Open Access
Translator Disclaimer
30 June 2021 On records of Dryophilus forticornis Abeille de Perrin, 1875 (Coleoptera: Ptinidae) from Italy and Croatia
Gianluca Nardi
Author Affiliations +

Dryophilus forticornis Abeille de Perrin, 1875 (Coleoptera: Ptinidae) is excluded from the faunas of Italy and Croatia, from where it was erroneously reported in the past.

The object of this note is to discuss the occurrence of Dryophilus forticornis Abeille de Perrin, 1875 in Italy and Croatia, chiefly in view of the forthcoming new edition of the checklist of Italian Bostrichoidea (Nardi, in preparation).

Author's interpolations are given in square brackets.

Dryophilus forticornis Abeille de Perrin, 1875

  • Dryophilus [(Dryophilus)] forticornis Abeille de Perrin, 1875: Abeille de Perrin 1875: 212.

  • Dryophilus forticornis [Abeille de Perrin, 1875]: Kiesenwetter 1877: 94.

  • Dryophilus (Homophthalmus) forticornis Abeille [de Perrin]: Reitter 1883: 122; Heyden 1891: 459.

  • Dryophilus forticornis Abeille [de Perrin]: Ragusa 1896: 90, footnote.

  • Dryophilus [(Dryophilus)] forticornis Abeille [de Perrin]: Schilsky 1898: 12–12a.

  • Dryophilus [(Dryophilus)] forticornis: Schilsky 1899: 36 U.

  • Dryophilus (Dryophilus) forticornis Ab.: Reitter 1901: 9; Winkler 1927: 799; Luigioni 1929: 644; Porta 1929: 431.

  • Dryophilus (Dryophilus in. sp.) forticornis Ab.: Heyden 1906: 427.

  • Dryophilus forticornis Ab.: Pic 1912: 11; Bodenheimer 1932: 58, 1937: 132; Novak 1952: 198; EspaÑOl 1977: 6.

  • Dryophilus (in. sp. [= Dryophilus]) forticornis Ab.-Perr.: Jacobson 1915: 818.

  • Dryophilus forticornis Abeille de Perrin: Halperin & Español 1978: 7.

  • Dryophilus forticornis Abeille de Perrin, 1875: Audisio et al. 1995: 8; Sparacio 1997: 83; Zahradník et al. 2000: 309; Zahradník 2007: 348; Toskina 2013: 39; Nardi 2014: 45, 2015: 90; Audisio et al. 2015; Zahradník 2016: 102; Lompe 2020.

  • Dryophilus forticornis was described on the basis of about fifteen female syntypes from “Jaffa, Beyrouth, Mersina (Syrie)” (Abeille de Perrin 1875: 212); these cities now belong, respectively, to Israel, Lebanon and Asiatic Turkey. Since a lectotype has not been designated, the type locality encompasses the three above cities (ICZN 1999: Article 73.2.3).

  • This species was, until present, recorded from Croatia (cf. Zahradník 2016), Italy, Cyprus, Israel, Lebanon, Syria, and Asiatic Turkey (Zahradník et al. 2000; Zahradník 2007, 2016), whereas the old generic record for “M [= Méditerranéen = Mediterranean Sea]” by Bodenheimer (1937) is arbitrarily extensive.

  • Adults of this species have been collected by beating dead branches and flowering shrubs (Abeille de Perrin 1875) and with light traps (Bodenheimer 1932). No other data are available on its ecology.

  • Kiesenwetter (1877: 94) provided the sole precise Italian locality for this species: “Messina [Sicily]”. Nevertheless, as stated by Ragusa (1896), Kiesenwetter (1877) simply listed the toponyms given in the original description of the species (Abeille de Perrin 1872), but erroneously wrote “Messina” instead of “Mersina (Syrie)”.

  • This correction by Ragusa (1896) was later overlooked, with few exceptions (i.e., Schilsky 1899; Reitter 1901; Bertolini 1904; Jacobson 1915; Halperin & Español 1978; Zahradník 1998), and the species was repeatedly mentioned generically from Sicily (Schilsky 1898; Heyden 1906; PIC 1912; Winkler 1927; Luigioni 1929; Porta 1929; Audisio et al. 1995; Sparacio 1997; Zahradník et al. 2000; Audisio et al. 2015) or Italy (Zahradník 2002; Toskina 2013; Lompe 2020), and it was consequently included also in the Red Lists of Italian saproxylic beetles (Nardi 2014, 2015). In reality, D. forticornis must be excluded from the Italian fauna, since no specimen from Italy has so far been recorded (see above) or examined (F. Angelini & G. Nardi, unpublished data).

  • Zahradník (2016) listed this species also from Croatia, but only on the basis of the catalogues of Reitter (1883) and Heyden (1891), and based on Novak (1952). Reitter (1883) and Heyden (1891) recorded the species only from Dalmatia (without further details); this historical geographic region is currently divided between Croatia, Montenegro and, for a small part, Bosnia and Herzegovina. Nevertheless, in a following edition of one of the above catalogues (Heyden 1906), D. forticornis was listed only from Sicily. Novak (1952) referred to an old (1885) record from the island of Hvar (= Lesina) in Croatian Dalmatia based on a colleague's card index. However, this record of D. forticornis appears incorrect, as this species is absent in other parts of the Balkans (Zahradník 2015, 2016), and the island of Hvar (Croatia) would be a north-western station disjoined from the rest of its range; zoogeography suggests that this species is endemic to the Near East (cf. Vigna Taglianti et al. 1999: 38).

  • Acknowledgements

    I would like to thank Davide Badano (Sapienza Università di Roma, Rome) for the revision of the manuscript, Paolo Audisio (Sapienza Università di Roma, Rome), Leonardo Latella (Museo Civico di Storia Naturale di Verona, Verona), Franco Mason (Verona), Angelo Nardi (Cisterna di Latina, Latina), Roberto Poggi (Museo Civico di Storia Naturale “G. Doria”, Genoa), Laura Spada (Centro Nazionale Carabinieri Biodiversità “Bosco Fontana”, Marmirolo, Mantua) and Dmitry Telnov (Natural History Museum, London), for their constant support. Finally, a special thank you to the recently departed Fernando Angelini (Francavilla Fontana, Brindisi) for his support and encouragement over many years.



    Abeille De Perrin, E. (1875): Synopsis des Dryophilus du bassin de la Méditerranée. – Annales de la Société Entomologique de France (5) 5 : 207–212. Google Scholar


    Audisio, P., Gobbi, G., Liberti, G. & Nardi, G. (1995): Coleoptera Polyphaga IX (Bostrichoidea, Cleroidea, Lymexyloidea). – In: Minelli, A., Ruffo, S. & La Posta, S. (eds.), Checklist delle specie della fauna italiana 54, 27 pp.; Bologna (Calderini). [Available from: Scholar


    Audisio, P., Alonso Zarazaga, M., Slipinski, A., Nilsson, A., Jelínek, J., Vigna Taglianti, A., Turco, F., Otero, C., Canepari, C., Kral, D., Liberti, G., Sama, G., Nardi, G., Löbl, I., Horak, J., Kolibac, J., Háva, J., Sapiejewski, M. (†), Jäch, M., Bologna, M., Biondi, M., Nikitsky, N., Mazzoldi, P., Zahradník, P., Wegrzynowicz, P., Constantin, R., Gerstmeier, R., Zhantiev, R., Fattorini, S., Tomaszewska, W., Rücker, W., Vazquez-Albalate, X., Cassola, F., Angelini, F., Johnson, C., Schawaller, W., Regalin, R., Baviera, C., Rocchi, S., Cianferoni, F., Beenen, R., Schmitt, M., Sassi, D., Kippenberg, H., Zampetti, M., Trizzino, M., Chiari, S., Carpaneto, G., Sabatelli, S. & De Jong, Y. (2015): Fauna Europaea: Coleoptera, 2, (excl. Series Elateriformia, Scarab-aeiformia, Staphyliniformia and superfamily Curculio-noidea). – Biodiversity Data Journal 3: e4750.  Google Scholar


    Bertolini, S. (1904): Catalogo dei Coleotteri d'Italia, 144 pp.; Siena (Rivista Italiana di Scienze Naturali). Google Scholar


    Bodenheimer, F. S. (1932): Studies on the ecology of Palestinean Coleoptera: I. Coleoptera at light-traps. – Bulletin de la Société Royale Entomologique d'Égypte 16 (1–2): 52–65. Google Scholar


    Bodenheimer, F. S. (1937): Prodromus faunae Palestinae. Essai sur les éléments zoogéographiques et historiques du sud-ouest du sous-règne Paléarctique. – Mémoires de l'Institut d'Égypte 33: 1–286. Google Scholar


    Español, F. (1977): Sobre algunos Anobiidae (Col.) de Turquía recogidos por el Dr. W. Wittmer (nota 78). – Mediterránea. Serie de Estudios sobre Biología Terrestre Mediterránea 2: 5–11. Google Scholar


    Halperin, J. & Español, F. (1978): The Anobiidae (Coleoptera) of Israel and Near East. – Israel Journal of Entomology 12: 5–18. Google Scholar


    Heyden, L. Von (1891): Byrrhidae. – In: Heyden, L. Von, Reitter, E. & Weise, J. (eds.): Catalogus coleopterorum Europae, Caucasi et Armeniae rossicae. Auctoribus Dr. L. v. Heyden, E. Reitter & J. Weise cum aliis sociis coleop-terologicis. Edidit Edmund Reitter, pp. 459–466; Berlin (R. Friedländer & Sohn); Mödling (Edmund Reitter); Caen (Revue d'Entomologie). Google Scholar


    Heyden, L. Von (1906): Anobiidae. – In: Heyden, L. Von, Reitter, E. & Weise, J. (eds.): Catalogus coleopterorum Europae, Caucasi et Armeniae rossicae. Auctoribus Dr. L. v. Heyden, E. Reitter & J. Weise cum aliis sociis coleoptero-logicis. Editio secunda. Edidit Edmund Reitter, pp. 423–432; Berlin (R. Friedländer & Sohn); Paskau (Edmund Reitter); Caen (Revue d'Entomologie). Google Scholar


    International Commission on Zoological Nomenclature [ICZN] (1999): International Code of Zoological Nomenclature. Fourth edition adopted by the International Union of Biological Sciences, XXIX+ 306 pp.; London (International Trust for Zoological Nomenclature). Google Scholar


    Jacobson, G. G. (1915): Zhuki Rosii i zapodnoj Evropy, 1024 pp.; St.-Petersburg (A.F. Devrien) (1905–1915). [In Russian.] Google Scholar


    Kiesenwetter, E. A. H. Von (1877): Naturgeschichte der Insecten Deutschlands begonnen von Dr. W.F. Erichson, fortgesetzt von Prof. Dr. H. Schaum, Dr. G. Kraatz, H.v. Kiesenwetter, Julius Weise, Edm. Reitter und Dr. G. Seidlitz. Erste Abthei-lung. Coleoptera. Fünfter Band. Erste Hälfte, 200 pp.; Berlin (Nicolai). Google Scholar


    Lompe, A. (2020): Die Käfer Europas. Ein Bestimmungswerk im Internet. Version: 3.6.2 (27.05.2020). Available from: (accessed 8 June 2020). Google Scholar


    Luigioni, P. (1929): I Coleotteri d'Italia. Catalogo sinonimico-topografico-bibliografico. – Memorie della Pontificia Acca-demia delle Scienze, I Nuovi Lincei (2) 13 : [4 +]1–1159 [+ 1]. Google Scholar


    Nardi, G. (2014): Famiglia Ptinidae (including Anobiinae), appendice 1 e schede tecniche on line ( – In: Audisio, P., Baviera, C., Carpaneto, G. M., Biscaccianti, A. B., Battistoni, A., Teofili, C. & Rondinini, C. (eds.): Lista Rossa IUCN dei Coleotteri saproxilici italiani, pp. 120–122; Rome (Comitato italiano IUCN e Ministero dell'Ambiente e della Tutela del Territorio e del Mare). Google Scholar


    Nardi, G. (2015): Ptinidae. – In: Carpaneto, G. M., Baviera, C., Biscaccianti, A. B., Brandmayr, P., Mazzei, A., Mason, F., Battistoni, A., Teofili, C., Rondinini, C., Fattorini, S. & Audisio, P. (eds.), A Red List of Italian Saproxylic Beetles: taxonomic overview, ecological features and conservation issues (Coleoptera). Fragmenta Entomologica 47 (2): 89–91. Google Scholar


    Novak, P. (1952): Korniaši Jadranskog Primorja (Coleoptera), 521 pp.; Zagreb (Jugoslavenska Akademija Znanosti i Umjetnosti). Google Scholar


    Pic, M. (1912): Anobiidae. – In: Schenkling, S. (ed.), Coleopterorum Catalogus, vol. 10 (1910–1938) pars 48, pp. 1–91; Berlin (W. Junk). Google Scholar


    Porta, A. (1929): Fauna Coleopterorum Italica. III. Diversicornia, 466 pp.; Piacenza (Stabilimento Tipografico Piacen-tino). Google Scholar


    Ragusa, E. (1896): Catalogo ragionato dei Coleotteri di Sicilia. – Il Naturalista Siciliano (N. S.) 1 (1–7): 69–106. Google Scholar


    Reitter, E. (1883): Byrrhidae. – In: Heyden, L. Von, Reitter, E. & Weise, J. (eds.): Catalogus Coleopterorum Europae et Caucasi. Editio Tertia, pp. 122–124; London (E. Janson), Berlin (Nicolai), Paris (L. Buquet). Google Scholar


    Reitter, E. (1901): Analytische Uebersicht der palaearctischen Gattungen und Arten der Coleopteren – Familien: Byrrhidae (Anobiidae) und Cioidae. – Verhandlungen des naturfor-schenden Vereines in Brünn 40, 3–64. Google Scholar


    Schilsky, J. (1898): Die Käfer Europa's. Nach der Natur beschrieben von Dr. H.C. Küster und Dr. G. Kraatz. Fortgesetzt von J. Schilsky. Fünfunddreissigstes Heft, VIII + 100 nrs. [each on 1 to 4 sheets] + [43] pp. [= 355 pp.]; Nürnberg (Bauer und Raspe [Emil Küster]). Google Scholar


    Schilsky, J. (1899): Die Käfer Europa's. Nach der Natur beschrieben von Dr. H.C. Küster und Dr. G. Kraatz. Fortgesetzt von J. Schilsky. Sechsunddreissigstes Heft, IV + 100 nrs. [each on 1 to 5 sheets] + [54] pp. [= 354 pp.]; Nürnberg (Bauer und Raspe [Emil Küster]). Google Scholar


    Sparacio, I. (1997): Mediterraneo 4. Coleotteri di Sicilia. Parte seconda, 208 pp.; Palermo (L'Epos). Google Scholar


    Toskina, I. N. (2013): A new Palaearctic species and keys to species of two genera of the subfamily Dryophilinae (Coleoptera: Ptinidae). – Byulleten' Moskovskogo Obshchestva Ispytatelei Prirody Otdel Biologicheskii 118 (1): 37–42. [In Russian.] Google Scholar


    Vigna Taglianti, A., Audisio, P. A., Biondi, M., Bologna, M. A., Carpaneto, G. M., De Biase, A., Fattorini, S., Piattella, E., Sindaco, R., Venchi, A. & Zapparoli, M. (1999): A proposal for a chorotype classification of the Near East fauna, in the framework of the Western Paleartic region. Biogeo-graphia (N. S.) 20: 31–59. Google Scholar


    Winkler, A. (1927): Anobiidae. – In: Winkler, A. (ed.), Catalogus Coleopterorum regionis palaearcticae. I. A. Cara-boidea, B. Palpicornia, Staphylinoidea, C. Diversicornia. Pars 7, pp. 798–809; Wien (Albert Winkler). Google Scholar


    Zahradník, P. (1998): Anobiidae of Turkey (Coleoptera). – Klapalekiana 34: 263–286. Google Scholar


    Zahradník, P. (2007): Ptinidae (without Gibbiinae and Ptini-nae). – In: Löbl, I. & Smetana, A. (eds.): Catalogue of Palaearctic Coleoptera. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Volume 4, pp. 339–362 Stenstrup (Apollo Books). Google Scholar


    Zahradník, P. (2015): Ptinidae of Greece (Coleoptera: Bostrichoidea) - a commented checklist with descriptions of new species. Folia Heyrovskyana, series A 23 (2): 151–200. Google Scholar


    Zahradník, P. (2016): A check-list of Ptinidae (Coleoptera: Bostrichoidea) of the Balkan Peninsula. Folia Heyrovskyana, Series A 24 (2): 91–140. Google Scholar


    Zahradník, P., Chikatunov, V. & Pavlíček, T. (2000): Catalogue of beetles (Coleoptera) in Israel and adjacent areas. 2. Anobiidae. Klapalekiana 36: 307–319. Google Scholar
    Gianluca Nardi "On records of Dryophilus forticornis Abeille de Perrin, 1875 (Coleoptera: Ptinidae) from Italy and Croatia," Integrative Systematics: Stuttgart Contributions to Natural History 4(1), 215-217, (30 June 2021).
    Received: 21 May 2021; Accepted: 9 June 2021; Published: 30 June 2021
    3 PAGES

    Near East
    Get copyright permission
    Back to Top