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1 January 2000 The Vegetation of Mount Nyiru (Samburu District, Kenya): A Checklist and Syntaxonomical Survey
Benny Bytebier, Rainer W. Bussmann
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Abstract

The forests of Northern Kenya, and particularly of Mt Nyiru, have not been studied in detail, although they are extensive and play an important role in the life of the pastoralist communities surrounding them. Here, we present a checklist of the plants of Mount Nyiru and a syntaxonomic survey of the forest. Four hundred and thirty three taxa belonging to 284 genera and 102 families of higher plants are recorded. Thirty-nine taxa are new for the floral region K1. A syntaxonomic survey of the area shows that most forests belonged to the Juniperetea procerae (Montane Xeromorphic Cedar Forests). The Juniperion procerae, with the Faureo salignae-Ilicetum mitis on the wet mountain tops, and the Myrsino africanae-Juniperetum procerae in drier areas, were most commonly encountered. Interestingly, the top of Mt. Nyiru was covered with rather large stands of the Hagenietea abyssinicae (Subalpine Elfin Forests) but lacking Hagenia abyssinica itself, and extensive Sinarundinarietea alpinae (Bamboo Forests).

THE VEGETATION OF MOUNT NYIRU (SAMBURU DISTRICT, KENYA): A CHECKLIST AND SYNTAXONOMICAL SURVEYBenny Bytebier*East African Herbarium, National Museums of KenyaP.O. Box 45166, Nairobi, KenyaRainer W. BussmannLehrstuhl für Pflanzenphysiologie, Universität BayreuthD-95440 Bayreuth, Germanyceja.andina@gmx.deABSTRACTThe forests of Northern Kenya, and particularly of Mt Nyiru, have not been studied in detail, although they are extensive and play an important role in the life of the pastoralist communities surrounding them. Here, we present a checklist of the plants of Mount Nyiru and a syntaxonomic survey of the forest. Four hundred and thirty three taxa belonging to 284 genera and 102 families of higher plants are recorded. Thirty-nine taxa are new for the floral region K1. A syntaxonomic survey of the area shows that most forests belonged to the Juniperetea procerae (Montane Xeromorphic Cedar Forests). The Juniperion procerae, with the Faureo salignae- Ilicetum mitis on the wet mountain tops, and the Myrsino africanae-Juniperetum procerae in drier areas, were most commonly encountered. Interestingly, the top of Mt. Nyiru was covered with rather large stands of the Hagenietea abyssinicae (Subalpine Elfin Forests) but lacking Hagenia abyssinica itself, and extensive Sinarundinarietea alpinae (Bamboo Forests).INTRODUCTIONThe Northern Region of Kenya occupies nearly 50% of the land surface area of the country, yet it has only received marginal biological attention. For a long time, it was basically only visited by adventurers and big game hunters. The colonial name "Northern Frontier District" clearly illustrates this situation.It is still sparsely populated, and large parts are only accessible with difficulty. The few scientific studies mat have been carried out in the North, mainly focused on its geological features (Shackleton, 1946; Dixey, 1948; Dodson, 1963; Randel, 1967). The first, more comprehensive scientific research project, was the Unesco-IPAL study of the 1970's.* Current Address: University of Stellenbosch. Biochemistry Department, Private Bag XI, 7602 Matieland, South Africa, bytebier@maties.sun.ac.zaGeologyThe Northern Region consists of vast alluvial inland plains, inclining from altitudes of about 1200 m to the North of Mt. Kenya to barely 400 m around Lake Turkana. On the Southwest, a chain of mountains consisting of old cristallinic Precambrian basement rocks, mainly extremely durable gneisses and granites, borders the plains. For this reason, the steep Ndoto and Nyiru Ranges, reaching up to 2752 m, were left standing during the different erosion cycles influencing the region (Shackleton, 1946). Next to these, a series of Quarternary volcanic peaks, like Mt. Kulal (2285 m), Mt. Marsabit (1707 m) and the Huri Hills (1479 m), tower over the inland plains. In contrast to the soils of the plains, which consist mainly of Vertisols, Regosols, Lithosols and Cambisols, the mountain slopes are mainly covered with humic Acrisols over the basement formations, and deep, humic Andosols in volcanic areas (Mäckel, 1986; Mäckel & Schultka, 1983; Mäckel & Walter, 1983).ClimateAccording to the climatological classification of Jätzold (1977, 1981), the northern plains are part of the hot, arid tropical climate, with two short sub-humid seasons. Mean monthly temperatures range from 2026°C in the plains, to 17-19°C in the mountains (Gatab, 1657 m). The average annual rainfall can be as low as 100-150 mm in the Hedad plain and Chalbi desert, rising to 500 mm in the valleys of the Nyiru and Ndoto mountains. In the mountain forest zone, a rainfall of about 1200 mm can be reached (Edwards et al., 1979). The main rainfall is concentrated in two wet seasons, from March to May and from October to December. However, extreme rainfall occurs, e.g. 175 mm in 6 hours in Gatab on Mt. Kulal (Mäckel & Walter, 1983).VegetationMost mountain areas in Northern Kenya, located between 36°40'-38°00'E and 01°40'- 03°40'N, are covered with evergreen montane forest. They owe their existence to the humidity received from mist condensation and frequent cloud formation in the peak areas. Neumann (1898) after visiting the southern Turkana Region wrote: "The western face of Nyiru is.... topped with dark forests...., and here and there hang waterfalls., filled with the outpourings of the heavy clouds which often cap the summit." Due to their enormous importance for water catchment (Synott, 1979), most mountain areas are gazetted as forest reserves. The extent of these reserves, however, does not really reflect the amount of land actually covered with forest. The Mt. Nyiru forest reserve measures a total of 45,496 ha of which barely 7,890 ha are covered with true forest (Beentje, 1990).The main reasons for forest destruction in the area are fires caused by honey-hunters and pastoralists, who burn the old grass at the start of the wet season. Others are overgrazing in the forest, and serving the firewood needs of a fast-growing population.Despite their importance, few studies have been conducted on the montane forests of Northern Kenya. Herlocker (1979) gives some general remarks on the vegetation of the area and Synnott (1979) reports briefly on their status, importance and protection. Only Mount Kulal has received marginally more attention (Hepper, 1983), and a plant checklist for this area was produced (Hepper et al., 1981). Based on 20 relevés on this mountain, Schultka & Hilger (1983) distinguish mainly Olea hochstetteri-Cassipourea malosana and Olea africana-Juniperus procera forest.Synnot (1979) reports that much less is known on the Nyiru and Ndoto forests than on the Kulal and Marsabit forests. He provides a checklist for the trees and shrubs of Mount Nyiru, containing 35 species. White (1983) includes a short comment on the Marsabit region of Kenya and its afromontane forests, however mainly based on Synnott's observations from Mt. Kulal and Mt. Marsabit, without specific regard to Mt. Nyiru. Beentje (1990) mentions that the vegetation of Mount Nyiru is "mostly unknown".Although several collectors have visited the area, like J. Adamson (1947 and 1955), O. Kerfoot (1960), P.G. Archer (1971), J.B.C. Cameron (1972), G. Bono (1977), M. Ichikawa (1977) and M.G. Gilbert, F.N. Gachati and G.W. Gatheri (1978), no concerted effort has been made to compile the existing information.In the work presented here, the forests of Mt. Nyiru were studied in greater detail. A checklist is produced compiling our own with previous collections, and a detailed description of the vegetation texture of the forests, resulting in their syntaxonomic description according to Barkman et al. (1986), is given.MATERIALS AND METHODSPlant CollectionsThe majority of plants were collected between 29 March 1995 and 2 April 1995 in the Collector Series Bytebier B., Mwangangi O.M., Kirika P., Waiganjo T., Newton M. & Bussmann R.W., abbreviated to Bytebier et al. in the checklist. All specimens were deposited at the East African Herbarium in Nairobi (EA), with duplicates at the Royal Botanic Gardens, Kew (K) and the National Botanic Garden of Belgium, Meise (BR). Bussmann returned several times at a later date and his specimens (Collector Series Bussmann R.W.) are deposited at the Bayreuth University Herbarium. Based on identification lists from the EA archives, we retrieved herbarium specimens previously collected. All specimens were entered in the Brahms (Botanical Research And Herbarium Management System) database.NomenclatureThe nomenclature of plant families follows Bamps (1976), and in particular the available parts of the Flora of Tropical East Africa (FTEA, Turril et al., 1952-1998). The nomenclature of genera and species of Pteridophytes, Monocotyledones and Dicotyledones follows the new edition of "Upland Kenya Wild Flowers" (Agnew & Agnew 1994). The genus Sinarundinaria was treated according to Chao & Renvoize (1989). Cyperaceae and Juncaceae are named according to Haines & Lye (1983), Gramineae according to Phillips (1995). The nomenclature of trees and shrubs is according to "Kenya Trees, Shrubs and Lianas" (Beentje, 1995).RelevésDuring 1995-1996, 48 relevés were established and analyzed according to the methods of Braun-Blanquet (1964) and Mueller-Dombois & Ellenberg (1974), as slightly modified by Hammen et al. (1989). For a comprehensive description of the sampling methodology see Bussmann (1994), Bussmann & Beck (1995a) and Hammen et al. (1989).RESULTS AND DISCUSSIONHigher plants checklistThe checklist presented in Appendix 1 is based on the following collections and a few scattered collections leading to a total of 679 specimens, plus a few sight records by Bussmann RW.Adamson J28 specimens1947 & 1955/56Archer PG20 specimens1971Bono G39 specimens1977Bussmann RW45 specimens1995/96Bytebier et al.325 specimens1995Cameron JBC37 specimens1972Gilbert MG, Gachati FN and Gatheri GW18 specimens1978Ichikawa M5 specimens1977Kerfoot O156 specimens1960A total of 433 taxa belonging to 284 genera and 102 families are now on record. Amongst the 433 taxa there are 40 pteridophytes, 2 gymnosperms, 335 dicots and 56 monocots. Thirty-nine taxa from our own collections were previously not recorded from the floral region K1. They are indicated in the checklist as First Record for K1. The families best represented were Compositae (37 species), Leguminosae (34 species), Gramineae (20 species), Labiatae (18 species), Rubiaceae (16 species), Adiantaceae (12 species), Acanthaceae (12 species), Malvaceae (12 species) and Aspleniaceae (11 species).Forest types on Mt. NyiruA detailed phytosociological description of the forests of Mt. Nyiru, as well as other Northern Kenyan forest areas is included in Bussmann (in press), thus we present here an abbreviated floristical description of the forest types encountered.Montane Xeromorphic Cedar Forests (Juniperetea /-etalia BUSSMANN 1994)In the dry regions of Northern Kenya, closed forests are always restricted to mountain areas and hilltops, where mist condensation leads to more humid conditions.All forests encountered clearly belong to the Montane Xeromorphic Cedar Forests. Species such as Geranium arabicum and Achyranthes aspera were commonly growing in the ground layer, together with the grass Brachypodium flexum. Of the differential species of the Cedar-forests, Juniperus procera itself was common on Nyiru. In the herbal vegetation the tall grass Stipa dregeana, as well as Sanicula elata, were encountered regularly.Many forests of the dry Kenyan North belong to the pure Cedar forests (Juniperion procerae BUSSMANN 1994). The Myrsine-Juniperus forest (Myrsino africanae- Juniperetum procerae BUSSMANN 1994), with its pronounced fire-cycle, as the most typical association of the alliance, occurred less often; and the Faurea-Ilex forest (Faureo salignae-Ilicetum mitis BUSSMANN 1994) was the association found most commonly. Of the characteristic Cedar forest species, the East African Olive, Olea europaea ssp. africana was found only in few areas, particularly on southern Mt. Nyiru, forming sometimes almost exclusively the about 10 m high canopy. Other differential species were rarely found, and only in some places Rapanea melanophloeos, with its dark-green, shiny leaves appeared in the forest. The grass Ehrharta erecta, differential species for the Ehrharto erectae-Juniperetum procerae BUSSMANN 1994, occurred in many places, forming partly dense tufts on the forest floor on Mt. Nyiru. The floristic composition of the forests, however, indicated without doubt that most stands belonged to the pure Faureo salignae-Ilicetum mitis BUSSMANN 1994.This association, first described from the northeastern slopes of Mt. Kenya (Bussmann & Beck, 1995a), clearly dominated most of the mountain areas studied. In the type locality, forests of this type were encountered only as remnants on very steep slopes with relatively shallow soils, whereas in the North in a very striking contrast, the Faureo-Ilicetum was encountered under almost all slope conditions. Only the Proteacean Faurea saligna, with its leathery leaves shining red in the green canopy, grew with high abundance, often dominating the canopy, while Ilex mitis was completely absent. In contrast to the lack of differential species, the accompanying flora encountered was very characteristic, with e.g. Piper capense, the leathery fern Arachnoides foliosa or the stinging nettle Laportea alatipes, among many other species, the typical companions of the association were found abundantly. Nevertheless, in comparison to the stands on Mt. Kenya, the floristic composition of the Faureo-Ilicetum showed many differences, especially with regard to the shrub stratum, and therefore the northern stands have to be regarded as a distinct subassociation of their own, differentiated by the very abundant occurrence of Xymalos monospora, often forming a dense shrub layer, a fact which was never observed in the typical Faureo-Ilicetum. In addition, Xymalos, normally found only as a small shrub, reaches up into the lower canopy in many places forming a second, lower tree stratum of its own. In the shrub layer Brucea antidysenterica and the climbing Clerodendrum johnstonii, as well as Clausena anisata, were also encountered as differential species. The ferns Pteris quadriaurita and Doryopteris kirkii, together with the otherwise rare liana Clematis brachiata, and especially the Acanthacean Dicliptera colorata, in places covering large areas of the forest floor, are also differential, and so are the epiphytic orchids Polystachya piersii and Aerangis thomsonii, the latter often dangling in carpets from its sustaining branches. The most striking feature of these forests however, is the abundance of very old, and therefore extremely tall and thick specimens, of Faurea saligna, some of which reached a girth of more than 2 m at breast height indicating, that these forests have never been disturbed by logging.Differences in humidity lead to two variants of these Xymalos-dominated forests. The most commonly encountered wet variety, growing mainly on higher altitudes, clearly receives more moisture, which was shown by the many fern species in the undergrowth. Of them, Amauropeltis bergiana and Stenogramma pozoi were regarded as differential species. Further differential taxa, all requiring high humidity were the creeping Droguetia iners and Crassula alsinoides, Drymaria cordata, Dicrocephala integrifolia, together with the Urticacean Pilea johnstonii, and Aneilema pedunculosum. In the higher strata Solanum schumannianum and the tree Schefflera volkensii appeared. Of the companions, the rare Tarenna graveolens, Chionanthes battiscombei and Turraea holstii, appearing mainly in the shrub stratum, as well as Pavetta gardeniifolia deserve special note.At the lower limit of the Faurea-Ilex forests, in transition to the Myrsine-Juniperus- Cadia forest (Myrsino-Junipertum cadietosum purpureae, Bussmann, in press), almost all of the less drought resistant species, especially Pteridophytes, disappeared due to the much drier conditions. Whereas in the previous variant the forest floor was always covered with living herbs and also with an often dense grass layer even at the peak of the dry season, in the lower forests the ground was often found bare, as all vegetation had disappeared due to the drought. Differential species encountered were the tall Plectranthus barbatus, Stellaria sennii and the Cyperacean Schoenoxiphium lehmannii. Because livestock often grazes these areas at the beginning of the dry season, weed-like species like Solatium incanum, Pupalia lappacea and Pteridium aquilinum (the latter being an indicator of fire), also occurred as differential species. Important companions were Crassocephalum montuosum, Desmodium repandum, Hypoestes forskahlii, Leonotis nepetifolia, Microglossa pyridifolia and Mikaniopsis bambuseti.The drier lower slopes of Mt. Nyiru were covered by a vegetation belonging to the Myrsino africanae-Juniperetum procerae BUSSMANN 1994, forming the transition zone to the savanna areas. Fires occur regularly in this area. They are lit by pastoralists to improve the grass growth before the start of the rains or by honey hunters smoking out bees. At long intervals they also occur naturally. Therefore the Myrsino-Juniperetum shows the fire cycle of the Cedar-forests best (Bussmann & Beck, 1995b). Consequently Myrsine afiicana, a differential species for these forests, and one particularly indicating the influence of fire, was found with very high cover/abundance in most areas, often forming a second, lower shrub stratum. On Mt. Nyiru, Juniperus procera occurred with high cover also. Of the characteristic species, only Rhamnus prinoides was found although much less abundant. Interestingly, Nuxia congesta appeared often with high cover in the tree stratum, and the Acanthacean Justitia striata formed patches on the forest floor. The high abundance of Teclea nobilis in the shrub layer and the lower tree stratum is also worth mentioning. The presence of a rather different flora, in comparison to other areas, e.g. Mt. Kenya (Bussmann & Beck, 1995a), description of a new subassociation (Bussmann, in press).In transition to a dense thorny bushland, formed mainly by species of the genera Commiphora, Grewia and partly Acacia, this vegetation type was encountered on the steep rocky lower slopes of Mt. Nyiru. With an often very dry ground layer, leaving many areas of the rocks exposed, forests of this type showed a very open appearance and due to the frequent fires even the higher shrub stratum had been nearly completely destroyed. Many Juniperus trees in these areas were found dead or dying, and due to the frequent fires young specimens were rarely observed. In the open shrub stratum, Cadia purpurea and Vangueria apiculata occurred as differential species of this subassociation, together with the small tree Cordia monoica in the canopy. The set of differential species was completed by a high number of drought resistant ferns, especially of the genus Cheilanthes, growing among the rocks. Namely Pellea alchemilloides, Cheilanthes bergiana, C. multifida, C. tecta and C. hirta were encountered, whereas the very rare Asplenium trichomanes occurred on some high, shady cliffs. The companions, the Euphorbiacean Croton megalocarpus in the canopy and the Rutacean Teclea simplicifolia in the shrub stratum, deserve special note, as they showed clear links to the Brachylaenion huillensis BUSSMANN 1994, an alliance mainly found in Central and Southern Kenya. Scadoxus multiflorus, with its large bright-red inflorescence was also found as a companion, among many other species.Subalpine Elfin forests (Hagenietea abyssinicae BUSSMANN 1994.)Large grassy clearings were only found on top of Mt. Nyiru. Many huts used by Samburu pastoralists during the dry season indicated that these areas are heavily grazed. At the borders of the grasslands, and partly as islands in-between, dense thickets of St. John's Wort Hypericum revolutum) were growing together with young specimens of Juniperus procera. The high cover/abundance of Hypericum indicated that these forests belong to the Hagenia-Hypericum forest (Hagenio abyssinicae-Hypericetum revoluti BUSSMANN 1994). Whether Hagenia abyssinica itself has ever grown in these areas remains an enigma as no koso trees were found. The high cover of young Juniperus trees, all of the same age, has to be regarded as a sign of a very large fire about 10 years ago (judged by the size of the trees). Only few dead specimens of old Cedars were found. It is assumed that in the successional process the Hagenio-Hypericetum on Nyiru will probably be replaced by the Myrsino-Juniperetum, especially with regard to the more frequent use of the area as dry season pasture, leading to more frequent burning.At present, despite the growing influence of Juniperus, the Thymeleaceae Gnidia glauca, with its yellow flowers appearing in dense clusters in the wet season, still dominates the canopy of these forests, forming often closed stands about 8 m tall. Therefore, the topmost forests of Mt. Nyiru clearly belong to the Gnidietum glaucae BUSSMANN 1994.Montane Bamboo Forests (Sinarundinarietea alpinae BUSSMANN 1994)The East African Bamboo, Sinarundinaria alpina, is known to cover vast areas especially of the wet southern and south-eastern slopes of Mt. Kenya and the Aberdare and Mau Ranges in Central Kenya. In other areas of the country, however, the species occurs only rarely and in Northern Kenya bamboo is nearly absent. In some areas of Nyiru, probably in places with the highest amount of condensing mist, Sinarundinaria alpina was encountered forming dense stands up to 6-8 m tall. These bamboo forests were very similar to the ones described from Western Mt. Kenya (Bussmann & Beck, 1995a), with Podocarpus latifolius frequently protruding from the closed bamboo stands. All stands studied were very dense, with only a small amount of light reaching the ground, which was therefore often bare of vegetation. Trails of large game, e.g. elephants and buffaloes, very frequent in other areas where bamboo grows, were not found on Nyiru. This made the bamboo forests nearly impenetrable. The presence of Sinarundinaria on Mt. Nyiru has to be regarded as of special importance with respect to the biogeography of the species. The Nyiru population can be interpreted as a link between the main growing area of the species and the stands on the southern slopes of the Bale mountains in Southern Ethiopia, about 500 km further North (Bussmann, 1997).ACKNOWLEDGEMENTSThe authors would like to thank Onesmus Mwangangi, Paul Kirika, Titus Waiganjo and Mike Newton, who joined the initial expedition; Elizabeth Muthuma, David Kones and Joyce Chege, who assisted in preparing the checklist; and Emma and Yoav Chen from Desert Rose for logistical support in the field. Rainer Bussmann acknowledges the financial support for this work by the Deutsche Forschungsgemeinschaft (DFG) and wishes to thank the National Research Council of Kenya for granting permission for research. The expedition to Mount Nyiru was funded by the Foundation for the Furtherance of Scientific Research in Africa (Belgium).REFERENCESAgnew, A.D.Q. & S. Agnew (1995). Upland Kenya Wild Flowers. East Africa Natural History Society, Nairobi.Baker, B.H. (1960). Geology of the Baragoi Area. Geological Survey of Kenya, Nairobi.Bamps, P. (1976). Catalogue of the phanerogamic families dealt with in the main floras of Tropical Africa. Boissiera 24: 667-686.Barkman, J.J., J. Moravec & S. Rauschert (1986). Code of phytosociological nomenclature 2nd ed. Vegetatio 67: 145-195.Beentje, H.J. (1990). The forests of Kenya. Mitteilungen des Instituts fuer Allgemeine Botanik Hamburg 23a: 265-286.Beentje, H.J. (1995). Kenya Trees, Shrubs and Lianas. National Museums of Kenya, Nairobi.Braun-Blanquet, J. (1964). Pflanzensoziologie. Third edition. Springer, Wien, New York.Bussmann, R.W. (1994). The forests of Mount Kenya (Kenya). Vegetation, ecology, destruction and management of a tropical mountain forest ecosystem. Ph.D. thesis, Universität Bayreuth.Bussmann, R.W. (1997). The forest vegetation of the Harenna Escarpment (Bale Province, Ethiopia)syntaxonomy and phytogeographical affinities. Phytocoenologia 27(1): 1-23.Bussmann, R.W. (in press). 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Hogg (1979). A prelimnary analysis of climatological data from the Marsabit District of Northern Kenya. IPAL Technical Report B-l. Unesco, Nairobi.Haines, R.W. & K.A. Lye (1983). The Sedges and Rushes of East Africa. East Africa Natural History Society, Nairobi.Hammen, T.v.d., D. Mueller-Dombois & M.A. Little (1989). Manual of Methods for Mountain Transect Studies. IUBS.Hepper, F.N. (1983). The phytogeography of Mt. Kulal, Kenya, with special reference to Compositae, Leguminosae and Gramineae. Bothalia 14: 534-551.Hepper, F.N., P.M.L. Jaeger, J.B. Gillett & M.G. Gilbert (1981). Annotated checklist of the plants of Mount Kulal. IPAL Technical. Report D-3. Unesco, Nairobi.Herlocker, D. (1979). Vegetation of southwestern Marsabit district, Kenya. IPAL Technical Report D-l. Unesco, Nairobi.Jätzold, R. (1977). Klimageographie, Serie E, Blatt 5, 1:1000000. Bornträger, Berlin, Stuttgart.Jätzold, R. (1981). Klimageograpgie Ostafrika-Afrika Kartenwerk, Serie E, Beiheft zu Blatt 5. 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Edwards, (eds.), Flora of Ethiopia and Eritrea, Vol. 7. National Herbarium, Addis Ababa.Randel, R.P. (1967). Geology of the Laisamis Area. Government Printer, Nairobi.Schultka, W. & H.H. Hilger (1983). Epizoochore Verbreitung in der Krautschicht beweideter Bergwälder des Mt. Kulal (Nordkenia). Beitraege zur Biologie der Pflanzen 58: 333-356.Shackleton, R.M. (1946). Geology of the country between Nanyuki and Maralal. Geological Survey of Kenya Report 11. Survey of Kenya, Nairobi.Survey of Kenya (1959, 1970). National Atlas of Kenya. Survey of Kenya, Nairobi.Survey of Kenya (1966). East African mean monthly rainfall in millimetres (North sheet: Kenya and Uganda, Scale 1:2.000.000). Survey of Kenya, Nairobi.Synott, T.J. (1979). A report on the status, importance and protection of the montane forests. IPAL Technical Report D-2a. Unesco, Nairobi.Turril W.B., C.E. Hubbard, E. Milne-Redhead, R.M. Polhill & H.J. Beentje. (eds.) (1952-1998): Flora of Tropical East Africa. Crown Agents, London, and A.A. Balkema, Rotterdam.White, F. (1983). The vegetation of Africa. Unesco, Paris.APPENDIX 1: Checklist of the plants of Mount NyiruPTERIDOPHYTAACTINIOPTERIDACEAEActiniopteris semiflabellata Pic.Serm.Bytebier B et al. 344; Bussmann R 974316002450 mADIANTACEAEAdiantum capillusveneris L.Bytebier B et al. 314; Cameron JBC 151;Gilbert MG, Gachathi FN & Gatheri GW5209; Archer PG 689; Bussmann R 973412192450 mAdiantum poiretii Wikstr.Bytebier B et al. 7624002400 mCheilanthes bergiana KunzeBussmann R 981123502550 mFirst record for K1Cheilanthes farinosa (Forssk.) Kaulf.Bytebier B et al. 252; Bussmann R 980922002450 mCheilanthes hirta Sw.Bussmann R 981223502550 mCheilanthes marantae (L.) DominCameron JBC 14816761676 mCheilanthes multifida (Sw.) Sw.Bytebier B et al. 253, 329, 336; GilbertMG, Gachathi FN & Gatheri GW 5196;Bussmann R 981816002450 mCheilanthes tecta F.M.Jarret ined.Bussmann R., Sight record23502550 mDoriopteris kirkii (Hook.) AlstonBussmann R 975223502550 mPellaea adiantoides (Willd.) J.Sm.Bussmann R 979723502550 mPellaea calomelanos (Sw.) LinkBytebier B et al. 33916001600 mPellaea longipilosa Bonap.Bytebier B et al. 327; Bussmann R 982216002450 mASPLENIACEAEAsplenium abyssinicum FéeCameron JBC 14924392439 mAsplenium adiantumnigrum L.Cameron JBC 13325152515 mAsplenium aethiopicum (Burm.f.) Bech.Bytebier B et al. 72; Gilbert MG, GachathiFN & Gatheri GW 5193; Cameron JBC146; Bussmann R 969816002450 mAsplenium elliottii C.H.WrightBytebier B et al. 17A; Bussmann R 970124002450 mAsplenium erectum Willd. var.usambarense (Hieron.) SchelpeBytebier B et al. 38, 251; Bussmann R 970822002450 mAsplenium friesiorum C.Chr.Bytebier B et al. 6, 17B; Cameron JBC144; Bussmann R 971522862450 mAsplenium loxoscaphoides BakerBussmann R 974023502550 mAsplenium monanthes L.Bytebier B et al. 73, Cameron JBC 137;Bussmann R 970421342450 mAsplenium strangeanum Pic.Serm.Bytebier B et al. 250; Cameron JBC 13622002286 mAsplenium theciferum (Kunth) Mett.Bytebier B et al. 74; Cameron JBC 134;Bussmann R 970621342450 mAsplenium trichomanes L.Cameron JBC 135; Gilbert MG, GachathiFN & Gatheri GW 5195; Bussmann R 971016002450 mDENNSTAEDTIACEAEHypolepis goetzei ReimersCameron JBC 13921342134 mPteridium aquilinum (L.)Kuhn Bytebier B et al. 19425002500 mDRYOPTERIDACEAEArachniodes foliosa (C.Chr.) SchelpeCameron JBC 13024392439 mPolystichum fuscopaleaceum AlstonBytebier B et al. 50; Cameron JBC 147;Bono G 23; Bussmann R 9803, 979624002450 mPOLYPODIACEAELoxogramme abyssinica (Baker)M.G.PriceBytebier B et al. 4624002400 mPleopeltis macrocarpa (Bory ex Willd.)Kaulf.Cameron JBC 131, 14524392439 mPTERIDACEAEPteris catoptera KunzeBytebier B et al. 6924002400 mPteris dentata Forssk.Bono G 24, 38, 211; Cameron JBC 143, 15212192439 mPteris quadriaurita Retz.Bussmann R 981423502550 mSCHIZAEACEAEMohria vestita BakerBytebier B et al. 25422002200 mFirst record for K1SELAGINELLACEAESelaginella dregei (C.Presl) Hieron.Alexander EAH 11859THELYPTERIDACEAEAmauropelta bergiana (Schltdl.) HolttumBussmann R 979923502550 mAmauropelta oppositiformis (C.Chr.)HolttumBytebier B et al. 22225002500 mFirst record for K1Stegnogramma pozoi (Lag.) K.Iwats.Bytebier B et al. 48; Bussmann R 9791, 979224002450 mWOODSIACEAECystopteris diaphanum (Bory) BlasdellBytebier B et al. 25; Cameron JBC 15024002439 mCystopteris fragilis (L.) Bernh.Bussmann R 979523502550 mGYMNOSPERMAECUPRESSACEAEJuniperus procera Endl.Adamson J 393, Adamson J B 6162;Bytebier B et al. 18824392500 mPODOCARPACEAEPodocarpus latifolius (Thunb.) Mirb.Bytebier B et al. 96; Cameron JBC 128;Adamson J 39224392500 mANGIOSPERMAEDICOTYLEDONAEACANTHACEAEAcanthopale pubescens C.B.ClarkeKerfoot O 206724392439 mCrossandra massaica Mildbr.Bytebier B et al. 32316001600 mDicliptera colorata C.B.ClarkeBussmann R 978623502550 mDicliptera laxata C.B.ClarkeKerfoot O 206827432743 mDyschoriste radicans NeesBytebier B et al. 18925002500 mHypoestes forskahlii (Vahl) R.Br.Bytebier B et al. 269Kerfoot O 200123502439 mHypoestes triflora (Forssk.) Roem. & Schult.Bytebier B et al. 210Kerfoot O 207624392500 mIsoglossa gregorii (S.Moore) LindauBytebier B et al. 58Kerfoot O 206924002743 mJusticia glabra Koen. ex Roxb.Bytebier B et al. 31216001600 mJusticia lorata EnsermuBytebier B et al. 341,35016001600 mFirst record for K1Justicia striata (K1.) BullockBussmann R, Sight record23502550 mThunbergia alata Bojer ex SimsBytebier B et al. 258; Kerfoot O 209023502591 mAMARANTHACEAEAchyranthes aspera L.Bytebier B et al. 4424002400 mCelosia anthelminthica Asch.Kerfoot O 202424392439 mCelosia schweinfurthiana SchinzBytebier B et al. 30916001600 mCyathula polycephala BakerBytebier B et al. 6624002400 mFirst record for K1Pupalia lappacea (L.) A.Juss.Bytebier B et al. 24822002200 mSericocomopsis hildebrandtii SchinzKerfoot O 208027132713 mANACARDIACEAERhus natalensis KraussBytebier B et al. 278; Kerfoot O 1948, 1949, 195021342350 mRhus ruspolii Engl.Bytebier B et al. 276; Kerfoot O 194521342350 mAPOCYNACEAECarissa edulis (Forssk.) VahlKerfoot O 202321342134 mARALIACEAECussonia holstii Engl. var. holstiiBytebier B et al. 262; Kerfoot O 1942;Bono G 32321342350 mSchefflera volkensii (Engl.) HarmsBussmann R, Sight record23502550 mASCLEPIADACEAECeropegiaBytebier B et al. 20125002500 mCeropegia ballyana BullockBytebier B et al. 24922002200 mFirst record for K1Cynanchum altiscandens K.Schum.Bytebier B et al. 26323502350 mFirst record for K1Gomphocarpus fruticosus (L.) W.T.AitonBytebier B et al. 240; Kerfoot O 197922002591 mPergularia daemia (Forssk.) Blatt. & MacOwanKerfoot O 2021; Bono G 3318001829 mPeriploca linearifolia Quart.Dill. & A.Rich.Kerfoot O 199824392439 mSecamone punctulata Decne.Bytebier B et al. 294, 36316001600 mBALSAMINACEAEImpatiens hochstetteri Warb.Bussmann R 978523502550 mImpatiens meruensis GilgCameron JBC 13822862286 mImpatiens meruensis Gilg ssp.septentrionalis GreyWilsonBytebier B et al. 51; Kerfoot O 208224002743 mImpatiens sodenii Engl.Bytebier B et al. 325; Kerfoot O 2083;Bono G 202; Adamson TG 18; BussmannR 978316002743 mBEGONIACEAEBegoniaBytebier B et al. 32416001600 mBERBERIDACEAEBerberis holstii Engl.Kerfoot O 193925912591 mBORAGINACEAECordia monoica Roxb.Kerfoot O 1938; Bono G 2621342300 mCynoglossum coeruleum A.DC.Bytebier B et al. 41, 17624002500 mCynoglossum coeruleum A.DC. ssp.johnstonii (Baker) Verde.Kerfoot O 209527432743 mLithospermum afromontanum Weim.Kerfoot O 200622862286 mCALLITRICHACEAECallitriche stagnalis Scop.Bytebier B et al. 231; Bono G 20824502500 mCAMPANULACEAECampanula edulis Forssk.Archer PG 695; Kerfoot O 209722862743 mWahlenbergia abyssinica (A.Rich.) Thulin ssp. abyssinicaBytebier B et al. 92; Cameron JBC 108;Kerfoot O 2063; Adamson J 541;Bussmann R 977821342743 mWahlenbergia capillacea (L.f.) A.DC. ssp.tenuior (Engl.) ThulinBytebier B et al. 9325002500 mWahlenbergia lobelioides (L.f.) A.DC. ssp.nutabunda (Guss.) Murb.Archer PG 69722862286 mWahlenbergia virgata Engl.Bytebier B et al. 11; Adamson J 540;Archer PG 69424002439 mCAPPARACEAECapparis tomentosa Lam.Bytebier B et al. 334; Kerfoot O 198516002591 mCleomeKerfoot O 206221342134 mCleome usambarica PaxBytebier B et al. 31816001600 mCrateva adansonii DC.Kerfoot O 199015241524 mGynandropsis gynandra (L.) Briq.Bytebier B et al. 28823502350 mMaerua angolensis DC.Kerfoot O 193119811981 mMaerua triphylla A.Rich.Bytebier B et al. 34516001600 mThylachium africanum Lour.Kerfoot O 195824392439 mCARYOPHYLLACEAECerastium indicum Wight & Arn.Bytebier B et al. 164; Kerfoot O 207325002743 mDrymaria cordata (L.) Willd. ex Roem. & Schult.Bytebier B et al. 1; Kerfoot O 209624002743 mPollichia campestris AitonBytebier B et al. 238Kerfoot O 209922002743 mSilene burchellii Otth ex DC.Adamson J 55227432743 mSilene macrosolen Steud. ex A.Rich.Kerfoot O 207427432743 mStellaria sennii Chiov.Bytebier B et al. 21525002500 mCELASTRACEAEMaytenus heterophylla (Eckl. & Zeyh.)N.RobsonBytebier B et al. 27923502350 mMystroxylon aethiopicum (Thunb.) Loes.Bytebier B e/a/. 31916001600 mCHENOPODIACEAEChenopodium schraderianum Schult.Bytebier B et al.289 23502350 mCOMPOSITAEAdenostemma perrottetii DC.Gilbert MG, Gachathi FN & Gatheri GW5210; Kerfoot O 205613502743 mBerkheya spekeana Oliv.Archer PG 70525602560 mBidens flagellata (Sherff) MesfinBytebier Bet al. 21325002500 mBidens hildebrandtii O.Hoffm.Bytebier B et al. 31716001600 mBidens kilimandscharica (O.Hoffm.)SherffKerfoot O 205724392439 mBothriocline longipes (Oliv. & Hiern)N.E.Br.Bytebier B et al. 20; Kerfoot O 204921342400 mCarduus nyassanus (S.Moore) R.E.Fr.Bytebier B et al. 23225002500 mFirst record for K1Cineraria deltoidea Sond.Kerfoot O 204621342134 mConyza newii Oliv. & HiernBytebier B et al. 19; Kerfoot O 204324002439 mConyza steudelii Sch.Bip. ex A.Rich.Bytebier B et al. 22425002500 mConyza stricta Willd.Bytebier B et al. 22, 83, 197; Kerfoot O 204521342500 mConyza sumatrensis (Retz.) E.WalkerBytebier B et al. 22525002500 mCrassocephalum montuosum (S.Moore)MilneRedh.Bussmann R, Sight record23502550 mDichrocephala chrysanthemifolia (Blume)DC.Kerfoot O 205225912591 mDichrocephala integrifolia (L.f.) KuntzeBytebier B et al. 3524002400 mEmilia discifolia (Oliv.) C.JeffreyBytebier B et al. 245; Kerfoot O 2055;Gilbert MG, Gachathi FN & Gatheri GW 519816002439 mEmilia somalensis (S.Moore) C.JeffreyBytebier B et al. 8025002500 mGerbera viridifolia (DC.) Sch.Bip.Bytebier B et al. 17825002500 mGnaphalium rubriflorum HilliardBytebier B et al. 179,22625002500 mGutenbergia cordifolia Benth. ex Oliv.Bytebier B et al. 28723502350 mHelichrysum argyranthum O.Hoffm.Adamson J 56427432743 mHelichrysum forskahlii (J.F.Gmel.)Hilliard & B.L.Burtt var. forskahliiBytebier B et al. 62, 87; Adamson J 55624002500 mHelichrysum kilimanjari Oliv.Bytebier B et al. 2624002400 mFirst record for K1Helichrysum nudifolium (L.) Less. var. nudifoliumBytebier B et al. 204; Archer PG 702;Bono G 21322862500 mHelichrysum odoratissimum (L.) Less.Bussmann R 975923502550 mHirpicium diffusum (O.Hoffm.) RoesslerBytebier B et al. 239, 36216002200 mKleinia odora (Forssk.) DC.Gilbert MG, Gachathi FN & Gatheri GW 518713501350 mL agger a elatior R.E.Fr.Bytebier B et al. 5724002400 mFirst record for K1Microglossa pyrifolia (Lam.) KuntzeBytebier B et al. 8; Kerfoot O 204721342450 mMikaniopsis bambuseti (R.E. Fries)C.Jeffrey Bussmann R, Sight record23502550 mOsteospermum vail I ant ii (Decne.) Norl.Kerfoot O 205424392439 mSenecio hadiensis Forssk.Bono G 33017001700 mSenecio syringifolius O.Hoffm.Bytebier B et al. 18025002500 mSonchus afromontanus R.E.Fr.Bytebier B et al. 168,21625002500 mFirst record for K1Vernonia galamensis (Cass.) Less. ssp.nairobiensis M.G.GilbertBytebier B et al. 19625002500 mVernonia hymenolepis A.Rich.Kerfoot O 2041; Bono G 2024002439 mVernonia syringifolia O.Hoffm.Bytebier B et al. 9, 16924002500 mFirst record for K1CONVOLVULACEAEIpomoea spathulata Hallier f.Bytebier B et al. 30016001600 mIpomoea wightii (Wall.) ChoisyBytebier B et al. 27323502350 mCRASSULACEAECrassula alba Forssk.Bytebier B et al. 217; Archer PG 69922862500 mCrassula alsinoides (Hook.f.) Engl.Kerfoot O 209127432743 mCrassula nodulosa SchOnl. var. nodulosaArcher PG 698; Kerfoot O 202521342286 mCrassula schimperi Fisch. & C.A.Mey.ssp. schimperiBytebier B et al. 85, 24322002500 mKalanchoe citrina Schweinf.Bytebier B et al. 340, 356; Kerfoot O 208616002134 mKalanchoe densiflora Rolfe var. densifloraBytebier B et al. 220; Kerfoot O 208725002743 mCRUCIFERAEArabis glabra (L.) Bernh.Bytebier B et al. 17525002500 mFirst record for K1Cardamine africana L.Archer PG 69024392439 mCUCURBITACEAEGerrardanthus lobatus (Cogn.) C.JeffreyBytebier B et al. 30716001600 mLagenariaBytebier B et al. 35516001600 mLagenaria abyssinica (Hook.f.) C.JeffreyBytebier B et al. 5224002400 mFirst record for K1DIPSACACEAEDipsacus pinnatifidus A.Rich.Bytebier B et al. 184; Archer PG 704;Kerfoot O 2003; Bussmann R 982122862500 mScabiosa columbaria L.Archer PG 696; Cameron JBC 123;Adamson J 555; Kerfoot O 207522862743 mERICACEAEAgauria salicifolia (Lam.) Oliv.Kerfoot O 197027432743 mErica arborea L.Bytebier B et al. 171 ; Kerfoot O 2018;Bono G 214; Cameron JBC 11821342500 mErica mannii (Hook.f.) Beentje ssp.usambarensis (Aim & T.C.E.Fr.) BeentjeBytebier B et al. 18725002500 mFirst record for K1EUPHORBIACEAEAcalypha fruticosa Forssk. var.eglandulosa Radcl.Sm.Kerfoot O 199422862286 mAcalypha fruticosa Forssk. var. fruticosaBono G 12114001400 mAcalypha volkensii PaxBytebier B et al. 260; Kerfoot O 199521342350 mBridelia micrantha (Hochst.) Baill.Ichikawa M 906; Kerfoot O 194317681829 mClutia abyssinica Jaub. & Spach var. abyssinicaBytebier B et al. 193; Bono G 14219002500 mCroton dichogamus PaxBono G 2718001800 mCroton megalocarpus Hutch.Kerfoot O 196121342134 mEuphorbiaBytebier B et al. 36116001600 mEuphorbia brevicornu PaxBytebier B et al. 5624002400 mFirst record for K1Euphorbia depauperata A.Rich, var. depauperataBytebier B et al. 17025002500 mEuphorbia nyikae Pax var. nyikaeBytebier B et al. 35916001600 mFirst record for K1Phyllanthus flscheri PaxKerfoot O 201721342134 mPhyllanthus sepialis Mull.Arg.Bono G 12214001400 mFLACOURTIACEAEDovyalis abyssinica (A.Rich.) Warb.Bytebier B et al. 158; Kerfoot O 196524392500 mTrimeria grandifolia (Burkill) Sleumer ssp. tropicaBono G 3924002400 mGERANIACEAEGeranium aculeolatum Oliv.Kerfoot O 203824392439 mGeranium arabicum Forssk.Bytebier B et al. 190A; Kerfoot O 2036;Bono G 20121342500 mPelargonium alchemilloides (L.) Ait.f. ssp.multibracteatum (A.Rich.) KokwaroBytebier B et al. 244; Bono G 205;Kerfoot O 2039; Bussmann R 975722002450 mPelargonium whytei BakerKerfoot O 203724392439 mGUTTIFERAEGarcinia livingstonei T.AndersonBytebier B et al 33716001600 mHypericum kiboense Oliv.Adamson J 55827432743 mHypericum revolutum VahlBytebier B et al. 191; Kerfoot O 1932;Adamson J 562; Cameron JBC 11725002743 mHypericum roeperanum A.Rich.Kerfoot O 193424392439 mHALORAGACEAEGunner a per pens a L.Bytebier B et al. 230; Bussmann R 978824502500 mLABIATAEAeollanthus repens Oliv.Bytebier B et al. 25923502350 mFirst record for K1Becium decumbens (Guerke) A.J.PatonBytebier B et al. 7825002500 mBecium obovatum (E.Mey. ex Benth.)N.E.Br, var. capitatumArcher PG 700; Kerfoot O 209422862743 mLeonotis nepetifolia (L.) R.Br.Bytebier B et al. 6024002400 mLeucas grandis GiirkeKerfoot O 2089; Gilbert MG, Gachathi FN & Gatheri GW 521413502743 mLeucas urticifolia (Vahl) R.Br. var. annulata SebaldBytebier B et al. 28323502350 mOcimum suave Willd.Bytebier B et al. 281; Kerfoot O 200223502439 mPlectranthus barbatus Andr.Kerfoot O 199221342134 mPlectranthus edulis (Vatke) AgnewBytebier B et al. 23325002500 mFirst record for K1Plectranthus grandicalyx E.A.BruceBytebier B et al. 8425002500 mPlectranthus igniriaus (Schweinf.) AgnewGilbert MG, Gachathi FN & Gatheri GW 521616501650 mPlectranthus sylvestris GiirkeBytebier B et al. 7024002400 mSalvia nilotica (Juss.) Jacq.Bytebier B et al. 23; Kerfoot O 206624002439 mSaturejaBytebier B et al. 29, 19224002500 mSatureja abyssinica (Benth.) Briq.Archer PG 691; Kerfoot O 2060; CameronJBC 112; Bono G 11521342600 mSatureja biflora (D.Don) Benth.Bytebier B et al. 181; Kerfoot O 2029;Cameron JBC 10921342500 mSatureja pseudosimensis BrenanBytebier B et al. 20525002500 mFirst record for K1Tinnea aethiopica Kotschy ex Hook.f.Bytebier B et al. 32616001600 mLEGUMINOSAEAcacia etbaica Schweinf.Bytebier B et al. 30816001600 mAcacia hockii De Wild.Kerfoot O 190321342134 mAcacia Senegal (L.) WildBytebier B et al. 35710001000 mAmphicarpa qfricana (Hook.f.) HarmsKerfoot O 191526522652 mArgyrolobium fischeri Taub.Archer PG 703; Bono G 30; Cameron JBC 11320002286 mCadiapurpurea (Pice.) AitonBytebier B et al. 333; Ichikawa M 899;Kerfoot O 1905; JexBlake M 11777; Jex-Blake AJ H69/5116002286 mChamaecrista usambarensis (Taub.)StandleyBytebier B et al. 199; Adamson J 56525002743 mCrotalariaBytebier B et al. 21225002500 mCrotalaria fascicularis PolhillBono G 2819001900 mCrotalaria incana L. ssp. purpurascens(Lam.) MilneRedh.Bytebier B et al. 152, 26523502500 mCrotalaria keniensis Baker f.Kerfoot O 191322862286 mCrotalaria lachnocarpoides Engl.Bytebier B et al. 156; Kerfoot O 1906, 190724392500 mCrotalaria natalitia Meisn. var. natalitiaBytebier B et al. 274; Bono G 132;Kerfoot O 191421342500 mDesmodium repandum (Vahl) DC.Kerfoot O 191826522652 mDolichos sericeus E.Mey. ssp. sericeusBono G 32020002000 mGlycine wightii (Wight & Arn.) Verde, ssp.petitiana (A.Rich.) Verde, var. meamsii(De Wild.) Verde.Kerfoot O 191718291829 mGlycine wightii (Wight & Arn.) Verde, ssp.wightii var. longicauda (Schweinf.)Verde.Ichikawa M 90516761676 mIndigofera atriceps Hook.f. ssp. atricepsKerfoot O 190921342134 mIndigofera lupatana Baker f.Kerfoot O 191221342134 mIndigofera schimperi Jaub. & Spach var. schimperiGilbert MG, Gachathi FN & Gatheri GW 519213501350 mIndigofera swaziensis Bolus var. swaziensisKerfoot O 191024392439 mIndigofera volkensii Taub.Bytebier B et al. 37; Gilbert MG, GachathiFN & Gatheri GW 5190A13502400 mKotschya recurvifolia (Taub.) F.White ssp.keniensis Verde.Cameron JBC 11924392439 mLablab purpureus (L.) Sweet ssp.uncinatus Verde.Kerfoot O 192224392439 mLotus goetzei HarmsKerfoot O 191121342134 mMucuna gigantea (Willd.) DC. ssp.quadrialata (Baker) Verde.Bytebier B et al. 29816001600 mOrmocarpum trachycarpum (Taub.)HarmsKerfoot O 193721342134 mPsoraleafoliosa Oliv.Adamson J 53727432743 mPterolobium stellatum (Forssk.) BrenanKerfoot O 193518291829 mTephrosia interrupta Engl. ssp. interruptaAdamson J 559; Kerfoot O 1908; Bono G133; Gilbert MG, Gachathi FN & GatheriGW 520416002652 mTeramnus labialis (L.f.) Spreng. ssp.labialis var. abyssinicus (A.Rich.) Verde.Kerfoot O 192024392439 mTrifolium semipilosum Fresen. var. semipilosumBono G 20425002500 mVigna schimperi BakerBytebier B et al. 34; Kerfoot O 191624002439 mZornia setosa Baker f. ssp. obovata (Baker f.) J.Léon. & MilneRedh.Bytebier B et al. 9125002500 mLINACEAELinum keniense T.C.E.Fr.Bussmann R 978023502550 mLinum volkensii Engl.Bytebier B et al. 95; Adamson J 54725002743 mLOBELIACEAELobelia giberroa Hemsl.Bytebier B et al. 23625502550 mLobelia holstii Engl.Bytebier B et al. 207; Adamson J 566;Kerfoot O 2085; Cameron JBC 12422862743 mMonopsis stellarioides (C.Presl) Urb. ssp.schimperiana (Urb.) ThulinBytebier B et al. 20225002500 mFirst record for K1LOGANIACEAENuxia congesta Fresen.Bytebier B et al. 182; Bono G 2523002500 mLORANTHACEAEAgelanthus elegantulus (Engl.) Polhill & WiensBussmann R 975823502550 mFirst record for K1Englerina woodfordioides (Schweinf.)BalleBytebier B et al. 40, 20024002500 mMALVACEAEAbutilon hirtum (Lam.) SweetKerfoot O 200826522652 mAbutilon longicuspe A.Rich.Kerfoot O 198627432743 mAbutilon mauritianum (Jacq.) SweetKerfoot O 2010, 201321342439 mHibiscusBono G 13024002400 mHibiscus fuscus GarckeBytebier B et al. 28623502350 mHibiscus ludwigii Eckl. & Zeyh.Bono G 2921002100 mHibiscus vitifolius L.Kerfoot O 200927432743 mPavonia kilimandscharica GiirkeKerfoot O 201127432743 mPavonia patens (Andr.) Chiov.Bytebier B et al. 8225002500 mPavonia wens Cav.Bytebier B et al. 9725002500 mSida rhombifolia L.Kerfoot O 201424392439 mSida tenuicarpa VollesenBytebier B et al. 167; Bussmann R 975624502500 mMELIACEAETurraea abyssinica A.Rich.Bytebier B et al. 28023502350 mFirst record for K1Turraea holstii GiirkeBussmann R, Sight record23502350 mMENISPERMACEAECocculus pendulus (J.R.Forst. & G.Forst.)DielsBono G 20023502350 mStephania abyssinica (Quart.Dill. & A.Rich.) Walp.Kerfoot O 210127432743 mMONIMIACEAEXymalos monospora (Harv.) Warb.Bytebier B et al. 4; Cameron JBC 129;Ichikawa M 90220002400 mMORACEAEFicus scassellatii Pamp. ssp. scassellatiiBytebier B et al. 33816001600 mFicus vallischoudae DelileBytebier B et al. 29516001600 mMYRSINACEAEMaesa lanceolata Forssk.Kerfoot O 195227432743 mMyrsine africana L.Bytebier B et al. 237; Cameron JBC 120;Bono G 2218502200 mRapanea melanophloeos (L.) MezBytebier B et al. 224002400 mMYRTACEAESyzygium cordatum Hochst.Kerfoot O 194418291829 mNYCTAGINACEAECommicarpus helenae (Roem. & Schult.)MeikleBytebier B et al. 31016001600 mOCHNACEAEOchna insculpta SleumerBytebier B et al. 26823502350 mOLEACEAEChionanthus battiscombei (Hutch.) StearnBussmann R, Sight record23502550 mJasminum abyssinicum Hochst. ex DC.Bytebier B et al. 2424002400 mJasminum floribundum R.Br, ex Fresen.Kerfoot O 197827432743 mJasminum fluminense Veil. ssp. holstii(Gilg) TurrillKerfoot O 1976; Bono G 12923002743 mOlea capensis L.Bytebier B et al. 10, 9824002500 mOlea europaea L. ssp. africana (Mill.)P.GreenBytebier B et al. 88; Kerfoot O 1962,1964; Adamson J 395; Adamson J B 616423782500 mSchrebera alata (Hochst.) Welw.Bytebier B et al. 29223502350 mONAGRACEAEEpilobium hirsutum L.Bytebier B et al. 31616001600 mFirst record for K1OPILIACEAEOpilia amentacea Roxb.BonoG 13623002300 mOROBANCHACEAEOrobanche minor Sm.Bytebier B et al. 3124002400 mOXALIDACEAEOxalis corniculata L.Bytebier B et al. 16325002500 mFirst record for K1PIPERACEAEPeperomia abyssinica Miq.Bytebier B et al. 6724002400 mPiper capense L.Bytebier B et al. 64, 173; Kerfoot O 202224002743 mPLUMBAGINACEAEPlumbago dawei RolfeKerfoot O 206127432743 mPOLYGALACEAEPolygala sphenoptera Fresen.Bytebier B et al. 94, 242; Bussmann R 977922002500 mPOLYGONACEAEPolygonum amphibium L.Bussmann R 978723502550 mFirst record for K1Polygonum setosulum A.Rich.Bytebier B et al. 227; Kerfoot O 2034, 203525002743 mRumex bequaertii De Wild.Kerfoot O 203021342134 mRumex steudelii A.Rich.Bytebier B et al. 6324002400 mPRIMULACEAELysimachia volkensii Engl.Bytebier B et al. 1524002400 mPROTEACEAEFaurea saligna Harv.Bytebier B et al. 190B; Kerfoot O 193625002743 mProtea caffra Meisn. ssp.kilimandscharica (Engl.) Chisumpa & BrummittCameron JBC 12721342134 mRANUNCULACEAEClematis brachiata Thunb.Bytebier B et al. 5524002400 mClematis simensis Fresen.Bytebier B et al. 320; Bono G 41; Kerfoot O 200416002591 mRanunculus multifidus Forssk.Bytebier B et al. 223; Kerfoot O 2032, 203318292743 mThalictrum rhynchocarpum Quart.Dill. & A.Rich.Bytebier B et al. 2724002400 mFirst record for K1RHAMNACEAERhamnus prinoides L'HéritBytebier B et al. 32; Kerfoot O 197124002743 mRhamnus staddo A.Rich.Kerfoot O 194621342134 mScutia myrtina (Burm.f.) KurzBytebier B et al. 17425002500 mZiziphus mucronata Willd. ssp. mucronataIchikawa M 898; Kerfoot O 194018901981 mRHIZOPHORACEAECassipourea malosana (Baker) AlstonBytebier B et al. 1124002400 mROSACEAEAlchemilla cryptantha A.Rich.Bytebier B et al. 3324002400 mPrunus africana (Hook.f.) KalkmanBytebier B et al. 155; Adamson J 39024392500 mRubus adolfifriederici Engl.Kerfoot O 198327432743 mRubus apetalus Poir.Bytebier B et al. 7; Kerfoot O 198424002743 mRUBIACEAEAnthospermum usambarense K.Schum.Kerfoot O 199922862286 mGaliniera saxifraga (Hochst.) BridsonBytebier B et al. 4724002400 mFirst record for K1Galium aparinoides Forssk.Bytebier B et al. 3924002400 mFirst record for K1Pavetta abyssinica Fresen. var. abyssinicaBytebier B et al. 27223502350 mPavetta gardeniifolia A.Rich, var. gardeniifoliaCameron JBC 12121342134 mPentas lanceolata (Forssk.) DeflersBytebier B et al. 27123502350 mPentas parvifolia HiernBytebier B et al. 304; Kerfoot O 2031;Gilbert MG, Gachathi FN & Gatheri GW 521313502439 mPsychotria kirkii Hiern var. tarambassica(Bremek.) Verde.Bytebier B et al. 241; Kerfoot O 1966,1967, 1968; Gilbert MG, Gachathi FN & Gatheri GW 519716002591 mPsychotria orophila PetitBytebier B et al. 14; Kerfoot O 195321342400 mPsydrax schimperiana (A.Rich.) Bridsonssp. schimperianaBytebier B et al. 31516001600 mRytigynia uhligii (K.Schum. & K.Krause)Verde.Bytebier B et al. 27723502350 mTarenna graveolens (S.Moore) Bremek.var. graveolensKerfoot O 195522862286 mTarenna graveolens (S.Moore) Bremek.var. impolita BridsonGilbert MG, Gachathi FN & Gatheri GW 519013501350 mVangueria apiculata K.Schum.Gilbert MG, Gachathi FN & Gatheri GW 521516501650 mVangueria madagascariensis J.F.Gmel.Bytebier B et al. 301, 346; Kerfoot O 195416001829 mVangueria volkensii K.Schum. var.volkensiiBytebier B et al. 275, 32816002350 mRUTACEAEClausena anisata (Willd.) Benth.Bytebier B et al. 5; Kerfoot O 1993;Adamson J 39624002652 mTeclea nobilis DelileBytebier B et al. 42, 172; Kerfoot O 195724002500 mTeclea simplicifolia (Engl.) Verd.Kerfoot O 198022862286 mVepris samburuensis KokwaroBytebier B et al. 31316001600 mSALVADORACEAESalvador a persica L. var. persicaBono G 21220002000 mSANTALACEAEOsyridicarpos schimperianus A.DC.Bytebier B et al. 297,35116001600 mOsyris abyssinica (Hochst) A.Rich.Kerfoot O 201622862286 mOsyris lanceolata Hochst. & Steud.Bytebier B et al. 18525002500 mSAPINDACEAEAllophylus abyssinicus (Hochst.) Radlk.Bytebier B et al. 1224002400 mAllophylus griseotomentosus GilgKerfoot O 195121342134 mCardiospermum halicacabum L.Bytebier B et al. 31116001600 mDodonaea viscosa (L.) Jacq.Kerfoot O 198125912591 mSAPOTACEAEManilkara mochisia (Baker) DubardAdamson TG 17; Kerfoot O 19479142134 mMimusops kummel A. DC.Bytebier B et al. 330, 34916001600 mSCROPHULARIACEAEBartsia longiflora Hochst. ex Benth.Adamson J 54627432743 mCycnium tenuisectum (Standi.)O.J.HansenBytebier B et al. 20825002500 mHalleria lucida L.Bytebier B et al. 86; Adamson J 536;Kerfoot O 197525002743 mHebenstretia angolensis RolfeBytebier B et al. 162; Cameron JBC 126;Kerfoot O 202024392500 mMisopates orontium (L.) Raf.Archer PG 69222862286 mSelago thomsonii RolfeBytebier B et al. 186, 364; Adamson J 55316002743 mVerbascum brevipedicellatum (Engl.)Hub.Mor.Bytebier B et al. 30, 161; Adamson J 56024002743 mVeronica anagallisaquatica L.Bytebier B et al. 23525502550 mSIMAROUBACEAEBrucea antidysenterica Lam.Bytebier B et al. 16; Bono G 3724002450 mSOLANACEAESolarium aculeatissimum Jacq.Bytebier B et al. 65; Kerfoot O 198724002743 mSolarium benderianum Engl.Kerfoot O 200025912591 mSolarium incanum L.Bussmann R, Sight record23502550 mSolarium indicum L.Bytebier B et al. 13,26623502400 mSolarium indicum L. ssp. adoense(Hochst.) BitterKerfoot O 1988, 198924392743 mSolanum indicum L. ssp. grandifronsBitterBytebier B et al. 7524002400 mSolanum nigrum L.Bytebier B et al. 221; Bono G 13424002500 mSolanum renschii VatkeBytebier B et al. 30516001600 mSolanum schumannianum DammerBussmann R, Sight record23502550 mSolanum terminale Forssk.Bytebier B et al. 2824002400 mSTERCULIACEAEDombeya goetzenii K.Schum.Kerfoot O 194124392439 mDombeya rotundifolia Planch.Gilbert MG, Gachathi FN & Gatheri GW519916001600 mTHYMELAEACEAEGnidia glauca (Fresen.) GilgBytebier B et al. 165; Kerfoot O 2005Cameron JBC 12222862500 mStruthiola thomsonii Oliv.Bytebier B et al. 203; Archer PG 706;Adamson J 54525002743 mTILIACEAESparrmannia ricinocarpa (Eckl. & Zeyh.)KuntzeKerfoot O 201524392439 mUMBELLIFERAEAlepidea pedunculahs A.Rich.Bytebier B et al. 206; Kerfoot O 2064;Cameron JBC 125; Archer PG 688;Adamson J 53924392743 mAnthriscus sylvestris (L.) Hoffm. var. sylvestrisBytebier B et al. 45, 20924002500 mDiplolophium africanum Turcz.Adamson J 56124392439 mFerula communis L.Kerfoot O 202621342134 mHeteromorpha trifoliata (H.L.Wendl.)Eckl. & Zeyh.Bytebier B et al. 291; Bono G 21; Kerfoot O 2027, 202822862439 mSanicula elata Buch.Ham. ex D.DonBytebier B et al. 1824002400 mTorilis arvensis (Huds.) LinkBytebier B et al. 264; Bussmann R 978423502450 mURTICACEAEDroguetia iners (Forssk.) Schweinf.Bussmann R, Sight recordGirardinia diversifolia (Link) FriisBytebier B et al. 28223502350 mLaportea alatipes Hook.f.Bytebier B et al. 324002400 mFirst record for K1Parietaria debilis G.Forst.Bytebier B et al. 4324002400 mPileajohnstonii Oliv.Bussmann R, Sight record23502550 mUrera hypselodendron (A.Rich.) Wedd.Bytebier B et al. 10025002500 mFirst record for K1VERBENACEAEClerodendrum johnstonii Oliv.Kerfoot O 196024392439 mClerodendrum myricoides (Hochst.) VatkeBytebier B et al. 322; Gilbert MG,Gachathi FN & Gatheri GW 520016001600 mVIOLACEAEViola abyssinica Oliv.Bytebier B et al. 54; Kerfoot O 210224002743 mVISCACEAEViscum triflorum DC.Bytebier B et al. 29023502350 mViscum tuberculatum A.Rich.Bussmann R 974923502550 mVITACEAECyphostemma bambuseti (Gilg & Brandt)Wild & R.B.Drumm.Bytebier B et al. 28523502350 mFirst record for K1Cyphostemma kilimandschahcum (Gilg)Wild & R.B.Drumm.Bytebier B et al. 4924002400 mRhoicissus revoilii Planch.Bytebier B et al. 29316001600 mMONOCOTYLEDONAEALOACEAEAloeBytebier B et al. 36016001600 mAMARYLLIDACEAEScadoxus multiflorus (Martyn) Raf.Bytebier B et al. 25623502350 mASPARAGACEAEAsparagus qfricanus Lam.Bytebier B et al. 18325002500 mAsparagus falcatus L. var. ternifoliusJessopBytebier B et al. 27023502350 mASPHODELACEAEBulbine abyssinica A.Rich.Bytebier B et al. 151,219; Bussmann R 982024502500 mCOMMELINACEAEAneilema leicocaule K.Schum.Bussmann R, Sight record23502550 mCommelina africana L.Bytebier B et al. 15425002500 mCommelina benghalensis Wall.Kerfoot O 209227432743 mCommelina foliacea Chiov.Bytebier B et al. 3624002400 mCyanotis foecunda Hassk.Bytebier B et al. 343; Bussmann R 978116002450 mCYPERACEAECyperusBytebier B et al. 6824002400 mCyperus comosipes Mattf. & Kük. ssp. comosipesBytebier B et al. 25723502350 mCyperus dichroostachyus A.Rich.Bytebier B et al. 234; Kerfoot O 192625002743 mCyperus impubes Steud.Bytebier B et al. 26123502350 mCyperus niveus Retz. var. leucocephalus(Kunth) FossbergKerfoot O 192427432743 mCyperus rigidifolius Steud.Bytebier B et al. 59, 15924002500 mCyperus rohlfsii Boeck.Bytebier B et al. 34216001600 mCyperus sesquiflorus (Torn) Mattf. & Kük.ssp. appendiculatus (K.Schum.) LyeBytebier B et al. 7124002400 mCyperus niger Ruiz & Pav. ssp.elegantulus (Steud.) LyeKerfoot O 192527432743 mSchoenoxiphium lehmannii (Nees) Steud.Bussmann R 981923502550 mDRACAENACEAEDracaena afromontana Mildbr.Bytebier B et al. 2124002400 mDracaena ellenbeckiana Engl.Bytebier B et al. 35816001600 mGRAMINEAEAgrostis schimperana Steud.Bytebier B et al. 22825002500 mAndropogonBytebier B et al. 15325002500 mBrachypodium flexum NeesBussmann R, Sight recordBromus leptoclados NeesCameron JBC 11625912591 mChloris roxburghiana Schult.Bytebier B et al. 29916001600 mCynodon nlemfuensis Vanderyst var.nlemfuensisBytebier B et al. 28423502350 mDigitaria velutina (Forssk.) P.Beauv.Bytebier B et al. 29616001600 mEhrharta erecta Lam. var. abyssinica(Hochst.) Pilg.Bytebier B et al. 61; Kerfoot O 193024002743 mEleusine multiflora A.Rich.Bytebier B et al. 16625002500 mFirst record for K1Enteropogon macrostachyus K.Schum. ex Engl.Bytebier B et al. 30616001600 mEragrostis schweinfurthii Chiov.Bytebier B et al. 19525002500 mFirst record for K1Eragrostis tenuifolia (A.Rich.) Steud.Bytebier B et al. 8925002500 mExotheca abyssinica (A.Rich.) AnderssonBytebier B et al. 21125002500 mLeptochloa rupestris C.E.Hubb.Bytebier B et al. 30316001600 mFirst record for K1Panicum deustum Thunb.Bytebier B et al. 30216001600 mPanicum hymeniochilum NeesBytebier B et al. 22925002500 mFirst record for K1Rhynchelytrum repens (Willd.) C.E.Hubb.Bytebier B et al. 35316001600 mSinarundinaria alpina (K.Schum.)C.S.Chao & RenvoizeBytebier B et al. 5324002400 mSporobolus africanus (Poir.) Robyns & ToumayBytebier B et al. 9925002500 mStipa dregeana Steud. var. elongata (Nees)StapfBytebier B et al. 26723502350 mHYACINTHACEAEAlbuca abyssinica Jacq.Bytebier B et al. 15725002500 mHYPOXIDACEAEHypoxis kilimanjarica Baker ssp.kilimanjaricaBytebier B et al. 90; Bussmann R 977624502500 mFirst record for K1Hypoxis obtusa Burch.Bytebier B et al. 21425002500 mIRIDACEAEAristea alata BakerBytebier B et al. 79; Adamson J 53525002743 mGladiolus goetzei HarmsArcher PG 70124392439 mGladiolus newii Bak.Bytebier B et al. 21825002500 mFirst record for K1JUNCACEAEJuncus oyxcarpus KunthBytebier et al. 81, Bussmann R976024502500 mORCHIDACEAEAerangis thomsonii (Rolfe) Schltr.Adamson TG 534; Bytebier B et al. 10223502743 mBrownleea parviflora Lindl.Kerfoot O 210027432743 mDiaphananthe rohrii (Rchb.f.) Summerh.Bytebier B et al. 13125002500 mFirst record for K1Eulophia petersii Rchb.f.Bytebier B et al. 14416001600 mPolystachya confusa RolfeArcher PG 713; Bytebier B et al. 110, 113, 132,13324002652 mPolystachya piersii P.J.CribbBytebier B et al. 101; Archer PG 70923502350 mRangaeris amaniensis (Kraenzl.)Summerh.Bytebier B et al. 14222502250 m

Benny Bytebier and Rainer W. Bussmann "The Vegetation of Mount Nyiru (Samburu District, Kenya): A Checklist and Syntaxonomical Survey," Journal of East African Natural History 89(1), 45-71, (1 January 2000). https://doi.org/10.2982/0012-8317(2000)89[45:TVOMNS]2.0.CO;2
Published: 1 January 2000
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