Bing-Bing Liu,^1,2,* Bin Tian,^3,* Hui Ma,¹ Zhi-Qiang Lu,¹ Qiang Qiu,¹ Kang-Shan Mao,² Jian-Quan Liu<sup>1,**

1</sup>2State Key Laboratory of Grassland Agro-Ecosystem, School of Life Sciences, Lanzhou University, Lanz
²3Key Laboratory for Bio-resources and Eco-environment, College of Life Sciences, Sichuan University,
³4Key Laboratory of Biodiversity Conservation in Southwest China, State Forestry Administration, Sout

^* These authors contributed equally to this work.
^** Author for correspondence: liujq@nwipb.ac.cn