Anachemmis Chamberlin is revised, and contains five species from California, Arizona, and Sonora that are hypothesized to represent the sister group of the Appalachian genus Liocranoides Keyserling. The male of the southern Californian type species, A. sober Chamberlin, is described for the first time, as are both sexes of four new species: A. linsdalei from northern and central California, A. jungi from caves in Inyo County, California, A. aalbui from in and near caves in San Bernardino County, California, and A. beattyi from Arizona and Sonora.