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30 November 2023 The First Fossil of Ceratophyus (Coleoptera, Geotrupidae) from the Middle Pleistocene Shiobara Group of Nasushiobara City, Tochigi Prefecture, Japan
Hiroaki Aiba, David Král
Author Affiliations +
Paleontological Research, 28(3):265-272 (2023). https://doi.org/10.2517/PR230013
Figure 1.
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Photographs of Ceratophyus yatagaii sp. nov., holotype, KYFSI209a (female). A, dorsal view; B, enlarged head; C, enlarged right antenna. Abbreviations: aI–VI, antennomere I–VI.
Figure 2.
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Photographs of Ceratophyus yatagaii sp. nov., holotype, KYFSI209 (female). A, ventral view; B, enlarged right hindleg; C, enlarged left mandible; D, enlarged right protibia. A, D, KYFSI209b, conterpart; B, C, KYFSI209a. Abbreviations: T I, apical protibial teeth; T II, subapical protibial teeth; T III, third protibial teeth; T IV, fourth protibial teeth; t I–V, tarsomere I–V.
Figure 3.
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Line drawing of Ceratophyus yatagaii sp. nov., female holotype.
Figure 4.
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Comparison of pronotum and protibia of some females of genus Ceratophyus. A, Ceratophyus yatagaii sp. nov., this study; B, C. sinicus after Zicha et al. (2015); C, C. mesasiaticus after Zoological Institute RAS (2015–2020); D, C. schaffrathi after Hillert and Král (2013); E, C. maghrebinicus after Hillert and Král (2013); F, C. polyceros after Peter (2007–2023); G, C. hoffmannseggi after LMDI (2002–2023); H, C. alloini after LMDI (2002–2023).
Figure 5.
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Map of world distribution of Ceratophyus species. circle, after Král et al. (2020); inverted triangle, after Prunier et al. (2016); square, after Cartwright (1966); star, this study; triangle, after Hillert and Král (2013).
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ARTICLE SOURCE
Paleontological Research
Vol. 28 • No. 3
July 2024
KEYWORDS
Ceratophyus
fossil insect
geotrupidae
Middle Pleistocene
Siobara Group
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