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1 March 2010 Lee Denmar Miller (1935–2008): His Life and a Brief History of the Allyn Museum of Entomology
Jacqueline Y. Miller
Author Affiliations +

Dr. Lee D. Miller, a life member of the Lepidopterists' Society, passed away on 5 April 2008 at his home near Gainesville, Florida, after a long illness. He was 72 years old. He is survived by his wife, Jacqueline Y. Miller, and two daughters, Kathryn Lee Angeli of Saratoga, California, and Laura Sue Langford of New York, New York, and one granddaughter, Rowan Langford.

Fig. 1.

Lee Miller, Allyn Museum collections in 1998.


Born on 1 June 1935 in Des Moines, Iowa, Lee was the son of a lawyer, Guy Denmar Miller and a hospital administrator, Anabel Lee Smith. He grew up and attended schools in Des Moines, Iowa. Lee attended Iowa State University and worked with Dr. Jean L. Laffoon, who was a professor in the Departments of Zoology and Entomology. Dr. Laffoon served as the curator of the Iowa State Entomology Collection, a specialist in systematics of fungus gnats and was also involved in mosquito control. Active in the Entomological Society of America and Iowa Academy of Science, among other scientific organizations, he was an excellent mentor and had a major impact on Lee's life. It was here that Lee began to consider a possible career in Entomology. However, Lee had multiple scientific interests, and later transferred to the University of Iowa where he had a triple major in Geology, Biology, and English, until three days prior to graduation. Since he was required to choose only one, Lee selected and graduated with a degree in Biology in 1960. Lee began his graduate program at University of Pittsburgh while also working at the Carnegie Museum of Natural History in the then Section of Insects and Spiders (now Invertebrate Zoology). Lee completed his M. S. degree (1963) on a review of the genus Osmodes Holland (Lepidoptera: Hesperiidae) (1964a). In 1965, he completed his Ph. D. dissertation entitled: The Higher Classification, Phylogeny, and Zoogeography of the Satyridae (Lepidoptera) with publication in 1968.

During his professional career, Lee served as a Research Assistant Professor and Assistant Professor (1965–1968) at the Department of Biology, Catholic University of America, Washington, D.C. He was subsequently the Curator of Allyn Foundation, Inc. (1968–1972), later the Allyn Museum of Entomology (1972–1981). When the Director, Arthur C. Allyn, donated the collection, facilities and property to the Florida State Museum (now Florida Museum of Natural History) in 1981, Lee served as a Curator in the Department of Natural History, Florida Museum of Natural History, Allyn Museum of Entomology, University of Florida (1981–2004) (Fig. 1). With the development of the McGuire Center of Lepidoptera and Biodiversity, the Allyn Museum collections were moved to the new Center in 2004. Lee served as the Allyn Curator of Lepidoptera in the McGuire Center at the Florida Museum of Natural History from 2004 until his death. He was also an Adjunct Professor in the Departments of Zoology (1981–2008) and Entomology and Nematology (1995–2008) at the University of Florida. In addition to the above, Lee was a Visiting Associate Professor in the Department of Biology, University of Florida, Tampa (1973–1977) and a Research Scholar and Adjunct Faculty Member in the Division of Natural Sciences, at New College of Florida (State Honors College), University of South Florida, Sarasota (1995–2004), where he taught Entomology, Zoogeography and Phylogenetics. A Research Associate of the Department of Zoology-Entomology, Field Museum of Natural History, Chicago, Illinois (1971–1979), Lee was also appointed as a Research Associate of the Section of Invertebrate Zoology, Carnegie Museum of Natural History, Pittsburgh, Pennsylvania (1965–2008). Lee taught courses and seminars, including three workshops on the collection and identification of insects at the College of the Bahamas, Nassau, Bahamas (1988–1993) along with Mark Simon and me.

Over the years Lee served the Lepidopterists' Society in various capacities, including: Zone Coordinator, Season Summary (1964–1968); Secretary-elect (1971); Secretary (1972–1976); Editor, Supplements of the Lepidopterists' Society (1971–1973); Member, Editorial Committee, Journal of the Lepidopterists' Society (1972–1995); Editor, Clench Memorial issue (vol. 37 (2) 1980); President-elect (1982–83); President (1983–84); and Immediate Past-President (1984–1985). He served as a Member at Large on several occasions. Lee conceived and implemented the idea for the Karl Jordan Medal honoring and recognizing lepidopterists for publication of original research of exceptional quality on morphology, taxonomy, systematies, zoogeography and “natural history.” He was also a member of the Association for Tropical Biology, Inc., The Society for the Study of Evolution, Society of Systematic Biology, The American Entomological Society, Entomological Society of America, Southern Lepidopterists' Society, Willi Hennig Society, Fellow, Royal Entomological Society (London), and a Fellow, Linnean Society (London). He was the Editor of the Bulletin of the Allyn Museum (1971–2007), during which he oversaw the development, external review, and production of this series, which now numbers more than 160 issues and is still in production at the McGuire Center. He also served as an Associate Editor of the Journal of Research on the Lepidoptera (1971–1978). In addition to the above, Lee was an accredited judge of the American Orchid Society, and served as Judges Co-training Coordinator, Florida North Central Judging Center in Tampa (1986–1996).

As a child, Lee was introduced to natural history at an early age as both of his parents were avid bird watchers and interested in native plants, insects, and mammals. Denmar had had a butterfly and moth collection as a child, and he and Lee collected in both Iowa and Minnesota throughout Lee's childhood. Denmar's collection was eventually donated to Lee's grade school in Des Moines. In addition, there was a group of boys in east Des Moines, the “35th St. Boys”, who were pals and went fishing, hunting, and collected insects among other things. They included B. C. Johnson, Ron Royer, and Norris Young among others. Lee was the oldest, and taught them how to collect and prepare butterflies and moths. When his parents moved to Franklin and 39th St. in West Des Moines, these friends would still get together on occasion. They would often go collecting at various state parks including Pilot Knob State Park (Hancock Co.), and Waubonsie State Park (Fremont Co.) in southwestern Iowa, in search of new additions to their collections. A couple of these trips culminated in Lee's first paper published in the Journal (1962a) in which he reported observations on nine Iowa butterfly species, including four species new to the state. Later, Lee joined John Downey and others in revisiting a few of these sites (1975–1978), to see if some of the uncommon species were still extant, and contributed to the recently published The Butterflies of Iowa (Schlicht et al. 2007). Lee's personal collection was donated to the Carnegie Museum about the time that he completed his Ph. D. in May, 1965.

In addition to sharing their love for natural history, the Miller family also shared a passion for fishing and hunting. Summers were spent fishing along the Raccoon River near Des Moines or in lakes near their cabin or farm in Minnesota. Fall activities included collecting wild rice and cranberries, hunting ducks or grouse and watching the Springer Spaniels work the fields for birds. Of course, collecting and watching Lepidoptera were always part of these trips. Lee and Denmar coveted their Master Angler trophies for their fishing prowess in Canada (Fig. 2), but butterfly nets were always included with the fishing gear.

Fig. 2–3.

(2) Lee with one of his Master Angler fish about 1974. (3) Lee and Denmar Miller upon Lee winning the Iowa State High School tournament in 1953.


Lee and Denmar also shared a love for the game of golf. Denmar had won nearly all of the state's top titles and had represented the state of Iowa at the Western Open (1936–1939). Lee in turn was a competitive golfer in high school, won the Iowa State High School title in 1953 (Fig. 3), and went on to play golf at both Iowa State University and the University of Iowa. He once competed against Jack Nicklaus in a Big 10 tournament, but Nicklaus was a formidable competitor even then. However, golf took more of a recreational role later in Lee's life.

Following his graduation from the University of Iowa in 1960, Lee and his then wife, Susan, with daughter Kathryn, went to Casa Grande, Arizona, near Tucson, in search of employment as an entomologist. It wasn't long before Lee was out in the field and saw some Megathymus. These skippers seemed unusual compared to other Hesperiidae that he had encountered in the Midwest. Soon after, Lee had the opportunity to meet Kilian Roever, who was then in the Department of Entomology at the University of Arizona. Kilian took him to some of his special collecting sites, and Lee had numerous opportunities to observe and collect immature Megathymus. He accumulated tents of seven species, some from two or more localities, and began a comparative study of their emergence patterns. When some potential job opportunities arose in Des Moines later that year, the family traveled back east with these immature Megathymus in the car. Lee immediately contacted “the 35 St. Boys” about the Megathymus, and they eagerly came to see these new treasures and caught up on their recent collecting experiences. Lee recorded the number of males versus females and made observations on other aspects of their emergence patterns. He later reported back to Kilian in a letter of “the megs emerging thick and fast for about a month.” Thus began an exchange of information on skippers between Kilian and Lee, who shared a passion for all butterflies, especially Hesperioidea. Years later after moving to Sarasota in 1969, Kilian arrived unannounced the following spring to collect Megathymus cofaqui on Longboart Key and further south to Venice. At that time, Longboat was mostly undeveloped, and there were stands of Yucca aloifolia all along the key. Following visits to various spots in the Sarasota area, Kil continued to collect M. cofaqui all the way up the Gulf coast into the Florida panhandle.

The last Megathymus from Arizona emerged in Des Moines early on 11 November 1960, the day that Lee and the family headed off to Meadville, Pennsylvania, where he had secured a job as a plant pest control inspector for USDA (1960–1961). Prior to this trip, Lee had already corresponded with Harry Clench and took the opportunity to visit the Carnegie Museum of Natural History in Pittsburgh on several occasions. Here he met the Curator in the Division of Insects and Spiders, Dr. Richard M. Fox, who held a teaching appointment in the Department of Biological Sciences at the University of Pittsburgh. Following a successful collecting expedition to Liberia, Fox had recently taken this position at Carnegie. Given Lee's interest in Hesperiidae, he was intrigued with Fox's Liberian material. Arthur W. Lindsey was already working on the skippers from Liberia, but due to his declining health, he needed some assistance in completing this portion of the proposed volume. Lee was also exploring the possibility of continuing his education and obtaining advanced degrees with Fox. In addition to completing the Liberian monograph, Fox was in the process of organizing a two month expedition to Baja California (October–December, 1961). This trip was financially supported by a distinguished scientist research fellow at the Academy of Natural Sciences, Philadelphia, Margaret J. Cary. Cary was a noted specialist on the Sphingidae and at that time, the southern half of the Baja peninsula was largely unexplored. There were excellent opportunities for studying the biodiversity and discovering new species in a number of phyla. Lee joined the expedition as an entomologist while arrangements were in progress for his admission to graduate school. For Lee, who until then had encountered such difficulty in trying to secure an entomological position, it was a surprising turn of events.

As is with such expeditions, not everything went as planned. Harry Clench (lepidopterist), Neil Richmond (herpetologist), and John Bauer (preparator) drove from Pittsburgh and met Lee at Richmond, Indiana. The itinerary scheduled the above personnel to travel and collect along the west coast of Mexico for two weeks in the newly acquired International Carryall and to meet Dr. Fox and his wife, Jean, in Mazatlan (Fig. 4a). However, the vehicle was soon nicknamed “Pariah” as the gas mileage was a little over 14 mi/gal., and within four days of starting the trip from Pittsburgh, the battery had died. The unexpected problems with the truck provided constant delays, immense frustration, and entertainment throughout the trip. The field team flew from Mazatlan into southern Baja on November 4, 1961, while the vehicle and other equipment arrived three days later by ferry. The group was separated into three field teams, who would collect herps, fossils, plants, and insects. With Lee on a horse and Harry on a mule, they headed up into the Boca de la Sierra for a week of collecting and camping (4b, 4c). After the first day, Lee decided that he would rather walk and collect than ride. A few days later, Harry's mule fell down a hill. In the end, both animals carried gear, and Lee and Harry walked out of the forest.

Fig. 4.

Margaret J. Cary field expedition to Baja California. October–November, 1961: a) Field team at Guaycura, La Paz (left to right), Lee Miller, Jean Fox, Harry Clench, Neil Richmond, Richard Fox, and John Bauer; b) Harry and Lee heading up into the Boca de la Sierra; c) Camping area in La Cienaga I, Sierra Laguna; d) Lee and Harry examining Agave for Megathymus larvae near San Ignacio; e) Rocky road about 30 mi. north of Bahia, S. L. Gonzaga; f) Sand dunes approximately 29 mi. N. of San Felipe with Lee and Harry (two future presidents of the Lepidopterists' Society) hard at work celebrating the end of the trip.


Once the surveys had been completed in southern Baja, Dick and Jean Fox flew directly back to Pittsburgh. The rest of the crew drove the 600 miles north to Tijuana, collecting along the way over the primitive trails and “rocky roads” (Fig. 4e) into a variety of habitats (4f). Lee was heavily involved in the collecting of insects during this expedition, especially Lepidoptera and primarily the Hesperioidea (4d). His efforts resulted in a collaboration with Don MacNeil and the description of two new subspecies, Polites sabuleti margaretae and Cogia hippalus peninsularis (1969). This experience was the impetus for taking four additional collecting trips to continental Mexico with an emphasis on different states (1966–1973) (Tamaulipas, Veracruz, Morelos, Guererro, Chiapas, Michoacan, Sinaloa, Nayarit, Durango), and has provided a historical timeline and background for the more thorough biodiversity surveys in progress and completed by Jorge Llorente, Amando Luis, and other members in their working group at Universidad Nacional Autónoma de México (UNAM) in addition to Carmen Pozo De la Tijera, El Colegio de la Frontera Sur (ECOSUR).

Following the Baja trip, Lee entered graduate school in the Department of Biological Sciences, University of Pittsburgh in January, 1962. He was a member of the first Organization of Tropical Studies class at La Selva in 1963 where he met Dan Janzen and Jay Savage and made new friendships that would last throughout his life. It was also here that his interest in all aspects of the Neotropics and the associated fauna and flora intensified. During his stay, Mount Irazú, the tallest volcano in Central America, erupted. Lee took pictures of this natural event and noted how the insects attempted to adapt to the situation. For a geologist/biologist, Lee continued to have a keen interest over the years on the succession of plants and insects following such catastrophic events and the associated evolutionary changes.

Following the completion of his Ph. D., Lee accepted a position as a Research Assistant Professor in the Department of Biology at Catholic University of America in Washington, D. C. in 1965. There he developed and taught five courses in addition to mentoring more than six graduate students in just three years. In late 1967, Lee was presented with an opportunity to curate a private Lepidoptera collection. He interviewed in November with Arthur C. Allyn, an avid collector, private businessman, and co-owner of the Chicago White Sox. Allyn had recently acquired the W. J. Kaye collection, an international collection that contained a number of types. Mr. Allyn wanted someone to fill this position immediately, but Lee had already signed a teaching contract and had three graduate students, who were in the process of trying to finish degrees. So he turned down the position with regret. Allyn then placed an advertisement in Science and received more than 300 applications for the position. Following Christmas vacation, there was message from Allyn requesting that Lee return his call. However, when he attempted to do so, there was some confusion with another Miller, the representative for the baseball players' union. In the end, Allyn reoffered the curatorial position to Lee and said that he was willing to wait until July 1, 1968, for him to begin. Lee gladly accepted Allyn's offer, but with a few caveats. Allyn stated that the position would be finite and last for seven years, but he had a long list of things that he would like to see accomplished during this period. To fulfill these requirements and initiate a state of the art scientific collection, Lee would need a curatorial assistant for preparation of specimens, general collection maintenance, etc. Lee and I had met at the University of Pittsburgh and had worked together on various courses at Pitt and at Catholic University. Neither of us could deny the natural compatibility and magic behind what would become the “Miller team.” Mr. Allyn was supportive of an assistant position which I accepted along with Lee's proposal.

Initially our goals were to expand both the taxonomic and geographic representation of the collection worldwide, especially of butterflies and some moths. For the latter, we would focus on Saturniidae, Sphingidae, and Arctiidae as there were significant holdings in the W. J. Kaye collection. In addition, we also wanted to establish the Allyn collection as one of the most significant scientific research resources for future lepidopterists. Allyn was very open to these suggestions. In the interim, Mr. Allyn wanted Lee to represent him at an auction of Lepidoptera in Paris in early February, 1968. The auction included the LeMoult collection among others and was conducted by Claude Lemaire. Although Allyn was intrigued by the more showy Morpho and Papilionidae, Lee was interested in obtaining unprepared specimens, especially of the Charaxes, Euphaedra, and Euriphene (African Nymphalidae) and also in filling some major voids in Allyn's collection, especially in the Riodinidae, Hesperioidea, and Lycaenidae that were each represented by a single drawer. In addition to some types and a number of prepared specimens, Lee eventually purchased more than 300 “LeMoult” boxes of unprepared specimens.

Lee's interest in the African Lepidoptera expanded when we joined Arthur Allyn. Allyn was interested in a number of African genera, especially Charaxes and Colotis (Pieridae). Although the Carnegie Museum of Natural History and American Museum of Natural History had significant African collections, we sought to increase holdings from other African countries that were not well represented in other U.S. collections. Fortunately, Allyn already had business interests in South Africa with excellent contacts. We supported a number of collectors in the field and obtained a number of collections during 1968–1977. In addition, Lee had purchased material earlier from Father Theo Maessen, a priest in Ghana, beginning in 1963, and this continued until Maessen left the country in 1987. Over the years Lee managed to make three trips into South Africa and one into Kenya. This provided him with additional information not only about the habitats and ecology of these butterflies but more insight concerning the historical geology and biogeography of this continent.

Another major geographic area of interest was, of course, the Neotropics due to Lee's research on the Hesperiidae and Nymphalidae, and in particular the Satyrinae. Lee was particularly interested in the higher altitudinal Satyrinae as there was more promise for obtaining endemic species and new taxa to be described. He had made contacts with a number of collectors and/or colleagues throughout the Neotropics. Through F. Martin Brown, who had conducted surveys various in countries in South America in the late 1920's and lived for some time in Ecuador, Lee contacted Rosario de Lafebre and the Velastigui family. In 1969, Rosario, her family, and their cadre of collectors were charged with collecting selected lepidopteran groups on all of the volcanic peaks in Ecuador. There were occasional forays into lower elevations, such as the Rio Coca, as opportunities for travel into some of these isolated areas for biodiversity surveys became available. Through Rosario, we also met Nadia Venedictoff, who had collected both butterflies and moths throughout Ecuador, often accompanied by Rosario. Nadia donated her large Lepidoptera collection to the Allyn Museum in 1985. Both of these collections among others were significant additions to the neotropical holdings as they not only provided new material for description but increased our taxonomic and geographic representations in the collection.

Due to the expansion of the collections, library and additional space for new equipment, the Allyn Museum soon outgrew the original Florida facilities at the Sarasota Bank and Trust building and was moved to a new 5500 square foot building on Bay Shore Road. We hosted the Lepidopterists' Society meetings there in 1973. Research visits to the collections increased markedly following the meetings, and a few of these visitors were attracted to the Sarasota area. In 1977, Steve Steinhauser moved to Sarasota to work in the collections on a regular basis, and he was soon followed by Dale and Joanne Jenkins in 1979. Nadia Venedictoff also moved to Sarasota and worked there 1986–1991. Arthur Allyn had purchased a scanning electron microscope to enable his detailed morphological studies on Lepidoptera. Beginning in 1981, John Downey spent summers working and collaborating with Arthur Allyn on a number of butterfly ultrastructure studies using the SEM as well as morphological studies on immature stages and pupal sound production. John moved permanently to Sarasota in 1988. There were also a number of New College students, who worked on undergraduate theses, and others who assisted with the preparation of specimens. The Museum was a hub of activity with the integration of new specimens and revisionary and other studies in progress. Lee always enjoyed working in the collections with visitors. He had a humorous bent and was known for not only being able to discuss the systematics, biogeography and life history of various Lepidoptera but also for having a long list of the latest jokes. There was never a dull moment at the Allyn Museum; however, the combination of Steve Steinhauser and Lee made for a truly comical team. Together there was always some bizarre occurrence, interesting puns or new humor to be shared.

It is difficult to summarize all of the significant highlights of Lee's life and accomplishments as a lepidopterist here. Some are listed above, but a few additional accounts are noteworthy. Over a number of years, F. Martin Brown had located, documented, and designated type specimens originally described by William Henry Edwards and published these for separate taxonomic groups. In 1974, Brownie enlisted Lee to assist him with the Hesperioidea to complete the project. Over the next six years, they visited various museums throughout the U. S. and Canada and published four papers (1975, 1977, 1980, 1987) and in all, designated 53 Lectotypes and 23 Neotypes. In addition to the above, Lee and Brownie co-authored A Catalogue/Checklist of the Butterflies of America North of Mexico published by the Lepidopterists' Society in 1981. Initially they reworked the original dos Passos checklist and updated the nomenclature including the higher classification, which has been subsequently refined. There were many discussions between the authors about generic versus subgeneric designations, and often these resulted in compromise. In order to keep costs down and through the kindness of Jack Serbin, Serbin Printing Inc., we were allowed to typeset some of the volume on his new electronic equipment on the weekends or at night. The Lepidopterists' Society benefited from this effort as the volume sold well and the original publication investment was returned within five years.

In 1980, Dr. David Spencer Smith, who I had met at the University of Miami Medical School in 1965, visited the Allyn Museum along with Dennis Leston and Barbara Lenczewski. They were actively working on variation in Eurema daira, especially palmira, in south Florida. David was also a friend of Mark Simon, and he knew that Lee and Mark had been collecting in the Bahamas on a regular basis. David was interested in Bahamian butterflies and had also been collecting there along with Dennis Knowles. Over the next few years, we developed a potential project on the Caribbean butterflies, which culminated in The Butterflies of the West Indies and South Florida. At the time, David was the Hope Professor at Oxford University Museum and a Professor in the Department of Zoology. He approached Oxford University Press about their potential interest in the project, and we obtained a contract in 1987. Although the text was initiated in 1988, there were so many voids in our knowledge, especially of the butterfly fauna of the Virgin Islands and Lesser Antilles. Thus, we initiated surveys over the next six years making several trips each year in order to obtain current information on the butterfly biodiversity of these islands. We did not want this work to be based solely on specimens in museum and private collections. Along with the superb illustrations of Richard Lewington and through the assistance and kindness of many colleagues, the volume was finally published in 1994 after years of editing, annotations, and descriptions of new taxa.

Lee's primary research interests included the systematies, taxonomy, and biogeography of Lepidoptera, especially the Hesperiidae, Nymphalidae, and Lyeaenidae. He was an authority on the Satyrinae worldwide and published a number of revisionary studies on the group. His dissertation work on The Higher Classification, Phylogeny, and Zoogeography of the Satyridae (Lepidoptera) (1968a) represents a seminal work within the field, and his studies on Pindis, Megisto, and Paramacera set the standard for future revisionary studies. He had initiated several revisionary treatments of the Euptyehiina, especially the genus Taygetis; these unfinished studies are in progress and will be completed by various collaborators in conjunction with me in the future. Lee had also continued his interest within the Hesperiidae and was especially enamored with the African fauna (Miller & Collins, 1997; Douglass & Miller 2003) and the close alignment with certain taxa in the Neotropics. Our studies in the West Indies further fueled Lee’s interest in the historical biogeography and relative age of Lepidoptera and are summarized in three papers (1989, 1998, 2001).

Figs. 5–6.

(5) Lee sharing some humorous moments at the Hope Museum in 1986. Left to right: Lee Miller. Chris O'Toole, and David Spencer Smith; (6) Collecting in the Sierra Maestre in eastern Cuba. 1995. Left to right: Mark Simon, Lee, and Jackie Miller.


In Lee's opinion, one of his greatest accomplishments was the implementation of a phylogenetic arrangement for museum Lepidoptera on a worldwide basis. Lee realized that such an arrangement provided researchers and students with the opportunity to review higher level taxa in one place as opposed to curating taxa according to a biogeographic region. He believed that natural history collections are not only a historical document but also an evolutionary learning tool. Amateurs and professionals can learn the characteristic appearance or gestalt of a group through such an arrangement.

However, with all of these interests and accomplishments, Lee would probably state here that his greatest contribution was his interaction with students of Lepidoptera at every level and especially with amateurs in the field. The Lepidopterists' Society has always been a very unique organization in the respect that it includes both amateurs and professionals and provides the opportunity for both to interact, learn, and collaborate with one another. Lee's Presidential address in 1984 was a tribute to the amateur lepidopterist during which he recounted the accomplishments of a number of well known amateur lepidopterists through time, including Pieter Cramer, Dru Drury, Jacob Hübner, Frederick DuCane Godman, Osbert Salvin, Walter Rothschild, James Joicey, William Barnes, Lionel Higgins, Cyril F. dos Passos, Roy and Connie Kendall, Dick Dominick, Arthur C. Allyn, and today, Lee would have added William McGuire. These individuals, among nameless others, were or are amateurs, who worked in other occupations but have enjoyed the sense of discovery and learning about all aspects of Lepidoptera. Lee always had time to talk with amateurs, professionals, and students at all levels of their careers, review specimens with them in the collection, and discuss various topics. Together, we followed the careers of a number of students, some from middle school through their graduate degrees and into their professional careers. Over the years, Lee was proud of the accomplishments of so many students, perhaps best exemplified by Ron Royer, now at Minot State University, who was one of the original “35th St. Boys” from Des Moines, and with whom he had spent so many enjoyable trips in the field as a teenager. As Ron Royer so aptly stated about Lee, “he has given countless others such indelible memories to cherish as he gave us (the 35th St. Boys).” To Lee, this was always time well spent, as these efforts were repaid with the enthusiasm which comes with new discoveries, long-term friendships, and sometimes, even new additions to the Museum collections. For Lee, life was to be enjoyed, and he felt that he was one of the luckiest people in the world — working on Lepidoptera and actually getting paid for it!


The study of Lepidoptera is steeped in the “history” of this unique insect order, from the study of systematics to observations on and the first publication of that special life history of a particular species. In essence, lepidopterists are a composite drawn from their life experiences in the field and with people whom they have met and shared information along the way. This obituary is a historical retrospect and features some, but not all, of Lee's contacts over the years. There are numerous stories and events, but it would have been impossible to have included everyone and every detail here.

I would like to thank Drs. Deborah L. Matthews, Andrew D. Warren, and Rebecca B. Simmons for providing comments on this manuscript, many of which have been incorporated into the final version. I would like to especially thank Dr. Matthews for her invaluable assistance in helping me organize materials for this work and to Dr. Ron Royer for providing me with additional insight into Lee's early life as a collector in Des Moines. I would also like to thank here the hundreds of lepidopterists, colleagues, and friends throughout the world who phoned, sent emails, or cards with personal messages and remembrances upon Lee's death. He was a very private person and would have been somewhat taken aback by all of your kind words.


  • Adelpha milleri Beutelspacher 1976. Beutelspacher, C. R. 1976. Estudios sobre género Adelpha Hübner en Mexico (Lepidoptera: Nymphalidae). Revista de la Sociedad Mexicana de Lepidopterología 2(1): 8–14.

  • Bungalotis milleri Freeman 1977. Freeman, H. A. 1977. Six new species of Hesperiidae from Mexico. Journal of the Lepidopterists' Society, 31(2): 89–99.

  • Pachyneuria milleri Steinhauser 1989. Steinhauser, S. R. 1989. Taxonomic notes and descriptions of new taxa in the neotropical Hesperiidae. Part I. Pyrginae. Bulletin of the Allyn Museum 127: 1–70.

  • Nesiostrymon milleri K. Johnson. 1991. Johnson, K. 1991. Cladistics and the biogeography of two trans-Caribbean hairstreak butterfly genera: Nesiostrymon and Terra (Lepidoptera, Lycaenidae). American Novitates (3011): 1–43.

  • Celaenorrhinus milleri Collins & Larsen. 2003. Collins, S. C., T. .B. Larsen, & H. Warren-Gash. 2003. Notes on Afrotropical butterflies with description of eleven new species and four new subspecies (Lepidoptera: Rhopalocera). ABRI Research Paper, No. 3. Metamorphosis 14: 63–110.

  • Piruna millerorum Steinhauser 1991. Steinhauser, S.R. 1991. Six new species of skippers from Mexico (Lepidoptera: Hesperiidae: Pyrginae and Heteropterinae). Insecta Mundi. 5(1): 25–44. (named for L. D. and J. Y. Miller)

  • Callophrys (Mitoura) millerorum Clench 1981. Clench, H. K. 1981. New Callophrys (Lycaenidae) from North and Middle America. Bulletin of the Allyn Museum (64): 21–23. (named for L. D. and J. Y. Miller)

  • Mycelia cyaniris millerorum Jenkins 1984. Jenkins, D. W. 1984. Neotropical Nymphalidae II. Revision of Myscelia. Bulletin of the Allyn Museum, 87: 1–64. (named for L. D. and J. Y. Miller)

  • Eurema agave millerorum Llorente & Luis 1987. Llorente, J. E., & M. A. Luis. 1987. Una nueva subespecie de Eurema agave Cramer (Lepidoptera: Pieridae: Coliadinae) Folia Entomológica Mexicana 71: 17–25. (named for L. D. and J. Y. Miller)

Bibiography: Lee D. Miller


L. D. Miller 1962a. Notes on nine Iowa butterfly species, including four new to the state. Journal of the Lepidopterists' Society 15: 97–98. Google Scholar


L. D. Miller 1962b. A new Hesperia from Arizona (Lepid.: Hesperiidae). Entomological News 73: 85–90. (Hesperia susanaeGoogle Scholar


L. D. Miller 1964a. A review of the genus Osmodes Holland (Lepidoptera: Hesperiidae). Transactions of the American Entomological Society 89: 277–304. (Osmodes lindseyiGoogle Scholar


L. D. Miller 1964b. African Hesperiidae: notes and descriptions (Lepidoptera). Bulletin de la Institute francais d'Afrique Noire (A) 26: 640–647. (Celaenorrhinus bakolo, Sarangesa brigida sanaga, Gorgyra bule, Ceratrichia flava benitoensis, Ceratrichia weberi, Greta bugoma, stat. nov.Google Scholar


K. W. Cummins , L. D. Miller , N .A. Smith & R. M. Fox 1965. Experimental Entomology. New York, Reinhold: xi + 176 pp.; ill. Google Scholar


A. W. Lindsey & L. D. Miller . 1965. Family Hesperiidae, in R. M. Fox , et al., The Butterflies of Liberia. Memoirs of the American Entomological Society 19: 47–146. (Gyrogra, new genus, Meza leucophaea bassa, Andronymus antonius, Gretna balenge zowa, Caenides naGoogle Scholar


L. D. Miller & S. M. Miller . 1966. Chromosomes from testicular preparations of Lepidoptera. Science 152: 529–530. Google Scholar


L. D. Miller & S. M. Miller . 1966a. Systematics and zoogeography of the genus Phanus (Hesperiidae). Journal of Research on the Lepidoptera 4: 115–130. (Phanus obscurior prestoni, P. australisGoogle Scholar


L. D. Miller & S. M. Miller . 1966b. A review of the West Indian “Choranthus”. Journal of Research on the Lepidoptera 4: 259–274. (Choranthus richmondi, ParachoranthusGoogle Scholar


L. D. Miller & S. M. Miller . 1966c. (Abstract). The evolution, phylogeny and zoogeography of the higher taxa of the Satyridae (Insecta: Lepidoptera). Dissertation Abstracts 27(4): 1–2. Google Scholar


L. D. Miller & S. M. Miller . 1966d. A new Staphylus from Costa Rica (Lepid.: Hesperiidae). Entomological News, 77: 261–264. (Staphylus esmeraldusGoogle Scholar


R. S. Monroe & L. D. Miller . 1967. Report on a collection of Hesperiidae from Honduras. Journal of the Lepidopterists' Society 21: 243–247. Google Scholar


L. D. Miller & H. K. Clench . 1968. Some aspects of mating behavior in butterflies. Journal of the Lepidopterists' Society 22: 125–132. Google Scholar


L. D. Miller & H. K. Clench . 1968a. On Mexican Satyridae, with description of a new species. Journal of Research on the Lepidoptera 7: 51–55. (Cyllopsis dospassosiGoogle Scholar


L. D. Miller & H. K. Clench . 1968b. The Higher Classification, Phylogeny and Zoogeography of the Satyridae (Lepidoptera). Memoirs of the American Entomological Society, (24): iii + 174 pp.; ill. (Antirrhini, Euptychiini, new tribes) Google Scholar


L. D. Miller & H. K. Clench . & C. D. MacNeill . 1969. Reports on the Margaret M. Cary-Carnegie Museum expedition to Baja California, Mexico, 1961. 5. Two new subspecies of Hesperiidae (Lepidoptera) from the Cape region, Baja California Sur, Mexico. Annals of Carnegie Museum 41: 19–24. (Cogia hippalus peninsularis, Polites sabuleti margaretaeGoogle Scholar


L. D. Miller & H. K. Clench . 1969. An anomalous foreleg in a male of Dodonidia helmsii (Satyridae). Journal of the Lepidopterists' Society 23: 201–202. Google Scholar


L. D. Miller & H. K. Clench . 1970a. Multiple capture of Caria ino melicerta (Riodinidae) at light. Journal of the Lepidopterists' Society. 24: 13–15. Google Scholar


L. D. Miller & H. K. Clench . 1970b. Two new Mexican Hesperiidae. Journal of the Lepidopterists' Society 24: 120–124. (Vinius freemani, Virga clenchiGoogle Scholar


L. D. Miller & H. K. Clench . 1970c. Nomenclature of wing veins and cells. Journal of Research on the Lepidoptera 8: 37–48. Google Scholar


L. D. Miller & H. K. Clench . 1970d. Report on the Margaet M. Cary-Carnegie Museum expedition to Baja California, Mexico, 1961. 7. The family Hesperiidae (Lepidoptera). Annals of Carnegie Museum 41: 169–200. (Neotype, Hesperia zampa (=Systacea zampa) (Edwards)) Google Scholar


L. D. Miller & H. K. Clench . & J. Y. Miller . 1970a. Pieris protodice and Urbanus dorantes in southern Florida. Journal of the Lepidopterists' Society 24: 244–247. Google Scholar


L. D. Miller & H. K. Clench . 1970b. Notes on two rare Mexican Adelpha and related Central American species (Nymphalidae). Journal of the Lepidopterists' Society 24: 292–297. Google Scholar


L. D. Miller & H. K. Clench . 1971. Descriptions of new species and notes on other Hesperiidae of Africa. Bulletin of the Allyn Museum 2: 1–17.(Celaennorrhinus illustroides, C. beni jacquelinae, Abantis lucretia etombiensis, A. maesseni, Ceratrichia phocion camerona, C. maesseni, Osmodes adonides, O. maesseni, O. lindseyi occidentalis, Meza cybeutes volta, Caenides allyni, Fresna maesseniGoogle Scholar


L. D. Miller & H. K. Clench . & T. C. Emmel . 1971. The Brazilian “Cercyonis” (Satyridae). Journal of the Lepidopterists' Society 25: 12–19. ( Cercyeuptychia, PseudocercyonisGoogle Scholar


L. D. Miller & H. K. Clench . & S. S. Nicolay . 1971. Two new Charaxinae from Panama and the Canal Zone (Nymphalidae). Bulletin of the Allyn Museum 1: 1–5. ( Agrias zenodorus smalli, Anaea kingiGoogle Scholar


L. D. Miller & H. K. Clench . 1972a. (Review of): Australian Butterflies by Charles McCubbin, and Butterflies of the Australian Region by Bernard D'Abrera. Journal of the Lepidopterists' Society 26: 200–202. Google Scholar


L. D. Miller & H. K. Clench . 1972b. Revision of the Euptychiini (Satyridae). 1. Introduction and Paramacera Butler. Bulletin of the Allyn Museum 8: 1–18. (Paramacera chinanteca, P. copiosa, P. xicaque rubrosuffusa, P. allyniGoogle Scholar


L. D. Miller & H. K. Clench . 1972c. The Karl Jordan Medal. Journal of the Lepidopterists' Society 26: 207–209. Google Scholar


L. D. Miller & H. K. Clench . & J. Y. Miller . 1972a. A new riodinid from northern Argentina (Riodinidae). Bulletin of the Allyn Museum 4: 1–5. (Eiseleia pichanalensisGoogle Scholar


L. D. Miller & H. K. Clench . 1972b. New high-altitude Hesperiinae from Mexico and Ecuador (Hesperiidae). Bulletin of the Allyn Museum 7: 1–9. (Dalla roeveri, Nylla (new genus), N. cordillera, N. allynorum, Paratrytone capta, P. deceptaGoogle Scholar


L. D. Miller & H. K. Clench . 1973a. (Review of): Butterflies of Australia by I. F. B. Common and D. F. Waterhouse. Journal of the Lepidopterists' Society 27: 310–311. Google Scholar


L. D. Miller & H. K. Clench . 1973b. First Karl Jordan Medal awarded to Henri Stempffer. Journal of the Lepidopterists' Society 27: 311–312. Google Scholar


L. D. Miller & H. K. Clench . 1974a. (Review of): Butterflies of the World by H. L. Lewis. Journal of the Lepidopterists' Society 28: 178–179. Google Scholar


L. D. Miller & H. K. Clench . 1974b. (Review of): Hewitson on Butterflies, 1867–1877, with preface by Dr. L. G. Higgins. Journal of the Lepidopterists' Society 28: 179–180. Google Scholar


L. D. Miller & H. K. Clench . 1974c. (Review of): An Amateur's Guide to the Study of the Genitalia of Lepidoptera by the A. E. S. Journal of the Lepidopterists' Society 28: 180. Google Scholar


L. D. Miller & H. K. Clench . 1974d. Revision of the Euptychiini (Satyridae). 2. Cyllopsis R. Felder. Bulletin of the Allyn Museum 20: 1–98. (Cyllopsis hedemanni tamaulipensis, C. jacquelinae, C. steinhauserorum, C. guatemalena, C. diazi, C. schausi, C. pallens, C. parvimaculata, C. suivalenoides, C. suivalens escalantei, C. pyracmon nabokovi, C. henshawi, C. pertepida intermedia, C. windi, C. perplexaGoogle Scholar


L. D. Miller & H. K. Clench . 1974e. Comment in support of a proposal by N. D. Riley and L. G. Higgins on Papilio actaeon Fabricius, 1775 v. Papilio actaeon von Rottemburg, 1775. Z. N. (S.) 1937. Bulletin of Zoological Nomenclature 31(3): 104–105. Google Scholar


L. D. Miller & H. K. Clench . 1975a. Threatened status for two butterflies. Field Museum Bulletin, Sept.: 15–18, 22. Google Scholar


L. D. Miller & H. K. Clench . 1975b. Butterfly conservation: the right way, the wrong way, the government way. Insect World Digest, Sept./Oct.: 2–9. Google Scholar


F. M. Brown & L. D. Miller . 1975. The types of the hesperiid butterflies named by William Henry Edwards. Part I, Hesperiidae; Pyrginae. Transactions of the American Entomological Society 101: 597–649. (Lectotype, Eudamus dorus (=Zestusa dorus ) (Edwards), Lectotype, Eudamus drusius (= Thorybes drusius) (Edwards), Lectotype, Eudamus mochus (=Cogia caicus moschus) (Edwards), Lectotype, Eudamus hippalus (=Cogia hippalus hippalus) (Edwards), Lectotype, Hesperia hayhurstii (=Staphylus mazans hayhurstii) (Edwards), Lectotype, Pholisora ceos (= Staphylus ceos) (Edwards), Lectotype, Hesperia tamenund (=Achlyoldes thraso tamenund) (Edwards), Lectotype, Thanaos clitus (=Erynnis juvenalis clitus) (Edwards), Lectotype, Pyrgus xanthus (Edwards), Lectotype, Hesperia ricara (=Pyrgus ruralis) (Edwards), Neotype, Syrichtus petreius (=Pyrgus ruralis) (Edwards), Lectotype, Hesperia wyandot (=Pyrgus centaureae wyandot) (Edwards), Lectotype, Syrichtus alba (=Heliopetes ericetorum) (Boisduval), Lectotype, Pyrgus nearchus (=Heliopetes domicella domicella) (Erichson), Lectotype, Pholisora nessus (=Celotes nessus) (Edwards), Lectotype, Thananos alpheus (=Pholisora alpheus alpheus) (Edwards), Lectotype, Pholisora oricus (=Pholisora alpheus oricus) (Edwards), Lectotype, Ancyloxpha lena (=Pholisora libya lena) (Edwards)) Google Scholar


L. D. Miller & F. M. Brown . 1975. Notes on the rare Mexican pierid Prestonia clarki (Pieridae). Journal of the Lepidopterists' Society 29(4): 256–258. Google Scholar


T. C. Emmel , L. D. Miller & H. K. Clench . 1975. The neotropical metalmark Hermathena oweni (Riodinidae): new records and major extension of the known range from Costa Rica to El Salvador and Mexico. Journal of the Lepidopterists' Society 29(2): 108–111. Google Scholar


H. K. Clench & L. D. Miller . 1976. How to prepare slides of sclerotized parts of Lepidoptera. Section of Insects and Spiders, Carnegie Museum of Natural History, Pittsburgh, Pennsylvania. 6 pp. Google Scholar


L. D. Miller 1976. Revision of the Euptychiini (Satyridae). 3. Megisto Hübner. Bulletin of the Allyn Museum 33: 1–23. ( Megisto rubricata pseudocleophes, M. r. anabelaeGoogle Scholar


L. D. Miller & J. Y. Miller . 1976. Notes and descriptions on Mexican Charaxinae (Nymphalidae). Bulletin of the Allyn Museum 41: 1–3. ( Prepona (Prepona) brooksiana diaziana, Anaea (Memphis) wellingiGoogle Scholar


L. D. Miller 1977. Appendix in Fisher, M. S. The taxonomy and identity of Papilio nitra W. H. Edwards in Colorado (Papilionidae). Bulletin of the Allyn Museum 47: 7–8. Google Scholar


F. M. Brown & L. D. Miller . 1977. The types of the hesperiid butterflies by William Henry Edwards. Part II; Hesperiidae: Hesperiinae, Section I. Transactions of the American Entomological Society 103: 259–302. (Neotype, Hesperia mandon (=Carterocephalus palaemon mandan) (Edwards), Lectotype, Pholisora pirus (=Piruna pirus) (Edwards), Lectotype, Hesperia nortonii (Edwards) (=Lerema accius accius) (Smith), Neotype, Heteropterus arene (=Ancyloxypha arene) (Edwards), Lectotype, Thymeticus (sic) hylax (Edwards) (=Oarisma garita) (Reakirt), Neotype, Hesperia waco (Edwards) (=Copaeodes aurantiaca) (Hewitson), Lectotype, Heteropterus procris Edwards (=Copaeodes aurantiaca) (Hewitson), Lectotype, Pamphila rhesus (=Yvretta rhesus) (Edwards), Lectotype, Pamphila carus (=Yvretta carus) (Edwards), Copaeodes eunus (=Pseudocopaeodes eunus) (Edwards), Lectotype, Copaeodes wrightii (=Pseudocopaeodes eunus) (Edwards), Lectotype, Pamphila Morrisoni (=Stinga morrisoni) (Edwards), Lectotype, Pamphila attalus (=Hesperia atalus atalus) (Edwards), Lectotype, Pamphila straton (=Hesperia meskei) (Edwards), Lectotype, Pamphila harpalus (=Hesperia comma harpalus) (Edwards), Lectotype, Pamphila cabelus (=Hesperia comma harpalus) (Edwards), Lectotype, [Pamphila] oregonia (=Hesperia comma oregonia) (Ed- (Ed- wards), Lectotype, [Pamphila] idaho (=Hesperia comma harpalus) (Edwards), Neotype, Hesperia uncas uncas Edwards)) Google Scholar


S. R. Steinhauser & L D. Miller . 1977. Three new species of Adelpha (Nymphalidae) from Mexico and Colombia. Bulletin of the Allyn Museum 46: 1–10. ( Adelpha levona, A. escalantei, A. jacquelinaeGoogle Scholar


L. D. Miller 1978a. (Review of): The Butterflies and Moths of Hampshire and the Isle of Wight (being and account of the whole of the Lepidoptera) By B. Goater. Journal of the Lepidopterists' Society 32(1): 63. Google Scholar


L. D. Miller 1978b. (Review of): Butterflies of West Malaysia and Singapore by W. A. Fleming. Journal of the Lepidopterists' Society 32(1): 63–64. Google Scholar


L. D. Miller 1978c. Notes and descriptions of Euptychiini (Lepidoptera: Satyridae) from the Mexican Region. Journal of the Lepidopterists' Society 32(2): 75–85. (Taygetis mermeria griseomarginata, Splendeuptychia kendalli, Cyllopsis wellingiGoogle Scholar


L. D. Miller 1978. Electrostrymon angelia angelia (Lycaenidae) The oldest Florida record? Journal of the Lepidopterists' Society 32(2): 139–140. Google Scholar


L. D. Miller & J. Y. Miller . 1978. “East Indian” butterflies: notes and descriptions. 1. Satyridae and Danaidae. Bulletin of the Allyn Museum 49: (1–23). (Mycalesis sitius procyon, M. duponcheli allynorum, M. mucia aruana, Acrophtalmia artemis windorum, Ypthima pandocus microocellata, Y. pandocoides, Danaus philene jimiensis, D. albata dabreraiGoogle Scholar


L. D. Miller 1978. Revision of the Euptychiini (Satyridae). 4. Pindis R. Felder. Bulletin of the Allyn Museum 50: 1–12. Google Scholar


L. D. Miller 1979. (Review of): Pennington's butterflies of Southern Africa. edited by C. G. C. Dickson with D. M. Kroon. Journal of the Lepidopterists' Society 33(1): 69–71. Google Scholar


J. Y. Miller & L. D. Miller . 1979 [1978]. Notes on Mexican Actinote (Nymphalidae: Acraeinae) and their relatives, with description of a new subspecies. Journal of the Lepidopterists' Society 32(4): 261–272. ( Actinote stratonice oaxacaGoogle Scholar


L. D. Miller & F. M. Brown . 1979. Studies in the Lycaeninae (Lycaenidae). 4. The higher classification of the American coppers. Bulletin of the Allyn Museum 51: 1–30. ( Hyllolycaena, Hermelycaena)  Google Scholar


L. D. Miller & B. Rotger . 1979. Two new Chlosyne (Nymphalidae) from Mexico. Bulletin of the Allyn Museum 54: 1–4. ( Chlosyne gaudialis wellingi, C. mazarumGoogle Scholar


L. D. Miller 1980a. Obituary: Harry Kendon Clench (1925–1979). Journal of the Lepidopterists' Society 34(2): 81–85. Google Scholar


L. D. Miller 1980b. A review of the Erora laeta group with description of a new species (Lycaenidae). Journal of the Lepidopterists' Society 34(2): 209–216. (Erora caudataGoogle Scholar


F. M. Brown & L. D. Miller . 1980. The types of hesperiid butterflies named by William Henry Edwards. Part II, Hesperiidae: Hesperiinae, section II. Transactions of the American Entomological Society 106 (1): 43–88. (Lectotype, Pamphila siris (=Polites sonora siris (Edwards), Lectotype, Pamphila draco (=Polites draco) (Edwards), Neotype, Hesperia mystic (=Polites mystic mystic) (Edwards), Lectotype, Pamphila mardon (=Polites mardon) (Edwards), Lectotype, Pamphila mardon (=Polites mardon) (Edwards), Neotype, Pamphila verna (=Pompeius verna) (Edwards), Neotype, Hesperia huron (=Atalopedes campestris) (Boisduval), Neotype, Hesperia logan (=Atrytone logan logan) (Edwards), Neotype, Hesperia delaware (=Anatrytone logan delaware) (Edwards), Lectotype, Pamphila lagus (=Anatrytone logan lagus) (Edwards), Lectotype, Pamphila byssus (=Problema byssus) (Edwards), Neotype, Hesperia yuma (=Ochlodes yuma) (Edwards), Lectotype, Pamphila versus (=Ochlodes agricola versus) (Edwards), Neotype, Hesperia napa (=Ochlodes sylvanoides napa) (Edwards), Neotype, Poanes viator) (Edwards), Lectotype, Pamphila taxiles (=Poanes taxiles) (Edwards), Lectotype, Hesperia melane (=Paratrytone melane melane) (Edwards), Neotype, Pamphila rurea (=Euphyes metacomet) (Harris), Lectotype, Hesperia oskya (=Euphyes metacomet) (Harris), Neotype, Hesperia conspicua (=Euphyes conspicua conspicua) (Edwards), Neotype, Pamphila pontiac (=Euphyes conspicua conspicua) (Edwards), Lectotype, Pamphila dion (=Euphyes dion dion) (Edwards)) Google Scholar


H. K. Clench & L. D. Miller . 1980. Papilio ladon Cramer vs. Argus pseudargiolus Boisduval and LeConte (Lycaenidae): a nomenclatorial nightmare. Journal of the Lepidopterists' Society 34(2): 103–119. Google Scholar


L. D. Miller 1981a [1980]. (Review of) A revision of the genus Hipparchia Fabricius, by Otakar Kudrna. Journal of the Lepidopterists Society 34 (3) : 326. Google Scholar


L. D. Miller 1981b [1980]. (Review of) Butterflies of the Australian region, second ed. by Bernard D'Abrera. Journal of the Lepidopterists' Society 34(4): 377–378. Google Scholar


L. D. Miller 1981c. External Anatomy, Lycaeninae, and Satyridae. In Butterflies of the Rocky Mountain States ( C. D. Ferris & F. M. Brown , eds.) Univ. of Oklahoma Press, Norman: 442 pp. Google Scholar


L. D. Miller & F. M. Brown . 1981. A Catalogue/Checklist of the Butterflies of America North of Mexico ( C. V. Covell Jr , ed. ) The Lepidopterist's Society, Memoir No. 2: 280 pp. Google Scholar


L. D. Miller 1982. Insects and Arachnids, in Reader's Digest North American Wildlife ( J. Wernert Susan , ed.) Reader's Digest Association, Inc. pp. 263–282. Google Scholar


L. D. Miller 1983a. (Review of) Butterflies of Oman by Torben Larsen. Journal of the Lepidopterists' Society 37 (1): 69. Google Scholar


L. D. Miller 1983b. (Review of) The Butterflies of the Malay Peninsula by A. S. Corbet and H. M. Pendlebury, 3rd ed. revised by J. N. Eliot. Journal of the Lepidopterists' Society 37(1): 95–96. Google Scholar


L. D. Miller & F. M. Brown . 1983a. Butterfly taxonomy: a reply. Journal of Research on the Lepidoptera 20: 193–198. Google Scholar


L. D. Miller 1983b. Hesperioidea and Papilionoidea in Checklist of The Lepidoptera of America North of Mexico ( R. W. Hodges , ed.) E. W. Classey Ltd. and Wedge Ent. Res. Found., Washington, D. C. pp.42–65. Google Scholar


L. D. Miller 1984a [1983]. Chermock, Hovanitz and Weber collections donated to Allyn Museum. Journal of the Lepidopterists' Society 37(4): 317–318. Google Scholar


L. D. Miller 1984b. “Butterfly” in World Book Encyclopedia, Chicago pp. 618–629. Google Scholar


L. D. Miller & J. De La Maza E. 1984. Notes on Cyllopsis, especially from Mexico, with description of a new species (Lepidoptera: Satyridae). Bulletin of the Allyn Museum 88:1–7. ( Cyliopsis whiteorumGoogle Scholar


L. D. Miller 1985. A review of the genus Ebusus Evans (Lepidoptera: Hesperiidae), with the description of a new subspecies from Mexico. Insecta Mundi 1: 43–45. ( Ebusus ebusus nigriorGoogle Scholar


L. D. Miller , D. J. Harvey & J. Y. Miller . 1985. Notes on the genus Euphyes with description of a new subspecies (Lepidoptera: Hesperiidae). Florida Entomologist 68(2); 323–335. (Euphyes pilatka klotsiGoogle Scholar


L. D. Miller & J.Y. Miller . 1985. Arthur C. Allyn (1913–1985). Bulletin of the Allyn Museum (97): 1–6. Google Scholar


L. D. Miller 1985. (Review of) Species Group Names in the Hesperioidea by Charles Bridges. Journal of the Lepidopterists' Society 39: 51. Google Scholar


L. D. Miller & J. Y. Miller . 1986. A New Thecline from Sulawesi (Lepidoptera: Lycaenidae: Theclinae). Insecta Mundi 2: 143–145. ( Deudorix cleoraGoogle Scholar


L. D. Miller 1986a. Relationship between Pedaliodes perperna and petronius with the description of a new subspecies. Journal of the Lepidopterists' Society 39: 187–195. Google Scholar


L. D. Miller 1986b. Presidential address, 1984: a tribute to the amateur. Journal of the Lepidopterists' Society 40: 1–7. Google Scholar


L. D. Miller 1986c. (review of) Milkweed Butterflies by P. R. Ackery & R. I. Vane-Wright. Journal of the Lepidopterists' Society 40: 72–73. Google Scholar


L. D. Miller & J. Y. Miller . 1986. A. C. Allyn. Journal of the Lepidopterists' Society 40: 75–76. Google Scholar


F. M. Brown & L. D. Miller . 1987. The types of hesperiid butterflies described by William Henry Edwards. Part II — Hesperiidae: Hesperiinae, Section III and Megathymidae. Transactions of the American Entomological Society 113: 28–71. (Lectotype, Pamphila pittacus (=Atrytonopsis pittacus) (Edwards), Lectotype, Pamphila regulus (=Atrytonopsis loammi), Lectotype, Pam- phila lunus (=Atrytonopsis lunus) (Edwards), Neotype, Hesperia deva (=Atrytonopsis deva) (Edwards), Neotype, Hesperia vialis (Amblyscirtes vialis) (Edwards), Lectotype, Hesperia eos (=Amblyscirtes eos) (Edwards), Neotype, Hesperis comus (=Amblyscirtes eos) (Edwards), Lectotype, Amblyscirtes nysa Edwards, Lectotype, Hesperia wakulla Edwards (=Amblyscirtes aesculapius (Fabricus), Neotype, Hesperia nemoris Edwards (=Amblyscirtes hegon) (Scudder), Lectotype, Amblyscirtes aenus Edwards, Lectotype, Ambyscirtes cassus Edwards, Lectotype, Amblyscirtes simius Edwards, Pamphila arabus (=Lerodea arabus) (Edwards), Neotype, Hesperia eufala (=Lerodea eufala) (Edwards), Neotype, Hesperia maculata (= Oligoria maculata) (Edwards), Neotype or Lectotype, Hesperia ocola (=Panoquina ocola) (Edwards), Neotype, Hesperia ophis (=Panoquin panoquin) (Scudder), Lectotype, Megathymus neumoegeni (=Agathymus neumoegeni) (Edwards)) Google Scholar


L. D. Miller 1987. A new subspecies of Heraclides aristodemus from Crooked Island, Bahamas, with a discussion of the distribution of the species. Bulletin of the Allyn Museum 113: 1–8. (Heraclides a. majasiGoogle Scholar


D. S. Smith , S. J. Ramos , F. McKenzie , E. Munroe & L. D. Miller . 1988. Biogeographical affinities of the butterflies of a “forgotten” island: Mona (Puerto Rico). Bulletin of the Allyn Museum 121: 1–35. Google Scholar


L. D. Miller & J. Y. Miller . 1988. A new Euptychia species from northwestern Mexico (Satyridae). Journal of the Lepidopterists' Society 42: 276–280. (Euptychia rubrofasciataGoogle Scholar


L. D. Miller 1989. Satyridae and Hesperiidae (part) in C. D. Ferris (ed.) Supplement to a Catalogue/Checklist of the butterflies of America north of Mexico. Memoir of the Lepidopterists' Society (3): 103 pages. Google Scholar


L. D. Miller & J. Y. Miller . 1989. The biogeography of West Indian butterflies (Lepidoptera: Papilionoidea, Hesperioidea): a vicariance model. Pp. 229”262 in C. A. Woods (ed.) Biogeography of the West Indies. Sandhill Crane Press, Gainesville, Florida. Google Scholar


L. D. Miller 1990. Nearctic Aglais and Nymphalis: Laurasia revisited? The Entomologist (London): 109: 106–115. Google Scholar


L. D. Miller 1990a. (Review of) The Charaxinae Butterflies of Africa by S. F. Henning. Journal of the Lepidopterists' Society 44: 100–101. Google Scholar


L. D. Miller 1990b. (Review of) Mariposas Mexicanas by R. de la Maza Ramirez. Journal of the Lepidopterists' Society 44: 101–102. Google Scholar


S. R. Steinhauser , L. D. Miller , J. Y. Miller & C. A. Bridges III . 1990. Proposal to suppress Eumesia C. and R. Felder [1865] and validate Dalla Mabille 1904 (Lepidoptera: Hesperiidae). Bulletin of Zoological Nomenclature 47(3): 184–186. Google Scholar


L. D. Miller 1990. (Review of) The Butterflies of Egypt by T. B. Larsen. Journal of the Lepidopterists' Society 44: 205–206. Google Scholar


K. Johnson & L. D. Miller . 1991. A new Ministrymon from Chile (Lepidoptera: Lycaenidae). Revista Chilena de Entomologia 16: 183–192. ( Ministrymon quebradivagaGoogle Scholar


L. D. Miller 1991. A new species of Lymanopoda from Southern Ecuador. Deutsche entomologische Zeitscrift 38: 103–107. (Lymanopoda hannemanniGoogle Scholar


D. S. Smith , L. D. Miller , & F. McKenzie . 1991. The butterflies of Anegada, British Virgin Islands, with descriptions of a new Calisto (Satyridae) and a new Copaeodes (Hesperiidae) endemic to the island. Bulletin of the Allyn Museum 133: 1–25. ( Calisto anegadensis, Copeodes eoaGoogle Scholar


L. D. Miller , M. J. Simon & D. J. Harvey . 1992. The butterflies of Crooked, Acklins and Mayaguana, Bahamas, with a biogeographic analysis of the butterflies of the southern Bahamas. Annals of Carnegie Museum 61: 1–31. ( Memphis intermedia venus, M. i. mayaguana, Hemiargus thomasi clenchi, Eurema chamberlaini clenchiGoogle Scholar


L. D. Miller & S. R. Steinhauser . 1992. Butterflies of the Cayman Islands, with the description of a new subspecies. Journal of the Lepidopterists' Society 46: 119–127. ( Dryas iulia zoeGoogle Scholar


K. Johnson and L. D. Miller . 1992. Additions to the Chilean fauna. Pp. 6–9 in Reports of the Museum of Natural History, University of Wisconsin (Stevens Point) ( K. Johnson ,ed.) 23: 1–20, figs. (Heoda (new genus) atacamaGoogle Scholar


J. Y. Miller & L. D. Miller . 1993. The butterflies of the Tonga Islands and Niue, Cook Islands, with the descriptions of two new subspecies. Bishop Museum Occasional Papers 34: 1–24. ( Jamides carissma thomasi, Catochrysops taitensis hopkinsiGoogle Scholar


L. D. Miller 1993. (review of) Butterflies of the Bulolo-Wau Valley by Michael Parsons. Annals of the Entomological Society of America 86: 506–507. Google Scholar


D. S. Smith , L. D. Miller & J. Y. Miller . 1994. The Butterflies of the West Indies and South Florida. Oxford Univ. Press, Oxford: x + 264 pp., 33 pls. Google Scholar


L. D. Miller , J. Y. Miller , & R. A. Anderson . 1997. You caught what in your backyard? Journal of the Lepidopterists' Society 51: 184–187. Google Scholar


L. D. Miller & M. J. Simon . 1997. New and notable records of Bahamian butterflies and moths (Lepidoptera: Lycaenidae, Hesperiidae, Sphingidae). Bahamian Journal of Science 4: 20–23. Google Scholar


L. D. Miller & S. C. Collins . 1997. A review of the genus Gamia Holland 1896 (Lepidoptera: Hesperiidae: Hesperiinae) with description of a new species. Metamorphosis 8:147–159. ( Gamia abriGoogle Scholar


J. Y. Miller & L. D. Miller . 1997. Guide to the Butterflies of Central Florida. 15pp., 9 pl., 178 figs. Google Scholar


L. D. Miller & J. Y. Miller . 1998. Gondwanan Butterflies: The African-South American Connection. Metamorphosis, Occasional Supplement 3: 42–51. Google Scholar


L. R. Hernandez , L. D. Miller , J. Y. Miller , M. J. Simon & T. W. Turner . 1998. New Records and Range Extensions of Butterflies from Eastern Cuba. Caribbean Journal of Science 34: 324–327. Google Scholar


L. D. Miller & M. J. Simon . 1998a. Rediscovery of the Rare, “Endemic” Cuban Butterfly, Achlyodes munroei, with Notes on Its Behavior and Possible Origin. Caribbean Journal of Science 34: 327–329. Google Scholar


L. D. Miller 1998b. A preliminary annotated list of the butterflies (Lepidoptera: Rhopalocera) of Cat Island, Bahamas. Bahamas Journal of Science 6: 35–42. Google Scholar


L. D. Miller & G. Lamas . 1999. A new Taygetis from Subandean Eastern Peru and Western Brazil (Lepidoptera: Satyrinae). Revista Peruana de Entomologia 41:31–35. ( Taygetis inambariGoogle Scholar


A. O. Debrot , J. Y. Miller , B. T. Leysner & L. D. Miller . 1999. The Butterfly Fauna of Curacao, West Indies: 1996 Status and Long-Term Species Turnover. Caribbean Journal of Science 35: 184–194. Google Scholar


J. Y. Miller & L. D. Miller . 2001. The biogeography of the West Indian butteflies (Lepidoptera): An application of a Vicarian/Dispersalist Model. Pp. 127–155 in Biogeography of the West Indies, Patterns and Perspectives ( C. A. Woods , F. E. Sergile , eds.). 2nd edition. CRC Press, Boca Raton, Florida. Google Scholar


J. F. Douglass & L. D. Miller . 2003. Afrotropical skippers (Lepidoptera: Hesperioidea) and the emergence of the combined refugium theory. Bulletin of the Allyn Museum 143: 1–18.  Google Scholar


J. Y. Miller , A. O. Debrot , & L. D. Miller . 2003. Butterflies of Aruba and Bonaire with new species records for Curacao. Caribbean Journal of Science 39: 170–175. Google Scholar


Á. L. Viloria , L. D. Miller & J. Y. Miller . 2003. Pedaliodes pheretias (Hewitson) form griseola Weymer (Nymphalidae: Satyrinae) : Its identity and availability, with description of a new species. Journal of the Lepidopterists' Society 57:62–67. ( Pedalioides gustaviGoogle Scholar


J. V. Calhoun , L. D. Miller & J. Y. Miller . 2004. Melitaea nycteis Doubleday & Hewitson [1847] (currently Chlosyne nycteis, Insecta, Lepidoptera): Proposed conservation of the specific name. Submitted to the ICZN for consideration, action, and publication. (ICZN website) Google Scholar


L. D. Miller 2004. Taygetomorpha, new genus. P. 285 in Atlas of Neotropical Lepidoptera. Checklist: 4A Hesperioidea — Papilionoidea ( G. Lamas , ed.) (Taygetomorpha). Association for Tropical Lepidoptera, Scientific Publishers, Gainesville, Florida. Google Scholar


Á. Viloria , L. D. Miller , & J. Y. Miller . 2004. Neopedaliodes, new genus. P. 284 in Atlas of Neotropical Lepidoptera. Checklist: 4A. Hesperioidea — Papilionoidea ( G. Lamas , ed.) (Neopedaliodes). Association for Tropical Lepidoptera, Scientific Publishers, Gainesville, Florida. Google Scholar


L. D. Miller & J. Y. Miller . 2004. The Butterfly Handbook. Quarto Publishing and New Burlington Books, London, England, 256 pp. Google Scholar


J. V. Calhoun , L. D. Miller & J. Y. Miller . 2005. Melitaea nycteis Doubleday, 1847 (currently Chlosyne nycteis; Insecta, Lepidoptera): proposed conservation of the specific name. Bulletin of Zoological Nomenclature 62(2): 79–83 Google Scholar


L. D. Miller & J. Y. Miller 2005. 2005 Abbot Award to Lee D. Miller and Jacqueline Y. Miller. Southern Lepidopterists' News, 27(4): 99–101. Google Scholar


J. Y. Miller & L. D. Miller . 2006. Report on Field Work and Observations on Cat Island, Bahamas. Bahamas National Trust. 5 pp. Google Scholar


G. T. Austin , L. D. Miller & J. Y. Miller . 2007. Taxonomic comments on Pseudolycaena Wallengren (Lepidoptera: Lycaenidae: Theclinae: Eumaeini). Bulletin of the Allyn Museum 149: 1–22. Google Scholar


L. D. Miller , M. J. Simon & L. Wills . 2007. A new species of Delias (Lepidoptera: Pieridae) from New Ireland Island, Bismarck Islands, Papua New Guinea. Bulletin of the Allyn Museum 144: 1–5. ( Delias laknekeiGoogle Scholar


J. Y. Miller & L. D. Miller . 2008. Obituary Stephen Rogers Steinhauser (1921–2007). Journal of the Lepidopterists' Society 62(4): 232–236. Google Scholar


Á. L. Viloria , L. D. Miller , J. Y. Miller & T. W. Pyrcz . 2008. A reassessment of Parapedaliodes Forster and Neopedaliodes Viloria, L. D. Miller & J. Y. Miller with descriptions of new taxa (Nymphalidae: Satyrinae). Bulletin of the Allyn Museum 151: 1–43. ( Neopedaliodes p. proculeja, new comb., N. juba jubilosa, N. michaeli, N. michaeli citrica, N. lafebreae, N. margaretha allyni, N. nora stepheniGoogle Scholar

Literature Cited


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Jacqueline Y. Miller "Lee Denmar Miller (1935–2008): His Life and a Brief History of the Allyn Museum of Entomology," The Journal of the Lepidopterists' Society 64(1), 38-49, (1 March 2010).
Received: 8 February 2010; Accepted: 15 February 2010; Published: 1 March 2010
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