Eurypterid setae and fragments of cuticle are encountered in palynological samples from different areas in Gondwana. In the literature published in the past seven decades, eurypterid remains are recorded as a part of Paleozoic palynological assemblages without giving significance to their biostratigraphic potential. This is because of the absence of a taxonomic classification that allows tracking the occurrence of similar specimens from different samples and regions. In the present study, a rich assemblage of eurypterids is found in the Devonian Ora Formation in the Akkas-3 well in Iraq. The assemblage includes isolated setae and cuticular sheets that still have attached setae. The detached setae show great variation in dimensions including the length, base width and length/width ratios. This is in addition to other important morphological characteristics such as segmentation and flexible ends. The cuticular sheets, in contrast, bear many setae that are identical in shape and size. The observation of cuticular sheets bearing many identical setae is an indication that individual eurypterids produce certain classes of similar setae. This can be used as a basis to establish a taxonomy for the setae after more extensive study. Such a taxonomy will facilitate tracking the setae throughout successive sampling intervals, and their biostratigraphic classification. At least 23 types of setae are documented here from the studied samples in the Ora Formation, with some stratigraphic variations documented for five of them. The results of this study establish a basis for taxonomic nomenclature of eurypterid remains that may benefit biostratigraphic studies in the Paleozoic of Gondwana. The main morphological characteristics of the eurypterid cuticles and setae recorded in the present study are compared to those from other parts of Gondwana. The morphological differences and similarities are documented.
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11 March 2025
Eurypterid Setae and Cuticle Fragments from the Ora Formation (Upper Devonian) of Iraq
Walid Ahmed Makled,
Rahma Sael Al-Auqadi,
Ali Ismail Al-Juboury,
Mohamed Mahmoud El Garhy,
Nasir Alarifi,
Nagham Omar,
Abdelaziz Mahmoud
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Palynology
Vol. 49 • No. 2
May 2025
Vol. 49 • No. 2
May 2025
eurypterid cuticles
eurypterid setae
Morphology of eurypterids
Ora Formation