Open Access
How to translate text using browser tools
1 December 2012 Siluro-Devonian of Podolia, Ukraine: Paleobiological, Biostratigraphic, and Geochemical Aspects
Hubert Szaniawski
Author Affiliations +

In the southern part of Podolia (southwestern Ukraine), on the steep escarpments of the Dniester River and its tributaries many prominent exposures of the Siluro-Devonian deposits are located. Because of the very complicated political history of the region investigations of them have been conducted irregularly and with long interruptions. Geologists of different nationalities have been involved but for long periods of time they did not have access to the whole territory. Initially the area was mainly studied by Polish geologists (see Szajnocha 1889; Kozłowski 1929), but only in western part of the region. Later on the investigations were conducted by geologists from the Soviet Union, including researchers from the Ukrainian Republic (see Nikifirova and Predtechenskij 1968; Nikiforova et al. 1972; Tsegelnyuk et al. 1983; Drygant 1984). Since the independence of the country (1991) the research has been carried out by Ukrainian workers (see Tsegelnyuk et al. 1983; Gritsenko et al. 1999; Drygant 2000, 2010; Voichyshyn 2011) who in recent years have often been cooperating with geologists from other countries (Uhman et al. 2004; Kaljo et al. 2007, 2012; Skompski et al. 2008; Malkowski et al. 2009; Olempska et al. 2011).

The Siluro-Devonian deposits exposed in Podolia have a total thickness in excess of 900 m, are rich in fossils and represent one of the most complete and continuous sequence of that age in the world. They provide a full record of the transition from the open marine conditions (Silurian) through marginal marine (Lochkovian) to alluvial (Old Red facies).

The main purpose of the reported project was to refine their biostratigraphy and to recognize the regional biotic response to the environmental changes influenced by the global changes in the oceanic biogeochemical cycles. However, exceptionally good preservation of the fossils in some layers also enabled some new, unexpected paleobiological observations.

Three expeditions of the Polish and Ukrainian participants of the project under the field guidance of Daniel Drygant allowed us to make the field investigations and collect samples for the geochemical, paleontological, microfacies and palynofacies investigations from fourteen sections. These samples were used in combination with materials collected during an earlier project involving the same participants sponsored by a NATO grant. Some of the results of the projects were published earlier (Malkowski et al. 2009; Balinski 2010; Olempska et al. 2011) and some are still in preparation.

Despite the great progress in the geological and paleontological recognition of the Podolian Siluro-Devonian, the region still needs comprehensive research of many intervals and still offers very attractive paleobiological perspectives.

The investigations conducted by Grzegorz Racki and co-authors were focused on the biotic response to the global isotopic events—Ireviken (early Wenlock) and Klonk (Silurian—Devonian transition). The authors came to the conclusion that the environmental evolution during the first event was not significantly influenced by the geochemical changes but was caused rather by a regional tectonic regime and eustatic sea-level fluctuations. In contrast, the perturbations of the Klonk event are reflected in carbonate crisis and significant increase of eutrophication, as well as the oxygen deficiency. The S—D environmental changes caused turnover in both groups of fauna—the benthic and pelagic. Some of the stratigraphically important brachiopod and conodont species became extinct. However, after this high stress episode, the Devonian carbonate ecosystem quickly recovered and new lineages of the both groups evolved.

The paper of Daniel Drygant and Hubert Szaniawski is devoted to the evolution of the early Devonian conodonts and the refinement of the Lochkovian biostratigraphy based on the fossils. Especially thoroughly investigated were the lowermost and the uppermost parts of the marine Lochkovian sequence, representing the S—D transitional beds and the uppermost marine Devonian beds preceeding ingression of the Old Red facies. The investigations enabled conodont zonation of the Podolian Lochkovian and its correlation with other regions, as well as the description of two new conodont species of potential importance for stratigraphy.

Paweł Filipiak and co-authors have made palynological and microfacies investigations of the upper part of the two sections in which the uppermost marine deposits occur. The authors stated that the taxonomic diversity of acritarchs decrease along the upper part of the section and that the chitinozoans gradually disappear, while the frequency of leiospheres increase. This shows a regressive environmental change, toward more brackish conditions. Also the changes in limestone microfacies demonstrates progressive transition from the shrinking, marine basin toward the brackish, muddy lagoon.

Victor Voichyshyn and Hubert Szaniawski described, for the first time from Podolia, a dentigerous jaw bones of the earliest jawed fishes—the Acanthodi. One new genus and one new species were established. Besides these, three species are also described in open nomenclature. Certain morphological features are recognized as important for generic and specific diagnosis. Good preservation of the jaw bones is probably caused by secondary mineralization.

Andrzej Baliński presents the first detailed description of brachiopods from the classic and well dated section across the Silurian—Devonian boundary at Dnistrove. The author stated that the latest Silurian brachiopods represent an impoverished and relatively deep-water assemblages whereas the early Lochkovian reveals progressively improved conditions for these sessile shelly faunas. Two species, Duyiu bohemicu and Dnestrina gutta can be regarded as characteristic for the uppermost Silurian. Two new species and one new subspecies are established.

Ewa Olempska described the three-dimensionally preserved colonies of boring ctenostome bryozoans and micro-borings of “fungi”. This is the first record of soft-tissue fossilization of boring bryozoans. The exceptional preservation is resulted by secondary phosphatization. New genus and species are established. Besides, described are phosphatized fungi-like endoliths co-occuring with the bryozoans. The material has been obtained by processing of the limestone samples in acetic acid.

All the results came into being thanks to the cooperation of the Institute of Paleobiology of the Polish Academy of Sciences and State Museum of Natural History of National Academy of Sciences of Ukraine and have been supported by the Polish Ministry of Science and Higher Education (grant no. N N307 057834).



A. Baliński 2010. First colour-patterned strophomenide brachiopod from the earliest Devonian of Podolia, Ukraine. Acta Palaeontologica Polonica 55: 695–700. Google Scholar


D.M. Drygant 1984. Korreluciâ i konodonty silurijskih-nižniedevonskih otloženij Volyno-Podolii. 192 pp. Naukova Dumka, Kiyev. Google Scholar


D.M. Drygant 2000. Lower and Middle Paleozoic of the Volyn'-Podillja margin of the East European Platform and Carpathian Foredeep [in Ukrainian]. Proceedings of the State Natural History Museum, Lviv 15: 24–130. Google Scholar


D.M. Drygant 2010. Devonian Conodonts from South−West Margin of the East European Platform (Volyn'−Podolian Ukraine) [in Ukrainian]. 156 pp. Academperiodyka, Kyiv. Google Scholar


V.P. Gritsenko , A.A. Istchenko , L.I. Konstantinenko , and P.D. Tsegelnjuk 1999. Animal and plant communities of Podolia. In : A.J. Boucot and J.D. Lawson (eds.), Paleocommunities—A Case Study from the Silurian and Lower Devonian , 462–487. Cambridge University Press, Cambridge. Google Scholar


D. Kaljo , V. Grytsenko , T. Martma , and M.A. Mõtus 2007. Three global carbon isotope shifts in the Silurian of Podolia (Ukraine): stratigraphical implications. Estonian Journal of Earth Sciences 56: 205–220. Google Scholar


D. Kaljo , T. Martma , V. Grytsenko , A. Brazauskas , and D. Kaminskas 2012. Přídolí carbon isotope trend and upper Silurian to lowermost De− vonian chemostratigraphy based on sections in Podolia (Ukraine) and the East Baltic area. Estonian Journal of Earth Sciences 61: 162–180. Google Scholar


R. Kozłowski 1929. Les Brachiopodes gothlandiens de la Podolie Polo− naise. Palaeontologia Polonica 1: 1–254. Google Scholar


K. Małkowski , G. Racki , D. Drygant , and H. Szaniawski 2009. Carbon isotope stratigraphy across the Silurian-Devonian transition in Podolia, Ukraine: evidence for a global biogeochemical perturbation. Geologi− cal Magazine 146: 674–689. Google Scholar


O.I. Nikiforova and N.N. Predtechensky 1968. A Guide to the Geological Excursion on Silurian and Lower Devonian Deposits oof Podolia (Mid− dle Dnestr River). Third Interenational Symposium On Silurian—Devo− nian Boundary and Lower and Middle Devonian Stratigraphy. 58 pp. Ministry of Geology of the USSR, Leningrad. Google Scholar


O.I. Nikiforova , N.N. Priedtiechiensky [Priedtiečensky, N.N.], A.F. Abushik [Abušik, A.F.], M.M. Ignatovich [Ignatovič, M.M], T.L. Modza− levskaya [Modzalevskaâ, T.L.], A.Ya. Berger [Berger, A. Â], L.S. Novoselova , Yu.K. Burkov [Burkov, Û.K.] 1972. Opornyj raz− riez silura i nižniego devona Podolii. 262 pp. Nauka, Leningrad. Google Scholar


E. Olempska , D.J. Horne , and H. Szaniawski 2011. First record of preserved soft parts in Paleozoic podocopid (Metacopina) ostracod, Cytherellina submagna: phylogenetic implications. Proceedings of the Royal Society Biological Sciences 279: 564–570. Google Scholar


S. Skompski , P. Łuczyński , D. Drygant , and W. Kozłowski 2008. High− energy sedimentary events in lagoonal successions of the Upper Silu− rian of Podolia, Ukraine. Facies 54: 277–296. Google Scholar


W. Szajnocha 1889. O stratygrafii pokładów sylurskich galicyjskiego Podola. Sprawozdania Komisyi Fizyograficznej Akademii Umiejętno− ści 23: 185–200. Google Scholar


P. Tsegelnyuk , V. Gritsenko , L. Konstantinenko , A. Ishchenko , A. Abushik , N. Kadlets , D. Bogoyavlenskaya, Drygant , K. Zaika−Novatsky , N. Kadlets , G. Kiselev , and V. Sytova 1983. The Silurian of Podolia. The Guide to Excursion. 224 pp. Naukova Dumka, Kiev. Google Scholar


A. Uchman , D. Drygant , M. Paszkowski , S.J. Porêbski , and E. Turnau 2004. Early Devonian trace fossils in marine to non−marine redbeds in Podolia, Ukraine: palaeoenvironmental implications. Palaeogeogra− phy, Palaeoclimatology, Palaeoecology 214: 67–83. Google Scholar


V.K. Voichyshyn 2011. The Early Devonian Armoured Agnathans of Podolia, Ukraine. Palaeontologia Polonica 66: 1–211. Google Scholar
© 2012 H. Szaniawski. This is an open-access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Hubert Szaniawski "Siluro-Devonian of Podolia, Ukraine: Paleobiological, Biostratigraphic, and Geochemical Aspects," Acta Palaeontologica Polonica 57(4), 793-794, (1 December 2012).
Published: 1 December 2012
Back to Top