Although the large white-headed gull group (genus Larus) has long been a model in speciation studies, the systematic status and evolutionary relationships of many of its taxa remain unresolved. We used mitochondrial DNA control region and cytochrome-b gene sequences in an attempt to resolve some of those uncertainties. In contrast to previously published results based on nuclear markers, mitochondrial DNA was found to be strongly structured among species, indicating that mitochondrial gene flow is very low. Phylogenetic relationships remain largely unresolved, mainly because of the low amount of variation between species. Horizontal transfer of mitochondrial lineages is demonstrated or suspected between most taxa and obscured the reconstruction of the history of the group. The Mediterranean form michahellis was as differentiated from the other western European species as these are from each others, confirming it is neither conspecific with L. fuscus nor with L. argentatus. The forms fuscus and graellsii do not show any significant differences in haplotypes frequencies, arguing for their subspecific status.

The phylogeny of all known Xiphorhynchus (Dendrocolaptidae) species and many of its subspecies was reconstructed to evaluate species limits in this taxonomically challenging genus and investigate the possible role played by the Amazonian “várzea” (floodplain forest)–“terra-firme” (upland forest) ecotone in its diversification. Phylogenies were inferred based on 2,430 bp of the mitochondrial DNA genes ND2, ND3, and cytochrome b. All phylogeny estimates supported the monophyly of all extant Xiphorhynchus species to the exclusion of the sibling species pair Straight-billed (X. picus) and Zimmer's (X. kienerii) woodcreeper. Confirming findings of previous molecular and anatomical studies, strong support was found to include the Lesser Woodcreeper (Lepidocolaptes fuscus) in Xiphorhynchus. Levels of sequence divergence among some subspecies of Buff-throated (X. guttatus), Ocellated (X. ocellatus), and Spix's (X. spixii) woodcreepers reached or exceeded those found between closely related, undisputed biological species of Xiphorhynchus. High levels of sequence differentiation and the paraphyly of some Xiphorhynchus species indicated that the following taxa should be recognized as species: Lafresnaye's (X. guttatoides), Tschudi's (X. chunchotambo), and Elegant (X. elegans) woodcreepers. All Xiphorhynchus species restricted to terra-firme forest in lowland Amazonia formed a well supported monophyletic group, whereas species restricted to várzea forest were either basal to a clade containing species found in a wide variety of habitats (Striped Woodcreeper [X. obsoletus]) or belonged to a distinct lineage likely to be regarded as a separate genus (X. kienerii). These findings falsified an anticipated sister relationship between várzea and terra-firme species, as expected if the várzea–terra-firme ecotone had played a decisive role in population differentiation and speciation within Xiphorhynchus. Instead, phylogeny estimates suggested that the várzea–terra-firme habitat specialization evolved early on in the evolutionary history of Xiphorhynchus and that subsequent differentiation occurred mostly within the terra-firme habitat.

We examined genetic population structure of five putative subspecies of Song Sparrows (Melospiza melodia) in the San Francisco Bay region (M. m. samuelis, M. m. maxillaris, M. m. pusillula, M. m. gouldii, and M. m. heermanni) at nine microsatellite loci to assist the development of Song Sparrow conservation and management strategies. We sampled nine populations from five putative subspecies and found low estimates of differentiation between populations within subspecies and between. Despite low estimates of divergence, genetic structure at the subspecies level was indicated by the larger amount of variance accounted for by subspecies than populations. We propose that a management unit encompassing the range of M. m. pusillula be given priority for conservation on the basis of the extent of genetic divergence shown by Cavalli-Sforza and Edward's chord distance, and the topology of an unweighted pair group cluster analysis supported by 100% of bootstrap replicates across loci. Although M. m. samuelis and M. m. maxillaris appear undifferentiated from M. m. heermanni, it remains possible that adaptive differences between those types were not identified with neutral loci.

Vertical stratification of bird species has been intensively studied in temperate as well as tropical forest bird communities. One question that has received less attention is the variation of foraging stratum breadth. Hypotheses trying to explain that variation are based on differences in perception, evasion, adaptation, competition, and environmental and geometric boundary constraints. Although some predict that foraging strata near the ground should be narrower than those near the canopy, others predict that strata in the midstory should be broadest. These hypotheses were tested using observational data of 92 rainforest bird species. Observations were made from or near a 40 m high canopy crane situated close to the Surumoni River (3°10′N, 65°40′W) near Esmeralda, Estado Amazonas, southern Venezuela (Surumoni-Crane-Project). I found a highly significant parabolic relationship between mean foraging height and its standard deviation showing that midstory bird species foraged in a broader stratum than either understory or canopy species. Using a conservative comparative method, I also showed that the relationship was not the result of the phylogenetic relatedness of the species involved. Analysis of previously published data supports the relationship for two other tropical avifaunas. Because the ground and the outer canopy represent geometric boundaries to the movement of most forest birds, two simulation models were used to test whether midstory species may simply have a broader stratum because they are less constrained in their movements than species near the boundaries. The “random model” allowed individuals to move randomly between the two boundaries. It illustrated that boundary constraints alone cannot explain the observed parabolic relationship. In the “preference model,” individuals tended to move back to their preferred height within the forest column. That model simulated the observed parabolic relationship once a sufficient number of time steps had been reached, suggesting that a simple combination of geometric boundaries and preference for a specific foraging height may generate narrower strata near the boundaries. The other hypotheses only explain the parabolic relationship if other factors (e.g. vegetation density) are vertically distributed in such a way as to generate the observed parabolic pattern. Finally, I briefly discuss the advantages and disadvantages of using raw data in across-species analyses versus corrected data in phylogenetically-controlled analyses.

Little is known of the eastern North American population of Barrow's Goldeneyes (Bucephala islandica), which was recently listed as “of special concern” in Canada. In 1998 and 1999, we marked 18 adult males wintering along the St. Lawrence River, Québec, with satellite transmitters to document their breeding, molting, and wintering distribution and phenology, and to describe timing and routes of their spring, molt, and fall migrations. Thirteen males moved inland from the St. Lawrence River to breed; the spring migration averaged 5.9 days, and birds arrived on breeding areas on average 9 May. All breeding areas were inland, on the north shore of the St. Lawrence River estuary and gulf. Breeding areas averaged 64.8 km from the St. Lawrence corridor. Males stayed on their respective breeding area a mean of 34.5 days, and left on average 11 June. Twelve males were tracked to their molting areas, one of which stayed on its wintering area until 5 June and flew directly to its molting area. Their molt migration averaged 18.6 days, and the mean arrival date on molting areas was 30 June. All molting areas were located north and averaged 986 km from breeding areas. Four males molted in Hudson Bay, four in Ungava Bay, two in northern Labrador, one on Baffin Island, and one inland, near the Québec–Labrador border. The mean length of stay on the molting areas was 105.3 days, and the mean date of departure from molting areas was 4 October. All goldeneyes for which the radio still functioned during fall migration returned to winter in the St. Lawrence River estuary, on average 6 November. Our results refute the idea that the main breeding area of the eastern North American population of Barrow's Goldeneyes is located in northern Québec and Labrador and rather indicate that it is in the boreal forest just north of the St. Lawrence River estuary and gulf. They also indicate that Barrow's Goldeneye males undertake a genuine molt migration, and highlight the importance of molting areas because birds stayed there approximately four months each year.

Routine silvicultural practices continue to alter the structure and composition of forests after logging by removing deciduous vegetation from regenerating coniferous forests. We identified nest trees and surveyed vegetation in a 5 m radius surrounding songbird nests (nest patch) and compared the nest patches to available habitat in nine 11–22 year old conifer plantations (22–47 ha) where 90–96% of deciduous stems were removed by two treatments: manual thinning, and manual thinning plus application of glyphosate (herbicide). The control and two treatments were replicated three times. We characterized the nest patches of five species: Warbling Vireo (Vireo gilvus), Dusky Flycatcher (Empidonax oberholseri), Swainson's Thrush (Catharus ustulatus), American Robin (Turdus migratorius), and Chipping Sparrow (Spizella passerina). During three post-treatment years, areas treated with thinning plus herbicide remained depauperate of deciduous vegetation whereas thinned sites experienced deciduous regrowth. Despite variation in the density of deciduous trees and in the type of tree used for nesting in the control and treatments, nest patches were positively correlated with the amount of remaining deciduous vegetation, representing habitat that either escaped or recovered from silvicultural treatments. That relationship was stronger in areas with the fewest deciduous trees. Nests were more likely to be successful in areas with more willow. Within a mosaic of managed forest stands, birds appear to use the same proximate habitat cues for selecting a nest patch despite varying fitness consequences across different silviculture regimes. Although birds appeared to compensate for changes in stand habitat by finding patches of untreated vegetation or altering the type of tree they nested in, there was a reproductive cost for some species.

The wood-warblers (family Parulidae) fall within a radiation of passerine birds commonly known as the New World nine-primaried oscines. Defining familial relationships within that radiation has previously been challenging because of its extremely high diversity, a paucity of phylogenetically informative morphological characters, and an apparent high rate of cladogenesis early in the radiation's history. Here, analyses of mitochondrial and nuclear DNA sequences demonstrate that the 25 extant genera traditionally placed in the Parulidae do not form a monophyletic group. Instead, all reconstructions identify a well-resolved clade of 19 genera (Vermivora, Parula, Dendroica, Catharopeza, Mniotilta, Setophaga, Protonotaria, Helmitheros, Limnothlypis, Seiurus, Oporornis, Geothlypis, Wilsonia, Cardellina, Ergaticus, Myioborus, Euthlypis, Basileuterus, and Phaeothlypis) that are all morphologically typical wood-warblers traditionally placed in the Parulidae. Six genera traditionally assigned to the Parulidae—Microligea, Teretistris, Zeledonia, Icteria, Granatellus, and Xenoligea—fall outside this highly supported clade in all mtDNA-based and nuclear DNA-based reconstructions, and each is probably more closely allied to taxa traditionally placed in other nine-primaried oscine families. The long, well-supported, and independently confirmed internode at the base of this wood-warbler clade provides the opportunity to define a monophyletic Parulidae using several complementary molecular phylogenetic criteria. Support for those relationships comes from reconstructions based on a range of nucleotide-intensive (from 894 to 3,638 nucleotides per taxon) and taxon-intensive (45 to 128 species) analyses of mtDNA sequences, as well as independent reconstructions based on nucleotide substitutions in the nuclear-encoded c-mos gene. Furthermore, the 19 typical wood-warbler genera share a synapomorphic one-codon c-mos deletion not found in other passerines. At a slightly deeper phylogenetic level, our mtDNA-based reconstructions are consistent with previous morphologic and genetic studies in suggesting that many nine-primaried oscine taxa have unanticipated affinities, that many lineages arose during an early and explosive period of cladogenesis, and that the generation of a robust nine-primaried oscine phylogeny will require robust taxonomic sampling and extensive phylogenetic information.

Previously published morphological and behavioral data for the tyrant flycatchers (Tyrannidae) were reanalyzed using cladistic techniques. Several additional characters, including two putative synapomorphies of the Tyrannidae were incorporated. Nearly all of the ∼100 traditional tyrannid genera were included. Results of the analysis of this set of 68 characters support three previously proposed tyrannid assemblages: the kingbird assemblage and slightly restricted Empidonax and Myiarchus assemblages. Characters of the nasal septum that past workers have considered conservative and phylogenetically informative are supported as synapomorphies of the kingbird and restricted Empidonax assemblages. Several small monophyletic groups of genera are supported. The monophyly of neither the Elaenia assemblage nor the flatbill and tody-tyrant assemblage is supported in any most-parsimonious tree, but such trees are not significantly better than trees supporting their monophyly. A Tyrannidae exclusive of the Cotingidae and Pipridae is monophyletic in some most-parsimonious trees, but not in others in which flatbills and tody-tyrants are basal to a clade containing Cotingidae, Pipridae, and the remaining tyrannid genera. There is some evidence that characters describing plumage color pattern are more homoplasious than the other character suites I examined.

Age-related trends in breeding performance of female Eurasian Sparrowhawks (Accipiter nisus) were examined in south Scotland. Over a 22 year period, most individuals were banded and of known age. Previous analyses had shown that average number of young raised per breeding attempt (or year) increased progressively in successive age-groups to midlife (around five years) and declined thereafter. Here we analyze different components of breeding performance (such as clutch-size, hatching and fledging success) to find which components contribute most importantly to the overall age-related trend in productivity. Most aspects of breeding showed a similar upward–downward trend through the life span, but the downward trend in productivity in later life could be attributed mainly to poorer nestling survival. In general, in each aspect of breeding, the mean trend of the population as a whole was followed by individuals studied at different ages. Such individual trends were sufficient to account for the overall mean trend, and there was no need to postulate that high-performance or low-performance birds entered and left the breeding population at different ages. Only one aspect of breeding showed no trend with age, namely proportion of birds that laid having built a nest, but that type of failure affected only a small proportion of individuals.

Rapid assessment surveys of tropical bird communities are increasingly used to estimate species richness and to determine conservation priorities, but results of different studies are often not comparable due to the lack of standardization. On the basis of computer simulations and six years of field testing, we evaluated the recently proposed “20-species-list” survey method and statistical estimators for assessing species richness of tropical bird communities. This method generates a species-accumulation curve by subdividing consecutive observations of birds into lists of 20 species, thus relating cumulative species richness to the number of observations rather than time or space and thereby accounting for moderate differences in observer qualification and field conditions. Species accumulation curves from computer-simulated communities and two empirical data sets from Bolivia were analyzed with nine species richness estimators to evaluate estimator accuracy with respect to variations in species-list size, sample size, species-pool size, and community structure. For empirical and most simulated data sets, the MMMEAN estimator performed best, but it was more sensitive to differences in community structure than most other estimators. The CHAO 2 estimator, which was recommended by previous studies, performed reasonably well but was considerably more sensitive to sample size than MMMEAN. The bootstrap and first- and second-order jackknife estimators performed poorly. We recommend using MMMEAN or, when standard deviations of richness estimates are indispensable, CHAO 2 with 10-species lists for estimating species richness of tropical bird communities and propose a set of standard survey rules. Careful examination of estimator accumulation curves is required, however, and a technique based on the ratio between estimator and species accumulation curve is suggested to control for the confounding effects of sampling effort. Overall, the species-list method combined with statistical richness estimation is doubtlessly much more standardized and valuable than simple comparisons of one-dimensional locality lists and represents a promising tool for conservation assessment and the study of avian diversity patterns in the tropics.

We observed courtship and copulatory behavior in two populations of Phainopeplas (Phainopepla nitens) with contrasting social systems and used multilocus DNA fingerprinting to assess rates of extrapair fertilization (EPF). Phainopeplas from both territorial and loosely colonial populations copulated infrequently (≤2 times per day). No extrapair copulations (EPCs) were observed in either population. We found no evidence of EPFs in 48 nestlings from 25 nests. Thus, Phainopeplas appear to be both socially and genetically monogamous. However, Phainopeplas are not monogamous over their lifetimes—they do not retain mates from year to year and appear to switch mates between breeding localities within the same year. Compressed breeding seasons, geographically shifting food resources, and widespread breeding failures may favor monogamy during any given breeding attempt while allowing opportunities to switch mates between breeding localities and years.

Annual survival probability estimates for songbirds are generally biased low because dispersal is not differentiated from mortality. Presently, knowledge of between-year breeding dispersal is lacking for most songbirds. To assess adult survival probabilities and dispersal, we color-banded and resighted 436 Yellow Warblers (Dendroica petechia) over five breeding seasons at 11 study sites in the Bitterroot Valley, Montana. During the last two of those seasons, we searched extensively for marked warblers between and surrounding those sites. We compared survival probabilities estimated with and without this added dispersal information and assessed the effectiveness of adjusting survival probabilities with transient-type models. Survival probabilities were calculated using open population models, and model selection was based on Akaike's Information Criterion (AIC) within program MARK. The best model indicated that survival probabilities differed between males and females and varied among years. We found that dispersal off the study site was common (in 1999, 30% of resighted birds were found off their original study site), and survival probabilities increased by 6.5–22.9% (0.02 ± 0.07−0.106 ± 0.06) with the inclusion of dispersed birds. We suggest that emigration can have substantial effect on survival probabilities and advise against the use of return rates from small study areas or spatial or temporal comparisons of return rates because of spatial and temporal variation in extent of emigration. In addition, our results suggest that additional assessments of the reliability of transient-type models under some sampling schemes may be warranted.

The Carrion Crow (Corvus corone) is almost invariably reported as a species breeding as unassisted pairs, but a population of Carrion Crows (Corvus c. corone) from northern Spain that we have been studying since 1995 regularly exhibits cooperative breeding. The Spanish population thus demonstrates that cooperative breeding can vary greatly across populations. Most of the breeding territories (73.3%) in Spain were held by cohesive groups, which consisted of up to nine birds (mode = 3 birds). The proportion of fledglings that delayed dispersal for one year varied between 12.2 and 47.5% for different cohorts. The corresponding values for juveniles postponing dispersal for two years were 4.9 and 19.2%. Philopatry was skewed toward males. Some social groups contained immigrants. Therefore, delayed juvenile dispersal was not the only route towards sociality. Up to three helpers provided food to the nestling and at least one helper was found in all the groups surveyed. However, some group members refrained from visiting nests and thus the contribution to nestling care was highly variable among individuals.

Evolutionary theory predicts that heritability of fitness-related traits should be low. Egg size in birds is a fitness-related trait, but exhibits high heritability. One possible explanation for the apparent failure of natural selection to exhaust variability is that variation in egg size is mediated by female condition. In this study, we estimated repeatability of egg size within and between successive clutches in a wild, double-brooded population of House Wrens (Troglodytes aedon) in central Illinois, and examined the relationship of egg size with female size and condition. Repeatability of egg volume and mass in individual birds was high within years and between successive years (>0.77), suggesting a substantial heritable component to variation in egg size. However, egg mass was also significantly correlated with female condition. We suggest that the high repeatability values largely reflect permanent but nongenetic (ontogenetic) variation in egg size among females, which is reflected in the positive correlation of egg size with female condition. If variation in egg size is attributable to a combination of nonheritable, ontogenetic variation and variation due to current environmental conditions among females, then selection may not produce an evolutionary response in egg size.

A new species from Brazil—Pionopsitta aurantiocephala—is described, which is easily distinguished from the other species of that genus by its completely bare, intensely orange colored head. Specimens of this species have been historically identified as immatures of P. vulturina, which occur in simpatry with P. aurantiocephala on the Middle Tapajós River and possibly on the Lower Madeira River. The description of a new species of Psittacidae stresses the importance of new studies in the regions of the Madeira and Tapajós rivers.

Pitman and Jehl (1998) recently argued that Masked Boobies (formerly Sula dactylatra granti) breeding on the Nazca Plate in the eastern Pacific Ocean are morphologically and ecologically distinct from other Masked Boobies and may represent a full species. The American Ornithologists' Union subsequently elevated that subspecies to a full species: the Nazca Booby (S. granti). To evaluate that change in classification, we compared sequence variation in the mitochondrial cytochrome-b gene among 75 Nazca Boobies and 37 Masked Boobies representing three subspecies from the central and eastern Pacific and Atlantic oceans. Results indicated strong differentiation of cytochrome-b variation among taxa. Sequences constituted three distinct groups: Nazca Boobies, Masked Boobies from the central and eastern Pacific (S. d. personata and S. d. californica), and Masked Boobies (S. d. dactylatra) from the Caribbean and Atlantic. Those three groups probably diverged within a very short period, 400,000–500,000 years ago. Our results support the proposal that S. granti represents a distinct species.

In many avian species, reproductive success increases with parental age and experience, and declines seasonally. However, it is difficult to evaluate how parental attributes affect reproduction independently of date effects, because young, inexperienced pairs generally lay later than older, experienced pairs. We examined how parental experience and timing affected nestling growth in the Thick-billed Murre (Uria lomvia), an Arctic seabird that lays a single-egg clutch, by taking advantage of a natural contrast made possible by marked within-group synchrony. That is a common feature at murre colonies, and enabled us to monitor the breeding performance of experienced pairs over an extended period that overlapped with breeding by inexperienced pairs late in the season. Whereas growth of offspring raised by experienced parents was unaffected by their hatching date, offspring raised by inexperienced parents grew more slowly than those raised concurrently by experienced parents. Therefore, parental experience influenced nestling growth rates whereas timing did not, a result that accords with previous studies on this species. Absence of direct effects of timing of egg-laying on breeding success of Thick-billed Murres stands in sharp contrast to many other avian species, and seems surprising for an Arctic-nesting species often assumed to be strictly time-constrained in its breeding.

We satellite-tracked Steller's Sea Eagles (Haliaeetus pelagicus), Black-faced Spoonbills (Platalea minor), and White-naped Cranes (Grus vipio) to determine migration routes and stopover tactics of adult and immature birds. During spring migration, there were no significant differences between adult and immature birds in total migration distance and distance between stopover sites, but the migration period and resting period at stopover sites were significantly longer in immature individuals. There are two possible tactics for migrating without expending fat stores: longer rests or shorter migration distances between stopover sites. Our results suggest that immature birds try to increase fat stores by increasing time available for foraging at stopover sites.

Eleonora's Falcon (Falco eleonorae) breeds in the Mediterranean region and is highly adapted for catching small birds on passage migration between Eurasia and their African winter quarters, which they feed their young. We studied gliding flight behavior of Eleonora's Falcon at a breeding colony located on a small island southwest of Sardinia, Italy. Gliding and soaring flight performance was measured using an optical range finder and evaluated against flight mechanical theory. The male falcon does the majority of hunting and usually sets off from the colony to hunting areas located at high altitude over the open sea to catch prey. To lower the cost of transport and maximize the energy gain from hunting, we show that the birds use vertical winds for soaring when available. The occurrence of rising air changes with wind direction. At north-northwesterly winds (on-shore), slope lift is available outside the nesting cliffs, and at south-southeasterly winds thermals that form over the island drift out over the sea. Our observations demonstrated the flexibility of flight behavior in relation to the wind situation, and birds thereby make full use of available soaring conditions.

Maternal condition can affect reproductive investment in one or more ways. Captive female Red Junglefowl (Gallus gallus) that were heavier, controlling for tarsus length, produced larger eggs and initiated laying earlier, but did not produce more eggs per clutch. Maternal mass was also a positive predictor of offspring mass. That appears to result from the strong positive relationships between egg size and offspring mass. When the relationship between egg size and offspring mass was statistically controlled, maternal mass no longer predicted offspring mass. Mothers in better condition produced male-biased broods, as predicted when offspring condition depends on maternal condition and when male reproductive success is more dependent than female reproductive success on condition.

We continuously recorded temperatures of the nest cup (T_n) and air (T_a) just outside the nest box throughout development of Tree Swallow (Tachycineta bicolor) chicks in east-central Minnesota. Brood size was manipulated (three or six chicks per nest) to study the effect of number of chicks in the nest on its thermal environment. From day 0 to day 4, T_n paralleled T_a, approximately 2–7°C higher, with nocturnal T_n maintained at 24°C and daytime T_n maintained at 26°C. From day 6 to day 12, T_n was relatively constant throughout the day, maintained at 32°C (day) and 29°C (night). The T_n became increasingly independent of T_a, as determined by regression analysis of T_n versus T_a with age. Nocturnal T_n of larger clutches (six chicks) was ∼2°C warmer than smaller clutches (three chicks), and T_n of larger clutches exhibited greater independence from T_a at day 10 than in smaller clutches. The occurrence of nest homeothermy at day 10 in six-chick nests correlates with near-maximal body mass of chicks and completion of feather insulation. Small (three-chick) clutches showed greater dependence of T_n on T_a at 10–12 days of age than large (six-chick) clutches; we propose that development of thermoregulatory capacity may have proceeded more slowly in chicks from those nests.

Hughes (1997) hypothesized that the blue coloration of Black-billed (Coccyzus erythropthalmus) and Yellow-billed (C. americanus) cuckoo eggs evolved in response to egg discrimination by their hosts. From that hypothesis, we predicted that cuckoo hosts would eject ancestral nonmimetic (white) eggs, but be more likely to accept mimetic (blue) eggs. American Robins (Turdus migratorius) and Gray Catbirds (Dumetella carolinensis) were used to test that hypothesis because they are ejecter species that lay blue eggs and have been parasitized by Coccyzus cuckoos. Hughes' egg-mimicry hypothesis was not supported because robins and catbirds did not significantly eject white cuckoo eggs more frequently than blue cuckoo eggs. Indeed, most robins accepted both egg types. Furthermore, the observational evidence for egg mimicry is not convincing because only 5 of the 15 host species that Hughes (1997) described as having eggs that “fully or nearly match” the egg color of Black-billed and Yellow-billed cuckoos actually do so.

We experimentally parasitized Northern Mockingbird (Mimus polyglottos) nests to determine whether that species is more likely to eject immaculate eggs of the Bronzed Cowbird (Molothrus aeneus) or the spotted eggs of the Brown-headed Cowbird (M. ater); the latter species lays eggs that more closely resemble mockingbird eggs. Mockingbirds ejected 69% of model Bronzed Cowbird eggs, indicating that contrary to previous evidence Northern Mockingbirds eject Bronzed Cowbirds eggs at a high frequency. Mockingbirds also ejected 60% of model and real Brown-headed Cowbird eggs. Bronzed Cowbird eggs were ejected faster than Brown-headed Cowbird eggs (1.3 and 2.1 days, respectively). Ejection by mockingbirds may account for the lack of observed parasitism at our study sites in southern Texas, but it is more likely that cowbirds did not parasitize mockingbirds.

Numerous explanations exist for the evolution of reversed size dimorphism in raptorial species. A recent study concluded that reversed size dimorphism in skuas and jaegers was probably not attributable to breeding-role specialization, but that there was evidence for sexual selection, and in particular intrasexual competition by females for males. Our study tested the applicability of those conclusions for Southern (or Brown) Skuas (Stercorarius skua lonnbergi) breeding in South Georgia. Clutch volume was related positively to size and condition of females and negatively to condition of males, but there was no evidence of assortative mating for size or condition within pairs. Potential explanations for the discrepancy between this and previous studies are that size is less closely correlated with individual quality because of highly diverse foraging strategies, territory quality is a confounding factor, or because lower aggression in Southern Skuas reduces the necessity for small females to avoid large males.