Neuwirth, G. & Aptroot, A. 2011. Recognition of four morphologically distinct species in the Graphis scripta complex in Europe. — Herzogia 24: 207–230.

Four morphologically distinct species can be distinguished in the Graphis scripta complex in Europe, namely Graphis betulina, G. macrocarpa, G. pulverulenta and G. scripta. Graphis betulina is characterized by its raised thalline margin, G. macrocarpa has open, brownish discs, G. pulverulenta has open, white-pruinose discs, and G. scripta has apothecia that remain closed. The species are recognizable in the field, but there are also microscopical differences. All species are sympatric and occasionally co-occur on the same tree.