A new monotypic genus and species of predaceous midge from Costa Rica, Spinellihelea brevicosta Borkent, Grogan, and Picado is described and illustrated. Phylogenetic interpretation of the group of Ceratopogonini in which females have their clypei fused laterally to the eyes, indicates that Spinellihelea is the sister group of Fittkauhelea Wirth and Blanton and that it likely arose in the Neotropical Region. The genus Leptohelea Wirth and Blanton is newly placed in a monophyletic group with Fittkauhelea, Spinellihelea, Diaphanobezzia Ingram and Macfie and Parabezzia Malloch.
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Vol. 110 • No. 3