The hind wings of all known families and most subfamilies of Coleoptera are illustrated, annotated and discussed utilising the terminology of Kukalová-Peck and Lawrence (), with a few changes in nomenclature suggested by the senior author. The beetle families are discussed in 21 groups, based on recent classifications of Coleoptera. For each of these groups, the most recent works on phylogeny and classification are reviewed, and the wing characters are discussed to determine if some of the wing features might support or refute relationships based on recent molecular and morphological analyses. Part 1 includes a general discussion of wing structure divided into the following sections: hind wing fields, veinal systems (including the history of wing nomenclature), wing folding, wing edge and embayments, hinges and bending zones, cross-veins and braces, cells and other landmarks. It is followed by discussion of the first 14 groups (Archostemata to Elateroidea), 15 figures supporting general discussions, and 426 labelled wing images of the discussed groups, representing 380 genera.

This paper is the third in a series on the genus Habroloma Thomson (1864) from China. Twelve new species are described and illustrated: Habroloma brevium sp. nov., H. duploarcum sp. nov., H. contractiformis sp. nov., H. longiusculum sp. nov., H. dilatipennis sp. nov., H. chishuiensis sp. nov., H. fusiformis sp. nov., H. subfusiformis sp. nov., H, semiparallelum sp. nov., H. chongrenensis sp. nov., H. yiyangensis sp. nov., and H. changtingensis sp. nov.

Seven species of Carpophilinae: Urophorus humeralis (Fabricius), Carpophilus (Carpophilus) jelineki Audisio & Kirejtshuk, Carpophilus (Carpophilus) obsoletus Erichson, Carpophilus (Carpophilus) flavipes Murray, Carpophilus (Myothorax) nepos Murray, Carpophilus (Ecnomorphus) plagiatipennis Motschulsky, and Carpophilus (Ecnomorphus) venkataramani Dasgupta, Pal & Powell sp. nov., are recorded from Tripura state of India. The genera and species are (re)described and a key to the genera and species of Carpophilinae of Tripura is given.

A new heterotardigrade genus, Zealandiscus gen. nov., is erected for the species Echiniscus palmai. Dorsal plates' sculpturing with sparsely distributed large hemispherical protuberances and very tiny granules distinguish this genus from all known members of the genus Echiniscus. The new genus is most similar to the genus Kristenseniscus by the shape of claws and some aspects of dorsal cuticle i.e. presence of sparsely distributed quite large hemispherical protuberances. However, in the newly erected genus dorsal plates' sculpture is “double-like” (i.e. it is composed of two types of granulation visible in different focal planes), hemispherical protuberances are larger and not so dense, dorsal plates are not subdivided into smaller subplates, median plate m3 and ventral subcephalic plates are present. Such autapomorphies fully justify erection of the new genus, even if the genetic data are at present unavailable.

The genus Phosphaenopterus encompasses two species restricted to Europe, both underrepresented in the literature and entomological collections. Phosphaenopterus metzneri is the most widely distributed within the genus, being recorded in France, Portugal, and Spain. Since its century-old original description based only on the male, the biology of this species was not further investigated. By collecting larvae of Phosphaenina where male of Phosphaenopterus metzneri occur, we reared them to adults, resulting in male and females of Phosphaenopterus metzneri – the latter hitherto unknown. The male-female association was possible due to copula recorded ex-situ. Here, we describe for the first time the neotenic female of Phosphaenopterus metzneri, a diurnal firefly, and redescribed the male. We provide illustration of diagnostic features of the adults, and an updated distribution map for Phosphenopterus metzneri. Considering the morphological resemblance between all the three species within Phosphaenina, we also redescribe the male of Phosphaenus hemipterus (), a species also presents in Portugal, providing important diagnostic features for both taxa. By comparing the morphology of males of Phosphaenus hemipterus and Phosphaenopterus metzneri, we could not find any difference other than proportions of elytra, and wings, which may be a result of the brachypterous condition of the former. This study stresses and discusses a putative case of wing polymorphism in Lampyridae, a rare condition in fireflies. This is also the first study to report both species in sympatric distribution.