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1Institute of Zoology, Chinese Academy of Sciences, No. 1, Beichen West Road, Chaoyang District, Beijing 100101, China; Graduate University of Chinese Academy of Sciences, No. 19, Yuquan Road, Shijingshan District, Beijing 100049, China. E-mail: chenj 2Institute of Zoology, Chinese Academy of Sciences, No. 1, Beichen West Road, Chaoyang District, Beijing 100101, China. Corresponding author. E-mail: email@example.com
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