Translator Disclaimer
1 June 2012 Coccoloba howardii (Polygonaceae), a new species from Cuba
Author Affiliations +

Coccoloba howardii, a new species from the north-central region of Cuba, is described and illustrated. It appears to be closely related to C. subcordata, from which it differs primarily in the spiral arrangement of the short branches and leaf features such as the petiole inserted at the base of the ochrea, a membranous blade and alternate primary veins not grouped at the base.

See the PDF.


  1. Alain [bro.] 1960: Novedades de la flora cubana XL — Candollea 17: 99–111. Google Scholar

  2. A. Borhidi & O. Muñiz 1971: Combinationes novae florae cubanae I. — Bot. Közlem. 58: 175–177. Google Scholar

  3. A. Borhidi & O. Muñiz 1976 [“1975”]: New plants in Cuba IV. — Acta Bot. Acad. Sci. Hung. 21: 221–230. Google Scholar

  4. A. Borhidi & O. Muñíz 1977 [“1976”]: Plantas nuevas en Cuba V. — Acta Bot. Acad. Sci. Hung. 22: 295–320. Google Scholar

  5. J. Brandbyge 1990: The diversity of micromorphological features in the genus Coccoloba (Polygonaceae). — Nordic J. Bot. 10: 25–44. Google Scholar

  6. J. Brandbyge 1993: Polygonaceae. — Pp. 531–544 in: K. Kubitzki (ed.), The families and genera of vascular plants 2. — Berlin, etc.: Springer. Google Scholar

  7. J. Burke, A. Sánchez, K. Kron & M. Luckow 2010: Placing the woody tropical genera of Polygonaceae: a hypothesis of character evolution and phylogeny. —  Amer. J. Bot. 97: 1377–1390. Google Scholar

  8. R. A. Howard 1949: The genus Coccoloba in Cuba. — J. Arnold Arbor. 30: 388–424. Google Scholar

  9. R. A. Howard 1957a: Studies in the genus Coccoloba, III. The Jamaican species. — J. Arnold Arbor. 38: 81–106. Google Scholar

  10. R. A. Howard 1957b: Studies in the genus Coccoloba, IV. The species from Puerto Rico and the Virgin Islands and from the Bahamas. — J. Arnold Arbor. 38: 211–242. Google Scholar

  11. R. Howard 1958: Studies in the genus Coccoloba, V. The genus in Haiti and the Dominican Republic. — J. Arnold Arbor. 39: 1–48. Google Scholar

  12. IUCN 2001: IUCN Red List categories and criteria: version 3.1. — IUCN Species Survival Commission. IUCN, Gland, Switzerland, and Cambridge, UK, also published at Scholar

  13. León [bro.] & Alain [bro.] 1951: Flora de Cuba 2, Dicotiledóneas: Casuarinaceas a Meliaceas. — Contr. Ocas. Mus. Hist. Nat. Colegio “De La Salle” 10. Google Scholar

  14. D. Mabberley 2008: Mabberley's plant-book, ed. 3. — Cambridge: Cambridge University. Google Scholar

  15. A. Sánchez, T. Schuster & K. Kron 2009: A large-scale phylogeny of Polygonaceae based on molecular data. — Int. J. Pl. Sci. 170: 1044–1055. Google Scholar

© 2012 BGBM Berlin-Dahlem.
Idelfonso Castañeda Noa "Coccoloba howardii (Polygonaceae), a new species from Cuba," Willdenowia 42(1), 95-98, (1 June 2012).
Published: 1 June 2012

Back to Top