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1 November 2005 Phylogenetic Studies of the Subgenus Nialoe (s. lat.) (Coleoptera, Carabidae, Genus Pterostichus), Part 2: The Asymmetricus Species Group
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Abstract

Pterostichus (Nialoe) asymmetricus Bates and its allies are revised based on the membranous parts of their genitalia. Four new taxa, P. (N.) ovaliphallus sp. nov., P. (N.) fujimurai ibukiyamanus ssp. nov., P. (N.) shotaroi kiimontanus ssp. nov., and P. (N.) basilobatus sp. nov. are described. A cladistic analysis based on 31 morphological characters reveals that Daisenilaoe (s. str.) Nakane and Straneo is included as one clade within Nialoe (s. str.). The speciation and dispersal process of this species group are also discussed.

Kôji Sasakawa "Phylogenetic Studies of the Subgenus Nialoe (s. lat.) (Coleoptera, Carabidae, Genus Pterostichus), Part 2: The Asymmetricus Species Group," Zoological Science 22(11), 1217-1228, (1 November 2005). https://doi.org/10.2108/zsj.22.1217
Received: 24 March 2005; Accepted: 1 September 2005; Published: 1 November 2005
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