Makoto Iwasaki,^1,* Yusuke Sakai,^1,* Taishi Shirata,² Tomohiro Sugihara,¹ Mitsumasa Koyanagi,^1,3 Akihisa Terakita<sup>1,3,**

1</sup>Department of Biology, Graduate School of Science, Osaka Metropolitan University 3-3-138 Sugimoto, Sumiyoshi-Ku Osaka 558-8585, Japan
²Department of Biology and Geosciences, Graduate School of Science, Osaka City University, 3-3-138 Sugimoto, Sumiyoshi-Ku, Osaka 558-8585, Japan
³The OMU Advanced Research Institute for Natural Science and Technology, Osaka Metropolitan University, Osaka, Japan

^*These authors contributed equally to this work.
^**Corresponding author. E-mail: terakita@omu.ac.jp