Shota Murakami,^1,*,** Kimiyuki Tsuchiya,² Keisuke Nakata,³ Mana Nishikata,⁴ Kazuhiro Kitada,⁴ Hitoshi Suzuki<sup>1

1</sup>Graduate School of Environmental Science, Hokkaido University, Sapporo, Hokkaido, Japan
²Laboratory of Bioresources, Applied Biology Co. Ltd, Minato-ku, Tokyo, Japan
³Forestry Research Institute, Hokkaido Research Organization, Bibai, Hokkaido, Japan
⁴Graduate School of Life Science, Hokkaido University, Sapporo, Hokkaido, Japan

^*To whom correspondence should be addressed. E-mail: shota-m@g.ecc.u-tokyo.ac.jp
^**Present address: Department of General Systems Studies, Graduate School of Arts and Sciences, The University of Tokyo, Meguro-ku, Tokyo, Japan