Price, M. J. & E. Maier (2011). Catalogue of the bryophyte types in G: the Grimmiaceae. Candollea 66: 285–309. In English, English and French abstracts.
In the bryophyte collection in G a total of 138 types have been located for 97 specific and infraspecific names either described under Coscinodon Spreng., Grimmia Hedw., Racomitrium Brid., Schistidium Brid., or later transferred into one of these genera. These represent 63 currently recognised taxa, 58 from the Grimmiaceae, two that are now in the Pottiaceae and three that are in the Orthotrichaceae, Ptychomitriaceae and Seligeriaceae. Of the 138 types identified 33 are holotypes or lectotypes, 75 are isotypes or isolectotypes, 19 are syntypes or paratypes, 1 is a neotype and 10 are of uncertain designation. Four original specimens for three nomina nuda were also located. A previously un-documented original Hedwig specimen for Trichostomum heterostichum Hedw. was found recently and details on it are given herein. Newly identified potential type specimens from the Nees von Esenbeck herbarium, for 11 names described in Bryologia Germanica, are included. A list of problematic material and types that are cited in the literature as present in G but that were not located in the herbarium has been compiled. This catalogue includes the basionym and current name, protologue type citation, specimen label information, G barcode and typification information for each of the 154 specimens treated.
Introduction
There are an estimated 5000 specimens of Grimmiaceae in G originating from Switzerland, Europe and North America as well as from other parts of the world such as Argentina, Australia, Bhutan, Bulgaria, Chile, Ethiopia, Kenya, Peru, Nepal, Syria, Turkey and Uganda. Recent taxonomic work by Maier (2010) on Grimmia Hedw., which incorporated the G collections, revealed that there were more Grimmia types than expected within the G collection. Cataloguing efforts were therefore extended to include Coscinodon Spreng., Racomitrium Brid. and Schistidium Brid. A total of 138 types for 97 specific or infraspecific names, either described under Coscinodon, Grimmia, Racomitrium, Schistidium or later transferred into one of these genera are now known to be present in G. These 97 names represent 63 currently recognised taxa. For all types basionyms and their current synonyms are listed in Appendix 1.
During the course of this work an original Hedwig herbarium sheet for Trichostomum heterostichum Hedw., a name that had previously been lectotypified with non-Hedwig material, was located (Fig. 1). Previously overlooked material from the Nees von Esenbeck herbarium in G was also discovered, representing potential types for 11 names described by Nees and Hornschuch in Bryologia Germanica of Nees & al. (1827): Grimmia affinis var. macrocarpa Nees & Hornsch., G. affinis var. ramosissima Nees & Hornsch., G. apocarpa f. atrofusca Nees & Hornsch., G. apocarpa var. bicolor Nees & Hornsch., G. apocarpa var. longifolia Nees & Hornsch., G. apocarpa var. nana Nees & Hornsch., G. apocarpa var. nigroviridis Nees & Hornsch., G. apocarpa var. tenerrima Nees & Hornsch., G. cylindrica Nees & Hornsch., G. fusca Nees & Hornsch., and G. urceolaris Nees & Hornsch.
This catalogue presents information on the types deposited in G of Coscinodon, Grimmia, Racomitrium, Schistidium, or types that were described under these names but later transferred to genera in other families. The aim of this work is to make information on these specimens more accessible and contribute to the long-term conservation of the G bryophyte collection through the cataloguing, and eventual digitilization, of type specimens. This project revealed that G contains, from the 138 catalogued Grimmiaceae types, 33 holotypes or lectotypes, 75 isotypes or isolectotypes, 19 syntypes or paratypes, 1 neotype and 10 types that were of uncertain designation as well as four original specimens for three nomina nuda.
Catalogue of Grimmiaceae types in G
The names associated with the G Grimmiaceae types are listed here alphabetically by basionym. The protologue information is given alongside the label information from each type specimen. The designation of the G types (lecto- = lectotype, holo- = holotype, isolecto- = isolectotype, iso- = isotype, syn-= syntype, para- = paratype) are given and the term “type” is used where the designation is undetermined. Published typifications for names (author, publication year and page number), including the location of the holotype/lectotype, are given. Herbaria acronyms are according to Index Herbariorum. The G specimen barcode is provided as an identifier and the current name used for each basionym is listed (see Appendix 1). Author names are abbreviated according to Brummitt & Powell (1992). Publications are abbreviated according to Crosby (1999) and Taxonomic Literature 2 (Stafleu & Cowan, 1976–1988). Reference works used included the invaluable World Synopsis of the Genus Grimmia of Muñoz & Pando (2000) and a recent treatment of the genus Grimmia by Maier (2010).
Coscinodon humilis Milde in Bot. Zeitung (Berlin) 22, Beil. 3: 13. 1864.
Type citation. — [Süd-Tirol] s.l.c.
= Coscinodon cribrosus (Hedw.) Spruce, fide Ignatov & Afonina (1992: 29).
Label information. — “Grimmia humilis Milde, Passeyr, Milde.” Lecto-: G00114122.
Typification. — Maier & Geissler (1995: 76), lecto-: G.
Comments. — In the introduction by Milde (1864: 3) the following locality information is given: “Oberhalb von Forst an der Etsch, ½ Stunde nördlich von Meran, zieht sich etwa 200 Fuss über dem Niveau der Etsch, am Abhange des Marlinger Berges, in der Richtung von Nordwest nach Südwest eine schmale Wasserleitung hin, der sogenannte Marlinger-Waal.” This locality information is presumed to be applicable to the species that he cites in his work.
Dicranum aciculare Hedw., Sp. Musc. Frond.: 135. 1801.
= Racomitrium aciculare (Hedw.) Brid., Muscol. Recent. Suppl. 4: 80. 1818.
Type citation. — In Helvetia, Gallia, Carniolia, Saxonia, Thuringia, Hercynia, Anglia, Scotia ad saxa fluviorum montanorum et ad rupes humidas.
Label information. — “Dicranum aciculare St. Crypt. III. p.79. tab 33. Bryum aciculare Linn. tenus Poenig ad Muldas, lect.” Lecto-: G00040271. See Price (2005: 57).
Typification. — Bednarek-Ochyra & Ochyra (1994 : 106), lecto-: G.
Dicranum ovale Hedw., Sp. Musc. Frond.: 140. 1801.
≡ Grimmia ovalis (Hedw.) Lindb. in Acta Soc. Sci. Fenn. 10: 75. 1871.
Type citation. — In rupestribus montosis Saxoniae, (Thuringiae prope Isenacum, in granite montis piniferi Franconiae, Austriae prope Engelhardszell).
Label information. — “Grimmia commutata B&S. Specimina ab Hedw. ad iconem Dicrani ovalis adhibita.” Lecto-: G00040349. See Price (2005: 77).
Typification. — Geissler & Maier (1995: 502), lecto-: G.
Dicranum phascoidum P. Beauv., Prodr. Aethéogam.: 54. 1805.
Type citation. — Cette espèce, qui a été découverte, si je ne me trompe, par M. Dufour, se trouve sur un des murs du bois de Boulogne, près Auteuil.
= Grimmia crinita Brid., fide Muñoz & Pando (2000: 24).
Label information. — “Dicranum phascoides P-B.” Lecto-: G00042967.
Typification. — Muñoz (2000a: 289), lecto-: G.
Fissidens pulvinatus Hedw., Sp. Musc. Frond.: 158, tab. 40, fig. 1–3. 1801. ≡ Grimmia pulvinata (Hedw.) Sm. in Sm. & Sowerby, Engl. Bot.: sub tab. 1728. 1807.
Type citation. — In tectis et muris vulgaris, vere maturans.
Label information. — “Bryum pulvinatum α communis antheris subrotundis, pedunculis reflexis, foliis piliferis Linn. Spec. pl. 2. p.1586. n.28. Sp. M. 158. t.40 f. 1–3. Bryum orbiculare pulvinatum, hirsutie canescens, capsulis immersis Dill. H. Musc 395. t.50. f. 65. [b. Grimmia gracilis (illegible word) ab ipso Richardo non differt].” Lecto-: G00040350. See Price (2005: 102).
Typification. — Cao & Vitt (1986: 191), lecto-: G.
Comments. — In Hedwig (1801: 158) the name “Fissidens pulvinatus α communis” is indicated in the margin under the epithet “pulvinatus” but to avoid nomenclatural confusion it is treated as an illegal name by Muñoz & Pando (2000: 58) and the name Fissidens pulvinatus Hedw. is accepted.
Fissidens pulvinatus var. africanus Hedw., Sp. Musc. Frond.: 159, tab. 40, fig. 4–6. 1801.
Type citation. — Ad Cap. B. Spei lecta specimina mihi communicavit O. Swartz.
= Grimmia pulvinata (Hedw.) Sm., fide Magill (1981: 275).
Label information. — “Fissidens pulvinatus β africanus Spec. M. Ad Cap. bonae spei lecta specimina misit O. Swartz.” Lecto-: G00040351. See Price (2005: 103).
Typification. — Muñoz (1999: 168), lecto-: G.
Grimmia aethiopica Müll. Hal. in Bot. Zeitung (Berlin) 13: 765. 1855.
Type citation. — 781 — Schistidium pulvinatum Br. & Sch. in Musc. Abyss. Schimper. Patria Abyssinia, in rupibus montium Simensium, inde a 9000 pedibus supra mare usque ad 13 000 pedes ascendens: W. Schimper 1838, 6 Januar legit.
= Schistidium pulvinatum (Hedw.) Brid., fide Bremer (1980a: 104).
Label information. — “Schimper iter Abyssinicum Sectio secunda. 465. Schistidium pulvinatum Brid. In rupibus montium Simensium indea 9000 pedibus supra mare U. i. 1842. usque ad 13000 pedes. d. 6 Jun. 1838.” Isolecto-: G00114203, G00114204.
Typification. — Muñoz & Pando (2000: 93), lecto-: NY.
Grimmia alpicola Hedw., Sp. Musc. Frond.: 77, tab. 15, fig. 1–5. 1801 [nom. rej.].
Type citation. — Suecia. O. Swartz. In saxis, ad rivulos alpinos.
= Schistidium agassizii Sull. & Lesq., fide Bremer (1980b: 2).
Label information. — “Grimmia alpicola Spec Musc. p. 77. t. 15. f.1–5. accepta ab O. Swartz.” Type: G00040295. See Price (2005: 114).
Typification. — Bremer (1980b: 2), type: G (information communicated by Deguchi).
Comments. — Rejected name (St. Louis Code Article 56, Appendix IV, pg. 396). See discussion in Bremer (1980b: 2–3).
Grimmia anodon var. sinaitica Renauld & Cardot in Bull. Herb. Boissier 2: 33. 1894.
Type citation. — Sinaï, Djebel Senah. Leg. F. Grote, decemb. 12. 1891.
= Grimmia anodon Bruch & Schimp., fide Muñoz & Pando (2000: 16).
Label information. — “F. Grote. Herbar. sinaiticum Djebel Senah. Grimmia anodon var. sinaitica Renauld & Cardot in Bull. Herb. Boiss. II (1894). 33. 12 / XII 1891.” Isolecto-: G00042980.
Typification. — Muñoz & Pando (2000: 16), lecto-: PC.
Grimmia antillarum Thér. in Rev. Bryol. Lichénol. 14: 13. 1944.
Type citation. — Santo-Domingo: Cordillera Central, prov. de Azua, Los Vallecitos de Yaque, 2.500 m. (no. 13.630).
= Grimmia longirostris Hook., fide Muñoz (1998a: 354).
Label information. — “Plantae Indiae Occidentalis No. H 13630. Grimmia antillarum Thér. sp. nov. on stones. det. Thér. Hispaniola. Civ. Santo Domingo: Cordillera Central, prov. de Azua, Los Vallecitos de Yaque, common, c. 2500 m. 2. X. 1929. E.L. Ekman.” Isolecto-: G00042973.
Typification. — Muñoz (1998a: 354), lecto-: PC.
Grimmia apocarpa Hedw., Sp. Musc. Frond.: 76. 1801.
≡ Schistidium apocarpum (Hedw.) Bruch & Schimp., Bryol. Eur. 3: 99. 1845.
Type citation. — Saxa depressiorum, roridorum locorum praeprimis incolit. Lipsiae ad piscinam Lindenthalensem, Gothanam versus Ileburgum, Dresdae ad rivulum Weiseriz ubertim.
Label information. — “Grimmia apocarpa Hedw. St. Cr.” Lecto-: G00040296. See Price (2005: 115).
Typification. — Bremer (1980a: 90), lecto-: G.
Grimmia apocarpa var. grisea Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 91, tab. 16, fig. 4b. 1827.
Type citation. — Wächst auf Felsen bei Berneck auf dem Fichtelgebirge (Funck!, Laurer!).
= Schistidium apocarpum (Hedw.) Bruch & Schimp., fide Bremer (1980a: 92).
Label information. — “Herb. Nees v. Esenb. Grimmia apocarpa α b. grisea.” Type: G00282001.
Typification. — Blom (1996: 218), lecto-: M.
Comments. — This name is not mentioned in Index Muscorum (Wijk & al., 1962). Bremer (1980a: 92) cites a type of this name as being in M (Herb. Funck), a specimen that was later used by Blom (1996) as the lectotype for this name.
Grimmia apocarpa var. nigrescens Molendo in Ber. Naturhist. Vereins Augsburg 18: 146. 1865. ≡ Schistidium apocarpum var. nigrescens (Molendo) Loeske, Laubm. Eur. 1: 29. 1913.
Type citation. — Auf Schieferplatten der a. Region selten: Nebelhorn 6500', Kratzerkoblach 6400', Höfatsplätt 5100'.
Label information. — “Grimmia apoc. nigrescens m. Algäuer Alpen: Auf einem Kalkschieferfels der Südseite des Kratzers im Koblach, 63-6400' 1864. L. Molendo.” Type: G00061667.
Typification. — None found.
Grimmia apocarpa var. pumila Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 111, tab. 18, fig. 4q. 1827.
Type citation. — Beide Formen wachsen auf Felsblöcken der höchsten Alpen, um Heiligenblut (Hsch.), auf dem Wildkrachkogel in der Gastein (Funck!); die var. pumila erhielten wir von Herrn Schleicher aus der Schweiz.
= Schistidium confertum (Funck) Bruch & Schimp., fide Bremer (1980a: 91).
Label information. — “Grimmia apocarpa η tenerrima α pumila. Herb. Nees von Esenb. Herrn Sturm.” Type: G000 48200.
Typification. — Blom (1996: 167), lecto-: M.
Comments. — Bremer (1980a: 91) indicated the presence of a potential type in M for the name “Grimmia apocarpa c. pumila Swartz ex Schleicher nom. nud. 1815. p. 33” which was later used as the lectotype for the name “Grimmia apocarpa var. pumila” by Blom (1996).
Grimmia apocarpa var. stricta Müll. Hal. in Paris, Index Bryol.: 518. 1896 [nom. nud.].
Type citation. — s.l.c.
= Schistidium apocarpum (Hedw.) Bruch & Schimp., fide Bremer (1980a: 94).
Label information. — “Nr. 132. Grimmia apocarpa var. stricta Rehm. Orange Free State, Kadziberg. s.d., Rehmann s.n.” Specimens: G00048186, G00048188.
Grimmia apocaula DC. in Lam. & DC., Fl. Franç. ed 3, 2: 458. 1805.
Type citation. — Elle croît sur les murs et les pierres.
= Grimmia plagiopodia Hedw., fide Blom (1996: 261). Label information. — “Grimmia apocaula Hoffmann [Hedwig].” Lecto-: G00042910.
Typification. — Blom (1996: 261), lecto-: G.
Grimmia apophysata Gangulee in Nova Hedwigia 12: 428, tab. 7. 1967.
Type citation. — Kurz 2344 lectus ad Phalut prope Darjeeling ad alt. 3,500 m. die 16.X.1868. et servatus in CAL.
= T rigonodictyon indicum Dixon & P. de la Varde in Ann. Cryptog. Exot. 1: 40. 1928. See Maier (2010: 354).
Label information. — “2344. Grimmia apophysata Hpe. Index Bryologicus 1894–1898, p. 519. Sikkim 11000' J. Kurz.” Iso-: G00042974.
Typification. — Gangulee (1967: 428), holo-: CAL.
Grimmia arenaria Hampe in Linnaea 10: 405. 1836.
Type citation. — In rupibus et saxis arenosis ad arcis pedem Regenstein, aestate fructu maturo.
= Grimmia donniana Sm., fide Maier (2010: 116). Label information. — “Grimmia arenaria Hpe. ad saxa aren. Regenstein Hercyniae.” Isolecto-: G00042959 ; “Grimmia arenaria m Linnaea X 4. curvula Br. & S. bryol. ad rup arenos. d. Regensteins pr Blankenburg Hercyniae Hampe.” Syn-: G00042960; “No. 1. Grimmia arenaria m trimorphula ad rup arenos. Regenstein Hercyn. Jul. Aug. c.fr. mat 700' (Hampe 34).” Syn-: G00042961; “Grimmia arenaria Hamp. (three illegible words) 43 Regenstein a/H 12 März 1843.” Syn-: G00042962.
Typification. — Muñoz (1998b: 376), lecto-: BM.
Grimmia aspera Müll. Hal. in Nuovo Giorn. Bot. Ital. ser. 2, 4: 261. 1897.
Type citation. — China interior, prov. Schen-si septentr., Hua-san, sterilis, Aug. 1895.
= Grimmia elatior Bals.-Criv. & De Not., fide CAO & Vitt (1986: 201).
Label information. — “Bryotheca E. Levier. Grimmia aspera C. Müll. Nuovo Giornale botanico italiano 1897, p. 261. China interior, provincia Schen-si sept. in monte Hua-san 25 Aug. 1895, legit Rev. Jos. Giraldi, determ. Prof. C. Müller sub. no. 1448.” Isolecto-: G00042956.
Typification. — Cao & Vitt (1986: 201), lecto-: H.
Grimmia benoistii Thér. in Rev. Bryol. Lichénol. 9: 10, fig. 4. 1936.
Type citation. — Massif du Pichincha, rochers verticaux exposés au Nord, face au Condorguachana, 3,900 m. (no. 3116, 3117, 3114 pp) [leg. R. Benoist].
= Grimmia austrofunalis Müll. Hal., fide Muñoz (1999: 127).
Label information. — “Grimmia Benoistii Thér. Equateur. Rochers verticaux exposés au Nord, en face du Condor - guachana (massif du Pichincha), 3900 m Leg. R. Benoist, 30.10.1930. No. 3116. Det. I. Thériot.” Isolecto-: G00042970; ibidem “no. 3117. Det. I. Thériot.” Syn-: G00042971.
Typification. — Muñoz (1999: 127), lecto-: PC.
Grimmia berliniana Schultz in Syll. Pl. Nov. 2: 132. 1828.
Type citation. — In agro Gottingensi?
=Bryoerythrophyllum recurvirostrum (Hedw.) P. C. Chen, fide Wijk & al. (1962: 375).
Label information. — “Grimmia berliniana Schultz (illegible) 1820 Sylloge (illegible). Grimmia striata Schrad.” Type: G00048145.
Typification. — None found.
Grimmia brachyodon Austin in Bull. Torrey Bot. Club 6: 45. 1875.
Type citation. — Pl. masc. ignota - Grimmia montana, var. Sulliv. & Lesqx. Exsic. Ed. 2, n. 215. pro parte.
= Grimmia montana Bruch & Schimp., fide Maier (2010: 224).
Label information. — “215. Grimmia montana Bryol. Europ. Grimm. Monogr. p. 26, t. 14. var. foliis brevius pilifero-acuminatis, operculo peristomiique dentibus brevioribus, etc. Hab: in rupibus schistosis montis Diablo Californiae (Bolander); etiam in alpibus Sierra Nevada montium (Brewer).” Isolecto-: G00042924, G00042925, G00042926.
Typification. — Muñoz (1998b: 398), lecto-: FH.
Grimmia brachypus Müll. Hal. in Nuovo Giorn. Bot. Ital. ser. 2, 4 : 129. 1897. ≡ Racomitrium brachypus (Müll. Hal.) Paris, Index Bryol.: 1070. 1898.
Type citation. — s.l.c.
Label information. — “Rhacomitrium brachypus C. Mull. Bolivia. Prov. Cochabamba, prope Choquecamata, Jun 1889, legit Germain. Det C. Müller sub. No. 1227.” Type: G000 48196.
Typification. — None found.
Comments. — Under the description of this taxon in MÜLLER (1897: 129) no locality or collector information is given, although in the introduction to this work he mentions collections by Germain from Choquecamata in Bryotheca E. Levier, information which corresponds to the specimen label.
Grimmia caffra Müll. Hal. in Hedwigia 38: 118. 1899. ≡ Schistidium caffrum (Müll. Hal.) Ochyra in Fragm. Florist. Geobot. 43: 105. 1998.
Type citation. — Orange Free State, Witteberge supra Kadziberg: Dr. A. Rehmann, 1875 cum Syrrhopodonte erectiolio.
Label information. — “No. 130. Grimmia caffra Rehm. n. sp. Orange Free State: Witteberge supra Kadziberg.” Type: G00114444.
Typification. — None found, types in BM, G, L cited in Bremer (1980a: 95).
Comments. — Under the description of this taxon (Müller, 1899: 118) noted that is was described by Rehmann in his Musci austro-africanis collection under no. 130. The G specimen was annotated “Schistidium apocarpum (Hedw.) Bruch & Schimp.” by Bremer in 1980.
Grimmia cernua f. atra Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 178. 1827.
Type citation. — Wächst in kleinen meist runden, lockeren polsterförmigen Räschen auf Felsblöcken und wurde zuerst auf dem Rathhausberg 1816 aufgefunden (Mielichhofer! Hsch.! Funck! Laurer!); später auch auf Zirknitz-Alpe bei Döllach (Laurer!), und auf andern hohen Alpen von Ober-Kärnthen (Funck! Laurer!).
= Grimmia torquata Drumm., fide Muñoz & Pando (2000: 76).
Label information. — “Grimmia cernua β spiralis α atra Herrn Sturm [Herb. Nees v Esenb.].” Lecto-: G00042921.
Typification. — MUÑOZ & Pando (2000: 76), lecto-: G.
Grimmia cinclidodontea Müll. Hal. in Bot. Centralbl. 44: 387. 1890. ≡ Schistidium cinclidodonteum (Müll. Hal.) B. Bremer in Lindbergia 6: 108. 1980.
Type citation. — Washington: Ellensburgh, ad rupes irroratas prope Thorp, I Junius 1888 c. fr. maturis.
Label information. — “Grimmia cinclidodontea n. sp. C. Müller. 1 Juni 1888. Thorp b/ Ellenburg, Dr. Julius Röll. Washington. No. 940.” Isolecto-: G00053135.
Typification. — Bremer (1980a: 108), lecto-: H-BR, iso-: G. See also Robinson & Hermann (1964: 170–174).
Comments. — One of the G specimens of this taxon was on loan and was thus not seen for this catalogue.
Grimmia cribrodontia Herzog in Repert. Spec. Nov. Regni Veg. 41: 287, tab. 235, fig. d-f. 1937. ≡ Schistidium cribro dontium (Herzog) Ochyra in Acta Bot. Acad. Sci. Hung. 28: 373. 1982.
Type citation. — Africa centralis: Ruwenzori, alpine Stufe, 4650 m, leg I. Hauman VII/VIII, 32 (no. 854).
Label information. — “Exped. Sc. Belge. au Ruwenzori 1952. Grimmia cribodontia [sic] Herz. Type. Ruwenzori, MontStanley 4650- det Herzog, Leg. Hauman VII-1932 – no. 854.” Iso-: G00042968.
Typification. — Muñoz & Pando (2000: 93), holo-: JE.
Grimmia cribrosa Hedw., Sp. Musc. Frond.: 76. 1801. ≡ Coscinodon cribrosus (Hedw.) Spruce in Ann. Mag. Nat. Hist. ser. 2, 3: 491. 1849.
Type citation. — In petris Goslariae, nec non Hassiae montis Meisner elegantem speciem primi dexerunt Persoon et Schrader.
Label information. — “Grimmia cribrosa Hedw. St. Crypt. Vol.III.p.73.t.31. A [a β Hedw. B. a Goslar c. Gogendbrug am Ochsenkopf c. An der Mühle unter der Salbergstraβe Schwarzenbg. (illegible word) e Scotia Arnott 1839].” ? Syn-: G00040083 (see Price, 2005: 116); “Herbier Hedwig. Grimmia cribrosa Hedw. Goslar, June 99.” ? Holo-: G00040082.
Typification. — Deguchi (1978: 244), holo-: G.
Comments. — Price (2005:116) erroneously noted that there was no type designation for this species. The specimen that has been annotated as the holotype in G by Deguchi [G000 40082] is non-original Hedwig material. The original Hedwig material for this name [G00040083] should be re-examined as it is likely that this herbarium sheet contains the true lectotype material for the name Grimmia cribrosa Hedw. (Price & Maier, 2011).
Grimmia dimorphula Müll. Hal. in Nuovo Giorn. Bot. Ital. ser. 2, 3: 108. 1896.
Type citation. — Prope cacumen montis Iu-kia-po ad saxa, Jun.
= Grimmia khasiana Mitt., fide Deguchi (1980: 259). Label information. — “Bryotheca E. Levier. Grimmia (Eugrimmia) dimorphula C. Müll. Nuovo Giornale botanico italiano 1896 p. 108. 1898. p. 189. China interior, provincia Schen-si sept. In monte prope Iu-Kia-po 27 Jun. 1894. legit Rev. Jos. Giraldi. determ. Prof. C. Müller sub. no. 877.” Isolecto-: G00042948, G00042949.
Typification. — Cao & Vitt (1986: 170), lecto-: H.
Grimmia dissimulata E. Maier in Candollea 56: 282, figs. 1, 5: 1–3. 2002.
Type citation. — Flora von Zypern. Grimmia trichophylla Grev. v. meridionalis Schpr. f. rhacomitrioides. Bez. Kyrenia, Kalkfelsen am Pass über Lapithos, an der Strasse bei 400 m, 10.V.1972. F. Koppe 10.5.1972(1).
Label information. — “Flora von Zypern Grimmia trichophylla Grev. v. meridionalis Schpr. f. rhacomitrioides. Bez. Kyrenia. Kalkfelsen am Pass über Lapithos an der Strasse bei 400 m. 10. 5. 1972, Koppe (1).” Iso-: G00042965.
Typification. — Maier (2002a: 282), holo-: STU.
Grimmia elatior var. squarrifolia Dixon & Thér. in Hong Kong Naturalist, Suppl. 2: 10. 1933.
Type citation. — Exposed granite rock, 400–500 ft. alt., Amoy I., 8th July, 1931; coll. G. A. C. Herklots (B.8 A, type). Vertical face of rock, ibidem (B.15 B, B.16 B).
= Grimmia pilifera P. Beauv., fide Deguchi (1978: 201).
Label information. — “Grimmia elatior Bruch var. nov. squarrifolia Dix. & Ther. China, Amoy Island, 450–500 ft. alt., 8 July 1931, G. A. C. Herklots. B.8 A.” Iso-: G00042909.
Typification. — Deguchi (1978: 201), holo-: BM.
Grimmia funalis f. epilifera J. E. Zetterst. in Revis. Grimm. Scand. 74. 1861 [nom. illeg., homon. post.].
Type citation. — Habitat in Norvegia media multis in locis, præcipue copiose in Dovre prope Kongsvold (ubi Kindberg perpauca specimina fructifera legit) et in monte Lauvhöen paroeciæ Lom; in Valders et in Voss eam legit Blytt.
= Grimmia funalis (Schwägr.) Bruch & Schimp., fide Muñoz & Pando (2000: 30).
Label information. — “Grimmia funalis Schwägr. forma epilifera Zetterst. Norwegiae: Zettersted leg. In rupibus juxta Kongsvol. Alpium Dovensium.” Isolecto-: G00061665.
Typification. — MUÑOZ & Pando (2000: 30), lecto-: S.
Grimmia ganderi Limpr. in Jahresber. Schles. Ges. Vaterl. Cult. 61: 215. 1884.
Type citation. —Wurde in Tirol bei Innervillgraten an glatten Schieferwänden am Wege nach «Kalchstein» bei 4000 Fuss vom Pfarrer Hieronymus Gander am 15. März 1884 gesammelt.
= Grimmia donniana Sm., fide Maier (2010: 116). Label information. — “Grimmia ganderi Limpr. n. sp. 61. Jahresbericht der schles. Gesellschaft 1884. Tirol : Innervillgraten an Schieferwänden 15/ 3 1884. Gander.” Isolecto-: G00042963.
Typification. — Muñoz (1998b: 381), lecto-: BP (BP-37547).
Grimmia gracilis Schwägr., Sp. Musc. Frond., Suppl. 1(1): 98, tab. 23. 1811.
Type citation. — Crescit ad lapides siccos montium et alpium Helvetia et in monte pinifero Franconiae auctore Funckio.
= Schistidium strictum (Turner) Mårtensson, fide Cao & Vitt (1986: 214).
Label information. — “14 Grimmia gracilis S. Frequens ad rupes subalpinas Schleicher.” Lecto-: G00048231; isolecto-: G00048233, G00048234.
Typification. — Deguchi (1978: 220), lecto-: G.
Comments. — According to Index Muscorum (Wijk & al., 1962), the name Grimmia gracilis Schleich. ex Schwägr. was replaced by Grimmia apocarpa var. gracilis Röhl., Deutschl. Fl. (ed. 2), Kryptog. Gew. 3: 47. 1813.
Grimmia incurva Schwägr., Sp. Musc. Frond., Suppl. 1(1): 90. 1811.
Type citation. — Legi in summis alpibus Carinthiae, Glockner aliisque similibus primaevis, in Sudetum fossis nivalibus sterilem observavit C. Ludwig.
Label information. — “Grimmia incurva Schwgr. Glockner.” Holo-: G00042984.
Typification. — Deguchi (1978: 190), holo-: G.
Grimmia itatiaiensis Broth. in Bull. Herb. Boissier 6: 108. 1898.
Type citation. — Brasilia, Serra Itatiaia, ad rupes, 2100 m. alta, Martio 1894: E. Ule, Coll. No. 1913.
= Grimmia longirostris Hook., fide Muñoz (1998a: 354).
Label information. — “E. Ule Bryotheca brasiliensis. 124. Grimmia itatiaiensis Broth n. sp. Serra do Itatiaia, 2100 m., ad rupes, m. Martii 1894. leg. E. Ule [1895].” Isolecto-: G00042946, G00042947.
Typification. — Muñoz (1998a: 354), lecto-: H-BR.
Grimmia mammillaris Poech in Lotos 1: 177. 1851.
Type citation. — Gemein bei St. Prokop und Kuchelbad nächst Prag.
= Grimmia orbicularis Wilson, fide Muñoz & Pando (2000: 58).
Label information. — “Musc. boh. num. 29. Grimmia mammilaris [sic] Poech Bemerk. Laub. in Mspt. Hab: In rupibus calcareis prope “Pragam”. Mat: Vere. - Dur: [perennial] Dr. Poech.” Lecto-: G00042931.
Typification. — Muñoz & Pando (2000: 58), lecto-: G. 292 – Candollea 66, 2011
Grimmia meteorae C. C. Towns. in Kit Tan, The Davis & Hedge Festschrift: 48., fig. 3. 1989.
Type citation. — Greece. Thessaly: on damp stony bank near the Varlaam Monastery, the Meteora, near Kalabaka, apparently calcareous, 10 May 1987, C. C. Townsend 87/125. E holotype, Herb. Townsend, isotype.
= Grimmia nutans Bruch, fide Greven (1995: 98).
Label information. — “Grimmia meteorae C.C. Townsend c. fr. Greece: Thessaly; on damp stony bank near the Varlaam Monastery, the Meteora, near Kalabaka, apparently calcareous. 10 May 1987. C.C. Townsend 87/125.” Iso-: G00042932.
Typification. — Townsend (1989: 48), holo-: E.
Grimmia microovata Müll. Hal. in Nuovo Giorn. Bot. Ital. ser. 2, 4: 128. 1897.
Type citation. — Ex habitu Grimmiae ovatae Europaeae, sed humilitate atque teneritate pertium omnium jam refugiens. A. Gr. longirostri Hook. montis Chimborazo humilitate simili foliis longe pilosis longe differt. Gr. Peruviana Sulliv. caule multo graciliore distincte refugit.
= Grimmia longirostris Hook., fide Muñoz (1998a: 354).
Label information. — “Bryotheca E. Levier. Grimmia micro-ovata C. Müll. n.sp. (Nuov. Giorn. bot. ital. 1897). Bolivia, prov. Cochabamba, prope Choquecamata. Jun. 1889 legit. Germain det. C. Müller sub. No. 1142.” Isolecto-: G00042935.
Typification. — Muñoz (1998a: 354), lecto-: JE.
Grimmia montana var. abnoba H. Schmidt in Mitt. Bad. Landesvereins Naturk. Naturschutz Freiburg 2: 121. 1927.
Type citation. — Schauinsland: Gneisfelsen nahe dem höchsten Gipfel und bei den unteren Häusern von Hofsgrund (S).
= Grimmia montana Bruch & Schimp., fide Muñoz (1998b: 390).
Label information. — “Bauer, Musci europ. et amer. exsiccati. 1918. Grimmia montana Br. eur. nova var. abnoba H. Schmidt Frib. Südlicher Schwarzwald: auf sonnigen Felsen nahe dem Gipfel des Schauinsland, um 1200 m s.m. und etwa 400 m tiefer bei Hofsgrund, Nov. 1925, legit Hermann Schmidt.” Isolecto-: G00042933.
Typification. — Muñoz (1998b: 390), lecto-: JE.
Grimmia montana var. idahensis Renauld & Cardot in Bot. Gaz. 30: 18. 1900.
Type citation. — North Idaho: west end of Lake Pend d'Oreille (J. B. Leiberg, 1892. Ren. & Card., Musci Amer. sept. exsicc. no. 289).
= Grimmia montana Bruch & Schimp., fide Maier (2010: 226).
Label information. — “No. 289. Grimmia montana BS var. idahensis Ren. & Card. Idaho: Lac Pend d'Oreille. Leg. J. B. Leiberg, 1892.” Isolecto-: G00046973.
Typification. — Muñoz (1998b: 398), lecto-: PC.
Grimmia nigricans DC. in Lam. & DC., Fl. Franç. ed. 3, 2: 459. 1805.
Type citation. — Cette espèce m'a été envoyée des Alpes voisines du Léman: elle a été trouvée dans les Pyrénées, par le C. Dufour, et dans les montagnes d'Auvergne, par le C. Dubois.
= Grimmia ovalis (Hedw.) Lindb., fide Muñoz & Pando (2000: 60).
Label information. — “Grim-ovata Pyrenées.” Lecto-: G00042927.
Typification. — Muñoz & Pando (2000: 60), lecto-: G.
Grimmia nigroviridis Müll. Hal. in Hedwigia 38: 121. 1899.
Type citation. — Prom. bonae spei, Mte. Tabulari et in montibus supra Worcester, Nov. 1875: Dr. A. Rehmann in Musc. austro-afr. No. 139.
= Racomitrium lamprocarpum (Müll. Hal.) A. Jaeger, fide Ochyra & al. (1988: 240).
Label information. — “139 b. Grimmia nigro-viridis C. M. n. sp. In montibus supra Worcester. [Rehmann].” Isolecto-: G00048210.
Typification. — De Sloover (1977: 170), lecto-: BM.
Grimmia nigroviridis var. robusticula Müll. Hal. in Hedwigia 38: 121. 1899.
Type citation: Devilspeak: Dr. A. Rehmann, Musc. a. a. No. 139 c.
= Racomitrium lamprocarpum (Müll. Hal.) A. Jaeger, fide Ochyra & al. (1988: 240).
Label information. — “139 c Grimmia nigro-viridis C. M. n. sp. Devilspeak. [A. Rehmann].” Iso-: G00048199, G000 48201.
Typification. — Ochyra & al. (1988: 240), holo-: BM.
Grimmia obliqua Hornsch. in Flora 2: 84. 1819.
Type citation: Auf Felsenblöcken in den Kärnthner u. Salzburger Alpen und auf dem Heiligenbl. Tauern.
= Grimmia longirostris Hook., fide Muñoz (1998a: 353).
Label information. — “Grimmia obliqua Hornsch.: caule basi fasciculate, apice ramoso fastigiato, foliis erectopatulis lanceolatis, pilo imposito, capsula alte emersa elliptica operculoque conico-rostellato obliquo. Ex alpp. Salisb. et Carinth. Hornschuch. ” Isolecto-: G00048229.
Typification. — Muñoz (1998a: 353), lecto-: BM.
Grimmia obtusa Schwägr., Sp. Musc. Frond., Suppl. 1(1): 88, tab. 25. 1811 [nom. illeg., non Brid. 1801.].
Type citation. — Legi in alpe Carinthiae Brennkogel, summon monti Glockner vicina, Augusto frutificantem.
= Grimmia donniana Sm., fide Deguchi (1978: 181).
Label information. — “Grimm. obtusa. No 33.” Type ?: G00052458; [sub-packet 1] “Grimmia obtusa (illegible word) Bad Gastein (illegible word) 57.”; [sub-packet 2] “Grimmia obtusa.” Type ?: G00052459.
Typification. — Deguchi (1978: 181), holo-: G.
Comments. — Grimmia obtusa was synonymised with G. donniana by Deguchi (1978: 181), who designated the holotype as being in G, although no information on the specimen label corresponding with this designation was given. Specimen 1 has been determined as G. donniana and G. sessitana and later as G. sessitana. Specimen 2 contains two sub-packets, one with a mixture of G. donniana and G. sessitana while the other contains G. sessitana. Label information given in Muñoz & Pando (2000: 25) “Glockner. Kaulfuss” does not correspond with either of the labels of the specimens in G under this name.
Grimmia obtusolinealis Müll. Hal. in Flora 73: 484. 1890.
Type citation. — Africa or. trop., monte Kilima-Ndscharo, 3000–4800 m: Dr. Hans Meyer 1889.
= Grimmia ovalis (Hedw.) Lindb., fide Maier (2010: 262).
Label information. — “Grimmia obtusolinealis Müll. Hal. isoneotype: Kenya. North Nyeri district: on wet rocks by the Sirimon stream, Mount Kenya, alt. c. 3570 m, 4 April 1975, C. C. Townsend 75/749.” Iso-: G00047359.
Typification. — Greven (2003: 172), neo-: E.
Grimmia ochyriana J. Muñoz in Nova Hedwigia 66: 235, fig. 1–11. 1998.
Type citation. — Nepal, Près du camp de base, direction de Lobuje, 5100 m, 31-V-1964, Zimmermann 558 (G-holotype).
= Grimmia atrata Hornsch., fide Maier (2002b: 164).
Label information. — “Expédition scientifique Genevoise au Népal 1952. No 558 31 V 1952 Grimmia subdoniana Noguchi Loc. Près du camp de Base; direction de Lobuje. Alt. 5100 m. Leg. A. Zimmermann. Det A. Noguchi.” Holo-: G00042972.
Typification. — Muñoz (1998c: 235), holo-: G.
Grimmia olneyi Sull., Musci Hep. U. S. (repr.): 37. 1856.
Type citation. — Rocks, Rhode Island, S. T. Olney.
= Grimmia ovalis (Hedw.) Lindb., fide Maier (2010: 262).
Label information. — “141. Grimmia olneyi. Hab. in rupibus maritimis Rhodensis Insulae (cl. S. T. Olney legit).” Isolecto-: G00053031, G00053039, G00053040.
Typification. — Muñoz & Pando (2000: 58), lecto-: FH.
Grimmia orbicularis var. persica Schiffn. in Oesterr. Bot. Z. 47: 129. 1897.
Type citation. — Persiae australis prov. Faristan [sic]: in rupestribus calcareis montium ad Schiras. 1650 –1700 m s. m. 25. X. 1892. c. fr. [Bornm. Iter Pers. — turc. Nr. 4448, 4448b.]. — Persiae australis prov. Faristan: Persepolis in decliv. apricis ± 1800 m s. m. 3. XI. 1892; c. fr. [Bornm. Iter Pers. — turc. Nr. 4458]. — Persia borealis: ad Teheran in planitie. ± 1200 m s. m. 31. I. 1892; c. fr. [Bornm. Iter Pers-.turc. Nr. 4452].
= Grimmia orbicularis Wilson, fide Muñoz (2000b: 101).
Label information. — “J. Bornmüller: Iter Persico-turcicum 1892-93 No. 4448. Grimmia orbicularis Bruch var. persica Schiffner (var. nov.) Persiae australis prov. Farsistan: in rupestribus calcar. montium ad Schiras. 1650–1700 m.s.m. 1892. X. 25. legit et determ. J. Bornmüller.” Isolecto-: G0004 2928, G00042929, G00042930.
Typification. — Muñoz (2000b: 101), lecto-: JE.
Grimmia ovata var. praecox A. Kern., Fl. Exsicc. Austro-Hungar.: n° 316. 1881 [in sched.].
Type citation. — Tirolia centralis. In saxis schist. ad. Trins in valle Gschnitz; 1200 Monts. m. copiose.
= Grimmia longirostris Hook., fide Muñoz (1998a: 353).
Label information. — “316. Grimmia ovata var. praecox [Austria] Tirolia centralis. In saxis schist. ad Trins in valle Gschnitz; 1200 m s.m. copiose. [A. Kerner, No. 316].” Isolecto-: G00046109, G00046110, G00048225.
Typification. — Muñoz (1998a: 353), lecto-: BM.
Comments. — In TROPICOS the reference given is Laubm. —Fl. Oesterr. —Ung. 167. 1882.
Grimmia pensilvanica Schwägr., Sp. Musc. Frond., Suppl. 1(1): 91. 1811.
Type citation. — Lecta in Pensilvaniae rupibus a rev. Mühlenbergio.
= Grimmia pilifera P. Beauv., fide Maier (2010: 273).
Label information. — “Grimmia Pensilvanica Mhlbg. Februar n. 26.” Lecto-: G00042908; “Grimmia Pensylvanica mihi Arn. 80. rupestris H. fil. e Pensylvania.” Syn-: G00046170.
Typification. — Cao & Vitt (1986: 144), lectotype ammended by Maier (2010: 273), lecto-: G (G00042908).
Grimmia pitardii Corb. in Bull. Soc. Bot. France 56: LVI. 1910. ≡ Campylostelium pitardii (Corb.) E. Maier in Candollea 53: 301. 1998.
Type citation. — Tunisie-Sud: sol calcaire, au milieu des rochers humides de l'oued Jir, dans les Matmata, au S de Gabès; alt. 500–600 m.
Label information. — “[195 Grimmia pitardi Corb.] Plantae Tunetanae. Grimmia pitardii. Corbière . Maturata: Oued Jir, in humidis, II.1907, J. Pitard.” Iso-: G00113988.
Typification. — Maier (1998: 307), holo-: CHE.
Grimmia plagiopodia Hedw., Sp. Musc. Frond.: 78, tab. 15, fig. 6–13. 1801.
Type citation. — In Saxoniae rupibus detexit Cl. Flügge.
Label information. — “Grimmia plagiopus a Hg s. n. b a Flörkio.” Lecto-: G00040348. See Price (2005: 117).
Typification. — Deguchi (1984: 22), holo-: G. Maier (2010: 282), lecto-: G.
Grimmia pulla Cardot in Rev. Bryol. 36: 106. 1909.
Type citation. — Etat de Hidalgo: Honey-Station, sur troncs d'arbres, associé à Zygodon circinatus Sch., 1907 (n. 10423) [C. G. Pringle].
Label information. — “C. G. Pringle. Plantae Mexicanae. State of Hidalgo. 10423 Grimmia pulla, Card n. sp. River ledges, near Honey Station, 7,000 ft. 12, Dec 1907. Det. Cardot.” Isolecto-: G00042911.
Typification. — Muñoz (1999: 167), lecto-: PC.
Grimmia pulvinata var. asphaltita Renauld & Cardot in Bull. Soc. Roy. Bot. Belgique 33: 118. 1895.
Type citation. — Judée: In deserto lacûs Asphaltitae inter Hebron et Zuweirat-el-Faga (Dr. G. E. Post, 1892. Herb. Boissier).
= Grimmia pulvinata (Hedw.) Sm., fide Muñoz & Pando (2000: 69).
Label information. — “417. Plantae Deserti, Maris Mortui Ex Herb. Postian apud Colleg. Syriens. Protest. Entre Hebron et Zuweirat-el-foga, le 22 août, 1892.” Isolecto-: G00042912.
Typification. — Muñoz & Pando (2000: 69), lecto-: PC.
Grimmia recurvata Hedw., Sp. Musc. Frond.: 75. 1801.
≡ Seligeria recurvata (Hedw.) Bruch & Schimp., Bryol. Europ. 2: 12. 1846.
Type citation. — In saxis montis calvariae Clagenfurthi primus detexit rev. Abb. Wulfen. (In Anglia D., in Hercynia Schrader v. Journ. D. Bot.II. P.58
Label information. — “Grimmia recurvata Hedw. St. Cr. 1. 107. t.38. ab Abbate Wulffen.” Holo-: G00040302. See Price (2005: 118).
Typification. — Vitt (1976: 257), holo-: G.
Grimmia robusta Nees & Hornsch., Bryol. Germ. 2(1): 125, tab. 20, fig. 9. 1827.
Type citation. — Wurde von Herrn Funck auf Felsen bei Heiligenblut in Oberkärnthen gesammelt und uns gütigst mitgetheilt.
= Schistidium apocarpum (Hedw.) Bruch & Schimp., fide Bremer (1980a: 93).
Label information. — “Grimmia robusta. Herb. Nees v. Esenb.” Type: G00282003.
Typification. — Possible type cited as being in M by Bremer (1980a: 93).
Grimmia ryanii Bryhn in Nyt Mag. Naturvidensk. 34: 73. 1893.
Type citation. — Grimmia ryani er opdaget i August 1890 af Fabrikbestyrer Ryan ved Leirungsboden paa Sydsiden af Gjendin i Jotunfjeldene, hvor den er samlet, i rigelige Exemplarer af Ryan og Kaurin saavel i 1890 som i 1891.
= Grimmia funalis (Schwägr.) Bruch & Schimp., fide Muñoz & Pando (2000: 30).
Label information. — “Norvegia, Christians AMontc. fr. Grimmia Ryani Limpr. Jotunheimen, prope Leirungsboden in rup. siccis. Alt. metr. 1150. Aug 1890. Chr. Kaurin & E. Ryan.” Isolecto-: G00042955.
Typification. — Muñoz & Pando (2000: 30), lecto-: NY.
Grimmia ryanii f. tenuis J. Amann in Bauer, Musci Eur. & Amer. Exsicc.: n° 2215. 1932 [in sched.].
Type citation. — 2115. Grimmia Ryani Limpr. forma tenuis Am. in sch. Schweiz: Graubünden, an Gneisfelsen vor dem Morteratschgletscher, 1950 m.s.m., mit Grimmia torquata und Holleri, 10. September 1930 legit J. Amann.
= Grimmia funalis (Schwägr.) Bruch & Schimp., fide Muñoz & Pando (2000: 31).
Label information. — “2115. Grimmia Ryani Limpr. forma tenuis Am. in sch. Schweiz: Graubünden, an Gneisfelsen vor dem Morteratschgletscher, 1950 m.s.m., mit Grimmia torquata und Holleri, 10. September 1930 legit. J. Amann.” Isolecto-: G00046113.
Typification. — Muñoz & Pando (2000: 31), lecto-: MO. See Maier (2010: 142) for an explanation of the exsiccatae numbers.
Grimmia sciuroides f. elongata Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 159, tab. 22, fig. 17c. 1827.
Type citation. — Auf den Kärnthner-Alpen, ohne nähere Bezeichnung des Standortes (Laurer!, Hoppe!).
= Grimmia longirostris Hook., fide Muñoz & Pando (2000: 44).
Label information. — “Herb. Nees v Esenb. Grimmia sciuroides δ laxa α elongata.” Lecto-: G00042939.
Typification. — Muñoz & Pando (2000: 44), lecto-: G.
Comments. — The figure is cited in the text (pg. 159) as “e” but is correctly labelled as “c” in the illustration.
Grimmia sciuroides var. foliosa Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 158, tab. 22, fig. 17a. 1827.
Type citation. — In der Gastein (Hsch. 1816), bei Döllach (Funck! Laurer!), auf der Pasterze (Laurer! Hoppe!).
= Grimmia longirostris Hook., fide Muñoz & Pando (2000: 44).
Label information. — “Herb. Nees v Esenb. Grimmia sciuroides β foliosa.” Lecto-: G00042940.
Typification. — Muñoz & Pando (2000: 44), lecto-: G.
Grimmia sciuroides var. laxa Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 159, tab. 22, fig. 17d. 1827.
Type citation. — Im Anlauf- und Kötschachthal in der Gastein (Funck! Laurer!), bei dem Tauernhaus und am Geschlöss in der Windischmattrey (Laurer! Funck!), auf dem Malnitzer-Tauern (Mielichhofer ! Laurer!).
= Grimmia longirostris Hook., fide Muñoz & Pando (2000: 44).
Label information. — “Herb. Nees v Esenb. Grimmia sciuroides δ laxa. ” Lecto-: G00042941.
Typification. — Muñoz & Pando (2000: 44), lecto-: G.
Grimmia sciuroides var. rigida Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 158, tab. 22, fig. 17b. 1827.
Type citation. — Auf den Alpen um Heiligenblut (Hoppe! Funck! Laurer! Hsch.).
= Grimmia longirostris Hook., fide Muñoz & Pando (2000: 45).
Label information. — “Herb. Nees v Esenb. Grimmia sciuroides γ. rigida.” Isolecto-: G00042942.
Typification. — Muñoz & Pando (2000: 45), lecto-: GZU.
Grimmia sessitana De Not. in Atti Reale Univ. Genova 1: 704. 1869.
Type citation. — In praeruptis infra Hospitium della Valdobbia, in valle Sessitana, legit Carestia.
Label information. — “Erbar. Crittogam. Ital. - Ser. II. 55. Grimmia sessitana. Frane alle scaturigini del Vogna, sotto l'ospizio della Valdobbia, in Val Sesia. Carestia.” Isolecto-: G00052457.
Typification. — Cao & Vitt (1986: 164), lecto-: RO.
Grimmia sinensiapocarpa Müll. Hal. in Nuovo Giorn. Bot. Ital. ser. 2, 5: 187. 1898. ≡ Schistidium sinensiapocarpum (Müll. Hal.) Ochyra in Fragm. Florist. Geobot. 43: 107. 1998.
Type citation. — Bryotheca E. Levier. China interior, prov. Schen-si sept., prope locum « Zu-lu», Oct. 1896 c. fr. deoperculatis; quoque forma liliputana simplicicaulis ibidem.
Label information. — “Grimmia (Platystoma) sinensi-apocarpa C. Müll. China interior, provincia Schen-si sept. in loco dicto Zu-lu. 28 Oct. 1896, legit. Rev. Jos. Giraldi, determ. Prof. C. Müller sub N. 2051.” Iso-: G00046111.
Typification. — Muñoz & Pando (2000: 91), iso-: PC.
Grimmia stenopyxis Thér. in Rev. Bryol. Lichénol. 9: 8. 1936.
Type citation. — Base du Pichincha, sur rochers d'andésite, 3.200 m. (No. 3076, 2276) [M. R. Benoist]; Rumipamba, à terre dans les broussailles, 3.000 m (2957) [M. R. Benoist]; Environs de Quito, 2.800 m. (2281) [M. R. Benoist].
= Grimmia longirostris Hook., fide Muñoz (1998a: 354).
Label information. — “Grimmia stenopyxis Thér. Equateur.: sur un rocher, base du Pichincha (3200 m) Leg. R. Benoist, 24. 10. 1930 No. 3076. Det. I. Thériot.” Isolecto-: G00042943; “Grimmia stenopyxis Thér. Equateur.: sur rocher d'andesite, Quito. Leg. R. Benoist, 27 mars 1930 No. 2281. Det. I. Thériot.” Syn-: G00042944.
Typification. — Muñoz (1998a: 354), lecto-: PC.
Grimmia streptophylla Sull. in U.S. Expl. Exped., Musci: 8, tab. 6B. 1859. ≡ Schistidium streptophyllum (Sull.) Herzog in Biblioth. Bot. 87: 52. 1916.
Type citation. — Andes of Peru, near Baños.
Label information. — “Leg. Wilkes, Grimmia.” Iso-: G0011 4436.
Typification. — Bremer (1980a: 113), holo-: FH.
Grimmia subconferta Broth. in Symb. Sin. 4: 45. 1929.
≡ Schistidium subconfertum (Broth.) Deguchi in J. Sci. Hiro shima Univ., Ser. B, Div. 2, Bot. 16: 240. 1978. Type citation. — S.: An nassen Schieferfelsen der Hg. St. auf dem Gonschiga sw von Muli gegen Dschungdien, 4650 m, c. sp. 6. VIII. 1915 (7493).
Label information. — “Nr. 7493. Grimmia subconferta Broth. sp. nov. [China] Prov. Setschwan austro-occid.: In montis Gonschiga a monasterio Muli ad septentr. pagi Yünnanensis Yungning versus Dschungdien siti regione alpina, ad rupes udas. Substr. schistaceo alt. s. m. ca. 4650 m. Leg. 6 VIII. 1915. Dr. Heinr. Frh. v. Handel-Mazzetti (Diar. Nr. 1353).” Iso-: G00046108.
Typification. — Deguchi (1978: 240), holo-: H-BR.
Grimmia subdoniana Nog. in Candollea 19: 175, tab. 2, fig. 1–8. 1964.
Type citation. — Nepal: Moraine, near Lobuje (27°57′, 86°49′), alt. 4870 m, April 2, 1952, Zimmermann 301-type, 304; moraine, near the base camp at Lobuje, alt. 5000 m, April 24, 1952, Zimmermann 325; near the base camp, in the direction of Lobuje, alt. 5100 m, May 31, 1952, Zimmermann 558.
= Grimmia fuscolutea Hook., fide Maier (2010: 144). Label information. — “Grimmia subdoniana Nog. Candollea 19: 175 (1964). Nepal. Moraine près de Lobuje, 4870 m. Apr. 2, 1952, coll A. Zimmermann 301a.” Iso-: G00042950, G00042952, G00042957; “Expédition scientifique Genevoise au Népal 1952. No. 558, 31 V 1952. Grimmia subdoniana Noguchi Loc. Près du camp de base; direction de Lobuje. Alt. 5100 m. Leg. A. Zimmermann.” Para-: G00042951; “Expédition scientifique Genevoise au Népal 1952. No 304. 21 IV 1952 Grimmia subdoniana Noguchi Loc. Moraine, près de Lobuje. Alt. 4870 m. Leg. A. Zimmermann. Det. A. Noguchi.” Para-: G00042953; “Expédition scientifique Genevoise au Népal 1952. No 325 24 IV 1952. Grimmia subdoniana Nog. Loc. Moraine droite du camp de Base à Lobuje. Alt. 5000 m. Leg. A. Zimmermann.” Para-: G00042954.
Typification. — NOGUCHI (1964: 175), holo-: NICH (365224).
Grimmia subovata Müll. Hal. in Nuovo Giorn. Bot. Ital. ser. 2, 4: 128. 1897.
Type citation. —Bolivia, prov. Larecaja, Sorata, Ancouma, supra muros de la Hacienda Peñas, reg. alpina, 3700–4000 m. alta, Aprili 1860: G. Mandon.
= Grimmia longirostris Hook., fide Muñoz (1998a: 354). Label information. — Specimen 1, “G. Mandon. Plantae Andium Boliviensium. No. 1634. Grimmia subovata Schpr. Hab. Prov. Larecaja. Viciniis Sorata, Ancouma, supra muros de la hacienda, Peñas Alt. Reg. subalpina et alpina 3700–4000 m, April 1860.” Isolecto-: G00042936, G00042937, G00042938.
Typification. — Muñoz (1998a: 354), lecto-: BM.
Grimmia subsulcata Limpr., Laubm. Deutschl. 1: 757. 1889 [nom. illeg.]. citation. — s.l.c.
= Grimmia sessitana De Not., see discussion in Maier (2010: 316, 318).
Label information. — “Steiermark: Speiereben bei Schöder. Glimmerschiefer, 2300 m, 9.VIII.1888, J. Breidler s.n.” Iso-: G00052460.
Typification. — Muñoz (1998b), holo-: BP. Muñoz & Pando (2000: 98) treated G. subsulcata as having the same type as G. alpestris var. microstoma Bruch & Schimp.
Grimmia tenera J. E. Zetterst. in Kongl. Svenska Vetenskapsakad. Handl. 13(13): 17. 1876. ≡ Schistidium tenerum (J. E. Zetterst.) Nyholm, Moss Fl. Fennoscandia. II. Musci: 775. 1969.
Type citation. —In fissures rupium prope mare, sat raro ut videtur: Alten, ad pedem montis Kongshavnsfjellet; Talvig, in montibus juxta Talvigfjord praecipue prope amenm Isbärelfven.
Label information. — “Grimmia tenera Zett. nov. spec. Norge, Finmarken, in montibus juxta Talvigfjord. 25/7 1868. J. E. Zetterstedt.” Isolecto-: G00048192.
Typification. — Bremer (1980a: 107), lecto-: S.
Grimmia torngakiana Brassard & Hedd. in Mem. New York Bot. Gard. 45: 216. 1987.
Type citation. — Canada. Labrador: Nachvak Fiord, Minaret Plateau, 58°55′N 64°00′W, 1700 m elevation, turfs on mineral soil in seepage from late snowbeds, 22 Jul 1983, Hedderson 1893.
= Grimmia incurva Schwägr., fide Muñoz & Pando (2000: 27).
Label information. — “Bryophytes of Labrador, Canada, Salermiut Mountain Range, Torngats. Grimmia torngakiana Brassard & Hedderson. Mineral soil in seepage from late snowbeds. Extensive plateau with felsenmeer mantling. Snow beds restricted to N facing gullies. Occasional seepage areas with Racomitrium lanuginosum mats. 1700 m alt. Minuret Plateau. 58°55′N 64°00′W’ Leg. et Det. Terry A. Hedderson 1892. July 22–24 1983.” Para-: G00042983.
Typification. — Brassard & Hedderson (1987: 216), holo-: NFLD (no. 1893).
Grimmia torta Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 179, tab. 24, fig. 24. 1827.
Type citation. — [Austria] Wächst auf Felsblöcken der höchsten Alpen, auf dem Rathhausberg in der Gastein (Laurer!, Funck!, Hrnsch.), auf der Gösnitz-Alpe (Hrsch.), auf dem Naßfelder-Tauern (Funck! Laurer!), in der Nähe des Rauriser Tauernhauses (Laurer!), in den Schweizer Alpen (Fr. Nees v. Esenb.).
= Anoectangium aestivum (Hedw.) Mitt., fide Maier (2010: 332).
Label information. — “Grimmia? torquata mihi ex alpibus” (second label) “Grimmia torquata Herrn Sturm. Herb. Nees v. Esenb.” Lecto-: G00113898.
Typification. — Muñoz & Pando (2000: 76), lecto-: G. See comments in Maier (2010: 332).
Grimmia trichophylla var. vivariensis Husnot, Musci Gallicae 18: n° 861. 1897 [nom. nud.].
Type citation. — (from Exsiccata label) “861. Grimmia trichophylla Grev. var. vivariensis. Sur des pierres volcaniques à Lasbastide. (Ardèche). Mai 1892. Philibert.”
= Grimmia lisae De Not. syn. nov.
Label information. — “861. Grimmia trichophylla Grev. var. vivariensis. Sur des pierres volcaniques à Lasbastide. (Ardèche). Mai 1892. Philibert.” Specimen: G00046107.
Comments. — Muñoz & Pando (2000: 80) indicated that original material was present in FH.
Grimmia zahlbruckneri Garov., Bryol. Austr. Excurs.: 26. 1840.
Type citation. — No specific locality but with «Austrio Impericum» given in the title of the work.
= Grimmia donniana Sm., fide Maier (2010: 116).
Label information. — “Grimmia zahlbruckneri Gav. In Mont-Lineone ad rupes leg. Garov. (Gr. curvula Br. et Sch.).” Lecto-: G00042964.
Typification. — Maier (2010: 116), lecto-: G.
Guembelia abyssinica Müll. Hal., Syn. Musc. Frond. 1: 772. 1849. ≡ Grimmia abyssinica (Müll. Hal.) Mitt. in J. Proc. Linn. Soc., Bot. 7: 152. 1864.
Type citation. — Abyssinia, monte alpestri Silke, 18 Febr. 1840: W. Schimper.
Label information. — “Schimperi iter Abyssinicum, Sectio secunda. 435. Grimmia abyssinica Bruch & Schimp. In siccis ad rupes inde a limitibus arbornm (sic) et fructicum usque ad cacumen montis Silke. d. 18 Febr. 1840 (U. i. 1842)” Isolecto-: G00042975, G00042976, G000 42977, G00042978, G0004 2979.
Typification. — Muñoz & Pando (2000: 32), lecto-: MO.
Guembelia tenella Müll. Hal. in Bot. Centralbl. 44: 388. 1890.
Type citation. — Idaho, Coeur d'Alene (sic), 6 Aug. 1888. Dr. Röll cum fructibus vetustis legit.
= Grimmia montana Bruch & Schimp., fide Jones (1933: 29).
Label information. — “1153. Grimmia tenella C. Müller n.sp. 6 Aug. 1888 Coeur d'Alène Dr. Julius Röll. Idaho.” Lecto-: G00042934.
Typification. — Muñoz (1998b: 390), lecto-: G.
Hedwigia nervosa P. Beauv., Prodr. Aethéogam.: 60. 1805. ≡ Schistidium nervosum (P. Beauv.) Brid., Bryol. Univ. 1: 123. 1826.
Type citation. — Cette mousse m'a été envoyée depuis long-temps par Dickson… Angleterre.
= Schistidium strictum (Turn.) Mårtensson, fide Ochyra (1989: 101).
Label information. — “Hedwigia nervosa P. B. differ de l'a[noectangium]. ciliatum par les feuilles lanceolées marquées d'une côte très sensible, et privées de poil. Envoyée de Londres par M. Dickson.” Holo-: G00114445.
Typification. — Blom (1996: 34), holo-: G.
Comments. — Palisot de Beauvois (1805) described the taxon as follows: “C'est la seule de son genre dont les feuilles soient garnies d'une côte, ou du moins dont la côte soit aussi forte et aussi apparente.” The label citation for the G specimen given in Ochyra (1989: 101), and recopied by Blom (1996: 34), is incorrect (see above for corrected text).
Pterigynandrum ramondii DC. in Lam. & DC., Fl. Franç. ed. 3, 2: 462. 1805. ≡ Grimmia ramondii (DC.) Margad. in Lindbergia 1: 128. 1972.
Type citation. — Cette belle mousse a été découverte dans les Pyrénées, par le C. Ramond, qui le premier a remarqué ses caracterès distinctifs.
Label information. — “Pterigynandrum Ramondii fl. fr. Trichostomum patens ex Arnott 1825. Grimmia Pyrenées Ramond.” Lecto-: G00046112.
Typification. — Muñoz & Pando (2000: 71), lecto-: B (the lectotype specimen was not found in B and Maier (2010: 306, 312) indicated that the lectotype was from G and that the “B” was likely a misprint for G in the previous publication).
Racomitrium cataractarum Brid., Bryol. Univ. 1: 776. 1826.
Type citation. — In Hercynia Suevica superiore ad cataractas rivulosque in saxis perpetuo roridis caespitose habitat. Diligentissimus Braun detexit et communicavit.
= Racomitrium aquaticum (Schrad.) Brid., fide MÜLLER (1849: 800 under “Grimmia aquatica C. Müll.”).
Label information. — “Dryptodon cataractarum Br. S. Cataracta Geroldsau in Silva nigra Braun ad Brid.” Iso-: G001 28741.
Typification. — Bednarek-Ochyra (2006: 111), holo-: B-Bridel.
Racomitrium dicarpum Broth. in Symb. Sin. 4: 47. 1929. Type citation. —NW-Y.: Schieferfelsen der Hg. St. im birm. Mons. unter dem Doker-la in der Mekong-Salwin-Schiedekette, 28°15′, 4225m, c. sp. 17.IX. 1915 (8067) [Handel-Mazzetti].
= Racomitrium himalayanum (Mitt.) A. Jaeger, fide Frisvoll (1988: 191).
Label information. — “Nr. 8067. Rhacomitrium dicarpum Broth. sp. nova. Prov. Yünnan bor. —occid.: Ad confines Tibeticas sub jugo Doker-la, 28°15′, in regionis alpinae rupibus. Substr. Schistacco, alt. s. m. ca. 4225 m. Leg. 17. IX. 1915. Dr. Heinr. Frh. v. Handel-Mazzetti (Diar. Nr. 1499).” Isolecto-: G00048208.
Typification. — Frisvoll (1988: 191), lecto-: H-BR.
Racomitrium elongatum Frisvoll in Gunneria 41: 74, figs. 1: a, 2: e, 5–6, 9: b, 10: a, 12: a, 16: c, 18: a, c-d, 20: b, 23, 26, 38: a-b, 45: c, 49: c, 50. 1983.
Type citation. — Norway. Sør-Trøndelag: Skaun, Djupsjøåsen ca. 500 m SE of the cottage, sloping rock in front of a fen soak, 31. V. 1982. Frisvoll s.n. (TRH).
Label information. —“Racomitrium elongatum Frisvoll. sp. nov. Norvegia: Sør-Trøndelag: Skaun. Kbl. 1521 I. UTM NR 56 15. alt. 360 m. loc. Djupsjøåsen, ca. 500 m SE of cottage. Hab. Sloping rock in front of fen soak. 31. 5. 1982. leg. Arne A. Frisvoll.” Iso-: G00114448.
Typification. — Frisvoll (1983: 74), holo-: TRH.
Racomitrium javanicum var. brachyphyllum Cardot & P. de la Varde in Rev. Bryol. 50: 17. 1923.
Type citation. — [Madeira] no specific locality information given in protologue.
= Racomitrium subsecundum (Hook. & Grev. ex Harv.) Mitt., fide Frisvoll (1988: 208).
Label information. — “No. 30. Rhacomitrium javanicum Dozy + Molk. var. brachyphyllum Card. et P. de la V. v. nova. Loc. Kodaikanal Pacubas Torrent. Alt. 2300 m. Leg. G. F et G. R.” Isolecto-: G00048209.
Typification. — Frisvoll (1988: 208), lecto-: PC.
Comments. — The original description of this taxon by Cardot & Potier de la Varde in Potier de la Varde (1923) does not cite a specific specimen as the type of this name.
Racomitrium javanicum var. incanum Broth. in Symb. Sin. 4: 46. 1929.
Type citation. — NW-Y.: Im birm. Mons. Im Wald unter dem Doker-la an der tibetischen Grenze (8031). S.: Lungdschuschan bei Huili (967). Lose-schan s von Ningyüen, massenhaft (1446). Tschahungnyotscha jenseits des Yalung n von Yenyüen, 28°15′ (2664). Hieher wohl auch die Notzien vom Passe Döko sw Muli und im birm. Mons. Auf sandigen Wiesenmoorboden beim See im Hintergrund des Doyon-lumba in der Mekong-Salwin-Kette, massenhaft. [Handel-Mazzetti].
= Racomitrium subsecundum (Harv.) Mitt., fide Frisvoll (1988: 208).
Label information. — “8031. Rhacomitrium javanicum Dz. et Mbr. var incanum Broth. Prov. Yünnan bor. —occid.: Ad confines Tibeticas sub jugo Doker-la, 28° 15′, in regionis frigide temperatae saxis in silva. Substr. granitico. alt. s. m. ca. 3600–3700 m. Leg. 16. IX. 1915. Dr. Heinr. Frh. v. Handel-Mazzetti. (Diar. Nr. 1491).” Isolecto-: G00048194.
Typification. — Frisvoll (1988: 208), lecto-: H-BR.
Racomitrium subheterostichum A. Jaeger in Ber. Thätigk. St. Gallischen Naturwiss. Ges. 1872/73: 95. 1874 [nom. nud.].
Type citation. — Asia, Sikkim-Himalaya 9–10,000’ (S. Kurz Nr. 2263).
= Racomitrium subsecundum (Harv.) Mitt., fide Frisvoll (1988: 207).
Label information. — “2263. Rhacomitrium subheterostichum CM. Sikkim 9-10000’ S. Kurz.” Specimen: G00 048224.
Schistidium agassizii Sull. & Lesq. in Sull., Musci Hep. U.S. (repr.): 104. 1856.
Type citation. — Wet rocks, northern shore of Lake Superior, Agassiz [Musc. Bor. —Amer., No. 137].
Label information. — “137. Schistidium agassizii (n. sp). Priori affine, densius caespitosum, nigrescente-nitidum; foliis angustioribus longius linguaeformibus apice obtusa dentatis, perichaetio elongato exserto facile distinguendum. A. Sch. maritime foliis obtusis appressis nec crispulis distat. Hab. ad rupes irriguas Lacus Superioris (cl. Agassiz legit).” Isolecto-: G00114197, G00114198, G00114440.
Typification. — Bremer (1980b: 2), lecto-: FH, isolecto-: G.
Schistidium bryhnii I. Hagen in Kongel. Norske Vidensk. Selsk. Skr. (Trondheim) 1897(2): 6. 1898. ≡ Schistidium apocarpum var. bryhnii (I. Hagen) Loeske, Laubm. Eur. 1: 36. 1913.
Type citation. — Habitat ad rupes apricas porphyraceas nec non ad lapides saxaque porphyracea delapsa inordinate aggregata totorum laterum meridionalis occidentaliqsque jugi montis Bragernesåsen oppidi Drammen ab altitudine ca. 30 m. supra mare usque ad cacumen (280 m.), ubi immensa copia crescens fructusque vetustos gerens mense augusti 1894 detexit iterumque 8/5 1896 fructibus aliis operculatis calyptratisque aliis deoperculatis instructum legit. N. Bryhn, qui mecum nomine Schistidii conferti communicavit. Label information. — “Schistidium Bryhnii Hagen – Spec. orig.! Norvegia Buskerudsamt, Bragernesåsen opp. Drammen. — 8/5 1896. leg. Bryhn. comm. Hagen.” Syn-: G00128739, G00114443.
Typification. — Bremer (1980a: 94), lecto-: TRH.
Schistidium spinosum H. H. Blom & Lüth in Lindbergia 27: 122. 2002.
Type citation. — Holotype: Germany. Baden-Württemberg, Black Forest, Yachtal by Elzach, rock quarry facing Schneider-bauernhof, 600 m a.s.l., 14.XII. 1998. leg. M. Lüth. 2368 (STU). Isotypes G, TRH.
Label information. — “Schistidium spinosum Blom & Lüth spec. nov. LINDBERGIA 27: 122–126, Lund 2002. Isotypus! Deutschland, Baden-Württemberg, Schwarzwald. Yachtal bei Elzach, Vorderer Zinken Steinbruch gegenüber Schneiderbauernhof, unterer, stillgelegter Teil offener und sonniger, angesprengter Granitfels TK7814NO, 600m. leg. M. Lüth 14 Dezember 1998.” Iso-: G00114202.
Typification. — Blom & Lüth (2002: 122), holo-: STU.
Trichostomum affine F. Weber & D. Mohr., Bot. Taschenb.: 127. 1807. ≡ Racomitrium affine (F. Weber & D. Mohr) Lindb. in Acta Soc. Sci. Fenn. 10: 552. 1875.
Type citation. — Hae duae forte posthac merid. Germ.
Label information. — “Apotheker in Bex. [III. 18. trichost. affine S] ?dorf am 25 May 1803.” Syn-: G00048244; “Trichostomum affine T. heterostich ex Arnott. Ms. Schleicher.” Syn-: G00048247; “Trichostomum affine S. In valle Servan. T. heterostich ex Arnott.” Syn-: G00048246; “Trichost. affine Schl. 810.” Syn-: G00048245; “18
(Schl.) Lindb. 
In vallée de Servan. Racomitrium 
” Syn-: G00048248.
Typification. — Frisvoll (1984: 302), lecto-: LAU, syn-: G, G-Hedw./Schwaegr., G-DC.
Comments. — In specimen 1, the label has been cut and it is impossible to read what precedes “dorf”. The indication contained in the [ ] is not in the same hand as the label itself.
Trichostomum arcuatum DC. in Lam. & DC., Fl. Franç. ed. 3, 5: 218. 1815.
Type citation. — Il croît sur les pierres et les rochers. Il a été trouvé dans les Alpes par M. Schleicher; dans les Cévennes, par M. Grateloup.
= Grimmia elatior Bals.-Criv. & De Not., fide Maier (2010: 123).
Label information. — “Dicranum arcuatum Schl. Trichostomum patens β Schw. Trichostomum arcuatum DC ms Schleicher 51. P. DC. Dicranum arcuatum Trichost. patens var. ex. Arnott.” Lecto-: G00042958.
Typification. — Maier (2010: 123), lecto-: G (see comments in Maier, 2010: 126).
Trichostomum canadense Michx., Fl. Bor.-Amer. 2: 296. 1803.
Type citation. — Hab: In pinetis Canadae.
= Racomitrium microcarpon (Hedw.) Brid., fide Frisvoll (1984: 308).
Label information. — “Trichostomum canadense Mx. Snz”. Lecto-: G00048230.
Typification. — Frisvoll (1984: 308), lecto-: G.
Trichostomum canescens Hedw., Sp. Musc. Frond.: 111. 1801. ≡ Racomitrium canescens (Hedw.) Brid., Muscol. Recent. Suppl. 4: 78. 1818.
Type citation. — Lata grege inhabitat loca elatiora, aprica, arenosa, siccitate sterilia, maxime montosarum regionum Europae.
Label information. — “Trichostomum canescens (illegible word) 1797'.” Neo-: G00040272. See PRICE (2005: 338).
Typification. — Frisvoll (1983: 108), neo-: G.
Comments. — The designated neotype material originates from the Duby-Boissier herbarium and is composed of a cut out label and some plants that have been mounted directly on to a typical herbarium sheet from that collection. Two original Hedwig herbarium sheets for this taxon were seen by Frisvoll in 1982 [G00048235, G00048236] but were not considered by him in the type designation (Frisvoll, 1983: 108, 111–117). These original specimens should be re-examined.
Trichostomum decipiens Schultz, Prodr. Fl. Starg. Suppl.: 70. 1819. ≡ Grimmia decipiens (Schultz) Lindb. in Hartm., Handb. Skand. Fl. ed. 8: 386. 1861.
Type citation. — Habitat ad saxa granitosa rarius. - ad pagum Gevezien an der Landstrasse. Junio c. operculo.
Label information. — “Trichostomum decipiens (mihi) Tr. pulvinatum subdecumbens fastigiato-ramosum; foliis recurvato-patentibus lanceolatis complicatis acutis, acumine piliformi (breviore); sporangio e seta arcuata (brevi) immerso. Ad saxa granitosa Duc. Megapol. = Stargard Junio caps. operc. ad deoperculat.” Lecto-: G00042966.
Typification. — Geissler & Maier (1995: 499), lecto-: G.
Trichostomum funale Schwägr., Sp. Musc. Frond., Suppl. 1(1): 150. 37. 1811. ≡ Grimmia funalis (Schwägr.) Bruch & Schimp., Bryol. Eur. 25–28, Grimmia: 17. 1845.
Type citation. — Legit in subalpinis Sudeticis C. Ludwig, aestate fructiferum.
Label information. — “Trichostomum funale Suppl 1. 150 t. 37. Um die Schneekoppe Lg ex parte.” Lecto-: G00046106.
Typification. — Geissler & Maier (1995: 500), lecto-: G.
Trichostomum heterostichum Hedw., Sp. Musc. Frond.: 109. 1801. ≡ Racomitrium heterostichum (Hedw.) Brid., Muscol. Recent. Suppl. 4: 79. 1818.
Type citation. — Locis saxosis, lapidosis, montanarum regionum Germaniae.
Label information. — “Trichostomum heterostichum Hedw. Musc. St. Vol. II. t.25.”, “E Herb. Patriae e ducata Megapolitana.” Lecto-: G00040274. See Price (2005: 343). “Bryum hypnoides Linn. ex rupe nach Erwenschlag (?)enet Cheminitz.” Syn-: G00040275.
Typification. — Frisvoll (1984: 305), lecto-: G.
Comments. — The designated lectotype type material originates from the Duby-Boissier herbarium and is composed of three sections that have been cut out and mounted on a typical herbarium sheet from that herbarium. The small element at the bottom “ex. Herb. Patriae e ducata Megapolitana,” contrary to the indications of G. Colomb- Duplan, is in the hand of Hedwig. An original Hedwig sheet is present in G for this taxon, see Fig. 1 [G000 48237]. This herbarium sheet is labelled “Trichostomum heterostichum Hedw. St. Crypt. II. p. 70. t. 25. [b. alopercurum Brid.2 Schx.1 hb.3] versus Tschopa(illegible) in Sax. lectus” which corresponds with the locality information given in Hedwig (1789: 71), where this taxon was first described. Frisvoll (1984: 305) states that the G specimens seen by him are listed by their “reference number” but the numbers given correspond to G loan numbers and have nothing to do with the referencing or identification of specimens in G.
Trichostomum lanuginosum Hedw., Sp. Musc. Frond.: 109. 1801. ≡ Racomitrium lanuginosum (Hedw.) Brid., Muscol. Recent. Suppl. 4: 79. 1818.
Type citation. — Montana, arida, arenosa, lapidosa loca; (maxime in frustis granitis montium subalpinorum, Bructeri, Sudetum.).
Label information. — “Trichostomum lanuginosum Hedw. St. Crypt. Vol. III. p.3. t.2.” Lecto-: G00040358; “Bryum hypnoides α Linn a Dr. Ehrhart.” Syn-: G00040359. See Price (2005: 344).
Typification. — Frisvoll (1986: 83), lecto-: G. Erroneously noted in Price (2005: 344) as “type in G”.
Trichostomum microcarpon Hedw., Sp. Musc. Frond.: 112, tab. 23, fig. 8–12: b-c. 1801. ≡ Racomitrium microcarpum (Hedw.) Brid., Muscol. Recent. Suppl. 4: 79. 1818.
Type citation. — Locus alpestris, praesertim in fissuris rupium Angliae, Helvetiae, Sudetum auf der Schneekoppe (Austriae, Clagenfurthi, in granite Bructeri et Franconiae montis piniferi. S.).
Label information. — “Trichostomum microcarpon Spec. Musc. p.112. t. 23. f. 1–5 e Sudeticum Schnekuppe intrafissures rupium Ludwig hortus.” Lecto-: G00040276. See Price (2005: 345).
Typification. — Frisvoll (1984: 308), lecto-: G.
Trichostomum pulvinatum var. alpestris F. Weber & D. Mohr, Bot. Taschenbuch: 110. 1807. ≡ Grimmia alpestris (F. Weber & D. Mohr) Schleich., Cat. Pl. Helv. ed. 2: 29. 1808.
Type citation. — Grimmia alpestris Schleich. Cent. 2. n. 13 (in monte Sylvia ad rupes a Schleichero lecta) et Dicr. piliferum ej. l. c. 4. n. 12. (in calidiorr. Vallesiae ad saxa inventum….In subalp., in rup. (in summo Bruct. ipsi legimus).
Label information. — “Grimmia alpestris. Schl. donniana W et M. sudetica Schw. 1. p. 82. [a. p. DC.].” “Grimmia alpestris [Schleicher].” “13 Grimmia alpestris S. in M. Sylvio ad rupes. [m. Schleicher.].” Isolecto-: G00042915.
Typification. — Muñoz (1997: 518), lecto-: BM.
Comments. — The isolectotype material cited here was previously used as the lectotype of Grimmia sudetica Schwägr. by Geissler & Maier (1995: 503).
Possible types types for Nees and Hornschuch names described in Bryologica Germanica
The Christian Gottfried Daniel Nees von Esenbeck (1776–1858) herbarium collection was split up after his death. The Nees collections came to G as part of the Duby herbarium that was purchased by William Barbey for the Boissier herbarium (Briquet, 1940). Nees bryophyte specimens may also be found in B (mosses, some of which are preserved in herb. Bridel), E, L, PC (hepatics), S (hepatics) and W (hepatics). Frahm (2009) discusses the life and major bryological works of Nees, including the publication of Bryologia Germanica (see also Margadant, 1968: 188–189).
Grimmia affinis var. macrocarpa Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 145, tab. 21, fig. 13c. 1827.
Type citation. — … die var γ wächst auf verwitterten Urgebirgsfelsen in der Gastein (Funck!).
Label information. — “Grimmia affinis γ macrocarpa Herb. Nees v. Esenb.” Type: G00282011.
Typification. — None found.
Grimmia affinis var. ramosissima Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 144, tab. 21, fig. 13b. 1827.
Type citation. — … die var ß wächst auf den genannten höchsten Alpen um Heiligenblut, aber immer höher als α, z. B. am Fuße der Leiterköpfe auf der Pasterze, bei dem sogenannten Schlaflerloch auf der Leiteralpe, dem Malnitzer-Tauern u.s.w. (Hsch.).
Label information. — “Grimmia affinis ß ramosissima. Herb. Nees v. Esenb.” Type: G00282010.
Typification. — None found.
Grimmia apocarpa f. atrofusca Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 99, tab. 17, fig. 4i. 1827.
Type citation. — Wächst mehr auf hohen Alpen, auf den Voggesen (Nestler!), auf den Schweizergebirgen (Schleicher!, Fr. Nees v. Esenb!), am Fuße des Geisberges bei Salzburg (Laurer!), auf dem Rathhausberg in der Gastein und auf den Alpen um Heiligenblut (Laurer!, Hsch.).
Label information. — “Grimmia apocarpa γ gracilis α atrofusca. Herb. Nees v. Esenb.” Type: G00282012.
Typification. — None found.
Comments. — Described as “Grimmia apocarpa γ gracilis α atrofusca.” It is mentioned in Index Muscorum (Wijk & al., 1962) alongside the later homonym “atrofusca (Schimp.) Husn., Musc. Gall. 123. 1886 (Grimmia, 1876) hom. illeg. [non G. a. subvar. atrofusca Nees & Hornsch., 1827].”
Grimmia apocarpa var. bicolor Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 94, tab. 17, fig. 4f. 1827.
Type citation. — Wächst auf Steinen bei der Kasereck-Capelle auf dem Heiligenbluter-Tauern (Funck!), auf Kalkfelsen auf den Voggesen (Nestler!).
= Schistidium apocarpum (Hedw.) Bruch & Schimp., fide Bremer (1980a: 92).
Label information. — “Herb. Nees v. Esenb. Grimmia apocarpa α (illeg. symbol) bicolor.” Type: G00282004.
Typification. — None found, types not seen by Bremer (1980a: 92).
Comments. — Described as “Grimmia apocarpa α bicolor” in Nees & Hornschuch. Name not in Index Muscorum (Wijk & al., 1962).
Grimmia apocarpa var. longifolia Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 91, tab. 16, fig. 4c. 1827.
Type citation. — Wächst auf den Salzburger-Alpen (Laurer!) bei Erlangen (Maerker!).
= Schistidium apocarpum (Hedw.) Bruch & Schimp., fide Bremer (1980a: 92).
Label information. — “Grimmia apocarpa α longifolia. Herrn Sturm. Herb. Nees v. Esenb.” Type: G00282005.
Typification. — None found, types not seen by Bremer (1980a: 92).
Comments. — Name not in Index Muscorum (Wijk & al., 1962).
Grimmia apocarpa var. nana Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 112, tab. 19, fig. 4s. 1927.
Type citation. — Wächst auf Felsenblöcken, die auf der Alpe Pasterze am Glockner herumliegen (Hsch.).
= Schistidium apocarpum (Hedw.) Bruch & Schimp., fide Bremer (1980a: 92).
Label information. — “Herb. Nees von Esenb. Grimmia apocarpa τ nana.” Type: G00048206.
Typification. — None found, types not seen by Bremer (1980a: 92).
Comments. — Name not in Index Muscorum (Wijk & al., 1962).
Grimmia apocarpa var. nigroviridis Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 111, tab. 19, fig. 4r. 1827.
Type citation. — Wächst auf höchsten Alpen um Heiligenblut, z. B. bei der Salmshöhe am Glockner, auf Gösnitz- und Leiter-Alpe auf verwittertem Glimmerschiefer.
= Schistidium apocarpum (Hedw.) Bruch & Schimp., fide Bremer (1980a: 92).
Label information. — “Herb. Nees von Esenb. Grimmia apocarpa δ nigro-viridis.” Type: G00048204.
Typification. — None found.
Grimmia apocarpa var. tenerrima Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 110, tab. 18, fig. 4p. 1827.
Type citation. — Beide Formen wachsen auf Felsblöcken der höchsten Alpen, um Heiligenblut (Hsch.), auf dem Wildkrachkogel in der Gastein (Funck!); die var. pumila erhielten wir von Herrn Schleicher aus der Schweiz.
= Schistidium apocarpum (Hedw.) Bruch & Schimp., fide Bremer (1980a: 92).
Label information. — “Grimmia apocarpa η tenerrima. Herrn Sturm. Herb. Nees von Esenb.” Type: G00048202.
Typification. — None found.
Grimmia cylindrica Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 161, tab. 23, fig. 18. 1827.
Type citation. — Wächst auf kiesigen und sandigen Stellen im Anlaufthal bei Böckstein in der Gastein (Funck!).
= Grimmia ovalis (Hedw.) Lindb., fide Wijk & al. (1962: 381).
Label information. — “Herb. V. Esenb. Grimmia cylindrica.” Type: G00282009.
Typification. — None found.
Grimmia fusca Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 120, tab. 19, fig. 7. 1827.
Type citation. — Wächst auf Felsblöcken auf den Alpen Salzburgs (Laurer! Hsch.).
= Schistidium apocarpum (Hedw.) Bruch & Schimp., fide Bremer (1980a: 93).
Label information. — “Grimmia fusca. Herb. Nees v. Esenb.” Type: G00282006.
Typification. — None found, type cited as not seen by Bremer (1980a: 93).
Grimmia urceolaris Nees & Hornsch. in Nees & al., Bryol. Germ. 2(1): 121, tab. 19, fig. 8. 1827.
Type citation. — Wächst auf feuchten Kalkfelsen in der Schweiz (Schleicher!).
= Schistidium apocarpum (Hedw.) Bruch & Schimp., fide Bremer (1980a: 93).
Label information. — “Grimmia urceolaris Herb. Nees v. Esenb.” Type: G00282008.
Typification. — None found, type cited as not seen by Bremer (1980a: 93).
Cited types not found in G
Four types that are cited in the literature as being in G were not located in the collection.
Schistidium crassipilum H. H. Blom in Bryophyt. Biblioth. 49: 224, fig. 90. 1996.
Isotype cited as being in G – specimen not found.
Schistidium elegantulum H. H. Blom in Bryophyt. Biblioth. 49: 233, fig. 93. 1996.
Isotype cited as being in G – specimen not found.
Schistidium elegantulum subsp. wilsonii H. H. Blom in Bryophyt. Biblioth. 49: 239, fig. 96. 1996.
Isotype cited as being in G – specimen not found.
Schistidium trichodon var. nutans H. H. Blom in Bryophyt. Biblioth. 49: 70, fig. 13. 1996.
Isotype cited as being in G – specimen not found.
Problematic material
Two Hedwig moss names lack original material in G and have been typified using material from other herbaria.
Gymnostomum pulvinatum Hedw., Sp. Musc. Frond.: 36,.tab. 3, fig. 1–7. 1801.
Type citation. — “Ad pagum Reinhausen, prope Goettingam, legit misitque Hoffmann & Persoon.”
= Schistidium pulvinatum (Hedw.) Brid., Muscol. Recent. Suppl. 4: 21. 1818.
Label information. — Original Hedwig specimen not present in G, see Price (2005: 376).
Typification. — Ochyra (1989: 91), lecto-: LE-Mertens.
Trichostomum fasciculare Hedw., Sp. Musc. Frond.: 110. 1801. ≡ Racomitrium fasciculare (Hedw.) Brid., Muscol. Recent. Suppl. 4: 80. 1818.
Type citation. — “Habitat ad saxa in Bructero, Achtermannshoehe aliisque Hercyniae montibus.”
Label information. — “Trichostomum fasciculare e rupibus Brucht. Schw.” G00040273.
Typification. — Bednarek-Ochyra & Ochyra (2005: 1070), lecto-: [icon] “Dillenius (1741): f. 28 on pl. 47,” epi-: OXF-Dillenius.
Comments. — The specimen in G originates from the Duby-Boissier herbarium and is composed of a cut out label and some plants that have been glued onto a typical herbarium sheet from the Duby-Boissier herbarium. The handwriting on the label is of two origins. The word “fasciculare” has been written by Schwägrichen whereas the rest is by an unknown hand.
Acknowledgements
This project was supported by Conservatoire et Jardin botaniques de la Ville de Genève (G). We would like to thank Paola Emery for her patient and thorough help in locating some of the literature sources needed for this work and Philippe Clerc for translating the French abstract. We would like to thank Patrick Perret for reviewing this work.
