Fay-Wei Li,^1,2,*,** Li-Yaung Kuo,^3,** Yi-Han Chang,^4,5 Tian-Chuan Hsu,⁴ Hsin-Chie Hung,⁶ Wen-Liang Chiou,^4,7 Carl J. Rothfels,¹ Yao-Moan Huang<sup>4,*

1</sup>1University Herbarium and Department of Integrative Biology, University of California, Berkeley, Cal
²2Department of Biology, Duke University, Durham, North Carolina 27708, U. S. A.
³3Institute of Ecology and Evolutionary Biology, National Taiwan University, Taipei 10617, Taiwan.
⁴4Taiwan Forestry Research Institute, Taipei 10066, Taiwan.
⁵5Department of Biological Sciences, National Sun Yat-Sen University, Kaohsiung 80424, Taiwan.
⁶6Taiwan Society of Plant Systematics, Kaohsiung 80424, Taiwan.
⁷7Dr. Cecilia Koo Botanic Conservation Center, Pingtung 906, Taiwan.

^*Authors for Correspondence (fl43@duke.edu, huangym@tfri.gov.tw)
^**These authors contributed equally