Takema Saitoh,^1,* Kazuto Kawakami,² Yaroslav A. Red'kin,³ Isao Nishiumi,⁴ Chang-Hoe Kim,⁵ Alexey P. Kryukov<sup>6

1</sup>Yamashina Institute for Ornithology, Division of Natural History, 115 Konoyama, Abiko, Chiba 270-1145, Japan
²Forestry and Forest Products Research Institute, 1 Matsunosato, Tsukuba, Ibaraki 305-8687, Japan
³Zoological Museum, Moscow State University, Bolshaya Nikitskaya Str., 2, Moscow 125009, Russia
⁴Department of Zoology, National Museum of Nature and Science, Tokyo, 4-1-1 Amakubo, Tsukuba, Ibaraki 305-0005, Japan
⁵Team of National Ecosystem Survey, National Institute of Ecology, 1210 Geumgang-ro, Maseo-myeon, Seocheon 33657, Republic of Korea
⁶Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, 159, Pr-t 100-letiya Vladivostoka, Vladivostok 690022, Russia

^* Corresponding author. E-mail: saitoh@yamashina.or.jp