Rebecca J. Welchhttps://orcid.org/0000-0002-4182-7878,^1,2,*,** Melissa H. Schmitthttps://orcid.org/0000-0002-5544-7673,^1,3 Thando Mendela,⁴ Ric T.F. Bernard,^1,4 Dan M. Parkerhttps://orcid.org/0000-0001-7555-5674<sup>1,4

1</sup>School of Biology and Environmental Sciences, University of Mpumalanga, Private Bag X11283, Nelspruit, 1200 South Africa
²South African Institute for Aquatic Biodiversity (SAIAB), Private Bag 1015, Grahamstown, 6140 South Africa
³Department of Ecology, Evolution, and Marine Biology, University of California Santa Barbara, Santa Barbara, CA 93101, U.S.A.
⁴Wildlife and Reserve Management Research Group, Department of Zoology & Entomology, Rhodes University, Makhanda, South Africa

^*To whom correspondence should be addressed. E-mail: welchrebecca83@gmail.com
^**Rebecca Welch and Melissa Schmitt contributed equally to this