Acaricis Beard & Gerson, 2009: 2; Beard & Ochoa, 2011: 32.

Type species: Acaricis plana Beard & Gerson, 2009

Diagnosis. Body elongate, more than twice as long as wide; palpus usually 4-segmented. Prodorsum with a rostral shield and 2 or 3 pairs of setae (sc₁ and sc₂ or v₂, sc₁ and sc₂); setae sc₂ lanceolate. Opisthosoma with 9 pairs of dorsal setae; dorsosublateral setae absent; dorsocentral setae c₁ absent, d₁ and e₁ present; humeral setae c₃ and dorsolateral setae 6 pairs (d₃, e₃, f₂, f₃, h₂ and h₁); setae h₂ long and attenuate. Genua IV nude.

The original concept of Acaricis was based on two Australian species (Beard & Gerson, 2009). The new species from New Zealand with new morphological features allows revision of the generic diagnosis as presented above.

Diagnosis. The adult female of Acaricis urigersoni sp. nov. is easily distinguishable from those of the two other species by the absence of the first pair of prodorsal setae v₂, setae sc₁ bifurcated, setae e₃ lanceolate, cylinder-shaped spermathecal vesicle, and chaetotaxy (legs I–IV): femora 4-4-2-1, genua 2-2-0-0, tibiae 5-5-3-2.

The adult male of Acaricis urigersoni sp. nov. is differentiated from those of the two other species by the absence of the first pair of prodorsal setae v₂, setae e₃ lanceolate, setae 4a₂ elongate and flagelliform, and Chaetotaxy (legs I–IV): femora 4-4-2-1, genua 2-2-0-0, tibiae 5-5-3-2, tarsi 7+2ω-7+2ω-5+ω-5+ω.

Type specimens. Holotype ♀. New Zealand, Auckland, Kepa Reserve, 5 Aug, 2013, by Nicholas A. Martin, ex. Gahnia lacera (Cyperaceae). Paratypes. 6 females, 5 males, 7 deutonymphs, 4 protonymphs, 3 larvae, same data as holotype. 1 female, 1 male, 2 protonymphs, 3 larvae, Wenderholm Regional Park, Auckland, New Zealand, 27 Sep, 2013, ex. Gahnia lacera (Cyperaceae), by Nicholas A. Martin. 8 females, 4 males, 4 deutonymphs, 2 protonymphs, Kepa Reserve, Auckland, New Zealand, 9 Oct, 2013, ex. Gahnia lacera (Cyperaceae), by Nicholas A. Martin. The holotype and 39 paratypes (12 females, 8 males, 9 deutonymphs, 6 protonymphs, 4 larvae) will be deposited in the New Zealand Arthropod Collection, Landcare Research, Auckland, New Zealand (NZAC); 10 paratypes (2 females, 2 males, 2 deutonymphs, 2 protonymphs, and 2 larvae) will be deposited in the Natural History Museum, London (BMNH).

Adult Female (n=5)

Gnathosoma. (Fig. 3A) Rostrum reaching proximal end of femur I, subcapitular seta m setiform, m=16–18 (18), m-m=16–18 (18). Palp 4-segmented, setal formula: 0, 0, 2, 2; tarsus with 2 eupathidia 4, 4.

Idiosoma. (Figs. 1A, 2A, 20A, 21A) 445–470 (470) long, 195–205 (200) wide. Rostral shield pitted. Idiosoma widest at posterior margin of propodosoma, narrowing abruptly at base of opisthosoma, then expanding posteriorly. PRODORSUM covered with weak longitudinal sculpturing and faint broad band of transverse striations between setae sc₁ and d₁ (Figs. 1A, 20A); bearing 2 pairs of setae (sc₁ and sc₂), setae sc₁ bifurcated, sc₂ lanceolate, barbed; lengths: sc₁ 5–6 (5), sc₂ 31–36 (35); distances: sc₁-sc₁ 70–74 (73), sc₁-sc₂ 40–49 (43), sc₂-sc₂ 155–165 (155). HYSTEROSOMA smooth, covered with oblique and longitudinal wrinkles as shown in Fig. 21A, a pair of distinct pores anterior to e₁; with 1 pair of humeral setae c₃, 2 pairs of dorsocentral setae d₁ and e₁, and 6 pairs of dorsolateral setae. Setae c₃, d₁, d₃ and e₁ setiform, dorsolateral setae e₃, f₂, f₃ and h₁ lanceolate, barbed, and h₂ elongate, ending in a minute club. Lengths: d₁ 4–6 (4), e₁ 4, c₃ 4, d₃ 3–4 (3), e₃ 15–20 (20), f₂ 23–25 (25), f₃ 23–25 (25), h₂ 117–142 (117), h₁ 21–25 (21); distances: d₁-d₁ 24–28 (24), e₁-e₁ 10–20 (17), c₃-c₃ 160–170 (165), d₃-d₃ 120–130 (125), d₃-e₃ 110–130 (125), e₃-e₃ 120–135 (125), e₃-f₂ 23–24 (24), f₂-f₂ 115–125 (115), f₂-f₃ 19–22 (21), f₃-f₃ 95–105 (95), f₃-h₂ 21–27 (22), h₂-h₂ (34), h₂-h₁ 13–14 (13), h₁-h₁ 33–42 (34).

Venter. (Figs.1B, 2B, 3B, 22A, 23A, B) Venter with few broken oblique striae laterally and fine transverse striations between coxae II and IV. All coxal setae setiform. Setae 1a and posterior medioventral setae 4a₁ and 4a₂ flagelliform, anterior medioventral setae 3a setiform; setae 4a₁ and 4a₂ subequal, more than 8 times as long as 3a (Fig. 2B). Lengths: 1a 130–145 (135), 1b 10–11 (11), 1c 10–12 (11), 2b 17–19 (18), 2c 18–20 (18), 3a 13–15 (13), 3b 16–20 (18), 4a₁ 120–130 (125), 4a₂ 115–125 (115), 4b 15–20 (18). Distances: 1a-1a 25–33 (30), 3a-3a 30–32 (30), 4a₁-4a₁ 9–13 (9), 4a₁-4a₂ 11–15 (12), 4a₂-4a₂ 29–39 (29). Genital and ventral plates with flaps of cuticle as shown in Fig. 1B, bearing 1 pair of aggenital setae ag and 2 pairs of gential setae (g₁ and g₂), setiform; setae g₁ and g₂ subequal. Anal plate with 2 pairs of pseudanal setae (ps₁ and ps₂). Seta lengths: ag 13–16 (14), g₁ 15–18 (16), g₂ 15–18 (16), ps₁ 12–14 (12), ps₂ 10–12 (10). Distances: ag-ag 22–28 (22), g₁- g₁ 20–29 (24), g₁-g₂ 10–12 (11), g₂-g₂ 40–44 (40), ps₁-ps₂ 36–37 (37).

Spermatheca. (Figs. 3B, 22A, 23B) A short, narrow, unsclerotised tube extending from genital opening (mesad setae ps2) and ending in a cylinder-shaped vesicle.

Legs. (Figs. 4, 5) Lengths of legs I–IV: 125–135 (130), 100–110 (105), 82–87 (85), 80–84 (84). Chaetotaxy: coxae 2-2-1-1; trochanters 1-1-1-0; femora 4-4-2-1; genua 2-2-0-0, tibiae 5-5-3-2, tarsi 7+ω-7+ω-5-5. Most dorsal and lateral setae on femora, genua and tibiae lanceolate, lateral setae l′ on tibiae pectinate; ventral setae bv″ on femur I pectinate and on femur II lanceolate; seta ev′ on femur III setiform; ventral setae v pectinate. Setae ft″ on tarsi I–II and ft′ζ on tarsi III–IV flagelliform; unguinal setae u pectinate and equal in length; proral setae p′ζ and p″ζ eupathidial; tectal setae tc setiform except on tarsus IV pectinate. Lengths of solenidia: I ω″ 4–5 (4), II ω″ 4–5 (4). Claws developed with tenent hairs on each side.

Male (n=5)

Gnathosoma. (Fig. 7C) Rostrum reaching proximal end of femur I, and rostral shield pitted, subcapitular setae m setiform, m= 14–18, m-m= 14–18. Palp 4-segmented, setal formula: 0, 0, 2, 2; tibia with 2 bare setae, tarsus with 2 eupathidia, lengthes 3–4, 3–4.

Idiosoma. (Figs. 6A, 7A, 20B, 21B) 325–340 long, 140–155 wide. Rostral shield pitted. Idiosoma widest at posterior margin of propodosoma, narrowing abruptly at base of opisthosoma, then expanding posteriorly. PRODORSUM covered with faint broad band of transverse striations between setae sc₁ and d₁ (Figs. 6A, 20B); bearing 2 pairs of setae (sc₁ and sc₂), setae sc₁ setiform, sc₂ lanceolate, barbed; lengths: sc₁ 4–5, sc₂ 25–30; distances: sc₁-sc₁ 56–65, sc₁-sc₂ 34–38, sc₂-sc₂ 125–135. HYSTEROSOMA divided into metapodosoma and opisthosoma by narrow band of horizontal striations (Figs. 6A, 21B); with 1 pair of humeral setae c₃, 2 pairs of dorsocentral setae d₁ and e₁, and 6 pairs of dorsolateral setae. Setae c₃, d₁, d₃ and e₁ setiform, dorsolateral setae e₃, f₂, f₃ and h₁ lanceolate, barbed, and h2 elongate, ending in a minute club. Lengths: d₁ 3–4, e₁ 3–4, c₃ 3– 4, d₃ 3–4, e₃ 7–10, f₂ 18–19, f₃ 19–21, h₂ 117–122, h₁ 18–21; distances: d₁-d₁ 15–22, e₁-e₁ 10–13, c₃-c₃ 120–130, d₃-d₃ 80–91, d₃-e₃ 90–100, e₃-e₃ 70–81, e₃-f₂ 15–20, f₂-f₂ 70–82, f₂-f₃ 12–18, f₃- f₃ 66–78, f₃-h₂ 12–18, h₂-h₂ 45–53, h₂-h₁ 10, h₁-h₁ 24–36.

Venter. (Figs. 6B, 7B, 8A, 22B, 23D) Venter with fine transverse striation between coxae II and IV, longituidinal striations in the genital area. All coxal setae setiform. Setae 1a and posterior medioventral setae 4a₁ and 4a₂ flagelliform, anterior medioventral setae 3a setiform; setae 4a₁ and 4a₂ subequal, more than 9 times as long as 3a (Fig. 7B). Lengths: 1a 135–150, 1b 10–11, 1c 9–12, 2b 15–20, 2c 16–20, 3a 10–12, 3b 18–20, 4a₁ 100–110, 4a₂ 105–110, 4b 14–17. Distances: 1a-1a 25–30, 3a-3a 23–28, 4a₁-4a₁ 9–11, 4a₁-4a₂ 8–10, 4a₂-4a₂ 25–30. Genital and ventral plates with an aedeagus and seminal vesicle, bearing one pair of aggenital setae ag and 2 pairs of gential setae (g₁ and g₂), setiform; setae g₁ and g₂ subequal. Anal plate with 2 pairs of pseudanal setae (ps₁ and ps₂). Seta lengths: ag 15–18, g₁ 27–29, g₂ 27–30, ps₁ 11–12, ps₂ 7–8. Distances: ag-ag 11–18, g₁- g₁ 15–19, g₁-g₂ 10–11, g₂-g₂ 35–38, ps₁-ps₂ 7–14.

Aedeagus. (Figs. 8, 22B, 23D) A narrow, elongate, sclerotised aedeagus tapering to a point posteriorly (at genital opening); membranous duct running from sclerotised aedeagus to flared, lightly sclerotised, cone-shaped cup distally, appearing to open into a soft membranous vesicle.

Legs. (Figs. 9, 10) Lengths of legs I–IV: 110–120, 90–96, 72–85, 72–83. Chaetotaxy: coxae 2- 2-1-1; trochanters 1-1-1-0; femora 4-4-2-1; genua 2-2-0-0, tibiae 5-5-3-2, tarsi 7+2ω-7+2ω-5+ω- 5+ω. Most dorsal and lateral setae on femora, genua and tibiae lanceolate, lateral setae l′ on tibiae I and II setiform; ventral setae bv″ on femur I pectinate and on femur II lanceolate; seta ev′ on femur III setiform; ventral setae v pectinate except on trochanter II setiform. Setae ft″ on tarsi I–II and ft′ζ on tarsi III–IV flagelliform; unguinal setae u pectinate and equal in length; proral setae p′ζ and p″ζ eupathidial; tectal setae tc setiform except tc′ on tarsi I and II bifurcated. Lengths of solenidia: I ω′ 8–9, ω″ 6–8, II ω′ 7–8, ω″ 6, III ω′ 7, IV ω′ 7. Claws developed with tenent hairs on each side.

Deutonymph (n=5)

Gnathosoma. (Fig. 12A) Rostrum reaching proximal end of femur I, subcapitular seta m setiform, m=13–15, m-m=13–15. Palp 4-segmented, setal formula: 0, 0, 2, 2; tibia with 2 bare setae, tarsus with 2 eupathidia 3–4, 3–4.

Idiosoma. (Figs. 11A, 12C, 20C, 21C) 310–365 long, 140–160 wide. Rostral shield pitted. Idiosoma widest at posterior margin of propodosoma, narrowing abruptly at base of opisthosoma, then expanding posteriorly. PRODORSUM covered with faint broad band of transverse striations between setae sc₁ and d₁ (Figs. 11A, 20C), bearing 2 pairs of setae, setae sc₁ bifurcated, sc₂ lanceolate, barbed; lengths: sc₁ 3, sc₂ 21–28; distances: sc₁-sc₁ 56–63, sc₁-sc₂ 32–39, sc₂-sc₂ 115–130. HYSTEROSOMA smooth, covered with oblique and longitudinal wrinkles as shown in Fig.11A. Setae c₃, d₁, d₃ and e₁ setiform, subequal, dorsolateral setae e₃, f₂, f₃ and h₁ lanceolate, barbed, and h₂ elongate, ending in minute club. Lengths: d₁ 2–3, e₁ 2–4, c₃ 2–3, d₃ 2–3, e₃ 5–7, f₂ 12–16, f₃ 12–17, h₂ 86–105, h₁ 12–17; distances: d₁-d₁ 15–21, e₁-e₁ 6–10, c₃-c₃ 110–125, d₃-d₃ 81–100, d₃-e₃ 57–86, e₃-e₃ 73–95, e₃-f₂ 11–16, f₂-f₂ 72–95, f₂-f₃ 11–16, f₃-f₃ 60–85, f₃-h₂ 12–20, h₂- h₂ 40–51, h₂-h₁ 10–13, h₁-h₁ 19–26.

Venter. (Figs. 11B, 12C, 14C, 22C) Venter with fine transverse striations between coxae II and IV. All coxal setae setiform. Setae 1a and posterior medioventral setae 4a₁ flagelliform, anterior medioventral setae 3a setiform; setae 4a₁ about 6 times as long as 3a (Fig. 12C). Lengths: 1a 125–140, 1b 8–10, 1c 8–10, 2b 9–13, 2c 11–14, 3a 10–14, 3b 14–16, 4a₁ 79–90, 4b 12–13. Distances: 1a-1a 21–23, 3a-3a 21–24, 4a₁-4a₁ 12–19. Genital and ventral plates with oblique cuticles anterior to ag and transverse cuticles between ag and g₁, and oblique striations laterad seta g₁ as shown in Fig. 22C, bearing one pair of aggenital setae ag and one pair of gential setae g₁, setiform. Anal plate with 2 pairs of pseudanal setae (ps₁ and ps₂). Seta lengths: ag 10–14, g₁ 11–14, ps₁ 8–9, ps₂ 5–7. Distances: ag-ag 14–23, g₁-g₁ 13–23, ps₁-ps₂ 18–30.

Legs. (Figs. 13, 14A, B) Lengths of legs I–IV: 84–96, 66–77, 53–61, 48–55. Chaetotaxy: coxae 2-2-1-1; trochanters 1-1-1-0; femora 4-4-2-1; genua 2-2-0-0, tibiae 5-5-3-2, tarsi 7+ω-7+ω-5-5. Most dorsal and lateral setae on femora, genua and tibiae lanceolate, lateral setae l″ on genua I pectinate setiform, lateral setae l′ on tibiae I and II setiform. Ventral seta bv″ on femur I setiform and on femur II lanceolate; ventral seta v and ev′ setiform. Setae ft″ on tarsi I–II and ft′ζ on tarsi III–IV flagelliform; unguinal setae u setiform and equal in length; proral setae p′ζ and p″ζ eupathidial; tectal setae tc setiform. Lengths of solenidia: I ω″ 4, II ω″ 4. Claws developed with tenent hairs on each side.

Protonymph (n=4)

Gnathosoma. (Fig. 16D) Rostrum reaching proximal end of femur I,and rostral shield pitted, subcapitular seta m setiform, m=14–16, m-m=14–16. Palp 4-segmented, setal formula: 0, 0, 2, 2; tibia with 2 bare setae, tarsus with 2 eupathidia 3–4, 3–4.

Idiosoma. (Figs. 15A, 16B, 20D, 21D) 279–305 long, 120–130 wide. Rostral shield pitted. Idiosoma widest at posterior margin of propodosoma, narrowing abruptly at base of opisthosoma, then expanding posteriorly. PRODORSUM covered with faint broad band of transverse striations between setae sc₁ and d₁ (Figs. 16B), bearing 2 pairs of setae, setae sc₁ setiform, sc₂ lanceolate, barbed; lengths: sc₁ 3, sc₂ 20–21; distances: sc₁-sc₁ 45–57, sc₁-sc₂ 28–33, sc₂-sc₂ 105–115. HYSTEROSOMA smooth (Figs. 15A, 21D), with 1 pair of humeral seta c₃, 2 pairs of dorsocentral setae d₁ and e₁, and 6 pairs of dorsolateral setae. Setae c₃, d₁, d₃ and e₁ setiform, subequal, dorsolateral setae e₃, f₂, f₃ and h₁ lanceolate, barbed, and h₂ elongate, ending in minute club. Lengths: d₁ 2, e₁ 2–4, c₃ 2, d₃ 2, e₃ 3–5, f₂ 9–13, f₃ 10–12, h₂ 68–82, h₁ 9–11; distances: d₁-d₁ 15–24, e₁-e₁ 6–9, c₃-c₃ 93–99, d₃-d₃ 68–75, d₃-e₃ 55–58, e₃-e₃ 63–73, e₃-f₂ 8–15, f₂-f₂ 59–66, f₂-f₃ 9–13, f₃-f₃ 46–54, f₃-h₂ 8–14, h₂-h₂ 29–37, h₂-h₁ 9–10, h₁-h₁ 11–20.

Venter. (Figs. 15B, 16A, E, 22D) Venter similar to deutonymph. All coxal setae setiform. Setae 1a flagelliform and anterior medioventral setae 3a setiform (Fig. 16A). Lengths: 1a 98–105, 1b 8–10, 1c 8–9, 2c 11–12, 3a 9–10, 3b 11–14. Distances: 1a-1a 19–22, 3a-3a 21–25. Similar to deutonymph, bearing one pair of aggenital setae ag, setiform. Anal plate with 2 pairs of pseudanal setae (ps1 and ps2). Setae lengths: ag 7–12, ps1 6–7, ps2 5–6. Distances: ag-ag 15–16, ps1-ps2 14–17.

Legs. (Fig. 17) Lengths of legs I–IV: 67–74, 55–62, 48–50, 41–44. Chaetotaxy: coxae 2-1-1-0; trochanters 0-0-1-0; femora 3-3-2-1; genua 1-1-0-0, tibiae 5-5-3-2, tarsi 7+ω-7+ω-5-3. Dorsal setae on femora, genua and tibiae lanceolate; ventral seta bv″ on femur I setiform and on femur II lanceolate; lateral setae l′ on tibiae I and II setiform and l″ on tibiae I and II lanceolate; ventral seta v setiform. Setae ft″ on tarsi I–II and ft′ζ on tarsi III–IV flagelliform; unguinal seta u and tectal seta tc setiform; proral setae p′ζ and p″ζ eupathidial. Lengths of solenidia: I ω″ 3, II ω″ 3. Claws developed with tenent hairs on each side.

Larva (n=5)

Gnathosoma. (Fig. 19D) Rostrum reaching middle of femur I, subcapitular seta m setiform, m=14–17, m-m=14–17. Palp 4-segmented, setal formula: 0, 0, 2, 2; tibia with 2 bare setae, tarsus with 2 eupathidia 3–4, 3–4.

Idiosoma. (Figs. 18A,19F, 20E, 21E) 240–275 long, 85–110 wide. PRODORSUM covered with faint broad band of transverse striations between setae sc₁ and d₁ (Figs. 18A, 20E), bearing 2 pairs of setae, setae sc1 setiform, sc2 lanceolate, barbed; lengths: sc₁ 2–4, sc₂ 12–16; distances: sc₁- sc₁ 45–47, sc₁-sc₂ 20–25, sc₂-sc₂ 82–96. HYSTEROSOMA smooth (Figs. 18A, 21E), with 1 pair of humeral seta c₃, 2 pairs of dorsocentral setae d₁ and e₁, and 6 pairs of dorsolateral setae. Setae c₃, d₁, d₃, e₁ and e₃ setiform, subequal, dorsolateral setae f₂, f₃ and h₁ lanceolate, barbed, and h₂ elongate, ending in minute club. Lengths: d₁ 3–4, e₁ 2, c₃ 2, d₃ 2, e₃ 4, f₂ 8–9, f₃ 8–9, h₂ 89–98, h₁ 8; distances: d₁-d₁ 15–18, e₁-e₁ 6–8, c₃-c₃ 70–82, d₃-d₃ 45–53, d₃-e₃ 35–43, e₃-e₃ 50–60, e₃-f₂ 7–9, f₂-f₂ 45–55, f₂-f₃ 7–8, f₃-f₃ 35–44, f₃-h₂ 8–10, h₂-h₂ 22–27, h₂-h₁ 7–8, h₁-h₁ 8–11.

Venter. (Figs. 18B, 19E, 22E) Venter similar to deutonymph. All coxal setae setiform. Setae 1a flagelliform and anterior medioventral setae 3a setiform (Fig. 19E). Lengths: 1a 82–93, 1b 7–8, 3a 8–9. Distances: 1a-1a 18–25, 3a-3a 21–26. Anal plate with 2 pairs of pseudanal setae (ps₁and ps₂). Setae lengths: ps₁ 5–6, ps₂ 4–5. Distances: ps₁-ps₂ 8–10.

Legs. (Figs. 19A, B, C) Lengths of legs I–III: 50–55, 44–48, 41–44. Chaetotaxy: coxae 1-0-0; trochanters 0-0-0; femora 3-3-2; genua 1-1-0, tibiae 5-5-3, tarsi 5+ω-5+ω-3. Most dorsal setae on femora, genua and tibiae lanceolate except on tibiae I and II nude. Ventral setae bv” on femora and lateral seta l′ on tibiae setiform; lateral seta l″ on tibiae I and II lanceolate; ventral seta v setiform. Setae ft″ on tarsi I–II and ft′ζ on tarsus III flagelliform; unguinal seta u and tectal seta tc setiform; proral setae p′ζ and p″ζ eupathidial. Lengths of solenidia: I ω″ 2, II ω″ 2. Claws developed with tenent hairs on each side.

Etymology. The species is named after Prof Uri Gerson, who is an acarologist from Israel and has made significant contributions to the systematics and biology of the Tenuipalpidae.

Remarks. In live specimens, the body of this new species looks glossy with a translucent appearance, and is yellow-greenish with a series of minute black spots; legs I–II are obviously orange, and legs III–IV are pale orange; the eyes are red (Fig. 24). In one adult male, one dorsolateral setae h2 was duplicated (h2 and h2x in Fig. 23C), and shortened by three times; the other seta h2 was normal.

The ontogeny of Acaricis was reported by Beard and Gerson (2009). Based on the observation of Acaricis plana and Acaricis urigersoni sp. nov., the dorsal idiosomal setae (sc₁, sc₂, d₁, e₁, c₃, d₃, e₃, f₂, f₃, h₂ and h₁) and pseudanal setae (ps₁ and ps₂) are present throughout all stages.

Ventral setae 1c, 2c and 3b are added in the protonymph, 2b, 4a₁ and 4b added in the deutonymph, and 4a₂ added in the adult. Aggenital seta ag emerges in the protonymph; the first pair of genital setae g₁ are formed in the deutonymph and the second pair appear in the adult. The palpal chaetotaxy (0, 0, 2, 2) remains unchanged from larva to adult. The changes in the chaetotaxy of leg segments are listed in Table 1. All dorsal setae d of legs lanceolate except on tibiae I and II in larva nude; ventral setae v in larva, protonymph and deutonymph setiform, in female pectinate, in male on leg I–II pectinate and on leg III–IV setiform; ventral seta ev′ setiform; ventral seta bv′ in adult pectinate and in all immature stages setiform; lateral setae l″ in adult lanceolate, in larva, protonymph and deutonymph lanceolate except on genua I in deutonymph pectinate; unguinal seta u setiform in immature stages and pectinate in adult; tectal seta tc setiform and prorals setae p′ζ and p″ζ eupathidial.