Open Access
1 December 2013 William E. Miller (1930–2013)
Marc E. Epstein, Susan J. Weller, Roger D. Moon, William D. Hutchison, William J. Mattson, Ian V. Macrae, Todd M. Gilligan
Author Affiliations +

William E. Miller was curious about what made the natural world tick, particularly when it came to Lepidoptera. Although he lived much of his life in the North, he never lost the warm, southern charm acquired from his birthplace in Texas and early years in Louisiana. This tall, thin gentleman with beard and jaunty beret was a fixture on the St. Paul Campus at the University of Minnesota. Spring on campus brought him the pleasures of seeing the magnolia trees in full bloom. Little did the students in the Student Center nearby know that this emeritus professor had once sung in a barbershop quartet!

Bill's academic career began at Louisiana State University where he obtained his B.S. in Zoology in 1950. He then migrated north to study, receiving both his M.Sc. in 1951 and his Ph.D. in 1955 at The Ohio State University under the guidance of his major professor Alvah Peterson, who is best known for his seminal two-volume work Larvae of Insects. Bill enjoyed the rural setting of the Wooster Campus while working on his dissertation research titled The biology and control of the European pine shoot moth. He was a research assistant at the Ohio Agricultural Experiment Station in Wooster until 1954, when he joined the U.S. Army Division of Communicable Diseases at the Walter Reed Army Institute of Research.

October 1971


A fellowship from the West German Government (1957–1958) at the Forest Zoology Institute of Göttingen University, Hann. Münden coincided with Dr. Miller's appointment as Project Leader at the Lake States Forest Experiment Station in East Lansing, Michigan (1956–1964) on the campus of Michigan State University. Upon his return state-side, he worked at the experiment station while obtaining a second M.Sc. in Forestry from Michigan State (1961).

Dr. Miller was a Project Leader at the North Central Forest Experiment Station on the St. Paul campus of the University of Minnesota from 1964 until he retired in 1982. He became an adjunct (and later emeritus) professor in the Department of Entomology, Fisheries and Wildlife (now the Department of Entomology) in 1976, where he generously contributed to teaching, mentoring of graduate students, and departmental service for more than 20 years. Bill was reserved but influential to the intellectual and social fabric of the department. He was a major professor and favorite mentor of graduate students even after his “official” retirement. His thoughtful and positive encouragement was not limited to students, as he mentored faculty as well. Many a faculty member sought his advice on everything from statistical analyses to balancing the demands of a personal life with work. Bill was a favorite editor of manuscript drafts, leaving his carefully penned comments, which were rigorous, thorough, and never harsh, in green ink. For many years, he taught a special course on Lepidoptera he titled Lepidopterology, which included a potpourri of information such as atypical examples of strange biologies, life histories undergirded by his copious knowledge of evolution, and systematics of this prolific insect order. When Bill stepped away from daily university activities in the early 2000s due to health issues, he remained active in his research and contributions to the Minnesota Department of Entomology.

May 2005


During his career, Dr. Miller authored 137 publications, including several monographs; his complete bibliography follows this article. In the realm of systematics, Bill was a specialist on the Tortricidae, subfamily Olethreutinae. He described many new taxa (2 genera and 43 species), proposed new synonymies, and elucidated features of their life histories, becoming recognized as an authority on the North American members of this difficult group. His in-depth knowledge of systematics dovetailed nicely with his work responsibilities in the U.S. Forest Service, particularly for economic pests such as the European pine shoot moth, a species he published on extensively. Bill was especially interested in documenting or describing tortricid species that were introduced into North America, and many of his publications focused on these taxa. He also had an interest in documenting the “true” distribution of Holarctic species, many of which were considered different species in the Palaearctic and Nearctic regions. His interest in past North American tortricid workers resulted in detailed catalogs of the species described by J. B. Clemens and C. H. Fernald, which became foundations for stabilizing the names proposed by these two early lepidopterists. Dr. Miller's vast knowledge of Olethreutinae was partially summarized in his monograph Guide To the Olethreutine Moths of Midland North America (1987), a publication that greatly influenced many of the current Olethreutinae experts in North America. However, Bill did not limit his studies to tortricids. In his ecological work on Lepidoptera he researched and published on a diversity of topics such as body size and its relation to geography and altitude, the correlation between adult feeding and oviposition, the diversity and life-histories of gallforming Lepidoptera, and many others. His research explored and documented the interplay of systematics, ecology, evolution, and behavior in an experimental framework.

Dr. Miller was an avid reader, and he enjoyed fleshing out information from the literature by reanalyzing published data. For example, in his work on the population dynamics of Lepidoptera, he and his graduate student used published data on populations of multiple species from the same locality over at least a seven-year period to find that their fluctuations were often correlated. In more recent years, Bill compiled various literature sources of day-degree data and reanalyzed them using a more accurate analytical method that had been previously published but largely ignored. Just before his death, he worked to exhume and correct Professor Alexander Hodson's data set that detailed the past 51 years of spring development by woody plants on the St. Paul campus, and to make this data available to phenology researchers through the University of Minnesota's Digital Conservancy.

As an academic advisor and experienced editor, Dr. Miller stressed the importance of writing every day and challenged students to make the most of their creative faculties. He did this by example with his 1978 publication of The facile mind: an annotated bibliography for enhancing creativity and problem solving. Some of his other favorite topics included the use of multiple induction in hypothesis testing and various ideas on how to explain low probability events.

Bill was a dedicated member of the Lepidopterists' Society. Among his contributions to the Society was his service on the Executive Council. He served as editor of the Society's journal from 1985 to 1988, an important period of time where he was able to steer the journal back on track in his collected and methodical way. His thoughtful approach to giving critical but kind reviews and his organized system for keeping manuscripts moving served as a model for subsequent editors of the Journal. Bill became a life member of the Society and served as editor of the Society's Memoirs for over a decade, which culminated in an all-consuming effort to complete David Winter's Basic Techniques for Observing and Studying Moths & Butterflies in 2000. Bill put a number of his own projects on hold in order to complete this important manual and ensure the book was published. Thankfully, he was able to complete many of his postponed projects. The last paper for which he read galleys was published just prior to his death in the March 11, 2013 issue of the Journal of the Lepidopterists' Society and another is in press.

Bill encouraged students and young professionals to go to the Lepidopterists' Society annual meetings, which he himself was able to attend for many decades uninterrupted. Bill would often converse with students and new members, taking an interest in their passion for Lepidoptera and making them feel welcome to the Society he loved. For all of these reasons and more, he received of one of the Society's highest honors, the William D. Winter Service Award, at the 2007 meeting in Bakersfield, California.

In the last couple of years of his life, Bill continued to be a presence in the Department of Entomology as often as his health permitted and worked on his manuscripts from home and at his beloved cabin in Ely, Minnesota. His wife Jocelyn persuaded him to substitute his habitual walk with a car ride from their home in a nearby neighborhood. Bill is survived by Jocelyn M. Muggli, his wife of 35 years, and his six adult children. We were privileged and thankful to have the talents, contributions, and scholarly spirit of William E. Miller in the service of entomology and lepidopterology for over six decades. His legacy will live on in his publications, students, and friends.



W. E. Miller 1953. Biological notes of five hymenopterous parasites of pine bud and stem moth in Ohio. Ohio J. Sci. 53: 59–63. Google Scholar


W. E. Miller & R. B. Neiswander . 1955. Biology and control of the European pine shoot moth. Ohio Ag. Exper. Stat. Res. Bull. 760, 31 pp . Google Scholar


W. E. Miller 1955. Biology of Anacampsis innocuella (Zeller), a leafroller on aspen. J. Econ. Entomol. 48: 622–623. Google Scholar


W. E. Miller 1955. Notes on the life cycles of three parasites of the pitch twig moth. Ohio J. Sci. 55: 317–319. Google Scholar


W. E. Miller 1956. The pitch twig moth and its occurrence in Ohio. Ohio Ag. Exper. Stat. Res. Bull. 79, 24 pp. Google Scholar


W. E. Miller 1958. Control of the European pine shoot moth with concentrated DDT sprays. U.S. Dept. Ag. Lake States For. Exp. Stat. Tech. Note 543, 2 pp. Google Scholar


W. E. Miller 1958. Ecological notes on the pine midges Retinodiplosis resinicola (Osten Sacken) and R. inopis (Osten Sacken) in Southern Ohio (Diptera, Itonididae). Ohio J. Sci. 58: 231–235. Google Scholar


W. E. Miller & S. A. Altmann . 1958. Ecological observations on the Virginia pitch-nodule moth, Petrova wenzeli (Kearfott), including a note on its nomenclature. Ohio J. Sci. 58: 273–281. Google Scholar


W. E. Miller & D. L. Haynes . 1958. Timing DDT sprays in the spring for European pine shoot moth control. U.S. Dept. Ag. Lake States For. Exp. Stat. Tech. 542, 2 pp. Google Scholar


W. E. Miller 1959. A unique new North American species of pinecone-feeding Laspeyresia related to L. ingens Heinrich (Lepidoptera, Olethreutidae). Florida Entomol. 42: 131–134. Google Scholar


W. E. Miller 1959. Natural history notes on the goldenrod ball gall fly, Eurosta solidaginis (Fitch), and on its parasites Eurytoma obtusiventris Gahan and E. gigantea. J. Tenn. Acad. Sci. 34: 246–251. Google Scholar


W. E. Miller 1959. Petrova houseri, a new pitch-nodule moth from eastern North America. Ohio J. Sci. 59: 230–232. Google Scholar


W. E. Miller 1959. Preliminary study of European pine shoot moth parasitism in Lower Michigan. J. Econ. Entomol. 52: 768–769. Google Scholar


W. E. Miller 1959. Spread of forest Lepidoptera. Proceedings of the 14th Annual Meeting of the N. Cen. Bran. Entomol. Soc. Amer. 14: 11. [abstract] Google Scholar


W. E. Miller & R. B. Neiswander . 1959. The pitch pine tip moth and its occurrence in Ohio. Ohio Ag. Exper. Stat. Res. Bull 840, 23 pp. Google Scholar


W. E. Miller & H. J. Heikkenen . 1959. The relative susceptibility of eight pine species to European pine shoot moth attack in Michigan. J. Forest. 57: 912–914. Google Scholar


W. E. Miller 1960. A new pine tip moth (Olethreutidae) from the Gulf of Mexico region. J. Lepid. Soc. 14: 231–236. Google Scholar


W. E. Miller 1960. Discovery in Michigan of Orgilus obscurator (Nees), a foreign parasite of the European pine shoot moth. J. Econ. Entomol. 53: 318. Google Scholar


H. J. Heikkenen & W. E. Miller . 1960. European pine shoot moth damage as related to red pine growth. U.S. Dept. Ag., Lake States For. Exp. Stat. Tech. Note 83, 12 pp. Google Scholar


W. E. Miller & J. L. Arend . 1960. European pine shoot moth damage reduced to sheared Christmas trees. U.S. Dept. Ag., Lake States For. Exp. Stat. Tech. Note 592, 2 pp. Google Scholar


W. E. Miller 1960. The European pine shoot moth: relationship between proportion of trees infested and number of insects per tree. J. Forest. 58: 647–648. Google Scholar


W. E. Miller 1960. Unusual occurrence of the Virginia pitch-nodule moth, Petrova wenzeli (Kearfott) in Canada (Lepidoptera: Olethreutidae). Can. Entomol. 92: 879. Google Scholar


W. E. Miller & D. L. Haynes . 1961. Experiments with concentrated DDT sprays for European pine shoot moth suppression in forest plantations. J. Econ. Entomol. 54: 1014–1018. Google Scholar


W. E. Miller 1962. Differential population levels of the European pine shoot moth, Rhyacionia buoliana, between Europe and North America. Ann. Entomol. Soc. Amer. 55: 672–675. Google Scholar


R. L. Talerico , H. J. Heikkenen & W. E. Miller . 1963. Growth of red pine trees after chemical suppression of the European pine shoot moth. Can. Entomol. 95: 522–524. Google Scholar


W. E. Miller & C. H. Schallau . 1963. Problems in identifying old European pine shoot moth damage. J. Forest. 61: 677–678. Google Scholar


W. E. Miller 1963. The goldenrod gall moth Gnorimoschema gallaesolidaginis (Riley) and its parasites in Ohio. Ohio J. Sci. 63: 65–75. Google Scholar


W. E. Miller 1963. The shortleaf pitch-blister moth, Petrova houseri Miller. Ohio J. Sci. 63: 297–301. Google Scholar


W. E. Miller & L. F. Wilson . 1964. Composition and diagnosis of the pine tip moth infestations in the Southeast. J. Econ. Entomol. 57: 722–726. Google Scholar


W. E. Miller 1965. Number of branchlets on red pine in young plantations. For. Sci. 11: 42–49. Google Scholar


W. E. Miller 1965. Pine tip moths in Central America. Turrialba 15: 58. Google Scholar


W. E. Miller 1965. Protecting Christmas tree plantations. J. Forest. 63: 849–852. Google Scholar


W. E. Miller 1966. A new species of moth destructive to pine cones in Mexico (Tortricoidea). J. Lepid. Soc. 20: 251–253. Google Scholar


C. H. Schallau & W. E. Miller . 1966. Intercept site-index equations for red pine damaged by European pine shoot moth. J. Forest. 64: 538–540. Google Scholar


W. E. Miller 1966. Spiders in old insect galls on goldenrod. Ohio J. Sci. 66: 618–619. Google Scholar


W. E. Miller , L. F. Wilson & R. B. Buchman . 1967. Relative infestation of European pine shoot moth on interplanted ponderosa and red pines in Michigan. J. Forest. 65: 818–819. Google Scholar


W. E. Miller 1967. Taxonomic review of the Rhyaciona frustrana group of pine-tip moths, with description of a new species (Olethreutidae). Can. Entomol. 99: 590–596. Google Scholar


W. E. Miller 1967. The European pine shoot moth: ecology and control in the Lake States. For. Sci. Mon. 14, 72 pp. Google Scholar


P. J. Pointing & W. E. Miller . 1967. European pine shoot moth Rhyacionia buoliana (Schiff.), pp. 163–166. In A. G. Davidson & R. M. Prentice (eds.), Important forest insects and diseases of mutual concern to Canada, the United States and Mexico. Department of Forestry and Rural Development, Ottawa, Canada, no. 1180. Google Scholar


L. F. Wilson & W. E. Miller . 1968. How to control European pine shoot moth and pine root collar weevil. U.S. Dept. Ag. For. Serv. North Central Forest Exp. Stat., 6 pp. Google Scholar


W. E. Miller & A. H. Taylor . 1968. How to control red pine cone beetle. U.S. Dept. Ag. For. Serv. North Cen. For. Exp. Stat., 5 pp. Google Scholar


W. E. Miller , A. R. Hastings & V. M. Carolin . 1970. European pine shoot moth. U.S Dept. Ag. For. Serv. North. Area State Priv. Forest. Forest Insect & Disease Leaflet 59. Google Scholar


W. E. Miller 1970. Fernald types of North American Olethreutinae (Lepidoptera: Tortricidae). Proc. Entomol. Soc. Wash. 72: 288–294. Google Scholar


W. E. Miller 1971. Annual variation in starting date of spring feeding by the European pine shoot moth in a Michigan planting. Michigan Entomol. 4: 28–31. Google Scholar


W. E. Miller 1971. Identity of Phaneta refusana (Walker) with a description of a new species (Tortricidae). J. Lepid. Soc. 25: 284–287. Google Scholar


N. D. Addy , H. O. Batzer, W. J. Mattson & W. E. Miller . 1971. Impact of insects on multiple-use values of North Central forests. U.S. Department of Agriculture, Forest Service, North Central Forest Exp. Stat., Research Paper NC-57, 8 pp. Google Scholar


W. E. Miller 1972. Insect interlude in Michigan dune fixation. J. Forest. 70: 387. Google Scholar


W. E. Miller 1973. Clemens types of Olethreutinae (Lepidoptera, Tortricidae). Trans. Amer. Entomol. Soc. 99: 205–234. Google Scholar


W. E. Miller 1973. Insects as related to wood and nut production. U.S. Dept. Ag. Forest Serv. Gen. Tech. Report NC-4, 114 pp . Google Scholar


W. E. Miller 1973. Two previously unrecognized scientific names for the strawberry leafroller (Lepidoptera: Tortricidae). Ann. Entomol. Soc. Amer. 66: 553–554. Google Scholar


W. E. Miller 1974. Identities of taxonomically confused moths of the Eucosma agricolana group and description of a new species (Lepidoptera: Tortricidae). Ann. Entomol. Soc. Amer. 67: 601–604. Google Scholar


W. E. Miller 1976. Biology and taxonomy of three gall forming species of Epiblema (Olethreutidae). J. Lepid. Soc. 30: 50–58. Google Scholar


W. E. Miller 1977. A new species of Laspeyresia from Michigan (Lepidoptera: Olethreutidae). Great Lakes Entomol. 9: 171–172. Google Scholar


W. E. Miller 1977. Two new species of Petrova moths from pine in Southeast Asia (Tortricidae: Olethreutinae). J. Lepid. Soc. 31: 135–138. Google Scholar


W. E. Miller 1977. Uniform genitalia among wing color morphs of olethreutid moths. J. Lepid. Soc. 31: 118. Google Scholar


W. E. Miller 1977. Weights of Polia grandis pupae reared at two constant temperatures (Lepidoptera: Noctuidae). Great Lakes Entomol. 10: 47–49. Google Scholar


W. E. Miller 1977. Wing measure as a size index in Lepidoptera: the family Olethreutidae. Ann. Entomol. Soc. Amer. 70: 253–256. Google Scholar


W. E. Miller , R. F. Wambach & R. A. Anfang . 1978. Effect of past European pine shoot moth infestations on volume yield of polesized red pine. For. Sci. 24: 543–550. Google Scholar


W. E. Miller 1978. Larisa subsolana, a new genus and species of moth from eastern North America (Olethreutidae). J. Lepid. Soc. 32: 256–260. Google Scholar


J. A. Powell & W. E. Miller . 1978. Nearctic pine tip moths of the genus Rhyacionia: Biosystematic Review (Lepidoptera: Tortricidae, Olethreutinae). Forest Service, U.S. Department of Agriculture, Agriculture Handbook No. 514, 51 pp. Google Scholar


W. E. Miller 1978. Petrova pitch-blister moths of North America and Europe: two new species and synopsis (Olethreutidae). Ann. Entomol. Soc. Amer. 71: 329–340. Google Scholar


W. E. Miller 1978. The facile mind: an annotated bibliography for enhancing creativity and problem-solving. U.S. Dept. Ag. For. Serv. North Central Forest Exp. Stat., 13 pp. Google Scholar


W. E. Miller , K. J. Kessler , J. H. Ohman & J. T. Eschle . 1978. Timber quality of northern hardwood regrowth in the Lake States. For. Sci. 24: 247–259. Google Scholar


W. E. Miller 1978. Use of prescribed burning in seed production areas to control red pine cone beetle. Environ. Entomol. 7: 698–702. Google Scholar


W. E. Miller 1979. Fire as an insect management tool. Bull. Entomol. Soc. Amer. 25: 137–140. Google Scholar


W. E. Miller 1979. Identity corrections for two North American Apotomis moths (Tortricidae: Olethreutinae). Great Lakes Entomol. 12: 115–118. Google Scholar


W. E. Miller 1979. Prior name for a Petrova pine moth (Tortricidae). J. Lepid. Soc. 33: 204. Google Scholar


W. E. Miller 1979. The genus Olethreutes: identity corrections and description of a new species (Lepidoptera: Tortricidae: Olethreutinae). Ann. Entomol. Soc. Amer. 72: 232–236. Google Scholar


J. M. Muggli & W. E. Miller . 1980. Instar head widths, individual biomass, and development rate of forest tent caterpillar, Malacosoma disstria (Lepidoptera: Lasiocampidae), at two densities in the laboratory. Great Lakes Entomol. 13: 207–209. Google Scholar


W. E. Miller 1981. Composition of goldenrod (Solidago: Compositae) populations by species in the Upper Midwest. J. Minn. Acad. Sci. 47: 15–17. Google Scholar


W. E. Miller 1982. Grapholita delineana (Walker), a Eurasian hemp moth, discovered in North America. Ann. Entomol. Soc. Amer. 75: 184–186. Google Scholar


W. E. Miller 1983. Decomposition rates of aspen bole and branch litter. For. Sci. 29: 351–356. Google Scholar


W. E. Miller 1983. Eucosmomorpha albersana (Hübner), a Palaearctic species, collected in North America (Tortricidae, Grapholitini). J. Lepid. Soc. 37: 88–89. Google Scholar


W. E. Miller 1983. Genus Phaneta: new synonymies and a new species (Lepidoptera: Tortricidae). Ann. Entomol. Soc. Amer. 76: 98–103. Google Scholar


W. E. Miller 1983. Nearctic Endothenia species: a new synonymy, a misidentification, and a revised status (Lepidoptera: Tortricidae). Great Lakes Entomol. 16: 5–12. Google Scholar


W. E. Miller 1983. New synonymies in Nearctic Dichrorampha (Lepidoptera: Tortricidae). Proc. Entomol. Soc. Wash. 85: 727–733. Google Scholar


R. L. Brown & W. E. Miller . 1983. Valid names of the spruce seed moth and a related Cydia species (Lepidoptera: Tortricidae). Ann. Entomol. Soc. Amer.76: 110–111. Google Scholar


W. E. Miller & M. G. Pogue . 1984. Ragweed borer (Lepidoptera: Tortricidae: Eucosmini): taxonomic implications of an allometric analysis of adult characters. Ann. Entomol. Soc. Amer. 77: 227–231. Google Scholar


W. E. Miller 1984. Review of Dear Lord Rothschild: birds, butterflies, and history, by Miriam Rothschild . Univ. Minn. Dept. Entomol. News. Google Scholar


W. E. Miller 1985. Comments on the proposed conservation of Laspeyresia Hübner, 1825. Z.N. (S.) 2421. Bull. Zool. Nomen. 42: 8–10. Google Scholar


W. E. Miller 1985. Nearctic Epiblema: a new synonymy, a revised identity, and two new species (Lepidoptera: Tortricidae). Great Lakes Entomol. 18: 33–38. Google Scholar


W. E. Miller 1985. Nearctic Eucosma (Lepidoptera: Tortricidae) four new species and three new synonymies. Ann. Entomol. Soc. Amer. 78: 240–247. Google Scholar


W. E. Miller 1985. Nearctic Olethreutes: five new synonymies, two revised statuses, and notes (Lepidoptera: Tortricidae). Proc. Entomol. Soc. Wash. 87: 408–417. Google Scholar


W. E. Miller 1985. Nearctic Rhyacionia pine tip moths: a revised identity and a new species (Lepidoptera: Tortricidae). Great Lakes Entomol. 18: 119–122. Google Scholar


W. E. Miller 1985. Pammene perstructana (Walker) (Lepidoptera: Tortricidae) identified after more than a century. Great Lakes Entomologist 18: 145–147. Google Scholar


W. E. Miller 1985. Review of Forest entomology: ecology and management, by R. N. Coulson and J. A. Witter. For. Sci. 31: 517–518. Google Scholar


W. E. Miller 1986. Epinotia nisella: an unrecorded host and mode of feeding (Lepidoptera: Tortricidae). Great Lakes Entomol. 19: 205–207. Google Scholar


W. E. Miller 1986. New species of the genus Cydia that attack seeds of Mexican conifers (Lepidoptera: Tortricidae), pp. 5–7. In D. Cibrián-Tovar , B. H. Ebel , H. O. Yates & J. T. Mendez-Montiel (eds.), Insectos de conos y semillas de las coniferas de Mexico/Cone and seed insects of the Mexican conifers. U.S. Department of Agriculture, Southeastern Forest Experiment Station (Asheville, N.C.)/Universidad Autónoma Chapingo, SE-40. Google Scholar


W. E. Miller & M. E. Epstein . 1986. Synchronous population fluctuations among moth species (Lepidoptera). Environ. Entomol. 15: 443–447. Google Scholar


W. E. Miller 1986. The species of Pseudexentera (Tortricidae). J. Lepid. Soc. 40: 218–237. Google Scholar


W. E. Miller 1987. A new species of Gretchena (Tortricidae) injurious to planted Neotropical walnut J. Lepid. Soc. 41: 151–153. Google Scholar


W. E. Miller 1987. Change in nutritional quality of detached aspen and willow foliage used as insect food in the laboratory. Great Lakes Entomol. 20: 41–45. Google Scholar


W. E. Miller 1987. Guide to the Olethreutine moths of midland North America (Tortricidae). U.S. Dept. Ag. For. Serv. Ag. Hand. 660, 104 pp. Google Scholar


W. E. Miller 1987. Spruce budworm (Lepidoptera: Tortricidae): role of adult imbibing in reproduction. Environ. Entomol. 16: 1292–1295. Google Scholar


W. E. Miller 1988. European corn borer reproduction: effects of honey in imbibed water. J. Lepid. Soc. 42: 138–143. Google Scholar


W. E. Miller 1988. Two new species of Rhyacionia pine moths from Mexico (Tortricidae: Olethreutinae). J. Lepid. Soc. 42: 236–239. Google Scholar


W. E. Miller 1989. Reproductive enhancement by adult feeding: effects of honeydew in imbibed water on spruce budworm. J. Lepid. Soc. 43: 167–177. Google Scholar


W. E. Miller 1990. A new species of Sonia from eastern North America (Tortricidae). J. Lepid. Soc. 44: 88–90. Google Scholar


W. E. Miller 1990. Body size and diet quality in the genus Cydia (Tortricidae). J. Lepid. Soc. 44: 113–142. Google Scholar


W. E. Miller 1990. Differences between Nearctic Pammene perstructana and its most similar Palearctic relatives (Tortricidae). J. Lepid. Soc. 44: 288–289. Google Scholar


W. E. Miller 1991. Body size in North American Lepidoptera as related to geography. J. Lepid. Soc. 45: 158–168. Google Scholar


G. G. Grant , B. Pendrel , K. N. Slessor, M. Z. Meng & W. E. Miller . 1991. Identification of sex pheromone components for two lepidopteran defoliators, the oak olethreutid leafroller, Pseudexentera spoiliana (Clemens), and the aspen leafroller, Pseudexentera oregonana (Walsingham). Can. Entomol. 123: 1209–1218. Google Scholar


W. E. Miller 1991. Positive relation between body size and altitude of capture site in tortricid moths (Tortricidae). J. Lepid. Soc. 45: 66–67. Google Scholar


W. E. Miller 1993. Review of: Systematics of the Chrysoxena group of genera (Lepidoptera: Tortricidae: Euliini). Ann. Entomol. Soc. Amer. 86: 216–217. Google Scholar


W. E. Miller 1993. Review of: Tortricid Pests: their biology, natural enemies and control. Ann. Entomol. Soc. Amer. 39: 45–46. Google Scholar


L. D. Gibson & W. E. Miller . 1994. Two new synonymies in Nearctic Eucosma (Tortricidae: Olethreutinae). J. Lepid. Soc. 48: 69–71. Google Scholar


W. E. Miller 1995. Beringian Tortricidae: a new synonymy in Epiblema and a recount of Holarctic species. J. Lepid. Soc. 49: 250–252. Google Scholar


W. E. Miller 1995. Jerapowellia burnsorum, a new genus and species of moth from the southwestern United States (Tortricidae: Olethreutinae). J. Lepid. Soc. 49: 24–30. Google Scholar


G. G. Grant & W. E. Miller . 1995. Larval images on lepidopteran Wings—an unrecognized defense mechanism? Amer. Entomol. 41: 44–48. Google Scholar


H. O. Batzer , M. P. Martin , W. J. Mattson & W. E. Miller . 1995. The forest tent caterpillar in aspen strands: distribution and density estimation of four life stages in four vegetation strata. For. Sci. 41: 99–121. Google Scholar


W. E. Miller 1996. Population behavior and adult feeding capability in Lepidoptera. Environ. Entomol. 25: 213–226. Google Scholar


W. E. Miller 1997. Body weight as related to wing measure in hawkmoths (Sphingidae). J. Lepid. Soc. 51: 91–92. Google Scholar


W. E. Miller 1997. Diversity and evolution of tongue length in hawkmoths (Sphingidae). J. Lepid. Soc. 51: 9–31. Google Scholar


W. E. Miller 1997. Intoxicated lepidopterans: how is their fitness affected, and why do they tipple? J. Lepid. Soc. 51: 277–287. Google Scholar


M. Sabourin , R. J. Priest & W. E. Miller . 1997. Nearctic Acleris: resurrection of A. stadiana and a revised identity for A. semiannula (Lepidoptera: Tortricidae). Great Lakes Entomol. 30: 71–78. Google Scholar


J. Jalava & W. E. Miller . 1998. Boreal Olethreutini 1 (Lepidoptera: Tortricidae): new synonymies and Holarctic records. Entomol. Fennica 9: 137–142. Google Scholar


W. E. Miller & B. K. Nayak . 1998. The body size-altitude correlation in Lepidoptera: new findings from tropical Antheraea silk moths (Saturniidae). Bull. Indian Acad. Seric. 2: 16–20. Google Scholar


W. E. Miller 1999. A new synonymy in Dichrorampha that reveals an overlooked immigrant record for North America (Tortricidae). J. Lepid. Soc. 53: 74–75. Google Scholar


W. E. Miller 2000. A comparative taxonomic-natural history study of eight Nearctic species of Gnorimoschema that induce stem galls on Asteraceae, including descriptions of three new species (Lepidoptera: Gelechiidae). Entomol. Soc. Amer., Thomas Say Pub. Entomol., 80 pp. Google Scholar


W. E. Miller & J. Jalava . 2000. Boreal Olethreutini 2 (Tortricidae): wing and genitalia illustrations, a new synonymy and a new Holarctic addition. J. Lepid. Soc. 54: 47–51. Google Scholar


W. E. Miller , R. L. Brown & K. R. Tuck . 2000. Taxonomic clarification of Notocelia rosaecolana (Doubleday) and N. trimaculana (Haworth) (Tortricidae). J. Lepid. Soc. 54: 111–115. Google Scholar


W. E. Miller 2001. A new species of Eucosmomorpha from North America (Tortricidae). J. Lepid. Soc. 55: 81–84. Google Scholar


W. E. Miller & E. H. Lagasa . 2001. First report of Gypsonoma aceriana (Duponchel) (Lepidoptera: Tortricidae), an Old World poplar pest, in North America. Proc. Entomol. Soc. Wash. 103: 1020–1022. Google Scholar


M. Sabourin , W. E. Miller & P. T. Dang . 2002. Hedya salicella (L.), a Palaearctic species, collected in North America (Tortricidae). J. Lepid. Soc. 56: 172–173. Google Scholar


M. Sabourin , W. E. Miller , E. H. Metzler & J. T. Vargo . 2002. Revised identities and new species of Aethes from Midwestern North America (Tortricidae). J. Lepid. Soc. 56: 216–233. Google Scholar


G. G. Grant , P. de Groot , D. Langevin , S. A. Katovich , K. N. Slessor & W. E. Miller . 2002. Sex attractants and seasonal flight patterns for three Eucosoma (Lepidoptera: Tortricidae) species sympatric in eastern pine seed orchards and plantations. Can. Entomol. 134: 391–401. Google Scholar


M. A. Roberts & W. E. Miller . 2003. First report of the Palaearctic Dichrorampha acuminatana (Lienig & Zeller) in North America (Tortricidae). J. Lepid. Soc. 57: 152–153. Google Scholar


B. J. Cooke , W. E. Miller & J. Roland . 2003. Survivorship bias in tree-ring reconstructions of forest tent caterpillar outbreaks using trembling aspen. Tree-Ring Res. 59: 29–36. Google Scholar


W. E. Miller 2004. Host breadth and voltinism in gall-inducing Lepidoptera. J. Lepid. Soc. 58: 44–47. Google Scholar


W. E. Miller 2005. Gall-inducing Lepidoptera, pp. 431–466. In A. Raman , C. W. Schaefer & T. M. Withers (eds.), Biology, ecology, and evolution of gall-inducing arthropods, Volume 2. Science Publishers Inc., Enfield, NH, USA. Google Scholar


W. E. Miller 2005. Extrinsic effects on fecundity-maternal weight relations in capital-breeding Lepidoptera. J. Lepid. Soc. 59: 143–160. Google Scholar


W. E. Miller 2005. Reproductive bulk in capital-breeding Lepidoptera. J. Lepid. Soc. 59: 229–233. Google Scholar


W. E. Miller 2006. Forest tent caterpillar mating, oviposition, and adult congregation at town lights during a northern Minnesota outbreak. J. Lepid. Soc. 60: 156–160. Google Scholar


J. B. Sullivan & W. E. Miller . 2007. Intraspecific body size variation in macrolepidoptera as related to altitude of capture site and seasonal generation. J. Lepid. Soc. 61: 72–77. Google Scholar


W. E. Miller 2011. Temperature-dependent development in capitalbreeding Lepidoptera. J. Lepid. Soc. 65: 227–248. Google Scholar


W. E. Miller 2013. Smallness and bigness: relation of underlying cell size and number to lepidopteran body size. J. Lepid. Soc. 67: 67–69. Google Scholar


W. E. Miller (in press) Phenotypic correlates of genome size in Lepidoptera. J. Lepid. Soc. Google Scholar
Marc E. Epstein, Susan J. Weller, Roger D. Moon, William D. Hutchison, William J. Mattson, Ian V. Macrae, and Todd M. Gilligan "William E. Miller (1930–2013)," The Journal of the Lepidopterists' Society 67(3), 234-239, (1 December 2013).
Published: 1 December 2013
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