Attagenus pseudorobustior n. sp. from Namibia is described, illustrated and compared with the similar species Attagenus robustior (Pic, 1951). A list of all Attagenus species so far recorded from Namibia is given.
Currently the dermestid genus Attagenus Latreille, 1802 contains more than 200 known species and subspecies worldwide (Háva 2015), the majority of them belonging into the subgenus Attagenus (s. str.). The present paper considers the preceding taxonomical publications on the species of Attagenus from the Afrotropical region (e. g. Thunberg 1795, 1815; Reitter 1881; Péringuey 1885, 1886; Kalík 1955; Kalík & Háva 2005; Háva 2003a, 2003b; Herrmann & Háva 2014). During the determination of some material from South Africa deposited in the collection of the SMNS the authors found a new species of the genus Attagenus which already was recognised by the deceased coleopterist Vladimir Kalík (1918–2010) as being new to science (see Remarks). This new species is described below.
Acronyms of type depositories
Private collection of Andreas Herrmann, Stade, Germany
Private Entomological Laboratory and Collection Jiří Háva, Prague, Czech Republic
Staatliches Museum für Naturkunde, Stuttgart, Germany
2 Attagenus pseudorobustior n. sp.
Nomen nudum: Attagenus roeri Kalík: Schawaller 1994: 27 (see Remarks).
Holotype (♂): “SWA / Namibia, Nyanga, 14.–22.I.1985, leg. H. Roer”.
Paratypes (not sexed): 39 specimens, same data as holo type.
All specimens of the newly described species are provided with the following labels: “HOLOTYPE” [or “PARATYPE” respectively]”, “Attagenus pseudorobustior sp. n., A. Herrmann & J. Háva det. 2015”.
The holotype is stored in SMNS, paratypes are deposited in SMNS, AHEC and JHAC.
The name of the new species refers to its similarity to Attagenus robustior (Pic, 1951).
Holotype (♂): Body longish ovate, slightly arched. Measurements: TL 3.5 mm, EW 2.1 mm. Head slightly broader than long, sparsely and finely punctate, densely covered with long grey, decumbent hairs; frons flattened and shiny; palpi light brown; Ocellus present on front. Eyes large with hardly visible microsetae. Antennae 11-segmented; antennal club consisting of 3 antennomeres, combined shorter than the shaft (Fig. 2); terminal segment nearly as long as the preceding two segments combined; antennal club brown to darkish brown, shaft light brown. Pronotum twice as broad as long, arcuately narrowed anteriorly, finely and sparsely punctate (like head), cuticula shiny, entirely covered by uniform long grey, decumbent hairs; anterior angles rounded, not visible from above; posterior angles acute with rounded edge, middle of the base convexly extended. Scutellum small and triangular. Punctation and pubescence of the elytra similar to that of head and pronotum, but punctation somewhat more rough and dense; cuticle reddish brown with several indistinct black spots and fasciae, hardly visible because of the dense pubescence as shown in Fig. 1. Legs brown with long and coarse yellow brown pubescence; tarsi roughly as long as tibiae, the latter with strong spines on the exterior edge arranged in two irregular lines. Mesosternum, metasternum and all visible abdominal sternites with the same kind of punctation and pubescence as in elytra. Male genitalia as shown in Fig. 3.
Female: Habitually quite similar to male, slightly more elongated. The antennal club somewhat smaller (as usual in this genus), the spots and fasciae much more distinct than in male, the hairs located on the black elytral parts are darkened (Fig. 4).
Variability: Measurements of paratypes: TL 2.9–3.8 mm, EW 1.8–2.3 mm. Dimension, shape and position of spots and fascia on the elytra are slightly variable.
The new species strongly resembles Attagenus robustior (Pic, 1951) (figure of habitus see http://www.dermestidae.com/Attagenusrobustior.html), but differs in the following characters:
A. robustior: Antennal club black, broad; body narrower, pubescence of pronotum with several spots and patches, legs dark brown to black, male genitalia almost circular in dorsal aspect (Congo, Namibia, Senegal, Tanzania).
A. pseudorobustior n. sp.: Antennal club brown and more narrow; body broader, pubescence of pronotum uniform without any spots or patches, legs brown, male genitalia elongated in dorsal aspect (Namibia).
The new species differs from all other species of the genus occurring in Namibia in shape and arrangement of the elytral spots.
Kalík's dermestids are stored in several institutions. Some of them are new to science and already got a name by him and are labelled as types, but they were never published as valid descriptions. This concerns also a couple of beetles labelled as “Attagenus roeri sp. n.” which are deposited in the collection of the SMNS. This species is now described in the present paper. Schawaller (1994) cited the material from SMNS erroneously under the name Attagenus roeri Kalík in his type catalogue of Coleoptera from SMNS.
3 List of the Attagenus species from Namibia
Attagenus aurofasciatus Háva, 2005. — Distribution: Namibia, South Africa.
Attagenus cinereus (Thunberg, 1815). — Distribution: Namibia, South Africa.
Attagenus fasciatus (Thunberg, 1795). — Distribution: cosmopolitan.
Attagenus havai Kadej, 2006. — Distribution: Angola, Namibia.
Attagenus heinigi Herrmann & Háva, 2007. — Distribution: Namibia.
Attagenus kaniai Háva & Kadej, 2008. — Distribution: Namibia, Zimbabwe.
Attagenus muehlei Herrmann & Háva, 2014. — Distribution: Namibia.
Attagenus pardus Arrow, 1915. — Distribution: Botswana, Namibia, South Africa, Tanzania, Zimbabwe.
Attagenus pellio (Linnaeus, 1758). — Distribution: cosmopolitan.
Attagenus pseudorobustior n. sp. — Distribution: Namibia.
Attagenus robustior (Pic, 1951). — Distribution: Congo, Namibia, Senegal, Tanzania.
The authors are deeply indebted to Johannes Reibnitz (SMNS) for taking photos of the new species, and to Dr. Wolfgang Schawaller (SMNS) for the loan of specimens.