: 427 (catalog; Argentina).

  • — Portevin, 1929: 371 (key to French species).

  • — Roubal, 1930: 362 (catalog; Slovakia).

  • — Cameron, 1932: 22 (characters; key to species of British India).

  • — Joy, 1932: 128 (key to British species).

  • — Scheerpeltz, 1933: 1307 (world catalog supplement).

  • — Normand, 1935: 369 (list of species of Tunisia).

  • — Paulian, 1941: 212 (larval characters).

  • — Blackwelder, 1943: 473 (characters; type species: fulgidus; key to species of West Indies).

  • — Blackwelder, 1944: 146 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Tottenham, 1945: 70 (type species: Staphylinus linearis Olivier, 1794 = Staphylinus longiceps Gravenhorst).

  • — Kloet and Hincks, 1945: 165 (list of British species).

  • — Tottenham, 1949: 369, 416 (type species: linearis Olivier = longiceps Gravenhorst; notes; checklist of British species).

  • — Blackwelder, 1952: 404 (type species: fulgidus).

  • — V. Hansen, 1952: 7, 12 (characters; key to Danish species).

  • — Coiffait, 1956a: 44, 57 (characters; subgeneric classification; key to subgenera and French species).

  • — Kocher, 1958: 138 (checklist of species; Morocco).

  • — Smetana, 1958: 99 (characters; habitus of one species; type species: linearis; key to Czechoslovakian species).

  • — Smetana, 1959a: 208 (list of species collected in Albania).

  • — Coiffait, 1962b: 390, 394 (characters; key to North African species).

  • — Ferreira, 1962a: 16 (list of Portuguese species).

  • — Székessy, 1963: 50 (characters).

  • — Palm, 1963b: 49 (characters; key to species of Sweden).

  • — Lohse, 1964: 163 (characters; key to central European species).

  • — Pototskaia, 1967: 47 (larval characters).

  • — Coiffait, 1968: 128, 151 (characters; notes).

  • — Scheerpeltz, 1968a: 61 (catalog of Austrian species).

  • — Kasule, 1970: 74 (larval characters).

  • — Bordoni, 1971: 685 (checklist of species of Turkey).

  • — Coiffait, 1972: 218 (characters; larval characters; type species: linearis; key to species of western Palaearctic region; references).

  • — Bordoni, 1972: 154, 159 (genitalic anatomy; key to subgenera).

  • — Bordoni, 1973 (list of species of Hungary).

  • — Bordoni, 1973c (list of species of Romania).

  • — Bordoni, 1973j: 694 (list of species from islands near Sicily).

  • — Tikhomirova, 1973a: 181 (checklist of species of USSR).

  • — Bordoni, 1975: 88 (subgenus of Xantholinus; distributional notes).

  • — Pope, 1977: 28 (list of British species).

  • — Topp, 1978: 311 (larval characters).

  • — Muona, 1979: 16 (list of Scandinavian species).

  • — Uhlig, 1979: 251 (some species collected in Germany).

  • — M. Dvořák, 1979: 114 (some species collected in Slovakia).

  • — Smetana, 1979: 50 (requested that International Commission on Zoological Nomenclature set aside all type species designations and designate Staphylinus linearis Olivier as type species and place Xantholinus on the Official list of Generic Names in Zoology).

  • — Uhlig, Vogel, and Sieber, 1980: 245 (some species collected in Germany).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 14 (catalog; Poland).

  • — Bordoni, 1980 (importance of internal sac for taxonomic studies).

  • — Uhlig and Vogler, 1981: 108 (list of some species collected in Germany; habitat and collecting notes).

  • — Spahr, 1981: 97 (references to specimens from amber and copal).

  • — Smetana, 1982: 264 (characters; habitus; revision of and key to species of North America north of Mexico).

  • — Bordoni, 1982: 240 (characters; habitus; key to subgenera and species of Italy).

  • — ICZN, 1983: 85 (placed on Official List of Generic Names in Zoology and Staphylinus linearis is fixed as the type species).

  • — Shibata, 1983: 76 (checklist of species of Japan).

  • — Hammond, 1984: 207 (checklist of species of Borneo).

  • — Muona and Viramo, 1986: 14 (list of species of northeastern Finland).

  • — Costa, Vanin, and Neto, 1986: 101 (larval description; bioluminescence reported in larva; Brazil).

  • — Segers, 1986: 40 (checklist of species; Belgium).

  • — Dettner, 1987: 30 (defensive glands).

  • — Tóth, 1989: 19 (characters; key to species of Hungary).

  • — Drugmand, 1990a: 85 (characters; revision of and key to species of Belgium).

  • — Angelini, 1991: 194 (list of species from high elevations of Calabria, Italy).

  • — Silfverberg, 1992: 18 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Welch, 1993: 228 (ovariole structure and development).

  • — Janák, 1993a: 47 (list of species of Czech Republic and Slovakia).

  • — Hernández, Outerelo, and Gamarra, 1994: 190 (checklist of species; Canary Islands).

  • — Ciceroni and Zanetti, 1995: 25 (list of species of Italy).

  • — M. Hansen, 1996: 109 (list of Danish species).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 243 (additions to list of Danish species).

  • — Downie and Arnett, 1996: 402 (characters).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 74 (list of species collected in 1997; Denmark).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 392 (2 Nearctic species; notes; characters in key).

Idiolinus Casey, 1906: 368, 375 (species included: crassicornis). Type species: Xantholinus crassicornis Hochhuth, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 300 (synonym of Xantholinus).

  • — Cameron, 1932: 22 (subgenus of Xantholinus).

  • — Blackwelder, 1943: 473 (subgenus of Xantholinus; type species: crassicornis).

  • — Tottenham, 1949: 369, 416 (type species: crassicornis Hochhuth; subgenus of Xantholinus; checklist of British species).

  • — Blackwelder, 1952: 197 (type species: crassicornis).

  • — Coiffait, 1972: 218 (synonym of Xantholinus).

  • — Smetana, 1978c: 339 (valid subgenus of Xantholinus).

  • — Bordoni, 1982: 240 (synonym of Xantholinus).

Milichilinus Reitter, 1908: 117 [= 1908a: 20] (subgenus of Xantholinus; species included: decorus). Type species: Xantholinus decorus Erichson, fixed by monotypy.

  • — Reitter, 1909: 137 (subgenus of Xantholinus).

  • — Bernhauer and Schubert, 1914: 300 (subgenus of Xantholinus).

  • — Cameron, 1932: 22 (subgenus of Xantholinus).

  • — Blackwelder, 1943: 473 (subgenus of Xantholinus; type species: decorus).

  • — Blackwelder, 1952: 247 (type species: decorus).

  • — Coiffait, 1956a: 57, 72 (characters).

  • — Smetana, 1958: 99, 114 (subgenus of Xantholinus; characters).

  • — Coiffait, 1972: 219, 222, 223 (subgenus of Xantholinus; characters; type species: decorus; key to species of western Palaearctic region).

  • — Bordoni, 1972: 160 (subgenus of Xantholinus; characters).

  • — Bordoni, 1975: 90 (subgenus of Xantholinus; distributional notes).

Typhlolinus Reitter, 1908: 122 [= 1908a: 25] (subgenus of Xantholinus; species included: corallinus, azuganus, maritimus, hungaricus, haematodes, crassicornis). Type species: Xantholinus (Typhlolinus) hungaricus Reitter, fixed by subsequent designation by Blackwelder, 1943: 474 [Note: Erroneously stated to be monobasic by Blackwelder].

  • — Reitter, 1909: 138 (subgenus of Xantholinus; cited as new; characters).

  • — Bernhauer and Schubert, 1914: 300 (subgenus of Xantholinus).

  • — Cameron, 1932: 22 (subgenus of Xantholinus).

  • — Scheerpeltz, 1933: 1307 (subgenus of Xantholinus).

  • — Blackwelder, 1943: 473 (subgenus of Xantholinus; type species: hungaricus).

  • — Tottenham, 1949c: 41 (type species: crassicornis).

  • — Tottenham, 1949: 369 (synonym of Idiolinus; type species: crassicornis).

  • — Blackwelder, 1952: 401 (type species).

  • — Smetana, 1958: 99 (synonym of Xantholinus).

  • — Coiffait, 1972: 218 (characters; synonym of Xantholinus; type species: corallinus).

  • — Smetana, 1978c: 340 (valid subgenus of Xantholinus; notes).

  • — Bordoni, 1982: 240 (synonym of Xantholinus).

Echinophallus Coiffait, 1956a: 58, 60 (subgenus of Xantholinus; species included: translucidus, devillei, oronticus; preoccupied by Schumacker, 1914). Type species: Xantholinus translucidus Scriba, fixed by original designation.

  • — Coiffait, 1962b: 394, 395 (subgenus of Xantholinus; characters).

  • — Coiffait, 1972: 219, 223, 224 (subgenus of Xantholinus; characters; type species: translucidus; key to species of western Palaearctic region).

  • — Bordoni, 1972: 160, 166 (subgenus of Xantholinus; characters).

  • — Bordoni, 1975: 87 (subgenus of Xantholinus; distributional notes).

  • — Smetana, 1978c: 339 (synonym of Xantholinus subgenus Idiolinus).

  • — Bordoni, 1982: 240, 252 (subgenus of Xantholinus; key to species of Italy).

  • — Bordoni, 1999: 2 (synonym of Xantholinus subgenus Idiolinus).

Calolinus Coiffait, 1956a: 58, 61 (subgenus of Xantholinus; species included: piochardi, sidonensis). Type species: Xantholinus (Calolinus) piochardi Coiffait, fixed by original designation.

  • — Coiffait, 1962b: 394, 395 (subgenus of Xantholinus; characters).

  • — Coiffait, 1972: 219, 222, 226 (subgenus of Xantholinus; characters; type species: rufipennis; key to species of western Palaearctic region).

  • — Bordoni, 1972: 162 (subgenus of Xantholinus; characters).

  • — Bordoni, 1975: 87 (subgenus of Xantholinus; distributional notes).

  • — Bordoni, 1982: 240 (subgenus of Xantholinus; rufipennis cited as type species).

Purrolinus Coiffait, 1956a: 58, 65 (subgenus of Xantholinus; species included: colasi, tricolor, ibericus, inuus, jarrigei). Type species: Staphylinus tricolor Fabricius, fixed by original designation.

  • — Coiffait, 1962b: 394 (subgenus of Xantholinus).

  • — Coiffait, 1972: 219, 223, 238 (subgenus of Xantholinus; characters; type species: tricolor; key to species of western Palaearctic region).

  • — Bordoni, 1972: 161, 168 (subgenus of Xantholinus; characters).

  • — Bordoni, 1975: 87 (subgenus of Xantholinus; distributional notes).

  • — Bordoni, 1982: 240, 255 (subgenus of Xantholinus; key to species).

Helicophallus Coiffait, 1956: 58, 70 (subgenus of Xantholinus; species included: distans, rufipes, libanicus, maritimus; [Note: Coiffait stated that this subgenus may be a synonym of Typhlolinus]). Type species: Xantholinus rufipes P. Lucas, fixed by original designation.

  • — Coiffait, 1962b: 394 (subgenus of Xantholinus; characters).

  • — Coiffait, 1972: 219, 222, 249 (subgenus of Xantholinus; characters; type species: rufipes; key to species of western Palaearctic region).

  • — Bordoni, 1972: 164, 214 (subgenus of Xantholinus; characters).

  • — Bordoni, 1975: 90 (subgenus of Xantholinus; species group characters and classification; distributional notes).

  • — Smetana, 1978c: 340 (synonym of Xantholinus subgenus Typhlolinus).

  • — Bordoni, 1982: 241, 317 (subgenus of Xantholinus; key to species of Italy).

Acanthophallus Coiffait, 1956a: 59, 67 (subgenus of Xantholinus; species included: raffrayi, judaeus, gridellii, clairei; preoccupied by Cope, 1895, and Luche, 1903). Type species: Xantholinus raffrayi Sainte-Claire Deville, fixed by original designation.

  • — Coiffait, 1972: 219, 223, 243 (subgenus of Xantholinus; cited as Acanthopallus and Acanthophallus; characters; type species: appenninicola; key to species of western Palaearctic region).

  • — Bordoni, 1972: 161, 179 (subgenus of Xantholinus; cited as Acantophallus [p.161]; characters).

  • — Bordoni, 1975: 88 (subgenus of Xantholinus; distributional notes).

  • — Bordoni, 1982: 240, 269 (subgenus of Xantholinus; key to species of Italy; appenninicola, cited as apenninicola, cited as type species).

  • — Janák, 1993a: 47 (synonym of Xantholinus subgenus Typhlolinus).

Afrolinus Coiffait, 1962c: 73 (subgenus of Xantholinus species included: algericus). Type species: Xantholinus (Afrolinus) algericus Coiffait, fixed by original designation and monotypy.

  • — Coiffait, 1962b: 394, 395, 408 (subgenus of Xantholinus; characters).

  • — Coiffait, 1972: 219, 223, 238 (subgenus of Xantholinus; characters; type species: algericus; key to species of western Palaearctic region).

Heterolinus Coiffait, 1966f: 199 (subgenus of Xantholinus; species included: khnzoriani; preoccupied by Sharp, 1885). Type species: Xantholinus (Heterolinus) khnzoriani Coiffait, fixed by original designation.

  • — Coiffait, 1972: 219, 222, 249 (subgenus of Xantholinus; characters; type species: khnzoriani; key to species of western Palaearctic region).

  • — Bordoni, 1972: 162, 205 (subgenus of Xantholinus; characters).

  • — Bordoni, 1975: 89 (subgenus of Xantholinus; distributional notes).

  • — Bordoni, 1982: 241, 316 (subgenus of Xantholinus).

Stenophallus Bordoni, 1972: 160 (subgenus of Xantholinus; species included: laniger). Type species: Xantholinus laniger Fauvel, fixed by monotypy.

Toxophallus Bordoni, 1972: 160 (subgenus of Xantholinus; species included: heinzi and one named but undescribed species). Type species: Xantholinus heinzi Coiffait, fixed by monotypy since the second included species was unavailable.

  • — Bordoni, 1972a: 128 (subgenus of Xantholinus; [Note: Cited as a new subgenus of Xantholinus but the name was published in an earlier article. See the notes attached to Bordoni, 1972 and 1972a in the bibliography]; characters).

  • — Bordoni, 1999a: 119 (designation of type species: Xantholinus [Toxophallus] heinzi).

Polydontophallus Bordoni, 1972: 161, 176 (subgenus of Xantholinus; species included: ibericus, meridionalis, fageli, besucheti). Type species: Gyrohypnus meridionalis Nordmann, fixed by original designation.

  • — Bordoni, 1982: 240, 263 (subgenus of Xantholinus; key to species of Italy; jarrigei cited as type species).

Paraphallus Bordoni, 1972: 162, 197 (subgenus of Xantholinus; species included: longiventris, audrasi, rhenanus, sardous, gallicus, strandi, morandi, infimus, kocheri, tingitanus, trispinus, armeniacus, microtophilus, achemenius, massinissae). Type species: Xantholinus longiventris Heer, fixed by original designation [Note: Bordoni, 1972: 197, Xantholinus linearis Olivier was designated as the type species of Paraphallus. That designation would make Paraphallus an isogenotypic synonym of Xantholinus. The included species were stated to be linearis, rossicus, sublinearis, coiffaiti, ochraceus, multipunctatus, and elongatus (pp. 162, 197). However an “Errata corrige” (Bordoni, 1972c) was issued after publication of Bordoni's 1972 revisionary article. Bordoni pointed out and corrected errors that resulted in the transposition of Paraphallus and Xantholinus on pages 162, 187, and 197].

  • — Bordoni, 1972c (correction of type species designation and list of included species cited in Bordoni, 1972).

  • — Bordoni, 1975: 88 (subgenus of Xantholinus; distributional notes).

  • — Smetana, 1982: 264 (synonym of Xantholinus).

  • — Bordoni, 1982: 241, 283 (subgenus of Xantholinus; key to species of Italy; type species is cited as longiventris).

  • — Janák, 1993a: 47 (synonym of Xantholinus subgenus Xantholinus).

  • — Bordoni, 1999: 3 (subgenus of Xantholinus; characters; in original description of Paraphallus Bordoni, 1972, it and Xantholinus were erroneously inverted in key on page 162, and again on pages 187 and 197; type species of Paraphallus is Xantholinus [Paraphallus] longiventris, not X. [P.] linearis; error was corrected in “errata corrige” issued after publication of 1972 article and inserted in some copies of journal but not all).

  • — Herman, 2001: 27 (synonymy).

Tetralinus Bordoni, 1975: 69, 89 (subgenus of Xantholinus; species included: haematodes). Type species: Xantholinus haematodes Kolenati, by original designation.

  • — Bordoni, 1972a: 131 (subgenus of Xantholinus; characters; two species included, haematodes and comellinii; [Note: Bordoni cited the subgenus as new in this article but the name was unavailable because no type species was designated (Article 13.3). He described the subgenus as new again in 1975 but designated a type species thereby making the name available]).

Paracyclinus Bordoni, 1975: 74, 89 (subgenus of Xantholinus; species included: procerus). Type species: Xantholinus procerus Erichson, fixed by original designation.

  • — Bordoni, 1982: 241, 313 (subgenus of Xantholinus).

Liodelinus Marcuzzi, 1977: 11 (subgenus of Xantholinus; species included: marcuzzii; [Note: Nomen nudum, Article 13.1.1; genus and species both undescribed]).

Acanthophus Coiffait, 1983b: 345 (replacement name for Acanthophallus Coiffait). Type species: Xantholinus raffrayi Sainte-Claire Deville, fixed by objective synonymy with Acanthophallus for which a type had been fixed.

  • — Janák, 1993a: 47 (synonym of Xantholinus subgenus Typhlolinus).

  • — Bordoni, 1999: 2 (synonym of Xantholinus subgenus Typhlolinus).

Heterolius Coiffait, 1983b: 345 (replacement name for Heterolinus Coiffait). Type species: Xantholinus khnzoriani Coiffait, fixed by objective synonymy with Heterolinus for which a type had been fixed.

Echinophius Coiffait, 1983b: 345 (replacement name for Echinophallus Coiffait). Type species: Xantholinus translucidus Scriba, fixed by objective synonymy with Echinophallus for which a type had been fixed.

Neoleptophallus Bordoni, 1985a: 84 (subgenus of Xantholinus; species included: anatolicus, minutus). Type species: Xantholinus minutus Coiffait, fixed by original designation.

Phallolinus Ádám, 1987: 135, 148 (named for subgenus “Xantholinus” of Bordoni [1972: 187]; species included: longiventris; [Note: This genus-group name is an objective synonym of Xantholinus subgenus Paraphallus]). Type species: Xantholinus longiventris Heer, fixed by original designation.

  • — Herman, 2001: 27 (synonym of Xantholinus [Paraphallus]).

Meneidophallus Bordoni, 1999a: 119 ([Note: See Bordoni, 1972: 162, for description and list of species included). Type species: Xantholinus roubali Coiffait, fixed by original designation.

  • — Bordoni, 1972: 162 (subgenus of Xantholinus; species included: dissimilis, dvoraki, roubali, schweigeri, alaiensis and two undescribed species; [Note: An unavailable name here, without type species designation (Article 13.3)]).

  • — Bordoni, 1971: 686 (subgenus of Xantholinus; without description; with one species).

abessinus Bernhauer, 1931d: 582 (Xantholinus; subgenus Gyrohypnus; Type locality: Abyssinia: Mt. Chillalo, ca. 10,000 ft.).

  • — Scheerpeltz, 1933: 1307 (Xantholinus; catalog).

  • Distribution: Ethiopia.

achemenius Bordoni, 1973f: 72 (Xantholinus; Type locality: Persia: Teheran).

  • — Bordoni, 1972c (Xantholinus; subgenus Paraphallus; correction of subgeneric assignment in Bordoni, 1972: 162, 188).

  • Distribution: Iran.

aculeatus Coiffait, 1975, see: Nepalinus.

adustus Ushakov, 1986: 1090 (Xantholinus; subgenus Calolinus; Type locality: Armeniia: pos. Shikakhokh).

  • Distribution: Armenia.

aeneiceps Sharp, 1876, see: rufescens Erichson, 1839.

aeneipennis Bernhauer, 1934g: 235 (Xantholinus; Type locality: Kivu: Lulenga, Ende).

  • Distribution: Congo.

aeneotinctus Sharp, 1885: 484 (Xantholinus; Type locality: Guatemala, San Isidro 1600 feet, El Tumbador 2500 feet, Zapote; Panama, David, Bugaba).

  • — Bernhauer and Schubert, 1914: 300 (Xantholinus; catalog).

  • Distribution: Guatemala, Panama.

aeneus Cameron, 1943: 127 (Xantholinus; Type locality: Darjeeling: Ghum district).

  • Distribution: India.

aequalis Fauvel, 1898, see: linearis Olivier, 1795.

afghanicus Coiffait, 1982b: 75 (Xantholinus; subgenus Leptophallus; Type locality: S. Afghanistan: Kandahar, 950 m).

  • Distribution: Afghanistan.

aksekianus Bordoni, 1994: 27 (Xantholinus; subgenus Xantholinus; Type locality: Asia Minore: Akseki, 1000–1200 m).

  • Distribution: Turkey.

alaiensis Coiffait, 1966a: 125 (Xantholinus; Type locality: Kirgistan, Monts Alai, region de Sakh-i-Mardan).

  • — Coiffait, 1972: 281 (Xantholinus; subgenus Xantholinus; characters; Kirghistan).

  • — Bordoni, 1972: 162 (Xantholinus; subgenus Meneidophallus).

  • — Bordoni, 1975: 68 (Xantholinus; subgenus Meneidophallus; notes; Caucasus, Russia).

  • Distribution: Kyrgyzstan, Russia, Caucasus.

albertisi Fauvel, 1877, see: Thyreocephalus.

albicornis Heller, 1900, see: Thyreocephalus.

albocinctus Fauvel, 1905, see: Gauropterus.

algarvensis Coiffait, 1972, see: morandi Coiffait, 1958.

algericus Coiffait, 1962c: 73 (Xantholinus; subgenus Afrolinus; Type locality: environs d'Alger).

  • — Coiffait, 1962b: 398, 403, 408 (Xantholinus; subgenus Afrolinus; characters; Morocco; Algeria).

  • — Coiffait, 1972: 281 (Xantholinus; subgenus Afrolinus; characters; references; Algeria; Morocco).

  • Distribution: Algeria, Morocco.

alluaudi Fauvel, 1907, see: Eulissus.

alpinus Bordoni, 1972, see: bordonii Coiffait, 1970.

alticola Sharp, 1885: 481 (Xantholinus; Type locality: Guatemala, Quiche Mountains 7000 to 9000 feet, Quezaltenango 7800 feet).

  • — Bernhauer and Schubert, 1914: 300 (Xantholinus; catalog).

  • Distribution: Guatemala.

altissimus Bernhauer, 1939a: 78 (Xantholinus; Type locality: Kenya: Cratère de l'Elgon, pied du Koitobbos, 4000 m).

  • Distribution: Kenya.

altivagans Bernhauer, 1927e: 244 (Xantholinus; Type locality: Argentinien: Prov. Jujuy, Hornadita, 3400 m).

  • — Scheerpeltz, 1933: 1307 (Xantholinus; catalog).

  • Distribution: Argentina.

alverniae Bordoni, 1972, see: cerrutii Coiffait, 1964.

amabilis Boheman, 1848, see: Eulissus.

amazonicus Sharp, 1876, see: Homalolinus.

amissus Coquerel, 1860, see: Phacophallus.

anachoreta Erichson, 1839, see: Thyreocephalus.

anatolicus Coiffait, 1965e: 121 (Xantholinus; subgenus Leptophallus; Type locality: Turquie d'Asie, Monts d'Abant près de Bolu).

  • — Bordoni, 1971: 685 (Xantholinus; subgenus Leptophallus; Turkey).

  • — Coiffait, 1972: 258 (Xantholinus; subgenus Leptophallus; characters; Turkey).

  • — Bordoni, 1972: 160 (Xantholinus; subgenus Leptophallus).

  • — Bordoni, 1985a: 84 (Xantholinus; subgenus Neoleptophallus).

  • Distribution: Turkey.

andinus Fauvel, 1866, see: Neohypnus.

anglicus Lesne, 1926, see: linearis Olivier, 1795.

angularis Ganglbauer, 1895, see: Hypnogyra.

angulatus Küster, 1848, see: Gauropterus.

angusticeps Sharp, 1889: 251 (Xantholinus; Type locality: Nagasaki).

  • — Bernhauer and Schubert, 1914: 300 (Xantholinus; catalog).

  • Distribution: Japan.

angusticollis Fauvel, 1874, see: Vulda.

annulatus Fauvel, 1895, see: Thyreocephalus.

anthracinus Broun, 1880, see: Notolinus.

anticus Sharp, 1876: 200 (Xantholinus; Type locality: Rio Purus).

  • — Bernhauer and Schubert, 1914: 300 (Xantholinus; catalog).

  • Distribution: Brazil.

appenninicola Steel, 1946a: 192 (Xantholinus; replacement name for raffrayi Sainte-Claire Deville).

  • — Coiffait, 1972: 287 (Xantholinus; subgenus Acanthophallus; characters; Italy).

  • — Bordoni, 1972: 161, 182 (Xantholinus; subgenus Acanthophallus; cited as apennicola [p. 161]; characters; Italy).

  • — Bordoni, 1976c: 94 (Xantholinus; cited as apenninicola; Italy).

  • — Bordoni, 1982: 273 (Xantholinus; subgenus Acanthophallus; characters; Italy).

  • — Bordoni, 1989b: 519 (Xantholinus; subgenus Acanthophus; cited as apenninicola; characters; Italy).

  • — Ciceroni, 1993: 73 (Xantholinus; cited as apenninicola; Italy).

  • — Ciceroni and Zanetti, 1995: 25 (Xantholinus; Italy).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus).

  • Distribution: Italy.

raffrayi Sainte-Claire Deville, 1926: 119 (Xantholinus; [preoccupied]; Type locality: Apennin, notamment à Gubbio; Riofreddo, Monti Albani, Monte Terminillo).

  • — Coiffait, 1956a: 69 (Xantholinus; subgenus Acanthophallus; characters in key; Italy; Corsica).

  • — Coiffait, 1972: 287 (Xantholinus; synonym of appenninicola).

  • — Bordoni, 1972: 182 (Xantholinus; synonym of appenninicola, cited as apenninicola).

  • — Bordoni, 1982: 273 (Xantholinus; synonym of appenninicola, cited as apenninicola).

devilleanus Scheerpeltz, 1956d: 267 (Xantholinus; subgenus Acanthophallus; replacement name for raffrayi Sainte-Claire Deville).

  • — Coiffait, 1972: 287 (Xantholinus; synonym of appenninicola).

  • — Bordoni, 1972: 182 (Xantholinus; synonym of appenninicola, cited apenninicola).

apterus Bernhauer, 1939a: 81 (Xantholinus; Type locality: Kenya: Mt. Kinangopp, prairies alpines, 3500–3700 m).

  • Distribution: Kenya.

arambourgi Bernhauer, 1939a: 80 (Xantholinus; Type locality: Kenya: Camp III de Elgon, zone des Bruyères, Est, 3500 m).

  • Distribution: Kenya.

araxis Reitter, 1898: 116 (Xantholinus; Type locality: Araxesthal bei Ordubad; [Note: See lectotype designation by Bordoni, 1999]).

  • — Reitter, 1908: 121 (Xantholinus; characters; Arax Valley).

  • — Bernhauer and Schubert, 1914: 300 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1308 (Xantholinus; catalog).

  • — Jarrige, 1971: 493 (Xantholinus; Iran).

  • — Coiffait, 1972: 300 (Xantholinus; subgenus Helicophallus; characters; references; Caucasus; Armenia).

  • — Korge, 1973: 10 (Xantholinus; subgenus Helicophallus; notes; Armenia).

  • — Bordoni, 1975: 81 (Xantholinus; subgenus Helicophallus; characters; notes).

  • — Bordoni, 1999: 4 (Xantholinus; subgenus Helicophallus; lectotype designation).

  • Distribution: Azerbaijan, Iran.

arecae Broun, 1880, see: Paracorynus.

armatus Wollaston, 1877, see: Neohypnus.

armeniacus Coiffait, 1956a: 62 (Xantholinus; Type locality: Arménie Russe: Delhizan).

  • — Coiffait, 1972: 270 (Xantholinus; subgenus Xantholinus; characters; Armenia).

  • — Bordoni, 1972c (Xantholinus; subgenus Paraphallus; correction of subgeneric assignment in Bordoni, 1972: 162, 188).

  • — Coiffait, 1973: 109 (Xantholinus; Turkey).

  • — Bordoni, 1975: 65 (Xantholinus; subgenus Paraphallus; characteres; notes; Armenia; Georgia; Ukraine; Crimea).

  • — Coiffait, 1978d: 163 (Xantholinus; Turkey).

  • Distribution: Turkey, Armenia, Georgia, Ukraine.

assamensis Cameron, 1932: 27 (Xantholinus; Type locality: Assam: Naga Hills, Laimatak).

  • — Scheerpeltz, 1933: 1308 (Xantholinus; catalog).

  • Distribution: India.

atratus Heer, 1839, see: Gyrohypnus.

atriceps W. J. MacLeay, 1873, see: Zeteotomus.

atrocephalus Cameron, 1950d: 17 (Xantholinus; Type locality: Selangor: The Gap).

  • Distribution: Malaysia.

attarum Bernhauer, 1912e: 175 (Xantholinus; Type locality: Argentinien: Prov. Buenos Aires: Carhue).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • Distribution: Argentina.

attenuatus Erichson, 1839, see: Neohypnus.

audanti Blackwelder, 1943: 483 (Xantholinus; Type locality: Haiti: Port-au-Prince. Also cited from Dominican Republic).

  • Distribution: West Indies.

audrasi Coiffait, 1956a: 63 (Xantholinus; Type locality: France: Ain: Ambérieu).

  • — Smetana, 1958: 111 (Xantholinus; subgenus Xantholinus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 165 (Xantholinus; subgenus Xantholinus; characters; central Europe).

  • — Horion, 1965: 101 (Xantholinus; subgenus Xantholinus; France; Czech Republic).

  • — Smetana, 1966: 330 (Xantholinus; characters).

  • — Coiffait, 1972: 273 (Xantholinus; subgenus Xantholinus; characters; references; Czechoslovakia; Austria; France).

  • — Bordoni, 1972: 191 (Xantholinus; subgenus Xantholinus; characters; Italy).

  • — Bordoni, 1972c (Xantholinus; subgenus Paraphallus; correction of subgeneric assignment in Bordoni, 1972: 162, 188, 191).

  • — Bordoni, 1973c: 152 (Xantholinus; notes; Romania).

  • — Bordoni, 1982: 296 (Xantholinus; subgenus Paraphallus; characters; Italy).

  • — Drugmand and Haghebaert, 1987: 325 (Xantholinus; Belgium).

  • — Lucht, 1987: 103 (Xantholinus; checklist; Germany; Czechoslovakia; Austria; Denmark; Sweden).

  • — Tóth, 1989: 29 (Xantholinus; characters).

  • — Drugmand, 1990a: 90 (Xantholinus; characters; Belgium).

  • — Ciceroni, 1993: 74 (Xantholinus; Italy).

  • — Janák, 1993a: 47 (Xantholinus; subgenus Xantholinus; Czech Republic; Slovakia).

  • — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 28 (Xantholinus; additional records to Danish list of species).

  • — Ciceroni and Zanetti, 1995: 25 (Xantholinus; Italy).

  • Distribution: Europe.

strandi Coiffait, 1958: 19 (Xantholinus; Type locality: Norvège: Idd, près de la ville de Halden, lat. 59°10′ Nord, long. 11°25′ Est).

  • — Lohse, 1964: 166 (Xantholinus; subgenus Xantholinus; characters; central Europe).

  • — Horion, 1965: 102 (Xantholinus; subgenus Xantholinus; Norway; Sweden).

  • — Coiffait, 1972: 273 (Xantholinus; subspecies of audrasi; characters; Denmark; Scandinavia).

  • — Bordoni, 1972c (Xantholinus; subgenus Paraphallus; correction of subgeneric assignment in Bordoni, 1972: 162, 188).

  • — Lucht, 1987: 103 (Xantholinus; checklist; Sweden).

  • — Lohse and Lucht, 1989: 159 (Xantholinus; synonym of audrasi).

aulicus Erichson, 1840: 923 (Xantholinus; Type locality: Brasilia).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1308 (Xantholinus; catalog).

  • Distribution: Brazil.

auriceps Fauvel, 1878f: 244 (Xantholinus; Type locality: Nouvelle-Guinée, Hatam).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • Distribution: New Guinea.

azarbaijanicus Korge, 1973: 6 (Xantholinus; subgenus Echinophallus; Type locality: Azarbaijan, Iran, Charasu-dagh östlich Heroabad, 2000–2300 m).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Idiolinus; cited as azerbajanicus).

  • Distribution: Iran.

azuganus Reitter, 1908: 122 [= 1908a: 25] (Xantholinus; subgenus Typhlolinus; Type locality: Moldau: transsylvanische Alpen bei Azuga).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1308 (Xantholinus; catalog).

  • — Szujecki, 1969: 482 (Xantholinus; subgenus Acanthophallus; characters; Poland; Transylvania).

  • — Coiffait, 1972: 292 (Xantholinus; subgenus Acanthophallus; characters; references; Romania).

  • — Bordoni, 1972: 161 (Xantholinus; subgenus Acanthophallus).

  • — Bordoni, 1973: 49 (Xantholinus; subgenus Acanthophallus; Hungary).

  • — Bordoni, 1973c: 153 (Xantholinus; subgenus Acanthophallus; characters; notes; Romania).

  • — Tóth, 1985: 177 (Xantholinus; notes; not recorded in Hungary).

  • — Tóth, 1989: 25 (Xantholinus; characters).

  • — Bordoni, 1989a: 504 (Xantholinus; subgenus Acanthophus; characters; Romania).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus).

  • Distribution: Europe.

hungaricus Reitter, 1908: 122 [= 1908a: 25] (Xantholinus; subgenus Typhlolinus; Type locality: Hungaria bor.: Marmatos; [Note: See lectotype designation by Bordoni, 1999]. Originally cited from Nord- und Südeuropa).

  • — Reitter, 1909: 138 (Xantholinus; subgenus Typhlolinus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1311 (Xantholinus; catalog).

  • — Steel, 1946a: 192 (Xantholinus; synonym of distans).

  • — Smetana, 1958: 107 (Xantholinus; subgenus Xantholinus; characters; ecology).

  • — Szujecki, 1969: 481 (Xantholinus; synonym of azuganus).

  • — Coiffait, 1972: 302 (Xantholinus; synonym of distans).

  • — Bordoni, 1973c: 153 (Xantholinus; synonym of azuganus).

  • — Tóth, 1989: 25 (Xantholinus; synonym of azuganus).

  • — Bordoni, 1999: 3 (Xantholinus; lectotype designation: Hungaria bor.: Marmatos).

trellai Szujecki, 1969: 483 (Xantholinus; subgenus Acanthophallus; subspecies of azuganus; Type locality: Polonina: Krzemien, 1320 m).

  • — Coiffait, 1972: 292 (Xantholinus; subgenus Acanthophallus; subspecies of azuganus; characters; Poland).

  • — Bordoni, 1972: 161 (Xantholinus; subgenus Acanthophallus; subspecies of azuganus).

  • — Bordoni, 1975: 63 (Xantholinus; subgenus Acanthophallus; subspecies of azuganus; notes; Ukraine).

  • — Janák, 1993a: 47 (Xantholinus; subgenus Typhlolinus; subspecies of azuganus; Slovakia).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus; subspecies of azuganus).

azureus Lynch, 1884, see: Scytalinus.

badariottii Wasmann, 1900, see: Eulissus.

baeticus Bordoni, 1975b: 659 (Xantholinus; subgenus Acanthophallus; Type locality: Spagna: prov. di Granada, Pinos Genil).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus; cited as baethicus).

  • Distribution: Spain.

baicalensis Fauvel, 1875, see: Hypnogyra.

balaton Bordoni, 1973: 47 (Xantholinus; subgenus Meneidophallus; Type locality: Zalavár presso il Lago Balaton).

  • — Tóth, 1989: 31 (Xantholinus; characters; Hungary).

  • — Janák, 1993a: 47 (Xantholinus; subgenus Meneidophallus; Czech Republic; Slovakia).

  • Distribution: Hungary, Czech Republic, Slovakia.

balearicus Coiffait, 1962a: 274 (Xantholinus; subgenus Acantholinus; Type locality: Baléares: Majorque, Cala S. Vicente).

  • — Coiffait, 1972: 289 (Xantholinus; subgenus Acanthophallus; characters; Balearic Islands).

  • — Bordoni, 1972: 161 (Xantholinus; subgenus Acanthophallus).

  • — Bordoni, 1976c: 94 (Xantholinus; Majorca).

  • — Pons and Palmer, 1996: 166 (Xantholinus; Balearic endemic; notes; habitat; Mallorca; Menorca).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus).

  • Distribution: Balearic Islands.

barbarus Fauvel, 1873, see: Vulda.

barretoi Bernhauer, 1934b: 116 (Xantholinus; Type locality: Argentinien: Jujuy).

  • Distribution: Argentina.

batesi Sharp, 1876: 202 (Xantholinus; Type locality: Not cited).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • Distribution: Brazil.

beattyi Blackwelder, 1943: 485 (Xantholinus; Type locality: St. Croix: Butler Bay Estate. Also cited from Cuba, Haiti, and Dominican Republic).

  • — Ramos, 1946: 33 (Xantholinus; Mona Island, West Indies).

  • Distribution: West Indies.

belisarius Abeille, 1901, see: Vulda.

besucheti Bordoni, 1972b: 127 (Xantholinus; subgenus Polydontophallus; Type locality: Tunisia: Belif, presso Cap Negro).

  • — Bordoni, 1976c: 91 (Xantholinus; notes; Tunisia).

  • Distribution: Tunisia.

bhutanicus Coiffait, 1982a: 232, 243 (Xantholinus; Type locality: Nepal: Phulchoki, 2600 m. Also cited from Bhutan).

  • Distribution: Nepal, Bhutan.

bicolor Sharp, 1876: 199 (Xantholinus; Type locality: Ega).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • Distribution: Brazil.

bicoloripennis Scheerpeltz, 1974: 129 (Xantholinus; Type locality: Cape Prov.: Saldanhabaai, Malagas Island).

  • Distribution: South Africa.

bilimeki Bernhauer, 1904a: 233 (Xantholinus; Type locality: Mexiko, Orizaba).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • Distribution: Mexico.

birmanus Cameron, 1932: 24 (Xantholinus; Type locality: Burma: Ruby Mines).

  • — Scheerpeltz, 1933: 1308 (Xantholinus; catalog).

  • — Coiffait, 1977b: 206 (Xantholinus; Bhutan).

  • — Coiffait, 1982a: 232 (Xantholinus; India).

  • Distribution: India, Myanmar, Bhutan.

bonariensis Gemminger and Harold, 1868: 601 (Xantholinus; replacement name for gracilis Boheman).

  • — Lynch, 1884: 191 (Xantholinus; subgenus Gyrohypnus; characters; Argentina).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1308 (Xantholinus; catalog).

  • — Frank, 1982: 32 (Xantholinus; list of parasites).

  • Distribution: Argentina.

gracilis Boheman, 1858: 27 (Xantholinus; [preoccupied]; Type locality: Buenos Ayres).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; synonym of bonariensis).

bonomettoi Bordoni, 1972: 171 (Xantholinus; subgenus Purrolinus; Type locality: Alpi Cadorine, Lozzo, Pian dei Buoi, Belluno, 1800 m).

  • — Bordoni, 1982: 259 (Xantholinus; subgenus Purrolinus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 25 (Xantholinus; probable synonym of tricolor; Italy).

  • Distribution: Italy.

bordonii Coiffait, 1970: 292 (Xantholinus; subgenus Helicophallus, cited as Helicophalus; Type locality: Cansiglio, Vénétie).

  • — Coiffait, 1972: 303 (Xantholinus; subgenus Helicophallus; characters; references; Italy).

  • — Bordoni, 1972: 164, 228 (Xantholinus; subgenus Helicophallus; characters; Italy).

  • — Bordoni, 1982: 329 (Xantholinus; subgenus Helicophallus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 25 (Xantholinus; Italy).

  • — Bordoni, 1999: 6 (Xantholinus; subgenus Helicophallus).

  • Distribution: Italy.

alpinus Bordoni, 1972: 227 (Xantholinus; subgenus Helicophallus; Type locality: Alpi Cadorine, Monsampiano sul Monte Pavione, Feltre, 2000 m).

  • — Bordoni, 1982: 327 (Xantholinus; subgenus Helicophallus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 25 (Xantholinus; probable synonym of bordonii; Italy).

  • — Bordoni, 1999: 6 (Xantholinus; synonym of bordonii).

carnicus Bordoni, 1972: 223 (Xantholinus; subgenus Helicophallus; Type locality: Prealpi Carniche: Monte Chiampion, Gemona, 1710 m).

  • — Bordoni, 1982: 330 (Xantholinus; subgenus Helicophallus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 25 (Xantholinus; probable synonym of bordonii; Italy).

  • — Bordoni, 1999: 6 (Xantholinus; synonym of bordonii).

sontianus Bordoni, 1972: 225 (Xantholinus; subgenus Helicophallus; Type locality: Goriziano, Rubbia in Val l'Isonzo).

  • — Bordoni, 1982: 341 (Xantholinus; subgenus Helicophallus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; probable synonym of cerrutii; Italy).

  • — Bordoni, 1999: 6 (Xantholinus; synonym of bordonii).

brachypterus Sharp, 1885: 479 (Xantholinus; Type locality: Guatemala, Totonicapam 8500 to 10,000 feet).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • Distribution: Guatemala.

brasilianus Wendeler, 1956a: 226 (Xantholinus; Type locality: Brasilien: Nova Teutonia).

  • Distribution: Brazil.

brasiliensis Steel, 1949a: 270 (Xantholinus; replacement name for episcopalis Fauvel).

  • Distribution: Brazil.

episcopalis Fauvel, 1877: xxvi (Xantholinus; [preoccupied]; Type locality: Brésil, Thérézopolis).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • — Steel, 1949a: 270 (Xantholinus; synonym of brasiliensis).

breviventer Sperk, 1835, see: Gabrius.

bruchi Bernhauer, 1912e: 173 (Xantholinus; Type locality: Argentinien: Prov. Buenos Aires).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • Distribution: Argentina.

brunneipennis Erichson, 1839b: 327 (Xantholinus; Type locality: Columbia).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • Distribution: Colombia.

bulgaricus Coiffait, 1972: 305 (Xantholinus; subgenus Helicophallus; Type locality: Midzur, 2150 m, Stara Planina).

  • — Korge, 1973: 11 (Xantholinus; notes; Greece).

  • Distribution: Bulgaria, Greece.

bythinicus Coiffait, 1972: 286 (Xantholinus; subgenus Acanthophallus; Type locality: Abant Dag, Anatolie occidentale).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus).

  • Distribution: Turkey.

cachemiricus Coiffait, 1983d: 162 (Xantholinus; subgenus Lepidophallus; Type locality: Cachemire: Zagipal, près Pahalgan, 3500–3600 m).

  • Distribution: India.

cadaverinus Lacordaire, 1835, see: Megalinus.

caecus Broun, 1893, see: Pachycorynus.

calcidicus Bordoni, 1973, see: graecus Kraatz, 1858.

calidus Erichson, 1839b: 321 (Xantholinus; Type locality: Columbia).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • Distribution: Colombia.

canaliculatus Erichson, 1839, see: Homalolinus.

capensis Boheman, 1858, see: fallax Sachse, 1852.

carinatus Sharp, 1885: 480 (Xantholinus; Type locality: Mexico, Durasnal).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • Distribution: Mexico.

carius Coiffait, 1972, see: gridellii Coiffait, 1956.

carnicus Bordoni, 1972, see: bordonii Coiffait, 1970.

cassagnaui Coiffait, 1984e: 375 (Xantholinus; Type locality: Nepal oriental: Jaljale Himal, 2440 m).

  • Distribution: Nepal.

castanopterus Erichson, 1839b: 321 (Xantholinus; Type locality: Brasilia).

  • — Lynch, 1884: 184 (Xantholinus; subgenus Megalinus; characters; Argentina).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog; Paraguay).

  • — Scheerpeltz, 1933: 1308 (Xantholinus; catalog).

  • Distribution: Brazil, Paraguay, Argentina.

castigatus Erichson, 1839b: 315 (Xantholinus; Type locality: Columbia).

  • — Fauvel, 1891: 105 (Xantholinus; Venezuela).

  • — Fauvel, 1901: 82 (Xantholinus; Colombia; Mexico; Guatemala; Panama; Venezuela; New Granada).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • Distribution: Venezuela, Colombia, Panama, Guatemala, Mexico.

caucasicus Bordoni, 1975: 73 (Xantholinus; subgenus Heterolinus; Type locality: Teberda nel Caucaso occidentale).

  • Distribution: Caucasus.

cavicola Cameron, 1938, see: Nudobius.

cephalus Say, 1830, see: Nudobius.

cervinipennis W. J. MacLeay, 1873, see: erythropterus Erichson, 1839.

cerrutii Coiffait, 1964a: 506 (Xantholinus; subgenus Helicophallus; Type locality: Italie, Abruzzes, Massif du Grand Sasso).

  • — Coiffait, 1972: 304 (Xantholinus; characters; references; Italy).

  • — Bordoni, 1972: 164, 219 (Xantholinus; subgenus Helicophallus; characters; Italy).

  • — Bordoni, 1976c: 97 (Xantholinus; Italy).

  • — Bordoni, 1976d: 121 (Xantholinus; Italy).

  • — Bordoni, 1982: 331 (Xantholinus; subgenus Helicophallus; characters; Italy).

  • — Ciceroni, 1993: 75 (Xantholinus; Italy).

  • — Ciceroni and Zanetti, 1995: 25 (Xantholinus; Italy).

  • — Bordoni and Magrini, 1996: 177 (Xantholinus; notes; Italy).

  • — Bordoni, 1999: 7 (Xantholinus; subgenus Helicophallus).

  • Distribution: Italy.

alverniae Bordoni, 1972: 214 (Xantholinus; subgenus Helicophallus; Type locality: Toscana, M.te Verna).

  • — Bordoni, 1982: 326 (Xantholinus; subgenus Helicophallus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 25 (Xantholinus; probable synonym of cerrutii; Italy).

  • — Bordoni and Magrini, 1996: 178 (Xantholinus; synonym of cerrutii).

magrinii Bordoni, 1987: 25 (Xantholinus; subgenus Helicophallus; Type locality: Campania: M.te Cervialto, Salerno).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; probable synonym of cerrutii; Italy).

  • — Bordoni, 1999: 7 (Xantholinus; synonym of cerrutii).

samniticus Bordoni, 1988: 43 (Xantholinus; subgenus Helicophallus; Type locality: Molise, M.ti del Matese, M.te Miletto, 1500 m).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; probably synonym of cerrutii; Italy).

  • — Bordoni, 1999: 7 (Xantholinus; synonym of cerrutii).

chalcopterus Erichson, 1839, see: Eulissus.

championi Sharp, 1885: 482 (Xantholinus; Type locality: Guatemala, Capetillo).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • Distribution: Guatemala.

chloropterus Erichson, 1839, see: Eulissus.

christophi Lokay, 1919: 23 (Xantholinus; subgenus Metacyclinus; Type locality: Sibiria, Amur).

  • — Scheerpeltz, 1933: 1308 (Xantholinus; catalog).

  • Distribution: Russia.

cicatricosus Fauvel, 1881: 163 (Xantholinus; Type locality: Lebong, Sumatra).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • Distribution: Indonesia.

ciliciae Bordoni, 1971: 680 (Xantholinus; subgenus Echinophallus; Type locality: Cilicia, Adana, 1200 m).

  • — Bordoni, 1972: 160 (Xantholinus; subgenus Echinophallus).

  • — Bordoni, 1976c: 84 (Xantholinus; notes).

  • — Bordoni, 1994: 29 (Xantholinus; subgenus Helicophallus; Turkey).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Idiolinus).

  • — Assing and Wunderle, 2001: 36 (Xantholinus; Cyprus).

  • Distribution: Turkey.

cinctus Walker, 1858, see: Pachycorynus.

cinctus Fauvel, 1904, see: Gauropterus.

clairei Coiffait, 1956, see: laevigatus Jacobsen, 1849.

claviger Fauvel, 1905, see: Gauropterus.

coelestis Fauvel, 1878, see: Thyreocephalus.

coeruleipennis Quedenfeldt, 1881, see: Thyreocephalus.

coiffaiti H. Franz, 1966: 175 (Xantholinus; Type locality: nördlich der Südbahnstation Sollenau im Steinfeld, Niederösterreich).

  • — Coiffait, 1972: 272 (Xantholinus; subgenus Xantholinus; characters; Austria).

  • — Bordoni, 1972: 162 (Xantholinus; subgenus Paraphallus).

  • — Tóth, 1989: 28 (Xantholinus; characters).

  • — Lohse and Lucht, 1989: 159 (Xantholinus; characters).

  • Distribution: Austria.

coiffaitianus Bordoni, 1975: 84 (Xantholinus; subgenus Helicophallus; proposed for Coiffait's, 1971: 436 use of fortepunctatus “Reitter”; Type locality: Hamarat nel Caucaso).

  • — Coiffait, 1971: 436 (Xantholinus; subgenus Helicophallus; cited as fortepunctatus Reitter [Note: Misidentification according to Bordoni, 1975: 84]; characters).

  • Distribution: Caucasus.

colasi Jarrige, 1941: 48 (Xantholinus; Type locality: France: l'entrée de la grotte des sources de la Nive, Béhérobie, Basses-Pyrénées).

  • — Gridelli, 1950: 6 (Xantholinus; characters; France).

  • — Coiffait, 1956a: 65 (Xantholinus; subgenus Purrolinus; characters in key).

  • — Coiffait, 1972: 283 (Xantholinus; subgenus Purrolinus; characters; references; Pyrenees; France).

  • — Bordoni, 1972: 161 (Xantholinus; subgenus Purrolinus).

  • Distribution: France.

collaris Erichson, 1839, see: Nudobius.

coloratus Karsch, 1881, see: Lepidophallus.

coloratus Brancsik, 1893, see: nossibeanus Bernhauer and Schubert, 1914.

comellinii Bordoni, 1972a: 131 (Xantholinus; subgenus Tetralinus; Type locality: Marocco, Oued Tiflet).

  • — Bordoni, 1976c: 96 (Xantholinus; subgenus Tetralinus; notes; Morocco).

  • Distribution: Morocco.

confusus Mulsant and Rey, 1853, see: Gyrohypnus.

conicollis Wendeler, 1956a: 228 (Xantholinus; Type locality: Brasilien: Nova Teutonia).

  • Distribution: Brazil.

corallinus Reitter, 1901a: 68 (Xantholinus; Type locality: Buchara: Karatak; [Note: See lectotype designation by Bordoni, 1999]).

  • — Reitter, 1908: 122 (Xantholinus; subgenus Typhlolinus; characters; Turkestan).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1308 (Xantholinus; catalog).

  • — Coiffait, 1972: 265 (Xantholinus; subgenus Calolinus; characters; references; Uzbekistan; Turkestan).

  • — Bordoni, 1975: 59 (Xantholinus; subgenus Calolinus; characters; notes; Buchara; Turkestan).

  • — Bordoni, 1999: 2, 3 (Xantholinus; subgenus Calolinus; lectotype designation).

  • Distribution: Uzbekistan.

corallipes Fauvel, 1891: 105 (Xantholinus; Type locality: Colonie Tovar).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • Distribution: Venezuela.

cordieri Boieldieu, 1859, see: rufipes P. Lucas, 1846.

coriaceus Cameron, 1926b: 344 (Xantholinus; Type locality: Simla Hills; Theog, Kotgarh, 7000 to 7600 feet).

  • — Cameron, 1932: 29 (Xantholinus; characters; India).

  • — Scheerpeltz, 1933: 1308 (Xantholinus; catalog).

  • Distribution: India.

corsicus Coiffait, 1972, see: sardous Gridelli, 1947.

costaricensis Bernhauer, 1942: 23 (Xantholinus; Type locality: Kostarika: La Caja bei San José).

  • Distribution: Costa Rica.

crassiceps Sharp, 1885: 484 (Xantholinus; Type locality: Mexico: Guanajuato; Oaxaca, Cerro de Plumas. Guatemala: Capetillo).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • Distribution: Mexico, Guatemala.

crassicornis Hochhuth, 1851: 7 (Xantholinus; Type locality: Caucasien, Abastuman).

  • — Casey, 1906: 375 (Idiolinus).

  • — Reitter, 1908: 123 (Xantholinus; subgenus Typhlolinus; characters; Caucasus).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1309 (Xantholinus; catalog).

  • — Bordoni, 1971: 684 (Xantholinus; subgenus Megalinus; characters; Turkey).

  • — Coiffait, 1972: 261 (Xantholinus; subgenus Echinophallus; characters; references; Caucasus; Anatolia).

  • — Bordoni, 1972: 160 (Xantholinus; subgenus Echinophallus).

  • — Bordoni, 1973c: 155 (Xantholinus; subgenus Megalinus; notes; Romania).

  • — Korge, 1973: 5 (Xantholinus; characters; notes).

  • — Bordoni, 1975: 58 (Xantholinus; subgenus Echinophallus; notes).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Idiolinus).

  • Distribution: Turkey, Georgia.

crateris Bernhauer, 1939, see: Notolinus.

cribratus Fauvel, 1878, see: Zeteotomus.

cribripennis Fauvel, 1873, see: laevigatus Jacobsen, 1849.

cubensis Blackwelder, 1943: 480 (Xantholinus; Type locality: Cuba: Sierra Bonilla).

  • Distribution: West Indies.

cultus Broun, 1880, see: Pseudocorynus.

cunctator Sharp, 1889: 251 (Xantholinus; Type locality: Miyanoshita).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • Distribution: Japan.

cyaneipennis W. J. MacLeay, 1873, see: Eulissus.

cyanopterus Erichson, 1839, see: Eulissus.

dahli Blackwelder, 1944, see: Lepidophallus.

dalmatinus Rambousek, 1914, see: Vulda.

darlingtoni Blackwelder, 1943: 481 (Xantholinus; Type locality: Haiti: La Visite and vicinity, in the La Selle Range, elevation 5000 to 7000 feet).

  • Distribution: West Indies.

decorus Erichson, 1839b: 324 (Xantholinus; Type locality: Austria; Hungaria montana).

  • — Redtenbacher, 1849: 692 (Xantholinus; characters; Austria).

  • — Kraatz, 1857d: 639 (Xantholinus; characters; Germany).

  • — Redtenbacher, 1857: 181 (Xantholinus; characters; Austria).

  • — Redtenbacher, 1874: 220 (Xantholinus; characters; Austria).

  • — Ganglbauer, 1895: 483 (Xantholinus; characters; eastern Europe; Greece).

  • — Reitter, 1908: 117 (Xantholinus; subgenus Milichilinus; characters; eastern and southern Europe).

  • — Montandon, 1908: 79 (Xantholinus; Romania).

  • — Reitter, 1909: 137 (Xantholinus; subgenus Milichilinus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1309 (Xantholinus; catalog).

  • — Coiffait, 1956a: 72 (Xantholinus; subgenus Milichilinus; characters).

  • — Smetana, 1958: 115 (Xantholinus; subgenus Milichilinus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 164 (Xantholinus; subgenus Milichilinus; characters; central Europe).

  • — Horion, 1965: 97 (Xantholinus; subgenus Milichilinus, cited as Milichinus; Greece; Bosnia Hercegovina; Croatia; Italy; Romania; Hungary; Czech Republic; Slovakia; Ukraine; Austria).

  • — Coiffait, 1972: 256 (Xantholinus; subgenus Milichilinus; characters; references; central Europe; Italy; Balkans).

  • — Bordoni, 1972: 160 (Xantholinus; subgenus Milichilinus).

  • — Bordoni, 1973: 51 (Xantholinus; subgenus Milichilinus; Hungary).

  • — Bordoni, 1973c: 156 (Xantholinus; subgenus Milichilinus; Romania).

  • — Bordoni, 1975: 84 (Xantholinus; subgenus Milichilinus; notes).

  • — Bordoni, 1986b: 386 (Xantholinus; Slovakia; Serbia).

  • — Lucht, 1987: 103 (Xantholinus; checklist; Czechoslovakia; Austria).

  • — Tóth, 1989: 21 (Xantholinus; characters; Hungary).

  • — Janák, 1993a: 47 (Xantholinus; subgenus Milichilinus; Czech Republic; Slovakia).

  • Distribution: Europe.

densicephalus Bernhauer, 1938b: 51 (Xantholinus; Type locality: Chine-Nord).

  • Distribution: China.

densiceps Bernhauer, 1933: 28 (Xantholinus; Type locality: Szetschwan: Mukue Tatsienlu).

  • Distribution: China.

densipennis Bernhauer, 1934: 7 (Xantholinus; Type locality: Mittel-Szechuan: Hwa-Yin-Shan, 1600 m).

  • Distribution: China.

detryi Drugmand, 1990a: 87 (Xantholinus; subgenus Xantholinus; Type locality: Belgique: Antheit, Corphalie).

  • Distribution: Belgium.

devilleanus Scheerpeltz, 1956, see: appenninicola Steel, 1946.

devillei Coiffait, 1956a: 61 (Xantholinus; subgenus Echinophallus; Type locality: Tende, Alpes Maritimes).

  • — Coiffait, 1958: 23 (Xantholinus; subgenus Echinophallus; notes).

  • — Coiffait, 1972: 260 (Xantholinus; subgenus Echinophallus; characters; references; Italy).

  • — Bordoni, 1972: 160, 167 (Xantholinus; subgenus Echinophallus; characters; Italy).

  • — Bordoni, 1982: 253 (Xantholinus; subgenus Echinophallus; characters; Italy).

  • — Ciceroni, 1993: 72 (Xantholinus; Italy).

  • — Ciceroni and Zanetti, 1995: 25 (Xantholinus; Italy).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Idiolinus).

  • Distribution: Italy.

diabolicus Cameron, 1926, see: Mitomorphus.

difficilis Sharp, 1885: 482 (Xantholinus; Type locality: Guatemala, El Tumbador 2500 feet).

  • — Sharp, 1887: 791 (Xantholinus; Mexico).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • Distribution: Guatemala, Mexico.

dimidiatus LeConte, 1880, see: Neohypnus.

discretus Sharp, 1885: 481 (Xantholinus; Type locality: Guatemala, Volcan de Agua between 8500 and 10,500 feet; Panama, Volcan de Chiriqui 4000 to 6000 feet).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • Distribution: Guatemala, Panama.

dispilus Erichson, 1839, see: Thyreocephalus.

dissimilis Coiffait, 1956b: 143 (Xantholinus; Type locality: Tchécoslovaquie: Bysbrica [Note: Correct spelling probably Bystrica).

  • — Smetana, 1958: 113 (Xantholinus; subgenus Xantholinus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 166 (Xantholinus; subgenus Xantholinus; characters; central Europe).

  • — Horion, 1965: 102 (Xantholinus; subgenus Xantholinus; Slovakia; Germany).

  • — Smetana, 1966: 330 (Xantholinus; characters).

  • — Coiffait, 1972: 278 (Xantholinus; subgenus Xantholinus; characters; references; Czechoslovakia; Germany).

  • — Bordoni, 1972: 162 (Xantholinus; subgenus Meneidophallus).

  • — Lucht, 1987: 103 (Xantholinus; checklist; Germany; Czechoslovakia).

  • — Tóth, 1989: 30 (Xantholinus; characters).

  • — Janák, 1993a: 47 (Xantholinus; subgenus Meneidophallus; Czech Republic ?; Slovakia).

  • Distribution: Czech Republic ?, Slovakia, Germany.

distans Mulsant and Rey, 1853: 58 [= 1853a: 71] (Xantholinus; variety of tricolor; Type locality: les montagnes de Lyonnais et de la Loire).

  • — Mulsant and Rey, 1853: 58 (Xantholinus; variety of tricolor; characters; France).

  • — Kraatz, 1855: 23 (Xantholinus; variety of tricolor; notes).

  • — Kraatz, 1857d: 639 (Xantholinus; characters; Germany).

  • — Redtenbacher, 1857: 983 (Xantholinus; characters; Austria).

  • — Thomson, 1860: 191 (Xantholinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 302, cited linearis Thomson as a synonym of distans; Thomson did not describe the species as new, he attributed it to Olivier]; characters; Scandinavia).

  • — Fauvel, 1873b: 105 [= 1874a: 391] (Xantholinus; characters; France; Britain; Germany; Switzerland; Austria; Italy; Russia).

  • — Redtenbacher, 1874: 219 (Xantholinus; characters; Austria).

  • — Seidlitz, 1875: 281 (Xantholinus; characters; Baltic region).

  • — J. Sahlberg, 1876: 35 (Xantholinus; characters; Finland).

  • — Mulsant and Rey, 1877a: 279 (Xantholinus; characters; France).

  • — Ganglbauer, 1895: 484 (Xantholinus; characters; middle Europe).

  • — Reitter, 1908: 120 (Xantholinus; characters; central Europe).

  • — Reitter, 1909: 138 (Xantholinus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • — Portevin, 1929: 373 (Xantholinus; characters; France).

  • — Scheerpeltz, 1933: 1309 (Xantholinus; catalog).

  • — Steel, 1946a: 193 (Xantholinus; subgenus Idiolinus; characters; Britain).

  • — V. Hansen, 1952: 19 (Xantholinus; subgenus Xantholinus; characters).

  • — Coiffait, 1956a: 71 (Xantholinus; subgenus Helicophallus; characters in key).

  • — Smetana, 1958: 107 (Xantholinus; subgenus Xantholinus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 165 (Xantholinus; subgenus Helicophallus; characters; central Europe).

  • — Horion, 1965: 102 (Xantholinus; subgenus Helicophallus; France; Belgium; Belgium; Czech Republic; Germany; Austria).

  • — Coiffait, 1972: 302 (Xantholinus; subgenus Helicophallus; characters; references; central Europe; Czechoslovakia; Romania; Britain; France).

  • — Bordoni, 1973c: 155 (Xantholinus; subgenus Helicophallus; notes).

  • — Bordoni, 1975: 80 (Xantholinus; subgenus Helicophallus; notes).

  • — Lucht, 1987: 103 (Xantholinus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).

  • — Tóth, 1989: 26 (Xantholinus; characters).

  • — Janák, 1993a: 47 (Xantholinus; subgenus Helicophallus; Czech Republic).

  • Distribution: Europe, Russia.

schuleri Coiffait, 1958: 21 (Xantholinus; subgenus Helicophallus; Type locality: Bas-Rhin: Champ du Feu, 1099 m).

  • — Lohse, 1960a: 37, 38 (Xantholinus; characters; notes).

  • — Lohse, 1964: 165 (Xantholinus; subgenus Helicophallus; characters; central Europe).

  • — Horion, 1965: 103 (Xantholinus; subgenus Helicophallus; France; Germany).

  • — Coiffait, 1972: 302 (Xantholinus; subgenus Helicophallus; subspecies of distans; France).

  • — Bordoni, 1972: 164 (Xantholinus; subgenus Helicophallus).

  • — Bordoni, 1976d: 122 (Xantholinus; subspecies of distans; Netherlands).

  • — Lucht, 1987: 103 (Xantholinus; checklist; Germany; France; Benelux).

  • — Gerend and Braunert, 1997: 197 (Xantholinus; cited as valid species; notes; Luxembourg).

  • — Assing, Frisch, Kahlen, et al., 1998: 136 (Xantholinus; subspecies of distans; notes).

divisus Erichson, 1839, see: Ehomalolinus.

doderoi Rambousek, 1914, see: Vulda.

dubius W. J. MacLeay, 1873: 139 (Xantholinus; Type locality: Gayndah).

  • — Fauvel, 1877: 247 (Xantholinus; characters; Australia).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • Distribution: Australia.

dvoraki Coiffait, 1956: 163 (Xantholinus; Type locality: Tchélékovica [Note: The correct spelling is probably Čelákovice]).

  • — Smetana, 1958: 112 (Xantholinus; subgenus Xantholinus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 166 (Xantholinus; subgenus Xantholinus; characters; central Europe).

  • — Horion, 1965: 102 (Xantholinus; subgenus Xantholinus; Czech Republic; Slovakia; Austria).

  • — Smetana, 1966: 330 (Xantholinus; characters).

  • — Coiffait, 1972: 280 (Xantholinus; subgenus Xantholinus; characters; references; Czechoslovakia; Austria).

  • — Bordoni, 1972: 162 (Xantholinus; subgenus Meneidophallus).

  • — Bordoni, 1973: 46 (Xantholinus; subgenus Meneidophallus; Hungary).

  • — Bordoni, 1975: 68 (Xantholinus; subgenus Meneidophallus; notes; Siberia).

  • — Tóth, 1985: 176 (Xantholinus; notes; not recorded in Hungary).

  • — Lucht, 1987: 103 (Xantholinus; checklist; Czechoslovakia; Austria).

  • — Tóth, 1989: 29 (Xantholinus; characters).

  • — Janák, 1993a: 47 (Xantholinus; subgenus Meneidophallus; Czech Republic; Slovakia).

  • Distribution: Austria, Czechoslovakia, Russia.

lichtneckerti Tóth, 1985: 173 (Xantholinus; subgenus Meneidophallus; Type locality: Hungaria occidentalis: Velencei-tó: Sukoró).

  • — Lohse, 1987: 138 (Xantholinus; synonym of dvoraki).

  • — Tóth, 1989: 29 (Xantholinus; cited as valid species; [Note: I assume Tóth overlooked the synonymy by Lohse]; characters; Hungary).

ebneri Scheerpeltz, 1925, see: Gyrohypnus.

eichelbaumi Steel, 1949a: 270 (Xantholinus; replacement name for puncticeps Eichelbaum).

  • Distribution: Tanzania.

puncticeps Eichelbaum, 1913: 129 (Xantholinus; [preoccupied]; Type locality: Amani).

  • — Scheerpeltz, 1933: 1313 (Xantholinus; catalog).

  • — Steel, 1949a: 270 (Xantholinus; synonym of eichelbaumi).

elegans Olivier, 1795: 19 (Staphylinus; Type locality: forêt de Saint-Germain, Seine-et-Oise, France, Yvelines [Note: See neotype designation by Drugmand, 1994]. Originally cited from Paris).

  • — Gravenhorst, 1802: 46 (Staphylinus; [Note: Bernhauer and Schubert, 1914: 308, listed elegans Gravenhorst as a synonym of tricolor, but Gravenhorst did not describe elegans as new, he attributed it to Olivier]; characters).

  • — Latreille, 1804: 331 (Staphylinus; characters; France).

  • — Gravenhorst, 1806: 103 (Staphylinus; characters).

  • — Gyllenhal, 1810: 355 (Staphylinus; synonym of tricolor).

  • — C. Sahlberg, 1830: 334 (Staphylinus; synonym of tricolor).

  • — Griffith and Pidgeon, 1832: 292 (Xantholinus).

  • — Erichson, 1839a: 428 (Xantholinus; synonym of tricolor).

  • — Erichson, 1839b: 332 (Xantholinus; synonym of tricolor).

  • — Kolenati, 1846: 15 (Xantholinus; synonym of tricolor).

  • — Küster, 1848: 10 (Xantholinus; synonym of tricolor).

  • — Rosenhauer, 1856: 70 (Xantholinus; notes).

  • — Fairmaire and Laboulbène, 1856: 501 (Xantholinus; characters; France).

  • — Fairmaire and Laboulbène, 1856: 502 (Xantholinus; synonym of tricolor).

  • — Kraatz, 1857d: 638 (Xantholinus; synonym of tricolor).

  • — Redtenbacher, 1857: 182 (Xantholinus; characters; Germany).

  • — Fauvel, 1873b: 103 [= 1873a: 389] (Xantholinus; characters; France; Spain).

  • — Mulsant and Rey, 1877a: 271 (Xantholinus; characters; France).

  • — Ganglbauer, 1895: 483 (Xantholinus; synonym of tricolor).

  • — Reitter, 1908: 120 (Xantholinus; synonym of meridionalis).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; synonym of tricolor).

  • — Portevin, 1929: 372 (Xantholinus; characters; France).

  • — C. Koch, 1937: 87 (Xantholinus; [Note: Cited as meridionalis Nordmann but according to Drugmand, 1994: 245, a misidentification of elegans Olivier]; characters; Italy).

  • — Steel, 1946a: 194 (Xantholinus; [Note: Cited as tricolor but according to Drugmand, 1994: 245, a misidentification of elegans]; characters; Britain).

  • — Smetana, 1958: 109 (Xantholinus; synonym of tricolor).

  • — Lohse, 1964: 164 (Xantholinus; subgenus Purrolinus; [Note: Cited as semirufus but according to Drugmand, 1994: 245, a misidentification of elegans]; characters; central Europe).

  • — Bordoni, 1982: 256 (Xantholinus; synonym of tricolor).

  • — Drugmand, 1994: 245 (Xantholinus; neotype designation: forêt de Saint-Germain, Seine-et-Oise, France, Yvelines; valid species; characters; Great Britain; France; Belgium; Luxembourg; Italy; Germany; Austria; Poland).

  • — Ciceroni and Zanetti, 1995: 25 (Xantholinus; Italy).

  • Distribution: Europe.

meridionalis Lacordaire, 1835: 413 (Xantholinus; Type locality: Paris).

  • — Erichson, 1839b: 323 (Xantholinus; synonym of elegans).

  • — Rosenhauer, 1856: 70 (Xantholinus; synonym of elegans).

  • — Fairmaire and Laboulbène, 1856: 501 (Xantholinus; synonym of elegans).

  • — Fauvel, 1873b: 104 [= 1874a: 391] (Xantholinus; synonym of tricolor).

  • — Mulsant and Rey, 1877a: 273 (Xantholinus; synonym of tricolor).

  • — Ganglbauer, 1895: 483 (Xantholinus; synonym of tricolor).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; synonym of tricolor).

  • — Smetana, 1958: 109 (Xantholinus; synonym of tricolor).

  • — Coiffait, 1972: 284 (Xantholinus; synonym of tricolor).

  • — Bordoni, 1972: 168 (Xantholinus; synonym of tricolor).

  • — Bordoni, 1975: 62 (Xantholinus; synonym of tricolor).

  • — Bordoni, 1982: 256 (Xantholinus; synonym of tricolor).

  • — Drugmand, 1994: 245 (Xantholinus; synonym of elegans).

paskoviensis Reitter, 1908: 120 [= 1908a: 23] (Xantholinus; variety of meridionalis; Type locality: Mähren).

  • — Reitter, 1909: 138 (Xantholinus; variety of meridionalis; characters).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; variety of meridionalis; cited as pascoviensis).

  • — Scheerpeltz, 1933: 1312 (Xantholinus; variety of meridionalis).

  • — Steel, 1950c: 141 (Xantholinus; synonym of semirufus).

  • — Smetana, 1958: 110 (Xantholinus; synonym of jarrigei).

  • — Coiffait, 1972: 282 (Xantholinus; subgenus Purrolinus; variety of jarrigei).

  • — Bordoni, 1982: 265 (Xantholinus; synonym of jarrigei).

  • — Drugmand, 1994: 245 (Xantholinus; synonym of elegans).

jarrigei Coiffait, 1956a: 67 (Xantholinus; subgenus Purrolinus; proposed for Erichson's use of elegans and Steel's use of semirufus; Type locality: France; Angleterre; Espagne: environs de Madrid; Catalogne; Italie).

  • — Erichson, 1839b: 323 (Xantholinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 305, cited elegans Erichson as a synonym of meridionalis Nordmann; Erichson did not describe the species as new, he attributed it to Olivier]; characters; France; Spain).

  • — Last, 1950a: 139, 141 (Xantholinus; [Note: Misidentification of meridionalis according to an editor's note on page 138; Steel, 1950c, regarded Last's record of meridionalis to be semirufus]; characters; Britain; also listed from Holland, France, Italy, Moravia).

  • — Last, 1950a: 139, 140 (Xantholinus; [Note: Misidentification of tricolor according to an editor's note on page 138; Steel, 1950c, regarded Last's record of tricolor to be semirufus]; characters; Britain; also listed from northern and central Europe).

  • — Steel, 1950c: 141 (Xantholinus; [Note: Misidentification of semirufus and regarded by Coiffait, 1956a: 67, as a synonym of jarrigei]; valid species; discussion; list of synonyms and misidentification and found in Britain and France).

  • — Coiffait, 1958: 23 (Xantholinus; subgenus Purrolinus; France).

  • — Smetana, 1958: 110 (Xantholinus; subgenus Xantholinus; characters; ecology; Czechoslovakia).

  • — Lohse, 1960a: 37, 38 (Xantholinus; cited as semirufus Steel with jarrigei and pascoviensis as synonyms; characters; notes).

  • — Lohse, 1964: 164 (Xantholinus; synonym of semirufus).

  • — M. Dvořák, 1965: 88 (Xantholinus; Slovakia).

  • — Coiffait, 1972: 282 (Xantholinus; subgenus Purrolinus; characters; references; France; England; Iberia; Italy).

  • — Bordoni, 1972: 176 (Xantholinus; synonym of meridionalis).

  • — Bordoni, 1976d: 121 (Xantholinus; cited as meridionalis Nordmann sensu Gridelli with jarrigei listed as synonym; Netherlands).

  • — Pope, 1977: 28 (Xantholinus; subgenus Polydontophallus; Britain).

  • — Bordoni, 1977a: 15 (Xantholinus; Spain).

  • — Bordoni, 1982: 265 (Xantholinus; subgenus Polydontophallus; characters; Italy).

  • — Bordoni, 1985a: 62 (Xantholinus; anatomical studies of labrum, aedeagus, and male genital segment).

  • — Lucht, 1987: 103 (Xantholinus; cited as “semirufus [Rtt.] Steel, 1950”; checklist; Germany; Poland; Czechoslovakia; Austria; Benelux).

  • — Schülke and Uhlig, 1988: 11 (Xantholinus; cited as semirufus Steel and as a valid species; notes; Germany).

  • — Tóth, 1989: 24 (Xantholinus; characters; Hungary).

  • — Janák, 1993a: 47 (Xantholinus; subgenus Polydontophallus; Czech Republic; Slovakia).

  • — Assing, 1993: 26 (Xantholinus; bionomics; Germany).

  • — Ciceroni, 1993: 73 (Xantholinus; Italy).

  • — Drugmand, 1994: 245 (Xantholinus; synonym of elegans).

  • — Hodge and Jones, 1995: 38 (Xantholinus; characters; Britain).

  • — Ciceroni and Zanetti, 1995: 25 (Xantholinus; synonym of elegans).

elianae Jarrige, 1941, see: Lemiganus.

elongatus Heer, 1834 [Note: Nomen nudum], see: longiventris Heer, 1839.

elongatus Fauvel, 1866, see: Lepidophallus.

enigmaticus Coiffait, 1968: 151 (Xantholinus; Type locality: Gabon: Belinga).

  • Distribution: Gabon.

episcopalis Lacordaire, 1835, see: Leptacinus.

episcopalis Fauvel, 1877, see: brasiliensis Steel, 1949.

eppelsheimi Bernhauer and Schubert, 1914, see: Thyreocephalus.

eppelsheimi Cameron, 1925, see: Thyreocephalus.

erythropterus Erichson, 1839b: 320 (Xantholinus; Type locality: Nova Hollandia).

  • — Fauvel, 1877: 240 (Xantholinus; characters; Australia: New South Wales; Queensland).

  • — Fauvel, 1878g: 538 (Xantholinus; Australia).

  • — Olliff, 1887: 480 (Xantholinus; characters; Australia).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1309 (Xantholinus; catalog).

  • Distribution: Australia.

cervinipennis W. J. MacLeay, 1873: 138 (Xantholinus; Type locality: Gayndah).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; synonym of erythropterus).

  • — Lea, 1925: 219 (Eulissus; synonym of erythopterus).

euboicus Korge, 1973: 7 (Xantholinus; subgenus Purrolinus; Type locality: Euböa, Griechenland, Dirphys bei Steni, 1000–1200 m).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus).

  • Distribution: Greece.

excelsus Bernhauer, 1939, see: Nudobius.

fageli Coiffait, 1971: 432 (Xantholinus; subgenus Purrolinus; Type locality: Liban: Karabata, 1200–1400 m).

  • — Coiffait, 1972: 283 (Xantholinus; subgenus Purrolinus; characters; Lebanon).

  • — Bordoni, 1972: 161 (Xantholinus; subgenus Polydontophallus).

  • Distribution: Lebanon.

fallaciosus Gridelli, 1947, see: Vulda.

fallax Sachse, 1852: 125 (Xantholinus; Type locality: Vorgeb. d. g. Hoffnung).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • Distribution: South Africa.

capensis Boheman, 1858: 27 (Xantholinus; Type locality: Promontorium Bonae Spei).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; synonym of fallax).

fasciatus Hochhuth, 1849, see: Calontholinus.

feae Fauvel, 1895, see: Thyreocephalus.

fenyesi Bernhauer, 1910b: 376 (Xantholinus; Type locality: Mexiko: Vera Cruz, Cordoba).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • Distribution: Mexico.

ferox Harold, 1881, see: Eulissus.

festivus Erichson, 1839, see: Agerodes.

flagellifer Sharp, 1885: 485 (Xantholinus; Type locality: Panama, Bugaba 800 to 1500 feet).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • Distribution: Panama.

flavipennis Erichson, 1839, see: Homalolinus.

flavipennis Redtenbacher, 1849, see: Hypnogyra.

flavocinctus Hochhuth, 1849, see: Leptophius.

floridae Duvivier, 1883, see: Neohypnus.

formosae Cameron, 1949, see: Megalinus.

forsteri Scheerpeltz, 1960c: 96 (Xantholinus; Type locality: Bolivien: Yungas de Arepucho, Sihuencas, 2200–2500 m).

  • Distribution: Bolivia.

fortepunctatus Motschulsky, 1860: 564 (Xantholinus; Type locality: Georgie: Piatigonk; [Note: See lectotype designation by Bordoni, 1999]. Originally cited from les plaines qui avoisinent le Caucase).

  • — Hochhuth, 1862: 66 (Xantholinus; characters; Caucasus).

  • — Reitter, 1908: 121 (Xantholinus; characters; Caucasus; Armenia; Talysh).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1309 (Xantholinus; catalog).

  • — Jarrige, 1971: 493 (Xantholinus; Iran).

  • — Bordoni, 1975: 72 (Xantholinus; characters; synonym of semirufus).

  • — Bordoni, 1999: 1, 4 (Xantholinus; lectotype designation: Georgie: Piatigonk).

  • Distribution: Armenia, Azerbaijan, Iran.

semirufus Reitter, 1901a: 68 (Xantholinus; Type locality: Samarkand; [Note: See lectotype designation by Bordoni, 1999]).

  • — Reitter, 1908: 120 (Xantholinus; synonym of meridionalis).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • — Horion, 1965: 99 (Xantholinus; subgenus Purrolinus; Britain; Holland; Belgium; France; Spain; Italy; Bosnia Hercegovina; Germany; Austria; Czech Republic).

  • — Szujecki, 1969a: 254 (Xantholinus; notes; Poland).

  • — Coiffait, 1972: 293 (Xantholinus; subgenus Heterolinus; characters; Uzbekistan).

  • — Bordoni, 1975: 70 (Xantholinus; subgenus Heterolinus; notes; Tadjkistan; Crimea).

  • — Nowosad, 1990: 97 (Xantholinus; collected from mole nest; notes; Poland).

  • — Drugmand, 1994: 245, 248 (Xantholinus; cited as valid species (p. 248) and synonym of elegans (p. 245); characters; Uzbekistan).

  • — Assing and Schülke, 1999: 8 (Xantholinus; not a synonym of jarrigei).

  • — Bordoni, 1999: 1, 4 (Xantholinus; subgenus Heterolius; lectotype designation; synonym of fortepunctatus).

  • Distribution: Tajikistan, Uzbekistan, Ukraine, Poland.

franzi Bordoni, 1973c: 154 (Xantholinus; subgenus Acanthophallus; Type locality: Roumanie: Azuga).

  • — Tóth, 1989: 26 (Xantholinus; characters).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus).

  • Distribution: Romania.

frigidus Dufour, 1843, see: Atrecus.

fucosus Casey, 1885, see: Neohypnus.

fuenteanus Reitter, 1901a: 68 (Xantholinus; Type locality: Spanien: Pozuelo de Caletrava; [Note: See lectotype designation by Drugmand, 1994]).

  • — Reitter, 1908: 120 (Xantholinus; variety of meridionalis; characters; Spain).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1312 (Xantholinus; variety of meridionalis).

  • — Coiffait, 1972: 284 (Xantholinus; subgenus Purrolinus; valid species; characters; references; Iberia).

  • — Bordoni, 1976c: 93 (Xantholinus; valid species; Spain).

  • — Drugmand, 1994: 249 (Xantholinus; valid species; lectotype designation; characters; Spain).

  • Distribution: Spain.

fuliginosus Sharp, 1885: 478 (Xantholinus; Type locality: Mexico).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; catalog).

  • Distribution: Mexico.

fumatus Sharp, 1885: 480 (Xantholinus; Type locality: Mexico, San Andres, Guanajuato, Las Vigas, Oaxaca).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; catalog).

  • Distribution: Mexico.

furvus Wendeler, 1956a: 227 (Xantholinus; Type locality: Brasilien: Nova Teutonia).

  • Distribution: Brazil.

fusciceps LeConte, 1880, see: Neohypnus.

fuscipennis Cameron, 1922: 115 (Xantholinus; variety of vilis; Type locality: Jamaica).

  • — Scheerpeltz, 1933: 1314 (Xantholinus; variety of pusillus).

  • — Blackwelder, 1943: 477 (Xantholinus; valid species; characters; Jamaica).

  • Distribution: West Indies.

gagatinus Erichson, 1839, see: Gauropterus.

gallicus Coiffait, 1956a: 64 (Xantholinus; Type locality: Basses-Pyrénées: Col du Somport. Also cited from Spain).

  • — Smetana, 1958: 113 (Xantholinus; subgenus Xantholinus; characters; ecology).

  • — Lohse, 1960a: 38 (Xantholinus; characters; notes).

  • — Brakman, 1962: 47 (Xantholinus; characters; Holland).

  • — Lohse, 1964: 165 (Xantholinus; subgenus Xantholinus; characters; central Europe).

  • — Coiffait, 1972: 274 (Xantholinus; subgenus Xantholinus; characters; references; France; Belgium; Spain).

  • — Bordoni, 1972c (Xantholinus; subgenus Paraphallus; correction of subgeneric assignment in Bordoni, 1972: 162, 188).

  • — Bordoni, 1976d: 121 (Xantholinus; Netherlands).

  • — Pope, 1977: 28 (Xantholinus; Britain).

  • — Drugmand and Haghebaert, 1987: 325 (Xantholinus; Belgium).

  • — Lucht, 1987: 103 (Xantholinus; checklist; Germany; Benelux).

  • — Drugmand, 1990a: 91 (Xantholinus; characters; Belgium).

  • — Hodge and Jones, 1995: 38 (Xantholinus; notes; Britain).

  • — Outerelo, Gamarra, and Salgado, 1998: 129 (Xantholinus; subgenus Xantholinus; collected from cave; Spain).

  • Distribution: Europe.

garganicus Bernhauer, 1908, see: Vulda.

gastraeus Gistel, 1857: 60 (Xantholinus; [Note: Nomen dubium]; Type locality: Helvetia).

  • Distribution: Switzerland.

gestroi Fauvel, 1895, see: Thyreocephalus.

glabroides Coiffait, 1962, see: Hypnogyra.

goudoti Fauvel, 1905, see: Eulissus.

gracilis Sperk, 1835, see: Scopaeus.

gracilis Boheman, 1858, see: bonariensis Gemminger and Harold, 1868.

graecus Kraatz, 1858a: 64 (Xantholinus; Type locality: Attica; Nauplia, Creta; Athen).

  • — Saulcy, 1864: 641 (Xantholinus; characters; notes; Lebanon).

  • — Fauvel, 1878: 118 [= 1878a: 38] (Xantholinus; Algeria).

  • — Fauvel, 1886: 50 [= 1886a: 42] (Xantholinus; Algeria).

  • — Fauvel, 1902b: 102 (Xantholinus; Morocco; Algeria; Egypt).

  • — Reitter, 1908: 22 (Xantholinus; variety of linearis; characters; Greece; Caucasus; Sicily).

  • — Bernhauer and Schubert, 1914: 303 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1310 (Xantholinus; catalog).

  • — Gridelli, 1947: 79 (Xantholinus; characters; Greece; Cyprus; Sicily; Macedonia).

  • — Scheerpeltz, 1964a: 299 (Xantholinus; subgenus Acanthophallus; Crete).

  • — Scheerpeltz, 1965b: 400 (Xantholinus; Sicily).

  • — Bordoni, 1971: 682 (Xantholinus; subgenus Acanthophallus; Turkey).

  • — Coiffait, 1972: 293 (Xantholinus; subgenus Acanthophallus; characters; references; Greece; Crete; Cyprus; Turkey; Sicily).

  • — Bordoni, 1972: 161, 180 (Xantholinus; subgenus Acanthophallus; characters; Italy).

  • — Bordoni, 1975e: 145 (Xantholinus; notes; Corfu).

  • — Bordoni, 1976c: 96 (Xantholinus; Greece; Cyprus; Corfu).

  • — Bordoni, 1976d: 121 (Xantholinus; Corfu).

  • — Bordoni, 1982: 276 (Xantholinus; subgenus Acanthophallus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; Italy).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus).

  • — Assing and Wunderle, 2001: 36 (Xantholinus; Cyprus).

  • Distribution: Lebanon, Israel, Italy, Albania, Macedonia, Greece, Cyprus, Turkey, Caucasus.

judaeus Coiffait, 1956a: 68 (Xantholinus; subgenus Acanthophallus; Type locality: Jérusalem. Also cited from Turkey).

  • — Bordoni, 1971: 687 (Xantholinus; subgenus Acanthophallus; Turkey).

  • — Coiffait, 1972: 293 (Xantholinus; subgenus Acanthophallus; subspecies of graecus; characters; Lebanon; Palestine; Turkey).

  • — Bordoni, 1972: 161 (Xantholinus; subgenus Acanthophallus).

  • — Bordoni, 1975e: 146 (Xantholinus; subspecies of graecus; notes; Rodi).

calcidicus Bordoni, 1973f: 75 (Xantholinus; subgenus Acanthophallus; subspecies of graecus; Type locality: Grecia: Macedonia, Penisola di Akti Monte Athos).

  • — Bordoni, 1975e: 145 (Xantholinus; subspecies of graecus; notes; Crete).

  • — Bordoni, 1976c: 96 (Xantholinus; subspecies of graecus; notes; Albania; Turkey).

  • — Bordoni, 1982: 278 (Xantholinus; subgenus Acanthophallus; subspecies of graecus; characters; Italy).

  • — Ciceroni, 1993: 73 (Xantholinus; subspecies of graecus; notes; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; subspecies of graecus; cited as “sottospecie dubbia”; Italy).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus; subspecies of graecus).

gridellii Coiffait, 1956a: 68 (Xantholinus; subgenus Acanthophallus; Type locality: Liban: Saïda).

  • — Scheerpeltz, 1956c: 1099 (Xantholinus; subgenus Acanthophallus; Cyprus).

  • — Coiffait, 1970e: 62 (Xantholinus; Cyprus).

  • — Coiffait, 1972: 290 (Xantholinus; subgenus Acanthophallus; characters; Lebanon; Palestine; Cyprus).

  • — Bordoni, 1972: 161 (Xantholinus; subgenus Acanthophallus).

  • — Bordoni, 1976c: 94 (Xantholinus; Syria; Lebanon; Algeria).

  • — Bordoni, 1976d: 121 (Xantholinus; Algeria).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus).

  • Distribution: Algeria, Cyprus, Turkey, Lebanon, Israel, Syria.

carius Coiffait, 1972: 290 (Xantholinus; subgenus Acanthophallus; subspecies of gridellii; Type locality: Marmaris, Anatolie méridionale).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus; subspecies of gridellii).

guatemalenus Sharp, 1885: 485 (Xantholinus; Type locality: Guatemala, San Geronimo).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • Distribution: Guatemala.

gularis LeConte, 1880, see: Hypnogyra.

haematodes Kolenati, 1846: 14 (Xantholinus; Type locality: Elisabethopol, Karabagh Transcaucasiae; [Note: See lectotype designation by Bordoni, 1999]. Originally cited from Elisabethopol et Karabagh Transcaucasiae).

  • — Hochhuth, 1849: 108 (Xantholinus; characters; provincia Elisabethopol and Karabagh, Transcaucasia).

  • — Reitter, 1908: 122 (Xantholinus; subgenus Typhlolinus; characters).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1310 (Xantholinus; catalog).

  • — Coiffait, 1972: 309 (Xantholinus; characters; references; Caucasus).

  • — Bordoni, 1972a: 131 (Xantholinus; subgenus Tetralinus).

  • — Bordoni, 1975: 69 (Xantholinus; subgenus Tetralinus; type species of Tetralinus; characters; Daghestan).

  • — Bordoni, 1999: 2, 4 (Xantholinus; subgenus Tetralinus; lectotype designation: Elisabethopol, Karabagh Transcaucasiae).

  • Distribution: Russia, Azerbaijan.

haemorrhous Fauvel, 1878, see: Eulissus.

haitius Blackwelder, 1943: 484 (Xantholinus; Type locality: Haiti: Étang Lachaux on the southwest peninsula).

  • Distribution: West Indies.

hamatus Say, 1830, see: Neohypnus.

hanstroemi Scheerpeltz, 1974: 127 (Xantholinus; Type locality: Natal: Karkloof).

  • Distribution: South Africa.

haroldi Reitter, 1873, see: Gyrohypnus.

hebraicus Reiche and Saulcy, 1856, see: Leptophius.

heinzi Coiffait, 1970: 289 (Xantholinus; subgenus Echinophallus; Type locality: Col sud Ikizdere, 2000–2600 m, Anatolie septentrionale).

  • — Bordoni, 1971: 685 (Xantholinus; subgenus Echinophallus; Turkey).

  • — Coiffait, 1972: 261 (Xantholinus; subgenus Echinophallus characters; references; Anatolia).

  • — Bordoni, 1972a: 128 (Xantholinus; subgenus Toxophallus).

  • — Bordoni, 1972: 160 (Xantholinus; subgenus Toxophallus).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Idiolinus).

  • Distribution: Turkey.

hesperius Erichson, 1839, see: Lepidophallus.

heterocephalus Fauvel, 1905b: 171 (Xantholinus; Type locality: Diego-Suarez).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • Distribution: Madagascar.

himalayensis Coiffait, 1984e: 375 (Xantholinus; Type locality: Nepal oriental: Jaljale Himal, 2680 m).

  • Distribution: Nepal.

hochhuthi Bordoni, 1975: 63 (Xantholinus; subgenus Acanthophallus; Type locality: Caucaso occidentale, Krasnaja Poljana).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus).

  • Distribution: Russia (Caucasus).

hoffmanni Bernhauer, 1928, see: Hypnogyra.

holdhausi Bernhauer, 1908, see: Vulda.

holomelas Perroud and Montrousier, 1864, see: Thyreocephalus.

hongkongensis Redtenbacher, 1867, see: Thyreocephalus.

hottentottus Sachse, 1852, see: Notolinus.

hova Fauvel, 1905, see: Gauropterus.

humeralis Erichson, 1839b: 327 (Xantholinus; Type locality: Americae meridionalis ins. St. Johannus).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Blackwelder, 1943: 479 (Xantholinus; characters; Cuba; Haiti; Puerto Rico; St. John; St. Croix; Antigua; Montserrat; Dominica; St. Vincent).

  • Distribution: West Indies.

puncticeps Jacquelin du Val, 1857a: 35 (Xantholinus; [preoccupied]; Type locality: Not cited, presumably Cuba).

  • — Chevrolat and Fauvel, 1863: 451 (Xantholinus; characters; Cuba).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; synonym of punctiger).

  • — Blackwelder, 1943: 479 (Xantholinus; synonym of humeralis).

punctiger Gemminger and Harold, 1868: 603 (Xantholinus; replacement name for puncticeps Jacquelin du Val).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Blackwelder, 1943: 479 (Xantholinus; synonym of humeralis).

humerosus Bernhauer, 1934: 7 (Xantholinus; Type locality: Mittel-Szechuan: Hwa-Yin-Shan, 1600 m).

  • Distribution: China.

humidicola Gistel, 1857: 25 (Xantholinus; [Note: Nomen dubium]; Type locality: Styria).

  • Distribution: Austria.

humilis Sharp, 1885: 481 (Xantholinus; Type locality: Mexico, Jacale, Jalapa; Guatemala, Capetillo, Cerro Zunil 4000 to 5000 feet).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • Distribution: Mexico, Guatemala.

hungaricus Reitter, 1908, see: azuganus Reitter, 1908.

hydrocephalus Fauvel, 1901: 83 (Xantholinus; Type locality: Colombie: Bonda; Trinidad).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Blackwelder, 1943: 489 (Xantholinus; characters; Trinidad).

  • Distribution: Colombia, West Indies.

hypsibathus Bernhauer, 1915b: 51 (Xantholinus; Type locality: Himalaya: Kulu, 7000 Fuss).

  • — Cameron, 1932: 29 (Xantholinus; characters; India).

  • — Scheerpeltz, 1933: 1311 (Xantholinus; catalog).

  • — Coiffait, 1972: 308 (Xantholinus; characters; Himalaya).

  • Distribution: India.

iablokoffi Coiffait, 1975: 31 (Xantholinus; subgenus Heterolinus; Type locality: Dombai, Koubans).

  • Distribution: Georgia.

ibericus Steel, 1950c: 141 (Xantholinus; replacement name for meridionalis Nordmann. Species restricted to southern Spain).

  • — Coiffait, 1956a: 66 (Xantholinus; subgenus Purrolinus; characters in key; Spain).

  • — Coiffait, 1972: 284 (Xantholinus; synonym of fuenteanus).

  • — Bordoni, 1972: 161 (Xantholinus; subgenus Polydontophallus).

  • — Drugmand, 1994: 248 (Xantholinus; valid species; characters; Spain).

  • Distribution: Spain.

meridionalis Nordmann, 1837: 117 (Gyrohypnus; [preoccupied]; Type locality: Andalusia).

  • — Erichson, 1839b: 323 (Xantholinus; synonym of elegans).

  • — Reitter, 1908: 120 (Xantholinus; characters).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1312 (Xantholinus; catalog).

  • — Paulian, 1941: 213 (Xantholinus; larval characters).

  • — Gridelli, 1950: 7 (Xantholinus; characters; Italy).

  • — Coiffait, 1972: 284 (Xantholinus; synonym of fuenteanus).

  • — Bordoni, 1972: 161, 176 (Xantholinus; subgenus Polydontophallus; characters; Italy).

  • — Osella and Zanetti, 1975: 107 (Xantholinus; collected from nest of Talpa europaea; notes; Italy).

  • — Bordoni, 1976c: 92 (Xantholinus; notes; Spain).

  • — Drugmand, 1994: 248 (Xantholinus; synonym of ibericus).

ilgazensis Coiffait, 1966: 23 (Xantholinus; subgenus Helicophallus; Type locality: Monts Ilgaz, 2000–2300 m, Turquie d'Asie).

  • — Herman, 2001: 32 (Xantholinus; synonymic note).

  • Distribution: Turkey.

ilgazensis Coiffait, 1971: 434 (Xantholinus; subgenus Helicophallus; [preoccupied]; Type locality: Monts Ilgaz Dag, 2000–2300 m, Anatolie).

  • — Bordoni, 1971: 687 (Xantholinus; subgenus Helicophallus; Turkey).

  • — Bordoni, 1972: 164 (Xantholinus; subgenus Helicophallus).

  • — Coiffait, 1972: 306 (Xantholinus; subgenus Helicophallus; characters; Turkey).

  • — Herman, 2001: 32 (Xantholinus; synonym of ilgazensis).

ilgazensis Coiffait, 1971, see: ilgazensis Coiffait, 1966.

illucens Erichson, 1839b: 315 (Xantholinus; Type locality: Columbia).

  • — Scriba, 1855: 299 (Xantholinus; Venezuela).

  • — Fauvel, 1891: 105 (Xantholinus; Venezuela; Puerto Rico).

  • — Bernhauer and Schubert, 1914: 311 (Eulissus; catalog).

  • — Blackwelder, 1943: 488 (Xantholinus; characters; Puerto Rico; Montserrat; Guadeloupe; Dominica; St. Lucia; St. Vincent; Grenada; Tobago; Trinidad).

  • — Frank, 1982: 32 (Xantholinus; list of parasites).

  • Distribution: Colombia, Venezuela, West Indies.

impressifrons Guérin-Méneville, 1844, see: Agerodes.

impunctus Blackwelder, 1943: 487 (Xantholinus; Type locality: Cuba: Soledad near Cienfuegos).

  • Distribution: West Indies.

inclinans Walker, 1859, see: Gabronthus.

indianensis Blatchley, 1910, see: Xestolinus.

indicus Fauvel, 1895, see: Thyreocephalus.

infimus Coiffait, 1962a: 271 (Xantholinus; subgenus Xantholinus; Type locality: Grèce, environs de Trikala).

  • — Coiffait, 1965e: 120 (Xantholinus; Turkey).

  • — Bordoni, 1971: 686 (Xantholinus; subgenus Xantholinus; Turkey).

  • — Coiffait, 1972: 271 (Xantholinus; subgenus Xantholinus; characters; Greece; Turkey).

  • — Bordoni, 1972c (Xantholinus; subgenus Paraphallus; correction of subgeneric assignment in Bordoni, 1972: 162, 188).

  • — Bordoni, 1986a: 117 (Xantholinus; subgenus Paraphallus; Bulgaria).

  • Distribution: Bulgaria, Greece, Turkey.

infuscatus Bernhauer, 1910, see: rufipennis Erichson, 1839.

innocuus Fauvel, 1905b: 174 (Xantholinus; Type locality: forêt de Moramanga; Tananarive; forêt Tanala).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • Distribution: Madagascar.

inopinatus Cameron, 1951c: 400 (Xantholinus; Type locality: Uganda: Kampala).

  • Distribution: Uganda.

inoptatus Bernhauer, 1916g: 265 (Xantholinus; Type locality: Bolivien: Yuracarès).

  • — Scheerpeltz, 1933: 1311 (Xantholinus; catalog).

  • Distribution: Bolivia.

inseriatus Bernhauer, 1934b: 116 (Xantholinus; Type locality: Argentinien: Misiones, Loreto).

  • Distribution: Argentina.

insignis Erichson, 1839, see: Agerodes.

insularis Bernhauer, 1904a: 232 (Xantholinus; Type locality: Madagaskar).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1311 (Xantholinus; catalog).

  • Distribution: Madagascar.

insulatus Cameron, 1922: 115 (Xantholinus; Type locality: Grenada and Mustique).

  • — Scheerpeltz, 1933: 1311 (Xantholinus; [Note: Cited as insularis]; catalog).

  • — Blackwelder, 1943: 476 (Xantholinus; characters; Grenada; Grenadines).

  • Distribution: West Indies.

intermedius Küster, 1848, see: Gauropterus.

interocularis Eppelsheim, 1895, see: Thyreocephalus.

inuus Fauvel, 1900: 58 (Xantholinus; Type locality: Algérie: gorges de la Chiffa; Teniet-el-Had).

  • — Fauvel, 1902b: 101 (Xantholinus; Algeria).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Coiffait, 1956a: 67 (Xantholinus; subgenus Purrolinus; characters in key; Algeria).

  • — Coiffait, 1962b: 398, 404 (Xantholinus; subgenus Purrolinus; characters; Algeria).

  • — Coiffait, 1972: 285 (Xantholinus; subgenus Purrolinus; characters; references; Algeria).

  • — Bordoni, 1972: 161 (Xantholinus; subgenus Purrolinus).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus).

  • Distribution: Algeria.

istriensis Rambousek, 1924, see: Vulda.

jamaicensis Blackwelder, 1943: 487 (Xantholinus; Type locality: Jamaica: Kingston).

  • Distribution: West Indies.

japonicus Sharp, 1874, see: Lepidophallus.

jarrigei Coiffait, 1956, see: elegans Olivier, 1795.

javanicus Cameron, 1925a: 184 (Xantholinus; Type locality: Tjibodas, 1400 m).

  • — Scheerpeltz, 1933: 1311 (Xantholinus; catalog).

  • Distribution: Indonesia.

jeanneli Bernhauer, 1939a: 77 (Xantholinus; Type locality: Kenya: Mt. Elgon, prairies alpines, Vers. Est, 3900–4000 m).

  • Distribution: Kenya.

judaeus Coiffait, 1956, see: graecus Kraatz, 1858.

kambaitiensis Scheerpeltz, 1965: 194 (Xantholinus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).

  • Distribution: Myanmar.

kanalensis Fauvel, 1889: 261 (Xantholinus; Type locality: Kanala; Yahoué).

  • — Fauvel, 1903d: 267 (Xantholinus; New Caledonia).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • Distribution: New Caledonia.

kaszabi Bordoni, 1973: 49 (Xantholinus; subgenus Helicophallus; Type locality: Sátor nell'Ungheria settentrionale).

  • — Tóth, 1989: 26 (Xantholinus; characters; Hungary).

  • — Janák, 1993a: 47 (Xantholinus; subgenus Helicophallus; Slovakia).

  • Distribution: Hungary.

kazachstanicus Janák, 1979: 111 (Xantholinus; subgenus Acanthophallus; Type locality: U.R.S.S.: Kazakhstan: mont Medeo, 2000 m, près d'Alma-Ata).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus).

  • Distribution: Kazakhstan.

khnzoriani Coiffait, 1966f: 199 (Xantholinus; subgenus Heterolinus; Type locality: SSSR, Armeniia, Aparan).

  • — Coiffait, 1972: 294 (Xantholinus; subgenus Heterolinus; characters; Armenia).

  • — Bordoni, 1972: 162 (Xantholinus; subgenus Heterolinus).

  • — Bordoni, 1975: 70 (Xantholinus; subgenus Heterolinus; notes).

  • Distribution: Armenia.

kiesenwetteri Sachse, 1852, see: Nudobius.

kirghisicus Coiffait, 1966f: 200 (Xantholinus; subgenus Helicophallus; Type locality: SSSR, Kirgiziia, Sary-Chelek).

  • — Coiffait, 1972: 297 (Xantholinus; subgenus Helicophallus; characters; Kirghiztan).

  • — Bordoni, 1972: 164 (Xantholinus; subgenus Helicophallus; cited as kirgisicus).

  • — Bordoni, 1975: 80 (Xantholinus; subgenus Helicophallus; characters; notes; Kirghistan).

  • Distribution: Kyrgyzstan.

kirschenblati Bordoni, 1975: 82 (Xantholinus; subgenus Helicophallus; Type locality: Achti, Distr. Novo, Bajazet, Armenia, Eriwan presso il Lago Sevan).

  • Distribution: Armenia.

kocheri Coiffait, 1962c: 74 (Xantholinus; subgenus Xantholinus; Type locality: Maroc: Esmir).

  • — Coiffait, 1962b: 398, 403, 406 (Xantholinus; subgenus Xantholinus; characters; Morocco; Algeria).

  • — Coiffait, 1972: 277 (Xantholinus; characters; references; Morocco; Algeria).

  • — Bordoni, 1972c (Xantholinus; subgenus Paraphallus; correction of subgeneric assignment in Bordoni, 1972: 162, 188).

  • — Coiffait, 1973a: 273 (Xantholinus; Morocco).

  • Distribution: Morocco, Algeria.

kochi Bernhauer, 1940b: 634 (Xantholinus; Type locality: N.W. China: Tienmuschan).

  • Distribution: China.

korgei Coiffait, 1965e: 122 (Xantholinus; subgenus Helicophallus; Type locality: Turquie d'Asie: Kirklar Daglari près de Çamlik au sud de Rize, 2000 m).

  • — Bordoni, 1971: 687 (Xantholinus; subgenus Helicophallus; Turkey).

  • — Coiffait, 1972: 298 (Xantholinus; subgenus Helicophallus; characters; Turkey).

  • — Bordoni, 1972: 164 (Xantholinus; subgenus Helicophallus).

  • — Bordoni, 1976: 232 (Xantholinus; characters; notes; Turkey).

  • — Coiffait, 1978d: 163 (Xantholinus; Turkey).

  • Distribution: Turkey.

kraatzi Kirsch, 1865, see: Agerodes.

krejcii Coiffait, 1984e: 374 (Xantholinus; Type locality: Nepal oriental: Milke Himal, 2300–2600 m).

  • Distribution: Nepal.

labralis Broun, 1880: 104 (Xantholinus; Type locality: Whangarei Harbour).

  • — Duvivier, 1883: 156 (Xantholinus; cited as lateralis; catalog).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Emberson, 1998: 36 (Xantholinus; collecting notes; Chatham Islands).

  • Distribution: New Zealand, Chatham Islands.

lacertosus Sharp, 1885: 478 (Xantholinus; Type locality: Mexico, Oaxaca).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • Distribution: Mexico.

laeticeps Reitter, 1908 [Note: Nomen nudum], see: tricolor Fabricius, 1787.

laevigatus Jacobsen, 1849: 678 (Xantholinus; Type locality: Soro).

  • — Fowler, 1888: 290 (Xantholinus; cited as distans; [Note: Misidentification according to Steel, 1946a: 192]; characters; Britain).

  • — Reitter, 1908: 119 (Xantholinus; synonym of linearis variety longiventris).

  • — Reitter, 1908a: 23 (Xantholinus; [Note: Misidentification: Scheerpeltz, 1933: 1311, cited distans Reitter as a synonym of laevigatus; Reitter did not describe the species as new, he attributed it to Rey]; characters; central Europe).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; synonym of linearis).

  • — Johansen, 1914: 436 (Xantholinus; subgenus Xantholinus; characters; Denmark).

  • — Scheerpeltz, 1933: 1311 (Xantholinus; catalog).

  • — Steel, 1946a: 192 (Xantholinus; characters; England; Ireland; Scotland).

  • — V. Hansen, 1952: 18 (Xantholinus; subgenus Xantholinus; characters; Denmark).

  • — Coiffait, 1965e: 120 (Xantholinus; subgenus Acanthophallus; Turkey).

  • — Pototskaia, 1967: 48 (Xantholinus; larval characters).

  • — Bordoni, 1971: 687 (Xantholinus; subgenus Acanthophallus; Turkey).

  • — Coiffait, 1972: 288 (Xantholinus; characters; references; central Europe; Denmark; Scandinavia; France; Britain; Turkey).

  • — Bordoni, 1972: 161, 185 (Xantholinus; subgenus Acanthophallus; characters; Italy).

  • — Bordoni, 1973: 48 (Xantholinus; subgenus Acanthophallus; Hungary).

  • — Bordoni, 1973c: 155 (Xantholinus; subgenus Acanthophallus; notes; Romania).

  • — Bordoni, 1975: 63 (Xantholinus; subgenus Acanthophallus; notes; Russia).

  • — Bordoni, 1976: 232 (Xantholinus; notes; Turkey).

  • — Bordoni, 1976c: 94 (Xantholinus; France; Italy).

  • — Bordoni, 1976d: 121 (Xantholinus; Alps; Netherlands).

  • — Pope, 1977: 28 (Xantholinus; Britain).

  • — Bordoni, 1982: 271 (Xantholinus; subgenus Acanthophallus; characters; Italy).

  • — Tóth, 1989: 24 (Xantholinus; characters; Hungary).

  • — Bordoni, 1989b: 522 (Xantholinus; subgenus Acanthophus; characters; Italy).

  • — Assing, 1993: 27 (Xantholinus; bionomics; Germany).

  • — Janák, 1993a: 47 (Xantholinus; subgenus Typhlolinus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; Italy).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus).

  • Distribution: Europe, Russia, Caucasus, Turkey.

cribripennis Fauvel, 1873b: 104 [= 1873a: 390] (Xantholinus; Type locality: Aube; Italie; Caucase).

  • — Fauvel, 1873b: 131 [= 1873c: 24] (Xantholinus; notes).

  • — Mulsant and Rey, 1877a: 278 (Xantholinus; characters; France).

  • — Ganglbauer, 1895: 484 (Xantholinus; characters; France).

  • — Reitter, 1908: 120 (Xantholinus; characters; France; Italy).

  • — Bernhauer and Schubert, 1914: 302 (Xantholinus; catalog).

  • — Fiori, 1915: 13 (Xantholinus; Italy).

  • — Sainte-Claire Deville, 1926: 119 (Xantholinus; French specimens are laevigatus; Corsica).

  • — Portevin, 1929: 373 (Xantholinus; characters; France).

  • — Scheerpeltz, 1933: 1309 (Xantholinus; catalog).

  • — Steel, 1946a: 192 (Xantholinus; the French specimens reported as cribripennis are conspecific with laevigatus [see Sainte-Claire Deville, 1926: 119], the Italian ones were renamed as raffrayi Sainte-Claire Deville, and the Caucasus specimens are probably an undescribed species).

  • — Coiffait, 1972: 288 (Xantholinus; synonym of laevigatus).

  • — Bordoni, 1972: 185 (Xantholinus; synonym of laevigatus).

  • — Bordoni, 1975: 63 (Xantholinus; synonym of laevigatus).

  • — Pope, 1977: 29 (Xantholinus; synonym of laevigatus).

  • — Bordoni, 1982: 271 (Xantholinus; synonym of laevigatus).

clairei Coiffait, 1956a: 69 (Xantholinus; subgenus Acanthophallus; Type locality: le Jura Suisse: Nidau).

  • — Coiffait, 1958: 24 (Xantholinus; subgenus Acanthophallus; notes).

  • — Smetana, 1958: 108 (Xantholinus; subgenus Xantholinus; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 51 (Xantholinus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 165 (Xantholinus; subgenus Acanthophallus; characters; central Europe).

  • — Horion, 1965: 99 (Xantholinus; subgenus Acantophallus; Norway; Sweden; Finland; Denmark; Poland; Czechoslovakia; Switzerland; France; Belgium; Holland; Britain; Italy; Slovenia; Croatia; Germany; Austria).

  • — Coiffait, 1972: 288 (Xantholinus; synonym of laevigatus).

  • — Bordoni, 1972: 185 (Xantholinus; synonym of laevigatus).

  • — Bordoni, 1975: 63 (Xantholinus; synonym of laevigatus).

  • — Pope, 1977: 28 (Xantholinus; synonym of laevigatus).

  • — Bordoni, 1982: 271 (Xantholinus; synonym of laevigatus).

  • — Lucht, 1987: 103 (Xantholinus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Lohse and Lucht, 1989: 159 (Xantholinus; synonym of laevigatus).

  • — Nowosad, 1990: 97 (Xantholinus; cited as valid species [Note: Nowosad may have been unaware that clairei had been listed as a junior synonym]; collected from mole nest; notes; Poland).

  • — Janák, 1993a: 47 (Xantholinus; synonym of laevigatus).

laevipennis Sharp, 1885: 477 (Xantholinus; Type locality: Mexico, Tuxtla).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • Distribution: Mexico.

laevissimus Reitter, 1898, see: Megalinus.

laniger Fauvel, 1899c: 97 (Xantholinus; Type locality: Algérie: Bone; Philippeville: Djebel Taya; Tunisie: Teboursouk; camp de la Sante; Ain-Draham; El Fedja).

  • — Fauvel, 1902b: 102 (Xantholinus; Algeria; Tunisia).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Coiffait, 1962b: 397, 400 (Xantholinus; subgenus Calolinus; characters; Algeria; Tunisia).

  • — Coiffait, 1972: 285 (Xantholinus; subgenus Purrolinus; characters; references; Algeria; Tunisia).

  • — Bordoni, 1972: 160 (Xantholinus; subgenus Stenophallus).

  • Distribution: Algeria, Tunisia.

laticeps Erichson, 1839b: 308 (Xantholinus; Type locality: Java).

  • — Kraatz, 1859: 102 (Xantholinus; characters; Java).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • Distribution: Indonesia.

laxus Sharp, 1885: 482 (Xantholinus; Type locality: Guatemala, Aceituno; Panama, Volcan de Chiriqui up to 6000 feet, Peña Blanca 3000 to 4000 feet).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Marcuzzi, 1977: 11 (Xantholinus; Venezuela).

  • Distribution: Guatemala, Panama, Venezuela.

lecontei Duvivier, 1883, see: Neohypnus.

lederi Coiffait, 1966f: 197 (Xantholinus; subgenus Calolinus; Type locality: Kavkaz).

  • — Coiffait, 1972: 264 (Xantholinus; characters; Caucasus).

  • — Bordoni, 1975: 61 (Xantholinus; subgenus Calolinus; notes; Caucasus).

  • Distribution: Caucasus.

lessinicus Bordoni, 1972, see: tricolor Fabricius, 1787.

libanicus Coiffait, 1956a: 72 (Xantholinus; subgenus Helicophallus; Type locality: Liban: Tannourine-Faouka, 1500 m).

  • — Coiffait, 1972: 301 (Xantholinus; subgenus Helicophallus; characters; Lebanon).

  • — Bordoni, 1972: 164 (Xantholinus; subgenus Helicophallus).

  • — Bordoni, 1973a: 181 (Xantholinus; subgenus Helicophallus; characters; Lebanon).

  • — Bordoni, 1976c: 98 (Xantholinus; notes; Lebanon).

  • Distribution: Lebanon.

lichtneckerti Tóth, 1985, see: dvoraki Coiffait, 1956.

liebei Scheerpeltz, 1926, see: Gyrohypnus.

limbatus Waltl, 1835, see: Lepidophallus.

linearis Olivier, 1795:(42): 19 (Staphylinus; Type locality: Paris).

  • — Schrank, 1798: 649 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 305, cited punctulatus Schrank as a synonym of linearis; Schrank did not describe the species as new, he attributed it to Paykull]; characters; notes).

  • — Fabricius, 1798: 180 (Staphylinus; characters; France).

  • — Fabricius, 1801: 599 (Staphylinus; characters; France).

  • — Gravenhorst, 1802: 44 (Staphylinus; synonym of ochraceus Gravenhorst).

  • — Marsham, 1802: 516 (Staphylinus; characters; Britain).

  • — Gyllenhal, 1810: 350 (Staphylinus; synonym of longiceps Gravenhorst).

  • — Curtis, 1829: 27 (Gyrohypnus; catalog; Britain).

  • — Stephens, 1833: 261 (Gyrohypnus; characters; habitat; England).

  • — Erichson, 1839a: 428 (Xantholinus; characters; Germany).

  • — Heer, 1839: 246 (Xantholinus; characters; Switzerland).

  • — Erichson, 1839b: 332 (Xantholinus; characters; Europe).

  • — Stephens, 1839: 403 (Xantholinus; characters; England).

  • — P. Lucas, 1846: 107 (Xantholinus; notes; Algeria).

  • — Jacobsen, 1849: 679 (Xantholinus; characters; Denmark).

  • — Hochhuth, 1849: 109 (Xantholinus; characters; Tiflis).

  • — Redtenbacher, 1849: 693 (Xantholinus; characters; Austria).

  • — Hardy, 1851: 38 (Xantholinus; notes; England).

  • — Wollaston, 1854: 577 (Xantholinus; characters; notes; Madeira Islands).

  • — Scriba, 1855: 299 (Xantholinus; Venezuela).

  • — Fairmaire and Laboulbène, 1856: 502 (Xantholinus; characters; France).

  • — Wollaston, 1857: 188 (Xantholinus; habitat; Madeira Islands).

  • — Kraatz, 1857d: 641 (Xantholinus; characters; Germany).

  • — Redtenbacher, 1857: 182 (Xantholinus; characters; Austria).

  • — Peyron, 1858: 422 (Xantholinus; Turkey).

  • — Hochhuth, 1862: 67 (Xantholinus; Russia).

  • — Fauvel, 1865a: 298 [= 1865b: 48] (Xantholinus; habitat; France).

  • — Wollaston, 1865: 497 (Xantholinus; habitat; Madeira Islands).

  • — Crotch, 1867a: 383 (Xantholinus; Azores).

  • — Fauvel, 1873b: 106 [= 1874a: 392] (Xantholinus; characters; France; Europe; Mediterranean region; Georgia; Caucasus; Iran; Siberia; Madeira).

  • — Redtenbacher, 1874: 220 (Xantholinus; characters; Austria).

  • — Seidlitz, 1875: 281 (Xantholinus; characters; Baltic region).

  • — J. Sahlberg, 1876: 36 (Xantholinus; Finland).

  • — Mulsant and Rey, 1877a: 286 (Xantholinus; adult and larval characters; France).

  • — Fauvel, 1878: 117 [= 1878a: 37] (Xantholinus; Algeria).

  • — Fauvel, 1886: 49 [= 1886a: 41] (Xantholinus; Morocco; Algeria).

  • — Fowler, 1888: 291 (Xantholinus; characters; Britain).

  • — Ganglbauer, 1895: 484 (Xantholinus; characters; Palaearctic region).

  • — Fauvel, 1902b: 102 (Xantholinus; Barbary; Azores; Madeira).

  • — Poppius, 1905: 10 (Xantholinus; Shetland Islands).

  • — Sainte-Claire Deville, 1906: 92 (Xantholinus; Corsica).

  • — Reitter, 1908: 118 (Xantholinus; characters; Palaearctic region).

  • — Poppius, 1909: 23 (Xantholinus; Lena Valley, Siberia).

  • — Reitter, 1909: 138 (Xantholinus; characters; Germany).

  • — Montandon, 1908: 79 (Xantholinus; Romania).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • — Johansen, 1914: 437 (Xantholinus; subgenus Xantholinus; characters; Denmark).

  • — Portevin, 1929: 373 (Xantholinus; characters; France).

  • — Donisthorpe, 1930: 97 (Xantholinus; characters; Britain).

  • — Scheerpeltz, 1933: 1311 (Xantholinus; catalog).

  • — Bernhauer, 1940c: 4 (Xantholinus; Azores).

  • — C. Koch, 1941: 61 (Xantholinus; Morocco).

  • — Méquignon, 1942a: 19 (Xantholinus; Azores).

  • — Steel, 1946a: 194 (Xantholinus; characters; Britain).

  • — Gridelli, 1947: 65, 75 (Xantholinus; abdominal morphology; characters; summary of distribution).

  • — Scheerpeltz and Höfler, 1948: 156 (Xantholinus; fungus inhabitant).

  • — V. Hansen, 1952: 19 (Xantholinus; subgenus Xantholinus; characters; Denmark).

  • — Fagel, 1955a: 84 (Xantholinus; Greece).

  • — Coiffait, 1956a: 65 (Xantholinus; characters in key; western and central Europe; England; Spain; Italy).

  • — Hatch, 1957: 234 (Megalinus; characters; British Columbia; Washington; Oregon; Idaho).

  • — Smetana, 1958: 113 (Xantholinus; subgenus Xantholinus; characters; ecology; Czechoslovakia).

  • — Lohse, 1960a: 38 (Xantholinus; characters; notes).

  • — Scheerpeltz, 1961d: 127 (Xantholinus; subgenus Xantholinus; Italy).

  • — Coiffait, 1962b: 397, 401 (Xantholinus; subgenus Xantholinus; characters; Algeria; Tunisia).

  • — Brakman, 1962: 48 (Xantholinus; characters; Holland).

  • — Palm, 1963b: 51 (Xantholinus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 165 (Xantholinus; subgenus Xantholinus; characters; central Europe).

  • — Smetana, 1964d: 72 (Xantholinus; subgenus Xantholinus; Czechoslovakia).

  • — Raianu, 1964: 104 (Xantholinus; Romania).

  • — Scheerpeltz, 1965b: 400 (Xantholinus; Sicily).

  • — Scheerpeltz, 1965c: 504 (Xantholinus; Sicily).

  • — Horion, 1965: 100 (Xantholinus; subgenus Xantholinus; most all Europe).

  • — Szujecki, 1968a: 726 (Xantholinus; Poland).

  • — Kasule, 1970: 79 (Xantholinus; larval characters).

  • — Coiffait, 1972: 266 (Xantholinus; subgenus Xantholinus; characters; references; Holarctic region).

  • — Bordoni, 1972: 162, 197 (Xantholinus; subgenus Paraphallus; characters; Italy).

  • — Bordoni, 1972c (Xantholinus; subgenus Xantholinus; correction of subgeneric assignment in Bordoni, 1972: 162, 197).

  • — Bordoni, 1973c: 151 (Xantholinus; notes; Romania).

  • — Osella and Zanetti, 1975: 107 (Xantholinus; collected from nest of Talpa europaea; notes; Italy).

  • — Bordoni, 1975: 66 (Xantholinus; notes).

  • — Bordoni, 1976c: 88 (Xantholinus; France; Italy; Spain; Morocco; Algeria).

  • — Bordoni, 1976d: 120 (Xantholinus; Algeria; Germany; Netherlands).

  • — Pope, 1977: 29 (Xantholinus; Britain).

  • — Topp, 1979: 15 (Xantholinus; development; diapause).

  • — Smetana, 1979: 51 (requested that International Commission on Zoological Nomenclature place linearis on the Official List of Specific Names in Zoology).

  • — Outerelo, 1980a: 56 (Xantholinus; Spain).

  • — Hinton, 1981: 663 (Xantholinus; characters of egg).

  • — Smetana, 1982: 266 (Xantholinus; characters; British Columbia; California; Idaho; Massachusetts; Nevada; New York; Oregon; Pennsylvania; Utah; Washington).

  • — Bordoni, 1982: 300 (Xantholinus; subgenus Xantholinus; characters; Italy).

  • — Frank, 1982: 32 (Xantholinus; list of parasites).

  • — ICZN, 1983: 85 (Xantholinus; placed on Official List of Specific Names in Zoology and fixed as the type species of Xantholinus).

  • — Lucht, 1987: 103 (Xantholinus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Smetana, 1988a: 557 (Xantholinus; Nova Scotia; California; Idaho; New Hampshire).

  • — Tóth, 1989: 27 (Xantholinus; characters; Hungary).

  • — Drugmand, 1990a: 94 (Xantholinus; characters; Belgium).

  • — Borges, 1990: Table II (Xantholinus; Azores).

  • — Smetana, 1990c: 87 (Xantholinus; New Hampshire).

  • — Nowosad, 1990: 93 (Xantholinus; collected from mole nest; notes; Poland).

  • — Campbell and Davies, 1991: 124 (Xantholinus; checklist of species; Canada).

  • — Janák and Vysoky, 1992: 136 (Xantholinus; collected with Formica rufa; Bohemia).

  • — Ciceroni, 1993: 74 (Xantholinus; Italy).

  • — Assing, 1993: 24 (Xantholinus; bionomics; Germany).

  • — Janák, 1993a: 47 (Xantholinus; subgenus Xantholinus; Czech Republic; Slovakia).

  • — Welch, 1993: 228 (Xantholinus; ovariole number and ovary structure).

  • — Sparacio, 1995: 158 (Xantholinus; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; Italy).

  • — Downie and Arnett, 1996: 402 (Xantholinus; characters; USA).

  • — Mikat, Fremuth, and Prouza, 1997: 111 (Xantholinus; Czech Republic).

  • — Owen, 1997a: 149 (Xantholinus; collecting sites; Ireland).

  • — Owen, 1999a: 15 (Xantholinus; subterranean capture; Britain).

  • — Owen, 1999b: 61 (Xantholinus; captured underground at roots of old trees; Britain).

  • — Kashcheev, 1999a: 57 (Xantholinus; Kazakhstan).

  • — Owen, 2000: 251 (Xantholinus; pitfall trap collections from underground at roots of trees; Britain).

  • — Assing, 2001: 75 (Xantholinus; collected from xerothermous site; Germany).

  • Distribution: Tunisia, Algeria, Morocco, Madeira, Azores, Europe, Russia, Georgia, Turkey, Iran, Kazakhstan, Canada, USA, Venezuela.

ochraceus Gravenhorst, 1802: 43 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).

  • — Latreille, 1804: 329 (Staphylinus; characters; France).

  • — Gravenhorst, 1806: 97 (Staphylinus; characters).

  • — Panzer, 1808: 15 (Staphylinus; characters; illustrations).

  • — Gyllenhal, 1810: 350 (Staphylinus; synonym of longiceps Gravenhorst).

  • — Gyllenhal, 1827: 480 (Staphylinus; synonym of batychrus).

  • — Curtis, 1829: 27 (Gyrohypnus; catalog; Britain).

  • — C. Sahlberg, 1830: 332 (Staphylinus; synonym of batychrus Gyllenhal).

  • — Griffith and Pidgeon, 1832: 292 (Xantholinus).

  • — Stephens, 1833: 263 (Gyrohypnus; characters; England).

  • — Lacordaire, 1835: 416 (Xantholinus; characters; France).

  • — Erichson, 1839a: 428 (Xantholinus; synonym of linearis).

  • — Erichson, 1839b: 332 (Xantholinus; synonym of linearis).

  • — Laporte, 1840: 180 (Gyrohypnus; characters; France; Germany).

  • — P. Lucas, 1846: 107 (Xantholinus; synonym of linearis).

  • — Mulsant and Rey, 1877a: 286 (Xantholinus; variety of linearis).

  • — Ganglbauer, 1895: 485 (Xantholinus; synonym of linearis).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; synonym of linearis).

  • — Smetana, 1958: 114 (Xantholinus; synonym of linearis).

  • — Coiffait, 1972: 266 (Xantholinus; synonym of linearis).

  • — Bordoni, 1972: 197 (Xantholinus; synonym of linearis).

  • — Smetana, 1982: 266 (Xantholinus; synonym of linearis).

  • — Bordoni, 1982: 300 (Xantholinus; synonym of linearis).

longiceps Gravenhorst, 1802: 177 (Staphylinus; Type locality: Lusitania).

  • — Latreille, 1804: 334 (Staphylinus; characters; Portugal).

  • — Gyllenhal, 1810: 350 (Staphylinus; characters; Sweden).

  • — Gyllenhal, 1827: 480 (Staphylinus; characters).

  • — Zetterstedt, 1828: 80 (Xantholinus; characters; Lapland).

  • — Curtis, 1829: 27 (Gyrohypnus; catalog; Britain).

  • — C. Sahlberg, 1830: 331 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 33 (Gyrohypnus; Finland; Sweden).

  • — Mannerheim, 1831: 447 (Gyrohypnus; Finland; Sweden).

  • — Stephens, 1833: 262 (Gyrohypnus; characters; England).

  • — Runde, 1835: 11 (Gyrohypnus; characters; Germany).

  • — Nordmann, 1837: 114 (Gyrohypnus; characters for varieties; Europe).

  • — Zetterstedt, 1838: 66 (Xantholinus; characters; habitat; Lapland).

  • — Erichson, 1839a: 428 (Xantholinus; synonym of linearis).

  • — Heer, 1839: 246 (Xantholinus; synonym of linearis).

  • — Erichson, 1839b: 332 (Xantholinus; synonym of linearis).

  • — Stephens, 1839: 403 (Xantholinus; variety of linearis; characters; England).

  • — Redtenbacher, 1849: 693 (Xantholinus; synonym of linearis).

  • — Hardy, 1851: 38 (Xantholinus; synonym of linearis).

  • — Fairmaire and Laboulbène, 1856: 502 (Xantholinus; synonym of linearis).

  • — Kraatz, 1857d: 642 (Xantholinus; synonym of linearis).

  • — Redtenbacher, 1857: 182 (Xantholinus; synonym of linearis).

  • — J. Sahlberg, 1876: 36 (Xantholinus; synonym of linearis).

  • — Ganglbauer, 1895: 485 (Xantholinus; synonym of linearis).

  • — Reitter, 1908: 118 (Xantholinus; synonym of linearis).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; synonym of linearis).

  • — Coiffait, 1972: 266 (Xantholinus; synonym of linearis).

  • — Smetana, 1982: 266 (Xantholinus; synonym of linearis).

  • — Bordoni, 1982: 300 (Xantholinus; synonym of linearis).

punctulatus Gravenhorst, 1802: 177 (Staphylinus; [preoccupied]; Type locality: Lusitania).

  • — Gravenhorst, 1806: 98 (Staphylinus).

  • — Gyllenhal, 1810: 350 (Staphylinus; synonym of longiceps).

  • — C. Sahlberg, 1830: 331 (Staphylinus; synonym of longiceps).

  • — Erichson, 1839a: 428 (Xantholinus; synonym of linearis).

  • — Erichson, 1839b: 332 (Xantholinus; characters; synonym of linearis; [Note: Erichson cited the Gravenhorst species as a subsequent reference to the punctulatus mentioned by Schrank]).

  • — Mulsant and Rey, 1853: 58 (Xantholinus; characters; notes; with Formica fuliginosa).

  • — Kraatz, 1857d: 642 (Xantholinus; [Note: Kraatz cited Schrank as the author of punctulatus]; synonym of linearis).

  • — Hochhuth, 1862: 65 (Xantholinus; Russia).

  • — Ganglbauer, 1895: 485 (Xantholinus; synonym of linearis).

multipunctatus Thomson, 1860: 191 (Xantholinus; Type locality: Gottland).

  • — Fauvel, 1873b: 106 [= 1874a: 392] (Xantholinus; synonym of linearis).

  • — J. Sahlberg, 1876: 36 (Xantholinus; synonym of linearis).

  • — Mulsant and Rey, 1877a: 286 (Xantholinus; synonym of linearis).

  • — Reitter, 1908: 118 (Xantholinus; synonym of linearis).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; synonym of linearis).

  • — Smetana, 1958: 114 (Xantholinus; synonym of linearis).

  • — Coiffait, 1972: 266 (Xantholinus; synonym of linearis).

  • — Bordoni, 1972: 197 (Xantholinus; synonym of linearis).

  • — Smetana, 1982: 266 (Xantholinus; synonym of linearis).

  • — Bordoni, 1982: 300 (Xantholinus; synonym of linearis).

aequalis Fauvel, 1898: 98 (Xantholinus; Type locality: Algérie).

  • — Fauvel, 1902b: 102 (Xantholinus; Algeria; Tunisia).

  • — Bernhauer and Schubert, 1914: 300 (Xantholinus; catalog).

  • — Coiffait, 1962b: 397, 401 (Xantholinus; subgenus Xantholinus; variety of linearis; characters; Morocco; Algeria).

  • — Coiffait, 1972: 266, 267 (Xantholinus; subspecies of linearis; characters).

  • — Smetana, 1982: 266 (Xantholinus; synonym of linearis).

anglicus† Lesne, 1926: 8 (Xantholinus; subspecies of linearis; Pliocene subfossil; Type locality: Castle Eden, Angleterre septentrionale).

  • — Scheerpeltz, 1933: 1311 (Xantholinus; subspecies of linearis).

  • — Coiffait, 1972: 266, 267 (Xantholinus; variety of linearis; fossil; characters).

rossicus Bordoni, 1975: 66 (Xantholinus; subgenus Xantholinus; subspecies of linearis; Type locality: Amur, Simonovsk, 75 km ad Ovest di Svobodryi).

linearis Blatchley, 1910, see: Neohypnus.

lineatus Wendeler, 1924, see: Neohypnus.

lividipennis Coiffait, 1972: 299 (Xantholinus; subgenus Helicophallus; Type locality: Bey Dag, Anatolie méridionale).

  • Distribution: Turkey.

loebli Bordoni, 1976, see: longiventris Heer, 1839.

longiceps Gravenhorst, 1802, see: linearis Olivier, 1795.

longipennis Cameron, 1950g: 32 (Xantholinus; Type locality: Belgian Congo: Vitshumbi, S. lac Edouard, alt. 925 m).

  • Distribution: Congo.

longiventris Heer, 1839: 247 (Xantholinus; Type locality: Switzerland; [Note: See lectotype designation by Assing, 1995a]. Originally cited from Matt; Lägeren).

  • — Hochhuth, 1849: 109 (Xantholinus; characters; Caucasus).

  • — Kraatz, 1857d: 641 (Xantholinus; characters; Germany).

  • — Redtenbacher, 1857: 983 (Xantholinus; characters; Austria).

  • — Thomson, 1860: 191 (Xantholinus; characters; Scandinavia).

  • — Hochhuth, 1862: 67 (Xantholinus; Russia).

  • — Baudi, 1870: 388 (Xantholinus; characters; Italy).

  • — Fauvel, 1873b: 106 [= 1874a: 392] (Xantholinus; synonym of linearis).

  • — Redtenbacher, 1874: 220 (Xantholinus; characters; Austria).

  • — Mulsant and Rey, 1877a: 283 (Xantholinus; characters; France).

  • — Fowler, 1888: 291 (Xantholinus; characters; Britain).

  • — Ganglbauer, 1895: 485 (Xantholinus; synonym of linearis).

  • — Reitter, 1908: 119 (Xantholinus; characters; Europe).

  • — Reitter, 1909: 138 (Xantholinus; variety of linearis; characters; Germany).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; synonym of linearis).

  • — Donisthorpe, 1930: 97 (Xantholinus; characters; Britain).

  • — Scheerpeltz, 1933: 1312 (Xantholinus; catalog).

  • — C. Koch, 1936a: 75 (Xantholinus; variety of linearis; Libya).

  • — Tottenham, 1949: 416 (Xantholinus; checklist; Britain).

  • — V. Hansen, 1952: 20 (Xantholinus; subgenus Xantholinus; characters; Denmark).

  • — Coiffait, 1956a: 63 (Xantholinus; characters in key; Belgium; Czechoslovakia; France; Spain; Pyrenees).

  • — Coiffait, 1958: 24 (Xantholinus; subgenus Xantholinus).

  • — Lohse, 1960a: 38, 39 (Xantholinus; characters; notes).

  • — Brakman, 1962: 48 (Xantholinus; characters).

  • — Smetana, 1963b: 36 (Xantholinus; subgenus Xantholinus; Madeira).

  • — Palm, 1963b: 52 (Xantholinus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 166 (Xantholinus; subgenus Xantholinus; characters; central Europe).

  • — Horion, 1965: 100 (Xantholinus; subgenus Xantholinus; Madeira; Finland; Sweden; Norway; Britain; Holland; Belgium; France; Spain; Switzerland; Czechoslovakia; Poland; Romania; Dalmatia; Germany; Austria).

  • — Smetana, 1966: 330 (Xantholinus; characters).

  • — Pototskaia, 1967: 48 (Xantholinus; larval characters).

  • — Scheerpeltz, 1968a: 61 (Xantholinus; catalog; Austria).

  • — Smetana, 1970: 58 (Xantholinus; distributional summary; Azores).

  • — Coiffait, 1972: 268 (Xantholinus; subgenus Xantholinus; characters; references; Palaearctic region).

  • — Bordoni, 1972: 188 (Xantholinus; subgenus Xantholinus; characters; Italy).

  • — Bordoni, 1972c (Xantholinus; subgenus Paraphallus; correction of subgeneric assignment in Bordoni, 1972: 162, 188).

  • — Bordoni, 1973: 46 (Xantholinus; subgenus Xantholinus; Hungary).

  • — Bordoni, 1973c: 152 (Xantholinus; subgenus Xantholinus; notes; Romania).

  • — Bordoni, 1975: 64 (Xantholinus; subgenus Paraphallus; notes; Russia).

  • — Osella and Zanetti, 1975: 107 (Xantholinus; collected from nest of Talpa europaea; notes; Italy).

  • — Bordoni, 1976c: 88 (Xantholinus; Libya; Spain).

  • — Bordoni, 1976d: 121 (Xantholinus; Netherlands).

  • — Pope, 1977: 29 (Xantholinus; Britain).

  • — Muona, 1979: 16 (Xantholinus; Finland; Russia; Sweden; Norway; Denmark; Baltic States).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 20 (Xantholinus; catalog; Poland).

  • — Smetana, 1982: 271 (Xantholinus; characters; California; Oregon; Washington).

  • — Bordoni, 1982: 285 (Xantholinus; subgenus Paraphallus; characters; Italy).

  • — Ádám, 1987: 148 (Phallolinus; collecting notes; Hungary).

  • — Lucht, 1987: 103 (Xantholinus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Smetana, 1988a: 557 (Xantholinus; California).

  • — Tóth, 1989: 28 (Xantholinus; characters; Hungary).

  • — Drugmand, 1990a: 92 (Xantholinus; characters; Belgium).

  • — Borges, 1990: Table II (Xantholinus; Azores).

  • — Silfverberg, 1992: 18 (Xantholinus; Finland; Russia; Sweden; Norway; Denmark; Estonia; Lithuania).

  • — Assing, 1993: 27 (Xantholinus; bionomics; Germany).

  • — Ciceroni, 1993: 74 (Xantholinus; Italy).

  • — Welch, 1993: 228 (Xantholinus; ovariole number and ovary structure).

  • — Janák, 1993a: 47 (Xantholinus; subgenus Xantholinus; Czech Republic; Slovakia).

  • — Assing, 1995a: 967 (Xantholinus; lectotype designation: Switzerland).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; Italy).

  • — Franzen, 1996: 206 (Xantholinus; collecting notes; Germany).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 243 (Xantholinus; Denmark).

  • — M. Hansen, 1996: 109 (Xantholinus; checklist; Denmark).

  • — Smoleński, 1997: 80 (Xantholinus; collecting notes; Poland).

  • — Eisinger, 1997: 170 (Xantholinus; collecting notes; Germany).

  • — Wenzel, 1997: 346 (Xantholinus; collecting notes; Germany).

  • — Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 40 (Xantholinus; Latvia).

  • — Anderson, Nash, and O'Connor, 1997: 20 (Xantholinus; Ireland).

  • — Owen, 1997a: 149 (Xantholinus; collecting sites; Ireland).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 74 (Xantholinus; Denmark).

  • — Assing, 2001: 75 (Xantholinus; collected from xerothermous site; Germany).

  • Distribution: Libya, Madeira, Azores, Europe, Russia, USA.

elongatus Heer, 1834: 75 (Xantholinus; [Note: Nomen nudum; published without characters]).

  • — Kraatz, 1857d: 641 (Xantholinus; synonym of longiventris).

  • — Mulsant and Rey, 1877a: 283 (Xantholinus; synonym of longiventris).

  • — Reitter, 1908: 119 (Xantholinus; variety of linearis variety longiventris).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; synonym of linearis).

  • — Bordoni, 1972: 197 (Xantholinus; synonym of linearis).

  • — Smetana, 1982: 271 (Xantholinus; synonym of longiventris).

  • — Bordoni, 1982: 285 (Xantholinus; synonym of longiventris).

miguelensis Méquignon, 1942: 9 (Xantholinus; Type locality: Açores: Sâo Miguel: Ponte Delgada).

  • — Méquignon, 1942a: 19 (Xantholinus; characters; Azores).

  • — Coiffait, 1972: 268 (Xantholinus; subspecies of longiventris; characters; Azores).

  • — Smetana, 1982: 271 (Xantholinus; synonym of longiventris).

  • — Borges, 1990: Table II (Xantholinus; subspecies of longiventris; Azores).

oregonus Fender, 1946: 66 (Xestolinus; Type locality: Oregon: Forest Grove).

  • — Hatch, 1957: 234 (Megalinus; synonym of linearis).

  • — Smetana, 1982: 271 (Xantholinus; [Note: Smetana, 1982: 266, 271, cited oregonus as though it were a synonym of linearis (p. 266) and as a synonym of longiventris (p. 271); the holotype of oregonus is conspecific with longiventris; the allotype was a misidentifation and is conspecific with linearis]; synonym of longiventris).

  • — Bordoni, 1999: 6 (Xantholinus; subgenus Xantholinus).

sejugatus G. Benick, 1953: 39 (Xantholinus; Type locality: Deutschland: Schellbruch bei Lübeck).

  • — Horion, 1956: 3 (Xantholinus; characters; notes).

  • — Lohse, 1960a: 38, 39 (Xantholinus; characters; notes).

  • — Lohse, 1964: 166 (Xantholinus; subgenus Xantholinus; characters; central Europe).

  • — Coiffait, 1972: 308 (Xantholinus; characters; references; Germany).

  • — Assing, 1995a: 965, 967 (Xantholinus; characters; notes; synonym of longiventris).

  • — Assing, Frisch, Kahlen, et al., 1998: 136 (Xantholinus; synonym of longiventris).

loebli Bordoni, 1976c: 88 (Xantholinus; subgenus Paraphallus; Type locality: Legnago, Veneto).

  • — Bordoni, 1982: 287 (Xantholinus; subgenus Paraphallus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; probable synonym of longiventris; Italy).

  • — Bordoni, 1999: 6 (Xantholinus; synonym of longiventris).

lorquini Fauvel, 1877, see: Thyreocephalus.

lubricus Erichson, 1839b: 321 (Xantholinus; Type locality: San Joâo des Rey Brasiliae).

  • — Lynch, 1884: 183 (Xantholinus; subgenus Megalinus; characters; Argentina).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1312 (Xantholinus; catalog).

  • Distribution: Brazil, Argentina.

lucidus Erichson, 1839, see: Agerodes.

luteipennis Coiffait, 1970: 293 (Xantholinus; subgenus Helicophallus; Type locality: environs de Pinarbasi, 1700 m, Ouest de Kayseri, Anatolie centrale).

  • — Bordoni, 1971: 687 (Xantholinus; subgenus Helicophallus; Turkey).

  • — Coiffait, 1972: 300 (Xantholinus; subgenus Helicophallus; characters; Anatolia).

  • — Bordoni, 1972: 164 (Xantholinus; subgenus Helicophallus).

  • Distribution: Turkey.

lynceus Erichson, 1839, see: Thyreocephalus.

lynchi Bernhauer, 1912e: 174 (Xantholinus; Type locality: Uruguay).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • Distribution: Uruguay.

madagascarensis Steel, 1949, see: Eulissus.

maghrebinus Quedenfeldt, 1882: 182 (Xantholinus; variety of linearis; Type locality: Marocco: Jebel Hadid prope urbem Mogador).

  • — Quedenfeldt, 1884: 376 (Xantholinus; variety of linearis; characters; Morocco).

  • — Fauvel, 1902b: 102 (Xantholinus; variety of linearis).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; synonym of linearis).

  • — Coiffait, 1962b: 397, 400 (Xantholinus; subgenus Calolinus; valid species; characters; Morocco; Algeria; Tunisia).

  • — Coiffait, 1972: 265 (Xantholinus; subgenus Calolinus; characters; references; North Africa).

  • — Bordoni, 1972: 162 (Xantholinus; subgenus Calolinus).

  • — Bordoni, 1976c: 88 (Xantholinus; Morocco).

  • Distribution: Tunisia, Algeria, Morocco.

magrinii Bordoni, 1987, see: cerrutii Coiffait, 1964.

magyaricus Tóth, 1985, see: schweigeri Coiffait, 1966.

malaisei Scheerpeltz, 1965: 192 (Xantholinus; Type locality: N.E. Burma, Kambaiti, 7000 ft.).

  • Distribution: Myanmar.

mandschuricus Bernhauer, 1923a: 124 (Xantholinus; Type locality: Süd-Mandschurei: Chikuanshan; [Note: See lectotype designation by Bordoni, 1999]).

  • — Scheerpeltz, 1933: 1312 (Xantholinus; catalog).

  • — Bordoni, 1975: 85 (Xantholinus; characters; notes; Manchuria).

  • — Bordoni, 1999: 5 (Xantholinus; lectotype designation).

  • Distribution: China.

marginalis Wollaston, 1862, see: Gyrohypnus.

maritimus Reitter, 1908: 122 [= 1908a: 25] (Xantholinus; subgenus Typhlolinus; Type locality: Alpes maritimes; [Note: See lectotype designation by Bordoni, 1999]).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1312 (Xantholinus; catalog).

  • — Coiffait, 1956a: 72 (Xantholinus; subgenus Helicophallus; characters in key).

  • — Coiffait, 1972: 303 (Xantholinus; subgenus Helicophallus; characters; references; Maritime Alps; Apennines).

  • — Bordoni, 1972: 164, 216 (Xantholinus; subgenus Helicophallus; characters; Italy).

  • — Bordoni, 1976c: 97 (Xantholinus; Italy).

  • — Bordoni, 1982: 323 (Xantholinus; subgenus Helicophallus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; Italy).

  • — Bordoni, 1999: 2, 3 (Xantholinus; subgenus Helicophallus; lectotype designation).

  • Distribution: France, Italy.

martensi Bordoni, 1983: 61 (Xantholinus; subgenus Calolinus; Type locality: Iran: prov. Masandaran, Mt. Elburz, 11 km ö. Alasht westliches Seitental des Talar, 1400 m).

  • Distribution: Iran.

massinissae Bordoni, 1973, see: morandi Coiffait, 1958.

maurus Bordoni, 1972a: 128 (Xantholinus; subgenus Toxophallus; Type locality: Marocco: Jebel Tazeka, Taza).

  • — Bordoni, 1972: 160 (Xantholinus; subgenus Toxophallus; [Note: Bordoni cited the name as new but published no characters so the name is unavailable, Article 13.1.1]).

  • — Bordoni, 1976c: 86 (Xantholinus; notes; Morocco).

  • Distribution: Morocco.

maykopensis Coiffait, 1966f: 201 (Xantholinus; subgenus Helicophallus; Type locality: Severnyi Kavkas, okrestnosti Maikopa).

  • — Pototskaia, 1967: 47 (Xantholinus; cited as maycopensis; larval characters).

  • — Coiffait, 1972: 305 (Xantholinus; subgenus Helicophallus; characters; Caucasus).

  • — Bordoni, 1975: 80 (Xantholinus; subgenus Helicophallus; characters; notes; Circassia).

  • Distribution: Russia (Caucasus).

mediocris Broun, 1880, see: Pseudocorynus.

melanarius Fauvel, 1872, see: Gyrohypnus.

mendozanus Bernhauer, 1912e: 174 (Xantholinus; Type locality: Argentinien: Mendoza).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • Distribution: Argentina.

meridionalis Lacordaire, 1835, see: elegans Olivier, 1795.

meridionalis Nordmann, 1837, see: ibericus Steel, 1950.

metallicus Fauvel, 1895, see: Lepidophallus.

mexicanus Sharp, 1885: 479 (Xantholinus; Type locality: Mexico, Jalapa).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • Distribution: Mexico.

meyeri Drugmand, 1994, see: tricolor Fabricius, 1787.

microphthalmus Bernhauer, 1939a: 79 (Xantholinus; Type locality: Kenya: Mt. Aberdare, vers. Ouest, 2600 m, mais. forest. Kinangop).

  • Distribution: Kenya.

microps Fauvel, 1907: 35 (Xantholinus; Type locality: Kiobscho, 3000 m).

  • — Bernhauer and Schubert, 1914: 305 (Xantholinus; catalog).

  • Distribution: Kenya.

micropterus Bernhauer, 1939a: 79 (Xantholinus; Type locality: Kenya: Mt. Aberdare, vers. Ouest, 2600 m, mais. forest. Kinangop).

  • Distribution: Kenya.

microtophilus Coiffait, 1970: 287 (Xantholinus; Type locality: Saidkly, 2300 m, environs de Sisian, S.S.R. d'Arménie).

  • — Coiffait, 1972: 269 (Xantholinus; subgenus Xantholinus; characters; Armenia).

  • — Bordoni, 1972c (Xantholinus; subgenus Paraphallus; correction of subgeneric assignment in Bordoni, 1972: 162, 188).

  • — Korge, 1973: 9 (Xantholinus; Armenia; Iran).

  • — Bordoni, 1975: 65 (Xantholinus; subgenus Paraphallus; notes; Armenia).

  • Distribution: Armenia, Iran.

miguelensis Méquignon, 1942, see: longiventris Heer, 1839.

mihoki Bernhauer, 1914, see: Vulda.

minimus Fiori, 1915, see: Vulda.

minutus Lacordaire, 1835, see: Leptacinus.

minutus Coiffait, 1962c: 73 (Xantholinus; subgenus Leptophallus; [preoccupied]; Type locality: Algérie: Nemours).

  • — Coiffait, 1962b: 395, 399, 404 (Xantholinus; subgenus Lepidophallus; characters; Algeria).

  • — Coiffait, 1972: 258 (Xantholinus; subgenus Leptophallus; characters; references; Algeria).

  • — Bordoni, 1972: 160 (Xantholinus; subgenus Leptophallus).

  • — Bordoni, 1976c: 84 (Xantholinus; Algeria).

  • — Bordoni, 1976d: 120 (Xantholinus; Algeria).

  • — Bordoni, 1985a: 84 (Xantholinus; subgenus Neoleptophallus).

  • — Herman, 2001: 53 (Xantholinus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: Algeria.

mitomorphoides Coiffait, 1984e: 376 (Xantholinus; Type locality: Nepal oriental: vallée de l'Arun).

  • Distribution: Nepal.

mixtus Sharp, 1874, see: Lepidophallus.

mocquerysi Fauvel, 1903, see: Thyreocephalus.

modestus Cameron, 1926b: 345 (Xantholinus; Type locality: Chakrata District; Khedar Khud, 7500 feet).

  • — Cameron, 1932: 28 (Xantholinus; characters; India).

  • — Scheerpeltz, 1933: 1312 (Xantholinus; catalog).

  • Distribution: India.

morandi Coiffait, 1958: 20 (Xantholinus; Type locality: Espagne: Province d'Alicante: Castaglia).

  • — Coiffait, 1972: 273 (Xantholinus; subgenus Xantholinus; characters; Iberia).

  • — Bordoni, 1972c (Xantholinus; subgenus Paraphallus; correction of subgeneric assignment in Bordoni, 1972: 162, 188).

  • — Bordoni, 1976c: 91 (Xantholinus; Spain).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; Italy).

  • Distribution: Algeria, Morocco, Spain, Portugal.

algarvensis Coiffait, 1972: 276 (Xantholinus; subgenus Xantholinus; Type locality: Porto de Lagos, Algarve, Portugal).

  • — Ciceroni, 1994: 117 (Xantholinus; synonym of morandi).

massinissae Bordoni, 1973f: 74 (Xantholinus; Type locality: Algeria: Laverdure).

  • — Bordoni, 1972c (Xantholinus; subgenus Paraphallus; correction of subgeneric assignment in Bordoni, 1972: 162, 188).

  • — Bordoni, 1976c: 89 (Xantholinus; cited as massinissai; characters; notes; Algeria).

  • — Ciceroni, 1994: 117 (Xantholinus; synonym of morandi).

poneli Coiffait, 1985a: 276 (Xantholinus; subgenus Xantholinus; Type locality: Maroc: Ben Slimane).

  • — Ciceroni, 1994: 117 (Xantholinus; synonym of morandi).

morio Motschulsky, 1858, see: Thyreocephalus.

morio Wollaston, 1871, see: Notolinus.

morio Reitter, 1872, see: Gyrohypnus.

morrisoni Blackwelder, 1943: 484 (Xantholinus; Type locality: Trinidad: River Estate, 8 miles northwest of Port of Spain).

  • Distribution: West Indies.

motschulskyi Bordoni, 1999: 4 (Xantholinus; subgenus Helicophallus; named for misidentification of fortepunctatus by Coiffait, 1972, and Korge, 1973; [Note: Since no characters were presented by Bordoni motschulskyi is unavailable unless we apply the characters published by Coiffait, 1972, and Korge, 1973, to the new name]; Type locality: Iran: Gilan).

  • — Coiffait, 1972: 296 (Xantholinus; subgenus Helicophallus; [Note: Misidentified as fortepunctatus according to Bordoni, 1999: 4]; characters; references; Caucasus; Armenia).

  • — Korge, 1973: 9 (Xantholinus; subgenus Helicophallus; [Note: Misidentified as fortepunctatus according to Bordoni, 1999: 4]; characters; notes; Iran).

  • Distribution: Iran, Armenia.

multipunctatus Thomson, 1860, see: linearis Olivier, 1795.

myops Fauvel, 1873, see: Vulda.

nairobiensis Fauvel, 1907: 34 (Xantholinus; Type locality: Nairobi [Wa Kikuyu et Masai]).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • Distribution: Kenya.

nanus LeConte, 1880, see: Linohesperus.

nasutus Harold, 1879, see: Gauropterus.

nicolasi Coiffait, 1972: 263 (Xantholinus; subgenus Calolinus; Type locality: Emin Agha, Épire, Grèce).

  • Distribution: Greece.

niger Cameron, 1926b: 343 (Xantholinus; Type locality: Kheri Rau, Siwaliks).

  • — Cameron, 1932: 25 (Xantholinus; characters; India).

  • — Scheerpeltz, 1933: 1313 (Xantholinus; catalog).

  • Distribution: India.

nigerrimus Kraatz, 1859, see: Thyreocephalus.

nigriceps Guérin-Méneville, 1844: 12 (Xantholinus; Type locality: Nouvelle Granade).

  • — Sharp, 1885: 476 (Xantholinus; Mexico; Guatemala, Panama, Colombia).

  • — Fauvel, 1901: 82 (Xantholinus; synonym of castigatus).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • Distribution: Colombia, Panama, Guatemala, Mexico.

nigropolitus Cameron, 1928g: 443 (Xantholinus; Type locality: Borneo: Songei Tutau).

  • — Scheerpeltz, 1933: 1313 (Xantholinus; catalog).

  • Distribution: Borneo.

nitidicollis Reitter, 1908, see: Gyrohypnus.

nossibeanus Bernhauer and Schubert, 1914: 306 (Xantholinus; replacement name for coloratus Brancsik).

  • Distribution: Madagascar.

coloratus Brancsik, 1893: 220 (Xantholinus; [preoccupied]; Type locality: Nossibé).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; synonym of nossibeanus).

nudobiiformis Tikhomirova, 1973, see: Megalinus.

oakleyi Blackwelder, 1943: 476 (Xantholinus; Type locality: Puerto Rico: 13 miles east of Maricao. Also cited from Dominican Republic).

  • Distribution: West Indies.

oasis Steel, 1948, see: Megalinus.

obscurus Erichson, 1839, see: Neohypnus.

obsidianus Melsheimer, 1844, see: Gyrohypnus.

obsoletus Schubert, 1911: 16 (Xantholinus; Type locality: Antananarivo, Madagascar).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • Distribution: Madagascar.

occidentalis Waltl, 1835, see: Megalinus.

occultans Gistel, 1857: 25 (Xantholinus; [Note: Nomen dubium]; Type locality: Styriae).

  • Distribution: Austria.

ochraceus Gravenhorst, 1802, see: linearis Olivier, 1795.

ochripennis Eppelsheim, 1892, see: Gyrohypnus.

oculatus Sharp, 1885, see: Eulissus.

olliffi Lea, 1895, see: Eulissus.

ophthalmicus Gridelli, 1947, see: Vulda.

oregonus Fender, 1946, see: longiventris Heer, 1839.

ornatus Smetana, 1948 [Note: Nomen nudum], see: Hypnogyra.

oronticus Coiffait, 1956a: 61 (Xantholinus; subgenus Echinophallus; Type locality: Liban: El Laboué).

  • — Coiffait, 1972: 260 (Xantholinus; subgenus Echinophallus; characters; Lebanon).

  • — Bordoni, 1972: 160 (Xantholinus; subgenus Echinophallus).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Idiolinus).

  • Distribution: Lebanon.

orthodoxus Olliff, 1887, see: Thyreocephalus.

osellai Bordoni, 1976: 230 (Xantholinus; subgenus Acanthophallus; Type locality: Turchia: Draganaz gecidi, 1320 m).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus).

  • Distribution: Turkey.

ottomanus Cameron, 1912, see: Vulda.

ovaliceps Scheerpeltz, 1965: 196 (Xantholinus; Type locality: S. Shan-States, Road 40 km e. of Taunggyi).

  • Distribution: Myanmar.

padus Coiffait, 1973: 107 (Xantholinus; subgenus Purrolinus; Type locality: Italie: Piacenza).

  • — Bordoni, 1982: 268 (Xantholinus; subgenus Polydontophallus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; probable synonym of jarrigei; Italy).

  • Distribution: Italy.

palliatus Melsheimer, 1844, see: Nudobius.

pallidipennis Kraatz, 1859, see: Thyreocephalus.

pallipennis Notman, 1921: 153 (Gyrohypnus; Type locality: New York: Chautauqua Co.: Westfield).

  • — Scheerpeltz, 1933: 1313 (Xantholinus; catalog).

  • Distribution: USA.

pampeanus Lynch, 1884, see: subtilis Boheman, 1858.

pamphylicus Coiffait, 1972, see: varnensis Coiffait, 1972.

panamensis Sharp, 1885: 485 (Xantholinus; Type locality: Panama, Volcan de Chiriqui 4000 to 6000 feet).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • Distribution: Panama.

pantokratoris Bordoni, 1975e: 140 (Xantholinus; subgenus Calolinus; Type locality: Corfù: M.te Pantokràtor, 900 m).

  • Distribution: Greece.

parajaponicus J. Li, 1993: 58 (Xantholinus; Type locality: West suburb of Dashiqiao, Liaoning).

  • Distribution: China.

parcipennis Bernhauer, 1933, see: Nepalinus.

parcus Eppelsheim, 1895b: 397 (Xantholinus; Type locality: Sikkim, Darjeling; Pegu).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Cameron, 1932: 26 (Xantholinus; characters; Sikkim; India; Burma).

  • — Scheerpeltz, 1933: 1313 (Xantholinus; catalog).

  • Distribution: India, Myanmar.

parisii C. Koch, 1937b: 268 (Xantholinus; subgenus Typhlolinus; Type locality: Garian, Gebel centrale della Tripolitania).

  • — Coiffait, 1972: 307 (Xantholinus; subgenus Helicophallus; characters; Libya).

  • Distribution: Libya.

paskoviensis Reitter, 1908, see: elegans Olivier, 1795.

pauper Sharp, 1889, see: Xestolinus.

pectoralis Erichson, 1839b: 322 (Xantholinus; Type locality: Columbia).

  • — Fauvel, 1891: 105 (Xantholinus; Venezuela).

  • — Fauvel, 1901: 82 (Xantholinus; Colombia; Venezuela).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Marcuzzi, 1977: 11 (Xantholinus; Venezuela).

  • Distribution: Colombia, Venezuela.

peliopterus Solsky, 1875, see: Allolinus.

pellegrinus Coiffait, 1970, see: tricolor Fabricius, 1787.

perezi Outerelo, 1976: 261 (Xantholinus; subgenus Heterolinus; Type locality: Valdemanco, Sierra de la Cabrera en Madrid, 1100 m).

  • Distribution: Spain.

pernitidus Cameron, 1925a: 185 (Xantholinus; Type locality: Pangrango, 2400–3000 m).

  • — Scheerpeltz, 1933: 1313 (Xantholinus; catalog).

  • Distribution: Indonesia.

phenicus Coiffait, 1971: 429 (Xantholinus; subgenus Calolinus; Type locality: Liban: Kartaba, 1200–1400 m).

  • — Coiffait, 1972: 264 (Xantholinus; subgenus Calolinus; characters; Lebanon).

  • — Bordoni, 1972: 162 (Xantholinus; subgenus Calolinus).

  • — Bordoni, 1975e: 143 (Xantholinus; subgenus Calolinus; cited as phenicius; notes; Rodi).

  • — Bordoni, 1976c: 86 (Xantholinus; cited as phenicius; Greece; Cyprus; Corfu).

  • — Bordoni, 1986a: 118 (Xantholinus; subgenus Calolinus; cited as phenicius; Syria).

  • — Bordoni, 1986b: 386 (Xantholinus; cited as phenicius; Greece).

  • — Bordoni, 1994: 26 (Xantholinus; subgenus Calolinus; cited as phenicius; Turkey).

  • Distribution: Greece, Cyprus, Turkey, Lebanon, Syria.

phoenicopterus Erichson, 1839, see: Eulissus.

piceus W. J. MacLeay, 1873, see: Zeteotomus.

piceus Cameron, 1926b: 345 (Xantholinus; [preoccupied]; Type locality: Dehra Dun district; Nim Nadi. Siwaliks; Mohan Rau).

  • — Cameron, 1932: 28 (Xantholinus; characters; India).

  • — Scheerpeltz, 1933: 1313 (Xantholinus; catalog).

  • — Coiffait, 1984e: 373 (Xantholinus; Nepal).

  • — Herman, 2001: 53 (Xantholinus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: India, Nepal.

picipennis LeConte, 1880, see: Neohypnus.

picipes Thomson, 1860, see: Gyrohypnus.

pictipennis Fauvel, 1904, see: Nudobius.

pilosus Roth, 1851, see: Eulissus.

piochardi Coiffait, 1956, see: rufipennis Erichson, 1839.

pleuralis Sharp, 1874, see: Nudobius.

politus Cameron, 1951c: 399 (Xantholinus; Type locality: Kenya Colony: Lukenia).

  • Distribution: Kenya.

poneli Coiffait, 1985, see: morandi Coiffait, 1958.

pradeshensis Coiffait, 1982a: 244 (Xantholinus; Type locality: Inde: Himachal Pradesh, Kothi, 2500–2900 m).

  • Distribution: India.

proceroides Coiffait, 1971: 433 (Xantholinus; subgenus Heterolinus; Type locality: Calabre).

  • — Coiffait, 1972: 295 (Xantholinus; subgenus Heterolinus; characters; Italy).

  • — Bordoni, 1972: 162, 205 (Xantholinus; subgenus Heterolinus; characters; Italy).

  • — Bordoni, 1982: 313 (Xantholinus; subgenus Paracyclinus; characters; Italy).

  • — Ciceroni, 1993: 75 (Xantholinus; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; Italy).

  • Distribution: Italy.

procerus Erichson, 1839b: 331 (Xantholinus; Type locality: Germania borealis; Hispania; Sardinia).

  • — Hochhuth, 1849: 108 (Xantholinus; characters; Lenkoran; Tiflis).

  • — Redtenbacher, 1849: 824 (Xantholinus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 503 (Xantholinus; characters; France).

  • — Kraatz, 1857d: 637 (Xantholinus; characters; Germany).

  • — Redtenbacher, 1857: 182 (Xantholinus; characters; Austria).

  • — Redtenbacher, 1874: 220 (Xantholinus; characters; Austria).

  • — Ganglbauer, 1895: 484 (Xantholinus; characters; Germany; Austria; Italy; Sardinia; Spain).

  • — Reitter, 1908: 119 (Xantholinus; characters; Germany; Austria; Southwestern Europe).

  • — Reitter, 1909: 138 (Xantholinus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1313 (Xantholinus; catalog).

  • — Scheerpeltz, 1961d: 126 (Xantholinus; subgenus Megalinus; Italy).

  • — Coiffait, 1972: 294 (Xantholinus; subgenus Heterolinus; characters; references; central Europe; Germany; Italy; Sardinia; Spain; Hungary).

  • — Bordoni, 1972: 162 (Xantholinus; subgenus Heterolinus).

  • — Bordoni, 1973c: 155 (Xantholinus; subgenus Heterolinus; characters; Romania).

  • — Bordoni, 1975: 75 (Xantholinus; subgenus Paracylinus; type species of subgenus; notes).

  • — Tóth, 1989: 22 (Xantholinus; characters).

  • Distribution: Romania, Georgia, Azerbaijan.

validus Petri, 1912: 68 (Xantholinus; Type locality: Schässburg).

  • — Scheerpeltz, 1933: 1315 (Xantholinus; catalog).

  • — Coiffait, 1972: 310 (Xantholinus; unknown species).

  • — Bordoni, 1973c: 155 (Xantholinus; synonym of procerus).

provincialis Giebel, 1856, see: westwoodianus Heer, 1856.

pseudobalaton Tóth, 1985: 175 (Xantholinus; subgenus Meneidophallus; Type locality: Siófok Umg.).

  • — Tóth, 1989: 32 (Xantholinus; characters; Hungary).

  • Distribution: Hungary.

pseudohesperius Reitter, 1908, see: Lepidophallus.

puncticeps Guérin-Méneville, 1844, see: Heterolinus.

puncticeps Jacquelin du Val, 1857, see: humeralis Erichson, 1839.

puncticeps Eichelbaum, 1913, see: eichelbaumi Steel, 1949.

punctiger Gemminger and Harold, 1868, see: humeralis Erichson, 1839.

punctiventris Sharp, 1889, see: Liotesba.

punctulatus Gravenhorst, 1802, see: linearis Olivier, 1795.

pusillus Sachse, 1852, see: Neohypnus.

puthzi Bordoni, 1979: 108 (Xantholinus; subgenus Calolinus; Type locality: Asia Minore: S-Anatolia, Kilikischer Taurus, Sertaval Pass, 1600 mt.).

  • Distribution: Turkey.

pygialis Sharp, 1876, see: Agerodes.

quadratus Stephens, 1833: 260 (Gyrohypnus; [Note: Nomen dubium]; Type locality: Suffolk).

  • — Stephens, 1839: 403 (Xantholinus; characters; England).

  • Distribution: Britain.

quadrimaculatus Eichelbaum, 1910, see: Nudobius.

radiosus Peyron, 1858, see: Phacophallus.

raffrayi Fauvel, 1879: 93 (Xantholinus; Type locality: Gilolo).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; catalog).

  • Distribution: Indonesia.

raffrayi Sainte-Claire Deville, 1926, see: appenninicola Steel, 1946.

reitteri Coiffait, 1966f: 198 (Xantholinus; subgenus Acanthophallus; Type locality: Kavkaz, Svanetiia).

  • — Coiffait, 1972: 286 (Xantholinus; subgenus Acanthophallus; characters; Caucasus).

  • — Bordoni, 1975: 64 (Xantholinus; subgenus Acanthophallus; notes; Azerbaijan).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus).

  • Distribution: Georgia, Azerbaijan.

remotus Eppelsheim, 1895: 208 (Xantholinus; Type locality: Ganale Gudda, Arussi Galla).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; catalog).

  • Distribution: Ethiopia.

resectus Sharp, 1885: 480 (Xantholinus; Type locality: Mexico).

  • — Sharp, 1887: 791 (Xantholinus; Mexico).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; catalog).

  • Distribution: Mexico.

rhaeticus Bordoni, 1972, see: tricolor Fabricius, 1787.

rhenanus Coiffait, 1962a: 272 (Xantholinus; subgenus Xantholinus; Type locality: Bas-Rhin, Forêt de Hagueneau).

  • — Lohse, 1964: 166 (Xantholinus; subgenus Xantholinus; characters; central Europe).

  • — Smetana, 1964d: 73 (Xantholinus; subgenus Xantholinus; Czechoslovakia).

  • — Horion, 1965: 101 (Xantholinus; subgenus Xantholinus; France; Germany; Austria; Czechoslovakia; Denmark).

  • — Smetana, 1966: 329, 330 (Xantholinus; characters; notes; Czechoslovakia).

  • — Coiffait, 1972: 270 (Xantholinus; subgenus Xantholinus; characters; references; France; Germany; Czechoslovakia; Denmark; Austria).

  • — Bordoni, 1972: 195 (Xantholinus; subgenus Xantholinus; characters; Italy).

  • — Bordoni, 1972c (Xantholinus; subgenus Paraphallus; correction of subgeneric assignment in Bordoni, 1972: 162, 188, 195).

  • — Bordoni, 1982: 293 (Xantholinus; subgenus Paraphallus; characters; Italy).

  • — Tóth, 1985: 176 (Xantholinus; notes; Hungary).

  • — Drugmand and Haghebaert, 1987: 325 (Xantholinus; Belgium).

  • — Lucht, 1987: 103 (Xantholinus; checklist; Germany; Poland; Czechoslovakia; Austria; France; Benelux; Denmark).

  • — Schülke and Uhlig, 1988: 11 (Xantholinus; notes; Germany).

  • — Tóth, 1989: 28 (Xantholinus; characters; Hungary).

  • — Drugmand, 1990a: 89 (Xantholinus; characters; Belgium).

  • — Assing, 1993: 20 (Xantholinus; bionomics; Germany).

  • — Janák, 1993a: 47 (Xantholinus; subgenus Xantholinus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; Italy).

  • — Mikat, Fremuth, and Prouza, 1997: 111 (Xantholinus; Czech Republic).

  • Distribution: Europe.

richteri Bernhauer, 1912e: 175 (Xantholinus; Type locality: Argentinien: Prov. Tucuman).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; catalog).

  • Distribution: Argentina.

rossicus Bordoni, 1975, see: linearis Olivier, 1795.

roubali Coiffait, 1956b: 142 (Xantholinus; Type locality: Tchécoslovaquie: Bratislava. Also cited from Russia).

  • — Smetana, 1958: 112 (Xantholinus; subgenus Xantholinus; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 159 (Xantholinus; characters; Sweden; Denmark).

  • — Lohse, 1964: 166 (Xantholinus; subgenus Xantholinus; characters; central Europe).

  • — Horion, 1965: 102 (Xantholinus; subgenus Xantholinus; Slovakia; Czech Republic; Ukraine; Germany).

  • — Smetana, 1966: 330 (Xantholinus; characters).

  • — Coiffait, 1972: 278 (Xantholinus; subgenus Xantholinus; characters; references; Czechoslovakia; Germany; Denmark; Sweden; Austria).

  • — Bordoni, 1972: 162 (Xantholinus; subgenus Meneidophallus).

  • — Bordoni, 1973c: 152 (Xantholinus; subgenus Meneidophallus; characters; notes; Roumania).

  • — Bordoni, 1975: 68 (Xantholinus; subgenus Meneidophallus; notes; Russia).

  • — Ádám, 1987: 148 (Meneidophallus; collecting notes; Hungary).

  • — Lucht, 1987: 103 (Xantholinus; checklist; Germany; Poland; Czechoslovakia; Austria; Denmark; Sweden).

  • — Schülke and Uhlig, 1988: 11 (Xantholinus; notes; Germany).

  • — Tóth, 1989: 32 (Xantholinus; characters; Hungary).

  • — Nowosad, 1990: 96 (Xantholinus; collected from mole nest; notes; Poland).

  • — Janák, 1993a: 47 (Xantholinus; subgenus Meneidophallus; Czech Republic; Slovakia).

  • Distribution: Europe, Ukraine.

rudis Sharp, 1885: 477 (Xantholinus; Type locality: Guatemala, near the city, San Geronimo, Mirandilla 1700 feet).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; catalog).

  • Distribution: Guatemala.

rudis Eppelsheim, 1895, see: Thyreocephalus.

rufescens Erichson, 1839b: 328 (Xantholinus; Type locality: Puerto Cabello, Columbiae).

  • — Fauvel, 1891: 106 (Xantholinus; Venezuela).

  • — Fauvel, 1901: 83 (Xantholinus; Venezuela; Colombia; Saint Thomas; Cayenne; Amazonian region; New Granada).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; catalog).

  • Distribution: Brazil, Colombia, Venezuela, French Guiana, West Indies.

aeneiceps Sharp, 1876: 202 (Xantholinus; Type locality: Ega).

  • — Fauvel, 1891: 106 (Xantholinus; synonym of rufescens).

  • — Fauvel, 1901: 83 (Xantholinus; synonym of rufescens).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; synonym of rufescens).

ruffoi Bordoni, 1972: 212 (Xantholinus; subgenus Megalinus; Type locality: Veneto, Monte Grappa, Vincenza).

  • — Bordoni, 1982: 316 (Xantholinus; subgenus Heterolinus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; probable synonym of bordonii; Italy).

  • Distribution: Italy.

ruficaudatus Cameron, 1932: 27 (Xantholinus; Type locality: Darjeeling: Debrepani, alt. 6000 feet).

  • — Scheerpeltz, 1933: 1314 (Xantholinus; catalog).

  • — Coiffait, 1977b: 206 (Xantholinus; Bhutan).

  • Distribution: India, Bhutan.

ruficollis P. Lucas, 1846, see: Nudobius.

rufipennis Erichson, 1839b: 322 (Xantholinus; Type locality: Sicilia; Syria).

  • — Redtenbacher, 1849: 692 (Xantholinus; characters; Austria).

  • — Rosenhauer, 1856: 70 (Xantholinus; notes).

  • — Kraatz, 1857d: 640 (Xantholinus; characters; Austria).

  • — Redtenbacher, 1857: 182 (Xantholinus; characters; Austria).

  • — Kraatz, 1858a: 65 (Xantholinus; Crete; Greece).

  • — Saulcy, 1864: 641 (Xantholinus; notes; Jerusalem).

  • — Baudi, 1870: 387, 388 (Xantholinus; characters).

  • — Ganglbauer, 1895: 482 (Xantholinus; characters; Dalmatia; Sicily; eastern Mediterranean region).

  • — Reitter, 1908: 121 (Xantholinus; characters; Europe; southern and eastern Dalmatia).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1314 (Xantholinus; catalog).

  • — Schatzmayr, 1943: 114 (Xantholinus; Albania).

  • — Scheerpeltz, 1961e: 136 (Xantholinus; subgenus Calolinus; Rhodes).

  • — Coiffait, 1966: 21 (Xantholinus; subgenus Calolinus; Turkey).

  • — Coiffait, 1970e: 62 (Xantholinus; Cyprus).

  • — Bordoni, 1971: 685 (Xantholinus; subgenus Calolinus; Turkey).

  • — Coiffait, 1972: 262 (Xantholinus; subgenus Calolinus; characters; references; Sicily; Italy; Greece; Cyprus; Asia Minor; Syria; Lebanon; Palestine).

  • — Bordoni, 1972: 162, 202 (Xantholinus; subgenus Calolinus; characters; Italy).

  • — Bordoni, 1973c: 151 (Xantholinus; subgenus Calolinus; notes; Romania; Palestine; Syria; Cyprus; Greece; Croatia; Italy).

  • — Bordoni, 1975e: 145 (Xantholinus; subgenus Calolinus; notes; Rodi).

  • — Bordoni, 1976c: 86 (Xantholinus; notes; Italy; Albania; Asia Minor; Palestine).

  • — Bordoni, 1976d: 120 (Xantholinus; notes; Italy).

  • — Bordoni, 1982: 310 (Xantholinus; subgenus Calolinus; characters; Italy).

  • — Bordoni, 1986b: 386 (Xantholinus; Greece).

  • — Tóth, 1989: 23 (Xantholinus; characters).

  • — Sparacio, 1995: 158 (Xantholinus; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; Italy).

  • — Assing and Wunderle, 2001: 36 (Xantholinus; Cyprus).

  • Distribution: Europe, Turkey, Lebanon, Israel, Syria.

titus Saulcy, 1864: 642 (Xantholinus; Type locality: Jérusalem, Beïroût).

  • — Fauvel, 1875a: XXVI [= 1875b: 228] (Xantholinus; synonym of rufipennis).

  • — Ganglbauer, 1895: 482 (Xantholinus; synonym of rufipennis).

  • — Reitter, 1908: 121 (Xantholinus; variety of rufipennis).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; synonym of rufipennis).

  • — Coiffait, 1972: 262 (Xantholinus; synonym of rufipennis).

  • — Bordoni, 1972: 202 (Xantholinus; cited as tytus; synonym of rufipennis).

  • — Bordoni, 1982: 310 (Xantholinus; synonym of rufipennis).

infuscatus Bernhauer, 1910a: 72 (Xantholinus; subspecies of rufipennis; Type locality: Dalmatien; Korfu; Albanien; Türkei; Herzegowina; Griechenland).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; variety of rufipennis).

  • — Coiffait, 1972: 262 (Xantholinus; variety of rufipennis; characters).

  • — Bordoni, 1972: 202 (Xantholinus; synonym of rufipennis).

  • — Bordoni, 1982: 310 (Xantholinus; synonym of rufipennis).

piochardi Coiffait, 1956a: 62 (Xantholinus; subgenus Calolinus; Type locality: Jérusalem. Also cited from Syria, Lebanon, Cyprus and Greece).

  • — Scheerpeltz, 1956c: 1099 (Xantholinus; subgenus Calolinus; Greece).

  • — Scheerpeltz, 1963a: 419 (Xantholinus; subgenus Calolinus; Greece).

  • — Scheerpeltz, 1964a: 299 (Xantholinus; subgenus Calolinus; Crete).

  • — Coiffait, 1972: 262 (Xantholinus; synonym of rufipennis).

  • — Bordoni, 1972: 202 (Xantholinus; synonym of rufipennis).

  • — Bordoni, 1982: 310 (Xantholinus; synonym of rufipennis).

rufipes P. Lucas, 1846: 106 (Xantholinus; Type locality: Algeria: environs de Bab-el-Ouad; environs du cercle de Lacalle, pres les marais du lac Tonga).

  • — Fauvel, 1878: 117 [= 1878a: 37] (Xantholinus; Algeria).

  • — Fauvel, 1886: 49 [= 1886a: 41] (Xantholinus; Tunisia; Algeria).

  • — Fauvel, 1902b: 102 (Xantholinus; Algeria; Tunisia; Tripolitania).

  • — Reitter, 1908: 119 (Xantholinus; characters; Spain; Sicily; Italy; Algeria).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1314 (Xantholinus; catalog).

  • — Coiffait, 1956a: 71 (Xantholinus; subgenus Helicophallus; characters in key; Tunisia; Algeria).

  • — Coiffait, 1962b: 398, 404 (Xantholinus; subgenus Helicophallus; characters; Algeria; Tunisia).

  • — Coiffait, 1972: 297 (Xantholinus; subgenus Helicophallus; characters; references; Algeria; Tunisia; Sicily; Tripolitania).

  • — Bordoni, 1972: 164, 229 (Xantholinus; subgenus Helicophallus; characters; Italy).

  • — Bordoni, 1976c: 96 (Xantholinus; Algeria).

  • — Bordoni, 1982: 321 (Xantholinus; subgenus Helicophallus; characters; Italy).

  • — Sparacio, 1995: 158 (Xantholinus; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; Italy).

  • Distribution: Libya, Tunisia, Algeria, Spain, Italy.

cordieri Boieldieu, 1859: 464 (Xantholinus; Type locality: Sicile).

  • — Fauvel, 1869: 489 (Xantholinus; synonym of rufipes).

  • — Fauvel, 1875a: XXVI [= 1875b: 228] (Xantholinus; synonym of rufipes).

  • — Fauvel, 1878: 117 [= 1878a: 37] (Xantholinus; synonym of rufipes).

  • — Reitter, 1908: 119 (Xantholinus; synonym of rufipes).

  • — Bernhauer and Schubert, 1914: 307 (Xantholinus; synonym of rufipes).

  • — Coiffait, 1972: 297 (Xantholinus; synonym of rufipes).

  • — Bordoni, 1982: 321 (Xantholinus; synonym of rufipes).

rufitarsis Fauvel, 1877, see: Thyreocephalus.

rufus Grimmer, 1841: 33 (Xantholinus; [Note: Nomen dubium]; Type locality: Steiermark).

  • Distribution: Austria.

rulomus Blackwelder, 1943: 482 (Xantholinus; Type locality: Dominican Republic: Jarabacoa, elevation 1500 to 4000 feet).

  • Distribution: West Indies.

salvini Sharp, 1885: 483 (Xantholinus; Type locality: Guatemala, near the city, San Geronimo).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • Distribution: Guatemala.

samniticus Bordoni, 1988, see: cerrutii Coiffait, 1964.

sanguinipennis Kolenati, 1846, see: Gauropterus.

sanguinipennis LeConte, 1880, see: Neohypnus.

sanguinipes Reitter, 1889, see: Gauropterus.

sanguinolentus Melsheimer, 1844, see: Neohypnus.

sardous Gridelli, 1947: 78 (Xantholinus; Type locality: Oristano).

  • — Coiffait, 1956a: 64 (Xantholinus; characters in key).

  • — Coiffait, 1962b: 397, 401 (Xantholinus; subgenus Xantholinus; characters; Morocco; Algeria).

  • — Coiffait, 1972: 275 (Xantholinus; subgenus Xantholinus; characters; references; Sicily; North Africa; Mallorca).

  • — Bordoni, 1972: 194 (Xantholinus; subgenus Xantholinus; characters; Italy).

  • — Bordoni, 1972c (Xantholinus; subgenus Paraphallus; correction of subgeneric assignment in Bordoni, 1972: 162, 188, 194).

  • — Bordoni, 1982: 292 (Xantholinus; subgenus Paraphallus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; Italy).

  • Distribution: Algeria, Morocco, Spain, France, Italy.

corsicus Coiffait, 1972: 276 (Xantholinus; subgenus Xantholinus; Type locality: Corse: Zonza, 800 m).

  • — Coiffait, 1973: 109 (Xantholinus; France).

  • — Bordoni, 1976c: 91 (Xantholinus; notes; Italy).

  • — Bordoni, 1982: 291 (Xantholinus; subgenus Paraphallus; characters; Italy).

  • — Ciceroni, 1994: 116 (Xantholinus; synonym of sardous).

schuelkei Bordoni, 1986a: 119 (Xantholinus; subgenus Paracyclinus ?; Type locality: Siria settentrionale: Zainie, 45 km NO Latakia).

  • Distribution: Syria.

schuleri Coiffait, 1958, see: distans Mulsant and Rey, 1853.

schweigeri Coiffait, 1966: 22 (Xantholinus; subgenus Xantholinus; Type locality: environs du Lac Abant près de Bolu, Turquie d'Asie).

  • — Tóth, 1985: 176 (Xantholinus; habitat notes; Hungary; Austria; Bulgaria).

  • — Tóth, 1989: 32 (Xantholinus; characters; Hungary).

  • — Lohse and Lucht, 1989: 159 (Xantholinus; characters).

  • — Herman, 2001: 32 (Xantholinus; synonymic note).

  • Distribution: Turkey, Bulgaria, Hungary, Austria.

schweigeri Coiffait, 1971: 431 (Xantholinus; subgenus Xantholinus; [preoccupied]; Type locality: environs d'Abant See près de Bolu, Turquie d'Asie).

  • — Bordoni, 1971: 686 (Xantholinus; subgenus Meneidophallus; Turkey).

  • — Bordoni, 1972: 162 (Xantholinus; subgenus Meneidophallus).

  • — Coiffait, 1972: 279 (Xantholinus; subgenus Xantholinus; characters; references; Anatolia).

  • — Herman, 2001: 32 (Xantholinus; synonym of schweigeri).

magyaricus Tóth, 1985: 174 (Xantholinus; subgenus Meneidophallus; Type locality: Hungaria occidentalis: Velencei-tó, Sukoró).

  • — Lohse, 1987: 138 (Xantholinus; synonym of schweigeri).

  • — Tóth, 1989: 29 (Xantholinus; cited as valid species; [Note: I assume Tóth overlooked the synonymy by Lohse]; characters; Hungary).

schweigeri Coiffait, 1971, see: schweigeri Coiffait, 1966.

scoticus Joy, 1913, see: Gyrohypnus.

scripticollis Hochhuth, 1849, see: Zeteotomus.

scutellaris Fauvel, 1900, see: Megalinus.

sejugatus G. Benick, 1953, see: longiventris Heer, 1839.

semipallidus Bernhauer, 1931c: 128 (Xantholinus; Type locality: Chinabor: Manch de Adams).

  • — Scheerpeltz, 1933: 1314 (Xantholinus; catalog).

  • — Bordoni, 2000: 127 (Xantholinus; characters; China).

  • Distribution: China.

semirufus Reitter, 1901, see: fortepunctatus Motschulsky, 1860.

serpens Sharp, 1885: 479 (Xantholinus; Type locality: Mexico, Oaxaca).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • Distribution: Mexico.

sexpunctatus Fauvel, 1905b: 171 (Xantholinus; Type locality: Madagascar).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • Distribution: Madagascar.

sharpi Broun, 1880, see: Pseudoxantholinus.

sideralis Fauvel, 1878g: 539 (Xantholinus; Type locality: Australie occidentale).

  • — Olliff, 1887: 486 (Xantholinus; characters; Australia).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • Distribution: Australia.

sidonensis Coiffait, 1956a: 62 (Xantholinus; subgenus Calolinus; Type locality: Liban: Saïda).

  • — Scheerpeltz, 1956c: 1099 (Xantholinus; subgenus Calolinus; Cyprus).

  • — Bordoni, 1971: 679, 685 (Xantholinus; subgenus Calolinus; characters; Turkey).

  • — Coiffait, 1972: 264 (Xantholinus; characters; Lebanon).

  • — Bordoni, 1972: 162 (Xantholinus; subgenus Calolinus).

  • — Bordoni, 1976c: 87 (Xantholinus; Turkey).

  • Distribution: Lebanon, Turkey, Cyprus.

sighisoarae Bordoni, 1973c: 151 (Xantholinus; subgenus Echinophallus; Type locality: Roumanie: Sighisoara).

  • — Tóth, 1989: 27 (Xantholinus; characters).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Idiolinus).

  • Distribution: Romania.

silvanus Peyerimhoff, 1917, see: Gyrohypnus.

similis Wendeler, 1956a: 227 (Xantholinus, cited as Xanthelinus; Type locality: Brasilien: Nova Teutonia).

  • Distribution: Brazil.

socius Fauvel, 1877, see: Notolinus.

solitarius Blackwelder, 1943: 486 (Xantholinus; Type locality: Guadeloupe: Basse Terre, Matouba, 3 kilometers northwest of St. Claude).

  • Distribution: West Indies.

sontianus Bordoni, 1972, see: bordonii Coffait, 1970.

speciosus Erichson, 1839, see: Agerodes.

strandi Coiffait, 1958, see: audrasi Coiffait, 1956.

strigiceps Sharp, 1885, see: Eulissus.

strigicollis Bernhauer, 1928e: 46 (Xantholinus; Type locality: S. Catharina: Hammonia).

  • — Scheerpeltz, 1933: 1314 (Xantholinus; catalog).

  • Distribution: Brazil.

styriacus Grimmer, 1841: 33 (Xantholinus; [Note: Nomen dubium]; Type locality: Steiermark).

  • Distribution: Austria.

subaequalis Lynch, 1884, see: subtilis Boheman, 1858.

sublaevis Fauvel, 1875, see: Allolinus.

sublinearis Coiffait, 1970: 288 (Xantholinus; Type locality: Chantove, 1300 m, près Verria, Beroia, Mecédoine grecque).

  • — Coiffait, 1972: 272 (Xantholinus; subgenus Xantholinus; characters; Greece; Bulgaria).

  • — Bordoni, 1972: 162, 201 (Xantholinus; subgenus Paraphallus; characters; Italy).

  • — Bordoni, 1972c (Xantholinus; subgenus Xantholinus; correction of subgeneric assignment in Bordoni, 1972: 162, 201).

  • — Bordoni, 1982: 303 (Xantholinus; subgenus Xantholinus; characters; Italy).

  • — Tóth, 1985: 176 (Xantholinus; notes; Italy; Hungary; Slovakia).

  • — Tóth, 1989: 27 (Xantholinus; characters; Hungary).

  • — Lohse and Lucht, 1989: 159 (Xantholinus; characters; notes).

  • — Ciceroni, 1993: 74 (Xantholinus; Italy).

  • — Janák, 1993a: 47 (Xantholinus; subgenus Xantholinus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; Italy).

  • Distribution: Greece, Bulgaria, Hungary, Slovakia, Italy.

substrigosus Joy, 1913, see: Leptacinus.

subtilis Boheman, 1858: 27 (Xantholinus; Type locality: Buenos Ayres).

  • — Lynch, 1884: 193 (Xantholinus; subgenus Gyrohypnus; characters; Argentina).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1314 (Xantholinus; catalog).

  • — Frank, 1982: 32 (Xantholinus; list of parasites).

  • Distribution: Argentina.

pampeanus Lynch, 1884: 185 (Xantholinus; subgenus Megalinus; Type locality: Chacabuco).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; synonym of subtilis).

subaequalis Lynch, 1884: 187 (Xantholinus; subgenus Megalinus; Type locality: Chacabuco et Baradero).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; variety of subtilis).

suffusus Sharp, 1874, see: Lepidophallus.

svanetskiensis Coiffait, 1973: 108 (Xantholinus; subgenus Acanthophallus; Type locality: Svanetskij Khrebet, Caucase).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus; cited as swanetskiensis).

  • Distribution: Georgia.

taitiensis Boheman, 1858, see: Thyreocephalus.

tauricus Bordoni, 1972b: 128 (Xantholinus; subgenus Helicophallus; Type locality: Turchia: Tauro, Anemas).

  • — Bordoni, 1976c: 98 (Xantholinus; notes; Turkey).

  • Distribution: Turkey.

temporalis Sharp, 1876, see: Agerodes.

temporalis LeConte, 1880, see: Neohypnus.

tenebrarius† Scudder, 1900: 58 (Xantholinus; fossil; Type locality: Florissant, Colorado).

  • — Handlirsch, 1907: 726 (Xantholinus; catalog; Miocene; Colorado).

  • Distribution: USA.

tener Waltl, 1835, see: Leptacinus.

tenuipes Baudi, 1870, see: Vulda.

tenuipunctus Fauvel, 1900, see: Megalinus.

thomsoni Schwarz, 1872, see: Gyrohypnus.

tingitanus Coiffait, 1962c: 73 (Xantholinus; subgenus Xantholinus; Type locality: Tanger).

  • — Coiffait, 1962b: 398, 401, 405 (Xantholinus; subgenus Xantholinus; characters; Morocco).

  • — Coiffait, 1972: 275 (Xantholinus; subgenus Xantholinus; characters; references; Morocco; Algeria).

  • — Bordoni, 1972c (Xantholinus; subgenus Paraphallus; correction of subgeneric assignment in Bordoni, 1972: 162, 188).

  • Distribution: Morocco, Algeria.

titus Saulcy, 1864, see: rufipennis Erichson, 1839.

tothi Bordoni, 1986: 68 (Xantholinus; subgenus Purrolinus; Type locality: Ungheria: Vértes hegység, Fáni völgy).

  • Distribution: Hungary.

toumayeffi Bordoni, 1986, see: tricolor Fabricius, 1787.

translucidus Scriba, 1870: 81 (Xantholinus; Type locality: Minas de Esploradora Sierrae Nevadae et Gibraltar).

  • — Fauvel, 1886: 49 [= 1886a: 41] (Xantholinus; Morocco; Algeria).

  • — Fauvel, 1902b: 101 (Xantholinus; Morocco; Algeria).

  • — Reitter, 1908: 121 (Xantholinus; characters; Spain; Gibraltar).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1314 (Xantholinus; catalog).

  • — Coiffait, 1956a: 60 (Xantholinus; subgenus Echinophallus; characters in key; Pyrenees; Algeria; Portugal).

  • — Coiffait, 1958: 23 (Xantholinus; subgenus Echinophallus; Pyrenees).

  • — Coiffait, 1962b: 395, 400 (Xantholinus; subgenus Echinophallus; characters; Morocco; Algeria).

  • — Coiffait, 1972: 259 (Xantholinus; subgenus Echinophallus; characters; references; North Africa; Iberia; Pyrenees).

  • — Bordoni, 1972: 160 (Xantholinus; subgenus Echinophallus).

  • — Coiffait, 1973a: 273 (Xantholinus; Morocco).

  • — Bordoni, 1976c: 84 (Xantholinus; Portugal; Spain).

  • — Bordoni, 1976d: 120 (Xantholinus; Algeria).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Idiolinus).

  • Distribution: Algeria, Morocco, Spain, Portugal.

trellai Szujecki, 1969, see: azuganus Reitter, 1908.

tricolor Fabricius, 1787: 221 (Staphylinus; Type locality: Dania).

  • — Paykull, 1789: 23 (Staphylinus; characters; Sweden).

  • — Gmelin, 1790: 2030 (Staphylinus; characters).

  • — Fabricius, 1793: 537 (Paederus; characters).

  • — Panzer, 1795: 363 (Paederus; characters; Germany).

  • — Schrank, 1798: 647 (Staphylinus; characters; notes).

  • — Paykull, 1800: 378 (Staphylinus; characters; Sweden).

  • — Fabricius, 1801: 609 (Paederus; characters).

  • — Marsham, 1802: 516 (Staphylinus; characters; Britain).

  • — Marsham, 1802: 517 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1914: 308, cited affinis Marsham as a synonym of tricolor; Marsham did not describe the species as new, he attributed it to Paykull]; characters; Britain).

  • — Turton, 1802: 523 (Staphylinus; characters; Denmark).

  • — Latreille, 1804: 330 (Staphylinus; characters; Germany; Sweden).

  • — Gyllenhal, 1810: 355 (Staphylinus; characters; Sweden).

  • — Curtis, 1829: 27 (Gyrohypnus; catalog; Britain).

  • — C. Sahlberg, 1830: 334 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 33 (Gyrohypnus; Finland; Sweden).

  • — Mannerheim, 1831: 447 (Gyrohypnus; Finland; Sweden).

  • — Stephens, 1833: 261 (Gyrohypnus; characters; England).

  • — Stephens, 1833: 261 (Gyrohypnus; [Note: Cited as affinis but probably a misidentification of tricolor; Stephens based his treatment of affinis on the specimens used by Marsham and the specimens identified as affinis by Marsham have been regarded as misidentification of tricolor]; characters; England).

  • — Runde, 1835: 11 (Gyrohypnus; characters; Germany).

  • — Nordmann, 1837: 117 (Gyrohypnus; Sweden; Germany).

  • — Westwood, 1838a: 16 (Gyrohypnus; type species).

  • — Zetterstedt, 1838: 66 (Xantholinus; characters; habitat; Lapland).

  • — Erichson, 1839a: 427 (Xantholinus; characters; Germany).

  • — Heer, 1839: 246 (Xantholinus; characters; Switzerland).

  • — Erichson, 1839b: 331 (Xantholinus; characters; Europe).

  • — Stephens, 1839: 402 (Xantholinus; characters; England).

  • — Märkel, 1845: 236 (Xantholinus; with Formica fuliginosa).

  • — Kolenati, 1846: 15 (Xantholinus; Caucasus; Iberia).

  • — Küster, 1848: 10 (Xantholinus; characters; Europe).

  • — Jacobsen, 1849: 679 (Xantholinus; characters; Denmark).

  • — Hochhuth, 1849: 108 (Xantholinus; characters; Caucasus; Iberia; Tiflis).

  • — Redtenbacher, 1849: 692 (Xantholinus; characters; Austria).

  • — Mulsant and Rey, 1853: 58 (Xantholinus; characters; notes; France).

  • — Hardy, 1851: 38 (Xantholinus; notes; England).

  • — Rosenhauer, 1856: 70 (Xantholinus; Granada).

  • — Fairmaire and Laboulbène, 1856: 502 (Xantholinus; characters; France).

  • — Kraatz, 1857d: 638 (Xantholinus; characters; Germany).

  • — Redtenbacher, 1857: 182 (Xantholinus; characters; Austria).

  • — Thomson, 1860: 191 (Xantholinus; characters; Scandinavia).

  • — Fauvel, 1865a: 298 [= 1865b: 48] (Xantholinus; habitat; France).

  • — Fauvel, 1873b: 104 [= 1873a: 390] (Xantholinus; characters; France; Europe; Georgia; Caucasus).

  • — Redtenbacher, 1874: 220 (Xantholinus; characters; Austria).

  • — Seidlitz, 1875: 281 (Xantholinus; characters; Baltic region).

  • — J. Sahlberg, 1876: 35 (Xantholinus; Finland).

  • — Mulsant and Rey, 1877a: 272 (Xantholinus; adult and larval characters; France).

  • — Fowler, 1888: 290 (Xantholinus; characters; Britain).

  • — Eppelsheim, 1892: 335 (Xantholinus; Turkestan).

  • — Ganglbauer, 1895: 483 (Xantholinus; characters; Europe).

  • — Xambeu, 1900: 39 (Xantholinus; larval characters).

  • — Reitter, 1909: 138 (Xantholinus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • — Johansen, 1914: 436 (Xantholinus; subgenus Xantholinus; characters; Denmark).

  • — Šulc, 1922 (Xantholinus; osmeterium; anatomy; histology; physiology).

  • — Portevin, 1929: 372 (Xantholinus; characters; France).

  • — Scheerpeltz, 1933: 1315 (Xantholinus; catalog).

  • — C. Koch, 1937: 88 (Xantholinus; characters; Italy).

  • — Gridelli, 1950: 2 (Xantholinus; characters; Italy, Austria, Slovenia, Croatia, Bosnia, Hungary).

  • — V. Hansen, 1952: 18 (Xantholinus; subgenus Xantholinus; characters; Denmark).

  • — Coiffait, 1956a: 66 (Xantholinus; subgenus Purrolinus; characters in key).

  • — Smetana, 1958: 109 (Xantholinus; subgenus Xantholinus; characters; ecology; Czechoslovakia).

  • — Lohse, 1960a: 37, 38 (Xantholinus; characters; notes).

  • — Palm, 1963b: 51 (Xantholinus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 164 (Xantholinus; subgenus Purrolinus; characters; central Europe).

  • — Horion, 1965: 98 (Xantholinus; subgenus Purrolinus; northern and central Europe south to northern Italy and northern Balkans).

  • — Szujecki, 1966: 169 (Xantholinus; characters of egg).

  • — Pototskaia, 1967: 48 (Xantholinus; larval characters).

  • — Szujecki, 1968a: 726 (Xantholinus; Poland).

  • — Coiffait, 1972: 283 (Xantholinus; subgenus Purrolinus; characters; references; central Europe; Alps; Hungary; Yugoslavia; Russia; Scandinavia; Britain; France).

  • — Bordoni, 1972: 161, 168 (Xantholinus; subgenus Purrolinus; characters; Italy).

  • — Bordoni, 1973: 45 (Xantholinus; subgenus Purrolinus; characters; Hungary).

  • — Bordoni, 1973c: 153 (Xantholinus; subgenus Purrolinus; notes; Romania).

  • — Osella and Zanetti, 1975: 106 (Xantholinus; collected from nest of Talpa europaea; notes; Italy).

  • — Bordoni, 1975: 62 (Xantholinus; subgenus Purrolinus; notes; Russia).

  • — Shilov, 1975a: 375 [= 1975b: 97] (Xantholinus; Komi region of Russia).

  • — Bordoni, 1976c: 91 (Xantholinus; notes; France).

  • — Bordoni, 1976d: 121 (Xantholinus; Alps; Germany; Netherlands).

  • — Pope, 1977: 29 (Xantholinus; Britain).

  • — Bordoni, 1982: 256 (Xantholinus; subgenus Purrolinus; characters; Italy).

  • — Waterston, Owen, Welch, Bacchus and Hammond, 1981: 274 (Xantholinus; Outer Hebrides).

  • — Frank, 1982: 33 (Xantholinus; list of parasites).

  • — Ádám, 1987: 148 (Purrolinus; collecting notes; Hungary).

  • — Lucht, 1987: 103 (Xantholinus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Tóth, 1989: 24 (Xantholinus; characters; Hungary).

  • — Nowosad, 1990: 96 (Xantholinus; collected from mole nest; notes; Poland).

  • — Janák, 1993a: 47 (Xantholinus; subgenus Purrolinus; Czech Republic; Slovakia).

  • — Assing, 1993: 25 (Xantholinus; bionomics; Germany).

  • — Ádám, 1996: 243 (Purrolinus; cited as misidentification; synonym of laeticeps).

  • — Herman, 2001: 60 (Xantholinus; discussion; traditional use of name as species of Xantholinus, not as species of Lithocharis should be continued).

  • Distribution: Europe, Russia, Georgia, Tajikistan.

laeticeps Reitter, 1908: 120 [= 1908a: 23] (Xantholinus; aberration of tricolor; [Note: Nomen nudum (Articles 1.3.4, 45.5)]).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; variety of tricolor).

  • — Gridelli, 1950: 2 (Xantholinus; synonym of tricolor).

  • — Smetana, 1958: 109 (Xantholinus; synonym of tricolor).

  • — Coiffait, 1972: 284 (Xantholinus; variety of tricolor; Czechoslovakia).

  • — Bordoni, 1972: 164 (Xantholinus; subgenus Helicophallus; variety of tricolor).

  • — Bordoni, 1982: 256 (Xantholinus; synonym of tricolor).

  • — Ádám, 1996: 243 (Purrolinus; valid name; Hungary).

pellegrinus Coiffait, 1970: 291 (Xantholinus; subgenus Acanthophallus; Type locality: Col San Pellegrino, Trentin).

  • — Coiffait, 1972: 288 (Xantholinus; subgenus Acanthophallus; characters; Italy).

  • — Bordoni, 1972: 161, 179 (Xantholinus; subgenus Acanthophallus; characters; Italy).

  • — Bordoni, 1982: 275 (Xantholinus; subgenus Acanthophallus; characters; Italy).

  • — Ciceroni, 1994: 117 (Xantholinus; synonym of tricolor).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; cited as valid name for “tricolor Auct. nec Fabricius, 1787”; Italy).

rhaeticus Bordoni, 1972: 172 (Xantholinus; subgenus Purrolinus; Type locality: Alpi Retiche, Val Livigno, Livigno, Sondrio, 1816 m).

  • — Bordoni, 1982: 261 (Xantholinus; subgenus Purrolinus; characters; Italy).

  • — Ciceroni, 1994: 117 (Xantholinus; synonym of tricolor).

lessinicus Bordoni, 1972: 174 (Xantholinus; subgenus Purrolinus; [Note: This species was originally named baldensis but was changed to lessinicus in the “errata corrige” and the type locality was also changed]; Type locality: Trentino, M.ti Lessini, Sega di Ala).

  • — Ciceroni, 1994: 117 (Xantholinus; synonym of tricolor).

toumayeffi Bordoni, 1986: 68 (Xantholinus; subgenus Purrolinus; Type locality: Giura svizzero, Rouges Terres presso Tramelan, Berna).

  • — Lohse and Lucht, 1989: 159 (Xantholinus; characters).

  • — Ciceroni, 1994: 117 (Xantholinus; synonym of tricolor).

  • — Assing, Frisch, Kahlen, et al., 1998: 136 (Xantholinus; synonym of tricolor).

meyeri Drugmand, 1994: 250 (Xantholinus; proposed for tricolor of authors, the true tricolor being Lithocharis ochracea; Type locality: Luxembourg: Grand-Duché de Luxembourg. Also cited from Germany, France, Belgium, Norway, Italy, Switzerland, Russia, Poland).

  • — Herman, 2001: 60 (Xantholinus; synonym of tricolor).

tripunctatus Cameron, 1951c: 399 (Xantholinus; Type locality: Kenya Colony: Naro Moru).

  • Distribution: Kenya.

trispinus Coiffait, 1962c: 74 (Xantholinus; subgenus Xantholinus; Type locality: Algérie: Philippeville).

  • — Coiffait, 1962b: 398, 403, 407 (Xantholinus; subgenus Xantholinus; characters; Algeria; Tunisia).

  • — Coiffait, 1972: 267 (Xantholinus; subgenus Xantholinus; characters; references; Algeria; Tunisia).

  • — Bordoni, 1972c (Xantholinus; subgenus Paraphallus; correction of subgeneric assignment in Bordoni, 1972: 162, 188).

  • — Bordoni, 1976c: 89 (Xantholinus; Spain).

  • Distribution: Tunisia, Algeria, Spain.

tubulus Sharp, 1889, see: Hypnogyra.

umbilicatus Fauvel, 1905, see: Gauropterus.

umbratus Motschulsky, 1860, see: Nudobius.

ussuricus Tikhomirova, 1973, see: Megalinus.

vafer Normand, 1947a: 7 (Xantholinus; Type locality: Tunisie: Le Kef, montagne du Dyr).

  • — Coiffait, 1972: 309 (Xantholinus; characters; references; Tunisia).

  • Distribution: Tunisia.

vagepunctus Gravenhorst, 1806, see: vagopunctatus Latreille, 1804.

vagopunctatus Latreille, 1804: 301 (Staphylinus; [Note: Nomen dubium]; Type locality: Not cited).

  • — Herman, 2001: 32 (Xantholinus; synonymic note).

  • Distribution: ?.

vagepunctus Gravenhorst, 1806: 88 (Staphylinus; [Note: Gravenhorst described this species using specimens in Latreille's collection but Latreille published the species first]; Type locality: Not cited).

  • — Erichson, 1839b: 318 (Xantholinus; characters).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; cited as vagepunctatus; catalog).

  • — Herman, 2001: 32 (Xantholinus; synonym of vagopunctatus).

vagus Erichson, 1839, see: Eulissus.

validus Petri, 1912, see: procerus Erichson, 1839.

vandalicus Bordoni, 1973f: 78 (Xantholinus; subgenus Polydontophallus; Type locality: Spagna meridionale, Algeciras, Cadice).

  • — Bordoni, 1976c: 93 (Xantholinus; notes; Spain).

  • Distribution: Spain.

varhegyanus Bordoni, 1973c: 152 (Xantholinus; subgenus Meneidophallus; Type locality: Roumanie: Alpes Transylvaniques, Varhegy, Brasov).

  • — Tóth, 1985: 176 (Xantholinus; notes; Hungary).

  • — Tóth, 1989: 30 (Xantholinus; characters; Hungary).

  • Distribution: Romania, Hungary.

variabilis Hochhuth, 1851: 8 (Xantholinus; Type locality: Armenien; Helenendorf).

  • — Hochhuth, 1862: 67 (Xantholinus; Russia).

  • — Reitter, 1908: 121 (Xantholinus; characters; Caucasus).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • — Scheerpeltz, 1933: 1315 (Xantholinus; catalog).

  • — Coiffait, 1972: 298 (Xantholinus; subgenus Helicophallus; characters; Caucasus; Armenia; Alps; Apennines).

  • — Bordoni, 1976d: 122 (Xantholinus; Caucasus).

  • Distribution: Russia, Georgia, Armenia.

varius H. Bauer, 1914, see: Leptacinus.

varnensis Coiffait, 1972: 290 (Xantholinus; subgenus Acanthophallus; Type locality: Staro Oriakovo près de Varna, Bulgarie).

  • — Bordoni, 1999: 2, 5 (Xantholinus; subgenus Typhlolinus; notes).

  • Distribution: Bulgaria, Turkey.

pamphylicus Coiffait, 1972: 291 (Xantholinus; subgenus Acanthophallus; Type locality: Alanya, dim Irmak, Anatolie méridionale).

  • — Bordoni, 1999: 5 (Xantholinus; synonym of varnensis).

vermiculus Normand, 1947a: 8 (Xantholinus; Type locality: Tunisie: Le Kef).

  • — Jung, 1970: 130 (Xantholinus; characters; Tunisia).

  • — Coiffait, 1972: 309 (Xantholinus; characters; Tunisia).

  • Distribution: Tunisia.

vilis Sharp, 1885, see: Neohypnus.

vinicolor Ushakov, 1989: 127 (Xantholinus; subgenus Helicophallus; Type locality: Checheno-Iugushetiia, Karachoi iugo-vostochnee Vedeno, 950 m).

  • Distribution: Russia.

vinsoni Cameron, 1947, see: Pachycorynus.

vitalei Bernhauer, 1943: 74 (Xantholinus; Type locality: Sizilien: Naso).

  • — Coiffait, 1972: 271 (Xantholinus; subgenus Xantholinus; characters; Sicily).

  • — Bordoni, 1972: 160, 167 (Xantholinus; subgenus Echinophallus; characters; Italy).

  • — Bordoni, 1982: 254 (Xantholinus; subgenus Echinophallus; characters; Italy).

  • — Sparacio, 1995: 158 (Xantholinus; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 26 (Xantholinus; Italy).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Idiolinus).

  • Distribution: Italy.

volvulus Erichson, 1839b: 317 (Xantholinus; Type locality: Brasilia).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • Distribution: Brazil.

wagneri Scheerpeltz, 1926, see: Gyrohypnus.

westwoodianus† Heer, 1856: 16 (Xantholinus; Fossil; Type locality: Aix, Provence).

  • — Oustalet, 1874: 151 (Xantholinus; characters; France).

  • — Scudder, 1891: 594 (Xantholinus; catalog; Ligurian; France).

  • — Handlirsch, 1907: 726 (Xantholinus; catalog; Oligocene; France).

  • Distribution: France.

provincialis† Giebel, 1856: 72 (Lathrobium; fossil; Type locality: France: Aix).

  • — Scudder, 1891: 545 (Lathrobium; catalog; Ligurian Horizon; France).

  • — Handlirsch, 1907: 726 (Xantholinus; catalog; synonym of westwoodianus).

wilhelmensis Last, 1980: 156 (Xantholinus; Type locality: New Guinea: Mount Wilhelm, Field Station).

  • Distribution: New Guinea.

wingelmuelleri Bernhauer, 1899, see: Nudobius.

wrasei Bordoni, 1989a: 501 (Xantholinus; subgenus Heterolius; Type locality: Romania orientale: Dobrugia, Mamaia [Constanta]).

  • Distribution: Romania.

wunderlei Bordoni, 1994: 27 (Xantholinus; subgenus Acanthophus; Type locality: Asia Minore: Antalya, Manavgat).

  • — Bordoni, 1999: 2 (Xantholinus; subgenus Typhlolinus).

  • Distribution: Turkey.

xanthogaster Solsky, 1875: 19 (Xantholinus; Type locality: Pérou, Hacienda Amable Maria et près du lac Junin).

  • — Bernhauer and Schubert, 1914: 308 (Xantholinus; catalog).

  • Distribution: Peru.

zalathnae Bordoni, 1973c: 155 (Xantholinus; subgenus Helicophallus; Type locality: Roumanie: Zalathna).

  • — Tóth, 1989: 26 (Xantholinus; characters).

  • Distribution: Romania.

Xestolinus [2 species; Nearctic and Palaearctic Regions]

Xestolinus Casey, 1906: 368, 397 (species included: abdominalis, oviceps). Type species: Xestolinus abdominalis Casey, fixed by subsequent designation by Blackwelder, 1952: 407 [Note: Blackwelder stated that R. Lucas, 1920: 673, designated the type species but the designation was ambiguous because two nominal species were cited as “abdominalis et oviceps”].

  • — Bernhauer and Schubert, 1914: 208 (world catalog; 2 species).

  • — Leng, 1920: 105 (catalog of North American species).

  • — Blackwelder, 1952: 407 (type species).

  • — Hatch, 1957: 234 (characters; species of Pacific Northwest).

  • — Moore, 1963a: 94 (characters).

  • — Moore and Legner, 1975: 93 (catalog; North America north of Mexico).

  • — Moore and Legner, 1979: 77 (characters; notes).

  • — Smetana, 1982: 279 (characters; habitus; revision of species of North America north of Mexico).

  • — Shibata, 1983: 81 (checklist of species of Japan).

  • — Downie and Arnett, 1996: 402 (characters; [Note: Erroneously cited as Stictolinus]).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 392 (1 Nearctic species; notes; characters in key).

abdominalis Casey, 1906: 398 (Xestolinus; Type locality: Arizona, Walnut).

  • — Bernhauer and Schubert, 1914: 308 (Xestolinus; catalog).

  • — Smetana, 1982: 279 (Xestolinus; characters; Manitoba; Ontario; Quebec; Saskatchewan; Arizona; California; Colorado; Massachusetts; Michigan; Montana; New Hampshire; Oregon; Utah).

  • — Campbell and Davies, 1991: 124 (Xestolinus; checklist of species; Alaska and Canada).

  • — Downie and Arnett, 1996: 402 (Xestolinus, cited as Stictolinus [erroneously]; characters; USA).

  • Distribution: Canada, USA.

oviceps Casey, 1906: 398 (Xestolinus; Type locality: southwestern Utah; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 308 (Xestolinus; catalog).

  • — Smetana, 1982: 279 (Xestolinus; lectotype designation; synonym of abdominalis).

indianensis Blatchley, 1910: 396 (Xantholinus; Type locality: Porter County, Indiana; [Note: See lectotype designation by Smetana, 1982]).

  • — Bernhauer and Schubert, 1914: 304 (Xantholinus; catalog).

  • — Smetana, 1982: 279 (Xestolinus; lectotype designation; synonym of abdominalis).

frontalis Hatch, 1957: 235 (Xestolinus; Type locality: Oregon: Golden, near Wold Creek).

  • — Smetana, 1982: 279 (Xestolinus; synonym of abdominalis).

frontalis Hatch, 1957, see: abdominalis Casey, 1906.

indianensis Blatchley, 1910, see: abdominalis Casey, 1906.

oregonus Fender, 1946, see: Xantholinus.

oviceps Casey, 1906, see: abdominalis Casey, 1906.

pauper Sharp, 1889: 250 (Xantholinus; Type locality: Chiuzenji; Nikko).

  • — Bernhauer and Schubert, 1914: 306 (Xantholinus; catalog).

  • — Nakane, 1963a: 90 (Xestolinus; characters; habitus photograph; Japan).

  • — Watanabe, 1996a: 12 (Xestolinus; cave collection; Japan).

  • Distribution: Japan.

Zenon [1 species; Nearctic Region]

Zenon Smetana, 1982: 171 (species included: armifer; habitus). Type species: Zenon armifer Smetana, fixed by original designation and monotypy.

  • — Newton, Thayer, Ashe, and Chandler, 2000: 392 (1 Nearctic species; notes; characters in key).

armifer Smetana, 1982: 172 (Zenon; Type locality: Colorado: Denver).

  • Distribution: USA.

Zeteotomus [29 species; Ethiopian, Palaearctic, Oriental, Australian, and Oceanic Regions]

Zeteotomus Jacquelin du Val, 1856: 33 (species included: brevicornis). Type species: Leptacinus brevicornis Erichson, fixed by monotypy.

  • — Schaum, 1859: 27 (synonym of Metoponcus).

  • — Gemminger and Harold, 1868: 606 (synonym of Metoponcus).

  • — Jacquelin du Val, 1868: 66 (subgenus of Leptacinus; catalog; European species).

  • — Fauvel, 1873b: 93 [= 1873a: 379] (synonym of Metoponcus).

  • — Mulsant and Rey, 1877a: 309 (synonym of Vulda).

  • — Ganglbauer, 1895: 489 (synonym of Metoponcus).

  • — Bernhauer and Schubert, 1914: 290 (synonym of Metoponcus).

  • — Blackwelder, 1952: 408 (type species: brevicornis).

  • — Smetana, 1958: 83 (characters; habitus of one species; type species: brevicornis; Czechoslovakian species).

  • — Székessy, 1963: 48 (characters).

  • — Lohse, 1964: 158 (synonym of Metoponcus).

  • — Coiffait, 1968: 128 (synonym of Metoponcus).

  • — Coiffait, 1972: 131 (synonym of Metoponcus; date of publication).

  • — Tikhomirova, 1973a: 181 (checklist of species of USSR).

  • — Pototskaia, 1976: 170 (larval characters).

  • — Nikitskii, 1976: 182 (larval characters).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 5 (catalog; Poland).

  • — Bordoni, 1982: 160 (synonym of Metoponcus).

  • — Shibata, 1983: 67 (checklist of species of Japan).

  • — Janák, 1993a: 47 (in list of species of Czech Republic and Slovakia).

  • — Ciceroni and Zanetti, 1995: 25 (list of species of Italy).

Metoponcus Kraatz, 1857d: 651 (species included: brevicornis, filarius). Type species: Leptacinus brevicornis Erichson, fixed by monotypy.

  • — Redtenbacher, 1857: 983 (characters).

  • — Schaum, 1859: 27 (catalog; European species).

  • — Gemminger and Harold, 1868: 606 (catalog).

  • — Fauvel, 1873b: 93 [= 1873a: 379] (characters; larval characters).

  • — Fauvel, 1875a: XXV [= 1875b: 227] (catalog).

  • — Redtenbacher, 1874: 220 (characters).

  • — Sharp, 1876: 205 (notes).

  • — Mulsant and Rey, 1877a: 309 (characters; key to French species).

  • — Duvivier, 1883: 159 (catalog).

  • — Sharp, 1885: 503 (characters; Mexican and Central American species).

  • — Olliff, 1887: 477 (characters).

  • — Heyden, Reitter, and Weise, 1891: 105 (list of species of Europe and Caucasus).

  • — Ganglbauer, 1895: 489 (adult and larval characters).

  • — Heyden, Reitter, and Weise, 1906: 159 (list of species of Europe and Caucasus).

  • — Reitter, 1908: 109, 110 (characters; key to species).

  • — Reitter, 1909: 134, 135 (characters).

  • — Bernhauer and Schubert, 1914: 290 (world catalog; 15 species).

  • — Winkler, 1925: 370 (catalog for Palaearctic region).

  • — Porta, 1926: 87 (characters; species of Italy).

  • — Portevin, 1929: 369 (key to French species).

  • — Roubal, 1930: 361 (catalog; Slovakia).

  • — Scheerpeltz, 1933: 1298 (world catalog supplement).

  • — Blackwelder, 1952: 241 (type species: brevicornis).

  • — Coiffait, 1956a: 43 (characters).

  • — Scheerpeltz, 1956b: 8 (key to African species).

  • — Smetana, 1958: 83 (synonym of Zeteotomus).

  • — Lohse, 1964: 158 (characters; key to central European species).

  • — Coiffait, 1968: 128, (characters; notes).

  • — Scheerpeltz, 1968a: 58 (catalog of Austrian species).

  • — Coiffait, 1972: 131 (characters; disputes date of publication; type species: brevicornis; key to species of western Palaearctic region; distribution).

  • — Bordoni, 1982: 160 (characters; habitus; species of Italy).

  • — Hammond, 1984: 206 (cited as valid genus; checklist of species of Borneo).

  • — Tóth, 1989: 6 (characters; key to species of Hungary).

  • — Boháč, 1993: 47 (synonym of Zeteotomus).

Cylindrocephalus Motschulsky, 1860c: 128 (species included: pictus). Type species: Cylindrocephalus pictus Motschulsky, fixed by monotypy.

  • — Gemminger and Harold, 1868: 606 (synonym of Metoponcus).

  • — Fauvel, 1873b: 93 [= 1873a: 379] (synonym of Metoponcus).

  • — Ganglbauer, 1895: 489 (synonym of Metoponcus).

  • — Bernhauer and Schubert, 1914: 290 (synonym of Metoponcus).

  • — Blackwelder, 1952: 114 (type species: pictus).

  • — Smetana, 1958: 83 (synonym of Zeteotomus).

  • — Coiffait, 1968: 128 (synonym of Metoponcus).

  • — Coiffait, 1972: 131 (synonym of Metoponcus).

apiciventris Lea, 1931: 369 (Metoponcus; Type locality: Papua: Mount Lamington).

  • — Scheerpeltz, 1933: 1298 (Metoponcus; catalog).

  • Distribution: New Guinea.

atriceps W. J. MacLeay, 1873: 138 (Xantholinus; Type locality: Gayndah).

  • — Fauvel, 1877: 237 (Metoponcus; characters; Australia).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; catalog).

  • Distribution: Australia.

piceus W. J. MacLeay, 1873: 138 (Xantholinus; Type locality: Gayndah).

  • — Fauvel, 1877: 239 (Metoponcus; characters; evidently a variety of atriceps).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; variety of atriceps).

  • — Lea, 1925: 213 (Metoponcus; variety of atriceps).

basalis Aubé, 1850, see: scripticollis Hochhuth, 1849.

basiflavus Lea, 1925: 214 (Metoponcus; variety of cyaneipennis; Type locality: Kuranda).

  • — Scheerpeltz, 1933: 1298 (Metoponcus; catalog).

  • Distribution: Australia.

basiventris Sharp, 1876, see: Neoxantholinus.

breviceps Lea, 1925: 214 (Metoponcus; Type locality: Victoria: Melbourne).

  • — Scheerpeltz, 1933: 1298 (Metoponcus; catalog).

  • Distribution: Australia.

brevicornis Erichson, 1839b: 334 (Leptacinus; Type locality: Austria).

  • — Redtenbacher, 1849: 693 (Leptacinus; characters; Austria).

  • — Kraatz, 1857d: 652 (Metoponcus; characters; Germany).

  • — Redtenbacher, 1857: 183 (Leptacinus; characters; Austria).

  • — Fauvel, 1873b: 94 [= 1873a: 380] (Metoponcus; characters; France; Austria).

  • — Redtenbacher, 1874: 220 (Metoponcus; characters; Austria).

  • — Seidlitz, 1875: 280 (Metoponcus; characters; Baltic region).

  • — Mulsant and Rey, 1877a: 312 (Vulda; characters; France).

  • — Seidlitz, 1891: 380 (Metoponcus; characters; Europe).

  • — Seidlitz, 1891a: 405 (Metoponcus; characters; Europe).

  • — Ganglbauer, 1895: 492 (Metoponcus; characters; Austria; Germany; France).

  • — Reitter, 1908: 110 (Metoponcus; characters; central Europe; Greece).

  • — Reitter, 1909: 135 (Metoponus; characters; Germany).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; catalog).

  • — Portevin, 1929: 369 (Metoponcus; characters; France).

  • — Scheerpeltz, 1933: 1298 (Metoponcus; catalog).

  • — Coiffait, 1956a: 44 (Metoponcus; characters).

  • — Smetana, 1958: 85 (Zeteotomus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 158 (Metoponcus; characters; central Europe).

  • — Horion, 1965: 85 (Metoponcus; Turkey; Greece; Bosnia; Slovakia; Czech Republic; Poland; Austria; Switzerland; Corsica; Germany).

  • — Coiffait, 1972: 133 (Metoponcus; characters; central Europe; Balkans; Asia Minor; France; Corsica).

  • — Legner and Moore, 1977: 176 (Zeteotomus; found under bark).

  • — Bordoni, 1982: 162 (Metoponcus; characters; Italy).

  • — Bordoni, 1983a: 521 (Metoponcus; discussion of distribution and relationship to Abies alba and Picea orientalis).

  • — Lucht, 1987: 102 (Metoponcus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France).

  • — Tóth, 1989: 6 (Metoponcus; characters; Hungary).

  • — Boháč, 1993: 47 (Zeteotomus; Slovakia; Czech Republic).

  • — Drugmand, 1994a (Metoponcus; characters; Belgium).

  • — Assing and Schülke, 1999: 8 (Metoponcus; notes).

  • Distribution: Europe, Turkey.

cephallenicus Bordoni, 1983a: 522 (Metoponcus; subspecies of brevicornis; Type locality: Cefalonia, Ainos, 1000 m).

brouni Sharp, 1876, see: Neoxantholinus.

cairnsensis Blackburn, 1895, see: Neoxantholinus.

cephallenicus Bordoni, 1983, see: brevicornis Erichson, 1839.

cingulatus Sharp, 1885, see: Neoxantholinus.

congoensis Scheerpeltz, 1956b: 9, 11 (Metoponcus; Type locality: Belgisch-Kongo: Yangambi [see Scheerpeltz, 1956a: 483]. Also cited from French Congo).

  • Distribution: Congo.

cribratus Fauvel, 1878g: 540 (Xantholinus; Type locality: Australie méridionale: Victoria).

  • — Olliff, 1887: 490 (Xantholinus; characters; Australia).

  • — Bernhauer and Schubert, 1914: 301 (Xantholinus; catalog).

  • — Lea, 1925: 213 (Metoponcus; characters; from nest of Ectatomma metallicum; Australia).

  • Distribution: Australia.

enervus Olliff, 1887: 478 (Metoponcus; Type locality: Lottah, Gould's Country, Tasmania).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; catalog).

  • — Lea, 1925: 213 (Metoponcus; synonym of cribratus).

dilatipennis Kirshenblat, 1948: 48 (Metoponcus; Type locality: Primorskii krai: dol. r. Maikhe, Shkotovskii raion; [Note: See lectotype designation by Smetana, 1982a]).

  • — Smetana, 1982a: 310 (Zeteotomus; lectotype designation; characters; Kuril Islands).

  • Distribution: Russia.

enervus Olliff, 1887, see: cribratus Fauvel, 1878.

erythrocephalus Lea, 1927: 275 (Metoponcus; Type locality: Fiji: Viti Levu).

  • — Scheerpeltz, 1933: 1298 (Metoponcus; catalog).

  • Distribution: Fiji.

exiguus Kraatz, 1859, see: Metolinus.

floridanus LeConte, 1880, see: Lithocharodes.

fugitivus Olliff, 1889: 81 (Metoponcus; Type locality: Lord Howe Island: summit of Mount Ledgbird, 2500 feet).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; catalog).

  • Distribution: Lord Howe Island.

fulvipes Broun, 1880, see: Pachycorynus.

gabonicus Coiffait, 1968: 129 (Metoponcus; Type locality: Gabon: Makokou).

  • Distribution: Gabon.

guatemalensis Sharp, 1885, see: Neoxantholinus.

holisoides Sharp, 1876, see: Neoxantholinus.

hoplocephalus Lea, 1927: 274 (Metoponcus; Type locality: Fiji: Viti Levu).

  • — Scheerpeltz, 1933: 1298 (Metoponcus; catalog).

  • Distribution: Fiji.

howensis Lea, 1925, see: Neoxantholinus.

iranicus Coiffait, 1976a: 61 (Metoponcus; Type locality: Iran: Gole Lovae).

  • Distribution: Iran.

leucocnemis Kraatz, 1859, see: Metolinus.

lineatus Schubert, 1909, see: Neoxantholinus.

longiceps Sharp, 1885, see: Neoxantholinus.

maximus Bernhauer, 1907: 384 (Metoponcus; Type locality: Nemoro, Insel Jesso).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; catalog).

  • — Watanabe and Shibata, 1972: 63 (Zeteotomus; Yaku-shima Island).

  • — Smetana, 1982a: 309 (Zeteotomus; characters; Kuril Islands).

  • — J. Li, 1993: 32 (Zeteotomus, cited as Leteotomus; China).

  • — Bordoni, 1997: 167 (Zeteotomus; characters; Japan).

  • Distribution: Japan, China, Russia.

minutus Sharp, 1885, see: Neoxantholinus.

niger Cameron, 1928f: 409 (Metoponcus; Type locality: Mt. Penrissen, 4000 feet).

  • — Scheerpeltz, 1933: 1298 (Metoponcus; catalog).

  • Distribution: Borneo.

nigricollis Bernhauer, 1920 [Note: Nomen nudum], see: Neoxantholinus.

norfolcensis Lea, 1925, see: Neoxantholinus.

philippinus Cameron, 1941b: 445 (Metoponcus; Type locality: Philippines: Palawan: Binaluan).

  • Distribution: Philippines.

piceus W. J. MacLeay, 1873, see: atriceps W. J. MacLeay, 1873.

pictus Motschulsky, 1860, see: scripticollis Hochhuth, 1849.

planatus Lea, 1925: 215 (Metoponcus; Type locality: Queensland: Cairns).

  • — Scheerpeltz, 1933: 1298 (Metoponcus; catalog).

  • Distribution: Australia.

platycephalus Lea, 1927: 275 (Metoponcus; Type locality: Fiji: Viti Levu).

  • — Scheerpeltz, 1933: 1298 (Metoponcus; catalog).

  • Distribution: Fiji.

privus Bordoni, 1997: 169 (Zeteotomus; Type locality: India: Chambaganoor, Madura).

  • Distribution: India.

pulchellus Perroud and Montrousier, 1864: 83 (Othius; Type locality: Nouvelle Calédonie: prise à Kanala).

  • — Fauvel, 1869: 490 (Metoponcus).

  • — Fauvel, 1874c: 435 (Metoponcus; characters; New Caledonia).

  • — Fauvel, 1889b: 260 (Metoponcus; New Caledonia).

  • — Fauvel, 1903d: 267 (Metoponcus; New Caledonia; Iles des Pins).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; catalog).

  • Distribution: New Caledonia.

punctatus Coiffait, 1980b: 478 (Metoponcus; Type locality: Tahiti: région du Lac Vaihiria, 600 m).

  • Distribution: Society Islands.

rufulus Broun, 1880, see: Neoxantholinus.

schedli Scheerpeltz, 1956b: 9 (Metoponcus; Type locality: Belgisch-Kongo: Yangambi).

  • — Scheerpeltz, 1956a: 483 (Metoponcus; citation of type locality]

  • Distribution: Congo.

scripticollis Hochhuth, 1849: 106 (Xantholinus; Type locality: Georgien).

  • — Hochhuth, 1862: 69 (Leptacinus; notes; Georgia; Caucasus).

  • — Reitter, 1908: 110 (Metoponcus; characters; Hungary; Caucasus).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; catalog).

  • — Scheerpeltz, 1933: 1299 (Metoponcus; catalog).

  • — Coiffait, 1956a: 44 (Metoponcus).

  • — Coiffait, 1972: 133 (Metoponcus; characters; references; Caucasus; Hungary).

  • — Tóth, 1989: 7 (Metoponcus; characters; Hungary).

  • Distribution: Georgia, Hungary.

basalis Aubé, 1850: 314 (Leptacinus; Type locality: environs de Batoum, Iméritie).

  • — Ganglbauer, 1895: 492 (Metoponcus; characters; Austria; Germany; France).

  • — Reitter, 1908: 110 (Metoponcus; synonym of scripticollis).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; synonym of scripticollis).

  • — Coiffait, 1972: 133 (Metoponcus; synonym of scripticollis).

pictus Motschulsky, 1860c: 130 (Cylindrocephalus, cited as Cylydrocephalus; Type locality: montagnes de la Kakhétie en Transcaucasie).

  • — Ganglbauer, 1895: 492 (Metoponcus; synonym of basalis).

  • — Reitter, 1908: 110 (Metoponcus; synonym of scripticollis).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; synonym of scripticollis).

  • — Coiffait, 1972: 134 (Metoponcus; synonym of scripticollis).

tricolor Brancsick, 1870: 418 (Metoponcus; Type locality: Kanizsa in Ungarn).

  • — Ganglbauer, 1895: 487 (Metoponcus; synonym of basalis).

  • — Reitter, 1908: 110 (Metoponcus; synonym of scripticollis).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; synonym of scripticollis).

  • — Coiffait, 1972: 134 (Metoponcus; synonym of scripticollis).

semiruber Fauvel, 1877: 238 (Metoponcus; Type locality: Viti Island).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; catalog).

  • — Cameron, 1927c: 263 (Metoponcus; Fiji).

  • Distribution: Fiji Islands.

sinuatus Sharp, 1885, see: Neoxantholinus.

striolatus Scheerpeltz, 1956b: 9 (Metoponcus; Type locality: Belgisch Kongo: Umbeb. Elisabethville).

  • Distribution: Congo.

superbus Bernhauer, 1915k: 298 (Metoponcus; Type locality: Insel Principe: Roca Inf. D. Henrique, 100–300 m).

  • — Scheerpeltz, 1933: 1299 (Metoponcus; catalog).

  • — Scheerpeltz, 1956b: 8 (Metoponcus; characters).

  • Distribution: São Tomé and Príncipe.

tahitiensis Coiffait, 1980b: 477 (Metoponcus; Type locality: Tahiti: Vallée Papenoo, 3 km de la mer).

  • Distribution: Society Islands.

tricolor Brancsick, 1870, see: scripticollis Hochhuth, 1849.

ukerewensis Bernhauer, 1937b: 607 (Metoponcus; Type locality: Ukerewe Insel: Muruntuguru).

  • — Scheerpeltz, 1956b: 9 (Metoponcus; characters).

  • Distribution: Tanzania.

varians Sharp, 1885, see: Neoxantholinus.

variegatus Fauvel, 1874c: 435 (Metoponcus; Type locality: Nouvelle Calédonie).

  • — Fauvel, 1889b: 260 (Metoponcus; New Caledonia).

  • — Fauvel, 1903d: 266 (Metoponcus; New Caledonia).

  • — Bernhauer and Schubert, 1914: 290 (Metoponcus; catalog).

  • Distribution: New Caledonia.

STAPHYLININAE: TRIBE INCERTAE SEDIS Cretoquedius† [1 species; Palaearctic Region]

Cretoquedius† Ryvkin, 1988: 105 (Cretaceous fossil; species included: oculatus). Type species: Cretoquedius oculatus Ryvkin, fixed by original designation and monotypy.

  • — Ryvkin, 1989: 103 (translation of Ryvkin, 1988: 105).

oculatus† Ryvkin, 1988: 106 (Cretoquedius; Upper Cretaceous Fossil; Type locality: Magadanskaia obl., Ten'kinskii r-n, srednee techenie ruch'ia Obeshchaiushchii [pravyi pritok r. Nil]).

  • — Ryvkin, 1989: 104 (Cretoquedius; translation of Ryvkin, 1988: 106; Russia).

  • Distribution: Russia.

Laasbium† [2 species; Nearctic Region]

Laasbium† Scudder, 1900: 49 (species included: agassizii, sectile; [Note: Scudder, 1900 thought this genus resembled Lathrobium but could not find characters to place it in the Paederinae. In his publication the name was placed between Quedius and Leistotrophus, so I have placed it in the Staphylininae. Handlirsch, 1907 cited the genus near the end of his list of fossil staphylinds]). Type species: Laasbium agassizii Scudder, fixed by subsequent designation by Cockerell, 1909: 85.

  • — Blackwelder, 1952: 207 (type species).

  • — Carpenter, 1992: 298 (checklist; similar to Lathrobium; type species designation: agassizii).

agassizii† Scudder, 1900: 49 (Laasbium; fossil; Type locality: Florissant, Colorado).

  • — Handlirsch, 1907: 730 (Laasbium; cited as agassizi; catalog; Miocene; Colorado).

  • Distribution: USA.

sectile† Scudder, 1900: 50 (Laasbium; fossil; Type locality: Florissant, Colorado).

  • — Handlirsch, 1907: 731 (Laasbium; catalog; Miocene; Colorado).

  • Distribution: USA.

Laostaphylinus† [2 species; Palaearctic Region]

Laostaphylinus† Zhang, 1988: 80, 83 (species included: nigritellus, fuscus). Type species: Laostaphylinus nigritellus Zhang, fixed by original designation.

fuscus† Zhang, 1988: 80, 83 (Laostaphylinus; Type locality: Jurassic; third member of Laiyang Formation; Nanligezhuang of Laiyang, Shandong Province, China).

  • Distribution: China.

nigritellus† Zhang, 1988: 80, 83 (Laostaphylinus; Type locality: Jurassic; third member of Laiyang Formation; Nanligezhuang of Laiyang, Shandong Province, China).

  • Distribution: China.

Mesostaphylinus† [2 species; Palaearctic Region]

Mesostaphylinus† Zhang, 1988: 79, 82 (species included: laiyangensis). Type species: Mesostaphylinus laiyangensis Zhang, fixed by original designation and monotypy.

fraternus† Zhang, Wang, and Xu, 1992: 277, 280 (Mesostaphylinus; Type locality: Late Jurassic of Laiyang Formation in Laiyang City, Shandong Province, China).

  • Distribution: China.

laiyangensis† Zhang, 1988: 79, 82 (Mesostaphylinus; Type locality: Jurassic; third member of Laiyang Formation; Nanligezhuang of Laiyang, Shandong Province, China).

  • Distribution: China.

STAPHYLINIDAE: SUBFAMILY INCERTAE SEDIS

Bembicidiodes† [1 species; Palaearctic Region]

Bembicidiodes† Schaufuss, 1888: 267 (Oligocene fossil; species included: inaequicollis; [Note: Compared to Phloeonaeus and Holisus]). Type species: Bembicidiodes inaequicollis Schaufuss, fixed by monotypy.

  • — Spahr, 1981: 94 (references to specimens from amber and copal).

  • — Carpenter, 1992: 298 (in checklist).

inaequicollis† Schaufuss, 1888: 267 (Bembicidiodes; Oligocene Baltic amber fossil; Type locality: Baltic region).

  • Distribution: Baltic region.

Protostaphylinus† [1 species; Palaearctic Region]

Protostaphylinus† Lin, 1976: 114 (Jurassic fossil; species included: mirus). Type species: Protostaphylinus mirus Lin, fixed by monotypy.

  • — Carpenter, 1992: 299 (checklist; Jurassic fossil; little known genus).

mirus† Lin, 1976: 114 (Protostaphylinus; Type locality: China: western Liaoning, Middle and Upper Jurassic sediments).

  • Distribution: China.

Sulcelytrinus† [1 species; Palaearctic Region]

Sulcelytrinus† Tikhomirova, 1968: 153 (Jurassic fossil; cited as Staphylinidae incertae sedis; species included: antiquus). Type species: Sulcelytrinus antiquus Tikhomirova, fixed by original designation and monotypy.

  • — Carpenter, 1992: 299 (cited as Subcelytrinus; checklist; similar to Staphylinus).

antiquus† Tikhomirova, 1968: 153 (Sulcelytrinus; Jurassic fossil; Type locality: Karatau, s. Michailovka).

  • Distribution: Kazhakstan.

Staphylinites† [1 species; Nearctic Region]

Staphylinites† Scudder, 1876: 78 (Oligocene fossil; genus is provisional; [Note: Nomen nudum]).

obsoletum† Scudder, 1876: 78 (Staphylinites; [Note: Nomen nudum] Oligocene fossil; Type locality: Green River).

  • Distribution: USA.

Tunicopterus† [1 species; Palaearctic Region]

Tunicopterus† Tikhomirova, 1968: 153 (Jurassic fossil; cited as Staphylinidae incertae sedis; species included: sigara). Type species: Tunicopterus sigara Tikhomirova, fixed by original designation.

  • — Carpenter, 1992: 299 (checklist; Jurassic fossil; similar to Staphylinus).

sigara† Tikhomirova, 1968: 154 (Tunicopterus; Type locality: Karatau, s. Michailovka).

  • Distribution: Kazakhstan.

PROTACTINAE† [2 species; Palaearctic Region]

Protactiden† Heer, 1847: 28 (genera included: Protactus). Type genus: Protactus, fixed by monotypy.

Protactidae†:

  • — Giebel, 1856: 75 (cited with Protactus).

Protactus† [2 species; Palaearctic Region]

Protactus† Heer, 1847: 28 (Miocene fossil; species included: erichsonii). Type species: Protactus erichsonii Heer, fixed by monotypy.

  • — Giebel, 1856: 75 (fossil species).

  • — Blackwelder, 1952: 323 (type species: erichsonii).

  • — Carpenter, 1992: 299 (checklist; family position uncertain).

erichsonii† Heer, 1847: 28 (Protactus; Miocene fossil; Type locality: Oeningen).

  • — Giebel, 1856: 75 (Protactus; characters; fossil species).

  • — Scudder, 1891: 570 (Protactus; catalog; Tortonian; Baden).

  • — Handlirsch, 1907: 730 (Protactus; catalog; Miocene; Germany).

  • Distribution: Germany.

minor† Heer, 1862: 45 (Protactus; Miocene fossil; Type locality: Oeningen).

  • — Scudder, 1891: 570 (Protactus; catalog; Tortonian; Baden).

  • Distribution: Germany.

Copyright © American Museum of Natural History 2001
Lee H. Herman "Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium.
VI. Staphylinine Group (Part 3)
Staphylininae: Staphylinini (Quediina, Staphylinina, Tanygnathinina, Xanthopygina), Xantholinini
Staphylinidae Incertae Sedis
Fossils, Protactinae†," Bulletin of the American Museum of Natural History 2001(265), 3021-3840, (18 July 2001). https://doi.org/10.1206/0003-0090.265.1.6
Published: 18 July 2001
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