Roland Grimm (Fig. 1) was born on 24 March 1948 in Ellwangen (Jagst), a Swabian town in the eastern part of Baden-Württemberg. Already as a child, he developed a strong interest in insects and started to collect butterflies at the age of 14. He finished high school in Ellwangen in 1968 and then carried out his military service until 1969.
Following his interests in natural history, he studied biology at Eberhard-Karls-University in Tübingen from 1970 on, with a focus on zoology and entomology. He dedicated his diploma thesis to mayflies, therein describing the modified internal reproductive organs of Leptophlebiidae, which possess immotile spermatozoa and a derived sperm pump. He accomplished his doctoral thesis in 1983 at the Zoological Institute of Tübingen University under the supervision of Dr. Gerhard Mickoleit. In this important contribution to the phylogeny of mayflies, he compared the abdominal musculature across different families of Ephemeroptera. He subsequently contributed to the faunistics of mayflies in Baden-Württemberg throughout the 1980s. He did not continue to work on mayflies after that, but he is still highly regarded among ephemeropterists for his substantial contributions to the field.
During his early scientific career, Roland had also become interested in darkling beetles, the group for which he would soon become renowned as an eminent taxonomist, with focus on the Palearctic and Oriental regions. His first contribution on Tenebrionidae, a treatise on the darkling beetles of the Greek island Santorini, was published in 1981 in this very journal (Stuttgarter Beiträge zur Naturkunde, Serie A).
From March 1984 to February 1986, ROLAND held a position as scientific trainee at the State Museum of Natural History in Stuttgart. During this time, in March 1985, he visited together with Wolfgang Schawaller the Hungarian Natural History Museum in Budapest, where Zoltán Kaszab, who was the museum's director and a leading world specialist on Tenebrionidae, taught them the first steps in tenebrionid taxonomy.
But Roland's professional career took a turn towards freshwater biology. From 1986 to 1990 he worked as a freelancer providing ecological assessments, followed in 1990–1991 by a position at the Institute of Hydraulic Engineering in Karlsruhe. From 1991 to 1995 he contributed his expertise to the Fisheries Research Centre in Langenargen. Finally, from 1995 until his retirement in 2013 he was education and conservation officer at the Baden-Württemberg State Fisheries Association, from 2005 as office head.
Roland's ultimate passion, however, was for darkling beetles. His first collecting trips led him to Mediterranean countries like southern Italy, Spain, Greece, Turkey, the Canary Islands, Madeira and the adjacent Porto Santo, Malta and the adjacent Gozo, and Cyprus. Subsequently, he visited the Saharan regions of Morocco, Algeria, Niger and Mali, as well as Gambia and Kenya. In recent years he focused his collecting activities in South East Asia and travelled several times to northern Thailand and Borneo (Sabah and Sarawak). Although he usually preferred to plan and carry out fieldwork alone, the three joint collecting trips with Wolfgang Schawaller in SE Asia (Thailand, Sabah, Sarawak) were enjoyable and very successful. When in the field, Roland also enjoyed hunting for good restaurants that served various seafood dishes.
Roland was a successful collector. He focused his collection on Tenebrionidae and collected only a few beetles from other families, mainly as exchange specimens for other coleopterists or as gifts for friends. On his collecting trips he would explore the area for suitable habitats during the day, often by car, particularly looking for old trees and forest remnants. He would turn stones, sift rotten wood and inspect barks and fungi, often also collecting by fogging with mosquito spray. At night he would usually set up a light trap or would walk in the dark with a torch to check the desert floor or trees in forests, parks or along roads.
Grimm's collection of Tenebrionidae is one of the largest private collections of this family and is stored in about 260 large drawers. It was not only expanded through his own collecting trips, but also through specimens obtained from colleagues who allowed him to keep duplicates in exchange for his thorough identifications of their material. Roland's beetle collection contains a large number of type specimens from his various publications. It is now deposited in the State Museum of Natural History in Stuttgart according to his wish. Also stored here is his aquatic collection, with about 30,000 specimens that mainly represent the local mayfly fauna of Baden-Württemberg.
Roland regularly visited the museum in Stuttgart for comparative studies with specimens in his collection, for literature searches in the library, and for discussions with Wolfgang Schawaller about taxonomic problems, doubtful taxa, or joint projects. He was also a regular visitor in Munich (H. J. Bremer collection) and Basel (H. Gebien collection). During the annual German meetings of coleopterists in Beutelsbach (“Deutsches Koleopterologentreffen”) and Munich (“Bayerischer Entomologentag”), he would receive numerous tenebrionid specimens for identification, on loan or as gifts, from museums as well as from amateur coleopterists.
Although he was a somewhat reserved person, Roland was fully integrated and respected in the international team of tenebrionid workers because of his high taxonomic expertise. He cooperated with all of them and was even visited at his home by R Aalbu from Sacramento (USA) and K Ando from Osaka (Japan). He also participated in all of the International Tenebrionoidea Meetings: Munich (2002), Lyon (2005), Phoenix (2013), Mendoza (2015) and Prague (2018). He only missed the last meeting in 2021, which was held as a video conference due to COVID-19 restrictions. ROLAND usually travelled with Wolfgang Sc to these meetings. In connection with the meetings in Phoenix, Arizona and Mendoza, Argentina, the two of them privately organised some additional fieldwork with American colleagues R Aalbu, Gustavo Flores, Warren Steiner and others in California, Utah and Arizona as well as in Argentina and Chile. These trips and the interesting habitats of the New World deserts and Andes mountains were an unforgettable experience for both Roland and Wolfgang.
It was most unfortunate to see the rapid progress of Parkinson's disease, which severely affected ROLAND in the last few years and which prevented him from travel-ling and studying and dissecting beetles. He passed away on September 11, 2021 in his home in Neuenbürg.
Acknowledgements
Thijlbert Strubelt (Neuenbürg) provided valuable information on Roland's life; Martin Lillig (Saarbrücken) kindly corrected and completed the list of publications.
Appendices
List of publications
Grimm, R. (1975): Zum Vorkommen von Carabus catenatus Panz. (Coleoptera, Carabidae) in Württemberg. – Jahreshefte der Gesellschaft für Naturkunde in Württemberg 130: 383–384.
Grimm, R. (1977): Die Sperma-Pumpe der Eintagsfliegen-Art Habrophlebia lauta (Ephemeroptera, Leptophlebiidae). – Entomologica Germanica 3: 337–343.
Grimm, R. (1980): Baetis digitatus Bengtsson, eine für Deutschland neue Eintagsfliegenart, mit weiteren Angaben zur Verbreitung einiger Arten der Familie Baetidae in Baden-Württemberg (Ephemeroptera, Baetidae). – Nachrichtenblatt der Bayerischen Entomologen 29: 118–125.
Grimm, R. (1981): Die Fauna der Ägäis-Insel Santorin. Teil 2. Tenebrionidae (Coleoptera). – Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie) 348: 1–14.
Grimm, R. (1982): Zur Flora und Fauna im Großen Lautertal bei den Riedwiesen. Die Eintagsfliegen. In: Münsingen. Geschichte, Landschaft, Kultur. Festschrift zum Jubiläum des Württembergischen Landeseinigungsvertrages von 1482, pp. 778–783; Sigmaringen (Thorbecke).
Grimm, R. (1982): Schwarzkäfer und andere Insekten. In: Dittrich, P. (ed.): Biologie der Sahara, pp. 104–130; Frankfurt (Chimaira).
Grimm, R. (1983): Schwarzkäfer und andere Insekten. In: Dittrich, P. (ed): Biologie der Sahara, 2. Erweiterte Auflage, pp. 107–141; Frankfurt (Chimaira).
Grimm, R. (1985a): Zur Kenntnis der Tenebrioniden aus Süditalien (Insecta: Coleoptera). – Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie) 379: 1–32.
Grimm, R. (1985b): Vergleichend-anatomische Untersuchung der abdominalen Muskulatur und der Gonoducte männlicher Ephemeroptera-Imagines (Insecta). – Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie) 384: 1–59.
Grimm, R. & Hellmann, V. (1985): Über Cybopiestes csikii Reitter 1917 (Coleoptera: Tenebrionidae). – Entomologische Zeitschrift 95: 299–301.
Grimm, R. (1986a): Tenebrionidae vom Maltesischen Archipel (Insecta: Coleoptera). – Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie) 392: 1–17.
Grimm, R. (1986b): Tenebrionidae aus Niger und Mali (Insecta, Coleoptera). – Mitteilungen der Münchner Entomologischen Gesellschaft 76: 71–78.
Grimm, R. (1986c): Eintagsfliegen aus Baden-Württemberg (Insecta, Ephemeroptera: Siphlonuridae + Caenidae). – Jahreshefte der Gesellschaft für Naturkunde in Württemberg 141: 201–206.
Grimm, R. (1987a): Pachychila denticollis sp. n. aus Südspanien (Coleoptera, Tenebrionidae). – Nachrichtenblatt der Bayerischen Entomologen 36: 21–24.
Grimm, R. (1987b): Beitrag zur Kenntnis der Eintagsfliegenfauna der Iller (Ephemeroptera). – Nachrichtenblatt der Bayerischen Entomologen 36: 95–102.
Grimm, R. (1988a): Zur Eintagsfliegenfauna der Donauzuflüsse Rot und Rauglen (Baden-Württemberg) (Insecta, Ephemeroptera). – Nachrichtenblatt der Bayerischen Entomologen 37: 73–83.
Grimm, R. (1988b): Die männliche Imago von Baetis vardaren-sis Ikonomov 1962 (Ephemeroptera: Baetidae). – Entomologische Zeitschrift 98: 325–328.
Grimm, R. (1989): Eintagsfliegen vom Neckar bei Rottenburg a. N. (Insecta, Ephemeroptera). – Jahreshefte der Gesellschaft für Naturkunde in Württemberg 144: 241–246.
Grimm, R. (1991a): Eine neue Art der Gattung Corinta Koch von der ostafrikanischen Küste (Coleoptera: Tenebrionidae). – Entomologische Zeitschrift 101: 226–229.
Grimm, R. (1991b): Tenebrioniden von der Insel Zypern (Insecta: Coleoptera). – Biocosme Mésogéen 8: 15–49.
Schawaller, W. & Grimm, R. (1992): 83. Familie: Tenebrionidae. Ergänzungen und Berichtigungen. – In LOHSE, G. A. & LUCHT, W. H. (eds.): Die Käfer Mitteleuropas, Suppl. 2, pp. 196–199; Krefeld (Goecke & Evers).
Grimm, R. (1993): Fische und Fischerei im Oberrhein. – Berichte zur Fischereiforschung 3, 66 pp.; Staatliche Lehr- und Versuchsanstalt Aulendorf (Fischereiforschungsstelle).
Hoffmann, R., Berg, R., Blank, S., Dehus, P., Grimm, R. & RÖsch, R. (1995): Fische in Baden-Württemberg – Gefährdung und Schutz, 92 pp.; Stuttgart (Ministerium für Ländlichen Raum, Ernährung und Forsten Baden-Württemberg).
Schawaller, W. & Grimm, R. (1995): Neatus picipes in der Paläarktis – eine Artengruppe (Coleoptera: Tenebrionidae). – Annales Historico-Naturales Musei Nationalis Hungarici 87: 67–73.
Grimm, R. (1995): Beitrag zur Kenntnis der Schwarzkäfer-fauna der Kanarischen Inseln. (Coleoptera, Tenebrionidae). – Mitteilungen der Münchner Entomologischen Gesellschaft 85: 33–50.
Grimm, R. (2000): Die Gattung Eurycaulus Fairmaire, 1868, in Algerien und Tunesien (Coleoptera: Tenebrionidae). – Entomologische Zeitschrift 110: 89–91.
Grimm, R. & Lillig, M. (2000): Erodius graniventris in der westlichen Sahara: eine Artengruppe (Coleoptera, Tenebrionidae). – Bulletin de la Société Entomologique de France 105: 31–35.
Grimm, R. & Schawaller, W. (2000): The genus Colpotus Mulsant & Rey (Coleoptera: Tenebrionidae) in the Eastern Mediterranean Region, with descriptions of two new species. – Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie) 615: 1–15.
Grimm, R. (2001a): Faunistik und Taxonomie einiger Arten der Gattung Tribolium Macleay, 1825, mit Beschreibung von drei neuen Arten aus Afrika (Coleoptera, Tenebrionidae). – Entomofauna 22: 393–404.
Grimm, R. (2001b): Zur Taxonomie von Eurycaulus Fairmaire, 1869, Subgenus Ammotrypes Fairmaire, 1879, stat. n. (Scle-ronimon Reitter, 1904, syn. n. (Col. Tenebrionidae). – Nachrichtenblatt der Bayerischen Entomologen 50: 88–90.
Grimm, R. (2002): Tenebrionidae von Gambia (Insecta, Coleoptera). – Entomofauna 23: 353–380.
Grimm, R. (2003): Die Arten der Gattung Palorinus Blair, mit Beschreibungen von vier neuen Arten (Coleoptera: Tenebrionidae). – Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie) 646: 1–9.
Grimm, R. (2004a): Die Arten der Gattung Ulomina Baudi di Selve, mit Beschreibungen von drei neuen Arten aus Laos und Thailand (Coleoptera: Tenebrionidae). – Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie) 664: 1–10.
Grimm, R. (2004b): Neue Arten der Gattungen Palorinus Blair, 1930 und Paloropsis Masumoto et Grimm, 2004 aus Indien und Thailand (Coleoptera, Tenebrionidae, Palorinae). – Entomologica Basiliensia 26: 293–298.
Masumoto, K. & Grimm, R. (2004): A New Genus and a species of the Palorinae (Coleopter: Tenebrionidae) from Japan. – Entomological Review of Japan 59: 127–130.
Grimm, R. (2005): Taxonomic and faunistic notes on the genus Eurycaulus, with descriptions of two new species from the Arabian Peninsula (Coleoptera: Tenebrionidae). – Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie) 672: 1–11.
Grimm, R. (2008a): A new species of Tribolium Macleay from Namibia. – Stuttgarter Beiträge zur Naturkunde A, Neue Serie 1: 371–374.
Aistleitner, E. & Grimm, R. (2007): Fragmenta entomofaunistica X. Schwarzkäfer vom Madeira-Archipel und von den Kanarischen Inseln (Coleoptera: Tenebrionidae). – Nachrichtenblatt der Bayerischen Entomologen 56: 98–101.
Grimm, R. (2008b): Guanobius borneensis n. gen., n. sp. from Borneo (Coleoptera: Tenebrionidae: Alphitobiini) – Stuttgarter Beiträge zur Naturkunde A, Neue Serie 1: 375–379.
Grimm,R.(2008c):PostandrosusKulzer,1951–sexualdimorphism in P. ater Kulzer, 1952 and new faunistic data (Coleoptera: Tenebrionidae, Stenochiinae, Cnodalonini). – Entomologische Zeitschrift 118: 117–118.
Grimm, R. (2009a): A new species of Postandrosus Kulzer, 1951 and new records of P. maculipennis Kulzer, 1951 from Sarawak (Malaysia, Borneo) (Coleoptera: Tenebrionidae, Stenochiinae, Cnodalonini). – Mitteilungen der Münchner Entomologischen Gesellschaft 99: 17–20.
Grimm, R. (2009b): Two new species of the genus Bradymerus Perroud, 1864 (Coleoptera: Tenebrionidae) from Borneo. – Caucasian Entomological Bulletin 5: 235–236, pl. 7, figs. 1–2.
Grimm, R. (2009c): Peltonotus nasutus Arrow, 1910, und Phaenochrous-Arten als Bestäuber von Amorphophallus paenoiifolius (Araceae) in Thailand (Coleoptera: Scarabaeidae). – Entomologische Zeitschrift 119: 167–168.
Grimm, R. (2010): New and little known species of Tenebrionidae (Coleoptera) from Borneo. – Stuttgarter Beiträge zur Naturkunde A, Neue Serie 3: 257–267.
Grimm, R. (2011a): New and little known species of Tenebrionidae (Coleoptera) from Borneo (2). – Stuttgarter Beiträge zur Naturkunde A, Neue Serie 4: 249–257.
Grimm, R. (2011b): Two new species of the genus Triplehornia Matthews & Lawrence from Borneo and the Philippines (Coleoptera: Tenebrionidae). – Stuttgarter Beiträge zur Naturkunde A, Neue Serie 4: 259–261.
Grimm, R. (2011c): Five new species and new records of Malaya-plamius Masumoto, 1986 (Coleoptera: Tenebrionidae: Cnodalonini). – Annales Zoologici 61: 229–235.
Grimm, R. (2011d): A new species of Palorinus (Blair, 1930) from Nilgiri Hills in southern India. – Mitteilungen der Münchner Entomologischen Gesellschaft 101: 69–71.
Grimm, R. (2012a): A new species and new records of Amaryg-mus Dalman from North Maluku (Coleoptera: Tenebrionidae). – Stuttgarter Beiträge zur Naturkunde A, Neue Serie 5: 239–241.
Grimm, R. (2012b): A new species of Ammobius Guérin-Méneville, 1844 from Oman (Coleoptera: Tenebrionidae: Opatrini). – Mitteilungen der Münchner Entomologischen Gesellschaft 102: 37–39.
Grimm, R. (2013a): New and little known species of Tenebrionidae (Coleoptera) from Borneo (3). – Stuttgarter Beiträge zur Naturkunde A, Neue Serie 6: 175–181.
Grimm, R. (2013b): A new species of Sclerum Dejean, 1834 from the United Arab Emirates and Oman (Coleoptera: Tenebrionidae: Opatrini). – Mitteilungen der Münchner Entomologischen Gesellschaft 103: 81–84.
Grimm, R. (2013c): The genus Cryphaeus Klug, 1833 in Australia, with description of a new species (Coleoptera: Tenebrionidae: Toxicini). – Australian Entomologist 49: 243–247.
Grimm, R. (2014a): New and little known species of Tenebrionidae (Coleoptera) from Borneo (4). – Stuttgarter Beiträge zur Naturkunde A, Neue Serie 7: 183–197.
Grimm, R. (2014b): The genus Neopachypterus Bouchard, Löbl & Merkl in Thailand, with descriptions of five new species (Coleoptera: Tenebrionidae: Opatrini). – Mitteilungen der Münchner entomologischen Gesellschaft 104: 85–92.
Schawaller, W. & Grimm, R. (2014): The genus Alphitobius Stephens (Coleoptera, Tenebrionidae, Alphitobiini) in Africa and adjacent islands. – ZooKeys 415: 169–190.
Grimm, R. (2015a): Four new species of the genus Sclerum Dejean (Coleoptera: Tenebrionidae: Opatrini) from South East Asia. – Stuttgarter Beiträge zur Naturkunde A, Neue Serie 8: 207–214.
Grimm, R. (2015b): New and little known species of Tenebrionidae (Coleoptera) from Borneo (5). – Stuttgarter Beiträge zur Naturkunde A, Neue Serie 8: 215–225.
Grimm, R. (2015c): A new species of Corticeus Piller & Mitterpacher from Ethiopia (Coleoptera: Tenebrionidae: Diaperinae: Hypophlaeini). – Mitteilungen der Münchner entomologischen Gesellschaft 105: 45–47.
Grimm, R. (2015d): Tenebrionidae (Insecta: Coleoptera) from Iran. – Vernate 34: 299–318.
Grimm, R. (2015e): Bradymerus triplehorni Grimm (Coleoptera: Tenebrionidae), new species from West Malaysia. – Coleopterists Society Monographs 14: 78–79.
Grimm, R. (2016a): A new species of Promethis Pascoe from West Papua with unusual head armature (Coleoptera: Tenebrionidae: Cnodalonini). – Stuttgarter Beiträge zur Naturkunde A, Neue Serie 9: 181–183.
Grimm, R. (2016b): New and little known species of Tenebrionidae (Coleoptera) from Borneo (6). – Stuttgarter Beiträge zur Naturkunde A, Neue Serie 9: 185–190.
Grimm, R. (2016c): A new species of Heterotarsus Latreille from Thailand (Coleoptera: Tenebrionidae: Opatrini). – Mitteilungen der Münchner Entomologischen Gesellschaft 106: 9–11.
Grimm, R. (2017a): New and little known species of Tenebrionidae (Coleoptera) from Borneo (7). – Stuttgarter Beiträge zur Naturkunde A, Neue Serie 10: 175–180.
Grimm, R. (2017b): Two new species of Ceropria Laporte & Brullé from West Malaysia and Sumatra (Coleoptera: Tenebrionidae: Diaperini). – Stuttgarter Beiträge zur Naturkunde A, Neue Serie 10: 181–183.
Grimm, R. (2017c): New taxonomic and faunistic data on the genus Cneocnemis Gebien, 1914 in Laos and Thailand, with descriptions of two new species (Coleoptera: Tenebrionidae: Ulomini). – Mitteilungen der Münchner Entomologischen Gesellschaft 107: 19–22.
Bremer, H. J. & Grimm, R. (2017): The species of Gonocnemis and Paragonocnemis from Borneo with notes on synonymy in the Oriental species (Coleoptera: Tenebrionidae). – Acta Entomologica Musei Nationalis Pragae 57: 331–343.
Grimm, R. (2018a): New and little known species of Tenebrionidae (Coleoptera) from Borneo (8). – Integrative Systematics 1: 59–70.
Grimm, R. (2018b): New synonyms of Ulomoides (Phayllidius syn. nov.= dispar (Gebien, 1921) (Martianus platypterus Gebien, 1927; syn. nov.) (Coleoptera: Tenebrionidae, Diaperini). – Entomologische Zeitschrift 128: 103–104.
Grimm, R. (2018c): Pedinus (Colpotus) skalei n. sp. (Insecta: Coleoptera: Pedinini) from southern Turkey. – Vernate 37: 307–308.
Nabozhenko, M. V. & Grimm, R. (2018): New and little known species of the tribe Helopini (Insecta: Coleoptera: Tenebrionidae) from Iran. – Vernate 37: 309–315.
Grimm, R. & Merkl, O. (2018): A new species of Thraustocolus Kraatz from Iran (Insecta: Coleoptera: Tentyriini). – Vernate 37: 317–319.
Grimm, R., Smith, A. D., Steiner, W. E. jr. & Wachtel, F. (2018): New synonymy and new combination of Hypogena (Coleoptera, Tenebrionidae). – Spixiana 41: 210.
Lillig, M. & Grimm, R. (2019): Discovery of the male and redescription of Cryphaeus camerunus Ardoin (Coleoptera: Tenebrionidae: Toxicini). – Annales Zoologici 69: 133–136.
Nabozhenko, M. V. & Grimm, R. (2019): New species and new records of darkling beetles of the tribe Helopini (Coleoptera: Tenebrionidae) from the Western Palaearctic. – Caucasian Bulletin of Entomology 15: 107–116.
Schawaller, W. & Grimm, R. (2019): New taxonomic and faunistic data on the genera Cneocnemis GEBIEN and Ulomimus Bates, with description of Cneocnemis baehri sp. n. (Coleoptera: Tenebrionidae: Ulomini). – Mitteilungen der Münchner Entomologischen Gesellschaft 109: 43–50.
Bremer, H. J. & Grimm, R. (2019): A new Corticeus from China (Coleoptera, Tenebrionidae, Hypophlaeini). – Mitteilungen der Münchner Entomologischen Gesellschaft 109: 51–54.
Grimm, R. & Schawaller, W. (2019): Palorus ruthmuellerae sp. n. from Zimbabwe, with distributional data of other African species of Palorus (Coleoptera: Tenebrionidae: Palorini). – Mitteilungen der Münchner Entomologischen Gesellschaft 109: 55–58.
Grimm, R., Lillig, M. & Mifsud, D. (2020): An update for the Tenebrionidae (Coleoptera) of the Maltese Archipelago. – Bulletin of the Entomological Society of Malta 11: 47–56.
Grimm, M. & Lillig (2020a): Description of a new Phaleria Latreille, 1802 from Bali and a new synonym of Phaleria atri-ceps (Lewis, 1894) (Insecta: Coleoptera: Tenebrionidae: Diaperinae: Phaleriini). – Vernate 39: 301– 305.
Grimm, R. & Lillig, M (2020b): Arabammobius gen. n. buetti-keri sp. n. from Oman (Insecta: Coleoptera: Tenebrionidae: Opatrini). – Vernate 39: 307–312.
Grimm, R. & Schawaller, W. (2021): Checklist for the Darkling Beetles of Borneo (Coleoptera: Tenebrionidae s.str.). – Integrative Systematics 2 (2019): 61–82.
Grimm, R. (2021): New and little known species of Tenebrionidae (Coleoptera) from Borneo (9). – Integrative Systematics 4 (accepted).
Grimm, R. & Lillig, M. (2022): Hegeter (Hegeter) merkli sp. nov. (Coleoptera: Tenebrionidae: Tentyriini) from La Gomera (Canary Islands). – Annales Zoologici 72 (1) (accepted).
