Utah juniper (Juniperus osteosperma) has greatly expanded its historical range in the western United States. Management plans for the species have focused on curtailing its encroachment into sagebrush and grassland communities. These plans often include burning or other methods of elimination. These methods may result in subdivision or fragmentation of existing juniper stands. We initiated a study at Dugway Proving Ground, a U.S. Army facility, to examine the effects of fragmentation on the reproductive success of Utah juniper in isolated populations. We used enzyme electrophoresis to quantify genetic variability in isolated populations. We also determined population reproductive success by examining juniper fruits for evidence of seed abortion and/or presence of insect parasites. We compared reproductive and genetic variability in isolated populations at Dugway to 2 nonisolated and encroaching Utah juniper populations. The Dugway populations exhibited reduced seed set due to high seed abortion and/or insect seed parasitism, and a loss of genetic variability in comparison to the nonisolated populations. Additionally, there was a significant correlation between reproductive success and genetic variability.

Indian ricegrass (Achnatherum hymenoides [Roem. & Schult.] Barkworth) is a rangeland species native to western North America with populations that may exhibit distinct seed-size morphs borne on separate plants. These morphs are often associated with very different levels of seed dormancy. We compared RAPD profiles of the relatively infrequent and highly dormant jumbo seed morph to its more germinable small or large (non-jumbo) companion seed morphs collected from 9 sites where they were sympatric. Jumbo seed accessions from 9 additional sites that did not exhibit seed polymorphism were included to better sample this seed morph across the species' range. Sympatric seed morphs were consistently distinguishable by their RAPD profiles, and sympatric seed morphs were no more likely to cluster together than nonsympatric seed morphs. The ability of sympatric seed morphs to maintain their genetic identity is compatible with the hypothesis of a predominately autogamous breeding system. Furthermore, these data support the hypothesis that local polymorphic populations result from overlapping waves of migration with distinct geographical origins. The jumbo seed morphs exhibited moderate correlation between genetic distance and geographic distance, but the non-jumbo seed morphs exhibited no such correlation. Jumbo seed morphs that were collected at or east of the Continental Divide in New Mexico and Colorado are more genetically homogeneous than those west of the Continental Divide, and non-jumbo seed morphs from the east are more genetically variable than jumbo seed morphs from the same locations. This suggests that gene flow eastward across the Rocky Mountains has been more frequent for the non-jumbo seed morphs than for the jumbo seed morphs.

The purposes of this study were to develop a multivariate statistical model related to plant succession, to classify by seral stage, and to monitor succession in Wyoming big sagebrush shrubsteppe habitat (Artemisia tridentata ssp. wyomingensis/Agropyron smithii–Bouteloua gracilis) in Thunder Basin National Grassland, Wyoming. This model can be used by range and wildlife managers to evaluate management alternatives by assessing changes in plant species cover and composition within and between seral stages. Four ecological seral stages that represent early to late succession were quantitatively identified with an estimated 92% accuracy. Three key plant species provided the necessary information to define seral stages and monitor trends. Percent canopy cover and percent frequency (used to calculate index values: % canopy cover ×% frequency of occurrence) of Wyoming big sagebrush (Artemisia tridentata ssp. wyomingensis), western wheatgrass (Agropyron smithii), and blue grama (Bouteloua gracilis) are the only field measurements required for this model.

There is a paucity of information concerning livestock and vegetation responses to rotational grazing in short-grass steppe. We compared effects between 1995 and 2003 of time-controlled, short-duration rotational grazing against season-long continuous grazing, at a moderate stocking rate (1.95 ha ·animal unit month⁻¹), on livestock gains and on foliar and basal cover in short-grass steppe. Livestock average daily gains, grazing-season gains, and beef production did not differ between grazing systems. We observed no relationship between precipitation (annual or growing-season) and average daily gain. In contrast, both grazing-season gains and beef production exhibited a significant curvilin-ear response to both growing-season and annual precipitation. Basal and foliar cover of all plant functional groups (C₃ annual grasses, C₃ perennial grasses and grass-likes, C₄ perennial grasses, cactus, annual forbs, perennial forbs, and shrubs/subshrubs) did not differ between grazing systems. Litter and bare ground were also unaffected by grazing system, with litter increasing and bare ground decreasing over the duration of this experiment. Land managers in short-grass steppe who are implementing rotational grazing at moderate stocking rates should not expect increased livestock gains or changes in basal and foliar cover of the plant community. Rotational grazing may be a useful management tool for achieving conservation goals through modifications of seasonality and intensity of use in paddocks within larger management units to accommodate spatial variability in vegetation and temporal variation in precipitation.

We used live-trapping and foraging experiments to describe use of roadside vegetation by kangaroo rats (Dipodomys ordii) in short-grass prairie in Colorado, and to determine whether this species' perception of predation risk differed between roadside habitats, where predators were expected to be most active, and locations 120 m into adjacent grazed pastures. Giving-up densities (GUDs) were measured in paired seed trays placed beneath saltbush (Atriplex canescens) shrubs and in the open, 3 m from shrubs along transects in roadside and pasture locations. Trials were conducted on nights with a partial (1st-quarter), full, and new moon to assess how intensity of risk affected microhabitat use. Kangaroo rats were much more abundant in areas with saltbush cover than in grazed prairie, and were 4–6 times more numerous along roadsides in saltbush areas than in other locations. On dark nights and in pasture locations, foraging rates of kangaroo rats were similar in shrub trays and open trays. The fewest seeds were removed (i.e., GUDs were highest), however, in open trays along roads on moonlit nights, suggesting that kangaroo rats recognized potential risks associated with roadsides on bright nights. The high numbers of kangaroo rats along roads suggest that benefits associated with these habitats (ease of digging, dust bathing, higher soil seed banks) exceed the costs associated with higher risk of mortality from predators and vehicles. Our results demonstrate how foraging decisions differ depending on the spatial and temporal contexts in which behaviors are measured, and underscore the potential value of less common habitats such as road margins for increasing landscape-scale diversity and wildlife habitat in grazed grasslands.

Heteromyid rodents in the deserts of North America have been shown to harvest large quantities of seeds of both native and introduced plants from soil seed banks, but rarely has the impact of this seed removal been demonstrated experimentally. I used a series of fenced plots (some of which excluded rodents) to demonstrate that heteromyids at a western Nevada study site can measurably reduce seed banks and subsequent seedling establishment of Salsola paulsenii, an introduced invasive weed that has become a significant problem over much of the desert Southwest. The frequency of S. paulsenii seedlings in both 2004 and 2005 was significantly greater around the interior perimeters of plots that permitted access by rodents than in plots that excluded rodents. Density of S. paulsenii seedlings was significantly greater inside than outside rodent exclusion plots, but there was no such difference in seedling density inside versus outside plots that permitted rodent access. Salsola paulsenii has such a conspicuous presence in many desert environments that the effect of rodents in reducing its abundance may not be visually apparent; however, rodents may still ameliorate competitive effects of this weed on coexisting plants. Heteromyid rodents disperse seeds through caching, and they also consume them. Caching may enhance establishment of native plant seedlings, but is unlikely to benefit exotics such as S. paulsenii.

We documented black bears (Ursus americanus) consuming the seeds of Jeffery pine (Pinus jeffreyi) in the eastern Sierra Nevada from late August through September. Shells of pine seeds were the exclusive item in 20 of 27 scat samples collected in September and October and comprised >90% of the remaining 7 samples. Bears obtain seeds of Jeffrey pines by climbing trees, removing ripe, unopened cones from branches, dismantling cones on the ground, and consuming the seeds. Cone harvesting by bears can cause substantial damage to cone-bearing trees and might result in high predispersal mortality of seeds. In other regions, the use of pine seeds by bears has been associated with whitebark pines (P. albicaulis) from which bears obtain seeds by exploiting the cone-storing activities of squirrels. Despite the lack of an apparent pine-squirrel-bear interaction, harvesting unopened cones might be a common foraging strategy used by black bears to obtain seeds of Jeffrey pines in the Sierra Nevada.

Variation in kit fox (Vulpes macrotis) population parameters can be influenced by vegetative cover and the distribution and abundance of other predator and prey species. Dramatic changes to Great Basin Desert habitats, which can potentially impact mammalian species, have occurred in some areas in Utah. We examined kit fox demographics and prey populations from 1999 to 2001 on Dugway Proving Ground (DPG), a U.S. Army facility in Utah, and compared some parameters to historical levels (1956–1958–1966–1969). Adult survival rates were fairly consistent between 1999 and 2000 and between 1999 and 2001; however, survival was greater in 2001 than in 2000. Reproductive rates ranged from 1.0 to 3.8 pups per female in 1999–2000 and were similar to historical numbers (1.0–4.2 pups per female). We found a decrease in pre-whelping kit fox density from the 1960s (0.12 foxes · km⁻²) to 1999–2001 (0.04 foxes · km⁻²); however, densities were similar between the current study and the 1950s (0.08 foxes · km⁻²). Using 9 years of data, we found density dependence between reproductive rates of the current year and annual fox density from the previous year. Using 7 years of data, we found a slight correlation between kit fox annual density and a 1-year lag in leporid abundance, even though leporid abundance was lower during the present study than it was historically. Compared to historical levels, current small mammal abundance and species composition has changed in several habitats. Kit fox breeding density and annual density were inversely correlated with coyote (Canis latrans) density. Changes to the landscape at DPG, especially due to invasion of cheatgrass (Bromus tectorum) and addition of artificial water sources, have caused a change in available kit fox habitat and prey species, and have increased the abundance of coyotes, the kit fox's major competitor.

We studied 7 urban roosts occupied by Mexican free-tailed bats (Tadarida brasiliensis) in downtown Waco, Texas, from August 2001 to September 2002. We examined frequency of roost use, colony sizes, and roost fidelity in relation to gender. Use of roosts and colony sizes were highest in the fall and lowest in the winter, but fluctuated widely during the study. These roosts were not used by maternity colonies, and patterns of occupancy suggested that they served as stopover sites during migratory seasons. Individual bats exhibited high fidelity to discrete roosts included in the study. However, the bats commonly left the study area, suggesting that populations are defined at greater spatial scales than we included in this study. Male bats were recaptured more often and spent more days in the study area than female bats, suggesting a sexual difference in roosting behavior.

We used aerial surveys to study abundance and distribution of waterfowl (ducks, geese, swans, and coots) during spring in southern Oregon and northeastern California (SONEC). Total waterfowl-use days in SONEC during the 119-day, 5 January–3 May, spring period was similar during 2002 (127,977,700) and 2003 (128,076,200) and averaged 1,075,900 birds per day (bpd); these estimates should be adjusted upward 4%–10% to account for areas not surveyed. Waterfowl abundance peaked in mid-March in both years: 2,095,700 in 2002 and 1,681,700 in 2003. Northern Pintail (Anas acuta) was the most abundant species in both years, accounting for 25.6% of the 2002 and 24.5% of the 2003 waterfowl-use days. Pintail abundance peaked during the 13 March survey at 689,300 in 2002 and 532,100 in 2003. All other dabbling ducks accounted for 27.6% and 28.6%, diving ducks for 13.5% and 9.2%, geese for 24.6% and 29.3%, swans for 2.8% and 1.9%, and coots for 5.8% and 6.4% of the spring waterfowl-use days in SONEC during 2002 and 2003, respectively. Although use days changed little for total waterfowl ( 0.08%) and dabbling ducks (−0.1%), diving duck use was lower (−32%), and goose use days were greater ( 19%) in 2003 than in 2002. Distribution was similar in both years, with the most waterfowl use in the Lower (66%) and Upper (14%) Klamath subregions; 2%–6% occurred in each of the other subregions. Although the Lower Klamath subregion received the greatest overall waterfowl use, distribution among subregions varied among species and surveys, and all subregions were important during some part of the spring for 1 or more species. Peak spring abundance in SONEC during 2002 and 2003 averaged 50.3% of the midwinter abundance in California (all survey regions) and southern Oregon (69-3 survey region) for all waterfowl, 46.1% for dabbling ducks, 62.4% for diving ducks, 68.8% for geese, 109.4% for swans, and 43.8% for coots. Each spring, 75% of all waterfowl use in SONEC occurred on federal, state, or Nature Conservancy lands (i.e., protected areas). On protected areas there was a higher percentage of dabbling ducks (80.5%), geese (70.5%), and coots (81.5%) than diving ducks (60.4%) and swans (49%). Waterfowl use of Klamath Basin National Wildlife Refuge (NWR) averaged 42% greater during spring 2002–2003 (568,500 bpd) than during 1998–2001 (Gilmer et al. 2004). Numerous factors likely impacted magnitude and distribution of waterfowl use of SONEC during spring, including weather, waterfowl populations, SONEC habitat, and species ecology. SONEC is a critical spring staging area for waterfowl that winter in the Central Valley of California and other Pacific Flyway regions and should be a major focus area for waterfowl-habitat conservation efforts.

We investigated present indirect effects from a decade-old burn on the diets of stream fish. Based on soil instability and burn patterns from a 12–14-year-old wildfire complex, we separated 9 streams into 3 conditions: unburned, burned, and burned/scoured (i.e., experiencing a massive scour event 1 year post-burn). In the streams, we measured light levels and water temperatures, and we estimated fish density and biomass. We removed the digestive tracts from 9–15 rainbow trout (Oncorhynchus mykiss) from each stream and used a gravimetric procedure to analyze gut contents. Canopy cover development may dictate the composition of dietary items. Greater amounts of aquatic invertebrates and inorganic material were found in trout from streams with reduced overhead canopy. Further, these streams had lower amounts of terrestrial invertebrates and organic materials in the diets of fish. Although trout abundance was not different among the stream treatments, fire-induced, indirect effects on fish diets were still evident more than a decade after the burn. This suggests that recovery rates for trout assemblages may take longer than predicted and may depend on riparian recovery.

We sampled streams in the Upper Clear Creek Watershed in northwestern California in fall 2004 and fall 2005 to document assemblages of aquatic vertebrates and to provide resource managers with information on the importance of these assemblages in terms of regional biodiversity. We used single-pass backpack electrofishing to sample 15 sites in fall 2004 and the same 15 sites plus 4 new sites in fall 2005. We captured 10 fish taxa and 2 species of larval amphibians. Seven of the fish taxa were native species. Of the exotic species, only brook trout (Salvelinus fontinalis) occurred at more than 1 site. Ordinations by nonmetric multidimensional scaling indicated a gradient from sites with rainbow trout (Oncorhynchus mykiss), Pacific giant salamander (Dicamptodon tenebrosus), and tailed frog (Ascaphus truei) to sites dominated by riffle sculpin (Cottus gulosus), California roach (Hesperoleucas symmetricus), and Sacramento sucker (Catostomus occidentalis). The gradient in species composition was associated with changes in elevation, gradient, discharge, and substrate. The Upper Clear Creek Watershed represents a unique area of overlap between the North Coast California amphibian fauna and the Central Valley fish fauna with a notable paucity of exotic fishes and amphibians. Preservation of the integrity of native aquatic assemblages is an important goal for aquatic resource management in the region; our results provide a critcial baseline to gauge future management actions.

This study compared brood production of the spruce beetle (Dendroctonus rufipennis Kirby [Coleoptera: Curculionidae, Scolytinae]) in downed host material felled during summer and spring seasons on the Wasatch Plateau in south central Utah. Thirty-three matched pairs of Engelmann spruce (Picea engelmannii Parry ex Engelm.) trees were selected for study in spring 1996. One tree of each pair was cut during August 1996 (summer-felled), and the other tree was cut in early April 1997 (spring-felled), so that trees would be colonized by spruce beetles of the same flight period. Brood adults were collected and counted from bark samples, which were removed from the top, bottom, and sides of all sample trees in October 1998. The number of emergent adults produced in June 1999 was determined from exit holes counted in bark samples removed from these same locations. Mixed-model procedures were used to compare differences in the mean number of adults produced in summer-felled versus spring-felled trees in each year. The results indicated that significantly fewer spruce beetles were produced in summer-felled trees than in spring-felled trees. More brood adults were also present in, or emerged from, unexposed bole aspects (bottom, north, and east) of sample trees than exposed aspects (top, south, and west). These findings suggest that disturbances providing spruce beetles with an abundance of fresh host material in the spring result in the greatest potential for spruce beetle production, particularly beneath unexposed bark aspects. Examples of such disturbances include snow avalanches, blowdown, and snow and ice damage.

We report the discovery of established populations of Cnemidophorus neomexicanus in the Salt Lake City area. These are the 1st records of this species in Utah and in the Great Basin and are far from all other known populations of the species. We conclude that C. neomexicanus was introduced in the Salt Lake City area, perhaps within the last 30 years.