Toru Miura,^1,2,* Kohei Oguchi,^1,2 Mayuko Nakamura,¹ Naoto Jimi,³ Sakiko Miura,^1,2 Yoshinobu Hayashi,⁴ Shigeyuki Koshikawa,² M. Teresa Aguado<sup>5,6

1</sup>Misaki Marine Biological Station, School of Science, The University of Tokyo, Misaki, Miura, Kanagawa 238-0225, Japan
²Faculty of Environmental Earth Science, Hokkaido University, Sapporo, Hokkaido 060-0810, Japan
³Bioscience Group, National Institute of Polar Research, Tachikawa, Tokyo 190-0014, Japan
⁴Department of Biology, Keio University, Yokohama, Kanagawa 223-8521, Japan
⁵Animal Evolution and Biodiversity, Johann-Friedrich-Blumenbach Institute for Zoology and Anthropology, Georg-August-Universität Göttingen, Göttingen, Germany
⁶Centro de Investigación en Biodiversidad y Cambio Global (CIBC-UAM), Departamento de Biología, Facultad de Ciencias, Universidad Autónoma de Madrid, Cantoblanco, 28049, Madrid, Spain

^*Corresponding author. E-mail: miu@mmbs.s.u-tokyo.ac.jp