Takanori Nishino,^1,2,* Takahiro Hosokawa,^3,* Xian-Ying Meng,² Ryuichi Koga,² Minoru Moriyama,² Takema Fukatsu<sup>1,2,4,**

1</sup>Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba 305-8572, Japan
²Bioproduction Research Institute, National Institute of Advanced Industrial Science and Technology (AIST), Tsukuba 305-8566, Japan
³Department of Biology, Faculty of Science, Kyushu University, Fukuoka 819-0395, Japan
⁴Department of Biological Sciences, Graduate School of Science, University of Tokyo, Tokyo 113-0033, Japan

^* Contributed equally to the work
^** Corresponding author. E-mail: t-fukatsu@aist.go.jp