Sei-ichiro Takeda,^1,2,* Tatsuo Oshida,³ Masaharu Motokawa,⁴ Shin-ichiro Kawada,⁵ Hideki Endo<sup>1

1</sup>The University Museum, The University of Tokyo 7-3-1 Hongo, Bunkyo-ku, Tokyo 113-0032, Japan
²Graduate School of Agricultural and Life Sciences, The University of Tokyo, 1-1-1, Yayoi, Bunkyo-ku, Tokyo 113-0032, Japan
³Laboratory of Wildlife Biology, Obihiro University of Agriculture and Veterinary Medicine, Obihiro 080-8555, Japan
⁴The Kyoto University Museum, Kyoto University, Kyoto 606-8501, Japan
⁵Department of Zoology, National Museum of Nature and Science, 4-1-1, Amakubo, Tsukuba, Ibaraki 305-0005, Japan

^*To whom correspondence should be addressed. E-mail: seiichirotakeda@um.u-tokyo.ac.jp