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Arctic fox Alopex lagopus and red fox Vulpes vulpes are close relatives with similar niche demands in the Holarctic. Where they are sympatric, they compete for territories, dens and food. Scat analyses from Fennoscandia have shown different proportions of lemmings and voles in the diets of the two fox species suggesting food partitioning. However, it was not clear if this was due to different food preferences or distinct habitat use. Since the arctic fox is an endangered species in Fennoscandia, it is important to know whether the superior, north spreading red fox can oust it from the tundra habitat, or if food specialisation may prevent displacement. In a feeding experiment at the Lycksele Zoo in northern Sweden, we compared the food preferences of two arctic and two red foxes. Our results show that the four individuals responded similarly to a variety of food items, and particularly that the two species were not distinct in their food preferences concerning lemmings and voles. However, the foxes had considerable individual predilections. Therefore, in the wild, the unequal proportions of lemmings and voles found in scats may reflect different habitat use for hunting.
In this paper, I describe historical and present harvesting and population trends of pine marten Martes martes in Scandinavia, based on a literature review and analyses of harvest statistics and population indices. The pine marten population has experienced two periods of over-harvesting with subsequent large-scale declines in population density and local extinctions; in the 1500–1600s and in the early 1900s. The principal incentive for harvesting appears to have been economic (valuable pelt), but eradication efforts may have compounded the effect on the population. In the last decades, the pine marten population density has increased. At present, it is receding but pine martens are still harvested intensively. I discuss implications for management, and caution about over-harvesting of the Scandinavian pine marten population in a near future.
To assess how important reindeer Rangifer tarandus fences are as a mortality factor for ptarmigan Lagopus spp. we collected data during 1991–1994. Our fieldwork covered 12 different sections of reindeer fence (totalling 71.1 km) in the county of Finnmark, northern Norway. The sections consisted of steel wire, steel netting or a combination of these, and ranged in height from 100 to 250 cm. The fieldwork took the form of spring patrols during which dead birds and their remains were searched for along the fences immediately after snow melt. We covered a total of 179.9 km and found 253 collision victims belonging to at least 20 species. Of the 253 victims found, 215 were willow ptarmigan Lagopus lagopus and rock ptarmigan L. mutus; thus these two species comprised 85% of the victims. During the winters of 1992/93 and 1993/94, we carried out experiments with dummy willow ptarmigan which were placed along fence sections and monitored serving as artificial fencestrike victims. Our experiments showed that approximately 64% of the total number of ptarmigan killed by the fences during winter would be detected during spring patrols along the fences. The type and height of the fence had no effect on the ptarmigan collision rate. In contrast, both the fence section and year factor contributed significantly to the observed variation in collision rate. We estimate that on average 1.4 ± 0.5 (SE) ptarmigan are being killed per kilometre of reindeer fence in Finnmark annually, with a greater variation between fence sections than between years.
Traditionally, pink-footed geese Anser brachyrhynchus wintering in Denmark, the Netherlands and Belgium have used the Danish sites only during mild winters, rapidly moving southwards in case of cold spells. Since the 1980s, an increasing number of geese have remained on the Danish wintering grounds despite cold spells, foraging on pastures and winter wheat Triticum aestivum fields. We compare the daily time and energy budgets and the food quality in the two habitats during winter. Winter wheat fields were increasingly used by the geese as temperatures dropped. At temperatures around 0°C, the geese foraged in both habitats, spending on average 83.8% and 74.9% of the daytime foraging in pastures and winter wheat, respectively. The estimated daily energy expenditure was slightly higher on pastures than on winter wheat fields (1,076 vs 1,057 kJ). The estimated daily food intake determined using the ‘marker substance’ method was 148 and 157 g ash free dry weight (AFDW) in geese feeding on pastures and winter wheat fields, respectively, equivalent to a daily net energy intake of 1,109 kJ and 1,145 kJ. Daily food intake, estimated on the basis of oesophagus contents of collected birds, was 170 g AFDW in pasture feeding geese and varied within 159–229 g AFDW in winter wheat feeding geese. In the mild winter, the protein content in winter wheat and Poa did not differ, whereas in the severe winter the protein content remained high in winter wheat but decreased in Poa. During the winters of 1994–1996, the abdominal profile index, API, in individually neck-banded geese observed repeatedly, only changed significantly during late January 1996. Neither during cold spells was there any change in API. Since the 1980s, the area covered by winter wheat has increased markedly in Denmark. Because winter wheat represents a reliable and profitable food source even in severe winters, the recent change in agricultural practice has enhanced the development of a new wintering strategy of pink-footed geese, allowing a northward expansion of their winter range. Potentially, this will increase the crop damage conflict and may lead to further population growth.
Brent geese Branta bernicla spring fattening around Agerø, Denmark, alternate between feeding on saltmarshes and submerged Zostera beds in Limfjorden. It appeared from field observations that these alternations depended on the water level in Limfjorden. A model was developed to assess the impact of water level fluctuations on the habitat use. A second model was developed to estimate the impact of water level on Zostera availability. The first model was successful in demonstrating that fluctuations in water levels had considerable influence on habitat use by the brent geese, i.e. they fed on Zostera at low water levels and on saltmarshes during high water levels, particularly so in early spring, and that the switch between habitats occurred within a narrow water level span of ca 30 cm. The second model demonstrated that the switch between habitats could be explained by lowered availability of Zostera as water levels increased. By combining the output from the two models, differences between years could partly be explained by differences in Zostera availability in the early spring period (21 March - 25 April), whereas a more complicated situation was detected later in spring (26 April - 31 May). The models presented may be considered as tools in investigations of habitat use and carrying capacity of seagrass beds in non-tidal areas, where birds' access to feeding areas regularly may be hindered by high water levels.
Species/habitat relationships and their responses at different scales are important aspects of ecological and conservational research. We studied the occurrence of hazel grouse Bonasa bonasia males in a forest reserve over a 10-year period at varying scales and population densities, using two sets of habitat descriptions. Avoidance of pine Pinus sylvestris was the only habitat effect in the hazel grouse/habitat analyses that was significant through all scales, seasons and densities. Thus, in spite of long-term data on a well-known species and detailed vegetation descriptions, only a few clear patterns relating to hazel grouse habitat selection were found at the relatively small scales analysed. We conclude that the non-significant relationships were due to the generally suitable composition and small variation of habitats within the study area, and that significant results may not be expected within the scales analysed and with the methods used. Thus, to find associations between animal species and habitats, a suitable study area must include a certain degree of habitat variation and the relationships should probably be examined at a scale equal to or larger than the home-range of the species in question, or considerably larger if population data are available. The results are discussed in relation to other studies and the applicability of the habitat descriptions and census techniques for conservation of hazel grouse populations in managed forests are discussed.
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