In the morning of 23 November 2018 Volker Mahnert, former director of the Natural History Museum of Geneva (MHNG) and former editor-in-chief of the Revue suisse de Zoologie, passed away in a Genevan hospital following complications after surgery. His family was at his side when he closed his eyes forever.
Volker Mahnert was born in Innsbruck (Austria) on 3 December 1943, the second of three sons of Klaus and Hanna Mahnert. His father was a businessman in Innsbruck, his mother (from an old family with Anglo- Irish roots) a housewife with a degree in economics.
Volker Mahnert attended grammar school (Bundes- Realgymnasium) in Innsbruck and subsequently intended to study medicine at the University of Innsbruck. Before that, however, he wanted to explore another part of the world – as one does at that age. He went to Hamburg (Germany), found a berth on a cargo ship and worked his passage to the USA. By the time he got back to Innsbruck after four months, he found he had missed the deadline to enrol at the Faculty of Medicine and so he enrolled at the Faculty of Philosophy and studied biology instead. During his student years he greatly enjoyed skiing and playing football (he considered going professional), and he became Tyrolian pole vault champion as a member of the Innsbruck athletics club. He was still keen on exploring also other parts of the world, and between semesters in 1965 he had the opportunity to visit British relatives of his mother in Nairobi, Kenya, where they ran a business. This family contact proved useful for expeditions to Kenya in later years. Being a zoologist in the making, he wanted to know how best to make scientific use of his first visit to a tropical country. What kind of animals should he collect there? Bernd Hauser (then assistant professor at the Institute of Zoology of Innsbruck University) suggested the pseudoscopions, because Prof. Max Beier (1903–1979) at the Natural History Museum in Vienna would certainly be interested in them. Volker Mahnert followed this advice, and a fascination developed that lasted for the rest of his life.
In 1969 to 1970 he was employed as an assistant at the Institute of Zoology in Innsbruck whilst also working on a doctoral thesis on the ecto- and entoparasites of small mammals of his native Austrian province of Tyrol under the supervision of Prof. Heinz Janetschek (1913–1997).
Volker Mahnert in the park of Schloß Matzen (Matzen Castle; Tyrol, Austria), which was owned by his Anglo-Irish greatgrandmother Frances M. C. “Fanny” Grohman, née Reade (1831–1908), a distant relative of the 1st Duke of Wellington, famous for defeating Napoleon in the Battle of Waterloo (photo A. Mahnert).
This study also involved an examination of protozoan blood parasites, which made it necessary to repeatedly consult with Dr Heinz E. Krampitz at the Institute of Tropical Medicine in Munich (Germany). In 1971 Volker Mahnert received his doctorate from the Faculty of Science of Innsbruck University. At the beginning of 1971 he was engaged by the Natural History Museum of Geneva as the curator of the Department of Herpetology and Ichthyology, succeeding Villy Aellen (1926–2000) who then became director of the museum. Volker Mahnert moved to Geneva with his fiancée, Anka Fiedler, and they married here on 22 May 1971. Together they have a son and a daughter, Jan (born 1973) and Annick (born 1975). Upon arrival in Geneva, Volker Mahnert had no knowledge at all of the French language yet, but he learned it fast and he learned it well.
During his time as curator (1971–1989) he collected animals in the region and also went on several official scientific expeditions further away: Greece in 1971 (together with Bernd Hauser and Ivan Löbl), 1972 (with B. Hauser), 1973 (with B. Hauser and I. Löbl); Kenya in 1975, 1977 (with Jean-Luc Perret); Ivory Coast in 1980 (with J.-L. Perret); Paraguay in 1979 (with François Baud, Carlo Dlouhy, J.-L. Perret and Claude Vaucher), 1982 (with F. Baud, C. Dlouhy, J.-L. Perret and C. Vaucher), twice in 1985 (with F. Baud and C. Dlouhy; C. Vaucher and C. Dlouhy), 1990 (with Sonia Muller and C. Dlouhy). The Paraguayan expeditions were guided by Carlo J. Dlouhy (1933–2017), a Genevan who lived in that country for many years. Other trips (Italy in 1969, USA in 1998, Australia in 1992, Brazil in 1998, 2006) had different purposes (scientific meetings, Ph.D. defences), but he always took the opportunity to collect. During all these trips he collected not only amphibians, reptiles and fish, as was expected of the curator of the herpetological and ichthyological collections, but also arthropods, and especially pseudoscorpions.
Volker Mahnert on his first zoological expedition as curator at the MHNG; together with Ivan Löbl (left) on the Ionian island of Zakynthos in 1971 (photo B. Hauser).
The years (1971–1989) when he was curator were his most productive in terms of scientific output (see appendix 1), spurred on by an amicable competition with Ivan Löbl (former curator of the Department of Entomology 2) for who could get more publications. It is such a pitty that one of the contestants dropped out of the race much too early. Nevertheless the record is most impressive. Volker Mahnert has published 200 papers on various subjects and on a wide spectrum of animals, including blood-parasitic protozoans, endoparasitic worms and ectoparasitic arthropods, spiders (only peripherally), palpigrades, amphibians, reptilians, fish, mammals and, above all, pseudoscorpions. Two co-operations with close friends can be highlighted: on characiform fish with Jacques Géry (1917–2007) and on Brazilian pseudoscorpions with Joachim U. Adis (1950–2007).
Volker Mahnert chasing aquatic wildlife and being the attraction of village folks on Lefkada Island in 1971 (photo B. Hauser).
Volker Mahnert sorting specimens from a Winkler/Moczarski soil eclector in Paraguay (photo C. Vaucher).
In 1989 Volker Mahnert became director of the museum, again succeeding Villy Aellen, and stayed in office until his retirement in December 2005. He followed the tradition of his predecessors and also became professor of zoology and animal behaviour at the University of Geneva in 1990. He was in the examination committee of numerous students in Innsbruck, Neuchâtel, Geneva, Nancy, Rennes, Paris, São Paulo and Manaus, and he supervised the M. Sc. thesis of 20 students and the Ph. D. thesis of 12 students, three of which (Sonia Fisch-Muller, Raphaël Covain and Lionel Monod) were later employed by the MHNG and one (Juan Montoya-Burgos) by Geneva University.
He pursued this double role (director + professor) until his retirement. Indeed, at that moment he was also interim director of the “Musée d'ethnographie” (from April to December 2005). During all these years he continued to study pseudoscorpions and to publish on them.
Whilst serving as the director of the MHNG from 1989 to 2005, Volker Mahnert was also the editor-in-chief of the Revue suisse de Zoologie, and before that (from 1975 to 1989) its co-editor (together with V. Aellen and F. Baud). Between 1997 and 2005 he also edited the Instrumenta biodiversitatis, the monograph series of the Revue suisse de Zoologie. Additionally he edited the “Catalogue of Pseudoscorpionida to 1988″ by Mark S. Harvey (published by the Manchester University Press in 1990), the “Proceedings of the 13th International Congress of Arachnology, Geneva 3–8 September 1995″ (published as a two volume special issue of the Revue Suisse de Zoologie in 1996), and he co-edited the series “Fauna of Arabia” between 1996 and 2000.
Volker Mahnert was a member (in some cases a very active one) of several scientific societies and commissions: Federal Scientific Commission for the Surveillance of Trade in Animals (between 1975 and 1991); Cantonal Commission for the protection of animals; Cantonal Subcommittee for the Surveillance of Keeping and Trading Animals (vice-president between 1981 and 1991); Swiss Zoological Society (vice-president in 1977, central committee member since 1989); Swiss Entomological Society; Arachnologische Gesellschaft (member of the scientific committee of the Arachnologische Mitteilungen since its foundation in 1990); French Zoological Society; British arachnological Society; Biological Society of Washington; New York Academy of Sciences; European arachnological Society; European Association of Zoological Nomenclature (president between 1992 and 1996); American arachnological Society; International Society for Subterranean Biology (ISSB); Centre International de Documentation arachnologique (CIDA; president from 1989 to 1992); International Society of Arachnology; International Commission of Zoological Nomenclature (ICZN; commissioner from 1989 to 2007).
Volker Mahnert opening the 13th International Congress of Arachnology, which he organized in the MHNG, on 3 september 1995 (photo C. Ratton, MHNG).
A meeting of Tyrolean zoologists in the MHNG: V. Mahnert, Konrad Thaler (1940–2005; middle) and B. Hauser (right) (photo C. Ratton, MHNG).
Years before his retirement he and his wife bought a house on the French side of the border, in Douvaine, where he set up a private little “arachnology lab” on the upper floor and continued his studies on pseudoscorpions till the end. He then had the status of honorary director of the MHNG and associate scientist of the Department of Arthropods and Entomology 1. From time to time he came into the museum and brought pseudoscorpion specimens that he had identified and/or described and that we either placed in the collections of the museum or sent back to other institutions. He only kept a minimum of specimens and literature at home, which made the migration of his lab material to the museum after his death a quite easily manageable undertaking.
Volker Mahnert leaves behind an outstanding scientific legacy: 200 publications (see appendix 1), the descriptions of 20 genera (one genus of fleas, all others pseudoscorpiones) and 350 species/subspecies (6 fleas, 12 fish, all others pseudoscorpions; see appendix 2), and one of the most important pseudoscorpion collections in the world. Originally a fairly modest collection of central European pseudoscorpions composed by Roger de Lessert (1878–1945), V. Mahnert greatly expanded and enriched it with specimens from all over the world and with type specimens through his taxonomic work, through routine identifications and through his extensive scientific network. This excellent collection now contains 1255 nominal species (out of about 3800 species currently known worldwide), 250 of them represented by primary types, plus 87 species that are either new to science or the identity of which is unclear. Apart from this he contributed considerably to the expansion of the fish, mite, lice and flea collections. For the latter his contact with Fritz Peus (1904–1978), whose flea collection is now deposited in the MHNG, was of great importance.
Volker Mahnert in my office, bringing new pseudoscorpions for the MHNG collection (photo P. Schwendinger).
Volker Mahnert was a very gentle person, good-humoured and always with a joke on his lips, never refusing to give advice and information. I never saw him angry and I never met anybody who seriously complained about him. A boss and colleague as one could only wish to have. He was also much liked and respected by fellow scientists in his extensive international network, which is reflected by the fact that there are three animal genera (one genus of pseudoscorpions, the others mites), at least 56 animal species (in the triclads, nematods, tapeworms, earthworms, scorpions, spiders, opilionids, pseudoscorpions, mites, millipedes, diplurans, bugs, beetles, fish, amphibians and snakes) and one mineral named in his honour (see appendix 3). Many more will certainly follow. Volker Mahnert was taken from us in his still productive years – much too early – but his memory will live on far beyond our lifetimes in the scientific results he published and in the scientific collections he amassed.
Acknowledgements
Anka Mahnert, Danielle Decrouez, Sonia Fisch-Muller, Bernd Hauser, Claude Vaucher and Giulio Gardini provided information on and/or photos of V. Mahnert; Christina Lehmann-Graber compiled information on the pseudoscorpion collection of the MHNG; Raphaël Covain checked for fi sh types in the MHNG; Philippe Wagneur helped fi nd photos in the MHNG archive and scanned them; Christelle Mougin clarifi ed some obscure bibliographical references; Mark S. Harvey provided a complete list of pseudoscorpion species described by V. Mahnert; John Hollier checked the English text.
Peter J. Schwendinger
Geneva, 18 February 2019
Appendices
Appendix 1: List of publications
Parasitology
Mahnert V. 1970. Über Ento- und Ektoparasiten von Kleinsäugern der mittleren Ostalpen (Nordtirol). Ph. D. thesis, University of Innsbruck, 139 pp.
Mahnert V. 1970. Trypanosomen aus alpinen Kleinsäugern Tirols (Österreich). Berichte des Naturwissenschaftlich- Medizinischen Vereins in Innsbruck 58: 131–142.
Prokopic J., Mahnert V. 1970. Über Helminthen der Kleinsäuger (Insectivora, Rodentia) Tirols (Österreichs). Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck 58: 143–154.
Arthropoda
Martius C., Höfer H., Verhaagh M., Adis J., Mahnert V. 1994. Terrestrial arthropods colonizing an abandoned termite nest in a floodplain forest of the Amazon River during the flood. Andrias 13: 17–22.
Adis J., Bonaldo A.B., Brescovit A.D., Bertani R., Cokendolpher J.C., Condé B., Kury A.B., Lourenço W.R., Mahnert V., Pinto-da-Rocha R., Platnick N.I., Reddel J.R., Rheims C.A., Rocher L.S., Rowland J.M., Weygoldt P., Woas S. 2003. Arachnida at “Reserva Ducke”, Central Amazonia / Brazil. Amazoniana 17(1/2): 1–14.
Borges P.A.V., Aguiar L., Amaral J., Amorim I.R., André G., Arraiol A., Baz A., Dinis F., Enghoff H., Gaspar C., Ilharco F., Mahnert V., Melo C., Perlira F., Quartan J.A., Ribeira S.P., Ribes J., Serrano A.R.M., Sousa A.B., Strasser R.Z., Vieira L., Vireia V., Vitorinba A., Wunderlich J. 2005. Ranking protected areas in the Azores using standardised sampling of soil epigean arthropods. Biodiversity and Conservation 14: 2029–2060.
Baert L., Mahnert V. 2015. The distribution of the non-araneae and non-acari arachnids of Galapagos. Belgian Journal of Entomology 28: 1–76.
Borges P.A.V., Gaspar C., Crespo L., Rigal F., Cardoso P., Pereira F., Rego C., Amorim I.R., Melo C., Aguiar C., André G., Mendonça E., Ribeiro S.P., Hortal J., Santos A.M., Barcelos L., Enghoff H., Mahnert V., Pita M.T., Ribes J., Baz A., Sousa A.B., Vieira V., Wunderlich J., Parmakelis A., Whittaker R.A., Quartau J.A., Serrano A.R.M., Triantis K.A. 2016. New records and detailed distribution and abundance of selected arthropod species collected between 1999 and 2011 in Azorean native forests. Biodiversity Data Journal 4: 1–84.
Araneae
Mahnert V. 1988. Quelle horreur: des araignées sur timbresposte ! Musées de Genève 281: 6–12.
Palpigradi
Mahnert V., Janetschek H. 1970. Bodenlebende Palpenläufer in den Alpen (Arach., Palpigradida). Oecologia 4: 106–110.
Pseudoscorpiones
Mahnert V. 1972. Neobisium (Blothrus) kwartirnikovi nov. spec. (Pseudoscorpionidea) aus Bulgarien. Archives des Sciences 24: 383–389.
Mahnert V. 1972. Über griechische Pseudoskorpione I: Microcreagris leucadia nov. spec. (Arachnida: Pseudoscorpiones, Neobisiidae). Berichte des Naturwissenschaftlich- Medizinischen Vereins in Innsbruck 59: 51–56.
Mahnert V. 1973. Über griechische Pseudoskorpione II. Höhlenpseudoskorpione (Pseudoscorpiones, Neobisiidae) von Korfu. Revue suisse de Zoologie 80: 207–220.
Mahnert V. 1973. Drei neue Neobisiidae (Arachnida: Pseudoscorpiones) von den Ionischen Inseln (Über griechische Pseudoskorpione III). Berichte des Naturwissenschaftlich- Medizinischen Vereins in Innsbruck 60: 27–39.
Mahnert V. 1974. Über höhlenbewohnende Pseudoskorpione (Neobisiidae, Pseudoscorpiones) aus Süd- und Osteuropa. Revue suisse de Zoologie 81: 205–218.
Mahnert V. 1974. Einige Pseudoskorpione aus Israel. Revue suisse de Zoologie 81: 377–386.
Mahnert V. 1974. Roncus viti n. sp. (Arachnida: Pseudoscorpiones) aus dem Iran. Berichte des Naturwissenschaftlich- Medizinischen Vereins in Innsbruck 61: 87–91.
Mahnert V. 1974. Acanthocreagris nov. gen. mit Bemerkungen zur Gattung Microcreagris (Pseudoscorpiones, Neobisiidae) (Über griechische Pseudoskorpione IV). Revue suisse de Zoologie 81: 845–885.
Mahnert V. 1975. Griechische Höhlenpseudoskorpione. Revue suisse de Zoologie 82: 169–184.
Mahnert V. 1975. Pseudoscorpione von den Maltesischen Inseln. Fragmenta entomologica 11: 185–197.
Mahnert V. 1975. Pseudoskorpione der Insel Réunion und von T.F.A.I. (Djibouti). Revue suisse de Zoologie 82: 539–561.
Mahnert V. 1976. Zur Kenntnis der Gattungen Acanthocreagris und Roncocreagris (Arachnida, Pseudoscorpiones, Neobisiidae). Revue suisse de Zoologie 83: 193–214.
Mahnert V. 1976. Zwei neue Pseudoskorpion-Arten (Arachnida) aus griechischen Höhlen (Über griechische Pseudoskorpione VII). Berichte des Naturwissenschaftlich- Medizinischen Vereins in Innsbruck 63: 177–183.
Mahnert V. 1976. Pseudoscorpions des grottes de la Sardaigne. Fragmenta entomologica 12: 309–316.
Mahnert V. 1977. Zwei neue Pseudoskorpion-Arten (Arachnida, Pseudoscorpiones) aus marokkanischen Höhlen. International Journal of Speleology 8: 375–381.
Mahnert V. 1977. Über einige Atemnidae und Cheliferidae Griechenlands (Pseudoscorpiones). Bulletin de la Société Entomologique Suisse 50: 67–74.
Mahnert V. 1977. Etude comparative des trichobothries de pseudoscorpions au microscope électronique à balayage. Compte rendu des séances de la Société de physique et d'histoire naturelle de Genève (N. S.) 11: 96–99.
Mahnert V. 1977. Pseudoskorpione (Arachnida) aus dem Tien- Shan. Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck 64: 89–95.
Mahnert V. 1977. Zur Verbreitung höhlenbewohnender Pseudoskorpione der iberischen Halbinsel (pp. 21–23). In: Comunicacions del 6è simposium d'Espeleologia (Bioespeleologia), Terrassa, desembre del 1977. Centre excursionista de Terrassa, 160 pp.
Mahnert V. 1978. Spanische Höhlenpseudoskorpione. Miscelanea zoologica (Barcelona) 4: 61–104.
Mahnert V. 1978. Zwei neue Rhacochelifer-Arten aus dem westlichen Mediterrangebiet und Wiederbeschreibung von Chelifer heterometrus L. Koch. Compte rendu des séances de la Société de physique et d'histoire naturelle de Genève (N.S.) 12: 14–24.
Mahnert V. 1978. Pseudoskorpione (ausgenommen Olpiidae, Garypidae) aus Congo-Brazzaville (Arachnida, Pseudoscorpiones). Folia entomologica hungarica 31: 69–133.
Mahnert V. 1978. Zwei neue Dactylochelifer-Arten aus Spanien und Mallorca (Pseudoscorpiones). Eos (Madrid) 52: 149–157.
Mahnert V. 1978. Pseudoskorpione (Arachnida) aus der Höhle Sisco (Korsika). Revue suisse de Zoologie 85: 381–384.
Mahnert V. 1978. Die Pseudoskorpiongattung Toxochernes Beier, 1932. Symposia of the Zoological Society of London 42: 309–315.
Mahnert V. 1978. Weitere Pseudoskorpione (Arachnida Pseudoscorpiones) aus griechischen Höhlen. Annales Musei Goulandris 4: 273–298.
Mahnert V. 1978. Contribution à l'étude de la faune terrestre des îles granitiques de l'archipel des Séchelles (Mission P.L.G. Benoit & J.J. van Mol, 1972). Pseudoscorpiones. Revue zoologique africaine 92: 867– 888.
Mahnert V. 1979. Pseudoskorpione (Arachnida) aus Höhlen der Tü rkei und des Kaukasus. Revue suisse de Zoologie 86: 259–266.
Mahnert V. 1979. Zwei neue Chthoniiden-Arten aus der Schweiz (Pseudoscorpiones). Revue suisse de Zoologie 86: 501–507.
Mahnert V. 1979. Pseudoskorpione (Arachnida) aus dem Amazonas-Gebiet (Brasilien). Revue suisse de Zoologie 86: 719–810.
Mahnert V. 1979. The identity of Microcreagris gigas Balzan (Pseudioscorpiones, Neobisiidae). Bulletin of the British arachnological Society 4: 339–341.
Mahnert V. 1979. L'identité de Olpium chironomum L. Koch (Pseudoscorpions, Neobisiidae). Revue Arachnologique 2: 249–252.
Mahnert V. 1979. Pseudoskorpione (Arachnida) aus Höhlen Griechenlands, insbesondere Kretas. Archives des Sciences 32: 213–233.
Mahnert V. 1980. Zwei neue Chthonius-Arten (Pseudoscorpiones) aus Höhlen Marokkos. Bulletin de la Société Entomologique Suisse 53: 215–219.
Mahnert V. 1980. Pseudoscorpions from the Canary Islands. Entomologica Scandinavica 11: 259–264.
Mahnert V. 1980. Arachnids of Saudi Arabia. Pseudoscorpiones. Fauna of Saudi Arabia 2: 32–48.
Mahnert V. 1980. On the tracks of Balzan in Paraguay. Newsletter of the British arachnological Society 27: 1–12.
Mahnert V. 1981. Pseudoscorpiones (Arachnida) aus Höhlen Italiens, mit Bemerkungen zur Gattung Pseudoblothrus. Le Grotte d'Italia (4. ser.) 8: 21–38.
Mahnert V. 1981. Chthonius (C.) hungaricus sp. n., eine neue Afterskorpion-Art aus Ungarn (Arachnida). Folia entomologica hungarica 41: 279–282.
Mahnert V., Schuster R. 1981. Pachyolpium atlanticum n. sp., ein Pseudoskorpion aus der Gezeitenzone der Bermudas - Morphologie und Ökologie (Pseudoscorpiones: Olpiidae). Revue suisse de Zoologie 88: 265–273.
Mahnert V. 1981. Die Pseudoskorpione (Arachnida) Kenyas. I. Neobisiidae und Ideoroncidae. Revue suisse de Zoologie 88: 535–559.
Mahnert V. 1981. Höhlenpseudoskorpione aus Norditalien und der dalmatinischen Insel Krk. Atti e Memorie della Commissione Grotte „E. Boegan” 20: 95–100.
Mahnert V. 1981. Taxonomische Irrwege: Olpium savignyi Simon, O. kochi Simon, O. bicolor Simon (Pseudoscorpiones). Folia entomologica hungarica 42: 95– 99.
Mahnert V. 1982. Mesochelifer ressli n. sp., eine mit Chelifer cancroides (L.) verwechselte Art aus Mitteleuropa (Pseudoscorpiones, Cheliferidae). Veröffentlichungen des Tiroler Landesmuseums Ferdinandeum 61: 47–53.
Mahnert V. 1982. The pseudoscorpion genus Corosoma Karsch, 1879, with remarks on Dasychernes Chamberlin, 1929 (Pseudoscorpiones: Chernetidae). The Journal of Arachnology 10: 11–14.
Mahnert V. 1982. Die Pseudoskorpione (Arachnida) Kenyas II. Feaellidae; Cheiridiidae. Revue suisse de Zoologie 89: 115–134.
Mahnert V. 1982. Sigles trichobothriaux chez les pseudoscorpions (Arachnida) (pp. 185–192). In: Tongiorgi P., Balsamo M. (eds). VIe Colloque d'Arachnologie d'Expression Française. Colloque International Européen. Memorie Atti della Societa Toscana di Scienze Naturali, Pisa (Ser. B) Suppl. 88: 1–259.
Mahnert V. 1982. Die Pseudoskorpione (Arachnida) Kenyas. 3. Olpiidae. Monitore zoologico italiano Suppl. 16(11): 263– 304.
Mahnert V. 1982. Die Pseudoskorpione (Arachnida) Kenyas. IV. Garypidae. Annales historico-naturales Musei nationalis hungarici 74: 307–329.
Mahnert V. 1982. Die Pseudoskorpione (Arachnida) Kenyas V. Chernetidae. Revue suisse de Zoologie 89: 691–712.
Mahnert V. 1982. Neue höhlenbewohnende Pseudoskorpione aus Spanien, Malta und Griechenland (Arachnida, Pseudoscorpiones). Bulletin de la Société Entomologique Suisse 55: 297–304.
Mahnert V. 1983. Die Pseudoskorpione Kenyas. VI. Dithidae (Arachnida). Revue zoologique africaine 97: 141–157.
Mahnert V. 1983. Die Pseudoskorpione Kenyas. VII. Miratemnidae und Atemnidae. Revue suisse de Zoologie 90: 357–398.
Mahnert V. 1983. The genus Caffrowithius Beier, 1932, with the description of a new genus from South Africa (Arachnida, Pseudoscorpiones). Annals of the Natal Museum 25: 501–510.
Mahnert V. 1983. Pseudoscorpiones from the Hortobagy National Park (Arachnida) (pp. 361–363). In: Mahunka S. (ed.). The Fauna of the Hortobágy National Park, vol. 2. Akadémiai Kiadó, Budapest, 489 pp.
Mahnert V. 1984. Forschungen an der Somalilandkü ste. Am Strand und auf den Dü nen bei Sar Uanle. 36. Pseudoscorpiones (Arachnida). Monitore zoologico italiano Suppl. 19: 43–66.
Mahnert V. 1984. Beitrag zu einer besseren Kenntnis der Ideoroncidae (Arachnida: Pseudoscorpiones), mit Beschreibung von sechs neuen Arten. Revue suisse de Zoologie 91: 651–686.
Mahnert V. 1984. Pseudoscorpions (Arachnida) récoltés durant la mission spéologique espagnole au Pérou en 1977. Revue arachnologique 6: 17–28.
Mahnert V. 1985. Roncus (Parablothrus) comasi, espèce nouvelle d'une grotte de la Tunisie (Pseudoscorpiones, Neobisiidae). Speleon 26–27: 17–20.
Harvey M.S., Mahnert V. 1985. Olpium L. Koch, 1873 (Arachnida, Pseudoscorpionida Olpiidae): proposed designation of type species and related problems. Z.N. (S.) 2484. Bulletin of Zoological Nomenclature 42: 85–88.
Mahnert V. 1985. Pseudoscorpions (Arachnida) from the lower Amazon region. Revista Brasileira de Zoologia 29: 75–80.
Mahnert V. 1985. Weitere Pseudoskorpione (Arachnida) aus dem zentralen Amazonasgebiet (Brasilien). Amazoniana 9: 215–241.
Adis J., Mahnert V. 1986. On the natural history and ecology of Pseudoscorpiones (Arachnida) from an Amazonian blackwater inundation forest. Amazoniana 9: 297–314.
Mahnert V. 1986. Die Pseudoskorpione (Arachnida) Kenyas. VIII. Chthoniidae. Revue suisse de Zoologie 92: 823–843.
Mahnert V. 1986. Arthropodes épigés du massif de « San Juan de la Pena » (Jaca, Huesca). III. Pseudoscorpions. Pirineos 124: 73–86.
Mahnert V. 1986. Parachernes gracilimanus n. sp., espèce nouvelle de Pseudoscorpion (Arachnida, Chernetidae) de l'Equateur. Revue suisse de Zoologie 93: 813–816.
Mahnert V. 1986. Une nouvelle espèce du genre Tyrannochthonius Chamb. des îles Canaries, avec remarques sur les genres Apolpiolum Beier et Calocheirus Chamberlin (Arachnida, Pseudoscorpiones). Mémoires de la Société royale d'entomologie de Belgique 33: 143–153.
Mahnert V., Adis J. 1986. On the occurrence and habitat of pseudoscorpiones (Arachnida) from Amazonian forests of Brazil. Studies on Neotropical Fauna and Environment 20: 211–215.
Mahnert V., Aguiar N.O. 1986. Wiederbeschreibung von Neocheiridium corticum (Balzan, 1890) und Beschreibung von zwei neuen Arten der Gattung aus Sü damerika (Pseudoscorpiones, Cheiridiidae). Bulletin de la Société Entomologique Suisse 59: 499–509.
Morais J.W., Adis J., Mahnert V. 1986. Abundancia e distribuiçao de pseudoscorpiones do solo numa floresta neotropical nao inundada (p. 258). Abstracts, Congresso Brasileiro de Zoologia, Cuiaba-Mato Grosso, 2.-7.II.1986.
Halperin J., Mahnert V. 1987. On some bark-inhabiting Pseudoscorpiones (Arachnida) from Israel. Israel Journal of Entomology 21: 127–128.
Mahnert V. 1987. Neue oder wenig bekannte, vorwiegend mit Insekten vergesellschaftete Pseudoskorpione (Arachnida) aus Sü damerika. Bulletin de la Société Entomologique Suisse 60: 403–416.
Mahnert V., Adis J., Bührnheim P.F. 1987. Key to the families of Amazonian Pseudoscorpiones (Arachnida) (in English, German and Portuguese). Amazoniana 10: 21–40.
Adis J., Mahnert V., Morais J.W., Rodrigues J.M.G. 1988. Adaptation of an Amazonian pseudoscorpion (Arachnida) from dryland forests to inundation forests. Ecology 69: 287–291.
Leclerc P., Mahnert V. 1988. A new species of the genus Levigatocreagris Curcic (Pseudoscorpiones: Neobisiidae) from Thailand, with remarkable sexual dimorphism. Bulletin of the British arachnological Society 7(9): 273– 277.
Mahnert V. 1988. Die Pseudoskorpione (Arachnida) Kenyas. Familien Withiidae und Cheliferidae. Tropical Zoology 1: 39–89.
Mahnert V. 1988. Zwei neue Garypininae-Arten (Pseudoscorpiones: Olpiidae) aus Afrika und Bemerkungen zu den Gattungen Serianus Chamberlin und Paraserianus Beier. Stuttgarter Beiträge zur Naturkunde (Ser. A, Biol.) 420: 1–11.
Mahnert V. 1988. Neobisium carcinoides (Hermann, 1804) (Pseudoscorpionida, Neobisiidae) - une espèce polymorphe ? (pp. 161–174). In: Carard A. (ed.). Comptes rendu du Xe Colloque Européen d'Arachnologie (1988). Bulletin de la Société Scientifique de Bretagne, Rennes 59, Suppl. 1: 1–221.
Adis J., Mahnert V. 1989. Vertical distribution and abundance of Pseudoscorpiones (Arachnida) in the soil of a Neotropical secondary forest during the dry season and rainy season (p. 1). Abstracts, XI. International Congress of Arachnology, 1989, Turku.
Mahnert V. 1989. Les pseudoscorpions (Pseudoscorpiones, Arachnida) récoltés pendant la campagne biospéologique 1987 à Minorque. Endins 14–15: 85–87.
Mahnert V. 1989. Les pseudoscorpions (Arachnida) des grottes des Iles Canaries, avec description de deux espèces nouvelles du genre Paraliochthonius Beier. Mémoires de biospéologie 16: 41–46.
Adis J., Mahnert V. 1990. On the species composition of Pseudoscorpiones (Arachnida) from Amazonian dryland and inundation forests in Brazil. Revue suisse de Zoologie 97: 49–53
Adis J., Mahnert V. 1990. Vertical distribution and abundance of Pseudoscorpion species (Arachnida) in the soil of a Neotropical secondary forest during the dry and the rainy season. Acta zoologica fennica 190: 11–16.
Mahnert V. 1990. Deux nouvelles espèces du genre Pseudoblothrus Beier, 1931 (Pseudoscorpiones, Syarinidae) des Açores (Portugal). Vieraea 18: 167–170.
Mahnert V. 1991. Pseudoscorpiones of the Batorliget Nature Reserves (NE Hungary) (pp. 683–684). In: Mahunka S. (ed.). The Batorliget Nature Reserves - after forty years. Hungarian Natural History Museum, Budapest, 848 pp.
Mahnert V. 1991. Pseudoscorpions (Arachnida) from the Arabian Peninsula. Fauna of Saudi Arabia 12: 171–199.
Adis J., Mahnert V. 1993. Vertical distribution and abundance of pseudoscorpion species in the soil of two different Neotropical primary forests during the dry and rainy season. In: Raven R.R. (ed.). Proceedings of the XIIth International Congress of Arachnology, Brisbane, 1992. Memoirs of the Queensland Museum 33(2): 431–440.
Mahnert V. 1993. XXII. Els Pseudoscorpins (Arachnida, Pseudoscorpiones). In: Alcover J.A., Ballestros E., Fornes J.J. (eds). Historia Natural de l'Arxipelag de Cabrera. Monografies de la Societat d'Historia natural de les Balears 2: 355–360.
Mahnert V. 1993. Pseudoskorpione (Arachnida: Pseudoscorpiones) von Inselns des Mittelmeers und des Atlantiks (Balearen, Kanarische Inseln, Madeira, Ascension), mit vorwiegend subterraner Lebensweise. Revue suisse de Zoologie 100(4): 971–992.
Mahnert V. 1994. New chernetid pseudoscorpions (Pseudoscorpionida: Chernetidae) from Venezuela and Brazil, with remarks on the genus Ancalochernes Beier. Revue suisse de Zoologie 101(3): 829–838.
Mahnert V., Horak P. 1995. Distribution and ecology of pseudoscorpions (Arachnida: Pseudoscorpiones) in relictforests in Styria (Austria). Bolletino della Accademia Gioenia di Scienze Naturali Catania 26(345): 245–252.
Mahnert V. 1997. New species and records of pseudoscorpions (Arachnida, Pseudoscorpiones) from the Canary Islands. Revue suisse de Zoologie 104(3): 559–585.
Morais J.W., Adis J., Mahnert V., Berti-Filho E. 1997. Abundance and phenology of Pseudoscorpiones (Arachnida) from a mixed water inundation forest in Central Amazonia, Brazil. Revue suisse de Zoologie 104(3): 475–483.
Mahnert V., Andrade R. de 1998. Description of a new troglophilous species of the genus Maxchernes Feio, 1960 (Pseudoscorpiones, Chernetidae) from Brazil (Sao Paulo State). Revue suisse de Zoologie 105: 771–775.
Mahnert V. 2001. Cave-dwelling pseudoscorpions (Arachnida, Pseudoscorpiones) from Brazil. Revue suisse de Zoologie 108(1): 95–148.
Mahnert V., Oromi P. 2001. Pseudoscorpiones (pp. 149–150). In: Izquierdo J., Martin J.L., Zurita N., Arechavaleta M. (eds). Lista de especies silvestres de Canarias (hongos, plantes y animals terrestres). Consejeria de Politica Territorial y Medio Ambiente Gobierno de Canarias, 429 pp.
Mahnert V., Adis J. 2002. 4.6. Pseudoscorpiones (pp. 367–380). In: Adis J. (ed.). Amazonian Arachnida and Myriapoda. Pensoft, Sofia-Moscow, vi + 590 pp.
Mahnert V. 2002. Two new species of pseudoscorpions (Arachnida, Pseudoscorpiones) from caves on Tenerife and La Palma (Canary Islands, Spain), with some new records from the Canary Islands and the Azores (Portugal). Revue suisse de Zoologie 109(4): 777–784.
Andrade R. de, Mahnert V. 2003. Spelaeobochica muchmorei sp. n., a new cavernicolous pseudoscorpion (Pseudoscorpiones: Bochicidae) from Brazil (Sao Paulo State). Revue suisse de Zoologie 110(3): 541–546.
Mahnert V. 2003. Four new species of pseudoscorpions (Arachnida, Pseudoscorpiones: Neobisiidae, Chernetidae) from caves in Yunnan Province, China. Revue suisse de Zoologie 110(4): 739–748.
Mahnert V. 2004. Die Pseudoskorpione Österreichs (Arachnida, Pseudoscorpiones). Denisia 12: 459–471.
Mahnert V., Lopez H., Oromi P. 2004. Pseudoscorpiones (pp. 163–164). In: Izquierdo I., Martin J.L., Zurita N., Arechavaleta M. (eds). Lista de especies silvestres de Canarias (hongos, plantas y animales terrestres). Consejo de Medio Ambiente y Ordenacion Territorial, Gobierno de Canaries, La Laguna, 500 pp.
Mahnert V., Borges P.A.V. 2005. Pseudoscorpiones (p. 173). In: Borges P.A.V., Cunha R., Gabriel R., Frias Martins A., Silva L., Vieira V. (eds). Listagem da faune (Mollusca e Arthropoda) e flora (Bryophyta, Pteridophyta e Spermatophyta) terrestres dos Açores. Direcção Regional do Ambienta e Universidade dos Açores, Horta, Angra do Heroismo e Ponta Delgada, 317 pp.
Harvey M.S., Mahnert V. 2006. The systematic position of the Amazonian species of Albiorix (Pseudoscorpiones, Ideoroncidae). The Journal of Arachnology 34(1): 227–230.
Mahnert V. 2007. Pseudoscorpions (Arachnida: Pseudoscorpiones) from the Socotra Archipelago, Yemen. Fauna of Arabia 23: 271–307.
Gonzalez V.H., Mantilla B., Mahnert V. 2008. A new host record for Dasychernes inquilinus (Arachnida, Pseudoscorpiones, Chernetidae), with an overview of pseudoscorpion-bee relationships. The Journal of Arachnology 35: 470–474.
Mahnert V. 2008. Pseudoscorpiones (p. 279). In: Borges P.A.V., Abreu C., Aguiar A.M.F., Carvalho P., Jardim R., Melo I., Oliveira P., Sérgio C., Serranio A.R.M., Vieira P. (eds). A list of the terrestrial fungio, flora and fauna of Madeira and Selvagens archipelagos. Direcção Regional do Ambienta e Universidade dos Açores, Horta, Angra do Heroismo e Ponta Delgada, 440 pp.
Mahnert V. 2009. Order Pseudoscorpiones. Arthropod fauna of the UAE 2: 26–42.
Mahnert V. 2009. Pseudoscorpiones (Pseudoskorpione) (pp. 501–508). In: Rabitsch W., Essl F. (eds). Endemiten - Kostbarkeiten in Österreichs Pflanzen- und Tierwelt. Naturwissenschaftlicher Verein fü r Kärnten und Umweltbundesamt GmbH, Klagenfurt und Wien, 924 pp.
Mahnert V. 2009. New species of pseudoscorpions (Arachnida, Pseudoscorpiones: Chthoniidae, Chernetidae) from caves in China. Revue suisse de Zoologie 116: 185–201.
Mahnert V. 2009. Attaleachernes gen. nov., a new Chernetid genus from palm trees in the Brazilian Pantanal (Pseudoscorpiones, Chernetidae). Contributions to Natural History 12(2): 921–930.
Mahnert V. 2010. Pseudoscorpiones (p. 198). In: Borges P.A.V., Costa A., Cunha R., Gabriel R., Gonçalves V., Martins A.F., Melo I., Parente M., Raposeiro P., Santos R.S., Silva L., Vieira P., Vieira V. (eds.). A list of the terrestrial and marine biota from the Azores. Principia, Cascais, 432 pp.
Oromi P., Lopez H., Mahnert V. 2010. Diversity and allopatric distribution in the Canarian MSS: a case study in an old island (p. 29). In: Moskiric A., Trontelj P. (eds). Abstract Book, 20th International Conference on Subterranean Biology, Postojna, Slovenia, Aug. 29 - Sept. 3, 2010. Organizing committee, Postojna, 192 pp.
Turienzo P., Di Iorio O., Mahnert V. 2010. Global checklist of pseudoscorpions (Arachnida) found in birds' nests. Revue suisse de Zoologie 117(4): 557–598.
Harvey M.S., Mahnert V. 2011. Case 3533. Neobisium Chamberlin, 1930 (Arachnida, Pseudoscorpiones): proposed precedence over Blothrus Schiödte, 1847. Bulletin of Zoological Nomenclature 68(1): 47–53.
Mahnert V. 2011. A nature's treasury: Pseudoscorpion diversity of the Canary Islands, with the description of nine new species (Pseudoscorpiones, Chthoniidae, Cheiridiidae) and new records. Revista Iberica de Aracnologia 19: 27–45.
Mahnert V. 2011. Pseudoblothrus infernus sp. n. (Pseudoscorpiones, Syarinidae) from the Hölloch cave (Schwyz, Switzerland), with new records of Pseudoblothrus strinatii Vachon from Switzerland and France. Revue suisse de Zoologie 118(1): 11–15.
Mahnert V. 2011. New records of pseudoscorpions from the Juan Fernandez Islands (Chile), with the description of a new genus and three new species of Chernetidae (Arachnida: Pseudoscorpiones). Revue suisse de Zoologie 118(1): 17–29.
Mahnert V., Di Iorio O., Turienzo P., Porta A. 2011. Pseudoscorpions (Arachnida) from Argentina. New records of distributions and habitats, corrections and an identification key. Zootaxa 2881: 1–30.
Ratton P., Mahnert V., Lopes Ferreira R. 2012. A new cavedwelling species of Spelaeobochica (Pseudoscorpiones: Bochicidae) from Brazil. Journal of Arachnology 40(3): 274–280.
Mahnert V. 2013. Nachträge zur Pseudoskorpionfauna (Arachnida: Pseudoscorpiones) der Höhlen der Insel Santorin (Thera) (Kykladen, Griechenland). Revue suisse de Zoologie 120(1): 93–100.
Mahnert V. 2013. The identity of two pseudoscorpion species (Arachnida) described from Guyana by L. di Caporiacco. Arachnology 16(1): 16–20.
Mahnert V. & Weber D. 2013. Pseudoskorpione (Arachnida, Pseudoscorpiones) aus dem Grossherzogtum Luxemburg. Ferrantia 69: 108–114.
Mahnert V. 2014. Pseudoscorpions (Arachnida: Pseudoscorpiones) from the Galapagos Islands (Ecuador). Revue suisse de Zoologie 121(2): 135–210.
Mahnert V., Gardini G. 2014. Cave-inhabiting pseudoscorpion species of the genus Roncus (Pseudoscorpiones: Neobisiidae) from western Greece, including the Ionian Islands. Arachnologische Mitteilungen 48: 28–37.
Mahnert V., Sharaf M., Aldawood A.S. 2014. Further records of pseudoscorpions (Arachnida, Pseudoscorpiones) from Saudi Arabia. Zootaxa 3764(3): 387–393.
Monteiro von Schimonsky D., Bichuette M.E., Mahnert V. 2014. First record of the family Pseudochiridiidae (Arachnida, Pseudoscorpiones) from continental South America - a Pseudochiridium from a Brazilian cave. Zootaxa 3889(3): 442–446.
Harvey M.S., Mahnert V. 2015. A new species of Sphaerowithius (Pseudoscorpiones, Withiidae) from Namibia. African Invertebrates 56(2): 491–497.
Mahnert V. 2016. Re-description of eight species described by Luigi Balzan from South America (Argentina, Brazil, Paraguay) (Pseudoscorpiones: Chernetidae). Opuscula zoologica (Budapest) 47(1): 73–85.
Mahnert V., Li Y.-C. 2016. Cave-inhabiting Neobisiidae (Arachnida: Pseudoscorpiones) from China, with description of four new species of Bisetocreagris Curcic. Revue suisse de Zoologie 123(2): 259–268.
Battirola L.D., dos Santos G.B., Meurer E., Castilho A.C.C., Mahnert V., Brescovit A.D., Marquez M.I. 2017. Soil and canopy Pseudoscorpiones (Arthropoda, Arachnida) in a monodominant forest of Attalea phalerata (Arecaceae) in the Brazilian Pantanal. Studies of Neotropical Fauna and Environment 52: 1–8.
Battirola L.D., Rosado-Neto G.H., Batistella D.A., Mahnert V., Brescovit A.D., Marques M.I. 2017. Vertical and time distribution of Pseudoscorpiones (Arthropoda: Arachnida) in a floodplain forest in the Brazilian Pantanal. Revista de Biologia Tropical (Costa Rica) 65(2): 445–459.
Krajčovičová K., Christophoryová J., Mahnert V. 2017. Rhacochelifer disjunctus (Pseudoscorpiones: Cheliferidae) new to the fauna of Slovakia. Arachnologische Mitteilungen 53: 38–42.
Harvey M.S., Mahnert V. 2019. The status of the Central American pseudoscorpion genus Coprochernes Beier, 1976, a new synonym of Neoallochernes Hoff, 1947 (Pseudoscorpiones: Chernetidae). Arachnology 18(1): 7–13.
Acari
Mahnert V. 1971. Parasitologische Untersuchungen an alpinen Kleinsäugern: Ixodoidea (Acari). Bulletin de la Société Entomologique Suisse 44: 323–332.
Mahnert V. 1971. Parasitologische Untersuchungen an alpinen Kleinsäugern: Parasitische Milben. Revue suisse de Zoologie 78: 909–935.
Anoplura
Mahnert V. 1971. Parasitologische Untersuchungen an alpinen Kleinsäugern: Anoplura (Insecta). Bulletin de la Société Entomologique Suisse 44: 333–341.
Mahnert V. 1975. Quelques parasites (Anoplura, Siphonaptera) de micromammifères nouveaux pour la région de Genève. Bulletin de la Société Entomologique Suisse 47: 327–330.
Buettiker W., Mahnert V. 1978. Vorläufige Liste der Anoplura (Insecta) der Schweiz. Bulletin de la Société Entomologique Suisse 51: 299–306.
Siphonaptera
Mahnert V. 1969. Über Flöhe Tirols (Ins., Siphonaptera). Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck 57: 159–178.
Mahnert V. 1972. Zum Auftreten von Kleinsäuger-Flöhen auf ihren Wirten in Abhängigkeit von Jahreszeit und Höhenstufen. Oecologia 8: 400–418.
Mahnert V. 1976. Alectopsylla unisetosa nov. gen., nov. spec. (Siphonaptera, Ischnopsyllidae) from Argentina. Revue suisse de Zoologie 83: 769–776.
Mahnert V. 1977. A new Ctenophthalmus (Siphonaptera) from Ivory Coast. Revue suisse de Zoologie 84: 869–872.
Mahnert V. 1981. Einige interessante Flöhe (Insecta: Siphonaptera) aus Italien und Afghanistan, aufbewahrt im Museum A. Koenig, Bonn. Bonner zoologische Beiträge 31: 379–383.
Mahnert V. 1982. Two new flea species in the genera Plocopsylla Jordan and Hectopsylla Frauenfeld (Insecta, Siphonaptera) from Argentina. Revue suisse de Zoologie 89: 567–572.
Mahnert V., Beaucournu J.C. 1985. Une nouvelle sousespèce de Typhlopsylla poppei Wagner (Siphonaptera: Hystrichopsyllidae) d'Israël. Revue suisse de Zoologie 92: 205–209.
Beaucournu J.C., Mahnert V. 1988. Un Metastivalius nouveau de Nouvelle-Irlande (Siphonaptera, Pygiopsyllidae). Revue suisse de Zoologie 95: 533–538.
Mahnert V. 2012. 8.34. Ordre Siphonaptera (pp. 338–339). In: Merz B. (ed.). Liste annotée des insectes (Insecta) du Canton de Genève. Instrumenta Biodiversitatis 8: 1–532.
Pisces
Mahnert V., Géry J. 1977. La mâchoire supérieure de certains Barbus s.l. (Pisces, Cypriniformes) africains. Revue suisse de Zoologie 84: 229–236.
Mahnert V., Géry J. 1977. Description d'une espèce et d'une sous-espèce nouvelles d'Afrique occidentale du genre Brycinus (Pisces, Cypriniformes, Characoidei), avec une revue de la super-espèce Brycinus longipinnis (Gü nther, 1864). Revue suisse de Zoologie 84: 617–643.
Mahnert V. 1981. Les poissons des petits cours d'eau du canton de Genève (Suisse). Archives des Sciences 34: 113–124.
Mahnert V., Géry J. 1982. Poissons du bassin de l'Ivindo IX. Notes sur le genre Barbus (Cyprinidae). Revue suisse de Zoologie 89: 461–495.
Géry J., Mahnert V. 1984. Poisson Characoïdes (Ostariophysi) récoltés par Cl. Vaucher en Amazonie péruvienne. Archives des Sciences 37: 171–180.
Mahnert V., Géry J. 1984. Poisson Characoïdes du Paraguay I: Xenurobrycon macropus Myers et Miranda Ribeiro. Revue suisse de Zoologie 91: 497–513.
Mahnert V. 1985. Les poissons du lac. Bulletin de l'association des Amis des Musées de Nyon 8: 10–31.
Mahnert V. 1985. Les candirus ou le danger des baignades exotiques. Musées de Genève 253: 8–12.
Mahnert V. 1985. Families Acipenseridae and Osteoglossidae. Acipenser brevirostrum LeSueur, 1818 … Scleropages formosus Günther, 1864 (pp. 1–10). In: CITES (ed.). CITES identification manual, volume 3: Reptilia, Amphibia, Pisces. Secretariat of the convention, Gland.
Géry J., Mahnert V. 1986. A new rummy-nose tetra from the Rio Negro, Brazil: Hemigrammus bleheri n. sp. (Characidae: Tetragonopterinae), with comments on Paracheirodon. Tropical Fish Hobbyist Magazine 7: 37–52.
Mahnert V. 1986. Les Piranhas. Musées de Genève 269: 6–10.
Géry J., Mahnert V., Dlouhy C. 1987. Poissons Characoidea non Characidae du Paraguay (Pisces, Ostariophysi). Revue suisse de Zoologie 94: 357–464.
Mahnert V. 1987. Paraguay, das wiederentdeckte Fisch- “Paradies”. Das Aquarium 221: 564–570.
Mahnert V., Géry J. 1987. Deux nouvelles espèces du genre Hyphessobrycon (Pisces, Ostariophysi, Characidae) du Paraguay: H. guarani n. sp. et H. procerus n. sp. Bonner zoologische Beiträge 38: 307–314.
Géry J., Mahnert V. 1988. Hydrolycus Mü ller & Troschel, 1844 (Osteichthyes, Cypriniformes): confirmation proposée de Hydrocyon scomberoides Cuvier, 1819 comme espècetype. Bulletin of Zoological Nomenclature 45: 38–40.
Mahnert V., Géry J. 1988. Les genres Piabarchus Myers et Creagrutus Gü nther du Paraguay, avec la description de deux espèces nouvelles (Pisces, Ostraiophysi, Characidae). Revue française d'aquariologie-herpétologie 15: 1–8.
Mahnert V. 1989. Neoceratodus forsteri (Krefft, 1870) (pp. 1–2). In: CITES (ed.). CITES identification manual, volume 3: Reptilia, Amphibia, Pisces. Secretariat of the convention, Gland.
Mahnert V. 1990. Les poissons du Paraguay. Musées de Genève 301: 5–8.
Mahnert V. 1990. Les poissons du Lac. Aquariums du Musée du Léman, Cahier et Catalogues 3: 19–48.
Géry J., Mahnert V. 1992. Notes sur quelques Brycon des bassins de l'Amazone, du Parana-Paraguay et du Sud-Est brésilien (Pisces, Characiformes, Characidae). Revue suisse de Zoologie 99: 793–819.
Weber C., Muller S., Mahnert V. 1992. Harnischwelse Paraguays. DATZ-Sonderheft „Harnischwelse” IX.1992: 10–13.
Géry J., Mahnert V. 1993. Hyphessobrycon pytai n. sp., une nouvelle espèce endémique du bassin du Rio Monday au Paraguay (Pisces, Ostariophysi, Characidae). Revue française d'aquariologie-herpétologie 20(2): 33–36.
Géry J., Mahnert V. 1994. Case 2835. Alestes Mü ller & Troschel, 1844 (Osteichthyes, Characiformes): conservation proposed. Bulletin of Zoological Nomenclature 51(1): 35–40.
Mahnert V., Géry J., Muller S. 1997. Leporinus falcipinnis n. sp., a new species from the lower Tapajos basin, Para (Pisces, Characiformes, Anostomidae). Revue suisse de Zoologie 104(4): 837–844.
Amphibia
Mahnert V. 1986. Family Bufonidae. Atelopus varius zeteki Dunn, 1933 … Nectophrynoides spp. (pp. 1–10). In: CITES (ed.). CITES identification manual, volume 3: Reptilia, Amphibia, Pisces. Secretariat of the convention, Gland.
Keller A., Aellen V., Mahnert V. 1993. Atlas de répartition des amphibiens et reptiles du canton de Genève. Muséum d'histoire naturelle de Genève, 48 pp.
Reptilia
Mahnert V. 1973. Recherches zoologiques dans les îles Ioniennes. Musées de Genève 131: 2–7.
Mahnert V. 1981. Reconnaissez-vous votre sac à main ? Musées de Genève 213: 2–7.
Mahnert V. 1981. Snake skins. Identification aid to entire snake skins (pp. 1–3). In: CITES (ed.). CITES identification manual 5: parts and derivatives 2. Secretariat of the convention, Gland.
Mahnert V. 1981. Snake skins. Acrochordus javanicus Hornstedt, 1787 … Vipera russelli (Shaw,1797) (pp. 1–31). In: CITES (ed.). CITES identification manual 5: parts and derivatives 2. Secretariat of the convention, Gland.
Mahnert V. 1984. Le (petit) monstre du Parc de la Grange. Musées de Genève 243: 7–10.
Mammalia
Mahnert V. 1969. Funde von Myotis oxygnathus (Monticell, 1885) in Nordtirol (Österreich) und in der Schweiz. Revue suisse de Zoologie 76: 1039–1044.
Mahnert V. 1970. Das Vorkommen von Crocidura suaveolens mimula Miller, 1901 (Mammalia, Insectivora) in Nordtirol gesichert. Zeitschrift für Säugetierkunde 35: 104–106.
Museology
Mahnert V. 1976. Catalogue des types de poissons, amphibiens et reptiles du Muséum d'histoire naturelle de Genève. Revue suisse de Zoologie 83: 471–496.
Mahnert V. 1999. The importance, role and advantages of museum collections in scientific research. Abstracts, Conference on Public Collections and Science (World Congress on Science, UNESCO-ICSU-HAS), Budapest, 1–2 July 1999): 46–47.
Mahnert V. 1999. Les collections d'un Musée d'histoire naturelle et la recherche. Verband der Museen der Schweiz (VMS/AMS) - Informationen 63: 43–47.
Orbituaries
Mahnert V. 1980. Professor Dr Max Beier, 1903 - 1979. Bulletin of the British arachnological Society 5: 115–116.
Mahnert V. 1998. Charles-André Vaucher (25 août 1915 - 13 mars 1997). Archives des Sciences 51: 285–287.
Mahnert V. 2000. Villy Aellen (4 décembre 1926 - 22 janvier 2000). Archives des Sciences 53: 59–60.
Mahnert V. 2000. In Memoriam. Villy Aellen, 4 décembre 1926 - 22 janvier 2000. Karstologia 36(2): 57.
Le Bail P.-Y., Weber C., Fisch-Muller S., Mahnert V. 2008. In memoriam Jacques Géry (12 mars 1917 - 15 juin 2007). Cybium 41(4): 397–398.
Mahnert V. 2009. Konrad Thaler 19.12.1940 - 11.6.2005. Contributions to Natural History 12(1): 21–22.
Appendix 2: Taxa described by Volker Mahnert and depository of corresponding type specimens in MHNG. * = secondary types present; ** = primary types present; *** = primary and secondary types present
Genera and subgenera:
Acanthocreagris Mahnert, 1974 (Pseudoscorpiones)
Afroroncus Mahnert, 1981 (Pseudoscorpiones)
Alectopsylla Mahnert, 1976 (Siphonaptera)
Attaleachernes Mahnert, 2009 (Pseudoscorpiones)
Beierochelifer Mahnert, 1977 (Pseudoscorpiones)
Beierowithius Mahnert, 1979 (replacement name for Oligowithius Beier, 1937; Pseudoscorpiones)
Caecatemnus Mahnert, 1985 (Pseudoscorpiones)
Cardiolpium Mahnert 1986 (Pseudoscorpiones)
Ceratochernes Mahnert, 1994 (Pseudoscorpiones)
Halominniza Mahnert, 1975 (Pseudoscorpiones)
Leptocheiridium Mahnert & Schmidl, 2011 (Pseudoscorpiones)
Microblothrus Mahnert, 1985 (Pseudoscorpiones)
Phymatochernes Mahnert, 1979 (Pseudoscorpiones)
Pseudomenthus Mahnert, 2007 (Pseudoscorpiones)
Roncocreagris Mahnert, 1974 (originally a subgenus of Microcreagris; Pseudoscorpiones)
Selachochernes Mahnert, 2001 (Pseudoscorpiones)
Spelaeobochica Mahnert, 2001 (Pseudoscorpiones)
Spelaeochernes Mahnert, 2001 (Pseudoscorpiones)
Sphaerowithius Mahnert, 1988 (Pseudoscorpiones)
Xorilbia Harvey & Mahnert, 2006 (Pseudoscorpiones)
Species and subspecies:
Acanthocreagris aelleni Mahnert, 1978 (Pseudoscorpiones) **
Acanthocreagris beieri Mahnert, 1974 (Pseudoscorpiones) **
Acanthocreagris corcyraea Mahnert, 1976 (Pseudoscorpiones) **
Acanthocreagris corsa Mahnert, 1974 (Pseudoscorpiones) ***
Acanthocreagris leucadia epirensis Mahnert, 1974 (Pseudoscorpiones) ***
Acanthocreagris lycaonis Mahnert, 1978 (Pseudoscorpiones) ***
Acanthocreagris obtusa Mahnert, 1976 (Pseudoscorpiones) ***
Acanthocreagris relicta Mahnert, 1977 (Pseudoscorpiones) *
Acanthocreagris vachoni Mahnert, 1976 [currently in the synonymy of Acanthocreagris ronciformis (Redikorzev, 1949); Pseudoscorpiones] ***
Afrogarypus quadrimaculatus Mahnert, 2007 (Pseudoscorpiones) ***
Afroroncus kikuyu Mahnert, 1981 (Pseudoscorpiones)***
Afroroncus sulcatus Mahnert, 1981 (Pseudoscorpiones)*
Albiorix lamellifer Mahnert, 1985 (Pseudoscorpiones) Albiorix gracilis Mahnert, 1985 (currently in Xorilbia; Pseudoscorpiones)***
Alectopsylla unisetosa Mahnert, 1976 (Siphonaptera)***
Allochernes longepilosus Mahnert, 1997 (Pseudoscorpiones)***
Allochernes maroccanus Mahnert, 1976 (Pseudoscorpiones)***
Allochernes peregrinans Mahnert, 2009 (Pseudoscorpiones)**
Amblyolpium graecum Mahnert, 1976 (Pseudoscorpiones)**
Americhernes bethaniae Mahnert, 1979 (Pseudoscorpiones)*
Americhernes incertus Mahnert, 1979 (Pseudoscorpiones)*
Anaperochernes margaritifer Mahnert, 1985 (Pseudoscorpiones)
Apocheiridium cavicola Mahnert, 1993 (Pseudoscorpiones)**
Apocheiridium lienhardi Mahnert, 2011 (Pseudoscorpiones)***
Apocheiridium pallidum Mahnert, 1982 (Pseudoscorpiones)***
Apolpiolum gigas Mahnert, 1980 (currently in Calocheirus; Pseudoscorpiones)
Attaleachernes thaleri Mahnert, 2009 (Pseudoscorpiones)*
Barbus condei Mahnert & Géry, 1982 (currently in Enteromius; Pisces)***
Barbus prionacanthus Mahnert & Géry, 1982 (currently in Enteromius; Pisces)
Beierolpium benoiti Mahnert, 1978 (Pseudoscorpiones)*
Beierolpium flavum Mahnert, 1984 (Pseudoscorpiones)*
Beierolpium holmi Mahnert, 1982 (Pseudoscorpiones)*
Beierolpium kerioense Mahnert, 1982 (Pseudoscorpiones)***
Beierolpium tanense Mahnert, 1982 (Pseudoscorpiones)***
Beierolpium vanharteni Mahnert, 2007 (Pseudoscorpiones)***
Bisetocreagris baozinensis Mahnert & Li, 2016 (Pseudoscorpiones)*
Bisetocreagris cavernarum Mahnert & Li, 2016 (Pseudoscorpiones)*
Bisetocreagris chuanensis Mahnert & Li, 2016 (Pseudoscorpiones)*
Bisetocreagris juanxuae Mahnert & Li, 2016 (Pseudoscorpiones)*
Brycinus derhami Géry & Mahnert, 1977 (Pisces)***
Brycinus longipinnis bagbeensis Géry & Mahnert, 1977 [currently in the synonymy of Bryconalestes longipinnis (Günther, 1864); Pisces]
Caecatemnus setosipygus Mahnert, 1985 (Pseudoscorpiones)*
Caffrowithius biseriatus Mahnert, 1983 (Pseudoscorpiones)*
Caffrowithius planicola Mahnert, 1982 (Pseudoscorpiones)***
Calocheiridius gracilipalpus Mahnert, 1982 (Pseudoscorpiones)***
Calocheirus gracilis Mahnert, 1991 (Pseudoscorpiones)*
Calocheirus mirus Mahnert, 1986 (Pseudoscorpiones)***
Calocheirus tenerifae Mahnert 2002 (Pseudoscorpiones)
Catatemnus exiguus Mahnert, 1978 (Pseudoscorpiones)***
Catatemnus granulatus Mahnert, 1978 (Pseudoscorpiones)
Ceratochernes granulatus Mahnert, 1994 (Pseudoscorpiones)*
Ceratochernes guanophilus Mahnert, 1994 (Pseudoscorpiones)*
Ceriochernes amazonicus Mahnert, 1985 (Pseudoscorpiones)*
Cheiridium brasiliense Mahnert, 2001 (Pseudoscorpiones)*
Cheiridium perreti Mahnert, 1982 (Pseudoscorpiones)***
Cheiridium somalicum Mahnert, 1984 (Pseudoscorpiones)*
Cheiridium tumidum Mahnert 1982 (Pseudoscorpiones)***
Chelanops gracilipalpus Mahnert, 2011 (Pseudoscorpiones)***
Chelanops gracilipes Mahnert, 2011 (Pseudoscorpiones)**
Chelanops (Neochelanops) peruanus Mahnert, 1984 (currently in genus Neochelanops; Pseudoscorpiones)*
Chthonius (Chthonius) apollinis Mahnert, 1978 (Pseudoscorpiones)***
Chthonius (Chthonius) herbarii Mahnert, 1980 (Pseudoscorpiones)***
Chthonius (Chthonius) hungaricus Mahnert, 1981 (Pseudoscorpiones)*
Chthonius (Chthonius) imperator Mahnert, 1978 (Pseudoscorpiones)***
Chthonius (Chthonius) lucifugus Mahnert, 1977 (Pseudoscorpiones)
Chthonius (Chthonius) ponticoides Mahnert, 1975 (Pseudoscorpiones)**
Chthonius (Chthonius) sestasi Mahnert 1980 (Pseudoscorpiones)***
Chthonius (Chthonius) strinatii Mahnert, 1975 (Pseudoscorpiones)***
Chthonius (Chthonius) thessalus Mahnert, 1980 (Pseudoscorpiones)***
Chthonius (Chthonius) tzanoudakisi Mahnert, 1975 (Pseudoscorpiones)***
Chthonius (Ephippiochthonius) atlantis Mahnert, 1980 (currently in genus Ephippiochthonius; Pseudoscorpiones)**
Chthonius (Ephippiochthonius) balearicus Mahnert, 1977 (currently in genus Ephippiochthonius; Pseudoscorpiones)*
Chthonius (Ephippiochthonius) bellesi Mahnert, 1989 (currently in genus Ephippiochthonius; Pseudoscorpiones)*
Chthonius (Ephippiochthonius) cabreriensis Mahnert, 1993 (currently in genus Ephippiochthonius; Pseudoscorpiones)
Chthonius (Ephippiochthonius) corcyraeus Mahnert, 1976 (currently in genus Ephippiochthonius; Pseudoscorpiones)**
Chthonius (Ephippiochthonius) creticus Mahnert, 1980 (currently in genus Ephippiochthonius; Pseudoscorpiones)***
Chthonius (Ephippiochthonius) daedaleus Mahnert, 1980 (currently in genus Ephippiochthonius; Pseudoscorpiones)***
Chthonius (Ephippiochthonius) dubius Mahnert, 1993 (currently in Occidenchthonius; Pseudoscorpiones)**
Chthonius (Ephippiochthonius) girgentiensis Mahnert, 1982 (currently in genus Ephippiochthonius; Pseudoscorpiones)**
Chthonius (Ephippiochthonius) gracilimanus Mahnert, 1997 (currently in Occidenchthonius; Pseudoscorpiones)*
Chthonius (Ephippiochthonius) longesetosus Mahnert, 1976 (currently in genus Ephippiochthonius; Pseudoscorpiones)**
Chthonius (Ephippiochthonius) lopezi Mahnert, 2011 (currently in Occidenchthonius; Pseudoscorpiones)*
Chthonius (Ephippiochthonius) maltensis Mahnert, 1975 (currently in genus Ephippiochthonius; Pseudoscorpiones)*
Chthonius (Ephippiochthonius) maroccanus Mahnert, 1980 (currently in genus Ephippiochthonius; Pseudoscorpiones)**
Chthonius (Ephippiochthonius) minous minous Mahnert, 1980 (currently in genus Ephippiochthonius; Pseudoscorpiones)***
Chthonius (Ephippiochthonius) minous peramaea Mahnert, 1980 (currently in genus Ephippiochthonius; Pseudoscorpiones)*
Chthonius (Ephippiochthonius) nidicola Mahnert, 1979 (currently in genus Ephippiochthonius; Pseudoscorpiones)***
Chthonius (Ephippiochthonius) nudipes Mahnert, 1982 (currently in genus Ephippiochthonius; Pseudoscorpiones)*
Chthonius (Ephippiochthonius) platakisi Mahnert, 1980 (currently in genus Ephippiochthonius; Pseudoscorpiones)***
Chthonius (Ephippiochthonius) poeninus Mahnert 1979 (currently in Globochthonius; Pseudoscorpiones)***
Chthonius (Ephippiochthonius) ponsi Mahnert, 1993 (currently in genus Ephippiochthonius; Pseudoscorpiones)*
Chthonius (Ephippiochthonius) rimicola Mahnert, 1993 (currently in genus Ephippiochthonius; Pseudoscorpiones)*
Chthonius (Ephippiochthonius) samius Mahnert, 1982 (currently in genus Ephippiochthonius; Pseudoscorpiones)***
Chthonius (Ephippiochthonius) setosus Mahnert, 1993 (currently in Occidenchthonius; Pseudoscorpiones)**
(Ephippiochthonius) tamaran Mahnert, 2011 (currently in Occidenchthonius; Pseudoscorpiones)*
Chthonius (Ephippiochthonius) tenerifae Mahnert, 2011 (currently in Occidenchthonius; Pseudoscorpiones)
Creagrutus paraguayensis Mahnert & Géry, 1988 (Pisces)***
Creagrutus pearsoni Mahnert & Géry, 1988 (Pisces)
Cryptocheiridium confundens Mahnert, 2014 (Pseudoscorpiones)*
Ctenophthalmus (Ethioctenophthalmus) digitosignatus Mahnert, 1977 (Siphonaptera) ***
Cyclatemnus brevidigitatus Mahnert, 1978 (Pseudoscorpiones)
Dactylochelifer arabicus Mahnert, 1991 (Pseudoscorpiones)*
Dactylochelifer besucheti Mahnert, 1978 (Pseudoscorpiones)***
Dactylochelifer scaurus Mahnert, 1978 (Pseudoscorpiones)**
Dactylochelifer vtorovi Mahnert, 1977 (Pseudoscorpiones)***
Dasychernes panamensis Mahnert, 1987 (Pseudoscorpiones)***
Dasychernes roubiki Mahnert, 1987 (Pseudoscorpiones)***
Dasychernes trigonae Mahnert, 1987 (Pseudoscorpiones)***
Dhanus pohli Mahnert, 2007 (currently in Shravana; Pseudoscorpiones)
Dhanus socotraensis Mahnert, 2007 (currently in Shravana; Pseudoscorpiones)***
Dhanus taitii Mahnert, 2007 (currently in Shravana; Pseudoscorpiones)*
Dolichowithius (Dolichowithius) intermedius Mahnert, 1979 (Pseudoscorpiones)*
Dolichowithius (Dolichowithius) mediofasciatus Mahnert, 1979 (Pseudoscorpiones)*
Dolichowithius (Dolichowithius) minutus Mahnert, 1979 (Pseudoscorpiones)*
Elattogarypus cicatricosus Mahnert, 2007 (Pseudoscorpiones)*
Elattogarypus somalicus Mahnert 1984 (Pseudoscorpiones)*
Feaella (Tetrafeaella) perreti Mahnert, 1982 (Pseudoscorpiones)***
Garypus darsahensis Mahnert, 2007 (Pseudoscorpiones)
Garypinus nicolaii Mahnert, 1988 (Pseudoscorpiones)*
Garypus granosus Mahnert, 2014 (Pseudoscorpiones)*
Garypus marmoratus Mahnert, 1982 (Pseudoscorpiones)***
Garypus occultus Mahnert, 1982 (Pseudoscorpiones)***
Geogarypus (Afrogarypus) plumatus Mahnert, 1982 (currently in Afrogarypus; Pseudoscorpiones)***
Geogarypus (Afrogarypus) pseudocurtus Mahnert, 1982 (currently in Afrogarypus; Pseudoscorpiones)***
Geogarypus (Afrogarypus) stellifer Mahnert, 1982 (currently in Afrogarypus; Pseudoscorpiones)***
Geogarypus (Geogarypus) amazonicus Mahnert, 1979 (Pseudoscorpiones)*
Geogarypus (Geogarypus) ocellatus Mahnert, 1978 (Pseudoscorpiones)*
Geogarypus quadrimaculatus Mahnert, 2007 (Pseudoscorpiones)
Halominniza oromii Mahnert, 1997 (Pseudoscorpiones)
Halominniza parentorum Mahnert, 1975 (Pseudoscorpiones)**
Halominniza taitii Mahnert, 2007 (Pseudoscorpiones)
Haplochernes nanus Mahnert, 1975 (Pseudoscorpiones)***
Hectopsylla gracilis Mahnert, 1982 (Siphonaptera)***
Hemigrammus bleheri Géry & Mahnert, 1986 (Pisces)*
Hyphessobrycon guarani Mahnert & Géry, 1987 (currently in Diapoma; Pisces)***
Hyphessobrycon procerus Mahnert & Géry, 1987 (Pisces)***
Hyphessobrycon pytai Mahnert & Géry, 1993 (Pisces)***
Ideobisium schusteri Mahnert, 1985 (Pseudoscorpiones)***
Ideobisium amazonicum Mahnert, 1979 (currently in Ideoblothrus; Pseudoscorpiones)*
Ideobisium baloghi Mahnert, 1978 (currently in Ideoblothrus; Pseudoscorpiones)*
Ideobisium brasiliense Mahnert, 1979 (currently in Ideoblothrus; Pseudoscorpiones)*
Ideobisium caecum Mahnert, 1979 (currently in Ideoblothrus; Pseudoscorpiones)*
Ideobisium zicsii Mahnert, 1978 (currently in Ideoblothrus; Pseudoscorpiones)*
Ideoblothrus emigrans Mahnert, 2014 (Pseudoscorpiones)*
Ideoblothrus galapagensis Mahnert, 2014 (Pseudoscorpiones)***
Ideoblothrus levipalpus Mahnert, 1985 (Pseudoscorpiones)**
Ideoblothrus paraensis Mahnert, 1985 (Pseudoscorpiones)*
Ideoblothrus tenuis Mahnert, 1985 (Pseudoscorpiones)*
Ideoroncus anophthalmus Mahnert, 1984 (Pseudoscorpiones)***
Ideoroncus arboricola Mahnert, 1979 (currently in Xorilbia; Pseudoscorpiones)*
Ideoroncus beieri Mahnert, 1984 (Pseudoscorpiones)***
Ideoroncus cavicola Mahnert, 2001 (Pseudoscorpiones)*
Ideoroncus divisus Mahnert, 1984 (Pseudoscorpiones)*
Ideoroncus paranensis Mahnert, 1984 (Pseudoscorpiones)***
Ideoroncus setosus Mahnert, 1984 (Pseudoscorpiones)***
Lagynochthonius curvidigitatus Mahnert, 1997 (Pseudoscorpiones)
Lagynochthonius insulanus Mahnert, 2007 (Pseudoscorpiones)***
Lagynochthonius lopezi Mahnert, 2011 (Pseudoscorpiones)*
Lagynochthonius microdentatus Mahnert, 2011 (Pseudoscorpiones)
Lagynochthonius oromii Mahnert, 2011 (Pseudoscorpiones)*
Lagynochthonius subterraneus Mahnert, 2011 (Pseudoscorpiones)*
Lagynochthonius tenuimanus Mahnert, 2011 (Pseudoscorpiones)*
Lechytia dentata Mahnert, 1978 (Pseudoscorpiones)
Leporinus falcipinnis Mahnert, Géry & Muller, 1997 (currently in the synonymy of Leporinus altipinnis Borodin, 1929; Pisces)
Leptocheiridium pfeiferae Mahnert & Schmidl, 2011 (Pseudoscorpiones)***
Levigatocreagris hamatus Leclerc & Mahnert, 1988 (currently in Stenoya; Pseudoscorpiones)***
Lophochelifer gracilipes Mahnert, 1988 (currently in Lissochelifer; Pseudoscorpiones)***
Lophochelifer nairobiensis Mahnert, 1988 (currently in Lissochelifer; Pseudoscorpiones)***
Maxchernes iporangae Mahnert & Andrade, 1998 (Pseudoscorpiones)*
Megachernes glandulosus Mahnert, 2009 (Pseudoscorpiones)**
Megachernes tuberosus Mahnert, 2009 (Pseudoscorpiones)***
Mesochelifer ressli Mahnert, 1981 (Pseudoscorpiones)***
Metastivalius novaehiberniae Beaucournu & Mahnert, 1988 (Siphonaptera)***
Micratemnus crassipes Mahnert, 1983 (Pseudoscorpiones)***
Microblothrus trident Mahnert, 1985 (Pseudoscorpiones)*
Microchelifer dentatus Mahnert, 1988 (Pseudoscorpiones)**
Microcreagrina cavicola Mahnert, 1993 (Pseudoscorpiones)*
Microcreagrina gomerae Mahnert, 1993 [currently in the synonymy of Microcreagrina hispanica (Ellingsen, 1910); Pseudoscorpiones]**
Microcreagrina madeirensis Mahnert, 1993 (Pseudoscorpiones)**
Microcreagrina subterranea Mahnert, 1993 (Pseudoscorpiones)*
Microcreagris leucadia Mahnert, 1972 [currently in Acanthocreagris; Pseudoscorpiones]**
Minniza antonii Mahnert, 2009 (Pseudoscorpiones)***
Minniza barkhamae Mahnert, 1991 (Pseudoscorpiones)*
Minniza elegans Mahnert, 1991 (Pseudoscorpiones)*
Minniza gallagheri Mahnert, 1991 (Pseudoscorpiones)*
Minniza levisetosa Mahnert, 1991 (Pseudoscorpiones)*
Minniza monticola Mahnert, 1991 (Pseudoscorpiones)*
Minniza nigrimanus Mahnert, 1991 (Pseudoscorpiones)*
Miratemnus kenyaensis Mahnert, 1983 (Pseudoscorpiones)***
Myrmecowithius buettikeri Mahnert, 1980 (currently in Nannowithius; Pseudoscorpiones)*
Myrmecowithius paradoxus Mahnert, 1980 now in Nannowithius; Pseudoscorpiones)*
Nannobisium beieri Mahnert, 1979 (Pseudoscorpiones)*
Negroroncus azanius Mahnert, 1981 (Pseudoscorpiones)***
Negroroncus densedentatus Mahnert, 1981 (Pseudoscorpiones)
Negroroncus gregoryi Mahnert, 1981 (Pseudoscorpiones)***
Negroroncus kerenyaga Mahnert, 1981 (Pseudoscorpiones)**
Negroroncus silvicola Mahnert, 1981 (Pseudoscorpiones)*
Negroroncus tsavoensis Mahnert, 1981 (Pseudoscorpiones)***
Neobisium (Blothrus) cervelloi Mahnert, 1977 (Pseudoscorpiones)
Neobisium (Blothrus) hians Mahnert, 1979 (Pseudoscorpiones)
Neobisium (Blothrus) kwartirnikovi Mahnert, 1972 (Pseudoscorpiones)***
Neobisium (Blothrus) monasterii Mahnert, 1977 (Pseudoscorpiones)*
Neobisium (Neobisium) bosnicum ondriasi Mahnert, 1973 (Pseudoscorpiones)***
Neobisium (Neobisium) intermedium Mahnert, 1974 (Pseudoscorpiones)***
Neobisium (Neobisium) phitosi Mahnert, 1973 (Pseudoscorpiones)***
Neobisium (Neobisium) reductum Mahnert, 1977 (Pseudoscorpiones)*
Neobisium (Ommatoblothrus) gaditanum Mahnert, 1977 (Pseudoscorpiones)
Neobisium (Ommatoblothrus) patrizii romanum Mahnert, 1980 (Pseudoscorpiones)
Neobisium (Ommatoblothrus) paucedentatum Mahnert, 1982 (Pseudoscorpiones)*
Neobisium (Ommatoblothrus) phaeacum Mahnert, 1973 (Pseudoscorpiones)***
Neobisium (Ommatoblothrus) samniticum Mahnert, 1980 (Pseudoscorpiones)*
Neocheiridium africanum Mahnert, 1982 (Pseudoscorpiones)*
Neocheiridium pusillum Mahnert, 1982 (Pseudoscorpiones)***
Neocheiridium strinatii Mahnert & Aguiar, 1986 (Pseudoscorpiones)***
Neocheiridium triangulare Mahnert & Aguiar, 1986 (Pseudoscorpiones)*
Nesowithius dilatimanus Mahnert, 1988 (Pseudoscorpiones)***
Nhatrangia ceylonensis Mahnert, 1984 (currently in Shravana; Pseudoscorpiones)***
Nudochernes gracilimanus Mahnert, 1982 (Pseudoscorpiones)***
Nudochernes lipsae Mahnert, 2003 (Pseudoscorpiones)**
Nudochernes troglobius Mahnert, 2009 (Pseudoscorpiones)***
Olpium flavum Mahnert, 2007 (Pseudoscorpiones)***
Olpium halophilum Mahnert, 1982 (Pseudoscorpiones)***
Olpium kuriense Mahnert, 2007 (Pseudoscorpiones)*
Olpium milneri Mahnert, 2007 (Pseudoscorpiones)***
Olpium omanense Mahnert, 1991 (Pseudoscorpiones)
Olpium socotraense Mahnert, 2007 (Pseudoscorpiones)***
Olpium vanharteni Mahnert, 2007 (Pseudoscorpiones)***
Pachyolpium atlanticum Mahnert & Schuster, 1981 (Pseudoscorpiones)***
Pachyolpium irmgardae Mahnert, 1979 (Pseudoscorpiones)*
Parachernes adisi Mahnert, 1979 (Pseudoscorpiones)*
Parachernes gracilimanus Mahnert, 1986 (Pseudoscorpiones)***
Parachernes inpai Mahnert, 1979 (Pseudoscorpiones)*
Parachernes niger Mahnert 1987 (Pseudoscorpiones)*
Parachernes ovatus Mahnert, 1979 (Pseudoscorpiones)*
Parachernes pulcher Mahnert, 1979 (Pseudoscorpiones)
Parachernes setiger Mahnert, 1979 (Pseudoscorpiones)*
Paraliochthonius azanius Mahnert, 1986 (Pseudoscorpiones)***
Paraliochthonius galapagensis Mahnert, 2014 (Pseudoscorpiones)***
Paraliochthonius litoralis Mahnert, 2014 (Pseudoscorpiones)*
Paraliochthonius martini Mahnert, 1989 (Pseudoscorpiones)*
Paraliochthonius mirus Mahnert, 2002 (Pseudoscorpiones)*
Paraliochthonius pecki Mahnert, 2014 (Pseudoscorpiones)***
Paraliochthonius rupicola Mahnert, 2014 (Pseudoscorpiones)***
Paraliochthonius tenebrarum Mahnert, 1989 (Pseudoscorpiones)
Paramenthus nanus Mahnert, 2007 (Pseudoscorpiones)*
Parawithius (Victorwithius) coniger Mahnert, 1979 (currently in Victorwithius; Pseudoscorpiones)*
Parawithius (Victorwithius) gracilimanus Mahnert, 1979 (currently in Victorwithius; Pseudoscorpiones)*
Parobisium martii Mahnert, 2003 (currently in Bisetocreagris; Pseudoscorpiones)**
Parobisium scaurum Mahnert, 2003 (currently in Bisetocreagris; Pseudoscorpiones)**
Parobisium titanium Mahnert, 2003 (currently in Bisetocreagris; Pseudoscorpiones)**
Petterchernes tuberculatus Mahnert, 1994 (Pseudoscorpiones)*
Phymatochernes crassimanus Mahnert, 1979 (Pseudoscorpiones)
Piabarchus torrenticola Mahnert & Géry, 1988 (Pisces)***
Pilochelifer insularis gracilior Mahnert, 1975 (Pseudoscorpiones)***
Plesiochernes bergeri Mahnert, 1978 (currently in Caffrowithius; Pseudoscorpiones)**
Plocopsylla angusticeps Mahnert, 1982 (Siphonaptera)**
Progarypus gracilis Mahnert, 2001 (Pseudoscorpiones)*
Progarypus liliae Mahnert, 2001 (Pseudoscorpiones)
Progarypus nigrimanus Mahnert, 2001 (Pseudoscorpiones)*
Progarypus setifer Mahnert, 2001 (Pseudoscorpiones)*
Pseudoblothrus infernus Mahnert, 2011 (Pseudoscorpiones)**
Pseudoblothrus oromii Mahnert, 1990 (Pseudoscorpiones)*
Pseudoblothrus vulcanus Mahnert 1990 (Pseudoscorpiones)*
Pseudochernes arabicus Mahnert, 1991 (Pseudoscorpiones)
Pseudochiridium kenyense Mahnert, 1982 (Pseudoscorpiones)*
Pseudochthonius arabicus Mahnert, 2014 (Pseudoscorpiones)**
Pseudochthonius beieri Mahnert, 1978 (Pseudoscorpiones)*
Pseudochthonius biseriatus Mahnert, 2001 (Pseudoscorpiones)*
Pseudochthonius gracilimanus Mahnert, 2001 (Pseudoscorpiones)*
Pseudochthonius perreti Mahnert, 1986 (Pseudoscorpiones)*
Pseudochthonius ricardoi Mahnert, 2001 (Pseudoscorpiones)
Pseudochthonius tuxeni Mahnert, 1979 (Pseudoscorpiones)*
Pseudohorus embuensis Mahnert, 1982 (Pseudoscorpiones)***
Pseudohorus luscus Mahnert, 1982 (Pseudoscorpiones)***
Pseudohorus pilosus Mahnert, 1982 (Pseudoscorpiones)***
Pseudohorus vermis Mahnert, 1982 (Pseudoscorpiones)***
Pseudomenthus spinifer Mahnert, 2007 (Pseudoscorpiones)*
Pseudomenthus uniseriatus Mahnert, 2007 (Pseudoscorpiones)**
Pseudopilanus crassifemoratus Mahnert, 1985 (Pseudoscorpiones)*
Pseudorhacochelifer canariensis Mahnert, 1997 (Pseudoscorpiones)*
Pycnowithius cavernicola Mahnert, 1988 (Pseudoscorpiones)***
Rhacochelifer barkhamae Mahnert, 1980 (Pseudoscorpiones)
Rhacochelifer corcyrensis procerus Mahnert, 1978 (Pseudoscorpiones)***
Rhacochelifer euboicus Mahnert, 1977 (Pseudoscorpiones)**
Rhacochelifer gracilimanus Mahnert, 1993 (Pseudoscorpiones)***
Rhacochelifer sonyae Mahnert, 1991 (Pseudoscorpiones)*
Rhacochelifer spiniger Mahnert, 1978 (currently in Pseudorhacochelifer; Pseudoscorpiones)*
Roncocreagris beieri Mahnert, 1976 (Pseudoscorpiones)***
Roncus gasparoi Mahnert & Gardini, 2014 (Pseudoscorpiones)**
Roncus giachinoi Mahnert & Gardini, 2014 (Pseudoscorpiones)***
Roncus pieperi Mahnert & Gardini, 2014 (Pseudoscorpiones)***
Roncus (Parablothrus) boneti tarbenae Mahnert, 1977 (Pseudoscorpiones)*
Roncus (Parablothrus) comasi Mahnert, 1985 (Pseudoscorpiones)*
Roncus (Parablothrus) corcyraeus minor Mahnert, 1975 (Pseudoscorpiones)***
Roncus (Parablothrus) paolettii Mahnert, 1980 (Pseudoscorpiones)*
Roncus (Parablothrus) puddui Mahnert, 1976 (Pseudoscorpiones)***
Roncus (Roncus) giganteus Mahnert, 1973 (Pseudoscorpiones)***
Roncus (Roncus) viti Mahnert, 1974 (Pseudoscorpiones)**
Selachochernes allodentatus Mahnert, 2001 (Pseudoscorpiones)**
Serianus elongatus Mahnert 2014 (Pseudoscorpiones)***
Serianus maritimus Mahnert 2014 (Pseudoscorpiones)***
Serianus sahariensis Mahnert, 1988 (Pseudoscorpiones)***
Spelaeobochica allodentatus Mahnert, 2001 (Pseudoscorpiones)*
Spelaeobochica iuiu Ratton, Mahnert & Ferreira, 2012 (Pseudoscorpiones)*
Spelaeobochica muchmorei Andrade & Mahnert, 2003 (Pseudoscorpiones)*
Spelaeochernes altamirae Mahnert, 2001 (Pseudoscorpiones)*
Spelaeochernes armatus Mahnert, 2001 (Pseudoscorpiones)*
Spelaeochernes bahiensis Mahnert, 2001 (Pseudoscorpiones)*
Spelaeochernes dentatus Mahnert, 2001 (Pseudoscorpiones)*
Spelaeochernes dubius Mahnert, 2001 (Pseudoscorpiones)*
Spelaeochernes eleonorae Mahnert, 2001 (Pseudoscorpiones)*
Spelaeochernes gracilipalpus Mahnert, 2001 (Pseudoscorpiones)*
Spelaeochernes pedroi Mahnert, 2001 (Pseudoscorpiones)*
Sphaerowithius ansieae Harvey & Mahnert, 2015 (Pseudoscorpiones)***
Stenolpium fasciculatum Mahnert, 1984 (Pseudoscorpiones)
Thaumastogarypus mancus Mahnert, 1982 (Pseudoscorpiones)**
Thaumatowithius aberrans Mahnert, 1975 (Pseudoscorpiones)***
Titanatemnus tanensis Mahnert, 1983 (Pseudoscorpiones)***
Tridenchthonius beieri Mahnert, 1983 (Pseudoscorpiones)***
Tridenchthonius brasiliensis Mahnert, 1979 (Pseudoscorpiones)*
Typhloceras poppei orientalis Mahnert & Beaucournu, 1985 (Siphonaptera)**
Tyrannochthonius akaleus Mahnert, 2009 (Pseudoscorpiones)**
Tyrannochthonius (Tyrannochthonius) amazonicus Mahnert, 1979 (Pseudoscorpiones)*
Tyrannochthonius antridraconis Mahnert, 2009 (Pseudoscorpiones)***
Tyrannochthonius (Tyrannochthonius) brasiliensis Mahnert, 1979 (Pseudoscorpiones)
Tyrannochthonius confusus Mahnert, 1986 (Pseudoscorpiones)*
Tyrannochthonius ferox Mahnert, 1978 (currently in Lagynochthonius; Pseudoscorpiones)
Tyrannochthonius (Lagynochthonius?) flavus Mahnert, 1986 (currently in Lagynochthonius; Pseudoscorpiones)***
Tyrannochthonius ganshuanensis Mahnert, 2009 (Pseudoscorpiones)***
Tyrannochthonius (Tyrannochthonius) gomyi Mahnert, 1975 (Pseudoscorpiones)**
Tyrannochthonius (Lagynochthonius) guasirih Mahnert, 1988 (currently in Lagynochthonius; Pseudoscorpiones)***
Tyrannochthonius (Tyrannochthonius) irmleri Mahnert, 1979 (currently in Lagynochthonius; Pseudoscorpiones)*
Tyrannochthonius (Lagynochthonius?) kenyensis Mahnert, 1986 (currently in Lagynochthonius; Pseudoscorpiones)***
Tyrannochthonius mahunkai Mahnert, 1978 (Pseudoscorpiones)*
Tyrannochthonius (Tyrannochthonius) migrans Mahnert, 1979 (Pseudoscorpiones)*
Tyrannochthonius (Tyrannochthonius) minor Mahnert, 1979 (currently in Lagynochthonius; Pseudoscorpiones)*
Tyrannochthonius procerus Mahnert, 1978 (Pseudoscorpiones)*
Tyrannochthonius pugnax Mahnert, 1978 (currently in Lagynochthonius; Pseudoscorpiones)*
Tyrannochthonius riberai Mahnert, 1984 (Pseudoscorpiones)*
Tyrannochthonius (Tyrannochthonius) rotundimanus Mahnert, 1985 (Pseudoscorpiones)*
Tyrannochthonius setiger Mahnert, 1997 (currently in Paraliochthonius; Pseudoscorpiones)*
Tyrannochthonius (Tyrannochthonius) simulans Mahnert, 1986 (Pseudoscorpiones)***
Tyrannochthonius (Tyrannochthonius) superstes Mahnert, 1986 (currently in Paraliochthonius; Pseudoscorpiones)*
Tyrannochthonius zicsii Mahnert, 1978 (currently in Lagynochthonius; Pseudoscorpiones)
Withius (Allowithius) arabicus Mahnert, 1980 (Pseudoscorpiones)*
Withius brevidigitatus Mahnert, 1988 (Pseudoscorpiones)***
Withius fuscus Mahnert, 1988 (Pseudoscorpiones)***
Withius gracilipalpus Mahnert, 1988 (Pseudoscorpiones)***
Withius nanus Mahnert, 1988 (Pseudoscorpiones)***
Zaona cavicola Mahnert, 2001 (Pseudoscorpiones)
Appendix 3: Taxa dedicated to Volker Mahnert
Animal genera
Mahnertella Mahunka, 1997 (currently in the synonymy of Rugoppia Mahunka, 1986; Acari)
Mahnertius Harvey & Muchmore, 2013 (Pseudoscorpiones)
Mahnertozetes Mahunka & Mahunka-Papp, 2009 (Acari)
Animal species
Acanthocreagris mahnerti Dumitresco & Orghidan, 1986 (Pseudoscorpiones)
Acritus mahnerti Gomy, 1981 (Coleoptera)
Akyttara mahnerti Jocqué, 1987 (Araneae)
Allochernes mahnerti Georgescu & Căpuse, 1996 (Pseudoscorpiones)
Allolobophora handlirschi mahnerti Zicsi, 1973 (currently in Aporrectodea; Oligochaeta)
Americhernes mahnerti Harvey, 1990 (Pseudoscorpiones)
Apimela mahnerti Pace, 1996 (Coleoptera)
Atheta mahnerti Pace, 1995 (Coleoptera)
Ausobskya mahnerti Silhavý, 1976 (Opiliones)
Bothriechis mahnerti Schätti & Kramer, 1991 [currently in the synonymy of Bothrops punctatus (Garcia, 1896); Serpentes]
Camillina mahnerti Platnick & Murphy, 1987 (Araneae)
Catharosoma mahnerti Golovatch, 2005 (Diplopoda)
Centruroides mahnerti Lourenço, 1983 (Scorpiones)
Chactas mahnerti Lourenço, 1995 (Scorpiones)
Chthonius (Ephippiochthonius) mahnerti Zaragoza, 1984 (Pseudoscorpiones)
Ctenobelba mahnerti Mahunka, 1974 (Acari)
Cypha mahnerti Pace, 1994 (Coleoptera)
Delamarea mahnerti Leleup, 1983 (Coleoptera)
Dendrobaena mahnerti Zicsi, 1974 (Oligochaeta)
Dolicheremaeus mahnerti Mahunka & Mahunka-Papp, 2009 (Acari)
Drusilla mahnerti Pace, 1996 (Coleoptera)
Edaphus mahnerti Puthz, 1990 (Coleoptera)
Elgonidium mahnerti Bonadona, 1978 (Coleoptera)
Embuana mahnerti Heiss & Baňař, 2016 (Heteroptera)
Epipleuria mahnerti Fürsch, 2001 (Coleoptera)
Glossodrilus mahnerti Zicsi, 1989 (Oligochaeta)
Guaraniella mahnerti Baert, 1984 (Araneae)
Gyrophaena mahnerti Pace, 1994 (Coleoptera)
Helladocampa mahnerti Condé, 1984 (Diplura)
Hemigrammus mahnerti Uj & Géry, 1989 (Pisces)
Holoparasitus mahnerti Juvara-Bals, 2008 (Acari)
Leleupiozethus mahnerti Coulon, 1979 (Coleoptera)
Magellozetes mahnerti Mahunka, 1984 (currently in the synonymy of Ceratozetes platyrhinoides Hammer, 1961; Acari)
Megarthrus mahnerti Cuccodoro & Löbl, 1995 (Coleoptera)
Metanapis mahnerti Brignoli, 1981 (Araneae)
Microdipnites mahnerti Garetto & Giachino, 1999 (Coleoptera)
Microplana mahnerti Minelli, 1977 [currently in the synonymy of Microplana terrestris (O.F. Müller, 1773); Tricladida]
Neobisium mahnerti Heurtault, 1980 (Pseudoscorpiones) Occidenchthonius mahnerti Zaragoza, 2017 (Pseudoscorpiones)
Oedichirus mahnerti de Rougemont, 2018 (Coleoptera)
Oonops mahnerti Brignoli, 1974 (Araneae)
Paracoryza mahnerti Balkenohl, 2000 (Coleoptera)
Paratemnus mahnerti Sivaraman, 1981 (currently in Paratemnoides; Pseudoscorpiones)
Paratricommatus mahnerti Soares & Soares, 1985 (Opiliones)
Proteocephalus mahnerti de Chambrier & Vaucher, 1999 (Cestoda)
Ptychadena mahnerti Perret, 1996 (Amphibia)
Roncus mahnerti Ćurčić & Beron, 1981 (Pseudoscorpiones)
Scaphoxium mahnerti Löbl, 2010 (Coleoptera)
Scheloribates mahnerti Mahuka & Mahunka-Papp, 2008 (Acari)
Schistura mahnerti Kottelat, 1990 (Pisces)
Scutacarus mahnerti Mahunka, 1972 (Acari)
Sprentascaris mahnerti Petter & Cassone, 1984 [currently in Raphidascaris (Sprentascaris); Nematoda]
Stenohya mahnerti Schawaller, 1994 (Pseudoscorpiones)
Trichouropoda mahnerti Kontschán, 2015 (Acari)
Zodarion mahnerti Brignoli, 1984 [currently in the synonymy of Zodarion emarginatum (Simon, 1873); Araneae]
Zyras (Pycnodonia) mahnerti Pace, 1996 (Coleoptera)
