http://zoobank.org/8C34C897-2B57-465D-9CB3-280AA3F71AD7

Type species: Chandlerea donaldi sp. nov., here designated (monotypy)

DIAGNOSIS

The members of Chandlerea may be separated from other faronite genera by the following combination of characters: small, body length 1.8 mm (Fig. 1); male antennomere 7 enlarged and subquadrate with round depression (Fig. 1); frontal rostrum prominent and frontal sulcus linear, reaching apex of rostrum (Fig. 4); anterior and posterior frontal foveae present, anterior frontal fovea covered by rostrum (Fig. 4); prosternum with median and lateral procoxal foveae (Fig. 6); mesoventrite with promesocoxal foveae (Fig. 8); abdominal segment VI enlarged, at least twice longer than VII (Fig. 10); only known from northern South Island (Fig. 16: black squares).

DISTRIBUTION

New Zealand.

ETYMOLOGY

This genus is named for a world-renown beetle specialist and one of the most influential specialists of Pselaphinae during modern time, Donald S. Chandler.

REMARKS

Females are unknown.

 http://zoobank.org/ECB8FBC6-476F-4EFE-BD7E-E54D404A90B5

DESCRIPTION OF MALE

Length 1.8 mm. Body reddish brown, maxillary palpi, elytra, and legs paler (Fig. 1). Head. Male head bluntly transverse, widest across eyes (Fig. 4). Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3–5 subquadrate, 6 transverse, 7 enlarged and subquadrate with round depression, 8–10 transverse (Fig. 1). Frontal rostrum prominent and frontal sulcus linear reaching apex of rostrum (Fig. 4). Anterior and posterior frontal foveae present, anterior frontal fovea covered by rostrum (Fig. 4). Posterior frontal sulcus deep and round (Fig. 4). Eyes large and prominent, two–thirds length of temples (eye : temple = 2 : 3) (Fig. 4). Thorax. Prosternum as long as wide, widest at midpoint of prosternum (Fig. 6). Elytra longer than wide (Fig. 1). Hind wings fully developed. Meso- metaventrite trapezoidal, longer than wide (Fig. 8). Abdomen. Abdominal tergite IV with pair of transverse patches of microtrichia reaching middle (Fig. 1). Abdominal segment VI enlarged, at least twice longer than VII (Fig. 10). Abdominal sternite IV with distinct basolateral foveae, lacking on V–VI (Fig. 10). Aedeagus. Median lobe oval and dorsal process transverse (Fig. 12). Phallobase symmetrical and rounded (Fig. 12). Parameres asymmetrical and slender with many apical setae (Fig. 12).

TYPE MATERIAL

HOLOTYPE. NEW ZEALAND: Marlborough (MB): 1♂ (NZAC), “NEW ZEALAND: MB: Pelorus Bridge Scenic Reserve, 35m, 41″ 18.3′ S 173″ 34′ E, 27 xi 2005, mixed broadleaf (incl. Nothofagus spp.)-podocarp forest; FMHD#2005-042, berl., leaf & log litter, A. Newton, A. Solodovnikov & D. Clarke; ANMT site 1155”, “HOLOTYPE Chandlerea donaldi Park and Carlton des. 2013”. PARATYPE (1 male): New Zealand: Nelson (NN): 1♂ (slide-mounted), Dun Mt., 2000' 10-I-1942, E. S. Gourlay (NZAC).

DISTRIBUTION

Marlborough (MB), Nelson (NN) (Fig. 16: black circles).

HABITAT

The holotype was collected by sifting leaf and log litter in broadleaf and podocarp forests.

ETYMOLOGY

This species is named for a world-renowned beetle specialist and one of the most influential specialists of Pselaphinae during modern time, Donald S. Chandler.

 http://zoobank.org/6B361E5E-CF2D-4EDB-89EF-229411AB56E9

Type species: Nunnea johni sp. nov., here designated

DIAGNOSIS

The members of Nunnea may be separated from other faronite genera by the following combination of characters: body length 1.8–2.5 mm (Figs. 2 and 3); frontal rostrum prominent and frontal sulcus linear, reaching eyes (Fig. 5); anterior frontal fovea absent and posterior frontal fovea oval (Fig. 5); prosternum longer than wide with lateral procoxal foveae (Fig. 7); mesocoxae divided by distinctively prominent ventral process (Fig. 9: arrow); mesoventrite with promesocoxal foveae and large lateral mesosternal fovea (Fig. 9); abdominal segment VI at least twice larger than VII (Fig. 11); abdominal sternites IV–VI with distinctly large basolateral foveae (Fig. 11); only known from northern South Island (Fig. 16: black triangles and circles).

DISTRIBUTION

New Zealand.

ETYMOLOGY

This genus is named for the collector of the holotype, John T. Nunn, who also provided many valuable specimens for this study.

REMARKS

The members of this genus do not possess distinct external secondary sexual characters. Male abdominal sternite IX is fragile, and partially concealed by sternite VIII, rendering it simple and reduced in appearance. Females possess a more robust, rectangular abdominal sternite IX bearing a pair of long setae that are usually visible in ventral view. Female genitalia, including spermathecae, apparently are membranous and were not observable after clearing specimens using 10% potassium hydroxide.

 http://zoobank.org/2F02E262-A913-4F40-B52D-46164FF6989D

DIAGNOSIS

This species is separated from other species of Nunnea by the dimensions of antennomere 4, which is longer than wide (Fig. 2) and slender median lobe of genitalia (Fig. 13).

DESCRIPTION

Length 1.8–2.5 mm. Body reddish brown; antenna, elytra, legs, maxillary palpi paler (Fig. 7). Head. Male head round, widest across eyes (Fig. 5). Ventral head convex. Antennomere 2 longer than wide, 3 subquadrate, 4 longer than wide, 5–10 subquadrate (Fig. 5). Eyes prominent, approximately one-half length of temples (eye : temple = 1 : 2) (Fig. 5). Thorax. Prosternum longer than wide, widest at midpoint of prosternum (Fig. 7). Elytra as long as wide (Fig. 2). Hind wings rudimentary as small pads. Meso- metaventrite trapezoidal, longer than wide (Fig. 9). Abdomen. Abdominal tergite IV without patches of microtrichia (Fig. 2). Aedeagus. Median lobe slender and S-shaped with round dorsal process (Fig. 13). Phallobase symmetrical and rounded (Fig. 13). Parameres symmetrical with apical setae (Fig. 13). Female abdominal sternite IX rectangular, longer than wide and bearing pair of long and short setae from posterior margin (Fig. 15).

TYPE MATERIAL

HOLOTYPE. NEW ZEALAND: Nelson (NN): 1♂ (NZAC), “NEW ZEALAND: NN: Kahurangi N.P., Arthur Range, above Flora Saddle, 1000m, 41″11.351′S 172″44.456′E, 28 XI-19 XII 2005, Nothofagus-dominant forest; FMHD#2005-044, flight intercept trap, A. Newton & M. Thayer; ANMT site 1156”, “HOLOTYPE Nunnea johni Park and Carlton des. 2013”. PARATYPEs (n = 19; 8 males, 11 females). New Zealand: Buller (BR): 3♂♂ 2♀♀ (1♂, slide-mounted), Nelson Lakes NP, Mt. Robert, Speargrass Tr, 875 m, 41° 49′ 46″ S, 172° 48′ 31″ E, 30-XI-17-XII-2005, Nothofagus forest, FMHD#2005-061, pitfall trap, A. Newton, M. Thayer, ANMT site 1161 (FMNH); 1♀, Nelson Lakes NP, n slope Mt. Robert, Pinchgut Tr, 950 m, 14-XII-1984-6-I-1985, Nothofagus forest, A. Newton, M. Thayer 707, window trap (FMNH); Nelson (NN): 1♀, same data as holotype (FMNH); 1♂, Kahurangi NP, Cobb Dam Rd, Asbestos Tr, 450 m, 41° 06′ 33″ S, 172° 43′ 17″ E, 29-XI-18-XII-2005, mixed broadleaf (incl. Nothofagus fusca)-podocarp forest, FMHD#2005-111, litter, A. Solodovnikov, D. Clarke et. al, ANMT site 1160 (FMNH); 3♂♂ 2♀♀, Dun Mt., 31-VI-1996, A. K. Walker, litter 66/274 (NZAC); 1♂, Upper Maitai, 13-II-1957, E. S. Gourlay (NZAC); 1♀, Cobb Reservoir, 1037 m, 18-IX-1964, T. G. Wood, moss 64/100 (NZAC); Marlborough Sounds (SD): 3♀♀, Tennyson Inlet, west side Te Mako Bay, 125 m, 15-XII-1984-5-I-1985, Nothofagus-podo-hdwd, A. Newton, M. Thayer 710, FIT&window trap (FMNH); 1♀, 70 km ne Nelson, Tennyson Inlet, 480 m, 27-V-1982, FMHD#2005-604, Beech forest litter, S. Peck (FMNH).

DISTRIBUTION

Buller (BR), Nelson (NN), Marlborough Sounds (SD) (Fig. 16: black squares).

HABITAT

Specimens of this species were collected using pitfall, flight intercept, window traps, or by sifting beech and leaf litter in broadleaf, hardwood, podocarp or Nothofagus forests.

ETYMOLOGY

This species is named for the collector of the holotype, John T. Nunn, who also provided many valuable specimens for this study.

 http://zoobank.org/5C1DCB25-1702-462C-BE00-0CFF8C211D74

DIAGNOSIS

This species is separated from other species of this genus by the dimensions of antennomere 4, which are subquadrate (Fig. 3) and broader median lobe of genitalia (Fig. 14).

DESCRIPTION

Length 1.8–2.5 mm. Body reddish brown; antennae, elytra, legs, and maxillary palpi paler (Fig. 3). Head. Male head round, widest across eyes. Ventral head convex. Antennomere 2 longer than wide, 3–10 subquadrate (Fig. 3). Eyes prominent, approximately one-half length of temples (eye : temple = 1 : 2). Thorax. Prosternum longer than wide, widest at midpoint of prosternum. Elytra as long as wide (Fig. 3). Hind wings rudimentary as small pads. Meso- metaventrite trapezoidal, longer than wide. Abdomen. Abdominal tergite IV without patches of microtrichia (Fig. 3). Aedeagus. Median lobe broad and semicircular with round dorsal process (Fig. 14). Phallobase symmetrical and rounded (Fig. 14). Parameres symmetrical apical setae (Fig. 14). Female abdominal sternite IX rectangular, longer than wide and bearing pair of setae from posterior margin.

TYPE MATERIAL

HOLOTYPE. NEW ZEALAND: Nelson (NN): 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “New Zealand: NN: Devil River Rd, Tawhai SF 3km S of Reefton 17 IV 1972, 197m J. McBurney, litter, PB15”, “HOLOTYPE Nunnea kuscheli Park and Carlton des. 2013”. PARATYPES (n = 8; 4 males, 4 females). New Zealand: Buller (BR): 2♀♀, Fletchers Ck, Stoney Ck, 28-I-1972, J. S. Dugdale, PN15, litter 72/101 (NZAC); 1♂ 1♀, Fletchers Ck, 7-III-1972, J. S. Dugdale, litter 72/106 (NZAC); 2♂♂, Reefton, 12-IV-1977, J. A. Wightman, pit trap cutover pine (NZAC); 1♂, W Inangahua SF, 126 m, Fletchers Ck, 18-IV-1972, J. S. Dugdale, moss & litter on Beech forest floor, XB2 (NZAC); 1♀, 1.8 km n Punakaiki, 80 m, 19-XII-1984-20-I-1985, hardwood forest with nikau, A. Newton, M. Thayer 718, FIT & window trap (FMNH).

DISTRIBUTION

Buller (BR), Nelson (NN) (Fig. 16: black triangles).

HABITAT

Specimens of this species were collected using pitfall, flight intercept, window traps, or by sifting moss and leaf litter.

ETYMOLOGY

This species is named for one of the most influential New Zealand beetle specialists, Guillermo Kuschel.

The key is based on male specimens because female specimens are indistinguishable based on external morphology, and a female specimen of Chandlerea gen. nov. is unknown.

1.— Male antennomere 7 enlarged (Fig. 1); elytra longer than wide (Fig. 1) Chandlerea donaldi Park & Carlton sp. nov.

1′.— Antennomere 7 of both sexes slightly larger than 6 (Figs. 2–3); elytra as long as wide (Figs. 2 and 3) 2

2. (1) Antennomere 4 longer than wide; median lobe of genitalia slender and S-shaped (Fig. 13) Nunnea johni Park & Carlton sp. nov.

2′.— Antennomere 4 subquadrate; median lobe of genitalia broad and semicircular (Fig. 14) N. kuscheli Park & Carlton sp. nov.